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<document id="E22432B3E3EEEC8C6A43C0D2C263B72E" ID-DOI="10.37828/em.2020.35.10" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232490" ID-ZooBank="urn:lsid:zoobank.org:pub:F63DAA57-8D7A-4EC7-9FA0-602943FED1FB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722913961471" checkinUser="felipe" docAuthor="Georgiev, Dilian &amp; Glöer, Peter" docDate="2020" docId="C97AB10B4B33FFF662D27399FDC9683E" docLanguage="en" docName="EcolMontenegrina.35.129-137.pdf" docOrigin="Ecologica Montenegrina 35" docSource="http://dx.doi.org/10.37828/em.2020.35.10" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Grossuana beroni Georgiev &amp; Glöer, 2020, n. sp." docType="treatment" docUuid="F8C2DAFC-0535-4E8E-A891-6F9ECFFA88AC" docUuidSource="ZooBank" docVersion="2" lastPageNumber="133" masterDocId="3543C9734B31FFF262457045FFEE693E" masterDocTitle="Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855" masterLastPageNumber="137" masterPageNumber="129" pageNumber="131" updateTime="1722916995419" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="244A8646E3DBCA79B14EF294AAE9B9CF">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<mods:namePart id="44F9488400E0F3B2AFB5DB898A328BDD">Georgiev, Dilian</mods:namePart>
<mods:affiliation id="15388A250697CA8636710B2CFC93B514">Department of Ecology and Nature Conservation, Faculty of Biology, Paisii Hilendarski University of Plovdiv, 24 Tsar Assen Str., 4000 Plovdiv, Bulgaria</mods:affiliation>
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<mods:namePart id="804DBCEA468AD4C23C3C9C5A3587FE68">Glöer, Peter</mods:namePart>
<mods:affiliation id="8D22139D1E3ACF40B845FE0A67493175">Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany Corresponding author. E-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<mods:title id="EA4224FD79580A15A3E8A254298CCFB7">Ecologica Montenegrina</mods:title>
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<mods:date id="389F910ED91F8C785C3667798178DC51">2020</mods:date>
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<mods:identifier id="666956C5860A5BD7D13D6C44AD3C06F7" type="DOI">10.37828/em.2020.35.10</mods:identifier>
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<treatment id="C97AB10B4B33FFF662D27399FDC9683E" ID-DOI="http://doi.org/10.5281/zenodo.13232604" ID-Zenodo-Dep="13232604" LSID="urn:lsid:zoobank.org:act:F8C2DAFC-0535-4E8E-A891-6F9ECFFA88AC" httpUri="http://treatment.plazi.org/id/C97AB10B4B33FFF662D27399FDC9683E" lastPageId="4" lastPageNumber="133" pageId="2" pageNumber="131">
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<paragraph id="416C001D4B33FFF062D27399FD976AC8" blockId="2.[151,1159,987,1044]" box="[151,633,987,1014]" pageId="2" pageNumber="131">
<heading id="1A24B7714B33FFF062D27399FD976AC8" box="[151,633,987,1014]" fontSize="11" level="2" pageId="2" pageNumber="131" reason="2">
<emphasis id="73A7DC0F4B33FFF062D27399FE556AC8" bold="true" box="[151,443,988,1014]" pageId="2" pageNumber="131">
<taxonomicName id="86D37B9E4B33FFF062D27399FE9F6AC8" ID-CoL="7RG9W" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[151,369,988,1014]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="beroni" status="sp. nov.">
<emphasis id="73A7DC0F4B33FFF062D27399FE9F6AC8" bold="true" box="[151,369,988,1014]" italics="true" pageId="2" pageNumber="131">Grossuana beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B33FFF0633D7399FE556AC8" box="[376,443,988,1014]" pageId="2" pageNumber="131" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D9E81C984B33FFF0638E739EFDF36AC8" box="[459,541,987,1014]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="2" pageNumber="131">Figs. 2</figureCitation>
,
<figureCitation id="D9E81C984B33FFF0606F739EFDD46AC8" box="[554,570,987,1014]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="2" pageNumber="131">3</figureCitation>
,
<figureCitation id="D9E81C984B33FFF06002739EFD816AC8" box="[583,623,987,1014]" captionStart="Figures 6-10" captionStartId="3.[151,236,1972,1996]" captionTargetBox="[151,1444,1565,1953]" captionTargetId="figure-136@3.[151,1444,1565,1953]" captionTargetPageId="3" captionText="Figures 6-10. The Grossuana spp. 6-7: G. beroni n. sp. (paratypes), 8: penis of G. beroni n. sp. 9: G. thasia n. sp (holotype), 10: penis of G. thasia n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232500" httpUri="https://zenodo.org/record/13232500/files/figure.png" pageId="2" pageNumber="131">6-8</figureCitation>
)
</heading>
</paragraph>
<paragraph id="416C001D4B33FFF062D273B9FB696D2A" blockId="2.[151,1159,987,1044]" box="[151,1159,1020,1044]" pageId="2" pageNumber="131">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</subSubSection>
<subSubSection id="09C953964B33FFF062D2747AFBE26C94" pageId="2" pageNumber="131" type="materials_examined">
<paragraph id="416C001D4B33FFF062D2747AFE9E6D67" blockId="2.[151,1436,1087,1349]" box="[151,368,1087,1113]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D2747AFE9E6D67" bold="true" box="[151,368,1087,1113]" pageId="2" pageNumber="131">Material studied:</emphasis>
</paragraph>
<paragraph id="416C001D4B33FFF062B07425FB566D44" blockId="2.[151,1436,1087,1349]" box="[245,1208,1119,1146]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062B07425FE9F6D44" bold="true" box="[245,369,1120,1146]" pageId="2" pageNumber="131">
<typeStatus id="9E68BEBF4B33FFF062B07425FE826D44" box="[245,364,1120,1146]" pageId="2" pageNumber="131" type="holotype">Holotype</typeStatus>
:
</emphasis>
shell height
<quantity id="862BADF84B33FFF0604D741AFD8A6D44" box="[520,612,1119,1146]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="131" unit="mm" value="1.7">1.7 mm</quantity>
, width
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, from type locality (ZMH 140789).
</paragraph>
<paragraph id="416C001D4B33FFF062B074C4FEA86D80" blockId="2.[151,1436,1087,1349]" pageId="2" pageNumber="131">
<materialsCitation id="F1BB0A404B33FFF062B074C4FC9D6DA2" box="[245,883,1153,1180]" collectionCode="ZMH" pageId="2" pageNumber="131" specimenCode="ZMH 140790" specimenCount="43" typeStatus="paratype">
<emphasis id="73A7DC0F4B33FFF062B074C4FE926DA5" bold="true" box="[245,380,1153,1179]" pageId="2" pageNumber="131">
<typeStatus id="9E68BEBF4B33FFF062B074C4FE996DA5" box="[245,375,1153,1179]" pageId="2" pageNumber="131" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="57D5CB944B33FFF063CD74C4FDC46DA2" box="[392,554,1153,1180]" count="43" pageId="2" pageNumber="131" type="generic">43 specimens</specimenCount>
in ethanol (
<specimenCode id="1175A8664B33FFF0608574C4FC826DA2" box="[704,876,1153,1180]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="131" type="Museum">ZMH 140790</specimenCode>
)
</materialsCitation>
,
<specimenCount id="57D5CB944B33FFF061C674C4FBF86DA2" box="[899,1046,1153,1180]" count="5" pageId="2" pageNumber="131" type="generic">5 specimens</specimenCount>
in coll. Glöer,
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in coll. Georgiev.
</paragraph>
<paragraph id="416C001D4B33FFF062B07481FBEA6C7B" blockId="2.[151,1436,1087,1349]" pageId="2" pageNumber="131">
<materialsCitation id="F1BB0A404B33FFF062B07481FBEE6C7B" collectionCode="ZMH" collectorName="Gloer" country="Greece" county="Thassos Island" elevation="124" latitude="40.75822" location="Panagia village" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" pageId="2" pageNumber="131" specimenCode="ZMH 140792, ZMH 140791" specimenCount="50" stateProvince="Voreio Aigaio">
Additional localities: (1)
<collectingCountry id="39C4408D4B33FFF060587481FD9D6DE1" box="[541,627,1220,1247]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
,
<collectingCounty id="A80D78914B33FFF0603A7481FCDE6DE1" box="[639,816,1220,1247]" pageId="2" pageNumber="131">Thassos Island</collectingCounty>
, spring below
<collectingMunicipality id="A1089A674B33FFF061997481FB4F6DE1" box="[988,1185,1220,1247]" pageId="2" pageNumber="131">Drakotrypa cave</collectingMunicipality>
,
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<geoCoordinate id="24E766DA4B33FFF0672C7481FF1D6C3F" degrees="40" direction="north" minutes="44" orientation="latitude" pageId="2" pageNumber="131" precision="1" seconds="06.3" value="40.735085">N40 44 06.3</geoCoordinate>
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,
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a.s.l.
</elevation>
(
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:
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,
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coll.
<collectorName id="EC2665CB4B33FFF066CD74A3FB236C3F" box="[1160,1229,1254,1281]" pageId="2" pageNumber="131">Glöer</collectorName>
) and (2)
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,
<location id="440C56C64B33FFF062D2754DFEA96C1D" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF062D2754DFEA96C1D" box="[151,327,1288,1315]" country="Greece" county="Thassos Island" latitude="40.75822" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" name="Thassos Island" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Thassos Island</location>
,
<collectingRegion id="8317CEFF4B33FFF06311754DFE126C1D" box="[340,508,1288,1315]" country="Greece" name="Voreio Aigaio" pageId="2" pageNumber="131">North Aegean</collectingRegion>
, a big spring in the yard of the church of
<location id="440C56C64B33FFF061A1754DFB4E6C1D" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF061A1754DFB4E6C1D" box="[996,1184,1288,1315]" country="Greece" county="Thassos Island" latitude="40.75822" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" name="Rachoni village" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Rachoni village</location>
,
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<geoCoordinate id="24E766DA4B33FFF0670D754DFF236C7B" degrees="24" direction="east" minutes="38" orientation="longitude" pageId="2" pageNumber="131" precision="1" seconds="17.3" value="24.638138">E24 38 17.3</geoCoordinate>
,
<elevation id="CAFEE72E4B33FFF0629C756FFE8C6C7B" box="[217,354,1322,1349]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="131" unit="m" value="107.0">
<quantity id="862BADF84B33FFF0629C756FFECA6C7B" box="[217,292,1322,1349]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="131" unit="m" value="107.0">107 m</quantity>
a.s.l.
</elevation>
(
<specimenCode id="1175A8664B33FFF06334756FFDF96C7B" box="[369,535,1322,1349]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="131" type="Museum">ZMH 140791</specimenCode>
:
<specimenCount id="57D5CB944B33FFF06067756FFD5B6C7B" box="[546,693,1322,1349]" count="5" pageId="2" pageNumber="131" type="generic">5 specimens</specimenCount>
,
<specimenCount id="57D5CB944B33FFF06085756FFCBE6C7B" box="[704,848,1322,1349]" count="4" pageId="2" pageNumber="131" type="generic">4 specimens</specimenCount>
in coll.
<collectorName id="EC2665CB4B33FFF061F5756FFC166C7B" box="[944,1016,1322,1349]" pageId="2" pageNumber="131">Glöer</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="416C001D4B33FFF062D2752BFBE26C94" blockId="2.[151,1436,1389,1450]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D2752BFEAB6CB6" bold="true" box="[151,325,1390,1416]" pageId="2" pageNumber="131">Type locality:</emphasis>
<materialsCitation id="F1BB0A404B33FFF0630A7528FBE66C94" country="Greece" elevation="115" latitude="40.68364" location="Skala Potamia" longLatPrecision="1" longitude="24.771305" municipality="Thassos Island" pageId="2" pageNumber="131" specimenCount="1" stateProvince="Voreio Aigaio">
<collectingCountry id="39C4408D4B33FFF0630A7528FE4B6CB6" box="[335,421,1389,1416]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
,
<collectingMunicipality id="A1089A674B33FFF063F67528FD8B6CB6" box="[435,613,1389,1416]" pageId="2" pageNumber="131">Thassos Island</collectingMunicipality>
,
<collectingRegion id="8317CEFF4B33FFF060367528FCF36CB6" box="[627,797,1389,1416]" country="Greece" name="Voreio Aigaio" pageId="2" pageNumber="131">North Aegean</collectingRegion>
, a spring (water source) south of
<location id="440C56C64B33FFF066F17528FAB06CB6" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF066F17528FAB06CB6" box="[1204,1374,1389,1416]" country="Greece" latitude="40.68364" longLatPrecision="1" longitude="24.771305" municipality="Thassos Island" name="Skala Potamia" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Skala Potamia</location>
near the round road of the island,
<geoCoordinate id="24E766DA4B33FFF063AC75CAFD956C94" box="[489,635,1423,1450]" degrees="40" direction="north" minutes="41" orientation="latitude" pageId="2" pageNumber="131" precision="1" seconds="01.1" value="40.68364">N40 41 01.1</geoCoordinate>
<geoCoordinate id="24E766DA4B33FFF060C775CAFCFD6C94" box="[642,787,1423,1450]" degrees="24" direction="east" minutes="46" orientation="longitude" pageId="2" pageNumber="131" precision="1" seconds="16.7" value="24.771305">E24 46 16.7</geoCoordinate>
,
<elevation id="CAFEE72E4B33FFF0615A75CAFC496C94" box="[799,935,1423,1450]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="131" unit="m" value="115.0">
<quantity id="862BADF84B33FFF0615A75CAFC866C94" box="[799,872,1423,1450]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="131" unit="m" value="115.0">115 m</quantity>
a.s.l.
</elevation>
(
<figureCitation id="D9E81C984B33FFF061F275CAFC156C94" box="[951,1019,1423,1450]" captionStart="Figure 1" captionStartId="1.[151,226,1019,1043]" captionTargetBox="[393,1196,606,1000]" captionTargetId="figure-224@1.[390,1196,606,1000]" captionTargetPageId="1" captionText="Figure 1. Faksimile of the original description of Amnicola charpentieri Roth, 1855" figureDoi="http://doi.org/10.5281/zenodo.13232494" httpUri="https://zenodo.org/record/13232494/files/figure.png" pageId="2" pageNumber="131">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B33FFF062D27596FBD86F31" pageId="2" pageNumber="131" type="etymology">
<paragraph id="416C001D4B33FFF062D27596FBD86F31" blockId="2.[151,1436,1490,1551]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D27596FEC66CD3" bold="true" box="[151,296,1491,1517]" pageId="2" pageNumber="131">Etymology:</emphasis>
Named in honor of Prof. Petar Beron (National Natural History Museum -
<collectingRegion id="8317CEFF4B33FFF0669D7597FAF66CD3" box="[1240,1304,1490,1517]" country="Bulgaria" name="Sofiya-Grad" pageId="2" pageNumber="131">Sofia</collectingRegion>
) who paid attention on this island and inspired D. Georgiev in its young age by his books.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B33FFF162D2767DFECC683E" lastPageId="3" lastPageNumber="132" pageId="2" pageNumber="131" type="description">
<paragraph id="416C001D4B33FFF062D2767DFAD66E23" blockId="2.[151,1437,1591,1821]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D2767DFEDC6F6C" bold="true" box="[151,306,1592,1618]" pageId="2" pageNumber="131">Description:</emphasis>
Shell: The corneous, small shell is ovate with a prominent body whorl. The 4-4.5 whorls are convex with a weak suture, with smooth surface and fine growth lines. The aperture is rounded ovate with a sharp periostome thickened at the columella. The umbilicus is closed (
<figureCitation id="D9E81C984B33FFF061AB763EFBD46FA8" box="[1006,1082,1659,1686]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="2" pageNumber="131">Fig. 2</figureCitation>
). The operculum is dark red. Animal: The animal is mostly dirty whitish or gray colored. The mantle is white or gray with a black wide stripe in its middle. The snout and tentacles are whitish or gray. Small patches of black pigment are visible around eyes and at the base of the snout. Penis: The penis is nearly regularly broad with a long tapered penis-tip and a bilobed outgrowth in the middle part on the left side. There is a dark spot near the penis-tip.
</paragraph>
<paragraph id="416C001D4B33FFF162D27703FECC683E" blockId="2.[151,1436,1861,1990]" lastBlockId="3.[151,1436,195,256]" lastPageId="3" lastPageNumber="132" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D27703FE256E5E" bold="true" box="[151,459,1862,1888]" pageId="2" pageNumber="131">Differentiating features:</emphasis>
<taxonomicName id="86D37B9E4B33FFF063927703FD5E6E5E" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[471,688,1861,1888]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="beroni" status="sp. nov.">
<emphasis id="73A7DC0F4B33FFF063927703FD5E6E5E" box="[471,688,1861,1888]" italics="true" pageId="2" pageNumber="131">Grossuana beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B33FFF060F97703FCEE6E5E" box="[700,768,1862,1888]" pageId="2" pageNumber="131" rank="species">n. sp.</taxonomicNameLabel>
is most similar to
<taxonomicName id="86D37B9E4B33FFF061B57703FF326EBC" authority="(Westerlund, 1881)" baseAuthorityName="Westerlund" baseAuthorityYear="1881" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="marginata">
<emphasis id="73A7DC0F4B33FFF061B57703FB166E61" box="[1008,1272,1862,1888]" italics="true" pageId="2" pageNumber="131">Grossuana marginata</emphasis>
(Westerlund, 1881)
</taxonomicName>
from which it differs by more regularly growing whorls, sharper and thinner periostome and lighter soft body pigmentation. The new species is also similar to
<taxonomicName id="86D37B9E4B33FFF0612F77CFFE6A6EF8" authority="Gloer, Reuselaars &amp; Papavasileiou, 2018" authorityName="Gloer, Reuselaars &amp; Papavasileiou" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="sidironerensis">
<emphasis id="73A7DC0F4B33FFF0612F77CFFB756E9A" box="[874,1179,1929,1956]" italics="true" pageId="2" pageNumber="131">Grossuana sidironerensis</emphasis>
<bibRefCitation id="25427DEC4B33FFF066E377CCFE6A6EF8" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." pageId="2" pageNumber="131" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer, Reuselaars &amp; Papavasileiou, 2018</bibRefCitation>
</taxonomicName>
described from the area of Drama at the mainland of
<collectingCountry id="39C4408D4B33FFF0664177EEFBB86EF8" box="[1028,1110,1963,1990]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
(
<bibRefCitation id="25427DEC4B33FFF0662377EEFAD76EF8" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." box="[1126,1337,1963,1990]" pageId="2" pageNumber="131" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer et al., 2018</bibRefCitation>
) but the last species has a penis with shorter tip, black colored mantle, and its whorls are fast growing (as in
<taxonomicName id="86D37B9E4B32FFF1673A7081FEFF683E" baseAuthorityName="Westerlund" baseAuthorityYear="1881" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="marginata">
<emphasis id="73A7DC0F4B32FFF1673A7081FEFF683E" italics="true" pageId="3" pageNumber="132">G. marginata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="09C953964B32FFF662D2716CFDC9683E" lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132" type="distribution">
<paragraph id="416C001D4B32FFF162D2716CFCDE68B9" blockId="3.[151,1436,297,391]" pageId="3" pageNumber="132">
<emphasis id="73A7DC0F4B32FFF162D2716CFED4687D" bold="true" box="[151,314,297,323]" pageId="3" pageNumber="132">Distribution:</emphasis>
Known only from few springs on Thassos Island: a spring (water source) south of Skala Potamia near the round road of the island (
<typeStatus id="9E68BEBF4B32FFF160CB710FFD2C685B" box="[654,706,330,357]" pageId="3" pageNumber="132">type</typeStatus>
locality), spring below Drakotrypa cave, Panagia village, and a big spring in the yard of the church of Rachoni village.
</paragraph>
<caption id="15AC50954B32FFF162D275ECFD866CC0" ID-DOI="http://doi.org/10.5281/zenodo.13232498" ID-Zenodo-Dep="13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="3" pageNumber="132" startId="3.[151,236,1449,1473]" targetBox="[151,1439,404,1431]" targetPageId="3" targetType="figure">
<paragraph id="416C001D4B32FFF162D275ECFD866CC0" blockId="3.[151,1436,1449,1535]" pageId="3" pageNumber="132">
<emphasis id="73A7DC0F4B32FFF162D275ECFEF66CFF" bold="true" box="[151,280,1449,1473]" pageId="3" pageNumber="132">Figures 3-5</emphasis>
.
<emphasis id="73A7DC0F4B32FFF1636D75ECFEDB6CFF" bold="true" box="[296,309,1449,1473]" pageId="3" pageNumber="132">3</emphasis>
: Type locality of
<taxonomicName id="86D37B9E4B32FFF163BE75ECFD516CFF" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[507,703,1449,1473]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="beroni" status="sp. nov.">
<emphasis id="73A7DC0F4B32FFF163BE75ECFD516CFF" box="[507,703,1449,1473]" italics="true" pageId="3" pageNumber="132">Grossuana beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF1608D75ECFCEA6CFE" box="[712,772,1449,1472]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
, water source south of Skala Potamia near the round road of the island.
<emphasis id="73A7DC0F4B32FFF163557582FEF36CE1" bold="true" box="[272,285,1479,1503]" pageId="3" pageNumber="132">4</emphasis>
: Type locality of
<taxonomicName id="86D37B9E4B32FFF163A2758DFD486CE1" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[487,678,1479,1503]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
<emphasis id="73A7DC0F4B32FFF163A2758DFD486CE1" box="[487,678,1479,1503]" italics="true" pageId="3" pageNumber="132">Grossuana thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF160F5758DFD036CE1" box="[688,749,1480,1503]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
, a spring south of Kinira near the round road of the island.
<emphasis id="73A7DC0F4B32FFF167C27582FA7A6CE1" bold="true" box="[1415,1428,1479,1503]" pageId="3" pageNumber="132">5</emphasis>
: Type locality of
<taxonomicName id="86D37B9E4B32FFF1630D75A3FDC66CC0" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[328,552,1510,1534]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="rachonica" status="sp. nov.">
<emphasis id="73A7DC0F4B32FFF1630D75A3FDC66CC0" box="[328,552,1510,1534]" italics="true" pageId="3" pageNumber="132">Bythinella rachonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF1606A75A2FD866CC0" box="[559,616,1511,1534]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="15AC50954B32FFF162D277F1FDD46ED5" ID-DOI="http://doi.org/10.5281/zenodo.13232500" ID-Zenodo-Dep="13232500" httpUri="https://zenodo.org/record/13232500/files/figure.png" pageId="3" pageNumber="132" startId="3.[151,236,1972,1996]" targetBox="[151,1444,1565,1953]" targetPageId="3" targetType="figure">
<paragraph id="416C001D4B32FFF162D277F1FDD46ED5" blockId="3.[151,1436,1972,2027]" pageId="3" pageNumber="132">
<emphasis id="73A7DC0F4B32FFF162D277F1FEC06EF2" bold="true" box="[151,302,1972,1996]" pageId="3" pageNumber="132">Figures 6-10.</emphasis>
The
<taxonomicName id="86D37B9E4B32FFF1632977F0FDC16EF2" box="[364,559,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="undefined-6">
<emphasis id="73A7DC0F4B32FFF1632977F0FE0F6EF2" box="[364,481,1973,1996]" italics="true" pageId="3" pageNumber="132">Grossuana</emphasis>
spp.
<emphasis id="73A7DC0F4B32FFF1606777F1FDC16EF2" bold="true" box="[546,559,1972,1996]" pageId="3" pageNumber="132">6</emphasis>
</taxonomicName>
<emphasis id="73A7DC0F4B32FFF1606A77F1FDAB6EF2" bold="true" box="[559,581,1972,1996]" pageId="3" pageNumber="132">-7</emphasis>
:
<taxonomicName id="86D37B9E4B32FFF1601277F0FD2F6EF2" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[599,705,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="beroni" status="n. sp.">
<emphasis id="73A7DC0F4B32FFF1601277F0FD2F6EF2" box="[599,705,1972,1996]" italics="true" pageId="3" pageNumber="132">G. beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF1608E77F0FCE66EF2" box="[715,776,1973,1996]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
(paratypes),
<emphasis id="73A7DC0F4B32FFF161DE77F1FC466EF2" bold="true" box="[923,936,1972,1996]" pageId="3" pageNumber="132">8</emphasis>
: penis of
<taxonomicName id="86D37B9E4B32FFF1665877F0FB696EF2" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1053,1159,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="beroni" status="n. sp.">
<emphasis id="73A7DC0F4B32FFF1665877F0FB696EF2" box="[1053,1159,1972,1996]" italics="true" pageId="3" pageNumber="132">G. beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF166D477F0FB206EF2" box="[1169,1230,1973,1996]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="73A7DC0F4B32FFF1669D77F1FB0B6EF2" bold="true" box="[1240,1253,1972,1996]" pageId="3" pageNumber="132">9</emphasis>
:
<taxonomicName id="86D37B9E4B32FFF166B277F0FAB26EF2" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1271,1372,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="thasia">
<emphasis id="73A7DC0F4B32FFF166B277F0FAB26EF2" box="[1271,1372,1972,1996]" italics="true" pageId="3" pageNumber="132">G. thasia</emphasis>
</taxonomicName>
n. sp (holotype),
<emphasis id="73A7DC0F4B32FFF163517796FEC16ED5" bold="true" box="[276,303,2003,2027]" pageId="3" pageNumber="132">10</emphasis>
: penis of
<taxonomicName id="86D37B9E4B32FFF163DD7791FE146ED5" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[408,506,2003,2027]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="thasia" status="n. sp.">
<emphasis id="73A7DC0F4B32FFF163DD7791FE146ED5" box="[408,506,2003,2027]" italics="true" pageId="3" pageNumber="132">G. thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF160447791FDD46ED5" box="[513,570,2004,2027]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="416C001D4B35FFF662D27081FDC9683E" blockId="4.[151,1436,195,256]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D27081FEEF69E0" bold="true" box="[151,257,196,222]" pageId="4" pageNumber="133">Habitat:</emphasis>
Karst spring in a mixed broad leaf forest (mainly
<taxonomicName id="86D37B9E4B35FFF6611C7086FBD469E0" box="[857,1082,195,222]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="73A7DC0F4B35FFF6611C7086FBD469E0" box="[857,1082,195,222]" italics="true" pageId="4" pageNumber="133">Platanus orientalis</emphasis>
</taxonomicName>
and
<taxonomicName id="86D37B9E4B35FFF666337081FAF869E0" box="[1142,1302,195,222]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B35FFF666337081FB3469E0" box="[1142,1242,196,222]" italics="true" pageId="4" pageNumber="133">Quercus</emphasis>
spp.
</taxonomicName>
). Found on moss, dry leaves, sand and stones.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,193 @@
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<mods:title id="3ECF5FBB46A791C1440CEED722404864">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<mods:namePart id="E51E28E48F335018E4D39D15CAEFB751">Georgiev, Dilian</mods:namePart>
<mods:affiliation id="EF5F94E60E0DE74FC5A2343A1764E305">Department of Ecology and Nature Conservation, Faculty of Biology, Paisii Hilendarski University of Plovdiv, 24 Tsar Assen Str., 4000 Plovdiv, Bulgaria</mods:affiliation>
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<mods:affiliation id="286281576F24DA41163BD4981E5A355C">Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany Corresponding author. E-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<mods:title id="61DC7CECB133D8D7B5F433C0B8927152">Ecologica Montenegrina</mods:title>
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<mods:date id="613B341EAC1B933D01515EE4091E8171">2020</mods:date>
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<mods:number id="36D7B2FDB772C3A753DC689AC046E3CD">2020-10-22</mods:number>
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<mods:url id="BE5445F8317BE3852B4FCE2042E1E85A">http://dx.doi.org/10.37828/em.2020.35.10</mods:url>
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<mods:classification id="5BB9AF30E90EE98A54EB24D890F216A4">journal article</mods:classification>
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<subSubSection id="09C953964B34FFF762D276F8FB9D6FCB" pageId="5" pageNumber="134" type="nomenclature">
<paragraph id="416C001D4B34FFF762D276F8FC606FE9" blockId="5.[151,1139,1724,1781]" box="[151,910,1724,1751]" pageId="5" pageNumber="134">
<heading id="1A24B7714B34FFF762D276F8FC606FE9" box="[151,910,1724,1751]" fontSize="11" level="2" pageId="5" pageNumber="134" reason="2">
<emphasis id="73A7DC0F4B34FFF762D276F8FD3E6FE9" bold="true" box="[151,720,1725,1751]" pageId="5" pageNumber="134">
<taxonomicName id="86D37B9E4B34FFF762D276F8FD686FE9" ID-CoL="7QHRB" authority="Gloer &amp; Georgiev" authorityName="Gloer &amp; Georgiev" authorityYear="2020" box="[151,646,1725,1751]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="134" phylum="Mollusca" rank="species" species="reischuetzi" status="sp. nov.">
<emphasis id="73A7DC0F4B34FFF762D276F8FE706FE9" bold="true" box="[151,414,1725,1751]" italics="true" pageId="5" pageNumber="134">Bythinella reischuetzi</emphasis>
Glöer &amp; Georgiev
</taxonomicName>
<taxonomicNameLabel id="689461744B34FFF760CB76F8FD3E6FE9" box="[654,720,1725,1751]" pageId="5" pageNumber="134" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D9E81C984B34FFF760A476F9FCDD6FE9" box="[737,819,1724,1751]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="5" pageNumber="134">Figs. 2</figureCitation>
,
<figureCitation id="D9E81C984B34FFF7610576F9FC6B6FE9" box="[832,901,1724,1751]" captionStart="Figs 18-21" captionStartId="6.[151,198,807,831]" captionTargetBox="[151,1435,302,788]" captionTargetId="figure-59@6.[151,1435,302,788]" captionTargetPageId="6" captionText="Figs 18-21. Bythinella reischuetzi n. sp. 18: holotype, 19: original label, 20: paratype, 21: apical view." figureDoi="http://doi.org/10.5281/zenodo.13232504" httpUri="https://zenodo.org/record/13232504/files/figure.png" pageId="5" pageNumber="134">18-21</figureCitation>
)
</heading>
</paragraph>
<paragraph id="416C001D4B34FFF762D27698FB9D6FCB" blockId="5.[151,1139,1724,1781]" box="[151,1139,1757,1781]" pageId="5" pageNumber="134">
https://zoobank.org/
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<subSubSection id="09C953964B34FFF762D2775AFC566EFE" pageId="5" pageNumber="134" type="materials_examined">
<paragraph id="416C001D4B34FFF762D2775AFE956E07" blockId="5.[151,1243,1823,1916]" box="[151,379,1823,1849]" pageId="5" pageNumber="134">
<emphasis id="73A7DC0F4B34FFF762D2775AFE956E07" bold="true" box="[151,379,1823,1849]" pageId="5" pageNumber="134">Materials studied:</emphasis>
</paragraph>
<paragraph id="416C001D4B34FFF762B07705FB356E65" blockId="5.[151,1243,1823,1916]" box="[245,1243,1856,1883]" pageId="5" pageNumber="134">
<materialsCitation id="F1BB0A404B34FFF762B07705FB356E65" box="[245,1243,1856,1883]" collectionCode="ZSM" collectorName="Mol &amp; Roth." pageId="5" pageNumber="134" specimenCount="1" typeStatus="holotype">
<emphasis id="73A7DC0F4B34FFF762B07705FE896E64" bold="true" box="[245,359,1856,1882]" pageId="5" pageNumber="134">
<typeStatus id="9E68BEBF4B34FFF762B07705FE896E64" box="[245,359,1856,1882]" pageId="5" pageNumber="134" type="holotype">Holotype</typeStatus>
</emphasis>
: shell height
<quantity id="862BADF84B34FFF760437705FD8D6E65" box="[518,611,1856,1883]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="134" unit="mm" value="2.5">2.5 mm</quantity>
, shell width
<quantity id="862BADF84B34FFF760B27705FCBA6E65" box="[759,852,1856,1883]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="134" unit="mm" value="1.7">1.7 mm</quantity>
,
<collectionCode id="27C298D84B34FFF761257705FC726E65" box="[864,924,1856,1883]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="134" type="Museum">ZSM</collectionCode>
<collectorName id="EC2665CB4B34FFF761E67705FC3A6E65" box="[931,980,1856,1883]" pageId="5" pageNumber="134">Mol</collectorName>
20202395, coll.
<collectorName id="EC2665CB4B34FFF766DF7705FB356E65" box="[1178,1243,1856,1883]" pageId="5" pageNumber="134">Roth.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="416C001D4B34FFF762B07727FB8E6E42" blockId="5.[151,1243,1823,1916]" box="[245,1120,1889,1916]" pageId="5" pageNumber="134">
<materialsCitation id="F1BB0A404B34FFF762B07727FB8E6E42" box="[245,1120,1889,1916]" collectingDate="2100-01-20" collectionCode="ZSM" collectorName="Mol &amp; Roth." pageId="5" pageNumber="134" specimenCount="1" typeStatus="paratype">
<emphasis id="73A7DC0F4B34FFF762B07727FE9C6E42" bold="true" box="[245,370,1890,1916]" pageId="5" pageNumber="134">
<typeStatus id="9E68BEBF4B34FFF762B07727FE9C6E42" box="[245,370,1890,1916]" pageId="5" pageNumber="134" type="paratype">Paratypes</typeStatus>
</emphasis>
: numerous (&gt; 150) specimens,
<collectionCode id="27C298D84B34FFF760A07724FCCF6E42" box="[741,801,1889,1916]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="134" type="Museum">ZSM</collectionCode>
<collectorName id="EC2665CB4B34FFF7616D7724FCB76E42" box="[808,857,1889,1916]" pageId="5" pageNumber="134">Mol</collectorName>
<date id="356D26DD4B34FFF761247724FC376E42" box="[865,985,1889,1916]" pageId="5" pageNumber="134" value="2100-01-20">
<collectingDate id="2529DF354B34FFF761247724FC376E42" box="[865,985,1889,1916]" pageId="5" pageNumber="134" value="2100-01-20">20012100</collectingDate>
</date>
, coll.
<collectorName id="EC2665CB4B34FFF7665A7724FB8E6E42" box="[1055,1120,1889,1916]" pageId="5" pageNumber="134">Roth.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="416C001D4B34FFF762D277E3FC566EFE" blockId="5.[151,952,1957,1984]" box="[151,952,1957,1984]" pageId="5" pageNumber="134">
<emphasis id="73A7DC0F4B34FFF762D277E3FED76EFE" bold="true" box="[151,313,1958,1984]" pageId="5" pageNumber="134">Type locality</emphasis>
:
<collectingCountry id="39C4408D4B34FFF7630D77E0FE706EFE" box="[328,414,1957,1984]" name="Greece" pageId="5" pageNumber="134">Greece</collectingCountry>
, Párnis (Mountain) in Parnitha National Park.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B37FFF462D27081FE62681C" pageId="6" pageNumber="135" type="etymology">
<paragraph id="416C001D4B37FFF462D27081FE62681C" blockId="6.[151,1436,195,290]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D27081FEC669E0" bold="true" box="[151,296,196,222]" pageId="6" pageNumber="135">Etymology:</emphasis>
Named after Peter L. Reischütz (Vienna), outstanding expert on Hydrobiids of the Balkans, who already pointed out in 2008 (p. 35) that the
<taxonomicName id="86D37B9E4B37FFF460F570A0FC8D683E" box="[688,867,229,256]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B37FFF460F570A0FCC6683E" box="[688,808,229,256]" italics="true" pageId="6" pageNumber="135">Bythinella</emphasis>
spp.
</taxonomicName>
of the Peleponnese is much more diverse than believed at that time.
</paragraph>
</subSubSection>
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<paragraph id="416C001D4B37FFF462D27362FB346A00" blockId="6.[151,1242,806,831]" box="[151,1242,806,831]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D27362FEE26A01" bold="true" box="[151,268,807,831]" pageId="6" pageNumber="135">Figs 18-21</emphasis>
.
<taxonomicName id="86D37B9E4B37FFF4635C7363FDEE6A00" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[281,512,806,830]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="reischuetzi" status="sp. nov.">
<emphasis id="73A7DC0F4B37FFF4635C7363FDEE6A00" box="[281,512,806,830]" italics="true" pageId="6" pageNumber="135">Bythinella reischuetzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B37FFF460427362FDAE6A00" box="[519,576,807,830]" pageId="6" pageNumber="135" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="73A7DC0F4B37FFF460027362FD8C6A01" bold="true" box="[583,610,807,831]" pageId="6" pageNumber="135">18</emphasis>
: holotype,
<emphasis id="73A7DC0F4B37FFF4609E7362FD186A01" bold="true" box="[731,758,807,831]" pageId="6" pageNumber="135">19</emphasis>
: original label,
<emphasis id="73A7DC0F4B37FFF461DB7362FC576A01" bold="true" box="[926,953,807,831]" pageId="6" pageNumber="135">20</emphasis>
: paratype,
<emphasis id="73A7DC0F4B37FFF466757362FBA56A01" bold="true" box="[1072,1099,807,831]" pageId="6" pageNumber="135">21</emphasis>
: apical view.
</paragraph>
</caption>
<subSubSection id="09C953964B37FFF462D273CFFB656D51" pageId="6" pageNumber="135" type="description">
<paragraph id="416C001D4B37FFF462D273CFFEAD6D34" blockId="6.[151,1436,905,1034]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D273CFFEC66A9A" bold="true" box="[151,296,906,932]" pageId="6" pageNumber="135">Description</emphasis>
: The ovate shell is thick-walled with 4-4.5 slightly convex whorls, separated by a clear suture. The first whorl lies in a plane and is not visible from frontal view. The aperture is ovate with a slight angle at the top. The peristome is thickened at the columella. The umbilicus is closed. The shell is
<quantity id="862BADF84B37FFF466947388FAC36AD6" box="[1233,1325,973,1000]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="135" unit="mm" value="2.5">2.5 mm</quantity>
high and
<quantity id="862BADF84B37FFF462D273AAFF1F6D37" box="[151,241,1007,1034]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="135" unit="mm" value="1.7">1.7 mm</quantity>
broad.
</paragraph>
<paragraph id="416C001D4B37FFF462D27476FB656D51" blockId="6.[151,1435,1074,1135]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D27476FE306D73" bold="true" box="[151,478,1075,1101]" pageId="6" pageNumber="135">Differentiating characters</emphasis>
: The species differs from all other
<taxonomicName id="86D37B9E4B37FFF461CB7477FBAF6D73" box="[910,1089,1074,1101]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B37FFF461CB7477FBE86D73" box="[910,1030,1074,1101]" italics="true" pageId="6" pageNumber="135">Bythinella</emphasis>
spp.
</taxonomicName>
of
<collectingCountry id="39C4408D4B37FFF466287477FB516D73" box="[1133,1215,1074,1101]" name="Greece" pageId="6" pageNumber="135">Greece</collectingCountry>
by the first whorl which lies in a plane. Thus the compressed shell shape is characteristic of this species.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B37FFF462D274DDFD5C6D8C" box="[151,690,1175,1202]" pageId="6" pageNumber="135" type="distribution">
<paragraph id="416C001D4B37FFF462D274DDFD5C6D8C" blockId="6.[151,690,1175,1202]" box="[151,690,1175,1202]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D274DDFEDE6D8C" bold="true" box="[151,304,1176,1202]" pageId="6" pageNumber="135">Distribution</emphasis>
: Only known from
<typeStatus id="9E68BEBF4B37FFF4605C74DDFDA56D8C" box="[537,587,1176,1202]" pageId="6" pageNumber="135">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B37FFF462D2749EFABF6C29" pageId="6" pageNumber="135" type="discussion">
<paragraph id="416C001D4B37FFF462D2749EFABF6C29" blockId="6.[151,1436,1243,1303]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D2749EFF136DCB" bold="true" box="[151,253,1243,1269]" pageId="6" pageNumber="135">Remark</emphasis>
: In this region another
<taxonomicName id="86D37B9E4B37FFF4606C749EFD3E6DC8" box="[553,720,1243,1270]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B37FFF4606C749EFD4F6DC8" box="[553,673,1243,1270]" italics="true" pageId="6" pageNumber="135">Bythinella</emphasis>
sp.
</taxonomicName>
occurred (see
<figureCitation id="D9E81C984B37FFF461CF749EFC0A6DC8" box="[906,996,1243,1270]" captionStart="Figures 11-17" captionStartId="5.[151,236,1533,1557]" captionTargetBox="[151,1429,336,1515]" captionTargetId="figure-74@5.[151,1429,336,1515]" captionTargetPageId="5" captionText="Figures 11-17. Bythinella charpentieri, syntypes and others from colletions ZSM and SMF. 11: lectotype of B. charpentieri Roth, 1855 from Roths collection ZSM 20012106. 12: paralectotype, Hymettus, ZSM29912124, 13: topotype, Hymettus SMF141584, 14: Kessisos ZSM20012113, 15: Attika ZSM20012106, 16: Attika, ZSM20012110, 17: Parnes, SMF141838." figureDoi="http://doi.org/10.5281/zenodo.13232502" httpUri="https://zenodo.org/record/13232502/files/figure.png" pageId="6" pageNumber="135">Fig. 15</figureCitation>
), distinct from
<taxonomicName id="86D37B9E4B37FFF466ED7499FAA56DC8" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1192,1355,1243,1270]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="reischuetzi" status="n. sp.">
<emphasis id="73A7DC0F4B37FFF466ED7499FAA56DC8" box="[1192,1355,1243,1270]" italics="true" pageId="6" pageNumber="135">B. reischuetzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B37FFF4671D7499FA726DC8" box="[1368,1436,1244,1270]" pageId="6" pageNumber="135" rank="species">n. sp.</taxonomicNameLabel>
Unfortunately we do not know the exact locations because these have not been mentioned at the labels.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,270 @@
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<mods:title id="13E1A82AC3B88CDF931C18CA394CF627">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<mods:namePart id="511F3ED2410544374829877D9838B1E7">Glöer, Peter</mods:namePart>
<mods:affiliation id="815668B406789314C91802B08131532A">Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany Corresponding author. E-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<mods:title id="7507E27D5A4F5378617554DD8A0C5713">Ecologica Montenegrina</mods:title>
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<mods:date id="7FEF73E1F3B040D0AFD1D627E104ED36">2020</mods:date>
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<mods:number id="E06D0D9E34D2439BD176D4BD09EF2F79">2020-10-22</mods:number>
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<mods:url id="DBABE4A95BDCFE4923FB797B708284F1">http://dx.doi.org/10.37828/em.2020.35.10</mods:url>
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<mods:identifier id="1C4874D5AE3018752F83CBBC441E9849" type="DOI">10.37828/em.2020.35.10</mods:identifier>
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<treatment id="C97AB10B4B35FFF662D2710EFDC96F96" ID-DOI="http://doi.org/10.5281/zenodo.13232609" ID-Zenodo-Dep="13232609" LSID="urn:lsid:zoobank.org:act:019E5D1D-E033-4913-896D-4C779ACA9198" httpUri="http://treatment.plazi.org/id/C97AB10B4B35FFF662D2710EFDC96F96" lastPageNumber="133" pageId="4" pageNumber="133">
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<emphasis id="73A7DC0F4B35FFF662D2710EFE58685B" bold="true" box="[151,438,331,357]" pageId="4" pageNumber="133">
<taxonomicName id="86D37B9E4B35FFF662D2710EFE83685B" ID-CoL="7RG9Y" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[151,365,331,357]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
<emphasis id="73A7DC0F4B35FFF662D2710EFE83685B" bold="true" box="[151,365,331,357]" italics="true" pageId="4" pageNumber="133">Grossuana thasia</emphasis>
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</emphasis>
(
<figureCitation id="D9E81C984B35FFF66382710FFDF6685B" box="[455,536,330,357]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="4" pageNumber="133">Figs. 2</figureCitation>
,
<figureCitation id="D9E81C984B35FFF66060710FFDD8685B" box="[549,566,330,357]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="4" pageNumber="133">4</figureCitation>
,
<figureCitation id="D9E81C984B35FFF66007710FFD97685B" box="[578,633,330,357]" captionStart="Figures 6-10" captionStartId="3.[151,236,1972,1996]" captionTargetBox="[151,1444,1565,1953]" captionTargetId="figure-136@3.[151,1444,1565,1953]" captionTargetPageId="3" captionText="Figures 6-10. The Grossuana spp. 6-7: G. beroni n. sp. (paratypes), 8: penis of G. beroni n. sp. 9: G. thasia n. sp (holotype), 10: penis of G. thasia n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232500" httpUri="https://zenodo.org/record/13232500/files/figure.png" pageId="4" pageNumber="133">9-10</figureCitation>
)
</heading>
</paragraph>
<paragraph id="416C001D4B35FFF662D2712EFB9D68BD" blockId="4.[151,1139,330,387]" box="[151,1139,363,387]" pageId="4" pageNumber="133">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</subSubSection>
<subSubSection id="09C953964B35FFF662D271E8FC786BAC" pageId="4" pageNumber="133" type="materials_examined">
<paragraph id="416C001D4B35FFF662D271E8FE9E68F9" blockId="4.[151,1427,429,557]" box="[151,368,429,455]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D271E8FE9E68F9" bold="true" box="[151,368,429,455]" pageId="4" pageNumber="133">Material studied:</emphasis>
</paragraph>
<paragraph id="416C001D4B35FFF662B0718AFB5668D7" blockId="4.[151,1427,429,557]" box="[245,1208,462,489]" pageId="4" pageNumber="133">
<materialsCitation id="F1BB0A404B35FFF662B0718AFB5668D7" box="[245,1208,462,489]" collectionCode="ZMH" pageId="4" pageNumber="133" specimenCode="ZMH 140793" specimenCount="1" typeStatus="holotype">
<emphasis id="73A7DC0F4B35FFF662B0718AFE9F68D7" bold="true" box="[245,369,463,489]" pageId="4" pageNumber="133">
<typeStatus id="9E68BEBF4B35FFF662B0718AFE8268D7" box="[245,364,463,489]" pageId="4" pageNumber="133" type="holotype">Holotype</typeStatus>
:
</emphasis>
shell height
<quantity id="862BADF84B35FFF6604D718BFD8A68D7" box="[520,612,462,489]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="133" unit="mm" value="1.7">1.7 mm</quantity>
, width
<quantity id="862BADF84B35FFF660FF718BFCF968D7" box="[698,791,462,489]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="133" unit="mm" value="1.1">1.1 mm</quantity>
, from type locality (
<specimenCode id="1175A8664B35FFF66646718BFB4568D7" box="[1027,1195,462,489]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="133" type="Museum">ZMH 140793</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="416C001D4B35FFF662B071B4FEE26B13" blockId="4.[151,1427,429,557]" pageId="4" pageNumber="133">
<materialsCitation id="F1BB0A404B35FFF662B071B4FCA06B35" box="[245,846,496,523]" collectionCode="ZMH" pageId="4" pageNumber="133" specimenCode="ZMH 140794" specimenCount="5" typeStatus="paratype">
<emphasis id="73A7DC0F4B35FFF662B071B4FE926B35" bold="true" box="[245,380,497,523]" pageId="4" pageNumber="133">
<typeStatus id="9E68BEBF4B35FFF662B071B4FE996B35" box="[245,375,497,523]" pageId="4" pageNumber="133" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="57D5CB944B35FFF663C171B5FDFD6B35" box="[388,531,496,523]" count="5" pageId="4" pageNumber="133" type="generic">5 specimens</specimenCount>
in ethanol (
<specimenCode id="1175A8664B35FFF660DA71B5FCA86B35" box="[671,838,496,523]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="133" type="Museum">ZMH 140794</specimenCode>
)
</materialsCitation>
,
<specimenCount id="57D5CB944B35FFF6611F71B5FC076B35" box="[858,1001,496,523]" count="5" pageId="4" pageNumber="133" type="generic">5 specimens</specimenCount>
in coll. Glöer,
<specimenCount id="57D5CB944B35FFF666DF71B5FAD56B35" box="[1178,1339,496,523]" count="11" pageId="4" pageNumber="133" type="generic">11 specimens</specimenCount>
in coll. Georgiev.
</paragraph>
<paragraph id="416C001D4B35FFF662D27213FC786BAC" blockId="4.[151,1435,597,658]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D27213FEAD6B4E" bold="true" box="[151,323,598,624]" pageId="4" pageNumber="133">Type locality:</emphasis>
<materialsCitation id="F1BB0A404B35FFF6630E7210FC7C6BAC" country="Greece" elevation="144" latitude="40.635418" location="Kinira" longLatPrecision="1" longitude="24.7645" municipality="Thassos Island" pageId="4" pageNumber="133" specimenCount="1" stateProvince="Voreio Aigaio">
<collectingCountry id="39C4408D4B35FFF6630E7210FE4F6B4E" box="[331,417,597,624]" name="Greece" pageId="4" pageNumber="133">Greece</collectingCountry>
,
<collectingMunicipality id="A1089A674B35FFF663E87210FDB06B4E" box="[429,606,597,624]" pageId="4" pageNumber="133">Thassos Island</collectingMunicipality>
,
<collectingRegion id="8317CEFF4B35FFF6602F7210FCFD6B4E" box="[618,787,597,624]" country="Greece" name="Voreio Aigaio" pageId="4" pageNumber="133">North Aegean</collectingRegion>
, a spring (water source) south of
<location id="440C56C64B35FFF666DB7210FB066B4E" LSID="urn:lsid:plazi:treatment:C97AB10B4B35FFF662D2710EFDC96F96:440C56C64B35FFF666DB7210FB066B4E" box="[1182,1256,597,624]" country="Greece" latitude="40.635418" longLatPrecision="1" longitude="24.7645" municipality="Thassos Island" name="Kinira" pageId="4" pageNumber="133" stateProvince="Voreio Aigaio">Kinira</location>
near the round road of the island,
<geoCoordinate id="24E766DA4B35FFF663377232FDEA6BAC" box="[370,516,631,658]" degrees="40" direction="north" minutes="38" orientation="latitude" pageId="4" pageNumber="133" precision="1" seconds="07.5" value="40.635418">N40 38 07.5</geoCoordinate>
<geoCoordinate id="24E766DA4B35FFF6604E7232FD726BAC" box="[523,668,631,658]" degrees="24" direction="east" minutes="45" orientation="longitude" pageId="4" pageNumber="133" precision="1" seconds="52.2" value="24.7645">E24 45 52.2</geoCoordinate>
,
<elevation id="CAFEE72E4B35FFF660ED7232FCDE6BAC" box="[680,816,631,658]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="4" pageNumber="133" unit="m" value="144.0">
<quantity id="862BADF84B35FFF660ED7232FD1C6BAC" box="[680,754,631,658]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="4" pageNumber="133" unit="m" value="144.0">144 m</quantity>
a.s.l.
</elevation>
(
<figureCitation id="D9E81C984B35FFF661057232FC686BAC" box="[832,902,631,658]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="4" pageNumber="133">Fig. 3</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B35FFF662D272FEFC596BEB" box="[151,951,698,725]" pageId="4" pageNumber="133" type="etymology">
<paragraph id="416C001D4B35FFF662D272FEFC596BEB" blockId="4.[151,951,698,725]" box="[151,951,698,725]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D272FEFEC66BEB" bold="true" box="[151,296,699,725]" pageId="4" pageNumber="133">Etymology:</emphasis>
Named after Thassos Island on which the species lives.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B35FFF662D272BBFBC36CC1" pageId="4" pageNumber="133" type="description">
<paragraph id="416C001D4B35FFF662D272BBFEEB6ADD" blockId="4.[151,1436,766,995]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D272BBFEDC6A26" bold="true" box="[151,306,766,792]" pageId="4" pageNumber="133">Description:</emphasis>
Shell: The corneous shell is elongate ovate with a prolonged body whorl (
<figureCitation id="D9E81C984B35FFF666EA72BBFB146A27" box="[1199,1274,766,793]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="4" pageNumber="133">Fig. 2</figureCitation>
). The apex is rounded and often eroded. The 4-4.5 whorls are slightly convex with a weak suture, with rough irregular growth lines. The aperture is ovate slightly angled at the top with a sharp periostome thickened at the columella. The umbilicus is closed. The operculum is dark red. Animal: The animal is mostly black colored. The mantle is black with a thin white border. The snout and tentacles are black with white or grayish tips. Penis: The penis is nearly regularly broad with a long tapered penis-tip. There is a dark spot at the near the penis-tip.
</paragraph>
<paragraph id="416C001D4B35FFF662D27449FEB16C2D" blockId="4.[151,1436,1036,1535]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D27449FE246D18" bold="true" box="[151,458,1036,1062]" pageId="4" pageNumber="133">Differentiating features:</emphasis>
The new species is most similar to
<taxonomicName id="86D37B9E4B35FFF661C87448FF3C6D77" authority="Gloer &amp; Georgiev, 2009" authorityName="Gloer &amp; Georgiev" authorityYear="2009" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="angeltsekovi">
<emphasis id="73A7DC0F4B35FFF661C87448FB426D19" box="[909,1196,1036,1063]" italics="true" pageId="4" pageNumber="133">Grossuana angeltsekovi</emphasis>
<bibRefCitation id="25427DEC4B35FFF666F27449FF3C6D77" author="Gloer, P. &amp; Georgiev, D." pageId="4" pageNumber="133" pagination="123 - 136" refId="ref4427" refString="Gloer, P., &amp; Georgiev, D. (2009) New Rissooidea from Bulgaria (Gastropoda: Rissooidea). Mollusca, 27 (2), 123 - 136." type="journal article" year="2009">Glöer &amp; Georgiev, 2009</bibRefCitation>
</taxonomicName>
which is known to be widely distributed in
<collectingCountry id="39C4408D4B35FFF66092746BFCD26D77" box="[727,828,1070,1097]" name="Bulgaria" pageId="4" pageNumber="133">Bulgaria</collectingCountry>
and North
<collectingCountry id="39C4408D4B35FFF66187746BFBFB6D77" box="[962,1045,1070,1097]" name="Greece" pageId="4" pageNumber="133">Greece</collectingCountry>
(
<bibRefCitation id="25427DEC4B35FFF66661746BFB376D77" author="Georgiev, D." box="[1060,1241,1070,1097]" pageId="4" pageNumber="133" pagination="1 - 4" refId="ref4338" refString="Georgiev, D. (2013) First record of Grossuana angeltsekovi Gloer &amp; Georgiev, 2009 (Gastropoda: Risooidea) from Greece. ZooNotes, 37, 1 - 4." type="journal article" year="2013">Georgiev 2013</bibRefCitation>
,
<bibRefCitation id="25427DEC4B35FFF666A0746BFF3B6D54" author="Georgiev, D. &amp; Gloer, P. &amp; Dedov, I. &amp; Irikov, A." pageId="4" pageNumber="133" pagination="159 - 166" refId="ref4371" refString="Georgiev, D., Gloer, P., Dedov, I. &amp; Irikov, A. (2015) Review of the Genus Grossuana Radoman, 1973 (Gastropoda: Truncatelloidea) from Bulgaria, with a description of a new species. Acta Zoologica Bulgarica, 67, 159 - 166." type="journal article" year="2015">Georgiev et al., 2015</bibRefCitation>
, Falniowski et al. 2015,
<bibRefCitation id="25427DEC4B35FFF66041740AFD0E6D54" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." box="[516,736,1103,1130]" pageId="4" pageNumber="133" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer et al., 2018</bibRefCitation>
). From this species
<taxonomicName id="86D37B9E4B35FFF661937415FB476D54" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[982,1193,1103,1130]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
<emphasis id="73A7DC0F4B35FFF661937415FB476D54" box="[982,1193,1103,1130]" italics="true" pageId="4" pageNumber="133">Grossuana thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B35FFF666F07415FB196D54" box="[1205,1271,1104,1130]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
differs by its more elongated and slender shell with flatter whorls, which have rough surface with irregular growth lines, and by its more rounded, often eroded apex. The whorls of the shell of the new species are 4-4.5 while in
<taxonomicName id="86D37B9E4B35FFF667C574D1FEC76DF1" authorityName="Gloer &amp; Georgiev" authorityYear="2009" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="angeltsekovi">
<emphasis id="73A7DC0F4B35FFF667C574D1FEC76DF1" italics="true" pageId="4" pageNumber="133">G. angeltsekovi</emphasis>
</taxonomicName>
they are 5-5.5 and the later is larger (height of the
<typeStatus id="9E68BEBF4B35FFF661C574F1FC086DF1" box="[896,998,1204,1231]" pageId="4" pageNumber="133" type="holotype">holotype</typeStatus>
=
<quantity id="862BADF84B35FFF6664274F1FB886DF1" box="[1031,1126,1204,1231]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="133" unit="mm" value="2.1">2.1 mm</quantity>
). In addition
<taxonomicName id="86D37B9E4B35FFF6674474F0FA806DF1" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1281,1390,1204,1231]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia">
<emphasis id="73A7DC0F4B35FFF6674474F0FA806DF1" box="[1281,1390,1204,1231]" italics="true" pageId="4" pageNumber="133">G. thasia</emphasis>
</taxonomicName>
has a well visible bifurcated penis lobe, while in
<taxonomicName id="86D37B9E4B35FFF660877492FC956DCF" authorityName="Gloer &amp; Georgiev" authorityYear="2009" box="[706,891,1238,1265]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="angeltsekovi">
<emphasis id="73A7DC0F4B35FFF660877492FC956DCF" box="[706,891,1238,1265]" italics="true" pageId="4" pageNumber="133">G. angeltsekovi</emphasis>
</taxonomicName>
it is very small and hardly visible (
<bibRefCitation id="25427DEC4B35FFF667707493FEBD6C2D" author="Gloer, P. &amp; Georgiev, D." pageId="4" pageNumber="133" pagination="123 - 136" refId="ref4427" refString="Gloer, P., &amp; Georgiev, D. (2009) New Rissooidea from Bulgaria (Gastropoda: Rissooidea). Mollusca, 27 (2), 123 - 136." type="journal article" year="2009">Glöer &amp; Georgiev, 2009</bibRefCitation>
).
</paragraph>
<paragraph id="416C001D4B35FFF662B0755EFCED6C46" blockId="4.[151,1436,1036,1535]" pageId="4" pageNumber="133">
<taxonomicName id="86D37B9E4B35FFF662B0755EFE286C0B" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[245,454,1306,1333]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
<emphasis id="73A7DC0F4B35FFF662B0755EFE286C0B" box="[245,454,1306,1333]" italics="true" pageId="4" pageNumber="133">Grossuana thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B35FFF66388755EFDE26C0B" box="[461,524,1307,1333]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
is similar by its shell morphology to
<taxonomicName id="86D37B9E4B35FFF66181755EFE4C6C68" authority="Gloer, Reuselaars &amp; Papavasileiou, 2018" authorityName="Gloer, Reuselaars &amp; Papavasileiou" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="stenaensis">
<emphasis id="73A7DC0F4B35FFF66181755EFB2A6C0A" box="[964,1220,1307,1333]" italics="true" pageId="4" pageNumber="133">Grossuana stenaensis</emphasis>
<bibRefCitation id="25427DEC4B35FFF66689755FFE4C6C68" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." pageId="4" pageNumber="133" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer, Reuselaars &amp; Papavasileiou, 2018</bibRefCitation>
</taxonomicName>
described from the nearby mainland of
<collectingCountry id="39C4408D4B35FFF6613D757EFC246C68" box="[888,970,1339,1366]" name="Greece" pageId="4" pageNumber="133">Greece</collectingCountry>
(
<bibRefCitation id="25427DEC4B35FFF6619C757EFB446C68" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." box="[985,1194,1339,1366]" pageId="4" pageNumber="133" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer et al., 2018</bibRefCitation>
) but this species has a penis with broader base and shorter and thicker tip.
</paragraph>
<paragraph id="416C001D4B35FFF662B0753AFBC36CC1" blockId="4.[151,1436,1036,1535]" pageId="4" pageNumber="133">
Previously all
<taxonomicName id="86D37B9E4B35FFF663E275C5FDC76CA4" authorityName="Radoman" authorityYear="1983" box="[423,553,1408,1434]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="genus">
<emphasis id="73A7DC0F4B35FFF663E275C5FDC76CA4" box="[423,553,1408,1434]" italics="true" pageId="4" pageNumber="133">Grossuana</emphasis>
</taxonomicName>
of Thassos were assigned to
<taxonomicName id="86D37B9E4B35FFF661DD75C5FF316C82" authority="(Kuster, 1853) (Reischutz, 1983)" authorityName="Reischutz" authorityYear="1983" baseAuthorityName="Kuster" baseAuthorityYear="1853" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="hohenackeri">
<emphasis id="73A7DC0F4B35FFF661DD75C5FBBF6CA4" box="[920,1105,1407,1434]" italics="true" pageId="4" pageNumber="133">G. hohenackeri</emphasis>
(Küster, 1853) (
<bibRefCitation id="25427DEC4B35FFF66765753AFF386C82" author="Reischutz, P." pageId="4" pageNumber="133" pagination="133 - 146" refId="ref4865" refString="Reischutz, P. (1983) Ein Beitrag zur Molluskenfauna der Insel Thasos (Griechenland). Annalen des Naturhistorischen Museums in Wien, 85 B, 133 - 146." type="journal article" year="1983">Reischütz, 1983</bibRefCitation>
)
</taxonomicName>
. It was later re-described and split into two species:
<taxonomicName id="86D37B9E4B35FFF6611B75E4FA896C82" authority="(Bourguignat, 1852)" baseAuthorityName="Bourguignat" baseAuthorityYear="1852" box="[862,1383,1441,1468]" class="Gastropoda" family="Hydrobiidae" genus="Radomaniola" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tritonum">
<emphasis id="73A7DC0F4B35FFF6611B75E4FB826C85" box="[862,1132,1441,1468]" italics="true" pageId="4" pageNumber="133">Radomaniola tritonum</emphasis>
(Bourguignat, 1852)
</taxonomicName>
and
<taxonomicName id="86D37B9E4B35FFF662D27586FCF26CE3" authority="Boeters, Gloer &amp; Falniowski (2018)" authorityName="Boeters, Gloer &amp; Falniowski" authorityYear="2018" box="[151,796,1474,1501]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tembii">
<emphasis id="73A7DC0F4B35FFF662D27586FE876CE3" box="[151,361,1474,1501]" italics="true" pageId="4" pageNumber="133">Grossuana tembii</emphasis>
<bibRefCitation id="25427DEC4B35FFF663347586FCF26CE3" author="Boeters, H. D. &amp; Gloer, P. &amp; Falniowski, A." box="[369,796,1474,1501]" pageId="4" pageNumber="133" pagination="83 - 88" refId="ref3918" refString="Boeters, H. D., Gloer, P. &amp; Falniowski, A. (2018) A new species of Grossuana (Caenogastropoda: Truncatelloidea: Hydrobiidae). Folia Malacologica, 26 (2), 83 - 88. https: // doi. org / 10.12657 / folmal. 026.010" type="journal article" year="2018">Boeters, Glöer &amp; Falniowski (2018)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="25427DEC4B35FFF661697587FBF66CE3" author="Boeters, H. D. &amp; Gloer, P. &amp; Falniowski, A." box="[812,1048,1474,1501]" pageId="4" pageNumber="133" pagination="83 - 88" refId="ref3918" refString="Boeters, H. D., Gloer, P. &amp; Falniowski, A. (2018) A new species of Grossuana (Caenogastropoda: Truncatelloidea: Hydrobiidae). Folia Malacologica, 26 (2), 83 - 88. https: // doi. org / 10.12657 / folmal. 026.010" type="journal article" year="2018">Boeters et al., 2018</bibRefCitation>
).
<taxonomicName id="86D37B9E4B35FFF6666B7586FB746CE3" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1070,1178,1474,1501]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="n. sp.">
<emphasis id="73A7DC0F4B35FFF6666B7586FB746CE3" box="[1070,1178,1474,1501]" italics="true" pageId="4" pageNumber="133">G. thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B35FFF666E67586FB0C6CE3" box="[1187,1250,1475,1501]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="86D37B9E4B35FFF667C57586FF0E6CC1" authorityName="Boeters, Gloer &amp; Falniowski" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tembii">
<emphasis id="73A7DC0F4B35FFF667C57586FF0E6CC1" italics="true" pageId="4" pageNumber="133">G. tembii</emphasis>
</taxonomicName>
by its higher whorl number (
<quantity id="862BADF84B35FFF6607375A1FD566CC1" box="[566,696,1508,1535]" metricMagnitude="-2" metricUnit="m" metricValue="8.5725" metricValueMax="8.89" metricValueMin="8.255" pageId="4" pageNumber="133" unit="in" value="3.375" valueMax="3.5" valueMin="3.25">3.25-3.5 in</quantity>
<taxonomicName id="86D37B9E4B35FFF6608575A0FCC36CC1" authorityName="Boeters, Gloer &amp; Falniowski" authorityYear="2018" box="[704,813,1508,1535]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tembii">
<emphasis id="73A7DC0F4B35FFF6608575A0FCC36CC1" box="[704,813,1508,1535]" italics="true" pageId="4" pageNumber="133">G. tembii</emphasis>
</taxonomicName>
) and longer penis tip.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B35FFF662D2766DFDC96F96" pageId="4" pageNumber="133" type="distribution">
<paragraph id="416C001D4B35FFF662D2766DFD316F7D" blockId="4.[151,735,1576,1603]" box="[151,735,1576,1603]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D2766DFED46F7C" bold="true" box="[151,314,1576,1602]" pageId="4" pageNumber="133">Distribution:</emphasis>
Known only from the
<typeStatus id="9E68BEBF4B35FFF66003766CFD966F7D" box="[582,632,1577,1603]" pageId="4" pageNumber="133">type</typeStatus>
locality.
</paragraph>
<paragraph id="416C001D4B35FFF662D2762EFDC96F96" blockId="4.[151,1417,1643,1704]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D2762EFEEF6FBB" bold="true" box="[151,257,1643,1669]" pageId="4" pageNumber="133">Habitat:</emphasis>
Karst spring in a mixed broad leaf forest (mainly
<taxonomicName id="86D37B9E4B35FFF66108762EFBC36FB8" box="[845,1069,1643,1670]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="73A7DC0F4B35FFF66108762EFBC36FB8" box="[845,1069,1643,1670]" italics="true" pageId="4" pageNumber="133">Platanus orientalis</emphasis>
</taxonomicName>
and
<taxonomicName id="86D37B9E4B35FFF666237629FAEB6FB8" box="[1126,1285,1643,1670]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B35FFF666237629FB276FB8" box="[1126,1225,1644,1670]" italics="true" pageId="4" pageNumber="133">Quercus</emphasis>
spp.
</taxonomicName>
). Found on moss, dry leaves, sand and stones.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,128 @@
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<mods:title id="C5961D883E642991812E6581BDF94BB6">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<paragraph id="416C001D4B35FFF662D276B7FCE26E33" blockId="4.[151,780,1778,1805]" box="[151,780,1778,1805]" pageId="4" pageNumber="133">
<heading id="1A24B7714B35FFF662D276B7FCE26E33" box="[151,780,1778,1805]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="133" reason="2">
<taxonomicName id="86D37B9E4B35FFF662D276B7FDD16E32" ID-CoL="P2NT" authority="Roth, 1855" authorityName="Roth" authorityYear="1855" box="[151,575,1778,1804]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="charpentieri">
<emphasis id="73A7DC0F4B35FFF662D276B7FDD16E32" bold="true" box="[151,575,1778,1804]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D276B7FE5E6E32" bold="true" box="[151,432,1778,1804]" italics="true" pageId="4" pageNumber="133">Bythinella charpentieri</emphasis>
<bibRefCitation id="25427DEC4B35FFF663F276B7FDD16E32" author="Roth, J. R." box="[439,575,1778,1804]" pageId="4" pageNumber="133" pagination="17 - 58" refId="ref4948" refString="Roth, J. R. (1855) Spicilegium molluscorum orientalium annis 1852 et 1853 collectorum. Malakozoolosche Blatter, 2 (1), 17 - 58." type="journal article" year="1855">Roth, 1855</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation id="D9E81C984B35FFF6601576B7FD4E6E33" box="[592,672,1778,1805]" captionStart="Figure 1" captionStartId="1.[151,226,1019,1043]" captionTargetBox="[393,1196,606,1000]" captionTargetId="figure-224@1.[390,1196,606,1000]" captionTargetPageId="1" captionText="Figure 1. Faksimile of the original description of Amnicola charpentieri Roth, 1855" figureDoi="http://doi.org/10.5281/zenodo.13232494" httpUri="https://zenodo.org/record/13232494/files/figure.png" pageId="4" pageNumber="133">Figs. 1</figureCitation>
,
<figureCitation id="D9E81C984B35FFF660EA76B7FD2E6E33" box="[687,704,1778,1805]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="4" pageNumber="133">2</figureCitation>
,
<figureCitation id="D9E81C984B35FFF6608976B7FCEC6E33" box="[716,770,1778,1805]" captionStart-0="Figures 6-10" captionStart-1="Figures 11-17" captionStartId-0="3.[151,236,1972,1996]" captionStartId-1="5.[151,236,1533,1557]" captionTargetBox-0="[151,1444,1565,1953]" captionTargetBox-1="[151,1429,336,1515]" captionTargetId-0="figure-136@3.[151,1444,1565,1953]" captionTargetId-1="figure-74@5.[151,1429,336,1515]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 6-10. The Grossuana spp. 6-7: G. beroni n. sp. (paratypes), 8: penis of G. beroni n. sp. 9: G. thasia n. sp (holotype), 10: penis of G. thasia n. sp." captionText-1="Figures 11-17. Bythinella charpentieri, syntypes and others from colletions ZSM and SMF. 11: lectotype of B. charpentieri Roth, 1855 from Roths collection ZSM 20012106. 12: paralectotype, Hymettus, ZSM29912124, 13: topotype, Hymettus SMF141584, 14: Kessisos ZSM20012113, 15: Attika ZSM20012106, 16: Attika, ZSM20012110, 17: Parnes, SMF141838." figureDoi-0="http://doi.org/10.5281/zenodo.13232500" figureDoi-1="http://doi.org/10.5281/zenodo.13232502" httpUri-0="https://zenodo.org/record/13232500/files/figure.png" httpUri-1="https://zenodo.org/record/13232502/files/figure.png" pageId="4" pageNumber="133">9-15</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09C953964B35FFF762D27773FE9A687A" lastPageId="5" lastPageNumber="134" pageId="4" pageNumber="133" type="materials_examined">
<paragraph id="416C001D4B35FFF662D27773FD3E6E6F" blockId="4.[151,720,1846,1873]" box="[151,720,1846,1873]" pageId="4" pageNumber="133">
<materialsCitation id="F1BB0A404B35FFF662D27773FD266E6F" box="[151,712,1846,1873]" location="Athens" municipality="Hymettos" pageId="4" pageNumber="133" specimenCount="1" stateProvince="Attiki" typeStatus="holotype">
<emphasis id="73A7DC0F4B35FFF662D27773FED76E6E" bold="true" box="[151,313,1846,1872]" pageId="4" pageNumber="133">
<typeStatus id="9E68BEBF4B35FFF662D27773FF386E6E" box="[151,214,1846,1872]" pageId="4" pageNumber="133">Type</typeStatus>
locality
</emphasis>
:
<collectingMunicipality id="A1089A674B35FFF6630D7772FE536E6F" box="[328,445,1847,1873]" pageId="4" pageNumber="133">Hymettos</collectingMunicipality>
, east of
<location id="440C56C64B35FFF6605B7773FD9D6E6F" LSID="urn:lsid:plazi:treatment:C97AB10B4B35FFF762D276B7FE9A687A:440C56C64B35FFF6605B7773FD9D6E6F" box="[542,627,1846,1873]" municipality="Hymettos" name="Athens" pageId="4" pageNumber="133" stateProvince="Attiki">Athens</location>
,
<collectingRegion id="8317CEFF4B35FFF6603A7773FD266E6F" box="[639,712,1846,1873]" country="Greece" name="Attiki" pageId="4" pageNumber="133">Attika</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="416C001D4B35FFF662D2773CFE306EE6" blockId="4.[151,1436,1913,2008]" pageId="4" pageNumber="133">
<materialsCitation id="F1BB0A404B35FFF662D2773CFB556EAA" box="[151,1211,1913,1940]" collectionCode="ZSM" county="There" location="Hymettos" municipality="Kephesos" pageId="4" pageNumber="133" specimenCount="1" stateProvince="Attiki" typeStatus="syntype">
<collectingCounty id="A80D78914B35FFF662D2773CFF356EAA" box="[151,219,1913,1940]" pageId="4" pageNumber="133">There</collectingCounty>
are four samples (
<collectingMunicipality id="A1089A674B35FFF66383773CFDD26EAA" box="[454,572,1913,1940]" pageId="4" pageNumber="133">Kephesos</collectingMunicipality>
,
<collectingRegion id="8317CEFF4B35FFF66009773CFD766EAA" box="[588,664,1913,1940]" country="Greece" name="Attiki" pageId="4" pageNumber="133">Attika</collectingRegion>
, 2x
<location id="440C56C64B35FFF66097773CFCAB6EAA" LSID="urn:lsid:plazi:treatment:C97AB10B4B35FFF762D276B7FE9A687A:440C56C64B35FFF66097773CFCAB6EAA" box="[722,837,1913,1940]" county="There" municipality="Kephesos" name="Hymettos" pageId="4" pageNumber="133" stateProvince="Attiki">Hymettos</location>
) marked as
<typeStatus id="9E68BEBF4B35FFF661A4773CFBA26EAA" box="[993,1100,1913,1940]" pageId="4" pageNumber="133" type="syntype">Syntypes</typeStatus>
in
<collectionCode id="27C298D84B35FFF6663E773CFB556EAA" box="[1147,1211,1913,1940]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="133" type="Museum">ZSM</collectionCode>
</materialsCitation>
.
<materialsCitation id="F1BB0A404B35FFF6668F773CFE346EE6" collectingDate="2106-01-20" collectionCode="ZSM" location="Thus" municipality="Only" pageId="4" pageNumber="133" specimenCount="1" stateProvince="Attiki" typeStatus="lectotype">
<collectingMunicipality id="A1089A674B35FFF6668F773CFAEA6EAA" box="[1226,1284,1913,1940]" pageId="4" pageNumber="133">Only</collectingMunicipality>
one sample contained a specimen which corresponds to the original description.
<location id="440C56C64B35FFF6619277DEFBFC6E88" LSID="urn:lsid:plazi:treatment:C97AB10B4B35FFF762D276B7FE9A687A:440C56C64B35FFF6619277DEFBFC6E88" box="[983,1042,1947,1974]" municipality="Only" name="Thus" pageId="4" pageNumber="133" stateProvince="Attiki">Thus</location>
we designate this species as the
<typeStatus id="9E68BEBF4B35FFF662D277F8FEEC6EE6" box="[151,258,1981,2008]" pageId="4" pageNumber="133" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="27C298D84B35FFF6635477F8FEA16EE6" box="[273,335,1981,2008]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="133" type="Museum">ZSM</collectionCode>
<date id="356D26DD4B35FFF6631277F8FE3C6EE6" box="[343,466,1981,2008]" pageId="4" pageNumber="133" value="2106-01-20">
<collectingDate id="2529DF354B35FFF6631277F8FE3C6EE6" box="[343,466,1981,2008]" pageId="4" pageNumber="133" value="2106-01-20">20012106</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="416C001D4B34FFF762D27081FE9A687A" blockId="5.[151,1436,195,324]" pageId="5" pageNumber="134">
<emphasis id="73A7DC0F4B34FFF762D27081FEA169E0" bold="true" box="[151,335,196,222]" pageId="5" pageNumber="134">Re-description</emphasis>
: The ovate-cylindrical shell has 4.5-5 regularly growing convex whorls with a deep suture. The aperture is ovate with a round angle at the top. The aperture height takes
<superScript id="B6A6AD554B34FFF7663B70A6FB6969CA" attach="right" box="[1150,1159,227,244]" fontSize="7" pageId="5" pageNumber="134">1</superScript>
/
<subScript id="DD5702584B34FFF766CA70AAFB76683E" attach="left" box="[1167,1176,239,256]" fontSize="7" pageId="5" pageNumber="134">3</subScript>
of shell height. The peristome is sharp, somewhat reflexed at the columella. The umbilicus is closed. The shell is
<quantity id="862BADF84B34FFF767457142FAB3681C" box="[1280,1373,263,290]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="134" unit="mm" value="3.3">3.3 mm</quantity>
high and
<quantity id="862BADF84B34FFF7628C716CFECD687D" box="[201,291,297,324]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="134" unit="mm" value="1.8">1.8 mm</quantity>
broad.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,227 @@
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<mods:title id="12781B7C9769986BE57AEB097AD67AF9">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<mods:namePart id="B5E422FF44DC696420B9F63ADA4672D8">Georgiev, Dilian</mods:namePart>
<mods:affiliation id="8F9C5D7D00C65F138181EB3B7560A929">Department of Ecology and Nature Conservation, Faculty of Biology, Paisii Hilendarski University of Plovdiv, 24 Tsar Assen Str., 4000 Plovdiv, Bulgaria</mods:affiliation>
</mods:name>
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<mods:namePart id="46E455BB4137120947A227168A891B55">Glöer, Peter</mods:namePart>
<mods:affiliation id="9A3D51FBEDEE21A5B1384915F03A2EF4">Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany Corresponding author. E-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<treatment id="C97AB10B4B37FFF562D27527FE956DDC" LSID="urn:lsid:zoobank.org:act:DD058BE9-160A-4CA0-8ECB-7F5E642420DC" httpUri="http://treatment.plazi.org/id/C97AB10B4B37FFF562D27527FE956DDC" lastPageId="7" lastPageNumber="136" pageId="6" pageNumber="135">
<subSubSection id="09C953964B37FFF462D27527FB6B6CA5" pageId="6" pageNumber="135" type="nomenclature">
<paragraph id="416C001D4B37FFF462D27527FD706C43" blockId="6.[151,1157,1378,1435]" box="[151,670,1378,1405]" pageId="6" pageNumber="135">
<heading id="1A24B7714B37FFF462D27527FD706C43" box="[151,670,1378,1405]" fontSize="11" level="2" pageId="6" pageNumber="135" reason="2">
<emphasis id="73A7DC0F4B37FFF462D27527FE0E6C42" bold="true" box="[151,480,1378,1404]" pageId="6" pageNumber="135">
<taxonomicName id="86D37B9E4B37FFF462D27527FE786C42" ID-CoL="7QHR8" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[151,406,1378,1404]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="rachonica" status="sp. nov.">
<emphasis id="73A7DC0F4B37FFF462D27527FE786C42" bold="true" box="[151,406,1378,1404]" italics="true" pageId="6" pageNumber="135">Bythinella rachonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B37FFF463DB7527FE0E6C42" box="[414,480,1378,1404]" pageId="6" pageNumber="135" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D9E81C984B37FFF463B47527FDAD6C43" box="[497,579,1378,1405]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="6" pageNumber="135">Figs. 2</figureCitation>
,
<figureCitation id="D9E81C984B37FFF460157527FD7B6C43" box="[592,661,1378,1405]" captionStart="Figures 22-25" captionStartId="7.[151,236,891,915]" captionTargetBox="[151,1446,270,872]" captionTargetId="figure-42@7.[151,1446,269,873]" captionTargetPageId="7" captionText="Figures 22-25. Bythinella rachonica n. sp., 22: shell, 23: penis in situ, 24: penis with penial appendix, 25: tubular gland." figureDoi="http://doi.org/10.5281/zenodo.13232506" httpUri="https://zenodo.org/record/13232506/files/figure.png" pageId="6" pageNumber="135">22-25</figureCitation>
)
</heading>
</paragraph>
<paragraph id="416C001D4B37FFF462D275C6FB6B6CA5" blockId="6.[151,1157,1378,1435]" box="[151,1157,1411,1435]" pageId="6" pageNumber="135">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="09C953964B37FFF462D27580FCF76FB6" pageId="6" pageNumber="135" type="materials_examined">
<paragraph id="416C001D4B37FFF462D27580FE956CE1" blockId="6.[151,1222,1477,1570]" box="[151,379,1477,1503]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D27580FE956CE1" bold="true" box="[151,379,1477,1503]" pageId="6" pageNumber="135">Materials studied:</emphasis>
</paragraph>
<paragraph id="416C001D4B37FFF462B075A3FB286F3F" blockId="6.[151,1222,1477,1570]" box="[245,1222,1510,1537]" pageId="6" pageNumber="135">
<materialsCitation id="F1BB0A404B37FFF462B075A3FB286F3F" box="[245,1222,1510,1537]" collectionCode="ZMH" pageId="6" pageNumber="135" specimenCode="ZMH 140795" specimenCount="1" typeStatus="holotype">
<emphasis id="73A7DC0F4B37FFF462B075A3FE9F6F3E" bold="true" box="[245,369,1510,1536]" pageId="6" pageNumber="135">
<typeStatus id="9E68BEBF4B37FFF462B075A3FE826F3E" box="[245,364,1510,1536]" pageId="6" pageNumber="135" type="holotype">Holotype</typeStatus>
:
</emphasis>
shell height
<quantity id="862BADF84B37FFF4604D75A3FD8A6F3F" box="[520,612,1510,1537]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="135" unit="mm" value="3.1">3.1 mm</quantity>
, width
<quantity id="862BADF84B37FFF460FF75A3FCCB6F3F" box="[698,805,1510,1537]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="135" unit="mm" value="1.75">1.75 mm</quantity>
, from type locality (
<specimenCode id="1175A8664B37FFF4665775A3FB576F3F" box="[1042,1209,1510,1537]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="135" type="Museum">ZMH 140795</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="416C001D4B37FFF462B0764DFB666F1C" blockId="6.[151,1222,1477,1570]" box="[245,1160,1543,1570]" pageId="6" pageNumber="135">
<materialsCitation id="F1BB0A404B37FFF462B0764DFCA06F1C" box="[245,846,1543,1570]" collectionCode="ZMH" pageId="6" pageNumber="135" specimenCode="ZMH 140796" specimenCount="3" typeStatus="paratype">
<emphasis id="73A7DC0F4B37FFF462B0764DFE926F1C" bold="true" box="[245,380,1544,1570]" pageId="6" pageNumber="135">
<typeStatus id="9E68BEBF4B37FFF462B0764DFE996F1C" box="[245,375,1544,1570]" pageId="6" pageNumber="135" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="57D5CB944B37FFF463C17642FDFD6F1C" box="[388,531,1543,1570]" count="3" pageId="6" pageNumber="135" type="generic">3 specimens</specimenCount>
in ethanol (
<specimenCode id="1175A8664B37FFF460DA7642FCA86F1C" box="[671,838,1543,1570]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="135" type="Museum">ZMH 140796</specimenCode>
)
</materialsCitation>
,
<specimenCount id="57D5CB944B37FFF4611F7642FC336F1C" box="[858,989,1543,1570]" count="1" pageId="6" pageNumber="135" type="generic">1 specimen</specimenCount>
in coll. Glöer.
</paragraph>
<paragraph id="416C001D4B37FFF462D2760EFCF76FB6" blockId="6.[151,1435,1611,1672]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D2760EFEA86F5B" bold="true" box="[151,326,1611,1637]" pageId="6" pageNumber="135">Type locality:</emphasis>
<materialsCitation id="F1BB0A404B37FFF46317760EFCFB6FB6" country="Greece" elevation="107" latitude="40.75822" location="Rachoni village" longLatPrecision="1" longitude="24.638138" municipality="Thassos Island" pageId="6" pageNumber="135" specimenCount="1" stateProvince="Voreio Aigaio">
<collectingCountry id="39C4408D4B37FFF46317760EFE466F58" box="[338,424,1611,1638]" name="Greece" pageId="6" pageNumber="135">Greece</collectingCountry>
,
<collectingMunicipality id="A1089A674B37FFF463FD760EFD826F58" box="[440,620,1611,1638]" pageId="6" pageNumber="135">Thassos Island</collectingMunicipality>
,
<collectingRegion id="8317CEFF4B37FFF46039760EFCC66F58" box="[636,808,1611,1638]" country="Greece" name="Voreio Aigaio" pageId="6" pageNumber="135">North Aegean</collectingRegion>
, a big spring in the yard of the church of
<location id="440C56C64B37FFF4677C760EFF076FB6" LSID="urn:lsid:plazi:treatment:C97AB10B4B37FFF562D27527FE956DDC:440C56C64B37FFF4677C760EFF076FB6" country="Greece" latitude="40.75822" longLatPrecision="1" longitude="24.638138" municipality="Thassos Island" name="Rachoni village" pageId="6" pageNumber="135" stateProvince="Voreio Aigaio">Rachoni village</location>
,
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<geoCoordinate id="24E766DA4B37FFF463CA7628FDCE6FB6" box="[399,544,1645,1672]" degrees="24" direction="east" minutes="38" orientation="longitude" pageId="6" pageNumber="135" precision="1" seconds="17.3" value="24.638138">E24 38 17.3</geoCoordinate>
,
<elevation id="CAFEE72E4B37FFF460697628FD5D6FB6" box="[556,691,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="135" unit="m" value="107.0">
<quantity id="862BADF84B37FFF460697628FD986FB6" box="[556,630,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="135" unit="m" value="107.0">107 m</quantity>
a.s.l.
</elevation>
(
<figureCitation id="D9E81C984B37FFF460867628FCE26FB6" box="[707,780,1645,1672]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="6" pageNumber="135">Fig. 4</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B37FFF462D276F4FBD66FF5" box="[151,1080,1712,1739]" pageId="6" pageNumber="135" type="etymology">
<paragraph id="416C001D4B37FFF462D276F4FBD66FF5" blockId="6.[151,1080,1712,1739]" box="[151,1080,1712,1739]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D276F4FEC66FF5" bold="true" box="[151,296,1713,1739]" pageId="6" pageNumber="135">Etymology:</emphasis>
Named after village of Rachoni where the new species was found.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B37FFF562D276B1FC15683E" lastPageId="7" lastPageNumber="136" pageId="6" pageNumber="135" type="description">
<paragraph id="416C001D4B37FFF462D276B1FCC26E4A" blockId="6.[151,1436,1780,1908]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D276B1FEDC6E30" bold="true" box="[151,306,1780,1806]" pageId="6" pageNumber="135">Description:</emphasis>
Shell ivory, cylindrical with 4.5 whorls, separated by a deep suture. Surface silky and finely striated. Apex obtuse, umbilicus closed. Aperture oval, with a sharp peristome, angled at the top. Shell height
<quantity id="862BADF84B37FFF462D27772FECD6E6C" box="[151,291,1847,1874]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="6" pageNumber="135" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9-3.1 mm</quantity>
, width
<quantity id="862BADF84B37FFF4633F7772FE376E6C" box="[378,473,1847,1874]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="135" unit="mm" value="1.8">1.8 mm</quantity>
. Penial appendix twice longer than the penis, tubular gland medium-sized, with 5 half loops, tapering from distal to proximal part (
<figureCitation id="D9E81C984B37FFF46090771CFCF36E4A" box="[725,797,1881,1908]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="6" pageNumber="135">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="416C001D4B37FFF462D277D8FA756EC5" blockId="6.[151,1435,1948,2043]" pageId="6" pageNumber="135">
<emphasis id="73A7DC0F4B37FFF462D277D8FE296E89" bold="true" box="[151,455,1949,1975]" pageId="6" pageNumber="135">Differentiating features:</emphasis>
From this region no
<taxonomicName id="86D37B9E4B37FFF4608D77D9FC966E89" box="[712,888,1948,1975]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B37FFF4608D77D9FCAE6E89" box="[712,832,1948,1975]" italics="true" pageId="6" pageNumber="135">Bythinella</emphasis>
spp.
</taxonomicName>
are known (
<bibRefCitation id="25427DEC4B37FFF4664F77D9FAA06E89" author="Gloer, P. &amp; Hirschfelder, H. - J." box="[1034,1358,1948,1975]" pageId="6" pageNumber="135" pagination="60 - 67" refId="ref4521" refString="Gloer, P. &amp; Hirschfelder, H. - J. (2020) Some new Bythinella spp. from southern Greece (Gastropoda: Bythinellidae). Ecologica Montenegrina, 30, 60 - 67. https: // doi. org / 10.37828 / em. 2020.30.5" type="journal article" year="2020">Glöer &amp; Hirschfelder 2020</bibRefCitation>
,
<bibRefCitation id="25427DEC4B37FFF4671C77D9FE776EE7" author="Gloer, P. &amp; Reuselaars, R." pageId="6" pageNumber="135" pagination="20 - 33" refId="ref4571" refString="Gloer, P. &amp; Reuselaars, R. (2020 a) The Bythinella spp. of Greece (Gastropoda: Hydrobioidea: Bythinelliidae). Ecologica Montenegrina, 29, 20 - 33. https: // doi. org / 10.37828 / em. 2020.29.3" type="journal article" year="2020">Glöer &amp; Reuselaars 2020a</bibRefCitation>
, b, c), except from the neighbouring Samothrake Island, from where Reischütz described
<taxonomicName id="86D37B9E4B37FFF4635577A5FC6F6EC5" authority="P. L. Reischutz 1988" authorityName="P. L. Reischutz" authorityYear="1988" box="[272,897,2016,2043]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="135" phylum="Mollusca" rank="subSpecies" species="charpentieri" subSpecies="cabirius">
<emphasis id="73A7DC0F4B37FFF4635577A5FD676EC5" box="[272,649,2016,2043]" italics="true" pageId="6" pageNumber="135">Bythinella charpentieri cabirius</emphasis>
P.L. Reischütz 1988
</taxonomicName>
(considered as a separate species by
<bibRefCitation id="25427DEC4B37FFF4677F77A5FA756EC5" author="Gloer, P. &amp; Georgiev, D." box="[1338,1435,2016,2043]" pageId="6" pageNumber="135" pagination="278 - 282" refId="ref4714" refString="Gloer, P. &amp; Georgiev, D. (2012) Three new gastropod species from Greece and Turkey (Mollusca: Gastropoda: Rissooidea) with notes on the anatomy of Bythinella charpentieri cabirius Reischutz 1988. North-Western Journal of Zoology, 8 (2), 278 - 282." type="journal article" year="2012">Glöer &amp;</bibRefCitation>
</paragraph>
<paragraph id="416C001D4B36FFF562D27086FC15683E" blockId="7.[151,1435,195,256]" pageId="7" pageNumber="136">
Georgiev 2012), which is a small and slim species (
<quantity id="862BADF84B36FFF561667086FC5A69E0" box="[803,948,195,222]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="7" pageNumber="136" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5-2.7 mm</quantity>
high and
<quantity id="862BADF84B36FFF5667D7086FB2869E0" box="[1080,1222,195,222]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="7" pageNumber="136" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2-1.3 mm</quantity>
broad) (
<bibRefCitation id="25427DEC4B36FFF567717086FEA6683E" author="Gloer, P. &amp; Georgiev, D." pageId="7" pageNumber="136" pagination="278 - 282" refId="ref4714" refString="Gloer, P. &amp; Georgiev, D. (2012) Three new gastropod species from Greece and Turkey (Mollusca: Gastropoda: Rissooidea) with notes on the anatomy of Bythinella charpentieri cabirius Reischutz 1988. North-Western Journal of Zoology, 8 (2), 278 - 282." type="journal article" year="2012">Glöer &amp; Georgiev 2012</bibRefCitation>
) and in this way different from
<taxonomicName id="86D37B9E4B36FFF560FB70A0FC5B683E" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[702,949,229,256]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="136" phylum="Mollusca" rank="species" species="rachonica" status="sp. nov.">
<emphasis id="73A7DC0F4B36FFF560FB70A0FC5B683E" box="[702,949,229,256]" italics="true" pageId="7" pageNumber="136">Bythinella rachonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B36FFF561F870A3FC15683E" box="[957,1019,230,256]" pageId="7" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="15AC50954B36FFF562D2733EFF376A8F" ID-DOI="http://doi.org/10.5281/zenodo.13232506" ID-Zenodo-Dep="13232506" httpUri="https://zenodo.org/record/13232506/files/figure.png" pageId="7" pageNumber="136" startId="7.[151,236,891,915]" targetBox="[151,1446,270,872]" targetPageId="7" targetType="figure">
<paragraph id="416C001D4B36FFF562D2733EFF376A8F" blockId="7.[151,1436,891,945]" pageId="7" pageNumber="136">
<emphasis id="73A7DC0F4B36FFF562D2733EFED26AAD" bold="true" box="[151,316,891,915]" pageId="7" pageNumber="136">Figures 22-25.</emphasis>
<taxonomicName id="86D37B9E4B36FFF56302733EFDC56AAD" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[327,555,891,915]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="136" phylum="Mollusca" rank="species" species="rachonica" status="sp. nov.">
<emphasis id="73A7DC0F4B36FFF56302733EFDC56AAD" box="[327,555,891,915]" italics="true" pageId="7" pageNumber="136">Bythinella rachonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B36FFF560737339FD9B6AAD" box="[566,629,892,915]" pageId="7" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
,
<emphasis id="73A7DC0F4B36FFF560C1733EFD716AAD" bold="true" box="[644,671,891,915]" pageId="7" pageNumber="136">22</emphasis>
: shell,
<emphasis id="73A7DC0F4B36FFF560B0733EFCFE6AAD" bold="true" box="[757,784,891,915]" pageId="7" pageNumber="136">23</emphasis>
: penis in situ,
<emphasis id="73A7DC0F4B36FFF561FE733EFC386AAD" bold="true" box="[955,982,891,915]" pageId="7" pageNumber="136">24</emphasis>
: penis with penial appendix,
<emphasis id="73A7DC0F4B36FFF56761733EFAD16AAD" bold="true" box="[1316,1343,891,915]" pageId="7" pageNumber="136">25</emphasis>
: tubular gland.
</paragraph>
</caption>
<subSubSection id="09C953964B36FFF562D273B8FE956DDC" pageId="7" pageNumber="136" type="distribution">
<paragraph id="416C001D4B36FFF562D273B8FD6A6D43" blockId="7.[151,1435,1020,1149]" pageId="7" pageNumber="136">
<emphasis id="73A7DC0F4B36FFF562D273B8FED46D29" bold="true" box="[151,314,1021,1047]" pageId="7" pageNumber="136">Distribution:</emphasis>
Known only from the
<typeStatus id="9E68BEBF4B36FFF5601573B8FD6C6D29" box="[592,642,1021,1047]" pageId="7" pageNumber="136">type</typeStatus>
locality. The spring at the Agora of Limenas was checked but no living snails were found. Some empty, eroded shells of a
<taxonomicName id="86D37B9E4B36FFF5610A745BFBC86D07" box="[847,1062,1054,1081]" pageId="7" pageNumber="136">
<emphasis id="73A7DC0F4B36FFF5610A745BFC296D07" box="[847,967,1054,1081]" italics="true" pageId="7" pageNumber="136">Bythinella</emphasis>
species
</taxonomicName>
were found by D. Georgiev in sand deposits in a spring below Panagia village (
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<geoCoordinate id="24E766DA4B36FFF561C57405FBF86D65" box="[896,1046,1088,1115]" degrees="24" direction="east" minutes="44" orientation="longitude" pageId="7" pageNumber="136" precision="1" seconds="26.6" value="24.740723">E24 44 26.6</geoCoordinate>
,
<quantity id="862BADF84B36FFF566667405FB8E6D65" box="[1059,1120,1088,1115]" metricMagnitude="1" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="136" unit="m" value="49.0">49 m</quantity>
a.s.l.) but it is not clear if they belong to
<taxonomicName id="86D37B9E4B36FFF563037427FDD36D43" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[326,573,1122,1149]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="136" phylum="Mollusca" rank="species" species="rachonica" status="sp. nov.">
<emphasis id="73A7DC0F4B36FFF563037427FDD36D43" box="[326,573,1122,1149]" italics="true" pageId="7" pageNumber="136">Bythinella rachonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B36FFF560007426FD6A6D43" box="[581,644,1123,1149]" pageId="7" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="416C001D4B36FFF562D274E0FE956DDC" blockId="7.[151,1435,1189,1250]" pageId="7" pageNumber="136">
<emphasis id="73A7DC0F4B36FFF562D274E0FEEF6D81" bold="true" box="[151,257,1189,1215]" pageId="7" pageNumber="136">Habitat:</emphasis>
Found in a spring (captured under the church of Rachoni, with a concrete bed) on stones and submerged mosses.
</paragraph>
</subSubSection>
</treatment>
</document>