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<mods:title id="BDF8EBEAE2591B04FF88CDE5D31543A5">Notes on the family Macrodinychidae (Acari: Uropodina) with description of two new species</mods:title>
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<subSubSection id="32E8652FFFE8FFB82A57FAAA5295FA81" box="[334,911,1312,1336]" pageId="2" pageNumber="1620" type="nomenclature">
<paragraph id="7A4D36A4FFE8FFB82A57FAAA5295FA81" blockId="2.[334,911,1312,1336]" box="[334,911,1312,1336]" pageId="2" pageNumber="1620">
<heading id="210581C8FFE8FFB82A57FAAA5295FA81" box="[334,911,1312,1336]" centered="true" fontSize="36" level="2" pageId="2" pageNumber="1620" reason="7">
Family
<taxonomicName id="BDF24D27FFE8FFB82ABAFAAA5295FA81" ID-CoL="CB5" ID-ENA="122016" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" box="[419,911,1312,1336]" class="Arachnida" family="Macrodinychidae" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="1620" phylum="Arthropoda" rank="family">
<emphasis id="4886EAB6FFE8FFB82ABAFAAA53A3FA81" bold="true" box="[419,697,1312,1336]" pageId="2" pageNumber="1620">MACRODINYCHIDAE</emphasis>
<bibRefCitation id="1E634B55FFE8FFB829D9FAAA5295FA81" author="Hirschmann W" box="[704,911,1312,1336]" pageId="2" pageNumber="1620" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="32E8652FFFE8FFB82B89FAFA5019FA61" pageId="2" pageNumber="1620" type="reference_group">
<paragraph id="7A4D36A4FFE8FFB82B89FAFA5399FA31" blockId="2.[144,643,1392,1416]" box="[144,643,1392,1416]" pageId="2" pageNumber="1620">
<treatmentCitationGroup id="5AE2118AFFE8FFB82B89FAFA5399FA31" box="[144,643,1392,1416]" pageId="2" pageNumber="1620">
<treatmentCitation id="FB5310B5FFE8FFB82B89FAFA5365FA31" author="Hirschmann W" box="[144,639,1392,1416]" page="70" pageId="2" pageNumber="1620" year="1979">
<taxonomicName id="BDF24D27FFE8FFB82B89FAFA5365FA31" ID-CoL="CB5" authority="Hirschmann, 1979: 58 - 70" authorityName="Hirschmann" authorityPageNumber="58 - 70" authorityYear="1979" box="[144,639,1392,1416]" class="Arachnida" family="Macrodinychidae" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="1620" phylum="Arthropoda" rank="family">
Macrodinychidae
<bibRefCitation id="1E634B55FFE8FFB82A45FAFA5365FA31" author="Hirschmann W" box="[348,639,1392,1416]" pageId="2" pageNumber="1620" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979: 58–70</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
<paragraph id="7A4D36A4FFE8FFB82B89FA2A5019FA61" blockId="2.[144,1101,1440,1496]" pageId="2" pageNumber="1620">
<treatmentCitationGroup id="5AE2118AFFE8FFB82B89FA2A5019FA61" pageId="2" pageNumber="1620">
<taxonomicName id="BDF24D27FFE8FFB82B89FA2A50F5FA01" ID-CoL="CB5" authority="Hirschmann" authorityName="Hirschmann" authorityYear="1979" box="[144,495,1440,1464]" class="Arachnida" family="Macrodinychidae" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="1620" phylum="Arthropoda" rank="family">Macrodinychidae Hirschmann</taxonomicName>
in
<bibRefCitation id="1E634B55FFE8FFB8293BFA2A53F6FA01" author="Wisniewski J" box="[546,748,1440,1465]" pageId="2" pageNumber="1620" pagination="68 - 74" refId="ref8086" refString="Wisniewski J. 1979. Gangsystematik der Parasitiformes Teil 339. Zur Kenntnis der Uropodidae- Fauna Polens. Acarol. Schrift. Vergl. Milbenkunde. 26: 68 - 74." type="journal article" year="1979">Wiśniewski, 1979</bibRefCitation>
(sic!)
<treatmentCitation id="FB5310B5FFE8FFB82826FA2A51E4FA61" author="Farrier MH &amp; Hennessey MK" page="162" pageId="2" pageNumber="1620" year="1996">
<bibRefCitation id="1E634B55FFE8FFB82826FA2A51E4FA61" author="Farrier MH &amp; Hennessey MK" pageId="2" pageNumber="1620" pagination="408" refId="ref7012" refString="Farrier MH, Hennessey MK. 1996. Soil-inhabiting and free-living Mesostigmata (Acari- Parasitiformes) from North America. An annotated checklist with bibliography and index. North Carolina Agricultural Service. North Carolina State University Raleigh. North Carolina Technical Bulletin 302. pp. 408" type="journal article" year="1996">Farrier and Hennessey 1996: 162</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="32E8652FFFE8FFB92B89F988531EFB92" lastPageId="3" lastPageNumber="1621" pageId="2" pageNumber="1620" type="diagnosis">
<paragraph id="7A4D36A4FFE8FFB82B89F98851E6F9A0" blockId="2.[144,1101,1538,1688]" box="[144,252,1538,1561]" pageId="2" pageNumber="1620">
<emphasis id="4886EAB6FFE8FFB82B89F98851E6F9A0" box="[144,252,1538,1561]" italics="true" pageId="2" pageNumber="1620">Diagnosis</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE8FFB92B89F9AB50DCFF47" blockId="2.[144,1101,1538,1688]" lastBlockId="3.[213,1170,166,254]" lastPageId="3" lastPageNumber="1621" pageId="2" pageNumber="1620">Idiosoma large, oval, posterior margin rounded. Colour yellowish-brown. Legs short and can be found on the apical part of the idiosoma. All legs with welldeveloped ambulacra. Genital shield of female small and oval, similar to genital shield of male. Tritosternum with narrow basis and trifurcated. Gnathosoma: hypostomal setae long, with pilose margins. Corniculi horn-like. Internal malae pilose. Epistome marginally pilose. Chelicerae with internal sclerotized nodes, fixed digit without process.</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFEA0502DFEFB" blockId="3.[213,1030,298,354]" box="[213,311,298,322]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCFEA0502DFEFB" box="[213,311,298,322]" italics="true" pageId="3" pageNumber="1621">Remarks</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFEC3551CFED8" blockId="3.[213,1030,298,354]" box="[213,1030,329,354]" pageId="3" pageNumber="1621">
Only one genus (
<taxonomicName id="BDF24D27FFE9FFB92A8BFEC053C9FED8" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[402,723,329,354]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFE9FFB92A8BFEC05320FEDB" box="[402,570,330,354]" italics="true" pageId="3" pageNumber="1621">Macrodinychus</emphasis>
<bibRefCitation id="1E634B55FFE9FFB92959FEC353C9FED8" author="Berlese A" box="[576,723,329,353]" pageId="3" pageNumber="1621" pagination="125 - 177" refId="ref6947" refString="Berlese A. 1916. Centuria seconda di Acari nuovi. Redia. 12: 125 - 177." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
</taxonomicName>
) is known from this family.
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFE045040FE1F" blockId="3.[213,854,398,453]" box="[213,346,398,422]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCFE045040FE1F" box="[213,346,398,422]" italics="true" pageId="3" pageNumber="1621">Distribution</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFE27524CFE7C" blockId="3.[213,854,398,453]" box="[213,854,429,453]" pageId="3" pageNumber="1621">Circum-tropical, European, Australian and Middle-East.</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFE78500FFDB0" blockId="3.[213,1170,498,744]" box="[213,277,498,521]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCFE78500FFDB0" box="[213,277,498,521]" italics="true" pageId="3" pageNumber="1621">Notes</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFD985385FD51" blockId="3.[213,1170,498,744]" pageId="3" pageNumber="1621">
In the last comprehensive work (
<bibRefCitation id="1E634B55FFE9FFB92940FD9B5260FD90" author="Krantz GW &amp; Walter DE" box="[601,890,529,553]" pageId="3" pageNumber="1621" refId="ref7781" refString="Krantz GW, Walter DE. 2009. A manual of acarology. 3 rd edition. Lubbock (TX): Texas University Press. pp. 807." type="book" year="2009">Krantz and Walter 2009</bibRefCitation>
),
<taxonomicName id="BDF24D27FFE9FFB9288BFD9B558BFD90" box="[914,1169,529,553]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4886EAB6FFE9FFB9288BFD9B5520FD90" box="[914,1082,529,553]" italics="true" pageId="3" pageNumber="1621">Macrodinychus</emphasis>
species
</taxonomicName>
were mentioned as members of the family
<taxonomicName id="BDF24D27FFE9FFB929F0FDBB5595FDF0" authority="(Lindquist et al. 2009)" baseAuthorityName="Lindquist" baseAuthorityYear="2009" box="[745,1167,561,585]" class="Arachnida" family="Dinychidae" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="family">
Dinychidae (
<bibRefCitation id="1E634B55FFE9FFB92899FDBB559DFDF0" author="Lindquist EE &amp; Krantz GW &amp; Walter DE" box="[896,1159,561,585]" pageId="3" pageNumber="1621" pagination="124 - 232" refId="ref7888" refString="Lindquist EE, Krantz GW, Walter DE. 2009. Order Mesostigmata. In: Krantz GW, Walter DE. editors. A manual of acarology. 3 rd edition. Lubbock (TX): Texas University Press, pp. 124 - 232." type="book chapter" year="2009">Lindquist et al. 2009</bibRefCitation>
)
</taxonomicName>
. Unfortunately this statement [fixed cheliceral digit typically with a rounded or acuminate apical “finger” (see (
<bibRefCitation id="1E634B55FFE9FFB9295CFDFB5226FD30" author="Lindquist EE &amp; Krantz GW &amp; Walter DE" box="[581,828,625,649]" pageId="3" pageNumber="1621" pagination="124 - 232" refId="ref7888" refString="Lindquist EE, Krantz GW, Walter DE. 2009. Order Mesostigmata. In: Krantz GW, Walter DE. editors. A manual of acarology. 3 rd edition. Lubbock (TX): Texas University Press, pp. 124 - 232." type="book chapter" year="2009">Lindquist et al. 2009</bibRefCitation>
) on page 197], which can be found in the key to the family
<taxonomicName id="BDF24D27FFE9FFB92929FD1B53A9FD10" box="[560,691,657,681]" class="Arachnida" family="Dinychidae" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="family">Dinychidae</taxonomicName>
, is not true for the
<taxonomicName id="BDF24D27FFE9FFB92896FD1B5594FD10" box="[911,1166,657,681]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4886EAB6FFE9FFB92896FD1B552DFD10" box="[911,1079,657,681]" italics="true" pageId="3" pageNumber="1621">Macrodinychus</emphasis>
species
</taxonomicName>
, where this character is lacking on their chelicerae (
<bibRefCitation id="1E634B55FFE9FFB9281DFD3A52C8FD71" author="Hirschmann W" box="[772,978,688,712]" pageId="3" pageNumber="1621" pagination="292 - 370" refId="ref7436" refString="Hirschmann W. 1993. Gangsystematik der Parasitiformes Teil 550. Bestimmungstabellen der Uropodiden der Erde, Atlas der Ganggattungen der Atrichopygiina. Acarol Schrift Vergl Milbenkunde. 40: 292 - 370." type="journal article" year="1993">Hirschmann 1993</bibRefCitation>
). For this reason, I discuss these mites as a separate family.
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFC9E5030FC95" blockId="3.[213,1170,788,1067]" box="[213,298,788,812]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCFC9E5030FC95" box="[213,298,788,812]" italics="true" pageId="3" pageNumber="1621">Biology</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFCBE531EFB92" blockId="3.[213,1170,788,1067]" pageId="3" pageNumber="1621">
Currently there is little information about the biology and ecology of these mites.
<bibRefCitation id="1E634B55FFE9FFB92F72FCBE50B0FCD5" author="Bal DA &amp; Ozkan M" pageId="3" pageNumber="1621" pagination="125 - 132" refId="ref6903" refString="Bal DA, Ozkan M. 2005. A new viviparous Uropodid mite (Acari: Gamasida: Uropodina) for the Turkish Fauna, Macrodinychus (Monomacrodinychus) bregetovaae Hirschmann, 1975. Turk J Zool. 25: 125 - 132." type="journal article" year="2005">Bal and Özkan (2005)</bibRefCitation>
reported viviparity in
<taxonomicName id="BDF24D27FFE9FFB929A4FCDE5010FC35" authority="Hirschmann, 1975" authorityName="Hirschmann" authorityYear="1975" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="species" species="bregetovaae">
<emphasis id="4886EAB6FFE9FFB929A4FCDE52E8FCD5" box="[701,1010,852,876]" italics="true" pageId="3" pageNumber="1621">Macrodinychus bregetovaae</emphasis>
Hirschmann, 1975
</taxonomicName>
because they found about
<specimenCount id="6CF4FD2DFFE9FFB92926FCFE53BCFC35" box="[575,678,884,908]" pageId="3" pageNumber="1621" type="larva">30 larvae</specimenCount>
in the female’s body, but this phenomenon has not been recorded in other Macrodinychid species. One species of this family (
<taxonomicName id="BDF24D27FFE9FFB92BC7FC3E52D8FC75" authority="Hirschmann and Zirngiebl-Nicol, 1975" authorityName="Hirschmann and Zirngiebl-Nicol" authorityYear="1975" box="[222,962,948,972]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="species" species="sellnicki">
<emphasis id="4886EAB6FFE9FFB92BC7FC3E50F6FC75" box="[222,492,948,972]" italics="true" pageId="3" pageNumber="1621">Macrodinychus sellnicki</emphasis>
Hirschmann and Zirngiebl-Nicol, 1975
</taxonomicName>
) is a well-known parasite of ant pupae (
<bibRefCitation id="1E634B55FFE9FFB92AC2FC5953D2FC52" author="Gonzales VE &amp; Gomez LA &amp; Mesa NC" box="[475,712,979,1003]" pageId="3" pageNumber="1621" pagination="143 - 149" refId="ref7060" refString="Gonzales VE, Gomez LA, Mesa NC. 2004. Observations on the biology and behaviour of the mite Macrodinychus sellnicki (Mesostigmata: Uropodidae) ectoparasite of the crazy ant Paratrechina fulva (Hymenoptera: Formicidae). Rev Colomb Entomol. 30 (2): 143 - 149." type="journal article" year="2004">Gonzales et al. 2004</bibRefCitation>
;
<bibRefCitation id="1E634B55FFE9FFB929CCFC5E52BEFC52" author="Vargas GAO &amp; Diaz AP &amp; Lastra LAB &amp; Meas NCC &amp; de Polania IZ &amp; Gomez LL" box="[725,932,979,1004]" pageId="3" pageNumber="1621" pagination="225 - 232" refId="ref7961" refString="Vargas GAO, Diaz AP, Lastra LAB, Meas NCC, de Polania IZ, Gomez LL. 2004. Reconocimiento de enemigos naturales de la hormiga loca, Paratrechina fulva (Hymenoptera: Formicidae), en el municipio de El Colegio (Cundinamarca) y en el valle del rio Cauca. Rev Colomb Entomol. 30 (2): 225 - 232." type="journal article" year="2004">Vargas et al. 2004</bibRefCitation>
;
<bibRefCitation id="1E634B55FFE9FFB928ABFC5E559CFC52" author="Krantz GW &amp; Gomez LA &amp; Gonzalez VE" box="[946,1158,979,1003]" pageId="3" pageNumber="1621" pagination="29 - 38" refId="ref7811" refString="Krantz GW, Gomez LA, Gonzalez VE. 2007. Parasitism in the Uropodina: a case history from Colombia. In: Morales-Malacara JB, Behan-Pelletier V, Ueckermann EA, Perez TM, Estrada-Venegas EG, Badii MH. editors. Acarology XI: Proceedings of the International Congress. Instituto de Biologia and Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, Mexico, City, pp. 29 - 38." type="book chapter" year="2007">Krantz et al. 2007</bibRefCitation>
), and presumably other species are associated with ant species or live in anthills, but we have no other information.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="8FBC0E250967171C26DE5FD8992CDFD2">Notes on the family Macrodinychidae (Acari: Uropodina) with description of two new species</mods:title>
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<heading id="210581C8FFE9FFB92AF5FBD75261FBCC" box="[492,891,1116,1141]" centered="true" fontSize="36" level="2" pageId="3" pageNumber="1621" reason="7">
Genus
<taxonomicName id="BDF24D27FFE9FFB92922FBD65261FBCC" authority="Berlese, 1917" authorityName="Berlese" authorityYear="1917" box="[571,891,1116,1141]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFE9FFB92922FBD653F9FBCC" bold="true" box="[571,739,1116,1141]" italics="true" pageId="3" pageNumber="1621">Macrodinychus</emphasis>
<bibRefCitation id="1E634B55FFE9FFB929F0FBD75261FBCC" author="Berlese A" box="[745,891,1117,1141]" pageId="3" pageNumber="1621" pagination="7 - 16" refId="ref6966" refString="Berlese A. 1917. Intorno agli Uropodidae. Redia. 13: 7 - 16." type="journal article" year="1917">Berlese, 1917</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="32E8652FFFE9FFB92BCCFB2B5240FAB1" pageId="3" pageNumber="1621" type="reference_group">
<paragraph id="7A4D36A4FFE9FFB92BCCFB2B535FFB01" blockId="3.[213,581,1184,1209]" box="[213,581,1184,1209]" pageId="3" pageNumber="1621">
<treatmentCitationGroup id="5AE2118AFFE9FFB92BCCFB2B535FFB01" box="[213,581,1184,1209]" pageId="3" pageNumber="1621">
<treatmentCitation id="FB5310B5FFE9FFB92BCCFB2B535BFB01" author="Berlese A" box="[213,577,1184,1209]" page="12" pageId="3" pageNumber="1621" year="1917">
<taxonomicName id="BDF24D27FFE9FFB92BCCFB2B535BFB01" authority="Berlese, 1917: 12" authorityName="Berlese" authorityPageNumber="12" authorityYear="1917" box="[213,577,1184,1209]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFE9FFB92BCCFB2B5067FB00" box="[213,381,1185,1209]" italics="true" pageId="3" pageNumber="1621">Macrodinychus</emphasis>
<bibRefCitation id="1E634B55FFE9FFB92A9DFB2A535BFB01" author="Berlese A" box="[388,577,1184,1208]" pageId="3" pageNumber="1621" pagination="7 - 16" refId="ref6966" refString="Berlese A. 1917. Intorno agli Uropodidae. Redia. 13: 7 - 16." type="journal article" year="1917">Berlese, 1917: 12</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFB5B5240FAB1" blockId="3.[213,1170,1232,1289]" pageId="3" pageNumber="1621">
<treatmentCitationGroup id="5AE2118AFFE9FFB92BCCFB5B5240FAB1" pageId="3" pageNumber="1621">
<treatmentCitation id="FB5310B5FFE9FFB92BCCFB5B525EFB51" author="Wisniewski J &amp; Hirschmann W" box="[213,836,1232,1257]" page="32" pageId="3" pageNumber="1621" year="1993">
<taxonomicName id="BDF24D27FFE9FFB92BCCFB5B525EFB51" authority=": Wisniewski and Hirschmann 1993: 32" authorityName=": Wisniewski and Hirschmann" authorityPageNumber="32" authorityYear="1993" box="[213,836,1232,1257]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1621" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFE9FFB92BCCFB5B5067FB50" box="[213,381,1233,1257]" italics="true" pageId="3" pageNumber="1621">Macrodinychus</emphasis>
:
<bibRefCitation id="1E634B55FFE9FFB92A97FB5A525EFB51" author="Wisniewski J &amp; Hirschmann W" box="[398,836,1232,1257]" pageId="3" pageNumber="1621" pagination="1 - 220" refId="ref8187" refString="Wisniewski J, Hirschmann W. 1993. Gangsystematik der Parasitiformes Teil 548. Katalog der Ganggattungen, Untergattungen, Gruppen und Arten der Uropodiden der Erde. Acarol Schrift Vergl Milbenkunde. 40: 1 - 220." type="journal article" year="1993">Wiśniewski and Hirschmann 1993: 32</bibRefCitation>
</taxonomicName>
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,
<treatmentCitation id="FB5310B5FFE9FFB9284BFB5A5594FB51" author="Hirschmann W" box="[850,1166,1232,1256]" page="330" pageId="3" pageNumber="1621" year="1993">
<bibRefCitation id="1E634B55FFE9FFB9284BFB5A5594FB51" author="Hirschmann W" box="[850,1166,1232,1256]" pageId="3" pageNumber="1621" pagination="292 - 370" refId="ref7436" refString="Hirschmann W. 1993. Gangsystematik der Parasitiformes Teil 550. Bestimmungstabellen der Uropodiden der Erde, Atlas der Ganggattungen der Atrichopygiina. Acarol Schrift Vergl Milbenkunde. 40: 292 - 370." type="journal article" year="1993">Hirschmann 1993: 330–331</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="FB5310B5FFE9FFB92BCCFB7A50C2FAB1" author="Wisniewski J" box="[213,472,1264,1289]" page="383" pageId="3" pageNumber="1621" year="1993">
<bibRefCitation id="1E634B55FFE9FFB92BCCFB7A50C2FAB1" author="Wisniewski J" box="[213,472,1264,1289]" pageId="3" pageNumber="1621" pagination="371 - 429" refId="ref8158" refString="Wisniewski J. 1993 b. Alphabetisches Verzeichnis der Uropodiden (Gattungen, Arten, Synonyma). Acarol Schrift Vergl Milbenkunde. 40: 371 - 429." type="journal article" year="1993">Wiśniewski 1993b: 383</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="FB5310B5FFE9FFB92AFAFB7A524CFAB1" author="Farrier MH &amp; Hennessey MK" box="[483,854,1264,1289]" page="162" pageId="3" pageNumber="1621" year="1996">
<bibRefCitation id="1E634B55FFE9FFB92AFAFB7A524CFAB1" author="Farrier MH &amp; Hennessey MK" box="[483,854,1264,1289]" pageId="3" pageNumber="1621" pagination="408" refId="ref7012" refString="Farrier MH, Hennessey MK. 1996. Soil-inhabiting and free-living Mesostigmata (Acari- Parasitiformes) from North America. An annotated checklist with bibliography and index. North Carolina Agricultural Service. North Carolina State University Raleigh. North Carolina Technical Bulletin 302. pp. 408" type="journal article" year="1996">Farrier and Hennessey 1996: 162</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="32E8652FFFE9FFBE2BCCFAB05030FC22" lastPageId="4" lastPageNumber="1622" pageId="3" pageNumber="1621" type="diagnosis">
<paragraph id="7A4D36A4FFE9FFB92BCCFAB0505BFAE8" blockId="3.[213,414,1338,1393]" box="[213,321,1338,1361]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCFAB0505BFAE8" box="[213,321,1338,1361]" italics="true" pageId="3" pageNumber="1621">Diagnosis</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFAD35084FAC8" blockId="3.[213,414,1338,1393]" box="[213,414,1369,1393]" pageId="3" pageNumber="1621">See under Family.</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFA175040FA0C" blockId="3.[213,414,1437,1493]" box="[213,346,1437,1461]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCFA175040FA0C" box="[213,346,1437,1461]" italics="true" pageId="3" pageNumber="1621">Distribution</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCFA375084FA6C" blockId="3.[213,414,1437,1493]" box="[213,414,1469,1493]" pageId="3" pageNumber="1621">See under Family.</paragraph>
<paragraph id="7A4D36A4FFE9FFB92BCCF98B509EF9A0" blockId="3.[213,1170,1537,1689]" box="[213,388,1537,1561]" pageId="3" pageNumber="1621">
<emphasis id="4886EAB6FFE9FFB92BCCF98B509EF9A0" box="[213,388,1537,1561]" italics="true" pageId="3" pageNumber="1621">Systematic note</emphasis>
</paragraph>
<paragraph id="7A4D36A4FFE9FFBE2BCCF9AB51CCFF67" blockId="3.[213,1170,1537,1689]" lastBlockId="4.[144,1101,166,923]" lastPageId="4" lastPageNumber="1622" pageId="3" pageNumber="1621">
<bibRefCitation id="1E634B55FFE9FFB92BCCF9AB50ABF980" author="Hirschmann W" box="[213,433,1569,1593]" pageId="3" pageNumber="1621" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann (1979)</bibRefCitation>
used three different categories in his specific Uropodina-system: “Ganggattung”, “Stadiengattung” and “Stadiengruppen”. However, Hirschmann’s categories are not recognized by the International Code of Zoological Nomenclature (
<bibRefCitation id="1E634B55FFE9FFB92BC4F90A5076F921" author="ICZN" box="[221,364,1664,1688]" pageId="3" pageNumber="1621" refId="ref7519" refString="ICZN 1999. International Code of Zoological Nomenclature, 4 th edition. London: International Trust for Zoological Nomenclature. pp. 306." type="book" year="1999">ICZN 1999</bibRefCitation>
) so in the present work “Ganggattung” is regarded as genus, “Stadiengattung” as subgenus and “Stadiengruppen” as species-group (see
<bibRefCitation id="1E634B55FFEEFFBE28CBFF2C51D0FF67" author="Kontschan J" pageId="4" pageNumber="1622" pagination="55 - 66" refId="ref7738" refString="Kontschan J. 2008. Trigonuropoda (Baloghiatrigon) dominicana sp. nov. from the Dominican Republic, with notes on the subgenus Baloghiatrigon Hirschmann, 1979 (Acari: Uropodina: Trigonuropodidae). Zootaxa 1856: 55 - 66." type="journal article" year="2008">Kontschán 2008</bibRefCitation>
).
</paragraph>
<paragraph id="7A4D36A4FFEEFFBE2BA2FF6C50F4FE07" blockId="4.[144,1101,166,923]" pageId="4" pageNumber="1622">
First,
<bibRefCitation id="1E634B55FFEEFFBE2A12FF6C50E5FF47" author="Hirschmann W" box="[267,511,230,254]" pageId="4" pageNumber="1622" pagination="35 - 36" refId="ref7238" refString="Hirschmann W. 1975 a. Gangsystematik der Parasitiformes Teil 204. Die Gattung Macrodinychus (Berlese, 1917) und die Untergattung Monomacrodinychus novum subgenus (Trichouropodini, Uropodinae). Acarol Schrift Vergl Milbenkunde. 21: 35 - 36." type="journal article" year="1975">Hirschmann (1975a)</bibRefCitation>
subdivided the genus
<taxonomicName id="BDF24D27FFEEFFBE2831FF6C52CAFF47" authorityName=": Wisniewski and Hirschmann" authorityYear="1993" box="[808,976,230,254]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2831FF6C52CAFF47" box="[808,976,230,254]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
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into two “Stadiengattungen”,
<taxonomicName id="BDF24D27FFEEFFBE2A65FE8C533EFEA7" authorityName=": Wisniewski and Hirschmann" authorityYear="1993" box="[380,548,262,286]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2A65FE8C533EFEA7" box="[380,548,262,286]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
</taxonomicName>
and
<taxonomicName id="BDF24D27FFEEFFBE2943FE8C5224FEA7" box="[602,830,262,286]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="subGenus" subGenus="Monomacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2943FE8C5224FEA7" box="[602,830,262,286]" italics="true" pageId="4" pageNumber="1622">Monomacrodinychus</emphasis>
</taxonomicName>
, on the basis of the denticulation of chelicerae, which was often used in his catalogue (see
<bibRefCitation id="1E634B55FFEEFFBE286CFEAF555BFE84" author="Hirschmann W" box="[885,1089,293,317]" pageId="4" pageNumber="1622" pagination="292 - 370" refId="ref7436" refString="Hirschmann W. 1993. Gangsystematik der Parasitiformes Teil 550. Bestimmungstabellen der Uropodiden der Erde, Atlas der Ganggattungen der Atrichopygiina. Acarol Schrift Vergl Milbenkunde. 40: 292 - 370." type="journal article" year="1993">Hirschmann 1993</bibRefCitation>
). The shape of the chelicerae is diverse within a genus, because modification of chelicerae is the result of an adaptation of feeding habits (
<bibRefCitation id="1E634B55FFEEFFBE29E0FEEF5265FEC4" author="Evans GO" box="[761,895,357,381]" pageId="4" pageNumber="1622" pagination="193 - 206" refId="ref6983" refString="Evans GO. 1972. Leg chaetotaxy and the classification of the Uropodina (Acari: Mesostigmata). J Zool Lond. 167: 193 - 206." type="journal article" year="1972">Evans 1972</bibRefCitation>
), so its taxonomic value is questionable. For this reason, this difference is insufficient to justify the separation of these two groups.
</paragraph>
<paragraph id="7A4D36A4FFEEFFBE2BA2FE4F51D2FD45" blockId="4.[144,1101,166,923]" pageId="4" pageNumber="1622">
Some years later
<bibRefCitation id="1E634B55FFEEFFBE2AADFE4F53BBFE64" author="Hirschmann W" box="[436,673,453,477]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann (1979)</bibRefCitation>
mentioned
<taxonomicName id="BDF24D27FFEEFFBE2855FE4F52EEFE64" authorityName=": Wisniewski and Hirschmann" authorityYear="1993" box="[844,1012,453,477]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2855FE4F52EEFE64" box="[844,1012,453,477]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
</taxonomicName>
as a “Ganggattung”, with
<taxonomicName id="BDF24D27FFEEFFBE2AA8FE6F5204FE44" authority="Berlese, 1917" authorityName="Berlese" authorityYear="1917" box="[433,798,485,509]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2AA8FE6F5343FE44" box="[433,601,485,509]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE296DFE6F5204FE44" author="Berlese A" box="[628,798,485,509]" pageId="4" pageNumber="1622" pagination="7 - 16" refId="ref6966" refString="Berlese A. 1917. Intorno agli Uropodidae. Redia. 13: 7 - 16." type="journal article" year="1917">Berlese, 1917</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BDF24D27FFEEFFBE2827FE6F507BFDA4" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Bregetovamacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2827FE6F5556FE44" box="[830,1100,485,509]" italics="true" pageId="4" pageNumber="1622">Bregetovamacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2B89FD8F507BFDA4" author="Hirschmann W" box="[144,353,517,541]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BDF24D27FFEEFFBE2A75FD8F523CFDA4" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" box="[364,806,517,541]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Loksamacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2A75FD8F5348FDA4" box="[364,594,517,541]" italics="true" pageId="4" pageNumber="1622">Loksamacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2941FD8F523CFDA4" author="Hirschmann W" box="[600,806,517,541]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BDF24D27FFEEFFBE2844FD8F507EFD84" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Baloghmacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2844FD8F5557FDA4" box="[861,1101,517,541]" italics="true" pageId="4" pageNumber="1622">Baloghmacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2B89FDAF507EFD84" author="Hirschmann W" box="[144,356,549,573]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
as “Stadiengattungen”. Although
<bibRefCitation id="1E634B55FFEEFFBE29E3FDAF52C3FD84" author="Hirschmann W" box="[762,985,549,573]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann (1979)</bibRefCitation>
discussed these groups as subgenera of the genus
<taxonomicName id="BDF24D27FFEEFFBE2967FDCF523CFDE4" authorityName="Berlese" authorityYear="1917" box="[638,806,581,605]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2967FDCF523CFDE4" box="[638,806,581,605]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
</taxonomicName>
, in the same paper (on pages 69 and 70) he refers to
<taxonomicName id="BDF24D27FFEEFFBE2AF1FDEF522AFDC5" authority="Berlese, 1917" authorityName="Berlese" authorityYear="1917" box="[488,816,612,637]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2AF1FDEF538AFDC4" box="[488,656,613,637]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2980FDEE522AFDC5" author="Berlese A" box="[665,816,612,636]" pageId="4" pageNumber="1622" pagination="7 - 16" refId="ref6966" refString="Berlese A. 1917. Intorno agli Uropodidae. Redia. 13: 7 - 16." type="journal article" year="1917">Berlese, 1917</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BDF24D27FFEEFFBE2827FDEF507BFD25" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Bregetovamacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2827FDEF5556FDC4" box="[830,1100,613,637]" italics="true" pageId="4" pageNumber="1622">Bregetovamacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2B89FD0E507BFD25" author="Hirschmann W" box="[144,353,644,668]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BDF24D27FFEEFFBE2A75FD0E523CFD25" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" box="[364,806,644,668]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Loksamacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2A75FD0E5348FD25" box="[364,594,644,668]" italics="true" pageId="4" pageNumber="1622">Loksamacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2941FD0E523CFD25" author="Hirschmann W" box="[600,806,644,668]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BDF24D27FFEEFFBE2844FD0E507EFD05" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Baloghmacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2844FD0E5557FD25" box="[861,1101,644,668]" italics="true" pageId="4" pageNumber="1622">Baloghmacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2B89FD2E507EFD05" author="Hirschmann W" box="[144,356,676,700]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
as genera as well. This subdivision was based on the shape of the peritreme, of which
<specimenCount id="6CF4FD2DFFEEFFBE2AB1FD4E5307FD64" box="[424,541,708,733]" pageId="4" pageNumber="1622" type="generic" typeStatus="types">four types</specimenCount>
are known in the genus
<taxonomicName id="BDF24D27FFEEFFBE285AFD4E52F1FD65" authorityName="Berlese" authorityYear="1917" box="[835,1003,708,732]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE285AFD4E52F1FD65" box="[835,1003,708,732]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
</taxonomicName>
(see the key).
</paragraph>
<paragraph id="7A4D36A4FFEEFFBE2BA2FC8E5030FC22" blockId="4.[144,1101,166,923]" pageId="4" pageNumber="1622">
According to my assumption
<taxonomicName id="BDF24D27FFEEFFBE2900FC8E53DBFCA5" authorityName="Berlese" authorityYear="1917" box="[537,705,772,796]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE2900FC8E53DBFCA5" box="[537,705,772,796]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
</taxonomicName>
is a Uropodina genus distributed worldwide and comprising four subgenera; accordingly I consider
<taxonomicName id="BDF24D27FFEEFFBE28BCFCAE5028FCE2" authority="Berlese, 1917" authorityName="Berlese" authorityYear="1917" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1622" phylum="Arthropoda" rank="genus">
<emphasis id="4886EAB6FFEEFFBE28BCFCAE5557FC85" box="[933,1101,804,828]" italics="true" pageId="4" pageNumber="1622">Macrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2B89FCC95028FCE2" author="Berlese A" box="[144,306,835,859]" pageId="4" pageNumber="1622" pagination="7 - 16" refId="ref6966" refString="Berlese A. 1917. Intorno agli Uropodidae. Redia. 13: 7 - 16." type="journal article" year="1917">Berlese, 1917</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BDF24D27FFEEFFBE2A52FCCE5254FCE2" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" box="[331,846,835,860]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Bregetovamacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2A52FCCE5343FCE5" box="[331,601,836,860]" italics="true" pageId="4" pageNumber="1622">Bregetovamacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2977FCC95254FCE2" author="Hirschmann W" box="[622,846,835,859]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BDF24D27FFEEFFBE287FFCCE507BFCC2" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Loksamacrodinychus">
<emphasis id="4886EAB6FFEEFFBE287FFCCE5556FCE5" box="[870,1100,836,860]" italics="true" pageId="4" pageNumber="1622">Loksamacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE2B89FCE9507BFCC2" author="Hirschmann W" box="[144,353,867,891]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BDF24D27FFEEFFBE2A84FCEE527DFCC2" authority="Hirschmann, 1979" authorityName="Hirschmann" authorityYear="1979" box="[413,871,867,892]" class="Arachnida" family="Macrodinychidae" genus="Macrodinychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1632" phylum="Arthropoda" rank="subGenus" subGenus="Baloghmacrodinychus">
<emphasis id="4886EAB6FFEEFFBE2A84FCEE5397FCC5" box="[413,653,868,892]" italics="true" pageId="4" pageNumber="1622">Baloghmacrodinychus</emphasis>
<bibRefCitation id="1E634B55FFEEFFBE298FFCE9527DFCC2" author="Hirschmann W" box="[662,871,867,891]" pageId="4" pageNumber="1622" pagination="57 - 70" refId="ref7363" refString="Hirschmann W. 1979. Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann, 1979. Acarol Schrift Vergl Milbenkunde. 26: 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</taxonomicName>
as subgenera in the present work.
</paragraph>
</subSubSection>
</treatment>
</document>

147
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<paragraph id="B5842367A5F76075DB579262AE33D1AD" pageNumber="1297">
<taxonomicName id="0ADD77A34FC1F938446C205851FC1A9E" ID-CoL="LN2J" authority="(Thomas 1916)" baseAuthorityName="Thomas" baseAuthorityYear="1916" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" pageNumber="1297" phylum="Chordata" rank="species" species="mackenziei">
<emphasis id="F8BEFEF47F0A6C6A0B562FAD63E43971" italics="true" pageNumber="1297">Berylmys mackenziei</emphasis>
(Thomas 1916)
</taxonomicName>
</paragraph>
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<paragraph id="BEC0DE0AE822A7450DA155C416107198" pageNumber="1297">
<treatmentCitation id="1C383EBB9547FFD0CF4B8502A2A55E5A" author="Thomas" authority="(Thomas 1916)" baseAuthorityName="Thomas" baseAuthorityYear="1916" class="Mammalia" family="Muridae" genus="Berylmys" journal="J. Bombay Nat. Hist. Soc." kingdom="Animalia" order="Rodentia" page="40" pageNumber="1297" phylum="Chordata" rank="species" species="mackenziei" title="[Berylmys] mackenziei" volume="24" year="1916">
<taxonomicName id="E449DCF72C8B2131F5CA12CB0DE52001" ID-CoL="LN2J" authority="(Thomas 1916)" baseAuthorityName="Thomas" baseAuthorityYear="1916" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" pageNumber="1297" phylum="Chordata" rank="species" species="mackenziei">
<emphasis id="C2D34E01D174AAE37D4B0465051AD35E" italics="true" pageNumber="1297">[Berylmys] mackenziei</emphasis>
(Thomas 1916)
</taxonomicName>
,
<bibCitation id="ABC45028B7AD3298713A62C71E49DB72" author="Thomas" journal="J. Bombay Nat. Hist. Soc." pageNumber="1297" pagination="40" title="[Berylmys] mackenziei" volume="24" year="1916">
J.
<collectingCountry id="61670A3E27F7E27DF3FF703A25B54E44" name="India">Bombay</collectingCountry>
Nat. Hist. Soc., 24: 40
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5809C8D1FA019710EB8F1B4CEBDF3D70" pageNumber="1297" type="materials_examined">
<paragraph id="FF004D230950786CD5C634E32B8656BE" pageNumber="1297">
<emphasis id="A72BC32F57F52CF2518B4C66E4A4739A" inLineHeading="true" pageNumber="1297">Type Locality:</emphasis>
<materialsCitation id="118C53024AE59A1C0BA2B82503F2B482" ID-GBIF-Occurrence="4867324364" country="Myanmar" pageNumber="1297" stateProvince="Chin">
<collectingCountry id="18A302F1E4E11CE1C61E7CDDD3AB9D15" name="Myanmar">Burma</collectingCountry>
,
<collectingRegion id="247FA3CC1E7C933722558E6AF37BE366" country="Myanmar" name="Chin">Chin</collectingRegion>
Hills,
<quantity id="D2C89B09B5BF2700718C93E38FBE6E0B" metricMagnitude="4" metricUnit="m" metricValue="8.04672" unit="mi" value="50.0">50 mi</quantity>
(
<quantity id="6DB8AF89EE654F986E8AAEA636CF8745" metricMagnitude="4" metricUnit="m" metricValue="8.0" unit="km" value="80.0">80 km</quantity>
) west of Kindat.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C22032F6FFE6E73ACC4175EBDDFCF5C7" pageNumber="1297" type="vernacular_names">
<paragraph id="6B38B597BFFA2F16D5D806917FB27C5C" pageNumber="1297">
<emphasis id="29D8A2D1F768CB600F72F1639AACB436" inLineHeading="true" pageNumber="1297">Vernacular Names:</emphasis>
<vernacularName id="F15DB15296FBB1D3AC27E8E048C5BDEA" language="eng" pageNumber="1297">Mackenzie's Berylmys</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="3D02E838D462A057FD4D71A2D923E247" pageNumber="1297" type="synonym_list">
<paragraph id="9E1ACC9459D0828561587B7656E94422" pageNumber="1297">
<emphasis id="06098151173E9067CDA593F3800C519A" inLineHeading="true" pageNumber="1297">Synonyms:</emphasis>
<taxonomicName id="3178371AF00A2C434AAF8683344D5B62" authority="(Thomas 1916)" baseAuthorityName="Thomas" baseAuthorityYear="1916" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" pageNumber="1297" phylum="Chordata" rank="species" species="fea">
<emphasis id="C1567FD7C10B2311ADAE597E610E1283" italics="true" pageNumber="1297">Berylmys fea</emphasis>
(Thomas 1916)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1A761AE9C0B43B17CDC3400A65EDBBDD" pageNumber="1297" type="distribution">
<paragraph id="B4D725E2E67BBC050136FDDB62AB56CD" pageNumber="1297">
<emphasis id="01FB588C0D5A2BEA08E6E798EB0F978A" inLineHeading="true" pageNumber="1297">Distribution:</emphasis>
NE
<collectingCountry id="DC8E49AB7E20B170F51050B6F0BA8724" name="India">India</collectingCountry>
in
<collectingRegion id="1AB0FE9B9AA6EF3129A7F2A1B130F8E7" country="India" name="Meghalaya">Meghalaya</collectingRegion>
(Shillong and Cherrapunji),
<collectingRegion id="2C154BA38C363AE24E3C52C81D01CF7D" country="India" name="Nagaland">Nagaland</collectingRegion>
,
<collectingRegion id="D7E64F3859052A26D0843B0E1BBA62F8" country="India" name="Mizoram">Mizoram</collectingRegion>
(Lushai Hills), and
<collectingRegion id="FB549DF647CCCDFAA239709C816AC591" country="India" name="Manipur">Manipur</collectingRegion>
(Bishenpur and Tamenglong); C and S
<collectingCountry id="09B257D4D66B90078E94366682046C33" name="Myanmar">Burma</collectingCountry>
,
<collectingCountry id="173D3E7C61587BF44674B17FF3B773EB" name="China">China</collectingCountry>
(
<collectingRegion id="94FAE131E338A39C88232C8F28905AA8" country="China" name="Sichuan">Sichuan</collectingRegion>
), and S
<collectingCountry id="DBDF72E7B00040C74D4494FE1E391F3A" name="Vietnam">Vietnam</collectingCountry>
; see
<bibRefCitation id="B66A008EF1AD5097BA836DECBC0AC2CC" author="Musser, G. G. &amp; C. Newcomb" refId="ref252768" refString="Musser, G. G., and C. Newcomb. 1983. Malaysian murids and the giant rat of Sumatra. Bulletin of the American Museum of Natural History, 174: 327 - 598." year="1983">Musser and Newcomb (1983)</bibRefCitation>
and
<bibRefCitation id="F75BB70EDDC054B6D1C65D3DBB4FC74B" author="Agrawal, V. C." refId="ref2022" refString="Agrawal, V. C. 2000. Taxonomic studies on Indian Muridae and Hystricidae (Mammalia: Rodentia). Records of the Zoological Survey of India, Occasional Paper No. 180, 186 pp." year="2000">Agrawal (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31C199B240C40C35AE68DFE22243D3DA" pageNumber="1297" type="conservation">
<paragraph id="C9B5DBA2490749F26C80F964560CAFAD" pageNumber="1297">
<emphasis id="D9A25569FD6521F41D1371A05F41210B" inLineHeading="true" pageNumber="1297">Conservation:</emphasis>
<collectionCode id="D3372FABD1A32E58E130355456C6F6CF">IUCN</collectionCode>
– Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="5EDCA4BC2A790003C0FB25189E4087DD" pageNumber="1297" type="discussion">
<paragraph id="7FB037B34DD8052AD2736492C05FCF8B" pageNumber="1297">
<emphasis id="28E558B9E2036EBB37138288599EB167" inLineHeading="true" pageNumber="1297">Discussion:</emphasis>
Specimens of this distinctive species are usually misidentified as
<taxonomicName id="81CC2DEAA85684DE7F91D6EEA187B66E" baseAuthorityName="Zheng" baseAuthorityYear="1993" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bowersi">
<emphasis id="3F5C5A6C781A0CA4D27E8707C41C385F" italics="true" pageNumber="1297">B. bowersi</emphasis>
</taxonomicName>
; see detailed comparisons in
<bibRefCitation id="9DD832C35A50CFEDDBE1DD48AF8FDC7D" author="Musser, G. G. &amp; C. Newcomb" refId="ref252768" refString="Musser, G. G., and C. Newcomb. 1983. Malaysian murids and the giant rat of Sumatra. Bulletin of the American Museum of Natural History, 174: 327 - 598." year="1983">Musser and Newcomb (1983)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.727.20222" ID-GBIF-Dataset="13853cb6-522e-4f94-b633-c4a51ba63624" ID-PMC="PMC5799789" ID-Pensoft-Pub="1313-2970-727-1" ID-PubMed="29416388" ID-ZBK="12B663F7190040788E1EEF8BAC4DF81B" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-727-1" ModsDocOrigin="ZooKeys 727" ModsDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" checkinTime="1515696595036" checkinUser="pensoft" docAuthor="Wood, Hannah M. &amp; Scharff, Nikolaj" docDate="2017" docId="F25BD83EA8857DD76C9835B9DD9DDE41" docLanguage="en" docName="ZooKeys 727: 1-96" docOrigin="ZooKeys 727" docSource="http://dx.doi.org/10.3897/zookeys.727.20222" docTitle="Eriauchenius pauliani Legendre 1970" docType="treatment" docVersion="4" lastPageNumber="1" masterDocId="FFEBD959ED2FFFAA9450FF8B1957B348" masterDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="1" updateTime="1668165276717" updateUser="ExternalLinkService">
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<mods:title>A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)</mods:title>
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<mods:namePart>Scharff, Nikolaj</mods:namePart>
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<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>727</mods:number>
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<treatment ID-GBIF-Taxon="140245890" LSID="urn:lsid:plazi:treatment:F25BD83EA8857DD76C9835B9DD9DDE41" httpUri="http://treatment.plazi.org/id/F25BD83EA8857DD76C9835B9DD9DDE41" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Legendre, 1970" authorityName="Legendre" authorityYear="1970" class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriauchenius pauliani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pauliani">Eriauchenius pauliani (Legendre, 1970)</taxonomicName>
Figs 18, 32
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Archaeidae" genus="Archaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaea pauliani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pauliani">Archaea pauliani</taxonomicName>
Legendre, 1970: 21, figs 6-7, plate 3, fig. 4.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Male holotype: as
<taxonomicName family="Archaeidae" genus="Archaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaea pauliani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pauliani">Archaea pauliani</taxonomicName>
Legendre, 1970: Madagascar, Andohahelo (Fort Dauphin), 1800 m., Jan /1954, R. Paulian (examined, deposited in MNHN; MNHN 13/1970).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">No other material examined.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Male is distinguished from other &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea bourgini" pageId="0" pageNumber="1" rank="species" species="bourgini">bourgini</taxonomicName>
group&quot; species by the distinctly shaped cephalon, that is triangular with a post-ocular pointed
<normalizedToken originalValue="“head”">&quot;head&quot;</normalizedToken>
(Fig. 18A), and also by the complete fusion of the sternum and carapace. Male is distinguished from other &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea bourgini" pageId="0" pageNumber="1" rank="species" species="bourgini">bourgini</taxonomicName>
group&quot; species except
<taxonomicName lsidName="E. milloti" pageId="0" pageNumber="1" rank="species" species="milloti">E. milloti</taxonomicName>
sp. n. and
<taxonomicName lsidName="E. lukemacaulayi" pageId="0" pageNumber="1" rank="species" species="lukemacaulayi">E. lukemacaulayi</taxonomicName>
sp. n. (and presumably
<taxonomicName lsidName="E. milajaneae" pageId="0" pageNumber="1" rank="species" species="milajaneae">E. milajaneae</taxonomicName>
sp. n., although the male is unknown) by the presence of a lateral process on the conductor (Figs 14
<normalizedToken originalValue="G–I">G-I</normalizedToken>
, 17
<normalizedToken originalValue="D–I">D-I</normalizedToken>
, 18
<normalizedToken originalValue="F–G">F-G</normalizedToken>
, asterisk), which in only
<taxonomicName lsidName="E. pauliani" pageId="0" pageNumber="1" rank="species" species="pauliani">E. pauliani</taxonomicName>
is as wide as long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Male holotype (MNHN 13/1970, from Andohahelo, Fort Dauphin, Madagascar). Total length 2.39, carapace 0.97 long, 0.82 wide. Abdomen 1.30 long, 1.40 high. Carapace tilt angle 62.38°, tilt height (CtH) 2.57, constriction 0.54, head length 1.22, neck length 1.29 (Fig. 2). CtH divided by carapace length 2.65. Cephalon with AME on large bulge, and with postocular pointed head (Fig. 18
<normalizedToken originalValue="A–B">A-B</normalizedToken>
). Cephalon with 4 small post-ocular spines at the crown, not on protrusions. Chelicerae 2.07 long, and with a short spine 0.31 from base of chelicerae that projects perpendicular to the cheliceral cuticle. Femur I 2.99 long. Sternum 0.41 long, 0.65 wide, but completely fused to the carapace; measurements were taken based on approximations of where the original sternum edge would have been. Carapace and sternum reddish brown with white setae, chelicera light reddish brown. Legs beige with reddish brown annulations on tibia and metatarsus. Abdomen beige, mottled with brown and with tufts of white setae (Fig. 18A). Pedipalpal bulb with a membranous sac above the base of the embolus, with a greatly exposed embolus that is encircled by the conductor. Conductor with basal part an uneven ridge shape (see the basal
<normalizedToken originalValue="“c”">&quot;c&quot;</normalizedToken>
in Fig. 18
<normalizedToken originalValue="F–G">F-G</normalizedToken>
), with distal part that is narrow as it encircles the embolus, with a large, broad, bulge before tapering off. Conductor with lateral process that is as wide as long (Fig. 18
<normalizedToken originalValue="F–G">F-G</normalizedToken>
, asterisk). Embolus broad but tapering at tip to a long, fine wire (Fig. 18I); the embolus tip is broken off of the left pedipalp (Fig. 18G, arrow), but is entire in the right pedipalp (Fig. 18I).
</paragraph>
<paragraph pageId="0" pageNumber="1">Female: The female paratype at MNHN was not available for examination.</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 18.
<taxonomicName class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriauchenius pauliani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pauliani">Eriauchenius pauliani</taxonomicName>
(Legendre, 1970), holotype, MNHN.
<normalizedToken originalValue="A–B">A-B</normalizedToken>
male A habitus, lateral view, image reversed B anterior view
<normalizedToken originalValue="C–H">C-H</normalizedToken>
male left pedipalpal bulb C, F prolateral view D, G ventral view, arrow showing the broken tip of the embolus in the left pedipalp E, H retrolateral view I male right pedipalpal bulb, embolus tip close-up showing the unbroken embolus, image reversed. Scale bars: 1 mm (A); 0.25 mm (C).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">No other known material.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="natural history">
<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="1">Specimens collected at 1800 m elevation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Known only from the type locality, Andohahelo, in southeast Madagascar (Fig. 32). The distribution point in Fig. 32 is an approximation since the holotype label does not have latitude/longitude data.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
In
<taxonomicName lsidName="E. pauliani" pageId="0" pageNumber="1" rank="species" species="pauliani">E. pauliani</taxonomicName>
the sternum is fused to the carapace, so the sternum measurements were based on an approximation of where the sternum border would be if this fusion was absent. In the type specimen the embolus tip is broken off in the left pedipalp (Fig. 18G - arrow), however the right pedipalp shows that the embolus tip is curved and much longer (Fig. 18I).
</paragraph>
</subSubSection>
</treatment>
</document>