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<document id="B4632AD4BB8AFFF6E7D66256EE50EB92" ID-DOI="10.48156/1388.2024.1917256" ID-ISSN="2159-6719" ID-Zenodo-Dep="15555034" checkinTime="1748616411025" checkinUser="felipe" docAuthor="Tennessen, Kenneth J." docDate="2024" docId="03A2E731F016FF88FCBF79EAFC359F88" docLanguage="en" docName="IntJOdonatol.27.19-25.pdf" docOrigin="International Journal of Odonatology 27" docSource="https://doi.org/10.48156/1388.2024.1917256" docStyle="DocumentStyle:BF61EFB44F4363273DD5896762672266:IntJOdonatol.2022-.journal_article" docStyleId="BF61EFB44F4363273DD5896762672266" docStyleName="IntJOdonatol.2022-.journal_article" docTitle="Gomphomacromia signata Tennessen, 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="FF9B9F49F017FF8DFF927938FFF49918" masterDocTitle="Gomphomacromia signata sp. n. from the Andes Mountains in Ecuador (Odonata: Synthemistidae)" masterLastPageNumber="25" masterPageNumber="19" pageNumber="20" updateTime="1748616742094" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B4F38CB5838364D01931C440230D6BCA">Gomphomacromia signata sp. n. from the Andes Mountains in Ecuador (Odonata: Synthemistidae)</mods:title>
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<mods:namePart id="D1035DFFD0E898613B5AB18701B35CB2">Tennessen, Kenneth J.</mods:namePart>
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<mods:title id="1C97AD648BD14DC66186BD0C69F7F103">International Journal of Odonatology</mods:title>
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<mods:date id="E4C47702EB2ADD0FB5677AF44CE2AD86">2024</mods:date>
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<mods:number id="36F9CB5C30881DD43262DA2080CAFB8E">2024-01-25</mods:number>
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<mods:identifier id="4E0D1B5CF31BC99128D8603E5C228D2C" type="DOI">10.48156/1388.2024.1917256</mods:identifier>
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<subSubSection id="C31105ACF016FF8CFCBF79EAFACD99EA" box="[813,1337,210,242]" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="8BB45627F016FF8CFCBF79EAFACD99EA" blockId="1.[813,1337,210,275]" box="[813,1337,210,242]" pageId="1" pageNumber="20">
<heading id="D0FCE14BF016FF8CFCBF79EAFACD99EA" box="[813,1337,210,242]" fontSize="11" level="2" pageId="1" pageNumber="20" reason="7">
<emphasis id="B97F8A35F016FF8CFCBF79EAFACD99EA" bold="true" box="[813,1337,210,242]" pageId="1" pageNumber="20">
<taxonomicName id="4C0B2DA4F016FF8CFCBF79EAFB9A99F7" authorityName="Tennessen" authorityYear="2024" box="[813,1134,210,239]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="signata" status="sp. nov.">Gomphomacromia signata</taxonomicName>
<emphasis id="B97F8A35F016FF8CFBE779EEFACD99EA" bold="true" box="[1141,1337,214,242]" italics="true" pageId="1" pageNumber="20">
<taxonomicNameLabel id="A24C374EF016FF8CFBE779EEFB4299EA" box="[1141,1206,214,242]" pageId="1" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
Tennessen
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31105ACF016FF88FCBF79CFFC359F88" lastPageId="5" lastPageNumber="24" pageId="1" pageNumber="20" type="description">
<paragraph id="8BB45627F016FF8CFCBF79CFFBFB980B" blockId="1.[813,1337,210,275]" box="[813,1039,247,275]" pageId="1" pageNumber="20">
(
<figureCitation id="13304AA2F016FF8CFCA179CFFC35980B" box="[819,961,247,275]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 46" captionStartId-0="1.[121,189,2083,2108]" captionStartId-1="2.[121,189,1041,1066]" captionStartId-2="2.[121,189,2082,2107]" captionStartId-3="3.[813,880,2018,2043]" captionTargetBox-0="[127,1458,1007,2045]" captionTargetBox-1="[283,1304,144,1001]" captionTargetBox-2="[283,1303,1142,2042]" captionTargetBox-3="[814,1417,1652,1980]" captionTargetId-0="figure-328@1.[121,1467,989,2060]" captionTargetId-1="figure-24@2.[283,1304,144,1001]" captionTargetId-2="figure-44@2.[283,1304,1135,2045]" captionTargetId-3="figure-652@3.[810,1450,1621,1990]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionText-0="Figure 1. Gomphomacromia signata sp. n., holotype male." captionText-1="Figure 2. Gomphomacromia signata sp. n., holotype male, showing living eye color and postfrons pattern in anterior view." captionText-2="Figure 3. Gomphomacromia signata sp. n., holotype male accessory genitalia, ventral view." captionText-3="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi-0="http://doi.org/10.5281/zenodo.15555036" figureDoi-1="http://doi.org/10.5281/zenodo.15555038" figureDoi-2="http://doi.org/10.5281/zenodo.15555040" figureDoi-3="http://doi.org/10.5281/zenodo.15555042" httpUri-0="https://zenodo.org/record/15555036/files/figure.png" httpUri-1="https://zenodo.org/record/15555038/files/figure.png" httpUri-2="https://zenodo.org/record/15555040/files/figure.png" httpUri-3="https://zenodo.org/record/15555042/files/figure.png" pageId="1" pageNumber="20">Figures 16</figureCitation>
,
<figureCitation id="13304AA2F016FF8CFC5E79CFFC2A980B" box="[972,990,247,275]" captionStart="Figure 9" captionStartId="5.[121,189,2054,2079]" captionTargetBox="[123,770,1388,2002]" captionTargetId="figure-186@5.[118,775,1648,1765]" captionTargetPageId="5" captionText="Figure 9. Lef posterior hamule of Gomphomacromia signata sp. n. and G. fallax, mediolateral view." figureDoi="http://doi.org/10.5281/zenodo.15555048" httpUri="https://zenodo.org/record/15555048/files/figure.png" pageId="1" pageNumber="20">9</figureCitation>
,
<figureCitation id="13304AA2F016FF8CFC7B79CFFBFD980B" box="[1001,1033,247,275]" captionStart="Figure 10" captionStartId="6.[121,189,958,983]" captionTargetBox="[122,770,151,917]" captionTargetId="figure-302@6.[121,775,141,923]" captionTargetPageId="6" captionText="Figure 10. Lamina of anterior hamule of Gomphomacromia signata sp. n. and G. fallax, lateral view at slight anterior angle." figureDoi="http://doi.org/10.5281/zenodo.15555050" httpUri="https://zenodo.org/record/15555050/files/figure.png" pageId="1" pageNumber="20">10</figureCitation>
)
</paragraph>
<paragraph id="8BB45627F016FF8CFCBF7802FC5E984D" blockId="1.[813,938,314,341]" box="[813,938,314,341]" pageId="1" pageNumber="20">
<emphasis id="B97F8A35F016FF8CFCBF7802FC5E984D" box="[813,938,314,341]" italics="true" pageId="1" pageNumber="20">Etymology</emphasis>
</paragraph>
<paragraph id="8BB45627F016FF8CFCBF7844FB059B27" blockId="1.[813,1467,380,575]" pageId="1" pageNumber="20">
Name formed from the Latin
<emphasis id="B97F8A35F016FF8CFB3F7844FAFF988F" box="[1197,1291,380,407]" italics="true" pageId="1" pageNumber="20">signum</emphasis>
(adj., bearing marks, signs, or flags), referring collectively to the yel- low spots on the frons, the isolated oval yellow spot on the anterolateral part of the mesepimeron and the forked metepisternal and metepimeral yellow stripes, a unique combination within the genus.
</paragraph>
<paragraph id="8BB45627F016FF8CFCBF7B5EFC409B99" blockId="1.[813,948,613,641]" box="[813,948,613,641]" pageId="1" pageNumber="20">
<emphasis id="B97F8A35F016FF8CFCBF7B5EFC6C9B99" box="[813,920,614,641]" italics="true" pageId="1" pageNumber="20">
<typeStatus id="54B0E885F016FF8CFCBF7B5EFC6C9B99" box="[813,920,614,641]" pageId="1" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D0D9DAEF016FF8CFC0C7B5DFC409B99" box="[926,948,613,641]" count="1" pageId="1" pageNumber="20" type="male"></specimenCount>
</paragraph>
<paragraph id="8BB45627F016FF8CFCBF7B90FC559A52" blockId="1.[813,1467,680,842]" pageId="1" pageNumber="20">
<collectingCountry id="F31C16B7F016FF8CFCBF7B90FC649BDC" box="[813,912,680,708]" name="Ecuador" pageId="1" pageNumber="20">Ecuador</collectingCountry>
,
<collectingRegion id="49CF98C5F016FF8CFC097B90FBBB9BDC" box="[923,1103,680,708]" country="Ecuador" name="Napo" pageId="1" pageNumber="20">Napo Province</collectingRegion>
, hillside seep along Hwy. E20, about
<quantity id="4CF3FBC2F016FF8CFCEA7BF2FC339BFE" box="[888,967,714,742]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="20" unit="km" value="5.5">5.5 km</quantity>
E of Papallacta in a straight line (
<geoCoordinate id="EE3F30E0F016FF8CFAAC7BF2FA4C9BFE" box="[1342,1464,714,742]" degrees="0.37436" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="1" value="-0.37436">0.37436 S</geoCoordinate>
,
<geoCoordinate id="EE3F30E0F016FF8CFCBF7BD3FC499A1F" box="[813,957,747,775]" degrees="78.07705" direction="west" orientation="longitude" pageId="1" pageNumber="20" precision="1" value="-78.07705">78.07705 W</geoCoordinate>
,
<quantity id="4CF3FBC2F016FF8CFC557BD3FBEB9A1F" box="[967,1055,747,775]" metricMagnitude="3" metricUnit="m" metricValue="2.716" pageId="1" pageNumber="20" unit="m" value="2716.0">2716 m</quantity>
a.s.l.),
<date id="FFB570E7F016FF8CFBE17BD3FABF9A1F" box="[1139,1355,747,775]" pageId="1" pageNumber="20" value="2013-11-05">5 November 2013</date>
, K. J. Ten- nessen leg. Deposited in Florida State Collection of Ar- thropods.
</paragraph>
<caption id="DF7406AFF016FF8CFFEB711BFD1B9127" ID-DOI="http://doi.org/10.5281/zenodo.15555036" ID-Zenodo-Dep="15555036" box="[121,751,2083,2111]" httpUri="https://zenodo.org/record/15555036/files/figure.png" pageId="1" pageNumber="20" startId="1.[121,189,2083,2108]" targetBox="[127,1458,1007,2045]" targetPageId="1" targetType="figure">
<paragraph id="8BB45627F016FF8CFFEB711BFD1B9127" blockId="1.[121,751,2083,2111]" box="[121,751,2083,2111]" pageId="1" pageNumber="20">
<emphasis id="B97F8A35F016FF8CFFEB711BFF2C9124" bold="true" box="[121,216,2083,2108]" pageId="1" pageNumber="20">Figure 1.</emphasis>
<taxonomicName id="4C0B2DA4F016FF8CFF4C711EFE0A9126" authorityName="Tennessen" authorityYear="2024" box="[222,510,2086,2110]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="signata" status="sp. nov.">
<emphasis id="B97F8A35F016FF8CFF4C711EFE0A9126" box="[222,510,2086,2110]" italics="true" pageId="1" pageNumber="20">Gomphomacromia signata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C374EF016FF8CFD96711EFDB59127" box="[516,577,2086,2111]" pageId="1" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
, holotype male.
</paragraph>
</caption>
<caption id="DF7406AFF015FF8FFFEB7D29FA689D35" ID-DOI="http://doi.org/10.5281/zenodo.15555038" ID-Zenodo-Dep="15555038" box="[121,1436,1041,1069]" httpUri="https://zenodo.org/record/15555038/files/figure.png" pageId="2" pageNumber="21" startId="2.[121,189,1041,1066]" targetBox="[283,1304,144,1001]" targetPageId="2" targetType="figure">
<paragraph id="8BB45627F015FF8FFFEB7D29FA689D35" blockId="2.[121,1436,1041,1069]" box="[121,1436,1041,1069]" pageId="2" pageNumber="21">
<emphasis id="B97F8A35F015FF8FFFEB7D29FF2C9D32" bold="true" box="[121,216,1041,1066]" pageId="2" pageNumber="21">Figure 2.</emphasis>
<taxonomicName id="4C0B2DA4F015FF8FFF4C7D2CFE0A9D34" authorityName="Tennessen" authorityYear="2024" box="[222,510,1044,1068]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="signata" status="sp. nov.">
<emphasis id="B97F8A35F015FF8FFF4C7D2CFE0A9D34" box="[222,510,1044,1068]" italics="true" pageId="2" pageNumber="21">Gomphomacromia signata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C374EF015FF8FFD967D2CFDB59D35" box="[516,577,1044,1069]" pageId="2" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, showing living eye color and postfrons pattern in anterior view.
</paragraph>
</caption>
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<paragraph id="8BB45627F015FF8FFFEB711AFBBB9126" blockId="2.[121,1103,2082,2110]" box="[121,1103,2082,2110]" pageId="2" pageNumber="21">
<emphasis id="B97F8A35F015FF8FFFEB711AFF2C9123" bold="true" box="[121,216,2082,2107]" pageId="2" pageNumber="21">Figure 3.</emphasis>
<taxonomicName id="4C0B2DA4F015FF8FFF4C711DFE0A9125" authorityName="Tennessen" authorityYear="2024" box="[222,510,2085,2109]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="signata" status="sp. nov.">
<emphasis id="B97F8A35F015FF8FFF4C711DFE0A9125" box="[222,510,2085,2109]" italics="true" pageId="2" pageNumber="21">Gomphomacromia signata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C374EF015FF8FFD96711DFDB59126" box="[516,577,2085,2110]" pageId="2" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
, holotype male accessory genitalia, ventral view.
</paragraph>
</caption>
<paragraph id="8BB45627F014FF8EFFEB79ABFE3A99B6" blockId="3.[121,462,147,174]" box="[121,462,147,174]" pageId="3" pageNumber="22">
<emphasis id="B97F8A35F014FF8EFFEB79ABFE3A99B6" box="[121,462,147,174]" italics="true" pageId="3" pageNumber="22">
Description of
<typeStatus id="54B0E885F014FF8EFEB679ABFE7899B6" box="[292,396,147,174]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="8BB45627F014FF8EFFEB79EEFCF3980B" blockId="3.[121,775,214,275]" pageId="3" pageNumber="22">
A dark brown medium-sized dragonfly with yellow markings on the postfrons, thorax and abdomen (
<figureCitation id="13304AA2F014FF8EFD2A79CFFD01980B" box="[696,757,247,275]" captionStart="Figure 1" captionStartId="1.[121,189,2083,2108]" captionTargetBox="[127,1458,1007,2045]" captionTargetId="figure-328@1.[121,1467,989,2060]" captionTargetPageId="1" captionText="Figure 1. Gomphomacromia signata sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.15555036" httpUri="https://zenodo.org/record/15555036/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BB45627F014FF8EFFEB7802FE3698E4" blockId="3.[121,776,314,508]" pageId="3" pageNumber="22">
Head. Eyes in life green dorsally, brown anteriorly and laterally, pale gray ventrally (
<figureCitation id="13304AA2F014FF8EFE587863FDFA986F" box="[458,526,347,375]" captionStart="Figure 2" captionStartId="2.[121,189,1041,1066]" captionTargetBox="[283,1304,144,1001]" captionTargetId="figure-24@2.[283,1304,144,1001]" captionTargetPageId="2" captionText="Figure 2. Gomphomacromia signata sp. n., holotype male, showing living eye color and postfrons pattern in anterior view." figureDoi="http://doi.org/10.5281/zenodo.15555038" httpUri="https://zenodo.org/record/15555038/files/figure.png" pageId="3" pageNumber="22">Fig. 2</figureCitation>
); labrum orange-tan, clypeus and antefrons dark brown, postfrons mostly yel- low with brown medially, vertex brown; labium mostly tan, darker distally; mandibles and maxillae orange-tan; rear of head orange-brown.
</paragraph>
<paragraph id="8BB45627F014FF8EFFEB7B1BFD169D4C" blockId="3.[121,776,547,1108]" pageId="3" pageNumber="22">
Thorax. Total surface closely beset with long dark setae. Prothorax brown, middle lobe light brown. Pterothorax: middorsal carina narrowly yellow, mesepisternum dark brown but with a small, narrow anterior and posterior yellow spot (
<figureCitation id="13304AA2F014FF8EFE847B90FEAA9BDC" box="[278,350,680,708]" captionStart="Figure 2" captionStartId="2.[121,189,1041,1066]" captionTargetBox="[283,1304,144,1001]" captionTargetId="figure-24@2.[283,1304,144,1001]" captionTargetPageId="2" captionText="Figure 2. Gomphomacromia signata sp. n., holotype male, showing living eye color and postfrons pattern in anterior view." figureDoi="http://doi.org/10.5281/zenodo.15555038" httpUri="https://zenodo.org/record/15555038/files/figure.png" pageId="3" pageNumber="22">Fig. 2</figureCitation>
); mesepimeron dark brown with a large oval yellow spot on anteroventral (lower) portion; metepisternum brown with a wide yellow stripe, narrow ventrally, widening dorsally and forked at upper end, the dorsal prong short, blunt and extending upward onto mesepisternum; metepimeron brown with a yel- low rectangular stripe with an upper extension (
<figureCitation id="13304AA2F014FF8EFD277A48FD009A94" box="[693,756,880,908]" captionStart="Figure 1" captionStartId="1.[121,189,2083,2108]" captionTargetBox="[127,1458,1007,2045]" captionTargetId="figure-328@1.[121,1467,989,2060]" captionTargetPageId="1" captionText="Figure 1. Gomphomacromia signata sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.15555036" httpUri="https://zenodo.org/record/15555036/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
); mesinfraepisternum and metinfraepimeron brown; venter of pterothorax gray-brown. Legs varying shades of medium to dark brown, coxae partly yellow, femora yellow at bases, darker apically; keel on prothoracic tibia extending along distal 0.57 of tibia; metathoracic femur
<quantity id="4CF3FBC2F014FF8EFF557D00FED59D4C" box="[199,289,1080,1108]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="22" unit="mm" value="4.7">4.7 mm</quantity>
long, metathoracic tibia
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long.
</paragraph>
<paragraph id="8BB45627F014FF8EFFEB7D43FD769C04" blockId="3.[121,775,1147,1308]" pageId="3" pageNumber="22">
Wings. Hyaline, venation brown; dark brown spot in basal cells between costa,subcosta and RA;yellow wash in subcostal space (
<figureCitation id="13304AA2F014FF8EFEF07D86FE569DC2" box="[354,418,1214,1242]" captionStart="Figure 1" captionStartId="1.[121,189,2083,2108]" captionTargetBox="[127,1458,1007,2045]" captionTargetId="figure-328@1.[121,1467,989,2060]" captionTargetPageId="1" captionText="Figure 1. Gomphomacromia signata sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.15555036" httpUri="https://zenodo.org/record/15555036/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
); triangles free; hw anal loop 5-celled; fw Ax 9, hw Ax 6; fw Px 8, hw Px 9; pterostigma orange-brown, surmounting 1.6 to 1.9 cells.
</paragraph>
<paragraph id="8BB45627F014FF8EFFEB7C7BFAC3984D" blockId="3.[121,776,1347,2108]" lastBlockId="3.[813,1466,146,341]" pageId="3" pageNumber="22">
Abdomen. Color pattern dark brown, paler laterally, dorsum with yellow spots: S1 with a posteromedial yel- low spot; S2 with a pair of anterolateral yellow spots and another pair of submedial yellow spots just past midlength; S37 with a pair of submedial yellow spots divided by dark transverse carina, anterior part of yel- low spot larger than posterior part, the spots largest on S3 &amp; S4, diminished in size on S5 &amp; S6, very small on S7; S8 with pair of large, round submedial yellow spots; S9 &amp; 10 dark brown (
<figureCitation id="13304AA2F014FF8EFE137F57FE369F93" box="[385,450,1647,1675]" captionStart="Figure 1" captionStartId="1.[121,189,2083,2108]" captionTargetBox="[127,1458,1007,2045]" captionTargetId="figure-328@1.[121,1467,989,2060]" captionTargetPageId="1" captionText="Figure 1. Gomphomacromia signata sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.15555036" httpUri="https://zenodo.org/record/15555036/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
); posteroventral corner of S1 tergum broadly curved, lacking denticulate process; auricles brown, prominent, 68 black denticles on pos- terior margin; S10 deeper than S
<quantity id="4CF3FBC2F014FF8EFD997FEBFDC39FF7" box="[523,567,1747,1775]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="22" unit="in" value="9.0">9 in</quantity>
lateral view (
<figureCitation id="13304AA2F014FF8EFD4E7FEBFF789E08" captionStart="Figure 46" captionStartId="3.[813,880,2018,2043]" captionTargetBox="[814,1417,1652,1980]" captionTargetId="figure-652@3.[810,1450,1621,1990]" captionTargetPageId="3" captionText="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi="http://doi.org/10.5281/zenodo.15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22">Fig. 5</figureCitation>
). Anterior hamule laminate with a square notch; pos- terior hamule in ventral view slightly curved, a very short, blunt process at midlength directed medially and a longer, darker, rounded, denticulate distal lobe
<quantity id="4CF3FBC2F014FF8EFD417E60FF5C9E8E" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="22" unit="mm" value="0.45">0.45 mm</quantity>
long (
<figureCitation id="13304AA2F014FF8EFF7D7E42FECC9E8E" box="[239,312,1914,1942]" captionStart="Figure 3" captionStartId="2.[121,189,2082,2107]" captionTargetBox="[283,1303,1142,2042]" captionTargetId="figure-44@2.[283,1304,1135,2045]" captionTargetPageId="2" captionText="Figure 3. Gomphomacromia signata sp. n., holotype male accessory genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.15555040" httpUri="https://zenodo.org/record/15555040/files/figure.png" pageId="3" pageNumber="22">Figs 3</figureCitation>
,
<figureCitation id="13304AA2F014FF8EFED17E42FEA19E8E" box="[323,341,1914,1942]" captionStart="Figure 9" captionStartId="5.[121,189,2054,2079]" captionTargetBox="[123,770,1388,2002]" captionTargetId="figure-186@5.[118,775,1648,1765]" captionTargetPageId="5" captionText="Figure 9. Lef posterior hamule of Gomphomacromia signata sp. n. and G. fallax, mediolateral view." figureDoi="http://doi.org/10.5281/zenodo.15555048" httpUri="https://zenodo.org/record/15555048/files/figure.png" pageId="3" pageNumber="22">9</figureCitation>
). Anal appendages dark brown, with many dark setae; cercus in dorsal view nearly straight, apical fourth angled inward so that the pair converges distally, tips narrowed (
<figureCitation id="13304AA2F014FF8EFE0B7EE6FE169EE2" box="[409,482,2014,2042]" captionStart="Figure 46" captionStartId="3.[813,880,2018,2043]" captionTargetBox="[814,1417,1652,1980]" captionTargetId="figure-652@3.[810,1450,1621,1990]" captionTargetPageId="3" captionText="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi="http://doi.org/10.5281/zenodo.15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22">Fig. 4</figureCitation>
), in lateral view slightly arched, ventrolateral margin with a minute sharplytipped denticle at 3/10 length (
<figureCitation id="13304AA2F014FF8EFE427118FDE29124" box="[464,534,2080,2108]" captionStart="Figure 46" captionStartId="3.[813,880,2018,2043]" captionTargetBox="[814,1417,1652,1980]" captionTargetId="figure-652@3.[810,1450,1621,1990]" captionTargetPageId="3" captionText="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi="http://doi.org/10.5281/zenodo.15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22">Fig. 5</figureCitation>
); epiproct about 0.7 times length of cerci, in dorsal view of the appendages with tips visible beyond lateral margins of cerci (
<figureCitation id="13304AA2F014FF8EFAE2798CFA4799C8" box="[1392,1459,180,208]" captionStart="Figure 46" captionStartId="3.[813,880,2018,2043]" captionTargetBox="[814,1417,1652,1980]" captionTargetId="figure-652@3.[810,1450,1621,1990]" captionTargetPageId="3" captionText="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi="http://doi.org/10.5281/zenodo.15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22">Fig. 4</figureCitation>
) in normal repose, in ventral view as wide distally as ba- sally, with lateral margins slightly incurved, distal mar- gin sinuous, tips protruding posterolaterally (
<figureCitation id="13304AA2F014FF8EFADB7820FA7B982C" box="[1353,1423,280,308]" captionStart="Figure 46" captionStartId="3.[813,880,2018,2043]" captionTargetBox="[814,1417,1652,1980]" captionTargetId="figure-652@3.[810,1450,1621,1990]" captionTargetPageId="3" captionText="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi="http://doi.org/10.5281/zenodo.15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22">Fig. 6</figureCitation>
), in lateral view tips recurved, tooth-like (
<figureCitation id="13304AA2F014FF8EFB757801FADF984D" box="[1255,1323,313,341]" captionStart="Figure 46" captionStartId="3.[813,880,2018,2043]" captionTargetBox="[814,1417,1652,1980]" captionTargetId="figure-652@3.[810,1450,1621,1990]" captionTargetPageId="3" captionText="Figure 46. Gomphomacromia signata sp. n., holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus." figureDoi="http://doi.org/10.5281/zenodo.15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BB45627F014FF8EFCBF7844FBE198C2" blockId="3.[813,1466,380,474]" pageId="3" pageNumber="22">Measurements (mm). Total L 39.4, head W 5.3, hind wing L 26.3, abdomen L (excluding cercus) 26.9, cercus L 1.7, pt L 1.671.8.</paragraph>
<paragraph id="8BB45627F014FF8EFCBF7B39FC779B04" blockId="3.[813,899,513,540]" box="[813,899,513,540]" pageId="3" pageNumber="22">
<emphasis id="B97F8A35F014FF8EFCBF7B39FC779B04" box="[813,899,513,540]" italics="true" pageId="3" pageNumber="22">Female</emphasis>
</paragraph>
<paragraph id="8BB45627F014FF8EFCBF7B7CFA4E9A1E" blockId="3.[813,1467,580,774]" pageId="3" pageNumber="22">
A female of the new species was photographed by Cary Kerst at the same location and on the same day that I collected the male (
<figureCitation id="13304AA2F014FF8EFBA27BBEFB839BBA" box="[1072,1143,646,674]" captionStart="Figure 7" captionStartId="4.[121,189,2080,2105]" captionTargetBox="[121,1467,801,2041]" captionTargetId="figure-25@4.[121,1467,798,2044]" captionTargetPageId="4" captionText="Figure 7. Gomphomacromia signata sp. n., female, in situ. Photo: Cary Kerst." figureDoi="http://doi.org/10.5281/zenodo.15555044" httpUri="https://zenodo.org/record/15555044/files/figure.png" pageId="3" pageNumber="22">Fig. 7</figureCitation>
); it was not collected. The image shows that the color pattern is similar to that of the male
<typeStatus id="54B0E885F014FF8EFC337BF1FBFB9BFD" box="[929,1039,713,741]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
, although the eyes have only slight greenish reflections dorsally, the yellow spots on S27
</paragraph>
<paragraph id="8BB45627F014FF8EFA357F55FA4F9F8B" blockId="3.[1447,1467,1645,1683]" box="[1447,1467,1645,1683]" pageId="3" pageNumber="22">
<emphasis id="B97F8A35F014FF8EFA357F55FA4F9F8B" bold="true" box="[1447,1467,1645,1683]" pageId="3" pageNumber="22">6</emphasis>
</paragraph>
<caption id="DF7406AFF014FF8EFCBF7EDAFA669123" ID-DOI="http://doi.org/10.5281/zenodo.15555042" ID-Zenodo-Dep="15555042" httpUri="https://zenodo.org/record/15555042/files/figure.png" pageId="3" pageNumber="22" startId="3.[813,880,2018,2043]" targetBox="[814,1417,1652,1980]" targetPageId="3" targetType="figure">
<paragraph id="8BB45627F014FF8EFCBF7EDAFA669123" blockId="3.[812,1468,2018,2107]" pageId="3" pageNumber="22">
<emphasis id="B97F8A35F014FF8EFCBF7EDAFC529EE3" bold="true" box="[813,934,2018,2043]" pageId="3" pageNumber="22">Figure 46.</emphasis>
<taxonomicName id="4C0B2DA4F014FF8EFC3F7EDDFB3D9EE5" authorityName="Tennessen" authorityYear="2024" box="[941,1225,2021,2045]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="signata" status="sp. nov.">
<emphasis id="B97F8A35F014FF8EFC3F7EDDFB3D9EE5" box="[941,1225,2021,2045]" italics="true" pageId="3" pageNumber="22">Gomphomacromia signata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C374EF014FF8EFB437EDDFAF99EE6" box="[1233,1293,2021,2046]" pageId="3" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, anal appendages in dorsal, lateral and ventral views; arrow in Figure 5 indicates minute ventrolateral denticle on cercus.
</paragraph>
</caption>
<paragraph id="8BB45627F013FF89FFEB79ABFD2C98E4" blockId="4.[121,775,147,508]" pageId="4" pageNumber="23">
are larger and S8 is totally brown. The yellow spots on the postfrons and large yellow spot on the lower mesepimeron are evident in the photograph (females of
<taxonomicName id="4C0B2DA4F013FF89FFEB79CFFF2B980A" authorityName="McLachlan" authorityYear="1881" box="[121,223,247,274]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FFEB79CFFF2B980A" box="[121,223,247,274]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
have a completely dark postfrons and mesepimeron, and the paired yellow spots on the abdomen are smaller than in
<taxonomicName id="4C0B2DA4F013FF89FECC7802FE23984D" authorityName="Tennessen" authorityYear="2024" box="[350,471,314,341]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FECC7802FE23984D" box="[350,471,314,341]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
). The tip of the abdomen of
<taxonomicName id="4C0B2DA4F013FF89FF0B7863FEE6986E" authorityName="Tennessen" authorityYear="2024" box="[153,274,347,374]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FF0B7863FEE6986E" box="[153,274,347,374]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
as shown in dorsolateral view in the pho- tograph (
<figureCitation id="13304AA2F013FF89FF777844FEDE9880" box="[229,298,380,408]" captionStart="Figure 7" captionStartId="4.[121,189,2080,2105]" captionTargetBox="[121,1467,801,2041]" captionTargetId="figure-25@4.[121,1467,798,2044]" captionTargetPageId="4" captionText="Figure 7. Gomphomacromia signata sp. n., female, in situ. Photo: Cary Kerst." figureDoi="http://doi.org/10.5281/zenodo.15555044" httpUri="https://zenodo.org/record/15555044/files/figure.png" pageId="4" pageNumber="23">Fig. 7</figureCitation>
), does not allow a definitive view of the vulvar lamina. However, the lobes of the vulvar lamina appear to be short, possibly much shorter than in
<emphasis id="B97F8A35F013FF89FD527887FF6F98E3" italics="true" pageId="4" pageNumber="23">G. fal- lax</emphasis>
(see
<figureCitation id="13304AA2F013FF89FF4878D8FED598E4" box="[218,289,480,508]" captionStart="Figure 2" captionStartId="2.[121,189,1041,1066]" captionTargetBox="[283,1304,144,1001]" captionTargetId="figure-24@2.[283,1304,144,1001]" captionTargetPageId="2" captionText="Figure 2. Gomphomacromia signata sp. n., holotype male, showing living eye color and postfrons pattern in anterior view." figureDoi="http://doi.org/10.5281/zenodo.15555038" httpUri="https://zenodo.org/record/15555038/files/figure.png" pageId="4" pageNumber="23">Fig. 2i</figureCitation>
in
<bibRefCitation id="EF9A2BD6F013FF89FED178D8FD3C98E4" author="von Ellenrieder, N. &amp; Garrison, R. W." box="[323,712,480,508]" pageId="4" pageNumber="23" pagination="83 - 98" refId="ref3528" refString="von Ellenrieder, N. &amp; Garrison, R. W. (2005). A synopsis of the South American genus Gomphomacromia (Odonata: Libellulidae). International Journal of Odonatology, 8 (1), 83-98. doi: 10.1080/1 3887890.2005. 9748245" type="journal article" year="2005">von Ellenrieder &amp; Garrison, 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8BB45627F013FF89FFEB7B1BFF189B26" blockId="4.[121,236,547,574]" box="[121,236,547,574]" pageId="4" pageNumber="23">
<emphasis id="B97F8A35F013FF89FFEB7B1BFF189B26" box="[121,236,547,574]" italics="true" pageId="4" pageNumber="23">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB45627F013FF89FFEB7B5EFBD8982C" blockId="4.[121,775,614,742]" lastBlockId="4.[813,1467,147,742]" pageId="4" pageNumber="23">
<taxonomicName id="4C0B2DA4F013FF89FFEB7B5EFE6A9B99" authorityName="McLachlan" authorityYear="1881" box="[121,414,614,641]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FFEB7B5EFE6A9B99" box="[121,414,614,641]" italics="true" pageId="4" pageNumber="23">Gomphomacromia fallax</emphasis>
</taxonomicName>
is a darker species than
<emphasis id="B97F8A35F013FF89FD2F7B5EFF449BBA" italics="true" pageId="4" pageNumber="23">G. sig- nata</emphasis>
, lacking yellow on the postfrons and having less yellow on the thorax and abdomen. In particular, the thoracic pattern of
<taxonomicName id="4C0B2DA4F013FF89FECB7BF2FE4E9BFD" authorityName="McLachlan" authorityYear="1881" box="[345,442,714,741]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FECB7BF2FE4E9BFD" box="[345,442,714,741]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
differs in lacking an oval yel- low spot on the lower part of the mesepimeron and the metepisternal and metepimeral stripes are unforked. The abdominal pattern of
<taxonomicName id="4C0B2DA4F013FF89FBFB79EEFB3999E9" authorityName="McLachlan" authorityYear="1881" box="[1129,1229,214,241]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FBFB79EEFB3999E9" box="[1129,1229,214,241]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C0B2DA4F013FF89FCBF79CFFC5F980A" authorityName="Tennessen" authorityYear="2024" box="[813,939,247,274]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FCBF79CFFC5F980A" box="[813,939,247,274]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
, although the submedial yellow spots are much smaller (
<figureCitation id="13304AA2F013FF89FC497820FBD4982C" box="[987,1056,280,308]" captionStart="Figure 8" captionStartId="5.[121,189,1299,1324]" captionTargetBox="[121,1467,144,1261]" captionTargetId="figure-210@5.[121,1467,141,1261]" captionTargetPageId="5" captionText="Figure 8. Gomphomacromia fallax, male, Imbabura Province, Ecuador. Photo: KJT." figureDoi="http://doi.org/10.5281/zenodo.15555046" httpUri="https://zenodo.org/record/15555046/files/figure.png" pageId="4" pageNumber="23">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BB45627F013FF89FCD97802FBAE98C3" blockId="4.[813,1467,147,742]" pageId="4" pageNumber="23">
In
<taxonomicName id="4C0B2DA4F013FF89FCF87802FC10984D" authorityName="Tennessen" authorityYear="2024" box="[874,996,314,341]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FCF87802FC10984D" box="[874,996,314,341]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
, small dark brown spots are present in the base of all four wings and there is a yellow wash in the rows of cells between the costa and vein RA (
<figureCitation id="13304AA2F013FF89FAF87844FA5C9880" box="[1386,1448,380,408]" captionStart="Figure 1" captionStartId="1.[121,189,2083,2108]" captionTargetBox="[127,1458,1007,2045]" captionTargetId="figure-328@1.[121,1467,989,2060]" captionTargetPageId="1" captionText="Figure 1. Gomphomacromia signata sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.15555036" httpUri="https://zenodo.org/record/15555036/files/figure.png" pageId="4" pageNumber="23">Fig. 1</figureCitation>
); the wings bases and anterior rows of cells in males of
<taxonomicName id="4C0B2DA4F013FF89FCBF7887FC7A98C2" authorityName="McLachlan" authorityYear="1881" box="[813,910,447,474]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FCBF7887FC7A98C2" box="[813,910,447,474]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
are clear (
<figureCitation id="13304AA2F013FF89FB987887FBBA98C3" box="[1034,1102,447,475]" captionStart="Figure 8" captionStartId="5.[121,189,1299,1324]" captionTargetBox="[121,1467,144,1261]" captionTargetId="figure-210@5.[121,1467,141,1261]" captionTargetPageId="5" captionText="Figure 8. Gomphomacromia fallax, male, Imbabura Province, Ecuador. Photo: KJT." figureDoi="http://doi.org/10.5281/zenodo.15555046" httpUri="https://zenodo.org/record/15555046/files/figure.png" pageId="4" pageNumber="23">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BB45627F013FF88FCD978D8FC3C9CD0" blockId="4.[813,1467,147,742]" lastBlockId="5.[813,1467,1385,1680]" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
The distal lobe of the posterior hamule is longer in
<taxonomicName id="4C0B2DA4F013FF89FCDF7B3AFC3D9B05" authorityName="Tennessen" authorityYear="2024" box="[845,969,514,541]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FCDF7B3AFC3D9B05" box="[845,969,514,541]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
(
<quantity id="4CF3FBC2F013FF89FC4B7B3AFBBE9B06" box="[985,1098,514,542]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="23" unit="mm" value="0.45">0.45 mm</quantity>
) compared to
<taxonomicName id="4C0B2DA4F013FF89FA927B3AFA919B05" authorityName="McLachlan" authorityYear="1881" box="[1280,1381,514,541]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FA927B3AFA919B05" box="[1280,1381,514,541]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
(
<quantity id="4CF3FBC2F013FF89FAE47B3AFC689B27" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="4" pageNumber="23" unit="mm" value="0.285" valueMax="0.32" valueMin="0.25">0.25 0.32 mm</quantity>
, n = 9 from
<collectingCountry id="F31C16B7F013FF89FBBC7B1BFB7B9B27" box="[1070,1167,547,575]" name="Ecuador" pageId="4" pageNumber="23">Ecuador</collectingCountry>
and
<collectingCountry id="F31C16B7F013FF89FB5E7B1BFAE99B27" box="[1228,1309,547,575]" name="Bolivia" pageId="4" pageNumber="23">Bolivia</collectingCountry>
) (
<figureCitation id="13304AA2F013FF89FAA17B1BFA8D9B27" box="[1331,1401,547,575]" captionStart="Figure 9" captionStartId="5.[121,189,2054,2079]" captionTargetBox="[123,770,1388,2002]" captionTargetId="figure-186@5.[118,775,1648,1765]" captionTargetPageId="5" captionText="Figure 9. Lef posterior hamule of Gomphomacromia signata sp. n. and G. fallax, mediolateral view." figureDoi="http://doi.org/10.5281/zenodo.15555048" httpUri="https://zenodo.org/record/15555048/files/figure.png" pageId="4" pageNumber="23">Fig. 9</figureCitation>
). The anterior hamule of the
<taxonomicName id="4C0B2DA4F013FF89FBAE7B7CFB479B47" authorityName="Tennessen" authorityYear="2024" box="[1084,1203,580,607]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FBAE7B7CFB479B47" box="[1084,1203,580,607]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
<typeStatus id="54B0E885F013FF89FB2A7B7CFAD69B78" box="[1208,1314,580,608]" pageId="4" pageNumber="23" type="holotype">holotype</typeStatus>
has a square notch whereas the notch in
<taxonomicName id="4C0B2DA4F013FF89FB1F7B5EFB079B99" authorityName="McLachlan" authorityYear="1881" box="[1165,1267,614,641]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FB1F7B5EFB079B99" box="[1165,1267,614,641]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
is usually much less conspicuous (
<figureCitation id="13304AA2F013FF89FB937BBFFBA19BBB" box="[1025,1109,647,675]" captionStart="Figure 10" captionStartId="6.[121,189,958,983]" captionTargetBox="[122,770,151,917]" captionTargetId="figure-302@6.[121,775,141,923]" captionTargetPageId="6" captionText="Figure 10. Lamina of anterior hamule of Gomphomacromia signata sp. n. and G. fallax, lateral view at slight anterior angle." figureDoi="http://doi.org/10.5281/zenodo.15555050" httpUri="https://zenodo.org/record/15555050/files/figure.png" pageId="4" pageNumber="23">Fig. 10</figureCitation>
), but at least
<specimenCount id="9D0D9DAEF013FF89FB677BBFFA6F9BBB" box="[1269,1435,647,675]" count="1" pageId="4" pageNumber="23" type="generic">one specimen</specimenCount>
of
<taxonomicName id="4C0B2DA4F013FF89FCBF7B90FC799BDB" authorityName="McLachlan" authorityYear="1881" box="[813,909,680,707]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F013FF89FCBF7B90FC799BDB" box="[813,909,680,707]" italics="true" pageId="4" pageNumber="23">G. fallax</emphasis>
</taxonomicName>
from
<collectingCountry id="F31C16B7F013FF89FC437B90FBC69BDC" box="[977,1074,680,708]" name="Ecuador" pageId="4" pageNumber="23">Ecuador</collectingCountry>
had a square notch. The epiproct is rectangular in
<taxonomicName id="4C0B2DA4F013FF89FC657BF2FB869BFD" authorityName="Tennessen" authorityYear="2024" box="[1015,1138,714,741]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F013FF89FC657BF2FB869BFD" box="[1015,1138,714,741]" italics="true" pageId="4" pageNumber="23">G. signata</emphasis>
</taxonomicName>
compared to elongate and tapered in
<taxonomicName id="4C0B2DA4F012FF88FC2A7C51FBEB9C9C" authorityName="McLachlan" authorityYear="1881" box="[952,1055,1385,1412]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F012FF88FC2A7C51FBEB9C9C" box="[952,1055,1385,1412]" italics="true" pageId="5" pageNumber="24">G. fallax</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9A2BD6F012FF88FBFA7C51FC9C9CBF" author="von Ellenrieder, N. &amp; Garrison, R. W." pageId="5" pageNumber="24" pagination="83 - 98" refId="ref3528" refString="von Ellenrieder, N. &amp; Garrison, R. W. (2005). A synopsis of the South American genus Gomphomacromia (Odonata: Libellulidae). International Journal of Odonatology, 8 (1), 83-98. doi: 10.1080/1 3887890.2005. 9748245" type="journal article" year="2005">von Ellenrieder &amp; Garrison, 2005</bibRefCitation>
). Epiproct shape in
<taxonomicName id="4C0B2DA4F012FF88FBCD7CB3FB2E9CBE" authorityName="Tennessen" authorityYear="2024" box="[1119,1242,1419,1446]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F012FF88FBCD7CB3FB2E9CBE" box="[1119,1242,1419,1446]" italics="true" pageId="5" pageNumber="24">G. signata</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="4C0B2DA4F012FF88FCBF7C94FC359CDF" authorityName="Ris" authorityYear="1928" box="[813,961,1452,1479]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="nodisticta">
<emphasis id="B97F8A35F012FF88FCBF7C94FC359CDF" box="[813,961,1452,1479]" italics="true" pageId="5" pageNumber="24">G. nodisticta</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF7406AFF013FF89FFEB7118FC469123" ID-DOI="http://doi.org/10.5281/zenodo.15555044" ID-Zenodo-Dep="15555044" box="[121,946,2080,2107]" httpUri="https://zenodo.org/record/15555044/files/figure.png" pageId="4" pageNumber="23" startId="4.[121,189,2080,2105]" targetBox="[121,1467,801,2041]" targetPageId="4" targetType="figure">
<paragraph id="8BB45627F013FF89FFEB7118FC469123" blockId="4.[121,946,2080,2107]" box="[121,946,2080,2107]" pageId="4" pageNumber="23">
<emphasis id="B97F8A35F013FF89FFEB7118FF2C9121" bold="true" box="[121,216,2080,2105]" pageId="4" pageNumber="23">Figure 7.</emphasis>
<taxonomicName id="4C0B2DA4F013FF89FF4C711AFE0A9122" authorityName="Tennessen" authorityYear="2024" box="[222,510,2082,2106]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="signata" status="sp. nov.">
<emphasis id="B97F8A35F013FF89FF4C711AFE0A9122" box="[222,510,2082,2106]" italics="true" pageId="4" pageNumber="23">Gomphomacromia signata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C374EF013FF89FD96711AFDB59123" box="[516,577,2082,2107]" pageId="4" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
, female,
<emphasis id="B97F8A35F013FF89FD30711AFD179122" box="[674,739,2082,2106]" italics="true" pageId="4" pageNumber="23">in situ</emphasis>
. Photo: Cary Kerst.
</paragraph>
</caption>
<caption id="DF7406AFF012FF88FFEB7C2BFC1E9C36" ID-DOI="http://doi.org/10.5281/zenodo.15555046" ID-Zenodo-Dep="15555046" box="[121,1002,1299,1326]" httpUri="https://zenodo.org/record/15555046/files/figure.png" pageId="5" pageNumber="24" startId="5.[121,189,1299,1324]" targetBox="[121,1467,144,1261]" targetPageId="5" targetType="figure">
<paragraph id="8BB45627F012FF88FFEB7C2BFC1E9C36" blockId="5.[121,1002,1299,1326]" box="[121,1002,1299,1326]" pageId="5" pageNumber="24">
<emphasis id="B97F8A35F012FF88FFEB7C2BFF2C9C34" bold="true" box="[121,216,1299,1324]" pageId="5" pageNumber="24">Figure 8.</emphasis>
<taxonomicName id="4C0B2DA4F012FF88FF4C7C2DFE1D9C35" authorityName="McLachlan" authorityYear="1881" box="[222,489,1301,1325]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F012FF88FF4C7C2DFE1D9C35" box="[222,489,1301,1325]" italics="true" pageId="5" pageNumber="24">Gomphomacromia fallax</emphasis>
</taxonomicName>
, male, Imbabura Province, Ecuador. Photo: KJT.
</paragraph>
</caption>
<paragraph id="8BB45627F012FF88FCD97CF5FC359F88" blockId="5.[813,1467,1385,1680]" pageId="5" pageNumber="24">
The length of the keel on the prothoracic tibia ex- ceeds half the length of the tibia in
<taxonomicName id="4C0B2DA4F012FF88FB7C7CD7FA9F9F12" authorityName="Tennessen" authorityYear="2024" box="[1262,1387,1519,1546]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B97F8A35F012FF88FB7C7CD7FA9F9F12" box="[1262,1387,1519,1546]" italics="true" pageId="5" pageNumber="24">G. signata</emphasis>
</taxonomicName>
(0.57) and
<taxonomicName id="4C0B2DA4F012FF88FCF27F28FC369F33" authorityName="McLachlan" authorityYear="1881" box="[864,962,1552,1579]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B97F8A35F012FF88FCF27F28FC369F33" box="[864,962,1552,1579]" italics="true" pageId="5" pageNumber="24">G. fallax</emphasis>
</taxonomicName>
(0.500.66); the keel is shorter (0.330.50 length of tibia) in the southern species,
<taxonomicName id="4C0B2DA4F012FF88FAB87F09FA479F54" authorityName="Martin" authorityYear="1921" box="[1322,1459,1585,1612]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B97F8A35F012FF88FAB87F09FA479F54" box="[1322,1459,1585,1612]" italics="true" pageId="5" pageNumber="24">G. chilensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B2DA4F012FF88FCBF7F6BFC309F76" authorityName="Ris" authorityYear="1928" box="[813,964,1619,1646]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="nodisticta">
<emphasis id="B97F8A35F012FF88FCBF7F6BFC309F76" box="[813,964,1619,1646]" italics="true" pageId="5" pageNumber="24">G. nodisticta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B2DA4F012FF88FB967F6BFB639F76" authorityName="Brauer" authorityYear="1864" box="[1028,1175,1619,1646]" class="Insecta" family="Corduliidae" genus="Gomphomacromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B97F8A35F012FF88FB967F6BFB639F76" box="[1028,1175,1619,1646]" italics="true" pageId="5" pageNumber="24">G. paradoxa</emphasis>
</taxonomicName>
(von Ellenrieder &amp; Gar- rison, 2005).
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="03CEA72AFF930D1AEE367646117B993A" LSID="urn:lsid:plazi:treatment:03CEA72AFF930D1AEE367646117B993A" httpUri="http://treatment.plazi.org/id/03CEA72AFF930D1AEE367646117B993A" lastPageId="5" lastPageNumber="67" pageId="1" pageNumber="63">
<subSubSection id="C37D45B7FF930D1EEE36764617439874" box="[121,594,1344,1375]" pageId="1" pageNumber="63" type="nomenclature">
@ -52,7 +53,7 @@
</subSubSection>
<subSubSection id="C37D45B7FF930D1EEE36766115CB98A8" box="[121,218,1380,1408]" pageId="1" pageNumber="63" type="description">
<paragraph id="8BD8163CFF930D1EEE36766115CB98A8" blockId="1.[121,594,1344,1408]" box="[121,218,1380,1408]" pageId="1" pageNumber="63">
<figureCitation id="135C0AB9FF930D1EEE36766115CB98A8" box="[121,218,1380,1408]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="2.[121,189,2048,2073]" captionStartId-1="3.[121,189,2049,2074]" captionTargetBox-0="[154,1442,144,2010]" captionTargetBox-1="[121,1467,713,2012]" captionTargetId-0="figure-24@2.[143,1442,144,2010]" captionTargetId-1="figure-23@3.[121,1467,713,2012]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." captionText-1="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." pageId="1" pageNumber="63">Figs 12</figureCitation>
<figureCitation id="135C0AB9FF930D1EEE36766115CB98A8" box="[121,218,1380,1408]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="2.[121,189,2048,2073]" captionStartId-1="3.[121,189,2049,2074]" captionTargetBox-0="[154,1442,144,2010]" captionTargetBox-1="[121,1467,713,2012]" captionTargetId-0="figure-24@2.[143,1442,144,2010]" captionTargetId-1="figure-23@3.[121,1467,713,2012]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." captionText-1="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." figureDoi-0="http://doi.org/10.5281/zenodo.15554898" figureDoi-1="http://doi.org/10.5281/zenodo.15554900" httpUri-0="https://zenodo.org/record/15554898/files/figure.png" httpUri-1="https://zenodo.org/record/15554900/files/figure.png" pageId="1" pageNumber="63">Figs 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D45B7FF930D1AEE3676A217B698CD" lastPageId="5" lastPageNumber="67" pageId="1" pageNumber="63" type="materials_examined">
@ -96,18 +97,18 @@ leg.
</paragraph>
<paragraph id="8BD8163CFF930D1EEE367413116F9DF8" blockId="1.[121,776,1814,2108]" lastBlockId="1.[813,1466,147,208]" pageId="1" pageNumber="63">
Head (
<figureCitation id="135C0AB9FF930D1EEE877413140C9A1A" box="[200,285,1814,1842]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." pageId="1" pageNumber="63">Fig. 1a</figureCitation>
<figureCitation id="135C0AB9FF930D1EEE877413140C9A1A" box="[200,285,1814,1842]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="1" pageNumber="63">Fig. 1a</figureCitation>
). Eyes pale brown, darker dorsally. Face uniformly yellow,marked as follows: labium light brown- ish, labrum with medial and postclypeus with paired dark yellow markings, and frons slightly greenish. La- brum 2.5 times as wide as long, frons expanded. Ocellar region depressed and black between lateral ocelli and frons. First antennal segment black at the base, remain- der yellow; second antennal segment black at the base, remainder orange-yellow; filament black. Median ocelli wider than lateral ocelli. Vertex and occipital ridge yel- low, black at their boundary.
</paragraph>
<paragraph id="8BD8163CFF930D1EED6273F211D99FAA" blockId="1.[813,1467,247,775]" pageId="1" pageNumber="63">
Thorax (
<figureCitation id="135C0AB9FF930D1EEDC373F211159C3B" box="[908,1028,247,275]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." pageId="1" pageNumber="63">Figs 1b, d</figureCitation>
<figureCitation id="135C0AB9FF930D1EEDC373F211159C3B" box="[908,1028,247,275]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="1" pageNumber="63">Figs 1b, d</figureCitation>
). Prothorax yellow, with a distinctive black marking, widely expanded on anterior lobe, cen- ter of median lobe with paired yellow spots.Pterothorax yellow, marked with black: anterior portion of mesostig- ma, faint L-shaped markings along dorsal carina, two nail-shaped markings pointing forward on mesepister- num, thin stripe across upper margin of mesepimeron, junction of metepimeron and metepisternum faintly black. Antealar ridge conspicuous, black, and lined with small black spines, junction of dorsal carina forming a small, sharply pointed spine approximately midway be- tween wing base and mesostigma.
</paragraph>
<paragraph id="8BD8163CFF930D1EED04718210AA9E2F" blockId="1.[813,1467,247,775]" pageId="1" pageNumber="63">Legs black, marked with blackish brown: stripes across tibiae, posteriorly, two stripes on femur, fainter on metafemora. Covered with long, black spines across mid-axis; spines on femur small and sparse. Claws black.</paragraph>
<paragraph id="8BD8163CFF930D1EED62702B11A699D3" blockId="1.[813,1467,814,1275]" pageId="1" pageNumber="63">
Wings (
<figureCitation id="135C0AB9FF930D1EEDC6702B11149E62" box="[905,1029,814,842]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." pageId="1" pageNumber="63">Figs 1c, e</figureCitation>
<figureCitation id="135C0AB9FF930D1EEDC6702B11149E62" box="[905,1029,814,842]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="1" pageNumber="63">Figs 1c, e</figureCitation>
) hyaline, membrane extended and yellow, pterostigma and costal margin pale brown, pterostigmal brace present. Antenodals (1
<superScript id="7C12BB74FF930D1EEB6B706B10219E55" attach="left" box="[1316,1328,878,893]" fontSize="6" pageId="1" pageNumber="63">st</superScript>
and 4
@ -122,12 +123,12 @@ throughout, two rows in lef HW near margin.
</paragraph>
<paragraph id="8BD8163CFF930D1EED62762710869BA3" blockId="1.[812,1467,1314,1675]" pageId="1" pageNumber="63">
Abdomen (
<figureCitation id="135C0AB9FF930D1EEDE57627110A9816" box="[938,1051,1314,1342]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." pageId="1" pageNumber="63">Figs 1b, d</figureCitation>
<figureCitation id="135C0AB9FF930D1EEDE57627110A9816" box="[938,1051,1314,1342]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="1" pageNumber="63">Figs 1b, d</figureCitation>
). Ground color on S12 yellow, S3 to anterior half of S7 bluish-gray,yellow from there. Auricles small, yellow; covered with black spines. Marked black as follow: S17 with annules at posterior margin, short on S1, longitudinal paired black markings on S26: mark- ings connected to black rings on S36, faint black spots on S710: near posterior border on S78, at anterior margin on S910. Dorsal section of S10 extended, lateral margin excavated. S10 is expanded between branches of cerci; a hood-like lamina is present, covering downwardly curved, horn-like, paired structures with black apices.
</paragraph>
<paragraph id="8BD8163CFF930D1EED6275B716659AF0" blockId="1.[813,1467,1714,2008]" pageId="1" pageNumber="63">
Vesica spermalis (
<figureCitation id="135C0AB9FF930D1EEA4875B711909BE6" box="[1031,1153,1714,1742]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." pageId="1" pageNumber="63">Figs 4ac</figureCitation>
<figureCitation id="135C0AB9FF930D1EEA4875B711909BE6" box="[1031,1153,1714,1742]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." figureDoi="http://doi.org/10.5281/zenodo.15554904" httpUri="https://zenodo.org/record/15554904/files/figure.png" pageId="1" pageNumber="63">Figs 4ac</figureCitation>
). Hamuli yellow. Anterior hamuli thin, long, ending in a shoulder and anteriorly pointed hook, apical spine black. Posterior hamuli long, saddle-shaped, and covered with bristles; anterior mar- gin curved.
<collectionCode id="ED768EF9FF930D1EEDF6743216C19A7B" box="[953,976,1847,1875]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="63" type="Museum">V</collectionCode>
1 rounded, junction of
@ -142,10 +143,10 @@ Vesica spermalis (
</paragraph>
<paragraph id="8BD8163CFF930D1CED6274FA17C79C3B" blockId="1.[813,1467,2047,2108]" lastBlockId="3.[121,775,147,275]" lastPageId="3" lastPageNumber="65" pageId="1" pageNumber="63">
Caudal appendages (
<figureCitation id="135C0AB9FF930D1EEA7C74FA11BE9533" box="[1075,1199,2047,2075]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." pageId="1" pageNumber="63">Figs 4df</figureCitation>
<figureCitation id="135C0AB9FF930D1EEA7C74FA11BE9533" box="[1075,1199,2047,2075]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." figureDoi="http://doi.org/10.5281/zenodo.15554904" httpUri="https://zenodo.org/record/15554904/files/figure.png" pageId="1" pageNumber="63">Figs 4df</figureCitation>
). Cerci extended 1.6 times as far as epiproct. Cerci blade-shaped, darker on inner side, with a small basal spine, lower margin curved slightly inwards. Epiproct fused at base, widely diver- gent, appearing triangular in lateral view, upper margin ridge-like; ending in upwardly pointed black spines.
</paragraph>
<caption id="DF1846B4FF900D1DEE367B0517CE9512" pageId="2" pageNumber="64" startId="2.[121,189,2048,2073]" targetBox="[154,1442,144,2010]" targetPageId="2" targetType="figure">
<caption id="DF1846B4FF900D1DEE367B0517CE9512" ID-DOI="http://doi.org/10.5281/zenodo.15554898" ID-Zenodo-Dep="15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="2" pageNumber="64" startId="2.[121,189,2048,2073]" targetBox="[154,1442,144,2010]" targetPageId="2" targetType="figure">
<paragraph id="8BD8163CFF900D1DEE367B0517CE9512" blockId="2.[121,1467,2048,2106]" pageId="2" pageNumber="64">
<emphasis id="B913CA2EFF900D1DEE367B0515CA9531" bold="true" box="[121,219,2048,2073]" pageId="2" pageNumber="64">Figure 1.</emphasis>
<taxonomicName id="4C676DBFFF900D1DEEAA7B0617A99534" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[229,696,2051,2076]" class="Insecta" family="Gomphidae" genus="Anormogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="heteropterus">
@ -167,24 +168,24 @@ Three different sources have provided illustrations of the caudal appendages of
, viz., 1)
<bibRefCitation id="EFF66BCDFF910D1CEF7C714117519F48" author="Selys Longchamps, E. de &amp; Hagen H. A." box="[307,576,580,608]" pageId="3" pageNumber="65" pagination="257 - 720" refId="ref4685" refString="Selys Longchamps, E. de &amp; Hagen H. A. (1858). Monographie des gomphines. Memoires de la Societe des Sciences de Liege, 11, 257-720 [in French]." type="journal article" year="1858">Selys &amp; Hagen (1858)</bibRefCitation>
, 2) unpublished watercolor painting by Severin (see Material and meth- ods for the link), and 3) Fraser (1926; republished in 1934). These three sources do not agree well with each other, and the first two are very schematic, not showing any finer details of the structure. Only Draw- ing 1 (
<figureCitation id="135C0AB9FF910D1CED3873F216DB9C3B" box="[887,970,247,275]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." pageId="3" pageNumber="65">Fig. 4a</figureCitation>
<figureCitation id="135C0AB9FF910D1CED3873F216DB9C3B" box="[887,970,247,275]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." figureDoi="http://doi.org/10.5281/zenodo.15554904" httpUri="https://zenodo.org/record/15554904/files/figure.png" pageId="3" pageNumber="65">Fig. 4a</figureCitation>
) depicts the structure of cerci accurately. In Drawing 2 (
<figureCitation id="135C0AB9FF910D1CED93721D11259C1C" box="[988,1076,280,308]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." pageId="3" pageNumber="65">Fig. 4b</figureCitation>
<figureCitation id="135C0AB9FF910D1CED93721D11259C1C" box="[988,1076,280,308]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." figureDoi="http://doi.org/10.5281/zenodo.15554904" httpUri="https://zenodo.org/record/15554904/files/figure.png" pageId="3" pageNumber="65">Fig. 4b</figureCitation>
), the basal spine of cerci is not figured at all, whereas in Drawing 3 (
<figureCitation id="135C0AB9FF910D1CEABB723F10579C7E" box="[1268,1350,314,342]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." pageId="3" pageNumber="65">Fig. 4c</figureCitation>
<figureCitation id="135C0AB9FF910D1CEABB723F10579C7E" box="[1268,1350,314,342]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." figureDoi="http://doi.org/10.5281/zenodo.15554904" httpUri="https://zenodo.org/record/15554904/files/figure.png" pageId="3" pageNumber="65">Fig. 4c</figureCitation>
) the bas- al spine of cerci seems exaggerated, and the
<typeStatus id="54DCA89EFF910D1CEB33725E10AB9C5F" box="[1404,1466,347,375]" pageId="3" pageNumber="65" type="paratype">para-</typeStatus>
procts are very thin and curved sharply upwards. In the
<typeStatus id="54DCA89EFF910D1CED38729B16F09C92" box="[887,993,414,442]" pageId="3" pageNumber="65" type="holotype">holotype</typeStatus>
male, the cerci are broken, which might explain the discrepancy in the illustrations by Selys, es- pecially with regard to the first two of the three illustra- tions mentioned above. The third drawing in
<figureCitation id="135C0AB9FF910D1CEB007107109E9F36" box="[1359,1423,514,542]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." pageId="3" pageNumber="65">Fig. 4</figureCitation>
<figureCitation id="135C0AB9FF910D1CEB007107109E9F36" box="[1359,1423,514,542]" captionStart="Figure 4" captionStartId="5.[121,189,836,861]" captionTargetBox="[121,775,154,798]" captionTargetId="figure-23@5.[121,775,144,798]" captionTargetPageId="5" captionText="Figure 4. Anormogomphus heteropterus Selys, 1854. A com- pilation of illustrations modified afer (a) Selys &amp; Hagen, 1958: i) dorsal view, ii) lateral view, iii) lateral view of broken appendages, iv) cerci; (b) unpublished drawing by Severin (see Material and methods); and (c) Fraser (1934)." figureDoi="http://doi.org/10.5281/zenodo.15554904" httpUri="https://zenodo.org/record/15554904/files/figure.png" pageId="3" pageNumber="65">Fig. 4</figureCitation>
a-iii seems to be showing the broken appendages of the
<typeStatus id="54DCA89EFF910D1CED62714116869F48" box="[813,919,580,608]" pageId="3" pageNumber="65" type="holotype">holotype</typeStatus>
(compare with
<figureCitation id="135C0AB9FF910D1CEA1D714111B29F48" box="[1106,1187,580,608]" captionStart="Figure 3" captionStartId="4.[121,189,2049,2074]" captionTargetBox="[236,1351,144,2011]" captionTargetId="figure-23@4.[236,1351,144,2011]" captionTargetPageId="4" captionText="Figure 3. Anormogomphus heteropterus Selys, 1854. Holotype Male. (a) dorsal habitus, (b) lateral habitus, (c) lateral, (d) fron- tal, and (e) dorsal views of caudal appendages." pageId="3" pageNumber="65">Fig. 3c</figureCitation>
<figureCitation id="135C0AB9FF910D1CEA1D714111B29F48" box="[1106,1187,580,608]" captionStart="Figure 3" captionStartId="4.[121,189,2049,2074]" captionTargetBox="[236,1351,144,2011]" captionTargetId="figure-23@4.[236,1351,144,2011]" captionTargetPageId="4" captionText="Figure 3. Anormogomphus heteropterus Selys, 1854. Holotype Male. (a) dorsal habitus, (b) lateral habitus, (c) lateral, (d) fron- tal, and (e) dorsal views of caudal appendages." figureDoi="http://doi.org/10.5281/zenodo.15554902" httpUri="https://zenodo.org/record/15554902/files/figure.png" pageId="3" pageNumber="65">Fig. 3c</figureCitation>
). The male described in the present study has a peculiar structure at the base of the caudal appendages (highlighted with an arrow in
</paragraph>
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<paragraph id="8BD8163CFF910D1CEE367B04117E9514" blockId="3.[121,1466,2049,2108]" pageId="3" pageNumber="65">
<emphasis id="B913CA2EFF910D1CEE367B0415CA9532" bold="true" box="[121,219,2049,2074]" pageId="3" pageNumber="65">Figure 2.</emphasis>
<taxonomicName id="4C676DBFFF910D1CEEAB7B0117A99535" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[228,696,2052,2077]" class="Insecta" family="Gomphidae" genus="Anormogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="heteropterus">
@ -194,7 +195,7 @@ Selys, 1854
. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm].
</paragraph>
</caption>
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<paragraph id="8BD8163CFF960D1BEE367B0417639513" blockId="4.[121,1466,2049,2107]" pageId="4" pageNumber="66">
<emphasis id="B913CA2EFF960D1BEE367B0415C99532" bold="true" box="[121,216,2049,2074]" pageId="4" pageNumber="66">Figure 3.</emphasis>
<taxonomicName id="4C676DBFFF960D1BEE927B0617B79534" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[221,678,2051,2076]" class="Insecta" family="Gomphidae" genus="Anormogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="heteropterus">
@ -230,7 +231,7 @@ also has a much more westerly distribution with no known records from east of Ka
<collectingCountry id="F37056ACFF970D1AED3772BA16A69CF3" box="[888,951,447,475]" name="India" pageId="5" pageNumber="67">India</collectingCountry>
.
</paragraph>
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<paragraph id="8BD8163CFF970D1AEE36704117899EF2" blockId="5.[121,776,836,986]" pageId="5" pageNumber="67">
<emphasis id="B913CA2EFF970D1AEE36704115C89E75" bold="true" box="[121,217,836,861]" pageId="5" pageNumber="67">Figure 4.</emphasis>
<taxonomicName id="4C676DBFFF970D1AEEAF704317BD9E77" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[224,684,838,863]" class="Insecta" family="Gomphidae" genus="Anormogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="heteropterus">
@ -241,11 +242,11 @@ Selys, 1854
</paragraph>
</caption>
<paragraph id="8BD8163CFF970D1AEE36773C17B698CD" blockId="5.[121,776,1081,1509]" pageId="5" pageNumber="67">
<figureCitation id="135C0AB9FF970D1AEE36773C15DA997D" box="[121,203,1081,1109]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." pageId="5" pageNumber="67">Fig. 2d</figureCitation>
<figureCitation id="135C0AB9FF970D1AEE36773C15DA997D" box="[121,203,1081,1109]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554900" httpUri="https://zenodo.org/record/15554900/files/figure.png" pageId="5" pageNumber="67">Fig. 2d</figureCitation>
), made up of two lobes with black apices. These structures appear to be sclerotized extensions of the S10, but they were not mentioned in any description or considered in any illustration of this species. From the examination of the
<typeStatus id="54DCA89EFF970D1AEF2877BB148D99F2" box="[359,412,1214,1242]" pageId="5" pageNumber="67">type</typeStatus>
specimen it is clear that they are present (pointed out with an arrow in
<figureCitation id="135C0AB9FF970D1AEC3977DA17D899D3" box="[630,713,1247,1275]" captionStart="Figure 3" captionStartId="4.[121,189,2049,2074]" captionTargetBox="[236,1351,144,2011]" captionTargetId="figure-23@4.[236,1351,144,2011]" captionTargetPageId="4" captionText="Figure 3. Anormogomphus heteropterus Selys, 1854. Holotype Male. (a) dorsal habitus, (b) lateral habitus, (c) lateral, (d) fron- tal, and (e) dorsal views of caudal appendages." pageId="5" pageNumber="67">Fig. 3d</figureCitation>
<figureCitation id="135C0AB9FF970D1AEC3977DA17D899D3" box="[630,713,1247,1275]" captionStart="Figure 3" captionStartId="4.[121,189,2049,2074]" captionTargetBox="[236,1351,144,2011]" captionTargetId="figure-23@4.[236,1351,144,2011]" captionTargetPageId="4" captionText="Figure 3. Anormogomphus heteropterus Selys, 1854. Holotype Male. (a) dorsal habitus, (b) lateral habitus, (c) lateral, (d) fron- tal, and (e) dorsal views of caudal appendages." figureDoi="http://doi.org/10.5281/zenodo.15554902" httpUri="https://zenodo.org/record/15554902/files/figure.png" pageId="5" pageNumber="67">Fig. 3d</figureCitation>
). Ad- ditionally, in Fraser (1926, p. 735), the figure for
<taxonomicName id="4C676DBFFF970D1AECFE7604149F9816" authority="Bartenev, 1913" authorityName="Bartenev" authorityYear="1913" class="Insecta" family="Gomphidae" genus="Anormogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kiritshenkoi">
<emphasis id="B913CA2EFF970D1AECFE760415C29815" italics="true" pageId="5" pageNumber="67">A. kiritshenkoi</emphasis>
@ -284,15 +285,15 @@ Selys, 1854
Selys, 1854
</taxonomicName>
. From these genera it is differentiated by: (a) single-celled anal tri- angle (
<figureCitation id="135C0AB9FF970D1AEE8875F014069A39" box="[199,279,1781,1809]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." pageId="5" pageNumber="67">Fig. 1c</figureCitation>
<figureCitation id="135C0AB9FF970D1AEE8875F014069A39" box="[199,279,1781,1809]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="5" pageNumber="67">Fig. 1c</figureCitation>
), (b) base of HW rounded (not excavated;
<figureCitation id="135C0AB9FF970D1AEE36741315DB9A1A" box="[121,202,1814,1842]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." pageId="5" pageNumber="67">Fig. 1c</figureCitation>
<figureCitation id="135C0AB9FF970D1AEE36741315DB9A1A" box="[121,202,1814,1842]" captionStart="Figure 1" captionStartId="2.[121,189,2048,2073]" captionTargetBox="[154,1442,144,2010]" captionTargetId="figure-24@2.[143,1442,144,2010]" captionTargetPageId="2" captionText="Figure 1. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). (a) head; (b) lateral habitus; (c) base of lef HW; (d) dorsal habitus; (e) right FW &amp; HW [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554898" httpUri="https://zenodo.org/record/15554898/files/figure.png" pageId="5" pageNumber="67">Fig. 1c</figureCitation>
), (c) lateral margin of S10 curved (
<figureCitation id="135C0AB9FF970D1AEC37741317C19A1A" box="[632,720,1814,1842]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." pageId="5" pageNumber="67">Fig. 2e</figureCitation>
<figureCitation id="135C0AB9FF970D1AEC37741317C19A1A" box="[632,720,1814,1842]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554900" httpUri="https://zenodo.org/record/15554900/files/figure.png" pageId="5" pageNumber="67">Fig. 2e</figureCitation>
), (d) shape of caudal appendages (
<figureCitation id="135C0AB9FF970D1AEF937432175E9A7B" box="[476,591,1847,1875]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." pageId="5" pageNumber="67">Figs 2df</figureCitation>
<figureCitation id="135C0AB9FF970D1AEF937432175E9A7B" box="[476,591,1847,1875]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554900" httpUri="https://zenodo.org/record/15554900/files/figure.png" pageId="5" pageNumber="67">Figs 2df</figureCitation>
), and (e) shape of vesica spermalis (
<figureCitation id="135C0AB9FF970D1AEF25745C14CE9A5D" box="[362,479,1881,1909]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." pageId="5" pageNumber="67">Figs 2ac</figureCitation>
<figureCitation id="135C0AB9FF970D1AEF25745C14CE9A5D" box="[362,479,1881,1909]" captionStart="Figure 2" captionStartId="3.[121,189,2049,2074]" captionTargetBox="[121,1467,713,2012]" captionTargetId="figure-23@3.[121,1467,713,2012]" captionTargetPageId="3" captionText="Figure 2. Anormogomphus heteropterus Selys, 1854. Male (IBC-BO403). Hamuli: (a) lateral, (b) ventral, (c) lateral view of hamuli; caudal appendages: (d) posterior, (e) lateral, and (f) dorsal views [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554900" httpUri="https://zenodo.org/record/15554900/files/figure.png" pageId="5" pageNumber="67">Figs 2ac</figureCitation>
). This species belongs to the genus
<taxonomicName id="4C676DBFFF970D1AEEB5747F14D69ABD" authorityName="Selys" authorityYear="1854" box="[250,455,1914,1941]" class="Insecta" family="Gomphidae" genus="Anormogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B913CA2EFF970D1AEEB5747F14D69ABD" box="[250,455,1914,1941]" italics="true" pageId="5" pageNumber="67">Anormogomphus</emphasis>

View file

@ -1,41 +1,42 @@
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<treatment id="03CEA72AFF970D17ED62775F106F953B" LSID="urn:lsid:plazi:treatment:03CEA72AFF970D17ED62775F106F953B" httpUri="http://treatment.plazi.org/id/03CEA72AFF970D17ED62775F106F953B" lastPageId="8" lastPageNumber="70" pageId="5" pageNumber="67">
<subSubSection id="C37D45B7FF970D1AED62775F1065995B" box="[813,1396,1111,1142]" pageId="5" pageNumber="67" type="nomenclature">
@ -53,7 +54,7 @@ Joshi &amp; Mital
</subSubSection>
<subSubSection id="C37D45B7FF970D17ED62777E147D9533" lastPageId="8" lastPageNumber="70" pageId="5" pageNumber="67" type="description">
<paragraph id="8BD8163CFF970D1AED62777E169C99BF" blockId="5.[813,1396,1111,1175]" box="[813,909,1147,1175]" pageId="5" pageNumber="67">
<figureCitation id="135C0AB9FF970D1AED62777E169C99BF" box="[813,909,1147,1175]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStartId-0="6.[121,189,2008,2033]" captionStartId-1="7.[121,189,1991,2016]" captionTargetBox-0="[121,1467,144,1945]" captionTargetBox-1="[121,1467,143,1952]" captionTargetId-0="figure-24@6.[121,1467,144,1970]" captionTargetId-1="figure-24@7.[121,1467,143,1952]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." captionText-1="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." pageId="5" pageNumber="67">Figs 56</figureCitation>
<figureCitation id="135C0AB9FF970D1AED62777E169C99BF" box="[813,909,1147,1175]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStartId-0="6.[121,189,2008,2033]" captionStartId-1="7.[121,189,1991,2016]" captionTargetBox-0="[121,1467,144,1945]" captionTargetBox-1="[121,1467,143,1952]" captionTargetId-0="figure-24@6.[121,1467,144,1970]" captionTargetId-1="figure-24@7.[121,1467,143,1952]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." captionText-1="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." figureDoi-0="http://doi.org/10.5281/zenodo.15554906" figureDoi-1="http://doi.org/10.5281/zenodo.15554908" httpUri-0="https://zenodo.org/record/15554906/files/figure.png" httpUri-1="https://zenodo.org/record/15554908/files/figure.png" pageId="5" pageNumber="67">Figs 56</figureCitation>
</paragraph>
<paragraph id="8BD8163CFF970D1AED6277BB168999F1" blockId="5.[813,920,1214,1241]" box="[813,920,1214,1241]" pageId="5" pageNumber="67">
<emphasis id="B913CA2EFF970D1AED6277BB168999F1" box="[813,920,1214,1241]" italics="true" pageId="5" pageNumber="67">
@ -109,10 +110,10 @@ male
</paragraph>
<paragraph id="8BD8163CFF970D17ED6274B9148F9C7E" blockId="5.[813,1467,1980,2108]" lastBlockId="8.[121,776,147,342]" lastPageId="8" lastPageNumber="70" pageId="5" pageNumber="67">
Head (
<figureCitation id="135C0AB9FF970D1AED3774B916DD9AF0" box="[888,972,1980,2008]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="5" pageNumber="67">Fig. 5a</figureCitation>
<figureCitation id="135C0AB9FF970D1AED3774B916DD9AF0" box="[888,972,1980,2008]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="5" pageNumber="67">Fig. 5a</figureCitation>
). Eyes turquoise blue. Labium yellow, face dark brown, covered with hairs on the sides, faint yel- low paired spots on labrum, anteclypeus paler brown, large paired blue spots covering almost the entire post- frons, postfrons with brown setae. Blue and brown bands on the first and second segments of antenna, filament black. Labrum 2.3 times as wide as long. Ocel- lar region sky-blue, junction of frons and vertex white, vertex black, occiput pale bluish yellow, occipital ridge black with a yellow spot.
</paragraph>
<caption id="DF1846B4FF940D19EE3674DD17C7953B" pageId="6" pageNumber="68" startId="6.[121,189,2008,2033]" targetBox="[121,1467,144,1945]" targetPageId="6" targetType="figure">
<caption id="DF1846B4FF940D19EE3674DD17C7953B" ID-DOI="http://doi.org/10.5281/zenodo.15554906" ID-Zenodo-Dep="15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="6" pageNumber="68" startId="6.[121,189,2008,2033]" targetBox="[121,1467,144,1945]" targetPageId="6" targetType="figure">
<paragraph id="8BD8163CFF940D19EE3674DD17C7953B" blockId="6.[121,1467,2008,2067]" pageId="6" pageNumber="68">
<emphasis id="B913CA2EFF940D19EE3674DD15C99AD9" bold="true" box="[121,216,2008,2033]" pageId="6" pageNumber="68">Figure 5.</emphasis>
<taxonomicName id="4C676DBFFF940D19EE9174DE14E99ADB" authorityName="Joshi" authorityYear="2022" box="[222,504,2011,2035]" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="benritarum" status="sp. nov.">
@ -122,7 +123,7 @@ Head (
Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm].
</paragraph>
</caption>
<caption id="DF1846B4FF950D18EE3674C21094952A" pageId="7" pageNumber="69" startId="7.[121,189,1991,2016]" targetBox="[121,1467,143,1952]" targetPageId="7" targetType="figure">
<caption id="DF1846B4FF950D18EE3674C21094952A" ID-DOI="http://doi.org/10.5281/zenodo.15554908" ID-Zenodo-Dep="15554908" httpUri="https://zenodo.org/record/15554908/files/figure.png" pageId="7" pageNumber="69" startId="7.[121,189,1991,2016]" targetBox="[121,1467,143,1952]" targetPageId="7" targetType="figure">
<paragraph id="8BD8163CFF950D18EE3674C21094952A" blockId="7.[121,1466,1991,2050]" pageId="7" pageNumber="69">
<emphasis id="B913CA2EFF950D18EE3674C215C89AC8" bold="true" box="[121,217,1991,2016]" pageId="7" pageNumber="69">Figure 6.</emphasis>
<taxonomicName id="4C676DBFFF950D18EE9074CF14EA9ACA" authorityName="Joshi" authorityYear="2022" box="[223,507,1994,2018]" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="69" phylum="Arthropoda" rank="species" species="benritarum" status="sp. nov.">
@ -134,12 +135,12 @@ Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap
</caption>
<paragraph id="8BD8163CFF9A0D17EE36727914D49F36" blockId="8.[121,776,380,1045]" pageId="8" pageNumber="70">
Thorax (
<figureCitation id="135C0AB9FF9A0D17EE99727914589CB0" box="[214,329,380,408]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="8" pageNumber="70">Figs 5ce</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EE99727914589CB0" box="[214,329,380,408]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="8" pageNumber="70">Figs 5ce</figureCitation>
). Prothorax yellow with median lobes. Pterothorax black, marked with greenish yellow as fol- lows: mesothoracic collar, dorsally with oval spots, stripe on mesepisternum, laterally broadly yellow with black stripes across sutures.
</paragraph>
<paragraph id="8BD8163CFF9A0D17EED8712614E7993A" blockId="8.[121,776,380,1045]" pageId="8" pageNumber="70">
Legs black, profemora internally marked yel- low, covered with black spines; coxae yellow. Ven- ter of thorax and base of legs slightly pruinose. Wings (
<figureCitation id="135C0AB9FF9A0D17EE9D718214369F8B" box="[210,295,647,675]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="8" pageNumber="70">Fig. 5b</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EE9D718214369F8B" box="[210,295,647,675]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="8" pageNumber="70">Fig. 5b</figureCitation>
). Base of wings and membranes yellow. Pterostigma dark brown, pterostigmal brace present. Antenodals (1
<superScript id="7C12BB74FF9A0D17EF6F71C2143D9FFE" attach="left" box="[288,300,711,726]" fontSize="6" pageId="8" pageNumber="70">st</superScript>
and 5
@ -150,7 +151,7 @@ and IRP
<subScript id="17E31479FF9A0D17EE9C770315CD993D" attach="right" box="[211,220,1030,1045]" fontSize="6" pageId="8" pageNumber="70">2</subScript>
, two rows near margin.
</paragraph>
<caption id="DF1846B4FF9A0D17ED62718816A49FCD" pageId="8" pageNumber="70" startId="8.[813,881,653,678]" targetBox="[813,1467,144,614]" targetPageId="8" targetType="figure">
<caption id="DF1846B4FF9A0D17ED62718816A49FCD" ID-DOI="http://doi.org/10.5281/zenodo.15554910" ID-Zenodo-Dep="15554910" httpUri="https://zenodo.org/record/15554910/files/figure.png" pageId="8" pageNumber="70" startId="8.[813,881,653,678]" targetBox="[813,1467,144,614]" targetPageId="8" targetType="figure">
<paragraph id="8BD8163CFF9A0D17ED62718816A49FCD" blockId="8.[813,1468,653,741]" pageId="8" pageNumber="70">
<emphasis id="B913CA2EFF9A0D17ED627188169F9F8E" bold="true" box="[813,910,653,678]" pageId="8" pageNumber="70">Figure 7.</emphasis>
Type locality of
@ -163,17 +164,17 @@ at the Rudra Padh Temple,Tezpur,Assam, India (Photo: Manisha Kumari).
</caption>
<paragraph id="8BD8163CFF9A0D17EE36773D171F98A8" blockId="8.[121,776,1080,1408]" pageId="8" pageNumber="70">
Abdomen (
<figureCitation id="135C0AB9FF9A0D17EEB4773D1461997C" box="[251,368,1080,1108]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="8" pageNumber="70">Figs 5ce</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EEB4773D1461997C" box="[251,368,1080,1108]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="8" pageNumber="70">Figs 5ce</figureCitation>
). Narrow up to anterior half of S7, S8 expanded, S910 tapering. Black marked with yel- low. S1 to base of S3 laterally greenish yellow, S37 with basal annules, broken on S3 and extensive on S7. Auricles triangular, pointed. S23 with a mediodorsal triangular marking pointing posteriorly, small faint dor- sal spots on S45. Markings gradually turning yellowish brown, especially from S7. S9 with paired dorsal mark- ings along lateral margins and a small marking on the posterior border; S10 dark brown.
</paragraph>
<paragraph id="8BD8163CFF9A0D17EE3676A214099BC7" blockId="8.[121,776,1447,1975]" pageId="8" pageNumber="70">
Vesica spermalis (
<figureCitation id="135C0AB9FF9A0D17EF0576A214CF98EB" box="[330,478,1447,1475]" captionStart="Figure 6" captionStartId="7.[121,189,1991,2016]" captionTargetBox="[121,1467,143,1952]" captionTargetId="figure-24@7.[121,1467,143,1952]" captionTargetPageId="7" captionText="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." pageId="8" pageNumber="70">Figs 6a, b, d</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EF0576A214CF98EB" box="[330,478,1447,1475]" captionStart="Figure 6" captionStartId="7.[121,189,1991,2016]" captionTargetBox="[121,1467,143,1952]" captionTargetId="figure-24@7.[121,1467,143,1952]" captionTargetPageId="7" captionText="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554908" httpUri="https://zenodo.org/record/15554908/files/figure.png" pageId="8" pageNumber="70">Figs 6a, b, d</figureCitation>
). Genital lobe expanded, brown. Anterior hamuli short, simple, rounded, and covered with setae on apices. Posterior hamuli longer, shaped like a wrench; apical anterior margin notched, apices anteriorly curved, ending in a spine. V2 simple, rounded at edges, ventrally marked white. V3 curved at base, with a ventral beak-like prepuce, broadening on the apices, V4 with protuberances ventrally, saddle-shaped extensions; apical filaments short, apices ante- riorly curved.
</paragraph>
<paragraph id="8BD8163CFF9A0D17EED875F117369A9F" blockId="8.[121,776,1447,1975]" pageId="8" pageNumber="70">
Caudal appendages (
<figureCitation id="135C0AB9FF9A0D17EFD675F117259A38" box="[409,564,1780,1808]" captionStart="Figure 6" captionStartId="7.[121,189,1991,2016]" captionTargetBox="[121,1467,143,1952]" captionTargetId="figure-24@7.[121,1467,143,1952]" captionTargetPageId="7" captionText="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." pageId="8" pageNumber="70">Figs 6c, e, f</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EFD675F117259A38" box="[409,564,1780,1808]" captionStart="Figure 6" captionStartId="7.[121,189,1991,2016]" captionTargetBox="[121,1467,143,1952]" captionTargetId="figure-24@7.[121,1467,143,1952]" captionTargetPageId="7" captionText="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554908" httpUri="https://zenodo.org/record/15554908/files/figure.png" pageId="8" pageNumber="70">Figs 6c, e, f</figureCitation>
). Black, epiproct brown at base and laterally. Cerci 1.2 times longer than epiproct. Cerci expanded, trilobate; lateral mar- gin forming a sharp spine at base, outer edge serrated, apices ending in two spines. Epiproct slightly divaricate, ending in an upwardly curved spine.
</paragraph>
<paragraph id="8BD8163CFF9A0D17EE3674DB147D9533" blockId="8.[121,776,2014,2075]" pageId="8" pageNumber="70">Measurements. Abdomen + caudal appendages = 32.9, FW = 32.6, HW = 31.</paragraph>
@ -188,7 +189,7 @@ This species is placed in the genus
<emphasis id="B913CA2EFF9A0D17EA95706D10959EAB" box="[1242,1412,872,899]" italics="true" pageId="8" pageNumber="70">Platygomphus</emphasis>
</taxonomicName>
due to the shape of the male genitalia (ending with di- vergent apical filaments,
<figureCitation id="135C0AB9FF9A0D17EA2170AE11D99EEF" box="[1134,1224,939,967]" captionStart="Figure 6" captionStartId="7.[121,189,1991,2016]" captionTargetBox="[121,1467,143,1952]" captionTargetId="figure-24@7.[121,1467,143,1952]" captionTargetPageId="7" captionText="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." pageId="8" pageNumber="70">Fig. 6d</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EA2170AE11D99EEF" box="[1134,1224,939,967]" captionStart="Figure 6" captionStartId="7.[121,189,1991,2016]" captionTargetBox="[121,1467,143,1952]" captionTargetId="figure-24@7.[121,1467,143,1952]" captionTargetPageId="7" captionText="Figure 6. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402). (a) hamuli lateral; (b) hamuli ventral; (c) caudal ap- pendages posterior; (d) vesica spermalis lateral view; caudal appendages: (e) lateral; and (f) dorsal [scale bar = 0.5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554908" httpUri="https://zenodo.org/record/15554908/files/figure.png" pageId="8" pageNumber="70">Fig. 6d</figureCitation>
), and anal triangle three-celled and curved along the margin. From
<taxonomicName id="4C676DBFFF9A0D17EB3770C9169E9921" authorityName="Selys" authorityYear="1854" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="70" phylum="Arthropoda" rank="species" species="dolabratus">
<emphasis id="B913CA2EFF9A0D17EB3770C9169E9921" italics="true" pageId="8" pageNumber="70">P. dolabratus</emphasis>
@ -202,13 +203,13 @@ and
<emphasis id="B913CA2EFF9A0D17EDEB773516E59963" box="[932,1012,1072,1099]" italics="true" pageId="8" pageNumber="70">P. feae</emphasis>
</taxonomicName>
but medial ridge prominent with the two lobes less furcated;
<figureCitation id="135C0AB9FF9A0D17EA29775711CA9946" box="[1126,1243,1106,1134]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="8" pageNumber="70">Figs 5ef</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EA29775711CA9946" box="[1126,1243,1106,1134]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="8" pageNumber="70">Figs 5ef</figureCitation>
), (b) markings on face (labrum, clypeus, antefrons brown with pale blu- ish frons in
<taxonomicName id="4C676DBFFF9A0D17ED8E779111749987" authorityName="Joshi" authorityYear="2022" box="[961,1125,1172,1199]" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="70" phylum="Arthropoda" rank="species" species="benritarum">
<emphasis id="B913CA2EFF9A0D17ED8E779111749987" box="[961,1125,1172,1199]" italics="true" pageId="8" pageNumber="70">P. benritarum</emphasis>
</taxonomicName>
vs. face extensively yellow;
<figureCitation id="135C0AB9FF9A0D17ED6277B3169099FA" box="[813,897,1206,1234]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="8" pageNumber="70">Fig. 5a</figureCitation>
<figureCitation id="135C0AB9FF9A0D17ED6277B3169099FA" box="[813,897,1206,1234]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="8" pageNumber="70">Fig. 5a</figureCitation>
), and (c) shape of apical flagellum of V4 (short and thick, vs. long in
<taxonomicName id="4C676DBFFF9A0D17EA6777D211D399DA" authorityName="Selys" authorityYear="1854" box="[1064,1218,1239,1266]" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="70" phylum="Arthropoda" rank="species" species="dolabratus">
<emphasis id="B913CA2EFF9A0D17EA6777D211D399DA" box="[1064,1218,1239,1266]" italics="true" pageId="8" pageNumber="70">P. dolabratus</emphasis>
@ -218,7 +219,7 @@ and
<emphasis id="B913CA2EFF9A0D17EAB177D2105C99DA" box="[1278,1357,1239,1266]" italics="true" pageId="8" pageNumber="70">P. feae</emphasis>
</taxonomicName>
;
<figureCitation id="135C0AB9FF9A0D17EB1477D210BF99DB" box="[1371,1454,1239,1267]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." pageId="8" pageNumber="70">Fig. 5d</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EB1477D210BF99DB" box="[1371,1454,1239,1267]" captionStart="Figure 5" captionStartId="6.[121,189,2008,2033]" captionTargetBox="[121,1467,144,1945]" captionTargetId="figure-24@6.[121,1467,144,1970]" captionTargetPageId="6" captionText="Figure 5. Platygomphus benritarum sp. nov. Holotype male (IBC-BO402) (a) head; (b) right FW &amp; HW; (c) lateral habitus (live); (d) dorsal habitus; (e) lateral habitus [scale bar = 5 mm]." figureDoi="http://doi.org/10.5281/zenodo.15554906" httpUri="https://zenodo.org/record/15554906/files/figure.png" pageId="8" pageNumber="70">Fig. 5d</figureCitation>
).
<taxonomicName id="4C676DBFFF9A0D17ED6277FD117B983B" authorityName="Joshi" authorityYear="2022" box="[813,1130,1272,1299]" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="70" phylum="Arthropoda" rank="species" species="benritarum">
<emphasis id="B913CA2EFF9A0D17ED6277FD117B983B" box="[813,1130,1272,1299]" italics="true" pageId="8" pageNumber="70">Platygomphus benritarum</emphasis>
@ -274,7 +275,7 @@ A single male was found resting on a large
<emphasis id="B913CA2EFF9A0D17EB5075E9104B9A2F" box="[1311,1370,1772,1799]" italics="true" pageId="8" pageNumber="70">Ficus</emphasis>
</taxonomicName>
tree sit- uated about 56 meters from the banks of Brahmaputra River near the Rudra Padh Temple at around 17.00 h. The habitat along the banks is dominated by grasses, sparse trees, paddy fields, and marshlands along with some forest patches and tree plantations (
<figureCitation id="135C0AB9FF9A0D17EB50749610729A87" box="[1311,1379,1939,1967]" captionStart="Figure 7" captionStartId="8.[813,881,653,678]" captionTargetBox="[813,1467,144,614]" captionTargetId="figure-46@8.[813,1467,141,615]" captionTargetPageId="8" captionText="Figure 7. Type locality of Platygomphus benritarum sp. nov. at the Rudra Padh Temple,Tezpur,Assam, India (Photo: Manisha Kumari)." pageId="8" pageNumber="70">Fig. 7</figureCitation>
<figureCitation id="135C0AB9FF9A0D17EB50749610729A87" box="[1311,1379,1939,1967]" captionStart="Figure 7" captionStartId="8.[813,881,653,678]" captionTargetBox="[813,1467,144,614]" captionTargetId="figure-46@8.[813,1467,141,615]" captionTargetPageId="8" captionText="Figure 7. Type locality of Platygomphus benritarum sp. nov. at the Rudra Padh Temple,Tezpur,Assam, India (Photo: Manisha Kumari)." figureDoi="http://doi.org/10.5281/zenodo.15554910" httpUri="https://zenodo.org/record/15554910/files/figure.png" pageId="8" pageNumber="70">Fig. 7</figureCitation>
).
<taxonomicName id="4C676DBFFF9A0D17EB39749616C69AF8" class="Insecta" family="Gomphidae" genus="Platygomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="70" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B913CA2EFF9A0D17EB397496168A9AE7" italics="true" pageId="8" pageNumber="70">Platygomphus</emphasis>

View file

@ -0,0 +1,201 @@
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<treatment id="03EC87C7FFD2D21EFDD891D97BABFDDA" LSID="urn:lsid:plazi:treatment:03EC87C7FFD2D21EFDD891D97BABFDDA" httpUri="http://treatment.plazi.org/id/03EC87C7FFD2D21EFDD891D97BABFDDA" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C35F655AFFD2D21DFDD891D97B32FE4A" box="[625,1321,451,475]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BFA36D1FFD2D21DFDD891D97B32FE4A" blockId="13.[625,1321,451,475]" box="[625,1321,451,475]" pageId="13" pageNumber="14">
<heading id="D0B281BDFFD2D21DFDD891D97B32FE4A" box="[625,1321,451,475]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B931EAC3FFD2D21DFDD891D97B32FE4A" bold="true" box="[625,1321,451,475]" pageId="13" pageNumber="14">
Key to the species of
<taxonomicName id="4C454D52FFD2D21DFCD891D97DE4FE4A" ID-CoL="6QKH" authority="Eigenmann and Eigenmann, 1888" box="[881,1023,451,475]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">Pimelodella</taxonomicName>
from the
<collectingCountry id="F3527641FFD2D21DFBD191D97AF9FE4A" box="[1144,1250,451,475]" name="Paraguay" pageId="13" pageNumber="14">Paraguai</collectingCountry>
basin
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F655AFFD2D21EFE4192147BABFDDA" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="key">
<key id="FFD461E0FFD2D21EFE4192147BABFDDA" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<keyStep id="30B12E74FFD2D21DFE4192147BABFC97" pageId="13" pageNumber="14">
<paragraph id="8BFA36D1FFD2D21DFE4192147BB4FD07" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4192147BB4FD07" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4192147C11FDB7" bold="true" box="[488,522,526,550]" pageId="13" pageNumber="14">1a.</emphasis>
Total vertebrae 46 (rarely 45); anal-fin adpressed terminus always anterior to adipose-fin terminus, reaching at least the vertical through three-fourths of adipose fin; adipose fin very long, slightly more than 2.0 to 2.5 times in SL .................................................................................................................
<taxonomicName id="4C454D52FFD2D21DFAF892657BB4FD07" ID-CoL="4HWDZ" baseAuthorityName="Valenciennes" baseAuthorityYear="1835" box="[1361,1455,638,662]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B931EAC3FFD2D21DFAF892657BB4FD07" box="[1361,1455,638,662]" italics="true" pageId="13" pageNumber="14">P. gracilis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36D1FFD2D21DFE4192B97BABFC97" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4192B97BABFC97" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4192B97C11FD2A" bold="true" box="[488,522,675,699]" pageId="13" pageNumber="14">1b.</emphasis>
Total vertebrae 3944; anal-fin adpressed terminus between verticals through adipose-fin terminus and slightly posterior to adipose-fin terminus; adipose fin 2.5 or more times in SL.............................................................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E74FFD2D21DFE4193097BABFB9A" pageId="13" pageNumber="14">
<paragraph id="8BFA36D1FFD2D21DFE4193097BB4FC0A" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4193097BB4FC0A" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4193097C11FCBA" bold="true" box="[488,522,787,811]" pageId="13" pageNumber="14">2a.</emphasis>
Openings of preoperculomandibular laterosensory canal at dentary large and conspicuous; posterior margin of pectoral-fin spine bearing 1420 small, retrorse blades along basal two-thirds (
<figureCitation id="137E2A54FFD2D21DFB9093447A83FCE7" box="[1081,1176,862,886]" captionStart="FIGURE 3" captionStartId="7.[488,561,1276,1296]" captionTargetBox="[496,1443,191,1219]" captionTargetId="figure-123@7.[488,1459,171,1242]" captionTargetPageId="7" captionText="FIGURE 3 | Pectoral-fin spine of A. Pimelodella guato, paratype, CIUnB 1772, 91.2 mm SL, length of spine 16.7 mm; B. P. taeniophora, ZUFMS-PIS 6320, 68.9 mm SL, length of spine 11.4 mm; C. P. mucosa, holotype, CAS 63720, 97.4 mm SL, length of spine 22.8 mm; D. P. serrata, holotype, FMNH 57979, 55.7 mm SL, length of spine 18.3 mm; and E. P. howesi, holotype, ANSP 69036, 79.3 mm SL, length of spine 17.8 mm." figureDoi="http://doi.org/10.5281/zenodo.13326473" httpUri="https://zenodo.org/record/13326473/files/figure.png" pageId="13" pageNumber="14">Fig. 3C</figureCitation>
); head roof heavily ornamented.....................................................................................................
<taxonomicName id="4C454D52FFD2D21DFAE4939E7BB4FC0A" ID-CoL="4HWF2" authorityName="Eigenmann &amp; Ward" authorityYear="1907" box="[1357,1455,900,923]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="mucosa">
<emphasis id="B931EAC3FFD2D21DFAE4939E7BB4FC0A" box="[1357,1455,900,923]" italics="true" pageId="13" pageNumber="14">P. mucosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36D1FFD2D21DFE4193B27BABFB9A" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4193B27BABFB9A" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4193B27C11FC51" bold="true" box="[488,522,936,960]" pageId="13" pageNumber="14">2b.</emphasis>
Openings of preoperculomandibular laterosensory canal at dentary not large or particularly conspicuous; posterior margin of pectoral-fin spine not as above; head roof ornamentation inconspicuous................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E74FFD2D21DFE4194027BABFAA7" pageId="13" pageNumber="14">
<paragraph id="8BFA36D1FFD2D21DFE4194027BABFB31" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4194027BABFB31" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4194027C11FBA1" bold="true" box="[488,522,1048,1072]" pageId="13" pageNumber="14">3a.</emphasis>
Supraoccipital process not reaching the anterior prenuchal plate; dorsal-fin spine small, approximately half or slightly more of second dorsal-fin ray total length; dorsal lobe of caudal fin notably longer than ventral caudal-lobe; hypural 5 variably fused to hypural 3+4......................................................................
<taxonomicName id="4C454D52FFD2D21DFA9294937BABFB31" ID-CoL="77JPH" authorityName="Miranda Ribeiro" authorityYear="1918" box="[1339,1456,1160,1184]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="megalura">
<emphasis id="B931EAC3FFD2D21DFA9294937BABFB31" box="[1339,1456,1160,1184]" italics="true" pageId="13" pageNumber="14">P. megalura</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36D1FFD2D21DFE4194B47BABFAA7" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4194B47BABFAA7" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4194B47C11FB57" bold="true" box="[488,522,1198,1222]" pageId="13" pageNumber="14">3b.</emphasis>
Supraoccipital process reaching the anterior prenuchal plate; dorsal-fin spine at least a third of second dorsal-fin total ray length (excepting the filamentuous portion, if present); caudal-fin lobes subequal or ventral lobe slightly longer than dorsal; hypural 5 completely free ...................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E74FFD2D21DFE4195597BABF9AA" pageId="13" pageNumber="14">
<paragraph id="8BFA36D1FFD2D21DFE4195597BB4FA5A" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
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<emphasis id="B931EAC3FFD2D21DFE4195597C11FACA" bold="true" box="[488,522,1347,1371]" pageId="13" pageNumber="14">4a.</emphasis>
Maxillary barbels always surpassing caudal-fin origin; posterior margin of pectoral-fin spine bearing 46 notably triangular, short and straight blades along basal two-thirds; dorsal fin distal third notably dark brown to black........................................................................................................
<taxonomicName id="4C454D52FFD2D21DFA9995AE7BB4FA5A" ID-CoL="4HWF4" authorityName="Eigenmann" authorityYear="1917" box="[1328,1455,1459,1483]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="notomelas">
<emphasis id="B931EAC3FFD2D21DFA9995AE7BB4FA5A" box="[1328,1455,1459,1483]" italics="true" pageId="13" pageNumber="14">P. notomelas</emphasis>
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</paragraph>
<paragraph id="8BFA36D1FFD2D21DFE4195C27BABF9AA" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
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<emphasis id="B931EAC3FFD2D21DFE4195C27C11FA61" bold="true" box="[488,522,1496,1520]" pageId="13" pageNumber="14">4b.</emphasis>
Maxillary barbels reaching between pelvic-fin origin and may extend beyond caudal-fin origin; posterior margin of the pectoral-fin spine not as above; dorsal fin not presenting a notably dark brown to black coloration ....................... 5
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<keyStep id="30B12E74FFD2D21DFE4196537BABF8D1" pageId="13" pageNumber="14">
<paragraph id="8BFA36D1FFD2D21DFE4196537BABF941" blockId="13.[488,1456,526,1931]" pageId="13" pageNumber="14">
<keyLead id="30B495E4FFD2D21DFE4196537BABF941" pageId="13" pageNumber="14">
<emphasis id="B931EAC3FFD2D21DFE4196537C11F9F0" bold="true" box="[488,522,1609,1633]" pageId="13" pageNumber="14">5a.</emphasis>
Dorsal profile straight from snout to dorsal fin; dorsal-fin spine robust, large, bearing small, straight spinules along three-fourths of its distal posterior margin; posterior margin of pectoral-fin spine bearing 1323 retrorse blades along basal two-thirds (
<figureCitation id="137E2A54FFD2D21DFD7196A27D36F940" box="[728,813,1720,1745]" captionStart="FIGURE 3" captionStartId="7.[488,561,1276,1296]" captionTargetBox="[496,1443,191,1219]" captionTargetId="figure-123@7.[488,1459,171,1242]" captionTargetPageId="7" captionText="FIGURE 3 | Pectoral-fin spine of A. Pimelodella guato, paratype, CIUnB 1772, 91.2 mm SL, length of spine 16.7 mm; B. P. taeniophora, ZUFMS-PIS 6320, 68.9 mm SL, length of spine 11.4 mm; C. P. mucosa, holotype, CAS 63720, 97.4 mm SL, length of spine 22.8 mm; D. P. serrata, holotype, FMNH 57979, 55.7 mm SL, length of spine 18.3 mm; and E. P. howesi, holotype, ANSP 69036, 79.3 mm SL, length of spine 17.8 mm." figureDoi="http://doi.org/10.5281/zenodo.13326473" httpUri="https://zenodo.org/record/13326473/files/figure.png" pageId="13" pageNumber="14">Fig. 3A</figureCitation>
) .................................................................................
<taxonomicName id="4C454D52FFD2D21DFAC996A37BABF941" authorityName="Pierre &amp; Slobodian" authorityYear="2024" box="[1376,1456,1721,1744]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guato">
<emphasis id="B931EAC3FFD2D21DFAC996A37BABF941" box="[1376,1456,1721,1744]" italics="true" pageId="13" pageNumber="14">P. guato</emphasis>
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<emphasis id="B931EAC3FFD2D21DFE4196C47C11F967" bold="true" box="[488,522,1758,1782]" pageId="13" pageNumber="14">5b.</emphasis>
Dorsal profile convex from snout to dorsal fin; dorsal-fin spine not particularly robust, nor bearing spinules at its posterior margin; posterior margin of pectoral-fin spine bearing 913 retrorse blades along basal two-thirds .................. 6
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<emphasis id="B931EAC3FFD2D21DFE4197547C11F8F7" bold="true" box="[488,522,1870,1894]" pageId="13" pageNumber="14">6a.</emphasis>
Maxillary barbels reaching at least anal-fin terminus when parallel to main body axis; adipose fin 2.5 to almost 3.0 times in SL ..............................
<taxonomicName id="4C454D52FFD2D21DFA88976E7BABF81A" ID-CoL="4HWFV" box="[1313,1456,1907,1931]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="taeniophora">
<emphasis id="B931EAC3FFD2D21DFA88976E7BABF81A" box="[1313,1456,1907,1931]" italics="true" pageId="13" pageNumber="14">P. taeniophora</emphasis>
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<paragraph id="8BFA36D1FFD1D21EFE4190A47BABFF6A" blockId="14.[488,1456,190,587]" pageId="14" pageNumber="15">
<keyLead id="30B495E4FFD1D21EFE4190A47BABFF6A" pageId="14" pageNumber="15">
<emphasis id="B931EAC3FFD1D21EFE4190A47C11FF47" bold="true" box="[488,522,190,214]" pageId="14" pageNumber="15">6b.</emphasis>
Maxillary barbels reaching between pelvic-fin origin and anal-fin terminus; adipose fin short, more than 3.5 times in SL............................................................. 7
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</keyStep>
<keyStep id="30B12E74FFD1D21EFE4191137BABFDDA" pageId="14" pageNumber="15">
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<keyLead id="30B495E4FFD1D21EFE4191137BB4FE27" pageId="14" pageNumber="15">
<emphasis id="B931EAC3FFD1D21EFE4191137C11FEB0" bold="true" box="[488,522,265,289]" pageId="14" pageNumber="15">7a.</emphasis>
Epiphyseal branch of supraorbital canal on the head (S6) with contralateral canals connecting at midline, proceeding posteriorly as a single canal and opening in a single pore (S6+S6 diaulic pore); total vertebrae 4142; dorsal region of body usually presenting a well-delimited darker stripe from supraoccipital process to first third of adipose-fin base ..........................................................................
<taxonomicName id="4C454D52FFD1D21EFAF891857BB4FE27" ID-CoL="6VJMV" authorityName="Eigenmann" authorityYear="1917" box="[1361,1455,414,438]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="griffini">
<emphasis id="B931EAC3FFD1D21EFAF891857BB4FE27" box="[1361,1455,414,438]" italics="true" pageId="14" pageNumber="15">P. griffini</emphasis>
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</paragraph>
<paragraph id="8BFA36D1FFD1D21EFE4191D97BABFDDA" blockId="14.[488,1456,190,587]" pageId="14" pageNumber="15">
<keyLead id="30B495E4FFD1D21EFE4191D97BABFDDA" pageId="14" pageNumber="15">
<emphasis id="B931EAC3FFD1D21EFE4191D97C11FE4A" bold="true" box="[488,522,451,475]" pageId="14" pageNumber="15">7b.</emphasis>
Epiphyseal branch of supraorbital canal on the head (S6) emerging as two separated pores; total vertebrae 3940; dorsal region of head and body slightly darkened, with a dark brown mark extending just between dorsal and adipose fins...............................................................................................
<taxonomicName id="4C454D52FFD1D21EFAE7922E7BABFDDA" ID-CoL="77JMZ" authorityName="Eigenmann" authorityYear="1917" box="[1358,1456,563,587]" family="Heptapteridae" genus="Pimelodella" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B931EAC3FFD1D21EFAE7922E7BABFDDA" box="[1358,1456,563,587]" italics="true" pageId="14" pageNumber="15">P. laticeps</emphasis>
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Genus
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species:
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<mods:roleTerm id="9489AA3DBD7C37A6EAA2C4EB5C3DC305">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="632B838755E21BCF12703129C339B3B8">Chandran, Reji</mods:namePart>
<mods:affiliation id="021ACF2B6D57D175EC107854F5B5E064">Society for Odonate Studies, Vellooparampil, Kuzhimattom PO, Kottayam, Kerala 686533, India</mods:affiliation>
</mods:name>
<mods:name id="64A77924E92E6786093D88E45CB0B042" type="personal">
<mods:role id="7A9CE15A249083AE11185E40017C8718">
<mods:roleTerm id="D3636D135A3454368CE7F88FC5ACAF49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="354EE4E3E88D4B89BD0C591C01008D02">Suraj, S. R.</mods:namePart>
<mods:affiliation id="D23862FE17032483229F5BC8B40F4F05">Shola Nature Society, SRS Bhavan, Choodal, Meenmutty PO, Palode, Thiruvananthapuram</mods:affiliation>
</mods:name>
<mods:name id="DB246920752ED39CC6640D60D4B18AFF" type="personal">
<mods:role id="14A3B06F671C9685EA068DD717645479">
<mods:roleTerm id="B6176906F41C48E5BCD0A4B8FF3966BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E1DEA0977988234E157ED4EEC03EB5A">Jose, Subin K.</mods:namePart>
<mods:affiliation id="9EC0467D65FCC5E80DC3E75A54CB3FF3">Aqua Research Lab, Department of Geology and Environmental Science, Christ College (Autonomous), Irinjalakuda, Thrissur, Kerala 680125, India</mods:affiliation>
</mods:name>
<mods:name id="6433D638F1AB3C77D4A248585AD3DFBE" type="personal">
<mods:role id="C13585AAFE8600E0B08EA098679C3DC5">
<mods:roleTerm id="9C67AA557C8B0BBBC0FDC6C896BBEEA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="922A2D2BF4802A0642FA83A04280F9B5">Koparde, Pankaj</mods:namePart>
<mods:affiliation id="3ED4173BCBC23B29B4C1A6D228A98A66">Department of Environmental Studies, Dr. Vishwanath Karad MIT World Peace University, Kothrud, Pune, Maharashtra, 411038, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8D54F049869E26ECBF5F057442AA84B3">text</mods:typeOfResource>
<mods:relatedItem id="668FEA551ADA996075CEE17C9B48AFBA" type="host">
<mods:titleInfo id="9611881C364397E6354DE113218F43A6">
<mods:title id="F8D8481C37756FD876CA0BC2B757A8EC">International Journal of Odonatology</mods:title>
</mods:titleInfo>
<mods:part id="1357E0A7792B54912DAE3AD77FF02034">
<mods:date id="FC02C50B7AC73FC4E468096EC19FDB95">2024</mods:date>
<mods:detail id="1FE4FEE262F34C66A36F980A8302CAE5" type="pubDate">
<mods:number id="C72DE2055370D199D3B766A66B93FE56">2024-02-13</mods:number>
</mods:detail>
<mods:detail id="67B0C4BE078AF864B9B11FA954C60B9D" type="volume">
<mods:number id="C3B6B13140BD641EDA6F6D2967955354">27</mods:number>
</mods:detail>
<mods:extent id="8BA9F8199783CAD19876D95F98715D63" unit="page">
<mods:start id="B976A84B605208149A484A47D1D7C48E">26</mods:start>
<mods:end id="24AD31A7774D4A0518D8DFB824125283">36</mods:end>
</mods:extent>
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<mods:location id="DEBCE563EF2D7573084CDA88DEEAFF05">
<mods:url id="BECC98D2D24FC35829C7BA7A90427E00">https://doi.org/10.48156/1388.2024.1917259</mods:url>
</mods:location>
<mods:classification id="25166B1B5F16E73B7936F00680E6E9BA">journal article</mods:classification>
<mods:identifier id="65CC91866AE4C316164056456F0ADF5A" type="DOI">10.48156/1388.2024.1917259</mods:identifier>
<mods:identifier id="7BD2194AE171CAD4BBD42883F443157F" type="ISSN">2159-6719</mods:identifier>
<mods:identifier id="9BF2C0FDA841BE8D16114E553AEA0DA8" type="Zenodo-Dep">15555054</mods:identifier>
</mods:mods>
<treatment id="2D25DB6FC044296DFFA9FBA15DC8FB9C" LSID="urn:lsid:plazi:treatment:2D25DB6FC044296DFFA9FBA15DC8FB9C" httpUri="http://treatment.plazi.org/id/2D25DB6FC044296DFFA9FBA15DC8FB9C" lastPageId="9" lastPageNumber="35" pageId="2" pageNumber="28">
<subSubSection id="ED9639F2C0442966FFA9FBA15988FB19" pageId="2" pageNumber="28" type="nomenclature">
<paragraph id="A5336A79C0442966FFA9FBA15988FB19" blockId="2.[121,775,1144,1242]" pageId="2" pageNumber="28">
<heading id="FE7BDD15C0442966FFA9FBA15988FB19" centered="true" fontSize="11" level="2" pageId="2" pageNumber="28" reason="7">
<emphasis id="97F8B66BC0442966FFA9FBA15988FB19" bold="true" pageId="2" pageNumber="28">
<taxonomicName id="628C11FAC0442966FFA9FBA15A34FB3B" authority="Chandran, Chandran and Jose" authorityName="Chandran, Chandran and Jose" authorityYear="2024" box="[121,775,1144,1175]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0442966FFA9FBA158B2FB38" bold="true" box="[121,385,1147,1175]" italics="true" pageId="2" pageNumber="28">Phylloneura rupestris</emphasis>
Chandran, Chandran and Jose
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0442966FFA9FB405988FB19" box="[121,187,1178,1206]" pageId="2" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ED9639F2C0442966FFA9FB6458A3FB75" box="[121,400,1214,1242]" pageId="2" pageNumber="28" type="description">
<paragraph id="A5336A79C0442966FFA9FB6458A3FB75" blockId="2.[121,775,1144,1242]" box="[121,400,1214,1242]" pageId="2" pageNumber="28">
(
<figureCitation id="3DB776FCC0442966FFAFFB6459F5FB75" box="[127,198,1214,1242]" captionStart="Figure 1" captionStartId="1.[121,189,2051,2076]" captionTargetBox="[327,1262,679,2010]" captionTargetId="figure-264@1.[321,1267,676,2013]" captionTargetPageId="1" captionText="Figure 1. Map of the southern Western Ghats showing the collection localities of Phylloneura rupestris sp. n. (Merchiston estate, Ponmudi) and P. westermanni (Thirunelli, Wayanad)." figureDoi="http://doi.org/10.5281/zenodo.15555058" httpUri="https://zenodo.org/record/15555058/files/figure.png" pageId="2" pageNumber="28">Figs 1</figureCitation>
,
<figureCitation id="3DB776FCC0442966FF03FB6459C4FB75" box="[211,247,1214,1242]" captionStart="Figure 2" captionStartId="2.[121,189,2052,2077]" captionTargetBox="[121,1467,1582,2014]" captionTargetId="figure-636@2.[118,1470,1579,2017]" captionTargetPageId="2" captionText="Figure 2. Terminal abdominal segments of (A) Phylloneura rupestris sp. n. and (B) P. westermanni, illustrating the difference in markings on S7." figureDoi="http://doi.org/10.5281/zenodo.15555062" httpUri="https://zenodo.org/record/15555062/files/figure.png" pageId="2" pageNumber="28">2A</figureCitation>
,
<figureCitation id="3DB776FCC0442966FED1FB645820FB75" box="[257,275,1214,1242]" captionStart="Figure 3" captionStartId="3.[121,189,2054,2079]" captionTargetBox="[121,1467,144,2012]" captionTargetId="figure-28@3.[118,1469,141,1538]" captionTargetPageId="3" captionText="Figure 3. Phylloneura rupestris sp. n.: (A) in its habitat, (B) pterostigma, (C) caudal appendages, (D) genital ligula; P. wester- manni: (E) in its habitat, (F) pterostigma, (G) caudal appendages, (H) genital ligula." figureDoi="http://doi.org/10.5281/zenodo.15555064" httpUri="https://zenodo.org/record/15555064/files/figure.png" pageId="2" pageNumber="28">3</figureCitation>
,
<figureCitation id="3DB776FCC0442966FECEFB645872FB75" box="[286,321,1214,1242]" captionStart="Figure 4" captionStartId="4.[121,189,517,542]" captionTargetBox="[207,1383,147,477]" captionTargetId="figure-28@4.[201,1383,140,489]" captionTargetPageId="4" captionText="Figure 4. Caudal appendages of (A) Phylloneura rupestris sp. nov. and (B) P. westermanni, traced from original photographs." figureDoi="http://doi.org/10.5281/zenodo.15555066" httpUri="https://zenodo.org/record/15555066/files/figure.png" pageId="2" pageNumber="28">4A</figureCitation>
,
<figureCitation id="3DB776FCC0442966FE9CFB6458BAFB75" box="[332,393,1214,1242]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStart-3="Figure 9" captionStart-4="Figure 10" captionStartId-0="5.[121,189,2023,2048]" captionStartId-1="6.[121,189,2054,2079]" captionStartId-2="7.[121,189,1213,1238]" captionStartId-3="7.[813,881,2084,2109]" captionStartId-4="8.[121,189,1965,1990]" captionTargetBox-0="[121,1467,1099,1984]" captionTargetBox-1="[121,1467,1124,2016]" captionTargetBox-2="[250,1336,145,1165]" captionTargetBox-3="[813,1467,1315,2045]" captionTargetBox-4="[121,1466,144,1927]" captionTargetId-0="figure-392@5.[121,1467,1099,1984]" captionTargetId-1="figure-379@6.[121,1467,1101,2019]" captionTargetId-2="figure-220@7.[250,1337,144,1175]" captionTargetId-3="figure-258@7.[813,1470,1315,2046]" captionTargetId-4="figure-27@8.[121,1467,144,1927]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="Figure 6. Photographs of Phylloneura rupestris sp. n., holotype ♂ in the wild: (A) in its habitat, (B) dorsofrontal view of head, (C) dorsal view of head, prothorax and pterothorax, (D) ventrofrontal view of head, (E) lateral view of head, prothorax and pterothorax, (F) dorsal view of terminal segments, (G) lateral view of caudal appendages." captionText-1="Figure 7. Photographs of preserved Phylloneura rupestris sp. n. holotype ♂: (A) habitus, (B) wings, (C) dorsal view of caudal appendages, (D) lateral view of caudal appendages." captionText-2="Figure 8. Phylloneura rupestris sp. n., paratype ♀: (A) habitus, (B) lateral view of the terminal three abdominal segments, (C) wings, (D) frontal view of the head, (E) ventrolateral view of the terminal abdominal segments." captionText-3="Figure 9. Habitat of Phylloneura rupestris sp. n." captionText-4="Figure 10. Reproductive behaviour of Phylloneura rupestris sp. n.: (A) in tandem, (B) intra-male sperm translocation, (C) copula, (D) oviposition." figureDoi-0="http://doi.org/10.5281/zenodo.15555072" figureDoi-1="http://doi.org/10.5281/zenodo.15555074" figureDoi-2="http://doi.org/10.5281/zenodo.15555076" figureDoi-3="http://doi.org/10.5281/zenodo.15555078" figureDoi-4="http://doi.org/10.5281/zenodo.15555080" httpUri-0="https://zenodo.org/record/15555072/files/figure.png" httpUri-1="https://zenodo.org/record/15555074/files/figure.png" httpUri-2="https://zenodo.org/record/15555076/files/figure.png" httpUri-3="https://zenodo.org/record/15555078/files/figure.png" httpUri-4="https://zenodo.org/record/15555080/files/figure.png" pageId="2" pageNumber="28">610</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ED9639F2C0442966FFA9FADA5C17FF5D" pageId="2" pageNumber="28" type="materials_examined">
<paragraph id="A5336A79C0442966FFA9FADA5812FAB3" blockId="2.[121,289,1280,1341]" box="[121,289,1280,1308]" pageId="2" pageNumber="28">
<emphasis id="97F8B66BC0442966FFA9FADA5812FAB3" bold="true" box="[121,289,1280,1308]" italics="true" pageId="2" pageNumber="28">Type material</emphasis>
</paragraph>
<paragraph id="A5336A79C0442966FFA9FAF85832FA92" blockId="2.[121,289,1280,1341]" box="[121,257,1313,1341]" pageId="2" pageNumber="28">
<emphasis id="97F8B66BC0442966FFA9FAF859D7FA92" box="[121,228,1314,1341]" italics="true" pageId="2" pageNumber="28">
<typeStatus id="7A37D4DBC0442966FFA9FAF859D7FA92" box="[121,228,1314,1341]" pageId="2" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="B38AA1F0C0442966FF3BFAFB5832FA92" box="[235,257,1313,1341]" count="1" pageId="2" pageNumber="28" type="male"></specimenCount>
</paragraph>
<paragraph id="A5336A79C0442966FFA9FABE583CF9A9" blockId="2.[121,776,1380,1542]" pageId="2" pageNumber="28">
<materialsCitation id="15E46024C0442966FFA9FABE5838F9A9" collectingDate="2023-06-28" collectionCode="WGRC, ZSI" collectorName="Coll. A. Vivek Chandran" country="India" elevation="850" latitude="8.74969" location="Labelled" longLatPrecision="1" longitude="77.1271" pageId="2" pageNumber="28" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<location id="A0533CA2C0442966FFA9FABE59EEFA2F" LSID="urn:lsid:plazi:treatment:2D25DB6FC044296DFFA9FBA15DC8FB9C:A0533CA2C0442966FFA9FABE59EEFA2F" box="[121,221,1380,1408]" country="India" latitude="8.74969" longLatPrecision="1" longitude="77.1271" name="Labelled" pageId="2" pageNumber="28" stateProvince="Kerala">Labelled</location>
<typeStatus id="7A37D4DBC0442966FF26FABE5854FA2F" box="[246,359,1380,1408]" pageId="2" pageNumber="28" type="holotype">Holotype</typeStatus>
<taxonomicName id="628C11FAC0442966FEA7FABE5808FA0D" authority="Chandran, Chandran, Jose" authorityName="Chandran, Chandran, Jose" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="rupestris">
<emphasis id="97F8B66BC0442966FEA7FABE5B4FFAD0" box="[375,636,1380,1407]" italics="true" pageId="2" pageNumber="28">Phylloneura rupestris</emphasis>
Chandran, Chandran, Jose
</taxonomicName>
,
<collectingCountry id="DD9B2AE9C0442966FE9FFA5C58BDFA0D" box="[335,398,1414,1442]" name="India" pageId="2" pageNumber="28">India</collectingCountry>
,
<collectingRegion id="6748A49BC0442966FE73FA5C58C2FA0D" box="[419,497,1414,1442]" country="India" name="Kerala" pageId="2" pageNumber="28">Kerala</collectingRegion>
, Thiruvananthapuram, Ponmudi [
<geoCoordinate id="C0B80CBEC0442966FF24FA7D58A0FA6C" box="[244,403,1447,1475]" degrees="8.749690" direction="north" orientation="latitude" pageId="2" pageNumber="28" precision="1" value="8.74969">8.749690° N</geoCoordinate>
,
<geoCoordinate id="C0B80CBEC0442966FE73FA7D5B78FA6C" box="[419,587,1447,1475]" degrees="77.127098" direction="east" orientation="longitude" pageId="2" pageNumber="28" precision="1" value="77.1271">77.127098° E</geoCoordinate>
],
<elevation id="2EA18D4AC0442966FDB1FA7D59A3FA4B" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="28" unit="m" value="850.0">
elevation
<quantity id="6274C79CC0442966FD0AFA7D59A3FA4B" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="28" unit="m" value="850.0">850 m</quantity>
</elevation>
above MSL,
<date id="D1324CB9C0442966FEEAFA1258F6FA4B" box="[314,453,1480,1508]" pageId="2" pageNumber="28" value="2023-06-28">
<collectingDate id="C176B551C0442966FEEAFA1258F6FA4B" box="[314,453,1480,1508]" pageId="2" pageNumber="28" value="2023-06-28">28 vi 2023</collectingDate>
</date>
,
<collectorName id="08790FAFC0442966FE04FA125BCCFA4B" box="[468,767,1480,1508]" pageId="2" pageNumber="28">Coll. A. Vivek Chandran</collectorName>
” (
<collectionCode id="C39DF2BCC0442966FFAFFA3059E1F9A9" box="[127,210,1514,1542]" pageId="2" pageNumber="28">WGRC</collectionCode>
,
<collectionCode id="C39DF2BCC0442966FF0BFA305836F9A9" box="[219,261,1514,1542]" pageId="2" pageNumber="28">ZSI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A5336A79C0442966FCFDFF495A90FF01" blockId="2.[813,931,147,174]" box="[813,931,147,174]" pageId="2" pageNumber="28">
<emphasis id="97F8B66BC0442966FCFDFF495A90FF01" box="[813,931,147,174]" italics="true" pageId="2" pageNumber="28">
<typeStatus id="7A37D4DBC0442966FCFDFF495A90FF01" box="[813,931,147,174]" pageId="2" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="A5336A79C0442966FCFDFF0C5C17FF5D" blockId="2.[813,1316,213,242]" box="[813,1316,213,242]" pageId="2" pageNumber="28">
<materialsCitation id="15E46024C0442966FCFDFF0C5C13FF5D" box="[813,1312,213,242]" collectingDate="2023-06-28" collectionCode="WGRC, ZSI" collectorName="Coll. A. Vivek Chandran" country="India" elevation="850" latitude="8.74969" location="Labelled" longLatPrecision="1" longitude="77.1271" pageId="2" pageNumber="28" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kerala" typeStatus="paratype">
<specimenCount id="B38AA1F0C0442966FCFDFF0C5A5DFF5E" box="[813,878,213,242]" count="2" pageId="2" pageNumber="28" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="B38AA1F0C0442966FC78FF0C5AE7FF5E" box="[936,980,213,242]" count="1" pageId="2" pageNumber="28" type="female">1 ♀</specimenCount>
, same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ED9639F2C0442963FCFDFEC2583DF93F" lastPageId="7" lastPageNumber="33" pageId="2" pageNumber="28" type="diagnosis">
<paragraph id="A5336A79C0442966FCFDFEC25A90FE9B" blockId="2.[813,931,280,308]" box="[813,931,280,308]" pageId="2" pageNumber="28">
<emphasis id="97F8B66BC0442966FCFDFEC25A90FE9B" bold="true" box="[813,931,280,308]" italics="true" pageId="2" pageNumber="28">Diagnosis</emphasis>
</paragraph>
<paragraph id="A5336A79C0442966FCFDFE815D3DFC60" blockId="2.[813,1467,347,1542]" pageId="2" pageNumber="28">
<taxonomicName id="628C11FAC0442966FCFDFE815D1BFED9" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[813,1064,347,374]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0442966FCFDFE815D1BFED9" box="[813,1064,347,374]" italics="true" pageId="2" pageNumber="28">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0442966FBE0FE815D42FED8" box="[1072,1137,347,375]" pageId="2" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished in the field from its only congener,
<emphasis id="97F8B66BC0442966FB4CFEA65C6DFE38" box="[1180,1374,380,407]" italics="true" pageId="2" pageNumber="28">
<taxonomicName id="628C11FAC0442966FB4CFEA65C6AFE38" box="[1180,1369,380,407]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">P. westermanni</taxonomicName>
,
</emphasis>
by the broad azure blue marking on S7 that extends to 3/4 of the segment dorsolaterally (S7 has only an azure blue apical ring occupying 1/5 of the segment in
<taxonomicName id="628C11FAC0442966FB2DFE3A5C81FE54" box="[1277,1458,480,507]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FB2DFE3A5C81FE54" box="[1277,1458,480,507]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
) (
<figureCitation id="3DB776FCC0442966FCE4FDD85A4FFDB1" box="[820,892,514,542]" captionStart="Figure 2" captionStartId="2.[121,189,2052,2077]" captionTargetBox="[121,1467,1582,2014]" captionTargetId="figure-636@2.[118,1470,1579,2017]" captionTargetPageId="2" captionText="Figure 2. Terminal abdominal segments of (A) Phylloneura rupestris sp. n. and (B) P. westermanni, illustrating the difference in markings on S7." figureDoi="http://doi.org/10.5281/zenodo.15555062" httpUri="https://zenodo.org/record/15555062/files/figure.png" pageId="2" pageNumber="28">Fig. 2</figureCitation>
). The following features become evident upon close inspection of specimens and, in combination, help to distinguish
<taxonomicName id="628C11FAC0442966FBDBFD9E5DB9FDF0" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[1035,1162,580,607]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="rupestris" status="sp. n.">
<emphasis id="97F8B66BC0442966FBDBFD9E5DB9FDF0" box="[1035,1162,580,607]" italics="true" pageId="2" pageNumber="28">P. rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0442966FB40FD9E5DFCFDCF" box="[1168,1231,580,608]" pageId="2" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
from
<emphasis id="97F8B66BC0442966FAC4FD9E5A70FD2E" italics="true" pageId="2" pageNumber="28">P. westerman- ni</emphasis>
(
<figureCitation id="3DB776FCC0442966FC83FDBC5AACFD2D" box="[851,927,614,642]" captionStart="Figure 3" captionStartId="3.[121,189,2054,2079]" captionTargetBox="[121,1467,144,2012]" captionTargetId="figure-28@3.[118,1469,141,1538]" captionTargetPageId="3" captionText="Figure 3. Phylloneura rupestris sp. n.: (A) in its habitat, (B) pterostigma, (C) caudal appendages, (D) genital ligula; P. wester- manni: (E) in its habitat, (F) pterostigma, (G) caudal appendages, (H) genital ligula." figureDoi="http://doi.org/10.5281/zenodo.15555064" httpUri="https://zenodo.org/record/15555064/files/figure.png" pageId="2" pageNumber="28">Figs 3</figureCitation>
,
<figureCitation id="3DB776FCC0442966FC7EFDBC5AF2FD2D" box="[942,961,614,642]" captionStart="Figure 4" captionStartId="4.[121,189,517,542]" captionTargetBox="[207,1383,147,477]" captionTargetId="figure-28@4.[201,1383,140,489]" captionTargetPageId="4" captionText="Figure 4. Caudal appendages of (A) Phylloneura rupestris sp. nov. and (B) P. westermanni, traced from original photographs." figureDoi="http://doi.org/10.5281/zenodo.15555066" httpUri="https://zenodo.org/record/15555066/files/figure.png" pageId="2" pageNumber="28">4</figureCitation>
): Pt rectangular with a length of
<quantity id="6274C79CC0442966FA8CFDBC5C88FD2D" box="[1372,1467,614,642]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="28" unit="mm" value="1.5">1.5 mm</quantity>
(Pt squarish with a length of
<quantity id="6274C79CC0442966FB59FD5D5DFCFD0C" box="[1161,1231,647,675]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="28" unit="mm" value="1.0">1 mm</quantity>
in
<taxonomicName id="628C11FAC0442966FB24FD5D5C99FD0D" box="[1268,1450,647,674]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FB24FD5D5C99FD0D" box="[1268,1450,647,674]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
). Cerci thicker and with broad ventral processes (cerci thinner and with narrower ventral processes in
<emphasis id="97F8B66BC0442966FAB8FD105AA9FCA9" italics="true" pageId="2" pageNumber="28">P. wes- termanni</emphasis>
). Tips of paraprocts rounded (tips of paraprocts taper to acute points in
<taxonomicName id="628C11FAC0442966FB74FCD65C68FC88" box="[1188,1371,780,807]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FB74FCD65C68FC88" box="[1188,1371,780,807]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
). In the apical part of the males genital ligula, the two outer processes are hammer-shaped, and the two inner processes are thicker (tips of the two outer processes end as small lobes and the two inner processes are thinner in
<taxonomicName id="628C11FAC0442966FC99FC695ACDFC61" box="[841,1022,947,974]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FC99FC695ACDFC61" box="[841,1022,947,974]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="A5336A79C0442961FC9BFC0E59F7FED8" blockId="2.[813,1467,347,1542]" lastBlockId="5.[121,776,147,375]" lastPageId="5" lastPageNumber="31" pageId="2" pageNumber="28">
It must be noted here that in the original description of
<taxonomicName id="628C11FAC0442966FC9FFC2C5D34FBBE" box="[847,1031,1014,1041]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FC9FFC2C5D34FBBE" box="[847,1031,1014,1041]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
by
<bibRefCitation id="C11D1788C0442966FBE8FC2C5DE3FBBD" author="Fraser, F. C." box="[1080,1232,1014,1042]" pageId="2" pageNumber="28" pagination="1 - 9" refId="ref4409" refString="Fraser, F. C. (1922). New and rare Odonata from Nilgiri Hills. Records of Indian Museum 24 (1), 1-9. doi: 10.26515/rzsi/v24/ i 1 / 1922 / 163465" type="journal article" year="1922">Fraser (1922</bibRefCitation>
,
<bibRefCitation id="C11D1788C0442966FB32FC2C5C2EFBBD" author="Fraser, F. C." box="[1250,1309,1014,1042]" pageId="2" pageNumber="28" refId="ref4455" refString="Fraser, F. C. (1933). The fauna of British India including Ceylon and Burma, Odonata, Volume I. London: Taylor and Francis Ltd." type="book" year="1933">1933</bibRefCitation>
), the “apical half” of S7 is said to be azure blue. This, however, is not correct, and as is evident from photographs published later on (
<bibRefCitation id="C11D1788C0442966FC48FB805DBEFBD9" author="Gopalan, S. V. &amp; Sherif, M. &amp; Chandran, A. V." box="[920,1165,1114,1142]" pageId="2" pageNumber="28" pagination="20654 - 20665" refId="ref4537" refString="Gopalan, S. V., Sherif, M., Chandran, A. V. (2022). A checklist of dragonflies &amp; damselflies (Insecta: Odonata) of Kerala, India. Journal of Threatened Taxa, 14 (2), 20654-20665. doi: 10.11609 / jott. 7504.14.2.20654 - 20665" type="journal article" year="2022">Gopalan et al., 2022</bibRefCitation>
;
<bibRefCitation id="C11D1788C0442966FB4DFB805C6EFBD9" author="Nair, V. P. &amp; Samuel, K. A. &amp; Palot, M. J. &amp; Sadasivan, K." box="[1181,1373,1114,1142]" pageId="2" pageNumber="28" pagination="185 - 238" refId="ref4753" refString="Nair, V. P., Samuel, K. A., Palot, M. J. &amp; Sadasivan, K. (2021). The dragonflies and damselflies (Odonata) of Kerala - Status and distribution. Entomon, 46 (3), 185-238. doi: 10.33307/entomon. v 46 i 3.609" type="journal article" year="2021">Nair et al., 2021</bibRefCitation>
;
<bibRefCitation id="C11D1788C0442966FABDFB805AFCFB38" author="Subramanian, K. A." pageId="2" pageNumber="28" refId="ref5046" refString="Subramanian, K. A. (2009). Dragonflies and damselflies of Peninsu- lar India - A field guide. Nodia: Vigyan Prasar." type="book" year="2009">Subramanian, 2009</bibRefCitation>
,
<bibRefCitation id="C11D1788C0442966FC0DFBA15D2BFB38" author="Subramanian, K. &amp; Emiliyamma, K. G. &amp; Babu, R. &amp; Radhakrishnan, C. &amp; Talmale, S. S." box="[989,1048,1147,1175]" pageId="2" pageNumber="28" refId="ref5130" refString="Subramanian, K. A, Emiliyamma, K. G., Babu, R., Radhakrishnan, C. &amp; Talmale, S. S. (2018). Atlas of Odonata (Insecta) of the Western Ghats, India. Kolkata: Zoological Survey of India." type="book" year="2018">2018</bibRefCitation>
;
<bibRefCitation id="C11D1788C0442966FBF7FBA15C63FB38" author="Subramanian, K. A. &amp; Palot, M. &amp; Raju, D. V. &amp; Chandran, A. V. &amp; Gopalan, S. V. &amp; Sherif, M." box="[1063,1360,1147,1175]" pageId="2" pageNumber="28" pagination="212 - 219" refId="ref5183" refString="Subramanian, K. A., Palot, M. J, Raju, D. V., Chandran, A. V., Gopalan, S. V. &amp; Sherif, M. (2022). Dragonflies and damselflies. In: P. M. Sureshan, K. A. Subramanian &amp; M. J. Palot (Eds.), Threatened animals of Kerala. pp. 212-219. Kozhikode: Zoological Survey of India." type="book chapter" year="2022">Subramanian et al., 2022</bibRefCitation>
; all from north of the Palghat Gap) and specimens collected in this study, only the apical fifh of S7 is azure blue. To verify this further, we obtained images of the
<typeStatus id="7A37D4DBC0442966FA80FB055C89FB54" box="[1360,1466,1247,1275]" pageId="2" pageNumber="28" type="holotype">holotype</typeStatus>
of
<taxonomicName id="628C11FAC0442966FC80FADA5D3AFAB4" box="[848,1033,1280,1307]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FC80FADA5D3AFAB4" box="[848,1033,1280,1307]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
from the Museum of Comparative Zoology, Harvard University, Cambridge,
<collectingCountry id="DD9B2AE9C0442966FAF1FAF85C61FA91" box="[1313,1362,1314,1342]" name="United States of America" pageId="2" pageNumber="28">USA</collectingCountry>
(
<figureCitation id="3DB776FCC0442966FAB5FAF85C9DFA91" box="[1381,1454,1314,1342]" captionStart="Figure 5" captionStartId="4.[121,189,1992,2017]" captionTargetBox="[204,1383,604,1952]" captionTargetId="figure-49@4.[204,1383,604,1952]" captionTargetPageId="4" captionText="Figure 5. Images of the holotype of Phylloneura westermanni: (A) habitus, (B) label, (C) dorsal view of terminal segments, (D) lateral view of terminal segments, (E) dorsal view of caudal appendages, (F) lateral view of caudal appendages. Photos: Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College. https://creativecommons. org/licenses/by-nc-sa/4.0/." figureDoi="http://doi.org/10.5281/zenodo.15555068" httpUri="https://zenodo.org/record/15555068/files/figure.png" pageId="2" pageNumber="28">Fig. 5</figureCitation>
). It was thus confirmed that only the apical fifh of S7 is azure blue in
<taxonomicName id="628C11FAC0442966FC3AFABE5DACFAD0" box="[1002,1183,1380,1407]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FC3AFABE5DACFAD0" box="[1002,1183,1380,1407]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
and Fraser had erroneously stated that the apical half of S7 is blue. This error was probably carried forward from the original description by
<bibRefCitation id="C11D1788C0442966FC5DFA125D13FA4B" author="Selys Longchamps, E." box="[909,1056,1480,1508]" pageId="2" pageNumber="28" pagination="431 - 462" refId="ref5002" refString="Selys Longchamps, E. (1860). Synopsis des Agrionines. Derniere legion: Protoneura. Bulletin Academie royale Belgique Serie 2 10 (9 / 10), 431-462. doi: 10.5962/bhl.title.66021" type="journal article" year="1860">Selys (1860)</bibRefCitation>
who stated, “moitié finale du 7e et les trois derniers bleus” [final half of the 7
<superScript id="52F9C731C0442966FA98FA3D5C64FA59" attach="left" box="[1352,1367,1511,1526]" fontSize="6" pageId="2" pageNumber="28">th</superScript>
and the last three segments blue]. The watercolour painting in the Selys collection states “après la description”, which indicates that the illustration was made based on the description and without revisiting the
<typeStatus id="7A37D4DBC0432961FDB2FF2D5BFFFEBC" box="[610,716,247,275]" pageId="5" pageNumber="31" type="holotype">holotype</typeStatus>
and thus explains why this illustration shows the apical half of segment 7 being blue as well (
<bibRefCitation id="C11D1788C0432961FDDEFEE05987FED8" author="Verspui, K. &amp; Wasscher, M. T." pageId="5" pageNumber="31" pagination="125 - 156" refId="ref5271" refString="Verspui, K. &amp; Wasscher, M. T. (2016). The damselfly and dragonfly watercolour collection of Edmond de Selys Longchamps: I Agrionines. International Journal of Odonatology, 19 (3), 125-156. doi: 10.1080/13887890.2016.1184718" type="journal article" year="2016">Verspui &amp; Wasscher, 2016</bibRefCitation>
).
</paragraph>
<caption id="F1F33AF1C0442966FFA9F7DE580CF791" ID-DOI="http://doi.org/10.5281/zenodo.15555062" ID-Zenodo-Dep="15555062" httpUri="https://zenodo.org/record/15555062/files/figure.png" pageId="2" pageNumber="28" startId="2.[121,189,2052,2077]" targetBox="[121,1467,1582,2014]" targetPageId="2" targetType="figure">
<paragraph id="A5336A79C0442966FFA9F7DE580CF791" blockId="2.[121,1466,2052,2110]" pageId="2" pageNumber="28">
<emphasis id="97F8B66BC0442966FFA9F7DE59EAF7B2" bold="true" box="[121,217,2052,2077]" pageId="2" pageNumber="28">Figure 2.</emphasis>
Terminal abdominal segments of (A)
<taxonomicName id="628C11FAC0442966FDA2F7DC5A64F7B1" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[626,855,2054,2078]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0442966FDA2F7DC5A64F7B1" box="[626,855,2054,2078]" italics="true" pageId="2" pageNumber="28">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0442966FC8EF7DC5AAAF7B0" box="[862,921,2054,2079]" pageId="2" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
and (B)
<taxonomicName id="628C11FAC0442966FC26F7DC5DA8F7B1" box="[1014,1179,2054,2078]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0442966FC26F7DC5DA8F7B1" box="[1014,1179,2054,2078]" italics="true" pageId="2" pageNumber="28">P. westermanni</emphasis>
</taxonomicName>
, illustrating the difference in markings on S7.
</paragraph>
</caption>
<caption id="F1F33AF1C0452967FFA9F7DC5ADEF7EF" ID-DOI="http://doi.org/10.5281/zenodo.15555064" ID-Zenodo-Dep="15555064" httpUri="https://zenodo.org/record/15555064/files/figure.png" pageId="3" pageNumber="29" startId="3.[121,189,2054,2079]" targetBox="[121,1467,144,2012]" targetPageId="3" targetType="figure">
<paragraph id="A5336A79C0452967FFA9F7DC5ADEF7EF" blockId="3.[121,1466,2054,2112]" pageId="3" pageNumber="29">
<emphasis id="97F8B66BC0452967FFA9F7DC59EAF7B0" bold="true" box="[121,217,2054,2079]" pageId="3" pageNumber="29">Figure 3.</emphasis>
<taxonomicName id="628C11FAC0452967FF31F7D358F5F78E" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[225,454,2057,2081]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0452967FF31F7D358F5F78E" box="[225,454,2057,2081]" italics="true" pageId="3" pageNumber="29">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0452967FE1DF7D35B38F78D" box="[461,523,2057,2082]" pageId="3" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
: (A) in its habitat, (B) pterostigma, (C) caudal appendages, (D) genital ligula;
<emphasis id="97F8B66BC0452967FA82F7D3598EF790" italics="true" pageId="3" pageNumber="29">P. wester- manni</emphasis>
: (E) in its habitat, (F) pterostigma, (G) caudal appendages, (H) genital ligula.
</paragraph>
</caption>
<caption id="F1F33AF1C0422960FFA9FDDF5C9FFD8E" ID-DOI="http://doi.org/10.5281/zenodo.15555066" ID-Zenodo-Dep="15555066" box="[121,1452,517,545]" httpUri="https://zenodo.org/record/15555066/files/figure.png" pageId="4" pageNumber="30" startId="4.[121,189,517,542]" targetBox="[207,1383,147,477]" targetPageId="4" targetType="figure">
<paragraph id="A5336A79C0422960FFA9FDDF5C9FFD8E" blockId="4.[121,1452,517,545]" box="[121,1452,517,545]" pageId="4" pageNumber="30">
<emphasis id="97F8B66BC0422960FFA9FDDF59EBFDB1" bold="true" box="[121,216,517,542]" pageId="4" pageNumber="30">Figure 4.</emphasis>
Caudal appendages of (A)
<taxonomicName id="628C11FAC0422960FE29FDD25BEEFD8F" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[505,733,520,544]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0422960FE29FDD25BEEFD8F" box="[505,733,520,544]" italics="true" pageId="4" pageNumber="30">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0422960FD33FDD25A06FD8E" box="[739,821,520,545]" pageId="4" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and (B)
<taxonomicName id="628C11FAC0422960FC5EFDD25D01FD8F" box="[910,1074,520,544]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0422960FC5EFDD25D01FD8F" box="[910,1074,520,544]" italics="true" pageId="4" pageNumber="30">P. westermanni</emphasis>
</taxonomicName>
, traced from original photographs.
</paragraph>
</caption>
<caption id="F1F33AF1C0422960FFA9F81258ACF7EF" ID-DOI="http://doi.org/10.5281/zenodo.15555068" ID-Zenodo-Dep="15555068" httpUri="https://zenodo.org/record/15555068/files/figure.png" pageId="4" pageNumber="30" startId="4.[121,189,1992,2017]" targetBox="[204,1383,604,1952]" targetPageId="4" targetType="figure">
<paragraph id="A5336A79C0422960FFA9F81258ACF7EF" blockId="4.[121,1468,1992,2112]" pageId="4" pageNumber="30">
<emphasis id="97F8B66BC0422960FFA9F81259E9F84E" bold="true" box="[121,218,1992,2017]" pageId="4" pageNumber="30">Figure 5.</emphasis>
Images of the holotype of
<taxonomicName id="628C11FAC0422960FDD9F8115A2DF84C" box="[521,798,1995,2019]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0422960FDD9F8115A2DF84C" box="[521,798,1995,2019]" italics="true" pageId="4" pageNumber="30">Phylloneura westermanni</emphasis>
</taxonomicName>
: (A) habitus, (B) label, (C) dorsal view of terminal segments, (D) lateral view of terminal segments, (E) dorsal view of caudal appendages, (F) lateral view of caudal appendages. Photos: Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College. https://creativecommons. org/licenses/by-nc-sa/4.0/.
</paragraph>
</caption>
<paragraph id="A5336A79C0432961FFA9FE655B75FE75" blockId="5.[121,582,447,474]" box="[121,582,447,474]" pageId="5" pageNumber="31">
<emphasis id="97F8B66BC0432961FFA9FE655B75FE75" box="[121,582,447,474]" italics="true" pageId="5" pageNumber="31">
Description of the
<typeStatus id="7A37D4DBC0432961FE82FE655889FE75" box="[338,442,447,474]" pageId="5" pageNumber="31" type="holotype">holotype</typeStatus>
(
<figureCitation id="3DB776FCC0432961FE19FE655B3EFE75" box="[457,525,447,474]" captionStart="Figure 6" captionStartId="5.[121,189,2023,2048]" captionTargetBox="[121,1467,1099,1984]" captionTargetId="figure-392@5.[121,1467,1099,1984]" captionTargetPageId="5" captionText="Figure 6. Photographs of Phylloneura rupestris sp. n., holotype ♂ in the wild: (A) in its habitat, (B) dorsofrontal view of head, (C) dorsal view of head, prothorax and pterothorax, (D) ventrofrontal view of head, (E) lateral view of head, prothorax and pterothorax, (F) dorsal view of terminal segments, (G) lateral view of caudal appendages." figureDoi="http://doi.org/10.5281/zenodo.15555072" httpUri="https://zenodo.org/record/15555072/files/figure.png" pageId="5" pageNumber="31">Figs 6</figureCitation>
&amp;
<figureCitation id="3DB776FCC0432961FDFFFE655B0DFE75" box="[559,574,447,474]" captionStart="Figure 7" captionStartId="6.[121,189,2054,2079]" captionTargetBox="[121,1467,1124,2016]" captionTargetId="figure-379@6.[121,1467,1101,2019]" captionTargetPageId="6" captionText="Figure 7. Photographs of preserved Phylloneura rupestris sp. n. holotype ♂: (A) habitus, (B) wings, (C) dorsal view of caudal appendages, (D) lateral view of caudal appendages." figureDoi="http://doi.org/10.5281/zenodo.15555074" httpUri="https://zenodo.org/record/15555074/files/figure.png" pageId="5" pageNumber="31">7</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="A5336A79C0432961FFA9FDD8584CFD90" blockId="5.[121,775,514,642]" pageId="5" pageNumber="31">
Male. Total length
<quantity id="6274C79CC0432961FEA3FDD858E7FDB1" box="[371,468,514,542]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="31" unit="mm" value="60.0">60 mm</quantity>
; abdomen
<quantity id="6274C79CC0432961FDBAFDD85BF8FDB1" box="[618,715,514,542]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="31" unit="mm" value="49.0">49 mm</quantity>
; FW
<quantity id="6274C79CC0432961FFA9FDF959D4FD90" box="[121,231,547,575]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="5" pageNumber="31" unit="mm" value="37.5">37.5 mm</quantity>
; HW
<quantity id="6274C79CC0432961FEF4FDF95848FD90" box="[292,379,547,575]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="31" unit="mm" value="36.0">36 mm</quantity>
.
</paragraph>
<paragraph id="A5336A79C0432961FF47FD9E59F8FD2D" blockId="5.[121,775,514,642]" pageId="5" pageNumber="31">Repository reference number: ZSI/WGRC/I.R.-INV 25601.</paragraph>
<paragraph id="A5336A79C0432961FFA9FD7259F7FBBD" blockId="5.[121,776,680,1042]" pageId="5" pageNumber="31">Head. Middle and lateral lobes of labium white, tipped with black; labrum, anteclypeus and postclypeus steely blue-black, outer margin of postclypeus projecting as short horns at both ends; bases of mandibles black, with a central blue spot; genae blue, with a black band which is confluent with the black of the clypeus; antefrons black, but the blue of the genae spreads onto it narrowly; postfrons black throughout; rest of the head black, including its ventral side; lower 2/3 of the eyes ultramarine blue, capped with black; ocelli translucent white.</paragraph>
<paragraph id="A5336A79C0432961FCFDFF495C05FEF9" blockId="5.[813,1467,147,342]" pageId="5" pageNumber="31">Prothorax. Posterior lobe black throughout; middle lobe almost completely black dorsally, but with a short ingress of azure blue from the anterior lobe, azure blue laterally; propleuron azure blue; anterior lobe almost completely azure blue, but with a short ingress of black from the middle lobe at two points dorsally.</paragraph>
<paragraph id="A5336A79C0432961FCFDFEA65A92FD2D" blockId="5.[813,1467,380,642]" pageId="5" pageNumber="31">Pterothorax. Jet black marked with azure blue as follows: middorsal carina and mesepisternum black; very narrow antehumeral blue stripes; mesepimeron black with a white spot at the anterior end on each side; metepisternum black with a broad blue stripe; mesinfraepisternum black, with only its lower end being blue; metepimeron blue; underside of thorax bluish white.</paragraph>
<paragraph id="A5336A79C0432961FCFDFD725DFAFD49" blockId="5.[813,1467,680,742]" pageId="5" pageNumber="31">Legs. Black, coxae and basal halves of trochanters blue; extensor face of tibiae pale yellow.</paragraph>
<paragraph id="A5336A79C0432961FCFDFCD65DE1FC5F" blockId="5.[813,1467,780,1008]" pageId="5" pageNumber="31">
Wings. Hyaline, lightly smoky; Ab complete, arched at origin, inclining towards the rear border of the wing distally; Pt dark reddish brown, rectangular,
<quantity id="6274C79CC0432961FA90FC955CAFFCC4" box="[1344,1436,847,875]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="31" unit="mm" value="1.5">1.5 mm</quantity>
in length, covering 3 cells, braced between thick, black nervures; Ax
<quantity id="6274C79CC0432961FC1BFC485AC4FC01" box="[971,1015,914,942]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="31" unit="in" value="2.0">2 in</quantity>
all wings; Px
<quantity id="6274C79CC0432961FB4AFC485DE6FC01" box="[1178,1237,914,942]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="5" pageNumber="31" unit="in" value="33.0">33 in</quantity>
lef FW,
<quantity id="6274C79CC0432961FAEFFC485C48FC01" box="[1343,1403,914,942]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="5" pageNumber="31" unit="in" value="32.0">32 in</quantity>
right FW,
<quantity id="6274C79CC0432961FCB4FC695A90FC60" box="[868,931,947,975]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="5" pageNumber="31" unit="in" value="29.0">29 in</quantity>
both HWs; many double cells in all wings, mostly towards the tips of the FWs.
</paragraph>
<caption id="F1F33AF1C0432961FFA9F83D5D0AF7EF" ID-DOI="http://doi.org/10.5281/zenodo.15555072" ID-Zenodo-Dep="15555072" httpUri="https://zenodo.org/record/15555072/files/figure.png" pageId="5" pageNumber="31" startId="5.[121,189,2023,2048]" targetBox="[121,1467,1099,1984]" targetPageId="5" targetType="figure">
<paragraph id="A5336A79C0432961FFA9F83D5D0AF7EF" blockId="5.[121,1468,2023,2112]" pageId="5" pageNumber="31">
<emphasis id="97F8B66BC0432961FFA9F83D59EBF7AF" bold="true" box="[121,216,2023,2048]" pageId="5" pageNumber="31">Figure 6.</emphasis>
Photographs of
<taxonomicName id="628C11FAC0432961FE5BF8305B5CF7AD" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[395,623,2026,2050]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0432961FE5BF8305B5CF7AD" box="[395,623,2026,2050]" italics="true" pageId="5" pageNumber="31">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0432961FDA5F8305B81F7AC" box="[629,690,2026,2051]" pageId="5" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
, holotype ♂ in the wild: (A) in its habitat, (B) dorsofrontal view of head, (C) dorsal view of head, prothorax and pterothorax, (D) ventrofrontal view of head, (E) lateral view of head, prothorax and pterothorax, (F) dorsal view of terminal segments, (G) lateral view of caudal appendages.
</paragraph>
</caption>
<paragraph id="A5336A79C0402962FFA9FF4959ECFD90" blockId="6.[121,775,147,575]" pageId="6" pageNumber="32">Abdomen. Black, marked with blue as follows: S1 broadly blue, with black basal and apical margins, the former turning into a median spot dorsally; S2 with a broad blue ventrolateral spot; S3 and S4 have very narrow basal annules; S5 has an almost invisible basal spot; S6 fully black; S7 with a broad black apical margin from where a large blue stripe extends to 3/4 of the segment towards its base, this stripe is split centrally from half its length; S8 and S9 entirely blue on dorsum, except for very narrow basal black annules, laterally almost fully blue, but with irregular infusions of black towards the bases and apices; S10 blue on dorsum, black laterally.</paragraph>
<paragraph id="A5336A79C0402962FFA9FDBC58AEFCE5" blockId="6.[121,776,614,842]" pageId="6" pageNumber="32">Caudal appendages. Black, 1.5 times the length of S10, covered irregularly with light brown hairs; cerci broad at their bases, with an upper obtuse tubercle at the base and another at the tip, a broad ventral process at its apical 3/4 portion; paraprocts broad at their bases, sloping downwards and tapering gradually, tips round- ed and directed upward.</paragraph>
<paragraph id="A5336A79C0402962FFA9FCAA59CAFC24" blockId="6.[121,249,880,907]" box="[121,249,880,907]" pageId="6" pageNumber="32">
<emphasis id="97F8B66BC0402962FFA9FCAA59CAFC24" box="[121,249,880,907]" italics="true" pageId="6" pageNumber="32">
<typeStatus id="7A37D4DBC0402962FFA9FCAA59D7FC24" box="[121,228,880,907]" pageId="6" pageNumber="32" type="paratype">Paratype</typeStatus>
1
</emphasis>
</paragraph>
<paragraph id="A5336A79C0402962FFA9FC69585AFC5F" blockId="6.[121,775,947,1075]" pageId="6" pageNumber="32">
Male. Total length
<quantity id="6274C79CC0402962FEA3FC6958E7FC60" box="[371,468,947,975]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="32" unit="mm" value="60.0">60 mm</quantity>
; abdomen
<quantity id="6274C79CC0402962FDBAFC695BF8FC60" box="[618,715,947,975]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="32" unit="mm" value="49.0">49 mm</quantity>
; FW
<quantity id="6274C79CC0402962FFA9FC0E59E3FC5F" box="[121,208,980,1008]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="32" unit="mm" value="37.0">37 mm</quantity>
; HW
<quantity id="6274C79CC0402962FEDEFC0E5856FC5F" box="[270,357,980,1008]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="32" unit="mm" value="35.0">35 mm</quantity>
.
</paragraph>
<paragraph id="A5336A79C0402962FF47FC2C59FFFB9C" blockId="6.[121,775,947,1075]" pageId="6" pageNumber="32">Repository reference number: ZSI/WGRC/I.R.-INV 25602.</paragraph>
<paragraph id="A5336A79C0402962FCFDFF495D99FF7F" blockId="6.[813,1467,147,208]" pageId="6" pageNumber="32">
Px
<quantity id="6274C79CC0402962FC9EFF495ABBFF00" box="[846,904,147,175]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="32" unit="in" value="30.0">30 in</quantity>
both FWs,
<quantity id="6274C79CC0402962FBDBFF495D76FF00" box="[1035,1093,147,175]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="6" pageNumber="32" unit="in" value="27.0">27 in</quantity>
lef HW and
<quantity id="6274C79CC0402962FB0DFF495C24FF00" box="[1245,1303,147,175]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="6" pageNumber="32" unit="in" value="26.0">26 in</quantity>
right HW; the rest agreeing with the
<typeStatus id="7A37D4DBC0402962FBE8FF6E5D95FF7F" box="[1080,1190,180,208]" pageId="6" pageNumber="32" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="A5336A79C0402962FCFDFF2D5C2BFE37" blockId="6.[813,1467,247,408]" pageId="6" pageNumber="32">
Genital ligula. As illustrated in
<figureCitation id="3DB776FCC0402962FB60FF2D5C39FEBC" box="[1200,1290,247,275]" captionStart="Figure 3" captionStartId="3.[121,189,2054,2079]" captionTargetBox="[121,1467,144,2012]" captionTargetId="figure-28@3.[118,1469,141,1538]" captionTargetPageId="3" captionText="Figure 3. Phylloneura rupestris sp. n.: (A) in its habitat, (B) pterostigma, (C) caudal appendages, (D) genital ligula; P. wester- manni: (E) in its habitat, (F) pterostigma, (G) caudal appendages, (H) genital ligula." figureDoi="http://doi.org/10.5281/zenodo.15555064" httpUri="https://zenodo.org/record/15555064/files/figure.png" pageId="6" pageNumber="32">Fig. 3D</figureCitation>
, first segment broad and curved; second segment short and featureless; tip of third segment projecting into four filaments, the two outer ones hammer-shaped, the two inner ones thick, conical, curved slightly at tips.
</paragraph>
<paragraph id="A5336A79C0402962FCFDFE655A9EFE75" blockId="6.[813,941,447,474]" box="[813,941,447,474]" pageId="6" pageNumber="32">
<emphasis id="97F8B66BC0402962FCFDFE655A9EFE75" box="[813,941,447,474]" italics="true" pageId="6" pageNumber="32">
<typeStatus id="7A37D4DBC0402962FCFDFE655AABFE75" box="[813,920,447,474]" pageId="6" pageNumber="32" type="paratype">Paratype</typeStatus>
2
</emphasis>
</paragraph>
<paragraph id="A5336A79C0402962FCFDFDD85D00FD90" blockId="6.[813,1467,514,642]" pageId="6" pageNumber="32">
Male. Total length
<quantity id="6274C79CC0402962FBF6FDD85DB4FDB1" box="[1062,1159,514,542]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="32" unit="mm" value="60.0">60 mm</quantity>
; abdomen
<quantity id="6274C79CC0402962FACEFDD85C4CFDB1" box="[1310,1407,514,542]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="32" unit="mm" value="49.0">49 mm</quantity>
; FW
<quantity id="6274C79CC0402962FCFDFDF95AB7FD90" box="[813,900,547,575]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="32" unit="mm" value="36.0">36 mm</quantity>
; HW
<quantity id="6274C79CC0402962FC11FDF95D1CFD90" box="[961,1071,547,575]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" pageId="6" pageNumber="32" unit="mm" value="34.5">34.5 mm</quantity>
.
</paragraph>
<paragraph id="A5336A79C0402962FC9BFD9E5A4CFD2D" blockId="6.[813,1467,514,642]" pageId="6" pageNumber="32">Repository reference number: ZSI/WGRC/I.R.-INV 25603.</paragraph>
<paragraph id="A5336A79C0402962FCFDFD725C63FD49" blockId="6.[813,1466,680,742]" pageId="6" pageNumber="32">
Px
<quantity id="6274C79CC0402962FC9FFD725AB9FD6B" box="[847,906,680,708]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="6" pageNumber="32" unit="in" value="31.0">31 in</quantity>
lef FW,
<quantity id="6274C79CC0402962FC22FD725D1FFD6B" box="[1010,1068,680,708]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="32" unit="in" value="30.0">30 in</quantity>
right FW,
<quantity id="6274C79CC0402962FB75FD725DECFD6B" box="[1189,1247,680,708]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="6" pageNumber="32" unit="in" value="29.0">29 in</quantity>
lef HW, and
<quantity id="6274C79CC0402962FA50FD725C89FD6B" box="[1408,1466,680,708]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="6" pageNumber="32" unit="in" value="26.0">26 in</quantity>
right HW; the rest agreeing with the
<typeStatus id="7A37D4DBC0402962FB0EFD105C7FFD49" box="[1246,1356,714,742]" pageId="6" pageNumber="32" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="A5336A79C0402962FCFDFCD65D36FC88" blockId="6.[813,1029,780,807]" box="[813,1029,780,807]" pageId="6" pageNumber="32">
<emphasis id="97F8B66BC0402962FCFDFCD65D36FC88" box="[813,1029,780,807]" italics="true" pageId="6" pageNumber="32">
<typeStatus id="7A37D4DBC0402962FCFDFCD65AABFC88" box="[813,920,780,807]" pageId="6" pageNumber="32" type="paratype">Paratype</typeStatus>
3 (
<figureCitation id="3DB776FCC0402962FC6DFCD65ACEFC88" box="[957,1021,780,807]" captionStart="Figure 8" captionStartId="7.[121,189,1213,1238]" captionTargetBox="[250,1336,145,1165]" captionTargetId="figure-220@7.[250,1337,144,1175]" captionTargetPageId="7" captionText="Figure 8. Phylloneura rupestris sp. n., paratype ♀: (A) habitus, (B) lateral view of the terminal three abdominal segments, (C) wings, (D) frontal view of the head, (E) ventrolateral view of the terminal abdominal segments." figureDoi="http://doi.org/10.5281/zenodo.15555076" httpUri="https://zenodo.org/record/15555076/files/figure.png" pageId="6" pageNumber="32">Fig. 8</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="A5336A79C0402962FCFDFC955D7AFC23" blockId="6.[813,1467,847,975]" pageId="6" pageNumber="32">
Female. Total length
<quantity id="6274C79CC0402962FBE7FC955DA7FCC4" box="[1079,1172,847,875]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="6" pageNumber="32" unit="mm" value="58.0">58 mm</quantity>
; abdomen
<quantity id="6274C79CC0402962FAF4FC955CB1FCC4" box="[1316,1410,847,875]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="32" unit="mm" value="47.0">47 mm</quantity>
; FW
<quantity id="6274C79CC0402962FCFDFCAA5AA9FC23" box="[813,922,880,908]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="6" pageNumber="32" unit="mm" value="36.5">36.5 mm</quantity>
; HW
<quantity id="6274C79CC0402962FC08FCAA5D76FC23" box="[984,1093,880,908]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" pageId="6" pageNumber="32" unit="mm" value="34.5">34.5 mm</quantity>
.
</paragraph>
<paragraph id="A5336A79C0402962FC9BFC485A4CFC60" blockId="6.[813,1467,847,975]" pageId="6" pageNumber="32">Repository reference number: ZSI/WGRC/I.R.-INV 25604.</paragraph>
<paragraph id="A5336A79C0402963FCFDFC2C5A35FAEC" blockId="6.[813,1466,1014,1075]" lastBlockId="7.[121,775,1319,1447]" lastPageId="7" lastPageNumber="33" pageId="6" pageNumber="32">Similar to the male, except for sexual dimorphic differences and blue markings on the terminal abdominal segments. Abdomen shorter and more robust; S7 with an</paragraph>
<caption id="F1F33AF1C0402962FFA9F7DC5B95F7EF" ID-DOI="http://doi.org/10.5281/zenodo.15555074" ID-Zenodo-Dep="15555074" httpUri="https://zenodo.org/record/15555074/files/figure.png" pageId="6" pageNumber="32" startId="6.[121,189,2054,2079]" targetBox="[121,1467,1124,2016]" targetPageId="6" targetType="figure">
<paragraph id="A5336A79C0402962FFA9F7DC5B95F7EF" blockId="6.[121,1467,2054,2112]" pageId="6" pageNumber="32">
<emphasis id="97F8B66BC0402962FFA9F7DC59EAF7B0" bold="true" box="[121,217,2054,2079]" pageId="6" pageNumber="32">Figure 7.</emphasis>
Photographs of preserved
<taxonomicName id="628C11FAC0402962FDD1F7D35BD5F78E" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[513,742,2057,2081]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0402962FDD1F7D35BD5F78E" box="[513,742,2057,2081]" italics="true" pageId="6" pageNumber="32">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0402962FD3DF7D35A14F78D" box="[749,807,2057,2082]" pageId="6" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
holotype ♂: (A) habitus, (B) wings, (C) dorsal view of caudal appendages, (D) lateral view of caudal appendages.
</paragraph>
</caption>
<caption id="F1F33AF1C0412963FFA9FB675DAEFB58" ID-DOI="http://doi.org/10.5281/zenodo.15555076" ID-Zenodo-Dep="15555076" httpUri="https://zenodo.org/record/15555076/files/figure.png" pageId="7" pageNumber="33" startId="7.[121,189,1213,1238]" targetBox="[250,1336,145,1165]" targetPageId="7" targetType="figure">
<paragraph id="A5336A79C0412963FFA9FB675DAEFB58" blockId="7.[121,1468,1213,1271]" pageId="7" pageNumber="33">
<emphasis id="97F8B66BC0412963FFA9FB6759E8FB79" bold="true" box="[121,219,1213,1238]" pageId="7" pageNumber="33">Figure 8.</emphasis>
<taxonomicName id="628C11FAC0412963FF33FB6558FAFB78" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[227,457,1215,1239]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0412963FF33FB6558FAFB78" box="[227,457,1215,1239]" italics="true" pageId="7" pageNumber="33">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0412963FE01FB655B23FB77" box="[465,528,1215,1240]" pageId="7" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
, paratype ♀: (A) habitus, (B) lateral view of the terminal three abdominal segments, (C) wings, (D) frontal view of the head, (E) ventrolateral view of the terminal abdominal segments.
</paragraph>
</caption>
<paragraph id="A5336A79C0412963FFA9FA925A34FACB" blockId="7.[121,775,1319,1447]" box="[121,775,1352,1380]" pageId="7" pageNumber="33">apical blue ring, thinner on the dorsum, expanding later-</paragraph>
<paragraph id="A5336A79C0412963FFA9FAB35A34FA2A" blockId="7.[121,775,1319,1447]" box="[121,775,1385,1413]" pageId="7" pageNumber="33">ally, but not joining ventrally; S8 and S9 with large trian-</paragraph>
<paragraph id="A5336A79C0412963FFA9FA515A33FA08" blockId="7.[121,775,1319,1447]" box="[121,768,1419,1447]" pageId="7" pageNumber="33">gular dorsal blue spots; dorsum of S10 completely blue.</paragraph>
<paragraph id="A5336A79C0412963FFA9FA175A34FA46" blockId="7.[121,775,1485,1613]" box="[121,775,1485,1513]" pageId="7" pageNumber="33">Caudal appendages. Black, covered with light brown</paragraph>
<paragraph id="A5336A79C0412963FFA9FA355A34F9A4" blockId="7.[121,775,1485,1613]" box="[121,775,1519,1547]" pageId="7" pageNumber="33">hairs; cerci short and conical; vulvar scale blue, tipped</paragraph>
<paragraph id="A5336A79C0412963FFA9F9CA5A34F983" blockId="7.[121,775,1485,1613]" box="[121,775,1552,1580]" pageId="7" pageNumber="33">with black, not reaching the end of the abdomen; ovi-</paragraph>
<paragraph id="A5336A79C0412963FFA9F9EB5846F9E2" blockId="7.[121,775,1485,1613]" box="[121,373,1585,1613]" pageId="7" pageNumber="33">positor black, robust.</paragraph>
<paragraph id="A5336A79C0412963FFA9F9AE583DF93F" blockId="7.[121,270,1652,1680]" box="[121,270,1652,1680]" pageId="7" pageNumber="33">
<emphasis id="97F8B66BC0412963FFA9F9AE583DF93F" bold="true" box="[121,270,1652,1680]" italics="true" pageId="7" pageNumber="33">Annotations</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="ED9639F2C0412963FFA9F96D5B39F7EE" pageId="7" pageNumber="33" type="description">
<paragraph id="A5336A79C0412963FFA9F96D5B39F7EE" blockId="7.[121,776,1719,2113]" pageId="7" pageNumber="33">
Male
<taxonomicName id="628C11FAC0412963FF10F96D588DF97D" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[192,446,1719,1746]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0412963FF10F96D588DF97D" box="[192,446,1719,1746]" italics="true" pageId="7" pageNumber="33">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0412963FE18F96D5B3FF97C" box="[456,524,1719,1747]" pageId="7" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
can at once be told apart in the field from male
<taxonomicName id="628C11FAC0412963FE32F9025BA8F95C" box="[482,667,1752,1779]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0412963FE32F9025BA8F95C" box="[482,667,1752,1779]" italics="true" pageId="7" pageNumber="33">P. westermanni</emphasis>
</taxonomicName>
by their extensive blue marking on S7, which in the latter is reduced to a thick apical ring. Female
<taxonomicName id="628C11FAC0412963FDE7F8C15B89F899" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[567,698,1819,1846]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestris" status="sp. n.">
<emphasis id="97F8B66BC0412963FDE7F8C15B89F899" box="[567,698,1819,1846]" italics="true" pageId="7" pageNumber="33">P. rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0412963FD14F8C15A34F898" box="[708,775,1819,1847]" pageId="7" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
have a distinctive apical blue ring on S7 as well, which is obscure in the female
<taxonomicName id="628C11FAC0412963FE71F8875B64F8D7" box="[417,599,1885,1912]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC0412963FE71F8875B64F8D7" box="[417,599,1885,1912]" italics="true" pageId="7" pageNumber="33">P. westermanni</emphasis>
</taxonomicName>
. When a male specimen is held in the hand, a close look at the caudal appendages with a magnifying lens helps with confirming its specifics identity, as
<taxonomicName id="628C11FAC0412963FE66F81B5B05F873" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[438,566,1985,2012]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestris" status="sp. n.">
<emphasis id="97F8B66BC0412963FE66F81B5B05F873" box="[438,566,1985,2012]" italics="true" pageId="7" pageNumber="33">P. rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0412963FDEDF81B5B4FF872" box="[573,636,1985,2013]" pageId="7" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
has considerably thicker cerci and rounded tips of its paraprocts. Moreover, as per current knowledge, the ranges of these two species do not overlap.
</paragraph>
</subSubSection>
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<paragraph id="A5336A79C0412963FCFDF7FE5C14F7EF" blockId="7.[813,1319,2084,2112]" box="[813,1319,2084,2112]" pageId="7" pageNumber="33">
<emphasis id="97F8B66BC0412963FCFDF7FE5ABFF792" bold="true" box="[813,908,2084,2109]" pageId="7" pageNumber="33">Figure 9.</emphasis>
Habitat of
<taxonomicName id="628C11FAC0412963FBD3F7FD5DD4F790" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[1027,1255,2087,2111]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC0412963FBD3F7FD5DD4F790" box="[1027,1255,2087,2111]" italics="true" pageId="7" pageNumber="33">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C0412963FB3DF7FD5C14F7EF" box="[1261,1319,2087,2112]" pageId="7" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="F1F33AF1C04E296CFFA9F877587EF847" ID-DOI="http://doi.org/10.5281/zenodo.15555080" ID-Zenodo-Dep="15555080" httpUri="https://zenodo.org/record/15555080/files/figure.png" pageId="8" pageNumber="34" startId="8.[121,189,1965,1990]" targetBox="[121,1466,144,1927]" targetPageId="8" targetType="figure">
<paragraph id="A5336A79C04E296CFFA9F877587EF847" blockId="8.[121,1466,1965,2024]" pageId="8" pageNumber="34">
<emphasis id="97F8B66BC04E296CFFA9F87759D6F869" bold="true" box="[121,229,1965,1990]" pageId="8" pageNumber="34">Figure 10.</emphasis>
Reproductive behaviour of
<taxonomicName id="628C11FAC04E296CFDC2F86A5BC5F867" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[530,758,1968,1992]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="rupestris" status="sp. nov.">
<emphasis id="97F8B66BC04E296CFDC2F86A5BC5F867" box="[530,758,1968,1992]" italics="true" pageId="8" pageNumber="34">Phylloneura rupestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CCB0B10C04E296CFD2DF86A5A09F866" box="[765,826,1968,1993]" pageId="8" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
: (A) in tandem, (B) intra-male sperm translocation, (C) copula, (D) oviposition.
</paragraph>
</caption>
<subSubSection id="ED9639F2C04F296DFFA9FF495B31FE9B" pageId="9" pageNumber="35" type="etymology">
<paragraph id="A5336A79C04F296DFFA9FF4959C9FF00" blockId="9.[121,250,147,175]" box="[121,250,147,175]" pageId="9" pageNumber="35">
<emphasis id="97F8B66BC04F296DFFA9FF4959C9FF00" bold="true" box="[121,250,147,175]" italics="true" pageId="9" pageNumber="35">Etymology</emphasis>
</paragraph>
<paragraph id="A5336A79C04F296DFFA9FF0C5B31FE9B" blockId="9.[121,775,214,308]" pageId="9" pageNumber="35">
The species epithet
<taxonomicName id="628C11FAC04F296DFEBAFF0C58E2FF5E" authorityName="Chandran &amp; Chandran &amp; Suraj &amp; Jose &amp; Koparde" authorityYear="2024" box="[362,465,214,241]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="rupestris">
<emphasis id="97F8B66BC04F296DFEBAFF0C58E2FF5E" box="[362,465,214,241]" italics="true" pageId="9" pageNumber="35">rupestris</emphasis>
</taxonomicName>
is derived from Latin and signifies “that lives on cliffs or rocks”, acknowledging this species habitat predilection.
</paragraph>
</subSubSection>
<subSubSection id="ED9639F2C04F296DFFA9FE815DC8FB9C" pageId="9" pageNumber="35" type="distribution">
<paragraph id="A5336A79C04F296DFFA9FE81583AFED8" blockId="9.[121,265,347,375]" box="[121,265,347,375]" pageId="9" pageNumber="35">
<emphasis id="97F8B66BC04F296DFFA9FE81583AFED8" bold="true" box="[121,265,347,375]" italics="true" pageId="9" pageNumber="35">Distribution</emphasis>
</paragraph>
<paragraph id="A5336A79C04F296DFFA9FE445B5EFDCF" blockId="9.[121,775,414,675]" pageId="9" pageNumber="35">
Presently, the new species is known only from the
<typeStatus id="7A37D4DBC04F296DFD02FE445A34FE15" box="[722,775,414,442]" pageId="9" pageNumber="35">type</typeStatus>
locality (Ponmudi Hills, southern Western Ghats), but it could be present in similar habitats (seasonal streams flowing over rocky cliffs at mid-altitudes) in the Agasthyamalai Hills, as these form a continuous chain of forested hills in the southern Western Ghats.
</paragraph>
<paragraph id="A5336A79C04F296DFF47FDBC5992FD0C" blockId="9.[121,775,414,675]" pageId="9" pageNumber="35">Temporal distribution. Recorded in June (30) and July (2).</paragraph>
<paragraph id="A5336A79C04F296DFFA9FD105802FD49" blockId="9.[121,305,714,742]" box="[121,305,714,742]" pageId="9" pageNumber="35">
<emphasis id="97F8B66BC04F296DFFA9FD105802FD49" bold="true" box="[121,305,714,742]" italics="true" pageId="9" pageNumber="35">Natural history</emphasis>
</paragraph>
<paragraph id="A5336A79C04F296DFFA9FCD65887FB9C" blockId="9.[121,776,780,1075]" pageId="9" pageNumber="35">
A population of 30 individuals was seen perched on vegetation around a steep rocky cliff over which rills of a seasonal stream flowed (
<figureCitation id="3DB776FCC04F296DFE13FC955B38FCC4" box="[451,523,847,875]" captionStart="Figure 9" captionStartId="7.[813,881,2084,2109]" captionTargetBox="[813,1467,1315,2045]" captionTargetId="figure-258@7.[813,1470,1315,2046]" captionTargetPageId="7" captionText="Figure 9. Habitat of Phylloneura rupestris sp. n." figureDoi="http://doi.org/10.5281/zenodo.15555078" httpUri="https://zenodo.org/record/15555078/files/figure.png" pageId="9" pageNumber="35">Fig. 9</figureCitation>
). The average depth of the stream, measured with a tape, was
<quantity id="6274C79CC04F296DFDA8FCAA5BFCFC23" box="[632,719,880,908]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="35" unit="m" value="0.14">0.14 m</quantity>
. The sky was overcast and there was light drizzle. Most of the observed individuals were males, and females were seen only while mating. They occasionally made short, slow flights, probably chasing flying insect prey, before returning to their perches.
</paragraph>
<paragraph id="A5336A79C04F296DFCFDFF495D7DFF00" blockId="9.[813,1102,147,175]" box="[813,1102,147,175]" pageId="9" pageNumber="35">
<emphasis id="97F8B66BC04F296DFCFDFF495D7DFF00" bold="true" box="[813,1102,147,175]" italics="true" pageId="9" pageNumber="35">Reproductive behaviour</emphasis>
</paragraph>
<paragraph id="A5336A79C04F296DFCFDFF0C5AD3FC23" blockId="9.[813,1467,214,1075]" pageId="9" pageNumber="35">
Oviposition was observed in six pairs seen, but photographs, and times of the entire sequence of reproductive activities could be taken for only one pair (
<figureCitation id="3DB776FCC04F296DFA8AFEC25C9CFE9B" box="[1370,1455,280,308]" captionStart="Figure 10" captionStartId="8.[121,189,1965,1990]" captionTargetBox="[121,1466,144,1927]" captionTargetId="figure-27@8.[121,1467,144,1927]" captionTargetPageId="8" captionText="Figure 10. Reproductive behaviour of Phylloneura rupestris sp. n.: (A) in tandem, (B) intra-male sperm translocation, (C) copula, (D) oviposition." figureDoi="http://doi.org/10.5281/zenodo.15555080" httpUri="https://zenodo.org/record/15555080/files/figure.png" pageId="9" pageNumber="35">Fig. 10</figureCitation>
). The duration of each activity is given in parenthesis: first, the male descended from its perch very slowly, hovered just above the stream, and dipped its abdomen in the water. Thereafer, it flew back in the direction of its former perch and caught a female in tandem (22 s). The male performed intra-male sperm translocation with the female held in tandem (10 s). Next, the pair flew around in tandem over the stream and slowly descended (18 s). The female oviposited on a submerged mat of moss in the stream, with the male holding it in tandem (10 min). The pair took to flight again, perched on another patch of moss in the stream about
<quantity id="6274C79CC04F296DFCAAFD105A95FD49" box="[890,934,714,742]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="35" unit="m" value="3.0">3 m</quantity>
away, and oviposited again (4 min). Thereafter, the male, with the female held in tandem, hung on streamside vegetation (41 s). Afer this, the pair separated and flew off in different directions. The entire reproductive activity of the observed pair thus lasted for 15.51 minutes.
</paragraph>
<paragraph id="A5336A79C04F296DFC9BFC485DC8FB9C" blockId="9.[813,1467,214,1075]" pageId="9" pageNumber="35">
All six observed pairs oviposited on moss growing in the seasonal rills in the rocky cliff. This oviposition technique is in sharp contrast to that of
<taxonomicName id="628C11FAC04F296DFB19FC0E5C4EFC40" box="[1225,1405,980,1007]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="97F8B66BC04F296DFB19FC0E5C4EFC40" box="[1225,1405,980,1007]" italics="true" pageId="9" pageNumber="35">P. westermanni</emphasis>
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, four pairs of which were observed by AVC ovipositing on the surface roots of riparian trees (
<figureCitation id="3DB776FCC04F296DFB4BFBCD5DDDFB9C" box="[1179,1262,1047,1075]" captionStart="Figure 11" captionStartId="9.[121,189,2084,2109]" captionTargetBox="[121,1467,1126,2045]" captionTargetId="figure-449@9.[118,1470,1124,2046]" captionTargetPageId="9" captionText="Figure 11. A pair of P. westermanni ovipositing on surface roots of a riparian tree, photographed in Wayanad." figureDoi="http://doi.org/10.5281/zenodo.15555082" httpUri="https://zenodo.org/record/15555082/files/figure.png" pageId="9" pageNumber="35">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,255 @@
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<mods:title id="175C78F26A4497064CFC47422FCA086A">The larva of Hetaerina duplex Selys, 1853 (Odonata: Calopterygidae) with a key to known Colombian larvae species</mods:title>
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<taxonomicName id="0199FBF1FFD7FFA19744FF16EB84559A" ID-CoL="3L6HV" authority="Selys, 1853" authorityName="Selys" authorityYear="1853" box="[121,470,211,242]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="duplex">
<emphasis id="F4ED5C60FFD7FFA19744FF16EB84559A" bold="true" box="[121,470,211,242]" pageId="3" pageNumber="19">
Hetaerina duplex
<bibRefCitation id="A208FD83FFD7FFA1966DFF13EB84559A" author="Selys Longchamps, E. de" box="[336,470,214,242]" pageId="3" pageNumber="19" pagination="1 - 73" refId="ref3431" refString="Selys Longchamps, E. de (1853) Synopsis des Calopterygines. Bul- letin de l'Academie Royale des sciences, des lettres et des beauxarts de Belgique, 20 (Annexe), 1-73" type="journal article" year="1853">
<emphasis id="F4ED5C60FFD7FFA1966DFF13EB84559A" box="[336,470,214,242]" italics="true" pageId="3" pageNumber="19">Selys, 1853</emphasis>
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</emphasis>
</taxonomicName>
(
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)
</heading>
</paragraph>
</subSubSection>
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<paragraph id="C6268072FFD7FFA19744FEDDEB0B545B" blockId="3.[121,345,280,307]" box="[121,345,280,307]" pageId="3" pageNumber="19">
<emphasis id="F4ED5C60FFD7FFA19744FEDDEB0B545B" box="[121,345,280,307]" italics="true" pageId="3" pageNumber="19">Material examined</emphasis>
</paragraph>
<paragraph id="C6268072FFD7FFA19744FE9EEB445757" blockId="3.[121,776,346,575]" pageId="3" pageNumber="19">
<materialsCitation id="76F18A2FFFD7FFA19744FE9EE8D754D2" collectingDate="2012-10-09" country="Colombia" elevation="1650" latitude="4.696944" location="Barbas Bremen Nature Reserve" longLatPrecision="1" longitude="-75.654724" municipality="Filandia Municipality" pageId="3" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Quindio">
<collectingCountry id="BE8EC0E2FFD7FFA19744FE9EEABC541F" box="[121,238,347,375]" name="Colombia" pageId="3" pageNumber="19">Colombia</collectingCountry>
,
<specimenCount id="D09F4BFBFFD7FFA197C0FE9EEB70541C" box="[253,290,346,375]" count="1" pageId="3" pageNumber="19" type="female">1♀</specimenCount>
,
<date id="B227A6B2FFD7FFA19608FE9EEBFA541F" box="[309,424,347,375]" pageId="3" pageNumber="19" value="2012-10-09">
<collectingDate id="A2635F5AFFD7FFA19608FE9EEBFA541F" box="[309,424,347,375]" pageId="3" pageNumber="19" value="2012-10-09">9 x 2012</collectingDate>
</date>
,
<collectingRegion id="045D4E90FFD7FFA1968AFE9EE8E4541F" box="[439,694,347,375]" country="Colombia" name="Quindio" pageId="3" pageNumber="19">Quindío Department</collectingRegion>
,
<collectingMunicipality id="26421A08FFD7FFA195F8FE9EEB1A54F0" pageId="3" pageNumber="19">Filandia Municipality</collectingMunicipality>
,
<location id="C346D6A9FFD7FFA1965DFEB9E95654F0" LSID="urn:lsid:plazi:treatment:4E303164FFD7FFA79744FF16EBD3525A:C346D6A9FFD7FFA1965DFEB9E95654F0" box="[352,772,380,408]" country="Colombia" latitude="4.696944" longLatPrecision="1" longitude="-75.654724" municipality="Filandia Municipality" name="Barbas Bremen Nature Reserve" pageId="3" pageNumber="19" stateProvince="Quindio">Barbas Bremen Nature Reserve</location>
,
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, -
<geoCoordinate id="A3ADE6B5FFD7FFA1967EFE5BE85554D2" box="[323,519,414,442]" degrees="75.65472222" direction="west" orientation="longitude" pageId="3" pageNumber="19" precision="1" value="-75.654724">75.65472222°W</geoCoordinate>
,
<quantity id="01612D97FFD7FFA19526FE5BE8D754D2" box="[539,645,414,442]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="3" pageNumber="19" unit="m" value="1650.0">
<elevation id="4DB46741FFD7FFA19526FE5BE8D754D2" box="[539,645,414,442]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="3" pageNumber="19" unit="m" value="1650.0">1650 m</elevation>
</quantity>
</materialsCitation>
.a.s.l., Leg: J. Montes, L. Perez;
<materialsCitation id="76F18A2FFFD7FFA1964FFE7AEB445757" collectingDate="2022-05-13" collectorName="M. Stand" country="Colombia" county="Pueblo Rico Municipality" latitude="5.2357125" location="Tatama National Park" longLatPrecision="1" longitude="-76.08712" municipality="Monte Bello Township" pageId="3" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Risaralda">
<specimenCount id="D09F4BFBFFD7FFA1964FFE7AEBC554B0" box="[370,407,446,475]" count="1" pageId="3" pageNumber="19" type="female">1♀</specimenCount>
,
<date id="B227A6B2FFD7FFA19694FE7AE87B54B3" box="[425,553,447,475]" pageId="3" pageNumber="19" value="2022-05-13">
<collectingDate id="A2635F5AFFD7FFA19694FE7AE87B54B3" box="[425,553,447,475]" pageId="3" pageNumber="19" value="2022-05-13">13 v 2022</collectingDate>
</date>
,
<collectingRegion id="045D4E90FFD7FFA1950BFE7AEAEE5494" country="Colombia" name="Risaralda" pageId="3" pageNumber="19">Risaralda Department</collectingRegion>
,
<collectingCounty id="2F47F8FEFFD7FFA197FBFE25EBBC5494" box="[198,494,480,508]" pageId="3" pageNumber="19">Pueblo Rico Municipality</collectingCounty>
,
<collectingMunicipality id="26421A08FFD7FFA196CAFE25E9565494" box="[503,772,480,508]" pageId="3" pageNumber="19">Monte Bello Township</collectingMunicipality>
,
<location id="C346D6A9FFD7FFA19744FDC7EBD55776" LSID="urn:lsid:plazi:treatment:4E303164FFD7FFA79744FF16EBD3525A:C346D6A9FFD7FFA19744FDC7EBD55776" box="[121,391,514,542]" country="Colombia" county="Pueblo Rico Municipality" latitude="5.2357125" longLatPrecision="1" longitude="-76.08712" municipality="Monte Bello Township" name="Tatama National Park" pageId="3" pageNumber="19" stateProvince="Risaralda">Tatamá National Park</location>
,
<geoCoordinate id="A3ADE6B5FFD7FFA196A5FDC7E86A5776" box="[408,568,514,542]" degrees="5.2357126" direction="north" orientation="latitude" pageId="3" pageNumber="19" precision="1" value="5.2357125">5.2357126°N</geoCoordinate>
, -
<geoCoordinate id="A3ADE6B5FFD7FFA19572FDC7E9565776" box="[591,772,514,542]" degrees="76.0871193" direction="west" orientation="longitude" pageId="3" pageNumber="19" precision="1" value="-76.08712">76.0871193°W</geoCoordinate>
,
<collectorName id="6B6CE5A4FFD7FFA19744FDE6EAB65757" box="[121,228,547,575]" pageId="3" pageNumber="19">M. Stand</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="C6268072FFD7FFA19410FF56EECB55C6" blockId="3.[813,1177,147,174]" box="[813,1177,147,174]" pageId="3" pageNumber="19">
<emphasis id="F4ED5C60FFD7FFA19410FF56EECB55C6" box="[813,1177,147,174]" italics="true" pageId="3" pageNumber="19">Description of final instar larva</emphasis>
</paragraph>
<paragraph id="C6268072FFD7FFA19410FF13E938545C" blockId="3.[813,1467,214,308]" pageId="3" pageNumber="19">Coloration of head and thorax mainly brownish with dark brownish dots, legs yellowish with dark brown dots.</paragraph>
<paragraph id="C6268072FFD7FFA69410FE9EEBF9543E" blockId="3.[813,1467,347,575]" lastBlockId="4.[121,775,147,342]" lastPageId="4" lastPageNumber="20" pageId="3" pageNumber="19">
Head. Brownish with dark brown pattern (
<figureCitation id="5EA29CF7FFD7FFA19208FE9EEFFC541F" box="[1333,1454,347,375]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="3" pageNumber="19">Figure 2a</figureCitation>
), with prominent and rounded posterolateral angles; antenna without setae. Labium: prementum-postmentum articulation reaching the anterior margin of mesocoxae; prementum with two setae on internal side of each lobe of ligula, external margins of ligular lobes serrate; labial palp ending in three well-developed and incurved hooks forming an 120° angle with it, median hook longest, inner one smallest; movable hook short- er than palp; inner margin of palp with small spines; four short apical palpal setae nearouter margin as in
<figureCitation id="5EA29CF7FFD0FFA69744FEDDEABF545C" box="[121,237,280,308]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="4" pageNumber="20">Figure 2b</figureCitation>
. Mandibles formula: L 12345 0 a(m
<superScript id="31EC2D3AFFD0FFA695A8FED3E881544D" attach="left" box="[661,723,278,293]" fontSize="6" pageId="4" pageNumber="20">1234567</superScript>
)b/
<collectionCode id="A08818B7FFD0FFA695CEFEDDE954545C" box="[755,774,280,308]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="20">R</collectionCode>
12345 y ab (
<figureCitation id="5EA29CF7FFD0FFA69636FEFFEBCD543E" box="[267,415,314,342]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="4" pageNumber="20">Figure 2cd</figureCitation>
).
</paragraph>
<caption id="92E6D0FAFFD7FFA19386F82DEF3D5D29" pageId="3" pageNumber="19" startId="3.[1211,1279,2024,2049]" targetBox="[125,1171,611,2106]" targetPageId="3" targetType="figure">
<paragraph id="C6268072FFD7FFA19386F82DEF3D5D29" blockId="3.[1211,1467,2024,2113]" pageId="3" pageNumber="19">
<emphasis id="F4ED5C60FFD7FFA19386F82DEF7A5D69" bold="true" box="[1211,1320,2024,2049]" pageId="3" pageNumber="19">Figure 3.</emphasis>
Distribution of
<taxonomicName id="0199FBF1FFD7FFA193E7F7CCEFCF5D4A" box="[1242,1437,2057,2082]" pageId="3" pageNumber="19">
<emphasis id="F4ED5C60FFD7FFA193E7F7CCEF175D49" box="[1242,1349,2057,2081]" italics="true" pageId="3" pageNumber="19">Hetaerina</emphasis>
species
</taxonomicName>
in Colombia, part I.
</paragraph>
</caption>
<paragraph id="C6268072FFD0FFA69744FEB9EE015757" blockId="4.[121,776,380,575]" lastBlockId="4.[813,1467,147,575]" pageId="4" pageNumber="20">
Thorax. Anterior margin of pronotum widely concave, with one prominent midlateral projection on each side pointing forward, posterior margin convex at middle; wing sheaths extending to posterior margin of S3; Legs: pale with three dark bands on femur, three protibial dark and faded bands; no setae or spines on femur and tibia.
<emphasis id="F4ED5C60FFD0FFA69452FF56E9B355C6" box="[879,993,147,174]" italics="true" pageId="4" pageNumber="20">Abdomen</emphasis>
: Lateral margins of S210 carinate, with well-developed posterolateral spine; posterolateral spine of S10 with five small spinules (
<figureCitation id="5EA29CF7FFD0FFA693C7FF13EF20559A" box="[1274,1394,214,242]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="4" pageNumber="20">Figure 2a</figureCitation>
); posterior margin of S10 with one mid-dorsal spine, and a group of four small spines each side of postero-ventral border (
<figureCitation id="5EA29CF7FFD0FFA694ADFEFFEE71543E" box="[912,1059,314,342]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="4" pageNumber="20">Figure 2ef</figureCitation>
). Female ovipositor not reaching distal margin of S10 (
<figureCitation id="5EA29CF7FFD0FFA6930BFE9EEEFF541F" box="[1078,1197,347,375]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="4" pageNumber="20">
Figure
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</figureCitation>
) and, in ventral view, with three ventral rows of spines (
<figureCitation id="5EA29CF7FFD0FFA693DCFEB9EF0754F0" box="[1249,1365,380,408]" captionStart="Figure 2" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[127,1461,1078,1978]" captionTargetId="figure-376@2.[121,1467,1062,1989]" captionTargetPageId="2" captionText="Figure 2. Details of morphology of Hetaerina duplex. (a) Final instar larva in dorsal view; (b) labium; (c) lef mandible; (d) right mandible, both mesal view; (e) S810 and caudal gills, dorsal view; (f) S810 and caudal gills, ventral view; (g) S810 showing the ovipositor, lateral view." pageId="4" pageNumber="20">Figure 2f</figureCitation>
); cercus conical and straight, 0.5 the length of S10. Caudal gills: lateral gills triquetral and lanceolate, with spines along lateral margins; central gill foliaceous, rounded at tip, with spines along dorsal and ventral margins about 0.7 the length of lateral gills.
</paragraph>
<caption id="92E6D0FAFFD0FFA69386F82DEF275D29" pageId="4" pageNumber="20" startId="4.[1211,1279,2024,2049]" targetBox="[125,1172,610,2105]" targetPageId="4" targetType="figure">
<paragraph id="C6268072FFD0FFA69386F82DEF275D29" blockId="4.[1211,1466,2024,2113]" pageId="4" pageNumber="20">
<emphasis id="F4ED5C60FFD0FFA69386F82DEF755D69" bold="true" box="[1211,1319,2024,2049]" pageId="4" pageNumber="20">Figure 4.</emphasis>
Distribution of
<taxonomicName id="0199FBF1FFD0FFA693E7F7CCEFCF5D4A" box="[1242,1437,2057,2082]" pageId="4" pageNumber="20">
<emphasis id="F4ED5C60FFD0FFA693E7F7CCEF175D49" box="[1242,1349,2057,2081]" italics="true" pageId="4" pageNumber="20">Hetaerina</emphasis>
species
</taxonomicName>
in Colombia, part II.
</paragraph>
</caption>
<paragraph id="C6268072FFD1FFA79744FF56EB2355C6" blockId="5.[121,369,147,174]" box="[121,369,147,174]" pageId="5" pageNumber="21">
<emphasis id="F4ED5C60FFD1FFA79744FF56EB2355C6" box="[121,369,147,174]" italics="true" pageId="5" pageNumber="21">Measurements (mm)</emphasis>
</paragraph>
<paragraph id="C6268072FFD1FFA79744FF13E8425708" blockId="5.[121,776,214,608]" pageId="5" pageNumber="21">Total body length (without caudal gills): 23.1. Head: maximum length: 2.8, maximum width: 4; prementum maximum length: 4.2, prementum maximum width: 2.9; clef length: 1.7; movable hook: 1.3. Thorax and legs: inner wing pads length: 6, outer wing pads length: 5.9; femur I: 4.5; femur II: 5.8; femur III: 6.8; tibia I: 4; tibia II: 5.1; tibia III: 5.9. Abdomen: total length: 13.7; S1 length: 1.7; S2 length: 1.7; S3 length: 1.4; S4 length: 1.7; S5 length: 1.6; S6 length: 1.2; S7 length: 1.4; S8 length: 1.2; S9 length: 1; S10 length: 0.8; cerci length: 0.4; female ovipositor length: 1.8; lateral caudal gills length: 6.7, median caudal gill length: 4.9.</paragraph>
<paragraph id="C6268072FFD1FFA79744FD42EABE57CA" blockId="5.[121,236,647,674]" box="[121,236,647,674]" pageId="5" pageNumber="21">
<emphasis id="F4ED5C60FFD1FFA79744FD42EABE57CA" box="[121,236,647,674]" italics="true" pageId="5" pageNumber="21">Diagnosis</emphasis>
</paragraph>
<paragraph id="C6268072FFD1FFA79744FD0FEB5E51B2" blockId="5.[121,776,714,1242]" pageId="5" pageNumber="21">
<taxonomicName id="0199FBF1FFD1FFA79744FD0FEB1B578D" authorityName="Selys" authorityYear="1853" box="[121,329,714,741]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="duplex">
<emphasis id="F4ED5C60FFD1FFA79744FD0FEB1B578D" box="[121,329,714,741]" italics="true" pageId="5" pageNumber="21">Hetaerina duplex</emphasis>
</taxonomicName>
occurs in
<collectingCountry id="BE8EC0E2FFD1FFA796EEFD0FE816578E" box="[467,580,714,742]" name="Colombia" pageId="5" pageNumber="21">Colombia</collectingCountry>
in the
<collectingRegion id="045D4E90FFD1FFA79595FD0FE955578E" box="[680,775,714,742]" country="Colombia" name="Quindio" pageId="5" pageNumber="21">Quindío</collectingRegion>
and
<collectingRegion id="045D4E90FFD1FFA7978FFD2EEB4C566F" box="[178,286,747,775]" country="Colombia" name="Risaralda" pageId="5" pageNumber="21">Risaralda</collectingRegion>
departments (
<figureCitation id="5EA29CF7FFD1FFA796E5FD2EE813566F" box="[472,577,747,775]" captionStart="Figure 3" captionStartId="3.[1211,1279,2024,2049]" captionTargetBox="[125,1171,611,2106]" captionTargetId="figure-176@3.[118,1176,602,2114]" captionTargetPageId="3" captionText="Figure 3. Distribution of Hetaerina species in Colombia, part I." pageId="5" pageNumber="21">Figure 3</figureCitation>
) and larvae for the present description were collected in fast-flowing streams, where they were found partially submerged among grassy vegetation. In Tatamá National Park, where
<specimenCount id="D09F4BFBFFD1FFA797F3FCB5EB2A56E4" box="[206,376,880,908]" count="1" pageId="5" pageNumber="21" type="generic">one specimen</specimenCount>
was collected,
<taxonomicName id="0199FBF1FFD1FFA7950BFCB5E8FA56E3" authorityName="Selys" authorityYear="1853" box="[566,680,880,907]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="duplex">
<emphasis id="F4ED5C60FFD1FFA7950BFCB5E8FA56E3" box="[566,680,880,907]" italics="true" pageId="5" pageNumber="21">H. duplex</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="0199FBF1FFD1FFA797C2FC57EBD356C5" authorityName="Selys" authorityYear="1873" box="[255,385,914,941]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="capitalis">
<emphasis id="F4ED5C60FFD1FFA797C2FC57EBD356C5" box="[255,385,914,941]" italics="true" pageId="5" pageNumber="21">H. capitalis</emphasis>
</taxonomicName>
and shares the following characters: lateral pronotal protuberance rounded, posterior border of S89 without row of spines on the dorsal surface, and two premental setae on each side of clef, but differs from
<taxonomicName id="0199FBF1FFD1FFA79637FBD2EBDE515A" authorityName="Selys" authorityYear="1873" box="[266,396,1047,1074]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="capitalis">
<emphasis id="F4ED5C60FFD1FFA79637FBD2EBDE515A" box="[266,396,1047,1074]" italics="true" pageId="5" pageNumber="21">H. capitalis</emphasis>
</taxonomicName>
in: lateral well-developed spine on each side of the lateral carina of S210 (one row of small spines in the lateral carina of S
<quantity id="01612D97FFD1FFA79513FB9FE826511E" box="[558,628,1114,1142]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="5" pageNumber="21" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">89 in</quantity>
<taxonomicName id="0199FBF1FFD1FFA79546FB9FE8AC511D" authorityName="Selys" authorityYear="1873" box="[635,766,1114,1141]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="capitalis">
<emphasis id="F4ED5C60FFD1FFA79546FB9FE8AC511D" box="[635,766,1114,1141]" italics="true" pageId="5" pageNumber="21">H. capitalis</emphasis>
</taxonomicName>
) and S10 with one well-developed mid-dorsal spine on the posterior border (S10 with three small spines in
<taxonomicName id="0199FBF1FFD1FFA79744FB7BEAA951B1" authorityName="Selys" authorityYear="1873" box="[121,251,1214,1241]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="capitalis">
<emphasis id="F4ED5C60FFD1FFA79744FB7BEAA951B1" box="[121,251,1214,1241]" italics="true" pageId="5" pageNumber="21">H. capitalis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8E83D3F9FFD1FFA79744FAC5EBD3525A" pageId="5" pageNumber="21" type="discussion">
<paragraph id="C6268072FFD1FFA79744FAC5EAB05074" blockId="5.[121,226,1280,1308]" box="[121,226,1280,1308]" pageId="5" pageNumber="21">
<emphasis id="F4ED5C60FFD1FFA79744FAC5EAB05074" bold="true" box="[121,226,1280,1308]" italics="true" pageId="5" pageNumber="21">Remarks</emphasis>
</paragraph>
<paragraph id="C6268072FFD1FFA79744FA86EBD3525A" blockId="5.[121,776,1347,1842]" pageId="5" pageNumber="21">
Like other
<taxonomicName id="0199FBF1FFD1FFA797C5FA86EB9E5037" box="[248,460,1347,1375]" pageId="5" pageNumber="21">
<emphasis id="F4ED5C60FFD1FFA797C5FA86EB3C5036" box="[248,366,1347,1374]" italics="true" pageId="5" pageNumber="21">Hetaerina</emphasis>
species
</taxonomicName>
inhabit lotic evironments, with or without abundant riparian vegetation (
<bibRefCitation id="A208FD83FFD1FFA79590FAA1EBE250CA" author="Guillermo-Ferreira, R &amp; Bispo, P." pageId="5" pageNumber="21" pagination="77 - 81" refId="ref3230" refString="Guillermo-Ferreira, R, &amp; Bispo, P. (2012). Description of the larva of Mnesarete pudica (Hagen in Selys, 1853) (Odonata: Calopterygidae) and notes on known genera of South American Calopterygidae larvae. Zootaxa, 3482, 77-81. doi: 10.11646/zootaxa.3482.1.5" type="journal article" year="2012">Guillermo-Ferreira &amp; Bispo, 2012</bibRefCitation>
),
<taxonomicName id="0199FBF1FFD1FFA796FAFA43E8C350C9" authorityName="Selys" authorityYear="1853" box="[455,657,1414,1441]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="duplex">
<emphasis id="F4ED5C60FFD1FFA796FAFA43E8C350C9" box="[455,657,1414,1441]" italics="true" pageId="5" pageNumber="21">Hetaerina duplex</emphasis>
</taxonomicName>
is record- ed in streams and small rivers with abundant vegetation, it has been found in sympatry with
<taxonomicName id="0199FBF1FFD1FFA7956FFA0DE886508B" authorityName="Selys" authorityYear="1873" box="[594,724,1480,1507]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="capitalis">
<emphasis id="F4ED5C60FFD1FFA7956FFA0DE886508B" box="[594,724,1480,1507]" italics="true" pageId="5" pageNumber="21">H. capitalis</emphasis>
</taxonomicName>
and
<taxonomicName id="0199FBF1FFD1FFA79744FA2FE86B536E" authority="(Bota-Sierra et al., 2024)" baseAuthorityName="Bota-Sierra" baseAuthorityYear="2024" box="[121,569,1514,1542]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="cruentata">
<emphasis id="F4ED5C60FFD1FFA79744FA2FEB5F536D" box="[121,269,1514,1541]" italics="true" pageId="5" pageNumber="21">H. cruentata</emphasis>
(Bota-Sierra et al., 2024)
</taxonomicName>
, interestingly, recent studies show that the surrounding vegetation determines the presence of the wing rubyspot (Stranding et al., 2022), the females have been seen ovipositing in dead wood inside river ponds (
<bibRefCitation id="A208FD83FFD1FFA796D9F9AAE8A453E3" author="Bota-Sierra, C. &amp; Sandoval-H, J. &amp; Ayala-Sanchez, D. &amp; Novelo-Gutierrez, R." box="[484,758,1647,1675]" pageId="5" pageNumber="21" refId="ref2887" refString="Bota-Sierra, C., Sandoval-H, J., Ayala-Sanchez, D. &amp; Novelo-Gutierrez, R. (2019). Libelulas de la Cordillera Occidental Colombiana, una mirada desde el Tatama. Panamericana Formas e Impresos, Bogota, 198 pp." type="book" year="2019">Bota-Sierra et al., 2019</bibRefCitation>
). Its distribution is limited to
<collectingCountry id="BE8EC0E2FFD1FFA796FCF955E86053C4" box="[449,562,1680,1708]" name="Colombia" pageId="5" pageNumber="21">Colombia</collectingCountry>
and
<collectingCountry id="BE8EC0E2FFD1FFA79550F955E88253C4" box="[621,720,1680,1708]" name="Ecuador" pageId="5" pageNumber="21">Ecuador</collectingCountry>
, and its conservation status, according to the International Union for Conservation of Nature (IUCN), is classified as Least Concern (
<bibRefCitation id="A208FD83FFD1FFA7966AF931E8E75278" author="Bota-Sierra, C. &amp; Palacino, F." box="[343,693,1780,1808]" pageId="5" pageNumber="21" refId="ref2933" refString="Bota-Sierra, C. &amp; Palacino, F. (2016). Hetaerina aurora. The IUCN Red List of Threatened Species 2016: e. T 49254606 A 49254949. Retrieved Ocotber, 23, 2024, from doi: 10.2305/IUCN.UK.2016-1. RLTS. T 49254606 A 49254949. en" type="book" year="2016">Bota-Sierra &amp; Palacino, 2016</bibRefCitation>
;
<bibRefCitation id="A208FD83FFD1FFA795FAF931EB22525A" author="Bota-Sierra, C. &amp; Rache, L." pageId="5" pageNumber="21" refId="ref2993" refString="Bota-Sierra, C. &amp; Rache, L. (2016). Hetaerina duplex. The IUCN Red List of Threatened Species 2016: e. T 49254256 A 49254969. Retrieved Ocotber, 23, 2024, from doi: 10.2305/IUCN.UK.2016-1. RLTS. T 49254256 A 49254969. en" type="book" year="2016">Bota-Sierra &amp; Rache, 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,300 @@
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<mods:title id="56B7F6893D5A04D08ABBF21BD51A3E6D">The final-instar larva of Bayadera strigata Davies &amp; Yang, 1996 from Yunnan, China (Odonata: Zygoptera: Euphaeidae)</mods:title>
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<mods:namePart id="8B3A3339E83F14088849E21AD2CFBDA6">Yang, Guo-Hui</mods:namePart>
<mods:affiliation id="173256B88242263EC39127F4079A98ED">College of Agriculture and Life Sciences, Dali University, Yunnan 671003, China &amp; Co-Innovation center for Cangshan Mountain and Erhai Lake Integrated Protection and Green Development of Yunnan Province, Dali University, Dali, Yunnan 671003, China &amp; Cangshan Forest Ecosystem Observation and Research Station of Yunnan Province, Dali University, Dali, Yunnan 671003, China</mods:affiliation>
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<treatment id="9B64EF42FFCCFFEFFF8B8567FB8BFE8D" LSID="urn:lsid:plazi:treatment:9B64EF42FFCCFFEFFF8B8567FB8BFE8D" httpUri="http://treatment.plazi.org/id/9B64EF42FFCCFFEFFF8B8567FB8BFE8D" lastPageId="4" lastPageNumber="15" pageId="1" pageNumber="12">
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<paragraph id="13725E54FFCCFFEAFF8B8567FDFAFAE6" blockId="1.[121,586,1344,1408]" box="[121,586,1344,1375]" pageId="1" pageNumber="12">
<heading id="483AE938FFCCFFEAFF8B8567FDFAFAE6" box="[121,586,1344,1375]" fontSize="11" level="2" pageId="1" pageNumber="12" reason="7">
<taxonomicName id="D4CD25D7FFCCFFEAFF8B8567FDFAFAE6" ID-CoL="L4P4" authority="Davies &amp; Yang, 1996" authorityName="Davies &amp; Yang" authorityYear="1996" box="[121,586,1344,1375]" class="Insecta" family="Euphaeidae" genus="Bayadera" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="12" phylum="Arthropoda" rank="species" species="strigata">
<emphasis id="21B98246FFCCFFEAFF8B8567FDFAFAE6" bold="true" box="[121,586,1344,1375]" pageId="1" pageNumber="12">
Bayadera strigata
<emphasis id="21B98246FFCCFFEAFEA08564FDFAFAE6" bold="true" box="[338,586,1347,1375]" italics="true" pageId="1" pageNumber="12">Davies &amp; Yang, 1996</emphasis>
</emphasis>
</taxonomicName>
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<subSubSection id="5BD70DDFFFCCFFEFFF8B8543FB8BFE8D" lastPageId="4" lastPageNumber="15" pageId="1" pageNumber="12" type="description">
<paragraph id="13725E54FFCCFFEAFF8B8543FECAFA39" blockId="1.[121,586,1344,1408]" box="[121,378,1380,1408]" pageId="1" pageNumber="12">
(
<figureCitation id="8BF642D1FFCCFFEAFF8D8543FF76FA39" box="[127,198,1380,1408]" captionStart="Figure 1" captionStartId="1.[813,881,2049,2074]" captionTargetBox="[816,1461,980,2011]" captionTargetId="figure-680@1.[813,1470,1452,1515]" captionTargetPageId="1" captionText="Figure 1. Caudal appendage of Bayadera strigata, (a) dorsal view; (b) lateral view." figureDoi="http://doi.org/10.5281/zenodo.15555016" httpUri="https://zenodo.org/record/15555016/files/figure.png" pageId="1" pageNumber="12">Figs 1</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFF218543FF55FA39" box="[211,229,1380,1408]" captionStart="Figure 2" captionStartId="2.[121,189,1178,1203]" captionTargetBox="[127,1456,155,1137]" captionTargetId="figure-26@2.[121,1467,144,1150]" captionTargetPageId="2" captionText="Figure 2. Larval habitus of Bayadera strigata, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555018" httpUri="https://zenodo.org/record/15555018/files/figure.png" pageId="1" pageNumber="12">2</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFF028543FEB2FA39" box="[240,258,1380,1408]" captionStart="Figure 3" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[124,1463,1271,1974]" captionTargetId="figure-28@2.[121,1467,1260,1978]" captionTargetPageId="2" captionText="Figure 3. Head of a larva of Bayadera strigata and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae." figureDoi="http://doi.org/10.5281/zenodo.15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="1" pageNumber="12">3</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFEFE8543FEAEFA39" box="[268,286,1380,1408]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="1" pageNumber="12">4</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFEDB8543FE8BFA39" box="[297,315,1380,1408]" captionStart="Figure 5" captionStartId="4.[121,189,2023,2048]" captionTargetBox="[285,1284,578,1986]" captionTargetId="figure-26@4.[278,1310,562,1992]" captionTargetPageId="4" captionText="Figure 5. Morphological details of the tibia and S810 of B. strigata: (a) tibial comb and tarsi, dorsal view; (b) male gonapophyses, ventral view; (c) male gonapophyses, lateral view; (d) female gonapophyses, ventral view; (e) female gonapophyses, lateral view; (f) lateral caudal gill, lateral view; (j) median caudal gill, lateral view; (h) caudal gills, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555024" httpUri="https://zenodo.org/record/15555024/files/figure.png" pageId="1" pageNumber="12">5</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFEB48543FEE8FA39" box="[326,344,1380,1408]" captionStart="Figure 6" captionStartId="6.[121,189,2024,2049]" captionTargetBox="[156,1440,625,1981]" captionTargetId="figure-243@6.[147,1443,624,1987]" captionTargetPageId="6" captionText="Figure 6. (a) Final instar of Bayadera strigata in the laboratory; (b) male imago emerging on 02.vi.2023; (c) emergence of a male on 04.xi.2023; (d) male of B. strigata perched on branches next to a stream; (e) numerous males of B. strigata taking females in tandem to lay eggs on wood in turbulent rivers." figureDoi="http://doi.org/10.5281/zenodo.15555026" httpUri="https://zenodo.org/record/15555026/files/figure.png" pageId="1" pageNumber="12">6</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFE918543FEC3FA39" box="[355,371,1380,1408]" captionStart="Figure 7" captionStartId="7.[121,189,671,696]" captionTargetBox="[121,775,144,634]" captionTargetId="figure-357@7.[121,775,144,634]" captionTargetPageId="7" captionText="Figure 7. Larval habitat of Bayadera strigata." figureDoi="http://doi.org/10.5281/zenodo.15555028" httpUri="https://zenodo.org/record/15555028/files/figure.png" pageId="1" pageNumber="12">7</figureCitation>
)
</paragraph>
<paragraph id="13725E54FFCCFFEAFF8B8580FDE6F88B" blockId="1.[121,345,1447,1474]" lastBlockId="1.[121,776,1513,1842]" pageId="1" pageNumber="12">
<emphasis id="21B98246FFCCFFEAFF8B8580FEE9FA7B" box="[121,345,1447,1474]" italics="true" pageId="1" pageNumber="12">Material examined</emphasis>
<materialsCitation id="A3A55409FFCCFFEAFCDF80B4FED4F9BF" collectingDate="2020-04-06" collectionCode="CVS, SPS" pageId="1" pageNumber="12" specimenCount="1" specimenCount-female="1">
Head. Broad and pentagonal in shape, flat dorsally, covered with numerous fine setae on the dorsum except on the vertex (
<figureCitation id="8BF642D1FFCCFFEAFC2780F1FBADFF4B" box="[981,1053,214,242]" captionStart="Figure 2" captionStartId="2.[121,189,1178,1203]" captionTargetBox="[127,1456,155,1137]" captionTargetId="figure-26@2.[121,1467,144,1150]" captionTargetPageId="2" captionText="Figure 2. Larval habitus of Bayadera strigata, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555018" httpUri="https://zenodo.org/record/15555018/files/figure.png" pageId="1" pageNumber="12">Figs 2</figureCitation>
,
<figureCitation id="8BF642D1FFCCFFEAFBDA80F1FBF9FF4B" box="[1064,1097,214,242]" captionStart="Figure 3" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[124,1463,1271,1974]" captionTargetId="figure-28@2.[121,1467,1260,1978]" captionTargetPageId="2" captionText="Figure 3. Head of a larva of Bayadera strigata and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae." figureDoi="http://doi.org/10.5281/zenodo.15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="1" pageNumber="12">3a</figureCitation>
). Labrum with long SS and scattered
<collectionCode id="75DCC691FFCCFFEAFC8480D0FC13FEAA" box="[886,931,247,275]" pageId="1" pageNumber="12">CVS</collectionCode>
on basal, and dense
<collectionCode id="75DCC691FFCCFFEAFB5980D0FB68FEAA" box="[1195,1240,247,275]" pageId="1" pageNumber="12">CVS</collectionCode>
and short SS at 4/5 of the base of the labrum (
<figureCitation id="8BF642D1FFCCFFEAFB9D813FFB57FE8D" box="[1135,1255,280,308]" captionStart="Figure 3" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[124,1463,1271,1974]" captionTargetId="figure-28@2.[121,1467,1260,1978]" captionTargetPageId="2" captionText="Figure 3. Head of a larva of Bayadera strigata and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae." figureDoi="http://doi.org/10.5281/zenodo.15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="1" pageNumber="12">Figs 3ab</figureCitation>
). Compound eyes broad and rounded, protruding anterolaterally. Occiput margin concave, with scattered
<collectionCode id="75DCC691FFCCFFEAFB44817CFB55FECE" box="[1206,1253,347,375]" pageId="1" pageNumber="12">SPS</collectionCode>
; postocular lobes strongly developed and rounded, but dense
<collectionCode id="75DCC691FFCCFFEAFAB3815BFADBFE21" box="[1345,1387,380,408]" pageId="1" pageNumber="12">SPS</collectionCode>
at the posterior corners create a seemingly squared profile; anterior margin with
<collectionCode id="75DCC691FFCCFFEAFBD48198FBE3FE62" box="[1062,1107,447,475]" pageId="1" pageNumber="12">CVS</collectionCode>
along the margins of the compound eyes, posterior margin intermingled with small spines, long SS and
<collectionCode id="75DCC691FFCCFFEAFBEA8225FBFAFDA7" box="[1048,1098,514,542]" pageId="1" pageNumber="12">CVS</collectionCode>
. Genae large, with rows of
<collectionCode id="75DCC691FFCCFFEAFA628225FA0AFDA7" box="[1424,1466,514,542]" pageId="1" pageNumber="12">SPS</collectionCode>
and
<collectionCode id="75DCC691FFCCFFEAFC978204FC22FD86" box="[869,914,547,575]" pageId="1" pageNumber="12">CVS</collectionCode>
along the ventral margins of the compound eyes; anterior margin of right gena with rows of blunt and distinct spines (78) and a row of
<collectionCode id="75DCC691FFCCFFEAFB018241FA90FD3B" box="[1267,1312,614,642]" pageId="1" pageNumber="12">CVS</collectionCode>
infused with
<collectionCode id="75DCC691FFCCFFEAFCDF82A0FCE7FD1A" box="[813,855,647,675]" pageId="1" pageNumber="12">SPS</collectionCode>
and SS (
<figureCitation id="8BF642D1FFCCFFEAFC4882A0FB80FD1A" box="[954,1072,647,675]" captionStart="Figure 3" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[124,1463,1271,1974]" captionTargetId="figure-28@2.[121,1467,1260,1978]" captionTargetPageId="2" captionText="Figure 3. Head of a larva of Bayadera strigata and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae." figureDoi="http://doi.org/10.5281/zenodo.15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="1" pageNumber="12">Figs 3bc</figureCitation>
); anterior margin of the lef gena with a row of 910 dominant spines and row of
<collectionCode id="75DCC691FFCCFFEAFA95828FFA24FD7D" box="[1383,1428,680,708]" pageId="1" pageNumber="12">CVS</collectionCode>
infused with
<collectionCode id="75DCC691FFCCFFEAFC4182EDFC6DFD5F" box="[947,989,714,742]" pageId="1" pageNumber="12">SPS</collectionCode>
and SS (
<figureCitation id="8BF642D1FFCCFFEAFBB182EDFB0BFD5F" box="[1091,1211,714,742]" captionStart="Figure 3" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[124,1463,1271,1974]" captionTargetId="figure-28@2.[121,1467,1260,1978]" captionTargetPageId="2" captionText="Figure 3. Head of a larva of Bayadera strigata and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae." figureDoi="http://doi.org/10.5281/zenodo.15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="1" pageNumber="12">Figs 3de</figureCitation>
). Antennae unusually long (1.3 × length of head measured from occipital margin to labrum), filiform, 7-segmented; third antennomere longest, length of antennomeres (mm): 0.67:0.8 6:1.23:0.83:0.46:0.23:0.09, giving a total length of 4.37 (
<figureCitation id="8BF642D1FFCCFFEAFCC68357FC37FC35" box="[820,903,880,908]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="1" pageNumber="12">Fig. 4a</figureCitation>
). Prementum 1.4 × longer than wide, subpenta-
<specimenCount id="05CB95DDFFCCFFEAFF8B85CDFF15F9BC" box="[121,165,1513,1542]" count="1" pageId="1" pageNumber="12" type="female">1 ♀</specimenCount>
larva,
<date id="67737894FFCCFFEAFF0485CDFED4F9BF" box="[246,356,1514,1542]" pageId="1" pageNumber="12" value="2020-04-06">
<collectingDate id="7737817CFFCCFFEAFF0485CDFED4F9BF" box="[246,356,1514,1542]" pageId="1" pageNumber="12" value="2020-04-06">6.iv.2020</collectingDate>
</date>
</materialsCitation>
,
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<collectorName id="BE383B82FFCCFFEAFE8285CDFE0CF9BF" box="[368,444,1514,1542]" pageId="1" pageNumber="12">ZiYang</collectorName>
stream (
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,
<geoCoordinate id="76F93893FFCCFFEAFD6F85CDFCB3F9BF" box="[669,771,1514,1542]" degrees="99.99" direction="east" orientation="longitude" pageId="1" pageNumber="12" precision="555" value="99.99">99.99° E</geoCoordinate>
,
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<quantity id="D435F3B1FFCCFFEAFF8B862CFF64F99E" box="[121,212,1547,1575]" metricMagnitude="3" metricUnit="m" metricValue="2.32" pageId="1" pageNumber="12" unit="m" value="2320.0">2320 m</quantity>
a.s.l.
</elevation>
),
<collectingCounty id="FA1326D8FFCCFFEAFED9862CFE69F99E" box="[299,473,1547,1575]" pageId="1" pageNumber="12">YangBi County</collectingCounty>
,
<location id="1612088FFFCCFFEAFE16862CFEB2F9F1" LSID="urn:lsid:plazi:treatment:9B64EF42FFCCFFEFFF8B8567FB8BFE8D:1612088FFFCCFFEAFE16862CFEB2F9F1" country="China" county="YangBi County" latitude="25.74" longLatPrecision="747" longitude="99.99" municipality="Mt Cangshan in Dali" name="Bai Autonomous Prefecture of Dali" pageId="1" pageNumber="12" stateProvince="Yunnan">Bai Autonomous Prefecture of Dali</location>
,
<collectingRegion id="D10990B6FFCCFFEAFEFF860BFE68F9F1" box="[269,472,1580,1608]" country="China" name="Yunnan" pageId="1" pageNumber="12">Yunnan Province</collectingRegion>
,
<collectingCountry id="6BDA1EC4FFCCFFEAFE10860BFD9AF9F1" box="[482,554,1580,1608]" name="China" pageId="1" pageNumber="12">China</collectingCountry>
,
<location id="1612088FFFCCFFEAFDC6860BFD27F9F1" LSID="urn:lsid:plazi:treatment:9B64EF42FFCCFFEFFF8B8567FB8BFE8D:1612088FFFCCFFEAFDC6860BFD27F9F1" box="[564,663,1580,1608]" country="China" county="YangBi County" latitude="25.74" longLatPrecision="747" longitude="99.99" municipality="Mt Cangshan in Dali" name="Guo-Hui" pageId="1" pageNumber="12" stateProvince="Yunnan">Guo-Hui</location>
<taxonomicName id="D4CD25D7FFCCFFEAFD6C860BFD66F9F1" box="[670,726,1580,1608]" class="Insecta" family="Buprestidae" genus="Yang" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="12" phylum="Arthropoda" rank="genus">Yang</taxonomicName>
leg.
<collectorName id="BE383B82FFCCFFEAFF8B8669FF4CF9D3" box="[121,252,1614,1642]" pageId="1" pageNumber="12">The ZiYang</collectorName>
is one of steams on
<collectingMunicipality id="F316C42EFFCCFFEAFDF48669FCB3F9D3" box="[518,771,1614,1642]" pageId="1" pageNumber="12">Mt Cangshan in Dali</collectingMunicipality>
,
<collectingRegion id="D10990B6FFCCFFEAFF8B8648FF66F932" box="[121,214,1647,1675]" country="China" name="Yunnan" pageId="1" pageNumber="12">Yunnan</collectingRegion>
.
<specimenCount id="05CB95DDFFCCFFEAFF108648FE95F933" box="[226,293,1646,1675]" count="2" pageId="1" pageNumber="12" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="05CB95DDFFCCFFEAFEC78648FECBF933" box="[309,379,1646,1675]" count="2" pageId="1" pageNumber="12" type="male">2 ♂♂</specimenCount>
larvae,
<date id="67737894FFCCFFEAFE298648FDE9F932" box="[475,601,1647,1675]" pageId="1" pageNumber="12" value="2023-04-27">
<collectingDate id="7737817CFFCCFFEAFE298648FDE9F932" box="[475,601,1647,1675]" pageId="1" pageNumber="12" value="2023-04-27">27.iv.2023</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="A3A55409FFCCFFEAFD968648FCB3F915" collectingDate="2023-06-02" collectorName="ZiYang &amp; The ZiYang" country="China" county="YangBi County" elevation="2320" latitude="25.74" location="Guo-Hui" longLatPrecision="747" longitude="99.99" municipality="Mt Cangshan in Dali" pageId="1" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
same locality,
<location id="1612088FFFCCFFEAFF8B86B7FF6CF915" LSID="urn:lsid:plazi:treatment:9B64EF42FFCCFFEFFF8B8567FB8BFE8D:1612088FFFCCFFEAFF8B86B7FF6CF915" box="[121,220,1680,1708]" name="Guo-Hui" pageId="1" pageNumber="12">Guo-Hui</location>
<taxonomicName id="D4CD25D7FFCCFFEAFF1786B7FEADF915" box="[229,285,1680,1708]" class="Insecta" family="Buprestidae" genus="Yang" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="12" phylum="Arthropoda" rank="genus">Yang</taxonomicName>
leg.
<specimenCount id="05CB95DDFFCCFFEAFEAB86B7FE37F915" box="[345,391,1680,1708]" count="1" pageId="1" pageNumber="12" type="male">1 ♂</specimenCount>
F-0 adult emergence
<date id="67737894FFCCFFEAFD6186B7FCB3F915" box="[659,771,1680,1708]" pageId="1" pageNumber="12" value="2023-06-02">
<collectingDate id="7737817CFFCCFFEAFD6186B7FCB3F915" box="[659,771,1680,1708]" pageId="1" pageNumber="12" value="2023-06-02">2.vi.2023</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="05CB95DDFFCCFFEAFF8B8695FF0AF974" box="[121,186,1713,1742]" count="2" pageId="1" pageNumber="12" type="male">2 ♂♂</specimenCount>
larvae,
<date id="67737894FFCCFFEAFEEA8695FE27F977" box="[280,407,1714,1742]" pageId="1" pageNumber="12" value="2023-09-27">27.ix.2023</date>
,
<materialsCitation id="A3A55409FFCCFFEAFE538695FDE2F88B" collectingDate="2023-11-04" collectorName="Dong-Dong Zi" country="China" county="HeQing County" elevation="2750" latitude="26.25" location="Mt Maer" longLatPrecision="746" longitude="100.13" pageId="1" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<location id="1612088FFFCCFFEAFE538695FDBBF977" LSID="urn:lsid:plazi:treatment:9B64EF42FFCCFFEFFF8B8567FB8BFE8D:1612088FFFCCFFEAFE538695FDBBF977" box="[417,523,1714,1742]" country="China" county="HeQing County" latitude="26.25" longLatPrecision="746" longitude="100.13" name="Mt Maer" pageId="1" pageNumber="12" stateProvince="Yunnan">Mt Maer</location>
(
<geoCoordinate id="76F93893FFCCFFEAFDEA8695FD34F977" box="[536,644,1714,1742]" degrees="26.25" direction="north" orientation="latitude" pageId="1" pageNumber="12" precision="555" value="26.25">26.25° N</geoCoordinate>
,
<geoCoordinate id="76F93893FFCCFFEAFD7D8695FCB3F977" box="[655,771,1714,1742]" degrees="100.13" direction="east" orientation="longitude" pageId="1" pageNumber="12" precision="555" value="100.13">100.13° E</geoCoordinate>
,
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<quantity id="D435F3B1FFCCFFEAFF8B86F4FF62F956" box="[121,210,1747,1775]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="12" unit="m" value="2750.0">2750 m</quantity>
a.s.l.
</elevation>
),
<collectingCounty id="FA1326D8FFCCFFEAFED586F4FE6DF956" box="[295,477,1747,1775]" pageId="1" pageNumber="12">HeQing County</collectingCounty>
, Bai Autonomous Prefecture of Dali,
<collectingRegion id="D10990B6FFCCFFEAFEFF86D3FE68F8A9" box="[269,472,1780,1808]" country="China" name="Yunnan" pageId="1" pageNumber="12">Yunnan Province</collectingRegion>
,
<collectingCountry id="6BDA1EC4FFCCFFEAFE1086D3FD9AF8A9" box="[482,554,1780,1808]" name="China" pageId="1" pageNumber="12">China</collectingCountry>
,
<collectorName id="BE383B82FFCCFFEAFDC186D3FD65F8A9" box="[563,725,1780,1808]" pageId="1" pageNumber="12">Dong-Dong Zi</collectorName>
leg.
<specimenCount id="05CB95DDFFCCFFEAFF8B8731FF14F888" box="[121,164,1813,1842]" count="1" pageId="1" pageNumber="12" type="male">1 ♂</specimenCount>
F-0 successful emergence
<date id="67737894FFCCFFEAFE138731FDE2F88B" box="[481,594,1814,1842]" pageId="1" pageNumber="12" value="2023-11-04">
<collectingDate id="7737817CFFCCFFEAFE138731FDE2F88B" box="[481,594,1814,1842]" pageId="1" pageNumber="12" value="2023-11-04">4.xi.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="13725E54FFCCFFEAFF8B875DFF4FF82C" blockId="1.[121,255,1914,1941]" box="[121,255,1914,1941]" pageId="1" pageNumber="12">
<emphasis id="21B98246FFCCFFEAFF8B875DFF4FF82C" box="[121,255,1914,1941]" italics="true" pageId="1" pageNumber="12">Description</emphasis>
</paragraph>
<paragraph id="13725E54FFCCFFEAFF8B879BFE40F785" blockId="1.[121,776,1980,2108]" pageId="1" pageNumber="12">
Habitus. A medium-sized larva, somewhat flattened, robust build, with long heavy legs; colour brownish black, lacking a distinct pattern, with dark saccoid gills with blunt and pale tips (
<figureCitation id="8BF642D1FFCCFFEAFE6D8807FE54F785" box="[415,484,2080,2108]" captionStart="Figure 2" captionStartId="2.[121,189,1178,1203]" captionTargetBox="[127,1456,155,1137]" captionTargetId="figure-26@2.[121,1467,144,1150]" captionTargetPageId="2" captionText="Figure 2. Larval habitus of Bayadera strigata, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555018" httpUri="https://zenodo.org/record/15555018/files/figure.png" pageId="1" pageNumber="12">Fig. 2</figureCitation>
).
</paragraph>
<caption id="47B20EDCFFCCFFEAFCDF8826FBA3F782" ID-DOI="http://doi.org/10.5281/zenodo.15555016" ID-Zenodo-Dep="15555016" httpUri="https://zenodo.org/record/15555016/files/figure.png" pageId="1" pageNumber="12" startId="1.[813,881,2049,2074]" targetBox="[816,1461,980,2011]" targetPageId="1" targetType="figure">
<paragraph id="13725E54FFCCFFEAFCDF8826FBA3F782" blockId="1.[813,1466,2049,2107]" pageId="1" pageNumber="12">
<emphasis id="21B98246FFCCFFEAFCDF8826FC3DF7A3" bold="true" box="[813,909,2049,2074]" pageId="1" pageNumber="12">Figure 1.</emphasis>
Caudal appendage of
<taxonomicName id="D4CD25D7FFCCFFEAFB718824FAF6F7A2" authorityName="Davies &amp; Yang" authorityYear="1996" box="[1155,1350,2051,2075]" class="Insecta" family="Euphaeidae" genus="Bayadera" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="12" phylum="Arthropoda" rank="species" species="strigata">
<emphasis id="21B98246FFCCFFEAFB718824FAF6F7A2" box="[1155,1350,2051,2075]" italics="true" pageId="1" pageNumber="12">Bayadera strigata</emphasis>
</taxonomicName>
, (a) dorsal view; (b) lateral view.
</paragraph>
</caption>
<caption id="47B20EDCFFCFFFE9FF8B84BDFD55FB0C" ID-DOI="http://doi.org/10.5281/zenodo.15555018" ID-Zenodo-Dep="15555018" box="[121,741,1178,1205]" httpUri="https://zenodo.org/record/15555018/files/figure.png" pageId="2" pageNumber="13" startId="2.[121,189,1178,1203]" targetBox="[127,1456,155,1137]" targetPageId="2" targetType="figure">
<paragraph id="13725E54FFCFFFE9FF8B84BDFD55FB0C" blockId="2.[121,741,1178,1205]" box="[121,741,1178,1205]" pageId="2" pageNumber="13">
<emphasis id="21B98246FFCFFFE9FF8B84BDFF68FB0A" bold="true" box="[121,216,1178,1203]" pageId="2" pageNumber="13">Figure 2.</emphasis>
Larval habitus of
<taxonomicName id="D4CD25D7FFCFFFE9FE6584BBFDE8FB0D" authorityName="Davies &amp; Yang" authorityYear="1996" box="[407,600,1180,1204]" class="Insecta" family="Euphaeidae" genus="Bayadera" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="strigata">
<emphasis id="21B98246FFCFFFE9FE6584BBFDE8FB0D" box="[407,600,1180,1204]" italics="true" pageId="2" pageNumber="13">Bayadera strigata</emphasis>
</taxonomicName>
, dorsal view.
</paragraph>
</caption>
<caption id="47B20EDCFFCFFFE9FF8B87C6FB38F783" ID-DOI="http://doi.org/10.5281/zenodo.15555020" ID-Zenodo-Dep="15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="2" pageNumber="13" startId="2.[121,189,2017,2042]" targetBox="[124,1463,1271,1974]" targetPageId="2" targetType="figure">
<paragraph id="13725E54FFCFFFE9FF8B87C6FB38F783" blockId="2.[121,1466,2017,2106]" pageId="2" pageNumber="13">
<emphasis id="21B98246FFCFFFE9FF8B87C6FF67F843" bold="true" box="[121,215,2017,2042]" pageId="2" pageNumber="13">Figure 3.</emphasis>
Head of a larva of
<taxonomicName id="D4CD25D7FFCFFFE9FE6F87C4FDEEF842" authorityName="Davies &amp; Yang" authorityYear="1996" box="[413,606,2019,2043]" class="Insecta" family="Euphaeidae" genus="Bayadera" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="strigata">
<emphasis id="21B98246FFCFFFE9FE6F87C4FDEEF842" box="[413,606,2019,2043]" italics="true" pageId="2" pageNumber="13">Bayadera strigata</emphasis>
</taxonomicName>
and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae.
</paragraph>
</caption>
<paragraph id="13725E54FFCEFFE8FF8B80B4FDC5FDD9" blockId="3.[121,776,147,842]" pageId="3" pageNumber="14">
gonal in shape, with a row of 1416 (lef side) and 13 17 (right side) strong broad spines and numerous small spines (each spine bearing SS) along the lateral margin of each side (
<figureCitation id="8BF642D1FFCEFFE8FEE480D0FE3BFEAA" box="[278,395,247,275]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="3" pageNumber="14">Figs 4bc</figureCitation>
); ligula convex, its anterior margin finely serrated, with a tiny median clef (
<figureCitation id="8BF642D1FFCEFFE8FD70813FFD66FE8D" box="[642,726,280,308]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="3" pageNumber="14">Fig. 4d</figureCitation>
); labial palp long, 0.54 × as long as the prementum, outer margin with scattered SS and CVS, while SS, CVS and SPS are dense in the basal half, weakly serrated on the inner margin, distal end with three processes, the outer process moderately incurved with a wavy outer margin, the middle process largest, weakly unciate, the inner process short and securiform, movable hook about 0.44 × as long as the labial palp, acuminate, bent slightly inwards (
<figureCitation id="8BF642D1FFCEFFE8FF3F8263FE91FDD9" box="[205,289,580,608]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="3" pageNumber="14">Fig. 4e</figureCitation>
). Mandibles brown, formula:
</paragraph>
<paragraph id="13725E54FFCEFFE8FF4B8241FD6CFCF3" blockId="3.[121,776,147,842]" pageId="3" pageNumber="14">
L 1+12340 a(m
<superScript id="E4B8F31CFFCEFFE8FE838244FE1DFDCB" attach="left" box="[369,429,611,626]" fontSize="6" pageId="3" pageNumber="14">1.2.3.4.5</superScript>
)b / R 1+1234 y a(m
<superScript id="E4B8F31CFFCEFFE8FD6C8244FD17FDCB" attach="left" box="[670,679,611,626]" fontSize="6" pageId="3" pageNumber="14">0</superScript>
)b, asymmetrical,fourincisors,welldeveloped,4&gt;3&gt;2&gt;1+1; molar crest with seven teeth (a&gt;1=2=3=4=5&gt;b) on lef mandible; a&gt;b, b very small on the right mandible (
<figureCitation id="8BF642D1FFCEFFE8FF8D82CCFF5EFCBE" box="[127,238,747,775]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="3" pageNumber="14">Figs 4fi</figureCitation>
); Maxilla with seven hooks, four long and three short, sharply pointed, basally with short setae; maxillary palp bearing dense, long setae (
<figureCitation id="8BF642D1FFCEFFE8FD918309FD60FCF3" box="[611,720,814,842]" captionStart="Figure 4" captionStartId="3.[121,189,2023,2048]" captionTargetBox="[132,1461,903,1983]" captionTargetId="figure-403@3.[121,1467,883,1992]" captionTargetPageId="3" captionText="Figure 4. Details of the head and mouthparts of the final instar of Bayadera strigata: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla." figureDoi="http://doi.org/10.5281/zenodo.15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="3" pageNumber="14">Figs 4jk</figureCitation>
).
</paragraph>
<paragraph id="13725E54FFCEFFE8FCDF80B4FBB9FDD9" blockId="3.[813,1468,147,608]" pageId="3" pageNumber="14">
Thorax. Prothorax trapezoidal, narrower than the head, with scattered CVS intermixed with SS. Procoxa arc-shaped, with a central protrusion, and supracoxal armament on prothorax circularly arc-shaped, with a depression in the middle, lateral margin of pronotal disc rounded (
<figureCitation id="8BF642D1FFCEFFE8FC6D811DFC59FEEF" box="[927,1001,314,342]" captionStart="Figure 2" captionStartId="2.[121,189,1178,1203]" captionTargetBox="[127,1456,155,1137]" captionTargetId="figure-26@2.[121,1467,144,1150]" captionTargetPageId="2" captionText="Figure 2. Larval habitus of Bayadera strigata, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555018" httpUri="https://zenodo.org/record/15555018/files/figure.png" pageId="3" pageNumber="14">Figs 2</figureCitation>
,
<figureCitation id="8BF642D1FFCEFFE8FC06811DFBA6FEEF" box="[1012,1046,314,342]" captionStart="Figure 3" captionStartId="2.[121,189,2017,2042]" captionTargetBox="[124,1463,1271,1974]" captionTargetId="figure-28@2.[121,1467,1260,1978]" captionTargetPageId="2" captionText="Figure 3. Head of a larva of Bayadera strigata and morphological details of the labrum and genae: (a) head, dorsal view; (b) labrum, dorsal view; (c) right genae; (d) lef genae; (e) detail of arrangement of the spines and setae along the margin of the right genae; (f) detail of arrangement of the spines and setae along the margin of the lef genae." figureDoi="http://doi.org/10.5281/zenodo.15555020" httpUri="https://zenodo.org/record/15555020/files/figure.png" pageId="3" pageNumber="14">3a</figureCitation>
). Synthorax nearly equal to head in width, covered with SS on its dorsal side. Wing sheaths divergent, forewing sheaths reaching the rear border of S5, hindwing sheaths extending to middle of S6. Legs long and robust; metafemur straight, reaching distal end of S7; all femorae and tibiae bearing SS and CVS; a row of claviform to rod-like setae on the distal end of the tibial comb; tarsal formula 3-3-3; claws small and simple (
<figureCitation id="8BF642D1FFCEFFE8FC7A8263FC61FDD9" box="[904,977,580,608]" captionStart="Figure 2" captionStartId="2.[121,189,1178,1203]" captionTargetBox="[127,1456,155,1137]" captionTargetId="figure-26@2.[121,1467,144,1150]" captionTargetPageId="2" captionText="Figure 2. Larval habitus of Bayadera strigata, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555018" httpUri="https://zenodo.org/record/15555018/files/figure.png" pageId="3" pageNumber="14">Figs 2</figureCitation>
,
<figureCitation id="8BF642D1FFCEFFE8FC298263FC4DFDD9" box="[987,1021,580,608]" captionStart="Figure 5" captionStartId="4.[121,189,2023,2048]" captionTargetBox="[285,1284,578,1986]" captionTargetId="figure-26@4.[278,1310,562,1992]" captionTargetPageId="4" captionText="Figure 5. Morphological details of the tibia and S810 of B. strigata: (a) tibial comb and tarsi, dorsal view; (b) male gonapophyses, ventral view; (c) male gonapophyses, lateral view; (d) female gonapophyses, ventral view; (e) female gonapophyses, lateral view; (f) lateral caudal gill, lateral view; (j) median caudal gill, lateral view; (h) caudal gills, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555024" httpUri="https://zenodo.org/record/15555024/files/figure.png" pageId="3" pageNumber="14">5a</figureCitation>
).
</paragraph>
<paragraph id="13725E54FFCEFFEFFCDF82A0FE80FE45" blockId="3.[813,1467,647,842]" lastBlockId="4.[121,776,147,508]" lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14">
Abdomen. Long, gradually narrowing from S1 to distal end, with scattered CVS and SS; seven pairs of pale lateroventral abdominal gills from S2 to S8. Male gonapophyses well developed, base stout, tips blunt, moderately divergent distally in ventral view, reaching more than halfway along S10, with a plate-like spine on the ventral side (
<figureCitation id="8BF642D1FFC9FFEFFEEE80B4FE28FF16" box="[284,408,147,175]" captionStart="Figure 5" captionStartId="4.[121,189,2023,2048]" captionTargetBox="[285,1284,578,1986]" captionTargetId="figure-26@4.[278,1310,562,1992]" captionTargetPageId="4" captionText="Figure 5. Morphological details of the tibia and S810 of B. strigata: (a) tibial comb and tarsi, dorsal view; (b) male gonapophyses, ventral view; (c) male gonapophyses, lateral view; (d) female gonapophyses, ventral view; (e) female gonapophyses, lateral view; (f) lateral caudal gill, lateral view; (j) median caudal gill, lateral view; (h) caudal gills, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555024" httpUri="https://zenodo.org/record/15555024/files/figure.png" pageId="4" pageNumber="15">Figs 5bc</figureCitation>
); female gonapophyses rising from the distal end of S8 and basal S9 and extending almost to the distal margin of S10 (
<figureCitation id="8BF642D1FFC9FFEFFDDE80F1FD17FF4B" box="[556,679,214,242]" captionStart="Figure 5" captionStartId="4.[121,189,2023,2048]" captionTargetBox="[285,1284,578,1986]" captionTargetId="figure-26@4.[278,1310,562,1992]" captionTargetPageId="4" captionText="Figure 5. Morphological details of the tibia and S810 of B. strigata: (a) tibial comb and tarsi, dorsal view; (b) male gonapophyses, ventral view; (c) male gonapophyses, lateral view; (d) female gonapophyses, ventral view; (e) female gonapophyses, lateral view; (f) lateral caudal gill, lateral view; (j) median caudal gill, lateral view; (h) caudal gills, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555024" httpUri="https://zenodo.org/record/15555024/files/figure.png" pageId="4" pageNumber="15">Figs 5de</figureCitation>
); lateral valvulae each with a nipple-shaped process at the tip and a plate-like spine on the ventral side; cerci in both sexes long, narrow, slightly curved with pointed tips, bearing claviform to rod-like setae; male paraprocts triangular. Caudal gills brownish black, swollen, saccoidshaped, with a yellowish white terminal filament, covered with SS; median gill about as long as the lateral gills (
<figureCitation id="8BF642D1FFC9FFEFFF4181C7FE95FE45" box="[179,293,480,508]" captionStart="Figure 5" captionStartId="4.[121,189,2023,2048]" captionTargetBox="[285,1284,578,1986]" captionTargetId="figure-26@4.[278,1310,562,1992]" captionTargetPageId="4" captionText="Figure 5. Morphological details of the tibia and S810 of B. strigata: (a) tibial comb and tarsi, dorsal view; (b) male gonapophyses, ventral view; (c) male gonapophyses, lateral view; (d) female gonapophyses, ventral view; (e) female gonapophyses, lateral view; (f) lateral caudal gill, lateral view; (j) median caudal gill, lateral view; (h) caudal gills, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15555024" httpUri="https://zenodo.org/record/15555024/files/figure.png" pageId="4" pageNumber="15">Figs 5fh</figureCitation>
).
</paragraph>
<caption id="47B20EDCFFCEFFE8FF8B87C0FAB6F7F9" ID-DOI="http://doi.org/10.5281/zenodo.15555022" ID-Zenodo-Dep="15555022" httpUri="https://zenodo.org/record/15555022/files/figure.png" pageId="3" pageNumber="14" startId="3.[121,189,2023,2048]" targetBox="[132,1461,903,1983]" targetPageId="3" targetType="figure">
<paragraph id="13725E54FFCEFFE8FF8B87C0FAB6F7F9" blockId="3.[121,1466,2023,2112]" pageId="3" pageNumber="14">
<emphasis id="21B98246FFCEFFE8FF8B87C0FF6AF7B9" bold="true" box="[121,218,2023,2048]" pageId="3" pageNumber="14">Figure 4.</emphasis>
Details of the head and mouthparts of the final instar of
<taxonomicName id="D4CD25D7FFCEFFE8FCA587CEFBAAF7B8" authorityName="Davies &amp; Yang" authorityYear="1996" box="[855,1050,2025,2049]" class="Insecta" family="Euphaeidae" genus="Bayadera" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="strigata">
<emphasis id="21B98246FFCEFFE8FCA587CEFBAAF7B8" box="[855,1050,2025,2049]" italics="true" pageId="3" pageNumber="14">Bayadera strigata</emphasis>
</taxonomicName>
: (a) right antenna; (b) labium, ventral view; (c) labium, dorsal view; (d) ligula, dorsal view; (e) labial palp, ventral view; (f) right mandible, ventral view; (g) right mandible, inner face; (h) lef mandible, ventral view; (i) lef mandible, inner face; (j) lef maxilla; (k) right maxilla.
</paragraph>
</caption>
<paragraph id="13725E54FFC9FFEFFCDF80B4FB8BFE8D" blockId="4.[813,1467,147,308]" pageId="4" pageNumber="15">Measurements (mm). Larvae (in alcohol, N = 7), body length (excluding antennae and caudal gills) 14.816.2; length of abdomen (including caudal gills) 1415.4; maximum width of head 5; length of hind femur 56.5; lateral caudal gills 78.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,71 +1,71 @@
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<treatment id="ED4EC161FFA7FF82FE62F913BD9BD26E" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E" httpUri="http://treatment.plazi.org/id/ED4EC161FFA7FF82FE62F913BD9BD26E" lastPageId="50" lastPageNumber="51" pageId="42" pageNumber="43">
<subSubSection id="2DFD23FCFFA7FF9AFE62F913BB24D2C7" box="[502,1086,1685,1712]" pageId="42" pageNumber="43" type="nomenclature">
@ -133,21 +133,20 @@ Male pedipalpal femur with a large intumescence on the subbasal prolateral surfa
<emphasis id="5793AC65FFA6FF9BFF7EFEB1BEBCD526" bold="true" box="[234,422,311,337]" pageId="43" pageNumber="44">Other material</emphasis>
</paragraph>
<paragraph id="65587077FFA6FF9BFF29FEDCBD12D652" blockId="43.[189,1399,311,549]" pageId="43" pageNumber="44">
<materialsCitation id="D58F7A2AFFA6FF9BFF29FEDCBCDED5CC" collectingDate="2024-01-14" collectorName="Jiaqi Zhao &amp; Jianzhou Sun &amp; Yiting Xu &amp; Ps." country="China" county="Autonomous Region" elevation="267" latitude="25.2475" location="Lingui Town" longLatPrecision="21" longitude="110.20083" municipality="Lingui District" pageId="43" pageNumber="44" specimenCode="MHBU-GX2024011401-18" specimenCount="18" specimenCount-female="8" specimenCount-male="10" stateProvince="Guangxi">
<materialsCitation id="D58F7A2AFFA6FF9BFF29FEDCBCDED5CC" collectedFrom="in leaf litter" collectingDate="2024-01-14" collectorName="Jiaqi Zhao &amp; Jianzhou Sun &amp; Yiting Xu" country="China" county="Lingui District" elevation="267" latitude="25.2475" location="Jiahe Garden" longLatPrecision="21" longitude="110.20083" municipality="Lingui Town" pageId="43" pageNumber="44" specimenCode="MHBU-GX2024011401, MHBU-GX2024011402, MHBU-GX2024011403, MHBU-GX2024011404, MHBU-GX2024011405, MHBU-GX2024011406, MHBU-GX2024011407, MHBU-GX2024011408, MHBU-GX2024011409, MHBU-GX2024011410, MHBU-GX2024011411, MHBU-GX2024011412. MHBU-GX2024011413, MHBU-GX2024011414, MHBU-GX2024011415, MHBU-GX2024011416, MHBU-GX2024011417, MHBU-GX2024011418" specimenCount="18" specimenCount-female="8" specimenCount-male="10" stateProvince="Guangxi Zhuang Autonomous Region">
<collectingCountry id="1DF030E7FFA6FF9BFF29FEDCBE00D503" box="[189,282,346,372]" name="China" pageId="43" pageNumber="44">CHINA</collectingCountry>
<specimenCount id="73E1BBFEFFA6FF9BFEA3FEDCBE96D502" box="[311,396,346,373]" count="10" pageId="43" pageNumber="44" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="73E1BBFEFFA6FF9BFE08FEDCBEF8D502" box="[412,482,346,373]" count="8" pageId="43" pageNumber="44" type="female">8 ♀♀</specimenCount>
;
<collectingRegion id="A723BE95FFA6FF9BFE67FEDCBDA6D503" box="[499,700,346,372]" country="China" name="Guangxi" pageId="43" pageNumber="44">Guangxi Zhuang</collectingRegion>
<collectingCounty id="8C3908FBFFA6FF9BFD50FEDCBCA4D503" box="[708,958,346,372]" pageId="43" pageNumber="44">Autonomous Region</collectingCounty>
<collectingRegion id="A723BE95FFA6FF9BFE67FEDCBCA4D503" box="[499,958,346,372]" pageId="43" pageNumber="44">Guangxi Zhuang Autonomous Region</collectingRegion>
,
<collectingMunicipality id="853CEA0DFFA6FF9BFC58FEDCBB9AD503" box="[972,1152,346,372]" pageId="43" pageNumber="44">Lingui District</collectingMunicipality>
<collectingCounty id="8C3908FBFFA6FF9BFC58FEDCBB9AD503" box="[972,1152,346,372]" pageId="43" pageNumber="44">Lingui District</collectingCounty>
,
<location id="603826ACFFA6FF9BFB1AFEDCBA31D502" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E:603826ACFFA6FF9BFB1AFEDCBA31D502" box="[1166,1323,346,373]" country="China" county="Autonomous Region" latitude="25.2475" longLatPrecision="21" longitude="110.20083" municipality="Lingui District" name="Lingui Town" pageId="43" pageNumber="44" stateProvince="Guangxi">Lingui Town</location>
<collectingMunicipality id="853CEA0DFFA6FF9BFB1AFEDCBA31D502" box="[1166,1323,346,373]" pageId="43" pageNumber="44">Lingui Town</collectingMunicipality>
,
<location id="603826ACFFA6FF9BFAAEFEDCBE0DD5E0" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E:603826ACFFA6FF9BFAAEFEDCBE0DD5E0" country="China" county="Autonomous Region" latitude="25.2475" longLatPrecision="21" longitude="110.20083" municipality="Lingui District" name="Jiahe Garden" pageId="43" pageNumber="44" stateProvince="Guangxi">Jiahe Garden</location>
<location id="603826ACFFA6FF9BFAAEFEDCBE0DD5E0" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E:603826ACFFA6FF9BFAAEFEDCBE0DD5E0" country="China" county="Lingui District" latitude="25.2475" longLatPrecision="21" longitude="110.20083" municipality="Lingui Town" name="Jiahe Garden" pageId="43" pageNumber="44" stateProvince="Guangxi Zhuang Autonomous Region">Jiahe Garden</location>
;
<geoCoordinate id="00D316B0FFA6FF9BFEB2FEFABEDAD5EF" box="[294,448,380,408]" degrees="25" direction="north" minutes="14" orientation="latitude" pageId="43" pageNumber="44" precision="15" seconds="51" value="25.2475">25°1451″ N</geoCoordinate>
,
@ -161,7 +160,9 @@ a.s.l.
<date id="115956B7FFA6FF9BFCB2FEF8BCDCD5EF" box="[806,966,382,408]" pageId="43" pageNumber="44" value="2024-01-14">
<collectingDate id="011DAF5FFFA6FF9BFCB2FEF8BCDCD5EF" box="[806,966,382,408]" pageId="43" pageNumber="44" value="2024-01-14">14 Jan. 2024</collectingDate>
</date>
; in leaf litter (
;
<collectedFrom id="D6FB338AFFA6FF9BFC42FEFBBB7CD5E0" box="[982,1126,381,407]" pageId="43" pageNumber="44">in leaf litter</collectedFrom>
(
<figureCitation id="FDDC6CF2FFA6FF9BFBEEFEFBBBCDD5EF" box="[1146,1239,381,408]" captionStart="Fig" captionStartId="6.[189,232,1756,1782]" captionTargetBox="[189,1399,450,1708]" captionTargetId="figure-126@6.[189,1399,450,1708]" captionTargetPageId="6" captionText="Fig. 2. Habitats of some species of Stenohya Beier, 1967. A. Stenohya polygonia Zhao, Guo &amp; Zhang sp. nov. B. Stenohya margarita Zhao, Guo &amp; Zhang sp. nov. C. Stenohya biverticis Zhao, Guo &amp; Zhang sp. nov. D. Stenohya cordata Zhao, Guo &amp; Zhang sp. nov. E. Stenohya harveyi Zhao, Guo &amp; Zhang sp. nov. F. Stenohya jiahensis Li &amp; Shi, 2023. G. Stenohya dongtianensis Li &amp; Shi, 2023." figureDoi="http://doi.org/10.5281/zenodo.15543367" httpUri="https://zenodo.org/record/15543367/files/figure.png" pageId="43" pageNumber="44">Fig. 2F</figureCitation>
);
<collectorName id="C81215A1FFA6FF9BFB64FEFBBA68D5E0" box="[1264,1394,381,407]" pageId="43" pageNumber="44">Jiaqi Zhao</collectorName>
@ -169,22 +170,20 @@ a.s.l.
<collectorName id="C81215A1FFA6FF9BFF29FE27BE41D5CC" box="[189,347,417,443]" pageId="43" pageNumber="44">Jianzhou Sun</collectorName>
and
<collectorName id="C81215A1FFA6FF9BFE00FE27BD10D5CC" box="[404,522,417,443]" pageId="43" pageNumber="44">Yiting Xu</collectorName>
leg.;
<collectorName id="C81215A1FFA6FF9BFDD9FE27BD76D5CC" box="[589,620,417,443]" pageId="43" pageNumber="44">Ps.</collectorName>
-
leg.; Ps.-
<specimenCode id="3541D80CFFA6FF9BFDE7FE27BCDED5CC" box="[627,964,417,443]" collectionCode="MHBU-GX" pageId="43" pageNumber="44">MHBU-GX202401140118</specimenCode>
</materialsCitation>
<materialsCitation id="D58F7A2AFFA6FF9BFC4AFE27BD12D652" collectingDate="2023-12-02" collectorName="Jiaqi Zhao &amp; Jianzhou Sun &amp; Tao Zheng &amp; Songtao Shi &amp; Ps." country="China" county="Autonomous Region" elevation="162" latitude="25.2475" location="Lingui Town" longLatPrecision="21" longitude="110.20111" municipality="Lingui District" pageId="43" pageNumber="44" specimenCode="MHBU-GX2023120201-05" specimenCount="1" stateProvince="Guangxi">
five tritonymphs;
<collectingRegion id="A723BE95FFA6FF9BFB24FE27BA6DD5CC" box="[1200,1399,417,443]" country="China" name="Guangxi" pageId="43" pageNumber="44">Guangxi Zhuang</collectingRegion>
<collectingCounty id="8C3908FBFFA6FF9BFF29FE42BEA7D5A9" box="[189,445,452,478]" pageId="43" pageNumber="44">Autonomous Region</collectingCounty>
<materialsCitation id="D58F7A2AFFA6FF9BFC4AFE27BD12D652" collectedFrom="in leaf litter" collectingDate="2023-12-02" collectorName="Jiaqi Zhao &amp; Jianzhou Sun &amp; Tao Zheng &amp; Songtao Shi" country="China" county="Lingui District" elevation="162" latitude="25.2475" location="Jiahe Garden" longLatPrecision="21" longitude="110.20111" municipality="Lingui Town" pageId="43" pageNumber="44" specimenCode="MHBU-GX2023120201, MHBU-GX2023120202, MHBU-GX2023120203, MHBU-GX2023120204, MHBU-GX2023120205" specimenCount="5" stateProvince="Guangxi Zhuang Autonomous Region">
<specimenCount id="73E1BBFEFFA6FF9BFC4AFE27BBBED5CC" box="[990,1188,417,443]" pageId="43" pageNumber="44">five tritonymphs</specimenCount>
;
<collectingRegion id="A723BE95FFA6FF9BFB24FE27BEA7D5A9" pageId="43" pageNumber="44">Guangxi Zhuang Autonomous Region</collectingRegion>
,
<collectingMunicipality id="853CEA0DFFA6FF9BFE46FE42BD91D5A9" box="[466,651,452,478]" pageId="43" pageNumber="44">Lingui District</collectingMunicipality>
<collectingCounty id="8C3908FBFFA6FF9BFE46FE42BD91D5A9" box="[466,651,452,478]" pageId="43" pageNumber="44">Lingui District</collectingCounty>
,
<location id="603826ACFFA6FF9BFD34FE42BC59D5A9" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E:603826ACFFA6FF9BFD34FE42BC59D5A9" box="[672,835,452,478]" country="China" county="Autonomous Region" latitude="25.2475" longLatPrecision="21" longitude="110.20111" municipality="Lingui District" name="Lingui Town" pageId="43" pageNumber="44" stateProvince="Guangxi">Lingui Town</location>
<collectingMunicipality id="853CEA0DFFA6FF9BFD34FE42BC59D5A9" box="[672,835,452,478]" pageId="43" pageNumber="44">Lingui Town</collectingMunicipality>
,
<location id="603826ACFFA6FF9BFCCCFE42BCE5D5A9" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E:603826ACFFA6FF9BFCCCFE42BCE5D5A9" box="[856,1023,452,478]" country="China" county="Autonomous Region" latitude="25.2475" longLatPrecision="21" longitude="110.20111" municipality="Lingui District" name="Jiahe Garden" pageId="43" pageNumber="44" stateProvince="Guangxi">Jiahe Garden</location>
<location id="603826ACFFA6FF9BFCCCFE42BCE5D5A9" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF82FE62F913BD9BD26E:603826ACFFA6FF9BFCCCFE42BCE5D5A9" box="[856,1023,452,478]" country="China" county="Lingui District" latitude="25.2475" longLatPrecision="21" longitude="110.20111" municipality="Lingui Town" name="Jiahe Garden" pageId="43" pageNumber="44" stateProvince="Guangxi Zhuang Autonomous Region">Jiahe Garden</location>
;
<geoCoordinate id="00D316B0FFA6FF9BFB80FE44BBA9D5A9" box="[1044,1203,450,478]" degrees="25" direction="north" minutes="14" orientation="latitude" pageId="43" pageNumber="44" precision="15" seconds="51" value="25.2475">25°1451″ N</geoCoordinate>
,
@ -198,7 +197,9 @@ a.s.l.
<date id="115956B7FFA6FF9BFEC1FE61BEFDD676" box="[341,487,487,514]" pageId="43" pageNumber="44" value="2023-12-02">
<collectingDate id="011DAF5FFFA6FF9BFEC1FE61BEFDD676" box="[341,487,487,514]" pageId="43" pageNumber="44" value="2023-12-02">2 Dec. 2023</collectingDate>
</date>
; in leaf litter;
;
<collectedFrom id="D6FB338AFFA6FF9BFE67FE61BD9AD676" box="[499,640,487,513]" pageId="43" pageNumber="44">in leaf litter</collectedFrom>
;
<collectorName id="C81215A1FFA6FF9BFD18FE61BC10D676" box="[652,778,487,513]" pageId="43" pageNumber="44">Jiaqi Zhao</collectorName>
,
<collectorName id="C81215A1FFA6FF9BFC82FE61BCACD676" box="[790,950,487,513]" pageId="43" pageNumber="44">Jianzhou Sun</collectorName>
@ -206,9 +207,7 @@ a.s.l.
<collectorName id="C81215A1FFA6FF9BFC56FE6EBB5AD676" box="[962,1088,487,514]" pageId="43" pageNumber="44">Tao Zheng</collectorName>
and
<collectorName id="C81215A1FFA6FF9BFBEDFE61BA1DD676" box="[1145,1287,487,513]" pageId="43" pageNumber="44">Songtao Shi</collectorName>
leg.;
<collectorName id="C81215A1FFA6FF9BFADEFE6EBA74D675" box="[1354,1390,488,514]" pageId="43" pageNumber="44">Ps.</collectorName>
-
leg.; Ps.-
<specimenCode id="3541D80CFFA6FF9BFF29FD8CBD12D652" box="[189,520,522,549]" collectionCode="MHBU-GX" pageId="43" pageNumber="44">MHBU-GX202312020105</specimenCode>
</materialsCitation>
</paragraph>

View file

@ -1,71 +1,71 @@
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<emphasis id="5793AC65FFA7FF9AFF29FBE4BEB0D00B" bold="true" box="[189,426,1122,1148]" pageId="42" pageNumber="43">Material examined</emphasis>
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<paragraph id="65587077FFA7FF9AFF7EFB16BDD8D099" blockId="42.[189,1399,1168,1262]" pageId="42" pageNumber="43">
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@ -146,7 +144,7 @@ Zhan
,
<collectingMunicipality id="853CEA0DFFA7FF9AFDD7FB37BDC2D0BC" box="[579,728,1201,1227]" pageId="42" pageNumber="43">Fuzhou City</collectingMunicipality>
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<location id="603826ACFFA7FF9AFD72FB37BCA5D0BC" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF9AFE7DFD2EBE3BD207:603826ACFFA7FF9AFD72FB37BCA5D0BC" box="[742,959,1201,1227]" country="China" latitude="26.066668" longLatPrecision="1243" longitude="119.35" municipality="Fuzhou City" name="Gushan Mountain" pageId="42" pageNumber="43" stateProvince="Fujian Province">Gushan Mountain</location>
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;
<collectorName id="C81215A1FFA7FF9AFF29FB52BE6CD099" box="[189,374,1236,1262]" pageId="42" pageNumber="43">Fusheng Huang</collectorName>
leg.;
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-
leg.; Ps.-
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,
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<location id="603826ACFFA7FF9AFCD3FAB0BB05D127" LSID="urn:lsid:plazi:treatment:ED4EC161FFA7FF9AFE7DFD2EBE3BD207:603826ACFFA7FF9AFCD3FAB0BB05D127" box="[839,1055,1334,1360]" country="China" latitude="26.094168" longLatPrecision="20" longitude="119.37444" municipality="Fuzhou City" name="Gushan Mountain" pageId="42" pageNumber="43" stateProvince="Fujian Province">Gushan Mountain</location>
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<collectorName id="C81215A1FFA7FF9AFDEFFADEBC3ED104" box="[635,804,1368,1395]" pageId="42" pageNumber="43">Weitong Wang</collectorName>
and
<collectorName id="C81215A1FFA7FF9AFCC1FADEBCD7D104" box="[853,973,1368,1395]" pageId="42" pageNumber="43">Xiao Zang</collectorName>
leg.;
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.

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<figureCitation id="FDDC6CF2FFACFF91FD1FF916BDE1D2DD" box="[651,763,1680,1706]" captionStart="Fig" captionStartId="2.[189,232,1575,1601]" captionTargetBox="[308,1288,265,1541]" captionTargetId="figure-256@2.[308,1288,265,1541]" captionTargetPageId="2" captionText="Fig. 1. Distribution map of species of Stenohya Beier, 1967 from China. 1. Stenohya xiningensis Zhao, Zhang &amp; Jia, 2011. 2. Stenohya setulosa Guo &amp; Zhang, 2016. 3. Stenohya bomica Zhao &amp; Zhang, 2011. 4. Stenohya spinata Zhan, Feng &amp; Zhang, 2023. 5. Stenohya qianlei Zhao, Guo &amp; Zhang sp. nov. 6. Stenohya gibba Zhao, Guo &amp; Zhang, 2024. 7. Stenohya papillata Zhao, Guo &amp; Zhang, 2024. 8. Stenohya margarita Zhao, Guo &amp; Zhang sp. nov. 9. Stenohya guangmingensis Zhao, Guo &amp; Zhang, 2024. 10. Stenohya jiaxiani Zhao, Guo &amp; Zhang sp. nov. 11. Stenohya huangi Hu &amp; Zhang, 2012. 12. Stenohya biverticis Zhao, Guo &amp; Zhang sp. nov. 13. Stenohya pengae Hu &amp; Zhang, 2012. 14. Stenohya hainanensis Guo &amp; Zhang, 2016. 15. Stenohya curvata Zhao, Zhang &amp; Jia, 2011. 16. Stenohya bicornuta Guo, Zang &amp; Zhang, 2019. 17. Stenohya harveyi Zhao, Guo &amp; Zhang sp. nov. 18. Stenohya arcuata Guo, Zang &amp; Zhang, 2019. 19. Stenohya tengchongensis Yang &amp; Zhang, 2013. 20. Stenohya meiacantha Yang &amp; Zhang, 2013. 21. Stenohya jiahensis Li &amp; Shi, 2023. 22. Stenohya cordata Zhao, Guo &amp; Zhang sp. nov. 23. Stenohya ungulata Zhao, Guo &amp; Zhang sp. nov. 24. Stenohya polygonia Zhao, Guo &amp; Zhang sp. nov. 25. Stenohya dongtianensis Li &amp; Shi, 2023." figureDoi="http://doi.org/10.5281/zenodo.15543363" httpUri="https://zenodo.org/record/15543363/files/figure.png" pageId="33" pageNumber="34">Figs 1.14</figureCitation>
,
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,
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,
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<location id="603826ACFFAFFF92FD6EFEDCBCFAD503" LSID="urn:lsid:plazi:treatment:ED4EC161FFACFF92FE50F9EABEF3D068:603826ACFFAFFF92FD6EFEDCBCFAD503" box="[762,992,346,372]" country="China" county="Lingshui County" latitude="18.666666" longLatPrecision="1274" longitude="109.916664" name="Diaoluo Mountains" pageId="34" pageNumber="35" stateProvince="Hainan Province">Diaoluo Mountains</location>
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@ -144,31 +144,27 @@ a.s.l.
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;
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leg.;
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-
leg.; Ps.-
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<typeStatus id="BA5CCED5FFAFFF92FF7EFE39BE7DD5AE" box="[234,359,447,473]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
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; same collection data as for holotype; Ps.-
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MHBU-HN
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06
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View file

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<taxonomicName id="A2E70BF4FFAEFF93FE3BFE8FBC6CD554" authority="Zhao, Guo &amp; Zhang" authorityName="Zhao, Guo &amp; Zhang" authorityYear="2025" box="[431,886,265,292]" class="Arachnida" family="Neobisiidae" genus="Stenohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="harveyi" status="sp. nov." subFamily="Microcreagrinae">
@ -79,14 +79,14 @@ Zhao, Guo &amp; Zhang
(哈氏狭伪$)
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<figureCitation id="FDDC6CF2FFAEFF93FDECFED6BDF2D51D" box="[632,744,336,362]" captionStart="Fig" captionStartId="2.[189,232,1575,1601]" captionTargetBox="[308,1288,265,1541]" captionTargetId="figure-256@2.[308,1288,265,1541]" captionTargetPageId="2" captionText="Fig. 1. Distribution map of species of Stenohya Beier, 1967 from China. 1. Stenohya xiningensis Zhao, Zhang &amp; Jia, 2011. 2. Stenohya setulosa Guo &amp; Zhang, 2016. 3. Stenohya bomica Zhao &amp; Zhang, 2011. 4. Stenohya spinata Zhan, Feng &amp; Zhang, 2023. 5. Stenohya qianlei Zhao, Guo &amp; Zhang sp. nov. 6. Stenohya gibba Zhao, Guo &amp; Zhang, 2024. 7. Stenohya papillata Zhao, Guo &amp; Zhang, 2024. 8. Stenohya margarita Zhao, Guo &amp; Zhang sp. nov. 9. Stenohya guangmingensis Zhao, Guo &amp; Zhang, 2024. 10. Stenohya jiaxiani Zhao, Guo &amp; Zhang sp. nov. 11. Stenohya huangi Hu &amp; Zhang, 2012. 12. Stenohya biverticis Zhao, Guo &amp; Zhang sp. nov. 13. Stenohya pengae Hu &amp; Zhang, 2012. 14. Stenohya hainanensis Guo &amp; Zhang, 2016. 15. Stenohya curvata Zhao, Zhang &amp; Jia, 2011. 16. Stenohya bicornuta Guo, Zang &amp; Zhang, 2019. 17. Stenohya harveyi Zhao, Guo &amp; Zhang sp. nov. 18. Stenohya arcuata Guo, Zang &amp; Zhang, 2019. 19. Stenohya tengchongensis Yang &amp; Zhang, 2013. 20. Stenohya meiacantha Yang &amp; Zhang, 2013. 21. Stenohya jiahensis Li &amp; Shi, 2023. 22. Stenohya cordata Zhao, Guo &amp; Zhang sp. nov. 23. Stenohya ungulata Zhao, Guo &amp; Zhang sp. nov. 24. Stenohya polygonia Zhao, Guo &amp; Zhang sp. nov. 25. Stenohya dongtianensis Li &amp; Shi, 2023." figureDoi="http://doi.org/10.5281/zenodo.15543363" httpUri="https://zenodo.org/record/15543363/files/figure.png" pageId="35" pageNumber="36">Figs 1.17</figureCitation>
,
@ -128,13 +128,11 @@ Male pedipalpal chelal hand has a projection with bifid horn-like bulge on prola
<emphasis id="5793AC65FFAEFF93FF29FD78BE76D76F" bold="true" box="[189,364,766,792]" pageId="35" pageNumber="36">Type material</emphasis>
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; in leaf litter (
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(
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