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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.948.51671" ID-GBIF-Dataset="9b2d84ae-218e-4cb1-88f2-76f1840067e9" ID-PMC="PMC7381483" ID-Pensoft-Pub="1313-2970-948-1" ID-Pensoft-UUID="52F291E3803D593EBF5741BFB13193FA" ID-PubMed="32765170" ID-ZooBank="20E7C61357714F328F6C44A7E84AFA68" ModsDocID="1313-2970-948-1" checkinTime="1594659732190" checkinUser="pensoft" docAuthor="Sutcharit, Chirasak, Thach, Phanara, Chhuoy, Samol, Ngor, Peng Bun, Jeratthitikul, Ekgachai, Siriwut, Warut, Srisonchai, Ruttapon, Ng, Ting Hui, Pholyotha, Arthit, Jirapatrasilp, Parin &amp; Panha, Somsak" docDate="2020" docId="A8BA2B42822F538E889B28DC9E7AA101" docLanguage="en" docName="ZooKeys 948: 1-46" docOrigin="ZooKeys 948" docSource="http://dx.doi.org/10.3897/zookeys.948.51671" docTitle="Lagocheilus klobukowskii" docType="treatment" docVersion="4" id="52F291E3803D593EBF5741BFB13193FA" lastPageNumber="1" masterDocId="52F291E3803D593EBF5741BFB13193FA" masterDocTitle="Annotated checklist of the land snail fauna from southern Cambodia (Mollusca, Gastropoda)" masterLastPageNumber="46" masterPageNumber="1" pageNumber="1" updateTime="1668169087223" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Annotated checklist of the land snail fauna from southern Cambodia (Mollusca, Gastropoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thach, Phanara</mods:namePart>
<mods:affiliation>Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3659-6577</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chhuoy, Samol</mods:namePart>
<mods:affiliation>Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ngor, Peng Bun</mods:namePart>
<mods:affiliation>Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia &amp; Wonders of the Mekong Project, c / o IFReDI, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3477-9548</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Siriwut, Warut</mods:namePart>
<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Ting Hui</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5123-0039</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pholyotha, Arthit</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>948</mods:number>
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<mods:start>1</mods:start>
<mods:end>46</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.948.51671</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.948.51671</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-948-1</mods:identifier>
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<treatment ID-GBIF-Taxon="165073725" LSID="urn:lsid:plazi:treatment:A8BA2B42822F538E889B28DC9E7AA101" httpUri="http://treatment.plazi.org/id/A8BA2B42822F538E889B28DC9E7AA101" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="A8BA2B42-822F-538E-889B-28DC9E7AA101" authority="(Morlet, 1885)" baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">Lagocheilus klobukowskii (Morlet, 1885)</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living snails of A Lagocheilus klobukowskii (Morlet, 1885) B Lagocheilus landesi (Morlet, 1885) C Pupina crosseana Morlet, 1883 D Valiguna siamensis (Martens, 1867) E, F Valiguna sp. E notum or dorsal view and F hyponotum or ventral view G Succinea tenuis Morelet, 1865 and H Hypselostoma cambodjense Benthem Jutting, 1962. All not to scale." figureDoi="10.3897/zookeys.948.51671.figure3" httpUri="https://binary.pensoft.net/fig/430960" pageId="0" pageNumber="1">Figs 3A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">, 7A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Morlet" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">Cyclophorus klobukowskii</taxonomicName>
Morlet, 1885[1884]: 391, 392, pl. 12, fig. 1. Type locality: Near the Kamchay rapids, around the
<normalizedToken originalValue="Kébal-Réméas">Kebal-Remeas</normalizedToken>
cave (Kampot-Hatien road); commonly found on mountains, in forests, up to Compong-Som, and on the banks of
<normalizedToken originalValue="Tap-Chéang">Tap-Cheang</normalizedToken>
.
<bibRefCitation author="Fischer, PH" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B26" refString="Fischer, PH, 1973a. Les mollusques testaces du Cambodge. Premier partie: Introduction et Gasteropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge. Premier partie: Introduction et Gasteropodes Prosobranches." volume="90" year="1973 a">Fischer 1973a</bibRefCitation>
: 46, 47.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">Lagocheilus klobukowskii</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
: 19, 20, figs 9b, c, 18c.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
Locality no. 9: CUMZ-CM044 (7 shells), CUMZ-CM045 (12 specimens in ethanol). Locality no. 10: CUMZ-CM068 (3 specimens in ethanol). Locality no. 11: CUMZ-CM079 (2 shells). Locality no. 13: CUMZ-CM128 (3 shells), CM129 (53 specimens in ethanol; Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living snails of A Lagocheilus klobukowskii (Morlet, 1885) B Lagocheilus landesi (Morlet, 1885) C Pupina crosseana Morlet, 1883 D Valiguna siamensis (Martens, 1867) E, F Valiguna sp. E notum or dorsal view and F hyponotum or ventral view G Succinea tenuis Morelet, 1865 and H Hypselostoma cambodjense Benthem Jutting, 1962. All not to scale." figureDoi="10.3897/zookeys.948.51671.figure3" httpUri="https://binary.pensoft.net/fig/430960" pageId="0" pageNumber="1">3A</figureCitation>
). Locality no. 17: CUMZ-CM137 (18 shells; Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">7A, B</figureCitation>
). The snails were found to live on the ground among leaf litter and decaying wood, on tree trunks and limestone wall.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Cambodia and Laos (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species was described from
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
grotte de
<normalizedToken originalValue="Kébal-Réméas">Kebal-Remeas</normalizedToken>
(route de Kampot
<normalizedToken originalValue="à">a</normalizedToken>
Hatien)
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
. We collected topotypic specimens that tend to have a variable shell colour from yellowish (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">7A</figureCitation>
, see fig. 9b in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
for the syntype) to purplish-black (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">7B</figureCitation>
). This limestone associated species has a wide distribution from southern Cambodia to eastern Laos (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
). The snails are commonly found in montane forest, living on decaying wood, on tree trunks and exposed limestone.
</paragraph>
<caption doi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">
<emphasis italics="true" pageId="0" pageNumber="1">Lagocheilus klobukowskii</emphasis>
</taxonomicName>
(Morlet, 1885)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
yellowish morph and
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
purplish-black morph
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName authorityName="Morlet" authorityYear="1883" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupina crosseana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crosseana">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina crosseana</emphasis>
</taxonomicName>
Morlet, 1883
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Morelet" authorityYear="1865" class="Gastropoda" family="Succineidae" genus="Succinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Succinea tenuis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tenuis">
<emphasis italics="true" pageId="0" pageNumber="1">Succinea tenuis</emphasis>
</taxonomicName>
Morelet, 1865 and
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma benetuitum" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="benetuitum">
<emphasis italics="true" pageId="0" pageNumber="1">Hypselostoma benetuitum</emphasis>
</taxonomicName>
Vermeulen et al., 2019
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
from locality no. 11 (type locality) and
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
from locality no. 17.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">
<emphasis italics="true" pageId="0" pageNumber="1">Lagocheilus klobukowskii</emphasis>
</taxonomicName>
was originally placed in the genus
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclophorus</emphasis>
</taxonomicName>
and later was transferred to the genus
<taxonomicName authorityName="W.T.Blanford" authorityYear="1864" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lagocheilus</emphasis>
</taxonomicName>
(see
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
). The distinguishing characters from the genus
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclophorus</emphasis>
</taxonomicName>
are a conic shell, an aperture thickened (not expanded), and a thick calcareous, multispiral and plate-like operculum, whereas
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclophorus</emphasis>
</taxonomicName>
has a turbinate shell, a thick and expanded lip, and a corneous multispiral operculum.
</paragraph>
</subSubSection>
</treatment>
</document>