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<document id="6F9F1BB31AD11710D507687F0522A230" ID-DOI="10.5281/zenodo.174523" ID-GBIF-Dataset="98f1f6af-5b4a-4375-aa8c-5e755359f5df" ID-ISSN="1175-5326" ID-Zenodo-Dep="174523" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459784389820" checkinUser="plazi" docAuthor="Hancock, E. Geoffrey, Marcos-Garcia, M-Angeles &amp; Rotheray, Graham E." docDate="2006" docId="F2211673D34DA42C5436FA5D0500134B" docLanguage="en" docName="zt01351p068.pdf" docOrigin="Zootaxa 1351" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Ptychoptera alexanderi Hancock, sp. nov." docType="treatment" docVersion="7" lastPageNumber="65" masterDocId="0E186E0BD34CA429553EFF9C010C1105" masterDocTitle="Ptychopteridae — a family of flies (Diptera) new to the Neotropical Region and description of a new species" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698218298708" updateUser="plazi">
<mods:mods id="385F6C68BEFAB1FB172649EA5882181C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3E5E9645C30F9CD1644B656282A968BF">
<mods:title id="BAFC64B2EEC80558E617BF5DBC8DF2A2">Ptychopteridae — a family of flies (Diptera) new to the Neotropical Region and description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="3056FC73084DBB9AED6D6080C8A59DEB" type="personal">
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<mods:namePart id="68C9E50741335FDE133CB07E8FA1361C">Hancock, E. Geoffrey</mods:namePart>
</mods:name>
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<mods:namePart id="F8CB862A1237D2F2D566084962DEEF57">Marcos-Garcia, M-Angeles</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="80554E2AA015D41EB923439721C9E619">Rotheray, Graham E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D455A9425950A5D88372081E3914E1A0">text</mods:typeOfResource>
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<mods:title id="A71BE4C16E074BD0EB496306A4692D23">Zootaxa</mods:title>
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<mods:part id="3687768186D3E9AAC29298C7D776609A">
<mods:date id="0068D19C01A4E476C7064CF029D87EFB">2006</mods:date>
<mods:detail id="808B2A8D490C174C70EE8FC136ECB090" type="volume">
<mods:number id="528B64A1102CA3FBFFEB3F6F0FD238F3">1351</mods:number>
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<mods:classification id="06243DCD3476CFBBBE55FF5343AF7F84">journal article</mods:classification>
<mods:identifier id="9E078D610A893F853CAEF8F550421C43" type="DOI">10.5281/zenodo.174523</mods:identifier>
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<treatment id="F2211673D34DA42C5436FA5D0500134B" ID-DOI="http://doi.org/10.5281/zenodo.6262000" ID-GBIF-Taxon="119348743" ID-Zenodo-Dep="6262000" LSID="urn:lsid:plazi:treatment:F2211673D34DA42C5436FA5D0500134B" httpUri="http://treatment.plazi.org/id/F2211673D34DA42C5436FA5D0500134B" lastPageId="5" lastPageNumber="65" pageId="1" pageNumber="62">
<subSubSection id="3292F4EED34DA4285436FA5D03F014DE" box="[264,764,1473,1499]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="7A37A765D34DA4285436FA5D03F014DE" blockId="1.[264,764,1473,1499]" box="[264,764,1473,1499]" pageId="1" pageNumber="62">
<heading id="217F1009D34DA4285436FA5D03F014DE" bold="true" box="[264,764,1473,1499]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="48FC7B77D34DA4285436FA5D03F014DE" bold="true" box="[264,764,1473,1499]" pageId="1" pageNumber="62">
<taxonomicName id="BD88DCE6D34DA4285436FA5D039B14DE" ID-CoL="78N7H" authority="Hancock" authorityName="Hancock" box="[264,663,1473,1499]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D34DA4285436FA5D032D14DE" bold="true" box="[264,545,1473,1499]" italics="true" pageId="1" pageNumber="62">Ptychoptera alexanderi</emphasis>
Hancock
</taxonomicName>
<taxonomicNameLabel id="53CFC60CD34DA42857A0FA5D03F014DE" box="[670,764,1473,1499]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3292F4EED34DA42B5436F99002E4128B" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="7A37A765D34DA42B5436F99002E4128B" blockId="1.[264,1323,1548,1897]" lastBlockId="2.[264,1323,284,1876]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
<emphasis id="48FC7B77D34DA4285436F99000841723" bold="true" box="[264,392,1548,1574]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
This species of
<taxonomicName id="BD88DCE6D34DA4285766F99003EB1720" box="[600,743,1548,1573]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="48FC7B77D34DA4285766F99003EB1720" box="[600,743,1548,1573]" italics="true" pageId="1" pageNumber="62">Ptychoptera</emphasis>
</taxonomicName>
shares the general morphological characters of other known species but exhibits a unique combination of a number of features. The most obvious is the slender male antennae being nearly the same length as the body. This is achieved by the basal flagellomere being twice the length of each of the three or four following flagellomeres, which are about 10 times as long as wide. By comparison, the Nearctic species have these same flagellomeres only two to three times their own width. The clypeus of both sexes is swollen.
<collectingCountry id="029FE7F5D34DA42857F2F96003E11613" box="[716,749,1788,1814]" name="American Samoa" pageId="1" pageNumber="62">As</collectingCountry>
in the majority of western Nearctic species, this new species has cross vein r-m meeting R4 + R5, whereas in the two eastern species,
<taxonomicName id="BD88DCE6D34DA428502BF8B800E4166C" authority="Say" authorityName="Say" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="quadrifasciata">
<emphasis id="48FC7B77D34DA428502BF8B800BF166D" italics="true" pageId="1" pageNumber="62">P. quadrifasciata</emphasis>
Say
</taxonomicName>
and
<taxonomicName id="BD88DCE6D34DA428571BF8D3022F166C" authority="Alexander" authorityName="Alexander" box="[549,803,1871,1897]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="osceola">
<emphasis id="48FC7B77D34DA428571BF8D30392166D" box="[549,670,1871,1896]" italics="true" pageId="1" pageNumber="62">P. osceola</emphasis>
Alexander
</taxonomicName>
, r-m meets Rs before the fork. The wing is strongly patterned with three dark curved bands and a short apical band that marks the upper internal edge of the apical patch. The central pair of bands is curved in opposing directions, having the effect of leaving an almost circular clear area in the middle of the wing centred over the base of the radial cells. There is no pigmentation proximally in the basal cells. Macrotrichia are extensive in all the radial cells (Fig. 1). A marked pattern on the dorsum is provided by a blackish outline to the deep mesonotal suture continued around the dorsal edge to above the wing base, giving a strong blackish W-shaped mark to the dorsum against a paler yellowish brown background. The black shiny anepimeron contrasting with the other pleurae is marked. There is no obvious pruinosity on the thorax; many other species of
<taxonomicName id="BD88DCE6D34EA42B5728FD1803A91398" box="[534,677,644,669]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="48FC7B77D34EA42B5728FD1803A91398" box="[534,677,644,669]" italics="true" pageId="2" pageNumber="63">Ptychoptera</emphasis>
</taxonomicName>
have silvery patterns changing with different viewing angles. There is no auxiliary copulatory organ on the third (visible) abdominal sternite, as in some Palaearctic species. In the male, the extreme length of the lateral arms of the ninth tergite, extending nearly the same distance as the gonostyles is noticeable. Another notable difference in the male genitalia of the Nearctic species
<taxonomicName id="BD88DCE6D34EA42B56AAFCB805511238" box="[916,1117,804,829]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="quadrifasciata">
<emphasis id="48FC7B77D34EA42B56AAFCB805511238" box="[916,1117,804,829]" italics="true" pageId="2" pageNumber="63">P. quadrifasciata</emphasis>
</taxonomicName>
and
<taxonomicName id="BD88DCE6D34EA42B51A6FCB804031238" box="[1176,1295,804,829]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="osceola">
<emphasis id="48FC7B77D34EA42B51A6FCB804031238" box="[1176,1295,804,829]" italics="true" pageId="2" pageNumber="63">P. osceola</emphasis>
</taxonomicName>
is the enlarged ninth sternite that accommodates the large spherical sperm pump. This character is not so strongly developed in
<taxonomicName id="BD88DCE6D34EA42B57D7FCE8028F1288" box="[745,899,884,909]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D34EA42B57D7FCE8028F1288" box="[745,899,884,909]" italics="true" pageId="2" pageNumber="63">P. alexanderi</emphasis>
</taxonomicName>
<emphasis id="48FC7B77D34EA42B56B5FCE802E4128B" bold="true" box="[907,1000,884,910]" pageId="2" pageNumber="63">
<taxonomicNameLabel id="53CFC60CD34EA42B56B5FCE802E4128B" box="[907,1000,884,910]" pageId="2" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3292F4EED34EA42B5406FC0000241789" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="7A37A765D34EA42B5406FC0000241789" blockId="2.[264,1323,284,1876]" pageId="2" pageNumber="63">
<emphasis id="48FC7B77D34EA42B5406FC0000FB12B3" bold="true" box="[312,503,924,950]" pageId="2" pageNumber="63">
<typeStatus id="A53319C7D34EA42B5406FC0000A612B3" box="[312,426,924,950]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Body length 8.0 mm, wing 8.8 mm, antenna 7.0 mm.
<emphasis id="48FC7B77D34EA42B51B2FC0005C312B3" bold="true" box="[1164,1231,924,950]" pageId="2" pageNumber="63">Head</emphasis>
: vertex shining black; occipital region and face brown; clypeus convex and same shade of brown as are palp, scape, pedicel, and basal flagellomere plus extreme base of second flagellomere, remainder of antenna brownish-black; 1st flagellomere twice length of succeeding flagellomere. Antennae of male elongated, nearly equal to body length.
<emphasis id="48FC7B77D34EA42B5436FBFB00691584" bold="true" box="[264,357,1127,1153]" pageId="2" pageNumber="63">Thorax</emphasis>
: shining and mainly yellowish-brown. Blackish in prescutellar area of dorsum, more obviously so when viewed from behind; black border to dorsal suture, joining anteriorly with broader black lateral markings that originate above wing base; poorly defined medial stripe visible within suture when viewed anteriorly. Anepimeron distinctly black, contrasting with remaining yellow pleurae. Halteres yellow in basal quarter, otherwise black. Scutellum yellow; mediotergite black. Mid and hind coxae with illdefined darker streaks of brown; femora yellowish-brown except for narrow apical ring; remaining leg segments brownish-black. Wing with partial black bands and apically darkened; typical wing fold rather obvious, extending from base of anal vein to bend in second cubital (Fig. 1).
<emphasis id="48FC7B77D34EA42B570FFA5303A714EC" bold="true" box="[561,683,1487,1513]" pageId="2" pageNumber="63">Abdomen</emphasis>
: shiny blackish-brown; laterobasal half of tergite 2, basal 3rd of tergite 3, and laterobasal corner of tergite 4 yellowish; sternites yellow. Genitalia with greatly extended lateral arms to tergite 9, as long as gonostyli (
<figureCitation id="E2B3BBE0D34EA42B51F1F9BE04101739" box="[1231,1308,1570,1596]" captionStart="FIGURE 2" captionStartId="3.[264,367,1564,1588]" captionTargetBox="[264,1316,324,1546]" captionTargetId="figure@3.[535,1069,727,1558]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia (dorsal) of Ptychoptera alexanderi sp. nov." httpUri="https://zenodo.org/record/174524/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
). Caudal margin of ninth sternite bearing clasper-like structure that protrudes dorsally (
<figureCitation id="E2B3BBE0D34EA42B51C5F9D600141789" captionStart="FIGURE 3" captionStartId="4.[264,367,727,751]" captionTargetBox="[466,1134,328,726]" captionTargetId="figure@4.[434,1177,288,735]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia (lateral, left side) of Ptychoptera alexanderi sp. nov." httpUri="https://zenodo.org/record/174525/files/figure.png" pageId="2" pageNumber="63">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3292F4EED34EA42B5406F90502501651" pageId="2" pageNumber="63" type="description">
<paragraph id="7A37A765D34EA42B5406F90502501651" blockId="2.[264,1323,284,1876]" pageId="2" pageNumber="63">
<emphasis id="48FC7B77D34EA42B5406F905009617B6" bold="true" box="[312,410,1689,1715]" pageId="2" pageNumber="63">Female.</emphasis>
body length 8.0 mm, wing 9.0 mm, antennae 2.5 mm. General appearance same as male except for short antennae and abdomen more broadly yellowish about base of tergites 3, 4, and 5; downcurved cerci totally yellow. Lighter wing markings may be due to specimen being recently emerged when collected, but degree of change from teneral condition to full pigmentation is unknown (
<figureCitation id="E2B3BBE0D34EA42B5638F8A602431651" box="[774,847,1850,1876]" captionStart="FIGURE 4" captionStartId="4.[264,367,1522,1546]" captionTargetBox="[273,1299,812,1492]" captionTargetId="figure@4.[265,1322,805,1498]" captionTargetPageId="4" captionText="FIGURE 4. Habitus of female Ptychoptera alexanderi sp. nov. The overall appearance is lighter than in the male, partly due to being teneral when collected." httpUri="https://zenodo.org/record/174526/files/figure.png" pageId="2" pageNumber="63">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="2EF7F7EDD34FA42A5436F98002F31731" box="[264,1023,1564,1588]" httpUri="https://zenodo.org/record/174524/files/figure.png" pageId="3" pageNumber="64" targetBox="[264,1316,324,1546]" targetPageId="3">
<paragraph id="7A37A765D34FA42A5436F98002F31731" blockId="3.[264,1023,1564,1588]" box="[264,1023,1564,1588]" pageId="3" pageNumber="64">
<emphasis id="48FC7B77D34FA42A5436F98000861731" bold="true" box="[264,394,1564,1588]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
Male genitalia (dorsal) of
<taxonomicName id="BD88DCE6D34FA42A5799F98102AE1731" box="[679,930,1565,1588]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D34FA42A5799F98102AE1731" box="[679,930,1565,1588]" italics="true" pageId="3" pageNumber="64">Ptychoptera alexanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CFC60CD34FA42A5697F98002F31731" box="[937,1023,1564,1588]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="3292F4EED34FA42A5406F9FB030E1677" pageId="3" pageNumber="64" type="materials_examined">
<paragraph id="7A37A765D34FA42A5406F9FB030E1677" blockId="3.[264,1323,1639,1906]" pageId="3" pageNumber="64">
<emphasis id="48FC7B77D34FA42A5406F9FB03201784" bold="true" box="[312,556,1639,1665]" pageId="3" pageNumber="64">Material examined.</emphasis>
<typeStatus id="A53319C7D34FA42A570AF9F403AD1787" box="[564,673,1640,1666]" pageId="3" pageNumber="64" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="029FE7F5D34FA42A57CCF9F402651787" box="[754,873,1640,1666]" name="Mexico" pageId="3" pageNumber="64">MEXICO</collectingCountry>
, Chiapas, Sierra Madre Mountains, El Triunfo (Biosfera Reserva):
<quantity id="BD700A80D34FA42A5741F90C021B17AF" box="[639,791,1680,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="3" pageNumber="64" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900-2000m</quantity>
,
<geoCoordinate id="1FBCC1A2D34FA42A560FF90C02BC17AF" box="[817,944,1680,1706]" direction="north" orientation="latitude" pageId="3" pageNumber="64" precision="15" value="5.6572223">5º39’26”N</geoCoordinate>
<geoCoordinate id="1FBCC1A2D34FA42A56FBF90C055417AF" box="[965,1112,1680,1706]" direction="west" orientation="longitude" pageId="3" pageNumber="64" precision="15" value="-92.80889">92º48’32”W</geoCoordinate>
, E.G. Hancock, collected by sweeping,
<date id="0E3681A5D34FA42A571AF92403A017D7" box="[548,684,1720,1746]" pageId="3" pageNumber="64" value="2003-07-30">30.vii.2003</date>
, (
<typeStatus id="A53319C7D34FA42A57FDF924022717D7" box="[707,811,1720,1746]" pageId="3" pageNumber="64" type="holotype">holotype</typeStatus>
male deposited in the Hunterian Museum (Zoology), Glasgow,
<collectingCountry id="029FE7F5D34FA42A5739F97C036117FF" box="[519,621,1760,1786]" name="United Kingdom" pageId="3" pageNumber="64">Scotland</collectingCountry>
bearing registration number
<collectionCode id="1C993FA0D34FA42A56E8F97C054A17FF" box="[982,1094,1760,1786]" httpUri="http://grbio.org/cool/p267-b3jh" name="University of Glasgow, Hunterian Museum" pageId="3" pageNumber="64">GLAHM</collectionCode>
: 127118.
<typeStatus id="A53319C7D34FA42A5185F97C042A17FF" box="[1211,1318,1760,1786]" pageId="3" pageNumber="64" type="paratype">Paratype</typeStatus>
: female: same data (also deposited in Hunterian Museum (Zoology), registration number
<collectionCode id="1C993FA0D34FA42A5436F8AC0074164F" box="[264,376,1840,1866]" httpUri="http://grbio.org/cool/p267-b3jh" name="University of Glasgow, Hunterian Museum" pageId="3" pageNumber="64">GLAHM</collectionCode>
: 127119). Both specimens were micropinned in the field and are staged on a polythene foam strip.
</paragraph>
</subSubSection>
<caption id="2EF7F7EDD348A42D5436FD4B056E13EA" box="[264,1122,727,751]" httpUri="https://zenodo.org/record/174525/files/figure.png" pageId="4" pageNumber="65" targetBox="[466,1134,328,726]" targetPageId="4">
<paragraph id="7A37A765D348A42D5436FD4B056E13EA" blockId="4.[264,1122,727,751]" box="[264,1122,727,751]" pageId="4" pageNumber="65">
<emphasis id="48FC7B77D348A42D5436FD4B008613EA" bold="true" box="[264,394,727,751]" pageId="4" pageNumber="65">FIGURE 3.</emphasis>
Male genitalia (lateral, left side) of
<taxonomicName id="BD88DCE6D348A42D5635FD44050A13EA" box="[779,1030,728,751]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D348A42D5635FD44050A13EA" box="[779,1030,728,751]" italics="true" pageId="4" pageNumber="65">Ptychoptera alexanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CFC60CD348A42D5133FD4B056E13EA" box="[1037,1122,727,751]" pageId="4" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="2EF7F7EDD348A42D5436FA6E028B1729" httpUri="https://zenodo.org/record/174526/files/figure.png" pageId="4" pageNumber="65" targetBox="[273,1299,812,1492]" targetPageId="4">
<paragraph id="7A37A765D348A42D5436FA6E028B1729" blockId="4.[264,1322,1522,1580]" pageId="4" pageNumber="65">
<emphasis id="48FC7B77D348A42D5436FA6E0080170F" bold="true" box="[264,396,1522,1546]" pageId="4" pageNumber="65">FIGURE 4.</emphasis>
Habitus of female
<taxonomicName id="BD88DCE6D348A42D575EFA6E0251170C" box="[608,861,1522,1545]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D348A42D575EFA6E0251170C" box="[608,861,1522,1545]" italics="true" pageId="4" pageNumber="65">Ptychoptera alexanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CFC60CD348A42D5658FA6E02B1170F" box="[870,957,1522,1546]" pageId="4" pageNumber="65" rank="species">
<emphasis id="48FC7B77D348A42D5658FA6E02B1170F" bold="true" box="[870,957,1522,1546]" pageId="4" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
The overall appearance is lighter than in the male, partly due to being teneral when collected.
</paragraph>
</caption>
<subSubSection id="3292F4EED348A42D5406F9C3027917CF" pageId="4" pageNumber="65" type="etymology">
<paragraph id="7A37A765D348A42D5406F9C3027917CF" blockId="4.[264,1323,1631,1898]" pageId="4" pageNumber="65">
<emphasis id="48FC7B77D348A42D5406F9C300C8177C" bold="true" box="[312,452,1631,1657]" pageId="4" pageNumber="65">Etymology.</emphasis>
<taxonomicName id="BD88DCE6D348A42D54EEF9FC03E4177C" box="[464,744,1632,1657]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D348A42D54EEF9FC03E4177C" box="[464,744,1632,1657]" italics="true" pageId="4" pageNumber="65">Ptychoptera alexanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CFC60CD348A42D57CBF9C3025B177C" box="[757,855,1631,1657]" pageId="4" pageNumber="65" rank="species">
<emphasis id="48FC7B77D348A42D57CBF9C3025B177C" bold="true" box="[757,855,1631,1657]" pageId="4" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
is named in memory of Charles Paul Alexander (
<date id="0E3681A5D348A42D54A0F914032E17A7" box="[414,546,1672,1698]" pageId="4" pageNumber="65" value="1889" valueMax="1982">1889-1982</date>
), the much admired and prolific New World nematoceran systematist, who included this family in his research.
</paragraph>
</subSubSection>
<subSubSection id="3292F4EED348A42C5406F94B0500134B" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65" type="biology_ecology">
<paragraph id="7A37A765D348A42C5406F94B0500134B" blockId="4.[264,1323,1631,1898]" lastBlockId="5.[264,1323,284,590]" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65">
<emphasis id="48FC7B77D348A42D5406F94B009017F4" bold="true" box="[312,412,1751,1777]" pageId="4" pageNumber="65">Biology.</emphasis>
The adults of
<taxonomicName id="BD88DCE6D348A42D5771F944026917F4" box="[591,869,1752,1777]" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
<emphasis id="48FC7B77D348A42D5771F944026917F4" box="[591,869,1752,1777]" italics="true" pageId="4" pageNumber="65">Ptychoptera alexanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CFC60CD348A42D5651F94B02C317F4" box="[879,975,1751,1777]" pageId="4" pageNumber="65" rank="species">
<emphasis id="48FC7B77D348A42D5651F94B02C317F4" bold="true" box="[879,975,1751,1777]" pageId="4" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
were swept from fairly rank streamside vegetation, where the stream margin comprised coarse sand grading to mud. Adult ptychopterids are generally found near running water or in marshy areas, and rest in dense waterside vegetation (
<bibRefCitation id="1E19DA94D348A42D576CF8CC03E5166F" author="Stubbs" box="[594,745,1872,1898]" pageId="4" pageNumber="67" refString="Stubbs, A. E. (1993) Provisional Atlas of the Ptychopterid Craneflies (Diptera, Ptychopteridae) of Britain and Ireland. Biological Records Centre, Institute of Ecology, Huntingdon, 34 pp." type="book" year="1993">Stubbs 1993</bibRefCitation>
). The larvae are associated with saturated mud or soils near water (
<bibRefCitation id="1E19DA94D349A42C54E7FE8003701033" author="Brindle" box="[473,636,284,310]" pageId="5" pageNumber="67" refString="Brindle, A. (1962) Taxonomic notes on the larvae of British Diptera, 9: the family Ptychopteridae. Entomologist, 95, 212 - 216." type="journal article" year="1962">Brindle 1962</bibRefCitation>
). They are considerably elongated caudally, presenting a superficial resemblance to the long-tailed larvae of the
<taxonomicName id="BD88DCE6D349A42C5698FED805B0105B" box="[934,1212,324,350]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Eristalinae">Eristalinae (Syrphidae)</taxonomicName>
. To cope with respiration after pupation, the right thoracic breathing horn is greatly elongated, up to twice the length of the pupa, whereas the other horn is atrophied. These and other observations (
<bibRefCitation id="1E19DA94D349A42C5494FE20036610D3" author="Alexander" box="[426,618,444,470]" pageId="5" pageNumber="67" refString="Alexander, C. P. (1920) The craneflies of New York. Part 2. Biology and phylogeny. Memoirs of the Cornell University Agricultural Experimental Station, 38, 695 - 1133." type="journal article" year="1920">Alexander 1920</bibRefCitation>
,
<bibRefCitation id="1E19DA94D349A42C5748FE2003B810D3" author="Alexander" box="[630,692,444,470]" pageId="5" pageNumber="67" refString="Alexander, C. P. (1981) Ptychopteridae, 22. In McAlpine, J. F., Peterson, B. V., Shewell, G. E., Teskey, H. J., Vockeroth, J. R. &amp; Wood, D. M. (Eds.), Manual of Nearctic Diptera. Volume 1. Monograph No. 27. Research Branch, Agriculture Canada, Ottawa. McAlpine, et al. (Editor). Manual of Nearctic Diptera, pp. 325 - 328." type="book chapter" year="1981">1981</bibRefCitation>
;
<bibRefCitation id="1E19DA94D349A42C5781FE20025010D3" author="Brindle" box="[703,860,444,470]" pageId="5" pageNumber="67" refString="Brindle, A. (1962) Taxonomic notes on the larvae of British Diptera, 9: the family Ptychopteridae. Entomologist, 95, 212 - 216." type="journal article" year="1962">Brindle 1962</bibRefCitation>
;
<bibRefCitation id="1E19DA94D349A42C5659FE20052010D3" author="Hodkinson" box="[871,1068,444,470]" pageId="5" pageNumber="67" refString="Hodkinson, I. D. (1973) The immature stages of Ptychoptera lenis lenis (Diptera, Ptychopteridae) with notes on their biology. Canadian Entomologist, 105, 1091 - 1099." type="journal article" year="1973">Hodkinson 1973</bibRefCitation>
;
<bibRefCitation id="1E19DA94D349A42C5106FE2005CF10D3" author="Stubbs" box="[1080,1219,444,470]" pageId="5" pageNumber="67" refString="Stubbs, A. E. (1993) Provisional Atlas of the Ptychopterid Craneflies (Diptera, Ptychopteridae) of Britain and Ireland. Biological Records Centre, Institute of Ecology, Huntingdon, 34 pp." type="book" year="1993">Stubbs1993</bibRefCitation>
) support the idea that this new species was found in a typical situation and probably was breeding in the stream margins near where the adults were found. Notwithstanding these conditions, no other specimens could be collected despite considerable effort.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">Figs 23</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">, 24</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31K, L</figureCitation>
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<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Pygofer (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23G</figureCitation>
) with apex slightly tapered and rounded. Subgenital plate (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23H</figureCitation>
) with small group of filiform setae on apex. Style, in lateral view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23K</figureCitation>
), blade with ventral margin not serrated; apex hook-shaped. Aedeagus (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23L, M</figureCitation>
) apodemal processes narrow basally, curved in right angle dorsally; external surface, in posterior view, bearing a broad process, rounded apically; shaft apical third broadened, posterior surface with several minute spiniform processes; apex bearing pair of long processes directed ventrally, with half length of shaft.
</paragraph>
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<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 7.9 mm; paratypes, males (n = 47) 7.5-7.9 mm, females (n = 3) 8.0-8.3 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23B</figureCitation>
), frons 1.2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus 1.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; lateral margins parallel; apex straight. Head, in lateral view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23C</figureCitation>
), crown-face transition thin, with 4 transverse carinae; clypeus not inflated. Profemur AV row with 4-5 setae; PV row with 1-2 setae. Protibia PD row with 3 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 22-23, 12 and 14-15 macrosetae respectively. Metatarsomere I 4
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 5 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23A-D</figureCitation>
,
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31K, L</figureCitation>
) reddish-brown. Head (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocelli. Face (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23B</figureCitation>
) without maculae; gena, lorum and clypeus yellowish. Pronotum (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23A</figureCitation>
) with black punctures; pair of circular black spots, near anterior margin, behind eye; proepimeron (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23C</figureCitation>
), almost entirely black; lateral carina yellowish. Forewing (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23D</figureCitation>
) with black maculae on apex of anal veins, cross veins of discal and apical cells; transverse brown stripe over subapical cells. Metatibia (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23E</figureCitation>
) 1.6
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin rounded, slightly more produced medially than near lateral angles. Valve (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23F</figureCitation>
) 2.5
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; dorsal margin slightly excavated; posterior margin rounded. Pygofer, in lateral view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23G</figureCitation>
), 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin, extending ventrally for 1/3 height of lateral lobe; anteroventral margin rounded, posterodorsal and posteroventral margins straight; apex slightly tapered, subacute. Subgenital plate, in lateral view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23G</figureCitation>
), short, not reaching pygofer apex; in ventral view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23H</figureCitation>
), elongated, 4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near median third; outer margin with minute setae, apex with small group of filiform setae; inner margin rounded; outer margin straight; apex rounded, weakly tapered. Connective (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23I</figureCitation>
) wider than long; anterior margin excavated; dorsal keel and stem reduced; stem wide. Style, in dorsal view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23J</figureCitation>
), with outer lobe reduced and rounded; in lateral view (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23K</figureCitation>
), blade thin and curved dorsally, slightly wider near base and tapered toward apex; ventral margin not serrated; apex acute, hook-shaped, curved ventrally. Aedeagus (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23L, M</figureCitation>
) preatrium reduced; dorsal apodeme with dorsal margin straight, lateral margins slightly produced laterally; apodemal processes narrow basally, curved in right angle dorsally in the basal third; external surface, in posterior view, bearing broad process, rounded apically; shaft symmetrical, strongly curved dorsally near base, with a basolateral expansion that surrounds the previous portion of the shaft; apical third broadened, posterior surface with several minute spiniform processes; apex bearing pair of long processes directed ventrally, with half length of shaft. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female terminalia">
<paragraph pageId="0" pageNumber="79">Female terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VII (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24A</figureCitation>
) 1.7
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterolateral angles slightly surpassing posterior margin; posterior margin weakly excavated each side of median lobe which occupies the median third and is V-shape notched medially. Internal sternite VIII membranous. Pygofer (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24A, B</figureCitation>
) 1.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; ventral margin slightly rounded; dorsal margin straight, with apical portion excavated; apex obliquely truncate. First valvifer (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24C</figureCitation>
) subquadrate, 1.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; anterior and dorsal margins straight; posterior margin rounded. First valvula (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24C</figureCitation>
) 7
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24D</figureCitation>
) tapered and acute. Second valvula (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24E</figureCitation>
) 4.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; wider after mid-length; dorsal protuberance developed and rounded; dorsal margin (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24F</figureCitation>
) teeth reduced, only an undulation present subapically; apex abruptly tapered, subacute. Second valvifer (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24G</figureCitation>
) 2.7
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Gonoplac (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24G</figureCitation>
) 3.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24H</figureCitation>
) rounded. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="A. C. Domahovski" country="Brazil" county="Brasil" elevation="365" latitude="-11.5841" location="Fazenda Continental" longLatPrecision="7" longitude="-55.3003" municipality="Claudia" specimenCount="1" stateProvince="Mato Grosso" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male:
<collectingCountry name="Brazil">Brazil</collectingCountry>
:
<collectingRegion country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
</emphasis>
: &quot;
<collectingCounty>Brasil</collectingCounty>
, MT,
<normalizedToken originalValue="Cláudia">
<collectingMunicipality>Claudia</collectingMunicipality>
</normalizedToken>
,\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:6F046C0EB413B7B5E801CA0874E45FD1" country="Brazil" county="Brasil" latitude="-11.5841" longLatPrecision="7" longitude="-55.3003" municipality="Claudia" name="Fazenda Continental" stateProvince="Mato Grosso">Fazenda Continental</location>
,\
<geoCoordinate degrees="11.5841" direction="south" orientation="latitude" precision="5" value="-11.5841">11.5841°S</geoCoordinate>
<geoCoordinate degrees="55.3003" direction="west" orientation="longitude" precision="5" value="-55.3003">55.3003°W</geoCoordinate>
,\
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.65" unit="m" value="365.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.65" unit="m" value="365.0">365m</elevation>
</quantity>
, sweep, 17-19.vi.\ 2017,
<collectorName>A.C. Domahovski</collectorName>
&quot; (DZUP)
</materialsCitation>
.
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="E. S. E. C. Rio Ronuro" country="Brazil" elevation="330" latitude="-13.1122" location="Nova Ubirata" longLatPrecision="7" longitude="-54.4436" municipality="Brasil" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Paratypes">
<typeStatus>Paratypes</typeStatus>
</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
, &quot;
<collectingMunicipality>Brasil</collectingMunicipality>
, MT,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:BC9CC6EF1C3EA614E3E5F9C7FBA68DC5" country="Brazil" latitude="-13.1122" longLatPrecision="7" longitude="-54.4436" municipality="Brasil" name="Nova Ubirata" stateProvince="Paratypes">
Nova
<normalizedToken originalValue="Ubiratã">Ubirata</normalizedToken>
</location>
,\
<collectorName>E.S.E.C. Rio Ronuro</collectorName>
,\
<geoCoordinate degrees="13.1122" direction="south" orientation="latitude" precision="5" value="-13.1122">13.1122°S</geoCoordinate>
<geoCoordinate degrees="54.4436" direction="west" orientation="longitude" precision="5" value="-54.4436">54.4436°W</geoCoordinate>
,\
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.3" unit="m" value="330.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.3" unit="m" value="330.0">330m</elevation>
</quantity>
, malaise, 11-16.vi.\ 2017, Cavichioli, Melo,\ Domahovski &amp; Muniz&quot; (DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2017-06-21" collectingDateMax="2017-06-25" collectingDateMin="2017-06-21" country="Brazil" county="Brasil" elevation="240" latitude="-9.4517" location="Est." longLatPrecision="7" longitude="-55.8396" municipality="Parque" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
, MT, Novo Mundo, Pq. [
<collectingMunicipality>Parque</collectingMunicipality>
]\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:F89B3D62E54C2F1134843775C17D5720" country="Brazil" county="Brasil" latitude="-9.4517" longLatPrecision="7" longitude="-55.8396" municipality="Parque" name="Est." stateProvince="Paratypes">Est.</location>
[
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:47901F90CEF9A5684FDD412AD18B2E78" country="Brazil" county="Brasil" latitude="-9.4517" longLatPrecision="7" longitude="-55.8396" municipality="Parque" name="Estadual" stateProvince="Paratypes">Estadual</location>
] do
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:8B2B987BF28D373AB3E85C3B72296ABD" country="Brazil" county="Brasil" latitude="-9.4517" longLatPrecision="7" longitude="-55.8396" municipality="Parque" name="Cristalino" stateProvince="Paratypes">Cristalino</location>
,
<geoCoordinate degrees="09.4517" direction="south" orientation="latitude" precision="5" value="-9.4517">09.4517°S</geoCoordinate>
<geoCoordinate degrees="55.8396" direction="west" orientation="longitude" precision="5" value="-55.8396">55.8396°W</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.4" unit="m" value="240.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.4" unit="m" value="240.0">240m</elevation>
</quantity>
, malaise,\
<collectingDate value="2017-06-21" valueMax="2017-06-25" valueMin="2017-06-21">21-25.vi.2017</collectingDate>
, Cavichioli,\ Melo, Domahovski &amp; Muniz&quot; (DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" county="Brasil" elevation="240" latitude="-9.4517" location="Pq. Est. do Cristalino" longLatPrecision="7" longitude="-55.8396" municipality="Novo Mundo" specimenCount="1" specimenCount-female="1" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
, MT,
<collectingMunicipality>Novo Mundo</collectingMunicipality>
,\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:08B035719A67CDC776110522D79A88C1" country="Brazil" county="Brasil" latitude="-9.4517" longLatPrecision="7" longitude="-55.8396" municipality="Novo Mundo" name="Pq. Est. do Cristalino" stateProvince="Rondonia">Pq. Est. do Cristalino</location>
,\
<geoCoordinate degrees="09.4517" direction="south" orientation="latitude" precision="5" value="-9.4517">09.4517°S</geoCoordinate>
<geoCoordinate degrees="55.8396" direction="west" orientation="longitude" precision="5" value="-55.8396">55.8396°W</geoCoordinate>
,\
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.4" unit="m" value="240.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.4" unit="m" value="240.0">240m</elevation>
</quantity>
,
<normalizedToken originalValue="luminárias">luminarias</normalizedToken>
- aloja-\ mento AC Domahovski&quot; (DZUP). -
<emphasis bold="true" pageId="0" pageNumber="79">
<normalizedToken originalValue="Rondônia">
<collectingRegion country="Brazil" name="Rondonia">Rondonia</collectingRegion>
</normalizedToken>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2011-03-28" collectingDateMax="2011-04-08" collectingDateMin="2011-03-28" country="Brazil" county="Brasil" latitude="-9.591389" location="Porto" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" specimenCount="12" specimenCount-male="12" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">6♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:AAE61452C537A9A72E50F0B12B6836A7" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Porto" stateProvince="Rondonia">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:1886EC7B6670DCD345B3B03116798494" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Velho" stateProvince="Rondonia">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:B029436530D08E90D13BD0BCF9A6E301" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Rio Madeira" stateProvince="Rondonia">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="29" value="-9.591389">09°35′29″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="57" value="-65.049164">65°02′57″W</geoCoordinate>
\ Malaise
<collectingDate value="2011-03-28" valueMax="2011-04-08" valueMin="2011-03-28">28.III-8.IV.2011</collectingDate>
\ Silva &amp; Albuquerque&quot; (
<specimenCount type="male">3♂</specimenCount>
DZRJ,
<specimenCount type="male">3♂</specimenCount>
MNRJ)
</materialsCitation>
;
<materialsCitation collectingDate="2011-01-05" collectingDateMax="2011-01-17" collectingDateMin="2011-01-05" country="Brazil" county="Brasil" latitude="-9.635" location="Porto" longLatPrecision="21" longitude="-65.45305" municipality="Rondonia" specimenCount="7" specimenCount-male="7" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">7♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:5F398A82F85521B2892FAEBB8AAE0E09" country="Brazil" county="Brasil" latitude="-9.635" longLatPrecision="21" longitude="-65.45305" municipality="Rondonia" name="Porto" stateProvince="Rondonia">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:87540B5EAF5499E01ADC7D761D9713FF" country="Brazil" county="Brasil" latitude="-9.635" longLatPrecision="21" longitude="-65.45305" municipality="Rondonia" name="Velho" stateProvince="Rondonia">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:25C40D966F1969BBEDA497C580BE24FA" country="Brazil" county="Brasil" latitude="-9.635" longLatPrecision="21" longitude="-65.45305" municipality="Rondonia" name="Rio Madeira" stateProvince="Rondonia">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="38" orientation="latitude" precision="15" seconds="06" value="-9.635">09°38′06″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="27" orientation="longitude" precision="15" seconds="11" value="-65.45305">65°27′11″W</geoCoordinate>
\ Malaise
<collectingDate value="2011-01-05" valueMax="2011-01-17" valueMin="2011-01-05">05-17.I.2011</collectingDate>
\ RRSilva &amp; RSProbst leg&quot; (MZSP)
</materialsCitation>
;
<materialsCitation collectingDate="2011-06-18" collectingDateMax="2011-06-29" collectingDateMin="2011-06-18" country="Brazil" county="Brasil" latitude="-9.598333" location="Porto" longLatPrecision="21" longitude="-65.04806" municipality="Rondonia" specimenCount="2" specimenCount-male="2" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">2♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:D093EB8EE77E4038005A9B7D62FFBE6C" country="Brazil" county="Brasil" latitude="-9.598333" longLatPrecision="21" longitude="-65.04806" municipality="Rondonia" name="Porto" stateProvince="Rondonia">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:F0512BDC47863A01AC89AFFAB71D0CC4" country="Brazil" county="Brasil" latitude="-9.598333" longLatPrecision="21" longitude="-65.04806" municipality="Rondonia" name="Velho" stateProvince="Rondonia">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:4B73466DBF64091A73A2A5AAE569F434" country="Brazil" county="Brasil" latitude="-9.598333" longLatPrecision="21" longitude="-65.04806" municipality="Rondonia" name="Rio Madeira" stateProvince="Rondonia">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="54" value="-9.598333">09°35′54″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="53" value="-65.04806">65°02′53″W</geoCoordinate>
\ Malaise
<collectingDate value="2011-06-18" valueMax="2011-06-29" valueMin="2011-06-18">18-29.VI.2011</collectingDate>
\ Albuquerque &amp; Ferreira&quot; (DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" county="Brasil" latitude="-9.591389" location="Porto" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" specimenCount="1" specimenCount-male="1" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="29" value="-9.591389">09°35′29″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="57" value="-65.049164">65°02′57″W</geoCoordinate>
(DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" county="Brasil" latitude="-9.435555" location="Porto" longLatPrecision="21" longitude="-64.802505" municipality="Rondonia" specimenCount="1" specimenCount-female="1" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, same data as preceding except
<geoCoordinate degrees="09" direction="south" minutes="26" orientation="latitude" precision="15" seconds="08" value="-9.435555">09°26′08″S</geoCoordinate>
<geoCoordinate degrees="64" direction="west" minutes="48" orientation="longitude" precision="15" seconds="9" value="-64.802505">64°48′9″W</geoCoordinate>
(DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2012-01-04" collectingDateMax="2012-01-17" collectingDateMin="2012-01-04" collectorName="W. Datilo, G. R. Mazao" country="Brazil" county="Brasil" latitude="-9.591389" location="Porto" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" specimenCount="32" specimenCount-male="32" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">16♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:D4104462AF2968F4CAC0F2901A2CBAE3" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Porto" stateProvince="Rondonia">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:A168CE8197E4F9A46074A7E69B7F7A5F" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Velho" stateProvince="Rondonia">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:3FEE604FBD4D23AFE787EDB28022CE61" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Rio Madeira" stateProvince="Rondonia">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="29" value="-9.591389">09°35′29″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="57" value="-65.049164">65°02′57″W</geoCoordinate>
\ Malaise
<collectingDate value="2012-01-04" valueMax="2012-01-17" valueMin="2012-01-04">04-17.I.2012</collectingDate>
\
<collectorName>W. Datilo</collectorName>
&amp;
<collectorName>
G. R.
<normalizedToken originalValue="Mazão”">Mazao&quot;</normalizedToken>
</collectorName>
(
<specimenCount type="male">6♂</specimenCount>
DZUP,
<specimenCount type="male">10♂</specimenCount>
MZSP)
</materialsCitation>
;
<materialsCitation collectingDate="2012-09-04" collectingDateMax="2012-09-18" collectingDateMin="2012-09-04" country="Brazil" county="Brasil" latitude="-9.4372225" location="Porto" longLatPrecision="21" longitude="-64.83278" municipality="Rondonia" specimenCount="8" specimenCount-male="8" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">8♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:C6FC9FEF44387C529B159CF9ABD87AE1" country="Brazil" county="Brasil" latitude="-9.4372225" longLatPrecision="21" longitude="-64.83278" municipality="Rondonia" name="Porto" stateProvince="Rondonia">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:A5DCEF65BD1F93C379C042604AA16F08" country="Brazil" county="Brasil" latitude="-9.4372225" longLatPrecision="21" longitude="-64.83278" municipality="Rondonia" name="Velho" stateProvince="Rondonia">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:F221A4B43983541CAD6C2BC262CEDB5F:4B7D001579997E1518CF7C918606CE7D" country="Brazil" county="Brasil" latitude="-9.4372225" longLatPrecision="21" longitude="-64.83278" municipality="Rondonia" name="Rio Madeira" stateProvince="Rondonia">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="26" orientation="latitude" precision="15" seconds="14" value="-9.4372225">09°26′14″S</geoCoordinate>
<geoCoordinate degrees="64" direction="west" minutes="49" orientation="longitude" precision="15" seconds="58" value="-64.83278">64°49′58″W</geoCoordinate>
\ Malaise
<collectingDate value="2012-09-04" valueMax="2012-09-18" valueMin="2012-09-04">04-18.IX.2012</collectingDate>
\ MA
<normalizedToken originalValue="Ullysséa">Ullyssea</normalizedToken>
&amp; LP Prado&quot; (MZSP)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" county="Brasil" latitude="-9.635" location="Porto" longLatPrecision="21" longitude="-65.45305" municipality="Rondonia" specimenCount="3" specimenCount-male="3" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">3♂</specimenCount>
, same data as preceding except
<geoCoordinate degrees="09" direction="south" minutes="38" orientation="latitude" precision="15" seconds="06" value="-9.635">09°38′06″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="27" orientation="longitude" precision="15" seconds="11" value="-65.45305">65°27′11″W</geoCoordinate>
(MZSP)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" county="Brasil" latitude="-9.591389" location="Porto" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" specimenCount="1" specimenCount-male="1" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="29" value="-9.591389">09°35′29″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="57" value="-65.049164">65°02′57″W</geoCoordinate>
(MZSP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the aedeagus with apodemal processes bearing projections at the mid-length of external surface resembling ears.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana oricula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="oricula">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana oricula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
) was strongly supported as sister to
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. taurina" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="taurina">
<emphasis italics="true" pageId="0" pageNumber="79">C. taurina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
(SR = 99). This relationship was supported by one synapomorphy, 137:0, aedeagal shaft with a conspicuous basolateral expansion that surrounds the previous portion of the shaft, and eight homoplastic characters, as for example, 87:0, valve more than 2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long and 133:1, apodemal processes of aedeagus, having a lateral process on external margin near mid-length.
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana oricula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="oricula">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana oricula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can be easily differentiated by the style (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23K</figureCitation>
) with a hook-shaped apex, the apodemal processes of aedeagus (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23L, M</figureCitation>
) with the process on external surface rounded, the aedeagal shaft with several minute spiniform processes on posterior surface, and the presence of a pair of long apical processes directed ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>

289
data/F2/21/F1/F221F15EDE11CB09FF6CFDE87618F876.xml Normal file
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<document id="334E2ECF37C34739ED88A333B1B92164" ID-DOI="10.5281/zenodo.176539" ID-GBIF-Dataset="86f7c9fb-723c-4561-818a-5fffd2d9307c" ID-ISSN="1175-5326" ID-Zenodo-Dep="176539" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459835377388" checkinUser="plazi" docAuthor="Rédei, Dávid" docDate="2007" docId="F221F15EDE11CB09FF6CFDE87618F876" docLanguage="en" docName="zt01465p053.pdf" docOrigin="Zootaxa 1465" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Enigmocephala Rédei, 2007, gen. n." docType="treatment" docVersion="8" lastPageNumber="51" masterDocId="0E188926DE10CB0DFFFBFFFA775EFF87" masterDocTitle="A new genus of tribelocephaline assassin bugs from Borneo (Hemiptera: Heteroptera: Reduviidae)" masterLastPageNumber="53" masterPageNumber="47" pageNumber="48" updateTime="1698223542698" updateUser="plazi">
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<mods:title id="4C297D67A4612C20D9182177F3C272A4">A new genus of tribelocephaline assassin bugs from Borneo (Hemiptera: Heteroptera: Reduviidae)</mods:title>
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<mods:namePart id="93FDCDDC9E4CF977E763C808369B3F6E">Rédei, Dávid</mods:namePart>
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<mods:date id="DEAD62E34495CC318B5FBFEB33AC0458">2007</mods:date>
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<treatment id="F221F15EDE11CB09FF6CFDE87618F876" ID-DOI="http://doi.org/10.5281/zenodo.6247065" ID-GBIF-Taxon="119358755" ID-Zenodo-Dep="6247065" LSID="urn:lsid:plazi:treatment:F221F15EDE11CB09FF6CFDE87618F876" httpUri="http://treatment.plazi.org/id/F221F15EDE11CB09FF6CFDE87618F876" lastPageId="4" lastPageNumber="51" pageId="1" pageNumber="48">
<subSubSection id="329213C3DE11CB0CFF6CFDE876F1FDAB" box="[151,431,530,556]" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="7A374048DE11CB0CFF6CFDE876F1FDAB" blockId="1.[151,431,530,556]" box="[151,431,530,556]" pageId="1" pageNumber="48">
<heading id="217FF724DE11CB0CFF6CFDE876F1FDAB" bold="true" box="[151,431,530,556]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="48FC9C5ADE11CB0CFF6CFDE876F1FDAB" bold="true" box="[151,431,530,556]" pageId="1" pageNumber="48">
<taxonomicName id="BD883BCBDE11CB0CFF6CFDE87608FDAB" ID-CoL="4BW2" box="[151,342,530,556]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="48FC9C5ADE11CB0CFF6CFDE87608FDAB" bold="true" box="[151,342,530,556]" italics="true" pageId="1" pageNumber="48">Enigmocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CF2121DE11CB0CFEA7FDE876F1FDAB" box="[348,431,530,556]" pageId="1" pageNumber="48" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE11CB0CFF6CFDA775F3FD68" pageId="1" pageNumber="48" type="diagnosis">
<paragraph id="7A374048DE11CB0CFF6CFDA775F3FD68" blockId="1.[151,1437,605,2031]" pageId="1" pageNumber="48">
<emphasis id="48FC9C5ADE11CB0CFF6CFDA77649FDF0" bold="true" box="[151,279,605,631]" pageId="1" pageNumber="48">Diagnosis.</emphasis>
Recognized among other genera of Tribelocephalinae by the combination of the following characters: labium very robust, being widest near middle of second visible segment and gradually narrowed apicad; apical labial segment bearing finger-like projections apically; legs short, stout, tibiae strongly curved apically; and abdominal sternites nearly entirely fused.
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE11CB0CFF3DFD0776D5F868" pageId="1" pageNumber="48" type="description">
<paragraph id="7A374048DE11CB0CFF3DFD0775E0FC90" blockId="1.[151,1437,605,2031]" box="[198,702,765,791]" pageId="1" pageNumber="48">
<emphasis id="48FC9C5ADE11CB0CFF3DFD077601FC90" bold="true" box="[198,351,765,791]" pageId="1" pageNumber="48">Description.</emphasis>
Based on an apterous female.
</paragraph>
<paragraph id="7A374048DE11CB0CFF3CFCDF76D5F868" blockId="1.[151,1437,605,2031]" pageId="1" pageNumber="48">
<emphasis id="48FC9C5ADE11CB0CFF3CFCDF7656FCB8" bold="true" box="[199,264,805,831]" pageId="1" pageNumber="48">Body</emphasis>
(
<figureCitation id="E2B35CCDDE11CB0CFEE0FCDF763FFCB8" box="[283,353,805,831]" captionStart="FIGURE 1" captionStartId="2.[151,254,1949,1973]" captionTargetBox="[309,1268,221,1895]" captionTargetId="figure@2.[281,1306,197,1923]" captionTargetPageId="2" captionText="FIGURE 1. Enigmocephala deinorhyncha gen. n., sp. n., female holotype." httpUri="https://zenodo.org/record/176540/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
) robust, heavily sclerotized, with rather short appendages; body surface nearly bare, with short, rather stiff, adpressed pilosity on legs and several distinct areas bearing short, dense pubescence.
<emphasis id="48FC9C5ADE11CB0CFAA2FCB772C2FCE0" bold="true" box="[1369,1436,845,871]" pageId="1" pageNumber="48">Head</emphasis>
(
<figureCitation id="E2B35CCDDE11CB0CFF64FC8F764EFC08" box="[159,272,885,911]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Figs. 2–5</figureCitation>
) oval, furnished with a transverse, collar-like pubescent area at base, provided with a pair of longitudinal fringe-like structures composed of long setae on ventral surface; with a deep interocular furrow surrounding a transverse, oval area between eyes; with two tubercle-like elevations in front of interocular furrow; area anterior to antenniferous tubercles strongly declivent, anteriorly flat, clypeus distinctly elevated, laterally somewhat flattened, declivent anteriorly; neck narrow (
<figureCitation id="E2B35CCDDE11CB0CFCD6FBEF7426FBA8" box="[813,888,1045,1071]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Fig. 4</figureCitation>
). Eyes relatively small, flat, barely projecting laterally from surface of head.
<emphasis id="48FC9C5ADE11CB0CFDFDFBC7752FFBD0" bold="true" box="[518,625,1085,1111]" pageId="1" pageNumber="48">Antenna</emphasis>
(
<figureCitation id="E2B35CCDDE11CB0CFD78FBC77594FBD0" box="[643,714,1085,1111]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Fig. 2</figureCitation>
) rather short; scape strongly widened, distinctly shorter than head, bearing a longitudinal pubescent area laterally; pedicel distinctly longer than scape, cylindrical, strongly curved near base; flagellum gracile, subdivided into six secondary segments (apparently basiflagellomere into two and distiflagellomere into four).
<emphasis id="48FC9C5ADE11CB0CFDA7FB4F75E1FB48" bold="true" box="[604,703,1205,1231]" pageId="1" pageNumber="48">Labium</emphasis>
(
<figureCitation id="E2B35CCDDE11CB0CFD28FB4F7418FB48" box="[723,838,1205,1231]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Figs. 3–6</figureCitation>
) very robust, with three visible segments (corresponding to segments II–IV), first visible segment (II) laterally flattened in basal half, wide and cylindrical in apical half, extending to about middle of eyes; second visible segment (III) laterally strongly flattened except its base, very strongly narrowing apically, extending between fore coxae; third visible segment (IV) extremely short, with short, dorsal and ventral apical projections (
<figureCitation id="E2B35CCDDE11CB0CFCC0FAAF74D8FAE8" box="[827,902,1365,1391]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Fig. 6</figureCitation>
).
<emphasis id="48FC9C5ADE11CB0CFC5BFAAF74A3FAE8" bold="true" box="[928,1021,1365,1391]" pageId="1" pageNumber="48">Thorax</emphasis>
with more or less wide stripes or patches of dense pubescence between tergum and pleuron, between pleurites, and between metapleura and abdomen (
<figureCitation id="E2B35CCDDE11CB0CFEE9FA5F7606FA38" box="[274,344,1445,1471]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Fig. 3</figureCitation>
). Pronotum (
<figureCitation id="E2B35CCDDE11CB0CFE0AFA5F7501FA38" box="[497,607,1445,1471]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Figs. 2, 3</figureCitation>
) transverse, bearing wide, rounded carinae and tubercles, with an anterior transverse pubescent area continuing onto anterior margin of propleuron. Meso- and metathoracic wings absent.
<emphasis id="48FC9C5ADE11CB0CFF09FA0F7672F988" bold="true" box="[242,300,1525,1551]" pageId="1" pageNumber="48">Legs</emphasis>
as in
<figureCitation id="E2B35CCDDE11CB0CFE8EFA0F76B8F988" box="[373,486,1525,1551]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="1" pageNumber="48">Figs. 7–9</figureCitation>
, rather short; femora robust, distinctly constricted before apex, with two ventral longitudinal pubescent areas; fore femur with one dorsal longitudinal pubescent area; tibiae strongly curved apically; tarsi two-segmented, segment I short; pretarsi with pair simple subequal claws.
<emphasis id="48FC9C5ADE11CB0CFB59F9BF7242F9D8" bold="true" box="[1186,1308,1605,1631]" pageId="1" pageNumber="48">Abdomen</emphasis>
nearly circular in dorsal aspect (
<figureCitation id="E2B35CCDDE11CB0CFE59F99776B9F900" box="[418,487,1645,1671]" captionStart="FIGURE 1" captionStartId="2.[151,254,1949,1973]" captionTargetBox="[309,1268,221,1895]" captionTargetId="figure@2.[281,1306,197,1923]" captionTargetPageId="2" captionText="FIGURE 1. Enigmocephala deinorhyncha gen. n., sp. n., female holotype." httpUri="https://zenodo.org/record/176540/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
), strongly convex ventrally (
<figureCitation id="E2B35CCDDE11CB0CFCBBF99774C9F900" box="[832,919,1645,1671]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="1" pageNumber="48">Fig. 10</figureCitation>
); dorsal laterotergites extremely wide; tergites I+II and III fused, together bearing pair large, oval pubescent areas; intersegmental suture between tergites IV and V strongly curved anteriad medially; a row of short carinulae present along anterior border of tergites IV–VII; orifices of dorsal abdominal glands present at posterior border of tergites IV and V; sternites largely fused, only shallow sulci present between them except between sternites VI and VII medially (
<figureCitation id="E2B35CCDDE11CB0CFA96F8F777E6F8C8" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="1" pageNumber="48">Fig. 10</figureCitation>
). Spiracles of segment I dorsal, located on distinct elevations of fused tergites I+II and III (
<figureCitation id="E2B35CCDDE11CB0CFB23F8CF7241F8C8" box="[1240,1311,1845,1871]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,269,1166,1190]" captionTargetBox="[174,1413,210,1125]" captionTargetId="figure@3.[155,1432,194,1142]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="1" pageNumber="48">Fig. 2</figureCitation>
); spiracles of segments II–VII ventral (
<figureCitation id="E2B35CCDDE11CB0CFE1DF8A77560F8F0" box="[486,574,1885,1911]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="1" pageNumber="48">Fig. 10</figureCitation>
); an additional pair of spiracles on separated sclerites derived from segment VIII (
<figureCitation id="E2B35CCDDE11CB0CFEE1F87F762CF818" box="[282,370,1925,1951]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="1" pageNumber="48">Fig. 12</figureCitation>
).
<emphasis id="48FC9C5ADE11CB0CFE7DF87F750DF818" bold="true" box="[390,595,1925,1951]" pageId="1" pageNumber="48">Female genitalia</emphasis>
determined mostly by tergite IX and gonocoxites VIII forming together a nearly vertical surface (
<figureCitation id="E2B35CCDDE11CB0CFE27F8577532F840" box="[476,620,1965,1991]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="1" pageNumber="48">Figs 10–12</figureCitation>
); tergite IX elongate, tongue-shaped, slightly narrowing apically; gonapophyses small.
</paragraph>
</subSubSection>
<caption id="2EF710C0DE12CB0FFF6CF86774E5F832" box="[151,955,1949,1974]" httpUri="https://zenodo.org/record/176540/files/figure.png" pageId="2" pageNumber="49" targetBox="[309,1268,221,1895]" targetPageId="2">
<paragraph id="7A374048DE12CB0FFF6CF86774E5F832" blockId="2.[151,955,1949,1974]" box="[151,955,1949,1974]" pageId="2" pageNumber="49">
<emphasis id="48FC9C5ADE12CB0FFF6CF8677646F832" bold="true" box="[151,280,1949,1973]" pageId="2" pageNumber="49">FIGURE 1.</emphasis>
<taxonomicName id="BD883BCBDE12CB0FFEE4F8647501F832" box="[287,607,1950,1973]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="deinorhyncha" status="gen. nov.">
<emphasis id="48FC9C5ADE12CB0FFEE4F8647501F832" box="[287,607,1950,1973]" italics="true" pageId="2" pageNumber="49">Enigmocephala deinorhyncha</emphasis>
</taxonomicName>
<emphasis id="48FC9C5ADE12CB0FFD9DF86775A4F832" bold="true" box="[614,762,1949,1974]" pageId="2" pageNumber="49">
<taxonomicNameLabel id="53CF2121DE12CB0FFD9DF86775F1F831" box="[614,687,1949,1974]" pageId="2" pageNumber="49" rank="genus">gen. n.</taxonomicNameLabel>
, sp. n.
</emphasis>
, female holotype.
</paragraph>
</caption>
<caption id="2EF710C0DE13CB0EFF6CFB747796FB6C" httpUri="https://zenodo.org/record/176541/files/figure.png" pageId="3" pageNumber="50" targetBox="[174,1413,210,1125]" targetPageId="3">
<paragraph id="7A374048DE13CB0EFF6CFB747796FB6C" blockId="3.[151,1436,1166,1259]" pageId="3" pageNumber="50">
<emphasis id="48FC9C5ADE13CB0EFF6CFB74761BFB21" bold="true" box="[151,325,1166,1190]" pageId="3" pageNumber="50">FIGURES 2–6.</emphasis>
<taxonomicName id="BD883BCBDE13CB0EFEB6FB7575D1FB21" box="[333,655,1167,1190]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="deinorhyncha" status="gen. nov.">
<emphasis id="48FC9C5ADE13CB0EFEB6FB7575D1FB21" box="[333,655,1167,1190]" italics="true" pageId="3" pageNumber="50">Enigmocephala deinorhyncha</emphasis>
</taxonomicName>
<emphasis id="48FC9C5ADE13CB0EFD6CFB74734FFB21" bold="true" box="[663,1041,1166,1190]" pageId="3" pageNumber="50">
<taxonomicNameLabel id="53CF2121DE13CB0EFD6CFB7475BAFB21" box="[663,740,1166,1190]" pageId="3" pageNumber="50" rank="genus">gen. n.</taxonomicNameLabel>
, sp. n., female holotype. 2
</emphasis>
, head, thorax and base of abdomen, dorsal view,
<emphasis id="48FC9C5ADE13CB0EFEE4FB4B7672FB4E" bold="true" box="[287,300,1201,1225]" pageId="3" pageNumber="50">3</emphasis>
, same, lateral view,
<emphasis id="48FC9C5ADE13CB0EFDF2FB4B7548FB4E" bold="true" box="[521,534,1201,1225]" pageId="3" pageNumber="50">4</emphasis>
, head, dorsal view,
<emphasis id="48FC9C5ADE13CB0EFD16FB4B75A4FB4E" bold="true" box="[749,762,1201,1225]" pageId="3" pageNumber="50">5</emphasis>
, same, anterior view,
<emphasis id="48FC9C5ADE13CB0EFC1CFB4B74AAFB4E" bold="true" box="[999,1012,1201,1225]" pageId="3" pageNumber="50">6</emphasis>
, apex of labium, lateral view. Scales in mm.
</paragraph>
</caption>
<subSubSection id="329213C3DE13CB0EFF3CFAE5734DFABE" box="[199,1043,1311,1337]" pageId="3" pageNumber="50" type="materials_examined">
<paragraph id="7A374048DE13CB0EFF3CFAE5734DFABE" blockId="3.[151,1437,1311,2017]" box="[199,1043,1311,1337]" pageId="3" pageNumber="50">
<emphasis id="48FC9C5ADE13CB0EFF3CFAE57637FABE" bold="true" box="[199,361,1311,1337]" pageId="3" pageNumber="50">
<typeStatus id="A533FEEADE13CB0EFF3CFAE5765AFABE" box="[199,260,1311,1337]" pageId="3" pageNumber="50">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="BD883BCBDE13CB0EFE8BFAE5758FFABF" box="[368,721,1311,1336]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="deinorhyncha" status="sp. nov.">
<emphasis id="48FC9C5ADE13CB0EFE8BFAE5758FFABF" box="[368,721,1311,1336]" italics="true" pageId="3" pageNumber="50">Enigmocephala deinorhyncha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CF2121DE13CB0EFD2CFAE57444FABE" box="[727,794,1311,1337]" pageId="3" pageNumber="50" rank="species">
<emphasis id="48FC9C5ADE13CB0EFD2CFAE57444FABE" bold="true" box="[727,794,1311,1337]" pageId="3" pageNumber="50">sp. n.</emphasis>
</taxonomicNameLabel>
, present designation.
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE13CB0EFF3CFABD763BFA36" pageId="3" pageNumber="50" type="etymology">
<paragraph id="7A374048DE13CB0EFF3CFABD763BFA36" blockId="3.[151,1437,1311,2017]" pageId="3" pageNumber="50">
<emphasis id="48FC9C5ADE13CB0EFF3CFABD760AFAE6" bold="true" box="[199,340,1351,1377]" pageId="3" pageNumber="50">Etymology.</emphasis>
The generic name from the Greek αɭνɭγμα (latinized as
<emphasis id="48FC9C5ADE13CB0EFC27FABD736CFAE7" box="[988,1074,1351,1376]" italics="true" pageId="3" pageNumber="50">enigma</emphasis>
) ‘riddle’ and κεφαλη (latinized as
<emphasis id="48FC9C5ADE13CB0EFF43FA95764DFA0F" box="[184,275,1391,1416]" italics="true" pageId="3" pageNumber="50">cephale</emphasis>
) ‘head’, the latter referring to
<taxonomicName id="BD883BCBDE13CB0EFD7BFA95746CFA0F" box="[640,818,1391,1416]" class="Insecta" family="Reduviidae" genus="Tribelocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="48FC9C5ADE13CB0EFD7BFA95746CFA0F" box="[640,818,1391,1416]" italics="true" pageId="3" pageNumber="50">Tribelocephala</emphasis>
</taxonomicName>
, the
<typeStatus id="A533FEEADE13CB0EFC95FA9574FFFA0E" box="[878,929,1391,1417]" pageId="3" pageNumber="50">type</typeStatus>
genus of the subfamily Tribelocephalinae. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE13CB09FF3CFA457618F876" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="distribution">
<paragraph id="7A374048DE13CB0EFF3CFA4575A9FA5E" blockId="3.[151,1437,1311,2017]" box="[199,759,1471,1497]" pageId="3" pageNumber="50">
<emphasis id="48FC9C5ADE13CB0EFF3CFA457639FA5E" bold="true" box="[199,359,1471,1497]" pageId="3" pageNumber="50">Distribution.</emphasis>
Known exclusively from Borneo.
</paragraph>
<paragraph id="7A374048DE13CB0EFF3CFA1D74EAF8C6" blockId="3.[151,1437,1311,2017]" pageId="3" pageNumber="50">
<emphasis id="48FC9C5ADE13CB0EFF3CFA1D7505F986" bold="true" box="[199,603,1511,1537]" pageId="3" pageNumber="50">Subfamily and tribal placement.</emphasis>
Although this new genus bears superficial resemblance to
<taxonomicName id="BD883BCBDE13CB0EFAEBFA1D779EF9AE" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Ectrichodiinae">Ectrichodiinae</taxonomicName>
(such as the heavily sclerotized body, the secondarily subdivided antennal flagellum, and the presence of short longitudinal carinulae at the anterior margin of the abdominal sternites),
<taxonomicName id="BD883BCBDE13CB0EFBD0F9CD73BDF9D7" box="[1067,1251,1591,1616]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="48FC9C5ADE13CB0EFBD0F9CD73BDF9D7" box="[1067,1251,1591,1616]" italics="true" pageId="3" pageNumber="50">Enigmocephala</emphasis>
</taxonomicName>
clearly belongs to Tribelocephalinae because of the following characters: third visible labial segment (IV) strongly shortened (synapomorphy of Tribelocephalinae); presence of basal collar-like pubescent area and a pair of longitudinal fringe-like structures on ventral surface of head (apomorphy occurring in many genera of Tribelocephalinae); absence of fossula spongiosa (invariably present on fore and mid legs in
<taxonomicName id="BD883BCBDE13CB0EFBF9F92D73EDF976" box="[1026,1203,1751,1777]" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Ectrichodiinae">Ectrichodiinae</taxonomicName>
); labrum not subdivided (subdivided in
<taxonomicName id="BD883BCBDE13CB0EFE6FF9057561F89E" box="[404,575,1791,1817]" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Ectrichodiinae">Ectrichodiinae</taxonomicName>
except in
<emphasis id="48FC9C5ADE13CB0EFD47F905743AF89F" box="[700,868,1791,1816]" italics="true" pageId="3" pageNumber="50">Ectrichodiella</emphasis>
Fracker &amp; Bruner, 1924 and
<emphasis id="48FC9C5ADE13CB0EFB3DF9057264F89F" box="[1222,1338,1791,1816]" italics="true" pageId="3" pageNumber="50">Schuhella</emphasis>
Dougherty, 1995); and ocelli absent (synapomorphy of Tribelocephalinae).
</paragraph>
<paragraph id="7A374048DE13CB09FF3CF8B57582FEAD" blockId="3.[151,1437,1311,2017]" lastBlockId="4.[151,1436,152,298]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
The fore wing venation is currently the single recognized character for differentiating Tribelocephalini from Opistoplatyini (
<bibRefCitation id="1E193DB9DE13CB0EFE6CF88D7566F816" author="Villiers" box="[407,568,1911,1937]" pageId="3" pageNumber="53" refString="Villiers, A. (1943) Morphologie et systematique des Tribelocephalitae africains (Hem. Reduviidae). Revue Francaise d'Entomologie, 10, 1 - 28." type="journal article" year="1943">Villiers 1943</bibRefCitation>
,
<bibRefCitation id="1E193DB9DE13CB0EFDB2F88D74D9F816" author="Maldonado" box="[585,903,1911,1937]" pageId="3" pageNumber="53" refString="Maldonado Capriles, J. (1996) New taxa and key to the tribes and genera in Tribelocephalinae Stal 1866 (Heteroptera: Reduviidae). Proceedings of the Entomological Society of Washington, 98, 138 - 144." type="journal article" year="1996">Maldonado Capriles 1996</bibRefCitation>
). This character cannot be examined in the apterous
<typeStatus id="A533FEEADE13CB0EFEFAF8657639F83E" box="[257,359,1951,1977]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
designated below for the new species of this genus. However, the new genus is very similar to the Bornean genera
<emphasis id="48FC9C5ADE13CB0EFE59F83D756AF867" box="[418,564,1991,2016]" italics="true" pageId="3" pageNumber="50">Gastrogyrus</emphasis>
Bergroth, 1921 and
<emphasis id="48FC9C5ADE13CB0EFCDCF83D7492F867" box="[807,972,1991,2016]" italics="true" pageId="3" pageNumber="50">Matangocoris</emphasis>
Miller, 1940, both clearly belonging to Tribelocephalini, in the anteriorly declivent head, the presence of a basal pubescent area and ventral fringelike pilosity of the head, and the robust labium. Some of these may be synapomorphies for the presumably monophyletic group composed of these genera (probably also
<emphasis id="48FC9C5ADE14CB09FC76FF127355FE86" box="[909,1035,232,257]" italics="true" pageId="4" pageNumber="51">Afrodecius</emphasis>
Jeannel, 1911). Therefore,
<taxonomicName id="BD883BCBDE14CB09FAA0FF127646FEAE" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="48FC9C5ADE14CB09FAA0FF127646FEAE" italics="true" pageId="4" pageNumber="51">Enigmocephala</emphasis>
</taxonomicName>
is placed in the tribe Tribelocephalini.
</paragraph>
<caption id="2EF710C0DE14CB09FF6CFB3B7231FB7C" httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="4" pageNumber="51" targetBox="[160,1428,369,1175]" targetPageId="4">
<paragraph id="7A374048DE14CB09FF6CFB3B7231FB7C" blockId="4.[151,1436,1217,1275]" pageId="4" pageNumber="51">
<emphasis id="48FC9C5ADE14CB09FF6CFB3B760CFB5E" bold="true" box="[151,338,1217,1241]" pageId="4" pageNumber="51">FIGURES 7–12.</emphasis>
<taxonomicName id="BD883BCBDE14CB09FEA0FB3875C2FB5E" box="[347,668,1218,1241]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="deinorhyncha" status="gen. nov.">
<emphasis id="48FC9C5ADE14CB09FEA0FB3875C2FB5E" box="[347,668,1218,1241]" italics="true" pageId="4" pageNumber="51">Enigmocephala deinorhyncha</emphasis>
</taxonomicName>
<emphasis id="48FC9C5ADE14CB09FD5EFB3B737FFB5E" bold="true" box="[677,1057,1217,1241]" pageId="4" pageNumber="51">
<taxonomicNameLabel id="53CF2121DE14CB09FD5EFB3B75ACFB5E" box="[677,754,1217,1241]" pageId="4" pageNumber="51" rank="genus">gen. n.</taxonomicNameLabel>
, sp. n., female holotype. 7
</emphasis>
, right fore leg,
<emphasis id="48FC9C5ADE14CB09FB35FB3B7385FB5E" bold="true" box="[1230,1243,1217,1241]" pageId="4" pageNumber="51">8</emphasis>
, right mid leg,
<emphasis id="48FC9C5ADE14CB09FA73FB3B72CBFB5E" bold="true" box="[1416,1429,1217,1241]" pageId="4" pageNumber="51">9</emphasis>
, right hind leg,
<emphasis id="48FC9C5ADE14CB09FECFFB197610FB7C" bold="true" box="[308,334,1251,1275]" pageId="4" pageNumber="51">10</emphasis>
, abdomen, lateral view,
<emphasis id="48FC9C5ADE14CB09FDAFFB197532FB7C" bold="true" box="[596,620,1251,1275]" pageId="4" pageNumber="51">11</emphasis>
, apex of abdomen, lateral view,
<emphasis id="48FC9C5ADE14CB09FC3CFB1974BCFB7C" bold="true" box="[967,994,1251,1275]" pageId="4" pageNumber="51">12</emphasis>
, same, posterior view. Scales in mm.
</paragraph>
</caption>
<paragraph id="7A374048DE14CB09FF3CFAD47381F9E6" blockId="4.[151,1437,1326,2033]" pageId="4" pageNumber="51">
<emphasis id="48FC9C5ADE14CB09FF3CFAD475D6FACF" bold="true" box="[199,648,1326,1352]" pageId="4" pageNumber="51">Differentiating from similar genera.</emphasis>
<taxonomicName id="BD883BCBDE14CB09FD6BFAD57416FACF" box="[656,840,1327,1352]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="48FC9C5ADE14CB09FD6BFAD57416FACF" box="[656,840,1327,1352]" italics="true" pageId="4" pageNumber="51">Enigmocephala</emphasis>
</taxonomicName>
is unique within Tribelocephalinae with respect to the characters mentioned under the Diagnosis except the tendency towards the fusion of the abdominal sternites which occurs at least in
<emphasis id="48FC9C5ADE14CB09FE27FA857504FA1F" box="[476,602,1407,1432]" italics="true" pageId="4" pageNumber="51">Afrodecius</emphasis>
Jeannel, 1919 and
<emphasis id="48FC9C5ADE14CB09FCC6FA85748FFA1F" box="[829,977,1407,1432]" italics="true" pageId="4" pageNumber="51">Homognetus</emphasis>
Bergroth, 1923.
<collectingCountry id="029F00D8DE14CB09FB62FA8573E4FA1E" box="[1177,1210,1407,1433]" name="American Samoa" pageId="4" pageNumber="51">As</collectingCountry>
pointed out above, the new genus is similar in several respects to
<emphasis id="48FC9C5ADE14CB09FD48FA5D741AFA47" box="[691,836,1447,1472]" italics="true" pageId="4" pageNumber="51">Gastrogyrus</emphasis>
and
<emphasis id="48FC9C5ADE14CB09FC86FA5D737FFA47" box="[893,1057,1447,1472]" italics="true" pageId="4" pageNumber="51">Matangocoris</emphasis>
. However, in the latter two genera the second visible labial segment is strongly narrowed in its apical 3/5–1/2 part, and the third visible segment is simple. In
<emphasis id="48FC9C5ADE14CB09FE78FA0D754AF997" box="[387,532,1527,1552]" italics="true" pageId="4" pageNumber="51">Gastrogyrus</emphasis>
, the antennal flagellum is not secondarily fragmented (
<bibRefCitation id="1E193DB9DE14CB09FB39FA0D7788F9BE" author="China" pageId="4" pageNumber="53" refString="China, W. E. &amp; Usinger, R. L. (1949) A new genus of Tribelocephalinae from Fernando Poo (Hemiptera Reduviidae). Annali del Museo Civico di Storia Naturale de Genova, 64, 43 - 47." type="journal article" year="1949">
<collectingCountry id="029F00D8DE14CB09FB39FA0D7254F996" box="[1218,1290,1527,1553]" name="China" pageId="4" pageNumber="51">China</collectingCountry>
&amp; Usinger, 1949
</bibRefCitation>
), and furthermore the basiflagellomere is much wider than the distiflagellomere. In
<emphasis id="48FC9C5ADE14CB09FB55F9E5720CF9BF" box="[1198,1362,1567,1592]" italics="true" pageId="4" pageNumber="51">Matangocoris</emphasis>
, setae of both the basal pubescent area and the ventral fringe-like pilosity of the head are very short.
</paragraph>
<paragraph id="7A374048DE14CB09FF3CF9957687F8AE" blockId="4.[151,1437,1326,2033]" pageId="4" pageNumber="51">
The projection-bearing third visible (IV) segment of the labium of
<taxonomicName id="BD883BCBDE14CB09FC29F99573D4F90F" box="[978,1162,1647,1672]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="48FC9C5ADE14CB09FC29F99573D4F90F" box="[978,1162,1647,1672]" italics="true" pageId="4" pageNumber="51">Enigmocephala</emphasis>
</taxonomicName>
is unique within
<taxonomicName id="BD883BCBDE14CB09FAAFF99577BCF936" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="family">Reduviidae</taxonomicName>
, even within Heteroptera. The only other genus with a more or less similar, strongly modified apical labial segment is
<emphasis id="48FC9C5ADE14CB09FE99F94576BEF95F" box="[354,480,1727,1752]" italics="true" pageId="4" pageNumber="51">Afrodecius</emphasis>
. However, in the latter, two beak-like, apparently movable lobes are present, both most probably having originated from segment IV. The exact origin and functional anatomy of these structures requires further study.
</paragraph>
<paragraph id="7A374048DE14CB09FF3CF8CD7618F876" blockId="4.[151,1437,1326,2033]" pageId="4" pageNumber="51">
The genera of the tribe Tribelocephalini can be distinguished with the key below.
<emphasis id="48FC9C5ADE14CB09FB3AF8CD72C3F8D7" box="[1217,1437,1847,1872]" italics="true" pageId="4" pageNumber="51">Acanthorhinocoris</emphasis>
Miller, 1940 was placed to Tribelocephalini by
<bibRefCitation id="1E193DB9DE14CB09FD25F8A5737CF8FE" author="Maldonado" box="[734,1058,1887,1913]" pageId="4" pageNumber="53" refString="Maldonado Capriles, J. (1990) Systematic Catalogue of the Reduviidae of the World (Insecta: Heteroptera). Caribbean Journal of Sciences, Special Edition, X + 694 pp." type="book" year="1990">Maldonado Capriles (1990</bibRefCitation>
,
<bibRefCitation id="1E193DB9DE14CB09FBC9F8A5732FF8FE" author="Capriles" box="[1074,1137,1887,1913]" pageId="4" pageNumber="53" refString="Maldonado Capriles, J. (1996) New taxa and key to the tribes and genera in Tribelocephalinae Stal 1866 (Heteroptera: Reduviidae). Proceedings of the Entomological Society of Washington, 98, 138 - 144." type="journal article" year="1996">1996</bibRefCitation>
). However, according to information received from M.D. Webb (The Natural History Museum, London), a distinct Cu-PCu cross vein is present in this genus; therefore,
<emphasis id="48FC9C5ADE14CB09FDD5F8557457F84F" box="[558,777,1967,1992]" italics="true" pageId="4" pageNumber="51">Acanthorhinocoris</emphasis>
is hereby removed from Tribelocephalini and placed in Opistoplatyini.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="33E8BB42CAC15B09379D7F20CB32ABFA">A new genus of tribelocephaline assassin bugs from Borneo (Hemiptera: Heteroptera: Reduviidae)</mods:title>
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<mods:identifier id="655FD7D6571617A2685B5959F69EFA51" type="DOI">10.5281/zenodo.176539</mods:identifier>
<mods:identifier id="3BF2C77AD6BB8796E29DE74169D56710" type="GBIF-Dataset">86f7c9fb-723c-4561-818a-5fffd2d9307c</mods:identifier>
<mods:identifier id="06B6D2461011ECA9DDF17C895C9373F1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="90A32986503C661AE752E9FA9EB3F36B" type="Zenodo-Dep">176539</mods:identifier>
</mods:mods>
<treatment id="F221F15EDE15CB0BFF6CF9A47245FCD8" ID-DOI="http://doi.org/10.5281/zenodo.6247067" ID-GBIF-Taxon="119358756" ID-Zenodo-Dep="6247067" LSID="urn:lsid:plazi:treatment:F221F15EDE15CB0BFF6CF9A47245FCD8" httpUri="http://treatment.plazi.org/id/F221F15EDE15CB0BFF6CF9A47245FCD8" lastPageId="6" lastPageNumber="53" pageId="5" pageNumber="52">
<subSubSection id="329213C3DE15CB08FF6CF9A47640F91C" pageId="5" pageNumber="52" type="nomenclature">
<paragraph id="7A374048DE15CB08FF6CF9A47510F9FF" blockId="5.[151,590,1630,1691]" box="[151,590,1630,1656]" pageId="5" pageNumber="52">
<heading id="217FF724DE15CB08FF6CF9A47510F9FF" bold="true" box="[151,590,1630,1656]" fontSize="11" level="1" pageId="5" pageNumber="52" reason="1">
<emphasis id="48FC9C5ADE15CB08FF6CF9A47510F9FF" bold="true" box="[151,590,1630,1656]" pageId="5" pageNumber="52">
<taxonomicName id="BD883BCBDE15CB08FF6CF9A4755AF9FF" ID-CoL="39V5X" box="[151,516,1630,1656]" class="Insecta" family="Reduviidae" genus="Enigmocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="deinorhyncha" status="sp. nov.">
<emphasis id="48FC9C5ADE15CB08FF6CF9A4755AF9FF" bold="true" box="[151,516,1630,1656]" italics="true" pageId="5" pageNumber="52">Enigmocephala deinorhyncha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53CF2121DE15CB08FDF7F9A47510F9FF" box="[524,590,1630,1656]" pageId="5" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7A374048DE15CB08FF6CF97B7640F91C" blockId="5.[151,590,1630,1691]" box="[151,286,1665,1691]" pageId="5" pageNumber="52">
(
<figureCitation id="E2B35CCDDE15CB08FF64F97B764BF91C" box="[159,277,1665,1691]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 6" captionStart-2="FIGURES 7 – 12" captionStartId-0="2.[151,254,1949,1973]" captionStartId-1="3.[151,269,1166,1190]" captionStartId-2="4.[151,269,1217,1241]" captionTargetBox-0="[309,1268,221,1895]" captionTargetBox-1="[174,1413,210,1125]" captionTargetBox-2="[160,1428,369,1175]" captionTargetId-0="figure@2.[281,1306,197,1923]" captionTargetId-1="figure@3.[155,1432,194,1142]" captionTargetId-2="figure@4.[155,1432,354,1193]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Enigmocephala deinorhyncha gen. n., sp. n., female holotype." captionText-1="FIGURES 2 – 6. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 2, head, thorax and base of abdomen, dorsal view, 3, same, lateral view, 4, head, dorsal view, 5, same, anterior view, 6, apex of labium, lateral view. Scales in mm." captionText-2="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri-0="https://zenodo.org/record/176540/files/figure.png" httpUri-1="https://zenodo.org/record/176541/files/figure.png" httpUri-2="https://zenodo.org/record/176542/files/figure.png" pageId="5" pageNumber="52">Figs 1–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE15CB08FF6CF92A7537F894" pageId="5" pageNumber="52" type="materials_examined">
<paragraph id="7A374048DE15CB08FF6CF92A7537F894" blockId="5.[151,1437,1744,2011]" pageId="5" pageNumber="52">
<emphasis id="48FC9C5ADE15CB08FF6CF92A7614F96D" bold="true" box="[151,330,1744,1770]" pageId="5" pageNumber="52">
<typeStatus id="A533FEEADE15CB08FF6CF92A778CF96D" box="[151,210,1744,1770]" pageId="5" pageNumber="52">Type</typeStatus>
material.
</emphasis>
<typeStatus id="A533FEEADE15CB08FEAFF92B769FF96C" box="[340,449,1745,1771]" pageId="5" pageNumber="52" type="holotype">Holotype</typeStatus>
(Ψ): “
<collectingCountry id="029F00D8DE15CB08FDF3F92B75F6F96C" box="[520,680,1745,1771]" name="Malaysia" pageId="5" pageNumber="52">MALAYSIA</collectingCountry>
:
<collectingCountry id="029F00D8DE15CB08FD49F92B7449F96C" box="[690,791,1745,1771]" name="Malaysia" pageId="5" pageNumber="52">Sarawak</collectingCountry>
\
<quantity id="BD70EDADDE15CB08FCD6F92B742EF96C" box="[813,880,1745,1771]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="52" unit="km" value="25.0">25km</quantity>
E Kapit \
<date id="0E366688DE15CB08FC13F92B7311F96C" box="[1000,1103,1745,1771]" pageId="5" pageNumber="52" value="1994-03">III. 1994</date>
\ leg. Kodada”; deposited in the Natural History Museum of Vienna.
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE15CB0BFF3DF8DA727DFC88" lastPageId="6" lastPageNumber="53" pageId="5" pageNumber="52" type="description">
<paragraph id="7A374048DE15CB08FF3DF8DA756EF8BC" blockId="5.[151,1437,1744,2011]" box="[198,560,1824,1851]" pageId="5" pageNumber="52">
<emphasis id="48FC9C5ADE15CB08FF3DF8DA7601F8BD" bold="true" box="[198,351,1824,1850]" pageId="5" pageNumber="52">Description.</emphasis>
Apterous female.
</paragraph>
<paragraph id="7A374048DE15CB08FF3CF8B277B4F85C" blockId="5.[151,1437,1744,2011]" pageId="5" pageNumber="52">
<emphasis id="48FC9C5ADE15CB08FF3CF8B2767DF8E5" bold="true" box="[199,291,1864,1890]" pageId="5" pageNumber="52">Colour.</emphasis>
Body rather uniformly brown; scape, pedicel, tibiae, and tarsi distinctly lighter; antennal flagellum pale yellow; thoracic pleuron and sternum dark blackish brown; supracoxal lobes brown. Dense pubescence of the body grayish brown; large paired oval pubescence near base of abdominal dorsum light yellowish brown.
</paragraph>
<paragraph id="7A374048DE16CB0BFF3CFF6D741BFE25" blockId="6.[151,1437,151,863]" pageId="6" pageNumber="53">
<emphasis id="48FC9C5ADE16CB0BFF3CFF6D76CEFF36" bold="true" box="[199,400,151,177]" pageId="6" pageNumber="53">Structure. Head</emphasis>
about 1.9 times as long as its width across eyes, diatone about 1.37 times greater than interocular distance.
<emphasis id="48FC9C5ADE16CB0BFE68FF457558FF5E" bold="true" box="[403,518,191,217]" pageId="6" pageNumber="53">Antenna.</emphasis>
Pedicel about 1.25 times longer than scape and about 1.2 times as long as width of head.
<emphasis id="48FC9C5ADE16CB0BFEBCFF1D76F4FE86" bold="true" box="[327,426,231,257]" pageId="6" pageNumber="53">Thorax.</emphasis>
Disc of pronotum with two pairs of tubercles surrounded by pair carinae laterally and posteriorly, posterior pair of these tubercles distinctly larger than anterior one; with pair large posterior protuberances at posterior border of pronotum; and with deep medial furrow between these protuberances. Meso- and metanota each with pair submedial tubercles and deep medial sulcus.
<emphasis id="48FC9C5ADE16CB0BFBF6FEA573D9FEFE" bold="true" box="[1037,1159,351,377]" pageId="6" pageNumber="53">Abdomen</emphasis>
about 1.2 times longer than its greatest width.
<emphasis id="48FC9C5ADE16CB0BFE5CFE7D752BFE26" bold="true" box="[423,629,391,417]" pageId="6" pageNumber="53">Female genitalia</emphasis>
as in
<figureCitation id="E2B35CCDDE16CB0BFD41FE72741FFE25" box="[698,833,392,418]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,269,1217,1241]" captionTargetBox="[160,1428,369,1175]" captionTargetId="figure@4.[155,1432,354,1193]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Enigmocephala deinorhyncha gen. n., sp. n., female holotype. 7, right fore leg, 8, right mid leg, 9, right hind leg, 10, abdomen, lateral view, 11, apex of abdomen, lateral view, 12, same, posterior view. Scales in mm." httpUri="https://zenodo.org/record/176542/files/figure.png" pageId="6" pageNumber="53">Figs 10–12</figureCitation>
.
</paragraph>
<paragraph id="7A374048DE16CB0BFF3CFE55727DFC88" blockId="6.[151,1437,151,863]" pageId="6" pageNumber="53">
<emphasis id="48FC9C5ADE16CB0BFF3CFE557620FE4E" bold="true" box="[199,382,431,457]" pageId="6" pageNumber="53">Measurements</emphasis>
(in mm). Body length 7.2. Length of head from apex of basal pubescent area to apex of clypeus 1.76, greatest width 0.93, interocular distance 0.68. Length of antennal segments I: II:
<date id="0E366688DE16CB0BFB07FE22727CFE71" box="[1276,1314,472,502]" pageId="6" pageNumber="53">III1</date>
:
<date id="0E366688DE16CB0BFAC2FE227200FE71" box="[1337,1374,472,502]" pageId="6" pageNumber="53">III2</date>
:
<date id="0E366688DE16CB0BFA8EFE2272C2FE71" box="[1397,1436,472,502]" pageId="6" pageNumber="53">IV1</date>
:
<date id="0E366688DE16CB0BFF5CFDF87790FDA6" box="[167,206,514,545]" pageId="6" pageNumber="53">IV2</date>
:
<date id="0E366688DE16CB0BFF1DFDF87652FDA6" box="[230,268,514,545]" pageId="6" pageNumber="53">IV3</date>
:
<date id="0E366688DE16CB0BFEDFFDF87615FDA6" box="[292,331,514,545]" pageId="6" pageNumber="53">IV4</date>
= 0.88: 1.10: 0.102: 0.186: 0.322: 0.153: 0.169: 0.339, greatest width of segments I: II: flagellum = 0.255: 0.085: 0.042. Length of visible labial segments I: II: III = 0.86: 1.61: 0.17. Length of pronotum from anterior margin of anterior pubescent area to level of posterior protuberances 1.27, greatest width of pronotum 1.61, greatest width of thorax (metapleuron) 1.88. Medial length of abdomen in dorsal aspect 3.8, greatest width 3.2. Lengths of femur, tibia, and tarsus (segments I and II) of fore leg 2.02, 1.95, and 0.44 (0.32 and 0.15); of mid leg 1.95, 1.81, and 0.36 (0.31 and 0.12); of hind leg 2.51, 2.48, and 0.46 (0.17 and 0.34), respectively; greatest widths of fore, mid, and hind femora 0.52, 0.46, and 0.50, respectively.
</paragraph>
</subSubSection>
<subSubSection id="329213C3DE16CB0BFF3CFCE77245FCD8" pageId="6" pageNumber="53" type="etymology">
<paragraph id="7A374048DE16CB0BFF3CFCE77245FCD8" blockId="6.[151,1437,151,863]" pageId="6" pageNumber="53">
<emphasis id="48FC9C5ADE16CB0BFF3CFCE7760AFCB0" bold="true" box="[199,340,797,823]" pageId="6" pageNumber="53">Etymology.</emphasis>
The specific name from the Greek δεɭνOς, δεɭνO- (latinized as
<emphasis id="48FC9C5ADE16CB0BFBABFCE473CFFCB0" box="[1104,1169,798,823]" italics="true" pageId="6" pageNumber="53">deino</emphasis>
-) ‘enormous, terrible’ and Greek ρυγχOς (latinized as
<emphasis id="48FC9C5ADE16CB0BFDF9FCBC7532FCD8" box="[514,620,838,863]" italics="true" pageId="6" pageNumber="53">rhynchos</emphasis>
) ‘beak’, referring to the very robust labium of the species.
</paragraph>
</subSubSection>
</treatment>
</document>