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<document id="586C32D9A53D4C75A5E3ED8F36ED9A7C" ID-DOI="10.37828/em.2020.36.1" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232063" ID-ZooBank="urn:lsid:zoobank.org:pub:1878F3DE-C9F8-4D7F-B562-73A99B9C552B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722913586256" checkinUser="felipe" docAuthor="Glöer, Peter &amp; Pešić, Vladimir" docDate="2020" docId="1B4D879A0759D02700F8FF7E277D4660" docLanguage="en" docName="EcolMontenegrina.36.1-5.pdf" docOrigin="Ecologica Montenegrina 36" docSource="http://dx.doi.org/10.37828/em.2020.36.1" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Bythinella reuselaarsi Glöer &amp; Pešić, 2020, n. sp." docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="E774FFE2075AD024006FFFBA25504265" masterDocTitle="Two new species of the genus Bythinella Moquin-Tandona, 1856 from the Greek island of Evia (Gastropoda: Bythinellidae)" masterLastPageNumber="5" masterPageNumber="1" pageNumber="4" updateTime="1722916243952" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3D183AED3F67785104604C35C601635C">Two new species of the genus Bythinella Moquin-Tandona, 1856 from the Greek island of Evia (Gastropoda: Bythinellidae)</mods:title>
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<mods:namePart id="DD4CC72B5485CEFE3FF36DCCD5FFA399">Glöer, Peter</mods:namePart>
<mods:affiliation id="0A9D424F3424F7F698861D804510DBCE">Biodiversity Research Lab, Schulstrasse 3, D- 25491 Hetlingen, Germany. E-mail: gloeer @ malaco. de</mods:affiliation>
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<mods:namePart id="701A58B85B33248D1F40850CE147C72E">Pešić, Vladimir</mods:namePart>
<mods:affiliation id="D9B1436B255986A0C8049B20A16AD4E3">Department of Biology, Faculty of Sciences, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro. E-mail: vladopesic @ gmail. com</mods:affiliation>
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<mods:title id="ECDC9327432FA2ED8F028442E169D59C">Ecologica Montenegrina</mods:title>
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<mods:date id="C6AC0E0D30A1F221CFA5DC948B137A56">2020</mods:date>
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<mods:number id="2FDF848310082FB7E9D54FE7138FB568">2020-11-01</mods:number>
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<mods:number id="D94315C418F6FF9C78F409CFC6E17AC5">36</mods:number>
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<mods:identifier id="DEEB3E509B318E4493EF0D895B688438" type="DOI">10.37828/em.2020.36.1</mods:identifier>
<mods:identifier id="B0D1885EB4CC4CE83EE0D236FE8E8AC2" type="ISSN">2336-9744</mods:identifier>
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<treatment id="1B4D879A0759D02700F8FF7E277D4660" ID-DOI="http://doi.org/10.5281/zenodo.13232173" ID-Zenodo-Dep="13232173" LSID="urn:lsid:plazi:treatment:1B4D879A0759D02700F8FF7E277D4660" httpUri="http://treatment.plazi.org/id/1B4D879A0759D02700F8FF7E277D4660" lastPageNumber="4" pageId="3" pageNumber="4">
<subSubSection id="DBFE65070759D02700F8FF7E212E4298" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="935B368C0759D02700F8FF7E24BC42BB" blockId="3.[151,1150,196,287]" box="[151,492,196,222]" pageId="3" pageNumber="4">
<heading id="C81381E00759D02700F8FF7E24BC42BB" box="[151,492,196,222]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<emphasis id="A190EA9E0759D02700F8FF7E24BC42BB" bold="true" box="[151,492,196,222]" pageId="3" pageNumber="4">
<taxonomicName id="54E44D0F0759D02700F8FF7E24F142BB" ID-CoL="7QHRD" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[151,417,196,222]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="reuselaarsi" status="sp. nov.">
<emphasis id="A190EA9E0759D02700F8FF7E24F142BB" bold="true" box="[151,417,196,222]" italics="true" pageId="3" pageNumber="4">Bythinella reuselaarsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50759D02701C6FF7E24BC42BB" box="[425,492,196,222]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="935B368C0759D02700F8FF5F212E4298" blockId="3.[151,1150,196,287]" box="[151,1150,229,253]" pageId="3" pageNumber="4">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="DBFE65070759D02700F8FEBE2457437A" box="[151,263,260,287]" pageId="3" pageNumber="4" type="description">
<paragraph id="935B368C0759D02700F8FEBE2457437A" blockId="3.[151,1150,196,287]" box="[151,263,260,287]" pageId="3" pageNumber="4">
<figureCitation id="0BDF2A090759D02700F8FEBE2457437A" box="[151,263,260,287]" captionStart="Figures 2-10. 2" captionStartId="2.[151,236,1245,1269]" captionTargetBox="[151,1439,337,1226]" captionTargetId="figure-70@2.[151,1439,336,1226]" captionTargetPageId="2" captionText="Figures 2-10. 2: Bythinella charpentieri (lectotype), 3-5: B. dimitrosensis Glöer &amp; Reuselaars, 2020a, 4: B. dimitrosensis from Evia Island, the region of Mount Ochi, Metochi spring; 5: B. dimitrosensis from Evia Island, the region of Mount Ochi, spring near Lenosei village; 6: B. kastanolongosensis n. sp. (holotype), 7: penis with tubular gland, 8: B. kastanolongosensis n. sp. (paratype), 9: B. reuselaarsi n. sp., 10: penis with tubular gland." figureDoi="http://doi.org/10.5281/zenodo.13232067" httpUri="https://zenodo.org/record/13232067/files/figure.png" pageId="3" pageNumber="4">Figs 9-10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DBFE65070759D02700F8FEF220C743A2" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="935B368C0759D02700F8FEF2269D43E1" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D02700F8FEF224DD4307" bold="true" box="[151,397,328,354]" pageId="3" pageNumber="4">Material examined</emphasis>
—
<materialsCitation id="238C3CD10759D02701A5FEFD269D43E1" collectionCode="ZMH" collectorName="Gloer." pageId="3" pageNumber="4" specimenCode="ZMH140807, ZMH140808" specimenCount="79" typeStatus="holotype">
<typeStatus id="4C5F882E0759D02701A5FEFD27674307" box="[458,567,327,354]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
(
<specimenCode id="C3429EF70759D0270220FEFD27BB4307" box="[591,747,327,354]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="4" type="Museum">ZMH140807</specimenCode>
): shell height
<quantity id="541C9B690759D02703C2FEFD21444307" box="[941,1044,327,354]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="4" unit="mm" value="3.1">3.1 mm</quantity>
, width
<quantity id="541C9B690759D0270415FEFD21B04307" box="[1146,1248,327,354]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="4" unit="mm" value="1.9">1.9 mm</quantity>
;
<typeStatus id="4C5F882E0759D027049AFEF2203A4307" box="[1269,1386,328,354]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
:
<specimenCount id="85E2FD050759D0270511FEFD244143E1" count="64" pageId="3" pageNumber="4" type="generic">64 specimens</specimenCount>
in isopropanol (
<specimenCode id="C3429EF70759D02701BFFED3273B43E1" box="[464,619,361,388]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="4" type="Museum">ZMH140808</specimenCode>
),
<specimenCount id="85E2FD050759D02702EBFED3267243E1" box="[644,802,361,388]" count="15" pageId="3" pageNumber="4" type="generic">15 specimens</specimenCount>
in coll.
<collectorName id="3E11535A0759D02703EDFED3269D43E1" box="[898,973,361,388]" pageId="3" pageNumber="4">Glöer.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="935B368C0759D0270098FE3120C743A2" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<materialsCitation id="238C3CD10759D0270098FE3120C743A2" collectingDate="2007-06-30" collectorName="Mount Ochi, GR &amp; Pesic." country="Greece" county="Melissonas village" latitude="38.090675" location="Evia Island" longLatPrecision="1" longitude="24.398603" municipality="Mount Ochi" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
<emphasis id="A190EA9E0759D0270098FE3124CE43C0" bold="true" box="[247,414,395,421]" pageId="3" pageNumber="4">
<typeStatus id="4C5F882E0759D0270098FE31246643C0" box="[247,310,395,421]" pageId="3" pageNumber="4">Type</typeStatus>
locality
</emphasis>
—
<collectingCountry id="EBF3761C0759D02701BCFE31277943C3" box="[467,553,395,422]" name="Greece" pageId="3" pageNumber="4">Greece</collectingCountry>
,
<location id="963B60570759D0270255FE31279643C3" LSID="urn:lsid:plazi:treatment:1B4D879A0759D02700F8FF7E277D4660:963B60570759D0270255FE31279643C3" box="[570,710,395,422]" country="Greece" county="Melissonas village" latitude="38.090675" longLatPrecision="1" longitude="24.398603" municipality="Mount Ochi" name="Evia Island" pageId="3" pageNumber="4">Evia Island</location>
, the region of
<collectorName id="3E11535A0759D02703EDFE36210243C3" box="[898,1106,395,422]" pageId="3" pageNumber="4">
<collectingMunicipality id="733FACF60759D02703EDFE36214943C3" box="[898,1049,395,422]" pageId="3" pageNumber="4">Mount Ochi</collectingMunicipality>
, GR
</collectorName>
4 limnocrene spring within
<taxonomicName id="54E44D0F0759D02700F8FE1625AF43A2" box="[151,255,428,455]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="A190EA9E0759D02700F8FE1625AF43A2" box="[151,255,428,455]" italics="true" pageId="3" pageNumber="4">Platanus</emphasis>
</taxonomicName>
grove along the road, near
<collectingCounty id="7A3A4E000759D027022FFE16264D43A2" box="[576,797,428,455]" pageId="3" pageNumber="4">Melissonas village</collectingCounty>
,
<geoCoordinate id="F6D0504B0759D0270345FE16269D43A2" box="[810,973,428,455]" degrees="38" direction="north" minutes="5" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="26.43" value="38.090675">38°5'26.43&quot;N</geoCoordinate>
,
<geoCoordinate id="F6D0504B0759D02703B6FE1621D843A2" box="[985,1160,428,455]" degrees="24" direction="east" minutes="23" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="54.97" value="24.398603">24°23'54.97&quot;E</geoCoordinate>
,
<date id="E75A104C0759D02704FCFE16204743A2" box="[1171,1303,428,455]" pageId="3" pageNumber="4" value="2007-06-30">
<collectingDate id="F71EE9A40759D02704FCFE16204743A2" box="[1171,1303,428,455]" pageId="3" pageNumber="4" value="2007-06-30">30.06.2007</collectingDate>
</date>
leg.
<collectorName id="3E11535A0759D027053DFE1720C743A2" box="[1362,1431,428,455]" pageId="3" pageNumber="4">Pešić.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DBFE65070759D027009AFE7527DE406E" pageId="3" pageNumber="4" type="etymology">
<paragraph id="935B368C0759D027009AFE7527DE406E" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D027009AFE75242C438C" bold="true" box="[245,380,463,489]" pageId="3" pageNumber="4">Etymology</emphasis>
— Named after Robert Reuselaars (Assen,
<collectingCountry id="EBF3761C0759D02703EAFE74211B438C" box="[901,1099,462,489]" name="Netherlands" pageId="3" pageNumber="4">The Netherlands</collectingCountry>
) in appreciation of his work in collecting
<taxonomicName id="54E44D0F0759D027015FFE4A24F9406E" authorityName="Moquin-Tandona" authorityYear="1856" box="[304,425,496,523]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="A190EA9E0759D027015FFE4A24F9406E" box="[304,425,496,523]" italics="true" pageId="3" pageNumber="4">Bythinella</emphasis>
</taxonomicName>
samples in
<collectingCountry id="EBF3761C0759D027025BFE4A27DA406E" box="[564,650,496,523]" name="Greece" pageId="3" pageNumber="4">Greece</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DBFE65070759D027009AFDA9271A41A7" pageId="3" pageNumber="4" type="description">
<paragraph id="935B368C0759D027009AFDA924D64048" blockId="3.[151,1436,327,1029]" box="[245,390,531,557]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D027009AFDA924D64048" bold="true" box="[245,390,531,557]" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="935B368C0759D027009AFD8924E040F7" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D027009AFD892460402B" box="[245,304,563,590]" italics="true" pageId="3" pageNumber="4">Shell</emphasis>
— Shell ovate-cylindrical with 4.5 slightly convex whorls separated by a deep suture. The aperture is ovate, somewhat oblique. The peristome is sharp. The umbilicus is closed. The shell is
<quantity id="541C9B690759D0270552FDEF20CA4015" box="[1341,1434,597,624]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="4" unit="mm" value="3.1">3.1 mm</quantity>
high and
<quantity id="541C9B690759D027016BFDCD240E40F7" box="[260,350,631,658]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="4" unit="mm" value="1.9">1.9 mm</quantity>
broad.
</paragraph>
<paragraph id="935B368C0759D0270098FD23269240B0" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D0270098FD23240F40D6" box="[247,351,665,691]" italics="true" pageId="3" pageNumber="4">Anatomy</emphasis>
— The penis is much shorter than the penial appendix. The tubular gland is thin at the proximal end and brod at the distal end (
<figureCitation id="0BDF2A090759D027021CFD00279840B0" box="[627,712,698,725]" captionStart="Figures 2-10. 2" captionStartId="2.[151,236,1245,1269]" captionTargetBox="[151,1439,337,1226]" captionTargetId="figure-70@2.[151,1439,336,1226]" captionTargetPageId="2" captionText="Figures 2-10. 2: Bythinella charpentieri (lectotype), 3-5: B. dimitrosensis Glöer &amp; Reuselaars, 2020a, 4: B. dimitrosensis from Evia Island, the region of Mount Ochi, Metochi spring; 5: B. dimitrosensis from Evia Island, the region of Mount Ochi, spring near Lenosei village; 6: B. kastanolongosensis n. sp. (holotype), 7: penis with tubular gland, 8: B. kastanolongosensis n. sp. (paratype), 9: B. reuselaarsi n. sp., 10: penis with tubular gland." figureDoi="http://doi.org/10.5281/zenodo.13232067" httpUri="https://zenodo.org/record/13232067/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
), with six half-loops.
</paragraph>
<paragraph id="935B368C0759D0270098FD67272E411B" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D0270098FD6727104092" bold="true" box="[247,576,733,759]" pageId="3" pageNumber="4">Differentiating characters</emphasis>
— The shell of
<taxonomicName id="54E44D0F0759D0270377FD6621704092" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[792,1056,732,759]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="reuselaarsi" status="sp. nov.">
<emphasis id="A190EA9E0759D0270377FD6621704092" box="[792,1056,732,759]" italics="true" pageId="3" pageNumber="4">Bythinella reuselaarsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50759D0270442FD6721214092" box="[1069,1137,733,759]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
is ovate-cylindrical, not elongated like in
<taxonomicName id="54E44D0F0759D0270104FD452771417C" authorityName="Roth" authorityYear="1855" box="[363,545,766,793]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="charpentieri">
<emphasis id="A190EA9E0759D0270104FD452771417C" box="[363,545,766,793]" italics="true" pageId="3" pageNumber="4">B. charpentieri</emphasis>
</taxonomicName>
. On the other hand,
<taxonomicName id="54E44D0F0759D0270342FD4526BD417C" authorityName="Gloer &amp; Reuselaars" authorityYear="2020" box="[813,1005,766,793]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="dimitrosensis">
<emphasis id="A190EA9E0759D0270342FD4526BD417C" box="[813,1005,766,793]" italics="true" pageId="3" pageNumber="4">B. dimitrosensis</emphasis>
</taxonomicName>
can be distinguished from the new species by a smaller size, ovate shell. The body whorl of
<taxonomicName id="54E44D0F0759D027032EFC9B26B6415E" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[833,998,800,827]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="reuselaarsi" status="n. sp.">
<emphasis id="A190EA9E0759D027032EFC9B26B6415E" box="[833,998,800,827]" italics="true" pageId="3" pageNumber="4">B. reuselaarsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50759D0270380FC9B217F415E" box="[1007,1071,801,827]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
is convex and not straight like in
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<emphasis id="A190EA9E0759D02700D7FCF824EF4139" box="[184,447,833,860]" italics="true" pageId="3" pageNumber="4">B. kastanolongosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50759D02701A6FCF8275A4139" box="[457,522,834,860]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
The tubular gland of
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<emphasis id="A190EA9E0759D0270379FCF8269B4139" box="[790,971,833,860]" italics="true" pageId="3" pageNumber="4">B. reuselaaarsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50759D02703B9FCF821474139" box="[982,1047,834,860]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
is longer (6 half-loops) vs. four half-loops in
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<emphasis id="A190EA9E0759D027015CFCDF2415411B" box="[307,325,869,894]" italics="true" pageId="3" pageNumber="4">B</emphasis>
.
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</taxonomicName>
<taxonomicNameLabel id="BAA357E50759D027022FFCDE272E411B" box="[576,638,868,894]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
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<emphasis id="A190EA9E0759D0270098FC3F240741FA" bold="true" box="[247,343,901,927]" pageId="3" pageNumber="4">Habitat</emphasis>
— Specimens were collected from a limnocrene spring with a well developed starwort (
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<emphasis id="A190EA9E0759D02700CEFC1C247041A4" box="[161,288,934,961]" italics="true" pageId="3" pageNumber="4">Callitriche</emphasis>
sp.
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) vegetation (
<figureCitation id="0BDF2A090759D027018CFC1D276941A7" box="[483,569,935,962]" captionStart="Figures 11-12" captionStartId="3.[151,236,1931,1955]" captionTargetBox="[151,1436,1063,1913]" captionTargetId="figure-326@3.[151,1436,1063,1913]" captionTargetPageId="3" captionText="Figures 11-12. Photographs of sampling site. 11: GR15 heleocrene spring in old-growth chestnut wood, Kastanolongos, locus typicus of Bythinella kastanolongosensis n. sp. 12: GR4 limnocrene spring near Melissonas, locus typicus of B. reuselaarsi n. sp. Photos: V. Pešić." figureDoi="http://doi.org/10.5281/zenodo.13232069" httpUri="https://zenodo.org/record/13232069/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
).
</paragraph>
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<paragraph id="935B368C0759D0270098FC73277D4660" blockId="3.[151,1436,327,1029]" pageId="3" pageNumber="4">
<emphasis id="A190EA9E0759D0270098FC7324C04186" bold="true" box="[247,400,969,995]" pageId="3" pageNumber="4">Distribution</emphasis>
—
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; Only known from the
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locality in the region of the Mt. Ochi in the southern part of the island of Evia.
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114
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<mods:title id="33723E89D6EB39832E9180EF0148F9C5">Two new species of the genus Bythinella Moquin-Tandona, 1856 from the Greek island of Evia (Gastropoda: Bythinellidae)</mods:title>
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<mods:affiliation id="C9EBF122BE1C4D99EC34147966D44FE3">Biodiversity Research Lab, Schulstrasse 3, D- 25491 Hetlingen, Germany. E-mail: gloeer @ malaco. de</mods:affiliation>
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<mods:affiliation id="E7FAB202A683DA51E9EDEB17730B63BB">Department of Biology, Faculty of Sciences, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro. E-mail: vladopesic @ gmail. com</mods:affiliation>
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<emphasis id="A190EA9E075BD02500F8FA6824EC4789" bold="true" box="[151,444,1490,1516]" italics="true" pageId="1" pageNumber="2">Bythinella dimitrosensis</emphasis>
Glöer &amp; Reuselaars, 2020
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<emphasis id="A190EA9E075BD02500F8F9AF2466444A" bold="true" box="[151,310,1557,1583]" pageId="1" pageNumber="2">New records</emphasis>
—
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<collectingCountry id="EBF3761C075BD025010BF9AF24EA4455" box="[356,442,1557,1584]" name="Greece" pageId="1" pageNumber="2">Greece</collectingCountry>
,
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, the region of
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: GR9 a big spring on road towards
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,
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,
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,
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leg.
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;
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GR16
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spring in
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,
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,
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,
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leg.
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<emphasis id="A190EA9E075BD02500F8F926245944D3" bold="true" box="[151,265,1692,1718]" pageId="1" pageNumber="2">Remarks</emphasis>
—
<taxonomicName id="54E44D0F075BD0250158F926270444D2" authorityName="Gloer &amp; Reuselaars" authorityYear="2020" box="[311,596,1692,1719]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="dimitrosensis">
<emphasis id="A190EA9E075BD0250158F926270444D2" box="[311,596,1692,1719]" italics="true" pageId="1" pageNumber="2">Bythinella dimitrosensis</emphasis>
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was recently described from the spring of Aghios Dimotros located in the southern part of Evia Island (
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<mods:namePart id="427068FC92329C82F93F34D6E3A460F9">Pešić, Vladimir</mods:namePart>
<mods:affiliation id="8612A8DF14DF8B99B7F1E00EFE0E51BB">Department of Biology, Faculty of Sciences, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro. E-mail: vladopesic @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="680D043C638E815A74FE5D79703EF7AF">text</mods:typeOfResource>
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<mods:titleInfo id="2037D2805CF44AD6C6B5313CFD018C4C">
<mods:title id="A2C4ABCA5CCD054D2B7382FF9794AC2D">Ecologica Montenegrina</mods:title>
</mods:titleInfo>
<mods:part id="9FFB2916091F2E6FB0E678542CBBB30F">
<mods:date id="80AD78368FB5CE1F18AD77450B272B66">2020</mods:date>
<mods:detail id="2D77F865C2C631A3D20DA4C31EE4746A" type="pubDate">
<mods:number id="AD62C27920F0ACECD48432D3F1A1AB5C">2020-11-01</mods:number>
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<mods:detail id="86237D56692820215EB92D2C0232AF90" type="volume">
<mods:number id="EE86B8CA3AAEC0B4E178370ECF523A0C">36</mods:number>
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<mods:start id="BB7B79D19E4F527D70C78F16D8A8578D">1</mods:start>
<mods:end id="19AAA215D8EB5275D2340A5EC020D716">5</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4F78670FB2CAE94A7AF699B862364192">
<mods:url id="8164896B300DA9E3F3EE74B3CA6C6A5D">http://dx.doi.org/10.37828/em.2020.36.1</mods:url>
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<mods:classification id="A0DD2D45440187284DE0748FABE46542">journal article</mods:classification>
<mods:identifier id="B2F4DF7B4E48691CED5DA9A7BFD8F831" type="DOI">10.37828/em.2020.36.1</mods:identifier>
<mods:identifier id="16C6E7C6F3F27C4AF947BD6518473465" type="ISSN">2336-9744</mods:identifier>
<mods:identifier id="BE7DF6EB20BAB20779EC7CC9933C5B0E" type="Zenodo-Dep">13232063</mods:identifier>
<mods:identifier id="15EF03C19ECC7D3A42E79EBF2113DADC" type="ZooBank">urn:lsid:zoobank.org:pub:1878F3DE-C9F8-4D7F-B562-73A99B9C552B</mods:identifier>
</mods:mods>
<treatment id="1B4D879A075BD02600F8F89920C645B7" LSID="urn:lsid:zoobank.org:act:A6B46268-5A30-4100-A33F-F08B7F28ABFB" httpUri="http://treatment.plazi.org/id/1B4D879A075BD02600F8F89920C645B7" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection id="DBFE6507075BD02500F8F8992126453E" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="935B368C075BD02500F8F89927014558" blockId="1.[151,1142,1827,1918]" box="[151,593,1827,1853]" pageId="1" pageNumber="2">
<heading id="C81381E0075BD02500F8F89927014558" box="[151,593,1827,1853]" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="A190EA9E075BD02500F8F89927014558" bold="true" box="[151,593,1827,1853]" pageId="1" pageNumber="2">
<taxonomicName id="54E44D0F075BD02500F8F89927574558" ID-CoL="7QHLX" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[151,519,1827,1853]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="kastanolongosensis" status="sp. nov.">
<emphasis id="A190EA9E075BD02500F8F89927574558" bold="true" box="[151,519,1827,1853]" italics="true" pageId="1" pageNumber="2">Bythinella kastanolongosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E5075BD0250261F89927014558" box="[526,593,1827,1853]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="935B368C075BD02500F8F8F92126453E" blockId="1.[151,1142,1827,1918]" box="[151,1142,1859,1883]" pageId="1" pageNumber="2">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="E7420C59075BD025021BF8F92126453E" box="[628,1142,1859,1883]" pageId="1" pageNumber="2">A6B46268-5A30-4100-A33F-F08B7F28ABFB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DBFE6507075BD02500F8F8D925A8451B" box="[151,248,1891,1918]" pageId="1" pageNumber="2" type="description">
<paragraph id="935B368C075BD02500F8F8D925A8451B" blockId="1.[151,1142,1827,1918]" box="[151,248,1891,1918]" pageId="1" pageNumber="2">
<figureCitation id="0BDF2A09075BD02500F8F8D925A8451B" box="[151,248,1891,1918]" captionStart="Figures 2-10. 2" captionStartId="2.[151,236,1245,1269]" captionTargetBox="[151,1439,337,1226]" captionTargetId="figure-70@2.[151,1439,336,1226]" captionTargetPageId="2" captionText="Figures 2-10. 2: Bythinella charpentieri (lectotype), 3-5: B. dimitrosensis Glöer &amp; Reuselaars, 2020a, 4: B. dimitrosensis from Evia Island, the region of Mount Ochi, Metochi spring; 5: B. dimitrosensis from Evia Island, the region of Mount Ochi, spring near Lenosei village; 6: B. kastanolongosensis n. sp. (holotype), 7: penis with tubular gland, 8: B. kastanolongosensis n. sp. (paratype), 9: B. reuselaarsi n. sp., 10: penis with tubular gland." figureDoi="http://doi.org/10.5281/zenodo.13232067" httpUri="https://zenodo.org/record/13232067/files/figure.png" pageId="1" pageNumber="2">Figs 6-8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DBFE6507075BD02600F8F81D261D4365" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="935B368C075BD02500F8F81D26FF4586" blockId="1.[151,1436,1958,2019]" pageId="1" pageNumber="2">
<emphasis id="A190EA9E075BD02500F8F81D24DD45A4" bold="true" box="[151,397,1959,1985]" pageId="1" pageNumber="2">Material examined</emphasis>
—
<materialsCitation id="238C3CD1075BD02501A5F81C26FF4586" collectionCode="ZMH" collectorName="Gloer." pageId="1" pageNumber="2" specimenCode="ZMH140809, ZMH140810" specimenCount="112" typeStatus="holotype">
<typeStatus id="4C5F882E075BD02501A5F81C276745A4" box="[458,567,1958,1985]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
(
<specimenCode id="C3429EF7075BD0250220F81C27BB45A4" box="[591,747,1958,1985]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="1" pageNumber="2" type="Museum">ZMH140809</specimenCode>
): shell height
<quantity id="541C9B69075BD02503C2F81C214445A4" box="[941,1044,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="2" unit="mm" value="3.1">3.1 mm</quantity>
, width
<quantity id="541C9B69075BD0250415F81C21B045A4" box="[1146,1248,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="2" unit="mm" value="1.8">1.8 mm</quantity>
;
<typeStatus id="4C5F882E075BD025049AF81D203A45A4" box="[1269,1386,1959,1985]" pageId="1" pageNumber="2" type="paratype">paratypes</typeStatus>
:
<specimenCount id="85E2FD05075BD0250511F81C24414586" count="96" pageId="1" pageNumber="2" type="generic">96 specimens</specimenCount>
in isopropanol (
<specimenCode id="C3429EF7075BD02501BFF872273B4586" box="[464,619,1992,2019]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="1" pageNumber="2" type="Museum">ZMH140810</specimenCode>
),
<specimenCount id="85E2FD05075BD02502EBF87226724586" box="[644,802,1992,2019]" count="16" pageId="1" pageNumber="2" type="generic">16 specimens</specimenCount>
coll.
<collectorName id="3E11535A075BD025030BF87226FF4586" box="[868,943,1992,2019]" pageId="1" pageNumber="2">Glöer.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="935B368C0758D026009AFF7E261D4365" blockId="2.[151,1435,195,324]" pageId="2" pageNumber="3">
<materialsCitation id="238C3CD10758D026009AFF7E261D4365" collectingDate="2007-01-07" collectorName="Mount Ochi, GR &amp; Pesic." country="Greece" county="Kastanolongos" latitude="38.05242" location="Evia Island" longLatPrecision="1" longitude="24.471064" municipality="Mount Ochi" pageId="2" pageNumber="3" specimenCount="1" typeStatus="holotype">
<emphasis id="A190EA9E0758D026009AFF7E24C942BB" bold="true" box="[245,409,196,222]" pageId="2" pageNumber="3">
<typeStatus id="4C5F882E0758D026009AFF7E246442BB" box="[245,308,196,222]" pageId="2" pageNumber="3">Type</typeStatus>
locality
</emphasis>
—
<collectingCountry id="EBF3761C0758D02601A4FF79277142BB" box="[459,545,195,222]" name="Greece" pageId="2" pageNumber="3">Greece</collectingCountry>
,
<location id="963B60570758D026025FFF7927EA42BB" LSID="urn:lsid:plazi:treatment:1B4D879A075BD02600F8F89920C645B7:963B60570758D026025FFF7927EA42BB" box="[560,698,195,222]" country="Greece" county="Kastanolongos" latitude="38.05242" longLatPrecision="1" longitude="24.471064" municipality="Mount Ochi" name="Evia Island" pageId="2" pageNumber="3">Evia Island</location>
, the region of
<collectorName id="3E11535A0758D0260300FF7E216D42BB" box="[879,1085,195,222]" pageId="2" pageNumber="3">
<collectingMunicipality id="733FACF60758D0260300FF7E215342BB" box="[879,1027,195,222]" pageId="2" pageNumber="3">Mount Ochi</collectingMunicipality>
, GR
</collectorName>
15
<collectingCounty id="7A3A4E000758D026040DFF79204342BB" box="[1122,1299,195,222]" pageId="2" pageNumber="3">Kastanolongos</collectingCounty>
, helocrenic spring,
<geoCoordinate id="F6D0504B0758D0260080FF5F24D34365" box="[239,387,229,256]" degrees="38" direction="north" minutes="3" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="8.72" value="38.05242">38°3'8.72&quot;N</geoCoordinate>
,
<geoCoordinate id="F6D0504B0758D02601E0FF5F276A4365" box="[399,570,229,256]" degrees="24" direction="east" minutes="28" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="15.83" value="24.471064">24°28'15.83&quot;E</geoCoordinate>
,
<date id="E75A104C0758D0260225FF5F279E4365" box="[586,718,229,256]" pageId="2" pageNumber="3" value="2007-01-07">
<collectingDate id="F71EE9A40758D0260225FF5F279E4365" box="[586,718,229,256]" pageId="2" pageNumber="3" value="2007-01-07">01.07.2007</collectingDate>
</date>
leg.
<collectorName id="3E11535A0758D0260368FF5F261D4365" box="[775,845,229,256]" pageId="2" pageNumber="3">Pešić.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DBFE65070758D026009AFEBD215E4321" pageId="2" pageNumber="3" type="etymology">
<paragraph id="935B368C0758D026009AFEBD215E4321" blockId="2.[151,1435,195,324]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D026009AFEBD242D4344" bold="true" box="[245,381,263,289]" pageId="2" pageNumber="3">Etymology</emphasis>
— Named after the beautiful chestnut tree forest (“Kastanolongos”) on the slopes of Mount Ochi (&quot;Oche&quot;) where the
<emphasis id="A190EA9E0758D0260277FE9327E04326" box="[536,688,297,324]" italics="true" pageId="2" pageNumber="3">locus typicus</emphasis>
of the new species is located.
</paragraph>
</subSubSection>
<caption id="C79B66040758D02600F8FB67218A4734" ID-DOI="http://doi.org/10.5281/zenodo.13232067" ID-Zenodo-Dep="13232067" httpUri="https://zenodo.org/record/13232067/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,236,1245,1269]" targetBox="[151,1439,337,1226]" targetPageId="2" targetType="figure">
<paragraph id="935B368C0758D02600F8FB67218A4734" blockId="2.[151,1437,1245,1361]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D02600F8FB6724654690" bold="true" box="[151,309,1245,1269]" pageId="2" pageNumber="3">Figures 2-10</emphasis>
.
<emphasis id="A190EA9E0758D026013DFB67240F4690" bold="true" box="[338,351,1245,1269]" pageId="2" pageNumber="3">2</emphasis>
:
<taxonomicName id="54E44D0F0758D0260111FB6727D64690" authorityName="Roth" authorityYear="1855" box="[382,646,1245,1269]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="charpentieri">
<emphasis id="A190EA9E0758D0260111FB6727D64690" box="[382,646,1245,1269]" italics="true" pageId="2" pageNumber="3">Bythinella charpentieri</emphasis>
</taxonomicName>
(lectotype),
<emphasis id="A190EA9E0758D0260340FB6726024690" bold="true" box="[815,850,1245,1269]" pageId="2" pageNumber="3">3-5</emphasis>
:
<taxonomicName id="54E44D0F0758D026031EFB6420C64690" authority="Gloer &amp; Reuselaars, 2020 a" authorityName="Gloer &amp; Reuselaars" authorityYear="2020" box="[881,1430,1245,1269]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="dimitrosensis">
<emphasis id="A190EA9E0758D026031EFB64217C4690" box="[881,1068,1245,1269]" italics="true" pageId="2" pageNumber="3">B. dimitrosensis</emphasis>
<bibRefCitation id="F7754B7D0758D026042BFB6720C64690" author="Gloer, P. &amp; Reuselaars, R." box="[1092,1430,1245,1269]" pageId="2" pageNumber="3" pagination="20 - 33" refId="ref2098" refString="Gloer, P. &amp; Reuselaars, R. (2020 a) The Bythinella spp. of Greece (Gastropoda: Hydrobioidea: Bythinelliidae). Ecologica Montenegrina, 29, 20 - 33. https: // doi. org / 10.37828 / em. 2020.29.3" type="journal article" year="2020">Glöer &amp; Reuselaars, 2020a</bibRefCitation>
</taxonomicName>
,
<emphasis id="A190EA9E0758D02600F8FB4625F44771" bold="true" box="[151,164,1276,1300]" pageId="2" pageNumber="3">4</emphasis>
:
<taxonomicName id="54E44D0F0758D02600DBFB4624314776" authorityName="Gloer &amp; Reuselaars" authorityYear="2020" box="[180,353,1275,1299]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="dimitrosensis">
<emphasis id="A190EA9E0758D02600DBFB4624314776" box="[180,353,1275,1299]" italics="true" pageId="2" pageNumber="3">B. dimitrosensis</emphasis>
</taxonomicName>
from Evia Island, the region of Mount Ochi, Metochi spring;
<emphasis id="A190EA9E0758D0260464FB4621484771" bold="true" box="[1035,1048,1276,1300]" pageId="2" pageNumber="3">5</emphasis>
:
<taxonomicName id="54E44D0F0758D0260447FB4621864776" authorityName="Gloer &amp; Reuselaars" authorityYear="2020" box="[1064,1238,1275,1299]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="dimitrosensis">
<emphasis id="A190EA9E0758D0260447FB4621864776" box="[1064,1238,1275,1299]" italics="true" pageId="2" pageNumber="3">B. dimitrosensis</emphasis>
</taxonomicName>
from Evia Island, the region of Mount Ochi, spring near Lenosei village;
<emphasis id="A190EA9E0758D0260283FAA027A94757" bold="true" box="[748,761,1306,1330]" pageId="2" pageNumber="3">6</emphasis>
:
<taxonomicName id="54E44D0F0758D0260366FAA126A74757" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[777,1015,1306,1330]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="kastanolongosensis" status="n. sp.">
<emphasis id="A190EA9E0758D0260366FAA126A74757" box="[777,1015,1306,1330]" italics="true" pageId="2" pageNumber="3">B. kastanolongosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50758D0260390FAA121694757" box="[1023,1081,1307,1330]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
(holotype),
<emphasis id="A190EA9E0758D02604D2FAA0219A4757" bold="true" box="[1213,1226,1306,1330]" pageId="2" pageNumber="3">7</emphasis>
: penis with tubular gland, 8:
<taxonomicName id="54E44D0F0758D0260094FA8024B84734" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[251,488,1337,1361]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="kastanolongosensis" status="n. sp.">
<emphasis id="A190EA9E0758D0260094FA8024B84734" box="[251,488,1337,1361]" italics="true" pageId="2" pageNumber="3">B. kastanolongosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50758D0260180FA8327784735" box="[495,552,1337,1360]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
(paratype),
<emphasis id="A190EA9E0758D02602C5FA8327E74734" bold="true" box="[682,695,1337,1361]" pageId="2" pageNumber="3">9</emphasis>
:
<taxonomicName id="54E44D0F0758D02602AAFA80260A4734" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[709,858,1337,1361]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="reuselaarsi" status="n. sp.">
<emphasis id="A190EA9E0758D02602AAFA80260A4734" box="[709,858,1337,1361]" italics="true" pageId="2" pageNumber="3">B. reuselaarsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50758D026030EFA8326CC4735" box="[865,924,1337,1360]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
,
<emphasis id="A190EA9E0758D02603C7FA8326934734" bold="true" box="[936,963,1337,1361]" pageId="2" pageNumber="3">10</emphasis>
: penis with tubular gland.
</paragraph>
</caption>
<subSubSection id="DBFE65070758D02600F8FA2624E745D4" pageId="2" pageNumber="3" type="description">
<paragraph id="935B368C0758D02600F8FA26247847D3" blockId="2.[151,1436,1436,2002]" box="[151,296,1436,1462]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D02600F8FA26247847D3" bold="true" box="[151,296,1436,1462]" pageId="2" pageNumber="3">Description</emphasis>
</paragraph>
<paragraph id="935B368C0758D0260098FA07204B4479" blockId="2.[151,1436,1436,2002]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D0260098FA07246247BD" box="[247,306,1469,1496]" italics="true" pageId="2" pageNumber="3">Shell</emphasis>
— Cylindrical with 4.5 slightly convex whorls with a deep suture. The body whorl has a straight vertical tangent line. The aperture is ovate, slightly angled at the top. The peristome is sharp with a brown periostracum. The umbilicus is closed. The shell is
<quantity id="541C9B690758D026032BF9BB269C4479" box="[836,972,1537,1564]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="2" pageNumber="3" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8-3.1 mm</quantity>
high and
<quantity id="541C9B690758D026042EF9BB21994479" box="[1089,1225,1537,1564]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="2" pageNumber="3" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8-1.9 mm</quantity>
broad.
</paragraph>
<paragraph id="935B368C0758D0260098F99826D54458" blockId="2.[151,1436,1436,2002]" box="[247,901,1570,1597]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D0260098F998241C4458" box="[247,332,1570,1597]" italics="true" pageId="2" pageNumber="3">Animal</emphasis>
— The head is light-brown with a white border.
</paragraph>
<paragraph id="935B368C0758D0260098F9FF263044E4" blockId="2.[151,1436,1436,2002]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D0260098F9FF2430443A" box="[247,352,1605,1631]" italics="true" pageId="2" pageNumber="3">Anatomy</emphasis>
— The penis is much shorter than the penial appendix (
<figureCitation id="0BDF2A090758D0260461F9FE2107443A" box="[1038,1111,1604,1631]" captionStart="Figures 2-10. 2" captionStartId="2.[151,236,1245,1269]" captionTargetBox="[151,1439,337,1226]" captionTargetId="figure-70@2.[151,1439,336,1226]" captionTargetPageId="2" captionText="Figures 2-10. 2: Bythinella charpentieri (lectotype), 3-5: B. dimitrosensis Glöer &amp; Reuselaars, 2020a, 4: B. dimitrosensis from Evia Island, the region of Mount Ochi, Metochi spring; 5: B. dimitrosensis from Evia Island, the region of Mount Ochi, spring near Lenosei village; 6: B. kastanolongosensis n. sp. (holotype), 7: penis with tubular gland, 8: B. kastanolongosensis n. sp. (paratype), 9: B. reuselaarsi n. sp., 10: penis with tubular gland." figureDoi="http://doi.org/10.5281/zenodo.13232067" httpUri="https://zenodo.org/record/13232067/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
). Tubular gland thin at the proximal end, broader at the distal part, with four half-loops.
</paragraph>
<paragraph id="935B368C0758D0260098F93227A1452E" blockId="2.[151,1436,1436,2002]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D0260098F932276B44C7" bold="true" box="[247,571,1672,1698]" pageId="2" pageNumber="3">Differentiating characters</emphasis>
— The new species resembles
<taxonomicName id="54E44D0F0758D02603C0F932219044C6" authorityName="Roth" authorityYear="1855" box="[943,1216,1672,1699]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="charpentieri">
<emphasis id="A190EA9E0758D02603C0F932219044C6" box="[943,1216,1672,1699]" italics="true" pageId="2" pageNumber="3">Bythinella charpentieri</emphasis>
</taxonomicName>
. The latter species was recently redescribed by
<bibRefCitation id="F7754B7D0758D026019CF913267944A1" author="Georgiev, D. &amp; Gloer, P." box="[499,809,1705,1732]" pageId="2" pageNumber="3" pagination="129 - 137" refId="ref1883" refString="Georgiev, D. &amp; Gloer, P. (2020) Three new hydrobiid species from Thassos Island (Greece) with a redescription of Amnicola charpentieri Roth, 1855. Ecologica Montenegrina, 35, 129 - 137. https: // doi. org / 10.37828 / em. 2020.35.10" type="journal article" year="2020">Georgiev &amp; Glöer (2020)</bibRefCitation>
on the basis of a
<typeStatus id="4C5F882E0758D0260462F910213744A1" box="[1037,1127,1706,1732]" pageId="2" pageNumber="3" type="syntype">syntype</typeStatus>
material from Hymettus,
<collectingRegion id="5120F86E0758D02600F8F971258E4483" box="[151,222,1739,1766]" country="Greece" name="Attiki" pageId="2" pageNumber="3">Attica</collectingRegion>
. The shell of
<taxonomicName id="54E44D0F0758D02601EDF97627644483" authorityName="Roth" authorityYear="1855" box="[386,564,1739,1766]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="charpentieri">
<emphasis id="A190EA9E0758D02601EDF97627644483" box="[386,564,1739,1766]" italics="true" pageId="2" pageNumber="3">B. charpentieri</emphasis>
</taxonomicName>
is elongated conical and not cylindrical as in
<taxonomicName id="54E44D0F0758D026043FF97620054483" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[1104,1365,1739,1766]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="kastanolongosensis" status="n. sp.">
<emphasis id="A190EA9E0758D026043FF97620054483" box="[1104,1365,1739,1766]" italics="true" pageId="2" pageNumber="3">B. kastanolongosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50758D0260532F97620CC4483" box="[1373,1436,1740,1766]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
Moreover the tangent line of the body whorl in
<taxonomicName id="54E44D0F0758D02602BEF95426D4456D" authorityName="Roth" authorityYear="1855" box="[721,900,1773,1800]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="charpentieri">
<emphasis id="A190EA9E0758D02602BEF95427B34562" box="[721,739,1774,1799]" italics="true" pageId="2" pageNumber="3">B</emphasis>
.
<emphasis id="A190EA9E0758D026029CF95726D4456D" box="[755,900,1773,1800]" italics="true" pageId="2" pageNumber="3">charpentieri</emphasis>
</taxonomicName>
is not straight as in the new species (
<figureCitation id="0BDF2A090758D0260529F95720DE456D" box="[1350,1422,1773,1800]" captionStart="Figures 2-10. 2" captionStartId="2.[151,236,1245,1269]" captionTargetBox="[151,1439,337,1226]" captionTargetId="figure-70@2.[151,1439,336,1226]" captionTargetPageId="2" captionText="Figures 2-10. 2: Bythinella charpentieri (lectotype), 3-5: B. dimitrosensis Glöer &amp; Reuselaars, 2020a, 4: B. dimitrosensis from Evia Island, the region of Mount Ochi, Metochi spring; 5: B. dimitrosensis from Evia Island, the region of Mount Ochi, spring near Lenosei village; 6: B. kastanolongosensis n. sp. (holotype), 7: penis with tubular gland, 8: B. kastanolongosensis n. sp. (paratype), 9: B. reuselaarsi n. sp., 10: penis with tubular gland." figureDoi="http://doi.org/10.5281/zenodo.13232067" httpUri="https://zenodo.org/record/13232067/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). The shell of
<taxonomicName id="54E44D0F0758D0260159F8AA24A7454F" authorityName="Gloer &amp; Reuselaars" authorityYear="2020" box="[310,503,1807,1834]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="dimitrosensis">
<emphasis id="A190EA9E0758D0260159F8AA24A7454F" box="[310,503,1807,1834]" italics="true" pageId="2" pageNumber="3">B. dimitrosensis</emphasis>
</taxonomicName>
, a species known from southern part of the island of Evia, is ovate and the species is smaller than
<taxonomicName id="54E44D0F0758D02601CAF88827FA452E" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[421,682,1840,1867]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="kastanolongosensis" status="n. sp.">
<emphasis id="A190EA9E0758D02601CAF88827FA452E" box="[421,682,1840,1867]" italics="true" pageId="2" pageNumber="3">B. kastanolongosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50758D02602DDF88B27A1452E" box="[690,753,1841,1867]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="935B368C0758D0260098F8E9203D4508" blockId="2.[151,1436,1436,2002]" box="[247,1389,1874,1901]" pageId="2" pageNumber="3">
Due to its size the new species is similar to
<taxonomicName id="54E44D0F0758D0260298F8E926CB4508" authorityName="Glöer &amp; Pešić" authorityYear="2020" box="[759,923,1874,1901]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="reuselaarsi" status="n. sp.">
<emphasis id="A190EA9E0758D0260298F8E926CB4508" box="[759,923,1874,1901]" italics="true" pageId="2" pageNumber="3">B. reuselaarsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BAA357E50758D02603CDF8E926B14508" box="[930,993,1875,1901]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
(see there for further discussion).
</paragraph>
<paragraph id="935B368C0758D0260098F8CE24E745D4" blockId="2.[151,1436,1436,2002]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D0260098F8CE240745EB" bold="true" box="[247,343,1908,1934]" pageId="2" pageNumber="3">Habitat</emphasis>
— Specimens were collected from the stones in a small seepage spring in old-growth chestnut wood (
<figureCitation id="0BDF2A090758D026013DF82C24F645D4" box="[338,422,1942,1969]" captionStart="Figures 11-12" captionStartId="3.[151,236,1931,1955]" captionTargetBox="[151,1436,1063,1913]" captionTargetId="figure-326@3.[151,1436,1063,1913]" captionTargetPageId="3" captionText="Figures 11-12. Photographs of sampling site. 11: GR15 heleocrene spring in old-growth chestnut wood, Kastanolongos, locus typicus of Bythinella kastanolongosensis n. sp. 12: GR4 limnocrene spring near Melissonas, locus typicus of B. reuselaarsi n. sp. Photos: V. Pešić." figureDoi="http://doi.org/10.5281/zenodo.13232069" httpUri="https://zenodo.org/record/13232069/files/figure.png" pageId="2" pageNumber="3">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DBFE65070758D0260098F80220C645B7" box="[247,1430,1975,2002]" pageId="2" pageNumber="3" type="distribution">
<paragraph id="935B368C0758D0260098F80220C645B7" blockId="2.[151,1436,1436,2002]" box="[247,1430,1975,2002]" pageId="2" pageNumber="3">
<emphasis id="A190EA9E0758D0260098F80224C045B7" bold="true" box="[247,400,1976,2002]" pageId="2" pageNumber="3">Distribution</emphasis>
—
<collectingCountry id="EBF3761C0758D02601D3F80D274245B7" box="[444,530,1975,2002]" name="Greece" pageId="2" pageNumber="3">Greece</collectingCountry>
; only known from the
<typeStatus id="4C5F882E0758D0260372F802261F45B7" box="[797,847,1976,2002]" pageId="2" pageNumber="3">type</typeStatus>
locality in the southern part of the island of Evia.
</paragraph>
</subSubSection>
</treatment>
</document>