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<document id="D66F83CC69046F9C87382693DDDDF63C" ID-DOI="10.11646/phytotaxa.395.3.8" ID-ISSN="1179-3163" ID-Zenodo-Dep="13716803" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725650516305" checkinUser="felipe" docAuthor="Al-Shehbaz, Ihsan A., Cano, Asunción &amp; Trinidad, Huber" docDate="2019" docId="03A41B4FFFD1FF9FFF239105F07155C2" docLanguage="en" docName="phytotaxa.395.3.8.pdf" docOrigin="Phytotaxa 395 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.395.3.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Draba ancashensis Al-Shehbaz, A. Cano" docType="treatment" docVersion="2" lastPageNumber="237" masterDocId="FF9D6337FFD1FF9DFFAB9469F4525320" masterDocTitle="Draba ancashensis (Brassicaceae; Cruciferae), a new species from Peru and a key to the Peruvian species of the genus" masterLastPageNumber="240" masterPageNumber="235" pageNumber="235" updateTime="1725657381358" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="489769860BABE8E409562617B417D711">Draba ancashensis (Brassicaceae; Cruciferae), a new species from Peru and a key to the Peruvian species of the genus</mods:title>
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<mods:namePart id="5314612D43D46BB4A604094B4D5089C2">Al-Shehbaz, Ihsan A.</mods:namePart>
<mods:affiliation id="BECB94929922C2C29280816EAA652A6B">Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, Missouri 63110, U. S. A.</mods:affiliation>
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<mods:namePart id="07A64E41F476A9D010733B89C02703BF">Cano, Asunción</mods:namePart>
<mods:affiliation id="75FDFD9715A6210D1711057FF4D5C0EB">Laboratorio de Florística, Departamento de Dicotiledóneas, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos (UNMSM), Av. Arenales 1256, Lima 11, Peru. &amp; Instituto de Investigación de Ciencias Biológicas Antonio Raimondi (ICBAR), Facultad de Ciencias Biológicas, (UNMSM), Calle Germán Amézaga 375, Lima 1, Peru.</mods:affiliation>
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<mods:namePart id="324CCA7E7EF16BA8708C8DBEBD1DDA1E">Trinidad, Huber</mods:namePart>
<mods:affiliation id="AB418A10E6233D9E9B8AB9B3CF11D45A">Laboratorio de Florística, Departamento de Dicotiledóneas, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos (UNMSM), Av. Arenales 1256, Lima 11, Peru.</mods:affiliation>
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<mods:title id="75A58D780A3B9308D5B35567B5CFE506">Phytotaxa</mods:title>
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<mods:date id="D6BD43ACBDFB801D85C6BAEAF85B0991">2019</mods:date>
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<mods:identifier id="E225AC7FB8DDB5EC3E05FFFF8BCF5A28" type="Zenodo-Dep">13716803</mods:identifier>
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<subSubSection id="C317F9D2FFD1FF9DFF239105F76A56A7" box="[136,824,1388,1415]" pageId="0" pageNumber="235" type="nomenclature">
<paragraph id="8BB2AA59FFD1FF9DFF239105F76A56A7" blockId="0.[136,824,1388,1415]" box="[136,824,1388,1415]" pageId="0" pageNumber="235">
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<taxonomicName id="4C0DD1DAFFD1FF9DFF239105F60356A7" ID-CoL="C6ZRN" authority="Al-Shehbaz, A. Cano" authorityName="Al-Shehbaz, A. Cano" box="[136,593,1388,1415]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="235" phylum="Tracheophyta" rank="species" species="ancashensis">
<emphasis id="B979764BFFD1FF9DFF239105F53656A7" bold="true" box="[136,356,1388,1415]" italics="true" pageId="0" pageNumber="235">Draba ancashensis</emphasis>
Al-Shehbaz, A.Cano
</taxonomicName>
&amp;
<collectingCountry id="F31AEAC9FFD1FF9DFDDF9107F68556A7" box="[628,727,1390,1415]" name="Trinidad and Tobago" pageId="0" pageNumber="235">Trinidad</collectingCountry>
. sp. nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C317F9D2FFD1FF9DFF2391FBF6A156D3" pageId="0" pageNumber="235" type="diagnosis">
<paragraph id="8BB2AA59FFD1FF9DFF2391FBF6A156D3" blockId="0.[136,1452,1426,2063]" pageId="0" pageNumber="235">
Diagnosis:—
<taxonomicName id="4C0DD1DAFFD1FF9DFEB791FBF5A0568B" box="[284,498,1426,1451]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="235" phylum="Tracheophyta" rank="species" species="ancashensis">
<emphasis id="B979764BFFD1FF9DFEB791FBF5A0568B" box="[284,498,1426,1451]" italics="true" pageId="0" pageNumber="235">Draba ancashensis</emphasis>
</taxonomicName>
is easily distinguished from the other Peruvian species of the genus by having woody lower stems, cauline leaves adaxially with simple trichomes and abaxially with stellate, 12-branched trichomes, fully bracteate racemes, white flowers, and persistent sepals.
</paragraph>
</subSubSection>
<subSubSection id="C317F9D2FFD1FF9DFF169197F5C4557F" pageId="0" pageNumber="235" type="materials_examined">
<paragraph id="8BB2AA59FFD1FF9DFF169197F5C4557F" blockId="0.[136,1452,1426,2063]" pageId="0" pageNumber="235">
<materialsCitation id="3B65A004FFD1FF9DFF169197F5C1557F" collectingDate="2017-08-05" collectorName="Asuncion Cano &amp; Huber Trinidad &amp; Sebastian Riva &amp; Niels Valencia" country="Peru" elevation="4761" latitude="-9.845361" location="Ichic Potrero" longLatPrecision="1" longitude="-77.09886" pageId="0" pageNumber="235" specimenCount="2" stateProvince="Ancash" typeStatus="holotype">
<emphasis id="B979764BFFD1FF9DFF169197F4AA5537" bold="true" box="[189,248,1534,1559]" pageId="0" pageNumber="235">Type</emphasis>
:—
<collectingCountry id="F31AEAC9FFD1FF9DFEB19197F5355537" box="[282,359,1534,1559]" name="Peru" pageId="0" pageNumber="235">PERU</collectingCountry>
.
<emphasis id="B979764BFFD1FF9DFEDF9197F5845537" bold="true" box="[372,470,1534,1559]" pageId="0" pageNumber="235">
<collectingRegion id="49C964BBFFD1FF9DFEDF9197F5835537" box="[372,465,1534,1559]" country="Peru" name="Ancash" pageId="0" pageNumber="235">Ancash</collectingRegion>
:
</emphasis>
Prov. Recuay,
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,
<date id="FFB38C99FFD1FF9DFC879197F7EA5537" box="[812,952,1534,1559]" pageId="0" pageNumber="235" value="2017-08-05">
<collectingDate id="EFF77571FFD1FF9DFC879197F7EA5537" box="[812,952,1534,1559]" pageId="0" pageNumber="235" value="2017-08-05">5 Aug 2017</collectingDate>
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,
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,
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,
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,
<emphasis id="B979764BFFD1FF9DFF23924BF784551B" box="[136,982,1570,1595]" italics="true" pageId="0" pageNumber="235">
<collectorName id="26F8CF8FFFD1FF9DFF23924BF568551B" box="[136,314,1570,1595]" pageId="0" pageNumber="235">Asunción Cano</collectorName>
,
<collectorName id="26F8CF8FFFD1FF9DFEE2924BF5AF551B" box="[329,509,1570,1595]" pageId="0" pageNumber="235">Huber Trinidad</collectorName>
,
<collectorName id="26F8CF8FFFD1FF9DFDA7924BF6E4551B" box="[524,694,1570,1595]" pageId="0" pageNumber="235">Sebastián Riva</collectorName>
&amp;
<collectorName id="26F8CF8FFFD1FF9DFD4A924BF7D7551B" box="[737,901,1570,1595]" pageId="0" pageNumber="235">Niels Valencia</collectorName>
22419
</emphasis>
(
<typeStatus id="54B614FBFFD1FF9DFC43924BF002551B" box="[1000,1104,1570,1595]" pageId="0" pageNumber="235" type="holotype">holotype</typeStatus>
: USM-304594!;
<typeStatus id="54B614FBFFD1FF9DFAB8924BF135551B" box="[1299,1383,1570,1595]" pageId="0" pageNumber="235" type="isotype">isotype</typeStatus>
, MO-6866608!).
<figureCitation id="1336B6DCFFD1FF9DFEA0922FF5C1557F" box="[267,403,1606,1631]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[136,229,2028,2049]" captionStartId-1="2.[136,229,549,570]" captionStartId-2="2.[136,229,1128,1149]" captionTargetBox-0="[326,1275,520,1975]" captionTargetId-0="figure-161@1.[297,1292,497,2003]" captionTargetPageId-0="1" captionText-0="FIGURE 1. Draba ancashensis.—A. Portion of plant.—B.A trichome from abaxial leaf surface.—C. Petal.—D. Infructescence after fruit dehiscence.—E. Fruit. Drawn by Al-Shehbaz from the isotype (Cano et al. 22419—MO-6866608)." captionText-1="FIGURE 2. Draba ancashensis. Scale = 7 mm." captionText-2="FIGURE 3. Draba ancashensis habitat." figureDoi-0="http://doi.org/10.5281/zenodo.13716805" httpUri-0="https://zenodo.org/record/13716805/files/figure.png" pageId="0" pageNumber="235">Figures 13</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="B979764BFFD1FF9DFF169203F50355A3" bold="true" box="[189,337,1642,1667]" pageId="0" pageNumber="235">Description:</emphasis>
—Plants small subshrubs, not scapose or pulvinate; caudex absent. Stems
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tall,
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in diam. above base, erect to ascending, several to many branched above, woody, naked at base and smooth bark exposed, covered at middle with remnants of outer stem layers and leaf remains, moderately pubescent with sessile, stellate trichomes with 2- or 3-branched rays and fewer simple trichomes to
<quantity id="4CF507BCFFD1FF9DFC4092BFF01055CF" box="[1003,1090,1750,1775]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="235" unit="mm" value="0.6">0.6 mm</quantity>
long; stem branches terminated in fruiting racemes of previous years, below which produced one or two branches of subsequent year. Basal leaves absent; leaves cauline, persistent, sessile, densely imbricate; petiole absent; leaf blade oblong to oblong-ovate, 510 ×
<quantity id="4CF507BCFFD1FF9DFF0B932BF547547B" box="[160,277,1858,1883]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="0" pageNumber="235" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">23.5 mm</quantity>
, abaxial surface densely pubescent with almost exclusively short-stalked stellate trichomes
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in diam., these with principal 4 rays each (2- or)3-branched and trichome appearing 12-branched, rarely few Y-shaped trichomes present, adaxial surface with antrorsely appressed simple trichomes
<quantity id="4CF507BCFFD1FF9DFC5E93E3F02E5483" box="[1013,1148,1930,1955]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="0" pageNumber="235" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
long, base cuneate, margin entire, ciliate near base with simple and sometimes fewer forked trichomes to
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long, apex obutse to subacute. Racemes bracteate throughout, 515-flowered, corymbose, dense, slightly elongated in fruit; peduncle
<quantity id="4CF507BCFFD1FF9DFAA993BBF13E54CB" box="[1282,1388,2002,2027]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="0" pageNumber="235" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
long; rachis pubescent as stem, straight; bracts similar to cauline leaves in shape and indumentum, gradually reduced in size upwards; lowermost fruiting pedicels divaricate to ascending, straight,
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long, densely hirsute all around with simple, Y-shaped, and fewer 3- or 4-rayed stalked trichomes. Sepals green, oblong,
<quantity id="4CF507BCFFD0FF9CFBD294ADF0BE53FD" box="[1145,1260,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="1" pageNumber="236" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">33.5 mm</quantity>
long, ascending, persistent through fruit maturity, base not saccate, densely stellate pubescent outside; petals white, spatulate, 44.5 ×
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, caducous, apex emarginate; claw only slightly differentiated from blade, ca.
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long; filaments white,
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long, slightly dilated at base; anthers ovate,
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long, obtuse at apex; nectar glands confluent, subtending bases of all filaments; ovules 1216 per ovary. Fruit oblong to oblong-ovate, 5.57 ×
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, strongly flattened, not twisted, sessile; valves puberulent with simple minutely staked or subsessile forked trichomes
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long, not or obscurely veined, obtuse at apex and base; style
<quantity id="4CF507BCFFD0FF9CFCCB95F5F7B55295" box="[864,999,412,437]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="236" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.40.7 mm</quantity>
long; stigma entire, narrower than style. Seeds brown, ovate, strongly compressed, 11.1 ×
<quantity id="4CF507BCFFD0FF9CFD6995A9F71952F9" box="[706,843,448,473]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="236" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, wingless.
</paragraph>
</subSubSection>
<caption id="DF72FAD1FFD0FF9CFF239385F0135B05" ID-DOI="http://doi.org/10.5281/zenodo.13716805" ID-Zenodo-Dep="13716805" httpUri="https://zenodo.org/record/13716805/files/figure.png" pageId="1" pageNumber="236" startId="1.[136,229,2028,2049]" targetBox="[326,1275,520,1975]" targetPageId="1" targetType="figure">
<paragraph id="8BB2AA59FFD0FF9CFF239385F0135B05" blockId="1.[136,1451,2028,2085]" pageId="1" pageNumber="236">
<emphasis id="B979764BFFD0FF9CFF239385F4AF5B21" bold="true" box="[136,253,2028,2049]" pageId="1" pageNumber="236">FIGURE 1.</emphasis>
<taxonomicName id="4C0DD1DAFFD0FF9CFEA99385F5EB5B21" box="[258,441,2028,2049]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="236" phylum="Tracheophyta" rank="species" species="ancashensis">
<emphasis id="B979764BFFD0FF9CFEA99385F5EB5B21" box="[258,441,2028,2049]" italics="true" pageId="1" pageNumber="236">Draba ancashensis</emphasis>
</taxonomicName>
.—A. Portion of plant.—B.A trichome from abaxial leaf surface.—C. Petal.—D. Infructescence after fruit dehiscence.—E. Fruit. Drawn by Al-Shehbaz from the isotype (
<emphasis id="B979764BFFD0FF9CFD409C79F7C55B05" box="[747,919,2064,2085]" italics="true" pageId="1" pageNumber="236">Cano et al. 22419</emphasis>
—MO-6866608).
</paragraph>
</caption>
<caption id="DF72FAD1FFD3FF9FFF23964CF606511A" box="[136,596,549,570]" pageId="2" pageNumber="237" startId="2.[136,229,549,570]" targetType="figure">
<paragraph id="8BB2AA59FFD3FF9FFF23964CF606511A" blockId="2.[136,596,549,570]" box="[136,596,549,570]" pageId="2" pageNumber="237">
<emphasis id="B979764BFFD3FF9FFF23964CF4AC511A" bold="true" box="[136,254,549,570]" pageId="2" pageNumber="237">FIGURE 2.</emphasis>
<emphasis id="B979764BFFD3FF9FFEA8964CF591511A" box="[259,451,549,570]" italics="true" pageId="2" pageNumber="237">
<taxonomicName id="4C0DD1DAFFD3FF9FFEA8964CF5ED511A" box="[259,447,549,570]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="ancashensis">Draba ancashensis</taxonomicName>
.
</emphasis>
Scale = 7 mm.
</paragraph>
</caption>
<caption id="DF72FAD1FFD3FF9FFF239001F659575D" box="[136,523,1128,1149]" pageId="2" pageNumber="237" startId="2.[136,229,1128,1149]" targetType="figure">
<paragraph id="8BB2AA59FFD3FF9FFF239001F659575D" blockId="2.[136,523,1128,1149]" box="[136,523,1128,1149]" pageId="2" pageNumber="237">
<emphasis id="B979764BFFD3FF9FFF239001F4AC575D" bold="true" box="[136,254,1128,1149]" pageId="2" pageNumber="237">FIGURE 3.</emphasis>
<taxonomicName id="4C0DD1DAFFD3FF9FFEA89001F5EE575D" box="[259,444,1128,1149]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="ancashensis">
<emphasis id="B979764BFFD3FF9FFEA89001F5EE575D" box="[259,444,1128,1149]" italics="true" pageId="2" pageNumber="237">Draba ancashensis</emphasis>
</taxonomicName>
habitat.
</paragraph>
</caption>
<subSubSection id="C317F9D2FFD3FF9FFF1690C4F48157CA" pageId="2" pageNumber="237" type="distribution">
<paragraph id="8BB2AA59FFD3FF9FFF1690C4F48157CA" blockId="2.[136,1452,1197,1762]" pageId="2" pageNumber="237">
<emphasis id="B979764BFFD3FF9FFF1690C4F50B57E6" bold="true" box="[189,345,1197,1222]" pageId="2" pageNumber="237">Distribution:</emphasis>
—Endemic to the Peruvian department
<collectingRegion id="49C964BBFFD3FF9FFCB690C4F72357E6" box="[797,881,1197,1222]" country="Peru" name="Ancash" pageId="2" pageNumber="237">Ancash</collectingRegion>
and known thus far only from the
<typeStatus id="54B614FBFFD3FF9FFAAF90C4F16657E6" box="[1284,1332,1197,1222]" pageId="2" pageNumber="237">type</typeStatus>
collection above.
</paragraph>
</subSubSection>
<subSubSection id="C317F9D2FFD3FF9FFF16909CF7185612" pageId="2" pageNumber="237" type="etymology">
<paragraph id="8BB2AA59FFD3FF9FFF16909CF7BD562E" blockId="2.[136,1452,1197,1762]" box="[189,1007,1269,1294]" pageId="2" pageNumber="237">
<emphasis id="B979764BFFD3FF9FFF16909CF51A562E" bold="true" box="[189,328,1269,1294]" pageId="2" pageNumber="237">Etymology:</emphasis>
—The species epithet is named after the department
<collectingRegion id="49C964BBFFD3FF9FFC3F909CF7BE562E" box="[916,1004,1269,1294]" country="Peru" name="Ancash" pageId="2" pageNumber="237">Ancash</collectingRegion>
.
</paragraph>
<paragraph id="8BB2AA59FFD3FF9FFF169170F7185612" blockId="2.[136,1452,1197,1762]" box="[189,842,1305,1330]" pageId="2" pageNumber="237">
<emphasis id="B979764BFFD3FF9FFF169170F5715612" bold="true" box="[189,291,1305,1330]" pageId="2" pageNumber="237">Habitat:</emphasis>
—Rocky crevices and outcrops at mountain tops.
</paragraph>
</subSubSection>
<subSubSection id="C317F9D2FFD3FF9FFF169154F07155C2" pageId="2" pageNumber="237" type="discussion">
<paragraph id="8BB2AA59FFD3FF9FFF169154F07155C2" blockId="2.[136,1452,1197,1762]" pageId="2" pageNumber="237">
<emphasis id="B979764BFFD3FF9FFF169154F5175676" bold="true" box="[189,325,1341,1366]" pageId="2" pageNumber="237">Discussion:</emphasis>
<taxonomicName id="4C0DD1DAFFD3FF9FFECA9154F6645676" box="[353,566,1341,1366]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="ancashensis">
<emphasis id="B979764BFFD3FF9FFECA9154F6645676" box="[353,566,1341,1366]" italics="true" pageId="2" pageNumber="237">Draba ancashensis</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F31AEAC9FFD3FF9FFD7B9154F7515676" box="[720,771,1341,1366]" name="Peru" pageId="2" pageNumber="237">Peru</collectingCountry>
and known thus far only from the
<typeStatus id="54B614FBFFD3FF9FFBDC9154F0F55676" box="[1143,1191,1341,1366]" pageId="2" pageNumber="237">type</typeStatus>
collection. It resembles the three Peruvian-endemics
<taxonomicName id="4C0DD1DAFFD3FF9FFE789108F6F7565A" box="[467,677,1377,1402]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="cajamarcaensis">
<emphasis id="B979764BFFD3FF9FFE789108F6F7565A" box="[467,677,1377,1402]" italics="true" pageId="2" pageNumber="237">D. cajamarcaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0DD1DAFFD3FF9FFD1E9108F723565A" box="[693,881,1377,1402]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="matthioloides">
<emphasis id="B979764BFFD3FF9FFD1E9108F723565A" box="[693,881,1377,1402]" italics="true" pageId="2" pageNumber="237">D. matthioloides</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C0DD1DAFFD3FF9FFC199108F015565A" box="[946,1095,1377,1402]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B979764BFFD3FF9FFC199108F015565A" box="[946,1095,1377,1402]" italics="true" pageId="2" pageNumber="237">D. peruviana</emphasis>
</taxonomicName>
(see key for authors of species names), in having sessile, exclusively cauline leaves, woody lower stems, leaves abaxially pubescent with stellate trichomes, and fully bracteate racemes. It differs from all of them in the persistent (vs. readily caducous) sepals. From
<taxonomicName id="4C0DD1DAFFD3FF9FFF2391A4F50A56C6" box="[136,344,1485,1510]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="cajamarcaensis">
<emphasis id="B979764BFFD3FF9FFF2391A4F50A56C6" box="[136,344,1485,1510]" italics="true" pageId="2" pageNumber="237">D. cajamarcaensis</emphasis>
</taxonomicName>
, it is easily distinguished by having stems and raceme rachis pubescent with simple and stellate (vs. malpighiaceous) trichomes, cauline leaves pubescent with stalked (vs. sessile) stellate trichomes with branched rays and appearing 12-branched (vs. cruciform trichomes with at least some rays unbranched), and oblong to oblong-ovate (vs. narrowly lanceolate) fruit. It differs from
<taxonomicName id="4C0DD1DAFFD3FF9FFD399250F7755572" box="[658,807,1593,1618]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B979764BFFD3FF9FFD399250F7755572" box="[658,807,1593,1618]" italics="true" pageId="2" pageNumber="237">D. peruviana</emphasis>
</taxonomicName>
in having adaxial leaf surface with primarily simple (vs. stellate) trichomes, entire (vs. dentate) leaves, smaller petals 44.5 × 1.41.7 (vs. 7l0 × 33.5) mm, and shorter styles 0.40.7 (vs.1.53) mm long. Finally, it separated from
<taxonomicName id="4C0DD1DAFFD3FF9FFD5692E8F7EB55BA" box="[765,953,1665,1690]" class="Magnoliopsida" family="Brassicaceae" genus="Draba" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="matthioloides">
<emphasis id="B979764BFFD3FF9FFD5692E8F7EB55BA" box="[765,953,1665,1690]" italics="true" pageId="2" pageNumber="237">D. matthioloides</emphasis>
</taxonomicName>
by its adaxial leaf surface with exclusively simple (vs. stellate) trichomes, white (vs. yellow, orange, or purple) petals 44.5 × 1.41.7 (vs. 610 × 2.53.5) mm, 1216 (vs. 2438) ovules per ovary, and styles 0.40.7 (vs. (0.5) 310) mm long.
</paragraph>
</subSubSection>
</treatment>
</document>