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<mods:title id="0F6691D0EAAA7753E467F546F2F2D0C7">A review of Scaphochlamys (Zingiberaceae) from Borneo, with description of eleven new species</mods:title>
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<treatment id="03D2ED3BEA03D45436D5FC8E7AEE4F71" LSID="urn:lsid:plazi:treatment:03D2ED3BEA03D45436D5FC8E7AEE4F71" httpUri="http://treatment.plazi.org/id/03D2ED3BEA03D45436D5FC8E7AEE4F71" lastPageNumber="276" pageId="45" pageNumber="276">
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<emphasis id="B90F803FEA03D45436D5FC8E7EFB4A7D" bold="true" box="[136,171,771,796]" pageId="45" pageNumber="276">24.</emphasis>
<taxonomicName id="4C7B27AEEA03D45436EAFC8C7DF04A7D" ID-CoL="6Y4X4" authority="Meekiong &amp; Hidir (2015: 34)" authorityName="Meekiong &amp; Hidir" authorityPageNumber="34" authorityYear="2015" box="[183,928,769,796]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="45" pageNumber="276" phylum="Tracheophyta" rank="species" species="samunsamensis">
<emphasis id="B90F803FEA03D45436EAFC8C7C7E4A7D" bold="true" box="[183,558,769,796]" italics="true" pageId="45" pageNumber="276">Scaphochlamys samunsamensis</emphasis>
<bibRefCitation id="EFEA21DCEA03D4543466FC8E7DF04A7D" author="Meekiong, K. &amp; Hidir, M. &amp; Yazid, K." box="[571,928,771,796]" pageId="45" pageNumber="276" pagination="31 - 44" refId="ref32559" refString="Meekiong, K., Hidir, M. &amp; Yazid, K. (2015) Three new Scaphochlamys species from Sarawak, Malaysia. Folia Malaysiana 16: 31 - 44." type="journal article" year="2015">Meekiong &amp; Hidir (2015: 34)</bibRefCitation>
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.
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:—MALYASIAN BORNEO.
<collectingCountry id="F36C1CBDEA03D454331EFC8E7BF84A7D" box="[1347,1448,771,796]" name="Malaysia" pageId="45" pageNumber="276">Sarawak</collectingCountry>
: Kuching Division, Samunsam Wildlife Sanctuary,
<date id="FFC57AEDEA03D45434E1FCAA7DC64A21" box="[700,918,807,832]" pageId="45" pageNumber="276" value="2014-11-12">12 November 2014</date>
,
<emphasis id="B90F803FEA03D45435FDFCAA7B3A4A21" box="[928,1386,807,832]" italics="true" pageId="45" pageNumber="276">K. Meekiong, Tasha &amp; M. Hidir MK2364</emphasis>
(holo SAR!, iso HUMS!)
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Terrestrial, perennial, rhizomatous herbs, to c.
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tall. Rhizome creeping, c.
<quantity id="4C83F1C8EA03D4543240FC1E7A304ACD" box="[1053,1120,915,940]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="276" unit="mm" value="2.0">2 mm</quantity>
in diameter, greenish yellow to light brown. Elements indistinct, 3- to 4-foliates, juvenile not seen; petiole
<quantity id="4C83F1C8EA03D4543259FC3A7AC14AB1" box="[1028,1169,951,976]" metricMagnitude="-2" metricUnit="m" metricValue="7.45" metricValueMax="10.3" metricValueMin="4.6" pageId="45" pageNumber="276" unit="cm" value="7.45" valueMax="10.3" valueMin="4.6">4.6–10.3 cm</quantity>
long,
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diameter, slender, canaliculate, glabrous; leaf sheath 3–5.6 ×
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, boat-shaped, reddish light brown, papery when dried, apex acute; lamina 8–15 ×
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, slightly asymmetric, lanceolate, margin slightly wavy, base attenuate, apex acute, adaxial surface light green to green yellow, abaxial surface pale green, sometimes greenish with very light reddish, dull appearance. Inflorescence exerting from leaf base,
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long, flowering in succession, a flower at a time; peduncle
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long, c.
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diameter, cylinder, yellowish green, paler toward base; bracts 3–5, 2.6–3.4 ×
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, stiff, boat-shaped, pale green yellow and very light red at the base, apex acute, each subtending a flower; flower purplish white, white when young; floral tube
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long, tubular, white; dorsal corolla lobe 6–8 ×
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, white, translucent; lateral corolla lobes 2, 4.5–8 ×
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, white with purple band at the base, tiny pubescent along the margin; labellum 10–15×
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, fan-shaped, white with purple streaks along the pale yellow median band, apex bilobed; anther obovate, crest c.
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, apex curved backwards, white with tinged purple at margin; ovary glabrous. Fruit and seeds not seen.
</paragraph>
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<emphasis id="B90F803FEA03D45436E0FAEA7FB94CE1" bold="true" box="[189,489,1383,1408]" pageId="45" pageNumber="276">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA03D45437ADFAEA7C594CE1" box="[496,521,1383,1408]" italics="true" pageId="45" pageNumber="276">—</emphasis>
Known only from Samunsam Widllife Sanctuary and Gunung Pueh, occurring in kerangas forest, sandy soil with thick humus at low elevations.
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<emphasis id="B90F803FEA03D45436E0FA227FFC4CA9" bold="true" box="[189,428,1455,1480]" pageId="45" pageNumber="276">Conservation status</emphasis>
:—Near Threatened (NT). This species is quite common within the protected area but it is only known from the
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locality and its vicinity.
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<emphasis id="B90F803FEA03D45436E0FA7A7F6B4F71" bold="true" box="[189,315,1527,1552]" pageId="45" pageNumber="276">Discussion</emphasis>
:—This species is closely similar to and possibly conspecific with
<taxonomicName id="4C7B27AEEA03D4543279FA7A7AE64F71" authorityName="Burtt &amp; Smith" authorityYear="1972" baseAuthorityName="K. Schum." box="[1060,1206,1527,1552]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="45" pageNumber="276" phylum="Tracheophyta" rank="species" species="polyphylla">
<emphasis id="B90F803FEA03D4543279FA7A7AE64F71" box="[1060,1206,1527,1552]" italics="true" pageId="45" pageNumber="276">S. polyphylla</emphasis>
</taxonomicName>
.
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</treatment>
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<mods:location id="BFDF1C1C488F168516E4A8C7F828A97A">
<mods:url id="D12B05904CE11353C75B772F4FC6F291">http://dx.doi.org/10.11646/phytotaxa.317.4.1</mods:url>
</mods:location>
<mods:classification id="8D751E900E62FF51269AEB9973D3E17F">journal article</mods:classification>
<mods:identifier id="D025500A1BECCDA33EA8FF0595C528D4" type="DOI">10.11646/phytotaxa.317.4.1</mods:identifier>
<mods:identifier id="A9824DBBEB57C29A581F3CAD8E998EE3" type="ISSN">1179-3163</mods:identifier>
</mods:mods>
<treatment id="03D2ED3BEA08D45F36D5FD8A7C494E19" LSID="urn:lsid:plazi:treatment:03D2ED3BEA08D45F36D5FD8A7C494E19" httpUri="http://treatment.plazi.org/id/03D2ED3BEA08D45F36D5FD8A7C494E19" lastPageNumber="269" pageId="38" pageNumber="269">
<subSubSection id="C3610FA6EA08D45F36D5FD8A7BFC4B41" box="[136,1452,517,544]" pageId="38" pageNumber="269" type="nomenclature">
<paragraph id="8BC45C2DEA08D45F36D5FD8A7BFC4B41" blockId="38.[136,1452,517,723]" box="[136,1452,517,544]" pageId="38" pageNumber="269">
<heading id="D08CEB41EA08D45F36D5FD8A7BFC4B41" box="[136,1452,517,544]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="269" reason="4">
<emphasis id="B90F803FEA08D45F36D5FD8A7EFB4B41" bold="true" box="[136,171,519,544]" pageId="38" pageNumber="269">20.</emphasis>
<taxonomicName id="4C7B27AEEA08D45F36EAFD887DFE4B41" ID-CoL="8X5G7" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[183,942,517,544]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="pseudoreticosa" status="sp. nov.">
<emphasis id="B90F803FEA08D45F36EAFD887C724B41" bold="true" box="[183,546,517,544]" italics="true" pageId="38" pageNumber="269">Scaphochlamys pseudoreticosa</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
</taxonomicName>
,
<emphasis id="B90F803FEA08D45F35E0FD8A7A474B41" box="[957,1047,519,544]" italics="true" pageId="38" pageNumber="269">
<taxonomicNameLabel id="A23C3D44EA08D45F35E0FD8A7A474B41" box="[957,1047,519,544]" pageId="38" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B90F803FEA08D45F327FFD8A7A0D4B41" bold="true" box="[1058,1117,519,544]" pageId="38" pageNumber="269">
<typeStatus id="54C0E28FEA08D45F327FFD8A7A0D4B41" box="[1058,1117,519,544]" pageId="38" pageNumber="269">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA08D45F36D5FDA67C604B09" pageId="38" pageNumber="269" type="description">
<paragraph id="8BC45C2DEA08D45F36D5FDA67BFC4B25" blockId="38.[136,1452,517,723]" box="[136,1452,555,580]" pageId="38" pageNumber="269">
<collectingCountry id="F36C1CBDEA08D45F36D5FDA67EBD4B25" box="[136,237,555,580]" name="Malaysia" pageId="38" pageNumber="269">Sarawak</collectingCountry>
: Kuching Division, Bau, Seromah,
<geoCoordinate id="EE4F3AEAEA08D45F34DAFDA67D784B25" box="[647,808,555,580]" degrees="01" direction="north" minutes="22" orientation="latitude" pageId="38" pageNumber="269" precision="1" seconds="36.5" value="1.3768055">01°22’36.5”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA08D45F356BFDA67D8F4B25" box="[822,991,555,580]" degrees="110" direction="east" minutes="11" orientation="longitude" pageId="38" pageNumber="269" precision="1" seconds="32.2" value="110.192276">110°11’32.2”E</geoCoordinate>
,
<quantity id="4C83F1C8EA08D45F35B3FDA67A7B4B25" box="[1006,1067,555,580]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="38" pageNumber="269" unit="m" value="40.0">40 m</quantity>
,
<date id="FFC57AEDEA08D45F3267FDA67AAB4B25" box="[1082,1275,555,580]" pageId="38" pageNumber="269" value="2014-01-12">12 January 2014</date>
,
<emphasis id="B90F803FEA08D45F3354FDA67BFC4B25" box="[1289,1452,555,580]" italics="true" pageId="38" pageNumber="269">I.H. Ooi &amp; K.</emphasis>
</paragraph>
<paragraph id="8BC45C2DEA08D45F36D5FDC27C604B09" blockId="38.[136,1452,517,723]" box="[136,560,591,616]" pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F36D5FDC27F774B09" box="[136,295,591,616]" italics="true" pageId="38" pageNumber="269">Jeland OIH83</emphasis>
(holo SAR),
<figureCitation id="134040A8EA08D45F37E3FDC27C604B09" box="[446,560,591,616]" captionStart="FIGURE 20" captionStartId="39.[136,229,1996,2017]" captionTargetBox="[162,1428,181,1968]" captionTargetId="figure-14@39.[159,1429,175,1971]" captionTargetPageId="39" captionText="FIGURE 20. Scaphochlamys pseudoreticosa. A. plant in habitat. B. inflorescence. C. lamina, adaxial surface. D. lamina, adaxial surface, note slightly raised tertiary veins. E. flower. F.bract and bracteoles. G. calyx and ovary. H. lamina, abaxial surface. I. stamen, note anther spurs. J. stamen, note spurless anthers.A–J from OIH83." pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F37E3FDC27C604B09" bold="true" box="[446,560,591,616]" pageId="38" pageNumber="269">Figure 20</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA08D45F36D5FDFB7BD14F71" pageId="38" pageNumber="269" type="diagnosis">
<paragraph id="8BC45C2DEA08D45F36D5FDFB7AC64BB2" blockId="38.[136,1452,517,723]" pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F36D5FDFB7EBB4BEA" bold="true" box="[136,235,630,651]" pageId="38" pageNumber="269">Diagnosis</emphasis>
:—
<taxonomicName id="4C7B27AEEA08D45F3754FDFB7C664BEA" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[265,566,630,651]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="pseudoreticosa">
<emphasis id="B90F803FEA08D45F3754FDFB7C664BEA" box="[265,566,630,651]" italics="true" pageId="38" pageNumber="269">Scaphochlamys pseudoreticosa</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C7B27AEEA08D45F34E6FDFB7D704BEA" authorityName="Smith" authorityYear="1987" baseAuthorityName="Ridley" baseAuthorityYear="1905" box="[699,800,630,651]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="reticosa">
<emphasis id="B90F803FEA08D45F34E6FDFB7D704BEA" box="[699,800,630,651]" italics="true" pageId="38" pageNumber="269">S. reticosa</emphasis>
</taxonomicName>
but differs by having the lamina abaxially sparsely pubescent (vs. glabrous), tertiary veins faintly visible but never raised (vs. prominently raised), sparsely pubescent light brownish green bracts (vs. glabrous and light brown), and light green moderately pubescent ovary (vs. white, densely pubescent).
</paragraph>
<paragraph id="8BC45C2DEA08D45F36D5FC8E7BD14F71" blockId="38.[136,1452,771,1912]" pageId="38" pageNumber="269">
Terrestrial, perennial, rhizomatous herbs, to c.
<quantity id="4C83F1C8EA08D45F34D6FC8E7C9F4A7D" box="[651,719,771,796]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="269" unit="cm" value="15.0">15 cm</quantity>
tall. Rhizome creeping on ground, c.
<quantity id="4C83F1C8EA08D45F3230FC8E7AFD4A7D" box="[1133,1197,771,796]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="269" unit="mm" value="5.0">5 mm</quantity>
in diameter, externally light to medium brown, internally yellowish white. Elements c.
<quantity id="4C83F1C8EA08D45F353EFCAA7D9F4A21" box="[867,975,807,832]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="2.5" metricValueMin="0.1" pageId="38" pageNumber="269" unit="mm" value="13.0" valueMax="25.0" valueMin="1.0">1–25 mm</quantity>
apart, 1-foliate, juvenile not seen; petiole
<quantity id="4C83F1C8EA08D45F36D5FCC67E8D4A05" box="[136,221,843,868]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="38" pageNumber="269" unit="cm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 cm</quantity>
long, canaliculate, light to yellowish green, glabrous, base pulvinate, white to light green; leafless sheath c. 3,
<quantity id="4C83F1C8EA08D45F36F8FCE27F734AE9" box="[165,291,879,904]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="3.2" metricValueMin="0.7" pageId="38" pageNumber="269" unit="cm" value="1.95" valueMax="3.2" valueMin="0.7">0.7–3.2 cm</quantity>
long, light green, glabrous, drying with age; leaf sheath c.
<quantity id="4C83F1C8EA08D45F359FFCE27A5F4AE9" box="[962,1039,879,904]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="269" unit="cm" value="1.5">1.5 cm</quantity>
long, membranous, white, glabrous, drying and shredding with age; ligule obscure; lamina 6–9 ×
<quantity id="4C83F1C8EA08D45F3579FC1E7DDE4ACD" box="[804,910,915,940]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="38" pageNumber="269" unit="cm" value="5.25" valueMax="6.0" valueMin="4.5">4.5–6 cm</quantity>
, ovate, semi-coriaceous, margin entire to slightly undulate, base cordate, apex acute; adaxial surface semi-glossy, light green to medium green, glabrous, midrib sunken, medium green, glabrous, main lateral veins slightly raised, tertiary veins slightly raised; abaxial surface light green, sparsely pubescent, midrib raised, medium green, pubescent, main lateral veins inconspicuous. Inflorescence c.
<quantity id="4C83F1C8EA08D45F33DFFC727EFA4D5D" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="38" pageNumber="269" unit="cm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 cm</quantity>
long, emerging from near leaf base inside leaf sheaths, tightly congested, comprised of c. 4, 2-flowered cincinni borne on a peduncle; peduncle c.
<quantity id="4C83F1C8EA08D45F3456FBCA7C304D01" box="[523,608,1095,1120]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="38" pageNumber="269" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
long, glabrous, greenish white, covered with marcescent leafless sheaths; bract c. 4, c. 18 ×
<quantity id="4C83F1C8EA08D45F3707FBE67FCE4DE5" box="[346,414,1131,1156]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="269" unit="mm" value="5.0">5 mm</quantity>
, spirally arranged, lanceolate, membranous, brownish light green, sparsely pubescent, apex acuminate, each bract subtending 2 flowers; bracteole 2, c.
<quantity id="4C83F1C8EA08D45F3542FB027D3E4DC9" box="[799,878,1167,1192]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="269" unit="mm" value="15.0">15 mm</quantity>
long, shorter than bracts but barely distinguishable from bract, brownish light green; flower
<quantity id="4C83F1C8EA08D45F3406FB3E7C954DAD" box="[603,709,1203,1228]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="38" pageNumber="269" unit="cm" value="3.8" valueMax="4.0" valueMin="3.6">3.6–4 cm</quantity>
long, floral odour reminiscent of torch-ginger (
<emphasis id="B90F803FEA08D45F32B8FB3E7BFB4DAD" box="[1253,1451,1203,1228]" italics="true" pageId="38" pageNumber="269">
<taxonomicName id="4C7B27AEEA08D45F32B8FB3E7BF34DAD" box="[1253,1443,1203,1228]" class="Liliopsida" family="Zingiberaceae" genus="Etlingera" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="elatior">Etlingera elatior</taxonomicName>
)
</emphasis>
tissues; calyx
<quantity id="4C83F1C8EA08D45F3778FB5A7FCE4D91" box="[293,414,1239,1264]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="38" pageNumber="269" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
long, sparsely pubescent, white, apex acute; floral tube
<quantity id="4C83F1C8EA08D45F324FFB5A7ADB4D91" box="[1042,1163,1239,1264]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="38" pageNumber="269" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">26–30 mm</quantity>
long, sparsely pubescent, white; corolla lobes
<quantity id="4C83F1C8EA08D45F3729FB767F834C75" box="[372,467,1275,1300]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="38" pageNumber="269" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long, lanceolate, yellowish white, glabrous, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA08D45F3311FB767BFC4C75" box="[1356,1452,1275,1300]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="38" pageNumber="269" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, light yellow, apex acute to rounded; labellum 8–10 ×
<quantity id="4C83F1C8EA08D45F3717FACE7FFA4C3D" box="[330,426,1347,1372]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="38" pageNumber="269" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white at base and violet till apex with a light yellow median band, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA08D45F3541FAEA7D0F4CE1" box="[796,863,1383,1408]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="269" unit="mm" value="3.0">3 mm</quantity>
, lobes sometimes slightly overlapping; stamen c. 4.5 ×
<quantity id="4C83F1C8EA08D45F36C2FA067EB24CC5" box="[159,226,1419,1444]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="269" unit="mm" value="2.0">2 mm</quantity>
, covered with glandular hairs, white; filament c.
<quantity id="4C83F1C8EA08D45F3554FA067D0F4CC5" box="[777,863,1419,1444]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="269" unit="mm" value="0.5">0.5 mm</quantity>
long; anther thecae c.
<quantity id="4C83F1C8EA08D45F3203FA067AE44CC5" box="[1118,1204,1419,1444]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="38" pageNumber="269" unit="mm" value="3.5">3.5 mm</quantity>
long, sometimes with very short basal spur, crest c.
<quantity id="4C83F1C8EA08D45F3791FA227C704CA9" box="[460,544,1455,1480]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="269" unit="mm" value="0.5">0.5 mm</quantity>
long; stigma less than
<quantity id="4C83F1C8EA08D45F3546FA227D0A4CA9" box="[795,858,1455,1480]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="269" unit="mm" value="1.0">1 mm</quantity>
long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA08D45F37CFFA5E7FB14C8D" box="[402,481,1491,1516]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="38" pageNumber="269" unit="mm" value="32.0">32 mm</quantity>
long, white, sparsely pubescent; ovary c.
<quantity id="4C83F1C8EA08D45F35EBFA5E7DA64C8D" box="[950,1014,1491,1516]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="269" unit="mm" value="2.0">2 mm</quantity>
long, 1-locular, light green, pubescent, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA08D45F343CFA7A7CF24F71" box="[609,674,1527,1552]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="269" unit="mm" value="1.0">1 mm</quantity>
long, free, needle-like, yellowish white. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA08D45F36E0F9967EB84F39" pageId="38" pageNumber="269" type="etymology">
<paragraph id="8BC45C2DEA08D45F36E0F9967EB84F39" blockId="38.[136,1452,771,1912]" pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F36E0F9967F6E4F55" bold="true" box="[189,318,1563,1588]" pageId="38" pageNumber="269">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C7B27AEEA08D45F3400F9967D554F55" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[605,773,1563,1588]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="pseudoreticosa">
<emphasis id="B90F803FEA08D45F3400F9967D554F55" box="[605,773,1563,1588]" italics="true" pageId="38" pageNumber="269">pseudoreticosa</emphasis>
</taxonomicName>
, alludes to the superficial similarity of this species to
<taxonomicName id="4C7B27AEEA08D45F33CBF9967EB14F39" authorityName="Smith" authorityYear="1987" baseAuthorityName="Ridley" baseAuthorityYear="1905" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="reticosa">
<emphasis id="B90F803FEA08D45F33CBF9967EB14F39" italics="true" pageId="38" pageNumber="269">S. reticosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA08D45F36E0F9EE7B2A4E6D" pageId="38" pageNumber="269" type="distribution">
<paragraph id="8BC45C2DEA08D45F36E0F9EE7DBA4FC1" blockId="38.[136,1452,771,1912]" pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F36E0F9EE7FB84F1D" bold="true" box="[189,488,1635,1660]" pageId="38" pageNumber="269">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA08D45F37B2F9EE7D374F1D" box="[495,871,1635,1660]" italics="true" pageId="38" pageNumber="269">
—
<taxonomicName id="4C7B27AEEA08D45F3451F9EE7D374F1D" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[524,871,1635,1660]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="pseudoreticosa">Scaphochlamys pseudoreticosa</taxonomicName>
</emphasis>
is only known from the
<typeStatus id="54C0E28FEA08D45F32DBF9EE7AE64F1D" box="[1158,1206,1635,1660]" pageId="38" pageNumber="269">type</typeStatus>
locality, occurring in mollisolic black soil on shaded forested Karst limestone of c.
<quantity id="4C83F1C8EA08D45F3561F90A7D254FC1" box="[828,885,1671,1696]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="38" pageNumber="269" unit="m" value="40.0">40 m</quantity>
elevation.
</paragraph>
<paragraph id="8BC45C2DEA08D45F36E0F9267B2A4E6D" blockId="38.[136,1452,771,1912]" pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F36E0F9267FE04FA5" bold="true" box="[189,432,1707,1732]" pageId="38" pageNumber="269">Conservation status</emphasis>
:—The conservation status of this species is Vulnerable (VU), meeting criteria B.1.a., B.1.b.(iii), B.2.a., B.2.b.(iii), and D.1. This species occurs at the margin of a protected area next to a village. The population of this limestone obligated species is small and potentially easily affected by nearby human activities.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA08D45F36E0F89A7C494E19" pageId="38" pageNumber="269" type="discussion">
<paragraph id="8BC45C2DEA08D45F36E0F89A7C494E19" blockId="38.[136,1452,771,1912]" pageId="38" pageNumber="269">
<emphasis id="B90F803FEA08D45F36E0F89A7F6B4E51" bold="true" box="[189,315,1815,1840]" pageId="38" pageNumber="269">Discussion</emphasis>
:—The anther spur may be present or absent in
<taxonomicName id="4C7B27AEEA08D45F3505F89A7A4D4E51" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[856,1053,1815,1840]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="pseudoreticosa">
<emphasis id="B90F803FEA08D45F3505F89A7A4D4E51" box="[856,1053,1815,1840]" italics="true" pageId="38" pageNumber="269">S. pseudoreticosa</emphasis>
</taxonomicName>
but is always absent in
<taxonomicName id="4C7B27AEEA08D45F3373F89A7BF44E51" authorityName="Smith" authorityYear="1987" baseAuthorityName="Ridley" baseAuthorityYear="1905" box="[1326,1444,1815,1840]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="reticosa">
<emphasis id="B90F803FEA08D45F3373F89A7BF44E51" box="[1326,1444,1815,1840]" italics="true" pageId="38" pageNumber="269">S. reticosa</emphasis>
</taxonomicName>
. The tertiary venation of the leaf laminae of
<taxonomicName id="4C7B27AEEA08D45F3426F8B67D114E35" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[635,833,1851,1876]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="pseudoreticosa">
<emphasis id="B90F803FEA08D45F3426F8B67D114E35" box="[635,833,1851,1876]" italics="true" pageId="38" pageNumber="269">S. pseudoreticosa</emphasis>
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is a faintly raised tessellate reticulum adaxially, never prominently raised as in
<taxonomicName id="4C7B27AEEA08D45F37C0F8D27C424E19" authorityName="Smith" authorityYear="1987" baseAuthorityName="Ridley" baseAuthorityYear="1905" box="[413,530,1887,1912]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="38" pageNumber="269" phylum="Tracheophyta" rank="species" species="reticosa">
<emphasis id="B90F803FEA08D45F37C0F8D27C424E19" box="[413,530,1887,1912]" italics="true" pageId="38" pageNumber="269">S. reticosa</emphasis>
</taxonomicName>
.
</paragraph>
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</treatment>
</document>

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<mods:title id="E5C26BD45388ECAC6DD00301097E49B5">A review of Scaphochlamys (Zingiberaceae) from Borneo, with description of eleven new species</mods:title>
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<subSubSection id="C3610FA6EA0CD45B36D5FC517BFC4A94" box="[136,1452,986,1013]" pageId="34" pageNumber="265" type="nomenclature">
<paragraph id="8BC45C2DEA0CD45B36D5FC517BFC4A94" blockId="34.[136,1452,986,1155]" box="[136,1452,986,1013]" pageId="34" pageNumber="265">
<heading id="D08CEB41EA0CD45B36D5FC517BFC4A94" box="[136,1452,986,1013]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="265" reason="4">
<emphasis id="B90F803FEA0CD45B36D5FC517EFC4A94" bold="true" box="[136,172,988,1013]" pageId="34" pageNumber="265">18.</emphasis>
<taxonomicName id="4C7B27AEEA0CD45B36EBFC577D6C4A94" ID-CoL="8X5G5" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[182,828,986,1013]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="nigra" status="sp. nov.">
<emphasis id="B90F803FEA0CD45B36EBFC577FE44A95" bold="true" box="[182,436,986,1013]" italics="true" pageId="34" pageNumber="265">Scaphochlamys nigra</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
</taxonomicName>
,
<emphasis id="B90F803FEA0CD45B3511FC517DF54A94" box="[844,933,988,1013]" italics="true" pageId="34" pageNumber="265">
<taxonomicNameLabel id="A23C3D44EA0CD45B3511FC517DF54A94" box="[844,933,988,1013]" pageId="34" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="B90F803FEA0CD45B35F2FC517DBA4A94" bold="true" box="[943,1002,988,1013]" pageId="34" pageNumber="265">
<typeStatus id="54C0E28FEA0CD45B35F2FC517DBA4A94" box="[943,1002,988,1013]" pageId="34" pageNumber="265">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA0CD45B331EFC517BF84A94" box="[1347,1448,988,1013]" name="Malaysia" pageId="34" pageNumber="265">Sarawak</collectingCountry>
:
</heading>
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<subSubSection id="C3610FA6EA0CD45B36D5FB8D7F8F4D5C" pageId="34" pageNumber="265" type="description">
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Kuching Division, Bau, Gunung Buan,
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,
<geoCoordinate id="EE4F3AEAEA0CD45B3554FB8D7DE44D78" box="[777,948,1024,1049]" degrees="110" direction="east" minutes="08" orientation="longitude" pageId="34" pageNumber="265" precision="1" seconds="35.2" value="110.14311">110°08’35.2”E</geoCoordinate>
,
<date id="FFC57AEDEA0CD45B359EFB8D7AF64D78" box="[963,1190,1024,1049]" pageId="34" pageNumber="265" value="2013-11-21">21 November 2013</date>
,
<emphasis id="B90F803FEA0CD45B32EBFB8D7BFB4D78" box="[1206,1451,1024,1049]" italics="true" pageId="34" pageNumber="265">I.H. Ooi &amp; T. Jepom</emphasis>
</paragraph>
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<emphasis id="B90F803FEA0CD45B36D5FBA97E864D5C" box="[136,214,1060,1085]" italics="true" pageId="34" pageNumber="265">OIH76</emphasis>
(holo SAR),
<figureCitation id="134040A8EA0CD45B3730FBA97F8F4D5C" box="[365,479,1060,1085]" captionStart="FIGURE 18" captionStartId="35.[136,229,2014,2035]" captionTargetBox="[156,1435,180,1984]" captionTargetId="figure-14@35.[152,1435,175,1989]" captionTargetPageId="35" captionText="FIGURE 18. Scaphochlamys nigra. A. plant in habitat. B. inflorescence. C. lamina, adaxial surface, note very dark colour. D. lamina, abaxial surface. E. flower. F.fruit. G. bract and bracteole. H. stamen, note anther spurs.A–H from OIH76." pageId="34" pageNumber="265">
<emphasis id="B90F803FEA0CD45B3730FBA97F8F4D5C" bold="true" box="[365,479,1060,1085]" pageId="34" pageNumber="265">Figure 18</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA0CD45B36D5FBC77FC2414C" pageId="34" pageNumber="265" type="diagnosis">
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<emphasis id="B90F803FEA0CD45B36D5FBC77EBB4D3E" bold="true" box="[136,235,1098,1119]" pageId="34" pageNumber="265">Diagnosis</emphasis>
:—
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<emphasis id="B90F803FEA0CD45B3754FBC77F8A4D3E" box="[265,474,1098,1119]" italics="true" pageId="34" pageNumber="265">Scaphochlamys nigra</emphasis>
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is readily distinguishable from the other Bornean species by the shortly broad ovate and coriaceous leaf laminae with very deep dark green adaxial surface (almost black), and a solitary flower per bract.
</paragraph>
<paragraph id="8BC45C2DEA0CD45B36D5FB397FC2414C" blockId="34.[136,1452,1204,2093]" pageId="34" pageNumber="265">
Terrestrial, perennial, rhizomatous herbs, to c.
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tall. Rhizome slightly below ground, c.
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in diameter, externally light brown to greenish light brown, internally brownish white. Elements c.
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apart, 1-foliate, juvenile not seen; petiole
<quantity id="4C83F1C8EA0CD45B3712FB717FF54C74" box="[335,421,1276,1301]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="34" pageNumber="265" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
long, canaliculate, glabrous, medium green, base pulvinate, white to light green; leafless sheath c. 3,
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long, brownish light green, sparsely pubescent, drying with age; leaf sheath c.
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long, membranous, white, glabrous, drying and shredding with age; ligule obscure; lamina 4–6 ×
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, short broad ovate, coriaceous, margin entire, base rounded, apex obtuse; adaxial surface semi-glossy, dark green to almost black, glabrous, midrib sunken, dark green, glabrous, main lateral veins inconspicuous; abaxial surface sparsely pubescent near midrib, glaucous green, midrib raised, dark green, glabrous, main lateral veins inconspicuous. Inflorescence c.
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long, emerging from near leaf base inside leaf sheaths, tightly congested, comprised of c. 3 solitary flowers borne on a peduncle, flowering spirally from base to apex; peduncle c.
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long, whitish light green, glabrous, covered with degraded leafless sheaths; bract c. 3, c. 15 ×
<quantity id="4C83F1C8EA0CD45B354AF9917D0A4F54" box="[791,858,1564,1589]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="265" unit="mm" value="3.0">3 mm</quantity>
, spirally arranged, lanceolate, membranous, sparsely pubescent, light brown, apex acuminate, each bract subtending a flower; bracteole 1, c.
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long, shorter than bracts but barely distinguishable from bract, light brown; flower
<quantity id="4C83F1C8EA0CD45B3556F9E97DD74F1C" box="[779,903,1636,1661]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="34" pageNumber="265" unit="cm" value="3.15" valueMax="3.4" valueMin="2.9">2.9–3.4 cm</quantity>
long, without noticeable floral odour; calyx
<quantity id="4C83F1C8EA0CD45B3329F9E97EE44FC0" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="34" pageNumber="265" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long, white, sparsely pubescent, apex acute; floral tube
<quantity id="4C83F1C8EA0CD45B3571F9057DF54FC0" box="[812,933,1672,1697]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="34" pageNumber="265" unit="mm" value="20.5" valueMax="23.0" valueMin="18.0">18–23 mm</quantity>
long, white, sparsely pubescent; corolla lobes
<quantity id="4C83F1C8EA0CD45B36D5F9217EA34FA4" box="[136,243,1708,1733]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="34" pageNumber="265" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long, lanceolate, glabrous, yellowish white, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA0CD45B320CF9217AFE4FA4" box="[1105,1198,1708,1733]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="34" pageNumber="265" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, light yellow, apex acute to rounded; labellum 9–11 ×
<quantity id="4C83F1C8EA0CD45B3329F95D7EE64E6C" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="34" pageNumber="265" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white at base and violet till apex with a light yellow median band, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA0CD45B3419F8957CD74E50" box="[580,647,1816,1841]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="265" unit="mm" value="4.0">4 mm</quantity>
, lobes sometimes slightly overlapping; stamen c. 4.5 ×
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, covered with glandular hairs, white; filament c.
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long; anther thecae c.
<quantity id="4C83F1C8EA0CD45B35FEF8B17DAA4E34" box="[931,1018,1852,1877]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="34" pageNumber="265" unit="mm" value="3.5">3.5 mm</quantity>
long, with very short basal spur, crest c.
<quantity id="4C83F1C8EA0CD45B36FEF8ED7EAA4E18" box="[163,250,1888,1913]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="265" unit="mm" value="0.5">0.5 mm</quantity>
long; stigma less than
<quantity id="4C83F1C8EA0CD45B37A0F8ED7C6F4E18" box="[509,575,1888,1913]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="265" unit="mm" value="1.0">1 mm</quantity>
long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA0CD45B36D5F8097E884EFC" box="[136,216,1924,1949]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="265" unit="mm" value="25.0">25 mm</quantity>
long, white, sparsely pubescent; ovary c.
<quantity id="4C83F1C8EA0CD45B34EEF8097CA54EFC" box="[691,757,1924,1949]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="265" unit="mm" value="2.0">2 mm</quantity>
long, 1-locular, densely pubescent, white, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA0CD45B37D2F8257F824EA0" box="[399,466,1960,1985]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="265" unit="mm" value="1.0">1 mm</quantity>
long, free, needle-like, white. Fruit c. 12 ×
<quantity id="4C83F1C8EA0CD45B3595F8257A5C4EA0" box="[968,1036,1960,1985]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="265" unit="mm" value="4.0">4 mm</quantity>
, dehiscent capsule, elongated ovoid, whitish green (greenish white translucent cuticle-like pericarp enclosing green seed), sparsely pubescent, subtended by marcescent bracts. Seeds c. 10 ×
<quantity id="4C83F1C8EA0CD45B3440F87D7C304168" box="[541,608,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="265" unit="mm" value="3.0">3 mm</quantity>
, slightly elongated ovoid, glabrous, green; aril c.
<quantity id="4C83F1C8EA0CD45B32D6F87D7A9E4168" box="[1163,1230,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="265" unit="mm" value="4.0">4 mm</quantity>
, lacerate, yellowish white; perisperm white.
</paragraph>
</subSubSection>
<caption id="DF040CA5EA0DD45A36D5F8537AD84176" pageId="35" pageNumber="266" startId="35.[136,229,2014,2035]" targetBox="[156,1435,180,1984]" targetPageId="35" targetType="figure">
<paragraph id="8BC45C2DEA0DD45A36D5F8537AD84176" blockId="35.[136,1452,2014,2071]" pageId="35" pageNumber="266">
<emphasis id="B90F803FEA0DD45A36D5F8537F5B4E92" bold="true" box="[136,267,2014,2035]" pageId="35" pageNumber="266">FIGURE 18.</emphasis>
<emphasis id="B90F803FEA0DD45A374FF8537FBA4E92" box="[274,490,2014,2035]" italics="true" pageId="35" pageNumber="266">
<taxonomicName id="4C7B27AEEA0DD45A374FF8537FB64E92" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[274,486,2014,2035]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="35" pageNumber="266" phylum="Tracheophyta" rank="species" species="nigra">Scaphochlamys nigra</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA0DD45A37ADF8537C584E92" bold="true" box="[496,520,2014,2035]" pageId="35" pageNumber="266">A.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA0DD45A34F2F8537C964E92" bold="true" box="[687,710,2014,2035]" pageId="35" pageNumber="266">B.</emphasis>
inflorescence.
<emphasis id="B90F803FEA0DD45A3507F8537D224E92" bold="true" box="[858,882,2014,2035]" pageId="35" pageNumber="266">C.</emphasis>
lamina, adaxial surface, note very dark colour.
<emphasis id="B90F803FEA0DD45A3318F8537B0D4E92" bold="true" box="[1349,1373,2014,2035]" pageId="35" pageNumber="266">D.</emphasis>
lamina, abaxial surface.
<emphasis id="B90F803FEA0DD45A3778F78F7F6B4176" bold="true" box="[293,315,2050,2071]" pageId="35" pageNumber="266">E.</emphasis>
flower.
<emphasis id="B90F803FEA0DD45A37D7F78F7FCF4176" bold="true" box="[394,415,2050,2071]" pageId="35" pageNumber="266">F.</emphasis>
fruit.
<emphasis id="B90F803FEA0DD45A3785F78F7FA14176" bold="true" box="[472,497,2050,2071]" pageId="35" pageNumber="266">G.</emphasis>
bract and bracteole.
<emphasis id="B90F803FEA0DD45A34E4F78F7C824176" bold="true" box="[697,722,2050,2071]" pageId="35" pageNumber="266">H.</emphasis>
stamen, note anther spurs. A–H from
<emphasis id="B90F803FEA0DD45A321DF78F7AD34176" box="[1088,1155,2050,2071]" italics="true" pageId="35" pageNumber="266">OIH76</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C3610FA6EA0AD45D36E0FF127B1349D9" box="[189,1347,159,184]" pageId="36" pageNumber="267" type="etymology">
<paragraph id="8BC45C2DEA0AD45D36E0FF127B1349D9" blockId="36.[136,1452,159,760]" box="[189,1347,159,184]" pageId="36" pageNumber="267">
<emphasis id="B90F803FEA0AD45D36E0FF127F6E49D9" bold="true" box="[189,318,159,184]" pageId="36" pageNumber="267">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C7B27AEEA0AD45D3410FF127CDA49D9" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[589,650,159,184]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="36" pageNumber="267" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="B90F803FEA0AD45D3410FF127CDA49D9" box="[589,650,159,184]" italics="true" pageId="36" pageNumber="267">nigra</emphasis>
</taxonomicName>
, refers to the deep dark green leaf lamina that is almost black.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA0AD45D36E0FF4E7FE34B25" pageId="36" pageNumber="267" type="distribution">
<paragraph id="8BC45C2DEA0AD45D36E0FF4E7ABB4861" blockId="36.[136,1452,159,760]" pageId="36" pageNumber="267">
<emphasis id="B90F803FEA0AD45D36E0FF4E7FB749BD" bold="true" box="[189,487,195,220]" pageId="36" pageNumber="267">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA0AD45D37B3FF4E7CAA49BD" box="[494,762,195,220]" italics="true" pageId="36" pageNumber="267">
—
<taxonomicName id="4C7B27AEEA0AD45D3456FF4E7CAA49BD" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[523,762,195,220]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="36" pageNumber="267" phylum="Tracheophyta" rank="species" species="nigra">Scaphochlamys nigra</taxonomicName>
</emphasis>
is known from the
<typeStatus id="54C0E28FEA0AD45D3586FF4E7A5B49BD" box="[987,1035,195,220]" pageId="36" pageNumber="267">type</typeStatus>
locality and its vicinity. The species occurs on podzolic black soil over sandstone in kerangas (tropical heath) forest,
<quantity id="4C83F1C8EA0AD45D3251FF6A7A254861" box="[1036,1141,231,256]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="36" pageNumber="267" unit="m" value="100.0" valueMax="130.0" valueMin="70.0">70–130m</quantity>
elevation.
</paragraph>
<paragraph id="8BC45C2DEA0AD45D36E0FE867E8848D5" blockId="36.[136,1452,159,760]" pageId="36" pageNumber="267">
<emphasis id="B90F803FEA0AD45D36E0FE867C944845" bold="true" box="[189,708,267,292]" pageId="36" pageNumber="267">
Additional specimen examined (
<typeStatus id="54C0E28FEA0AD45D3410FE867CEC4845" box="[589,700,267,292]" pageId="36" pageNumber="267" type="paratype">paratype</typeStatus>
)
</emphasis>
:—
<materialsCitation id="3B135670EA0AD45D34BEFE867BF74829" collectingDate="2014-02-11" collectorName="I. H. Ooi &amp; J. D. Mood &amp; T. Jepom" country="Malaysia" county="Gunung Buan" latitude="1.5580277" location="Kuching Division" longLatPrecision="1" longitude="110.14311" municipality="Bau" pageId="36" pageNumber="267" specimenCount="1">
MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA0AD45D327DFE867AD54845" box="[1056,1157,267,292]" name="Malaysia" pageId="36" pageNumber="267">Sarawak</collectingCountry>
:
<location id="8EA40AF6EA0AD45D32C8FE867B364845" LSID="urn:lsid:plazi:treatment:03D2ED3BEA0CD45D36D5FC517CCE4B99:8EA40AF6EA0AD45D32C8FE867B364845" box="[1173,1382,267,292]" country="Malaysia" county="Gunung Buan" latitude="1.5580277" longLatPrecision="1" longitude="110.14311" municipality="Bau" name="Kuching Division" pageId="36" pageNumber="267">Kuching Division</location>
,
<collectingMunicipality id="6BA0C657EA0AD45D332AFE867BF74845" box="[1399,1447,267,292]" pageId="36" pageNumber="267">Bau</collectingMunicipality>
,
<collectingCounty id="62A524A1EA0AD45D36D5FEA27F7A4829" box="[136,298,303,328]" pageId="36" pageNumber="267">Gunung Buan</collectingCounty>
,
<geoCoordinate id="EE4F3AEAEA0AD45D3765FEA27F894829" box="[312,473,303,328]" degrees="01" direction="north" minutes="33" orientation="latitude" pageId="36" pageNumber="267" precision="1" seconds="28.9" value="1.5580277">01°33’28.9”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA0AD45D37BAFEA27CC24829" box="[487,658,303,328]" degrees="110" direction="east" minutes="08" orientation="longitude" pageId="36" pageNumber="267" precision="1" seconds="35.2" value="110.14311">110°08’35.2”E</geoCoordinate>
,
<date id="FFC57AEDEA0AD45D34FDFEA27D3F4829" box="[672,879,303,328]" pageId="36" pageNumber="267" value="2014-02-11">
<collectingDate id="EF818305EA0AD45D34FDFEA27D3F4829" box="[672,879,303,328]" pageId="36" pageNumber="267" value="2014-02-11">11 February 2014</collectingDate>
</date>
,
<emphasis id="B90F803FEA0AD45D3520FEA27B004829" box="[893,1360,303,328]" italics="true" pageId="36" pageNumber="267">
<collectorName id="268E39FBEA0AD45D3520FEA27D8F4829" box="[893,991,303,328]" pageId="36" pageNumber="267">I.H. Ooi</collectorName>
,
<collectorName id="268E39FBEA0AD45D35B3FEA27A374829" box="[1006,1127,303,328]" pageId="36" pageNumber="267">J.D. Mood</collectorName>
&amp;
<collectorName id="268E39FBEA0AD45D32CCFEA27AA94829" box="[1169,1273,303,328]" pageId="36" pageNumber="267">T. Jepom</collectorName>
OIH90
</emphasis>
(SAR)
</materialsCitation>
;
<materialsCitation id="3B135670EA0AD45D36D5FEDE7C7548F1" collectingDate="2012-10-25" collectorName="P. C. Boyce &amp; S. Y. Wong &amp; T. Jepom" country="Malaysia" county="Mutud Waterfall" latitude="1.5531944" location="Kampung Segong" longLatPrecision="1" longitude="110.14389" municipality="Sungai Mutud" pageId="36" pageNumber="267" specimenCount="1">
<location id="8EA40AF6EA0AD45D36D5FEDE7F08480D" LSID="urn:lsid:plazi:treatment:03D2ED3BEA0CD45D36D5FC517CCE4B99:8EA40AF6EA0AD45D36D5FEDE7F08480D" box="[136,344,339,364]" country="Malaysia" county="Mutud Waterfall" latitude="1.5531944" longLatPrecision="1" longitude="110.14389" municipality="Sungai Mutud" name="Kampung Segong" pageId="36" pageNumber="267">Kampung Segong</location>
,
<collectingMunicipality id="6BA0C657EA0AD45D373AFEDE7C5D480D" box="[359,525,339,364]" pageId="36" pageNumber="267">Sungai Mutud</collectingMunicipality>
,
<collectingCounty id="62A524A1EA0AD45D3441FEDE7C8B480D" box="[540,731,339,364]" pageId="36" pageNumber="267">Mutud Waterfall</collectingCounty>
,
<geoCoordinate id="EE4F3AEAEA0AD45D34B7FEDE7DD9480D" box="[746,905,339,364]" degrees="01" direction="north" minutes="33" orientation="latitude" pageId="36" pageNumber="267" precision="1" seconds="11.5" value="1.5531944">01°33’11.5”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA0AD45D35C5FEDE7A13480D" box="[920,1091,339,364]" degrees="110" direction="east" minutes="08" orientation="longitude" pageId="36" pageNumber="267" precision="1" seconds="38.0" value="110.14389">110°08’38.0”E</geoCoordinate>
,
<date id="FFC57AEDEA0AD45D320FFEDE7B49480D" box="[1106,1305,339,364]" pageId="36" pageNumber="267" value="2012-10-25">
<collectingDate id="EF818305EA0AD45D320FFEDE7B49480D" box="[1106,1305,339,364]" pageId="36" pageNumber="267" value="2012-10-25">25 October 2012</collectingDate>
</date>
,
<emphasis id="B90F803FEA0AD45D337AFEDE7F8248F1" italics="true" pageId="36" pageNumber="267">
<collectorName id="268E39FBEA0AD45D337AFEDE7BF7480D" box="[1319,1447,339,364]" pageId="36" pageNumber="267">P.C. Boyce</collectorName>
,
<collectorName id="268E39FBEA0AD45D36D5FEFA7EA748F1" box="[136,247,375,400]" pageId="36" pageNumber="267">S.Y. Wong</collectorName>
&amp;
<collectorName id="268E39FBEA0AD45D3747FEFA7F2E48F1" box="[282,382,375,400]" pageId="36" pageNumber="267">T. Jepom</collectorName>
ZI-225
</emphasis>
(SAR)
</materialsCitation>
;
<materialsCitation id="3B135670EA0AD45D346DFEFA7E8148D5" collectingDate="2004-07-21" collectorName="Jeland K. &amp; T. Jepom" country="Malaysia" location="Kampung Segong" municipality="Ulu Sungai Adis" pageId="36" pageNumber="267" specimenCount="1">
<location id="8EA40AF6EA0AD45D346DFEFA7CAC48F1" LSID="urn:lsid:plazi:treatment:03D2ED3BEA0CD45D36D5FC517CCE4B99:8EA40AF6EA0AD45D346DFEFA7CAC48F1" box="[560,764,375,400]" country="Malaysia" municipality="Ulu Sungai Adis" name="Kampung Segong" pageId="36" pageNumber="267">Kampung Segong</location>
,
<collectingMunicipality id="6BA0C657EA0AD45D355AFEFA7D9348F1" box="[775,963,375,400]" pageId="36" pageNumber="267">Ulu Sungai Adis</collectingMunicipality>
,
<date id="FFC57AEDEA0AD45D3593FEFA7A0E48F1" box="[974,1118,375,400]" pageId="36" pageNumber="267" value="2004-07-21">
<collectingDate id="EF818305EA0AD45D3593FEFA7A0E48F1" box="[974,1118,375,400]" pageId="36" pageNumber="267" value="2004-07-21">21 July 2004</collectingDate>
</date>
<emphasis id="B90F803FEA0AD45D3238FEFA7BFB48F1" box="[1125,1451,375,400]" italics="true" pageId="36" pageNumber="267">
<collectorName id="268E39FBEA0AD45D3238FEFA7A8148F1" box="[1125,1233,375,400]" pageId="36" pageNumber="267">Jeland K.</collectorName>
&amp;
<collectorName id="268E39FBEA0AD45D32A9FEFA7B0848F1" box="[1268,1368,375,400]" pageId="36" pageNumber="267">T. Jepom</collectorName>
ZI-567
</emphasis>
(SAR)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC45C2DEA0AD45D36E0FE327FE34B25" blockId="36.[136,1452,159,760]" pageId="36" pageNumber="267">
<emphasis id="B90F803FEA0AD45D36E0FE327FFB48B9" bold="true" box="[189,427,447,472]" pageId="36" pageNumber="267">Conservation status</emphasis>
:—The conservation status of this species is Critically Endangered (CR), and meets criteria B.1.a., B.1.b.(iii), B.2.a., B.2.b.(iii), C.2.a.(i), and D. This species is only known from the
<typeStatus id="54C0E28FEA0AD45D32D6FE6E7AEB489D" box="[1163,1211,483,508]" pageId="36" pageNumber="267">type</typeStatus>
locality with a small population (Native Customary Rights land, non-protected area). Intensive logging activities on the other side of the mountain are encroaching.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA0AD45D36E0FDC27CCE4B99" pageId="36" pageNumber="267" type="discussion">
<paragraph id="8BC45C2DEA0AD45D36E0FDC27CCE4B99" blockId="36.[136,1452,159,760]" pageId="36" pageNumber="267">
<emphasis id="B90F803FEA0AD45D36E0FDC27F6B4B09" bold="true" box="[189,315,591,616]" pageId="36" pageNumber="267">Discussion</emphasis>
:—This is the only
<taxonomicName id="4C7B27AEEA0AD45D3448FDC27D4C4B09" box="[533,796,591,616]" pageId="36" pageNumber="267">
<emphasis id="B90F803FEA0AD45D3448FDC27C934B09" box="[533,707,591,616]" italics="true" pageId="36" pageNumber="267">Scaphochlamys</emphasis>
species
</taxonomicName>
on Borneo with a single flower per bract. Another species reported to have solitary flower is
<taxonomicName id="4C7B27AEEA0AD45D37A2FDFE7C314BED" authorityName="Holttum" authorityYear="1950" baseAuthorityName="Ridley" baseAuthorityYear="1893" box="[511,609,627,652]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="36" pageNumber="267" phylum="Tracheophyta" rank="species" species="biloba">
<emphasis id="B90F803FEA0AD45D37A2FDFE7C314BED" box="[511,609,627,652]" italics="true" pageId="36" pageNumber="267">S. biloba</emphasis>
</taxonomicName>
from South
<collectingCountry id="F36C1CBDEA0AD45D34B5FDFE7D1C4BED" box="[744,844,627,652]" name="Thailand" pageId="36" pageNumber="267">Thailand</collectingCountry>
and North Peninsular
<collectingCountry id="F36C1CBDEA0AD45D321CFDFE7AFB4BED" box="[1089,1195,627,652]" name="Malaysia" pageId="36" pageNumber="267">Malaysia</collectingCountry>
, which is differentiated from
<taxonomicName id="4C7B27AEEA0AD45D3695FD1A7F734BD1" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[200,291,663,688]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="36" pageNumber="267" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="B90F803FEA0AD45D3695FD1A7F734BD1" box="[200,291,663,688]" italics="true" pageId="36" pageNumber="267">S. nigra</emphasis>
</taxonomicName>
by having reddish-brown coriaceous bract, larger flower, white labellum sometimes with red streaks along yellow median band near base, lanceolate to narrowly elliptic leaf lamina, usually with white variegated bands, and sometimes flushed purple lamina abaxially.
</paragraph>
</subSubSection>
</treatment>
</document>

249
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@ -0,0 +1,249 @@
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<mods:title id="C0E19E7728F80DF04A4FC3CB6BA7EF15">A review of Scaphochlamys (Zingiberaceae) from Borneo, with description of eleven new species</mods:title>
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<mods:namePart id="5006B162CE19F4C50A609361692B36FF">Hin, Ooi Im</mods:namePart>
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<mods:role id="73CB34261C838735DAAFBA1C505352EA">
<mods:roleTerm id="B842CFC364F94DF53F64627AF76E8F01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE609C5C85EDD2EB1431965BEAA8F4D7">Yeng, Wong Sin</mods:namePart>
<mods:affiliation id="6AAFE9F9B08F6B9AA39A21CCAAC7ACB8">Research Associate, Harvard University Herbaria, 22 Divinity Avenue, Cambridge, MA 02138, United States.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="25649534C9711E839F82B5F606947751">text</mods:typeOfResource>
<mods:relatedItem id="0AABB6A2948E424752258467190B7231" type="host">
<mods:titleInfo id="968AB7009BD5D3901A288B9714AA27B6">
<mods:title id="09E78470DE7746AB456B7CAA3D4FD3C2">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="347716700FA18CFC706F67AA6E53AE1F">
<mods:date id="4591C754AB5921B3AA8708ACCED99A9A">2017</mods:date>
<mods:detail id="84254B6A70A53472C120A5A9E282D609" type="pubDate">
<mods:number id="D46A371CE04B73CB3ED3E9F2F8F416EA">2017-08-23</mods:number>
</mods:detail>
<mods:detail id="7FEA16AF37B08CBBDD1D11861AED1F3C" type="volume">
<mods:number id="CB07136AFFB52F9961D603C3F7CDECC2">317</mods:number>
</mods:detail>
<mods:detail id="812A035AAF9C4E1023DFB22202CE8BDF" type="issue">
<mods:number id="C72C89F314505125BC912D08E35710A0">4</mods:number>
</mods:detail>
<mods:extent id="31A8A1322493395DBDDD65D9982260C2" unit="page">
<mods:start id="53527C20AAF5F67B9F3A82C9312648B3">231</mods:start>
<mods:end id="B57B34C9B69BC77D75CBFE2BD06ECCE0">279</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="735220FD142A1EBC764F8834B65EC7D8">
<mods:url id="7EA9FA7E286A5965B1CDC76B7A0AAD0D">http://dx.doi.org/10.11646/phytotaxa.317.4.1</mods:url>
</mods:location>
<mods:classification id="EB2C4075667394A6F817A140329CB740">journal article</mods:classification>
<mods:identifier id="7B64594043760E484D83A98942F0E0DD" type="DOI">10.11646/phytotaxa.317.4.1</mods:identifier>
<mods:identifier id="10F804F4FA6DC991EB83B4DCACE5C3D5" type="ISSN">1179-3163</mods:identifier>
</mods:mods>
<treatment id="03D2ED3BEA22D47636D5F9797DD64DE4" LSID="urn:lsid:plazi:treatment:03D2ED3BEA22D47636D5F9797DD64DE4" httpUri="http://treatment.plazi.org/id/03D2ED3BEA22D47636D5F9797DD64DE4" lastPageId="15" lastPageNumber="246" pageId="12" pageNumber="243">
<subSubSection id="C3610FA6EA22D47536D5F9797BFC4E6C" box="[136,1452,1778,1805]" pageId="12" pageNumber="243" type="nomenclature">
<paragraph id="8BC45C2DEA22D47536D5F9797BFC4E6C" blockId="12.[136,1452,1778,1947]" box="[136,1452,1778,1805]" pageId="12" pageNumber="243">
<heading id="D08CEB41EA22D47536D5F9797BFC4E6C" box="[136,1452,1778,1805]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="243" reason="4">
<emphasis id="B90F803FEA22D47536D5F9797ECD4E6C" bold="true" box="[136,157,1780,1805]" pageId="12" pageNumber="243">7.</emphasis>
<taxonomicName id="4C7B27AEEA22D47536F8F97F7D1D4E6C" ID-CoL="8X5G8" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[165,845,1778,1805]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="scintillans" status="sp. nov.">
<emphasis id="B90F803FEA22D47536F8F97F7F874E6C" bold="true" box="[165,471,1778,1805]" italics="true" pageId="12" pageNumber="243">Scaphochlamys scintillans</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
</taxonomicName>
,
<emphasis id="B90F803FEA22D4753504F9797DFE4E6C" box="[857,942,1780,1805]" italics="true" pageId="12" pageNumber="243">
<taxonomicNameLabel id="A23C3D44EA22D4753504F9797DFE4E6C" box="[857,942,1780,1805]" pageId="12" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B90F803FEA22D47535EBF9797DA14E6C" bold="true" box="[950,1009,1780,1805]" pageId="12" pageNumber="243">
<typeStatus id="54C0E28FEA22D47535EBF9797DA14E6C" box="[950,1009,1780,1805]" pageId="12" pageNumber="243">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA22D475331EF9797BF84E6C" box="[1347,1448,1780,1805]" name="Malaysia" pageId="12" pageNumber="243">Sarawak</collectingCountry>
:
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA22D47536D5F8957FB84E34" pageId="12" pageNumber="243" type="description">
<paragraph id="8BC45C2DEA22D47536D5F8957BFC4E50" blockId="12.[136,1452,1778,1947]" box="[136,1452,1816,1841]" pageId="12" pageNumber="243">
Betong Division, Roban, Sebangkoi,
<geoCoordinate id="EE4F3AEAEA22D4753467F8957C8B4E50" box="[570,731,1816,1841]" degrees="01" direction="north" minutes="57" orientation="latitude" pageId="12" pageNumber="243" precision="1" seconds="29.7" value="1.95825">01°57’29.7”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA22D47534B6F8957DC44E50" box="[747,916,1816,1841]" degrees="111" direction="east" minutes="26" orientation="longitude" pageId="12" pageNumber="243" precision="1" seconds="16.9" value="111.43803">111°26’16.9”E</geoCoordinate>
,
<quantity id="4C83F1C8EA22D47535F9F8957DB34E50" box="[932,995,1816,1841]" metricMagnitude="1" metricUnit="m" metricValue="9.7" pageId="12" pageNumber="243" unit="m" value="97.0">97 m</quantity>
,
<date id="FFC57AEDEA22D47535AEF8957ACA4E50" box="[1011,1178,1816,1841]" pageId="12" pageNumber="243" value="2014-04-22">22 April 2014</date>
,
<emphasis id="B90F803FEA22D47532F4F8957BFC4E50" box="[1193,1452,1816,1841]" italics="true" pageId="12" pageNumber="243">I.H. Ooi &amp; F. Bakong</emphasis>
</paragraph>
<paragraph id="8BC45C2DEA22D47536D5F8B17FB84E34" blockId="12.[136,1452,1778,1947]" box="[136,488,1852,1877]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47536D5F8B17EB44E34" box="[136,228,1852,1877]" italics="true" pageId="12" pageNumber="243">OIH141</emphasis>
(holo SAR!),
<figureCitation id="134040A8EA22D47537D9F8B17FB84E34" box="[388,488,1852,1877]" captionStart="FIGURE 7" captionStartId="14.[136,229,1982,2003]" captionTargetBox="[172,1418,195,1955]" captionTargetId="figure-17@14.[170,1418,191,1956]" captionTargetPageId="14" captionText="FIGURE 7. Scaphochlamys scintillans. A. plant in habitat. B. inflorescence, artificially opened showing bracts and first bracteoles. C. lamina, adaxial surface. D. flower, top view. E. lamina, abaxial surface. F. stamen, note spurless anther thecae. G. flower, front view. H. lamina, adaxial surface, enlarged showing variegation.All from OIH141." pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47537D9F8B17FB84E34" bold="true" box="[388,488,1852,1877]" pageId="12" pageNumber="243">Figure 7</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA22D47636D5F8EF7F604AB0" lastPageId="15" lastPageNumber="246" pageId="12" pageNumber="243" type="diagnosis">
<paragraph id="8BC45C2DEA22D47536D5F8EF7CB94EFA" blockId="12.[136,1452,1778,1947]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47536D5F8EF7EBB4E16" bold="true" box="[136,235,1890,1911]" pageId="12" pageNumber="243">Diagnosis</emphasis>
:—
<taxonomicName id="4C7B27AEEA22D4753754F8EF7C584E16" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[265,520,1890,1911]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="scintillans">
<emphasis id="B90F803FEA22D4753754F8EF7C584E16" box="[265,520,1890,1911]" italics="true" pageId="12" pageNumber="243">Scaphochlamys scintillans</emphasis>
</taxonomicName>
is distinctive from all other
<taxonomicName id="4C7B27AEEA22D4753546F8EF7DAB4E16" box="[795,1019,1890,1911]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D4753546F8EF7DE04E16" box="[795,944,1890,1911]" italics="true" pageId="12" pageNumber="243">Scaphochlamys</emphasis>
species
</taxonomicName>
by the adaxially satin (glittering) dark green lamina with two refractive grey bands (feathered pattern).
</paragraph>
<caption id="DF040CA5EA23D47436D5F8537BF54E92" box="[136,1445,2014,2035]" pageId="13" pageNumber="244" startId="13.[136,229,2014,2035]" targetBox="[153,1436,175,1986]" targetPageId="13" targetType="figure">
<paragraph id="8BC45C2DEA23D47436D5F8537BF54E92" blockId="13.[136,1445,2014,2035]" box="[136,1445,2014,2035]" pageId="13" pageNumber="244">
<emphasis id="B90F803FEA23D47436D5F8537EAD4E92" bold="true" box="[136,253,2014,2035]" pageId="13" pageNumber="244">FIGURE 6.</emphasis>
<emphasis id="B90F803FEA23D474375EF8537C6B4E92" box="[259,571,2014,2035]" italics="true" pageId="13" pageNumber="244">
<taxonomicName id="4C7B27AEEA23D474375EF8537C684E92" authorityName="Meekiong, Ampeng &amp; Ipor" authorityYear="2011" box="[259,568,2014,2035]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="13" pageNumber="244" phylum="Tracheophyta" rank="species" species="salahuddiniana">Scaphochlamys salahuddiniana</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA23D474341DF8537C084E92" bold="true" box="[576,600,2014,2035]" pageId="13" pageNumber="244">A.</emphasis>
lithophytic form in habitat.
<emphasis id="B90F803FEA23D474353BF8537D2C4E92" bold="true" box="[870,892,2014,2035]" pageId="13" pageNumber="244">B.</emphasis>
inflorescence.
<emphasis id="B90F803FEA23D4743253F8537A754E92" bold="true" box="[1038,1061,2014,2035]" pageId="13" pageNumber="244">C.</emphasis>
infructescence. (
<bibRefCitation id="EFEA21DCEA23D4743295F8537BCA4E92" author="Meekiong, K. &amp; Ipor, I. B. &amp; Tawan, C. S. &amp; Ibrahim, H. &amp; Norhati, M. R. &amp; Lim, C. K. &amp; Ampeng, A." box="[1224,1434,2014,2035]" pageId="13" pageNumber="244" pagination="9 - 26" refId="ref32485" refString="Meekiong, K., Ipor, I. B., Tawan, C. S., Ibrahim, H., Norhati, M. R., Lim, C. K. &amp; Ampeng, A. (2011) Five new ginger species (Zingiberaceae) from the eastern part of Lanjak Entimau Wildlife Sanctuary, Sarawak, Borneo. Folia Malaysiana 12: 9 - 26." type="journal article" year="2011">
Meekiong
<emphasis id="B90F803FEA23D474336CF8537B324E92" box="[1329,1378,2014,2035]" italics="true" pageId="13" pageNumber="244">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</caption>
<caption id="DF040CA5EA20D47736D5F8337D16417A" pageId="14" pageNumber="245" startId="14.[136,229,1982,2003]" targetBox="[172,1418,195,1955]" targetPageId="14" targetType="figure">
<paragraph id="8BC45C2DEA20D47736D5F8337D16417A" blockId="14.[136,1452,1982,2075]" pageId="14" pageNumber="245">
<emphasis id="B90F803FEA20D47736D5F8337EAF4EB2" bold="true" box="[136,255,1982,2003]" pageId="14" pageNumber="245">FIGURE 7.</emphasis>
<taxonomicName id="4C7B27AEEA20D477375BF8337C564EB2" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[262,518,1982,2003]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="14" pageNumber="245" phylum="Tracheophyta" rank="species" species="scintillans">
<emphasis id="B90F803FEA20D477375BF8337C564EB2" box="[262,518,1982,2003]" italics="true" pageId="14" pageNumber="245">Scaphochlamys scintillans</emphasis>
</taxonomicName>
.
<emphasis id="B90F803FEA20D477344EF8337C7A4EB2" bold="true" box="[531,554,1982,2003]" pageId="14" pageNumber="245">A.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA20D477348CF8337CB84EB2" bold="true" box="[721,744,1982,2003]" pageId="14" pageNumber="245">B.</emphasis>
inflorescence, artificially opened showing bracts and first bracteoles.
<emphasis id="B90F803FEA20D47733C9F8337BFB4EB2" bold="true" box="[1428,1451,1982,2003]" pageId="14" pageNumber="245">C.</emphasis>
lamina, adaxial surface.
<emphasis id="B90F803FEA20D477372EF86F7FDB4E96" bold="true" box="[371,395,2018,2039]" pageId="14" pageNumber="245">D.</emphasis>
flower, top view.
<emphasis id="B90F803FEA20D4773467F86F7C004E96" bold="true" box="[570,592,2018,2039]" pageId="14" pageNumber="245">E.</emphasis>
lamina, abaxial surface.
<emphasis id="B90F803FEA20D477351FF86F7D074E96" bold="true" box="[834,855,2018,2039]" pageId="14" pageNumber="245">F.</emphasis>
stamen, note spurless anther thecae.
<emphasis id="B90F803FEA20D47732E6F86F7A844E96" bold="true" box="[1211,1236,2018,2039]" pageId="14" pageNumber="245">G.</emphasis>
flower, front view.
<emphasis id="B90F803FEA20D47733CEF86F7BFC4E96" bold="true" box="[1427,1452,2018,2039]" pageId="14" pageNumber="245">H.</emphasis>
lamina, adaxial surface, enlarged showing variegation. All from
<emphasis id="B90F803FEA20D47734ACF78B7D10417A" box="[753,832,2054,2075]" italics="true" pageId="14" pageNumber="245">OIH141</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BC45C2DEA21D47636D5FF2D7F604AB0" blockId="15.[136,1452,160,1157]" pageId="15" pageNumber="246">
Terrestrial, perennial, rhizomatous herbs, to c.
<quantity id="4C83F1C8EA21D47634C8FF2D7C8B49D8" box="[661,731,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="246" unit="cm" value="18.0">18 cm</quantity>
tall. Rhizome buried in ground, c.
<quantity id="4C83F1C8EA21D476323BFF2D7AF849D8" box="[1126,1192,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="246" unit="mm" value="5.0">5 mm</quantity>
in diameter, externally dark greenish brown, internally brownish white. Elements
<quantity id="4C83F1C8EA21D4763575FF497DC249BC" box="[808,914,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="15" pageNumber="246" unit="cm" value="0.55" valueMax="1.0" valueMin="0.1">0.1–1 cm</quantity>
apart, 1-foliate, juvenile not seen; petiole
<quantity id="4C83F1C8EA21D4763326FF497E884860" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="15" pageNumber="246" unit="cm" value="4.0" valueMax="4.5" valueMin="3.5">3.5– 4.5 cm</quantity>
long, canaliculate, glabrous, medium green, base pulvinate, yellowish green; leafless sheath c. 3,
<quantity id="4C83F1C8EA21D4763305FF657EFA4844" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="4.5" metricValueMin="0.7" pageId="15" pageNumber="246" unit="cm" value="2.6" valueMax="4.5" valueMin="0.7">0.7–4.5 cm</quantity>
long, light brown, glabrous, drying and shredding with age; leaf sheath c.
<quantity id="4C83F1C8EA21D476325BFE817A6F4844" box="[1030,1087,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="246" unit="cm" value="1.0">1 cm</quantity>
long, membranous, light green, glabrous, drying and shredding with age; ligule obscure; lamina 10–14 ×
<quantity id="4C83F1C8EA21D47635E7FEBD7A5F4828" box="[954,1039,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="15" pageNumber="246" unit="cm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 cm</quantity>
, elliptic to broad elliptic, coriaceous, margin entire, base cuneate to rounded, apex obtuse to mucronate; adaxial surface satiny (glittering-like), dark green, variegated (refractive grey bands, feathered pattern on each side), glabrous, midrib sunken, dark green, glabrous, main lateral veins inconspicuous; abaxial surface glaucous greenish white, glabrous, midrib raised, light green, pubescent, main lateral veins slightly visible. Inflorescence
<quantity id="4C83F1C8EA21D47634FCFE4D7CA248B8" box="[673,754,448,473]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="15" pageNumber="246" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
long, emerging from near leaf base inside leaf sheaths, slightly lax, comprised of c. 7, 2-flowered cincinni borne on a peduncle, flowering spirally from base to apex; peduncle c.
<quantity id="4C83F1C8EA21D4763328FE697BFB489C" box="[1397,1451,484,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="246" unit="cm" value="2.0">2 cm</quantity>
long, whitish light green, glabrous, covered with marcescent leafless sheath; bract c. 7, c. 4 ×
<quantity id="4C83F1C8EA21D47632C7FD857A8D4B40" box="[1178,1245,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="246" unit="mm" value="3.0">3 mm</quantity>
, spirally arranged, ovate, glabrous, brown, apex acute, each bract subtending c. 2 flowers; bracteole c. 2,
<quantity id="4C83F1C8EA21D476323BFDA17AB14B24" box="[1126,1249,556,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="15" pageNumber="246" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
long, longer than bracts but barely distinguishable from bract, brown; flower c.
<quantity id="4C83F1C8EA21D4763516FDDD7DC94B08" box="[843,921,592,617]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="15" pageNumber="246" unit="cm" value="2.4">2.4 cm</quantity>
long, without noticeable floral odour; calyx c.
<quantity id="4C83F1C8EA21D47636D5FDF97E9B4BEC" box="[136,203,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="246" unit="mm" value="9.0">9 mm</quantity>
long, light brown, sparsely pubescent, apex bidentate; floral tube
<quantity id="4C83F1C8EA21D4763599FDF97A6F4BEC" box="[964,1087,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="15" pageNumber="246" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
long, white to slightly greenish white, sparsely pubescent; corolla lobes
<quantity id="4C83F1C8EA21D476340AFD157CE74BD0" box="[599,695,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="15" pageNumber="246" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long, lanceolate, white to slightly brownish white, glabrous, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA21D47637EAFD317C444BB4" box="[439,532,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="15" pageNumber="246" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, light yellow, apex rounded; labellum 8–11 ×
<quantity id="4C83F1C8EA21D4763421FD6D7CB84B98" box="[636,744,736,761]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="15" pageNumber="246" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white with a light yellow median band (sometimes slightly tinged lilac), apex 2-lobed, slightly fan-shaped, indented c.
<quantity id="4C83F1C8EA21D4763338FC897BF74A7C" box="[1381,1447,772,797]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="246" unit="mm" value="4.0">4 mm</quantity>
, lobes sometimes slightly overlapping; stamen c. 4 ×
<quantity id="4C83F1C8EA21D476348AFCA57D7F4A20" box="[727,815,808,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="246" unit="mm" value="1.5">1.5 mm</quantity>
, covered with glandular hairs, white; filament c.
<quantity id="4C83F1C8EA21D476330BFCA57BFC4A20" box="[1366,1452,808,833]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="246" unit="mm" value="0.5">0.5 mm</quantity>
long; anther thecae c.
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long, spurless, crest c.
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long; stigma less than
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long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
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long, white, glabrous; ovary c.
<quantity id="4C83F1C8EA21D4763268FCFD7A254AE8" box="[1077,1141,880,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="246" unit="mm" value="2.0">2 mm</quantity>
long, 1-locular, light green, densely pubescent, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA21D4763560FC197D2E4ACC" box="[829,894,916,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="246" unit="mm" value="1.0">1 mm</quantity>
long, free, needle-like, brownish white. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA21D47636E0FC517A664A94" box="[189,1078,988,1013]" pageId="15" pageNumber="246" type="etymology">
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<emphasis id="B90F803FEA21D47636E0FC517F6E4A94" bold="true" box="[189,318,988,1013]" pageId="15" pageNumber="246">Etymology</emphasis>
:—The specific epithet refers to the glittering satiny lamina surface.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA21D47636E0FB8D7DD64DE4" pageId="15" pageNumber="246" type="distribution">
<paragraph id="8BC45C2DEA21D47636E0FB8D7D1F4D5C" blockId="15.[136,1452,160,1157]" pageId="15" pageNumber="246">
<emphasis id="B90F803FEA21D47636E0FB8D7FBB4D78" bold="true" box="[189,491,1024,1049]" pageId="15" pageNumber="246">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA21D47637AFFB8D7D674D78" box="[498,823,1024,1049]" italics="true" pageId="15" pageNumber="246">
—
<taxonomicName id="4C7B27AEEA21D4763452FB8D7D674D78" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[527,823,1024,1049]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="15" pageNumber="246" phylum="Tracheophyta" rank="species" species="scintillans">Scaphochlamys scintillans</taxonomicName>
</emphasis>
is known only from the
<typeStatus id="54C0E28FEA21D476323DFB8D7AC04D78" box="[1120,1168,1024,1049]" pageId="15" pageNumber="246">type</typeStatus>
locality where it occurs lowland forest red-yellow soil over shales, at c.
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elevation.
</paragraph>
<paragraph id="8BC45C2DEA21D47636E0FBC57DD64DE4" blockId="15.[136,1452,160,1157]" pageId="15" pageNumber="246">
<emphasis id="B90F803FEA21D47636E0FBC57FFF4D00" bold="true" box="[189,431,1096,1121]" pageId="15" pageNumber="246">Conservation status</emphasis>
:—Near Threatened (NT),
<bibRefCitation id="EFEA21DCEA21D47634B7FBC57DD84D00" author="IUCN" box="[746,904,1096,1121]" pageId="15" pageNumber="246" refId="ref32181" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. Second Edition, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
. The
<typeStatus id="54C0E28FEA21D4763593FBC57DAE4D00" box="[974,1022,1096,1121]" pageId="15" pageNumber="246">type</typeStatus>
locality of
<taxonomicName id="4C7B27AEEA21D47632D0FBC57B714D00" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[1165,1313,1096,1121]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="15" pageNumber="246" phylum="Tracheophyta" rank="species" species="scintillans">
<emphasis id="B90F803FEA21D47632D0FBC57B714D00" box="[1165,1313,1096,1121]" italics="true" pageId="15" pageNumber="246">S. scintillans</emphasis>
</taxonomicName>
is within a nominally protected recreational park although close to a main road.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E39A7E68F93FE82A8F22C05A79D5E0D5">A review of Scaphochlamys (Zingiberaceae) from Borneo, with description of eleven new species</mods:title>
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<subSubSection id="C3610FA6EA24D47336D5FAE57BFC4CE0" box="[136,1452,1382,1409]" pageId="10" pageNumber="241" type="nomenclature">
<paragraph id="8BC45C2DEA24D47336D5FAE57BFC4CE0" blockId="10.[136,1452,1382,1551]" box="[136,1452,1382,1409]" pageId="10" pageNumber="241">
<heading id="D08CEB41EA24D47336D5FAE57BFC4CE0" box="[136,1452,1382,1409]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="241" reason="4">
<emphasis id="B90F803FEA24D47336D5FAE57ECD4CE0" bold="true" box="[136,157,1384,1409]" pageId="10" pageNumber="241">5.</emphasis>
<taxonomicName id="4C7B27AEEA24D47336F8FAEB7D1B4CE0" ID-CoL="8X5G6" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[165,843,1382,1409]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="241" phylum="Tracheophyta" rank="species" species="penyamar" status="sp. nov.">
<emphasis id="B90F803FEA24D47336F8FAEB7F854CE0" bold="true" box="[165,469,1382,1409]" italics="true" pageId="10" pageNumber="241">Scaphochlamys penyamar</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
</taxonomicName>
,
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<taxonomicNameLabel id="A23C3D44EA24D4733505FAE57DFE4CE0" box="[856,942,1384,1409]" pageId="10" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B90F803FEA24D47335E8FAE57DA04CE0" bold="true" box="[949,1008,1384,1409]" pageId="10" pageNumber="241">
<typeStatus id="54C0E28FEA24D47335E8FAE57DA04CE0" box="[949,1008,1384,1409]" pageId="10" pageNumber="241">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA24D473331EFAE57BF84CE0" box="[1347,1448,1384,1409]" name="Malaysia" pageId="10" pageNumber="241">Sarawak</collectingCountry>
:
</heading>
</paragraph>
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<subSubSection id="C3610FA6EA24D47336D5FA017FDC4CA8" pageId="10" pageNumber="241" type="description">
<paragraph id="8BC45C2DEA24D47336D5FA017BFC4CC4" blockId="10.[136,1452,1382,1551]" box="[136,1452,1420,1445]" pageId="10" pageNumber="241">
Sarikei Division, Pakan, Berasok,
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,
<geoCoordinate id="EE4F3AEAEA24D47334E9FA017D0D4CC4" box="[692,861,1420,1445]" degrees="111" direction="east" minutes="38" orientation="longitude" pageId="10" pageNumber="241" precision="1" seconds="52.3" value="111.64786">111°38’52.3”E</geoCoordinate>
,
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,
<date id="FFC57AEDEA24D47335F2FA017A1D4CC4" box="[943,1101,1420,1445]" pageId="10" pageNumber="241" value="2014-04-21">21 April 2014</date>
,
<emphasis id="B90F803FEA24D4733205FA017BFC4CC4" box="[1112,1452,1420,1445]" italics="true" pageId="10" pageNumber="241">I.H. Ooi &amp; F. Bakong OIH137</emphasis>
</paragraph>
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(holo, SAR!),
<figureCitation id="134040A8EA24D4733775FA3D7FDC4CA8" box="[296,396,1456,1481]" captionStart="FIGURE 5" captionStartId="11.[136,229,1996,2017]" captionTargetBox="[165,1426,183,1963]" captionTargetId="figure-14@11.[159,1429,175,1971]" captionTargetPageId="11" captionText="FIGURE 5. Scaphochlamys penyamar. A. plant in habitat. B. rhizome. C. lamina, abaxial surface. D. flower, side view. E. flower, artificially opened showing ovary, epigynous glands, floral tube, and labellum. F.inflorescence with leafless sheath artificially opened. G. flower, front view. H. stamen, note spurless thecae. I. Calyx and ovary.All from OIH137." pageId="10" pageNumber="241">
<emphasis id="B90F803FEA24D4733775FA3D7FDC4CA8" bold="true" box="[296,396,1456,1481]" pageId="10" pageNumber="241">Figure 5</emphasis>
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<emphasis id="B90F803FEA24D47336D5FA5B7EBB4C8A" bold="true" box="[136,235,1494,1515]" pageId="10" pageNumber="241">Diagnosis</emphasis>
:—
<taxonomicName id="4C7B27AEEA24D4733754FA5B7C554C8A" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[265,517,1494,1515]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="241" phylum="Tracheophyta" rank="species" species="penyamar">
<emphasis id="B90F803FEA24D4733754FA5B7C554C8A" box="[265,517,1494,1515]" italics="true" pageId="10" pageNumber="241">Scaphochlamys penyamar</emphasis>
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is similar to
<taxonomicName id="4C7B27AEEA24D47334D9FA5B7D614C8A" box="[644,817,1494,1515]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="241" phylum="Tracheophyta" rank="species" species="salahuddiniana">
<emphasis id="B90F803FEA24D47334D9FA5B7D614C8A" box="[644,817,1494,1515]" italics="true" pageId="10" pageNumber="241">S. salahuddiniana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B27AEEA24D473353CFA5B7DFF4C8A" authorityName="Meekiong &amp; Ampeng" authorityYear="2011" box="[865,943,1494,1515]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="241" phylum="Tracheophyta" rank="species" species="iporii">
<emphasis id="B90F803FEA24D473353CFA5B7DFF4C8A" box="[865,943,1494,1515]" italics="true" pageId="10" pageNumber="241">S. iporii</emphasis>
</taxonomicName>
in the laxly arranged inflorescences and small white flower, but readily distinguishable by the creeping habit and variegated leaf laminae.
</paragraph>
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Terrestrial, perennial, rhizomatous herbs, to c.
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tall. Rhizome creeping above ground with stilt roots, c.
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in diameter, externally light greenish brown, internally brownish white. Elements
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apart, 1-foliate, juvenile not seen; petiole
<quantity id="4C83F1C8EA24D4733712F9057F9F4FC0" box="[335,463,1672,1697]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="10" pageNumber="241" unit="cm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5 cm</quantity>
long, canaliculate, medium green with scattered white dots, glabrous, base pulvinate, light green; leafless sheath c. 3,
<quantity id="4C83F1C8EA24D47337AAF9217C304FA4" box="[503,608,1708,1733]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="5.0" metricValueMin="0.8" pageId="10" pageNumber="241" unit="cm" value="2.9" valueMax="5.0" valueMin="0.8">0.8–5 cm</quantity>
long, light brown, glabrous, drying and shredding with age; leaf sheath c.
<quantity id="4C83F1C8EA24D47336D5F95D7E844F88" box="[136,212,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="241" unit="cm" value="0.4">0.4 cm</quantity>
long, membranous, light green, glabrous, drying and shredding with age; ligule obscure; lamina 9–12 ×
<quantity id="4C83F1C8EA24D4733330F95D7EFD4E6C" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="10" pageNumber="241" unit="cm" value="7.25" valueMax="8.0" valueMin="6.5">6.5–8 cm</quantity>
, ovate, chartaceous, margin slightly undulate, base attenuate, apex acute; adaxial surface matte, medium green, glabrous, variegated with two refractive grey broad longitudinal bands on each side of midrib, midrib sunken, medium green, glabrous, main lateral veins slightly visible; abaxial surface light green, sparsely pubescent, midrib raised, medium green, glabrous, main lateral veins inconspicuous. Inflorescence
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long, emerging from near leaf base inside leaf sheaths, laxly arranged, comprised of c. 5, 2-flowered cincinni borne on a peduncle; peduncle c.
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long, whitish green, glabrous, covered with marcescent leafless sheaths; bract c. 5, c. 4 ×
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, spirally arranged, ovate, brown, glabrous, apex acute, each bract subtending c. 2 flowers; bracteole c. 2,
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long, longer than bracts but barely distinguishable from bract, brown; flower c.
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long, floral odour reminiscent of torch-ginger (
<emphasis id="B90F803FEA24D473331FF87D7E8C414C" italics="true" pageId="10" pageNumber="241">
<taxonomicName id="4C7B27AEEA24D473331FF87D7E84414C" class="Liliopsida" family="Zingiberaceae" genus="Etlingera" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="241" phylum="Tracheophyta" rank="species" species="elatior">Etlingera elatior</taxonomicName>
)
</emphasis>
tissues; calyx c.
<quantity id="4C83F1C8EA24D47337F9F7997FA5414C" box="[420,501,2068,2093]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="241" unit="mm" value="10.0">10 mm</quantity>
long, light brown, sparsely pubescent, apex bidentate; floral tube
<quantity id="4C83F1C8EA24D47332B2F7997B3A414C" box="[1263,1386,2068,2093]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="10" pageNumber="241" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14–15 mm</quantity>
long, white to brownish white, sparsely pubescent; corolla lobes
<quantity id="4C83F1C8EA22D4753561FF2D7DFA49D8" box="[828,938,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="12" pageNumber="243" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long, lanceolate, white to slightly brownish white, glabrous, apex acute, hooded; staminodes
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long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, light yellow, apex rounded; labellum 8–11 ×
<quantity id="4C83F1C8EA22D475352DFF657D8E4860" box="[880,990,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="12" pageNumber="243" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white with a light yellow median band, apex 2-lobed, slightly fan-shaped, indented c.
<quantity id="4C83F1C8EA22D4753343FE817B334844" box="[1310,1379,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="243" unit="mm" value="4.0">4 mm</quantity>
, lobes sometimes slightly overlapping; stamen c. 4 ×
<quantity id="4C83F1C8EA22D47534C8FEBD7CBD4828" box="[661,749,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="243" unit="mm" value="1.5">1.5 mm</quantity>
, covered with glandular hairs, white; filament c.
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long; anther thecae c.
<quantity id="4C83F1C8EA22D4753719FED97FD7480C" box="[324,391,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="243" unit="mm" value="3.0">3 mm</quantity>
long, spurless, crest c.
<quantity id="4C83F1C8EA22D47534C8FED97CBD480C" box="[661,749,340,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="243" unit="mm" value="0.5">0.5 mm</quantity>
long; stigma less than
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long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA22D47534D8FEF57C8448F0" box="[645,724,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="243" unit="mm" value="24.0">24 mm</quantity>
long, white, glabrous; ovary c.
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long, 1-locular, light green, densely pubescent, placentation basal; epigynous glands 2, c.
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long, free, needle-like, brownish white. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<caption id="DF040CA5EA25D47236D5F8417DB54148" pageId="11" pageNumber="242" startId="11.[136,229,1996,2017]" targetBox="[165,1426,183,1963]" targetPageId="11" targetType="figure">
<paragraph id="8BC45C2DEA25D47236D5F8417DB54148" blockId="11.[136,1452,1996,2089]" pageId="11" pageNumber="242">
<emphasis id="B90F803FEA25D47236D5F8417F514E80" bold="true" box="[136,257,1996,2017]" pageId="11" pageNumber="242">FIGURE 5.</emphasis>
<emphasis id="B90F803FEA25D4723757F8417C5D4E80" box="[266,525,1996,2017]" italics="true" pageId="11" pageNumber="242">
<taxonomicName id="4C7B27AEEA25D4723757F8417C5A4E80" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[266,522,1996,2017]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="11" pageNumber="242" phylum="Tracheophyta" rank="species" species="penyamar">Scaphochlamys penyamar</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA25D4723448F8417C7D4E80" bold="true" box="[533,557,1996,2017]" pageId="11" pageNumber="242">A.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA25D4723481F8417CA24E80" bold="true" box="[732,754,1996,2017]" pageId="11" pageNumber="242">B.</emphasis>
rhizome.
<emphasis id="B90F803FEA25D4723504F8417D214E80" bold="true" box="[857,881,1996,2017]" pageId="11" pageNumber="242">C.</emphasis>
lamina, abaxial surface.
<emphasis id="B90F803FEA25D4723233F8417AD64E80" bold="true" box="[1134,1158,1996,2017]" pageId="11" pageNumber="242">D.</emphasis>
flower, side view.
<emphasis id="B90F803FEA25D4723315F8417B0F4E80" bold="true" box="[1352,1375,1996,2017]" pageId="11" pageNumber="242">E.</emphasis>
flower, artificially opened showing ovary, epigynous glands, floral tube, and labellum.
<emphasis id="B90F803FEA25D4723526F87D7DC04164" bold="true" box="[891,912,2032,2053]" pageId="11" pageNumber="242">F.</emphasis>
inflorescence with leafless sheath artificially opened.
<emphasis id="B90F803FEA25D47233CEF87D7BFC4164" bold="true" box="[1427,1452,2032,2053]" pageId="11" pageNumber="242">G.</emphasis>
flower, front view.
<emphasis id="B90F803FEA25D4723762F7997F094148" bold="true" box="[319,345,2068,2089]" pageId="11" pageNumber="242">H.</emphasis>
stamen, note spurless thecae.
<emphasis id="B90F803FEA25D4723427F7997CD94148" bold="true" box="[634,649,2068,2089]" pageId="11" pageNumber="242">I.</emphasis>
Calyx and ovary. All from
<emphasis id="B90F803FEA25D47235CDF7997DB04148" box="[912,992,2068,2089]" italics="true" pageId="11" pageNumber="242">OIH137</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C3610FA6EA22D47536E0FE697D7E4B40" pageId="12" pageNumber="243" type="etymology">
<paragraph id="8BC45C2DEA22D47536E0FE697D7E4B40" blockId="12.[136,1452,160,797]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47536E0FE697F6E489C" bold="true" box="[189,318,484,509]" pageId="12" pageNumber="243">Etymology</emphasis>
:—The specific epithet
<taxonomicName id="4C7B27AEEA22D4753409FE697C94489C" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[596,708,484,509]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="penyamar">
<emphasis id="B90F803FEA22D4753409FE697C94489C" box="[596,708,484,509]" italics="true" pageId="12" pageNumber="243">penyamar</emphasis>
</taxonomicName>
is a Malay word for masquerade, coined to reflect the overall similarity with
<taxonomicName id="4C7B27AEEA22D475376EFD857CC64B40" box="[307,662,520,545]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="salahuddiniana">
<emphasis id="B90F803FEA22D475376EFD857CC64B40" box="[307,662,520,545]" italics="true" pageId="12" pageNumber="243">Scaphochlamys salahuddiniana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B27AEEA22D4753490FD857D774B40" authorityName="Meekiong &amp; Ampeng" authorityYear="2011" box="[717,807,520,545]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="iporii">
<emphasis id="B90F803FEA22D4753490FD857D774B40" box="[717,807,520,545]" italics="true" pageId="12" pageNumber="243">S. iporii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA22D47536E0FDA17B004BD0" pageId="12" pageNumber="243" type="distribution">
<paragraph id="8BC45C2DEA22D47536E0FDA17B774B08" blockId="12.[136,1452,160,797]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47536E0FDA17F9A4B24" bold="true" box="[189,458,556,581]" pageId="12" pageNumber="243">Distribution &amp; ecology</emphasis>
:
<emphasis id="B90F803FEA22D475378CFDA17D5D4B24" box="[465,781,556,581]" italics="true" pageId="12" pageNumber="243">
—
<taxonomicName id="4C7B27AEEA22D47537B3FDA17D5D4B24" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[494,781,556,581]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="penyamar">Scaphochlamys penyamar</taxonomicName>
</emphasis>
is known only from the
<typeStatus id="54C0E28FEA22D4753249FDA17A144B24" box="[1044,1092,556,581]" pageId="12" pageNumber="243">type</typeStatus>
locality occurring along streams in shady kerangas (tropical heath) forest on podzolic black soil over sandstone, at an elevation of c.
<specimenCount id="9D7D97A4EA22D47532BAFDDD7B774B08" box="[1255,1319,592,617]" count="55" pageId="12" pageNumber="243" type="generic">
<quantity id="4C83F1C8EA22D47532BAFDDD7B734B08" box="[1255,1315,592,617]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="243" unit="m" value="55.0">55 m</quantity>
.
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</paragraph>
<paragraph id="8BC45C2DEA22D47536E0FDF97B004BD0" blockId="12.[136,1452,160,797]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47536E0FDF97FE14BEC" bold="true" box="[189,433,628,653]" pageId="12" pageNumber="243">Conservation status</emphasis>
:—Endangered, EN B2ab(iii). The
<typeStatus id="54C0E28FEA22D475350EFDF97DD34BEC" box="[851,899,628,653]" pageId="12" pageNumber="243">type</typeStatus>
locality is not within a protected area and is furthermore close to a logging site. Based on the criteria in
<bibRefCitation id="EFEA21DCEA22D475357FFD157DEB4BD0" author="IUCN" box="[802,955,664,689]" pageId="12" pageNumber="243" refId="ref32181" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. Second Edition, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
, the species is listed as Endangered.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA22D47536E0FD317C084A7C" pageId="12" pageNumber="243" type="discussion">
<paragraph id="8BC45C2DEA22D47536E0FD317C084A7C" blockId="12.[136,1452,160,797]" pageId="12" pageNumber="243">
<emphasis id="B90F803FEA22D47536E0FD317F6B4BB4" bold="true" box="[189,315,700,725]" pageId="12" pageNumber="243">Discussion</emphasis>
:—The adaxial surface of the lamina is variegated with two refractive grey broad longitudinal bands on each side of midrib. Several members in this group (
<taxonomicName id="4C7B27AEEA22D475348BFD6D7D354B98" box="[726,869,736,761]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="scintillans">
<emphasis id="B90F803FEA22D475348BFD6D7D354B98" box="[726,869,736,761]" italics="true" pageId="12" pageNumber="243">S. scintillans</emphasis>
</taxonomicName>
and occasionally
<taxonomicName id="4C7B27AEEA22D4753272FD6D7A994B98" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[1071,1225,736,761]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="243" phylum="Tracheophyta" rank="species" species="lanjakensis">
<emphasis id="B90F803FEA22D4753272FD6D7A994B98" box="[1071,1225,736,761]" italics="true" pageId="12" pageNumber="243">S. lanjakensis</emphasis>
</taxonomicName>
) also have a similar refractive variegation, but not as striking.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="C3610FA6EA31D46636D5FA3D7BFC4CA8" box="[136,1452,1454,1481]" pageId="31" pageNumber="262" type="nomenclature">
<paragraph id="8BC45C2DEA31D46636D5FA3D7BFC4CA8" blockId="31.[136,1452,1454,1623]" box="[136,1452,1454,1481]" pageId="31" pageNumber="262">
<heading id="D08CEB41EA31D46636D5FA3D7BFC4CA8" box="[136,1452,1454,1481]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="262" reason="4">
<emphasis id="B90F803FEA31D46636D5FA3D7EFB4CA8" bold="true" box="[136,171,1456,1481]" pageId="31" pageNumber="262">17.</emphasis>
<taxonomicName id="4C7B27AEEA31D46636EFFA237D004CA8" ID-CoL="8X5G4" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[178,848,1454,1481]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="multifolia" status="sp. nov.">
<emphasis id="B90F803FEA31D46636EFFA237FB04CA8" bold="true" box="[178,480,1454,1481]" italics="true" pageId="31" pageNumber="262">Scaphochlamys multifolia</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
</taxonomicName>
,
<emphasis id="B90F803FEA31D4663501FA3D7DE14CA8" box="[860,945,1456,1481]" italics="true" pageId="31" pageNumber="262">
<taxonomicNameLabel id="A23C3D44EA31D4663501FA3D7DE14CA8" box="[860,945,1456,1481]" pageId="31" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B90F803FEA31D46635EAFA3D7DA24CA8" bold="true" box="[951,1010,1456,1481]" pageId="31" pageNumber="262">
<typeStatus id="54C0E28FEA31D46635EAFA3D7DA24CA8" box="[951,1010,1456,1481]" pageId="31" pageNumber="262">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA31D466331EFA3D7BF84CA8" box="[1347,1448,1456,1481]" name="Malaysia" pageId="31" pageNumber="262">Sarawak</collectingCountry>
:
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA31D46636D5FA597FB84F70" pageId="31" pageNumber="262" type="description">
<paragraph id="8BC45C2DEA31D46636D5FA597BFB4C8C" blockId="31.[136,1452,1454,1623]" box="[136,1451,1492,1517]" pageId="31" pageNumber="262">
Kuching Division, Bau, Gunung Moi,
<geoCoordinate id="EE4F3AEAEA31D466341CFA597CB24C8C" box="[577,738,1492,1517]" degrees="01" direction="north" minutes="31" orientation="latitude" pageId="31" pageNumber="262" precision="1" seconds="52.7" value="1.5313056">01°31’52.7”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA31D46634B2FA597DCA4C8C" box="[751,922,1492,1517]" degrees="110" direction="east" minutes="07" orientation="longitude" pageId="31" pageNumber="262" precision="1" seconds="42.5" value="110.12847">110°07’42.5”E</geoCoordinate>
,
<quantity id="4C83F1C8EA31D46635FAFA597DA14C8C" box="[935,1009,1492,1517]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="31" pageNumber="262" unit="m" value="118.0">118 m</quantity>
,
<date id="FFC57AEDEA31D46635A2FA597AE14C8C" box="[1023,1201,1492,1517]" pageId="31" pageNumber="262" value="2014-03-30">30 March 2014</date>
,
<emphasis id="B90F803FEA31D46632E3FA597BFB4C8C" box="[1214,1451,1492,1517]" italics="true" pageId="31" pageNumber="262">I.H. Ooi &amp; T. Jepom</emphasis>
</paragraph>
<paragraph id="8BC45C2DEA31D46636D5FA757FB84F70" blockId="31.[136,1452,1454,1623]" box="[136,488,1528,1553]" pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D46636D5FA757EB04F70" box="[136,224,1528,1553]" italics="true" pageId="31" pageNumber="262">OIH111</emphasis>
(holo SAR),
<figureCitation id="134040A8EA31D466372AFA757FB84F70" box="[375,488,1528,1553]" captionStart="FIGURE 17" captionStartId="33.[136,229,1996,2017]" captionTargetBox="[161,1428,180,1967]" captionTargetId="figure-14@33.[159,1429,175,1971]" captionTargetPageId="33" captionText="FIGURE 17. Scaphochlamys multifolia. A. plant in habitat, side view. B. plant in habitat, top view. C. lamina, adaxial surface. D. lamina, abaxial surface. E. flower, side view. F. infructescence, artificially revealed fruit. G. young infructescence. H. flower, front view. I. stamen, note very short anther spurs. J. bract and bracteoles.All from OIH111." pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D466372AFA757FB84F70" bold="true" box="[375,488,1528,1553]" pageId="31" pageNumber="262">Figure 17</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA31D45B36D5F9937C514B08" lastPageId="34" lastPageNumber="265" pageId="31" pageNumber="262" type="diagnosis">
<paragraph id="8BC45C2DEA31D46636D5F9937AE14F36" blockId="31.[136,1452,1454,1623]" pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D46636D5F9937EBB4F52" bold="true" box="[136,235,1566,1587]" pageId="31" pageNumber="262">Diagnosis</emphasis>
:—
<taxonomicName id="4C7B27AEEA31D4663754F9937C544F52" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[265,516,1566,1587]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="multifolia">
<emphasis id="B90F803FEA31D4663754F9937C544F52" box="[265,516,1566,1587]" italics="true" pageId="31" pageNumber="262">Scaphochlamys multifolia</emphasis>
</taxonomicName>
is readily distinguishable from
<taxonomicName id="4C7B27AEEA31D4663560F9937DFC4F52" box="[829,940,1566,1587]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="petiolata">
<emphasis id="B90F803FEA31D4663560F9937DFC4F52" box="[829,940,1566,1587]" italics="true" pageId="31" pageNumber="262">S. petiolata</emphasis>
</taxonomicName>
by having polyphyllous elements, tightly congested terminal inflorescences within persistent leafless sheaths and lamina often with a yellowish green midrib.
</paragraph>
<paragraph id="8BC45C2DEA31D45B36D5F9057C514B08" blockId="31.[136,1459,1672,2093]" lastBlockId="34.[136,1452,160,941]" lastPageId="34" lastPageNumber="265" pageId="31" pageNumber="262">
Terrestrial, perennial, rhizomatous herbs, to c.
<quantity id="4C83F1C8EA31D46634F6F9057CA44FC0" box="[683,756,1672,1697]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="262" unit="cm" value="40.0">40 cm</quantity>
tall. Rhizome slightly below ground, c.
<quantity id="4C83F1C8EA31D466328EF9057B484FC0" box="[1235,1304,1672,1697]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="262" unit="mm" value="5.0">5 mm</quantity>
in diameter, externally brownish light green, internally brownish white. Elements c.
<quantity id="4C83F1C8EA31D46635F3F9217A494FA4" box="[942,1049,1708,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="31" pageNumber="262" unit="mm" value="12.5" valueMax="20.0" valueMin="5.0">5–20 mm</quantity>
apart, polyphyllous, up to 8-foliate, juvenile up to 8-foliate; petiole
<quantity id="4C83F1C8EA31D46637B7F95D7C094F88" box="[490,601,1744,1769]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" metricValueMax="2.4" metricValueMin="1.2" pageId="31" pageNumber="262" unit="cm" value="18.0" valueMax="24.0" valueMin="12.0">12–24 cm</quantity>
long, canaliculate, light to medium green, glabrous, base slightly pulvinate, white to light green; leafless sheath 0–3,
<quantity id="4C83F1C8EA31D4663402F9797C8D4E6C" box="[607,733,1780,1805]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="31" pageNumber="262" unit="cm" value="2.0" valueMax="3.5" valueMin="0.5">0.5–3.5 cm</quantity>
long, medium green, sparsely pubescent, drying with age; leaf sheath c.
<quantity id="4C83F1C8EA31D46636B2F8957F754E50" box="[239,293,1816,1841]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="262" unit="cm" value="1.0">1 cm</quantity>
long, membranous, white, glabrous, drying and shredding with age; ligule obscure; lamina 12–15 ×
<quantity id="4C83F1C8EA31D4663327F8957E874E34" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.5" metricValueMin="5.5" pageId="31" pageNumber="262" unit="cm" value="7.5" valueMax="9.5" valueMin="5.5">5.5– 9.5 cm</quantity>
, ovate to lanceolate, chartaceous, margin slightly undulate, base rounded, apex acute to obtuse; adaxial surface semi-glossy, light green to medium green, glabrous, midrib sunken, often yellowish green, glabrous, main lateral veins slightly raised; abaxial surface light green, sparsely pubescent, midrib raised, yellowish green, sparsely pubescent, main lateral veins inconspicuous. Inflorescence c.
<quantity id="4C83F1C8EA31D46634E6F8257D614EA0" box="[699,817,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="9.25" metricValueMax="12.5" metricValueMin="6.0" pageId="31" pageNumber="262" unit="cm" value="9.25" valueMax="12.5" valueMin="6.0">6–12.5 cm</quantity>
long, terminal within leafless sheaths, tightly congested, comprised of c. 5, 2–3-flowered cincinni borne on a peduncle; peduncle c.
<quantity id="4C83F1C8EA31D4663580F8417A164E84" box="[989,1094,1996,2021]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="31" pageNumber="262" unit="cm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 cm</quantity>
long, greenish white, glabrous, covered with persistent leafless sheaths; bract c. 5, c. 16 ×
<quantity id="4C83F1C8EA31D4663542F87D7D334168" box="[799,867,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="262" unit="mm" value="5.0">5 mm</quantity>
, spirally arranged, lanceolate, membranous, light to medium green, sparsely pubescent, apex acuminate, each bract subtending 2–3 flowers; bracteole 2–3, c.
<quantity id="4C83F1C8EA31D4663343F7997B3D414C" box="[1310,1389,2068,2093]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="262" unit="mm" value="14.0">14 mm</quantity>
long, shorter than bracts but barely distinguishable from bract, light green; flower
<quantity id="4C83F1C8EA0CD45B35BAFF2D7A1F49D8" box="[999,1103,160,185]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="34" pageNumber="265" unit="cm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 cm</quantity>
long, without noticeable floral odour; calyx
<quantity id="4C83F1C8EA0CD45B3744FF497FD249BC" box="[281,386,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="34" pageNumber="265" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
long, white, sparsely pubescent, apex tridentate; floral tube
<quantity id="4C83F1C8EA0CD45B3245FF497AC049BC" box="[1048,1168,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="34" pageNumber="265" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23–27 mm</quantity>
long, sparsely pubescent, white; corolla lobes
<quantity id="4C83F1C8EA0CD45B372CFF657F8D4860" box="[369,477,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="34" pageNumber="265" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
long, lanceolate, glabrous, yellowish white, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA0CD45B3310FF657BFC4860" box="[1357,1452,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="34" pageNumber="265" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, light yellow, apex acute to rounded; labellum 9–11 ×
<quantity id="4C83F1C8EA0CD45B371AFEBD7FE44828" box="[327,436,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="34" pageNumber="265" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white at base and violet till apex with a light yellow median band, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA0CD45B3541FED97D0E480C" box="[796,862,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="265" unit="mm" value="4.0">4 mm</quantity>
, lobes sometimes slightly overlapping; stamen c. 4 ×
<quantity id="4C83F1C8EA0CD45B36D5FEF57E9B48F0" box="[136,203,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="265" unit="mm" value="2.0">2 mm</quantity>
, covered with glandular hairs, white; filament c.
<quantity id="4C83F1C8EA0CD45B34B1FEF57D1248F0" box="[748,834,376,401]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="265" unit="mm" value="0.5">0.5 mm</quantity>
long; anther thecae c.
<quantity id="4C83F1C8EA0CD45B3260FEF57A2D48F0" box="[1085,1149,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="265" unit="mm" value="3.0">3 mm</quantity>
long, with very short basal spur, crest c.
<quantity id="4C83F1C8EA0CD45B3745FE117F3D48D4" box="[280,365,412,437]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="265" unit="mm" value="0.5">0.5 mm</quantity>
long; stigma less than
<quantity id="4C83F1C8EA0CD45B3435FE117CF748D4" box="[616,679,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="265" unit="mm" value="1.0">1 mm</quantity>
long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA0CD45B3683FE4D7F7D48B8" box="[222,301,448,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="265" unit="mm" value="25.0">25 mm</quantity>
long, white, sparsely pubescent; ovary c.
<quantity id="4C83F1C8EA0CD45B355EFE4D7D1448B8" box="[771,836,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="265" unit="mm" value="2.0">2 mm</quantity>
long, 1-locular, densely pubescent, white, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA0CD45B3781FE697C4F489C" box="[476,543,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="265" unit="mm" value="1.0">1 mm</quantity>
long, free, needle-like, brownish white. Fruit c. 12 ×
<quantity id="4C83F1C8EA0CD45B32D6FE697A80489C" box="[1163,1232,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="265" unit="mm" value="5.0">5 mm</quantity>
, dehiscent capsule, elongated ovoid, sparsely pubescent, whitish green (greenish white translucent cuticle-like pericarp enclosing green seed), subtended by persistent bracts. Seeds c. 2, c. 10 ×
<quantity id="4C83F1C8EA0CD45B34AAFDA17D694B24" box="[759,825,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="265" unit="mm" value="2.0">2 mm</quantity>
, elongated ovoid, glabrous, green; aril c.
<quantity id="4C83F1C8EA0CD45B335CFDA17B134B24" box="[1281,1347,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="265" unit="mm" value="4.0">4 mm</quantity>
, lacerate, brownish white; perisperm white.
</paragraph>
</subSubSection>
<caption id="DF040CA5EA0ED45936D5F8467C434149" pageId="32" pageNumber="263" startId="32.[136,229,1995,2016]" targetBox="[160,1430,181,1962]" targetPageId="32" targetType="figure">
<paragraph id="8BC45C2DEA0ED45936D5F8467C434149" blockId="32.[136,1452,1995,2088]" pageId="32" pageNumber="263">
<emphasis id="B90F803FEA0ED45936D5F8467F5C4E81" bold="true" box="[136,268,1995,2016]" pageId="32" pageNumber="263">FIGURE 16.</emphasis>
<emphasis id="B90F803FEA0ED459374EF8467FBB4E81" box="[275,491,1995,2016]" italics="true" pageId="32" pageNumber="263">
<taxonomicName id="4C7B27AEEA0ED459374EF8467FB74E81" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[275,487,1995,2016]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="32" pageNumber="263" phylum="Tracheophyta" rank="species" species="hasta">Scaphochlamys hasta</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA0ED45937AFF8467C5A4E81" bold="true" box="[498,522,1995,2016]" pageId="32" pageNumber="263">A.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA0ED45934EEF8467C994E81" bold="true" box="[691,713,1995,2016]" pageId="32" pageNumber="263">B.</emphasis>
plant before pressing.
<emphasis id="B90F803FEA0ED45935F1F8467D934E81" bold="true" box="[940,963,1995,2016]" pageId="32" pageNumber="263">C.</emphasis>
lamina, adaxial surface, note sunken main lateral veins.
<emphasis id="B90F803FEA0ED459369AF8627E8E4165" bold="true" box="[199,222,2031,2052]" pageId="32" pageNumber="263">D.</emphasis>
lamina, abaxial surface, note visible dark green main lateral veins.
<emphasis id="B90F803FEA0ED4593538F8627D2C4165" bold="true" box="[869,892,2031,2052]" pageId="32" pageNumber="263">E.</emphasis>
inflorescence.
<emphasis id="B90F803FEA0ED4593250F8627A724165" bold="true" box="[1037,1058,2031,2052]" pageId="32" pageNumber="263">F.</emphasis>
flower.
<emphasis id="B90F803FEA0ED4593232F8627AD84165" bold="true" box="[1135,1160,2031,2052]" pageId="32" pageNumber="263">G.</emphasis>
stamen, note spurless anthers.
<emphasis id="B90F803FEA0ED45936D5F79E7EF14149" bold="true" box="[136,161,2067,2088]" pageId="32" pageNumber="263">H.</emphasis>
bract and bracteoles. All from
<emphasis id="B90F803FEA0ED4593796F79E7C5D4149" box="[459,525,2067,2088]" italics="true" pageId="32" pageNumber="263">OIH80</emphasis>
.
</paragraph>
</caption>
<caption id="DF040CA5EA0FD45836D5F8417D2C4148" pageId="33" pageNumber="264" startId="33.[136,229,1996,2017]" targetBox="[161,1428,180,1967]" targetPageId="33" targetType="figure">
<paragraph id="8BC45C2DEA0FD45836D5F8417D2C4148" blockId="33.[136,1452,1996,2089]" pageId="33" pageNumber="264">
<emphasis id="B90F803FEA0FD45836D5F8417F594E80" bold="true" box="[136,265,1996,2017]" pageId="33" pageNumber="264">FIGURE 17.</emphasis>
<emphasis id="B90F803FEA0FD4583752F8417C5E4E80" box="[271,526,1996,2017]" italics="true" pageId="33" pageNumber="264">
<taxonomicName id="4C7B27AEEA0FD4583752F8417C5A4E80" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[271,522,1996,2017]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="33" pageNumber="264" phylum="Tracheophyta" rank="species" species="multifolia">Scaphochlamys multifolia</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA0FD458344FF8417C7A4E80" bold="true" box="[530,554,1996,2017]" pageId="33" pageNumber="264">A.</emphasis>
plant in habitat, side view.
<emphasis id="B90F803FEA0FD4583572F8417D154E80" bold="true" box="[815,837,1996,2017]" pageId="33" pageNumber="264">B.</emphasis>
plant in habitat, top view.
<emphasis id="B90F803FEA0FD458321FF8417A094E80" bold="true" box="[1090,1113,1996,2017]" pageId="33" pageNumber="264">C.</emphasis>
lamina, adaxial surface.
<emphasis id="B90F803FEA0FD458331AF8417B0F4E80" bold="true" box="[1351,1375,1996,2017]" pageId="33" pageNumber="264">D.</emphasis>
lamina, abaxial surface.
<emphasis id="B90F803FEA0FD4583776F87D7F114164" bold="true" box="[299,321,2032,2053]" pageId="33" pageNumber="264">E.</emphasis>
flower, side view.
<emphasis id="B90F803FEA0FD458345EF87D7C484164" bold="true" box="[515,536,2032,2053]" pageId="33" pageNumber="264">F.</emphasis>
infructescence, artificially revealed fruit.
<emphasis id="B90F803FEA0FD45835E5F87D7D814164" bold="true" box="[952,977,2032,2053]" pageId="33" pageNumber="264">G.</emphasis>
young infructescence.
<emphasis id="B90F803FEA0FD45832E4F87D7A824164" bold="true" box="[1209,1234,2032,2053]" pageId="33" pageNumber="264">H.</emphasis>
flower, front view.
<emphasis id="B90F803FEA0FD45833C1F87D7BFB4164" bold="true" box="[1436,1451,2032,2053]" pageId="33" pageNumber="264">I.</emphasis>
stamen, note very short anther spurs.
<emphasis id="B90F803FEA0FD45837B2F7997C514148" bold="true" box="[495,513,2068,2089]" pageId="33" pageNumber="264">J.</emphasis>
bract and bracteoles. All from
<emphasis id="B90F803FEA0FD4583576F7997D254148" box="[811,885,2068,2089]" italics="true" pageId="33" pageNumber="264">OIH111</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C3610FA6EA0CD45B36E0FDF97B444BEC" box="[189,1300,628,653]" pageId="34" pageNumber="265" type="etymology">
<paragraph id="8BC45C2DEA0CD45B36E0FDF97B444BEC" blockId="34.[136,1452,160,941]" box="[189,1300,628,653]" pageId="34" pageNumber="265">
<emphasis id="B90F803FEA0CD45B36E0FDF97F6E4BEC" bold="true" box="[189,318,628,653]" pageId="34" pageNumber="265">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C7B27AEEA0CD45B3410FDF97CEA4BEC" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[589,698,628,653]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="multifolia">
<emphasis id="B90F803FEA0CD45B3410FDF97CEA4BEC" box="[589,698,628,653]" italics="true" pageId="34" pageNumber="265">multifolia</emphasis>
</taxonomicName>
, refers to the polyphyllous element of the adult plant.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA0CD45B36E0FD157CBC4A7C" pageId="34" pageNumber="265" type="distribution">
<paragraph id="8BC45C2DEA0CD45B36E0FD157A514BB4" blockId="34.[136,1452,160,941]" pageId="34" pageNumber="265">
<emphasis id="B90F803FEA0CD45B36E0FD157FB84BD0" bold="true" box="[189,488,664,689]" pageId="34" pageNumber="265">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA0CD45B37B2FD157D7C4BD0" box="[495,812,664,689]" italics="true" pageId="34" pageNumber="265">
—
<taxonomicName id="4C7B27AEEA0CD45B3451FD157D7C4BD0" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[524,812,664,689]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="multifolia">Scaphochlamys multifolia</taxonomicName>
</emphasis>
is only known from the
<typeStatus id="54C0E28FEA0CD45B3211FD157A2C4BD0" box="[1100,1148,664,689]" pageId="34" pageNumber="265">type</typeStatus>
locality where it gown on podzolic black soil over sandstone in kerangas (tropical heath) forest, c.
<specimenCount id="9D7D97A4EA0CD45B35EEFD317A514BB4" box="[947,1025,700,725]" count="120" pageId="34" pageNumber="265" type="generic">
<quantity id="4C83F1C8EA0CD45B35EEFD317DAD4BB4" box="[947,1021,700,725]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="265" unit="m" value="120.0">120 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BC45C2DEA0CD45B36E0FD6D7CBC4A7C" blockId="34.[136,1452,160,941]" pageId="34" pageNumber="265">
<emphasis id="B90F803FEA0CD45B36E0FD6D7FFD4B98" bold="true" box="[189,429,736,761]" pageId="34" pageNumber="265">Conservation status</emphasis>
:—Near Threatened (NT). This species is common on the mountain but it is not within a protected area (subjected to conversion of land usage).
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA0CD45B36E0FCA57B1E4ACC" pageId="34" pageNumber="265" type="discussion">
<paragraph id="8BC45C2DEA0CD45B36E0FCA57B1E4ACC" blockId="34.[136,1452,160,941]" pageId="34" pageNumber="265">
<emphasis id="B90F803FEA0CD45B36E0FCA57F6B4A20" bold="true" box="[189,315,808,833]" pageId="34" pageNumber="265">Discussion</emphasis>
:—The leaf lamina of
<taxonomicName id="4C7B27AEEA0CD45B346CFCA57CEA4A20" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[561,698,808,833]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="multifolia">
<emphasis id="B90F803FEA0CD45B346CFCA57CEA4A20" box="[561,698,808,833]" italics="true" pageId="34" pageNumber="265">S. multifolia</emphasis>
</taxonomicName>
is reminiscent of that of
<taxonomicName id="4C7B27AEEA0CD45B3590FCA57A1D4A20" box="[973,1101,808,833]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="petiolata">
<emphasis id="B90F803FEA0CD45B3590FCA57A1D4A20" box="[973,1101,808,833]" italics="true" pageId="34" pageNumber="265">S. petiolata</emphasis>
</taxonomicName>
, but is retained in polyphyllous elements in adult plants. Inflorescences of
<taxonomicName id="4C7B27AEEA0CD45B3423FCC17D5B4A04" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[638,779,844,869]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="multifolia">
<emphasis id="B90F803FEA0CD45B3423FCC17D5B4A04" box="[638,779,844,869]" italics="true" pageId="34" pageNumber="265">S. multifolia</emphasis>
</taxonomicName>
are terminal within sheaths, not emerging from near leaf base like most of the other
<taxonomicName id="4C7B27AEEA0CD45B3797FCFD7C8A4AE8" box="[458,730,880,905]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90F803FEA0CD45B3797FCFD7C284AE8" box="[458,632,880,905]" italics="true" pageId="34" pageNumber="265">Scaphochlamys</emphasis>
species.
</taxonomicName>
<taxonomicName id="4C7B27AEEA0CD45B34B9FCFD7DAE4AE8" authorityName="R. J. Searle" authorityYear="2010" baseAuthorityName="Hallier f." baseAuthorityYear="1898" box="[740,1022,880,905]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="species" species="anomala">
<emphasis id="B90F803FEA0CD45B34B9FCFD7DAE4AE8" box="[740,1022,880,905]" italics="true" pageId="34" pageNumber="265">Scaphochlamys anomala</emphasis>
</taxonomicName>
is the only other species of Bornean
<taxonomicName id="4C7B27AEEA0CD45B36D5FC197F664ACC" authorityName="Baker" authorityYear="1892" box="[136,310,916,941]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="34" pageNumber="265" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F803FEA0CD45B36D5FC197F664ACC" box="[136,310,916,941]" italics="true" pageId="34" pageNumber="265">Scaphochlamys</emphasis>
</taxonomicName>
reported to have terminal inflorescences within sheaths but it belongs to the
<taxonomicName id="4C7B27AEEA0CD45B32CEFC197AA84ACC" authorityName="Samouelle" authorityYear="1819" box="[1171,1272,916,941]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B90F803FEA0CD45B32CEFC197AA84ACC" box="[1171,1272,916,941]" italics="true" pageId="34" pageNumber="265">Anomala</emphasis>
</taxonomicName>
Group.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="C3610FA6EA31D46636D5FF2D7BFC49D8" box="[136,1452,158,185]" pageId="31" pageNumber="262" type="nomenclature">
<paragraph id="8BC45C2DEA31D46636D5FF2D7BFC49D8" blockId="31.[136,1452,158,327]" box="[136,1452,158,185]" pageId="31" pageNumber="262">
<heading id="D08CEB41EA31D46636D5FF2D7BFC49D8" box="[136,1452,158,185]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="262" reason="4">
<emphasis id="B90F803FEA31D46636D5FF2D7EFB49D8" bold="true" box="[136,171,160,185]" pageId="31" pageNumber="262">16.</emphasis>
<taxonomicName id="4C7B27AEEA31D46636EBFF137D6C49D8" ID-CoL="8X5FY" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[182,828,158,185]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="hasta" status="sp. nov.">
<emphasis id="B90F803FEA31D46636EBFF137FE449D8" bold="true" box="[182,436,158,185]" italics="true" pageId="31" pageNumber="262">Scaphochlamys hasta</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
</taxonomicName>
,
<emphasis id="B90F803FEA31D4663511FF2D7DF549D8" box="[844,933,160,185]" italics="true" pageId="31" pageNumber="262">
<taxonomicNameLabel id="A23C3D44EA31D4663511FF2D7DF549D8" box="[844,933,160,185]" pageId="31" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B90F803FEA31D46635F2FF2D7DBA49D8" bold="true" box="[943,1002,160,185]" pageId="31" pageNumber="262">
<typeStatus id="54C0E28FEA31D46635F2FF2D7DBA49D8" box="[943,1002,160,185]" pageId="31" pageNumber="262">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA31D466331EFF2D7BF849D8" box="[1347,1448,160,185]" name="Malaysia" pageId="31" pageNumber="262">Sarawak</collectingCountry>
:
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA31D46636D5FF497F184860" pageId="31" pageNumber="262" type="description">
<paragraph id="8BC45C2DEA31D46636D5FF497BFB49BC" blockId="31.[136,1452,158,327]" box="[136,1451,196,221]" pageId="31" pageNumber="262">
Kuching Division, Bau, Bogag,
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,
<geoCoordinate id="EE4F3AEAEA31D46634FFFF497D1D49BC" box="[674,845,196,221]" degrees="110" direction="east" minutes="03" orientation="longitude" pageId="31" pageNumber="262" precision="1" seconds="44.5" value="110.06236">110°03’44.5”E</geoCoordinate>
,
<date id="FFC57AEDEA31D4663504FF497A4649BC" box="[857,1046,196,221]" pageId="31" pageNumber="262" value="2014-01-11">11 January 2014</date>
,
<emphasis id="B90F803FEA31D466327FFF497B3849BC" box="[1058,1384,196,221]" italics="true" pageId="31" pageNumber="262">I.H. Ooi &amp; K. Jeland OIH80</emphasis>
(holo
</paragraph>
<paragraph id="8BC45C2DEA31D46636D5FF657F184860" blockId="31.[136,1452,158,327]" box="[136,328,232,257]" pageId="31" pageNumber="262">
SAR),
<figureCitation id="134040A8EA31D466368BFF657F184860" box="[214,328,232,257]" captionStart="FIGURE 16" captionStartId="32.[136,229,1995,2016]" captionTargetBox="[160,1430,181,1962]" captionTargetId="figure-17@32.[158,1430,175,1971]" captionTargetPageId="32" captionText="FIGURE 16. Scaphochlamys hasta. A. plant in habitat. B. plant before pressing. C. lamina, adaxial surface, note sunken main lateral veins. D. lamina, abaxial surface, note visible dark green main lateral veins. E. inflorescence. F. flower. G. stamen, note spurless anthers. H. bract and bracteoles.All from OIH80." pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D466368BFF657F184860" bold="true" box="[214,328,232,257]" pageId="31" pageNumber="262">Figure 16</emphasis>
</figureCitation>
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<emphasis id="B90F803FEA31D46636D5FE837EBB4842" bold="true" box="[136,235,270,291]" pageId="31" pageNumber="262">Diagnosis</emphasis>
:—
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<emphasis id="B90F803FEA31D4663754FE837F884842" box="[265,472,270,291]" italics="true" pageId="31" pageNumber="262">Scaphochlamys hasta</emphasis>
</taxonomicName>
is readily distinguishable from
<taxonomicName id="4C7B27AEEA31D466355AFE837D234842" box="[775,883,270,291]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="petiolata">
<emphasis id="B90F803FEA31D466355AFE837D234842" box="[775,883,270,291]" italics="true" pageId="31" pageNumber="262">S. petiolata</emphasis>
</taxonomicName>
by its long lanceolate lamina and long slender petiole, dark green midrib, and adaxially slightly sunken main lateral veins.
</paragraph>
<paragraph id="8BC45C2DEA31D46636D5FEF57CB74DE4" blockId="31.[136,1452,376,1409]" pageId="31" pageNumber="262">
Terrestrial, perennial, rhizomatous herbs, to c.
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tall. Rhizome slightly below ground, c.
<quantity id="4C83F1C8EA31D466328EFEF57B4848F0" box="[1235,1304,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="262" unit="mm" value="5.0">5 mm</quantity>
in diameter, externally greenish light brown, internally yellow. Elements c.
<quantity id="4C83F1C8EA31D4663501FE117D9948D4" box="[860,969,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="15.0" metricValueMin="1.0" pageId="31" pageNumber="262" unit="mm" value="8.0" valueMax="15.0" valueMin="1.0">1–15 mm</quantity>
apart, 1-foliate, juvenile polyphyllous, up to 6-foliate; petiole
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long, canaliculate, light to medium green with white dots, glabrous, base pulvinate, greenish white to brownish green; leafless sheath c. 4,
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long, light green to light brown, glabrous, drying with age; leaf sheath c.
<quantity id="4C83F1C8EA31D4663707FD857FF74B40" box="[346,423,520,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="262" unit="cm" value="1.5">1.5 cm</quantity>
long, membranous, glabrous, white, drying and shredding with age; ligule obscure; lamina 17–22 ×
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, lanceolate, semi-coriaceous, margin entire to slightly undulate, base attenuate, apex acuminate; adaxial surface semi-glossy, light green to medium green, glabrous, midrib sunken, dark green, glabrous, main lateral veins slightly sunken; abaxial surface glabrous, glaucous green, midrib raised, dark green, glabrous, main lateral veins visible. Inflorescence c.
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long, emerging from near leaf base inside leaf sheaths, tightly congested, comprised of 3–5, 2-flowered cincinni borne on a peduncle; peduncle c.
<quantity id="4C83F1C8EA31D466351EFD317DFC4BB4" box="[835,940,700,725]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="31" pageNumber="262" unit="cm" value="5.25" valueMax="6.5" valueMin="4.0">4–6.5 cm</quantity>
long, greenish white, glabrous, covered with marcescent leafless sheaths; bract 3–5, c. 22 ×
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, spirally arranged, lanceolate, membranous, light brown, sparsely pubescent, apex acuminate, each bract subtending 2 flowers; bracteole 2,
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long, shorter than bracts but barely distinguishable from bract, light brown; flower
<quantity id="4C83F1C8EA31D46634A5FCA57D324A20" box="[760,866,808,833]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="31" pageNumber="262" unit="cm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 cm</quantity>
long, without noticeable floral odour; calyx
<quantity id="4C83F1C8EA31D466333BFCA57EE44A04" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="31" pageNumber="262" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
long, white, sparsely pubescent, apex acute; floral tube
<quantity id="4C83F1C8EA31D4663571FCC17DF54A04" box="[812,933,844,869]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="31" pageNumber="262" unit="mm" value="35.0" valueMax="40.0" valueMin="30.0">30–40 mm</quantity>
long, sparsely pubescent, white; corolla lobes
<quantity id="4C83F1C8EA31D46636D5FCFD7EB64AE8" box="[136,230,880,905]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="31" pageNumber="262" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
long, lanceolate, glabrous, yellowish white, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA31D4663210FCFD7AFB4AE8" box="[1101,1195,880,905]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="31" pageNumber="262" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, light yellow, apex acute to rounded; labellum 9–11 ×
<quantity id="4C83F1C8EA31D4663329FC197EE64AB0" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="31" pageNumber="262" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white at base and violet till apex with a light yellow median band, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA31D466341DFC517CD24A94" box="[576,642,988,1013]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="262" unit="mm" value="4.0">4 mm</quantity>
, lobes sometimes slightly overlapping; stamen c. 5 ×
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, covered with glandular hairs, white; filament c.
<quantity id="4C83F1C8EA31D4663458FB8D7C164D78" box="[517,582,1024,1049]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="262" unit="mm" value="1.0">1 mm</quantity>
long; anther thecae c.
<quantity id="4C83F1C8EA31D4663562FB8D7DC44D78" box="[831,916,1024,1049]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="31" pageNumber="262" unit="mm" value="3.5">3.5 mm</quantity>
long, spurless, crest c.
<quantity id="4C83F1C8EA31D46632CBFB8D7ABB4D78" box="[1174,1259,1024,1049]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="262" unit="mm" value="0.5">0.5 mm</quantity>
long; stigma less than
<quantity id="4C83F1C8EA31D46636E2FBA97F504D5C" box="[191,256,1060,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="262" unit="mm" value="1.0">1 mm</quantity>
long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA31D4663233FBA97AED4D5C" box="[1134,1213,1060,1085]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="31" pageNumber="262" unit="mm" value="42.0">42 mm</quantity>
long, white, sparsely pubescent; ovary c.
<quantity id="4C83F1C8EA31D4663738FBC57FF54D00" box="[357,421,1096,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="262" unit="mm" value="2.0">2 mm</quantity>
long, 1-locular, white, pubescent, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA31D46632A8FBC57B654D00" box="[1269,1333,1096,1121]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="262" unit="mm" value="1.0">1 mm</quantity>
long, free, needle-like, yellowish white. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA31D46636E0FB1D7B5A4DC8" box="[189,1290,1168,1193]" pageId="31" pageNumber="262" type="etymology">
<paragraph id="8BC45C2DEA31D46636E0FB1D7B5A4DC8" blockId="31.[136,1452,376,1409]" box="[189,1290,1168,1193]" pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D46636E0FB1D7F6E4DC8" bold="true" box="[189,318,1168,1193]" pageId="31" pageNumber="262">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C7B27AEEA31D4663410FB1D7CDA4DC8" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[589,650,1168,1193]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="hasta">
<emphasis id="B90F803FEA31D4663410FB1D7CDA4DC8" box="[589,650,1168,1193]" italics="true" pageId="31" pageNumber="262">hasta</emphasis>
</taxonomicName>
, spear in Latin, indicates the spear head-like leaf lamina.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA31D46636E0FB397A184C3C" pageId="31" pageNumber="262" type="distribution">
<paragraph id="8BC45C2DEA31D46636E0FB397DE64D90" blockId="31.[136,1452,376,1409]" pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D46636E0FB397F8F4DAC" bold="true" box="[189,479,1204,1229]" pageId="31" pageNumber="262">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA31D46637BAFB397CA04DAC" box="[487,752,1204,1229]" italics="true" pageId="31" pageNumber="262">
—
<taxonomicName id="4C7B27AEEA31D4663459FB397CA04DAC" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[516,752,1204,1229]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="hasta">Scaphochlamys hasta</taxonomicName>
</emphasis>
is only known from the
<typeStatus id="54C0E28FEA31D46635ABFB397A764DAC" box="[1014,1062,1204,1229]" pageId="31" pageNumber="262">type</typeStatus>
locality in shady kerangas (tropical heath) forest on podzolic black soil over sandstone, of c.
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elevation.
</paragraph>
<paragraph id="8BC45C2DEA31D46636E0FB717A184C3C" blockId="31.[136,1452,376,1409]" pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D46636E0FB717FE04C74" bold="true" box="[189,432,1276,1301]" pageId="31" pageNumber="262">Conservation status</emphasis>
:—The conservation status of this species is Vulnerable (VU), meeting criteria B.1.a., B.1.b.(iii), B.2.a., B.2.b.(iii), and D.1.
<taxonomicName id="4C7B27AEEA31D466346FFAAD7D744C58" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[562,804,1312,1337]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="31" pageNumber="262" phylum="Tracheophyta" rank="species" species="hasta">
<emphasis id="B90F803FEA31D466346FFAAD7D744C58" box="[562,804,1312,1337]" italics="true" pageId="31" pageNumber="262">Scaphochlamys hasta</emphasis>
</taxonomicName>
is only known from a single population. An assessment of Vulnerable is appropriate as the area is subjected to conversion into agricultural farm.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA31D46636E0FAE57D7A4CE0" box="[189,810,1384,1409]" pageId="31" pageNumber="262" type="discussion">
<paragraph id="8BC45C2DEA31D46636E0FAE57D7A4CE0" blockId="31.[136,1452,376,1409]" box="[189,810,1384,1409]" pageId="31" pageNumber="262">
<emphasis id="B90F803FEA31D46636E0FAE57F6B4CE0" bold="true" box="[189,315,1384,1409]" pageId="31" pageNumber="262">Discussion</emphasis>
:—Refer to discussion in
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<emphasis id="B90F803FEA31D4663407FAE57C8A4CE0" box="[602,730,1384,1409]" italics="true" pageId="31" pageNumber="262">S. petiolata</emphasis>
</taxonomicName>
below.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="38296B032071F885B0CE1696BCCF5B63">A review of Scaphochlamys (Zingiberaceae) from Borneo, with description of eleven new species</mods:title>
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<emphasis id="B90F803FEA39D46E36D5FCE77EFB4AE2" bold="true" box="[136,171,874,899]" pageId="23" pageNumber="254">12.</emphasis>
<taxonomicName id="4C7B27AEEA39D46E36E8FCE57D644AE2" ID-CoL="6XSWR" authority="Meekiong &amp; Yazid (2015: 33)" authorityName="Meekiong &amp; Yazid" authorityPageNumber="33" authorityYear="2015" box="[181,820,872,899]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="23" pageNumber="254" phylum="Tracheophyta" rank="species" species="limiana">
<emphasis id="B90F803FEA39D46E36E8FCE57F9C4AE2" bold="true" box="[181,460,872,899]" italics="true" pageId="23" pageNumber="254">Scaphochlamys limiana</emphasis>
<bibRefCitation id="EFEA21DCEA39D46E3788FCE77D644AE2" author="Meekiong, K. &amp; Hidir, M. &amp; Yazid, K." box="[469,820,874,899]" pageId="23" pageNumber="254" pagination="31 - 44" refId="ref32559" refString="Meekiong, K., Hidir, M. &amp; Yazid, K. (2015) Three new Scaphochlamys species from Sarawak, Malaysia. Folia Malaysiana 16: 31 - 44." type="journal article" year="2015">Meekiong &amp; Yazid (2015: 33)</bibRefCitation>
</taxonomicName>
.
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<typeStatus id="54C0E28FEA39D46E351CFCE77D2C4AE2" box="[833,892,874,899]" pageId="23" pageNumber="254">Type</typeStatus>
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA39D46E328FFCE77B674AE2" box="[1234,1335,874,899]" name="Malaysia" pageId="23" pageNumber="254">Sarawak</collectingCountry>
: Sematan, Samunsam Wildlife Sanctuary Extension, trail to Ranger Post,
<date id="FFC57AEDEA39D46E3512FC037A5F4AC6" box="[847,1039,910,935]" pageId="23" pageNumber="254" value="2014-10-25">25 October 2014</date>
,
<emphasis id="B90F803FEA39D46E3246FC037F7F4AAA" italics="true" pageId="23" pageNumber="254">K. Meekiong, K. Yazid, W. Yahud &amp; Dami MK2344</emphasis>
(holo SAR!, iso HUMS!),
<figureCitation id="134040A8EA39D46E3402FC3F7C814AAA" box="[607,721,946,971]" captionStart="FIGURE 12" captionStartId="24.[136,229,1960,1981]" captionTargetBox="[175,1415,179,1932]" captionTargetId="figure-17@24.[172,1417,175,1935]" captionTargetPageId="24" captionText="FIGURE 12. Scaphochlamys limiana. A. plant in habitat. B. plant with flower in habitat. C. inflorescence. D. bracts, side view. E. bracts, top view. F.bracts and a mature seed, noted paler bracts during fruiting G. flower, front view. H. flower, side view. I. stamen, noted anther spurs and purplish white crest. J. lamina, abaxial surface. K. roots, note thickened root. L. bracts and bracteoles. M. rhizome, note dark purplish red interior.A, C–F, I–J, L–M from OIH118; B &amp; K from OIH122; G–H from OIH121." pageId="23" pageNumber="254">
<emphasis id="B90F803FEA39D46E3402FC3F7C814AAA" bold="true" box="[607,721,946,971]" pageId="23" pageNumber="254">Figure 12</emphasis>
</figureCitation>
</paragraph>
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<paragraph id="8BC45C2DEA39D46E36D5FC777DB14EDA" blockId="23.[136,1453,1018,2051]" pageId="23" pageNumber="254">
Terrestrial, perennial, rhizomatous herbs, to c.
<quantity id="4C83F1C8EA39D46E34CDFC777C854D72" box="[656,725,1018,1043]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="254" unit="cm" value="19.0">19 cm</quantity>
tall. Rhizome creeping slightly below ground, c.
<quantity id="4C83F1C8EA39D46E32A0FC777B6E4D72" box="[1277,1342,1018,1043]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="254" unit="mm" value="5.0">5 mm</quantity>
diameter, externally medium brown, internally dark purplish red, roots sometimes pseudo-tuberous. Elements
<quantity id="4C83F1C8EA39D46E32A4FB937B344D56" box="[1273,1380,1054,1079]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="2.0" metricValueMin="0.2" pageId="23" pageNumber="254" unit="cm" value="1.1" valueMax="2.0" valueMin="0.2">0.2−2 cm</quantity>
apart, 1-foliate, juvenile not seen; petiole c.
<quantity id="4C83F1C8EA39D46E346AFBCF7C3F4D3A" box="[567,623,1090,1115]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="254" unit="cm" value="5.0">5 cm</quantity>
long, canaliculate, dark purplish red, sparsely pubescent, base pulvinate, dark pinkish white to dark purplish red; leafless sheath c. 4,
<quantity id="4C83F1C8EA39D46E3562FBEB7DFB4D1E" box="[831,939,1126,1151]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="6.0" metricValueMin="0.8" pageId="23" pageNumber="254" unit="cm" value="3.4" valueMax="6.0" valueMin="0.8">0.8−6 cm</quantity>
long, glabrous, dark purplish red with white near base, marcescent with age; leaf sheath c.
<quantity id="4C83F1C8EA39D46E34D3FB077C954DC2" box="[654,709,1162,1187]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="254" unit="cm" value="2.0">2 cm</quantity>
long, membranous, reddish white, glabrous, marcescent with age; ligule obscure, membranous, marcescent with age; lamina 11–14 ×
<quantity id="4C83F1C8EA39D46E35CCFB237DAF4DA6" box="[913,1023,1198,1223]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="23" pageNumber="254" unit="cm" value="5.75" valueMax="6.5" valueMin="5.0">5–6.5 cm</quantity>
, slightly ovate to elliptic, coriaceous, conduplicate, margin slightly undulate, base obtuse, apex acute; adaxial surface glossy, medium green to dark green, glabrous, midrib sunken, main lateral veins inconspicuous; abaxial surface, dark purplish red, glabrous, midrib raised, sparsely pubescent, main lateral veins slightly visible. Inflorescence c.
<quantity id="4C83F1C8EA39D46E35C5FA977D8C4C52" box="[920,988,1306,1331]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="254" unit="cm" value="18.0">18 cm</quantity>
long, emerging from near leaf base inside leaf sheaths, imbricating, rosette-like, comprised of 7–11, 2–3-flowered cincinni borne on a long peduncle; peduncle
<quantity id="4C83F1C8EA39D46E36D5FAEF7F5E4C1A" box="[136,270,1378,1403]" metricMagnitude="-1" metricUnit="m" metricValue="1.225" metricValueMax="1.45" metricValueMin="1.0" pageId="23" pageNumber="254" unit="cm" value="12.25" valueMax="14.5" valueMin="10.0">10–14.5 cm</quantity>
long, light reddish green to medium reddish green, glabrous, subtended by marcescent leafless sheaths at base; bract 7–11, 1.5–2 ×
<quantity id="4C83F1C8EA39D46E3796FA0B7C4B4CFE" box="[459,539,1414,1439]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="254" unit="cm" value="1.2">1.2 cm</quantity>
, spirally arranged, broadly obovate, coriaceous, glabrous, reddish medium green adaxially, dark purplish red abaxially, turn paler during fruiting, margin entire but slightly undulate at apex, base slightly incurved, apex mucronate, slightly recurved, each bract subtending 2–3 flowers; first bracteole c.
<quantity id="4C83F1C8EA39D46E3342FA437B3D4C86" box="[1311,1389,1486,1511]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="254" unit="mm" value="15.0">15 mm</quantity>
long, opposite bract, keeled, inflexed, reddish white; subsequent bracteoles 1–2, c.
<quantity id="4C83F1C8EA39D46E3258FA7F7A064F6A" box="[1029,1110,1522,1547]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="254" unit="mm" value="12.0">12 mm</quantity>
long, similar but smaller and not keeled; flower c.
<quantity id="4C83F1C8EA39D46E372BF99B7F954F4E" box="[374,453,1558,1583]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="254" unit="mm" value="30.0">30 mm</quantity>
long, floral odour very mild resinous; calyx c.
<quantity id="4C83F1C8EA39D46E358AF99B7A484F4E" box="[983,1048,1558,1583]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="254" unit="mm" value="7.0">7 mm</quantity>
long, tubular, white, glabrous, apex slightly tridentate; floral tube c.
<quantity id="4C83F1C8EA39D46E37B0F9B77C6B4F32" box="[493,571,1594,1619]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="254" unit="mm" value="19.0">19 mm</quantity>
long, white, glabrous; corolla lobes c.
<quantity id="4C83F1C8EA39D46E35B5F9B77A664F32" box="[1000,1078,1594,1619]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="254" unit="mm" value="10.0">10 mm</quantity>
long, lanceolate, white, glabrous, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA39D46E37B3F9D37C354F16" box="[494,613,1630,1655]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="23" pageNumber="254" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
long, linear to slightly oblanceolate, adaxial surface covered with glandular hairs, white, apex obtuse to rounded; labellum 12–15 ×
<quantity id="4C83F1C8EA39D46E3548F90F7D0C4FFA" box="[789,860,1666,1691]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="254" unit="mm" value="9.0">9 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white with violet streaks from base at both sides of light yellow median band, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA39D46E3339F92B7BF74FDE" box="[1380,1447,1702,1727]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="254" unit="mm" value="3.0">3 mm</quantity>
, recurved, lobes sometimes slightly overlapping; stamen c. 6.5 ×
<quantity id="4C83F1C8EA39D46E352CF9477D9C4F82" box="[881,972,1738,1763]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="254" unit="mm" value="1.5">1.5 mm</quantity>
, covered with glandular hairs, white with purplish at crest; filament c.
<quantity id="4C83F1C8EA39D46E3797F9637C5B4E66" box="[458,523,1774,1799]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="254" unit="mm" value="2.0">2 mm</quantity>
long; anther thecae c.
<quantity id="4C83F1C8EA39D46E3554F9637D1A4E66" box="[777,842,1774,1799]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="254" unit="mm" value="3.0">3 mm</quantity>
long, spurred, crest c.
<quantity id="4C83F1C8EA39D46E3217F9637AF04E66" box="[1098,1184,1774,1799]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="254" unit="mm" value="1.5">1.5 mm</quantity>
long, recurved, slightly 3-lobed; stigma less than
<quantity id="4C83F1C8EA39D46E37E8F89F7FA94E4A" box="[437,505,1810,1835]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="254" unit="mm" value="1.0">1 mm</quantity>
long, funnel-shaped, ostiole ciliate, forwards facing; style c.
<quantity id="4C83F1C8EA39D46E329BF89F7B494E4A" box="[1222,1305,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="254" unit="mm" value="24.0">24 mm</quantity>
long, white, glabrous; ovary c.
<quantity id="4C83F1C8EA39D46E370AF8BB7FC84E2E" box="[343,408,1846,1871]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="254" unit="mm" value="2.0">2 mm</quantity>
long, 1-locular, white, glabrous, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA39D46E3283F8BB7B644E2E" box="[1246,1332,1846,1871]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="254" unit="mm" value="3.5">3.5 mm</quantity>
long, free, needle-like, yellowish white. Fruit c.
<quantity id="4C83F1C8EA39D46E3470F8D77C2C4E12" box="[557,636,1882,1907]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="254" unit="mm" value="10.0">10 mm</quantity>
when dried, when mature pericarp splitting halfway into three revealing seeds, calyx persisting prior to maturity, subtended by persistent bracts; seeds c.
<quantity id="4C83F1C8EA39D46E3240F8F37A264EF6" box="[1053,1142,1918,1943]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="23" pageNumber="254" unit="mm" value="8.5">8.5 mm</quantity>
, obovoid, light brown with black dots when mature, aril c.
<quantity id="4C83F1C8EA39D46E37B5F82F7C104EDA" box="[488,576,1954,1979]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="254" unit="mm" value="0.7">0.7 mm</quantity>
, lacerate, perisperm yellowish white.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA39D46036E0F84B7F6F480C" lastPageId="25" lastPageNumber="256" pageId="23" pageNumber="254" type="distribution">
<paragraph id="8BC45C2DEA39D46E36E0F84B7A7B4162" blockId="23.[136,1453,1018,2051]" pageId="23" pageNumber="254">
<emphasis id="B90F803FEA39D46E36E0F84B7FBA4EBE" bold="true" box="[189,490,1990,2015]" pageId="23" pageNumber="254">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA39D46E37ACF84B7D484EBE" box="[497,792,1990,2015]" italics="true" pageId="23" pageNumber="254">
—
<taxonomicName id="4C7B27AEEA39D46E3453F84B7D484EBE" authorityName="Meekiong &amp; Yazid" authorityYear="2015" box="[526,792,1990,2015]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="23" pageNumber="254" phylum="Tracheophyta" rank="species" species="limiana">Scaphochlamys limiana</taxonomicName>
</emphasis>
is known only from north western
<collectingCountry id="F36C1CBDEA39D46E32EBF84B7B464EBE" box="[1206,1302,1990,2015]" name="Malaysia" pageId="23" pageNumber="254">Sarawak</collectingCountry>
occurring in kerangas (tropical heath) forest on quartz-rich soil over granite, between,
<quantity id="4C83F1C8EA39D46E359DF8677A764162" box="[960,1062,2026,2051]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="23" pageNumber="254" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">
20–
<specimenCount id="9D7D97A4EA39D46E35B7F8677A764162" box="[1002,1062,2026,2051]" count="40" pageId="23" pageNumber="254" type="generic">40 m</specimenCount>
</quantity>
.
</paragraph>
<caption id="DF040CA5EA36D46136D5F8257A754148" pageId="24" pageNumber="255" startId="24.[136,229,1960,1981]" targetBox="[175,1415,179,1932]" targetPageId="24" targetType="figure">
<paragraph id="8BC45C2DEA36D46136D5F8257A754148" blockId="24.[136,1452,1960,2089]" pageId="24" pageNumber="255">
<emphasis id="B90F803FEA36D46136D5F8257F594EDC" bold="true" box="[136,265,1960,1981]" pageId="24" pageNumber="255">FIGURE 12.</emphasis>
<emphasis id="B90F803FEA36D4613752F8257FAA4EDC" box="[271,506,1960,1981]" italics="true" pageId="24" pageNumber="255">
<taxonomicName id="4C7B27AEEA36D4613752F8257FA64EDC" authorityName="Meekiong &amp; Yazid" authorityYear="2015" box="[271,502,1960,1981]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="24" pageNumber="255" phylum="Tracheophyta" rank="species" species="limiana">Scaphochlamys limiana</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA36D46137A3F8257C464EDC" bold="true" box="[510,534,1960,1981]" pageId="24" pageNumber="255">A.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA36D46134EBF8257C9C4EDC" bold="true" box="[694,716,1960,1981]" pageId="24" pageNumber="255">B.</emphasis>
plant with flower in habitat.
<emphasis id="B90F803FEA36D46135BCF8257DA94EDC" bold="true" box="[993,1017,1960,1981]" pageId="24" pageNumber="255">C.</emphasis>
inflorescence.
<emphasis id="B90F803FEA36D46132D7F8257AF14EDC" bold="true" box="[1162,1185,1960,1981]" pageId="24" pageNumber="255">D.</emphasis>
bracts, side view.
<emphasis id="B90F803FEA36D461330DF8257B374EDC" bold="true" box="[1360,1383,1960,1981]" pageId="24" pageNumber="255">E.</emphasis>
bracts, top view.
<emphasis id="B90F803FEA36D46136B8F8417EAA4E80" bold="true" box="[229,250,1996,2017]" pageId="24" pageNumber="255">F.</emphasis>
bracts and a mature seed, noted paler bracts during fruiting
<emphasis id="B90F803FEA36D461356CF8417D1A4E80" bold="true" box="[817,842,1996,2017]" pageId="24" pageNumber="255">G.</emphasis>
flower, front view.
<emphasis id="B90F803FEA36D4613258F8417A4E4E80" bold="true" box="[1029,1054,1996,2017]" pageId="24" pageNumber="255">H.</emphasis>
flower, side view.
<emphasis id="B90F803FEA36D461328CF8417AB04E80" bold="true" box="[1233,1248,1996,2017]" pageId="24" pageNumber="255">I.</emphasis>
stamen, noted anther spurs and purplish white crest.
<emphasis id="B90F803FEA36D46137EBF87D7F984164" bold="true" box="[438,456,2032,2053]" pageId="24" pageNumber="255">J.</emphasis>
lamina, abaxial surface.
<emphasis id="B90F803FEA36D46134E6F87D7C844164" bold="true" box="[699,724,2032,2053]" pageId="24" pageNumber="255">K.</emphasis>
roots, note thickened root.
<emphasis id="B90F803FEA36D4613583F87D7DA44164" bold="true" box="[990,1012,2032,2053]" pageId="24" pageNumber="255">L.</emphasis>
bracts and bracteoles.
<emphasis id="B90F803FEA36D461328FF87D7ABE4164" bold="true" box="[1234,1262,2032,2053]" pageId="24" pageNumber="255">M.</emphasis>
rhizome, note dark purplish red interior. A, C–F, I–J, L–M from
<emphasis id="B90F803FEA36D461346BF7997CD34148" box="[566,643,2068,2089]" italics="true" pageId="24" pageNumber="255">OIH118</emphasis>
; B &amp; K from
<emphasis id="B90F803FEA36D4613557F7997D094148" box="[778,857,2068,2089]" italics="true" pageId="24" pageNumber="255">OIH122</emphasis>
; G–H from OIH121.
</paragraph>
</caption>
<paragraph id="8BC45C2DEA37D46036E0FF2D7ACE4860" blockId="25.[136,1452,160,545]" pageId="25" pageNumber="256">
<emphasis id="B90F803FEA37D46036E0FF2D7C1249D8" bold="true" box="[189,578,160,185]" pageId="25" pageNumber="256">Additional specimens examined</emphasis>
:—
<materialsCitation id="3B135670EA37D460343FFF2D7AAC49BC" collectingDate="2014-04-13" collectorName="I. H. Ooi &amp; S. Hyndman &amp; T. Jepom" country="Malaysia" county="Gunung Pueh" latitude="1.8343055" location="Kuching Division" longLatPrecision="1" longitude="109.6765" municipality="Lundu" pageId="25" pageNumber="256" specimenCount="1">
MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA37D46035C2FF2D7A5449D8" box="[927,1028,160,185]" name="Malaysia" pageId="25" pageNumber="256">Sarawak</collectingCountry>
:
<location id="8EA40AF6EA37D4603249FF2D7AB349D8" LSID="urn:lsid:plazi:treatment:03D2ED3BEA39D46036D5FCE77DA84B40:8EA40AF6EA37D4603249FF2D7AB349D8" box="[1044,1251,160,185]" country="Malaysia" county="Gunung Pueh" latitude="1.8343055" longLatPrecision="1" longitude="109.6765" municipality="Lundu" name="Kuching Division" pageId="25" pageNumber="256">Kuching Division</location>
,
<collectingMunicipality id="6BA0C657EA37D46032A9FF2D7B1049D8" box="[1268,1344,160,185]" pageId="25" pageNumber="256">Lundu</collectingMunicipality>
,
<collectingCounty id="62A524A1EA37D460330CFF2D7E9349BC" pageId="25" pageNumber="256">Gunung Pueh</collectingCounty>
,
<geoCoordinate id="EE4F3AEAEA37D4603690FF497F3E49BC" box="[205,366,196,221]" degrees="01" direction="north" minutes="50" orientation="latitude" pageId="25" pageNumber="256" precision="1" seconds="03.5" value="1.8343055">01°50’03.5”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA37D4603725FF497C7449BC" box="[376,548,196,221]" degrees="109" direction="east" minutes="40" orientation="longitude" pageId="25" pageNumber="256" precision="1" seconds="35.4" value="109.6765">109°40’35.4”E</geoCoordinate>
,
<date id="FFC57AEDEA37D4603473FF497C9B49BC" box="[558,715,196,221]" pageId="25" pageNumber="256" value="2014-04-13">
<collectingDate id="EF818305EA37D4603473FF497C9B49BC" box="[558,715,196,221]" pageId="25" pageNumber="256" value="2014-04-13">13 April 2014</collectingDate>
</date>
,
<emphasis id="B90F803FEA37D4603489FF497AF949BC" box="[724,1193,196,221]" italics="true" pageId="25" pageNumber="256">
<collectorName id="268E39FBEA37D4603489FF497D6249BC" box="[724,818,196,221]" pageId="25" pageNumber="256">I.H. Ooi</collectorName>
,
<collectorName id="268E39FBEA37D4603560FF497D9549BC" box="[829,965,196,221]" pageId="25" pageNumber="256">S. Hyndman</collectorName>
&amp;
<collectorName id="268E39FBEA37D46035BBFF497A1A49BC" box="[998,1098,196,221]" pageId="25" pageNumber="256">T. Jepom</collectorName>
OIH118
</emphasis>
(SAR)
</materialsCitation>
;
<geoCoordinate id="EE4F3AEAEA37D460335BFF497BF749BC" box="[1286,1447,196,221]" degrees="01" direction="north" minutes="49" orientation="latitude" pageId="25" pageNumber="256" precision="1" seconds="59.1" value="1.8330833">01°49’59.1”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA37D46036D5FF657F644860" box="[136,308,232,257]" degrees="109" direction="east" minutes="40" orientation="longitude" pageId="25" pageNumber="256" precision="1" seconds="27.2" value="109.67422">109°40’27.2”E</geoCoordinate>
,
<date id="FFC57AEDEA37D4603762FF657F8E4860" box="[319,478,232,257]" pageId="25" pageNumber="256" value="2014-04-13">13 April 2014</date>
,
<emphasis id="B90F803FEA37D46037B4FF657A164860" box="[489,1094,232,257]" italics="true" pageId="25" pageNumber="256">I.H. Ooi, S. Hyndman &amp; T. Jepom OIH121 &amp; OIH122</emphasis>
(SAR).
</paragraph>
<paragraph id="8BC45C2DEA37D46036E0FE817F6F480C" blockId="25.[136,1452,160,545]" pageId="25" pageNumber="256">
<emphasis id="B90F803FEA37D46036E0FE817FFE4844" bold="true" box="[189,430,268,293]" pageId="25" pageNumber="256">Conservation status</emphasis>
:—Near Threatened (NT). This species occurs within a protected area and its extension, which is not yet easily accessible. The populations near the boundary of the protected area are close to the adjacent oil palm plantation.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA37D46036E0FEF57DA84B40" pageId="25" pageNumber="256" type="discussion">
<paragraph id="8BC45C2DEA37D46036E0FEF57DA84B40" blockId="25.[136,1452,160,545]" pageId="25" pageNumber="256">
<emphasis id="B90F803FEA37D46036E0FEF57F1548F0" bold="true" box="[189,325,376,401]" pageId="25" pageNumber="256">Discussion:</emphasis>
—This species is unique among the Bornean
<taxonomicName id="4C7B27AEEA37D4603563FEF57DBC48F0" authorityName="Baker" authorityYear="1892" box="[830,1004,376,401]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="25" pageNumber="256" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F803FEA37D4603563FEF57DBC48F0" box="[830,1004,376,401]" italics="true" pageId="25" pageNumber="256">Scaphochlamys</emphasis>
</taxonomicName>
. The dark purplish red rhizome interior, lamina abaxially intensely purplish red, and by the long pedunculate rosette-like imbricating inflorescences are only known for species from Peninsular
<collectingCountry id="F36C1CBDEA37D4603444FE4D7CD248B8" box="[537,642,448,473]" name="Malaysia" pageId="25" pageNumber="256">Malaysia</collectingCountry>
, such as
<taxonomicName id="4C7B27AEEA37D46034B5FE4D7AC348B8" authority="Sam &amp; Saw (2005: 253)" authorityName="Sam &amp; Saw" authorityPageNumber="253" authorityYear="2005" box="[744,1171,448,473]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="25" pageNumber="256" phylum="Tracheophyta" rank="species" species="abdullahii">
<emphasis id="B90F803FEA37D46034B5FE4D7D2848B8" box="[744,888,448,473]" italics="true" pageId="25" pageNumber="256">S. abdullahii</emphasis>
<bibRefCitation id="EFEA21DCEA37D46035DDFE4D7AC348B8" author="Sam, Y. Y. &amp; Saw, L. G." box="[896,1171,448,473]" pageId="25" pageNumber="256" pagination="253 - 261" refId="ref33005" refString="Sam, Y. Y. &amp; Saw, L. G. (2005) Three new species of Scaphochlamys (Zingiberaceae) from Peninsular Malaysia. Gardens' Bulletin Singapore 57: 253 - 261." type="journal article" year="2005">Sam &amp; Saw (2005: 253)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B27AEEA37D4603296FE4D7FB4489C" authority="(Ridley 1893: 378) Holttum (1950: 91)" authorityName="Holttum" authorityPageNumber="91" authorityYear="1950" baseAuthorityName="Ridley" baseAuthorityPageNumber="378" baseAuthorityYear="1893" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="25" pageNumber="256" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="B90F803FEA37D4603296FE4D7B1F48B8" box="[1227,1359,448,473]" italics="true" pageId="25" pageNumber="256">S. sylvestris</emphasis>
(
<bibRefCitation id="EFEA21DCEA37D4603302FE4D7EAA489C" author="Ridley, H. N." pageId="25" pageNumber="256" pagination="267 - 408" refId="ref32788" refString="Ridley, H. N. (1893) On the flora of the Eastern coast of the Malay Peninsula. Transactions of the Linnean Society of London. ser. II 3: 267 - 408. https: // doi. org / 10.1111 / j. 1095 - 8339.1893. tb 00678. x" type="journal article" year="1893">Ridley 1893: 378</bibRefCitation>
)
<bibRefCitation id="EFEA21DCEA37D4603754FE697FB4489C" author="Holttum, R. E." box="[265,484,484,509]" pageId="25" pageNumber="256" pagination="1 - 249" refId="ref32053" refString="Holttum, R. E. (1950) The Zingiberaceae of the Malay Peninsula. Gardens' Bulletin Singapore 13: 1 - 249." type="journal article" year="1950">Holttum (1950: 91)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFEA21DCEA37D46037AFFE697CEC489C" author="Sam, Y. Y. &amp; Saw, L. G." box="[498,700,484,509]" pageId="25" pageNumber="256" pagination="253 - 261" refId="ref33005" refString="Sam, Y. Y. &amp; Saw, L. G. (2005) Three new species of Scaphochlamys (Zingiberaceae) from Peninsular Malaysia. Gardens' Bulletin Singapore 57: 253 - 261." type="journal article" year="2005">Sam &amp; Saw 2005</bibRefCitation>
;
<bibRefCitation id="EFEA21DCEA37D4603495FE697D02489C" author="Searle, R. J." box="[712,850,484,509]" pageId="25" pageNumber="256" pagination="75 - 121" refId="ref33212" refString="Searle, R. J. (2010) The genus Scaphochlamys (Zingiberaceae - Zingibereae): A Compendium for the field worker. Edinburgh Journal of Botany 67: 75 - 121. https: // doi. org / 10.1017 / S 0960428609990254" type="journal article" year="2010">Searle 2010</bibRefCitation>
).
<taxonomicName id="4C7B27AEEA37D4603539FE697A20489C" authorityName="Meekiong &amp; Yazid" authorityYear="2015" box="[868,1136,484,509]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="25" pageNumber="256" phylum="Tracheophyta" rank="species" species="limiana">
<emphasis id="B90F803FEA37D4603539FE697A20489C" box="[868,1136,484,509]" italics="true" pageId="25" pageNumber="256">Scaphochlamys limiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B27AEEA37D46032FBFE697B68489C" box="[1190,1336,484,509]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="25" pageNumber="256" phylum="Tracheophyta" rank="species" species="polyphylla">
<emphasis id="B90F803FEA37D46032FBFE697B68489C" box="[1190,1336,484,509]" italics="true" pageId="25" pageNumber="256">S. polyphylla</emphasis>
</taxonomicName>
are closer morphologically to species from Peninsular
<collectingCountry id="F36C1CBDEA37D4603425FD857C8F4B40" box="[632,735,520,545]" name="Malaysia" pageId="25" pageNumber="256">Malaysia</collectingCountry>
than to Bornean species.
</paragraph>
</subSubSection>
</treatment>
</document>

338
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@ -0,0 +1,338 @@
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<paragraph id="8BC45C2DEA3BD46C36D5FCC17BFC4A04" blockId="21.[136,1452,842,1011]" box="[136,1452,842,869]" pageId="21" pageNumber="252">
<heading id="D08CEB41EA3BD46C36D5FCC17BFC4A04" box="[136,1452,842,869]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="252" reason="4">
<emphasis id="B90F803FEA3BD46C36D5FCC17EF94A04" bold="true" box="[136,169,844,869]" pageId="21" pageNumber="252">11.</emphasis>
<taxonomicName id="4C7B27AEEA3BD46C36E8FCC77D6E4A04" ID-CoL="8X5G3" authority="Ooi, K. Meekiong &amp; S.Y.Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[181,830,842,869]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="lucens" status="sp. nov.">
<emphasis id="B90F803FEA3BD46C36E8FCC77F904A04" bold="true" box="[181,448,842,869]" italics="true" pageId="21" pageNumber="252">Scaphochlamys lucens</emphasis>
Ooi, K. Meekiong &amp; S.Y.Wong
</taxonomicName>
,
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<taxonomicNameLabel id="A23C3D44EA3BD46C3513FCC17DF74A04" box="[846,935,844,869]" pageId="21" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<typeStatus id="54C0E28FEA3BD46C35EFFCC17DBD4A04" box="[946,1005,844,869]" pageId="21" pageNumber="252">Type</typeStatus>
</emphasis>
:
<emphasis id="B90F803FEA3BD46C35A9FCC17A5D4A04" box="[1012,1037,844,869]" italics="true" pageId="21" pageNumber="252">—</emphasis>
MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA3BD46C331EFCC17BF84A04" box="[1347,1448,844,869]" name="Malaysia" pageId="21" pageNumber="252">Sarawak</collectingCountry>
:
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA3BD46C36D5FCFD7C6C4ACC" pageId="21" pageNumber="252" type="description">
<paragraph id="8BC45C2DEA3BD46C36D5FCFD7BFC4AE8" blockId="21.[136,1452,842,1011]" box="[136,1452,880,905]" pageId="21" pageNumber="252">
Kuching Division, Padawan, Gunung Temugan,
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,
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,
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,
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,
<emphasis id="B90F803FEA3BD46C3350FCFD7BFC4AE8" box="[1293,1452,880,905]" italics="true" pageId="21" pageNumber="252">I.H. Ooi &amp; K.</emphasis>
</paragraph>
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<emphasis id="B90F803FEA3BD46C36D5FC197F664ACC" box="[136,310,916,941]" italics="true" pageId="21" pageNumber="252">Jeland OIH125</emphasis>
(holo SAR),
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<emphasis id="B90F803FEA3BD46C3791FC197C6C4ACC" bold="true" box="[460,572,916,941]" pageId="21" pageNumber="252">Figure 11</emphasis>
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</paragraph>
</subSubSection>
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<emphasis id="B90F803FEA3BD46C36D5FC377EBB4AAE" bold="true" box="[136,235,954,975]" pageId="21" pageNumber="252">Diagnosis</emphasis>
:
<emphasis id="B90F803FEA3BD46C36ACFC377F564AAE" box="[241,262,954,975]" italics="true" pageId="21" pageNumber="252">—</emphasis>
The flower of
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<emphasis id="B90F803FEA3BD46C37CCFC377FB64AAE" box="[401,486,954,975]" italics="true" pageId="21" pageNumber="252">S. lucens</emphasis>
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is similar to
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<emphasis id="B90F803FEA3BD46C3438FC377CB64AAE" box="[613,742,954,975]" italics="true" pageId="21" pageNumber="252">S. graveolens</emphasis>
</taxonomicName>
, but
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<emphasis id="B90F803FEA3BD46C354AFC377D3C4AAE" box="[791,876,954,975]" italics="true" pageId="21" pageNumber="252">S. lucens</emphasis>
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is much smaller, up to
<quantity id="4C83F1C8EA3BD46C3210FC377AD84AAE" box="[1101,1160,954,975]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="252" unit="cm" value="25.0">25 cm</quantity>
tall (vs. up to 70), leaf lamina shorter,
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long (vs.
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long), and adaxially glossy green (vs. matte dull green).
</paragraph>
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Terrestrial, perennial, rhizomatous herbs, to c.
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tall. Rhizome creeping on ground, c.
<quantity id="4C83F1C8EA3BD46C32D9FBA97A954D5C" box="[1156,1221,1060,1085]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="252" unit="mm" value="5.0">5 mm</quantity>
diameter, externally light brown to brownish green, internally white. Elements
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apart, 1-foliate, juvenile up to 4-foliate; petiole
<quantity id="4C83F1C8EA3BD46C36D5FBE17EA94DE4" box="[136,249,1132,1157]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="21" pageNumber="252" unit="cm" value="13.0" valueMax="16.0" valueMin="10.0">10−16 cm</quantity>
long, canaliculate, medium green, pubescent, base pulvinate, white to yellowish light green; leafless sheath c. 3,
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long, light to medium green, glabrous, decaying with age; leaf sheath c.
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long, membranous, light green, glabrous, decaying with age; ligule obscure, membranous; lamina 9–12 ×
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, broadly ovate to broadly elliptic, coriaceous, margin entire, base obtuse, apex acute, often recurved; adaxial surface glossy, medium green to slightly dark green, glabrous, midrib sunken, medium green to slightly dark green, glabrous, main lateral veins slightly raised; abaxial surface whitish green, sparsely pubescent, midrib raised, medium green, densely pubescent, main lateral veins slightly visible. Inflorescence c.
<quantity id="4C83F1C8EA3BD46C3465FAC97C3F4C3C" box="[568,623,1348,1373]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="252" unit="cm" value="7.0">7 cm</quantity>
long, emerging from near leaf base inside leaf sheaths, tightly congested, bulb-like, comprised of c. 4, 2-flowered cincinni borne on a peduncle, flowering spirally from base to apex; peduncle
<quantity id="4C83F1C8EA3BD46C36D5FA017EA14CC4" box="[136,241,1420,1445]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="21" pageNumber="252" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5−2 cm</quantity>
long, greenish white, glabrous, exposed, slightly covered by the remaining decayed leafless sheaths; bract c. 4, c. 3.5 ×
<quantity id="4C83F1C8EA3BD46C36A4FA3D7F624CA8" box="[249,306,1456,1481]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="252" unit="cm" value="1.0">1 cm</quantity>
, spirally arranged, lanceolate, membranous, light green, sparsely pubescent, apex acuminate, each bract subtending c. 2 flowers; bracteole c. 2,
<quantity id="4C83F1C8EA3BD46C3460FA597CE54C8C" box="[573,693,1492,1517]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="21" pageNumber="252" unit="mm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 mm</quantity>
long, shorter than bracts but barely distinguishable from bract, light green; flower
<quantity id="4C83F1C8EA3BD46C3775FA757FC34F70" box="[296,403,1528,1553]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="21" pageNumber="252" unit="cm" value="6.75" valueMax="7.0" valueMin="6.5">6.5−7 cm</quantity>
long, floral odour reminiscent of liquid seeping from domestic waste; calyx c.
<quantity id="4C83F1C8EA3BD46C3346FA757B3B4F70" box="[1307,1387,1528,1553]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="252" unit="mm" value="18.0">18 mm</quantity>
long, brownish white, sparsely pubescent, apex acute; floral tube
<quantity id="4C83F1C8EA3BD46C357FF9917DCD4F54" box="[802,925,1564,1589]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="21" pageNumber="252" unit="mm" value="47.5" valueMax="50.0" valueMin="45.0">45−50 mm</quantity>
long, white, glabrous; corolla lobes
<quantity id="4C83F1C8EA3BD46C336EF9917BFC4F54" box="[1331,1452,1564,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="21" pageNumber="252" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">17–18 mm</quantity>
long, lanceolate, white to slightly yellowish white, glabrous, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA3BD46C32CEF9CD7B5B4F38" box="[1171,1291,1600,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="21" pageNumber="252" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">13–18 mm</quantity>
long, linear to slightly oblanceolate, white to light yellow, apex acute, adaxial surface covered with glandular hairs; labellum 20–25 ×
<quantity id="4C83F1C8EA3BD46C36D5F9057F544FC0" box="[136,260,1672,1697]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="21" pageNumber="252" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10−12 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white with violet along light yellow median band extending to lobes, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA3BD46C34D8F9217C984FA4" box="[645,712,1708,1733]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="252" unit="mm" value="5.0">5 mm</quantity>
, recurved, lobes sometimes slightly overlapping; stamen c. 11 ×
<quantity id="4C83F1C8EA3BD46C33C3F9217EE64F88" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="252" unit="mm" value="2.0">2 mm</quantity>
, covered with glandular hairs, white; filament c.
<quantity id="4C83F1C8EA3BD46C34BDF95D7D714F88" box="[736,801,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="252" unit="mm" value="5.0">5 mm</quantity>
long; anther thecae c.
<quantity id="4C83F1C8EA3BD46C327CF95D7A334F88" box="[1057,1123,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="252" unit="mm" value="5.0">5 mm</quantity>
long, spurless, crest c.
<quantity id="4C83F1C8EA3BD46C3337F95D7BFC4F88" box="[1386,1452,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="252" unit="mm" value="1.0">1 mm</quantity>
long; stigma less than
<quantity id="4C83F1C8EA3BD46C3722F9797FEF4E6C" box="[383,447,1780,1805]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="252" unit="mm" value="1.0">1 mm</quantity>
long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA3BD46C3342F9797B3D4E6C" box="[1311,1389,1780,1805]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="252" unit="mm" value="45.0">45 mm</quantity>
long, white, glabrous; ovary c.
<quantity id="4C83F1C8EA3BD46C37F8F8957FB64E50" box="[421,486,1816,1841]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="252" unit="mm" value="3.0">3 mm</quantity>
long, 1-locular, white, densely pubescent, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA3BD46C33C0F8957EE44E34" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="252" unit="mm" value="3.0">3 mm</quantity>
long, free, needle-like, yellowish white. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA3BD46C36E0F8ED7C5B4EFC" pageId="21" pageNumber="252" type="etymology">
<paragraph id="8BC45C2DEA3BD46C36E0F8ED7C5B4EFC" blockId="21.[136,1452,1060,2021]" pageId="21" pageNumber="252">
<emphasis id="B90F803FEA3BD46C36E0F8ED7F6E4E18" bold="true" box="[189,318,1888,1913]" pageId="21" pageNumber="252">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C7B27AEEA3BD46C3416F8ED7CC34E18" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[587,659,1888,1913]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="lucens">
<emphasis id="B90F803FEA3BD46C3416F8ED7CC34E18" box="[587,659,1888,1913]" italics="true" pageId="21" pageNumber="252">lucens</emphasis>
</taxonomicName>
, is derived of the Latin verb meaning “shining”, indicates the adaxially glossy light-reflecting leaf lamina.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA3BD46E36E0F8257ACB48F0" lastPageId="23" lastPageNumber="254" pageId="21" pageNumber="252" type="distribution">
<paragraph id="8BC45C2DEA3BD46C36E0F8257D914E84" blockId="21.[136,1452,1060,2021]" pageId="21" pageNumber="252">
<emphasis id="B90F803FEA3BD46C36E0F8257FB04EA0" bold="true" box="[189,480,1960,1985]" pageId="21" pageNumber="252">Distribution and ecology</emphasis>
:—
<taxonomicName id="4C7B27AEEA3BD46C345EF8257CAF4EA0" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[515,767,1960,1985]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="lucens">
<emphasis id="B90F803FEA3BD46C345EF8257CAF4EA0" box="[515,767,1960,1985]" italics="true" pageId="21" pageNumber="252">Scaphochlamys lucens</emphasis>
</taxonomicName>
is known from the
<typeStatus id="54C0E28FEA3BD46C3592F8257DAF4EA0" box="[975,1023,1960,1985]" pageId="21" pageNumber="252">type</typeStatus>
locality and the surrounding limestone areas occurring in deep litter humus pockets, between
<quantity id="4C83F1C8EA3BD46C34B4F8417D1C4E84" box="[745,844,1996,2021]" metricMagnitude="1" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="21" pageNumber="252" unit="m" value="47.5" valueMax="60.0" valueMin="35.0">35–60 m</quantity>
elevation.
</paragraph>
<caption id="DF040CA5EA38D46F36D5F8417A694148" pageId="22" pageNumber="253" startId="22.[136,229,1996,2017]" targetBox="[161,1427,181,1967]" targetPageId="22" targetType="figure">
<paragraph id="8BC45C2DEA38D46F36D5F8417A694148" blockId="22.[136,1452,1996,2089]" pageId="22" pageNumber="253">
<emphasis id="B90F803FEA38D46F36D5F8417F5B4E80" bold="true" box="[136,267,1996,2017]" pageId="22" pageNumber="253">FIGURE 11.</emphasis>
<taxonomicName id="4C7B27AEEA38D46F374EF8417FBE4E80" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[275,494,1996,2017]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="22" pageNumber="253" phylum="Tracheophyta" rank="species" species="lucens">
<emphasis id="B90F803FEA38D46F374EF8417FBE4E80" box="[275,494,1996,2017]" italics="true" pageId="22" pageNumber="253">Scaphochlamys lucens</emphasis>
</taxonomicName>
.
<emphasis id="B90F803FEA38D46F37A0F8417C454E80" bold="true" box="[509,533,1996,2017]" pageId="22" pageNumber="253">A.</emphasis>
Type,
<emphasis id="B90F803FEA38D46F3406F8417CFA4E80" box="[603,682,1996,2017]" italics="true" pageId="22" pageNumber="253">OIH125</emphasis>
(SAR).
<emphasis id="B90F803FEA38D46F34A2F8417D464E80" bold="true" box="[767,790,1996,2017]" pageId="22" pageNumber="253">B.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA38D46F359FF8417D894E80" bold="true" box="[962,985,1996,2017]" pageId="22" pageNumber="253">C.</emphasis>
petiole base and rhizome.
<emphasis id="B90F803FEA38D46F32BAF8417AAF4E80" bold="true" box="[1255,1279,1996,2017]" pageId="22" pageNumber="253">D.</emphasis>
inflorescence.
<emphasis id="B90F803FEA38D46F33C8F8417BFC4E80" bold="true" box="[1429,1452,1996,2017]" pageId="22" pageNumber="253">E.</emphasis>
lamina, adaxial surface.
<emphasis id="B90F803FEA38D46F372CF87D7FD64164" bold="true" box="[369,390,2032,2053]" pageId="22" pageNumber="253">F.</emphasis>
lamina, abaxial surface.
<emphasis id="B90F803FEA38D46F342FF87D7CDB4164" bold="true" box="[626,651,2032,2053]" pageId="22" pageNumber="253">G.</emphasis>
bract and bracteoles.
<emphasis id="B90F803FEA38D46F3506F87D7D244164" bold="true" box="[859,884,2032,2053]" pageId="22" pageNumber="253">H.</emphasis>
flower, side view.
<emphasis id="B90F803FEA38D46F3275F87D7A674164" bold="true" box="[1064,1079,2032,2053]" pageId="22" pageNumber="253">I.</emphasis>
flower, front view.
<emphasis id="B90F803FEA38D46F32AEF87D7B554164" bold="true" box="[1267,1285,2032,2053]" pageId="22" pageNumber="253">J.</emphasis>
flower, top view.
<emphasis id="B90F803FEA38D46F36D5F7997EF14148" bold="true" box="[136,161,2068,2089]" pageId="22" pageNumber="253">K.</emphasis>
calyx and ovary.
<emphasis id="B90F803FEA38D46F3717F7997F314148" bold="true" box="[330,353,2068,2089]" pageId="22" pageNumber="253">L.</emphasis>
stamen, note spurless anther.
<emphasis id="B90F803FEA38D46F34DDF7997CCC4148" bold="true" box="[640,668,2068,2089]" pageId="22" pageNumber="253">M.</emphasis>
anther crest and stigma. All from
<emphasis id="B90F803FEA38D46F35B9F7997A634148" box="[996,1075,2068,2089]" italics="true" pageId="22" pageNumber="253">OIH125</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BC45C2DEA39D46E36E0FF2D7A2B4844" blockId="23.[136,1452,160,509]" pageId="23" pageNumber="254">
<materialsCitation id="3B135670EA39D46E36E0FF2D7B1549BC" collectingDate="2014-10-05" collectorName="P. C. Boyce &amp; S. Y. Wong" country="Malaysia" county="Gua Tut" latitude="1.2853333" location="Kuching Division" longLatPrecision="1" longitude="110.25244" municipality="Padawan" pageId="23" pageNumber="254" specimenCount="2">
<emphasis id="B90F803FEA39D46E36E0FF2D7CBC49D8" bold="true" box="[189,748,160,185]" pageId="23" pageNumber="254">
<typeStatus id="54C0E28FEA39D46E36E0FF2D7C0049D8" box="[189,592,160,185]" isOtherTypeStatus="true" pageId="23" pageNumber="254">Additional specimens examined</typeStatus>
(
<typeStatus id="54C0E28FEA39D46E3437FF2D7CB449D8" box="[618,740,160,185]" pageId="23" pageNumber="254" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA39D46E3208FF2D7AEA49D8" box="[1109,1210,160,185]" name="Malaysia" pageId="23" pageNumber="254">Sarawak</collectingCountry>
:
<location id="8EA40AF6EA39D46E328DFF2D7BF749D8" LSID="urn:lsid:plazi:treatment:03D2ED3BEA3BD46E36D5FCC17C02489C:8EA40AF6EA39D46E328DFF2D7BF749D8" box="[1232,1447,160,185]" country="Malaysia" county="Gua Tut" latitude="1.2853333" longLatPrecision="1" longitude="110.25244" municipality="Padawan" name="Kuching Division" pageId="23" pageNumber="254">Kuching Division</location>
,
<collectingMunicipality id="6BA0C657EA39D46E36D5FF497EA049BC" box="[136,240,196,221]" pageId="23" pageNumber="254">Padawan</collectingMunicipality>
,
<collectingCounty id="62A524A1EA39D46E36A1FF497F0749BC" box="[252,343,196,221]" pageId="23" pageNumber="254">Gua Tut</collectingCounty>
<geoCoordinate id="EE4F3AEAEA39D46E3702FF497FAF49BC" box="[351,511,196,221]" degrees="01" direction="north" minutes="17" orientation="latitude" pageId="23" pageNumber="254" precision="1" seconds="07.2" value="1.2853333">01°17’07.2”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA39D46E3456FF497CE649BC" box="[523,694,196,221]" degrees="110" direction="east" minutes="15" orientation="longitude" pageId="23" pageNumber="254" precision="1" seconds="08.8" value="110.25244">110°15’08.8”E</geoCoordinate>
,
<date id="FFC57AEDEA39D46E349FFF497D2449BC" box="[706,884,196,221]" pageId="23" pageNumber="254" value="2014-10-05">
<collectingDate id="EF818305EA39D46E349FFF497D2449BC" box="[706,884,196,221]" pageId="23" pageNumber="254" value="2014-10-05">5 October 2014</collectingDate>
</date>
,
<emphasis id="B90F803FEA39D46E35DDFF497AA149BC" box="[896,1265,196,221]" italics="true" pageId="23" pageNumber="254">
<collectorName id="268E39FBEA39D46E35DDFF497DA949BC" box="[896,1017,196,221]" pageId="23" pageNumber="254">P.C. Boyce</collectorName>
&amp;
<collectorName id="268E39FBEA39D46E3242FF497ADE49BC" box="[1055,1166,196,221]" pageId="23" pageNumber="254">S.Y. Wong</collectorName>
OIH263
</emphasis>
(SAR)
</materialsCitation>
;
<materialsCitation id="3B135670EA39D46E330CFF497A9B4860" collectingDate="2014-04-14" collectorName="I. H. Ooi &amp; K. Jeland" country="Malaysia" latitude="1.2607223" location="Gunung Temugan" longLatPrecision="1" longitude="110.25872" pageId="23" pageNumber="254" specimenCount="1">
<location id="8EA40AF6EA39D46E330CFF497EA34860" LSID="urn:lsid:plazi:treatment:03D2ED3BEA3BD46E36D5FCC17C02489C:8EA40AF6EA39D46E330CFF497EA34860" country="Malaysia" latitude="1.2607223" longLatPrecision="1" longitude="110.25872" name="Gunung Temugan" pageId="23" pageNumber="254">Gunung Temugan</location>
,
<geoCoordinate id="EE4F3AEAEA39D46E375DFF657FF14860" box="[256,417,232,257]" degrees="01" direction="north" minutes="15" orientation="latitude" pageId="23" pageNumber="254" precision="1" seconds="38.6" value="1.2607223">01°15’38.6”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA39D46E37F2FF657C0A4860" box="[431,602,232,257]" degrees="110" direction="east" minutes="15" orientation="longitude" pageId="23" pageNumber="254" precision="1" seconds="31.4" value="110.25872">110°15’31.4”E</geoCoordinate>
,
<date id="FFC57AEDEA39D46E3435FF657D5C4860" box="[616,780,232,257]" pageId="23" pageNumber="254" value="2014-04-14">
<collectingDate id="EF818305EA39D46E3435FF657D5C4860" box="[616,780,232,257]" pageId="23" pageNumber="254" value="2014-04-14">14 April 2014</collectingDate>
</date>
,
<emphasis id="B90F803FEA39D46E3544FF657A254860" box="[793,1141,232,257]" italics="true" pageId="23" pageNumber="254">
<collectorName id="268E39FBEA39D46E3544FF657D294860" box="[793,889,232,257]" pageId="23" pageNumber="254">I.H. Ooi</collectorName>
&amp;
<collectorName id="268E39FBEA39D46E35FFFF657A404860" box="[930,1040,232,257]" pageId="23" pageNumber="254">K. Jeland</collectorName>
OIH126
</emphasis>
(SAR)
</materialsCitation>
;
<materialsCitation id="3B135670EA39D46E3284FF657A274844" collectingDate="2014-10-21" collectorName="S. Y. Wong &amp; P. C. Boyce" country="Malaysia" latitude="1.2597222" location="Gunung Temugan" longLatPrecision="1" longitude="110.29381" pageId="23" pageNumber="254" specimenCount="1">
<location id="8EA40AF6EA39D46E3284FF657BF84860" LSID="urn:lsid:plazi:treatment:03D2ED3BEA3BD46E36D5FCC17C02489C:8EA40AF6EA39D46E3284FF657BF84860" box="[1241,1448,232,257]" country="Malaysia" latitude="1.2597222" longLatPrecision="1" longitude="110.29381" name="Gunung Temugan" pageId="23" pageNumber="254">Gunung Temugan</location>
,
<geoCoordinate id="EE4F3AEAEA39D46E36D5FE817F794844" box="[136,297,268,293]" degrees="01" direction="north" minutes="15" orientation="latitude" pageId="23" pageNumber="254" precision="1" seconds="35.0" value="1.2597222">01°15’35.0”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA39D46E3769FE817F8F4844" box="[308,479,268,293]" degrees="110" direction="east" minutes="17" orientation="longitude" pageId="23" pageNumber="254" precision="1" seconds="37.7" value="110.29381">110°17’37.7”E</geoCoordinate>
,
<date id="FFC57AEDEA39D46E37B7FE817CF94844" box="[490,681,268,293]" pageId="23" pageNumber="254" value="2014-10-21">
<collectingDate id="EF818305EA39D46E37B7FE817CF94844" box="[490,681,268,293]" pageId="23" pageNumber="254" value="2014-10-21">21 October 2014</collectingDate>
</date>
,
<emphasis id="B90F803FEA39D46E34E9FE817A734844" box="[692,1059,268,293]" italics="true" pageId="23" pageNumber="254">
<collectorName id="268E39FBEA39D46E34E9FE817D734844" box="[692,803,268,293]" pageId="23" pageNumber="254">S.Y. Wong</collectorName>
&amp;
<collectorName id="268E39FBEA39D46E351AFE817D904844" box="[839,960,268,293]" pageId="23" pageNumber="254">P.C. Boyce</collectorName>
OIH264
</emphasis>
(SAR)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC45C2DEA39D46E36E0FEBD7ACB48F0" blockId="23.[136,1452,160,509]" pageId="23" pageNumber="254">
<emphasis id="B90F803FEA39D46E36E0FEBD7FF84828" bold="true" box="[189,424,304,329]" pageId="23" pageNumber="254">Conservation status</emphasis>
:—Near Threatened (NT).The populations of the species are quite large (200–300 individuals) at the localities. Some of the surrounding areas are popular for ecotourism activities; however, some areas are subjected to quarrying activities. Therefore, the listing is justified under the NT category (
<bibRefCitation id="EFEA21DCEA39D46E3258FEF57ADF48F0" author="IUCN" box="[1029,1167,376,401]" pageId="23" pageNumber="254" refId="ref32181" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. Second Edition, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA39D46E36E0FE117C02489C" pageId="23" pageNumber="254" type="discussion">
<paragraph id="8BC45C2DEA39D46E36E0FE117C02489C" blockId="23.[136,1452,160,509]" pageId="23" pageNumber="254">
<emphasis id="B90F803FEA39D46E36E0FE117F6B48D4" bold="true" box="[189,315,412,437]" pageId="23" pageNumber="254">Discussion</emphasis>
:—
<taxonomicName id="4C7B27AEEA39D46E3702FE117C0948D4" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[351,601,412,437]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="23" pageNumber="254" phylum="Tracheophyta" rank="species" species="lucens">
<emphasis id="B90F803FEA39D46E3702FE117C0948D4" box="[351,601,412,437]" italics="true" pageId="23" pageNumber="254">Scaphochlamys lucens</emphasis>
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appears related to
<taxonomicName id="4C7B27AEEA39D46E357CFE117DE548D4" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[801,949,412,437]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="23" pageNumber="254" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="B90F803FEA39D46E357CFE117DE548D4" box="[801,949,412,437]" italics="true" pageId="23" pageNumber="254">S. graveolens</emphasis>
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by sharing several remarkable characteristics, especially the tightly congested bulb-like inflorescence, white labellum with violet along light yellow median band extending to lobes, and foul floral odour.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="8BC45C2DEA3DD46A36D5FC717BFC4D74" blockId="19.[136,1452,1018,1188]" box="[136,1452,1018,1045]" pageId="19" pageNumber="250">
<heading id="D08CEB41EA3DD46A36D5FC717BFC4D74" box="[136,1452,1018,1045]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="250" reason="4">
<emphasis id="B90F803FEA3DD46A36D5FC717EFB4D74" bold="true" box="[136,171,1020,1045]" pageId="19" pageNumber="250">10.</emphasis>
<taxonomicName id="4C7B27AEEA3DD46A36ECFC777D044D74" ID-CoL="8X5FX" authority="Ooi, K. Meekiong &amp; S.Y. Wong" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[177,852,1018,1045]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="19" pageNumber="250" phylum="Tracheophyta" rank="species" species="graveolens" status="sp. nov.">
<emphasis id="B90F803FEA3DD46A36ECFC777FB84D74" bold="true" box="[177,488,1018,1045]" italics="true" pageId="19" pageNumber="250">Scaphochlamys graveolens</emphasis>
Ooi, K. Meekiong &amp; S.Y. Wong
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,
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<taxonomicNameLabel id="A23C3D44EA3DD46A3502FC717DE34D74" box="[863,947,1020,1045]" pageId="19" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B90F803FEA3DD46A35E4FC717DA44D74" bold="true" box="[953,1012,1020,1045]" pageId="19" pageNumber="250">
<typeStatus id="54C0E28FEA3DD46A35E4FC717DA44D74" box="[953,1012,1020,1045]" pageId="19" pageNumber="250">Type</typeStatus>
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:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA3DD46A331EFC717BF84D74" box="[1347,1448,1020,1045]" name="Malaysia" pageId="19" pageNumber="250">Sarawak</collectingCountry>
:
</heading>
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<subSubSection id="C3610FA6EA3DD46A36D5FBAD7FDA4D3C" pageId="19" pageNumber="250" type="description">
<paragraph id="8BC45C2DEA3DD46A36D5FBAD7BFB4D58" blockId="19.[136,1452,1018,1188]" box="[136,1451,1056,1081]" pageId="19" pageNumber="250">
Samarahan Division, Serian, Pichin,
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,
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,
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,
<date id="FFC57AEDEA3DD46A3597FBAD7A074D58" box="[970,1111,1056,1081]" pageId="19" pageNumber="250" value="2014-04-06">06 Apr 2014</date>
,
<emphasis id="B90F803FEA3DD46A323FFBAD7BFB4D58" box="[1122,1451,1056,1081]" italics="true" pageId="19" pageNumber="250">I.H. Ooi &amp; K. Jeland OIH116</emphasis>
</paragraph>
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(holo SAR),
<figureCitation id="134040A8EA3DD46A3745FBC97FDA4D3C" box="[280,394,1092,1117]" captionStart="FIGURE 10" captionStartId="20.[136,229,1996,2017]" captionTargetBox="[163,1428,179,1966]" captionTargetId="figure-17@20.[159,1429,175,1971]" captionTargetPageId="20" captionText="FIGURE 10. Scaphochlamys graveolens. A. Type, OIH116 (SAR). B. plant in habitat. C. inflorescence. D. lamina. E. flower, top view. F.flower, side view. G. labellum. H. stamen, note spurless anther and slightly purplish crest. I. bract, subtending two flowers. J. bract and bracteoles. K. calyx and ovary. L. petiole base and rhizome. A–E, H–L from OIH116; F–G from ZI568." pageId="19" pageNumber="250">
<emphasis id="B90F803FEA3DD46A3745FBC97FDA4D3C" bold="true" box="[280,394,1092,1117]" pageId="19" pageNumber="250">Figure 10</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA3DD46A36D5FBE67AF44142" pageId="19" pageNumber="250" type="diagnosis">
<paragraph id="8BC45C2DEA3DD46A36D5FBE67FDE4DC5" blockId="19.[136,1452,1018,1188]" pageId="19" pageNumber="250">
<emphasis id="B90F803FEA3DD46A36D5FBE67EBB4DE1" bold="true" box="[136,235,1131,1152]" pageId="19" pageNumber="250">Diagnosis</emphasis>
:—
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<emphasis id="B90F803FEA3DD46A3754FBE67C414DE1" box="[265,529,1131,1152]" italics="true" pageId="19" pageNumber="250">Scaphochlamys graveolens</emphasis>
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is distinguished from the rest Bornean species with its large paddle-shaped leaves and foul smelling floral odour.
</paragraph>
<paragraph id="8BC45C2DEA3DD46A36D5FB437AF44142" blockId="19.[136,1452,1230,2083]" pageId="19" pageNumber="250">
Terrestrial, perennial, rhizomatous herbs, to c.
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tall. Rhizome below ground, c.
<quantity id="4C83F1C8EA3DD46A3212FB437AD84D86" box="[1103,1160,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="250" unit="cm" value="1.0">1 cm</quantity>
diameter, externally light brown to medium brown, internally brownish white. Elements c.
<quantity id="4C83F1C8EA3DD46A3536FB7F7D984C6A" box="[875,968,1266,1291]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="19" pageNumber="250" unit="cm" value="3.0" valueMax="5.0" valueMin="1.0">1− 5 cm</quantity>
apart, 1-foliate, juvenile elements up to 3- foliate; petiole
<quantity id="4C83F1C8EA3DD46A376EFA9B7FF44C4E" box="[307,420,1302,1327]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="19" pageNumber="250" unit="cm" value="25.0" valueMax="32.0" valueMin="18.0">18−32 cm</quantity>
long, canaliculate, brownish green, pubescent, base pulvinate, white to light green; leafless sheath 3–4,
<quantity id="4C83F1C8EA3DD46A3749FAB77FDE4C32" box="[276,398,1338,1363]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="14.5" metricValueMin="3.0" pageId="19" pageNumber="250" unit="cm" value="8.75" valueMax="14.5" valueMin="3.0">3−14.5 cm</quantity>
long, brownish green to medium green, glabrous, margin membranous, decaying with age; leaf sheath c.
<quantity id="4C83F1C8EA3DD46A3774FAD37F334C16" box="[297,355,1374,1399]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="250" unit="cm" value="4.0">4 cm</quantity>
long, membranous, light green, glabrous, decaying with age; ligule obscure, pubescent; lamina
<quantity id="4C83F1C8EA3DD46A36D5FA0F7EAA4CFA" box="[136,250,1410,1435]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="19" pageNumber="250" unit="cm" value="27.5" valueMax="35.0" valueMin="20.0">20–35 cm</quantity>
×
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, ovate to oblong-elliptic, coriaceous, margin entire, base attenuate, apex mucronate, often slightly recurved; adaxial surface matte, medium green to dark green, glabrous, midrib sunken, brownish green to light green, sparsely pubescent at base, main lateral veins slightly raised; abaxial surface light green, pubescent, midrib raised, brownish green, densely pubescent, main lateral veins slightly visible. Inflorescence
<quantity id="4C83F1C8EA3DD46A32BBFA637B3B4F66" box="[1254,1387,1518,1543]" metricMagnitude="-1" metricUnit="m" metricValue="1.125" metricValueMax="1.25" metricValueMin="1.0" pageId="19" pageNumber="250" unit="cm" value="11.25" valueMax="12.5" valueMin="10.0">10–12.5 cm</quantity>
long, emerging from near leaf base inside sheaths, tightly congested, bulb-like, comprised of 4–6, 2-flowered cincinni borne on a peduncle; peduncle
<quantity id="4C83F1C8EA3DD46A37C2F9BB7C584F2E" box="[415,520,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="19" pageNumber="250" unit="cm" value="3.75" valueMax="5.0" valueMin="2.5">2.5–5 cm</quantity>
long, white, glabrous, exposed, slightly covered by the remaining decayed leafless sheaths; bract 4–6, c. 4 ×
<quantity id="4C83F1C8EA3DD46A37FDF9D77FBD4F12" box="[416,493,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="250" unit="cm" value="1.6">1.6 cm</quantity>
, spirally arranged, lanceolate, membranous except first and second bracts at outermost layer sometimes slightly coriaceous, light brown to greenish light brown, glabrous, apex acute, each bract subtending 2 flowers; bracteole c. 2,
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long, barely distinguishable from bracts, light brown to brownish white; flower c.
<quantity id="4C83F1C8EA3DD46A36D5F94B7E904FBE" box="[136,192,1734,1759]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="250" unit="cm" value="7.0">7 cm</quantity>
long, floral odour reminiscent of liquid seeping from domestic waste; calyx c.
<quantity id="4C83F1C8EA3DD46A321BF94B7AC64FBE" box="[1094,1174,1734,1759]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="19" pageNumber="250" unit="mm" value="18.0">18 mm</quantity>
long, white, apex acute, pubescent at base; floral tube
<quantity id="4C83F1C8EA3DD46A3788F9677C004E62" box="[469,592,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="19" pageNumber="250" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40−45 mm</quantity>
long, white, glabrous; corolla lobes c.
<quantity id="4C83F1C8EA3DD46A325FF9677A014E62" box="[1026,1105,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="250" unit="mm" value="22.0">22 mm</quantity>
long, lanceolate, white to light yellow, glabrous, apex acute, hooded; staminodes
<quantity id="4C83F1C8EA3DD46A3495F8837D134E46" box="[712,835,1806,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="19" pageNumber="250" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
long, linear to slightly oblanceolate, white to slightly light yellow, apex rounded to acute, adaxial surface covered with glandular hairs; labellum c. 25 ×
<quantity id="4C83F1C8EA3DD46A328AF8BF7B784E2A" box="[1239,1320,1842,1867]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="250" unit="mm" value="15.0">15 mm</quantity>
, spathulate, adaxial surface covered with glandular hairs, white with violet margined light yellow median band extending to lobes, apex 2-lobed, indented c.
<quantity id="4C83F1C8EA3DD46A37F3F8F77FA34EF2" box="[430,499,1914,1939]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="250" unit="mm" value="5.0">5 mm</quantity>
, recurved, lobes slightly overlapping; stamen c. 12 ×
<quantity id="4C83F1C8EA3DD46A320BF8F77ACA4EF2" box="[1110,1178,1914,1939]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="250" unit="mm" value="2.0">2 mm</quantity>
, covered with glandular hairs, white with tinged violet at crest; filament c.
<quantity id="4C83F1C8EA3DD46A3493F8137D414ED6" box="[718,785,1950,1975]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="250" unit="mm" value="5.0">5 mm</quantity>
long; anther thecae c.
<quantity id="4C83F1C8EA3DD46A324AF8137A0A4ED6" box="[1047,1114,1950,1975]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="250" unit="mm" value="5.0">5 mm</quantity>
long, spurless, crest c.
<quantity id="4C83F1C8EA3DD46A3334F8137BFC4ED6" box="[1385,1452,1950,1975]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="250" unit="mm" value="2.0">2 mm</quantity>
long; stigma less than
<quantity id="4C83F1C8EA3DD46A37D1F84F7F9F4EBA" box="[396,463,1986,2011]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="250" unit="mm" value="1.0">1 mm</quantity>
long, club-shaped with 2 dorsal knobs, ostiole ciliate, forward facing; style c.
<quantity id="4C83F1C8EA3DD46A3306F84F7BFC4EBA" box="[1371,1452,1986,2011]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="19" pageNumber="250" unit="mm" value="55.0">55 mm</quantity>
long, white, glabrous; ovary c.
<quantity id="4C83F1C8EA3DD46A37BFF86B7C734E9E" box="[482,547,2022,2047]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="250" unit="mm" value="4.0">4 mm</quantity>
long, 1-locular,white, sparsely pubescent but densely at apex, placentation basal; epigynous glands 2, c.
<quantity id="4C83F1C8EA3DD46A37D4F7877F9A4142" box="[393,458,2058,2083]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="250" unit="mm" value="6.0">6 mm</quantity>
long, free, needle-like, brownish white. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<caption id="DF040CA5EA3AD46D36D5F8417A3A4148" pageId="20" pageNumber="251" startId="20.[136,229,1996,2017]" targetBox="[163,1428,179,1966]" targetPageId="20" targetType="figure">
<paragraph id="8BC45C2DEA3AD46D36D5F8417A3A4148" blockId="20.[136,1452,1996,2089]" pageId="20" pageNumber="251">
<emphasis id="B90F803FEA3AD46D36D5F8417F5A4E80" bold="true" box="[136,266,1996,2017]" pageId="20" pageNumber="251">FIGURE 10.</emphasis>
<emphasis id="B90F803FEA3AD46D374CF8417C4D4E80" box="[273,541,1996,2017]" italics="true" pageId="20" pageNumber="251">
<taxonomicName id="4C7B27AEEA3AD46D374CF8417C494E80" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[273,537,1996,2017]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="20" pageNumber="251" phylum="Tracheophyta" rank="species" species="graveolens">Scaphochlamys graveolens</taxonomicName>
.
</emphasis>
<emphasis id="B90F803FEA3AD46D347FF8417C6A4E80" bold="true" box="[546,570,1996,2017]" pageId="20" pageNumber="251">A.</emphasis>
Type,
<emphasis id="B90F803FEA3AD46D3420F8417C994E80" box="[637,713,1996,2017]" italics="true" pageId="20" pageNumber="251">OIH116</emphasis>
(SAR).
<emphasis id="B90F803FEA3AD46D3546F8417D624E80" bold="true" box="[795,818,1996,2017]" pageId="20" pageNumber="251">B.</emphasis>
plant in habitat.
<emphasis id="B90F803FEA3AD46D358BF8417DBD4E80" bold="true" box="[982,1005,1996,2017]" pageId="20" pageNumber="251">C.</emphasis>
inflorescence.
<emphasis id="B90F803FEA3AD46D32DDF8417AC84E80" bold="true" box="[1152,1176,1996,2017]" pageId="20" pageNumber="251">D.</emphasis>
lamina.
<emphasis id="B90F803FEA3AD46D32B1F8417B524E80" bold="true" box="[1260,1282,1996,2017]" pageId="20" pageNumber="251">E.</emphasis>
flower, top view.
<emphasis id="B90F803FEA3AD46D36D5F87D7ECD4164" bold="true" box="[136,157,2032,2053]" pageId="20" pageNumber="251">F.</emphasis>
flower, side view.
<emphasis id="B90F803FEA3AD46D3713F87D7F374164" bold="true" box="[334,359,2032,2053]" pageId="20" pageNumber="251">G.</emphasis>
labellum.
<emphasis id="B90F803FEA3AD46D3791F87D7FB54164" bold="true" box="[460,485,2032,2053]" pageId="20" pageNumber="251">H.</emphasis>
stamen, note spurless anther and slightly purplish crest.
<emphasis id="B90F803FEA3AD46D35A2F87D7A5F4164" bold="true" box="[1023,1039,2032,2053]" pageId="20" pageNumber="251">I.</emphasis>
bract, subtending two flowers.
<emphasis id="B90F803FEA3AD46D3361F87D7B1E4164" bold="true" box="[1340,1358,2032,2053]" pageId="20" pageNumber="251">J.</emphasis>
bract and bracteoles.
<emphasis id="B90F803FEA3AD46D36A8F7997F5E4148" bold="true" box="[245,270,2068,2089]" pageId="20" pageNumber="251">K.</emphasis>
calyx and ovary.
<emphasis id="B90F803FEA3AD46D37E5F7997F9E4148" bold="true" box="[440,462,2068,2089]" pageId="20" pageNumber="251">L.</emphasis>
petiole base and rhizome. A–E, H–L from
<emphasis id="B90F803FEA3AD46D3536F7997DE84148" box="[875,952,2068,2089]" italics="true" pageId="20" pageNumber="251">OIH116</emphasis>
; F–G from
<emphasis id="B90F803FEA3AD46D3277F7997A344148" box="[1066,1124,2068,2089]" italics="true" pageId="20" pageNumber="251">ZI568</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C3610FA6EA3BD46C36E0FF2D7BCD49BC" pageId="21" pageNumber="252" type="etymology">
<paragraph id="8BC45C2DEA3BD46C36E0FF2D7BCD49BC" blockId="21.[136,1452,160,797]" pageId="21" pageNumber="252">
<emphasis id="B90F803FEA3BD46C36E0FF2D7F6E49D8" bold="true" box="[189,318,160,185]" pageId="21" pageNumber="252">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C7B27AEEA3BD46C3408FF2D7C8049D8" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[597,720,160,185]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="B90F803FEA3BD46C3408FF2D7C8049D8" box="[597,720,160,185]" italics="true" pageId="21" pageNumber="252">graveolens</emphasis>
</taxonomicName>
, was chosen based on the foul floral odour, which is unique to
<taxonomicName id="4C7B27AEEA3BD46C36D5FF497FE849BC" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[136,440,196,221]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="B90F803FEA3BD46C36D5FF497FE849BC" box="[136,440,196,221]" italics="true" pageId="21" pageNumber="252">Scaphochlamys graveolens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B27AEEA3BD46C37B2FF497C0349BC" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[495,595,196,221]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="lucens">
<emphasis id="B90F803FEA3BD46C37B2FF497C0349BC" box="[495,595,196,221]" italics="true" pageId="21" pageNumber="252">S. lucens</emphasis>
</taxonomicName>
. Many
<taxonomicName id="4C7B27AEEA3BD46C34F5FF497D0649BC" authorityName="Baker" authorityYear="1892" box="[680,854,196,221]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F803FEA3BD46C34F5FF497D0649BC" box="[680,854,196,221]" italics="true" pageId="21" pageNumber="252">Scaphochlamys</emphasis>
</taxonomicName>
have either no floral smell or pleasant floral smell).
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA3BD46C36E0FF657F284B40" pageId="21" pageNumber="252" type="distribution">
<paragraph id="8BC45C2DEA3BD46C36E0FF657F014828" blockId="21.[136,1452,160,797]" pageId="21" pageNumber="252">
<emphasis id="B90F803FEA3BD46C36E0FF657FBC4860" bold="true" box="[189,492,232,257]" pageId="21" pageNumber="252">Distribution and ecology</emphasis>
:
<emphasis id="B90F803FEA3BD46C37AEFF657D104860" box="[499,832,232,257]" italics="true" pageId="21" pageNumber="252">
—
<taxonomicName id="4C7B27AEEA3BD46C344DFF657D104860" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[528,832,232,257]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="graveolens">Scaphochlamys graveolens</taxonomicName>
</emphasis>
is known from the
<typeStatus id="54C0E28FEA3BD46C3273FF657A0E4860" box="[1070,1118,232,257]" pageId="21" pageNumber="252">type</typeStatus>
locality and from Kampung Segu, Padawan, occurring on moist mollisolic yellow-orange soil at the base of karst limestone along a forested stream, c.
<quantity id="4C83F1C8EA3BD46C36FEFEBD7E8C4828" box="[163,220,304,329]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="252" unit="m" value="90.0">90 m</quantity>
elevation.
</paragraph>
<paragraph id="8BC45C2DEA3BD46C36E0FED97AB748D4" blockId="21.[136,1452,160,797]" pageId="21" pageNumber="252">
<materialsCitation id="3B135670EA3BD46C36E0FED97B6B48F0" collectingDate="2004-08-20" collectorName="P. C. Boyce &amp; K. Simon" country="Malaysia" county="Pichin" latitude="1.1343611" location="Samarahan Division" longLatPrecision="1" longitude="110.45008" municipality="Serian" pageId="21" pageNumber="252" specimenCount="2">
<emphasis id="B90F803FEA3BD46C36E0FED97CB0480C" bold="true" box="[189,736,340,365]" pageId="21" pageNumber="252">
<typeStatus id="54C0E28FEA3BD46C36E0FED97C18480C" box="[189,584,340,365]" isOtherTypeStatus="true" pageId="21" pageNumber="252">Additional specimens examined</typeStatus>
(
<typeStatus id="54C0E28FEA3BD46C3403FED97C88480C" box="[606,728,340,365]" pageId="21" pageNumber="252" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—MALAYSIAN BORNEO.
<collectingCountry id="F36C1CBDEA3BD46C321CFED97AF6480C" box="[1089,1190,340,365]" name="Malaysia" pageId="21" pageNumber="252">Sarawak</collectingCountry>
:
<location id="8EA40AF6EA3BD46C32E5FED97BF7480C" LSID="urn:lsid:plazi:treatment:03D2ED3BEA3DD46C36D5FC717A1C4A7C:8EA40AF6EA3BD46C32E5FED97BF7480C" box="[1208,1447,340,365]" country="Malaysia" county="Pichin" latitude="1.1343611" longLatPrecision="1" longitude="110.45008" municipality="Serian" name="Samarahan Division" pageId="21" pageNumber="252">Samarahan Division</location>
,
<collectingMunicipality id="6BA0C657EA3BD46C36D5FEF57E8248F0" box="[136,210,376,401]" pageId="21" pageNumber="252">Serian</collectingMunicipality>
,
<collectingCounty id="62A524A1EA3BD46C36BFFEF57F7C48F0" box="[226,300,376,401]" pageId="21" pageNumber="252">Pichin</collectingCounty>
,
<geoCoordinate id="EE4F3AEAEA3BD46C3761FEF57F8D48F0" box="[316,477,376,401]" degrees="01" direction="north" minutes="08" orientation="latitude" pageId="21" pageNumber="252" precision="1" seconds="03.7" value="1.1343611">01°08’03.7”N</geoCoordinate>
,
<geoCoordinate id="EE4F3AEAEA3BD46C37B0FEF57CC848F0" box="[493,664,376,401]" degrees="110" direction="east" minutes="27" orientation="longitude" pageId="21" pageNumber="252" precision="1" seconds="00.3" value="110.45008">110°27’00.3”E</geoCoordinate>
,
<date id="FFC57AEDEA3BD46C34F5FEF57D3548F0" box="[680,869,376,401]" pageId="21" pageNumber="252" value="2004-08-20">
<collectingDate id="EF818305EA3BD46C34F5FEF57D3548F0" box="[680,869,376,401]" pageId="21" pageNumber="252" value="2004-08-20">20 August 2004</collectingDate>
</date>
,
<emphasis id="B90F803FEA3BD46C3528FEF57AB348F0" box="[885,1251,376,401]" italics="true" pageId="21" pageNumber="252">
<collectorName id="268E39FBEA3BD46C3528FEF57DA248F0" box="[885,1010,376,401]" pageId="21" pageNumber="252">P.C. Boyce</collectorName>
&amp;
<collectorName id="268E39FBEA3BD46C327DFEF57ADB48F0" box="[1056,1163,376,401]" pageId="21" pageNumber="252">K. Simon</collectorName>
ZI-568
</emphasis>
(SAR)
</materialsCitation>
;
<materialsCitation id="3B135670EA3BD46C3316FEF57AB348D4" collectingDate="1975-05-26" collectorName="B. L. Burtt" country="Malaysia" county="Kampung Segu" location="Kuching Division" municipality="Penrissen Road" pageId="21" pageNumber="252" specimenCount="1">
<location id="8EA40AF6EA3BD46C3316FEF57EB848D4" LSID="urn:lsid:plazi:treatment:03D2ED3BEA3DD46C36D5FC717A1C4A7C:8EA40AF6EA3BD46C3316FEF57EB848D4" country="Malaysia" county="Kampung Segu" municipality="Penrissen Road" name="Kuching Division" pageId="21" pageNumber="252">Kuching Division</location>
(‘First Division’),
<collectingMunicipality id="6BA0C657EA3BD46C3798FE117C2548D4" box="[453,629,412,437]" pageId="21" pageNumber="252">Penrissen Road</collectingMunicipality>
,
<collectingCounty id="62A524A1EA3BD46C34DDFE117D6048D4" box="[640,816,412,437]" pageId="21" pageNumber="252">Kampung Segu</collectingCounty>
,
<date id="FFC57AEDEA3BD46C3566FE117D8448D4" box="[827,980,412,437]" pageId="21" pageNumber="252" value="1975-05-26">
<collectingDate id="EF818305EA3BD46C3566FE117D8448D4" box="[827,980,412,437]" pageId="21" pageNumber="252" value="1975-05-26">26 May 1975</collectingDate>
</date>
,
<emphasis id="B90F803FEA3BD46C3582FE117AFC48D4" box="[991,1196,412,437]" italics="true" pageId="21" pageNumber="252">
<collectorName id="268E39FBEA3BD46C3582FE117A1E48D4" box="[991,1102,412,437]" pageId="21" pageNumber="252">B.L. Burtt</collectorName>
B. 8798
</emphasis>
(E!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC45C2DEA3BD46C36E0FE4D7F284B40" blockId="21.[136,1452,160,797]" pageId="21" pageNumber="252">
<emphasis id="B90F803FEA3BD46C36E0FE4D7FFB48B8" bold="true" box="[189,427,448,473]" pageId="21" pageNumber="252">Conservation status</emphasis>
:—Endangered, EN B2ab(iii). The localities of
<taxonomicName id="4C7B27AEEA3BD46C359DFE4D7A0848B8" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[960,1112,448,473]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="B90F803FEA3BD46C359DFE4D7A0848B8" box="[960,1112,448,473]" italics="true" pageId="21" pageNumber="252">S. graveolens</emphasis>
</taxonomicName>
are not within protected areas and the locality for
<emphasis id="B90F803FEA3BD46C373EFE697FAA489C" box="[355,506,484,509]" italics="true" pageId="21" pageNumber="252">Burtt B. 8798</emphasis>
collection is now an urban area. Based on the criteria in
<bibRefCitation id="EFEA21DCEA3BD46C3229FE697B5D489C" author="IUCN" box="[1140,1293,484,509]" pageId="21" pageNumber="252" refId="ref32181" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. Second Edition, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
, the species is listed as Endangered.
</paragraph>
</subSubSection>
<subSubSection id="C3610FA6EA3BD46C36E0FDA17A1C4A7C" pageId="21" pageNumber="252" type="discussion">
<paragraph id="8BC45C2DEA3BD46C36E0FDA17A1C4A7C" blockId="21.[136,1452,160,797]" pageId="21" pageNumber="252">
<emphasis id="B90F803FEA3BD46C36E0FDA17F6B4B24" bold="true" box="[189,315,556,581]" pageId="21" pageNumber="252">Discussion</emphasis>
:—
<taxonomicName id="4C7B27AEEA3BD46C3702FDA17CC24B24" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[351,658,556,581]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="B90F803FEA3BD46C3702FDA17CC24B24" box="[351,658,556,581]" italics="true" pageId="21" pageNumber="252">Scaphochlamys graveolens</emphasis>
</taxonomicName>
is, so far, the largest
<taxonomicName id="4C7B27AEEA3BD46C35CFFDA17A104B24" authorityName="Baker" authorityYear="1892" box="[914,1088,556,581]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F803FEA3BD46C35CFFDA17A104B24" box="[914,1088,556,581]" italics="true" pageId="21" pageNumber="252">Scaphochlamys</emphasis>
</taxonomicName>
in Borneo.
<emphasis id="B90F803FEA3BD46C3290FDA17B3B4B24" box="[1229,1387,556,581]" italics="true" pageId="21" pageNumber="252">Burtt B. 8798</emphasis>
from Kampung Segu, Penrissen Road is identified as
<taxonomicName id="4C7B27AEEA3BD46C34C4FDDD7D7F4B08" authorityName="Ooi, K. Meekiong &amp; S. Y. Wong" authorityYear="2017" box="[665,815,592,617]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="B90F803FEA3BD46C34C4FDDD7D7F4B08" box="[665,815,592,617]" italics="true" pageId="21" pageNumber="252">S. graveolens</emphasis>
</taxonomicName>
based on its much larger elements, attenuate lamina base, mucronate lamina apex, and long roots with very short tertiary roots.
<bibRefCitation id="EFEA21DCEA3BD46C35C6FDF97A614BEC" author="Smith, R. M." box="[923,1073,628,653]" pageId="21" pageNumber="252" pagination="203 - 232" refId="ref33328" refString="Smith, R. M. (1987) A review of Bornean Zingiberaceae: III (Hedychieae). Notes from the Royal Botanic Garden, Edinburgh 44: 203 - 232." type="journal article" year="1987">Smith (1987)</bibRefCitation>
and
<bibRefCitation id="EFEA21DCEA3BD46C3234FDF97B524BEC" author="Searle, R. J." box="[1129,1282,628,653]" pageId="21" pageNumber="252" pagination="75 - 121" refId="ref33212" refString="Searle, R. J. (2010) The genus Scaphochlamys (Zingiberaceae - Zingibereae): A Compendium for the field worker. Edinburgh Journal of Botany 67: 75 - 121. https: // doi. org / 10.1017 / S 0960428609990254" type="journal article" year="2010">Searle (2010)</bibRefCitation>
included
<emphasis id="B90F803FEA3BD46C332FFDF97EB14BD0" italics="true" pageId="21" pageNumber="252">Burtt B. 8798</emphasis>
as
<taxonomicName id="4C7B27AEEA3BD46C3754FD157CE84BD0" authority="Holttum (1950: 95)" authorityName="Holttum" authorityPageNumber="95" authorityYear="1950" box="[265,696,664,689]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" isUncertain="true" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="breviscapa">
<emphasis id="B90F803FEA3BD46C3754FD157F4E4BD0" box="[265,286,664,689]" italics="true" pageId="21" pageNumber="252">S.</emphasis>
aff.
<emphasis id="B90F803FEA3BD46C3708FD157F9F4BD0" box="[341,463,664,689]" italics="true" pageId="21" pageNumber="252">breviscapa</emphasis>
<bibRefCitation id="EFEA21DCEA3BD46C3785FD157CE84BD0" author="Holttum, R. E." box="[472,696,664,689]" pageId="21" pageNumber="252" pagination="1 - 249" refId="ref32053" refString="Holttum, R. E. (1950) The Zingiberaceae of the Malay Peninsula. Gardens' Bulletin Singapore 13: 1 - 249." type="journal article" year="1950">Holttum (1950: 95)</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="4C7B27AEEA3BD46C3564FD157D814BD0" authorityName="Holttum" authorityYear="1950" box="[825,977,664,689]" class="Liliopsida" family="Zingiberaceae" genus="Scaphochlamys" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="252" phylum="Tracheophyta" rank="species" species="breviscapa">
<emphasis id="B90F803FEA3BD46C3564FD157D814BD0" box="[825,977,664,689]" italics="true" pageId="21" pageNumber="252">S. breviscapa</emphasis>
</taxonomicName>
has a chartaceous acuminate lamina with the midrib pubescent abaxially, imbricating inflorescences with a short peduncle (
<quantity id="4C83F1C8EA3BD46C327EFD317AC14BB4" box="[1059,1169,700,725]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="21" pageNumber="252" unit="cm" value="2.25" valueMax="3.5" valueMin="1.0">1–3.5 cm</quantity>
), small coriaceous bracts (
<quantity id="4C83F1C8EA3BD46C36CDFD6D7EAD4B98" box="[144,253,736,761]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="21" pageNumber="252" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 cm</quantity>
), small flowers (less than
<quantity id="4C83F1C8EA3BD46C3441FD6D7C024B98" box="[540,594,736,761]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="252" unit="cm" value="4.0">4 cm</quantity>
long), white labellum with lilac streaks on both side of a light yellow median band near base, and was described from Kemaman,
<collectingRegion id="49BF92CFEA3BD46C3492FC897D064A7C" box="[719,854,772,797]" country="Malaysia" name="Terengganu" pageId="21" pageNumber="252">Terengganu</collectingRegion>
, Peninsular
<collectingCountry id="F36C1CBDEA3BD46C3583FC897A184A7C" box="[990,1096,772,797]" name="Malaysia" pageId="21" pageNumber="252">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>