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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.837.32057" ID-GBIF-Dataset="3b0408de-0dbe-4b70-81ae-82213844bbad" ID-PMC="PMC6477869" ID-Pensoft-Pub="1313-2970-837-1" ID-PubMed="31048963" ID-ZBK="4A18822935804DB7B1229F131F6A0AC8" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-837-1" ModsDocOrigin="ZooKeys 837" ModsDocTitle="A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini)" checkinTime="1555318524876" checkinUser="pensoft" docAuthor="Fabrizi, Silvia, Dalstein, Vivian &amp; Ahrens, Dirk" docDate="2019" docId="A8D4507147A33DE34531BB885E5A1DB7" docLanguage="en" docName="ZooKeys 837: 1-155" docOrigin="ZooKeys 837" docSource="http://dx.doi.org/10.3897/zookeys.837.32057" docTitle="Tetraserica cattienensis Fabrizi, Dalstein &amp; Ahrens, 2019, sp. n." docType="treatment" docUuid="244038AC-E7EE-4531-AA94-7439E68B0D81" docUuidSource="ZooBank" docVersion="5" lastPageNumber="56" masterDocId="677ECC24FFFF2100C748FFF7FFF0E909" masterDocTitle="A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini)" masterLastPageNumber="155" masterPageNumber="1" pageNumber="54" updateTime="1668167136029" updateUser="ExternalLinkService">
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<mods:title>A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fabrizi, Silvia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dalstein, Vivian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ahrens, Dirk</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>837</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>155</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.837.32057</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.837.32057</mods:identifier>
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<mods:identifier type="ZBK">4A18822935804DB7B1229F131F6A0AC8</mods:identifier>
<mods:identifier type="ZooBank">4A18822935804DB7B1229F131F6A0AC8</mods:identifier>
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<treatment ID-GBIF-Taxon="156200652" LSID="urn:lsid:zoobank.org:act:244038AC-E7EE-4531-AA94-7439E68B0D81" httpUri="http://treatment.plazi.org/id/A8D4507147A33DE34531BB885E5A1DB7" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
<subSubSection pageId="53" pageNumber="54" type="nomenclature">
<paragraph pageId="53" pageNumber="54">
<taxonomicName LSID="http://zoobank.org/244038AC-E7EE-4531-AA94-7439E68B0D81" class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica cattienensis" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cattienensis">Tetraserica cattienensis</taxonomicName>
<taxonomicNameLabel pageId="53" pageNumber="54">sp. n.</taxonomicNameLabel>
Figures 22, 46
</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="type material">
<paragraph pageId="54" pageNumber="55">
<pageBreakToken pageId="54" pageNumber="55" start="start">Type</pageBreakToken>
material examined.
</paragraph>
<paragraph pageId="54" pageNumber="55">
Holotype: ♂ &quot;Museum Leiden Viet Nam (Dong Nai Prov.)
<normalizedToken originalValue="Cát">Cat</normalizedToken>
<normalizedToken originalValue="Tiên">Tien</normalizedToken>
NP: Ficus trail. 9-26.iv.2007. leg. May Phu Quy, Nguyen Than Mahn/ humid lowland forest; in malaise traps; 250 m 11°26'20, 6&quot;N, 107°25'42&quot;E&quot; (RMNH). Paratypes: 1 ♂ &quot;Museum Leiden Viet Nam (Dong Nai Prov.)
<normalizedToken originalValue="Cát">Cat</normalizedToken>
<normalizedToken originalValue="Tiên">Tien</normalizedToken>
N.P.: Ficus trail. 9-26.iv.2007. leg. May Phu Quy, Nguyen Than Mahn/ humid lowland forest; in malaise traps; 250 m 11°26'20, 6&quot;N, 107°25'42&quot;E&quot; (ZFMK), 1 ♂ &quot;Museum Leiden Viet Nam (Dong Nai Prov.)
<normalizedToken originalValue="Cát">Cat</normalizedToken>
<normalizedToken originalValue="Tiên">Tien</normalizedToken>
N.P.: Ficus trail. 10.iv-15.v.2007. Leg. May Phu Quy, Nguyen Than Mahn/ humid lowland forest; in malaise traps; 250 m. 11°26'20, 6&quot;N, 107°25'42&quot;E&quot; (RMNH), 1 ♂ &quot;S Vietnam, 1-5.5.1994 Nam Cat Tien Nat. park P.
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
&amp; L.
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
leg./ coll. P.
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
/VS 118&quot; (CPPB).
</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="description">
<paragraph pageId="54" pageNumber="55">Description.</paragraph>
<paragraph pageId="54" pageNumber="55">Length of body: 8.4 mm; length of elytra: 6 mm; maximum width: 5 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large, ratio of diameter/interocular width: 0.67. Ratio of length of metepisternum/metacoxa: 1/1.61. Metatibia short and wide, ratio width/length: 1/2.35; basal group of dorsal spines of metatibia at first third of metatibial length.</paragraph>
<paragraph pageId="54" pageNumber="55">
Aedeagus: Fig. 22
<normalizedToken originalValue="A–C">A-C</normalizedToken>
. Habitus: Fig. 22D.
</paragraph>
<caption pageId="54" pageNumber="55">
<paragraph pageId="54" pageNumber="55">
Figure 22.
<normalizedToken originalValue="A–D">A-D</normalizedToken>
<taxonomicName class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica cattienensis" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cattienensis">Tetraserica cattienensis</taxonomicName>
sp. n. (holotype)
<normalizedToken originalValue="E–H">E-H</normalizedToken>
<taxonomicName lsidName="T. multiangulata" pageId="54" pageNumber="55" rank="species" species="multiangulata">T. multiangulata</taxonomicName>
sp. n. (holotype) A, E aedeagus, left side lateral view C, G aedeagus, right side lateral view B, F parameres, dorsal view D, H habitus. Scale bars: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph pageId="54" pageNumber="55">Female unknown.</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="variation">
<paragraph pageId="54" pageNumber="55">Variation.</paragraph>
<paragraph pageId="54" pageNumber="55">Length of body: 7.9-8.4 mm; length of elytra: 5.9-6 mm; maximum width: 4.9-5.0 mm.</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="diagnosis">
<paragraph pageId="54" pageNumber="55">Diagnosis.</paragraph>
<paragraph pageId="54" pageNumber="55">
<taxonomicName class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica cattienensis" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cattienensis">Tetraserica cattienensis</taxonomicName>
sp. n. differs from all other
<taxonomicName class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Tetraserica</taxonomicName>
species in having a straight or slightly convex posterior margin of the metafemur, small eyes, and the short median phallobasal lamina being half as long as phallobase. From the similar
<taxonomicName lsidName="T. bansanpakiana" pageId="54" pageNumber="55" rank="species" species="bansanpakiana">T. bansanpakiana</taxonomicName>
sp. n. and
<taxonomicName lsidName="T. margheritae" pageId="54" pageNumber="55" rank="species" species="margheritae">T. margheritae</taxonomicName>
sp. n.,
<taxonomicName lsidName="T. cattienensis" pageId="54" pageNumber="55" rank="species" species="cattienensis">T. cattienensis</taxonomicName>
sp. n. differs by the robust spines on the ventral margin of metatibia being subequal in length.
</paragraph>
</subSubSection>
<subSubSection pageId="55" pageNumber="56" type="etymology">
<paragraph pageId="55" pageNumber="56">
<pageBreakToken pageId="55" pageNumber="56" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="55" pageNumber="56">
The new species is named after the type locality,
<normalizedToken originalValue="Cát">Cat</normalizedToken>
<normalizedToken originalValue="Tiên">Tien</normalizedToken>
NP (adjective in the nominative singular).
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="823EEEEE5EE7A71F233F1C51102AE39E" pageId="0" pageNumber="11794">
<taxonomicName id="411104FF28FA89727F41F80C075ADF25" ID-CoL="89GW4" authority="The ́ el, 1879" authorityName="The ́ el" authorityYear="1879" class="Holothuroidea" family="Deimatidae" genus="Deima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deima validum" order="Synallactida" pageId="0" pageNumber="11794" phylum="Echinodermata" rank="species" species="validum" species_undef="cf.">Deima cf. validum Théel, 1879</taxonomicName>
</paragraph>
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<taxonomicName id="757CC093169E34F65F2BF8D05949B3CE" class="Holothuroidea" family="Deimatidae" genus="Deima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deima validum" order="Synallactida" pageId="0" pageNumber="11794" phylum="Echinodermata" rank="species" species="validum" species_undef="cf.">Deima cf. validum</taxonomicName>
In the &quot;Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone&quot; created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as &quot;
<taxonomicName id="4F463A6C790469A3AD7F1C396ED5FE0E" class="Holothuroidea" family="Deimatidae" genus="Deima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deima" order="Synallactida" pageId="0" pageNumber="11794" phylum="Echinodermata" rank="genus">Deima</taxonomicName>
morphotype&quot;.
</paragraph>
</subSubSection>
<subSubSection id="89C8AC6F4B8CED58E13AC99368E16B58" pageId="0" pageNumber="11794" type="materials_examined">
<paragraph id="AE029625B93204AFD4ADBC8E03D60979" pageId="0" pageNumber="11794">Materials</paragraph>
<paragraph id="7BB72B391E23FA32C01867C952BCB903" pageId="0" pageNumber="11794">
<materialsCitation id="15223C5E40107E6024803D053E028F48" collectingDate="2015-04-03" collectingMethod="Autonomous Underwater Vehicle" collectionCode="UHM" collectorName="Diva J Amon, Amanda F Ziegler" latitude="12.4977" location="UK Seabed Resources Ltd exploration contract area (UK- 1)" longitude="-116.6525" pageId="0" pageNumber="11794" specimenCount="1" specimenCount-adult="1" typeStatus="Other material">
Type status:
<typeStatus id="0B6A85BAA6DA7998A259FBA8F403E93D" pageId="0" pageNumber="11794">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName id="14D84E6C7B40FF476F37F0A1BD4A7219" pageId="0" pageNumber="11794">Diva J Amon, Amanda F Ziegler</collectorName>
; individualCount:
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; lifeStage:
<specimenType id="4EDB7DAF83BA4916ED86AED117B096E2" pageId="0" pageNumber="11794">Adult</specimenType>
; behavior: On seafloor; occurrenceStatus: present; preparations: Imaged only; Taxon: taxonConceptID: Deimacf.validum; scientificName: Deimavalidum; kingdom: Animalia; phylum: Echinodermata; class: Holothuroidea; order: Elasipodida; family: Deimatidae; genus: Deima; taxonRank: species; scientificNameAuthorship: Théel, 1879; Location: waterBody: Pacific Ocean; stateProvince: Clarion-Clipperton Zone; locality:
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; verbatimLocality: UK-1 Stratum B; maximumDepthInMeters: 4254; locationRemarks: RV Thompson Cruise TN319; decimalLatitude:
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; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 25; Identification: identifiedBy:
<determinerName id="0700ACF956776300FC20E936F470B896" pageId="0" pageNumber="11794">Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler</determinerName>
; dateIdentified: 2015; identificationRemarks: Identified only from imagery; identificationQualifier: cf.; Event: samplingProtocol:
<collectingMethod id="F7152C181072860037714819038C9F3C" pageId="0" pageNumber="11794">Autonomous Underwater Vehicle</collectingMethod>
; eventDate:
<collectingDate id="C35D5BD2C691D410576E540903EF0959" pageId="0" pageNumber="11794" value="2015-04-03">2015-03-04</collectingDate>
; eventTime: 4:01; habitat: Abyssal polymetallic-nodule field; fieldNumber: Dive 5 (AV05); Record Level: language: en; institutionCode:
<collectionCode id="DD97E67AED32F3156D8D2CF569C42EF6" pageId="0" pageNumber="11794">UHM</collectionCode>
; datasetName: ABYSSLINE; basisOfRecord: HumanObservation
</materialsCitation>
<materialsCitation id="11239AC6B087F579877BA06F223C95C7" collectingDate="2015-09-03" collectingMethod="Autonomous Underwater Vehicle" collectionCode="UHM" collectorName="Diva J Amon, Amanda F Ziegler" latitude="12.5814" location="UK Seabed Resources Ltd exploration contract area (UK- 1)" longitude="-116.7244" pageId="0" pageNumber="11794" specimenCount="1" specimenCount-adult="1" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; lifeStage:
<specimenType id="9EA6B4890CC0832F765B80A817EC2CB1" pageId="0" pageNumber="11794">Adult</specimenType>
; behavior: On seafloor; occurrenceStatus: present; preparations: Imaged only; Taxon: taxonConceptID: Deimacf.validum; scientificName: Deimavalidum; kingdom: Animalia; phylum: Echinodermata; class: Holothuroidea; order: Elasipodida; family: Deimatidae; genus: Deima; taxonRank: species; scientificNameAuthorship: Théel, 1879; Location: waterBody: Pacific Ocean; stateProvince: Clarion-Clipperton Zone; locality:
<location id="39118D4CB57F2ECE0B24E3037A1EC9D8" LSID="urn:lsid:plazi:treatment:A8D47E1D2D3901DB8F688B8C6FBAC93B:39118D4CB57F2ECE0B24E3037A1EC9D8" latitude="12.5814" longitude="-116.7244" name="UK Seabed Resources Ltd exploration contract area (UK- 1)" pageId="0" pageNumber="11794">UK Seabed Resources Ltd exploration contract area (UK-1)</location>
; verbatimLocality: UK-1 Stratum B; maximumDepthInMeters: 4226; locationRemarks: RV Thompson Cruise TN319; decimalLatitude:
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; decimalLongitude:
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; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 25; Identification: identifiedBy:
<determinerName id="28FB98DFB6C9805B515D0A404B7438A3" pageId="0" pageNumber="11794">Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler</determinerName>
; dateIdentified: 2015; identificationRemarks: Identified only from imagery; identificationQualifier: cf.; Event: samplingProtocol:
<collectingMethod id="1383D50E07A9D1ACC19DE4F6A1A3C562" pageId="0" pageNumber="11794">Autonomous Underwater Vehicle</collectingMethod>
; eventDate:
<collectingDate id="02B2E870476BDC85376A38D81330115C" pageId="0" pageNumber="11794" value="2015-09-03">2015-03-09</collectingDate>
; eventTime: 8:42; habitat: Abyssal polymetallic-nodule field; fieldNumber: Dive 6 (AV06); Record Level: language: en; institutionCode:
<collectionCode id="82A1DF757B09CA585E1B350A97A0E267" pageId="0" pageNumber="11794">UHM</collectionCode>
; datasetName: ABYSSLINE; basisOfRecord: HumanObservation
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9BE5EBD0595E2B067535AE0A4822D244" pageId="0" pageNumber="11794" type="notes">
<paragraph id="CFE2DD7D35AD5D4333B96A895F2DB5F9" pageId="0" pageNumber="11794">Notes</paragraph>
<paragraph id="01562BACD6AA21245C3798A5B132CFCF" pageId="0" pageNumber="11794">Fig. 40</paragraph>
</subSubSection>
</treatment>
</document>