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<mods:namePart id="127779E36966A8A36EF33D1F653B46F9">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="F4BE3C7B464FA619F65647B09223C7CA">2025</mods:date>
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3.4.1.
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<emphasis id="EABC5F3DFF836C4DFF7CFCC6FE0FFC71" bold="true" box="[225,436,870,897]" italics="true" pageId="13" pageNumber="514">Omalium alticola</emphasis>
<bibRefCitation id="BC59FEDEFF836C4DFE26FCC6FDC2FC70" author="Cameron, M." box="[443,633,870,896]" pageId="13" pageNumber="514" pagination="56 - 60" refId="ref30195" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). 3. The Proceedings of the Royal Entomological Society of London (B), 10, 56-60 + 142 - 147. https://doi.org/10.1111/j.1365-3113.1941.tb00709.x" type="journal article" year="1941">Cameron, 1941</bibRefCitation>
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(
<figureCitation id="40F39FAAFF836C4DFF02FC2BFEA4FC55" box="[159,287,907,933]" captionStart="FIGURES 3438" captionStartId="11.[151,264,1758,1782]" captionTargetBox="[160,1427,190,1725]" captionTargetId="figure-16@11.[151,1436,181,1734]" captionTargetPageId="11" captionText="FIGURES 3438. Aedeagus of Omalium (Figs 3436—O. pratense (holotype), Figs 3738—O. alticola (Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="13" pageNumber="514">Figs 3738</figureCitation>
)
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<taxonomicName id="1FC8F8ACFF836C4DFF0AFC6FFDE5FC17" authority="Cameron, 1941: 57" authorityName="Cameron" authorityPageNumber="57" authorityYear="1941" box="[151,606,975,999]" class="Insecta" family="Staphylinidae" genus="Omalium" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="EABC5F3DFF836C4DFF0AFC6FFF4EFC17" box="[151,245,975,999]" italics="true" pageId="13" pageNumber="514">Omalium</emphasis>
(s. str.)
<emphasis id="EABC5F3DFF836C4DFED5FC6FFE2EFC17" box="[328,405,975,999]" italics="true" pageId="13" pageNumber="514">alticola</emphasis>
<bibRefCitation id="BC59FEDEFF836C4DFE01FC6FFDE5FC17" author="Cameron, M." box="[412,606,975,999]" pageId="13" pageNumber="514" pagination="56 - 60" refId="ref30195" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). 3. The Proceedings of the Royal Entomological Society of London (B), 10, 56-60 + 142 - 147. https://doi.org/10.1111/j.1365-3113.1941.tb00709.x" type="journal article" year="1941">Cameron, 1941: 57</bibRefCitation>
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<taxonomicName id="1FC8F8ACFF836C4DFF0AFC4EFDCDFBF6" authority="Cameron, 1941: 58" authorityName="Cameron" authorityPageNumber="58" authorityYear="1941" box="[151,630,1006,1030]" class="Insecta" family="Staphylinidae" genus="Omalium" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="EABC5F3DFF836C4DFF0AFC4EFF4EFBF6" box="[151,245,1006,1030]" italics="true" pageId="13" pageNumber="514">Omalium</emphasis>
(s. str.)
<emphasis id="EABC5F3DFF836C4DFED5FC4EFE16FBF6" box="[328,429,1006,1030]" italics="true" pageId="13" pageNumber="514">monticola</emphasis>
<bibRefCitation id="BC59FEDEFF836C4DFE2EFC4EFDCDFBF6" author="Cameron, M." box="[435,630,1006,1030]" pageId="13" pageNumber="514" pagination="56 - 60" refId="ref30195" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). 3. The Proceedings of the Royal Entomological Society of London (B), 10, 56-60 + 142 - 147. https://doi.org/10.1111/j.1365-3113.1941.tb00709.x" type="journal article" year="1941">Cameron, 1941: 58</bibRefCitation>
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<taxonomicName id="1FC8F8ACFF836C4DFF0AFBACFEF2FBD4" authorityName="Cameron" authorityYear="1941" box="[151,329,1036,1060]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="EABC5F3DFF836C4DFF0AFBACFEF2FBD4" box="[151,329,1036,1060]" italics="true" pageId="13" pageNumber="514">Omalium alticola</emphasis>
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:
<bibRefCitation id="BC59FEDEFF836C4DFECAFBACFDA6FBD5" author="Shavrin, A. V." box="[343,541,1036,1061]" pageId="13" pageNumber="514" pagination="151 - 206" refId="ref30642" refString="Shavrin, A. V. (2023 a) A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region. Zootaxa, 5369 (2), 151-206. https://doi.org/10.11646/zootaxa.5369.2.1" type="journal article" year="2023">Shavrin 2023a: 172</bibRefCitation>
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Material examined.
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:
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:
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: Chung King Szechwan China centralis (
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).
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<emphasis id="EABC5F3DFF836C4DFF5AFBD7FE80FB61" bold="true" box="[199,315,1143,1169]" pageId="13" pageNumber="514">Remarks.</emphasis>
The Himalayan
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<emphasis id="EABC5F3DFF836C4DFE6EFBD7FDD0FB61" box="[499,619,1143,1169]" italics="true" pageId="13" pageNumber="514">O. alticola</emphasis>
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is known from
<collectingCountry id="A0DFC3BFFF836C4DFC85FBD7FCE9FB61" box="[792,850,1143,1169]" name="India" pageId="13" pageNumber="514">India</collectingCountry>
(Kashmir,
<collectingRegion id="1A0C4DCDFF836C4DFC50FBD7FBE2FB61" box="[973,1113,1143,1169]" country="India" name="Uttarakhand" pageId="13" pageNumber="514">Uttarakhand</collectingRegion>
) and western
<collectingCountry id="A0DFC3BFFF836C4DFB6EFBD7FA8DFB61" box="[1267,1334,1143,1169]" name="Nepal" pageId="13" pageNumber="514">Nepal</collectingCountry>
(
<bibRefCitation id="BC59FEDEFF836C4DFADEFBD7FF5BFB45" author="Shavrin, A. V." pageId="13" pageNumber="514" pagination="151 - 206" refId="ref30642" refString="Shavrin, A. V. (2023 a) A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region. Zootaxa, 5369 (2), 151-206. https://doi.org/10.11646/zootaxa.5369.2.1" type="journal article" year="2023">Shavrin 2023a</bibRefCitation>
). Aedeagus as in
<figureCitation id="40F39FAAFF836C4DFE34FB3BFD9CFB45" box="[425,551,1179,1205]" captionStart="FIGURES 3438" captionStartId="11.[151,264,1758,1782]" captionTargetBox="[160,1427,190,1725]" captionTargetId="figure-16@11.[151,1436,181,1734]" captionTargetPageId="11" captionText="FIGURES 3438. Aedeagus of Omalium (Figs 3436—O. pratense (holotype), Figs 3738—O. alticola (Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="13" pageNumber="514">Figs 3738</figureCitation>
and in
<figureCitation id="40F39FAAFF836C4DFD1DFB3BFCBBFB45" box="[640,768,1178,1205]" captionStart="FIGURES 4046" captionStartId="15.[151,264,1875,1899]" captionTargetBox="[159,1418,190,1844]" captionTargetId="figure-16@15.[151,1436,181,1852]" captionTargetPageId="15" captionText="FIGURES 4046. Aedeagus of Omalium (Figs 4043—O. cocleare (paratypes), Figs 4446—O. contiger (paratype)): 40, 4344—aedeagus, parameral view, 41, 45—aedeagus, lateral view, 42, 46—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896701" httpUri="https://zenodo.org/record/14896701/files/figure.png" pageId="13" pageNumber="514">
Figs
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in
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. It is here recorded from
<collectingCountry id="A0DFC3BFFF836C4DFA98FB3BFAF3FB45" box="[1285,1352,1179,1205]" name="China" pageId="13" pageNumber="514">China</collectingCountry>
for the first time.
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<mods:part id="66446FFEADE0E0C2BCDEFE09DAEE0AA7">
<mods:date id="BC1B4468F91EA85BD161FEDC2373EB55">2025</mods:date>
<mods:detail id="033069013B0AB4315181B0640732F869" type="pubDate">
<mods:number id="C6A13BC2C5E491937CB8BB728E3DAF81">2025-02-18</mods:number>
</mods:detail>
<mods:detail id="39CD4D9F5BC2B287181AFF0C2BF64ECC" type="volume">
<mods:number id="6E54289F0D6201B73A3D118430DC1711">5588</mods:number>
</mods:detail>
<mods:detail id="4AC16F6F28D180B85D3BD87247E66231" type="issue">
<mods:number id="169A9EBC15B474F1EC2263FDBED1E361">4</mods:number>
</mods:detail>
<mods:extent id="9BB4A3063A243B7A2057884E29FAFEC7" unit="page">
<mods:start id="2FF467C8A859B8C24AB98618AD96863A">501</mods:start>
<mods:end id="75F97F1462957CD3938C7182D76B513E">543</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="463A030F78202176D379ECF77D5C168C">
<mods:url id="FDA6DCE21B351BA767253B97866EA5BE">https://doi.org/10.11646/zootaxa.5588.4.1</mods:url>
</mods:location>
<mods:classification id="61D1AF0F1528360BC638A551DD101FE5">journal article</mods:classification>
<mods:identifier id="6628BB3DAB3D2E4F35CCA4E60ACD1627" type="DOI">10.11646/zootaxa.5588.4.1</mods:identifier>
<mods:identifier id="60D83DEF1BEC7BB3259C5E5026BC85B1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A943E4B95983192BA6756A61D14A808E" type="Zenodo-Dep">14896684</mods:identifier>
<mods:identifier id="2D4135DA8C6D49C3EFB91A88F139F33B" type="ZooBank">444F5F59-9228-4100-9135-DF824FF9BACA</mods:identifier>
</mods:mods>
<treatment id="50613239FF836C51FF0AFA8AFB79FE79" LSID="urn:lsid:plazi:treatment:50613239FF836C51FF0AFA8AFB79FE79" httpUri="http://treatment.plazi.org/id/50613239FF836C51FF0AFA8AFB79FE79" lastPageId="17" lastPageNumber="518" pageId="13" pageNumber="514">
<subSubSection id="90D2D0A4FF836C4DFF0AFA8AFD9FFAB5" box="[151,548,1322,1349]" pageId="13" pageNumber="514" type="nomenclature">
<paragraph id="D877832FFF836C4DFF0AFA8AFD9FFAB5" blockId="13.[151,548,1322,1385]" box="[151,548,1322,1349]" pageId="13" pageNumber="514">
<heading id="833F3443FF836C4DFF0AFA8AFD9FFAB5" bold="true" box="[151,548,1322,1349]" fontSize="11" level="1" pageId="13" pageNumber="514" reason="1">
<emphasis id="EABC5F3DFF836C4DFF0AFA8AFD9FFAB5" bold="true" box="[151,548,1322,1349]" pageId="13" pageNumber="514">
3.4.2.
<taxonomicName id="1FC8F8ACFF836C4DFF7CFA8AFE04FAB5" authorityName="Shavrin" authorityYear="2025" box="[225,447,1322,1349]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="cocleare" status="sp. nov.">
<emphasis id="EABC5F3DFF836C4DFF7CFA8AFE04FAB5" bold="true" box="[225,447,1322,1349]" italics="true" pageId="13" pageNumber="514">Omalium cocleare</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F18FE246FF836C4DFE5BFA8BFD9FFAB5" box="[454,548,1323,1349]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF836C4DFF0AFAEFFE42FA99" box="[151,505,1359,1385]" pageId="13" pageNumber="514" type="description">
<paragraph id="D877832FFF836C4DFF0AFAEFFE42FA99" blockId="13.[151,548,1322,1385]" box="[151,505,1359,1385]" pageId="13" pageNumber="514">
(
<figureCitation id="40F39FAAFF836C4DFF02FAEFFE9AFA99" box="[159,289,1359,1385]" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="13" pageNumber="514">Figs 1617</figureCitation>
, 32, 4043, 47, 49)
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF836C4EFF0AFA37FE11FD3D" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514" type="materials_examined">
<paragraph id="D877832FFF836C4DFF0AFA37FAEFFA09" blockId="13.[151,1437,1430,2033]" pageId="13" pageNumber="514">
<emphasis id="EABC5F3DFF836C4DFF0AFA37FE05FA40" bold="true" box="[151,446,1430,1457]" pageId="13" pageNumber="514">Type material examined.</emphasis>
<materialsCitation id="68A08972FF836C4DFE58FA37FB1EF9B1" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<typeStatus id="07733D8DFF836C4DFE58FA37FD96FA41" box="[453,557,1431,1457]" pageId="13" pageNumber="514" type="holotype">Holotype</typeStatus>
<specimenCount id="CECE48A6FF836C4DFDA9FA37FDF2FA41" box="[564,585,1431,1457]" count="1" pageId="13" pageNumber="514" type="male"></specimenCount>
(
<figureCitation id="40F39FAAFF836C4DFDC5FA37FD17FA41" box="[600,684,1431,1457]" captionStart="FIGURES 3133" captionStartId="10.[151,264,1388,1412]" captionTargetBox="[162,1423,536,1356]" captionTargetId="figure-96@10.[151,1435,525,1364]" captionTargetPageId="10" captionText="FIGURES 3133. Habitus of Omalium: 31—O. pratense (holotype), 32—O. cocleare (holotype), 33—O. contiger (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896692" httpUri="https://zenodo.org/record/14896692/files/figure.png" pageId="13" pageNumber="514">Fig. 32</figureCitation>
):
<emphasis id="EABC5F3DFF836C4DFD22FA36FC2CFA41" bold="true" box="[703,919,1430,1457]" pageId="13" pageNumber="514">
<collectingCountry id="A0DFC3BFFF836C4DFD22FA36FC9AFA40" box="[703,801,1430,1456]" name="China" pageId="13" pageNumber="514">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF836C4DFCB1FA36FC35FA40" box="[812,910,1430,1456]" country="China" name="Yunnan" pageId="13" pageNumber="514">YUNNAN</collectingRegion>
:
</emphasis>
CHINA: Yunnan, Dali | Cangshan ca
<quantity id="1F302ECAFF836C4DFAD5FA36FA27FA41" box="[1352,1436,1430,1457]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="514" unit="m" value="2500.0">2500 m</quantity>
| 10..
<date id="AC76A5EFFF836C4DFF54FA1BFE9DFA25" box="[201,294,1467,1493]" pageId="13" pageNumber="514" value="2003-04">
IV.
<quantity id="1F302ECAFF836C4DFF76FA1BFE9DFA25" box="[235,294,1467,1493]" metricMagnitude="1" metricUnit="m" metricValue="5.087619999999999" pageId="13" pageNumber="514" unit="in" value="2003.0">2003</quantity>
</date>
in moss | G. de. Rougemont leg. &lt;printed&gt;,
<taxonomicName id="1FC8F8ACFF836C4DFC83FA1BFC36FA25" authorityName="Gravenhorst" authorityYear="1802" box="[798,909,1467,1493]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">Omalium</taxonomicName>
| n. sp. | det. 2002 | G. de Rougemont [printed] &lt;handwritten&gt;, HOLOTYPE |
<taxonomicName id="1FC8F8ACFF836C4DFE66FA7FFDD8FA09" authorityName="Gravenhorst" authorityYear="1802" box="[507,611,1503,1529]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF836C4DFE66FA7FFDD8FA09" box="[507,611,1503,1529]" italics="true" pageId="13" pageNumber="514">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF836C4DFDEAFA7FFD6DFA09" authorityName="Shavrin" authorityYear="2025" box="[631,726,1503,1529]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="cocleare" status="sp. nov.">
<emphasis id="EABC5F3DFF836C4DFDEAFA7FFD6DFA09" box="[631,726,1503,1529]" italics="true" pageId="13" pageNumber="514">cocleare</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF836C4DFD40FA7FFC8CFA09" bold="true" box="[733,823,1503,1529]" pageId="13" pageNumber="514">
<taxonomicNameLabel id="F18FE246FF836C4DFD40FA7FFC8CFA09" box="[733,823,1503,1529]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025 &lt;printed&gt; (
<collectionCode id="BED91BEAFF836C4DFB45FA7FFAF2FA09" box="[1240,1353,1503,1529]" pageId="13" pageNumber="514">OUMNH</collectionCode>
).
<paragraph id="D877832FFF836C4EFF5AF9A3FE11FD3D" blockId="13.[151,1437,1430,2033]" lastBlockId="14.[151,1437,150,2013]" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">
<emphasis id="EABC5F3DFF836C4DFF5AF9A3FE84F9ED" bold="true" box="[199,319,1539,1565]" pageId="13" pageNumber="514">
<typeStatus id="07733D8DFF836C4DFF5AF9A3FE84F9ED" box="[199,319,1539,1565]" pageId="13" pageNumber="514" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="CECE48A6FF836C4DFED2F9A3FE24F9ED" box="[335,415,1539,1565]" count="16" pageId="13" pageNumber="514" type="male">16 ♂♂</specimenCount>
,
<specimenCount id="CECE48A6FF836C4DFE36F9A3FE53F9ED" box="[427,488,1539,1565]" count="9" pageId="13" pageNumber="514" type="female">9 ♀♀</specimenCount>
):
<emphasis id="EABC5F3DFF836C4DFD99F9A2FD67F9ED" bold="true" box="[516,732,1538,1565]" pageId="13" pageNumber="514">
<collectingCountry id="A0DFC3BFFF836C4DFD99F9A2FDDDF9EC" box="[516,614,1538,1564]" name="China" pageId="13" pageNumber="514">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF836C4DFDECF9A2FD68F9EC" box="[625,723,1538,1564]" country="China" name="Yunnan" pageId="13" pageNumber="514">YUNNAN</collectingRegion>
:
</emphasis>
<specimenCount id="CECE48A6FF836C4DFD7EF9A3FC98F9ED" box="[739,803,1539,1565]" count="2" pageId="13" pageNumber="514" type="male">2 ♂♂</specimenCount>
(
<specimenCount id="CECE48A6FF836C4DFCAFF9A3FC76F9ED" box="[818,973,1539,1565]" count="1" pageId="13" pageNumber="514" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="CECE48A6FF836C4DFBCFF9A3FBC5F9ED" box="[1106,1150,1539,1565]" count="1" pageId="13" pageNumber="514" type="female">1 ♀</specimenCount>
: CHINA:
<collectingRegion id="1A0C4DCDFF836C4DFB66F9A3FAEFF9ED" box="[1275,1364,1539,1565]" country="China" name="Yunnan" pageId="13" pageNumber="514">Yunnan</collectingRegion>
, Dali | Cangshan ca
<quantity id="1F302ECAFF836C4DFEB1F986FE3AF9B1" box="[300,385,1574,1601]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="514" unit="m" value="2500.0">2500 m</quantity>
|
<date id="AC76A5EFFF836C4DFE09F987FDABF9B1" box="[404,528,1575,1601]" pageId="13" pageNumber="514" value="2003-04-10">10.IV.2003</date>
stream moss | G. de. Rougemont leg. &lt;printed&gt; (
<specimenCount id="CECE48A6FF836C4DFBDFF987FBD6F9B1" box="[1090,1133,1575,1601]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
:
<collectionCode id="BED91BEAFF836C4DFBE5F987FB1EF9B1" box="[1144,1189,1575,1601]" pageId="13" pageNumber="514">cSh</collectionCode>
;
<materialsCitation id="68A08972FF836C4DFB2CF987FA23F9B1" box="[1201,1432,1575,1601]" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFB2CF987FB66F9B1" box="[1201,1245,1575,1601]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
,
<specimenCount id="CECE48A6FF836C4DFB74F987FAAEF9B1" box="[1257,1301,1575,1601]" count="1" pageId="13" pageNumber="514" type="female">1 ♀</specimenCount>
:
<collectionCode id="BED91BEAFF836C4DFABCF987FA2BF9B1" box="[1313,1424,1575,1601]" pageId="13" pageNumber="514">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFF0AF9EBFE82F995" box="[151,313,1611,1637]" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFF0AF9EBFF7BF995" box="[151,192,1611,1637]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
(dissected
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFEDEF9EBFC6DF979" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
left paramere missing): P.R. CHINA,
<collectingRegion id="1A0C4DCDFF836C4DFD76F9EBFCFFF995" box="[747,836,1611,1637]" country="China" name="Yunnan" pageId="13" pageNumber="514">Yunnan</collectingRegion>
, E | slope Cangshan at Dali, | N
<geoCoordinate id="BDFCE5E8FF836C4DFB00F9EAFAFFF995" box="[1181,1348,1610,1637]" degrees="25" direction="east" minutes="40" orientation="longitude" pageId="13" pageNumber="514" precision="1" seconds="15.1" value="25.67086">25°40´15.1´´ E</geoCoordinate>
100°07´| 39.9´´,
<date id="AC76A5EFFF836C4DFF6AF9CFFED3F979" box="[247,360,1647,1673]" pageId="13" pageNumber="514" value="2010-05-10">10.v.2010</date>
,
<quantity id="1F302ECAFF836C4DFEE9F9CFFE7FF979" box="[372,452,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="2.711" pageId="13" pageNumber="514" unit="m" value="2711.0">2711m</quantity>
, | sifting04, V. Grebennikov &lt;printed&gt; (
<collectionCode id="BED91BEAFF836C4DFC12F9CFFC75F979" box="[911,974,1647,1673]" pageId="13" pageNumber="514">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFC7EF9CFFD3DF921" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFC7EF9CFFB98F979" box="[995,1059,1647,1673]" count="2" pageId="13" pageNumber="514" type="male">2 ♂♂</specimenCount>
(
<specimenCount id="CECE48A6FF836C4DFBAFF9CFFB75F979" box="[1074,1230,1647,1673]" count="1" pageId="13" pageNumber="514" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="CECE48A6FF836C4DFAC9F9CEFA2DF979" box="[1364,1430,1646,1673]" count="5" pageId="13" pageNumber="514" type="female">5 ♀♀</specimenCount>
: (
<specimenCount id="CECE48A6FF836C4DFF02F933FEF2F95D" box="[159,329,1683,1709]" count="2" pageId="13" pageNumber="514" type="generic">two specimens</specimenCount>
dissected): CHINA:
<collectingRegion id="1A0C4DCDFF836C4DFDDAF933FD1AF95D" box="[583,673,1683,1709]" country="China" name="Yunnan" pageId="13" pageNumber="514">Yunnan</collectingRegion>
: Lijiang | Yushuizhai ca
<quantity id="1F302ECAFF836C4DFC22F933FBACF95D" box="[959,1047,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="514" unit="m" value="2600.0">2600 m</quantity>
|
<date id="AC76A5EFFF836C4DFBADF933FB17F95D" box="[1072,1196,1683,1709]" pageId="13" pageNumber="514" value="2003-04-14">14.IV.2003</date>
stream moss | G. de Rougemont leg. &lt;printed&gt; (
<specimenCount id="CECE48A6FF836C4DFE4AF917FDBAF921" box="[471,513,1719,1745]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
,
<specimenCount id="CECE48A6FF836C4DFD91F917FDF7F921" box="[524,588,1719,1745]" count="3" pageId="13" pageNumber="514" type="female">3 ♀♀</specimenCount>
:
<collectionCode id="BED91BEAFF836C4DFDC4F917FD3DF921" box="[601,646,1719,1745]" pageId="13" pageNumber="514">cSh</collectionCode>
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFD0CF917FC30F921" box="[657,907,1719,1745]" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFD0CF917FD07F921" box="[657,700,1719,1745]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
,
<specimenCount id="CECE48A6FF836C4DFD5AF917FCB3F921" box="[711,776,1719,1745]" count="2" pageId="13" pageNumber="514" type="female">2 ♀♀</specimenCount>
:
<collectionCode id="BED91BEAFF836C4DFC89F917FC38F921" box="[788,899,1719,1745]" pageId="13" pageNumber="514">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFC08F917FB93F905" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFC08F917FC7AF921" box="[917,961,1719,1745]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
: CHINA:
<collectingRegion id="1A0C4DCDFF836C4DFBA6F917FB2FF921" box="[1083,1172,1719,1745]" country="China" name="Yunnan" pageId="13" pageNumber="514">Yunnan</collectingRegion>
, Lijiang | Yushuizhai ca
<quantity id="1F302ECAFF836C4DFF0AF97AFF56F905" box="[151,237,1754,1781]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="514" unit="m" value="2500.0">2500 m</quantity>
|
<date id="AC76A5EFFF836C4DFE9EF97BFEC5F905" box="[259,382,1755,1781]" pageId="13" pageNumber="514" value="2003-04-14">14.IV.2003</date>
stream moss | G. de Rougemont leg. &lt;printed&gt; (
<collectionCode id="BED91BEAFF836C4DFC32F97BFBA4F905" box="[943,1055,1755,1781]" pageId="13" pageNumber="514">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFBA9F97BFC31F8CD" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFBA9F97BFBD9F905" box="[1076,1122,1755,1781]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
: CHINA
<date id="AC76A5EFFF836C4DFB4AF97BFA91F905" box="[1239,1322,1755,1781]" pageId="13" pageNumber="514" value="1986-10">X.1986</date>
|
<collectingRegion id="1A0C4DCDFF836C4DFAA2F97BFA22F905" box="[1343,1433,1755,1781]" country="China" name="Yunnan" pageId="13" pageNumber="514">Yunnan</collectingRegion>
: Kunming | G. de Rougemont &lt;printed&gt;,
<taxonomicName id="1FC8F8ACFF836C4DFDF8F95FFD40F8E9" box="[613,763,1791,1817]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="undetermined">Omalium sp.</taxonomicName>
[handwritten] |
<taxonomicName id="1FC8F8ACFF836C4DFC36F95FFB91F8E9" authorityName="Scheerpeltz" authorityYear="1965" box="[939,1066,1791,1817]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="malaisei">O. malaisei</taxonomicName>
[handwritten] | det. 198 [printed] Scheerp? [handwritten] | G. de. Rougemont &lt;printed&gt; (
<collectionCode id="BED91BEAFF836C4DFC8FF883FC39F8CD" box="[786,898,1827,1853]" pageId="13" pageNumber="514">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4DFC0BF883FC49F891" lastPageId="13" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFC0BF883FC7EF8CD" box="[918,965,1827,1853]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
: CHINA:
<collectingRegion id="1A0C4DCDFF836C4DFBDFF883FB20F8CD" box="[1090,1179,1827,1853]" country="China" name="Yunnan" pageId="13" pageNumber="514">Yunnan</collectingRegion>
, Lijiang | Yushuizai ca
<quantity id="1F302ECAFF836C4DFF0AF8E6FF56F891" box="[151,237,1862,1889]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="514" unit="m" value="2500.0">2500 m</quantity>
|
<date id="AC76A5EFFF836C4DFE9CF8E7FEC6F891" box="[257,381,1863,1889]" pageId="13" pageNumber="514" value="2003-04-14">14.IV.2003</date>
stream moss | G. de Rougemont &lt;printed&gt; (
<collectionCode id="BED91BEAFF836C4DFCE4F8E7FC52F891" box="[889,1001,1863,1889]" pageId="13" pageNumber="514">OUMNH</collectionCode>
)
</materialsCitation>
;
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<specimenCount id="CECE48A6FF836C4DFC60F8E7FB93F891" box="[1021,1064,1863,1889]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
(dissected): CHINA: N-Yunnan [C03-19B] | Dali Bai Nat. Aut. Pref., Diancang Shan, |
<quantity id="1F302ECAFF836C4DFD62F8CBFC83F875" box="[767,824,1899,1925]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="514" unit="km" value="3.0">3 km</quantity>
W Dali old town, | pine forest at “Cloud Road”, | right upper chairlift station, |
<geoCoordinate id="BDFCE5E8FF836C4DFE3FF82EFDA1F859" box="[418,538,1934,1961]" degrees="25" direction="north" minutes="41.1" orientation="latitude" pageId="13" pageNumber="514" precision="92" value="25.685">25°41.1´N</geoCoordinate>
,
<geoCoordinate id="BDFCE5E8FF836C4DFDB8F82FFD12F859" box="[549,681,1935,1961]" degrees="100" direction="east" minutes="06.8" orientation="longitude" pageId="13" pageNumber="514" precision="92" value="100.113335">100°06.8´E</geoCoordinate>
,
<quantity id="1F302ECAFF836C4DFD29F82EFCF5F859" box="[692,846,1934,1961]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.75" metricValueMin="2.65" pageId="13" pageNumber="514" unit="m" value="2700.0" valueMax="2750.0" valueMin="2650.0">26502750m</quantity>
, &lt;printed&gt;, [C03-19B] pine needles, moss | (dry) in ditches, mushrooms, tra[?] |
<date id="AC76A5EFFF836C4DFE4AF813FDF1F83D" box="[471,586,1971,1997]" pageId="13" pageNumber="514" value="2003-09-01">1.IX.2003</date>
, leg. M. Schülke &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke (
<collectionCode id="BED91BEAFF836C4DFEB3F877FED1F801" box="[302,362,2007,2033]" pageId="13" pageNumber="514">cSch</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF836C4EFEE0F877FF58FF25" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">
<specimenCount id="CECE48A6FF836C4DFEE0F877FE11F801" box="[381,426,2007,2033]" count="1" pageId="13" pageNumber="514" type="male">1 ♂</specimenCount>
: CHINA: N-Yunnan [C2005-11] | Dali Bai Nat. Aut. Pref., Diancang | Shan,
<quantity id="1F302ECAFF836C4DFA90F877FAFEF801" box="[1293,1349,2007,2033]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="514" unit="km" value="3.0">3 km</quantity>
W Dali old town, pine | forest at “Cloud Road”,
<geoCoordinate id="BDFCE5E8FF806C4EFDD2FF36FD7CFF41" box="[591,711,150,177]" degrees="25" direction="north" minutes="41.1" orientation="latitude" pageId="14" pageNumber="515" precision="92" value="25.685">25°41.1´N</geoCoordinate>
, |
<geoCoordinate id="BDFCE5E8FF806C4EFD41FF37FCDBFF41" box="[732,864,151,177]" degrees="100" direction="east" minutes="06.8" orientation="longitude" pageId="14" pageNumber="515" precision="92" value="100.113335">100°06.8´E</geoCoordinate>
,
<quantity id="1F302ECAFF806C4EFCF7FF36FBBBFF41" box="[874,1024,150,177]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.75" metricValueMin="2.65" pageId="14" pageNumber="515" unit="m" value="2700.0" valueMax="2750.0" valueMin="2650.0">26502750m</quantity>
, |
<date id="AC76A5EFFF806C4EFB88FF36FB2DFF41" box="[1045,1174,150,177]" pageId="14" pageNumber="515" value="2005-06-17">17.VI.2005</date>
, M. Schülke &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFF02FF1BFF60FF25" box="[159,219,187,213]" pageId="14" pageNumber="515">cSch</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFF6CFF1BFAA0FEED" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFF6CFF1BFEA5FF25" box="[241,286,187,213]" count="1" pageId="14" pageNumber="515" type="male">1 ♂</specimenCount>
(dissected): CHINA (
<collectingRegion id="1A0C4DCDFF806C4EFDBFFF1BFDC0FF25" box="[546,635,187,213]" country="China" name="Yunnan" pageId="14" pageNumber="515">Yunnan</collectingRegion>
) Dali Bai Aut. Pref., | mount range E Weishan,
<quantity id="1F302ECAFF806C4EFB3BFF1BFB54FF25" box="[1190,1263,187,213]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="515" unit="km" value="12.0">12 km</quantity>
NE | Weishan,
<quantity id="1F302ECAFF806C4EFF0AFF7FFE89FF09" box="[151,306,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.645" metricValueMax="2.66" metricValueMin="2.63" pageId="14" pageNumber="515" unit="m" value="2645.0" valueMax="2660.0" valueMin="2630.0">26302660 m</quantity>
(scrub with | pines and bamboo, litter sifted) | 25°17´02-15´´N / 100°22´23-30´´E |
<date id="AC76A5EFFF806C4EFB46FF7EFAE0FF09" box="[1243,1371,222,249]" pageId="14" pageNumber="515" value="2009-09-15">15.IX.2009</date>
D.W. Wrase [54A] &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFB79FEA3FAA8FEED" box="[1252,1299,259,285]" pageId="14" pageNumber="515">cSh</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFAB4FEA3FBEAFE95" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFAB4FEA3FAEEFEED" box="[1321,1365,259,285]" count="1" pageId="14" pageNumber="515" type="female">1 ♀</specimenCount>
(right antennomeres 311 missing): CHINA: N-Yunnan Diqing
<collectingRegion id="1A0C4DCDFF806C4EFCBCFE87FCE0FEB1" box="[801,859,295,321]" country="China" name="Xizang" pageId="14" pageNumber="515">Tibet</collectingRegion>
| Aut.Pr.Bitai Hai Lake area |
<quantity id="1F302ECAFF806C4EFB38FE87FB5EFEB1" box="[1189,1253,295,321]" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="515" unit="km" value="29.0">29km</quantity>
ESE Zhongdian
<geoCoordinate id="BDFCE5E8FF806C4EFF0AFEEAFEA0FE95" box="[151,283,330,357]" degrees="27" direction="north" minutes="43.65" orientation="latitude" pageId="14" pageNumber="515" precision="9" value="27.7275">27°43.65´N</geoCoordinate>
|
<geoCoordinate id="BDFCE5E8FF806C4EFEABFEEAFE0CFE95" box="[310,439,330,357]" degrees="99" direction="east" minutes="58.97" orientation="longitude" pageId="14" pageNumber="515" precision="9" value="99.982834">99°58.97´E</geoCoordinate>
<quantity id="1F302ECAFF806C4EFE5CFEEAFDB4FE95" box="[449,527,330,357]" metricMagnitude="3" metricUnit="m" metricValue="3.54" pageId="14" pageNumber="515" unit="m" value="3540.0">3540m</quantity>
<date id="AC76A5EFFF806C4EFD87FEEAFD30FE95" box="[538,651,330,357]" pageId="14" pageNumber="515" value="2005-06-01">1.VI.2005</date>
| A. Smetana [C147] &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFB87FEEBFBF2FE95" box="[1050,1097,331,357]" pageId="14" pageNumber="515">cSh</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFBFDFEEBFC8CFE5D" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFBFDFEEBFB2BFE95" box="[1120,1168,331,357]" count="1" pageId="14" pageNumber="515" type="female">1 ♀</specimenCount>
: (left antennomeres 7 11 missing): CHINA: N-Yunnan Diqing
<collectingRegion id="1A0C4DCDFF806C4EFDE0FECFFD00FE79" box="[637,699,367,393]" country="China" name="Xizang" pageId="14" pageNumber="515">Tibet</collectingRegion>
. | Aut.Pr.Zhongdian Co.
<quantity id="1F302ECAFF806C4EFC42FECEFB91FE79" box="[991,1066,366,393]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="515" unit="km" value="35.0">35 km</quantity>
ESE | Zhongdian
<geoCoordinate id="BDFCE5E8FF806C4EFA95FECFFA30FE79" box="[1288,1419,367,393]" degrees="27" direction="north" minutes="41.00" orientation="latitude" pageId="14" pageNumber="515" precision="9" value="27.683332">27°41.00´N</geoCoordinate>
|
<geoCoordinate id="BDFCE5E8FF806C4EFF0AFE33FE9DFE5D" box="[151,294,403,429]" degrees="100" direction="east" minutes="01.47" orientation="longitude" pageId="14" pageNumber="515" precision="9" value="100.0245">100°01.47´E</geoCoordinate>
<quantity id="1F302ECAFF806C4EFEB1FE32FEC1FE5D" box="[300,378,402,429]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="14" pageNumber="515" unit="m" value="3450.0">3450m</quantity>
|
<date id="AC76A5EFFF806C4EFE11FE32FE46FE5D" box="[396,509,402,429]" pageId="14" pageNumber="515" value="2005-06-03">3.VI.2005</date>
A. Smetana [C150] (
<collectionCode id="BED91BEAFF806C4EFD72FE33FC95FE5D" box="[751,814,403,429]" pageId="14" pageNumber="515">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFCDFFE32FEACFE05" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<emphasis id="EABC5F3DFF806C4EFCDFFE32FC0AFE5C" bold="true" box="[834,945,402,428]" pageId="14" pageNumber="515">
<collectingRegion id="1A0C4DCDFF806C4EFCDFFE32FC16FE5C" box="[834,941,402,428]" country="China" name="Sichuan" pageId="14" pageNumber="515">SICHUAN</collectingRegion>
:
</emphasis>
<specimenCount id="CECE48A6FF806C4EFC2AFE33FC5AFE5D" box="[951,993,403,429]" count="1" pageId="14" pageNumber="515" type="male">1 ♂</specimenCount>
(dissected): CHINA
<date id="AC76A5EFFF806C4EFB48FE33FA93FE5D" box="[1237,1320,403,429]" pageId="14" pageNumber="515" value="1986-10">X.1986</date>
|
<collectingRegion id="1A0C4DCDFF806C4EFAA7FE33FA22FE5D" box="[1338,1433,403,429]" country="China" name="Sichuan" pageId="14" pageNumber="515">Sichuan</collectingRegion>
: Emei Shan | G. de Rougemont &lt;printed&gt;,
<taxonomicName id="1FC8F8ACFF806C4EFD16FE17FC98FE21" box="[651,803,439,465]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="undetermined">Omalium sp.</taxonomicName>
[handwritten] | det. 198 | G. de Rougemont &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFF02FE7BFEB4FE05" box="[159,271,475,501]" pageId="14" pageNumber="515">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFEBFFE7BFEB8FDE9" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFEBFFE7BFEF5FE05" box="[290,334,475,501]" count="1" pageId="14" pageNumber="515" type="male">1 ♂</specimenCount>
: China S
<collectingRegion id="1A0C4DCDFF806C4EFE5EFE7BFDA6FE05" box="[451,541,475,501]" country="China" name="Sichuan" pageId="14" pageNumber="515">Sichuan</collectingRegion>
| S Xichang Mt. Luoji | 23002500 litter | 1624.07.[19]96 Kurbatov &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFF02FE5FFF40FDE9" box="[159,251,511,537]" pageId="14" pageNumber="515">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFE92FE5FFB31FDCD" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFE92FE5FFE87FDE9" box="[271,316,511,537]" count="1" pageId="14" pageNumber="515" type="female">1 ♀</specimenCount>
: CHINA:
<collectingRegion id="1A0C4DCDFF806C4EFE21FE5FFDA2FDE9" box="[444,537,511,537]" country="China" name="Sichuan" pageId="14" pageNumber="515">Sichuan</collectingRegion>
, |
<quantity id="1F302ECAFF806C4EFDAEFE5EFDC0FDE9" box="[563,635,510,537]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="515" unit="km" value="50.0">50 km</quantity>
W Dayi, Xiling | mts.,
<quantity id="1F302ECAFF806C4EFC1BFE5FFC65FDE9" box="[902,990,511,537]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="515" unit="m" value="2300.0">2300 m</quantity>
,
<date id="AC76A5EFFF806C4EFC76FE5EFBC8FDE9" box="[1003,1139,510,537]" pageId="14" pageNumber="515" value="2006-05-05" valueMax="2006-05-08" valueMin="2006-05-05">5.8.V.2006</date>
| S. Murzin &amp; I. Shokhin &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFBDBFD83FB39FDCD" box="[1094,1154,547,573]" pageId="14" pageNumber="515">cSch</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFB0AFD83FB24FD91" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFB0AFD83FB7CFDCD" box="[1175,1223,547,573]" count="1" pageId="14" pageNumber="515" type="male">1 ♂</specimenCount>
: CHINA:
<collectingRegion id="1A0C4DCDFF806C4EFADAFD83FA26FDCD" box="[1351,1437,547,573]" country="China" name="Yunnan" pageId="14" pageNumber="515">Yunnan</collectingRegion>
| above Dali,
<quantity id="1F302ECAFF806C4EFEAEFDE7FE6BFD91" box="[307,464,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="14" pageNumber="515" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">20002200 m</quantity>
|
<date id="AC76A5EFFF806C4EFE75FDE7FD3CFD91" box="[488,647,583,609]" pageId="14" pageNumber="515" value="1999-04-04" valueMax="1999-04-17" valueMin="1999-04-04">4.17.IV.1999</date>
| leg. W. SCHAWALLER &lt;printed&gt; (
<collectionCode id="BED91BEAFF806C4EFBD8FDE7FB2CFD91" box="[1093,1175,583,609]" pageId="14" pageNumber="515">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFB30FDE7FF53FD59" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFB30FDE7FB55FD91" box="[1197,1262,583,609]" count="2" pageId="14" pageNumber="515" type="male">2 ♂♂</specimenCount>
(
<specimenCount id="CECE48A6FF806C4EFB62FDE7FA27FD91" box="[1279,1436,583,609]" count="1" pageId="14" pageNumber="515" type="generic">one specimen</specimenCount>
dissected): CHINA:
<collectingRegion id="1A0C4DCDFF806C4EFE14FDCBFE64FD75" box="[393,479,619,645]" country="China" name="Yunnan" pageId="14" pageNumber="515">Yunnan</collectingRegion>
| above Dali,
<quantity id="1F302ECAFF806C4EFD1CFDCAFCA5FD75" box="[641,798,618,645]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="14" pageNumber="515" unit="m" value="2600.0" valueMax="2700.0" valueMin="2500.0">25002700 m</quantity>
|
<date id="AC76A5EFFF806C4EFCA9FDCBFC6FFD75" box="[820,980,619,645]" pageId="14" pageNumber="515" value="1999-04-08" valueMax="1999-04-18" valueMin="1999-04-08">8.18.IV.1999</date>
| leg. W. SCHAWALLER &lt;printed&gt; (
<specimenCount id="CECE48A6FF806C4EFA0DFDCBFF14FD59" count="1" pageId="14" pageNumber="515" type="male">1 ♂</specimenCount>
:
<collectionCode id="BED91BEAFF806C4EFF26FD2FFF53FD59" box="[187,232,655,681]" pageId="14" pageNumber="515">cSh</collectionCode>
</materialsCitation>
;
<materialsCitation id="68A08972FF806C4EFF68FD2FFE30FD59" box="[245,395,655,681]" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
<specimenCount id="CECE48A6FF806C4EFF68FD2FFE99FD59" box="[245,290,655,681]" count="1" pageId="14" pageNumber="515" type="male">1 ♂</specimenCount>
:
<collectionCode id="BED91BEAFF806C4EFEB3FD2FFEC4FD59" box="[302,383,655,681]" pageId="14" pageNumber="515">SMNS</collectionCode>
).
</materialsCitation>
<materialsCitation id="68A08972FF806C4EFE0FFD2FFE11FD3D" lastPageId="14" lastPageNumber="515" pageId="14" pageNumber="514">
All
<typeStatus id="07733D8DFF806C4EFE23FD2FFD92FD59" box="[446,553,655,681]" pageId="14" pageNumber="515" type="paratype">paratypes</typeStatus>
with additional red printed label:
<typeStatus id="07733D8DFF806C4EFC30FD2FFB86FD59" box="[941,1085,655,681]" pageId="14" pageNumber="515" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="1FC8F8ACFF806C4EFBCFFD2FFB01FD59" authorityName="Gravenhorst" authorityYear="1802" box="[1106,1210,655,681]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF806C4EFBCFFD2FFB01FD59" box="[1106,1210,655,681]" italics="true" pageId="14" pageNumber="515">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF806C4EFB53FD2FFA96FD59" authorityName="Shavrin" authorityYear="2025" box="[1230,1325,655,681]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="cocleare" status="sp. nov.">
<emphasis id="EABC5F3DFF806C4EFB53FD2FFA96FD59" box="[1230,1325,655,681]" italics="true" pageId="14" pageNumber="515">cocleare</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF806C4EFAA8FD2FFA34FD59" bold="true" box="[1333,1423,655,681]" pageId="14" pageNumber="515">
<taxonomicNameLabel id="F18FE246FF806C4EFAA8FD2FFA34FD59" box="[1333,1423,655,681]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025.
</materialsCitation>
</paragraph>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF806C50FF5AFD76FE48F9B5" lastPageId="16" lastPageNumber="517" pageId="14" pageNumber="515" type="description">
<paragraph id="D877832FFF806C4EFF5AFD76FEE8FCAD" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="515">
<emphasis id="EABC5F3DFF806C4EFF5AFD76FEE2FD00" bold="true" box="[199,345,726,752]" pageId="14" pageNumber="515">Description.</emphasis>
Measurements (n=26): HW: 0.490.52; HL: 0.360.40; OL: 0.130.15; TL: 0.07; AL (
<typeStatus id="07733D8DFF806C4EFAB5FD77FA2AFD01" box="[1320,1425,727,753]" pageId="14" pageNumber="515" type="holotype">holotype</typeStatus>
): 1.10; PL: 0.390.46; PWmax: 0.640.70; PWmin: 0.550.61; ESL: 0.850.95; EW: 0.880.96; MTbL (
<typeStatus id="07733D8DFF806C4EFAB5FD5BFA2AFCE5" box="[1320,1425,763,789]" pageId="14" pageNumber="515" type="holotype">holotype</typeStatus>
): 0.42; MTrL (
<typeStatus id="07733D8DFF806C4EFEB0FCBFFE2DFCC9" box="[301,406,799,825]" pageId="14" pageNumber="515" type="holotype">holotype</typeStatus>
): 0.25 (MTrL 14: 0.08; MTrL 5: 0.17); AW: 0.880.94; AedL: 0.580.62; BL: 2.703.56 (
<typeStatus id="07733D8DFF806C4EFF02FCE3FEBCFCAD" box="[159,263,835,861]" pageId="14" pageNumber="515" type="holotype">holotype</typeStatus>
: 3.55).
</paragraph>
<paragraph id="D877832FFF806C4EFF5AFCC7FB62FB19" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="515">
Habitus as in
<figureCitation id="40F39FAAFF806C4EFEFCFCC7FE0FFC71" box="[353,436,871,897]" captionStart="FIGURES 3133" captionStartId="10.[151,264,1388,1412]" captionTargetBox="[162,1423,536,1356]" captionTargetId="figure-96@10.[151,1435,525,1364]" captionTargetPageId="10" captionText="FIGURES 3133. Habitus of Omalium: 31—O. pratense (holotype), 32—O. cocleare (holotype), 33—O. contiger (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896692" httpUri="https://zenodo.org/record/14896692/files/figure.png" pageId="14" pageNumber="515">Fig. 32</figureCitation>
. Body brown to reddish-brown, with slightly darker head and abdomen (lateral and basal portions of abdomen, lateral and apical portions of elytra and paratergites of abdomen sometimes yellowish); antennomeres 611 or 711 brown; mouthparts, antennomeres 15 or 16 and legs yellowish. Punctation of head moderately dense, usually larger and deeper in middle, sometimes slightly finer in laterobasal portions of clypeus and on infraorbital portions, with interspaces between punctures in middle about as long as diameter of one-two nearest punctures; neck with dense and fine punctation, sparser in middle in several
<typeStatus id="07733D8DFF806C4EFBA4FBBBFB13FBC5" box="[1081,1192,1051,1077]" pageId="14" pageNumber="515" type="paratype">paratypes</typeStatus>
; pronotum with dense punctation about as that in middle part of head, distinctly sparser in mediobasal portion; punctation about as that on pronotum, sometimes slightly denser and coarser in basal portion and finer and sparser in middle, particularly along suture; abdomen with indistinct, fine and sparse or without visible punctation. Clypeus with distinct transverse microreticulation; scutellum with dense and fine isodiametric meshes; abdominal tergites with dense isodiametric microsculpture. Anterior and posterior margins of pronotum with indistinct short cuticular fringe.
</paragraph>
<paragraph id="D877832FFF806C4EFF5AFB53FE58F911" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="515">Head 1.3 times as broad as long, with strongly convex supra-antennal elevations and deep and relatively narrow anteriomedian depressions, reaching level of apical margins of eyes; posteriolateral margins of clypeus subdiagonally stretching posteriad and reaching level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with distinct diagonal elevations between punctures in basal portion of clypeus, with irregular elevations in middle and on postocular portions, sometimes forming indistinct four to six longitudinal wrinkles. Anteocellar foveae deep and moderately wide, suboval, indistinctly convergent latero-apicad toward level of posterior third or middle length of eyes. Temples 1.8 times to twice as long as longitudinal length of eye, from posterior margins of eyes gradually narrowed toward widely rounded hind angles. Apical part of neck sometimes narrowly depressed, with several short longitudinal elevations between punctures. Distance between ocelli about 1.8 times to twice as long as distance between ocellus and posterior margin of eye. Antenna with elongate antennomeres 57 and distinctly transverse 810; 4 short, about as long as broad, significantly shorter and slightly narrower than 3, 5 longer than 4, 6 slightly longer and broader than 5, 7 indistinctly broader than 6, 8 slightly shorter and distinctly broader than 7, 910 slightly broader than 8, apical antennomere about 1.3 times as long as 10.</paragraph>
<paragraph id="D877832FFF806C4EFF5AF94BFBC9F865" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="515">Pronotum 1.51.6 times as broad as long, 1.3 times as broad as head, widest in anterior third portion, distinctly more narrowed posteriad than anteriad. Apical angles not or slightly protruded anteriad. Lateral and laterobasal portions deeply and widely impressed. Surface of disc with two longitudinal depressions, significantly broadened basad; medioapical portion with indistinct or distinct and moderately shallow oval depression. Middle portions with indistinct or distinct irregular subdiagonal or longitudinal elevations between punctures.</paragraph>
<paragraph id="D877832FFF806C4EFF5AF83FFD0CF82D" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="515">Elytra indistinctly broader than long, twice as long as pronotum. Middle and apical portions of each elytron with irregular diagonal elevations between punctures.</paragraph>
<caption id="8CB7D3A7FF816C4FFF0AF8F3FAD2F87F" ID-DOI="http://doi.org/10.5281/zenodo.14896701" ID-Zenodo-Dep="14896701" httpUri="https://zenodo.org/record/14896701/files/figure.png" pageId="15" pageNumber="516" startId="15.[151,264,1875,1899]" targetBox="[159,1418,190,1844]" targetPageId="15" targetType="figure">
<paragraph id="D877832FFF816C4FFF0AF8F3FAD2F87F" blockId="15.[151,1436,1875,1936]" pageId="15" pageNumber="516">
<emphasis id="EABC5F3DFF816C4FFF0AF8F3FEECF89B" bold="true" box="[151,343,1875,1899]" pageId="15" pageNumber="516">FIGURES 4046.</emphasis>
Aedeagus of
<taxonomicName id="1FC8F8ACFF816C4FFE71F8F4FDF1F89C" authorityName="Gravenhorst" authorityYear="1802" box="[492,586,1876,1900]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF816C4FFE71F8F4FDF1F89C" box="[492,586,1876,1900]" italics="true" pageId="15" pageNumber="516">Omalium</emphasis>
</taxonomicName>
(Figs 4043—
<taxonomicName id="1FC8F8ACFF816C4FFD77F8F4FCD9F89C" authorityName="Shavrin" authorityYear="2025" box="[746,866,1876,1900]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="cocleare">
<emphasis id="EABC5F3DFF816C4FFD77F8F4FCD9F89C" box="[746,866,1876,1900]" italics="true" pageId="15" pageNumber="516">O. cocleare</emphasis>
</taxonomicName>
(paratypes), Figs 4446—
<taxonomicName id="1FC8F8ACFF816C4FFBE1F8F4FB49F89B" authorityName="Shavrin" authorityYear="2025" box="[1148,1266,1876,1899]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="contiger">
<emphasis id="EABC5F3DFF816C4FFBE1F8F4FB49F89B" box="[1148,1266,1876,1899]" italics="true" pageId="15" pageNumber="516">O. contiger</emphasis>
</taxonomicName>
(paratype)): 40, 4344—aedeagus, parameral view, 41, 45—aedeagus, lateral view, 42, 46—median lobe, lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="8CB7D3A7FF9E6C50FF0AFB95FBE2FBBD" ID-DOI="http://doi.org/10.5281/zenodo.14896705" ID-Zenodo-Dep="14896705" box="[151,1113,1077,1101]" httpUri="https://zenodo.org/record/14896705/files/figure.png" pageId="16" pageNumber="517" startId="16.[151,250,1077,1101]" targetBox="[151,1435,181,1052]" targetPageId="16" targetType="figure">
<paragraph id="D877832FFF9E6C50FF0AFB95FBE2FBBD" blockId="16.[151,1113,1077,1101]" box="[151,1113,1077,1101]" pageId="16" pageNumber="517">
<emphasis id="EABC5F3DFF9E6C50FF0AFB95FE9BFBBD" bold="true" box="[151,288,1077,1101]" pageId="16" pageNumber="517">FIGURE 47.</emphasis>
Distribution of
<taxonomicName id="1FC8F8ACFF9E6C50FE5FFB95FD99FBBD" authorityName="Gravenhorst" authorityYear="1802" box="[450,546,1077,1101]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="genus">Omalium</taxonomicName>
in China:
<taxonomicName id="1FC8F8ACFF9E6C50FD11FB95FCBAFBBD" authorityName="Shavrin" authorityYear="2025" box="[652,769,1077,1101]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="cocleare">
<emphasis id="EABC5F3DFF9E6C50FD11FB95FCBAFBBD" box="[652,769,1077,1101]" italics="true" pageId="16" pageNumber="517">O. cocleare</emphasis>
</taxonomicName>
(rounds),
<taxonomicName id="1FC8F8ACFF9E6C50FCF7FB95FC55FBBD" authorityName="Shavrin" authorityYear="2025" box="[874,1006,1077,1101]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="tibetanum">
<emphasis id="EABC5F3DFF9E6C50FCF7FB95FC55FBBD" box="[874,1006,1077,1101]" italics="true" pageId="16" pageNumber="517">O. tibetanum</emphasis>
</taxonomicName>
(squares).
</paragraph>
</caption>
<paragraph id="D877832FFF9E6C50FF5AFB3FFC9FFB49" blockId="16.[151,1437,1183,2038]" box="[199,804,1183,1210]" pageId="16" pageNumber="517">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="D877832FFF9E6C50FF5AFB63FEEDFA29" blockId="16.[151,1437,1183,2038]" pageId="16" pageNumber="517">
Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with moderately wide basal portion, slightly narrowed toward wide median lobe; median lobe spoon-shaped, from widest middle part gradually narrowed toward rounded apex; accessory plates narrow and relatively short; parameres moderately narrow, distinctly broadened in apical portions, with widely rounded apices, reaching preapical part of median lobe, each with two long and one short apical setae; internal sac wide and moderately short, with two elongate parallel and two shorter transverse sclerotized structures in basal portion (
<figureCitation id="40F39FAAFF9E6C50FB2EFAD7FA83FA61" box="[1203,1336,1399,1425]" captionStart="FIGURES 4046" captionStartId="15.[151,264,1875,1899]" captionTargetBox="[159,1418,190,1844]" captionTargetId="figure-16@15.[151,1436,181,1852]" captionTargetPageId="15" captionText="FIGURES 4046. Aedeagus of Omalium (Figs 4043—O. cocleare (paratypes), Figs 4446—O. contiger (paratype)): 40, 4344—aedeagus, parameral view, 41, 45—aedeagus, lateral view, 42, 46—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896701" httpUri="https://zenodo.org/record/14896701/files/figure.png" pageId="16" pageNumber="517">Figs 40, 43</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="40F39FAAFF9E6C50FE47FA3BFD96FA45" box="[474,557,1435,1461]" captionStart="FIGURES 4046" captionStartId="15.[151,264,1875,1899]" captionTargetBox="[159,1418,190,1844]" captionTargetId="figure-16@15.[151,1436,181,1852]" captionTargetPageId="15" captionText="FIGURES 4046. Aedeagus of Omalium (Figs 4043—O. cocleare (paratypes), Figs 4446—O. contiger (paratype)): 40, 4344—aedeagus, parameral view, 41, 45—aedeagus, lateral view, 42, 46—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896701" httpUri="https://zenodo.org/record/14896701/files/figure.png" pageId="16" pageNumber="517">Fig. 41</figureCitation>
; apical portion of median lobe (lateral view) wide, with truncate ventrolateral margin (
<figureCitation id="40F39FAAFF9E6C50FF68FA1FFEF1FA29" box="[245,330,1471,1497]" captionStart="FIGURES 4046" captionStartId="15.[151,264,1875,1899]" captionTargetBox="[159,1418,190,1844]" captionTargetId="figure-16@15.[151,1436,181,1852]" captionTargetPageId="15" captionText="FIGURES 4046. Aedeagus of Omalium (Figs 4043—O. cocleare (paratypes), Figs 4446—O. contiger (paratype)): 40, 4344—aedeagus, parameral view, 41, 45—aedeagus, lateral view, 42, 46—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896701" httpUri="https://zenodo.org/record/14896701/files/figure.png" pageId="16" pageNumber="517">Fig. 42</figureCitation>
).
</paragraph>
<paragraph id="D877832FFF9E6C50FF5AFA43FE48F9B5" blockId="16.[151,1437,1183,2038]" pageId="16" pageNumber="517">
Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite with wide basal portion and median part gradually narrowed toward rounded apex (
<figureCitation id="40F39FAAFF9E6C50FAF2F9A7FF0DF9B5" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="16" pageNumber="517">Fig. 16</figureCitation>
). Spermatheca as in
<figureCitation id="40F39FAAFF9E6C50FE00F98BFE4BF9B5" box="[413,496,1579,1605]" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="16" pageNumber="517">Fig. 17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF9E6C50FF5AF9EFFE0CF821" pageId="16" pageNumber="517" type="discussion">
<paragraph id="D877832FFF9E6C50FF5AF9EFFE0CF821" blockId="16.[151,1437,1183,2038]" pageId="16" pageNumber="517">
<emphasis id="EABC5F3DFF9E6C50FF5AF9EFFE0BF999" bold="true" box="[199,432,1615,1641]" pageId="16" pageNumber="517">Comparative notes.</emphasis>
Based on the length of the body, the presence of moderately strong longitudinal elevations on postocular portions of the head, the shape of the pronotum widest in apical third, and the general shape of the aedeagus,
<taxonomicName id="1FC8F8ACFF9E6C50FE90F938FE2BF941" authorityName="Shavrin" authorityYear="2025" box="[269,400,1687,1714]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="cocleare" status="sp. nov.">
<emphasis id="EABC5F3DFF9E6C50FE90F938FE2BF941" box="[269,400,1687,1714]" italics="true" pageId="16" pageNumber="517">O. cocleare</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF9E6C50FE05F937FE48F941" bold="true" box="[408,499,1687,1713]" pageId="16" pageNumber="517">
<taxonomicNameLabel id="F18FE246FF9E6C50FE05F937FE48F941" box="[408,499,1687,1713]" pageId="16" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to the Himalayan
<taxonomicName id="1FC8F8ACFF9E6C50FCA5F938FB2DF942" authority="Bernhauer, 1915" authorityName="Bernhauer" authorityYear="1915" box="[824,1174,1687,1714]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="EABC5F3DFF9E6C50FCA5F938FC6AF941" box="[824,977,1687,1714]" italics="true" pageId="16" pageNumber="517">O. altivagans</emphasis>
Bernhauer, 1915
</taxonomicName>
, known from Kashmir,
<collectingCountry id="A0DFC3BFFF9E6C50FF0AF91BFF6AF925" box="[151,209,1723,1749]" name="India" pageId="16" pageNumber="517">India</collectingCountry>
(
<bibRefCitation id="BC59FEDEFF9E6C50FF7FF91BFE32F925" author="Shavrin, A. V." box="[226,393,1723,1749]" pageId="16" pageNumber="517" pagination="151 - 206" refId="ref30642" refString="Shavrin, A. V. (2023 a) A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region. Zootaxa, 5369 (2), 151-206. https://doi.org/10.11646/zootaxa.5369.2.1" type="journal article" year="2023">Shavrin 2023a</bibRefCitation>
) and
<taxonomicName id="1FC8F8ACFF9E6C50FE50F91CFCADF925" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[461,790,1723,1750]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="latissimum">
<emphasis id="EABC5F3DFF9E6C50FE50F91CFDD1F925" box="[461,618,1723,1750]" italics="true" pageId="16" pageNumber="517">O. latissimum</emphasis>
Shavrin, 2023
</taxonomicName>
, described recently from
<collectingCountry id="A0DFC3BFFF9E6C50FBA6F91BFB36F925" box="[1083,1165,1723,1749]" name="Taiwan" pageId="16" pageNumber="517">Taiwan</collectingCountry>
(
<bibRefCitation id="BC59FEDEFF9E6C50FB03F91BFAF0F925" author="Shavrin, A. V." box="[1182,1355,1723,1749]" pageId="16" pageNumber="517" pagination="173 - 181" refId="ref30696" refString="Shavrin, A. V. (2023 b) Three new species of Omaliopsis Jeannel, 1940 and Omalium Gravenhorst, 1802 from Taiwan (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 5244 (2), 173-181. https://doi.org/10.11646/zootaxa.5244.2.5" type="journal article" year="2023">Shavrin 2023b</bibRefCitation>
). From
<taxonomicName id="1FC8F8ACFF9E6C50FF0AF940FE94F909" authorityName="Bernhauer" authorityYear="1915" box="[151,303,1759,1786]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="EABC5F3DFF9E6C50FF0AF940FE94F909" box="[151,303,1759,1786]" italics="true" pageId="16" pageNumber="517">O. altivagans</emphasis>
</taxonomicName>
it can be distinguished by the significantly denser punctation of the head and the pronotum, sligthly longer temples, slightly more protruded apical angles of the pronotum with the presence of medioapical depression, longer and narrower parameres, distinctly narrower median lobe without crenulation on lateral margins and the presence of elongate sclerotized structures in basal part of the internal sac. From
<taxonomicName id="1FC8F8ACFF9E6C50FBDFF8ECFB5BF895" authorityName="Shavrin" authorityYear="2023" box="[1090,1248,1867,1894]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="latissimum">
<emphasis id="EABC5F3DFF9E6C50FBDFF8ECFB5BF895" box="[1090,1248,1867,1894]" italics="true" pageId="16" pageNumber="517">O. latissimum</emphasis>
</taxonomicName>
it differs by the broader pronotum, longer elytra, narrower median lobe and sligthly longer and broader parameres. From both species it differs by the details of the external and internal morphology of the aedeagus, and different shape of the female accessory sclerite.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF9E6C50FF5AF87BFA21F805" box="[199,1434,2011,2038]" pageId="16" pageNumber="517" type="distribution">
<paragraph id="D877832FFF9E6C50FF5AF87BFA21F805" blockId="16.[151,1437,1183,2038]" box="[199,1434,2011,2038]" pageId="16" pageNumber="517">
<emphasis id="EABC5F3DFF9E6C50FF5AF87BFEE0F805" bold="true" box="[199,347,2011,2037]" pageId="16" pageNumber="517">Distribution.</emphasis>
<taxonomicName id="1FC8F8ACFF9E6C50FEFCF87BFD93F805" authorityName="Shavrin" authorityYear="2025" box="[353,552,2011,2037]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="cocleare" status="sp. nov.">
<emphasis id="EABC5F3DFF9E6C50FEFCF87BFD93F805" box="[353,552,2011,2037]" italics="true" pageId="16" pageNumber="517">Omalium cocleare</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF9E6C50FDB3F87BFD3FF805" bold="true" box="[558,644,2011,2037]" pageId="16" pageNumber="517">
<taxonomicNameLabel id="F18FE246FF9E6C50FDB3F87BFD3FF805" box="[558,644,2011,2037]" pageId="16" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from several localities in
<collectingRegion id="1A0C4DCDFF9E6C50FB9EF87CFBEEF806" box="[1027,1109,2012,2038]" country="China" name="Yunnan" pageId="16" pageNumber="517">Yunnan</collectingRegion>
and
<collectingRegion id="1A0C4DCDFF9E6C50FB17F87BFB58F805" box="[1162,1251,2011,2037]" country="China" name="Sichuan" pageId="16" pageNumber="517">Sichuan</collectingRegion>
,
<collectingCountry id="A0DFC3BFFF9E6C50FB73F87BFA95F805" box="[1262,1326,2011,2037]" name="China" pageId="16" pageNumber="517">China</collectingCountry>
(
<figureCitation id="40F39FAAFF9E6C50FAA0F87BFA35F805" box="[1341,1422,2011,2037]" captionStart="FIGURE 47" captionStartId="16.[151,250,1077,1101]" captionTargetBox="[151,1435,181,1052]" captionTargetId="figure-20@16.[151,1435,181,1052]" captionTargetPageId="16" captionText="FIGURE 47. Distribution of Omalium in China: O. cocleare (rounds), O. tibetanum (squares)." figureDoi="http://doi.org/10.5281/zenodo.14896705" httpUri="https://zenodo.org/record/14896705/files/figure.png" pageId="16" pageNumber="517">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF9F6C51FF5AFF37FF74FF25" pageId="17" pageNumber="518" type="etymology">
<paragraph id="D877832FFF9F6C51FF5AFF37FF74FF25" blockId="17.[151,1437,151,393]" pageId="17" pageNumber="518">
<emphasis id="EABC5F3DFF9F6C51FF5AFF37FEF5FF41" bold="true" box="[199,334,151,177]" pageId="17" pageNumber="518">Etymology.</emphasis>
The specific epithet is the Latin noun
<emphasis id="EABC5F3DFF9F6C51FD60FF37FCC0FF41" box="[765,891,151,177]" italics="true" pageId="17" pageNumber="518">coclearium</emphasis>
, -
<emphasis id="EABC5F3DFF9F6C51FC0EFF37FC20FF40" box="[915,923,151,176]" italics="true" pageId="17" pageNumber="518">i</emphasis>
(spoon). It alludes to the shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF9F6C51FF5AFF7EFB79FE79" pageId="17" pageNumber="518" type="biology_ecology">
<paragraph id="D877832FFF9F6C51FF5AFF7EFB79FE79" blockId="17.[151,1437,151,393]" pageId="17" pageNumber="518">
<emphasis id="EABC5F3DFF9F6C51FF5AFF7EFEF0FF08" bold="true" box="[199,331,222,248]" pageId="17" pageNumber="518">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="1F302ECAFF9F6C51FCD0FF7FFC45FF09" box="[845,1022,222,249]" metricMagnitude="3" metricUnit="m" metricValue="2.92" metricValueMax="3.54" metricValueMin="2.3" pageId="17" pageNumber="518" unit="m" value="2920.0" valueMax="3540.0" valueMin="2300.0">2300 to 3540 m</quantity>
a.s.l. and were sifted from leaf litter, mosses and mushrooms. Some specimens were collected by sifting wet moss and various debris around small seepages in an old, devastated mixed forest (
<taxonomicName id="1FC8F8ACFF9F6C51FD3DFE87FD65FEB1" box="[672,734,295,321]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="518" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF9F6C51FD3DFE87FD65FEB1" box="[672,734,295,321]" italics="true" pageId="17" pageNumber="518">Abies</emphasis>
</taxonomicName>
,
<taxonomicName id="1FC8F8ACFF9F6C51FD72FE87FC3DFEB1" authority=", Betula" authorityName="Betula" box="[751,902,295,321]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="518" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF9F6C51FD72FE87FC97FEB0" box="[751,812,295,320]" italics="true" pageId="17" pageNumber="518">Larix</emphasis>
,
<emphasis id="EABC5F3DFF9F6C51FCA0FE87FC3DFEB1" box="[829,902,295,321]" italics="true" pageId="17" pageNumber="518">Betula</emphasis>
</taxonomicName>
,
<taxonomicName id="1FC8F8ACFF9F6C51FC0AFE87FB87FEB1" box="[919,1084,295,321]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="17" pageNumber="518" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF9F6C51FC0AFE87FB87FEB1" box="[919,1084,295,321]" italics="true" pageId="17" pageNumber="518">Rhododendron</emphasis>
</taxonomicName>
) (C147) and in remnant of an original
<taxonomicName id="1FC8F8ACFF9F6C51FF6BFEEBFE36FE95" authority=", Betula" authorityName="Betula" box="[246,397,331,357]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="518" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF9F6C51FF6BFEEBFE8FFE95" box="[246,308,331,357]" italics="true" pageId="17" pageNumber="518">Abies</emphasis>
,
<emphasis id="EABC5F3DFF9F6C51FED9FEEBFE36FE95" box="[324,397,331,357]" italics="true" pageId="17" pageNumber="518">Betula</emphasis>
</taxonomicName>
,
<taxonomicName id="1FC8F8ACFF9F6C51FE01FEEBFDFAFE95" box="[412,577,331,357]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="17" pageNumber="518" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF9F6C51FE01FEEBFDFAFE95" box="[412,577,331,357]" italics="true" pageId="17" pageNumber="518">Rhododendron</emphasis>
</taxonomicName>
forest, treading of quite wet, mostly grassy vegetation in a seepage (C150). The photograph of the locality of
<specimenCount id="CECE48A6FF9F6C51FD8CFECFFD1AFE79" box="[529,673,367,393]" count="1" pageId="17" pageNumber="518" type="generic" typeStatus="paratype">one paratype</specimenCount>
in Diancang Shan Mts. (C03-19B) as in
<figureCitation id="40F39FAAFF9F6C51FBF6FECFFB79FE79" box="[1131,1218,367,393]" captionStart-0="FIGURE 49" captionStart-1="FIGURES 13" captionStartId-0="19.[151,250,1908,1932]" captionStartId-1="3.[151,264,1097,1121]" captionTargetBox-0="[151,1436,921,1885]" captionTargetBox-1="[151,1436,181,1073]" captionTargetId-0="figure-293@19.[151,1436,921,1885]" captionTargetId-1="figure-16@3.[151,1436,181,1073]" captionTargetPageId-0="19" captionTargetPageId-1="3" captionText-0="FIGURE 49. Type locality of Omalium cocleare in the Diancang Shan range in Yunnan, China (photograph: M. Schülke)." captionText-1="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14896711" figureDoi-1="http://doi.org/10.5281/zenodo.14896686" httpUri-0="https://zenodo.org/record/14896711/files/figure.png" httpUri-1="https://zenodo.org/record/14896686/files/figure.png" pageId="17" pageNumber="518">Fig. 49.</figureCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,268 @@
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<mods:title id="9FC6103B09959977A20BBB87C0AEF557">Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China</mods:title>
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<paragraph id="D877832FFF846C4AFF0AFF7FFD9DFF09" blockId="10.[151,550,223,285]" box="[151,550,223,250]" pageId="10" pageNumber="511">
<heading id="833F3443FF846C4AFF0AFF7FFD9DFF09" bold="true" box="[151,550,223,250]" fontSize="11" level="1" pageId="10" pageNumber="511" reason="1">
<emphasis id="EABC5F3DFF846C4AFF0AFF7FFD9DFF09" bold="true" box="[151,550,223,250]" pageId="10" pageNumber="511">
3.3.2.
<taxonomicName id="1FC8F8ACFF846C4AFF7CFF7FFE7AFF09" authorityName="Shavrin" authorityYear="2025" box="[225,449,223,250]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="pratense" status="sp. nov.">
<emphasis id="EABC5F3DFF846C4AFF7CFF7FFE7AFF09" bold="true" box="[225,449,223,250]" italics="true" pageId="10" pageNumber="511">Omalium pratense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F18FE246FF846C4AFE55FF7FFD9DFF09" box="[456,550,223,249]" pageId="10" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF846C4AFF0AFEA3FEC0FEED" box="[151,379,259,285]" pageId="10" pageNumber="511" type="description">
<paragraph id="D877832FFF846C4AFF0AFEA3FEC0FEED" blockId="10.[151,550,223,285]" box="[151,379,259,285]" pageId="10" pageNumber="511">
(
<figureCitation id="40F39FAAFF846C4AFF02FEA3FF4DFEED" box="[159,246,259,285]" captionStart="FIGURES 3133" captionStartId="10.[151,264,1388,1412]" captionTargetBox="[162,1423,536,1356]" captionTargetId="figure-96@10.[151,1435,525,1364]" captionTargetPageId="10" captionText="FIGURES 3133. Habitus of Omalium: 31—O. pratense (holotype), 32—O. cocleare (holotype), 33—O. contiger (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896692" httpUri="https://zenodo.org/record/14896692/files/figure.png" pageId="10" pageNumber="511">Figs 31</figureCitation>
,
<figureCitation id="40F39FAAFF846C4AFE9FFEA3FEF0FEED" box="[258,331,259,285]" captionStart="FIGURES 3438" captionStartId="11.[151,264,1758,1782]" captionTargetBox="[160,1427,190,1725]" captionTargetId="figure-16@11.[151,1436,181,1734]" captionTargetPageId="11" captionText="FIGURES 3438. Aedeagus of Omalium (Figs 3436—O. pratense (holotype), Figs 3738—O. alticola (Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="10" pageNumber="511">3436</figureCitation>
,
<figureCitation id="40F39FAAFF846C4AFECBFEA3FEC8FEED" box="[342,371,259,285]" captionStart="FIGURE 39" captionStartId="12.[151,250,1079,1103]" captionTargetBox="[151,1436,181,1055]" captionTargetId="figure-20@12.[151,1436,181,1055]" captionTargetPageId="12" captionText="FIGURE 39. Distribution of Omalium in China: O. kurbatovi (rounds), O. pratense (square), O. confertum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896697" httpUri="https://zenodo.org/record/14896697/files/figure.png" pageId="10" pageNumber="511">39</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF846C4AFF0AFEEBFF53FE05" pageId="10" pageNumber="511" type="materials_examined">
<paragraph id="D877832FFF846C4AFF0AFEEBFF53FE05" blockId="10.[151,1437,331,501]" pageId="10" pageNumber="511">
<emphasis id="EABC5F3DFF846C4AFF0AFEEBFE06FE95" bold="true" box="[151,445,331,357]" pageId="10" pageNumber="511">Type material examined.</emphasis>
<materialsCitation id="68A08972FF846C4AFE5FFEEBFF53FE05" pageId="10" pageNumber="511">
<typeStatus id="07733D8DFF846C4AFE5FFEEBFD91FE95" box="[450,554,331,357]" pageId="10" pageNumber="511" type="holotype">Holotype</typeStatus>
<specimenCount id="CECE48A6FF846C4AFDADFEECFDFEFE96" box="[560,581,332,358]" count="1" pageId="10" pageNumber="511" type="male"></specimenCount>
(
<figureCitation id="40F39FAAFF846C4AFDCEFEEBFD1EFE95" box="[595,677,331,357]" captionStart="FIGURES 3133" captionStartId="10.[151,264,1388,1412]" captionTargetBox="[162,1423,536,1356]" captionTargetId="figure-96@10.[151,1435,525,1364]" captionTargetPageId="10" captionText="FIGURES 3133. Habitus of Omalium: 31—O. pratense (holotype), 32—O. cocleare (holotype), 33—O. contiger (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896692" httpUri="https://zenodo.org/record/14896692/files/figure.png" pageId="10" pageNumber="511">Fig. 31</figureCitation>
; dissected; left apical part of paramere missing):
<emphasis id="EABC5F3DFF846C4AFB5FFEEBFA27FE95" bold="true" box="[1218,1436,331,357]" pageId="10" pageNumber="511">
<collectingCountry id="A0DFC3BFFF846C4AFB5FFEEBFA9FFE95" box="[1218,1316,331,357]" name="China" pageId="10" pageNumber="511">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF846C4AFAB3FEEBFA28FE95" box="[1326,1427,331,357]" country="China" name="Shaanxi" pageId="10" pageNumber="511">SHAANXI</collectingRegion>
:
</emphasis>
CHINA (S-Shaanxi) Qinling Shan | pass on r[oa]d Zhouzhi-Foping |
<quantity id="1F302ECAFF846C4AFC48FECFFB94FE79" box="[981,1071,367,394]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="10" pageNumber="511" unit="km" value="105.0">105 km</quantity>
SW Xi`an, N slope |
<quantity id="1F302ECAFF846C4AFAA6FECFFA2CFE7A" box="[1339,1431,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="10" pageNumber="511" unit="m" value="1990.0">1990 m</quantity>
,
<geoCoordinate id="BDFCE5E8FF846C4AFF0AFE33FF43FE5E" box="[151,248,403,430]" degrees="33" direction="north" minutes="44" orientation="latitude" pageId="10" pageNumber="511" precision="925" value="33.733334">33°44´N</geoCoordinate>
/
<geoCoordinate id="BDFCE5E8FF846C4AFE9CFE33FED0FE5E" box="[257,363,403,430]" degrees="107" direction="east" minutes="59" orientation="longitude" pageId="10" pageNumber="511" precision="925" value="107.98333">107°59´E</geoCoordinate>
| (small creek vall[ey]./mix[ed].decid[uous]. | for[est]./bamboo/small meadows) | 2./
<date id="AC76A5EFFF846C4AFABDFE33FA27FE5D" box="[1312,1436,403,429]" pageId="10" pageNumber="511" value="2001-07-04">4.VII.2001</date>
Wrase [01] &lt;printed&gt;, HOLOTYPE |
<taxonomicName id="1FC8F8ACFF846C4AFDFAFE17FD74FE21" authorityName="Gravenhorst" authorityYear="1802" box="[615,719,439,465]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF846C4AFDFAFE17FD74FE21" box="[615,719,439,465]" italics="true" pageId="10" pageNumber="511">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF846C4AFD71FE18FCF7FE21" authorityName="Shavrin" authorityYear="2025" box="[748,844,440,465]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="pratense" status="sp. nov.">
<emphasis id="EABC5F3DFF846C4AFD71FE18FCF7FE21" box="[748,844,440,465]" italics="true" pageId="10" pageNumber="511">pratense</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF846C4AFCC5FE17FC0CFE21" bold="true" box="[856,951,439,465]" pageId="10" pageNumber="511">
<taxonomicNameLabel id="F18FE246FF846C4AFCC5FE17FC0CFE21" box="[856,951,439,465]" pageId="10" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025 &lt;red, printed&gt; (
<collectionCode id="BED91BEAFF846C4AFF02FE7BFF60FE05" box="[159,219,475,501]" pageId="10" pageNumber="511">cSch</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="8CB7D3A7FF846C4AFF0AFACCFEECFA58" ID-DOI="http://doi.org/10.5281/zenodo.14896692" ID-Zenodo-Dep="14896692" httpUri="https://zenodo.org/record/14896692/files/figure.png" pageId="10" pageNumber="511" startId="10.[151,264,1388,1412]" targetBox="[162,1423,536,1356]" targetPageId="10" targetType="figure">
<paragraph id="D877832FFF846C4AFF0AFACCFEECFA58" blockId="10.[151,1437,1388,1448]" pageId="10" pageNumber="511">
<emphasis id="EABC5F3DFF846C4AFF0AFACCFEEFFA74" bold="true" box="[151,340,1388,1412]" pageId="10" pageNumber="511">FIGURES 3133.</emphasis>
Habitus of
<taxonomicName id="1FC8F8ACFF846C4AFE56FACCFD92FA74" authorityName="Gravenhorst" authorityYear="1802" box="[459,553,1388,1412]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF846C4AFE56FACCFD92FA74" box="[459,553,1388,1412]" italics="true" pageId="10" pageNumber="511">Omalium</emphasis>
</taxonomicName>
: 31—
<taxonomicName id="1FC8F8ACFF846C4AFDF4FACDFD64FA74" authorityName="Shavrin" authorityYear="2025" box="[617,735,1389,1412]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="pratense">
<emphasis id="EABC5F3DFF846C4AFDF4FACDFD64FA74" box="[617,735,1389,1412]" italics="true" pageId="10" pageNumber="511">O. pratense</emphasis>
</taxonomicName>
(holotype), 32—
<taxonomicName id="1FC8F8ACFF846C4AFC13FACDFBB8FA74" authorityName="Shavrin" authorityYear="2025" box="[910,1027,1388,1412]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="cocleare">
<emphasis id="EABC5F3DFF846C4AFC13FACDFBB8FA74" box="[910,1027,1388,1412]" italics="true" pageId="10" pageNumber="511">O. cocleare</emphasis>
</taxonomicName>
(holotype), 33—
<taxonomicName id="1FC8F8ACFF846C4AFB2EFACDFA9DFA74" authorityName="Shavrin" authorityYear="2025" box="[1203,1318,1389,1412]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="contiger">
<emphasis id="EABC5F3DFF846C4AFB2EFACDFA9DFA74" box="[1203,1318,1389,1412]" italics="true" pageId="10" pageNumber="511">O. contiger</emphasis>
</taxonomicName>
(holotype). Scale bar: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="90D2D0A4FF846C4DFF5AFA5AFE2AFE95" lastPageId="13" lastPageNumber="514" pageId="10" pageNumber="511" type="description">
<paragraph id="D877832FFF846C4AFF5AFA5AFE74F9AD" blockId="10.[151,1437,1530,1953]" pageId="10" pageNumber="511">
<emphasis id="EABC5F3DFF846C4AFF5AFA5AFEE2F9E4" bold="true" box="[199,345,1530,1556]" pageId="10" pageNumber="511">Description.</emphasis>
Measurements: HW: 0.50; HL: 0.35; OL: 0.17; TL: 0.03; AL: 0.85; PL: 0.41; PWmax: 0.62; PWmin: 0.42; ESL: 0.80; EW: 0.90; MTbL (
<typeStatus id="07733D8DFF846C4AFD05F9BEFCBAF9C8" box="[664,769,1566,1592]" pageId="10" pageNumber="511" type="holotype">holotype</typeStatus>
): 0.40; MTrL: 0.19 (MTrL 14: 0.07; MTrL 5: 0.12); AW: 0.82; AedL: 0.60; BL: 2.80.
</paragraph>
<paragraph id="D877832FFF846C4AFF5AF9C6FBBEF850" blockId="10.[151,1437,1530,1953]" pageId="10" pageNumber="511">
Habitus as in
<figureCitation id="40F39FAAFF846C4AFEF8F9C6FE03F971" box="[357,440,1638,1665]" captionStart="FIGURES 3133" captionStartId="10.[151,264,1388,1412]" captionTargetBox="[162,1423,536,1356]" captionTargetId="figure-96@10.[151,1435,525,1364]" captionTargetPageId="10" captionText="FIGURES 3133. Habitus of Omalium: 31—O. pratense (holotype), 32—O. cocleare (holotype), 33—O. contiger (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896692" httpUri="https://zenodo.org/record/14896692/files/figure.png" pageId="10" pageNumber="511">Fig. 31</figureCitation>
. Head, pronotum and abdomen dark-brown (laterobasal and basal margins of pronotum slightly and apical part of abdomen and paratergites distinctly paler); antennomeres 611 and elytra brown; mouthparts, antennomeres 15 and legs yellowish. Head with irregular and moderately large punctation, slightly denser in middle and on infraorbital portions; neck with sparse and fine punctation; punctation of pronotum irregular, larger and deeper than that in middle portion of head, finer and sparser in lateral and mediobasal portions; punctation of elytra about as that on pronotum, denser around scutellum, finer and sparser in middle; abdomen without visible punctation. Anterior portion of clypeus with fine transverse microreticulation, laterobasal portions of clypeus with fine diagonal meshes, portions between apical margins of eyes and clypeus with distinct elongate microsculpture; scutellum without meshes; abdominal tergites with dense transverse sculpture.
</paragraph>
<caption id="8CB7D3A7FF856C4BFF0AF97EFB51F8EA" ID-DOI="http://doi.org/10.5281/zenodo.14896694" ID-Zenodo-Dep="14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="11" pageNumber="512" startId="11.[151,264,1758,1782]" targetBox="[160,1427,190,1725]" targetPageId="11" targetType="figure">
<paragraph id="D877832FFF856C4BFF0AF97EFB51F8EA" blockId="11.[151,1437,1758,1819]" pageId="11" pageNumber="512">
<emphasis id="EABC5F3DFF856C4BFF0AF97EFEEEF906" bold="true" box="[151,341,1758,1782]" pageId="11" pageNumber="512">FIGURES 3438.</emphasis>
Aedeagus of
<taxonomicName id="1FC8F8ACFF856C4BFE7CF97EFD84F906" authorityName="Gravenhorst" authorityYear="1802" box="[481,575,1758,1782]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF856C4BFE7CF97EFD84F906" box="[481,575,1758,1782]" italics="true" pageId="11" pageNumber="512">Omalium</emphasis>
</taxonomicName>
(Figs 3436—
<taxonomicName id="1FC8F8ACFF856C4BFD44F97FFCF4F906" authorityName="Shavrin" authorityYear="2025" box="[729,847,1759,1782]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="pratense">
<emphasis id="EABC5F3DFF856C4BFD44F97FFCF4F906" box="[729,847,1759,1782]" italics="true" pageId="11" pageNumber="512">O. pratense</emphasis>
</taxonomicName>
(holotype), Figs 3738—
<taxonomicName id="1FC8F8ACFF856C4BFBC5F97FFB7FF906" authorityName="Cameron" authorityYear="1941" box="[1112,1220,1758,1782]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="EABC5F3DFF856C4BFBC5F97FFB7FF906" box="[1112,1220,1758,1782]" italics="true" pageId="11" pageNumber="512">O. alticola</emphasis>
</taxonomicName>
(Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="8CB7D3A7FF826C4CFF0AFB97FAFAFBBF" ID-DOI="http://doi.org/10.5281/zenodo.14896697" ID-Zenodo-Dep="14896697" box="[151,1345,1079,1103]" httpUri="https://zenodo.org/record/14896697/files/figure.png" pageId="12" pageNumber="513" startId="12.[151,250,1079,1103]" targetBox="[151,1436,181,1055]" targetPageId="12" targetType="figure">
<paragraph id="D877832FFF826C4CFF0AFB97FAFAFBBF" blockId="12.[151,1345,1079,1103]" box="[151,1345,1079,1103]" pageId="12" pageNumber="513">
<emphasis id="EABC5F3DFF826C4CFF0AFB97FE9BFBBF" bold="true" box="[151,288,1079,1103]" pageId="12" pageNumber="513">FIGURE 39.</emphasis>
Distribution of
<taxonomicName id="1FC8F8ACFF826C4CFE5FFB97FD9BFBBF" authorityName="Gravenhorst" authorityYear="1802" box="[450,544,1079,1103]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF826C4CFE5FFB97FD9BFBBF" box="[450,544,1079,1103]" italics="true" pageId="12" pageNumber="513">Omalium</emphasis>
</taxonomicName>
in China:
<taxonomicName id="1FC8F8ACFF826C4CFD16FB98FCB0FBBF" authorityName="Shavrin" authorityYear="2025" box="[651,779,1079,1103]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="kurbatovi">
<emphasis id="EABC5F3DFF826C4CFD16FB98FCB0FBBF" box="[651,779,1079,1103]" italics="true" pageId="12" pageNumber="513">O. kurbatovi</emphasis>
</taxonomicName>
(rounds),
<taxonomicName id="1FC8F8ACFF826C4CFCE9FB98FC51FBBF" authorityName="Shavrin" authorityYear="2025" box="[884,1002,1080,1103]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="pratense">
<emphasis id="EABC5F3DFF826C4CFCE9FB98FC51FBBF" box="[884,1002,1080,1103]" italics="true" pageId="12" pageNumber="513">O. pratense</emphasis>
</taxonomicName>
(square),
<taxonomicName id="1FC8F8ACFF826C4CFBCDFB98FB6DFBBF" authorityName="Shavrin" authorityYear="2025" box="[1104,1238,1079,1103]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="confertum">
<emphasis id="EABC5F3DFF826C4CFBCDFB98FB6DFBBF" box="[1104,1238,1079,1103]" italics="true" pageId="12" pageNumber="513">O. confertum</emphasis>
</taxonomicName>
(triangle).
</paragraph>
</caption>
<paragraph id="D877832FFF826C4CFF5AFB01FCF1F9B7" blockId="12.[151,1437,1185,2039]" pageId="12" pageNumber="513">Head 1.4 times as broad as long, with broad distinctly elevated clypeus and moderately wide and deep anteriomedian depressions, reaching level of apical margins of eyes; posteriolateral margins of clypeus relatively strongly narrowed posteriad and reaching level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with irregular transverse elevations between punctures in middle; postocular portions with two indistinct, narrow longitudinal wrinkles. Anteocellar foveae narrow, linear, reaching level of middle length of eyes. Temples very short, from posterior margins of eyes strongly narrowed toward neck. Apical part of neck without depression. Ocelli located slightly below level of posterior margins of eyes; distance between ocelli 1.4 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three and a half times as long as penultimate segment. Antenna with elongate antennomeres 47 and transverse 810; 3 distinctly narrower than 2, 4 distinctly shorter than 3, 5 broader than 4, 6 slightly longer and broader than 5, 7 indistinctly shorter and broader than 6, 8 broader than 7, 910 broader than 8, apical antennomere about twice as long as 10.</paragraph>
<paragraph id="D877832FFF826C4CFF5AF9F2FCEDF8EF" blockId="12.[151,1437,1185,2039]" pageId="12" pageNumber="513">Pronotum transverse, 1.5 times as broad as long, 1.2 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad. Anterior margin rounded, not protruded anteriad, indistinctly concave in middle. Laterobasal portions not concave in front of subacute hind angles. Laterobasal portions widely impressed and relatively deeply impressed. Surface of disc with two indistinct longitudinal depressions, broadened in mediobasal portion; medioapical portion without depression. Middle portion without longitudinal elevations between punctures; each latero-apical portion with distinct longitudinal elevations.</paragraph>
<paragraph id="D877832FFF826C4CFF5AF889FF40F87B" blockId="12.[151,1437,1185,2039]" pageId="12" pageNumber="513">Elytra 1.1 times as broad as long, 1.9 times as long as pronotum; hind margins straight. Dorsal surface of each elytron with irregular and moderately fine longitudinal elevations, more distinct in mediolateral and apical portions.</paragraph>
<paragraph id="D877832FFF826C4CFF5AF835FD39F85F" blockId="12.[151,1437,1185,2039]" box="[199,642,1941,1968]" pageId="12" pageNumber="513">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="D877832FFF826C4DFF5AF819FB95FEB1" blockId="12.[151,1437,1185,2039]" lastBlockId="13.[151,1437,151,717]" lastPageId="13" lastPageNumber="514" pageId="12" pageNumber="513">
Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII slightly sinuate.Aedeagus with wide and moderately short basal portion, slightly narrowed toward median lobe; median lobe elongate, from slightly broadened basal part indistinctly narrowed apically toward widely rounded apex; accessory plates relatively short and narrow; parameres moderately narrow, slightly narrowed apically, reaching preapical part of median lobe, each with three moderately short apical setae; internal sac wide and relatively short, with two small oval structures in basal part (
<figureCitation id="40F39FAAFF836C4DFE7FFEA3FD83FEED" box="[482,568,259,285]" captionStart="FIGURES 3438" captionStartId="11.[151,264,1758,1782]" captionTargetBox="[160,1427,190,1725]" captionTargetId="figure-16@11.[151,1436,181,1734]" captionTargetPageId="11" captionText="FIGURES 3438. Aedeagus of Omalium (Figs 3436—O. pratense (holotype), Figs 3738—O. alticola (Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="13" pageNumber="514">Fig. 34</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="40F39FAAFF836C4DFC76FEA3FB84FEED" box="[1003,1087,258,285]" captionStart="FIGURES 3438" captionStartId="11.[151,264,1758,1782]" captionTargetBox="[160,1427,190,1725]" captionTargetId="figure-16@11.[151,1436,181,1734]" captionTargetPageId="11" captionText="FIGURES 3438. Aedeagus of Omalium (Figs 3436—O. pratense (holotype), Figs 3738—O. alticola (Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="13" pageNumber="514">Fig. 35</figureCitation>
; apical portion of median lobe (lateral view) relatively wide, gradually narrowed toward subacute apex (
<figureCitation id="40F39FAAFF836C4DFC50FE87FB9AFEB1" box="[973,1057,295,321]" captionStart="FIGURES 3438" captionStartId="11.[151,264,1758,1782]" captionTargetBox="[160,1427,190,1725]" captionTargetId="figure-16@11.[151,1436,181,1734]" captionTargetPageId="11" captionText="FIGURES 3438. Aedeagus of Omalium (Figs 3436—O. pratense (holotype), Figs 3738—O. alticola (Sichuan)): 34, 37— aedeagus, parameral view, 35, 38—aedeagus, lateral view, 36—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896694" httpUri="https://zenodo.org/record/14896694/files/figure.png" pageId="13" pageNumber="514">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="D877832FFF836C4DFF5AFEEBFE2AFE95" blockId="13.[151,1437,151,717]" box="[199,401,331,357]" pageId="13" pageNumber="514">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF836C4DFF5AFECEFC8AFE05" pageId="13" pageNumber="514" type="discussion">
<paragraph id="D877832FFF836C4DFF5AFECEFC8AFE05" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="514">
<emphasis id="EABC5F3DFF836C4DFF5AFECEFE08FE79" bold="true" box="[199,435,366,393]" pageId="13" pageNumber="514">Comparative notes.</emphasis>
Based on the general shapes of the body and the the aedeagus,
<taxonomicName id="1FC8F8ACFF836C4DFB06FECFFA8DFE78" authorityName="Shavrin" authorityYear="2025" box="[1179,1334,367,393]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pratensum" status="sp. nov.">
<emphasis id="EABC5F3DFF836C4DFB06FECFFA8DFE78" box="[1179,1334,367,393]" italics="true" pageId="13" pageNumber="514">O. pratensum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF836C4DFADDFECFFA27FE79" bold="true" box="[1344,1436,367,393]" pageId="13" pageNumber="514">
<taxonomicNameLabel id="F18FE246FF836C4DFADDFECFFA27FE79" box="[1344,1436,367,393]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to the Palaearctic O.
<emphasis id="EABC5F3DFF836C4DFD99FE33FDE5FE5D" box="[516,606,403,429]" italics="true" pageId="13" pageNumber="514">littorale</emphasis>
Kraatz, 1857, widely distributed from Europe to Central Asia (
<bibRefCitation id="BC59FEDEFF836C4DFADEFE33FECEFE21" author="Shavrin, A. V. &amp; Khachikov, E. A." pageId="13" pageNumber="514" pagination="443 - 458" refId="ref30754" refString="Shavrin, A. V. &amp; Khachikov, E. A. (2023) Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 5227 (4), 443-458. https://doi.org/10.11646/zootaxa.5227.4.3" type="journal article" year="2023">Shavrin &amp; Khachikov 2023</bibRefCitation>
;
<bibRefCitation id="BC59FEDEFF836C4DFE1EFE17FDADFE21" author="Zanetti, A." box="[387,534,439,465]" pageId="13" pageNumber="514" refId="ref30978" refString="Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, xii + 472 pp." type="book" year="1987">Zanetti 1987</bibRefCitation>
,
<bibRefCitation id="BC59FEDEFF836C4DFDB9FE17FDE4FE21" author="Zanetti, A." box="[548,607,439,465]" pageId="13" pageNumber="514" pagination="45 - 63" refId="ref31009" refString="Zanetti, A. (2002) Studies on Omalium Gravenhorst, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). Bollettino del Museo Civivo di Storia Naturale di Verona. Botanica Zoologia, 26, 45-63." type="journal article" year="2002">2002</bibRefCitation>
, 2006, 2011). It can be distinguished from it by the significantly shorter temples and slightly broader apical part of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF836C4DFF5AFE5EFE09FDCD" pageId="13" pageNumber="514" type="distribution">
<paragraph id="D877832FFF836C4DFF5AFE5EFE09FDCD" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="514">
<emphasis id="EABC5F3DFF836C4DFF5AFE5EFEDAFDE8" bold="true" box="[199,353,510,536]" pageId="13" pageNumber="514">Distribution.</emphasis>
<taxonomicName id="1FC8F8ACFF836C4DFEFBFE5FFD8EFDE8" authorityName="Shavrin" authorityYear="2025" box="[358,565,511,537]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pratense" status="sp. nov.">
<emphasis id="EABC5F3DFF836C4DFEFBFE5FFD8EFDE8" box="[358,565,511,537]" italics="true" pageId="13" pageNumber="514">Omalium pratense</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF836C4DFDA6FE5FFD2FFDE9" bold="true" box="[571,660,511,537]" pageId="13" pageNumber="514">
<taxonomicNameLabel id="F18FE246FF836C4DFDA6FE5FFD2FFDE9" box="[571,660,511,537]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="07733D8DFF836C4DFC3CFE5FFC6AFDE9" box="[929,977,511,537]" pageId="13" pageNumber="514">type</typeStatus>
locality in Qinling Shan mountain range,
<collectingRegion id="1A0C4DCDFF836C4DFF0AFD83FF4FFDCD" box="[151,244,547,573]" country="China" name="Shaanxi" pageId="13" pageNumber="514">Shaanxi</collectingRegion>
,
<collectingCountry id="A0DFC3BFFF836C4DFF62FD83FEF9FDCD" box="[255,322,547,573]" name="China" pageId="13" pageNumber="514">China</collectingCountry>
(
<figureCitation id="40F39FAAFF836C4DFECFFD83FE1DFDCD" box="[338,422,547,573]" captionStart="FIGURE 39" captionStartId="12.[151,250,1079,1103]" captionTargetBox="[151,1436,181,1055]" captionTargetId="figure-20@12.[151,1436,181,1055]" captionTargetPageId="12" captionText="FIGURE 39. Distribution of Omalium in China: O. kurbatovi (rounds), O. pratense (square), O. confertum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896697" httpUri="https://zenodo.org/record/14896697/files/figure.png" pageId="13" pageNumber="514">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF836C4DFF5AFDE7FD2BFD75" pageId="13" pageNumber="514" type="etymology">
<paragraph id="D877832FFF836C4DFF5AFDE7FD2BFD75" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="514">
<emphasis id="EABC5F3DFF836C4DFF5AFDE7FEF5FD91" bold="true" box="[199,334,583,609]" pageId="13" pageNumber="514">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="EABC5F3DFF836C4DFCABFDE7FC26FD90" box="[822,925,583,608]" italics="true" pageId="13" pageNumber="514">pratensis</emphasis>
, -
<emphasis id="EABC5F3DFF836C4DFC2BFDE7FC72FD90" box="[950,969,583,608]" italics="true" pageId="13" pageNumber="514">is</emphasis>
, -
<emphasis id="EABC5F3DFF836C4DFC7CFDE7FC56FD90" box="[993,1005,583,608]" italics="true" pageId="13" pageNumber="514">e</emphasis>
(living in meadows). It alludes to the
<typeStatus id="07733D8DFF836C4DFF0AFDCBFF42FD75" box="[151,249,619,645]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
having been collected in a meadow.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF836C4DFF5AFD2EFE66FD3D" pageId="13" pageNumber="514" type="biology_ecology">
<paragraph id="D877832FFF836C4DFF5AFD2EFE66FD3D" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="514">
<emphasis id="EABC5F3DFF836C4DFF5AFD2EFEF0FD58" bold="true" box="[199,331,654,680]" pageId="13" pageNumber="514">Bionomics.</emphasis>
The
<typeStatus id="07733D8DFF836C4DFE15FD2FFE51FD59" box="[392,490,655,681]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
was collected at elevation
<quantity id="1F302ECAFF836C4DFCB9FD2FFCC0FD59" box="[804,891,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="13" pageNumber="514" unit="m" value="1990.0">1990 m</quantity>
a.s.l. by sifting in mixed deciduous forest with bamboo and small meadows.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="85A50367D376B26F77F405C20DD2532C">Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China</mods:title>
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<mods:name id="9F09FC3EAE4AEA7D7B5F0A63CE16908C" type="personal">
<mods:role id="5394AEB737BE204ADFF04989007C1C8C">
<mods:roleTerm id="D8F8F4AC84FF4A8E3DBBDA4CF70C62DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0068706E6DBEFBFE18E9236C34246B0">Shavrin, Alexey V.</mods:namePart>
<mods:nameIdentifier id="90A7886543810F1FB4A44DF436A209AD" type="ORCID">0000-0001-6487-4530</mods:nameIdentifier>
<mods:affiliation id="44CF5D69CB7B9F0497F0E33E1B82FA89">Institute of Life Sciences and Technologies, Daugavpils University, Vienibas 13, Daugavpils, LV- 5401, Latvia</mods:affiliation>
<mods:nameIdentifier id="E971FB697E1438ED22C35108E3D01CC6" type="email">ashavrin@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E7FA50C07EA6306023DCA12E2CF3A3D6">text</mods:typeOfResource>
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<mods:titleInfo id="B388924E74B11AD7A1D72553B003D333">
<mods:title id="15B904D90CF0E9BF1D395188F098D59E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F29AD4C68ECA05E228FFBDFEC20D177F">
<mods:date id="DB50C0BF9E26E77E5B9AA04A2738591C">2025</mods:date>
<mods:detail id="1919C6B15E10192467938C6D6C632775" type="pubDate">
<mods:number id="56BCC3F94EBDD99EA394493C1ECE3DE1">2025-02-18</mods:number>
</mods:detail>
<mods:detail id="FACB393AA543E925B229F15B3F2AF494" type="volume">
<mods:number id="F24E89375DC42D29197C8C45015AA860">5588</mods:number>
</mods:detail>
<mods:detail id="36DED461E6094A149AA779309D0B8049" type="issue">
<mods:number id="0E3E64EB43BB0EC0BF5BBBC1CC1935B6">4</mods:number>
</mods:detail>
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<mods:start id="684A1F702D912B3FFAD1B808E6D4F716">501</mods:start>
<mods:end id="9062B5945E08104DE6F4CC4565417F8F">543</mods:end>
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<mods:location id="DC40A7157BBD24EA8F299D1D6C7F6602">
<mods:url id="AF7BE2F5B085CD25BF5367770A4917B9">https://doi.org/10.11646/zootaxa.5588.4.1</mods:url>
</mods:location>
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<mods:identifier id="4288DDC0EEB0C9A1D283AB566CFFC1D9" type="DOI">10.11646/zootaxa.5588.4.1</mods:identifier>
<mods:identifier id="7170B04387A13A6395BFF5035979CA7C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B8CB69A7BEE259F18895AF0E16F8B1A4" type="Zenodo-Dep">14896684</mods:identifier>
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<treatment id="50613239FF866C49FF0AFCE2FBFCFC39" LSID="urn:lsid:plazi:treatment:50613239FF866C49FF0AFCE2FBFCFC39" httpUri="http://treatment.plazi.org/id/50613239FF866C49FF0AFCE2FBFCFC39" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509">
<subSubSection id="90D2D0A4FF866C48FF0AFCE2FDAFFCAC" box="[151,532,834,861]" pageId="8" pageNumber="509" type="nomenclature">
<paragraph id="D877832FFF866C48FF0AFCE2FDAFFCAC" blockId="8.[151,532,834,896]" box="[151,532,834,861]" pageId="8" pageNumber="509">
<heading id="833F3443FF866C48FF0AFCE2FDAFFCAC" bold="true" box="[151,532,834,861]" fontSize="11" level="1" pageId="8" pageNumber="509" reason="1">
<emphasis id="EABC5F3DFF866C48FF0AFCE2FDAFFCAC" bold="true" box="[151,532,834,861]" pageId="8" pageNumber="509">
3.2.3.
<taxonomicName id="1FC8F8ACFF866C48FF7CFCE2FE14FCAD" authorityName="Shavrin" authorityYear="2025" box="[225,431,834,861]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FF7CFCE2FE14FCAD" bold="true" box="[225,431,834,861]" italics="true" pageId="8" pageNumber="509">Omalium sectum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F18FE246FF866C48FE2BFCE2FDAFFCAC" box="[438,532,834,860]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF0AFCC6FED6FC70" box="[151,365,870,896]" pageId="8" pageNumber="509" type="description">
<paragraph id="D877832FFF866C48FF0AFCC6FED6FC70" blockId="8.[151,532,834,896]" box="[151,365,870,896]" pageId="8" pageNumber="509">
(
<figureCitation id="40F39FAAFF866C48FF02FCC6FF5CFC70" box="[159,231,870,896]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="8" pageNumber="509">Figs 3</figureCitation>
,
<figureCitation id="40F39FAAFF866C48FF69FCC6FE87FC70" box="[244,316,870,896]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="8" pageNumber="509">1012</figureCitation>
,
<figureCitation id="40F39FAAFF866C48FED5FCC6FEDEFC70" box="[328,357,870,896]" captionStart="FIGURE 30" captionStartId="9.[151,250,1849,1873]" captionTargetBox="[151,1435,993,1826]" captionTargetId="figure-318@9.[151,1435,993,1826]" captionTargetPageId="9" captionText="FIGURE 30. Distribution of Omalium in China: O. anguliferum (circles), O. rutilum (squares), O. sectum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896690" httpUri="https://zenodo.org/record/14896690/files/figure.png" pageId="8" pageNumber="509">30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF0AFC08FC66FBA1" pageId="8" pageNumber="509" type="materials_examined">
<paragraph id="D877832FFF866C48FF0AFC08FC66FBA1" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF0AFC08FE78FC31" bold="true" box="[151,451,935,962]" pageId="8" pageNumber="509">Type material examined.</emphasis>
<materialsCitation id="68A08972FF866C48FE51FC07FC66FBA1" pageId="8" pageNumber="509">
<typeStatus id="07733D8DFF866C48FE51FC07FD8FFC31" box="[460,564,935,961]" pageId="8" pageNumber="509" type="holotype">Holotype</typeStatus>
<specimenCount id="CECE48A6FF866C48FDA0FC08FDE9FC32" box="[573,594,936,962]" count="1" pageId="8" pageNumber="509" type="male"></specimenCount>
(
<figureCitation id="40F39FAAFF866C48FDFEFC07FD11FC32" box="[611,682,935,962]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="8" pageNumber="509">Fig. 3</figureCitation>
; dissected):
<emphasis id="EABC5F3DFF866C48FCA4FC07FBA8FC32" bold="true" box="[825,1043,935,962]" pageId="8" pageNumber="509">
<collectingCountry id="A0DFC3BFFF866C48FCA4FC07FC20FC31" box="[825,923,935,961]" name="China" pageId="8" pageNumber="509">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF866C48FC35FC07FBB1FC31" box="[936,1034,935,961]" country="China" name="Yunnan" pageId="8" pageNumber="509">YUNNAN</collectingRegion>
:
</emphasis>
CHINA: N-Yunnan [C2005-09] | Diqing
<collectingRegion id="1A0C4DCDFF866C48FF71FC6BFE91FC15" box="[236,298,971,997]" country="China" name="Xizang" pageId="8" pageNumber="509">Tibet</collectingRegion>
. Aut. Pref., | Deqin Co., Meili Xue Shan, | E-side,
<quantity id="1F302ECAFF866C48FCC2FC6CFC1DFC15" box="[863,934,971,998]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="509" unit="km" value="14.0">14 km</quantity>
W Deqin,
<quantity id="1F302ECAFF866C48FBBCFC6BFBC1FC16" box="[1057,1146,971,998]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="8" pageNumber="509" unit="m" value="2580.0">2580 m</quantity>
, &lt;printed&gt;,
<geoCoordinate id="BDFCE5E8FF866C48FA90FC6CFA23FC16" box="[1293,1432,972,998]" degrees="28" direction="north" minutes="27.47" orientation="latitude" pageId="8" pageNumber="509" precision="9" value="28.457834">28°27.47´N</geoCoordinate>
,
<geoCoordinate id="BDFCE5E8FF866C48FF0AFC4FFEA1FBFA" box="[151,282,1007,1034]" degrees="98" direction="east" minutes="46.35" orientation="longitude" pageId="8" pageNumber="509" precision="9" value="98.7725">98°46.35´E</geoCoordinate>
, creek | valley below glacier, mixed forest, | leaf litter, moss, dead wood, sifted, |
<date id="AC76A5EFFF866C48FB0CFC4FFAA8FBFA" box="[1169,1299,1007,1034]" pageId="8" pageNumber="509" value="2005-06-11">11.VI.2005</date>
, M. Schülke [C2005-09] &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke &lt;printed&gt;, HOLOTYPE |
<taxonomicName id="1FC8F8ACFF866C48FF0AFB97FF44FBA1" authorityName="Gravenhorst" authorityYear="1802" box="[151,255,1079,1105]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF866C48FF0AFB97FF44FBA1" box="[151,255,1079,1105]" italics="true" pageId="8" pageNumber="509">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF866C48FE8EFB98FEDAFBA1" authorityName="Shavrin" authorityYear="2025" box="[275,353,1080,1105]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FE8EFB98FEDAFBA1" box="[275,353,1080,1105]" italics="true" pageId="8" pageNumber="509">sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FEF5FB98FE7AFBA2" bold="true" box="[360,449,1080,1106]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FEF5FB98FE7AFBA2" box="[360,449,1080,1106]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025 &lt;red, printed&gt; (
<collectionCode id="BED91BEAFF866C48FC08FB97FC6AFBA1" box="[917,977,1079,1105]" pageId="8" pageNumber="509">cSch</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C49FF5AFBFBFE2AFD59" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509" type="description">
<paragraph id="D877832FFF866C48FF5AFBFBFEF9FB4E" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF5AFBFBFEE2FB85" bold="true" box="[199,345,1115,1141]" pageId="8" pageNumber="509">Description.</emphasis>
Measurements: HW: 0.62; HL: 0.43; OL: 0.17; TL: 0.12; AL: 1.35; PL: 0.55; PWmax: 0.72; PWmin: 0.67; ESL: 1.01; EW: 1.16; MTbL: 0.35; MTrL: 0.17 (MTrL 14: 0.07; MTrL 5: 0.10); AW: 1.02; AedL: 0.70; BL: 3.54.
</paragraph>
<paragraph id="D877832FFF866C48FF5AFB67FBEAFA65" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">
Habitus as in
<figureCitation id="40F39FAAFF866C48FEC3FB67FE1BFB12" box="[350,416,1223,1250]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="8" pageNumber="509">Fig. 3</figureCitation>
. Body reddish-brown, with slightly paler abdomen; antennomeres 611 brown; mouthparts, antennomeres 15 and legs yellow-brown. Punctation of head dense, denser and coarser in middle and on infraorbital portions; neck with moderately fine and sparse punctation; punctation of pronotum fine and sparse, sparser in medioapical, lateral and mediobasal portions; punctation of pronotum distinctly denser than that on pronotum, coarser in middle; abdominal tergites without punctation. Clypeus with fine transverse microsculpture; scutellum with fine isodiametric microreticulation; abdomen with dense isodiametric sculpture.
</paragraph>
<paragraph id="D877832FFF866C48FF5AFA3FFD6BF8B6" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">Head 1.4 times as broad as long, with broad and deep anteriomedian depressions; posteriolateral margins of clypeus stretching toward level of middle length of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with distinct transverse and longitudinal elevations between punctures in middle and irregular elongate elevations on postocular parts. Anteocellar foveae very wide and deep, somewhat elongate, reaching level of posterior third length of eyes. Temples 1.4 times as long as longitudinal length of eye, from posterior margin of eyes gradually narrowed posteriad, each with distinct subacute angle protruded basad. Apical part of neck relatively deeply depressed, with short irregular and narrow longitudinal elevations. Distance between ocelli 1.7 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with distinctly elongate antennomeres 28 and slightly elongate 910; antennomere 3 slightly longer and narrower than 2, 4 distinctly shorter than 3, 5 longer than 4, 67 longer and distinctly broader than 5, 8 slightly shorter than 7, 910 slightly shorter and distinctly broader than 8, apical antennomere slightly longer than 10.</paragraph>
<paragraph id="D877832FFF866C48FF5AF8F0FC80F809" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">Pronotum 1.3 times as broad as long, 1.1 times as broad as head, from widest anterior portion gradually narrowed posteriad toward widely rounded hind angles. Anterior angles acute and protruded anteriad. Anterior margin rounded, distinctly protruded anteriad. Each laterobasal portion not concave. Surface of disc with two wide and deep longitudinal and oval and deep anteriomedian depressions; surface between all depressions strongly elevated. Middle portion with irregular indistinct elevations.</paragraph>
<paragraph id="D877832FFF876C49FF5AFF37FD14FF09" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">Elytra 1.1 times as broad as long, 1.8 times as long as pronotum, from basal portion distinctly broadened posteriad; hind margins straight. Dorsal surface of each elytron with irregular elevations, diagonal in medioapical and median, and longitudinal in apical portions.</paragraph>
<paragraph id="D877832FFF876C49FF5AFEA3FC9FFEED" blockId="9.[151,1437,151,969]" box="[199,804,259,285]" pageId="9" pageNumber="510">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="D877832FFF876C49FF5AFE87FF07FE95" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">Abdomen slightly narrower than elytra, with wide intersegmental membranes between abdominal tergites V VI.</paragraph>
<paragraph id="D877832FFF876C49FF5AFECFFD63FD75" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
Male. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe moderately long, from narrow basal part significantly broadened slightly in front middle, strongly narrowed in preapical part and broadened in apical portion, with widely rounded apex; accessory plates narrow, curved and relatively long, rounded apically; parameres wide and long, reaching apical part of median lobe, from widest preapical parts gradully narrowed apically, each with two moderately short apical and two preapical setae; internal sac narrow and long, without sclerotized structures (
<figureCitation id="40F39FAAFF876C49FE08FDE7FE51FD91" box="[405,490,583,609]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="9" pageNumber="510">Fig. 10</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="40F39FAAFF876C49FC04FDE7FC50FD91" box="[921,1003,583,609]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="9" pageNumber="510">Fig. 11</figureCitation>
; apical portion of median lobe (lateral view) with crenulate ventrolateral margin (
<figureCitation id="40F39FAAFF876C49FDEAFDCBFD71FD75" box="[631,714,619,645]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="9" pageNumber="510">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="D877832FFF876C49FF5AFD2FFE2AFD59" blockId="9.[151,1437,151,969]" box="[199,401,655,681]" pageId="9" pageNumber="510">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFD12FA85FD01" pageId="9" pageNumber="510" type="discussion">
<paragraph id="D877832FFF876C49FF5AFD12FA85FD01" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFD12FE0BFD3D" bold="true" box="[199,432,690,717]" pageId="9" pageNumber="510">Comparative notes.</emphasis>
<taxonomicName id="1FC8F8ACFF876C49FE2BFD13FDC8FD3C" authorityName="Shavrin" authorityYear="2025" box="[438,627,691,717]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FE2BFD13FDC8FD3C" box="[438,627,691,717]" italics="true" pageId="9" pageNumber="510">Omalium sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FDE4FD13FD69FD3D" bold="true" box="[633,722,691,717]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FDE4FD13FD69FD3D" box="[633,722,691,717]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the remaining species of the acutangulum group by the shape of the pronotum with acute anterior angles protruded anteriad, and the shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFD5AFD31FCC9" pageId="9" pageNumber="510" type="distribution">
<paragraph id="D877832FFF876C49FF5AFD5AFD31FCC9" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFD5AFEDAFCE4" bold="true" box="[199,353,762,788]" pageId="9" pageNumber="510">Distribution.</emphasis>
<taxonomicName id="1FC8F8ACFF876C49FEF7FD5BFD91FCE4" authorityName="Shavrin" authorityYear="2025" box="[362,554,763,789]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FEF7FD5BFD91FCE4" box="[362,554,763,789]" italics="true" pageId="9" pageNumber="510">Omalium sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FDA9FD5BFD2BFCE5" bold="true" box="[564,656,763,789]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FDA9FD5BFD2BFCE5" box="[564,656,763,789]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from one locality in Meili Xue Shan and Diancang Shan mountain ranges in
<collectingRegion id="1A0C4DCDFF876C49FEE9FCBFFE76FCC9" box="[372,461,799,825]" country="China" name="Yunnan" pageId="9" pageNumber="510">Yunnan</collectingRegion>
,
<collectingCountry id="A0DFC3BFFF876C49FE4AFCBFFDA1FCC9" box="[471,538,799,825]" name="China" pageId="9" pageNumber="510">China</collectingCountry>
(
<figureCitation id="40F39FAAFF876C49FDB7FCBFFDC5FCC9" box="[554,638,799,825]" captionStart="FIGURE 30" captionStartId="9.[151,250,1849,1873]" captionTargetBox="[151,1435,993,1826]" captionTargetId="figure-318@9.[151,1435,993,1826]" captionTargetPageId="9" captionText="FIGURE 30. Distribution of Omalium in China: O. anguliferum (circles), O. rutilum (squares), O. sectum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896690" httpUri="https://zenodo.org/record/14896690/files/figure.png" pageId="9" pageNumber="510">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFCE3FF74FC71" pageId="9" pageNumber="510" type="etymology">
<paragraph id="D877832FFF876C49FF5AFCE3FF74FC71" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFCE3FEF5FCAD" bold="true" box="[199,334,835,861]" pageId="9" pageNumber="510">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="EABC5F3DFF876C49FD53FCE3FCA9FCAC" box="[718,786,835,860]" italics="true" pageId="9" pageNumber="510">sectus</emphasis>
, -
<emphasis id="EABC5F3DFF876C49FCB1FCE3FC81FCAC" box="[812,826,835,860]" italics="true" pageId="9" pageNumber="510">a</emphasis>
, -
<emphasis id="EABC5F3DFF876C49FCC9FCE3FCCDFCAC" box="[852,886,835,860]" italics="true" pageId="9" pageNumber="510">um</emphasis>
(divided). It alludes to the shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFC2AFBFCFC39" pageId="9" pageNumber="510" type="biology_ecology">
<paragraph id="D877832FFF876C49FF5AFC2AFBFCFC39" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFC2AFEF0FC54" bold="true" box="[199,331,906,932]" pageId="9" pageNumber="510">Bionomics.</emphasis>
Specimens were collected at elevation
<quantity id="1F302ECAFF876C49FC85FC2AFCCAFC55" box="[792,881,906,933]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="9" pageNumber="510" unit="m" value="2580.0">2580 m</quantity>
a.s.l. and was sifted from leaf litter, debris and mosses. It was collected together with
<taxonomicName id="1FC8F8ACFF876C49FDD5FC0FFD06FC39" authorityName="Shavrin" authorityYear="2025" box="[584,701,943,969]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FDD5FC0FFD06FC39" box="[584,701,943,969]" italics="true" pageId="9" pageNumber="510">O. rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FD59FC0FFCA5FC39" bold="true" box="[708,798,943,969]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FD59FC0FFCA5FC39" box="[708,798,943,969]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1FC8F8ACFF876C49FCC9FC0FFC5DFC39" authorityName="Shavrin" authorityYear="2025" box="[852,998,943,969]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FCC9FC0FFC5DFC39" box="[852,998,943,969]" italics="true" pageId="9" pageNumber="510">O. tibetanum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FC70FC0FFBFCFC39" bold="true" box="[1005,1095,943,969]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FC70FC0FFBFCFC39" box="[1005,1095,943,969]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,403 @@
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<mods:title id="7489F437EF67F40D2504A64FEC9F3994">Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China</mods:title>
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<emphasis id="EABC5F3DFF8B6C45FF0AFDE6FDA1FD91" bold="true" box="[151,538,582,609]" pageId="5" pageNumber="506">
3.2.2.
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<emphasis id="EABC5F3DFF8B6C45FF7CFDE6FE0EFD91" bold="true" box="[225,437,582,609]" italics="true" pageId="5" pageNumber="506">Omalium rutilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F18FE246FF8B6C45FE20FDE7FDA1FD91" box="[445,538,583,609]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8B6C45FF0AFDCBFE1EFD75" box="[151,421,618,645]" pageId="5" pageNumber="506" type="description">
<paragraph id="D877832FFF8B6C45FF0AFDCBFE1EFD75" blockId="5.[151,538,582,645]" box="[151,421,618,645]" pageId="5" pageNumber="506">
(
<figureCitation id="40F39FAAFF8B6C45FF02FDCBFF53FD75" box="[159,232,619,645]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="5" pageNumber="506">Figs 2</figureCitation>
,
<figureCitation id="40F39FAAFF8B6C45FF69FDCBFE9BFD75" box="[244,288,619,645]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="5" pageNumber="506">79</figureCitation>
,
<figureCitation id="40F39FAAFF8B6C45FEB1FDCAFECFFD75" box="[300,372,618,645]" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="5" pageNumber="506">1415</figureCitation>
, 30)
</paragraph>
</subSubSection>
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<emphasis id="EABC5F3DFF8B6C45FF0AFD13FE78FD3C" bold="true" box="[151,451,690,717]" pageId="5" pageNumber="506">Type material examined.</emphasis>
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<typeStatus id="07733D8DFF8B6C45FE56FD13FD88FD3D" box="[459,563,691,717]" pageId="5" pageNumber="506" type="holotype">Holotype</typeStatus>
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:
</emphasis>
CHINA: N-Yunnan [C2005-09] | Diqing
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. Aut. Pref., | Deqin Co., Meili Xue Shan, | E-side,
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W Deqin,
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, creek | valley below glacier, mixed forest, | leaf litter, moss, dead wood, sifted, |
<date id="AC76A5EFFF8B6C45FC6AFD5AFBC3FCE5" box="[1015,1144,762,789]" pageId="5" pageNumber="506" value="2005-06-11">11.VI.2005</date>
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|
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<emphasis id="EABC5F3DFF8B6C45FAD7FCBFFA27FCC9" box="[1354,1436,799,825]" italics="true" pageId="5" pageNumber="506">rutilum</emphasis>
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</emphasis>
| Shavrin A.V. des. 2025 &lt;red, printed&gt; (
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(
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,
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:
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(dissected; both antennes missing): same data as the
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(
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:
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: N-Yunnan Diqing
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| Aut.Pr. Deqin Co. Meili Xue | Shan E-side
<quantity id="1F302ECAFF8B6C45FBD5FC2BFB2BFC55" box="[1096,1168,907,933]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="506" unit="km" value="14.0">14 km</quantity>
W Deqin |
<geoCoordinate id="BDFCE5E8FF8B6C45FA8FFC2BFA23FC55" box="[1298,1432,907,933]" degrees="28" direction="north" minutes="27.47" orientation="latitude" pageId="5" pageNumber="506" precision="9" value="28.457834">28°27.47´N</geoCoordinate>
,
<geoCoordinate id="BDFCE5E8FF8B6C45FF0AFC0EFEA3FC39" box="[151,280,942,969]" degrees="98" direction="east" minutes="46.35" orientation="longitude" pageId="5" pageNumber="506" precision="9" value="98.7725">98°46.35´E</geoCoordinate>
<quantity id="1F302ECAFF8B6C45FEB9FC0EFEC9FC39" box="[292,370,942,969]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="5" pageNumber="506" unit="m" value="2580.0">2580m</quantity>
|
<date id="AC76A5EFFF8B6C45FE12FC0EFDB5FC39" box="[399,526,942,969]" pageId="5" pageNumber="506" value="2005-06-11">11.VI.2005</date>
A. Smetana [C158] (
<collectionCode id="BED91BEAFF8B6C45FC8BFC0FFCEEFC39" box="[790,853,943,969]" pageId="5" pageNumber="506">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF8B6C45FCF0FC0FFBA7FC39" box="[877,1052,943,969]" pageId="5" pageNumber="506">
<specimenCount id="CECE48A6FF8B6C45FCF0FC0FFC27FC39" box="[877,924,943,969]" count="1" pageId="5" pageNumber="506" type="male">1 ♂</specimenCount>
(dissected
</materialsCitation>
;
<materialsCitation id="68A08972FF8B6C45FBB1FC0FFE55FBA9" pageId="5" pageNumber="506">
right elytron missing):
<collectingCountry id="A0DFC3BFFF8B6C45FADDFC0FFA27FC39" box="[1344,1436,943,969]" name="China" pageId="5" pageNumber="506">CHINA</collectingCountry>
(
<collectingRegion id="1A0C4DCDFF8B6C45FF02FC73FF43FC1D" box="[159,248,979,1005]" country="China" name="Yunnan" pageId="5" pageNumber="506">Yunnan</collectingRegion>
) | Dali Bai Auton. Pref., | Diancang Shan W Dali |
<geoCoordinate id="BDFCE5E8FF8B6C45FCBBFC72FC0FFC1D" box="[806,948,978,1005]" degrees="25" direction="north" minutes="41" orientation="latitude" pageId="5" pageNumber="506" precision="15" seconds="20" value="25.688889">25°41´20´´N</geoCoordinate>
/
<geoCoordinate id="BDFCE5E8FF8B6C45FC23FC72FBE3FC1D" box="[958,1112,978,1005]" degrees="100" direction="east" minutes="06" orientation="longitude" pageId="5" pageNumber="506" precision="15" seconds="12" value="100.10333">100°06´12´´E</geoCoordinate>
|
<quantity id="1F302ECAFF8B6C45FBF7FC73FB05FC1D" box="[1130,1214,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="5" pageNumber="506" unit="m" value="3160.0">3160 m</quantity>
(small creek valley, | litter and debris sifted |
<date id="AC76A5EFFF8B6C45FE2DFC57FD99FBE1" box="[432,546,1015,1041]" pageId="5" pageNumber="506" value="2007-05-27">27.V.2007</date>
D.W.Wrase [02] &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke &lt;printed&gt; (
<collectionCode id="BED91BEAFF8B6C45FE1FFBBBFE05FBC5" box="[386,446,1051,1077]" pageId="5" pageNumber="506">cSch</collectionCode>
). All
<typeStatus id="07733D8DFF8B6C45FE63FBBBFDD2FBC5" box="[510,617,1051,1077]" pageId="5" pageNumber="506" type="paratype">paratypes</typeStatus>
with additional red printed label:
<typeStatus id="07733D8DFF8B6C45FC6AFBBBFB3CFBC5" box="[1015,1159,1051,1077]" pageId="5" pageNumber="506" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="1FC8F8ACFF8B6C45FB02FBBBFABCFBC5" authorityName="Gravenhorst" authorityYear="1802" box="[1183,1287,1051,1077]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF8B6C45FB02FBBBFABCFBC5" box="[1183,1287,1051,1077]" italics="true" pageId="5" pageNumber="506">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF8B6C45FA82FBBBFACAFBC5" authorityName="Shavrin" authorityYear="2025" box="[1311,1393,1051,1077]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF8B6C45FA82FBBBFACAFBC5" box="[1311,1393,1051,1077]" italics="true" pageId="5" pageNumber="506">rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8B6C45FAE6FBBBFF73FBA9" bold="true" pageId="5" pageNumber="506">
<taxonomicNameLabel id="F18FE246FF8B6C45FAE6FBBBFF73FBA9" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025.
</materialsCitation>
</paragraph>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8B6C47FF5AFBC2FD8BF806" lastPageId="7" lastPageNumber="508" pageId="5" pageNumber="506" type="description">
<paragraph id="D877832FFF8B6C45FF5AFBC2FE30FB19" blockId="5.[151,1437,690,2013]" pageId="5" pageNumber="506">
<emphasis id="EABC5F3DFF8B6C45FF5AFBC2FEE2FB8C" bold="true" box="[199,345,1122,1148]" pageId="5" pageNumber="506">Description.</emphasis>
Measurements (n=4): HW: 0.560.62; HL: 0.370.42; OL: 0.150.16; TL: 0.100.12; AL (
<typeStatus id="07733D8DFF8B6C45FF02FB27FEB3FB51" box="[159,264,1159,1185]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
): 1.15; PL: 0.370.49; PWmax: 0.660.76; PWmin: 0.600.72; ESL: 0.830.90; EW: 0.871.10; MTbL (
<typeStatus id="07733D8DFF8B6C45FF02FB0BFEB3FB35" box="[159,264,1195,1221]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
): 0.47; MTrL (
<typeStatus id="07733D8DFF8B6C45FE33FB0BFDACFB35" box="[430,535,1195,1221]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
): 0.20 (MTrL 14: 0.09; MTrL 5: 0.11); AW: 0.851.11; AedL: 0.47; BL: 2.57 3.10 (
<typeStatus id="07733D8DFF8B6C45FF4AFB6FFE84FB19" box="[215,319,1231,1257]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
: 2.70).
</paragraph>
<paragraph id="D877832FFF8B6C45FF5AFB53FB80F969" blockId="5.[151,1437,690,2013]" pageId="5" pageNumber="506">
Habitus as in
<figureCitation id="40F39FAAFF8B6C45FEC0FB53FE1AFAFD" box="[349,417,1267,1293]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="5" pageNumber="506">Fig. 2</figureCitation>
. Body reddish-brown, with distinctly darker head and abdomen; antennomeres 711 brown; mouthparts and antennomeres 16 yellow-brown; lateral portions of pronotum, lateral and apical portions of elytra, paratergites of abdomen and legs yellowish (tarsi slightly paler). Punctation of head dense, coarse and deep, denser and coarser in middle and on infraorbital portions, with interspaces between punctures in middle slightly less than diameter of nearest puncture; punctation of pronotum about as that in middle portion of head, but slightly larger, more irregular, slightly finer and sparser in middle and finer and sparser in mediobasal portion, with interspaces between punctures in middle about as long as diameter of nearest puncture; punctation of elytra slightly sparser and distinctly larger and deeper than that on pronotum, denser in mediobasal and mediolateral part, finer and sparser along suture and in medioapical portion. Anterior portion with distinct transverse microreticulation, middle part of clypeus with fine transverse and supra-antennal elevations with diagonal sculpture; mediobasal part of neck with transverse meshes; scutellum with dense and fine transverse microsculpture; abdomen with dense isodiametric sculpture. Apical and basal margins of pronotum with row of short cuticular fringe.
</paragraph>
<paragraph id="D877832FFF8B6C46FF5AF903FE47FF09" blockId="5.[151,1437,690,2013]" lastBlockId="6.[151,1436,150,249]" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">Head 1.41.5 times as broad as long, with strongly elevated supra-antennal elevations and wide and deep anteriomedian depressions. Dorsal surface with strong longitudinal elevations between punctures, distinctly diagonal in laterobasal parts of clypeus and on infraorbital portions; postocular parts with irregular longitudinal wrinkles. Anteocellar foveae narrow and deep, somewhat subparallel, almost reaching level of middle length of eyes. Temples 1.31.5 times as long as longitudinal length of eyes, each with subacute posterior angle protruded basad. Medioapical part of neck narrowly and slightly depressed, with short and narrow elevations. Distance between ocellus 1.51.7 times as long as distance between ocellus and posterior margin of eye. Apical palpomere about three and a half times as long as penultimate segment. Antenna with slightly elongate antennomeres 78 and distinctly transverse 910; antennomere 3 about as long as and distinctly narrower than 2, 4 distinctly shorter and slightly narrower than 3, 5 slightly longer than 4, 6 distinctly longer than 5, 7 indistinctly broader than 6, 8 slightly broader than 7, 910 distinctly broader than 8, apical antennomere slightly longer than preapical antennomere, from middle gradually narrowed toward rounded apex.</paragraph>
<caption id="8CB7D3A7FF886C46FF0AF9FEFEECF94D" ID-DOI="http://doi.org/10.5281/zenodo.14896688" ID-Zenodo-Dep="14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="6" pageNumber="507" startId="6.[151,264,1630,1654]" targetBox="[159,1428,281,1599]" targetPageId="6" targetType="figure">
<paragraph id="D877832FFF886C46FF0AF9FEFEECF94D" blockId="6.[151,1437,1630,1726]" pageId="6" pageNumber="507">
<emphasis id="EABC5F3DFF886C46FF0AF9FEFEF3F986" bold="true" box="[151,328,1630,1654]" pageId="6" pageNumber="507">FIGURES 412.</emphasis>
Aedeagus of
<taxonomicName id="1FC8F8ACFF886C46FE4AF9FEFD8EF986" authorityName="Gravenhorst" authorityYear="1802" box="[471,565,1630,1654]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF886C46FE4AF9FEFD8EF986" box="[471,565,1630,1654]" italics="true" pageId="6" pageNumber="507">Omalium</emphasis>
</taxonomicName>
(Figs 46—
<taxonomicName id="1FC8F8ACFF886C46FD25F9FEFCEFF986" authorityName="Shavrin" authorityYear="2025" box="[696,852,1630,1654]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="anguliferum">
<emphasis id="EABC5F3DFF886C46FD25F9FEFCEFF986" box="[696,852,1630,1654]" italics="true" pageId="6" pageNumber="507">O. anguliferum</emphasis>
</taxonomicName>
(paratype), Figs 79—
<taxonomicName id="1FC8F8ACFF886C46FBDEF9FEFB15F986" authorityName="Shavrin" authorityYear="2025" box="[1091,1198,1630,1654]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="rutilum">
<emphasis id="EABC5F3DFF886C46FBDEF9FEFB15F986" box="[1091,1198,1630,1654]" italics="true" pageId="6" pageNumber="507">O. rutilum</emphasis>
</taxonomicName>
(paratype), 1012—
<taxonomicName id="1FC8F8ACFF886C46FA19F9FEFF66F96A" authorityName="Shavrin" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="sectum">
<emphasis id="EABC5F3DFF886C46FA19F9FEFF66F96A" italics="true" pageId="6" pageNumber="507">O. sectum</emphasis>
</taxonomicName>
(holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="D877832FFF886C46FF5AF8B1FAD4F866" blockId="6.[151,1437,1808,1943]" pageId="6" pageNumber="507">Pronotum transverse, 1.51.7 times as broad as long, widest in anterior third, gradually narrowed posteriad. Laterobasal portions not concave in front of obtuse hind angles. Lateral and laterobasal portions deeply and widely impressed. Surface of disc with two long and deep longitudinal depression, distinctly broadened and deepened basad; medioapical part with deep oval depression; surface between all pronotal depressions strongly elevated.</paragraph>
<caption id="8CB7D3A7FF896C47FF0AFA26FEECF9FA" pageId="7" pageNumber="508" startId="7.[151,264,1414,1438]" subCaptionStartIDs="7.[619,664,1414,1438] 7.[1238,1283,1414,1438]" subCaptionStarts="Figs 13 &amp; Figs 15" targetBox="[159,1428,189,1382]" targetPageId="7">
<paragraph id="D877832FFF896C47FF0AFA26FEECF9FA" blockId="7.[151,1436,1414,1547]" pageId="7" pageNumber="508">
<emphasis id="EABC5F3DFF896C47FF0AFA26FEEFFA6E" bold="true" box="[151,340,1414,1438]" pageId="7" pageNumber="508">FIGURES 1329.</emphasis>
Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of
<taxonomicName id="1FC8F8ACFF896C47FEBEFA0AFE3AFA32" authorityName="Gravenhorst" authorityYear="1802" box="[291,385,1450,1474]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF896C47FEBEFA0AFE3AFA32" box="[291,385,1450,1474]" italics="true" pageId="7" pageNumber="508">Omalium</emphasis>
</taxonomicName>
: 13—
<taxonomicName id="1FC8F8ACFF896C47FE5CFA0BFDE0FA32" authorityName="Shavrin" authorityYear="2025" box="[449,603,1450,1474]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="anguliferum">
<emphasis id="EABC5F3DFF896C47FE5CFA0BFDE0FA32" box="[449,603,1450,1474]" italics="true" pageId="7" pageNumber="508">O. anguliferum</emphasis>
</taxonomicName>
, 1415—
<taxonomicName id="1FC8F8ACFF896C47FD22FA0BFC92FA32" authorityName="Shavrin" authorityYear="2025" box="[703,809,1450,1474]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="rutilum">
<emphasis id="EABC5F3DFF896C47FD22FA0BFC92FA32" box="[703,809,1450,1474]" italics="true" pageId="7" pageNumber="508">O. rutilum</emphasis>
</taxonomicName>
, 1617—
<taxonomicName id="1FC8F8ACFF896C47FC10FA0AFBBAFA32" authorityName="Shavrin" authorityYear="2025" box="[909,1025,1450,1474]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="517" phylum="Arthropoda" rank="species" species="cocleare">
O.
<emphasis id="EABC5F3DFF896C47FC36FA0AFBBAFA32" box="[939,1025,1450,1474]" italics="true" pageId="7" pageNumber="508">cocleare</emphasis>
</taxonomicName>
, 1819—
<taxonomicName id="1FC8F8ACFF896C47FBFBFA0BFB63FA32" authorityName="Shavrin" authorityYear="2025" box="[1126,1240,1451,1474]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="contiger">
<emphasis id="EABC5F3DFF896C47FBFBFA0BFB63FA32" box="[1126,1240,1451,1474]" italics="true" pageId="7" pageNumber="508">O. contiger</emphasis>
</taxonomicName>
, 20—
<taxonomicName id="1FC8F8ACFF896C47FA8AFA0BFA2DFA32" authorityName="Shavrin" authorityYear="2025" box="[1303,1430,1450,1474]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="kurbatovi">
<emphasis id="EABC5F3DFF896C47FA8AFA0BFA92FA32" box="[1303,1321,1451,1474]" italics="true" pageId="7" pageNumber="508">O</emphasis>
.
<emphasis id="EABC5F3DFF896C47FAA8FA0AFA2DFA32" box="[1333,1430,1450,1474]" italics="true" pageId="7" pageNumber="508">kurbatovi</emphasis>
</taxonomicName>
, 21—
<taxonomicName id="1FC8F8ACFF896C47FF57FA6FFEFDFA16" authorityName="Shavrin" authorityYear="2025" box="[202,326,1486,1510]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="EABC5F3DFF896C47FF57FA6FFEFDFA16" box="[202,326,1486,1510]" italics="true" pageId="7" pageNumber="508">O. schuelkei</emphasis>
</taxonomicName>
, 2223—
<taxonomicName id="1FC8F8ACFF896C47FE36FA6FFDD7FA16" authorityName="Shavrin" authorityYear="2025" box="[427,620,1486,1510]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="bambusaphilum">
<emphasis id="EABC5F3DFF896C47FE36FA6FFDD7FA16" box="[427,620,1486,1510]" italics="true" pageId="7" pageNumber="508">O. bambusaphilum</emphasis>
</taxonomicName>
, 2425—
<taxonomicName id="1FC8F8ACFF896C47FD4CFA6FFCEDFA17" authorityName="Shavrin" authorityYear="2025" box="[721,854,1487,1511]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="confertum">
<emphasis id="EABC5F3DFF896C47FD4CFA6FFCEDFA17" box="[721,854,1487,1511]" italics="true" pageId="7" pageNumber="508">O. confertum</emphasis>
</taxonomicName>
, 26—
<taxonomicName id="1FC8F8ACFF896C47FC08FA6FFC5DFA16" authorityName="Shavrin" authorityYear="2025" box="[917,998,1486,1510]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EABC5F3DFF896C47FC08FA6FFC1CFA16" box="[917,935,1487,1510]" italics="true" pageId="7" pageNumber="508">O</emphasis>
.
<emphasis id="EABC5F3DFF896C47FC2EFA6EFC5DFA16" box="[947,998,1486,1510]" italics="true" pageId="7" pageNumber="508">lama</emphasis>
</taxonomicName>
, 2728—
<taxonomicName id="1FC8F8ACFF896C47FBD6FA6FFB66FA16" authorityName="Shavrin" authorityYear="2025" box="[1099,1245,1487,1510]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="rougemonti">
<emphasis id="EABC5F3DFF896C47FBD6FA6FFB66FA16" box="[1099,1245,1487,1510]" italics="true" pageId="7" pageNumber="508">O. rougemonti</emphasis>
</taxonomicName>
, 29—
<taxonomicName id="1FC8F8ACFF896C47FA81FA6FFA2EFA16" authorityName="Shavrin" authorityYear="2025" box="[1308,1429,1487,1510]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="EABC5F3DFF896C47FA81FA6FFA2EFA16" box="[1308,1429,1487,1510]" italics="true" pageId="7" pageNumber="508">O. smetanai</emphasis>
</taxonomicName>
. Scale bar:
<quantity id="1F302ECAFF896C47FE9FFA53FEE8F9FA" box="[258,339,1523,1547]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="508" unit="mm" value="0.1">0.1 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="D877832FFF896C47FF5AF9EFFA3DF97D" blockId="7.[151,1436,1615,2038]" pageId="7" pageNumber="508">Elytra slightly less than twice or slightly more than twice as long as pronotum, lateral margins narrowly impressed; hind margins straight. Dorsal surface of each elytron with strong irregular elevations, finer in middle.</paragraph>
<paragraph id="D877832FFF896C47FF5AF937FF49F879" blockId="7.[151,1436,1615,2038]" pageId="7" pageNumber="508">
Male. Posterior margin of abdominal tergite VIII somewhat straight. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, slightly narrowed toward median lobe; median lobe moderately narrow, from widest middle gradually narrowed toward relatively small rounded apex; accessory plates narrow, moderately short and slightly curved apically; parameres wide, broadened in preapical part, each with two long and two short apical setae; internal sac moderately wide, long, without elongate sclerotized structures (
<figureCitation id="40F39FAAFF896C47FB6FF887FA8CF8B1" box="[1266,1335,1831,1857]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="7" pageNumber="508">Fig. 7</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="40F39FAAFF896C47FE4CF8EBFDAFF895" box="[465,532,1867,1893]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="7" pageNumber="508">Fig. 8</figureCitation>
; apical portion of median lobe elongate (lateral view), without ventrolateral teeth (
<figureCitation id="40F39FAAFF896C47FF02F8CFFF5FF879" box="[159,228,1903,1929]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="7" pageNumber="508">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="D877832FFF896C47FF5AF833FD8BF806" blockId="7.[151,1436,1615,2038]" pageId="7" pageNumber="508">
Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII widely rounded. Accessory sclerite with wide basal portion and short median part, strongly narrowed toward acute apex (
<figureCitation id="40F39FAAFF896C47FF02F87BFF48F805" box="[159,243,2011,2037]" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="7" pageNumber="508">Fig. 14</figureCitation>
). Spermatheca as in
<figureCitation id="40F39FAAFF896C47FE47F87BFD97F806" box="[474,556,2011,2038]" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="7" pageNumber="508">Fig. 15</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF5AFF36FDB4FE20" pageId="8" pageNumber="509" type="discussion">
<paragraph id="D877832FFF866C48FF5AFF36FDB4FE20" blockId="8.[151,1437,150,753]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF5AFF36FE0AFF40" bold="true" box="[199,433,150,176]" pageId="8" pageNumber="509">Comparative notes.</emphasis>
Based on the shape of postocular portions of head and the shape of the apical part of the pronotum with apical angles slightly protruded apicad,
<taxonomicName id="1FC8F8ACFF866C48FC94FF1BFCC4FF24" authorityName="Shavrin" authorityYear="2025" box="[777,895,186,213]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FC94FF1BFCC4FF24" box="[777,895,186,213]" italics="true" pageId="8" pageNumber="509">O. rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FC1AFF1AFC59FF24" bold="true" box="[903,994,186,212]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FC1AFF1AFC59FF24" box="[903,994,186,212]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to Nepalese
<taxonomicName id="1FC8F8ACFF866C48FB7AFF1BFA27FF24" authorityName="Shavrin" authorityYear="2025" box="[1255,1436,186,213]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="acutangulum">
<emphasis id="EABC5F3DFF866C48FB7AFF1BFA27FF24" box="[1255,1436,186,213]" italics="true" pageId="8" pageNumber="509">O. acutangulum</emphasis>
</taxonomicName>
and Chinese
<taxonomicName id="1FC8F8ACFF866C48FEB4FF7FFE6EFF08" authorityName="Shavrin" authorityYear="2025" box="[297,469,222,249]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="anguliferum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FEB4FF7FFE6EFF08" box="[297,469,222,249]" italics="true" pageId="8" pageNumber="509">O. anguliferum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FE40FF7EFD8CFF08" bold="true" box="[477,567,222,248]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FE40FF7EFD8CFF08" box="[477,567,222,248]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Based on general shapes of the body, the aedeagus and the female accessory sclerite, it is more similar to
<taxonomicName id="1FC8F8ACFF866C48FE52FEA3FD39FEEC" authorityName="Shavrin" authorityYear="2025" box="[463,642,258,285]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="acutangulum">
<emphasis id="EABC5F3DFF866C48FE52FEA3FD39FEEC" box="[463,642,258,285]" italics="true" pageId="8" pageNumber="509">O. acutangulum</emphasis>
</taxonomicName>
, from which it can be distinguished by the paler body, finer longitudinal elevations of median part of the head, denser punctation of the pronotum and the elytra, slightly shorter median lobe, broader parameres, and other details of the external and internal morphology of the aedeagus. From
<taxonomicName id="1FC8F8ACFF866C48FB6FFEEBFA27FE94" authorityName="Shavrin" authorityYear="2025" box="[1266,1436,330,357]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="anguliferum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FB6FFEEBFA27FE94" box="[1266,1436,330,357]" italics="true" pageId="8" pageNumber="509">O. anguliferum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FF0AFECEFF42FE78" bold="true" box="[151,249,366,392]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FF0AFECEFF48FE78" box="[151,243,366,392]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
it differs by the narrower antennomere 8, the shape of more transverse pronotum, widest in anterior third and with more pronounced dorsal (including medioapical) impressions, and different morphology of the aedeagus and the female accessory sclerite.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF5AFE7AFB6AFDE8" pageId="8" pageNumber="509" type="distribution">
<paragraph id="D877832FFF866C48FF5AFE7AFB6AFDE8" blockId="8.[151,1437,150,753]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF5AFE7AFEDAFE04" bold="true" box="[199,353,474,500]" pageId="8" pageNumber="509">Distribution.</emphasis>
<taxonomicName id="1FC8F8ACFF866C48FEF6FE7AFD94FE04" authorityName="Shavrin" authorityYear="2025" box="[363,559,474,500]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FEF6FE7AFD94FE04" box="[363,559,474,500]" italics="true" pageId="8" pageNumber="509">Omalium rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FDA7FE7AFD2CFE04" bold="true" box="[570,663,474,500]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FDA7FE7AFD2CFE04" box="[570,663,474,500]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from three localities in Qinling Shan mountain range in
<collectingRegion id="1A0C4DCDFF866C48FF0AFE5EFF4AFDE8" box="[151,241,510,536]" country="China" name="Shaanxi" pageId="8" pageNumber="509">Shaanxi</collectingRegion>
and in Meili Xue Shan and Diancang Shan mountain ranges in
<collectingRegion id="1A0C4DCDFF866C48FC27FE5FFBA8FDE9" box="[954,1043,511,537]" country="China" name="Yunnan" pageId="8" pageNumber="509">Yunnan</collectingRegion>
,
<collectingCountry id="A0DFC3BFFF866C48FB83FE5EFBDAFDE8" box="[1054,1121,510,536]" name="China" pageId="8" pageNumber="509">China</collectingCountry>
(
<figureCitation id="40F39FAAFF866C48FBEDFE5EFB7FFDE8" box="[1136,1220,510,536]" captionStart="FIGURE 30" captionStartId="9.[151,250,1849,1873]" captionTargetBox="[151,1435,993,1826]" captionTargetId="figure-318@9.[151,1435,993,1826]" captionTargetPageId="9" captionText="FIGURE 30. Distribution of Omalium in China: O. anguliferum (circles), O. rutilum (squares), O. sectum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896690" httpUri="https://zenodo.org/record/14896690/files/figure.png" pageId="8" pageNumber="509">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF5AFD82FF45FD90" pageId="8" pageNumber="509" type="etymology">
<paragraph id="D877832FFF866C48FF5AFD82FF45FD90" blockId="8.[151,1437,150,753]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF5AFD82FEF5FDCC" bold="true" box="[199,334,546,572]" pageId="8" pageNumber="509">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="EABC5F3DFF866C48FCB2FD82FCC3FDCC" box="[815,888,546,572]" italics="true" pageId="8" pageNumber="509">rutilus</emphasis>
, -
<emphasis id="EABC5F3DFF866C48FC0CFD83FC24FDCC" box="[913,927,547,572]" italics="true" pageId="8" pageNumber="509">a</emphasis>
, -
<emphasis id="EABC5F3DFF866C48FC2AFD83FC62FDCC" box="[951,985,547,572]" italics="true" pageId="8" pageNumber="509">um</emphasis>
(reddish). It alludes to the coloration of the body.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF5AFDCAFC97FD00" pageId="8" pageNumber="509" type="biology_ecology">
<paragraph id="D877832FFF866C48FF5AFDCAFC97FD00" blockId="8.[151,1437,150,753]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF5AFDCAFEF0FD74" bold="true" box="[199,331,618,644]" pageId="8" pageNumber="509">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="1F302ECAFF866C48FCD0FDCAFC45FD75" box="[845,1022,618,645]" metricMagnitude="3" metricUnit="m" metricValue="2.87" metricValueMax="3.16" metricValueMin="2.58" pageId="8" pageNumber="509" unit="m" value="2870.0" valueMax="3160.0" valueMin="2580.0">2580 to 3160 m</quantity>
a.s.l. and were sifted from leaf litter, debris and mosses.
<specimenCount id="CECE48A6FF866C48FEE9FD2EFDAEFD58" box="[372,533,654,680]" count="1" pageId="8" pageNumber="509" type="generic">One specimen</specimenCount>
was collected by sifting of wet leaf litter, debris and moss on and around small seepages and on edges of a small creek in a mixed broadleaved forest (C158). It was collected in Meili Xue Shan together with
<taxonomicName id="1FC8F8ACFF866C48FEAEFD77FE18FD00" authorityName="Shavrin" authorityYear="2025" box="[307,419,727,753]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FEAEFD77FE18FD00" box="[307,419,727,753]" italics="true" pageId="8" pageNumber="509">O. sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FE37FD76FDB8FD00" bold="true" box="[426,515,726,752]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FE37FD76FDB8FD00" box="[426,515,726,752]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1FC8F8ACFF866C48FDA7FD77FD77FD00" authorityName="Shavrin" authorityYear="2025" box="[570,716,726,753]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FDA7FD77FD77FD00" box="[570,716,726,753]" italics="true" pageId="8" pageNumber="509">O. tibetanum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FD4EFD76FC97FD00" bold="true" box="[723,812,726,752]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FD4EFD76FC97FD00" box="[723,812,726,752]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,556 @@
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<paragraph id="D877832FFF8C6C42FF0AFC55FDEDFBFF" blockId="2.[151,598,1012,1075]" box="[151,598,1012,1039]" pageId="2" pageNumber="503">
<heading id="833F3443FF8C6C42FF0AFC55FDEDFBFF" bold="true" box="[151,598,1012,1039]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="EABC5F3DFF8C6C42FF0AFC55FDEDFBFF" bold="true" box="[151,598,1012,1039]" pageId="2" pageNumber="503">
3.2.1.
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<emphasis id="EABC5F3DFF8C6C42FF7CFC54FE49FBFF" bold="true" box="[225,498,1012,1039]" italics="true" pageId="2" pageNumber="503">Omalium anguliferum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F18FE246FF8C6C42FE64FC55FDEDFBFF" box="[505,598,1013,1039]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8C6C42FF0AFBB9FEC0FBC3" box="[151,379,1049,1075]" pageId="2" pageNumber="503" type="description">
<paragraph id="D877832FFF8C6C42FF0AFBB9FEC0FBC3" blockId="2.[151,598,1012,1075]" box="[151,379,1049,1075]" pageId="2" pageNumber="503">
(
<figureCitation id="40F39FAAFF8C6C42FF02FBB9FF5DFBC3" box="[159,230,1049,1075]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="2" pageNumber="503">Figs 1</figureCitation>
,
<figureCitation id="40F39FAAFF8C6C42FF69FBB9FE9BFBC3" box="[244,288,1049,1075]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="2" pageNumber="503">46</figureCitation>
,
<figureCitation id="40F39FAAFF8C6C42FEB1FBB9FEF1FBC3" box="[300,330,1049,1075]" captionStart="FIGURES 1329" captionStartId="7.[151,264,1414,1438]" captionTargetBox="[159,1428,189,1382]" captionTargetId="figure-16@7.[151,1436,181,1390]" captionText="FIGURES 1329. Female accessory sclerite (Figs 1314, 16, 18, 2022, 24, 26, 28, 30) and spermatheca (Figs 15, 17, 19, 23, 25, 27, 29) of Omalium: 13—O. anguliferum, 1415—O. rutilum, 1617—O. cocleare, 1819—O. contiger, 20—O. kurbatovi, 21—O. schuelkei, 2223—O. bambusaphilum, 2425—O. confertum, 26—O. lama, 2728—O. rougemonti, 29—O. smetanai. Scale bar: 0.1 mm." pageId="2" pageNumber="503">13</figureCitation>
, 30)
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8C6C42FF0AFBC1FC71F864" pageId="2" pageNumber="503" type="materials_examined">
<paragraph id="D877832FFF8C6C42FF0AFBC1FC3FFB17" blockId="2.[151,1437,1121,1940]" pageId="2" pageNumber="503">
<emphasis id="EABC5F3DFF8C6C42FF0AFBC1FE04FB8B" bold="true" box="[151,447,1121,1147]" pageId="2" pageNumber="503">Type material examined.</emphasis>
<materialsCitation id="68A08972FF8C6C42FE5BFBC1FB15FAFB" pageId="2" pageNumber="503">
<typeStatus id="07733D8DFF8C6C42FE5BFBC1FD95FB8B" box="[454,558,1121,1147]" pageId="2" pageNumber="503" type="holotype">Holotype</typeStatus>
<specimenCount id="CECE48A6FF8C6C42FDA8FBC1FDF6FB8B" box="[565,589,1121,1147]" count="1" pageId="2" pageNumber="503" type="male"></specimenCount>
:
<emphasis id="EABC5F3DFF8C6C42FDC4FBC1FC8DFB8B" bold="true" box="[601,822,1121,1147]" pageId="2" pageNumber="503">
<collectingCountry id="A0DFC3BFFF8C6C42FDC4FBC1FD00FB8B" box="[601,699,1121,1147]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF8C6C42FD5AFBC1FC96FB8B" box="[711,813,1121,1147]" country="China" name="Sichuan" pageId="2" pageNumber="503">SICHUAN</collectingRegion>
:
</emphasis>
<emphasis id="EABC5F3DFF8C6C42FCDAFBC1FC1FFB8B" bold="true" box="[839,932,1121,1147]" pageId="2" pageNumber="503">
<collectingCountry id="A0DFC3BFFF8C6C42FCDAFBC1FC1FFB8B" box="[839,932,1121,1147]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
</emphasis>
:
<collectingRegion id="1A0C4DCDFF8C6C42FC2EFBC1FBE4FB8B" box="[947,1119,1121,1147]" country="China" name="Sichuan" pageId="2" pageNumber="503">SICHUAN PROV.</collectingRegion>
, | Emeishan Mt.,
<date id="AC76A5EFFF8C6C42FAB9FBC1FA23FB8B" box="[1316,1432,1121,1147]" pageId="2" pageNumber="503" value="2014-06-09">9.VI.2014</date>
, | LEIDONGPING env.,
<quantity id="1F302ECAFF8C6C42FE16FB25FE5EFB6F" box="[395,485,1157,1183]" metricMagnitude="3" metricUnit="m" metricValue="2.41" pageId="2" pageNumber="503" unit="m" value="2410.0">2410 m</quantity>
, | 29°32´40´´
<geoCoordinate id="BDFCE5E8FF8C6C42FD17FB25FC93FB6F" box="[650,808,1157,1183]" degrees="103" direction="east" minutes="20" orientation="longitude" pageId="2" pageNumber="503" precision="15" seconds="02" value="103.33389">103°20´02´´E</geoCoordinate>
, &lt;printed&gt;,
<emphasis id="EABC5F3DFF8C6C42FC5BFB25FB00FB6F" bold="true" box="[966,1211,1156,1183]" pageId="2" pageNumber="503">sift #8, mixed forest</emphasis>
with
<taxonomicName id="1FC8F8ACFF8C6C42FA9CFB25FA8DFB6F" box="[1281,1334,1157,1183]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF8C6C42FA9CFB25FA8DFB6F" box="[1281,1334,1157,1183]" italics="true" pageId="2" pageNumber="503">Acer</emphasis>
</taxonomicName>
, |
<taxonomicName id="1FC8F8ACFF8C6C42FACAFB25FF6CFB33" authority=", Picea" authorityName="Picea" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF8C6C42FACAFB25FA2EFB6F" box="[1367,1429,1157,1183]" italics="true" pageId="2" pageNumber="503">Abies</emphasis>
,
<emphasis id="EABC5F3DFF8C6C42FF0AFB0AFF6CFB33" box="[151,215,1194,1219]" italics="true" pageId="2" pageNumber="503">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="1FC8F8ACFF8C6C42FF78FB09FE31FB33" box="[229,394,1193,1219]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF8C6C42FF78FB09FE31FB33" box="[229,394,1193,1219]" italics="true" pageId="2" pageNumber="503">Rhododendron</emphasis>
</taxonomicName>
, | around limestone rocks, | J. Hájek &amp; J. Růžička leg. &lt;printed&gt;, HOLOTYPE |
<taxonomicName id="1FC8F8ACFF8C6C42FABAFB09FA34FB33" authorityName="Gravenhorst" authorityYear="1802" box="[1319,1423,1193,1219]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF8C6C42FABAFB09FA34FB33" box="[1319,1423,1193,1219]" italics="true" pageId="2" pageNumber="503">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF8C6C42FF0AFB6DFE9BFB17" authorityName="Shavrin" authorityYear="2025" box="[151,288,1229,1255]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="anguliferum" status="sp. nov.">
<emphasis id="EABC5F3DFF8C6C42FF0AFB6DFE9BFB17" box="[151,288,1229,1255]" italics="true" pageId="2" pageNumber="503">anguliferum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8C6C42FEBAFB6DFE3AFB17" bold="true" box="[295,385,1229,1255]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="F18FE246FF8C6C42FEBAFB6DFE3AFB17" box="[295,385,1229,1255]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025 &lt;printed&gt; (
<collectionCode id="BED91BEAFF8C6C42FCBFFB6DFCC3FB17" box="[802,888,1229,1255]" pageId="2" pageNumber="503">NMPC</collectionCode>
).
<paragraph id="D877832FFF8C6C42FF6AFB51FC71F864" blockId="2.[151,1437,1121,1940]" pageId="2" pageNumber="503">
<emphasis id="EABC5F3DFF8C6C42FF6AFB51FED4FAFB" bold="true" box="[247,367,1265,1291]" pageId="2" pageNumber="503">
<typeStatus id="07733D8DFF8C6C42FF6AFB51FED4FAFB" box="[247,367,1265,1291]" pageId="2" pageNumber="503" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="CECE48A6FF8C6C42FEE0FB51FE70FAFB" box="[381,459,1265,1291]" count="12" pageId="2" pageNumber="503" type="male">12 ♂♂</specimenCount>
,
<specimenCount id="CECE48A6FF8C6C42FE4BFB51FDA5FAFB" box="[470,542,1265,1291]" count="11" pageId="2" pageNumber="503" type="female">11 ♀♀</specimenCount>
):
<emphasis id="EABC5F3DFF8C6C42FDA5FB51FCAFFAFB" bold="true" box="[568,788,1265,1291]" pageId="2" pageNumber="503">
<collectingCountry id="A0DFC3BFFF8C6C42FDA5FB51FD21FAFB" box="[568,666,1265,1291]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF8C6C42FD38FB51FCB0FAFB" box="[677,779,1265,1291]" country="China" name="Sichuan" pageId="2" pageNumber="503">SICHUAN</collectingRegion>
:
</emphasis>
<specimenCount id="CECE48A6FF8C6C42FC84FB51FCFFFAFB" box="[793,836,1265,1291]" count="1" pageId="2" pageNumber="503" type="female">1 ♀</specimenCount>
: same data as the
<typeStatus id="07733D8DFF8C6C42FB9AFB51FBD2FAFB" box="[1031,1129,1265,1291]" pageId="2" pageNumber="503" type="holotype">holotype</typeStatus>
(
<collectionCode id="BED91BEAFF8C6C42FBEAFB51FB1DFAFB" box="[1143,1190,1265,1291]" pageId="2" pageNumber="503">cSh</collectionCode>
);
<materialsCitation id="68A08972FF8C6C42FB25FB51FF46FA87" pageId="2" pageNumber="503">
<specimenCount id="CECE48A6FF8C6C42FB25FB51FB4DFAFB" box="[1208,1270,1265,1291]" count="2" pageId="2" pageNumber="503" type="male">2 ♂♂</specimenCount>
(
<specimenCount id="CECE48A6FF8C6C42FA9EFB51FA27FAFB" box="[1283,1436,1265,1291]" count="1" pageId="2" pageNumber="503" type="generic">one specimen</specimenCount>
dissected):
<emphasis id="EABC5F3DFF8C6C42FEBDFAB5FEC6FADF" bold="true" box="[288,381,1301,1327]" pageId="2" pageNumber="503">
<collectingCountry id="A0DFC3BFFF8C6C42FEBDFAB5FEC6FADF" box="[288,381,1301,1327]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
</emphasis>
:
<collectingRegion id="1A0C4DCDFF8C6C42FE10FAB5FD80FADF" box="[397,571,1301,1327]" country="China" name="Sichuan" pageId="2" pageNumber="503">SICHUAN PROV.</collectingRegion>
, | Emeishan Mt.,
<quantity id="1F302ECAFF8C6C42FC9EFAB5FCE2FADF" box="[771,857,1301,1327]" metricMagnitude="3" metricUnit="m" metricValue="2.42" pageId="2" pageNumber="503" unit="m" value="2420.0">2420 m</quantity>
| JIEYINGDIAN TEMPLE, | 29°32´17´´
<geoCoordinate id="BDFCE5E8FF8C6C42FB69FAB5FA28FAC0" box="[1268,1427,1301,1328]" degrees="103" direction="east" minutes="19" orientation="longitude" pageId="2" pageNumber="503" precision="15" seconds="59" value="103.33305">103°19´59´´E</geoCoordinate>
, &lt;printed&gt;,
<date id="AC76A5EFFF8C6C42FE87FA99FE13FAA3" box="[282,424,1337,1363]" pageId="2" pageNumber="503" value="2014-06-10">10.VI. 2014</date>
,
<emphasis id="EABC5F3DFF8C6C42FE2EFA99FDAAFAA3" bold="true" box="[435,529,1336,1363]" pageId="2" pageNumber="503">sift #12,</emphasis>
| secondary
<emphasis id="EABC5F3DFF8C6C42FD03FA99FC89FAA3" bold="true" box="[670,818,1337,1363]" pageId="2" pageNumber="503">mixed forest</emphasis>
| above temple, | J. Hájek &amp; J. Růžička leg. &lt;printed&gt; (
<collectionCode id="BED91BEAFF8C6C42FF02FAFDFF4EFA87" box="[159,245,1373,1399]" pageId="2" pageNumber="503">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF8C6C42FE94FAFDFDF7FA4F" pageId="2" pageNumber="503">
<specimenCount id="CECE48A6FF8C6C42FE94FAFDFE8FFA87" box="[265,308,1373,1399]" count="1" pageId="2" pageNumber="503" type="female">1 ♀</specimenCount>
(dissected):
<emphasis id="EABC5F3DFF8C6C42FE52FAFDFD97FA87" bold="true" box="[463,556,1373,1399]" pageId="2" pageNumber="503">
<collectingCountry id="A0DFC3BFFF8C6C42FE52FAFDFD97FA87" box="[463,556,1373,1399]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
</emphasis>
:
<collectingRegion id="1A0C4DCDFF8C6C42FDA1FAFDFD51FA87" box="[572,746,1373,1399]" country="China" name="Sichuan" pageId="2" pageNumber="503">SICHUAN PROV.</collectingRegion>
, | Emeishan Mt.,
<quantity id="1F302ECAFF8C6C42FC2EFAFDFBB2FA87" box="[947,1033,1373,1399]" metricMagnitude="3" metricUnit="m" metricValue="2.48" pageId="2" pageNumber="503" unit="m" value="2480.0">2480 m</quantity>
| JIEYINGDIAN TEMPLE, | 29°32´01´´
<geoCoordinate id="BDFCE5E8FF8C6C42FF0AFA21FE8DFA6C" box="[151,310,1409,1436]" degrees="103" direction="east" minutes="19" orientation="longitude" pageId="2" pageNumber="503" precision="15" seconds="57" value="103.3325">103°19´57´´E</geoCoordinate>
, &lt;printed&gt;,
<date id="AC76A5EFFF8C6C42FE50FA21FDF4FA6B" box="[461,591,1409,1435]" pageId="2" pageNumber="503" value="2014-06-07">7.VI. 2014</date>
,
<emphasis id="EABC5F3DFF8C6C42FDC1FA21FD16FA6B" bold="true" box="[604,685,1408,1435]" pageId="2" pageNumber="503">sift #6,</emphasis>
|
<emphasis id="EABC5F3DFF8C6C42FD58FA21FC40FA6B" bold="true" box="[709,1019,1409,1435]" pageId="2" pageNumber="503">bamboo and mixed forest</emphasis>
| below temple near road, | J. Hájek &amp; J. Růžička leg. &lt;printed&gt; (
<collectionCode id="BED91BEAFF8C6C42FE73FA05FDFFFA4F" box="[494,580,1445,1471]" pageId="2" pageNumber="503">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF8C6C42FDC4FA05FD60FA13" pageId="2" pageNumber="503">
<specimenCount id="CECE48A6FF8C6C42FDC4FA05FD3EFA4F" box="[601,645,1445,1471]" count="1" pageId="2" pageNumber="503" type="male">1 ♂</specimenCount>
(dissected),
<specimenCount id="CECE48A6FF8C6C42FC85FA05FCFDFA4F" box="[792,838,1445,1471]" count="1" pageId="2" pageNumber="503" type="female">1 ♀</specimenCount>
:
<collectingCountry id="A0DFC3BFFF8C6C42FCC4FA05FC00FA4F" box="[857,955,1445,1471]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
:
<collectingRegion id="1A0C4DCDFF8C6C42FC5AFA05FB9AFA4F" box="[967,1057,1445,1471]" country="China" name="Sichuan" pageId="2" pageNumber="503">Sichuan</collectingRegion>
| Emei Shan, Wannian |
<quantity id="1F302ECAFF8C6C42FAA3FA05FA2CFA4F" box="[1342,1431,1445,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="503" unit="m" value="1050.0">1050 m</quantity>
,
<date id="AC76A5EFFF8C6C42FF0AFA69FEFDFA13" box="[151,326,1481,1507]" pageId="2" pageNumber="503" value="1999-03-19" valueMax="1999-03-30" valueMin="1999-03-19">19.30.III.1999</date>
| leg. P. JÄGER &lt;printed&gt; (
<collectionCode id="BED91BEAFF8C6C42FD1CFA69FD68FA13" box="[641,723,1481,1507]" pageId="2" pageNumber="503">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF8C6C42FD7BFA69FF5CF9DB" pageId="2" pageNumber="503">
<specimenCount id="CECE48A6FF8C6C42FD7BFA69FC98FA13" box="[742,803,1481,1507]" count="3" pageId="2" pageNumber="503" type="male">3 ♂♂</specimenCount>
(
<specimenCount id="CECE48A6FF8C6C42FCACFA69FC70FA13" box="[817,971,1481,1507]" count="1" pageId="2" pageNumber="503" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="CECE48A6FF8C6C42FBD0FA69FB36FA13" box="[1101,1165,1481,1507]" count="3" pageId="2" pageNumber="503" type="female">3 ♀♀</specimenCount>
: P.R.
<collectingCountry id="A0DFC3BFFF8C6C42FB48FA69FA8AFA13" box="[1237,1329,1481,1507]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
,
<collectingRegion id="1A0C4DCDFF8C6C42FAA6FA69FA23FA13" box="[1339,1432,1481,1507]" country="China" name="Sichuan" pageId="2" pageNumber="503">Sichuan</collectingRegion>
, | EmeiShan, N29°33´36.3´´ | E103°20´38.0´´, |
<date id="AC76A5EFFF8C6C42FD33FA4DFC93F9F8" box="[686,808,1517,1544]" pageId="2" pageNumber="503" value="2010-06-15">15.vi.2010</date>
,
<quantity id="1F302ECAFF8C6C42FCA8FA4DFC3DF9F7" box="[821,902,1517,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.9469999999999998" pageId="2" pageNumber="503" unit="m" value="1947.0">1947m</quantity>
, | sifting33, V. Grebennikov &lt;printed&gt; (
<specimenCount id="CECE48A6FF8C6C42FACBFA4DFA3AF9F7" box="[1366,1409,1517,1543]" count="1" pageId="2" pageNumber="503" type="male">1 ♂</specimenCount>
,
<specimenCount id="CECE48A6FF8C6C42FA13FA4DFF15F9DB" count="1" pageId="2" pageNumber="503" type="female">1 ♀</specimenCount>
:
<collectionCode id="BED91BEAFF8C6C42FF27F9B1FF5CF9DB" box="[186,231,1553,1579]" pageId="2" pageNumber="503">cSh</collectionCode>
</materialsCitation>
;
<materialsCitation id="68A08972FF8C6C42FF6EF9B1FE69F9DB" box="[243,466,1553,1579]" pageId="2" pageNumber="503">
<specimenCount id="CECE48A6FF8C6C42FF6EF9B1FE88F9DB" box="[243,307,1553,1579]" count="2" pageId="2" pageNumber="503" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CECE48A6FF8C6C42FEA2F9B1FEC4F9DB" box="[319,383,1553,1579]" count="2" pageId="2" pageNumber="503" type="female">2 ♀♀</specimenCount>
:
<collectionCode id="BED91BEAFF8C6C42FE11F9B1FE71F9DB" box="[396,458,1553,1579]" pageId="2" pageNumber="503">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68A08972FF8C6C42FE40F9B1FBA2F9BF" pageId="2" pageNumber="503">
<specimenCount id="CECE48A6FF8C6C42FE40F9B1FDB3F9DB" box="[477,520,1553,1579]" count="1" pageId="2" pageNumber="503" type="male">1 ♂</specimenCount>
,
<specimenCount id="CECE48A6FF8C6C42FD89F9B1FDEFF9DB" box="[532,596,1553,1579]" count="3" pageId="2" pageNumber="503" type="female">3 ♀♀</specimenCount>
: P.R.
<collectingCountry id="A0DFC3BFFF8C6C42FD00F9B1FD42F9DB" box="[669,761,1553,1579]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
,
<collectingRegion id="1A0C4DCDFF8C6C42FC99F9B1FCDAF9DB" box="[772,865,1553,1579]" country="China" name="Sichuan" pageId="2" pageNumber="503">Sichuan</collectingRegion>
, | Emei Shan, N29°33.6´| E103°20.6´, 27.vi.5.vii. | 2009,
<quantity id="1F302ECAFF8C6C42FF76F995FE39F9BF" box="[235,386,1589,1615]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="2" pageNumber="503" unit="m" value="2100.0" valueMax="2400.0" valueMin="1800.0">18002400m</quantity>
, sifti-| ng1117, V. Grebennikov &lt;printed&gt; (
<specimenCount id="CECE48A6FF8C6C42FC1CF995FC1EF9BF" box="[897,933,1589,1615]" count="1" pageId="2" pageNumber="503" type="male">1♂</specimenCount>
,
<specimenCount id="CECE48A6FF8C6C42FC2FF995FC64F9BF" box="[946,991,1589,1615]" count="1" pageId="2" pageNumber="503" type="female">1 ♀</specimenCount>
:
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: P.R.
<collectingCountry id="A0DFC3BFFF8C6C42FF0AF9F9FF48F983" box="[151,243,1625,1651]" name="China" pageId="2" pageNumber="503">CHINA</collectingCountry>
.
<collectingRegion id="1A0C4DCDFF8C6C42FF62F9F9FEE7F983" box="[255,348,1625,1651]" country="China" name="Sichuan" pageId="2" pageNumber="503">Sichuan</collectingRegion>
, | EmeiShan, N29°33´36.3´´ | E103°20´38.0´´, |
<date id="AC76A5EFFF8C6C42FCE2F9F9FC42F983" box="[895,1017,1625,1651]" pageId="2" pageNumber="503" value="2010-06-22">22.vi.2010</date>
,
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, | sifting39, V. Grebennikov &lt;printed&gt; (
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,
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:
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:
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)
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;
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:
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:
<collectingRegion id="1A0C4DCDFF8C6C42FCB3F9DDFC33F967" box="[814,904,1661,1687]" country="China" name="Sichuan" pageId="2" pageNumber="503">Sichuan</collectingRegion>
(2) | Qingcheng-Shan, | Rückseite,
<quantity id="1F302ECAFF8C6C42FA85F9DDFA34F968" box="[1304,1423,1661,1688]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="2" pageNumber="503" unit="m" value="675.0" valueMax="700.0" valueMin="650.0">650700m</quantity>
| 30.53.56N, 103.33.01E |
<date id="AC76A5EFFF8C6C42FE30F901FD95F94C" box="[429,558,1697,1724]" pageId="2" pageNumber="503" value="1997-05-18">18.05.1997</date>
, M. Schülke &lt;printed&gt;,
<taxonomicName id="1FC8F8ACFF8C6C42FCD8F901FC0FF94B" authorityName="Gravenhorst" authorityYear="1802" box="[837,948,1697,1723]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">Omalium</taxonomicName>
[handwritten] | spec. [handwritten] | det. M. Schülke 1999 &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke &lt;printed&gt; (
<collectionCode id="BED91BEAFF8C6C42FB78F965FA9AF92F" box="[1253,1313,1733,1759]" pageId="2" pageNumber="503">cSch</collectionCode>
)
</materialsCitation>
;
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:
<collectingRegion id="1A0C4DCDFF8C6C42FF0AF949FF47F8F3" box="[151,252,1769,1795]" country="China" name="Shaanxi" pageId="2" pageNumber="503">SHAANXI</collectingRegion>
:
</emphasis>
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(
<specimenCount id="CECE48A6FF8C6C42FEFCF949FE45F8F3" box="[353,510,1769,1795]" count="1" pageId="2" pageNumber="503" type="generic">one specimen</specimenCount>
dissected):
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: S-Shaanxi (Qinling Shan) | pass on rd. Zhouzhi—Foping, |
<quantity id="1F302ECAFF8C6C42FF0AF8ADFF56F8D7" box="[151,237,1805,1832]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="503" unit="km" value="105.0">105 km</quantity>
SW Xi`an, N-slope, |
<quantity id="1F302ECAFF8C6C42FE7AF8ADFD85F8D7" box="[487,574,1805,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="2" pageNumber="503" unit="m" value="1990.0">1990 m</quantity>
,
<geoCoordinate id="BDFCE5E8FF8C6C42FDD6F8ADFD15F8D7" box="[587,686,1805,1831]" degrees="33" direction="north" minutes="44" orientation="latitude" pageId="2" pageNumber="503" precision="925" value="33.733334">33°44´N</geoCoordinate>
,
<geoCoordinate id="BDFCE5E8FF8C6C42FD27F8ADFC9DF8D8" box="[698,806,1805,1832]" degrees="107" direction="east" minutes="59" orientation="longitude" pageId="2" pageNumber="503" precision="925" value="107.98333">107°59´E</geoCoordinate>
| leg. M. Schülke [C01-01] &lt;printed&gt;, 2./
<date id="AC76A5EFFF8C6C42FA96F8ADFA23F8D7" box="[1291,1432,1805,1831]" pageId="2" pageNumber="503" value="2001-07-04">4. VII. 2001</date>
, | small creek valley, mixed | deciduous forest, bamboo, | small meadows, dead wood, | mushrooms (sifted) [C01- 01] &lt;printed&gt;, Sammlung | M. Schülke | Berlin &lt;printed&gt; (
<collectionCode id="BED91BEAFF8C6C42FCD9F8F5FC3BF89F" box="[836,896,1877,1903]" pageId="2" pageNumber="503">cSch</collectionCode>
).
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<materialsCitation id="68A08972FF8C6C42FC0CF8F5FC71F864" pageId="2" pageNumber="503">
All
<typeStatus id="07733D8DFF8C6C42FC21F8F5FB9CF89F" box="[956,1063,1877,1903]" pageId="2" pageNumber="503" type="paratype">paratypes</typeStatus>
with additional red printed label:
<typeStatus id="07733D8DFF8C6C42FF06F8D9FE90F863" box="[155,299,1913,1939]" pageId="2" pageNumber="503" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="1FC8F8ACFF8C6C42FEA2F8D9FE1CF863" authorityName="Gravenhorst" authorityYear="1802" box="[319,423,1913,1939]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF8C6C42FEA2F8D9FE1CF863" box="[319,423,1913,1939]" italics="true" pageId="2" pageNumber="503">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="1FC8F8ACFF8C6C42FE26F8D9FDFFF863" authorityName="Shavrin" authorityYear="2025" box="[443,580,1913,1939]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="anguliferum" status="sp. nov.">
<emphasis id="EABC5F3DFF8C6C42FE26F8D9FDFFF863" box="[443,580,1913,1939]" italics="true" pageId="2" pageNumber="503">anguliferum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8C6C42FDD6F8D9FD1FF863" bold="true" box="[587,676,1913,1939]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="F18FE246FF8C6C42FDD6F8D9FD1FF863" box="[587,676,1913,1939]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025.
</materialsCitation>
</paragraph>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="8CB7D3A7FF8D6C43FF0AFBE9FEA1FB75" ID-DOI="http://doi.org/10.5281/zenodo.14896686" ID-Zenodo-Dep="14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="3" pageNumber="504" startId="3.[151,264,1097,1121]" targetBox="[151,1436,181,1073]" targetPageId="3" targetType="figure">
<paragraph id="D877832FFF8D6C43FF0AFBE9FEA1FB75" blockId="3.[151,1436,1097,1157]" pageId="3" pageNumber="504">
<emphasis id="EABC5F3DFF8D6C43FF0AFBE9FE81FB91" bold="true" box="[151,314,1097,1121]" pageId="3" pageNumber="504">FIGURES 13.</emphasis>
Habitus of
<taxonomicName id="1FC8F8ACFF8D6C43FE32FBE9FDB6FB91" authorityName="Gravenhorst" authorityYear="1802" box="[431,525,1097,1121]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF8D6C43FE32FBE9FDB6FB91" box="[431,525,1097,1121]" italics="true" pageId="3" pageNumber="504">Omalium</emphasis>
</taxonomicName>
: 1—
<taxonomicName id="1FC8F8ACFF8D6C43FDDDFBEAFD60FB91" authorityName="Shavrin" authorityYear="2025" box="[576,731,1097,1121]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="anguliferum">
<emphasis id="EABC5F3DFF8D6C43FDDDFBEAFD60FB91" box="[576,731,1097,1121]" italics="true" pageId="3" pageNumber="504">O. anguliferum</emphasis>
</taxonomicName>
(holotype). 2—
<taxonomicName id="1FC8F8ACFF8D6C43FCE1FBEAFC5DFB91" authorityName="Shavrin" authorityYear="2025" box="[892,998,1097,1121]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="rutilum">
<emphasis id="EABC5F3DFF8D6C43FCE1FBEAFC5DFB91" box="[892,998,1097,1121]" italics="true" pageId="3" pageNumber="504">O. rutilum</emphasis>
</taxonomicName>
(paratype), 3—
<taxonomicName id="1FC8F8ACFF8D6C43FB18FBEAFB51FB91" authorityName="Shavrin" authorityYear="2025" box="[1157,1258,1098,1121]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="sectum">
<emphasis id="EABC5F3DFF8D6C43FB18FBEAFB51FB91" box="[1157,1258,1098,1121]" italics="true" pageId="3" pageNumber="504">O. sectum</emphasis>
</taxonomicName>
(holotype). Scale bar: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="90D2D0A4FF8D6C44FF5AFB77FD25F985" lastPageId="4" lastPageNumber="505" pageId="3" pageNumber="504" type="description">
<paragraph id="D877832FFF8D6C43FF5AFB77FE31FAAD" blockId="3.[151,1437,1239,2022]" pageId="3" pageNumber="504">
<emphasis id="EABC5F3DFF8D6C43FF5AFB77FEE2FB01" bold="true" box="[199,345,1239,1265]" pageId="3" pageNumber="504">Description.</emphasis>
Measurements (n=24): HW: 0.510.57; HL: 0.340.41; OL: 0.120.17; TL: 0.070.10; AL (
<typeStatus id="07733D8DFF8D6C43FF02FB5BFEB3FAE5" box="[159,264,1275,1301]" pageId="3" pageNumber="504" type="holotype">holotype</typeStatus>
): 1.00; PL: 0.440.52; PWmax: 0.610.72; PWmin: 0.590.65; ESL: 0.760.92; EW: 0.840.97; MTbL (
<typeStatus id="07733D8DFF8D6C43FF02FABFFEB3FAC9" box="[159,264,1311,1337]" pageId="3" pageNumber="504" type="holotype">holotype</typeStatus>
): 0.42; MTrL (
<typeStatus id="07733D8DFF8D6C43FE2DFABFFDA2FAC9" box="[432,537,1311,1337]" pageId="3" pageNumber="504" type="holotype">holotype</typeStatus>
): 0.19 (MTrL 14: 0.07; MTrL 5: 0.12); AW: 0.840.90; AedL: 0.550.60; BL: 2.453.35 (
<typeStatus id="07733D8DFF8D6C43FE8BFAE3FEC4FAAD" box="[278,383,1347,1373]" pageId="3" pageNumber="504" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="D877832FFF8D6C43FF5AFAC7FC77F889" blockId="3.[151,1437,1239,2022]" pageId="3" pageNumber="504">
Habitus as in
<figureCitation id="40F39FAAFF8D6C43FEFDFAC7FE19FA72" box="[352,418,1383,1410]" captionStart="FIGURES 13" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="3" pageNumber="504">Fig. 1</figureCitation>
. Body brown to reddish-brown, with distinctly paler lateral and basal margins of pronotum and sometimes with slightly paler elytra and abdomen; antennomeres 511 or 611 brown; mouthparts, antennomeres 14 or 15 and legs yellow-brown; tarsi yellowish. Punctation of head dense, coarse and deep, sparser and finer in frontal and larger in middle portions, with interspaces between punctures in middle about as long as diameter of one nearest puncture or slightly shorter; neck with irregular, sparse and moderately fine punctation; punctation of pronotum regular, about as that in middle of head, distinctly finer and slightly sparser in mediobasal and significantly sparser in laterobasal portions; scutellum without punctures; punctation of elytra distinctly sparser, but larger and deeper than that on pronotum, coarser in medioapical part, finer and sparser along suture; abdominal tergites with fine and relatively dense punctation, indistinct on abdominal tergites VIVIII. Forebody glossy; anterior portion of clypeus with fine transverse microreticulation, supra-antennal elevations with fine and sometimes indistinct diagonal meshes; lateral sides of neck with diagonal sculpture; pronotum and elytra without microsculpture; scutellum without or with indistinct fine transverse sculpture; abdomen with dense microsculpture, somewhat transverse on abdominal tergites IVV and isodiametric on tergites VIVIII. Anterior part of head with several moderately long and erect setae; lateral margins of pronotum with several short setae; abdominal tergites with sparse and short setation; posterior margin of pronotum with row of short cuticular fringe.
</paragraph>
<paragraph id="D877832FFF8D6C44FF5AF823FE86FD3D" blockId="3.[151,1437,1239,2022]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="505" pageId="3" pageNumber="504">Head 1.31.5 times as broad as long, with broad clypeus and distinctly elevated supra-antennal elevations, with relatively narrow and deep anteriomedian depressions, almost reaching level of anterior margins or anterior third of eyes; posteriolateral margins of clypeus stretching posteriad toward level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus without semicircular notch. Dorsal surface with strong longitudinal and diagonal elevations between punctures in middle and on infraorbital portions; irregular elevations on postocular portions sometimes forming three to four longitudinal wrinkles.Anteocellar foveae moderately narrow and deep, distinctly convergent latero-apicad toward level of middle or anterior third of eyes. Temples long, 1.7 times as long as longitudinal length of eye, from posterior margins of eyes slightly narrowed posteriad, each with subacute angle distinctly protruded basad; postocular ridges missing. Apical part of neck narrowly and moderately deeply depressed, with several short and narrow elevations; each lateral margin with acute small projections, slightly protruded laterad. Eyes large and convex. Ocelli large, located significantly behind level of posterior margins of eyes; distance between ocelli short, 1.61.8 times as long as distance between ocellus and posterior margin of eye. Maxillary palpi long, apical palpomere about four times as long as small and transverse penultimate segment, from widest basal portion gradually narrowed toward acute apex. Antenna reaching basal portion of elytra when reclined, with slightly elongate antennomeres 46 and distinctly transverse antennomeres 710; antennomere 3 slightly shorter and distinctly narrower than 2, 4 1.4 times as long as and narrower than 3, 5 distinctly longer than 4, 6 slightly longer and broader than 5, 7 slightly broader than 6, 8 distinctly broader than 7, 910 slightly broader than 8, apical antennomere 1.31.4 times as long as preceding segment, from middle gradually narrowed toward subacute apex.</paragraph>
<paragraph id="D877832FFF8A6C44FF5AFD77FCDFFC1D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="505">
Pronotum convex, 1.3 times as broad as long, 1.11.2 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad. Apical angles slightly protruded anteriad. Anterior margin widely rounded, slightly protruded anteriad. Laterobasal portions sometimes slightly concave in front of obtuse hind angles. Lateral portions relatively deeply and narrowly impressed, deeper in middle. Lateral margins narrowly marginated, sometimes indistinctly crenulate in latero-apical part.Surface of disc with two long and wide longitudinal depressions, broadened basad; some
<typeStatus id="07733D8DFF8A6C44FE3AFC2BFDA9FC55" box="[423,530,907,933]" pageId="4" pageNumber="505" type="paratype">paratypes</typeStatus>
with indistinct oval depression in medioapical portion; each laterobasal portion widely and deeply depressed; surface between all pronotal depressions distinctly elevated. Middle portion with irregular longitudinal or diagonal elevations between punctures.
</paragraph>
<paragraph id="D877832FFF8A6C44FF5AFC57FC0AFB8D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="505">Elytra slightly broader than long, 1.7 times as long as pronotum, from basal portion slightly broadened posteriad; lateral margins marginated; lateral portions narrowly impressed; hind margins somewhat straight or widely rounded. Dorsal surface of each elytron with strong irregular elevations between punctures: transverse and subdiagonal in mediobasal and longitudinal in apical portions. Wings fully developed.</paragraph>
<paragraph id="D877832FFF8A6C44FF5AFB27FCF1FB51" blockId="4.[151,1437,151,2013]" box="[199,842,1159,1185]" pageId="4" pageNumber="505">Metatarsi slightly more than twice shorter than metatibia.</paragraph>
<paragraph id="D877832FFF8A6C44FF5AFB0BFC3EFAFD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="505">Abdomen convex, about as wide as elytra, with two small oval wing-folding patches in middle of abdominal tergite IV and distinct narrow palisade fringe on apical margin of abdominal tergite VII, sometimes with narrow to wide intersegmental membranes between abdominal tergites IIIV.</paragraph>
<paragraph id="D877832FFF8A6C44FF5AFAB7FC92F9F9" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="505">
Male. Posterior margin of abdominal tergite VIII slightly and sternite VIII widely and moderately deeply sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe wide, with deeply sinuate lateral parts in preapical portion and widely rounded apex; mediolateral portions with narrow and relatively short accessory plates rounded apically; parameres moderately narrow, distinctly broadened apically, reaching preapical part of median lobe, each with four moderately long apical setae; internal sac wide and relatively short, with two small oval structures in basal part (
<figureCitation id="40F39FAAFF8A6C44FD81FA6BFDDAFA15" box="[540,609,1483,1509]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="4" pageNumber="505">Fig. 4</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="40F39FAAFF8A6C44FB98FA6BFBF3FA15" box="[1029,1096,1482,1509]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="4" pageNumber="505">Fig. 5</figureCitation>
; apical portion of median lobe (lateral view) with three ventrolateral obtuse teeth (
<figureCitation id="40F39FAAFF8A6C44FD4BFA4FFCA0F9F9" box="[726,795,1519,1545]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="4" pageNumber="505">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="D877832FFF8A6C44FF5AF9B3FD25F985" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="505">
Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII truncate or rounded. Accessory sclerite with moderately narrow basal portion and relatively elongate median part, rounded apically (
<figureCitation id="40F39FAAFF8A6C44FF62F9FBFEFFF985" box="[255,324,1627,1653]" captionStart="FIGURES 412" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 412. Aedeagus of Omalium (Figs 46—O. anguliferum (paratype), Figs 79—O. rutilum (paratype), 1012—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="4" pageNumber="505">Fig. 9</figureCitation>
). Spermatheca not recognized.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8A6C45FF5AF9DEFEF6FF09" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="discussion">
<paragraph id="D877832FFF8A6C45FF5AF9DEFEF6FF09" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,501]" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<emphasis id="EABC5F3DFF8A6C44FF5AF9DEFE0BF969" bold="true" box="[199,432,1662,1689]" pageId="4" pageNumber="505">Comparative notes.</emphasis>
Based on the shape of temples,
<taxonomicName id="1FC8F8ACFF8A6C44FC88F9DFFC7BF969" authorityName="Shavrin" authorityYear="2025" box="[789,960,1663,1689]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anguliferum" status="sp. nov.">
<emphasis id="EABC5F3DFF8A6C44FC88F9DFFC7BF969" box="[789,960,1663,1689]" italics="true" pageId="4" pageNumber="505">O. anguliferum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8A6C44FC5BF9DFFBA4F969" bold="true" box="[966,1055,1663,1689]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="F18FE246FF8A6C44FC5BF9DFFBA4F969" box="[966,1055,1663,1689]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belong to the acutangulum group, recently established for two species (
<taxonomicName id="1FC8F8ACFF8A6C44FDA6F903FC2DF94D" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[571,918,1699,1725]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="acutangulum">
<emphasis id="EABC5F3DFF8A6C44FDA6F903FD4BF94D" box="[571,752,1699,1725]" italics="true" pageId="4" pageNumber="505">O. acutangulum</emphasis>
Shavrin, 2023
</taxonomicName>
and
<taxonomicName id="1FC8F8ACFF8A6C44FC52F903FAB7F94D" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[975,1292,1699,1725]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="amicorum">
<emphasis id="EABC5F3DFF8A6C44FC52F903FBDFF94C" box="[975,1124,1699,1725]" italics="true" pageId="4" pageNumber="505">O. amicorum</emphasis>
Shavrin, 2023
</taxonomicName>
) from
<collectingCountry id="A0DFC3BFFF8A6C44FAC4F903FA27F94D" box="[1369,1436,1699,1725]" name="Nepal" pageId="4" pageNumber="505">Nepal</collectingCountry>
(
<bibRefCitation id="BC59FEDEFF8A6C44FF02F967FEF7F911" author="Shavrin, A. V." box="[159,332,1735,1761]" pageId="4" pageNumber="505" pagination="151 - 206" refId="ref30642" refString="Shavrin, A. V. (2023 a) A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region. Zootaxa, 5369 (2), 151-206. https://doi.org/10.11646/zootaxa.5369.2.1" type="journal article" year="2023">Shavrin 2023a</bibRefCitation>
). Besides that, it related to Chinese
<taxonomicName id="1FC8F8ACFF8A6C44FD6BF967FCD5F911" authorityName="Shavrin" authorityYear="2025" box="[758,878,1735,1761]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF8A6C44FD6BF967FCD5F911" box="[758,878,1735,1761]" italics="true" pageId="4" pageNumber="505">O. rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8A6C44FCE4F967FC6DF911" bold="true" box="[889,982,1735,1761]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="F18FE246FF8A6C44FCE4F967FC6DF911" box="[889,982,1735,1761]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1FC8F8ACFF8A6C44FB89F967FB33F910" authorityName="Shavrin" authorityYear="2025" box="[1044,1160,1735,1761]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF8A6C44FB89F967FB33F910" box="[1044,1160,1735,1761]" italics="true" pageId="4" pageNumber="505">O. sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8A6C44FB0EF967FB4BF911" bold="true" box="[1171,1264,1735,1761]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="F18FE246FF8A6C44FB0EF967FB4BF911" box="[1171,1264,1735,1761]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Regarding the shape of postocular portions of head slightly narrowed posteriad and forming distinct hind angles, the shape of the pronotum with apical angles slightly protruded apicad,
<taxonomicName id="1FC8F8ACFF8A6C44FC98F8AFFC0AF8D9" authorityName="Shavrin" authorityYear="2025" box="[773,945,1807,1833]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anguliferum" status="sp. nov.">
<emphasis id="EABC5F3DFF8A6C44FC98F8AFFC0AF8D9" box="[773,945,1807,1833]" italics="true" pageId="4" pageNumber="505">O. anguliferum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8A6C44FC25F8AFFBA9F8D9" bold="true" box="[952,1042,1807,1833]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="F18FE246FF8A6C44FC25F8AFFBA9F8D9" box="[952,1042,1807,1833]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is more similar to
<taxonomicName id="1FC8F8ACFF8A6C44FB7AF8AFFA27F8D9" authorityName="Shavrin" authorityYear="2023" box="[1255,1436,1807,1833]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="acutangulum">
<emphasis id="EABC5F3DFF8A6C44FB7AF8AFFA27F8D9" box="[1255,1436,1807,1833]" italics="true" pageId="4" pageNumber="505">O. acutangulum</emphasis>
</taxonomicName>
and
<taxonomicName id="1FC8F8ACFF8A6C44FF5AF893FE87F8BD" authorityName="Shavrin" authorityYear="2025" box="[199,316,1843,1869]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF8A6C44FF5AF893FF60F8BD" box="[199,219,1843,1869]" italics="true" pageId="4" pageNumber="505">O</emphasis>
.
<emphasis id="EABC5F3DFF8A6C44FF77F893FE87F8BD" box="[234,316,1843,1869]" italics="true" pageId="4" pageNumber="505">rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8A6C44FED8F893FE24F8BD" bold="true" box="[325,415,1843,1869]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="F18FE246FF8A6C44FED8F893FE24F8BD" box="[325,415,1843,1869]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It can be distinguished from
<taxonomicName id="1FC8F8ACFF8A6C44FD71F893FC1AF8BD" authorityName="Shavrin" authorityYear="2023" box="[748,929,1843,1869]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="acutangulum">
<emphasis id="EABC5F3DFF8A6C44FD71F893FC1AF8BD" box="[748,929,1843,1869]" italics="true" pageId="4" pageNumber="505">O. acutangulum</emphasis>
</taxonomicName>
by the finer elevations between punctures in basal part of the head, and slightly narrower pronotum with finer and smaller depressions on the pronotal disc. From
<taxonomicName id="1FC8F8ACFF8A6C44FF0AF8DBFEB1F865" authorityName="Shavrin" authorityYear="2025" box="[151,266,1915,1941]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF8A6C44FF0AF8DBFF10F865" box="[151,171,1915,1941]" italics="true" pageId="4" pageNumber="505">O</emphasis>
.
<emphasis id="EABC5F3DFF8A6C44FF25F8DBFEB1F865" box="[184,266,1915,1941]" italics="true" pageId="4" pageNumber="505">rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8A6C44FE8CF8DBFED1F865" bold="true" box="[273,362,1915,1941]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="F18FE246FF8A6C44FE8CF8DBFED1F865" box="[273,362,1915,1941]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it differ by the shape of transverse antennomere 8 and the shape of narrower pronotum, widest in middle and with less pronounced dorsal impressions. From both species it can be distinguished by the darker body, significantly denser and coarser punctation of the pronotum, different shape of the broader median lobe and other details of internal and external morphology of the aedeagus, and different shape of the female accessory sclerite. From
<taxonomicName id="1FC8F8ACFF8B6C45FF46FF1BFEF1FF24" authorityName="Shavrin" authorityYear="2025" box="[219,330,187,213]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF8B6C45FF46FF1BFEF1FF24" box="[219,330,187,213]" italics="true" pageId="5" pageNumber="506">O. sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8B6C45FECCFF1BFE11FF25" bold="true" box="[337,426,187,213]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="F18FE246FF8B6C45FECCFF1BFE11FF25" box="[337,426,187,213]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it differs by the widely rounded anterior angles of the pronotum and different morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8B6C45FF5AFEA2FB79FEB1" pageId="5" pageNumber="506" type="distribution">
<paragraph id="D877832FFF8B6C45FF5AFEA2FB79FEB1" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="506">
<emphasis id="EABC5F3DFF8B6C45FF5AFEA2FEDAFEEC" bold="true" box="[199,353,258,284]" pageId="5" pageNumber="506">Distribution.</emphasis>
<taxonomicName id="1FC8F8ACFF8B6C45FEFAFEA3FDDCFEED" authorityName="Shavrin" authorityYear="2025" box="[359,615,259,285]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. nov.">
<emphasis id="EABC5F3DFF8B6C45FEFAFEA3FDDCFEED" box="[359,615,259,285]" italics="true" pageId="5" pageNumber="506">Omalium acutangulum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF8B6C45FDF3FEA3FD7CFEED" bold="true" box="[622,711,259,285]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="F18FE246FF8B6C45FDF3FEA3FD7CFEED" box="[622,711,259,285]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from several localities in Qinling Shan mountain range in
<collectingRegion id="1A0C4DCDFF8B6C45FF29FE87FEB5FEB1" box="[180,270,295,321]" country="China" name="Shaanxi" pageId="5" pageNumber="506">Shaanxi</collectingRegion>
and in Emeishan and Qingcheng-Shan mountain ranges in
<collectingRegion id="1A0C4DCDFF8B6C45FC3AFE87FBBFFEB1" box="[935,1028,295,321]" country="China" name="Sichuan" pageId="5" pageNumber="506">Sichuan</collectingRegion>
,
<collectingCountry id="A0DFC3BFFF8B6C45FB92FE87FBE9FEB1" box="[1039,1106,295,321]" name="China" pageId="5" pageNumber="506">China</collectingCountry>
(
<figureCitation id="40F39FAAFF8B6C45FBFFFE87FB0DFEB1" box="[1122,1206,295,321]" captionStart="FIGURE 30" captionStartId="9.[151,250,1849,1873]" captionTargetBox="[151,1435,993,1826]" captionTargetId="figure-318@9.[151,1435,993,1826]" captionTargetPageId="9" captionText="FIGURE 30. Distribution of Omalium in China: O. anguliferum (circles), O. rutilum (squares), O. sectum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896690" httpUri="https://zenodo.org/record/14896690/files/figure.png" pageId="5" pageNumber="506">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8B6C45FF5AFEEBFD98FE79" pageId="5" pageNumber="506" type="etymology">
<paragraph id="D877832FFF8B6C45FF5AFEEBFD98FE79" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="506">
<emphasis id="EABC5F3DFF8B6C45FF5AFEEBFEF5FE95" bold="true" box="[199,334,331,357]" pageId="5" pageNumber="506">Etymology.</emphasis>
The specific epithet (Latin adjective: that carries angles) refers to the shape of temples strongly narrowed and protruding posteriad.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF8B6C45FF5AFE32FEBEFE05" pageId="5" pageNumber="506" type="biology_ecology">
<paragraph id="D877832FFF8B6C45FF5AFE32FEBEFE05" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="506">
<emphasis id="EABC5F3DFF8B6C45FF5AFE32FEF0FE5C" bold="true" box="[199,331,402,428]" pageId="5" pageNumber="506">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="1F302ECAFF8B6C45FCA3FE32FC60FE5D" box="[830,987,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.565" metricValueMax="2.48" metricValueMin="0.65" pageId="5" pageNumber="506" unit="m" value="1565.0" valueMax="2480.0" valueMin="650.0">650 to 2480 m</quantity>
a.s.l. and were sifted from litter of mixed coniferous (
<taxonomicName id="1FC8F8ACFF8B6C45FEBDFE17FEEEFE21" box="[288,341,439,465]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="506" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF8B6C45FEBDFE17FEEEFE21" box="[288,341,439,465]" italics="true" pageId="5" pageNumber="506">Acer</emphasis>
</taxonomicName>
,
<taxonomicName id="1FC8F8ACFF8B6C45FEFBFE17FE4EFE20" authority=", Picea" authorityName="Picea" box="[358,501,439,465]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="506" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF8B6C45FEFBFE17FE1FFE21" box="[358,420,439,465]" italics="true" pageId="5" pageNumber="506">Abies</emphasis>
,
<emphasis id="EABC5F3DFF8B6C45FE28FE17FE4EFE20" box="[437,501,439,464]" italics="true" pageId="5" pageNumber="506">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="1FC8F8ACFF8B6C45FD98FE17FD11FE21" box="[517,682,439,465]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="506" phylum="Tracheophyta" rank="genus">
<emphasis id="EABC5F3DFF8B6C45FD98FE17FD11FE21" box="[517,682,439,465]" italics="true" pageId="5" pageNumber="506">Rhododendron</emphasis>
</taxonomicName>
), and deciduous forest with bamboo, dead wood and mushrooms (C01-01).
</paragraph>
</subSubSection>
</treatment>
</document>