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<paragraph id="682DD83E89256DE5BA570F34AE7535DB" pageId="12" pageNumber="143">Taxon classification Animalia Hygrophila Lymnaeidae</paragraph>
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<taxonomicName id="9F16C8EAF904318656ADB4D62600873A" authority="&quot; Ziegler &quot;" class="Gastropoda" family="Lymnaeidae" genus="Limnaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limnaeus opacus" pageId="12" pageNumber="143" phylum="Mollusca" rank="species" species="opacus">
opacus
<normalizedToken id="B9990D86CE5143AE05490A0EDF905B98" originalValue="“Ziegler”">&quot;Ziegler&quot;</normalizedToken>
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Fig. 62
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<paragraph id="4A3F4F2E463D51129EBAB96487575276" pageId="12" pageNumber="143">
<taxonomicName id="D490275C0317B3D9D8E934058D732ACC" class="Gastropoda" family="Lymnaeidae" genus="Limnaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limnaea opaca" pageId="12" pageNumber="143" phylum="Mollusca" rank="species" species="opaca">Limnaea opaca</taxonomicName>
<bibRefCitation id="DE9356A0DD1EA886B92D6DA02AC9D62B" author="Dupuy, D" journalOrPublisher="V. Masson, Paris" pageId="13" pageNumber="144" pagination="459 - 594" title="Histoire naturelle des mollusques terrestres et d'eau douce qui vivent en France" url="http://biodiversitylibrary.org/page/32249746" year="1851">Dupuy 1851</bibRefCitation>
: 473.
</paragraph>
<paragraph id="54824A5ABF9D97666F8ABCA6B2665259" pageId="12" pageNumber="143">
<taxonomicName id="AAD190A74B0C6032B08C9F59CDF04596" class="Gastropoda" family="Lymnaeidae" genus="Limnaeus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Limnaeus pereger var. opacus" pageId="12" pageNumber="143" phylum="Mollusca" rank="variety" species="pereger" variety="opacus">Limnaeus pereger var. opacus</taxonomicName>
von
<bibRefCitation id="18F439B3312875DDBAB91BB847B6BAC5" author="Gallenstein, M von" journalOrPublisher="Klagenfurt" pageId="13" pageNumber="144" title="Kaerntern's Land- und Suesswasser-Conchylien" url="10.5962/bhl.title.10673" year="1852">Gallenstein 1852</bibRefCitation>
: 43.
</paragraph>
<paragraph id="5B41C0250DCBDA4E5893ACDA5FA8AE60" pageId="12" pageNumber="143">
<taxonomicName id="F0CBD31F0409D078A0AC08FE98B69B84" class="Gastropoda" family="Lymnaeidae" genus="Limnaea" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Limnaea peregra var. opaca" pageId="12" pageNumber="143" phylum="Mollusca" rank="variety" species="peregra" variety="opaca">Limnaea peregra var. opaca</taxonomicName>
<bibRefCitation id="7DFB8EBC4E36596E1D521E92B717FA97" author="Moquin-Tandon, A" journalOrPublisher="J. - B. Bailliere, Paris, 2" pageId="14" pageNumber="145" title="Histoire naturelle des mollusques terrestres et fluviatiles de France contenant des etudes generales sur leur anatomie et leur physiologie et la description particuliere des genres, les especes et des varietes" url="10.5962/bhl.title.13098" year="1855">Moquin-Tandon 1855</bibRefCitation>
: 468.
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<paragraph id="42B0A8F9CF7E1CB82D59F34335D4F84B" pageId="12" pageNumber="143">Material.</paragraph>
<paragraph id="B3698A24BA5974111FDA2E881E67D0DF" pageId="12" pageNumber="143">Two shells collected in Carniolia and kept in ZMB under accession number 109755. The largest of two shells is 10.0 mm height.</paragraph>
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<paragraph id="1FDFE8BEAD465C92DA55EC7E2D52DBD5" pageId="12" pageNumber="143">Current taxonomic allocation.</paragraph>
<paragraph id="AE924C96664691B7F2D6E5A7B11A2769" pageId="12" pageNumber="143">
Possibly a synonym of
<taxonomicName id="728EB30573C719553708E0614E345E3D" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix (Peregriana) peregra" pageId="12" pageNumber="143" phylum="Mollusca" rank="species" species="peregra" subGenus="Peregriana">Radix (Peregriana) peregra</taxonomicName>
auct. =
<taxonomicName id="6B9EB793DCF0C0FC6001472BA8FA5987" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix labiata" pageId="12" pageNumber="143" phylum="Mollusca" rank="species" species="labiata">Radix labiata</taxonomicName>
sensu
<bibRefCitation id="A2B1FD02110F3DA9DC6C6B5BFA10E2D4" author="Falkner, G" journalOrPublisher="Collection Patrimoines Naturels" pageId="13" pageNumber="144" pagination="1 - 350" title="Mollusques continentaux de France. Liste de reference annotee et bibliographie." volume="52" year="2002">Falkner et al. 2002</bibRefCitation>
.
</paragraph>
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<subSubSection id="45EB66987B0DB9F9ADB051AEA6B874C0" pageId="12" pageNumber="143" type="remarks">
<paragraph id="B0189C43C1D655E6675F2DE5306E4DEB" pageId="12" pageNumber="143">Remark.</paragraph>
<paragraph id="8E6740E2D2B42D1BC3036F804116C167" pageId="12" pageNumber="143">
Though a few authors of the 19th century used this taxon name (
<bibRefCitation id="A3D177C31A1A4BC53CF0E3097FEFB619" author="Dupuy, D" journalOrPublisher="V. Masson, Paris" pageId="13" pageNumber="144" pagination="459 - 594" title="Histoire naturelle des mollusques terrestres et d'eau douce qui vivent en France" url="http://biodiversitylibrary.org/page/32249746" year="1851">Dupuy 1851</bibRefCitation>
; von
<bibRefCitation id="961E502AF6E157DA3C0197E31122E4ED" author="Gallenstein, M von" journalOrPublisher="Klagenfurt" pageId="13" pageNumber="144" title="Kaerntern's Land- und Suesswasser-Conchylien" url="10.5962/bhl.title.10673" year="1852">Gallenstein 1852</bibRefCitation>
;
<bibRefCitation id="4DA764B3B7785775EE91207969D84293" author="Moquin-Tandon, A" journalOrPublisher="J. - B. Bailliere, Paris, 2" pageId="14" pageNumber="145" title="Histoire naturelle des mollusques terrestres et fluviatiles de France contenant des etudes generales sur leur anatomie et leur physiologie et la description particuliere des genres, les especes et des varietes" url="10.5962/bhl.title.13098" year="1855">Moquin-Tandon 1855</bibRefCitation>
), I could not trace the source where it was described originally. Perhaps, it should be considered as one of those numerous &quot;manuscript names&quot; attributed to the authorship of Ziegler (see also comment to
<taxonomicName id="FCC04B7E8608014191A469AF7DB2A1DD" class="Gastropoda" family="Lymnaeidae" genus="Limnaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limnaeus compactus" pageId="12" pageNumber="143" phylum="Mollusca" rank="species" species="compactus">Limnaeus compactus</taxonomicName>
above).
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.954.54694" ID-GBIF-Dataset="9e9861de-a849-409a-bd85-108cc8c0c894" ID-PMC="PMC7406547" ID-Pensoft-Pub="1313-2970-954-1" ID-Pensoft-UUID="FDC60FB2C83C528A8A3B21F4C6178613" ID-PubMed="32821201" ID-ZooBank="3879B114B4034407B7B0ED5B8B826CC6" ModsDocID="1313-2970-954-1" checkinTime="1596086649365" checkinUser="pensoft" docAuthor="Liu, Weixin &amp; Golovatch, Sergei" docDate="2020" docId="16A678F3F4915D5E91AD7BEC646B7C22" docLanguage="en" docName="ZooKeys 954: 1-15" docOrigin="ZooKeys 954" docSource="http://dx.doi.org/10.3897/zookeys.954.54694" docTitle="Tonkinomeris huzhengkuni Liu &amp; Golovatch 2020, sp. nov." docType="treatment" docUuid="41BF7A61-D9E7-4B81-9685-4D5AB8B7C071" docUuidSource="ZooBank" docVersion="4" id="FDC60FB2C83C528A8A3B21F4C6178613" lastPageNumber="1" masterDocId="FDC60FB2C83C528A8A3B21F4C6178613" masterDocTitle="The first representatives of the millipede family Glomeridellidae (Diplopoda, Glomerida) recorded from China and Indochina" masterLastPageNumber="15" masterPageNumber="1" pageNumber="1" updateTime="1668169177999" updateUser="ExternalLinkService">
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<mods:title>The first representatives of the millipede family Glomeridellidae (Diplopoda, Glomerida) recorded from China and Indochina</mods:title>
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<mods:namePart>Liu, Weixin</mods:namePart>
<mods:affiliation>Department of Entomology, College of Agriculture, South China Agricultural University, 483 Wushanlu, Guangzhou 510642, China</mods:affiliation>
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<mods:namePart>Golovatch, Sergei</mods:namePart>
<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<taxonomicName LSID="16A678F3-F491-5D5E-91AD-7BEC646B7C22" authority="Liu &amp; Golovatch, 2020" authorityName="Liu &amp; Golovatch" authorityYear="2020" class="Diplopoda" family="Glomeridellidae" genus="Tonkinomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinomeris huzhengkuni" order="Glomerida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="huzhengkuni" status="sp. nov.">Tonkinomeris huzhengkuni</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tonkinomeris huzhengkuni sp. nov., ♀ paratype. A Gnathochilarium, ventral view, in situ B, C left mandible, mesal and subfrontal views, respectively D coxae 2 with vulvae. Abbreviations: ca: cardines of gnathochilarium, et: external tooth, gu: gula, iA: intermediate area, ip: inner palpus, it: inner tooth, ll: lamellae linguales, lp: lateral palpus, m: mentum, md: basal joint of mandible, mf: membranous fringe, mg: molar groove, mp: molar plate, pl: pectinate lamellae, st: stipites, vu: vulvae." figureDoi="10.3897/zookeys.954.54694.figure3" httpUri="https://binary.pensoft.net/fig/437444" pageId="0" pageNumber="1">, 3</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂ (SCAU TY01), China, Guizhou Province, Tongren City, Jiangkou County, Baishuidong Scenic Area,
<geoCoordinate degrees="27.652873" direction="north" orientation="latitude" precision="1" value="27.652872">27.652873N</geoCoordinate>
,
<geoCoordinate degrees="108.795223" direction="east" orientation="longitude" precision="1" value="108.79522">108.795223E</geoCoordinate>
, 450 m a.s.l., 25.XI.2019, Zhengkun Hu leg.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
: 1 ♀ (SCAU TY02), same data as for holotype.
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="name">
<paragraph pageId="0" pageNumber="1">Name.</paragraph>
<paragraph pageId="0" pageNumber="1">Honours Mr Huzhengkun, the collector and a millipede fan. A noun in genitive case.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Differs from
<taxonomicName lsidName="T. napoensis" pageId="0" pageNumber="1" rank="species" species="napoensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. napoensis</emphasis>
</taxonomicName>
Nguyen, Sierwald &amp; Marek, 2019, the sole other species of the genus (
<bibRefCitation author="Nguyen, AD" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="260 - 297" publicationUrl="https://lkcnhm.nus.edu.sg/app/uploads/2018/11/RBZ-2019-0020.pdf" refId="B15" refString="Nguyen, AD, Sierwald, P, Marek, PE, 2019. The pill millipedes of Vietnam: a key to genera and descriptions of five new species (Diplopoda: Glomerida: Glomeridae). Raffles Bulletin of Zoology 67: 260 - 297, https://lkcnhm.nus.edu.sg/app/uploads/2018/11/RBZ-2019-0020.pdf" title="The pill millipedes of Vietnam: a key to genera and descriptions of five new species (Diplopoda: Glomerida: Glomeridae)." url="https://lkcnhm.nus.edu.sg/app/uploads/2018/11/RBZ-2019-0020.pdf" volume="67" year="2019">Nguyen et al. 2019</bibRefCitation>
), by the larger body size (&gt; 11 mm vs 9.6 mm), the more numerous ommatidia (at least 6+1 vs 5+1), the glabrous, but caudomedially produced posterior margins of ♂ tergites 8-11 (vs unmodified in both sexes), the vivid, peculiar, sexually dimorphic colour pattern (vs even more vivid, but the same in both sexes), and some structural details of the telopods: the much higher central lobe and the much shorter horns of ♂ syncoxite 19 (vs the opposite), the shape and armament of both telopod pairs, the less strongly reduced, 4-segmented ♂ telopodites 17 (vs 2-segmented), etc. (see Discussion below).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Body length of holotype ♂ ca 12.5 mm, width of thoracic shield ca 8.0 mm (broadest), height of thoracic shield ca 4.2 mm (highest). Body length of paratype ♀, 11.2 mm, width of thoracic shield ca 7.1 mm (broadest), height of thoracic shield ca 4.0 mm (highest). Coloration (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1</figureCitation>
): body rather uniformly blackish, but with slightly yellowish edges. Dorsal pattern marbled yellow-brownish: collum with a small, rounded, central spot anteriorly and a paramedian pair of larger, transversely ovoid, lateral spots (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1D</figureCitation>
); thoracic shield and tergites 3-11 each with a similar, but much larger pair of lateral spots and a yellowish, but slightly purplish, median, narrow, axial stripe. Anal shield (= pygidium) of ♂ without stripe (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1F</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2D</figureCitation>
), but ♀ with a distinct, yellowish, median, triangular spot near caudal margin (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1C</figureCitation>
). Head largely dark brown, only labrum and
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
organ lighter, grey-yellowish. Antennomeres 3-6 brownish, slightly purplish, remaining antennomeres light yellow-brown. Venter and podomeres 1 and 2 entirely light grey-yellowish, remaining podomeres dark brown, slightly purplish (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1B, D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
Habitus of
<taxonomicName authorityName="Liu &amp; Golovatch" authorityYear="2020" class="Diplopoda" family="Glomeridellidae" genus="Tonkinomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinomeris huzhengkuni" order="Glomerida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="huzhengkuni">
<emphasis italics="true" pageId="0" pageNumber="1">Tonkinomeris huzhengkuni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Live coloration of ♂ holotype
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
body of ♀ paratype, ventral and dorsal views, respectively
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
body of ♂ holotype, ventral, lateral and dorsal views, respectively.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: mandibles (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tonkinomeris huzhengkuni sp. nov., ♀ paratype. A Gnathochilarium, ventral view, in situ B, C left mandible, mesal and subfrontal views, respectively D coxae 2 with vulvae. Abbreviations: ca: cardines of gnathochilarium, et: external tooth, gu: gula, iA: intermediate area, ip: inner palpus, it: inner tooth, ll: lamellae linguales, lp: lateral palpus, m: mentum, md: basal joint of mandible, mf: membranous fringe, mg: molar groove, mp: molar plate, pl: pectinate lamellae, st: stipites, vu: vulvae." figureDoi="10.3897/zookeys.954.54694.figure3" httpUri="https://binary.pensoft.net/fig/437444" pageId="0" pageNumber="1">3B, C</figureCitation>
) equal in both sexes, each with a large external tooth and a smaller internal tooth, the latter with four cusps. Molar plate with a long membranous fringe and a groove. At least seven rows of pectinate lamellae and a scaly intermediate area. Gnathochilarium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tonkinomeris huzhengkuni sp. nov., ♀ paratype. A Gnathochilarium, ventral view, in situ B, C left mandible, mesal and subfrontal views, respectively D coxae 2 with vulvae. Abbreviations: ca: cardines of gnathochilarium, et: external tooth, gu: gula, iA: intermediate area, ip: inner palpus, it: inner tooth, ll: lamellae linguales, lp: lateral palpus, m: mentum, md: basal joint of mandible, mf: membranous fringe, mg: molar groove, mp: molar plate, pl: pectinate lamellae, st: stipites, vu: vulvae." figureDoi="10.3897/zookeys.954.54694.figure3" httpUri="https://binary.pensoft.net/fig/437444" pageId="0" pageNumber="1">3A</figureCitation>
) equal in both sexes, unmodified, typical of
<taxonomicName authorityName="Nguyen, Sierwald &amp; Marek" authorityYear="2019" lsidName="" order="Glomerida" pageId="0" pageNumber="1" rank="order">Glomerida</taxonomicName>
. Left and right eyes asymmetrical, with 9+1/10+1 (♂) (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2A</figureCitation>
) or 8+1/6+1 (♀) (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1B</figureCitation>
) ommatidia.
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
organ transversely oval, ca 1.6 times wider than long (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2A</figureCitation>
). Lengths of antennomeres: 6&gt;&gt;3&gt;4=5&gt;1=2&gt;7 (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1B, D</figureCitation>
). Antennomere 6 ca 2.6 times as long as high. Antennomere 8 with four small, apical, sensory cones.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Collum</emphasis>
with two complete transverse striae (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1D</figureCitation>
). Thoracic shield with a narrow hyposchism extending past caudal tergal margin; about 12 or 13 superficial transverse striae laterally and dorsolaterally, but five or six confusedly arranged and incomplete. One or two starting below, one in front of schism, all others above schism, with three crossing the dorsum; mid-dorsal region with five additional, incomplete, broken, confused, mostly short striolae behind last regular stria (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1E</figureCitation>
). Tergites: surface smooth and shining, only paratergites with three or four short, incomplete, and superficial striae (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1E</figureCitation>
). Tergite 9 in ♂ drawn posteriad into a small, triangular, glabrous (non-dentate), median lobe (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2D</figureCitation>
), this being very week also in tergites 8, 10, and 11. Pygidium in both sexes uneven medially at caudal margin; in ♂ clearly impressed and concave centro-dorsally and with two very small, paramedian, flattened and rounded knobs (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1D-F</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2D</figureCitation>
), in ♀ only slightly flattened dorsocaudally (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Tonkinomeris huzhengkuni sp. nov. A Live coloration of ♂ holotype B, C body of ♀ paratype, ventral and dorsal views, respectively D-F body of ♂ holotype, ventral, lateral and dorsal views, respectively." figureDoi="10.3897/zookeys.954.54694.figure1" httpUri="https://binary.pensoft.net/fig/437442" pageId="0" pageNumber="1">1C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName authorityName="Liu &amp; Golovatch" authorityYear="2020" class="Diplopoda" family="Glomeridellidae" genus="Tonkinomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinomeris huzhengkuni" order="Glomerida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="huzhengkuni">
<emphasis italics="true" pageId="0" pageNumber="1">Tonkinomeris huzhengkuni</emphasis>
</taxonomicName>
sp. nov., ♂ holotype.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Right side of head
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
left leg 1
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
right leg 2 with gonopore
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
posterior part of body, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
left leg 9. Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="1">cx</emphasis>
: coxa,
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
: femur,
<emphasis bold="true" pageId="0" pageNumber="1">go</emphasis>
: gonopore,
<emphasis bold="true" pageId="0" pageNumber="1">O</emphasis>
: ommatidia,
<emphasis bold="true" pageId="0" pageNumber="1">ta</emphasis>
: tarsus,
<emphasis bold="true" pageId="0" pageNumber="1">To</emphasis>
:
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
organ,
<emphasis bold="true" pageId="0" pageNumber="1">8-11</emphasis>
refer to tergite numbers.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
long and slender. All podomeres densely setose, setae mostly being short. Coxae 1-16 each with a short, well-rounded, spinigerous, apico-mesal projection, this being especially evident in coxae 1 and 2 (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2B, C</figureCitation>
). Coxae 4-21 each with a similar apico-lateral process. Tarsi 1-16 each with two irregular transverse rows of 7-8+7-8 dorsal spines and 9-12+9-12 ventral spines (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2B, C, E</figureCitation>
). Femur 9, 2.4 times, tarsus 6.5 times longer than wide (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male sexual characters</emphasis>
: gonopore small, oval, with a few short setae around (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Right side of head B left leg 1 C right leg 2 with gonopore D posterior part of body, dorsal view E left leg 9. Abbreviations: cx: coxa, f: femur, go: gonopore, O: ommatidia, ta: tarsus, To: Toemoesvary's organ, 8 - 11 refer to tergite numbers." figureDoi="10.3897/zookeys.954.54694.figure2" httpUri="https://binary.pensoft.net/fig/437443" pageId="0" pageNumber="1">2C</figureCitation>
). Legs 17 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Left leg 17, oral view B right anterior telopod, oral view C right anterior telopod, caudal view D-F left posterior telopod, oral, mesal and caudal views, respectively G tip of syncoxital horn, caudal view. Abbreviations: cx: coxa, st: sternite, sx: syncoxite, t 1 - t 4: telopoditomeres 1 - 4. Scale bars: 0.5 mm (A-C, left); 0.5 mm (D-G, right)." figureDoi="10.3897/zookeys.954.54694.figure4" httpUri="https://binary.pensoft.net/fig/437445" pageId="0" pageNumber="1">4A</figureCitation>
) strongly reduced, very densely micropilose throughout. Coxae membranous, contiguous, but clearly separated medially. Each coxa with a very large, rather regularly rounded, outer lobe and a small, rounded, distomedial, setigerous finger. Telopodite 4-segmented, telopoditomere 2 largest, subrectangular, about twice as long as telopoditomere 1 or telopoditomeres 3 and 4 combined.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Anterior telopods</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Left leg 17, oral view B right anterior telopod, oral view C right anterior telopod, caudal view D-F left posterior telopod, oral, mesal and caudal views, respectively G tip of syncoxital horn, caudal view. Abbreviations: cx: coxa, st: sternite, sx: syncoxite, t 1 - t 4: telopoditomeres 1 - 4. Scale bars: 0.5 mm (A-C, left); 0.5 mm (D-G, right)." figureDoi="10.3897/zookeys.954.54694.figure4" httpUri="https://binary.pensoft.net/fig/437445" pageId="0" pageNumber="1">4B, C</figureCitation>
) also very densely micropilose throughout. Syncoxite (=?coxosternum) membranous, on either side with a small rounded lobule at base of telopoditomere 1. Telopodite 4-segmented, with a spine apically. Telopoditomere 1 subrectangular, 1.2 times longer than wide. Telopoditomere 2 largest, a little swollen ventro-parabasally, its apico-mesal tooth on caudal face bulged at base, sharp apically and extending to basal 1/4 telopoditomere 3. The latter subtrapezoid, its apico-mesal tooth on caudal face small, rounded, projecting above base of a subcylindrical telopoditomere 4.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Posterior telopods</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tonkinomeris huzhengkuni sp. nov., ♂ holotype. A Left leg 17, oral view B right anterior telopod, oral view C right anterior telopod, caudal view D-F left posterior telopod, oral, mesal and caudal views, respectively G tip of syncoxital horn, caudal view. Abbreviations: cx: coxa, st: sternite, sx: syncoxite, t 1 - t 4: telopoditomeres 1 - 4. Scale bars: 0.5 mm (A-C, left); 0.5 mm (D-G, right)." figureDoi="10.3897/zookeys.954.54694.figure4" httpUri="https://binary.pensoft.net/fig/437445" pageId="0" pageNumber="1">4D-G</figureCitation>
) particularly strongly incrassate, likewise very densely micropilose throughout. A large, high, thick, and roundish syncoxite placed on a large membranous sternite, with a high, roundish, median lobe closely flanked by two inconspicuous, short, spiniform, obliquely truncate, setose horns (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tonkinomeris huzhengkuni sp. nov., ♀ paratype. A Gnathochilarium, ventral view, in situ B, C left mandible, mesal and subfrontal views, respectively D coxae 2 with vulvae. Abbreviations: ca: cardines of gnathochilarium, et: external tooth, gu: gula, iA: intermediate area, ip: inner palpus, it: inner tooth, ll: lamellae linguales, lp: lateral palpus, m: mentum, md: basal joint of mandible, mf: membranous fringe, mg: molar groove, mp: molar plate, pl: pectinate lamellae, st: stipites, vu: vulvae." figureDoi="10.3897/zookeys.954.54694.figure3" httpUri="https://binary.pensoft.net/fig/437444" pageId="0" pageNumber="1">3G</figureCitation>
), each latter being much shorter than syncoxital lobe. Telopodite 4-segmented, with a spine apically. Telopoditomere 1 squarish, with a very small distomesal, setigerous cone (a strongly reduced trichostele). Telopoditomere 2 with a prominent, finger-shaped, distomesal process on caudal face, produced apically to ca 1/3 telopoditomere 3. The latter elongate, gently tapering distad and curved apically basad towards process on telopoditomere 2, with another, much smaller, caudad curved, caudolateral process. Telopoditomere 4 smallest, subcylindrical, erect, clearly shifted anteriad, subtended by and reaching the distal end of telopoditomere 3.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Vulva</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tonkinomeris huzhengkuni sp. nov., ♀ paratype. A Gnathochilarium, ventral view, in situ B, C left mandible, mesal and subfrontal views, respectively D coxae 2 with vulvae. Abbreviations: ca: cardines of gnathochilarium, et: external tooth, gu: gula, iA: intermediate area, ip: inner palpus, it: inner tooth, ll: lamellae linguales, lp: lateral palpus, m: mentum, md: basal joint of mandible, mf: membranous fringe, mg: molar groove, mp: molar plate, pl: pectinate lamellae, st: stipites, vu: vulvae." figureDoi="10.3897/zookeys.954.54694.figure3" httpUri="https://binary.pensoft.net/fig/437444" pageId="0" pageNumber="1">3D</figureCitation>
) densely setose, large, covering 1/2 coxa 2.
</paragraph>
<caption doi="10.3897/zookeys.954.54694.figure3" httpUri="https://binary.pensoft.net/fig/437444" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName authorityName="Liu &amp; Golovatch" authorityYear="2020" class="Diplopoda" family="Glomeridellidae" genus="Tonkinomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinomeris huzhengkuni" order="Glomerida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="huzhengkuni">
<emphasis italics="true" pageId="0" pageNumber="1">Tonkinomeris huzhengkuni</emphasis>
</taxonomicName>
sp. nov., ♀ paratype.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Gnathochilarium, ventral view,
<emphasis italics="true" pageId="0" pageNumber="1">in situ</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
left mandible, mesal and subfrontal views, respectively
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
coxae 2 with vulvae. Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="1">ca</emphasis>
: cardines of gnathochilarium,
<emphasis bold="true" pageId="0" pageNumber="1">et</emphasis>
: external tooth,
<emphasis bold="true" pageId="0" pageNumber="1">gu</emphasis>
: gula,
<emphasis bold="true" pageId="0" pageNumber="1">iA</emphasis>
: intermediate area,
<emphasis bold="true" pageId="0" pageNumber="1">ip</emphasis>
: inner palpus,
<emphasis bold="true" pageId="0" pageNumber="1">it</emphasis>
: inner tooth,
<emphasis bold="true" pageId="0" pageNumber="1">ll</emphasis>
: lamellae linguales,
<emphasis bold="true" pageId="0" pageNumber="1">lp</emphasis>
: lateral palpus,
<emphasis bold="true" pageId="0" pageNumber="1">m</emphasis>
: mentum,
<emphasis bold="true" pageId="0" pageNumber="1">md</emphasis>
: basal joint of mandible,
<emphasis bold="true" pageId="0" pageNumber="1">mf</emphasis>
: membranous fringe,
<emphasis bold="true" pageId="0" pageNumber="1">mg</emphasis>
: molar groove,
<emphasis bold="true" pageId="0" pageNumber="1">mp</emphasis>
: molar plate,
<emphasis bold="true" pageId="0" pageNumber="1">pl</emphasis>
: pectinate lamellae,
<emphasis bold="true" pageId="0" pageNumber="1">st</emphasis>
: stipites,
<emphasis bold="true" pageId="0" pageNumber="1">vu</emphasis>
: vulvae.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.954.54694.figure4" httpUri="https://binary.pensoft.net/fig/437445" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName authorityName="Liu &amp; Golovatch" authorityYear="2020" class="Diplopoda" family="Glomeridellidae" genus="Tonkinomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinomeris huzhengkuni" order="Glomerida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="huzhengkuni">
<emphasis italics="true" pageId="0" pageNumber="1">Tonkinomeris huzhengkuni</emphasis>
</taxonomicName>
sp. nov., ♂ holotype.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Left leg17, oral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
right anterior telopod, oral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
right anterior telopod, caudal view
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
left posterior telopod, oral, mesal and caudal views, respectively
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
tip of syncoxital horn, caudal view. Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="1">cx</emphasis>
: coxa,
<emphasis bold="true" pageId="0" pageNumber="1">st</emphasis>
: sternite,
<emphasis bold="true" pageId="0" pageNumber="1">sx</emphasis>
: syncoxite,
<emphasis bold="true" pageId="0" pageNumber="1">t1-t4</emphasis>
: telopoditomeres 1-4. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
, left); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">D-G</emphasis>
, right).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

225
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<taxonomicName id="6FA966A608DE993E2B65729B37F73341" ID-CoL="9G5YM" LSID="http://azoresbioportal.uac.pt/azores-species/polydesmus-coriaceus-8146/" authority="Porat, 1871" authorityName="Porat" authorityYear="1871" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus coriaceus" order="Polydesmida" pageId="0" pageNumber="10948" phylum="Arthropoda" rank="species" species="coriaceus">Polydesmus coriaceus Porat, 1871</taxonomicName>
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<paragraph id="C8CD003B08BACD1600E02A78E1BAADC6" pageId="0" pageNumber="10948">Ecological interactions</paragraph>
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<paragraph id="5A510633A299D416673D322A29771951" pageId="0" pageNumber="10948">Native status</paragraph>
<paragraph id="7C63011EF7ED51C3729F631F8CB418E4" pageId="0" pageNumber="10948">Introduced</paragraph>
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<paragraph id="25B421D85389BE5E4B9B1893286817CB" pageId="0" pageNumber="10948">Distribution</paragraph>
<paragraph id="00B72E5EF491DCA88ED75CE2080F69BF" pageId="0" pageNumber="10948">COR; FLO*; FAI; PIC; GRA; SJG*; TER; SMG; SMR</paragraph>
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<subSubSection id="C8BCF12095454B3902BEDDCFD5D76089" pageId="0" pageNumber="10948" type="notes">
<paragraph id="5313CBB11C93335F3D42DF56667E18E4" pageId="0" pageNumber="10948">Notes</paragraph>
<paragraph id="138BC54B5880A1B6285816B61A42325D" pageId="0" pageNumber="10948">Also present: CAN (Biogeographical Realm: Western Palearctic)</paragraph>
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</treatment>
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<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
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<mods:namePart>Sajan, Sheikh</mods:namePart>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
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<mods:namePart>Meng, Kaibaryer</mods:namePart>
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<mods:namePart>Asami, Takahiro</mods:namePart>
<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jaintiacus subsp. var. var. crassus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="jaintiacus" subSpecies="var." variety="crassus">Alycaeus jaintiacus var. crassus</taxonomicName>
Godwin-Austen, 1914: 375, pl. 137, figs 5, 5a.
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<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Dicharax) jaintiacus subsp. Var. var. crassa" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="jaintiacus" subGenus="Dicharax" subSpecies="Var." variety="crassa">Alycaeus (Dicharax) jaintiacus Var. crassa</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude 1921</bibRefCitation>
: 256-257.
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">&quot;in Nongjinghi, Jiantia, 4563 feet&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">Nonjinghi, Jiantia, coll. Godwin-Austen, NHMUK 1903.7.1.2752 (4 syntypes in 2 vials).</paragraph>
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<paragraph pageId="0" pageNumber="1">Protoconch matte, R1 smooth, no spiral lines visible (although the entire shell is somewhat weathered); R2 of normal length, the ribs are overall low, they are slightly curved towards the aperture.</paragraph>
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
species group
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<paragraph box="[118,208,679,697]" pageId="7" pageNumber="8" type="mainText">Diagnosis</paragraph>
<paragraph box="[118,1127,707,923]" pageId="7" pageNumber="8" type="mainText">Eleven-segmented antennae; anterior clypeal margin medially impressed; frontal carinae well-developed and usually running to posterior head margin; anterior face of mesosoma weakly developed; margination between lateral and dorsal mesosoma very weakly developed, sides round smoothly onto dorsum; propodeal spines always long to extremely long, and spinose; propodeal lobes short, triangular, and blunted or acute; petiolar node clublike, anterodorsal margin situated lower and much less pronounced than posterodorsal margin, in profile approximately as long as high, in dorsal view distinctly longer than wide; postpetiole globular to subglobular; mandibles strongly sculptured; sculpture on head, mesosoma, and waist segments distinct and predominantly reticulate-rugose; gaster unsculptured, smooth, and shiny; whole body covered with numerous, very long, fine, standing hairs; sting appendage spatulate.</paragraph>
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<paragraph box="[118,219,960,978]" pageId="7" pageNumber="8" type="mainText">Comments</paragraph>
<paragraph box="[118,1121,988,1064]" pageId="7" pageNumber="8" type="mainText">
The recently (Hita Garcia &amp; Fisher, 2012) proposed
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
species group contains
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
and the
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
T.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
. It is mainly distributed in the north and west of Madagascar, as well as on Nosy Be and Mayotte, and both species prefer forested habitats.
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Other
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
species from a different species group are unlikely to be confused with T.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
or
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri.</taxonomicName>
The possession of 11-segmented antennae and the very conspicuous petiolar node shape clearly distinguish them from all other Malagasy species groups.
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<paragraph box="[118,643,1213,1233]" pageId="7" pageNumber="8" type="mainText">
Key to species of the
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
species group (workers)
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1. Species with smaller eyes (OI 20); relatively longer antennal scapes (SI 101-104); and long propodeal spines (PSLI 35-38) (Figs. 1, 2).................................................................................. T.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
</paragraph>
<paragraph box="[175,1123,1314,1354]" pageId="7" pageNumber="8" type="mainText">
Species with larger eyes (OI 24-26); relatively shorter antennal scapes (SI 89-99); and extremely long propodeal spines (PSLI 49-68) (Figs. 3, 4)...............................................................................
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
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