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<document id="AECB24CEB7831F369CAC30CB8B5D6AF6" ID-DOI="10.37828/em.2020.34.1" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232071" ID-ZooBank="urn:lsid:zoobank.org:pub:D9ED9E59-18C6-4DD0-BC0E-EB3D91C259EC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722913617162" checkinUser="felipe" docAuthor="Yakovlev, Roman V." docDate="2020" docId="877AAD7B7F7FFF8F1481FDDFFBD4F832" docLanguage="en" docName="EcolMontenegrina.34.1-7.pdf" docOrigin="Ecologica Montenegrina 34" docSource="http://dx.doi.org/10.37828/em.2020.34.1" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Wittoecia Yakovlev, 2020, gen. n." docType="treatment" docUuid="C6EC2069-700E-44AF-8DE1-0CD9583CE1E1" docUuidSource="ZooBank" docVersion="2" lastPageNumber="2" masterDocId="7B43D5037F7EFF8E1416FFC9FFC7FFEA" masterDocTitle="Wittoecia - a new genus of Zeuzerinae (Lepidoptera, Cossidae) from the Federal Democratic Republic of Ethiopia" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1722916095877" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="94EBD551ADB5CFADBDC229B6AB133E12">Wittoecia - a new genus of Zeuzerinae (Lepidoptera, Cossidae) from the Federal Democratic Republic of Ethiopia</mods:title>
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<mods:namePart id="A70BDE885ED728E8FD0114314126D065">Yakovlev, Roman V.</mods:namePart>
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<mods:title id="4007E2A6809D8ADD50754652EAB99134">Ecologica Montenegrina</mods:title>
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<mods:date id="C139037CD260A0365C9E42E95433D944">2020</mods:date>
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<treatment id="877AAD7B7F7FFF8F1481FDDFFBD4F832" LSID="urn:lsid:zoobank.org:act:C6EC2069-700E-44AF-8DE1-0CD9583CE1E1" httpUri="http://treatment.plazi.org/id/877AAD7B7F7FFF8F1481FDDFFBD4F832" lastPageNumber="2" pageId="1" pageNumber="2">
<subSubSection id="47C94FE67F7FFF8F1481FDDFFBBCFDA4" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="0F6C1C6D7F7FFF8F1481FDDFFEA4FDDA" blockId="1.[151,1177,534,624]" box="[151,355,534,560]" pageId="1" pageNumber="2">
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<emphasis id="3DA7C07F7F7FFF8F1481FDDFFEA4FDDA" bold="true" box="[151,355,534,560]" pageId="1" pageNumber="2">
<taxonomicName id="C8D367EE7F7FFF8F1481FDDFFECFFDDA" authorityName="Yakovlev" authorityYear="2020" box="[151,264,534,560]" class="Insecta" family="Cossidae" genus="Wittoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3DA7C07F7F7FFF8F1481FDDFFECFFDDA" bold="true" box="[151,264,534,560]" italics="true" pageId="1" pageNumber="2">Wittoecia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26947D047F7FFF8F1506FDDFFEA4FDDA" box="[272,355,534,560]" pageId="1" pageNumber="2" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</subSubSection>
<subSubSection id="47C94FE67F7FFF8F1481FD9FFB5EFD9A" box="[151,1177,597,624]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="0F6C1C6D7F7FFF8F1481FD9FFB5EFD9A" blockId="1.[151,1177,534,624]" box="[151,1177,597,624]" pageId="1" pageNumber="2">
<typeStatus id="D068A2CF7F7FFF8F1481FD9FFF14FD9A" box="[151,211,598,624]" pageId="1" pageNumber="2">Type</typeStatus>
species (by monotypy)
<taxonomicName id="C8D367EE7F7FFF8F15FBFD9CFC3FFD9A" authority="Yakovlev &amp; Witt, 2016" authorityName="Yakovlev &amp; Witt" authorityYear="2016" box="[493,1016,597,624]" class="Insecta" family="Cossidae" genus="Azygophleps" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="brehmi">
<emphasis id="3DA7C07F7F7FFF8F15FBFD9CFD1BFD9A" box="[493,732,597,624]" italics="true" pageId="1" pageNumber="2">Azygophleps brehmi</emphasis>
<bibRefCitation id="6B42619C7F7FFF8F16F2FD9CFC3FFD9A" author="Yakovlev, R. V. &amp; Witt, Th." box="[740,1016,597,624]" pageId="1" pageNumber="2" pagination="71 - 73" refId="ref3283" refString="Yakovlev, R. V. &amp; Witt, Th. (2016). Azygophleps brehmi Yakovlev et Witt, sp. n. - a new Carpenter-Moth (Lepidoptera, Cossidae) from Ethiopia. Russian Entomological Journal, 25 (1), 71 - 73." type="journal article" year="2016">Yakovlev &amp; Witt, 2016</bibRefCitation>
</taxonomicName>
(
<figureCitation id="97E800E87F7FFF8F101EFD9CFBB5FD9A" box="[1032,1138,597,624]" captionStart="Figures 1−7" captionStartId="2.[151,237,1694,1718]" captionTargetBox="[159,1434,234,1623]" captionTargetId="figure-16@2.[151,1436,189,1644]" captionTargetPageId="2" captionText="Figures 1−7. Adult males of Zeuzerinae: 1. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., holotype (Museum Witt, Munich); 2. W. brehmi, paratype (Museum Witt, Munich); 3. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Museum Witt, Munich); 4. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (individual number: ANHRTUK: 00052313, African Natural History Research Trust, Leominster, GB); 5. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Museum Witt, Munich); 6. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Museum Witt, Munich); 7. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232073" httpUri="https://zenodo.org/record/13232073/files/figure.png" pageId="1" pageNumber="2">Figs 1−2</figureCitation>
,
<figureCitation id="97E800E87F7FFF8F1096FD9CFB48FD9A" box="[1152,1167,597,624]" captionStart="Figures 8−13" captionStartId="3.[151,236,1710,1734]" captionTargetBox="[235,1308,194,1696]" captionTargetId="figure-12@3.[228,1358,189,1703]" captionTargetPageId="3" captionText="Figures 8−13. Genitalia of males of Zeuzerinae: 8. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., paratype (Genitalpräparat Heterocera N 33.185, Museum Witt, Munich); 9. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Genitalpräparat Heterocera N 32.924, Museum Witt, Munich); 10. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (ANHRT Gen Slide 00075, African Natural History Research Trust, Leominster, GB); 11. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Genitalpräparat Heterocera N 33.161, Museum Witt, Munich); 12. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Genitalpräparat Heterocera N 32.709, Museum Witt, Munich); 13. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Genitalpräparat Heterocera N 33.167, Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232075" httpUri="https://zenodo.org/record/13232075/files/figure.png" pageId="1" pageNumber="2">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="47C94FE67F7FFF8F1481FD50FE00FBA3" pageId="1" pageNumber="2" type="description">
<paragraph id="0F6C1C6D7F7FFF8F1481FD50FA8DFC94" blockId="1.[151,1436,665,1097]" pageId="1" pageNumber="2">
<emphasis id="3DA7C07F7F7FFF8F1481FD50FEEFFD59" bold="true" box="[151,296,665,691]" pageId="1" pageNumber="2">Description</emphasis>
. Size medium (fore wing length
<quantity id="C82BB1887F7FFF8F16ACFD50FC85FD5E" box="[698,834,665,692]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="1" pageNumber="2" unit="mm" value="23.0" valueMax="24.0" valueMin="22.0">22−24 mm</quantity>
). Antenna goblet- like, bipectinate, very long crest processes in two proximal thirds of antenna (processes 4 times longer than rod diameter), very short crest processes in distal third (processes equal to antenna rod diameter). Thorax and abdomen covered with lightyellow hairs. Fore wing elongated, creamy, with reticulated pattern of black strokes: more dense and wide along costa, anal edge and in distal cell, significantly thinner submarginally. No pattern of strokes in cubital area. Hind wing creamy, with poorly expressed reticulated pattern of thin grey undulated strokes in submarginal and marginal areas (except for anal area). Fringe on all wings light-creamy, almost white.
</paragraph>
<paragraph id="0F6C1C6D7F7FFF8F14E3FC4CFAE4FBCD" blockId="1.[151,1436,665,1097]" pageId="1" pageNumber="2">Male genitalia (GenPrMWM 25284, 33185). Uncus short, triangle, apically sharp; tegumen wide; gnathos arms short, not fused, basally thick, distally narrowing gradually; valve narrow, long, with smooth parallel edges, apically rounded; juxta large, oval, with small semicircular cut on dorsal surface; saccus long, massive, tapered, apically sharp; aedeagus thin, 1/4 shorter than valve, basally thick, slightly narrowing to distal end, slightly curved throughout its length, small rod-like cornutus in lateral surface of vesica.</paragraph>
<paragraph id="0F6C1C6D7F7FFF8F14E3FBE7FE00FBA3" blockId="1.[151,1436,665,1097]" box="[245,455,1070,1097]" pageId="1" pageNumber="2">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="47C94FE67F7FFF8F1481FBBBFE56F96F" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="0F6C1C6D7F7FFF8F1481FBBBFED9F9CB" blockId="1.[151,1436,1137,1569]" pageId="1" pageNumber="2">
<emphasis id="3DA7C07F7F7FFF8F1481FBBBFED7FB66" bold="true" box="[151,272,1138,1164]" pageId="1" pageNumber="2">Diagnosis</emphasis>
. Externally, the new genus is most close to the genus
<taxonomicName id="C8D367EE7F7FFF8F179DFBB8FBE7FB66" authorityName="Hampson" authorityYear="1892" box="[907,1056,1137,1164]" class="Insecta" family="Cossidae" genus="Azygophleps" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3DA7C07F7F7FFF8F179DFBB8FBE7FB66" box="[907,1056,1137,1164]" italics="true" pageId="1" pageNumber="2">Azygophleps</emphasis>
</taxonomicName>
(
<figureCitation id="97E800E87F7FFF8F1039FBB8FB5BFB66" box="[1071,1180,1137,1164]" captionStart="Figures 1−7" captionStartId="2.[151,237,1694,1718]" captionTargetBox="[159,1434,234,1623]" captionTargetId="figure-16@2.[151,1436,189,1644]" captionTargetPageId="2" captionText="Figures 1−7. Adult males of Zeuzerinae: 1. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., holotype (Museum Witt, Munich); 2. W. brehmi, paratype (Museum Witt, Munich); 3. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Museum Witt, Munich); 4. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (individual number: ANHRTUK: 00052313, African Natural History Research Trust, Leominster, GB); 5. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Museum Witt, Munich); 6. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Museum Witt, Munich); 7. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232073" httpUri="https://zenodo.org/record/13232073/files/figure.png" pageId="1" pageNumber="2">Figs 3−4</figureCitation>
,
<figureCitation id="97E800E87F7FFF8F10BEFBB8FB2FFB66" box="[1192,1256,1137,1164]" captionStart="Figures 8−13" captionStartId="3.[151,236,1710,1734]" captionTargetBox="[235,1308,194,1696]" captionTargetId="figure-12@3.[228,1358,189,1703]" captionTargetPageId="3" captionText="Figures 8−13. Genitalia of males of Zeuzerinae: 8. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., paratype (Genitalpräparat Heterocera N 33.185, Museum Witt, Munich); 9. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Genitalpräparat Heterocera N 32.924, Museum Witt, Munich); 10. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (ANHRT Gen Slide 00075, African Natural History Research Trust, Leominster, GB); 11. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Genitalpräparat Heterocera N 33.161, Museum Witt, Munich); 12. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Genitalpräparat Heterocera N 32.709, Museum Witt, Munich); 13. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Genitalpräparat Heterocera N 33.167, Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232075" httpUri="https://zenodo.org/record/13232075/files/figure.png" pageId="1" pageNumber="2">9−10</figureCitation>
), from which it clearly differs in the male genitalia (in
<emphasis id="3DA7C07F7F7FFF8F1673FB5AFCC6FB44" box="[613,769,1171,1198]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="C8D367EE7F7FFF8F1673FB5AFD3AFB44" authorityName="Hampson" authorityYear="1892" box="[613,765,1171,1198]" class="Insecta" family="Cossidae" genus="Azygophleps" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Azygophleps</taxonomicName>
,
</emphasis>
the aedeagus is thick with a large cornutus in the lateral surface of the vesica, the gnathos arms are long and thin, not fused). From the genus
<taxonomicName id="C8D367EE7F7FFF8F10F0FB7DFEA2FB1B" authority="Schawerda, 1924" authorityName="Schawerda" authorityYear="1924" class="Insecta" family="Cossidae" genus="Phragmacossia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3DA7C07F7F7FFF8F10F0FB7DFA5BFB25" box="[1254,1436,1204,1231]" italics="true" pageId="1" pageNumber="2">Phragmacossia</emphasis>
Schawerda, 1924
</taxonomicName>
(
<typeStatus id="D068A2CF7F7FFF8F156FFB1FFE6AFB1B" box="[377,429,1238,1265]" pageId="1" pageNumber="2">type</typeStatus>
species, by original designation −
<taxonomicName id="C8D367EE7F7FFF8F1749FB1FFA53FB1B" authority="Pungeler, 1900" authorityName="Pungeler" authorityYear="1900" box="[863,1428,1238,1265]" class="Insecta" family="Cossidae" genus="Phragmatoecia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="3DA7C07F7F7FFF8F1749FB1FFBD3FB1B" box="[863,1044,1238,1265]" italics="true" pageId="1" pageNumber="2">Phragmatoecia</emphasis>
(sic)
<emphasis id="3DA7C07F7F7FFF8F1049FB1FFB15FB1B" box="[1119,1234,1238,1265]" italics="true" pageId="1" pageNumber="2">reticulata</emphasis>
Püngeler, 1900
</taxonomicName>
) (
<figureCitation id="97E800E87F7FFF8F1489FB31FF37FAF9" box="[159,240,1272,1299]" captionStart="Figures 1−7" captionStartId="2.[151,237,1694,1718]" captionTargetBox="[159,1434,234,1623]" captionTargetId="figure-16@2.[151,1436,189,1644]" captionTargetPageId="2" captionText="Figures 1−7. Adult males of Zeuzerinae: 1. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., holotype (Museum Witt, Munich); 2. W. brehmi, paratype (Museum Witt, Munich); 3. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Museum Witt, Munich); 4. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (individual number: ANHRTUK: 00052313, African Natural History Research Trust, Leominster, GB); 5. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Museum Witt, Munich); 6. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Museum Witt, Munich); 7. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232073" httpUri="https://zenodo.org/record/13232073/files/figure.png" pageId="1" pageNumber="2">Figs 5</figureCitation>
,
<figureCitation id="97E800E87F7FFF8F1514FB31FEE7FAF9" box="[258,288,1272,1299]" captionStart="Figures 8−13" captionStartId="3.[151,236,1710,1734]" captionTargetBox="[235,1308,194,1696]" captionTargetId="figure-12@3.[228,1358,189,1703]" captionTargetPageId="3" captionText="Figures 8−13. Genitalia of males of Zeuzerinae: 8. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., paratype (Genitalpräparat Heterocera N 33.185, Museum Witt, Munich); 9. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Genitalpräparat Heterocera N 32.924, Museum Witt, Munich); 10. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (ANHRT Gen Slide 00075, African Natural History Research Trust, Leominster, GB); 11. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Genitalpräparat Heterocera N 33.161, Museum Witt, Munich); 12. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Genitalpräparat Heterocera N 32.709, Museum Witt, Munich); 13. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Genitalpräparat Heterocera N 33.167, Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232075" httpUri="https://zenodo.org/record/13232075/files/figure.png" pageId="1" pageNumber="2">11</figureCitation>
) distributed in the south of Palaearctic and south-east Asia the new genus differs in a more variable pattern on the fore wing, the massive uncus and the short gnathos arms. In the male genital structure, it is most close to the genus
<taxonomicName id="C8D367EE7F7FFF8F15FDFAF2FC99FABC" authority="Newman, 1850" authorityName="Newman" authorityYear="1850" box="[491,862,1339,1366]" class="Insecta" family="Cossidae" genus="Phragmataecia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3DA7C07F7F7FFF8F15FDFAF2FD67FABC" box="[491,672,1339,1366]" italics="true" pageId="1" pageNumber="2">Phragmataecia</emphasis>
Newman, 1850
</taxonomicName>
(
<typeStatus id="D068A2CF7F7FFF8F1778FAF2FC65FABC" box="[878,930,1339,1366]" pageId="1" pageNumber="2">type</typeStatus>
species, by monotypy −
<taxonomicName id="C8D367EE7F7FFF8F10D9FAF4FEFBFA92" authority="Hubner, 1808" authorityName="Hubner" authorityYear="1808" class="Insecta" family="Noctuidae" genus="Noctua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="arundinis">
<emphasis id="3DA7C07F7F7FFF8F10D9FAF4FA5BFABC" box="[1231,1436,1339,1366]" italics="true" pageId="1" pageNumber="2">Noctua arundinis</emphasis>
Hübner, 1808
</taxonomicName>
) (
<figureCitation id="97E800E87F7FFF8F1541FA94FE61FA92" box="[343,422,1373,1400]" captionStart="Figures 1−7" captionStartId="2.[151,237,1694,1718]" captionTargetBox="[159,1434,234,1623]" captionTargetId="figure-16@2.[151,1436,189,1644]" captionTargetPageId="2" captionText="Figures 1−7. Adult males of Zeuzerinae: 1. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., holotype (Museum Witt, Munich); 2. W. brehmi, paratype (Museum Witt, Munich); 3. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Museum Witt, Munich); 4. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (individual number: ANHRTUK: 00052313, African Natural History Research Trust, Leominster, GB); 5. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Museum Witt, Munich); 6. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Museum Witt, Munich); 7. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232073" httpUri="https://zenodo.org/record/13232073/files/figure.png" pageId="1" pageNumber="2">Figs 6</figureCitation>
,
<figureCitation id="97E800E87F7FFF8F15A3FA94FE12FA92" box="[437,469,1373,1400]" captionStart="Figures 8−13" captionStartId="3.[151,236,1710,1734]" captionTargetBox="[235,1308,194,1696]" captionTargetId="figure-12@3.[228,1358,189,1703]" captionTargetPageId="3" captionText="Figures 8−13. Genitalia of males of Zeuzerinae: 8. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., paratype (Genitalpräparat Heterocera N 33.185, Museum Witt, Munich); 9. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Genitalpräparat Heterocera N 32.924, Museum Witt, Munich); 10. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (ANHRT Gen Slide 00075, African Natural History Research Trust, Leominster, GB); 11. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Genitalpräparat Heterocera N 33.161, Museum Witt, Munich); 12. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Genitalpräparat Heterocera N 32.709, Museum Witt, Munich); 13. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Genitalpräparat Heterocera N 33.167, Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232075" httpUri="https://zenodo.org/record/13232075/files/figure.png" pageId="1" pageNumber="2">12</figureCitation>
), widely spread in the Palaearctic and Paleotropics, from which it differs in the more variable fore wing pattern, larger size and very thick base of the uncus. Externally, the new genus resembles to the genus
<taxonomicName id="C8D367EE7F7FFF8F15A2FA6BFCC6FA56" authority="Gaede, 1930" authorityName="Gaede" authorityYear="1930" box="[436,769,1441,1468]" class="Insecta" family="Cossidae" genus="Zeuzeropecten" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3DA7C07F7F7FFF8F15A2FA6BFD98FA51" box="[436,607,1442,1467]" italics="true" pageId="1" pageNumber="2">Zeuzeropecten</emphasis>
Gaede, 1930
</taxonomicName>
(
<typeStatus id="D068A2CF7F7FFF8F1705FA68FC80FA56" box="[787,839,1441,1468]" pageId="1" pageNumber="2">type</typeStatus>
species, by original designation −
<taxonomicName id="C8D367EE7F7FFF8F10E7FA6BFE77FA37" authority="Gaede, 1930" authorityName="Gaede" authorityYear="1930" class="Insecta" family="Cossidae" genus="Zeuzeropecten" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="lactescens">
<emphasis id="3DA7C07F7F7FFF8F10E7FA6BFEC8FA37" italics="true" pageId="1" pageNumber="2">Zeuzeropecten lactescens</emphasis>
Gaede, 1930
</taxonomicName>
) (
<figureCitation id="97E800E87F7FFF8F15DEFA0BFDD1FA37" box="[456,534,1474,1501]" captionStart="Figures 1−7" captionStartId="2.[151,237,1694,1718]" captionTargetBox="[159,1434,234,1623]" captionTargetId="figure-16@2.[151,1436,189,1644]" captionTargetPageId="2" captionText="Figures 1−7. Adult males of Zeuzerinae: 1. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., holotype (Museum Witt, Munich); 2. W. brehmi, paratype (Museum Witt, Munich); 3. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Museum Witt, Munich); 4. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (individual number: ANHRTUK: 00052313, African Natural History Research Trust, Leominster, GB); 5. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Museum Witt, Munich); 6. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Museum Witt, Munich); 7. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232073" httpUri="https://zenodo.org/record/13232073/files/figure.png" pageId="1" pageNumber="2">Figs 7</figureCitation>
,
<figureCitation id="97E800E87F7FFF8F1632FA0BFD83FA37" box="[548,580,1474,1501]" captionStart="Figures 8−13" captionStartId="3.[151,236,1710,1734]" captionTargetBox="[235,1308,194,1696]" captionTargetId="figure-12@3.[228,1358,189,1703]" captionTargetPageId="3" captionText="Figures 8−13. Genitalia of males of Zeuzerinae: 8. Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n., paratype (Genitalpräparat Heterocera N 33.185, Museum Witt, Munich); 9. Azygophleps scalaris (Fabricius, 1775), Mali, Mopti region, Dogon Plateau, 400−800 m, Nov.−Dec. 2010, leg. Müller, Kravchenko, Traore &amp; al. (Genitalpräparat Heterocera N 32.924, Museum Witt, Munich); 10. A. inclusa (Walker, 1856), Zambia, Jiwundu Swamp, S 11°51′54′′ / E 25°33′20′′, 20−22.x.2014, light trap, leg. Smith, Takano &amp; Oram (ANHRT Gen Slide 00075, African Natural History Research Trust, Leominster, GB); 11. Phragmacossia ariana (Grum-Grshimailo, 1899), Tajikistan, Gissar Mts., Kondara, 38º50′N / 68º50′E, July 1998, ex coll. A. Schintlmeister (Genitalpräparat Heterocera N 33.161, Museum Witt, Munich); 12. Phragmataecia castaneae (Hübner, 1790), “S. Russland, Rostov/Don, Nedwigovka, 10.vi.1991, leg. A. Poltawsky, coll. [A.] Schitlm[eister]]” (Genitalpräparat Heterocera N 32.709, Museum Witt, Munich); 13. Zeuzeropecten combustus (Kenrick, 1913), Madagascar, Ranomafana, 40 km NE Fianarantsoa, 900 m, 30.xii.2003−3.i.2004, leg. S. Murzin &amp; A. Shamaev (Genitalpräparat Heterocera N 33.167, Museum Witt, Munich)." figureDoi="http://doi.org/10.5281/zenodo.13232075" httpUri="https://zenodo.org/record/13232075/files/figure.png" pageId="1" pageNumber="2">13</figureCitation>
), distributed in east Africa and
<collectingCountry id="77C45CFD7F7FFF8F17ABFA0BFB89FA37" box="[957,1102,1474,1501]" name="Madagascar" pageId="1" pageNumber="2">Madagascar</collectingCountry>
, but differs in its very short saccus (in
<taxonomicName id="C8D367EE7F7FFF8F1501FA2CFE28FA15" box="[279,495,1509,1535]" class="Insecta" family="Cossidae" genus="Zeuzeropecten" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3DA7C07F7F7FFF8F1501FA2CFE05FA14" box="[279,450,1509,1534]" italics="true" pageId="1" pageNumber="2">Zeuzeropecten</emphasis>
sp.
</taxonomicName>
, the saccus is very long) and longer uncus (in
<taxonomicName id="C8D367EE7F7FFF8F1033FA2CFB39FA15" box="[1061,1278,1509,1535]" class="Insecta" family="Cossidae" genus="Zeuzeropecten" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3DA7C07F7F7FFF8F1033FA2CFB16FA14" box="[1061,1233,1509,1534]" italics="true" pageId="1" pageNumber="2">Zeuzeropecten</emphasis>
sp.
</taxonomicName>
, the uncus is very short).
</paragraph>
<paragraph id="0F6C1C6D7F7FFF8F1481F983FE56F96F" blockId="1.[151,1436,1609,1670]" pageId="1" pageNumber="2">
<emphasis id="3DA7C07F7F7FFF8F1481F983FEF0F98E" bold="true" box="[151,311,1610,1636]" pageId="1" pageNumber="2">Composition</emphasis>
. The new genus is monotypic, it includes only one species –
<taxonomicName id="C8D367EE7F7FFF8F103EF982FED9F96C" authority="(Yakovlev &amp; Witt, 2016)" authorityName="Yakovlev" authorityYear="2020" baseAuthorityName="Yakovlev &amp; Witt" baseAuthorityYear="2016" class="Insecta" family="Cossidae" genus="Wittoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="brehmi" status="comb. nov.">
<emphasis id="3DA7C07F7F7FFF8F103EF982FB31F98E" box="[1064,1270,1609,1636]" italics="true" pageId="1" pageNumber="2">Wittoecia brehmi</emphasis>
(
<bibRefCitation id="6B42619C7F7FFF8F111EF980FED0F96C" author="Yakovlev, R. V. &amp; Witt, Th." pageId="1" pageNumber="2" pagination="71 - 73" refId="ref3283" refString="Yakovlev, R. V. &amp; Witt, Th. (2016). Azygophleps brehmi Yakovlev et Witt, sp. n. - a new Carpenter-Moth (Lepidoptera, Cossidae) from Ethiopia. Russian Entomological Journal, 25 (1), 71 - 73." type="journal article" year="2016">Yakovlev &amp; Witt, 2016</bibRefCitation>
)
</taxonomicName>
<emphasis id="3DA7C07F7F7FFF8F1530F9A2FE56F96F" bold="true" box="[294,401,1643,1669]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="26947D047F7FFF8F1530F9A2FE56F96F" box="[294,401,1643,1669]" pageId="1" pageNumber="2" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="47C94FE67F7FFF8F1481F966FAC0F8E7" pageId="1" pageNumber="2" type="distribution">
<paragraph id="0F6C1C6D7F7FFF8F1481F966FBDEF920" blockId="1.[151,1049,1711,1738]" box="[151,1049,1711,1738]" pageId="1" pageNumber="2">
<emphasis id="3DA7C07F7F7FFF8F1481F966FEF7F923" bold="true" box="[151,304,1711,1737]" pageId="1" pageNumber="2">Distribution</emphasis>
. Recorded only for Central
<collectingCountry id="77C45CFD7F7FFF8F166FF966FD1BF920" box="[633,732,1711,1738]" name="Ethiopia" pageId="1" pageNumber="2">Ethiopia</collectingCountry>
(
<collectingRegion id="CD17D28F7F7FFF8F16FDF966FC5DF920" box="[747,922,1711,1738]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="2">Oromia region</collectingRegion>
) (
<figureCitation id="97E800E87F7FFF8F17A4F966FBC0F920" box="[946,1031,1711,1738]" captionStart="Figure 14" captionStartId="4.[151,226,1087,1111]" captionTargetBox="[371,1215,222,1068]" captionTargetId="figure-16@4.[371,1215,222,1068]" captionTargetPageId="4" captionText="Figure 14. Map of distribution of Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n." figureDoi="http://doi.org/10.5281/zenodo.13232077" httpUri="https://zenodo.org/record/13232077/files/figure.png" pageId="1" pageNumber="2">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="0F6C1C6D7F7FFF8F1481F93BFAC0F8E7" blockId="1.[151,1287,1778,1805]" box="[151,1287,1778,1805]" pageId="1" pageNumber="2">
<emphasis id="3DA7C07F7F7FFF8F1481F93BFF3FF8E6" bold="true" box="[151,248,1778,1804]" pageId="1" pageNumber="2">Habitat</emphasis>
. Mountain forests on the alt. of
<quantity id="C82BB1887F7FFF8F1666F93BFCD3F8E7" box="[624,788,1778,1805]" metricMagnitude="3" metricUnit="m" metricValue="2.369" metricValueMax="2.388" metricValueMin="2.35" pageId="1" pageNumber="2" unit="m" value="2369.0" valueMax="2388.0" valueMin="2350.0">2350−2388 m</quantity>
(
<figureCitation id="97E800E87F7FFF8F1732F93BFCBCF8E7" box="[804,891,1778,1805]" captionStart="Figure 15" captionStartId="4.[151,226,1979,2003]" captionTargetBox="[151,1434,1213,1960]" captionTargetId="figure-33@4.[151,1434,1213,1960]" captionTargetPageId="4" captionText="Figure 15. Habitat of Wittoecia brehmi (Yakovlev &amp; Witt, 2016) comb. n. (photo by Harald Sulak)." figureDoi="http://doi.org/10.5281/zenodo.13232079" httpUri="https://zenodo.org/record/13232079/files/figure.png" pageId="1" pageNumber="2">Fig. 15</figureCitation>
). Adult flight period: April–May.
</paragraph>
</subSubSection>
<subSubSection id="47C94FE67F7FFF8F1481F8FFFBD4F832" pageId="1" pageNumber="2" type="etymology">
<paragraph id="0F6C1C6D7F7FFF8F1481F8FFFBD4F832" blockId="1.[151,1437,1846,2008]" pageId="1" pageNumber="2">
<emphasis id="3DA7C07F7F7FFF8F1481F8FFFED9F8BA" bold="true" box="[151,286,1846,1872]" pageId="1" pageNumber="2">Etymology</emphasis>
. The new genus is named after Thomas J. Witt (1947−2019) (
<figureCitation id="97E800E87F7FFF8F17E9F8FFFB93F8BB" box="[1023,1108,1846,1873]" captionStart="Figure 16" captionStartId="5.[151,226,2011,2035]" captionText="Figure 16. Thomas J. Witt in January 2019 (photo by R. Yakovlev)." pageId="1" pageNumber="2">Fig. 16</figureCitation>
) the world largest collector of
<taxonomicName id="C8D367EE7F7FFF8F14ABF89EFE8AF898" authorityName="Linnaeus" authorityYear="1758" box="[189,333,1879,1906]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
, founder of the Witt Museum (Munich,
<collectingCountry id="77C45CFD7F7FFF8F175EF89EFC7DF898" box="[840,954,1879,1906]" name="Germany" pageId="1" pageNumber="2">Germany</collectingCountry>
), organizer of the fund “Thomas- Witt- Stiftung zur Förderung der Wissenschaft und Forschung im Bereich der Zoologischen Systematik”. Thomas J. Witt was awarded with the Ritter-von-Spix-Medal of Zoologischen Staatssammlung Munchen in 2001. He also received an honorary doctorate from the University of Munich in 2013.
</paragraph>
</subSubSection>
</treatment>
</document>