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, by monotypy.
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The
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species-group
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Defined by
<bibRefCitation id="0879F3B9FF8675023662F8192161F81E" author="Platnick" box="[285,539,1937,1962]" pageId="2" pageNumber="323" refString="Platnick, N. I. &amp; Paz, N. (1979) On the Cryptocellus magnus group (Arachnida, Ricinulei). American Museum Novitates, 2677, 1 - 9." type="journal article" year="1979">Platnick &amp; Paz (1979)</bibRefCitation>
for species with a straight and massive accessory piece of the male copulatory apparatus. The following species are included in this group:
<taxonomicName id="ABE8F5CBFF8675023422F83D23A2F846" authority="Dumitresco &amp; Juvara-Bals, 1977" authorityName="Dumitresco &amp; Juvara-Bals" authorityYear="1977" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="5E9C525AFF8675023422F83D2735F87A" box="[861,1103,1973,1998]" italics="true" pageId="2" pageNumber="323">Cryptocellus bordoni</emphasis>
(
<bibRefCitation id="0879F3B9FF867502331EF83D23AAF846" author="Dumitresco" pageId="2" pageNumber="323" refString="Dumitresco, M. &amp; Juvara-Bals, I. (1977) Position systematique de Heteroricinoides bordoni n. g. n. sp. dans la famille Ricinuleididae (Arachnida). Boletin de la Sociedad Venezolana de Espeleologia, 7, 147 - 180." type="journal article" year="1977">Dumitresco &amp; Juvara-Bals, 1977</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="14FFCED8FF8675023790F8512212F846" box="[239,360,2009,2034]" name="Venezuela" pageId="2" pageNumber="323">Venezuela</collectingCountry>
: Zulia);
<taxonomicName id="ABE8F5CBFF86750236A9F85120ABF846" authority="Cokendolpher, 2000" authorityName="Cokendolpher" authorityYear="2000" box="[470,977,2009,2034]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="5E9C525AFF86750236A9F85121ACF846" box="[470,726,2009,2034]" italics="true" pageId="2" pageNumber="323">Cryptocellus brignolii</emphasis>
<bibRefCitation id="0879F3B9FF867502359AF85120ABF846" author="Cokendolpher" box="[741,977,2009,2034]" pageId="2" pageNumber="323" refString="Cokendolpher, J. C. (2000) First Cryptocellus from Suriname (Ricinulei). Memorie della Societa Entomologica Italiana, 78, 515 - 520." type="journal article" year="2000">Cokendolpher, 2000</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="14FFCED8FF8675023497F8512723F846" box="[1000,1113,2009,2034]" name="Suriname" pageId="2" pageNumber="323">Suriname</collectingCountry>
: Paramaribo);
<taxonomicName id="ABE8F5CBFF8675033272F851227BFF04" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" lastPageId="3" lastPageNumber="324" order="Ricinulei" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8675033272F851227BFF04" italics="true" lastPageId="3" lastPageNumber="324" pageId="2" pageNumber="323">Cryptocellus chimaera</emphasis>
</taxonomicName>
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<emphasis id="5E9C525AFF877503366AFF1F2206FF04" bold="true" box="[277,380,151,176]" pageId="3" pageNumber="324">sp. nov.</emphasis>
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(
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: Esmeraldas);
<taxonomicName id="ABE8F5CBFF87750335C5FF1F271CFF04" authority="Ewing, 1929" authorityName="Ewing" authorityYear="1929" box="[698,1126,151,176]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF87750335C5FF1F20CFFF04" box="[698,949,151,176]" italics="true" pageId="3" pageNumber="324">Cryptocellus magnus</emphasis>
<bibRefCitation id="0879F3B9FF87750334B6FF1F271CFF04" author="Ewing" box="[969,1126,151,176]" pageId="3" pageNumber="324" refString="Ewing, H. E. (1929) A synopsis of the American arachnids of the primitive order Ricinulei. Annals of the Entomological Society of America, 22, 583 - 600. http: // dx. doi. org / 10.1093 / aesa / 22.4.583" type="journal article" year="1929">Ewing, 1929</bibRefCitation>
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(
<collectingCountry id="14FFCED8FF87750333FDFF1F278DFF04" box="[1154,1271,151,176]" name="Colombia" pageId="3" pageNumber="324">Colombia</collectingCountry>
: Magdalena);
<taxonomicName id="ABE8F5CBFF87750337E8FF342100FF60" authority="Platnick &amp; Paz, 1979" authorityName="Platnick &amp; Paz" authorityYear="1979" box="[151,634,188,213]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="narino">
<emphasis id="5E9C525AFF87750337E8FF342201FF60" box="[151,379,188,213]" italics="true" pageId="3" pageNumber="324">Cryptocellus narino</emphasis>
<bibRefCitation id="0879F3B9FF87750336FBFF342100FF60" author="Platnick" box="[388,634,188,213]" pageId="3" pageNumber="324" refString="Platnick, N. I. &amp; Paz, N. (1979) On the Cryptocellus magnus group (Arachnida, Ricinulei). American Museum Novitates, 2677, 1 - 9." type="journal article" year="1979">Platnick &amp; Paz, 1979</bibRefCitation>
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(
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: Antioquia, Boyacá, Tolima);
<taxonomicName id="ABE8F5CBFF8775033323FF342249FF4C" authority="Roewer, 1952" authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="pseudocellatus">
<emphasis id="5E9C525AFF8775033323FF3426E1FF61" box="[1116,1435,188,213]" italics="true" pageId="3" pageNumber="324">Cryptocellus pseudocellatus</emphasis>
<bibRefCitation id="0879F3B9FF87750337E8FF682249FF4C" author="Roewer" box="[151,307,223,248]" pageId="3" pageNumber="324" refString="Roewer, C. F. (1952) Neotropische Arachnida Arthrogastra, zumeist aus Peru. Senckenbergiana Biologica, 33 (1 / 3), 37 - 58." type="journal article" year="1952">Roewer, 1952</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="14FFCED8FF877503363DFF682201FF4C" box="[322,379,224,248]" name="Peru" pageId="3" pageNumber="324">Peru</collectingCountry>
: Cajamarca).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,515 @@
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<mods:title id="4AAEB1D4C26FA7A79AB3273AC23DF520">A remarkable new species of the magnus species-group of Cryptocellus (Arachnida, Ricinulei) from Ecuador, with observations on the taxonomy of the New World genera</mods:title>
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<treatment id="E4413F5EFF87750937E8FEC8272EFD1C" ID-DOI="http://doi.org/10.5281/zenodo.6080964" ID-GBIF-Taxon="120620734" ID-Zenodo-Dep="6080964" LSID="urn:lsid:plazi:treatment:E4413F5EFF87750937E8FEC8272EFD1C" httpUri="http://treatment.plazi.org/id/E4413F5EFF87750937E8FEC8272EFD1C" lastPageId="9" lastPageNumber="330" pageId="3" pageNumber="324">
<subSubSection id="24F2DDC3FF87750337E8FEC822C0FEC9" pageId="3" pageNumber="324" type="nomenclature">
<paragraph id="6C578E48FF87750337E8FEC82176FEEF" blockId="3.[151,524,320,381]" box="[151,524,320,347]" pageId="3" pageNumber="324">
<heading id="371F3924FF87750337E8FEC82176FEEF" bold="true" box="[151,524,320,347]" fontSize="11" level="1" pageId="3" pageNumber="324" reason="1">
<emphasis id="5E9C525AFF87750337E8FEC82176FEEF" bold="true" box="[151,524,320,347]" pageId="3" pageNumber="324">
<taxonomicName id="ABE8F5CBFF87750337E8FEC822DDFEEE" ID-CoL="ZVPS" box="[151,423,320,346]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF87750337E8FEC822DDFEEE" bold="true" box="[151,423,320,346]" italics="true" pageId="3" pageNumber="324">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF87750336D0FEC92176FEEF" box="[431,524,321,347]" pageId="3" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6C578E48FF87750337E8FEEC22C0FEC9" blockId="3.[151,524,320,381]" box="[151,442,356,381]" pageId="3" pageNumber="324">
(
<figureCitation id="F4D392CDFF87750337E1FEEC224CFEC9" box="[158,310,356,381]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 10" captionStart-2="FIGURES 11 18" captionStart-3="FIGURES 19 34" captionStartId-0="5.[151,264,1976,1998]" captionStartId-1="6.[151,264,1940,1962]" captionStartId-2="7.[151,264,1604,1626]" captionStartId-3="8.[151,264,1746,1768]" captionTargetBox-0="[262,1306,201,1920]" captionTargetBox-1="[338,1229,194,1880]" captionTargetBox-2="[156,1431,187,1575]" captionTargetBox-3="[170,1411,201,1715]" captionTargetId-0="figure@5.[253,1312,193,1932]" captionTargetId-1="figure@6.[336,1232,193,1896]" captionTargetId-2="figure@7.[151,1436,183,1587]" captionTargetId-3="figure@8.[162,1414,193,1730]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." captionText-1="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." captionText-2="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." captionText-3="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri-0="https://zenodo.org/record/255268/files/figure.png" httpUri-1="https://zenodo.org/record/255269/files/figure.png" httpUri-2="https://zenodo.org/record/255270/files/figure.png" httpUri-3="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Figures 126</figureCitation>
,
<figureCitation id="F4D392CDFF877503363EFEEC22F0FEC8" box="[321,394,356,380]" captionStart="FIGURES 47 54" captionStartId="12.[151,264,1897,1919]" captionTargetBox="[270,1318,198,1853]" captionTargetId="figure@12.[268,1319,195,1862]" captionTargetPageId="12" captionText="FIGURES 47 54. Translucent areas on carapace of males of the two American ricinuleid genera: carapace on dorsal view (left) and detail of area (right). 47 48, Cryptocellus chimaera sp. nov. Holotype (MEPN 7813). 49 50 Cryptocellus becki Platnick &amp; Shadab, 1977. (MCN- 002). 51 52, Pseudocellus pearsei (Chamberlin &amp; Ivie, 1938). (CNAN-Rc 00032). 53 54 Pseudocellus chankin Valdez-Mondragón and Francke, 2011. Holotype (CNAN-T 0263)." httpUri="https://zenodo.org/record/255274/files/figure.png" pageId="3" pageNumber="324">4748</figureCitation>
,
<figureCitation id="F4D392CDFF87750336EAFEEC22C8FEC9" box="[405,434,356,381]" captionStart="FIGURE 55" captionStartId="13.[151,250,1294,1316]" captionTargetBox="[154,1435,194,1268]" captionTargetId="figure@13.[151,1436,193,1273]" captionTargetPageId="13" captionText="FIGURE 55. Map of Northwestern South America, plotting known locality records of Cryptocellus chimaera sp. nov., Cryptocellus magnus Ewing, 1929 and Cryptocellus leleupi Cooreman, 1976 (nomen dubium)." httpUri="https://zenodo.org/record/255275/files/figure.png" pageId="3" pageNumber="324">55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF87750337E8FE242272FDE1" pageId="3" pageNumber="324" type="materials_examined">
<paragraph id="6C578E48FF87750337E8FE242272FDE1" blockId="3.[151,1436,428,2013]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337E8FE242236FE71" bold="true" box="[151,332,428,453]" pageId="3" pageNumber="324">
<typeStatus id="B35330EAFF87750337E8FE2423A8FE71" box="[151,210,428,453]" pageId="3" pageNumber="324">Type</typeStatus>
material.
</emphasis>
<emphasis id="5E9C525AFF8775033625FE24211AFE71" box="[346,608,428,453]" italics="true" pageId="3" pageNumber="324">
<typeStatus id="B35330EAFF8775033625FE2422BBFE71" box="[346,449,428,453]" pageId="3" pageNumber="324" type="holotype">Holotype</typeStatus>
male (adult)
</emphasis>
(
<collectionCode id="0AF9168DFF8775033509FE2421BCFE70" box="[630,710,428,452]" pageId="3" pageNumber="324">MEPN</collectionCode>
7813):
<collectingCountry id="14FFCED8FF8775033453FE2420C2FE70" box="[812,952,428,452]" name="Ecuador" pageId="3" pageNumber="324">ECUADOR</collectingCountry>
: Esmeraldas, Reserva Ecológica Mache- Chindul, Estación Biológica Bilsa (aprox. 00º1932 to
<geoCoordinate id="09DCE88FFF8775033461FE5820CFFE53" box="[798,949,464,488]" direction="north" orientation="latitude" pageId="3" pageNumber="324" precision="15" value="0.38">00º2248 N</geoCoordinate>
; 79º4505 to
<geoCoordinate id="09DCE88FFF877503331DFE582787FE53" box="[1122,1277,464,488]" direction="west" orientation="longitude" pageId="3" pageNumber="324" precision="15" value="-79.683334">79º4100 W</geoCoordinate>
,
<quantity id="AB1023ADFF8775033275FE4726F5FE5C" box="[1290,1423,463,488]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="7.5" metricValueMin="3.0" pageId="3" pageNumber="324" unit="m" value="525.0" valueMax="750.0" valueMin="300.0">300750 m</quantity>
), Bosque maduro, pitfall trap,
<date id="1856A888FF877503357AFE7C210DFDB9" box="[517,631,500,525]" pageId="3" pageNumber="324" value="2007-10-16">16.x.2007</date>
, R. Espinosa, coll. (80% ethanol-preserved). The specimen has the opisthosoma detached from the prosoma; left copulatory apparatus was dissected and stored in a microvial with the specimen.
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF87750337B8FDD72390FD51" pageId="3" pageNumber="324" type="etymology">
<paragraph id="6C578E48FF87750337B8FDD72390FD51" blockId="3.[151,1436,428,2013]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8FDD72234FDCC" bold="true" box="[199,334,607,632]" pageId="3" pageNumber="324">Etymology.</emphasis>
<taxonomicName id="ABE8F5CBFF8775033627FDD722B2FDCC" box="[344,456,607,632]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="335" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8775033627FDD722B2FDCC" box="[344,456,607,632]" italics="true" pageId="3" pageNumber="324">Chimaera</emphasis>
</taxonomicName>
is the Latin word for ΧίΜαιρα (ancient Greek), which refers to one of the fabulous creatures of Greek mythology whose body was a puzzle of body parts of different animals. It is used here as a noun in apposition, due the display in the new species of morphological features present in relatives of both New World genera.
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF87750337B8FD672769FA24" pageId="3" pageNumber="324" type="diagnosis">
<paragraph id="6C578E48FF87750337B8FD67265DFC08" blockId="3.[151,1436,428,2013]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8FD67223BFCBC" bold="true" box="[199,321,751,776]" pageId="3" pageNumber="324">Diagnosis.</emphasis>
<taxonomicName id="ABE8F5CBFF8775033636FD672133FCBC" box="[329,585,751,776]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8775033636FD672133FCBC" box="[329,585,751,776]" italics="true" pageId="3" pageNumber="324">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF877503352EFD6721D1FCBC" box="[593,683,751,776]" pageId="3" pageNumber="324" rank="species">
<emphasis id="5E9C525AFF877503352EFD6721D1FCBC" bold="true" box="[593,683,751,776]" pageId="3" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
is unique among species in the
<taxonomicName id="ABE8F5CBFF877503336BFD792711FCBC" box="[1044,1131,753,776]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF877503336BFD792711FCBC" box="[1044,1131,753,776]" italics="true" pageId="3" pageNumber="324">magnus</emphasis>
</taxonomicName>
group for which males are known in the very incrassate femur of leg II of male (
<figureCitation id="F4D392CDFF877503358AFC9C2039FC98" box="[757,835,788,813]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Figs. 1</figureCitation>
,
<figureCitation id="F4D392CDFF877503342EFC9C200AFC99" box="[849,880,788,813]" captionStart="FIGURES 11 18" captionStartId="7.[151,264,1604,1626]" captionTargetBox="[156,1431,187,1575]" captionTargetId="figure@7.[151,1436,183,1587]" captionTargetPageId="7" captionText="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri="https://zenodo.org/record/255270/files/figure.png" pageId="3" pageNumber="324">15</figureCitation>
). The male of
<taxonomicName id="ABE8F5CBFF877503336AFC9D27A6FC99" box="[1045,1244,788,813]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="pseudocellatus">
<emphasis id="5E9C525AFF877503336AFC9D27A6FC99" box="[1045,1244,788,813]" italics="true" pageId="3" pageNumber="324">C. pseudocellatus</emphasis>
</taxonomicName>
has not yet been discovered; however, the female lacks cuticular pits on the cucullus (
<bibRefCitation id="0879F3B9FF87750334B9FCBF266CFCE4" author="Platnick" box="[966,1302,823,848]" pageId="3" pageNumber="324" refString="Platnick, N. I. &amp; Shadab, M. U. (1977) On Amazonian Cryptocellus (Arachnida, Ricinulei). American Museum Novitates, 2633, 1 - 17." type="journal article" year="1977">Platnick &amp; Shadab 1977: 15</bibRefCitation>
) which are present in
<taxonomicName id="ABE8F5CBFF8775033672FCD522E2FCC1" box="[269,408,860,885]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8775033672FCD522E2FCC1" box="[269,408,860,885]" italics="true" pageId="3" pageNumber="324">C. chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF87750336E0FCD42280FCC1" box="[415,506,860,885]" pageId="3" pageNumber="324" rank="species">
<emphasis id="5E9C525AFF87750336E0FCD42280FCC1" bold="true" box="[415,506,860,885]" pageId="3" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
Large carapacial translucent areas, similar to those of
<taxonomicName id="ABE8F5CBFF877503331DFCD52794FCC1" box="[1122,1262,860,885]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF877503331DFCD52794FCC1" box="[1122,1262,860,885]" italics="true" pageId="3" pageNumber="324">C. chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF877503338AFCD42635FCC1" box="[1269,1359,860,885]" pageId="3" pageNumber="324" rank="species">
<emphasis id="5E9C525AFF877503338AFCD42635FCC1" bold="true" box="[1269,1359,860,885]" pageId="3" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
, occur within the species-group at least in
<taxonomicName id="ABE8F5CBFF877503355FFC0821E2FC2C" box="[544,664,896,920]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF877503355FFC0821E2FC2C" box="[544,664,896,920]" italics="true" pageId="3" pageNumber="324">C. magnus</emphasis>
</taxonomicName>
(
<figureCitation id="F4D392CDFF87750335D9FCF72057FC2C" box="[678,813,895,920]" captionStart="FIGURES 35 40" captionStartId="10.[151,264,1954,1976]" captionTargetBox="[218,1343,193,1915]" captionTargetId="figure@10.[218,1343,193,1932]" captionTargetPageId="10" captionText="FIGURES 35 40. Cryptocellus magnus Ewing, 1929. Male and female from Villa Leonor (AMNH). 35, Male, carapace, dorsal view. 36, Female, carapace, dorsal view. 37, Male, prosoma, ventral view (sternal region). 38, Male, cucullus, frontal view. 39, Male, opisthosoma, dorsal view. 40, Female opisthosoma, dorsal view." httpUri="https://zenodo.org/record/255272/files/figure.png" pageId="3" pageNumber="324">Figs. 3536</figureCitation>
) and
<taxonomicName id="ABE8F5CBFF8775033415FC08209BFC2C" box="[874,993,895,920]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="5E9C525AFF8775033415FC08209BFC2C" box="[874,993,895,920]" italics="true" pageId="3" pageNumber="324">C. bordoni</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0879F3B9FF8775033363FCF72618FC2C" author="Salvatierra" box="[1052,1378,895,920]" pageId="3" pageNumber="324" refString="Salvatierra, L. &amp; Tourinho, A. L. (2016) The integumentary ultrastructure of Cryptocellus bordoni Dumitresco and Juvara-Bals, 1976 (Arachnida, Ricinulei). Micron, 81, 1 - 19. http: // dx. doi. org / 10.1016 / j. micron. 2015.10.003" type="journal article" year="2016">Salvatierra &amp; Tourinho 2016</bibRefCitation>
: figs. 2a, 3a). The area occupied by these structures on the carapace is much larger in the new species (
<figureCitation id="F4D392CDFF87750333ABFC2C2660FC08" box="[1236,1306,932,957]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="6C578E48FF87750337B8FC4F2769FA24" blockId="3.[151,1436,428,2013]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8FC4F2217FC54" bold="true" box="[199,365,967,992]" pageId="3" pageNumber="324">Comparisons.</emphasis>
Based on the shape of the male copulatory apparatus,
<taxonomicName id="ABE8F5CBFF877503348EFC4F278EFC54" box="[1009,1268,967,992]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF877503348EFC4F278EFC54" box="[1009,1268,967,992]" italics="true" pageId="3" pageNumber="324">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8775033381FC4F2621FC54" box="[1278,1371,967,992]" pageId="3" pageNumber="324" rank="species">
<emphasis id="5E9C525AFF8775033381FC4F2621FC54" bold="true" box="[1278,1371,967,992]" pageId="3" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
most closely resembles
<taxonomicName id="ABE8F5CBFF8775033617FC652298FBB0" box="[360,482,1005,1028]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8775033617FC652298FBB0" box="[360,482,1005,1028]" italics="true" pageId="3" pageNumber="324">C. magnus</emphasis>
</taxonomicName>
. Apart from the differences referred to above,
<taxonomicName id="ABE8F5CBFF877503337FFC6527F7FBB1" box="[1024,1165,1004,1029]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF877503337FFC6527F7FBB1" box="[1024,1165,1004,1029]" italics="true" pageId="3" pageNumber="324">C. chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF87750333EAFC64278BFBB1" box="[1173,1265,1004,1029]" pageId="3" pageNumber="324" rank="species">
<emphasis id="5E9C525AFF87750333EAFC64278BFBB1" bold="true" box="[1173,1265,1004,1029]" pageId="3" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
can be readily recognized from
<taxonomicName id="ABE8F5CBFF8775033626FB9822A8FB9C" box="[345,466,1040,1064]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8775033626FB9822A8FB9C" box="[345,466,1040,1064]" italics="true" pageId="3" pageNumber="324">C. magnus</emphasis>
</taxonomicName>
in several aspects, including:
<emphasis id="5E9C525AFF8775033454FB982047FB93" box="[811,829,1040,1063]" italics="true" pageId="3" pageNumber="324">i)</emphasis>
Tegument covered with bristle-like, translucent setae (
<figureCitation id="F4D392CDFF87750337E0FBBC239CFBF8" box="[159,230,1076,1101]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Fig. 1</figureCitation>
);
<emphasis id="5E9C525AFF877503367AFBBC2264FBFF" box="[261,286,1076,1099]" italics="true" pageId="3" pageNumber="324">ii)</emphasis>
cucullus with abundant pronounced surface tubercles on foremost two thirds (
<figureCitation id="F4D392CDFF87750333B2FBBC2644FBF8" box="[1229,1342,1076,1101]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Figs. 23</figureCitation>
,
<figureCitation id="F4D392CDFF877503322FFBBC261BFBF8" box="[1360,1377,1076,1100]" captionStart="FIGURES 4 10" captionStartId="6.[151,264,1940,1962]" captionTargetBox="[338,1229,194,1880]" captionTargetId="figure@6.[336,1232,193,1896]" captionTargetPageId="6" captionText="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="3" pageNumber="324">4</figureCitation>
);
<emphasis id="5E9C525AFF8775033204FBBC26E6FBFF" box="[1403,1436,1076,1099]" italics="true" pageId="3" pageNumber="324">iii)</emphasis>
opisthosoma ovate, with median plate of tergite XII clearly wider than long (
<figureCitation id="F4D392CDFF877503335DFBDF27EEFBC4" box="[1058,1172,1111,1136]" captionStart="FIGURES 4 10" captionStartId="6.[151,264,1940,1962]" captionTargetBox="[338,1229,194,1880]" captionTargetId="figure@6.[336,1232,193,1896]" captionTargetPageId="6" captionText="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="3" pageNumber="324">Figs. 6, 8</figureCitation>
);
<emphasis id="5E9C525AFF87750333D0FBD027B7FBDB" box="[1199,1229,1112,1135]" italics="true" pageId="3" pageNumber="324">iv)</emphasis>
basal segment of pygidium with shallow notch on dorsal posterior border (
<figureCitation id="F4D392CDFF8775033466FBF42024FB20" box="[793,862,1148,1173]" captionStart="FIGURES 4 10" captionStartId="6.[151,264,1940,1962]" captionTargetBox="[338,1229,194,1880]" captionTargetId="figure@6.[336,1232,193,1896]" captionTargetPageId="6" captionText="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="3" pageNumber="324">Fig. 7</figureCitation>
);
<emphasis id="5E9C525AFF877503340BFBF420F0FB27" box="[884,906,1148,1171]" italics="true" pageId="3" pageNumber="324">v)</emphasis>
legs I and II with ventral tubercles from femur to metatarsus (
<figureCitation id="F4D392CDFF877503363FFB1722B6FB0C" box="[320,460,1183,1208]" captionStart="FIGURES 11 18" captionStartId="7.[151,264,1604,1626]" captionTargetBox="[156,1431,187,1575]" captionTargetId="figure@7.[151,1436,183,1587]" captionTargetPageId="7" captionText="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri="https://zenodo.org/record/255270/files/figure.png" pageId="3" pageNumber="324">Figs. 1418</figureCitation>
);
<emphasis id="5E9C525AFF877503369CFB28217BFB03" box="[483,513,1184,1207]" italics="true" pageId="3" pageNumber="324">vi)</emphasis>
leg IV noticeably more slender than leg III, such that the width of femur IV is approximately half that of femur III (
<figureCitation id="F4D392CDFF877503353CFB4C21FCFB68" box="[579,646,1220,1245]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Fig. 1</figureCitation>
);
<emphasis id="5E9C525AFF87750335DEFB4C21BDFB6F" box="[673,711,1220,1243]" italics="true" pageId="3" pageNumber="324">vii)</emphasis>
tarsal process of the copulatory apparatus shallowly bifid (
<figureCitation id="F4D392CDFF8775033210FB4C23CCFAB4" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Fig. 26</figureCitation>
), with pronounced pro-ventral median ledge and array of distinct ventral longitudinal keels; L1 lobe shorter than L2 (
<figureCitation id="F4D392CDFF87750337B8FA842267FA91" box="[199,285,1292,1317]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Fig. 26</figureCitation>
);
<emphasis id="5E9C525AFF877503364BFA85221BFA90" box="[308,353,1293,1316]" italics="true" pageId="3" pageNumber="324">viii)</emphasis>
accessory piece of the copulatory apparatus with pronounced ventral sub-basal widening (
<figureCitation id="F4D392CDFF8775033210FA8423CDFAFC" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Fig. 20</figureCitation>
); L lobe subtly curved and thin (
<figureCitation id="F4D392CDFF877503353CFAA721AAFAFC" box="[579,720,1327,1352]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Figs. 2324</figureCitation>
); apex with spiniform proventral process and without pro- or retrodorsal processes (
<figureCitation id="F4D392CDFF87750336E0FADC2283FAD9" box="[415,505,1364,1389]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Fig. 25</figureCitation>
); retrolateral surface with series of protruding carinae (
<figureCitation id="F4D392CDFF87750333E6FADC2650FAD9" box="[1177,1322,1364,1389]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="3" pageNumber="324">Figs. 2425</figureCitation>
). See the 'comparative diagnosis' section of
<taxonomicName id="ABE8F5CBFF8775033567FAF021F5FA24" box="[536,655,1400,1424]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8775033567FAF021F5FA24" box="[536,655,1400,1424]" italics="true" pageId="3" pageNumber="324">C. magnus</emphasis>
</taxonomicName>
later in this paper for comparison.
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF87750937B8FA1422D8FF04" lastPageId="9" lastPageNumber="330" pageId="3" pageNumber="324" type="description">
<paragraph id="6C578E48FF87750337B8FA142166FA01" blockId="3.[151,1436,428,2013]" box="[199,540,1436,1461]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8FA142223FA01" bold="true" box="[199,345,1436,1461]" pageId="3" pageNumber="324">Description.</emphasis>
Male (
<typeStatus id="B35330EAFF87750336D8FA14216AFA01" box="[423,528,1436,1461]" pageId="3" pageNumber="324" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="6C578E48FF87750337B8FA3722BEF9F1" blockId="3.[151,1436,428,2013]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8FA372231FA6C" box="[199,331,1471,1496]" italics="true" pageId="3" pageNumber="324">Coloration:</emphasis>
<figureCitation id="F4D392CDFF877503362BFA3722C0FA6C" box="[340,442,1471,1496]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Figs 13</figureCitation>
. Cucullus, carapace, sternal region and legs II reddish brown; pedipalps, legs I, III, IV, and dorsal surface of opisthosoma reddish; opisthosoma ventrally with a darkened rounded area, covering the central part of sternites XI, XII, and XIII anteriorly. Longitudinal carapacial translucent areas yellow, contrasting with the background color.
</paragraph>
<paragraph id="6C578E48FF87750337B8F9D822DBF939" blockId="3.[151,1436,428,2013]" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8F9D82251F9D3" box="[199,299,1616,1639]" italics="true" pageId="3" pageNumber="324">Setation:</emphasis>
<figureCitation id="F4D392CDFF877503364CF9C722D1F9DC" box="[307,427,1615,1640]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 10" captionStart-2="FIGURES 11 18" captionStartId-0="5.[151,264,1976,1998]" captionStartId-1="6.[151,264,1940,1962]" captionStartId-2="7.[151,264,1604,1626]" captionTargetBox-0="[262,1306,201,1920]" captionTargetBox-1="[338,1229,194,1880]" captionTargetBox-2="[156,1431,187,1575]" captionTargetId-0="figure@5.[253,1312,193,1932]" captionTargetId-1="figure@6.[336,1232,193,1896]" captionTargetId-2="figure@7.[151,1436,183,1587]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." captionText-1="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." captionText-2="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri-0="https://zenodo.org/record/255268/files/figure.png" httpUri-1="https://zenodo.org/record/255269/files/figure.png" httpUri-2="https://zenodo.org/record/255270/files/figure.png" pageId="3" pageNumber="324">Figs. 118</figureCitation>
. Body and appendages entirely covered with fine, bristle-like, translucent setae, which are sparse in sternal region.
</paragraph>
<paragraph id="6C578E48FF87750437B8F9102060FF66" blockId="3.[151,1436,428,2013]" lastBlockId="4.[151,1437,151,1910]" lastPageId="4" lastPageNumber="325" pageId="3" pageNumber="324">
<emphasis id="5E9C525AFF87750337B8F9102244F91B" box="[199,318,1688,1711]" italics="true" pageId="3" pageNumber="324">Carapace:</emphasis>
<figureCitation id="F4D392CDFF8775033636F91F22CCF904" box="[329,438,1687,1712]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="3" pageNumber="324">Figs. 12</figureCitation>
,
<figureCitation id="F4D392CDFF87750336BAF9102175F904" box="[453,527,1688,1712]" captionStart="FIGURES 47 54" captionStartId="12.[151,264,1897,1919]" captionTargetBox="[270,1318,198,1853]" captionTargetId="figure@12.[268,1319,195,1862]" captionTargetPageId="12" captionText="FIGURES 47 54. Translucent areas on carapace of males of the two American ricinuleid genera: carapace on dorsal view (left) and detail of area (right). 47 48, Cryptocellus chimaera sp. nov. Holotype (MEPN 7813). 49 50 Cryptocellus becki Platnick &amp; Shadab, 1977. (MCN- 002). 51 52, Pseudocellus pearsei (Chamberlin &amp; Ivie, 1938). (CNAN-Rc 00032). 53 54 Pseudocellus chankin Valdez-Mondragón and Francke, 2011. Holotype (CNAN-T 0263)." httpUri="https://zenodo.org/record/255274/files/figure.png" pageId="3" pageNumber="324">4748</figureCitation>
. Trapezoidal in shape, with lateral margins not parallel (narrowing anteriorly); anterior margin straight in dorsal aspect, slightly re-curved in frontal aspect; posterior margin gently re-curved; carapace longer than wide, widest at level between coxae II and III. Longitudinal translucent areas dorsallydirected in the lateral edges of carapace, without well-defined borders, covering between one third and half the length of the lateral margins, from level of coxae I to level between coxae II and III; translucent areas mostly covered with setae as rest of carapace, glabrous regions as narrow areas at the margin of carapace at level of coxae II. Carapace with symmetrical arrangement of tubercle-containing cuticular pits, as follows: about 17 pits along median longitudinal axis (posterior-most three pits larger); posterior margin with about six pits on each side of the midline (lateral two pairs wider); short oblique rows posterior to the translucent areas, each with about eight pits, the anteriormost pit which penetrates into the translucent areas; few additional isolated pits found between the translucent areas and anterior to these. Carapace without any other pronounced depressions. Granules (apart from those of pits) only found in a row along posterior margin.
</paragraph>
<paragraph id="6C578E48FF80750437B8FF522747FEA1" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8FF522249FF47" box="[199,307,218,243]" italics="true" pageId="4" pageNumber="325">Cucullus:</emphasis>
<figureCitation id="F4D392CDFF8075043641FF5222FCFF47" box="[318,390,218,243]" captionStart="FIGURES 4 10" captionStartId="6.[151,264,1940,1962]" captionTargetBox="[338,1229,194,1880]" captionTargetId="figure@6.[336,1232,193,1896]" captionTargetPageId="6" captionText="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="4" pageNumber="325">Fig. 4</figureCitation>
. Wider than long, noticeably widest anteriorly; with abundant strong surface tubercles on distal two thirds; with few tubercle-containing cuticular pits and devoid of furrows.
</paragraph>
<paragraph id="6C578E48FF80750437B8FE952008FEC3" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8FE952231FE82" box="[199,331,285,310]" italics="true" pageId="4" pageNumber="325">Chelicerae:</emphasis>
<figureCitation id="F4D392CDFF807504362BFE9522E1FE82" box="[340,411,285,310]" captionStart="FIGURES 4 10" captionStartId="6.[151,264,1940,1962]" captionTargetBox="[338,1229,194,1880]" captionTargetId="figure@6.[336,1232,193,1896]" captionTargetPageId="6" captionText="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="4" pageNumber="325">Fig. 5</figureCitation>
. Movable finger twice the length of fixed finger and more robust; movable finger armed with one noticeably large sub-basal tooth and six small teeth progressively decreasing in size; fixed finger with five teeth, basal and distal ones slightly larger than three middle ones.
</paragraph>
<paragraph id="6C578E48FF80750437B8FE09210FFDAB" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8FE092214FE2D" box="[199,366,385,410]" italics="true" pageId="4" pageNumber="325">Sternal region:</emphasis>
<figureCitation id="F4D392CDFF807504360AFE0922C3FE2D" box="[373,441,385,410]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="4" pageNumber="325">Fig. 3</figureCitation>
. Coxae I meeting tritosternum; coxae IIIV meeting entirely, progressively decreasing in length; II and III sub-rectangular, IV pear-shaped; coxae II with anterior and posterior margins parallel, each forming a straight line perpendicular to the median axis; suture lines of coxae III and IV each about half the length of that of coxae II (coxae II are larger). Cuticle with minute tubercles along coxal margins (more abundant on pedipalp coxae) and without cuticular pits.
</paragraph>
<paragraph id="6C578E48FF80750437B8FDAF265FFC64" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8FDAF221EFDF4" box="[199,356,551,576]" italics="true" pageId="4" pageNumber="325">Opisthosoma:</emphasis>
<figureCitation id="F4D392CDFF8075043614FDAF22CDFDF4" box="[363,439,551,576]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="4" pageNumber="325">Figs. 1</figureCitation>
,
<figureCitation id="F4D392CDFF80750436BAFDA0217AFDF4" box="[453,512,552,576]" captionStart="FIGURES 4 10" captionStartId="6.[151,264,1940,1962]" captionTargetBox="[338,1229,194,1880]" captionTargetId="figure@6.[336,1232,193,1896]" captionTargetPageId="6" captionText="FIGURES 4 10. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 8 10, Opisthosoma, dorsal, ventral and left lateral views, respectively." httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="4" pageNumber="325">810</figureCitation>
. Oblong truncate, longer than wide, widest at level of tergite XII. Median plates of tergites XIXIII with paired antero-lateral depressions and lateral margins approximately parallel; median plate of tergites XI and XIII approximately as wide as long, that of XII clearly wider than long, that of X slit-like trapezoidal. Central region of tergite XI median plate only slightly elevated; tergite XIII median plate with rear corners pointy, protruding laterally. Dorsal and ventral surfaces with symmetrical arrangement of tuberclecontaining cuticular pits, as follows: tergite XI median plate with eleven along anterior margin, five on posterior margin, fourfive on lateral margins; tergite XII median plate with seven on anterior margin, five on posterior margin, four on each lateral margin; tergite XIII median plate with seven on anterior margin, three on each lateral margin. Internal margin of lateral plates of tergites X to XIII with onetwo, sixseven, sixseven and fiveseven pits, respectively. Lateral margins of ventral surface with a longitudinal row of pits consisting of one on each side of sternite X, sixseven on XI, seveneight on XII, and ten on each side of XIII. Ventral pits also present on paired antero-lateral depressions of XIXIII. Outside of the pits, minute tubercles only found on median plate of tergite X. Basal segment of pygidium with shallow notch on dorsal posterior border; ventral border without notch.
</paragraph>
<paragraph id="6C578E48FF80750437B8FC5126F2FBC3" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8FC512245FC46" box="[199,319,985,1010]" italics="true" pageId="4" pageNumber="325">Pedipalps:</emphasis>
<figureCitation id="F4D392CDFF8075043638FC5122E0FC46" box="[327,410,985,1010]" captionStart="FIGURES 11 18" captionStartId="7.[151,264,1604,1626]" captionTargetBox="[156,1431,187,1575]" captionTargetId="figure@7.[151,1436,183,1587]" captionTargetPageId="7" captionText="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri="https://zenodo.org/record/255270/files/figure.png" pageId="4" pageNumber="325">Fig. 13</figureCitation>
. Without cuticular pits; with few tubercles on ventral surface of trochanters I and II. Femur dorsally convex, widened in basal half. Tibia longer than femur, with dorsal surface straight, slightly widened ventrally in basal third, and with array of shallow elongated tubercles [='elevated tubercles' of
<bibRefCitation id="0879F3B9FF807504327EFB94222BFBE2" author="Salvatierra" pageId="4" pageNumber="325" refString="Salvatierra, L. &amp; Tourinho, A. L. (2016) The integumentary ultrastructure of Cryptocellus bordoni Dumitresco and Juvara-Bals, 1976 (Arachnida, Ricinulei). Micron, 81, 1 - 19. http: // dx. doi. org / 10.1016 / j. micron. 2015.10.003" type="journal article" year="2016">Salvatierra &amp; Tourinho (2016)</bibRefCitation>
] on all surfaces in distal half. Movable claw about twice the length of fixed claw and more robust; fixed claw armed with minute teeth, movable claw toothless or feebly serrate (any teeth might have worn down).
</paragraph>
<paragraph id="6C578E48FF80750437B8FB092166F9DF" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8FB09227CFB2C" box="[199,262,1153,1176]" italics="true" pageId="4" pageNumber="325">Legs:</emphasis>
<figureCitation id="F4D392CDFF8075043672FBF72220FB2C" box="[269,346,1151,1176]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1976,1998]" captionTargetBox="[262,1306,201,1920]" captionTargetId="figure@5.[253,1312,193,1932]" captionTargetPageId="5" captionText="FIGURES 1 3. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region)." httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="4" pageNumber="325">Figs. 1</figureCitation>
,
<figureCitation id="F4D392CDFF8075043615FB082172FB2C" box="[362,520,1152,1176]" captionStart="FIGURES 11 18" captionStartId="7.[151,264,1604,1626]" captionTargetBox="[156,1431,187,1575]" captionTargetId="figure@7.[151,1436,183,1587]" captionTargetPageId="7" captionText="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri="https://zenodo.org/record/255270/files/figure.png" pageId="4" pageNumber="325">1112, 1418</figureCitation>
. Without cuticular pits; leg segments with minute, inconspicuous granules in the very basal and distal boundaries facing the neighbouring segments. Leg II noticeably long and widened, especially on femur which very incrassate; other legs decrease in width in the order III, I, IV. Tibia of legs I and II with ventral notch, shallow on tibia I, more pronounced on tibia II. Legs I and II with ventral tubercles on femur, patella (on I only basally), tibia, and metatarsus, sparse and moderate in leg I, more abundant and sharp-tipped in leg II; the larger tubercles, found in femur and tibia of leg II, are mostly arranged in pro- and retroventral rows; legs III and IV devoid of tubercles. Leg I tarsus elevated, dorsally rounded, with claws sheltered in a cavity with disto-ventral opening; leg II distal tarsomere at least three times longer than the preceding tarsomeres, which are small and subequal in size and shape, with claws sheltered in a cavity with elongated ventral opening [like that of
<taxonomicName id="ABE8F5CBFF8075043277FA0223A7FA71" class="Arachnida" family="Ricinoididae" genus="Pseudocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="5E9C525AFF8075043277FA0223A7FA71" italics="true" pageId="4" pageNumber="325">Pseudocellus boneti</emphasis>
</taxonomicName>
(Bolívar y Pieltain, 1942) see
<bibRefCitation id="0879F3B9FF807504353EFA24205FFA71" author="Talarico" box="[577,805,1452,1477]" pageId="4" pageNumber="325" refString="Talarico, G., Palacios-Vargas, J. G, Silva, M. F. &amp; Alberti, G. (2006) Ultrastructure of tarsal sensilla and other integument structures of two Pseudocellus species (Ricinulei, Arachnida). Journal of Morphology, 267, 441 - 463. http: // dx. doi. org / 10.1002 / jmor. 10415" type="journal article" year="2006">
Talarico
<emphasis id="5E9C525AFF80750435DAFA25219AFA71" box="[677,736,1452,1477]" italics="true" pageId="4" pageNumber="325">et al.</emphasis>
2006
</bibRefCitation>
: fig. 3b]; leg III unmodified two tarsomeres sub-equal, slightly longer than the small tarsomeres of leg II; leg IV tarsomeres similar to those of leg II, except for terminal tarsomere which is at least twice longer than the preceding tarsomeres; terminal tarsomere of legs III and IV with distal dorsal and ventral V-shaped invaginations so the claws are exposed on either aspect. Leg III metatarsus not inflated, moderately excavated, with pro- and retrolateral subapical lobes; lamina cyathiformis longer than high, obtuse. Trochanter IV unmodified.
</paragraph>
<paragraph id="6C578E48FF80750437B8F9FC276DF8C2" blockId="4.[151,1437,151,1910]" pageId="4" pageNumber="325">
<emphasis id="5E9C525AFF80750437B8F9FC22BDF938" box="[199,455,1652,1677]" italics="true" pageId="4" pageNumber="325">Copulatory apparatus:</emphasis>
<figureCitation id="F4D392CDFF80750436B1F9FC212EF938" box="[462,596,1652,1677]" captionStart="FIGURES 11 18" captionStartId="7.[151,264,1604,1626]" captionTargetBox="[156,1431,187,1575]" captionTargetId="figure@7.[151,1436,183,1587]" captionTargetPageId="7" captionText="FIGURES 11 18. Cryptocellus chimaera sp. nov. Male Holotype (MEPN 7813). 11 12, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 16 17, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri="https://zenodo.org/record/255270/files/figure.png" pageId="4" pageNumber="325">Figs. 1112</figureCitation>
,
<figureCitation id="F4D392CDFF8075043520F9FC21D2F939" box="[607,680,1652,1677]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="4" pageNumber="325">1926</figureCitation>
. Tarsal process S-shaped on dorsal aspect, with moderate ipsilateral rotation; somewhat canoe-shaped on lateral aspect, with pronounced pro-ventral median ledge and array of distinct ventral longitudinal keels; dorsal pro- and retrolateral margins sinuous; apex shallowly bifid forming pro- (L2) and retrolateral (L1) lobes; L1 lobe rather horn-like, with irregular margins, shorter than L2. Accessory piece massive from base to apex, predominantly straight except for pronounced ventral sub-basal widening; apex with thin, dorsally-curved, retro-ventral hook-like lobe (L ), and spiniform pro-ventral process; L lobe not differentiated; dorsal aspect of the accessory piece with longitudinal sperm transfer groove ending in distal opening, retrolateral aspect with series of longitudinal carinae, one of which is especially protruding.
</paragraph>
<caption id="3897DEC0FF81750537E8F8302102F858" httpUri="https://zenodo.org/record/255268/files/figure.png" pageId="5" pageNumber="326" targetBox="[262,1306,201,1920]" targetPageId="5">
<paragraph id="6C578E48FF81750537E8F8302102F858" blockId="5.[151,1436,1976,2028]" pageId="5" pageNumber="326">
<emphasis id="5E9C525AFF81750537E8F8302246F87A" bold="true" box="[151,316,1976,1998]" pageId="5" pageNumber="326">FIGURES 13.</emphasis>
<taxonomicName id="ABE8F5CBFF817505363AF8302155F87A" box="[325,559,1976,1998]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF817505363AF8302155F87A" box="[325,559,1976,1998]" italics="true" pageId="5" pageNumber="326">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8175053548F83021F0F87A" box="[567,650,1976,1998]" pageId="5" pageNumber="326" rank="species">
<emphasis id="5E9C525AFF8175053548F83021F0F87A" bold="true" box="[567,650,1976,1998]" pageId="5" pageNumber="326">sp. nov.</emphasis>
</taxonomicNameLabel>
Male Holotype (MEPN 7813). 1, Habitus, dorsal view. 2, Carapace, dorsal view. 3, Prosoma, ventral view (sternal region).
</paragraph>
</caption>
<caption id="3897DEC0FF82750637E8F81C213EF853" httpUri="https://zenodo.org/record/255269/files/figure.png" pageId="6" pageNumber="327" targetBox="[338,1229,194,1880]" targetPageId="6">
<paragraph id="6C578E48FF82750637E8F81C213EF853" blockId="6.[151,1436,1940,2023]" pageId="6" pageNumber="327">
<emphasis id="5E9C525AFF82750637E8F81C223CF81E" bold="true" box="[151,326,1940,1962]" pageId="6" pageNumber="327">FIGURES 410.</emphasis>
<taxonomicName id="ABE8F5CBFF8275063633F81C2149F81E" box="[332,563,1940,1962]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8275063633F81C2149F81E" box="[332,563,1940,1962]" italics="true" pageId="6" pageNumber="327">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8275063546F81C21F3F81E" box="[569,649,1940,1962]" pageId="6" pageNumber="327" rank="species">
<emphasis id="5E9C525AFF8275063546F81C21F3F81E" bold="true" box="[569,649,1940,1962]" pageId="6" pageNumber="327">sp. nov.</emphasis>
</taxonomicNameLabel>
Male Holotype (MEPN 7813). 4, Cucullus, frontal view. 5, Chelicerae, dorsal view. 6, Opisthosoma, median plate of tergite XII. 7, Opisthosoma, pygidium, posterior view. 810, Opisthosoma, dorsal, ventral and left lateral views, respectively.
</paragraph>
</caption>
<caption id="3897DEC0FF83750737E8F9CC22BBF961" httpUri="https://zenodo.org/record/255270/files/figure.png" pageId="7" pageNumber="328" targetBox="[156,1431,187,1575]" targetPageId="7">
<paragraph id="6C578E48FF83750737E8F9CC22BBF961" blockId="7.[151,1436,1604,1749]" pageId="7" pageNumber="328">
<emphasis id="5E9C525AFF83750737E8F9CC2223F9EE" bold="true" box="[151,345,1604,1626]" pageId="7" pageNumber="328">FIGURES 1118.</emphasis>
<taxonomicName id="ABE8F5CBFF837507361AF9CC2129F9EE" box="[357,595,1604,1626]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF837507361AF9CC2129F9EE" box="[357,595,1604,1626]" italics="true" pageId="7" pageNumber="328">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF837507351FF9CC21CDF9EE" box="[608,695,1604,1626]" pageId="7" pageNumber="328" rank="species">
<emphasis id="5E9C525AFF837507351FF9CC21CDF9EE" bold="true" box="[608,695,1604,1626]" pageId="7" pageNumber="328">sp. nov.</emphasis>
</taxonomicNameLabel>
Male Holotype (MEPN 7813). 1112, Right leg III, prolateral and retrolateral views, respectively. 13, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 14, Right leg I, tibia, prolateral view. 15, Right leg II, femur, prolateral view. 1617, Right leg II, tibia, prolateral and ventral views, respectively. 18, Right leg II, metatarsus, ventral view. Abbreviations:
<emphasis id="5E9C525AFF837507350AF92A21F6F903" box="[629,652,1698,1719]" italics="true" pageId="7" pageNumber="328">ac</emphasis>
, accessory piece;
<emphasis id="5E9C525AFF8375073425F92A2009F903" box="[858,883,1698,1719]" italics="true" pageId="7" pageNumber="328">Lc</emphasis>
, lamina cyathiformis;
<emphasis id="5E9C525AFF8375073313F92A27F1F903" box="[1132,1163,1698,1719]" italics="true" pageId="7" pageNumber="328">mP</emphasis>
, metatarsal process;
<emphasis id="5E9C525AFF837507320BF92A26EFF903" box="[1396,1429,1698,1719]" italics="true" pageId="7" pageNumber="328">MT</emphasis>
, metatarsus;
<emphasis id="5E9C525AFF837507366FF948225EF961" box="[272,292,1728,1749]" italics="true" pageId="7" pageNumber="328">tP</emphasis>
, tarsal process.
</paragraph>
</caption>
<paragraph id="6C578E48FF83750737B8F976277DF85A" blockId="7.[151,1436,1788,2030]" pageId="7" pageNumber="328">
<emphasis id="5E9C525AFF83750737B8F9762142F8A0" box="[199,568,1788,1813]" italics="true" pageId="7" pageNumber="328">Measurements of male (in mm).</emphasis>
Body length, excluding pygidium 7.85; cucullus 1.13 long, greatest width 1.80; carapace 2.60 long, 2.33 wide at level between coxae II and III (where widest); opisthosoma 4.33 long (excluding pygidium), 3.07 wide at level of tergite XII (where widest); median plate of tergite XI 1.33 long, 1.60 wide (where widest); median plate of tergite XII 1.13 long, 1.60 wide; median plate of tergite XIII 1.33 long, 1.67 wide; suture line of coxae II 0.57 long, of coxae III 0.31 long; pedipalp femur 1.27 long, greatest depth 0.54; pedipalp tibia 1.87 long, greatest depth 0.26; femur I 1.47 long, greatest width 0.60; femur II 2.93 long, 1.47 greatest width, 1.87 greatest depth; femur III 0.80 width; femur IV 0.47 width.
</paragraph>
<caption id="3897DEC0FF8C750837E8F95A21B1F86A" httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="8" pageNumber="329" targetBox="[170,1411,201,1715]" targetPageId="8">
<paragraph id="6C578E48FF8C750837E8F95A21B1F86A" blockId="8.[151,1436,1746,2014]" pageId="8" pageNumber="329">
<emphasis id="5E9C525AFF8C750837E8F95A2223F95C" bold="true" box="[151,345,1746,1768]" pageId="8" pageNumber="329">FIGURES 1934.</emphasis>
<taxonomicName id="ABE8F5CBFF8C7508361BF95A212BF95C" box="[356,593,1746,1768]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8C7508361BF95A212BF95C" box="[356,593,1746,1768]" italics="true" pageId="8" pageNumber="329">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8C75083522F95A21C8F95C" box="[605,690,1746,1768]" pageId="8" pageNumber="329" rank="species">
<emphasis id="5E9C525AFF8C75083522F95A21C8F95C" bold="true" box="[605,690,1746,1768]" pageId="8" pageNumber="329">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="ABE8F5CBFF8C75083590F95A20B0F95C" box="[751,970,1746,1768]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8C75083590F95A20B0F95C" box="[751,970,1746,1768]" italics="true" pageId="8" pageNumber="329">Cryptocellus magnus</emphasis>
</taxonomicName>
, copulatory structures. 1926,
<taxonomicName id="ABE8F5CBFF8C75083264F95A238DF8B3" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8C75083264F95A238DF8B3" italics="true" pageId="8" pageNumber="329">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8C75083782F9792234F8B3" box="[253,334,1777,1799]" pageId="8" pageNumber="329" rank="species">
<emphasis id="5E9C525AFF8C75083782F9792234F8B3" bold="true" box="[253,334,1777,1799]" pageId="8" pageNumber="329">sp. nov.</emphasis>
</taxonomicNameLabel>
. Male Holotype (MEPN 7813), left copulatory apparatus. 1922, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 2733,
<taxonomicName id="ABE8F5CBFF8C750835ABF8A6273FF8F0" authority="Ewing, 1929" authorityName="Ewing" authorityYear="1929" box="[724,1093,1838,1860]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8C750835ABF8A620CAF8F0" box="[724,944,1838,1860]" italics="true" pageId="8" pageNumber="329">Cryptocellus magnus</emphasis>
Ewing, 1929
</taxonomicName>
. Male (AMNH), left copulatory apparatus. 2730, Dorsal, prolateral, ventral and retrolateral views, respectively. 3133, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34,
<taxonomicName id="ABE8F5CBFF8C750835C1F8E4275AF836" authority="Ewing, 1929" authorityName="Ewing" authorityYear="1929" box="[702,1056,1900,1922]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8C750835C1F8E420EFF836" box="[702,917,1900,1922]" italics="true" pageId="8" pageNumber="329">Cryptocellus magnus</emphasis>
Ewing, 1929
</taxonomicName>
. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations:
<emphasis id="5E9C525AFF8C75083546F8032137F814" box="[569,589,1931,1952]" italics="true" pageId="8" pageNumber="329">tP</emphasis>
, tarsal process;
<emphasis id="5E9C525AFF8C7508358BF8032071F814" box="[756,779,1931,1952]" italics="true" pageId="8" pageNumber="329">ac</emphasis>
, accessory piece (retrolateral carinae indicated by arrowheads);
<emphasis id="5E9C525AFF8C750837E8F82123D0F80B" box="[151,170,1961,1983]" italics="true" pageId="8" pageNumber="329">L </emphasis>
, lobe of
<emphasis id="5E9C525AFF8C75083671F822225CF80B" box="[270,294,1962,1983]" italics="true" pageId="8" pageNumber="329">ac</emphasis>
;
<emphasis id="5E9C525AFF8C75083648F8212203F80B" box="[311,377,1961,1983]" italics="true" pageId="8" pageNumber="329">L1L2</emphasis>
, lobes of
<emphasis id="5E9C525AFF8C75083698F8222287F80B" box="[487,509,1962,1983]" italics="true" pageId="8" pageNumber="329">tP</emphasis>
;
<emphasis id="5E9C525AFF8C75083571F822215EF80B" box="[526,548,1962,1983]" italics="true" pageId="8" pageNumber="329">sg</emphasis>
, sperm transfer groove;
<emphasis id="5E9C525AFF8C75083454F821203EF80B" box="[811,836,1961,1983]" italics="true" pageId="8" pageNumber="329">do</emphasis>
, distal opening of
<emphasis id="5E9C525AFF8C75083370F822275FF80B" box="[1039,1061,1962,1983]" italics="true" pageId="8" pageNumber="329">sg</emphasis>
;
<emphasis id="5E9C525AFF8C75083349F8222736F80B" box="[1078,1100,1962,1983]" italics="true" pageId="8" pageNumber="329">sp</emphasis>
, spiniform process;
<emphasis id="5E9C525AFF8C7508325AF8222656F80B" box="[1317,1324,1962,1983]" italics="true" pageId="8" pageNumber="329">t</emphasis>
, prodorsal tubercle;
<emphasis id="5E9C525AFF8C75083789F8402275F86A" box="[246,271,1992,2014]" italics="true" pageId="8" pageNumber="329">ml</emphasis>
, pro-ventral median ledge;
<emphasis id="5E9C525AFF8C75083558F8412142F86A" box="[551,568,1993,2014]" italics="true" pageId="8" pageNumber="329">st</emphasis>
, spermatheca.
</paragraph>
</caption>
<paragraph id="6C578E48FF8D750937B8FF1F22D8FF04" blockId="9.[151,1436,151,680]" box="[199,418,151,176]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8FF1F225FFF04" bold="true" box="[199,293,151,176]" pageId="9" pageNumber="330">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937B8FF3420A6FF61" box="[199,988,188,213]" pageId="9" pageNumber="330" type="distribution">
<paragraph id="6C578E48FF8D750937B8FF3420A6FF61" blockId="9.[151,1436,151,680]" box="[199,988,188,213]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8FF34221AFF61" bold="true" box="[199,352,188,213]" pageId="9" pageNumber="330">Distribution.</emphasis>
Known only from the
<typeStatus id="B35330EAFF8D7509351FFF3521EAFF61" box="[608,656,189,213]" pageId="9" pageNumber="330">type</typeStatus>
locality in
<collectingCountry id="14FFCED8FF8D75093470FF342016FF61" box="[783,876,188,213]" name="Ecuador" pageId="9" pageNumber="330">Ecuador</collectingCountry>
(
<figureCitation id="F4D392CDFF8D75093403FF3420AAFF61" box="[892,976,188,213]" captionStart="FIGURE 55" captionStartId="13.[151,250,1294,1316]" captionTargetBox="[154,1435,194,1268]" captionTargetId="figure@13.[151,1436,193,1273]" captionTargetPageId="13" captionText="FIGURE 55. Map of Northwestern South America, plotting known locality records of Cryptocellus chimaera sp. nov., Cryptocellus magnus Ewing, 1929 and Cryptocellus leleupi Cooreman, 1976 (nomen dubium)." httpUri="https://zenodo.org/record/255275/files/figure.png" pageId="9" pageNumber="330">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937B8FF572631FEF4" pageId="9" pageNumber="330" type="discussion">
<paragraph id="6C578E48FF8D750937B8FF572631FEF4" blockId="9.[151,1436,151,680]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8FF572241FF4C" bold="true" box="[199,315,223,248]" pageId="9" pageNumber="330">Remarks.</emphasis>
The
<typeStatus id="B35330EAFF8D7509360BFF6822DEFF4C" box="[372,420,224,248]" pageId="9" pageNumber="330">type</typeStatus>
locality of
<taxonomicName id="ABE8F5CBFF8D7509355CFF6821D4FF4C" box="[547,686,223,248]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8D7509355CFF6821D4FF4C" box="[547,686,223,248]" italics="true" pageId="9" pageNumber="330">C. chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8D750935CBFF572077FF4C" box="[692,781,223,248]" pageId="9" pageNumber="330" rank="species">
<emphasis id="5E9C525AFF8D750935CBFF572077FF4C" bold="true" box="[692,781,223,248]" pageId="9" pageNumber="330">sp. nov.</emphasis>
</taxonomicNameLabel>
(Bilsa Biological Station) is found in the lowlands west of the Andes, in North Western
<collectingCountry id="14FFCED8FF8D750936A3FE8C2146FEA9" box="[476,572,260,285]" name="Ecuador" pageId="9" pageNumber="330">Ecuador</collectingCountry>
. It is part of the Mache-Chindul Mointains, an area remarkable for its species diversity that has been proposed to be recognized as a Key Biodiversity Area (
<bibRefCitation id="0879F3B9FF8D7509337BFEAF2645FEF4" author="Ortega-Andrade" box="[1028,1343,295,320]" pageId="9" pageNumber="330" refString="Ortega-Andrade, H. M., Bermingham, J., Aulestia, C., &amp; Paucar, C. (2010) Herpetofauna of the Bilsa Biological Station, province of Esmeraldas, Ecuador. Check List, 6 (1), 119 - 154." type="journal article" year="2010">
Ortega-Andrade
<emphasis id="5E9C525AFF8D750933BCFEA12781FEF4" box="[1219,1275,295,320]" italics="true" pageId="9" pageNumber="330">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937B8FEC4272EFD1C" pageId="9" pageNumber="330" type="reference_group">
<paragraph id="6C578E48FF8D750937B8FEC4272EFD1C" blockId="9.[151,1436,151,680]" pageId="9" pageNumber="330">
<taxonomicName id="ABE8F5CBFF8D750937B8FEC422BDFED1" ID-CoL="ZVPS" box="[199,455,332,357]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8D750937B8FEC422BDFED1" box="[199,455,332,357]" italics="true" pageId="9" pageNumber="330">Cryptocellus chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8D750936B2FEC4215CFED1" box="[461,550,332,357]" pageId="9" pageNumber="330" rank="species">
<emphasis id="5E9C525AFF8D750936B2FEC4215CFED1" bold="true" box="[461,550,332,357]" pageId="9" pageNumber="330">sp. nov.</emphasis>
</taxonomicNameLabel>
, which is probably endemic to a restricted area, is the second ricinuleid species described from
<collectingCountry id="14FFCED8FF8D75093638FEE722DCFE3C" box="[327,422,367,392]" name="Ecuador" pageId="9" pageNumber="330">Ecuador</collectingCountry>
. The first species was
<taxonomicName id="ABE8F5CBFF8D750935DCFEE72731FE3C" authority="Cooreman, 1976" authorityName="Cooreman" authorityYear="1976" box="[675,1099,367,392]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="5E9C525AFF8D750935DCFEE720FFFE3C" box="[675,901,367,392]" italics="true" pageId="9" pageNumber="330">Cryptocellus leleupi</emphasis>
<bibRefCitation id="0879F3B9FF8D750934F2FEF82731FE3C" author="Cooreman" box="[909,1099,367,392]" pageId="9" pageNumber="330" refString="Cooreman, J. (1976) Description d'une stase nymphale d'un Ricinulei de l'Ecuador, Cryptocellus leleupi n. sp. In: Leleup, N. (Ed), Mission Zoologique Belge aux iles Galapagos et un Ecuador (N. et J. Leleup, 1964 - 1965), vol. 3. Koninklijk Museum voor Midden-Afrika: Bruxelles, 25 - 52." type="journal article" year="1976">Cooreman, 1976</bibRefCitation>
</taxonomicName>
, which
<bibRefCitation id="0879F3B9FF8D750933DDFEE726E6FE3C" author="Platnick" box="[1186,1436,367,392]" pageId="9" pageNumber="330" refString="Platnick, N. I. &amp; Paz, N. (1979) On the Cryptocellus magnus group (Arachnida, Ricinulei). American Museum Novitates, 2677, 1 - 9." type="journal article" year="1979">Platnick &amp; Paz (1979)</bibRefCitation>
regarded as a
<emphasis id="5E9C525AFF8D7509364DFE1D22ADFE19" box="[306,471,404,429]" italics="true" pageId="9" pageNumber="330">nomen dubium</emphasis>
due that it had been described on the basis of a protonymph, thus lacking all the adult features used for the taxonomy of the group. The
<typeStatus id="B35330EAFF8D75093587FE302052FE64" box="[760,808,440,464]" pageId="9" pageNumber="330">type</typeStatus>
locality of
<taxonomicName id="ABE8F5CBFF8D750934C5FE302754FE64" box="[954,1070,439,464]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="5E9C525AFF8D750934C5FE302754FE64" box="[954,1070,439,464]" italics="true" pageId="9" pageNumber="330">C. leleupi</emphasis>
</taxonomicName>
is “Oriente, Rio Negro,” with approximate geographic coordinates
<geoCoordinate id="09DCE88FFF8D75093541FE5421AAFE40" box="[574,720,476,500]" direction="south" orientation="latitude" pageId="9" pageNumber="330" precision="15" value="-1.4088888">01°2432 S</geoCoordinate>
<geoCoordinate id="09DCE88FFF8D750935A4FE542002FE40" box="[731,888,476,500]" direction="west" orientation="longitude" pageId="9" pageNumber="330" precision="15" value="-78.19111">78°1128 W</geoCoordinate>
in the eastern slopes of the Ecuadorian Andes, Tungurahua province. This is, some
<quantity id="AB1023ADFF8D7509354EFD8821FCFDAC" box="[561,646,511,536]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="330" unit="km" value="200.0">200 km</quantity>
SE of the collection locality of
<taxonomicName id="ABE8F5CBFF8D75093493FD88270DFDAC" ID-CoL="ZVPS" box="[1004,1143,511,536]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8D75093493FD88270DFDAC" box="[1004,1143,511,536]" italics="true" pageId="9" pageNumber="330">C. chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8D75093301FE7727A2FDAC" box="[1150,1240,511,536]" pageId="9" pageNumber="330" rank="species">
<emphasis id="5E9C525AFF8D75093301FE7727A2FDAC" bold="true" box="[1150,1240,511,536]" pageId="9" pageNumber="330">sp. nov.</emphasis>
</taxonomicNameLabel>
and separated by the Andes (
<figureCitation id="F4D392CDFF8D75093667FDAC2216FD89" box="[280,364,548,573]" captionStart="FIGURE 55" captionStartId="13.[151,250,1294,1316]" captionTargetBox="[154,1435,194,1268]" captionTargetId="figure@13.[151,1436,193,1273]" captionTargetPageId="13" captionText="FIGURE 55. Map of Northwestern South America, plotting known locality records of Cryptocellus chimaera sp. nov., Cryptocellus magnus Ewing, 1929 and Cryptocellus leleupi Cooreman, 1976 (nomen dubium)." httpUri="https://zenodo.org/record/255275/files/figure.png" pageId="9" pageNumber="330">Fig. 55</figureCitation>
). Adult ricinuleids from this area have not been reported; therefore,
<taxonomicName id="ABE8F5CBFF8D75093313FDAD27A3FD89" box="[1132,1241,548,573]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="5E9C525AFF8D75093313FDAD27A3FD89" box="[1132,1241,548,573]" italics="true" pageId="9" pageNumber="330">C. leleupi</emphasis>
</taxonomicName>
is better retained as
<emphasis id="5E9C525AFF8D750937CBFDC12220FDD4" box="[180,346,583,608]" italics="true" pageId="9" pageNumber="330">nomen dubium</emphasis>
until such material becomes available and allows the recognition of the species. Different locality records of ricinuleids usually yield different species, and the likelihood that the
<typeStatus id="B35330EAFF8D75093343FDE427D3FD31" box="[1084,1193,620,645]" pageId="9" pageNumber="330" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="ABE8F5CBFF8D750933AAFDE5263FFD31" box="[1237,1349,620,645]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="5E9C525AFF8D750933AAFDE5263FFD31" box="[1237,1349,620,645]" italics="true" pageId="9" pageNumber="330">C. leleupi</emphasis>
</taxonomicName>
and
<taxonomicName id="ABE8F5CBFF8D750932FDFDE5227BFD1C" ID-CoL="ZVPS" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8D750932FDFDE5227BFD1C" italics="true" pageId="9" pageNumber="330">C. chimaera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45AFEF21FF8D75093678FD07221BFD1C" box="[263,353,655,680]" pageId="9" pageNumber="330" rank="species">
<emphasis id="5E9C525AFF8D75093678FD07221BFD1C" bold="true" box="[263,353,655,680]" pageId="9" pageNumber="330">sp. nov.</emphasis>
</taxonomicNameLabel>
, on different sides of the Andes, could be conspecific is negligible.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,204 @@
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<mods:title id="68B419B2058257149423D68A4D8A8E8C">A remarkable new species of the magnus species-group of Cryptocellus (Arachnida, Ricinulei) from Ecuador, with observations on the taxonomy of the New World genera</mods:title>
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<treatment id="E4413F5EFF8D750937E8FD732792F8A9" ID-DOI="http://doi.org/10.5281/zenodo.6080966" ID-GBIF-Taxon="120620735" ID-Zenodo-Dep="6080966" LSID="urn:lsid:plazi:treatment:E4413F5EFF8D750937E8FD732792F8A9" httpUri="http://treatment.plazi.org/id/E4413F5EFF8D750937E8FD732792F8A9" lastPageNumber="330" pageId="9" pageNumber="330">
<subSubSection id="24F2DDC3FF8D750937E8FD73220EFC83" pageId="9" pageNumber="330" type="nomenclature">
<paragraph id="6C578E48FF8D750937E8FD73214CFCA2" blockId="9.[151,566,763,823]" box="[151,566,763,790]" pageId="9" pageNumber="330">
<heading id="371F3924FF8D750937E8FD73214CFCA2" bold="true" box="[151,566,763,790]" fontSize="11" level="1" pageId="9" pageNumber="330" reason="1">
<taxonomicName id="ABE8F5CBFF8D750937E8FD73214CFCA2" ID-CoL="ZVQC" authority="Ewing, 1929" authorityName="Ewing" authorityYear="1929" box="[151,566,763,790]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8D750937E8FD73214CFCA2" bold="true" box="[151,566,763,790]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937E8FD7322EFFCA1" bold="true" box="[151,405,763,789]" italics="true" pageId="9" pageNumber="330">Cryptocellus magnus</emphasis>
<bibRefCitation id="0879F3B9FF8D750936E2FD74214CFCA2" author="Ewing" box="[413,566,764,790]" pageId="9" pageNumber="330" refString="Ewing, H. E. (1929) A synopsis of the American arachnids of the primitive order Ricinulei. Annals of the Entomological Society of America, 22, 583 - 600. http: // dx. doi. org / 10.1093 / aesa / 22.4.583" type="journal article" year="1929">Ewing, 1929</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6C578E48FF8D750937E8FC96220EFC83" blockId="9.[151,566,763,823]" box="[151,372,798,823]" pageId="9" pageNumber="330">
(
<figureCitation id="F4D392CDFF8D750937E1FC96223EFC83" box="[158,324,798,823]" captionStart-0="FIGURES 19 34" captionStart-1="FIGURES 35 40" captionStart-2="FIGURES 41 46" captionStartId-0="8.[151,264,1746,1768]" captionStartId-1="10.[151,264,1954,1976]" captionStartId-2="11.[151,264,1146,1168]" captionTargetBox-0="[170,1411,201,1715]" captionTargetBox-1="[218,1343,193,1915]" captionTargetBox-2="[200,1376,181,1134]" captionTargetId-0="figure@8.[162,1414,193,1730]" captionTargetId-1="figure@10.[218,1343,193,1932]" captionTargetId-2="figure@11.[196,1382,180,1143]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." captionText-1="FIGURES 35 40. Cryptocellus magnus Ewing, 1929. Male and female from Villa Leonor (AMNH). 35, Male, carapace, dorsal view. 36, Female, carapace, dorsal view. 37, Male, prosoma, ventral view (sternal region). 38, Male, cucullus, frontal view. 39, Male, opisthosoma, dorsal view. 40, Female opisthosoma, dorsal view." captionText-2="FIGURES 41 46. Cryptocellus magnus Ewing, 1929. Male (AMNH). 41 42, Right leg III, prolateral and retrolateral views, respectively. 43, Left pedipalp tibia, movable (top) and fixed (bottom) claws. 44, Right leg I, tibia, prolateral view. 45, Left leg II, femur, prolateral view. 46, Right leg II, tibia, prolateral view. Abbreviations: ac, accessory piece; Lc, lamina cyathiformis; mP, metatarsal process; MT, metatarsus; tP, tarsal process." httpUri-0="https://zenodo.org/record/255271/files/figure.png" httpUri-1="https://zenodo.org/record/255272/files/figure.png" httpUri-2="https://zenodo.org/record/255273/files/figure.png" pageId="9" pageNumber="330">Figures 2746</figureCitation>
,
<figureCitation id="F4D392CDFF8D75093630FC962216FC83" box="[335,364,798,823]" captionStart="FIGURE 55" captionStartId="13.[151,250,1294,1316]" captionTargetBox="[154,1435,194,1268]" captionTargetId="figure@13.[151,1436,193,1273]" captionTargetPageId="13" captionText="FIGURE 55. Map of Northwestern South America, plotting known locality records of Cryptocellus chimaera sp. nov., Cryptocellus magnus Ewing, 1929 and Cryptocellus leleupi Cooreman, 1976 (nomen dubium)." httpUri="https://zenodo.org/record/255275/files/figure.png" pageId="9" pageNumber="330">55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937E8FCEE27CFFC2F" pageId="9" pageNumber="330" type="reference_group">
<paragraph id="6C578E48FF8D750937E8FCEE21D9FCC8" blockId="9.[151,1205,870,923]" box="[151,675,870,893]" pageId="9" pageNumber="330">
<treatmentCitationGroup id="4CF8A966FF8D750937E8FCEE21D9FCC8" box="[151,675,870,893]" pageId="9" pageNumber="330">
<taxonomicName id="ABE8F5CBFF8D750937E8FCEE2153FCC8" ID-CoL="ZVQC" authority="Ewing, 1929: 589" authorityName="Ewing" authorityPageNumber="589" authorityYear="1929" box="[151,553,870,893]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8D750937E8FCEE2217FCC9" box="[151,365,870,893]" italics="true" pageId="9" pageNumber="330">Cryptocellus magnus</emphasis>
<treatmentCitation id="ED49A859FF8D7509360BFCEE2153FCC8" author="Ewing" box="[372,553,870,892]" page="589" pageId="9" pageNumber="330" year="1929">
<bibRefCitation id="0879F3B9FF8D7509360BFCEE2282FCC8" author="Ewing" box="[372,504,870,892]" pageId="9" pageNumber="330" refString="Ewing, H. E. (1929) A synopsis of the American arachnids of the primitive order Ricinulei. Annals of the Entomological Society of America, 22, 583 - 600. http: // dx. doi. org / 10.1093 / aesa / 22.4.583" type="journal article" year="1929">Ewing, 1929</bibRefCitation>
: 589
</treatmentCitation>
</taxonomicName>
590, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6C578E48FF8D750937E8FC0D27CFFC2F" blockId="9.[151,1205,870,923]" box="[151,1205,901,923]" pageId="9" pageNumber="330">
<treatmentCitationGroup id="4CF8A966FF8D750937E8FC0D27CFFC2F" box="[151,1205,901,923]" pageId="9" pageNumber="330">
<taxonomicName id="ABE8F5CBFF8D750937E8FC0D2162FC2F" authority="Ewing, 1929: 591" authorityName="Ewing" authorityPageNumber="591" authorityYear="1929" box="[151,536,901,923]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="5E9C525AFF8D750937E8FC0D2227FC2F" box="[151,349,901,923]" italics="true" pageId="9" pageNumber="330">Cryptocellus manni</emphasis>
<treatmentCitation id="ED49A859FF8D7509361BFC0D2162FC2F" author="Ewing" box="[356,536,901,923]" page="591" pageId="9" pageNumber="330" year="1929">
<bibRefCitation id="0879F3B9FF8D7509361BFC0D2293FC2F" author="Ewing" box="[356,489,901,923]" pageId="9" pageNumber="330" refString="Ewing, H. E. (1929) A synopsis of the American arachnids of the primitive order Ricinulei. Annals of the Entomological Society of America, 22, 583 - 600. http: // dx. doi. org / 10.1093 / aesa / 22.4.583" type="journal article" year="1929">Ewing, 1929</bibRefCitation>
: 591
</treatmentCitation>
</taxonomicName>
592, figs 2, 5, 8 (synonymised by
<treatmentCitation id="ED49A859FF8D75093401FC0D27DDFC2F" author="Platnick" box="[894,1191,901,923]" page="5" pageId="9" pageNumber="330" year="1976">
<bibRefCitation id="0879F3B9FF8D75093401FC0D27F5FC2F" author="Platnick" box="[894,1167,901,923]" pageId="9" pageNumber="330" refString="Platnick, N. I. &amp; Shadab, M. U. (1976) On Colombian Cryptocellus (Arachnida, Ricinulei). American Museum Novitates, 2605, 1 - 8." type="journal article" year="1976">Platnick and Shadab, 1976</bibRefCitation>
: 5
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937E8FC4F2025FBC4" pageId="9" pageNumber="330" type="materials_examined">
<paragraph id="6C578E48FF8D750937E8FC4F22DEFBB1" blockId="9.[151,1436,967,1821]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937E8FC4F2231FC54" bold="true" box="[151,331,967,992]" pageId="9" pageNumber="330">
<typeStatus id="B35330EAFF8D750937E8FC4F23A8FC54" box="[151,210,967,992]" pageId="9" pageNumber="330">Type</typeStatus>
material.
</emphasis>
Female
<typeStatus id="B35330EAFF8D750936C7FC4F2160FC54" box="[440,538,967,992]" pageId="9" pageNumber="330" type="holotype">holotype</typeStatus>
from
<collectingCountry id="14FFCED8FF8D75093513FC40207EFC54" box="[620,772,968,992]" name="Colombia" pageId="9" pageNumber="330">COLOMBIA</collectingCountry>
, Magdalena, Cincinati; in National Museum of Natural History (not examined).
</paragraph>
<paragraph id="6C578E48FF8D750937B8FB872025FBC4" blockId="9.[151,1436,967,1821]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8FB8722CFFB9C" bold="true" box="[199,437,1039,1064]" pageId="9" pageNumber="330">Material examined.</emphasis>
<collectingCountry id="14FFCED8FF8D750936BFFB982120FB9C" box="[448,602,1040,1064]" name="Colombia" pageId="9" pageNumber="330">COLOMBIA</collectingCountry>
: Magdalena, Villa Leonor,
<quantity id="AB1023ADFF8D750934E4FB982089FB93" box="[923,1011,1040,1064]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="9" pageNumber="330" unit="m" value="1311.0">1311 m</quantity>
elev.,
<date id="1856A888FF8D7509333BFB8727C7FB9C" box="[1092,1213,1039,1064]" pageId="9" pageNumber="330" value="1975-04-12">12.iv.1975</date>
, J. A. Kochalka,
<specimenCount id="7AEE45C1FF8D750932F1FB9823AAFBF9" pageId="9" pageNumber="330" type="male">1 male</specimenCount>
,
<specimenCount id="7AEE45C1FF8D750937A2FBBC223AFBF9" box="[221,320,1076,1101]" pageId="9" pageNumber="330" type="female">1 female</specimenCount>
(
<collectionCode id="0AF9168DFF8D7509362FFBBC22D6FBF8" LSID="urn:lsid:biocol.org:col:34925" box="[336,428,1076,1100]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="330">AMNH</collectionCode>
). Magdalena,
<collectingCountry id="14FFCED8FF8D75093533FBBC21BAFBF9" box="[588,704,1076,1101]" name="Saint Pierre and Miquelon" pageId="9" pageNumber="330">San Pedro</collectingCountry>
de La Sierra,
<quantity id="AB1023ADFF8D7509341BFBBC20D7FBF8" box="[868,941,1076,1101]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="9" pageNumber="330" unit="m" value="960.0">960 m</quantity>
elev.,
<date id="1856A888FF8D75093486FBBC2713FBF9" box="[1017,1129,1076,1101]" pageId="9" pageNumber="330" value="1975-05-19">19.v.1975</date>
, J. A. Kochalka, 1 larva, 1 protonymph, 1 deutonymph, 2 tritonymphs,
<specimenCount id="7AEE45C1FF8D750935F8FBD02192FBC4" box="[647,744,1111,1136]" pageId="9" pageNumber="330" type="female">1 female</specimenCount>
(
<collectionCode id="0AF9168DFF8D75093588FBD02029FBC4" LSID="urn:lsid:biocol.org:col:34925" box="[759,851,1112,1136]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="330">AMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937B8FBF427DFFA6C" pageId="9" pageNumber="330" type="diagnosis">
<paragraph id="6C578E48FF8D750937B8FBF427DFFA6C" blockId="9.[151,1436,967,1821]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8FBF4229BFB21" bold="true" box="[199,481,1148,1173]" pageId="9" pageNumber="330">Comparative diagnosis.</emphasis>
<taxonomicName id="ABE8F5CBFF8D75093698FBF421AEFB20" box="[487,724,1148,1173]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8D75093698FBF421AEFB20" box="[487,724,1148,1173]" italics="true" pageId="9" pageNumber="330">Cryptocellus magnus</emphasis>
</taxonomicName>
can be readily recognized from
<taxonomicName id="ABE8F5CBFF8D75093341FBF527B3FB21" box="[1086,1225,1148,1173]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="chimaera" status="sp. nov.">
<emphasis id="5E9C525AFF8D75093341FBF527B3FB21" box="[1086,1225,1148,1173]" italics="true" pageId="9" pageNumber="330">C. chimaera</emphasis>
</taxonomicName>
<emphasis id="5E9C525AFF8D750933B0FBF42653FB21" bold="true" box="[1231,1321,1148,1173]" pageId="9" pageNumber="330">
<taxonomicNameLabel id="45AFEF21FF8D750933B0FBF42653FB21" box="[1231,1321,1148,1173]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in several aspects of the male morphology.
<emphasis id="5E9C525AFF8D7509356FFB28215BFB03" box="[528,545,1184,1207]" italics="true" pageId="9" pageNumber="330">i)</emphasis>
Tegument covered with lanceolate, iridescent setae (
<figureCitation id="F4D392CDFF8D75093303FB17267DFB0C" box="[1148,1287,1183,1208]" captionStart="FIGURES 35 40" captionStartId="10.[151,264,1954,1976]" captionTargetBox="[218,1343,193,1915]" captionTargetId="figure@10.[218,1343,193,1932]" captionTargetPageId="10" captionText="FIGURES 35 40. Cryptocellus magnus Ewing, 1929. Male and female from Villa Leonor (AMNH). 35, Male, carapace, dorsal view. 36, Female, carapace, dorsal view. 37, Male, prosoma, ventral view (sternal region). 38, Male, cucullus, frontal view. 39, Male, opisthosoma, dorsal view. 40, Female opisthosoma, dorsal view." httpUri="https://zenodo.org/record/255272/files/figure.png" pageId="9" pageNumber="330">Figs. 35, 39</figureCitation>
);
<emphasis id="5E9C525AFF8D75093262FB28264CFB03" box="[1309,1334,1184,1207]" italics="true" pageId="9" pageNumber="330">ii)</emphasis>
cucullus devoid of surface tubercles, surface granules restricted to anterior border (
<figureCitation id="F4D392CDFF8D7509336EFB4C2710FB68" box="[1041,1130,1220,1245]" captionStart="FIGURES 35 40" captionStartId="10.[151,264,1954,1976]" captionTargetBox="[218,1343,193,1915]" captionTargetId="figure@10.[218,1343,193,1932]" captionTargetPageId="10" captionText="FIGURES 35 40. Cryptocellus magnus Ewing, 1929. Male and female from Villa Leonor (AMNH). 35, Male, carapace, dorsal view. 36, Female, carapace, dorsal view. 37, Male, prosoma, ventral view (sternal region). 38, Male, cucullus, frontal view. 39, Male, opisthosoma, dorsal view. 40, Female opisthosoma, dorsal view." httpUri="https://zenodo.org/record/255272/files/figure.png" pageId="9" pageNumber="330">Fig. 38</figureCitation>
);
<emphasis id="5E9C525AFF8D750933FAFB4C27DCFB6F" box="[1157,1190,1220,1243]" italics="true" pageId="9" pageNumber="330">iii)</emphasis>
opisthosoma oblong elongate, with median plate of tergite XII approximately as long as wide (
<figureCitation id="F4D392CDFF8D750934A4FB6F274BFAB4" box="[987,1073,1255,1280]" captionStart="FIGURES 35 40" captionStartId="10.[151,264,1954,1976]" captionTargetBox="[218,1343,193,1915]" captionTargetId="figure@10.[218,1343,193,1932]" captionTargetPageId="10" captionText="FIGURES 35 40. Cryptocellus magnus Ewing, 1929. Male and female from Villa Leonor (AMNH). 35, Male, carapace, dorsal view. 36, Female, carapace, dorsal view. 37, Male, prosoma, ventral view (sternal region). 38, Male, cucullus, frontal view. 39, Male, opisthosoma, dorsal view. 40, Female opisthosoma, dorsal view." httpUri="https://zenodo.org/record/255272/files/figure.png" pageId="9" pageNumber="330">Fig. 39</figureCitation>
);
<emphasis id="5E9C525AFF8D7509333AFB60271EFB4B" box="[1093,1124,1256,1279]" italics="true" pageId="9" pageNumber="330">iv)</emphasis>
basal segment of pygidium without notch on either dorsal or ventral posterior borders;
<emphasis id="5E9C525AFF8D75093445FA84202AFA97" box="[826,848,1292,1315]" italics="true" pageId="9" pageNumber="330">v)</emphasis>
legs I and II without ventral tubercles;
<emphasis id="5E9C525AFF8D7509326BFA84264BFA97" box="[1300,1329,1292,1315]" italics="true" pageId="9" pageNumber="330">vi)</emphasis>
femur of legs III and IV similar in width;
<emphasis id="5E9C525AFF8D7509356AFAB82141FAF3" box="[533,571,1328,1351]" italics="true" pageId="9" pageNumber="330">vii)</emphasis>
tarsal process of the copulatory apparatus deeply bifid, spoon-like without median ledge, with no distinct ventral keels; L1 lobe longer than L2 (
<figureCitation id="F4D392CDFF8D75093499FADC2745FAD8" box="[998,1087,1364,1389]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="9" pageNumber="330">Fig. 33</figureCitation>
);
<emphasis id="5E9C525AFF8D75093325FADC27FDFADF" box="[1114,1159,1364,1387]" italics="true" pageId="9" pageNumber="330">viii)</emphasis>
accessory piece of the copulatory apparatus entirely straight, without sub-basal widening (
<figureCitation id="F4D392CDFF8D750934F2FAFF2098FA24" box="[909,994,1399,1424]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="9" pageNumber="330">Fig. 28</figureCitation>
); L lobe strongly curved and markedly robust (
<figureCitation id="F4D392CDFF8D7509378BFA1422FCFA00" box="[244,390,1436,1461]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="9" pageNumber="330">Figs. 3132</figureCitation>
); apex with spiniform retrodorsal process and pronounced prodorsal tubercle, without spiniform proventral process; retrolateral surface without conspicuous carinae (
<figureCitation id="F4D392CDFF8D75093370FA3727E2FA6C" box="[1039,1176,1471,1496]" captionStart="FIGURES 19 34" captionStartId="8.[151,264,1746,1768]" captionTargetBox="[170,1411,201,1715]" captionTargetId="figure@8.[162,1414,193,1730]" captionTargetPageId="8" captionText="FIGURES 19 34. Cryptocellus chimaera sp. nov. and Cryptocellus magnus, copulatory structures. 19 26, Cryptocellus chimaera sp. nov.. Male Holotype (MEPN 7813), left copulatory apparatus. 19 22, Dorsal, prolateral, ventral and retrolateral views, respectively. 23, Detail of distal half, prolateral view. 24, Idem, retrolateral view. 25, Accessory piece, apex, frontal view. 26, Tarsal process, ventro-frontal view. 27 33, Cryptocellus magnus Ewing, 1929. Male (AMNH), left copulatory apparatus. 27 30, Dorsal, prolateral, ventral and retrolateral views, respectively. 31 33, Detail of distal half on prolateral, retrolateral and ventro-frontal views, respectively. 34, Cryptocellus magnus Ewing, 1929. Female from San Pedro de La Sierra (AMNH), spermathecae. Abbreviations: tP, tarsal process; ac, accessory piece (retrolateral carinae indicated by arrowheads); L , lobe of ac; L 1 L 2, lobes of tP; sg, sperm transfer groove; do, distal opening of sg; sp, spiniform process; t, prodorsal tubercle; ml, pro-ventral median ledge; st, spermatheca." httpUri="https://zenodo.org/record/255271/files/figure.png" pageId="9" pageNumber="330">Figs. 3133</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937B8FA6C20C6FA49" box="[199,956,1508,1533]" pageId="9" pageNumber="330" type="distribution">
<paragraph id="6C578E48FF8D750937B8FA6C20C6FA49" blockId="9.[151,1436,967,1821]" box="[199,956,1508,1533]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8FA6C221AFA49" bold="true" box="[199,352,1508,1533]" pageId="9" pageNumber="330">Distribution.</emphasis>
Known only from Magdalena department,
<collectingCountry id="14FFCED8FF8D75093439FA6C20C2FA49" box="[838,952,1508,1533]" name="Colombia" pageId="9" pageNumber="330">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="24F2DDC3FF8D750937B8F9812792F8A9" pageId="9" pageNumber="330" type="description">
<paragraph id="6C578E48FF8D750937B8F98121BCF94C" blockId="9.[151,1436,967,1821]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8F9812142F9AB" box="[199,568,1543,1568]" italics="true" pageId="9" pageNumber="330">Measurements of male (in mm).</emphasis>
Body length, excluding pygidium 8.38; cucullus 1.33 long, greatest width 2.07; carapace 2.73 long, 2.60 wide at level of leg III (where widest); opisthosoma 5.00 long (excluding pygidium), 3.07 wide at level of tergite XII (where widest); median plate of tergite XI 1.60 long, 1.73 wide (where widest); median plate of tergite XII 1.40 long, 1.47 wide; median plate of tergite XIII 1.73 long, 1.73 wide; suture line of coxae II 0.56 long, of coxae III 0.38 long; pedipalp femur 1.33 long, greatest depth 0.57; pedipalp tibia 1.93 long, greatest depth 0.26; femur I 1.53 long, greatest width 0.67; femur II 2.73 long, 0.80 greatest width, 1.00 greatest depth; femur III 0.87 width; femur IV 0.66 width.
</paragraph>
<paragraph id="6C578E48FF8D750937B8F88C2792F8A9" blockId="9.[151,1436,967,1821]" box="[199,1256,1796,1821]" pageId="9" pageNumber="330">
<emphasis id="5E9C525AFF8D750937B8F88C226BF8A9" bold="true" box="[199,273,1796,1821]" pageId="9" pageNumber="330">Notes.</emphasis>
<taxonomicName id="ABE8F5CBFF8D75093667F88C217FF8A8" box="[280,517,1796,1821]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="5E9C525AFF8D75093667F88C217FF8A8" box="[280,517,1796,1821]" italics="true" pageId="9" pageNumber="330">Cryptocellus magnus</emphasis>
</taxonomicName>
was studied and described in detail by
<bibRefCitation id="0879F3B9FF8D750934C2F88C279EF8A9" author="Platnick" box="[957,1252,1796,1821]" pageId="9" pageNumber="330" refString="Platnick, N. I. &amp; Shadab, M. U. (1976) On Colombian Cryptocellus (Arachnida, Ricinulei). American Museum Novitates, 2605, 1 - 8." type="journal article" year="1976">Platnick &amp; Shadab (1976)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,190 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.681.13187" ID-GBIF-Dataset="ae626336-e3a1-46b6-a0b4-46336efd3a96" ID-PMC="PMC5523882" ID-Pensoft-Pub="1313-2970-681-119" ID-PubMed="28769722" ID-ZBK="3E0ADC40DD0843809542F2416A9A52A5" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-681-119" ModsDocOrigin="ZooKeys 681" ModsDocTitle="Taxonomic revision of imitating carpenter ants, CamponotussubgenusMyrmopytia (Hymenoptera, Formicidae) of Madagascar, using morphometry and qualitative traits" checkinTime="1502744592377" checkinUser="pensoft" docAuthor="Rasoamanana, Nicole, Csosz, Sandor &amp; Fisher, Brian L." docDate="2017" docId="E4414980EA2328492A28A772470D3A71" docLanguage="en" docName="ZooKeys 681: 119-152" docOrigin="ZooKeys 681" docSource="http://dx.doi.org/10.3897/zookeys.681.13187" docTitle="Camponotus karaha Rasoamanana, Csosz &amp; Fisher, 2017, sp. n." docType="treatment" docUuid="35B73CB2-30F5-4EAF-8321-F93DFADD95EB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="132" masterDocId="F977FFB9FF8EFF834D146F567237FFC9" masterDocTitle="Taxonomic revision of imitating carpenter ants, CamponotussubgenusMyrmopytia (Hymenoptera, Formicidae) of Madagascar, using morphometry and qualitative traits" masterLastPageNumber="152" masterPageNumber="119" pageNumber="128" updateTime="1668164553664" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of imitating carpenter ants, CamponotussubgenusMyrmopytia (Hymenoptera, Formicidae) of Madagascar, using morphometry and qualitative traits</mods:title>
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<mods:namePart>Rasoamanana, Nicole</mods:namePart>
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<mods:namePart>Csosz, Sandor</mods:namePart>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<treatment ID-GBIF-Taxon="132649180" LSID="urn:lsid:zoobank.org:act:35B73CB2-30F5-4EAF-8321-F93DFADD95EB" httpUri="http://treatment.plazi.org/id/E4414980EA2328492A28A772470D3A71" lastPageId="13" lastPageNumber="132" pageId="9" pageNumber="128">
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<paragraph pageId="9" pageNumber="128">
<taxonomicName LSID="http://zoobank.org/35B73CB2-30F5-4EAF-8321-F93DFADD95EB" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus karaha" order="Hymenoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="karaha">Camponotus karaha</taxonomicName>
<taxonomicNameLabel pageId="9" pageNumber="128">sp. n.</taxonomicNameLabel>
Figures 4B, 5B, 6B, 7B, 8A, 13, 14, 15, 18
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="128" type="type material">
<paragraph pageId="9" pageNumber="128">Type material investigated.</paragraph>
<paragraph pageId="9" pageNumber="128">Holotype worker: MADAGASCAR, Province Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, -14.435 49.76, 16.xi.2003, 775 m, rainforest, ex rotten log. (B.L. Fisher et al.). Collection code: BLF08983 Unique specimen identifier: CASENT0487715 (CASC).</paragraph>
<paragraph pageId="9" pageNumber="128">Paratype workers: Six workers with the same data as holotype, Collection code: BLF08983, Unique specimen identifiers: CASENT0487711 (2w), CASENT0487712 (2w), CASENT0487713 (2w), (CASC).</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="128" type="diagnosis">
<paragraph pageId="9" pageNumber="128">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="128">
Workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus karaha" order="Hymenoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="karaha">Camponotus karaha</taxonomicName>
can be differentiated from the other three species by the triangular form of the petiole in lateral view, and the protruding clypeus.
</paragraph>
<caption pageId="9" pageNumber="128">
<paragraph pageId="9" pageNumber="128">
Figure 13.
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus karaha" order="Hymenoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="karaha">Camponotus karaha</taxonomicName>
minor worker CASENT0067555. A Lateral view B Head in full-face view C Dorsal view.
</paragraph>
</caption>
<caption pageId="9" pageNumber="128">
<paragraph pageId="9" pageNumber="128">
Figure 14.
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus karaha" order="Hymenoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="karaha">Camponotus karaha</taxonomicName>
major worker CASENT0151921. A Lateral view B Head in full-face view C Dorsal view.
</paragraph>
</caption>
<caption pageId="9" pageNumber="128">
<paragraph pageId="9" pageNumber="128">
Figure 15. A Variant 1, a typical
<taxonomicName lsidName="C. karaha" pageId="9" pageNumber="128" rank="species" species="karaha">C. karaha</taxonomicName>
. CASENT0067555B Variant 2 from the north. CASENT0353274.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="129" pageId="9" pageNumber="128" type="description">
<paragraph pageId="9" pageNumber="128">Description of minor worker.</paragraph>
<paragraph pageId="9" pageNumber="128">Head longer than wide, anterior region of head elongated, caudate, sides of the head narrowed in front of eyes, CS 1.723 mm [1.35, 1.92] (n=47). Erect filiform setae on lateral margin of head and on gena. Eyes located anterior to the midlength of sides of head in full-face view, PoOC/CL 0.431 [0.404, 0.473]. Frontal carinae convex, FR/CS 0.238 [0.213, 0.267]. Scape distinctly surpassing posterior margin by almost half of its length, SL/CS 1.832 [1.651, 2.051]. Anterior margin of clypeus produced into an obtuse angle; in lateral view, clypeus protruding strongly, anterior portion may appear indented (Fig. 7B), masticatory margin with 6 teeth, palps long with respect to head size.</paragraph>
<paragraph pageId="9" pageNumber="128">Pronotum weakly undulant. Mesonotum straight and elongate, distinctly compressed laterally anterior to mesothoracic spiracles, MPD/CS 1.401 [1.196, 1.595]. Erect setae absent, appressed pubescence sparsely distributed. Mesothoracic spiracles prominent; propodeal dorsum protuberant, noticeably convex. ML/CS 2.203 [1.989, 2.405].</paragraph>
<paragraph pageId="9" pageNumber="128">Petiole conical, petiolar apex with an obtuse angle, posterior face of petiole flat to slightly convex, sides always distinct. PEW/CS 0.198 [0.169, 0.244].</paragraph>
<paragraph lastPageId="10" lastPageNumber="129" pageId="9" pageNumber="128">
Entire body generally black, pronotum may vary to brown, coxae a lighter color than mesosoma, basitarsus and up to 5 basal funiculi whitish. Mesonotum and pro
<pageBreakToken pageId="10" pageNumber="129" start="start">podeum</pageBreakToken>
smooth and glabrous to microsculptured with sparse appressed to subdecumbent setae.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="130" pageId="10" pageNumber="129" type="description">
<paragraph pageId="10" pageNumber="129">Description of major worker.</paragraph>
<paragraph pageId="10" pageNumber="129">Head subrectangular in full face view, occipital portion broad, sides of the head mostly parallel and weakly convex, CS 2.108 mm [1.92, 2.28] (n=9). Erect filiform setae present on entire head capsule. Eyes located at midlength of head capsule in full-face view. PrOc/CL 0.434 [0.417, 0.455]. Frontal carinae straight posterior to antennal insertion and curving smoothly toward the posterior margin of clypeus. FR/CS0.270 [0.255, 0.292]. Scape surpassing posterior margin by one fourth of its length. SL/ CS 0.971 [0.711, 1.331]. Anterior margin of clypeus projecting to an obtuse angle and, in lateral view, posterior portion of clypeus weakly produced dorsally (Fig. 7B) masticatory margin of mandible with 6 teeth, palps long with respect to head size.</paragraph>
<paragraph pageId="11" pageNumber="130">
<pageBreakToken pageId="11" pageNumber="130" start="start">Pronotum</pageBreakToken>
weakly undulant. Anterior region of mesonotum (immediately posterior to the pro-mesonotal suture) in profile rising above pronotum. Dorsal outline of mesonotum and propodeum form a continuous straight line interrupted by a shallow metanotal groove. Erect setae present, appressed pubescence sparsely distributed. Mesothoracic spiracle feebly produced laterally. ML/CS 1.872 [1.800, 2.009].
</paragraph>
<paragraph pageId="11" pageNumber="130">In profile, petiole conical, node summit acute. PEW/CS 0.222 [0.211, 0.250].</paragraph>
<paragraph pageId="11" pageNumber="130">Entire body black; femora and tibia dark brown, basitarsus of second and third legs light brown, funicular segments light brown, becoming dark apically. Pronotum and mesonotum microreticulate, sides of propodeum finely punctate, standing filiform setae present in all surfaces.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="130" type="distribution">
<paragraph pageId="11" pageNumber="130">Distribution and biology.</paragraph>
<paragraph pageId="11" pageNumber="130">
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus karaha" order="Hymenoptera" pageId="11" pageNumber="130" phylum="Arthropoda" rank="species" species="karaha">Camponotus karaha</taxonomicName>
is currently known from 14 localities along the eastern rainforest and montane rainforest of Madagascar at elevations ranging from 175 to 1325 m (Fig. 18B). Specimens have been collected on the ground in rotten logs and on low vegetation.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="130" type="etymology">
<paragraph pageId="11" pageNumber="130">Etymology.</paragraph>
<paragraph pageId="11" pageNumber="130">
The Malagasy word &quot;
<taxonomicName lsidName="karaha" pageId="11" pageNumber="130" rank="species" species="karaha">karaha</taxonomicName>
&quot; means similar, look-alike.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="131" pageId="11" pageNumber="130" type="notes on morphological variability">
<paragraph pageId="11" pageNumber="130">Notes on morphological variability.</paragraph>
<paragraph pageId="11" pageNumber="130">
Workers of
<taxonomicName lsidName="C. karaha" pageId="11" pageNumber="130" rank="species" species="karaha">C. karaha</taxonomicName>
exhibit morphological variability in qualitative traits such as sculpture, color of mesosoma, and profile of propodeum that differ between populations. This divergence is not, or barely, supported by multivariate analyses involving 19 quantitative traits. For this reason, we conclude that all populations examined represent a single species and ascribe the variation to intraspecific variability of populations occupying diverse sites, making geographic (e.g. elevation) or ecological factors as possible explanations for the variance. Populations from the north of its distribution range differ notably in shape of propodeal dorsum and petiolar node and further research on additional samples are needed to further evaluate species status of these populations. For now, we note the differences of the northern populations from the more typical
<taxonomicName lsidName="karaha" pageId="11" pageNumber="130" rank="species" species="karaha">karaha</taxonomicName>
.
</paragraph>
<paragraph pageId="11" pageNumber="130">
Variant 1. This variant is the typical
<taxonomicName lsidName="C. karaha" pageId="11" pageNumber="130" rank="species" species="karaha">C. karaha</taxonomicName>
, and is fairly widespread throughout the eastern rainforest of Madagascar. It can be recognized by having a propodeum dorsum smoothly convex in lateral view, declivitous face of petiole distinctly flat with defined lateral margins, petiolar apex forms a right angle (Fig. 16A).
</paragraph>
<paragraph pageId="12" pageNumber="131">
<pageBreakToken pageId="12" pageNumber="131" start="start">Variant</pageBreakToken>
2. Workers are known from montane rainforest of Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
and Parc National Masoala, Ambanizana, elevation 800-1100 m. It can be differentiated from the other morph by the propodeal dorsum almost flat in lateral view, declivitous face of petiole convex and not marginate laterally, petiolar apex acuminate (Fig. 16B).
</paragraph>
<caption pageId="12" pageNumber="131">
<paragraph pageId="12" pageNumber="131">
Figure 16.
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus longicollis" order="Hymenoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="longicollis">Camponotus longicollis</taxonomicName>
minor worker CASENT0191989. A Lateral view B Head in full-face view C Dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="13" pageNumber="132" type="materials_examined">
<paragraph pageId="13" pageNumber="132">
<pageBreakToken pageId="13" pageNumber="132" start="start">Additional</pageBreakToken>
material examined.
</paragraph>
<paragraph pageId="13" pageNumber="132">
Province Antsiranana: 6.5 km SSW Befingotra,
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Anjanaharibe-Sud, -14.75, 49.5, 875 m (B.L.Fisher) (CASC); 9.2 km WSW Befingotra,
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Anjanaharibe-Sud, -14.75, 49.46667, 1180 m (B.L.Fisher) (CASC); Betaolana Forest, along Bekona River, -14.52996, 49.44039, 880 m (B.L.Fisher et al.) (CASC); Betaolana forest, Ambodihazovolabe village along Ambolokopatrika river, -14.54484, 49.45163, 740 m (B.L.Fisher et al.) (CASC); Binara Forest, -13.26392, 49.59919, 1065 m (B.L.Fisher et al.) (CASC); Binara Forest, -13.26388, 49.60141, 900 m (B.L.Fisher et al.) (CASC); Galoko chain, Mont Kalabenono, -13.63999, 48.67374, 498 m (B.L.Fisher et al.) (CASC); Galoko chain, Mont Kalabenono, -13.64179, 48.67282, 643 m (B.L.Fisher et al.) (CASC); Galoko chain, Mont Kalabenono, -13.64609, 48.67732, 937 m (B.L.Fisher et al.) (CASC); Makirovana forest, -14.17066, 49.95409, 415 m (B.L.Fisher et al.) (CASC); Makirovana forest, -14.16044, 49.95216, 550 m (B.L.Fisher et al.) (CASC); Makirovana forest, -14.16666, 49.95, 715 m (B.L.Fisher et al.) (CASC); Makirovana forest, -14.16506, 49.9477, 900 m (B.L.Fisher et al.) (CASC); Parc National de Marojejy, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, -14.44333, 49.74333, 1325 m (B.L.Fisher) (CASC); Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, -14.435, 49.76, 775 m (B.L.Fisher et al.) (CASC); Parc National de Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, -14.43667, 49.775, 450 m (B.L.Fisher et al.) (CASC); RNI Marojejy, 10km NW Manantenina, -14.43333, 49.76667, 750 m (E.L. Quinter) (CASC). Province Fianarantsoa: 7.6 km 122
<normalizedToken originalValue="º">°</normalizedToken>
Kianjavato,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="Classée">Classee</normalizedToken>
Vatovavy, -21.4, 47.94, 175 m (B.L.Fisher et al.) (CASC); Foret
<normalizedToken originalValue="dAmbalagoavy">d'Ambalagoavy</normalizedToken>
Nord, Ikongo, Ambatombe, -21.8275, 47.33889, 625 m (R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
&amp; M.E. Irwin) (CASC);
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
de Vevembe, 66.6 km 293° Farafangana, -22.791, 47.18183, 600 m (B.L. Fisher et al.) (CASC). Province Toamasina: 6.9 km NE Ambanizana, Ambohitsitondroina, -13.56667, 50, 1080 m (B.L.Fisher) (CASC); Ambanizana, Parc National Masoala, -15.57167, 50.00611, 800-897 m (D. Andriamalala, D. Silva, et al.) (CASC); Ankerana, -18.4017, 48.80605, 1035 m (B.L.Fisher et al.) (CASC); Ankerana, -18.4061, 48.82029, 725 m (B.L.Fisher et al.) (CASC); Ankerana, -18.40062, 48.81311, 865 m (B.L.Fisher et al.) (CASC); Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 18.0 km 21° NNE Ambinanitelo, -15.18833, 49.615, 470 m (Fisher, Griswold et al.) (CASC); Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 19.5 km 27° NNE Ambinanitelo, -15.17833, 49.635, 1100 m (Fisher, Griswold et al.) (CASC); Parc National de Zahamena, Besaky River, -17.75244, 48.85321, 760 m (B.L.Fisher et al.) (CASC); Parc National de Zahamena, Sahavorondrano River, -17.75257, 48.85725, 765 m (B.L.Fisher et al.) (CASC); Parc National Mananara-Nord, 7.1 km 261° Antanambe, -16.455, 49.7875, 225 m (B.L.Fisher et al.) (CASC);
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
Nationale
<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
Betampona, Betampona 35.1 km NW Toamasina, -17.91801, 49.20074, 500 m (B.L.Fisher et al.) (CASC);
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
Naturelle Betampona, 34.1 km 332° Toamasina, -17.916135, 49.20185, 550 m (B.L.Fisher) (CASC).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,169 @@
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<subSubSection id="1493BB7D7675E453378A5D4BDDAD98C2" type="nomenclature">
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<taxonomicName id="97236595FB9FA0CD3ABD84036CFE5F9C" ID-CoL="33WM7" class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="variety" species="majalis" variety="majalis">
<emphasis id="CDA80602A5497B13D33274D6882B4074" italics="true">Dactylorhiza majalis</emphasis>
(Rchb.) P. F. Hunt &amp; Summerh. var. majalis
</taxonomicName>
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<subSubSection id="49C6E5896EBAD010E98F812B20077726" type="multiple">
<paragraph id="338D15C90B6DD3E1C7651041714D4DE0">
<normalizedToken id="DFB364316EE6B62E6DE37AE169A30001" originalValue="Varietät">Varietaet</normalizedToken>
ISFS: Checklist: 1014805
<taxonomicName id="CFE737332F323C8C7FA15AEA40CBB167" class="Liliopsida" family="Orchidaceae" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="family">Orchidaceae</taxonomicName>
<taxonomicName id="E5F23C094347AF94EE334ED8F1180225" class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="genus">Dactylorhiza</taxonomicName>
<taxonomicName id="5AA7D5BC5188747CBBAA42A067953A17" authority="(Rchb.) P. F. Hunt &amp; Summerh." authorityName="P. F. Hunt &amp; Summerh." baseAuthorityName="Rchb." class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="majalis">Dactylorhiza majalis (Rchb.) P. F. Hunt &amp; Summerh.</taxonomicName>
<taxonomicName id="59EFE94A2D226A1E347D6E5312AF58D3" class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="variety" species="majalis" variety="majalis">Dactylorhiza majalis (Rchb.) P. F. Hunt &amp; Summerh. var. majalis</taxonomicName>
</paragraph>
<paragraph id="3DA07625932245B3628C026FCC578FE6">
<heading id="40C6656BB258B8B31E2BE8CEFAEECE66">Zusammenfassung</heading>
KEINE ANGABE
</paragraph>
<paragraph id="2D9C1D898B811E960E67188B2A6A0E47">
<heading id="F85FB323C144CE771E02BA644D8304F3">
<normalizedToken id="71B3037E01ECF1C966C1C2644CE9B218" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="3B479FD1CF07EE9A5919B2DD88C108BB">
<heading id="F1EA341809410926F259F3FA76EA97E0">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="104DFF675129DDAB27CC8A7A96D0A978" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie Gefährdung Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie Gefährdung Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="661006E4FFB5B8BC4985FF5A89D5C75E">
<table id="1EE6E7BD33AFA18D03BEF9723D79F779" class="table" inLine="true">
<tr id="3DE68FE233701EEED20D615BBAEA4976">
<td id="2155B05A2C835CFA2BD1E71F5D988190">
<tbody id="1B866F602D135EA16D8E6881E8E4C566">KEINE ANGABE</tbody>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="6D93BBB90D4F539C872F05E5333030D9" type="biology_ecology">
<paragraph id="F910A5DDE11B0078C35CAB21E2BD225B">
<normalizedToken id="CDFE4EF874B563B4905070C9072BDF52" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="529950CE493168300EF081B0309DEA4E" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="22CCFD037FEB826151041FBE03B30649">
<table id="ED62756526548B089BD2FA4B2FED6B3B" class="table" inLine="true">
<tbody id="85436A2A5DBB076CB88547A51C33219C">
<tr id="15503E44ECF1543E580FFA719383D3B3">
<th id="E5B26D37E3EDD5B7D489FB033A04A4CE" colspan="2">
<emphasis id="69A751251BD5649F262983F09904C7B2" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="5D2DD2EA8809AD34B65BA0651EAD9EEE" colspan="2">
<emphasis id="920B217C7F0E1A32D795520984582AD7" bold="true">Klimafaktoren</emphasis>
</th>
<th id="AC1748F75C8A33760CF41380ACDC6A59" colspan="2">
<emphasis id="74C765021FA30CEEF8B643C7324E1CE9" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="9E60356DF36B7FD8BD1D860288EF0543">
<td id="C7520F3FE13C6D87D75F4884D35E65B2">Feuchtezahl F</td>
<td id="4E967F974720382D009375124230F022" style="text-align:left;width:100px">--</td>
<td id="121E79030EABF3546CDAC3B11F94CD2F">Lichtzahl L</td>
<td id="80AE8F4F0AEB8B16D52B488D75AAAEE7" style="text-align:left;width:60px">--</td>
<td id="80A898D1564F464014729CBE949A1D10">Salzzeichen</td>
<td id="8D9E07C5039C94DAC50170EF9BAD4A05" style="text-align:left;width:60px">--</td>
</tr>
<tr id="16CB7140F76B11D7377B70468EE8C892">
<td id="98AB02BC49B1FBDD50A718F670EB6C58">Reaktionszahl R</td>
<td id="245C1A5521552E6357EC976A4F1877A8" style="text-align:left">--</td>
<td id="078BD13695DAD1A68457BFF42C40FAD7">Temperaturzahl T</td>
<td id="10040AC7C58F8D1E226EFA5C3C5FF27B" style="text-align:left">--</td>
</tr>
<tr id="763C869BF8E4057F812C51D7F21D431C">
<td id="D25F863D5DC1D47670F7E323F5282A3B">
<normalizedToken id="9FB1D34DA005F59E035D91E936941DC4" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="BA6555856696D96966EB4592FED65C48" style="text-align:left">--</td>
<td id="F42CF3B6868EEBF12E7DED3CFF0CA19B">
<normalizedToken id="54102DB2908562F40FBA317125A58CAA" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="D674B8C535FE936E28BC09D1349644A7" style="text-align:left">--</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="8A476B39E771E490290563DBDDB4D4E3" type="reference_group">
<paragraph id="B8BBEEE6F57EB3044E36A76F77E687FA">
<heading id="5D1205FC723A1CCC014A2CD136334411">Nomenklatur</heading>
</paragraph>
<paragraph id="9256A86009BABB1A0533A73127F58B64">
<treatmentCitation id="F82A205BF62177F2882930511AFF9BCB">
<heading id="C50DE3510B28C4F694121E042FAE4D61">
<normalizedToken id="6732303CE626F29F7C824EC28C3E25F3" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="AABE25FCCBFA5C5AF9BCFA31C1076CAC" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="9D1EEAEF7240BED84CA037C6868501E0" ID-CoL="33WM7" class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="variety" species="majalis" variety="majalis">
<emphasis id="DA0DAA8A70A350B7DCC51C5953A86045" italics="true">Dactylorhiza majalis</emphasis>
(Rchb.) P. F. Hunt &amp; Summerh. var.
<emphasis id="AB7783DFDA7A6D8B1826894342839296" italics="true">majalis</emphasis>
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="0276A72BB8854FF5B5CBC059D90C6DB3" type="vernacular_names">
<paragraph id="22D35E5372DE2659B08578DC9C9BD7F8">Volksname</paragraph>
</subSubSection>
<subSubSection id="FF8F2C06C2014829B0EC11B3DE6A814F" type="multiple">
<paragraph id="34F9AF7DAC7FE6BE488029C10BC6DBB3">
Deutscher Name: -- Nom
<normalizedToken id="0096FBB346FF1BC5D16911F2AFB31BB1" originalValue="français">francais</normalizedToken>
: -- Nome italiano: --
</paragraph>
</subSubSection>
<subSubSection id="8F88444F3E65BC6E0289B3B78A08A1C2" type="conservation">
<paragraph id="BC2B1D9A8E5245D8CBCF5605DE43AE9F">
<heading id="DC041039284D8C7DDAFD221D6299ADE0">Status Indigenat</heading>
: -
</paragraph>
<paragraph id="8F61EC89BA4895923D5B68B105FDB386">
<emphasis id="9C221CD335562691EEF46F191A2F024A" bold="true">
<heading id="BD9430B72FB4246DCC6DFBC24D0491C5">
Liste der
<normalizedToken id="FB61FF80BF94D12D488BF000427294D5" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="89F70A43F64153DB7AF1FCE98F23F1CE" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
</subSubSection>
</treatment>
</document>