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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.872.35361" ID-GBIF-Dataset="ed1a1322-b118-4434-b9ff-503a0fb64792" ID-PMC="PMC6711934" ID-Pensoft-Pub="1313-2970-872-13" ID-Pensoft-UUID="A9BE568D9BF65A179464AB8C7CB8BEB2" ID-PubMed="31496884" ID-ZooBank="DCDEFCB1E33D476F90D7178BC08C311A" ModsDocID="1313-2970-872-13" checkinTime="1566364211746" checkinUser="pensoft" docAuthor="Astafurova, Yulia V., Proshchalykin, Maxim Yu. &amp; Schwarz, Maximilian" docDate="2019" docId="C6671BF631855F19A9D51ED3E1E89547" docLanguage="en" docName="ZooKeys 872: 13-40" docOrigin="ZooKeys 872" docSource="http://dx.doi.org/10.3897/zookeys.872.35361" docTitle="Sphecodes tadschicus Bluethgen 1935" docType="treatment" docVersion="4" id="A9BE568D9BF65A179464AB8C7CB8BEB2" lastPageNumber="33" masterDocId="A9BE568D9BF65A179464AB8C7CB8BEB2" masterDocTitle="The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010" masterLastPageNumber="40" masterPageNumber="13" pageNumber="33" updateTime="1668167741671" updateUser="ExternalLinkService">
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<mods:title>The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010</mods:title>
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<mods:namePart>Astafurova, Yulia V.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Proshchalykin, Maxim Yu.</mods:namePart>
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<mods:namePart>Schwarz, Maximilian</mods:namePart>
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<mods:date>2019</mods:date>
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<mods:number>872</mods:number>
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<mods:start>13</mods:start>
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<paragraph pageId="20" pageNumber="33">
<taxonomicName LSID="c6671bf6-3185-5f19-a9d5-1ed3e1e89547" authority="Bluethgen, 1935" authorityName="Bluethgen" authorityYear="1935" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes tadschicus" order="Hymenoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="tadschicus">
Sphecodes tadschicus
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1935
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</paragraph>
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<subSubSection pageId="20" pageNumber="33" type="reference_group">
<paragraph pageId="20" pageNumber="33">
<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes tadschicus" order="Hymenoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="tadschicus">Sphecodes tadschicus</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
in Popov, 1935: 366, ♂, ♀ (holotype: ♂, near Kulab [Tajikistan], 25.VII.1935, V. Popov leg.; ZISP).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="diagnosis">
<paragraph pageId="20" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="20" pageNumber="33">
See
<bibRefCitation DOI="https://doi.org/10.25221/fee.369.1" author="Astafurova, YuV" journalOrPublisher="Far Eastern Entomologist" pageId="22" pageNumber="35" pagination="1 - 47" refId="B13" refString="Astafurova, YuV, Proshchalykin, MYu, Engel, MS, 2018a. The cuckoo bee genus Sphecodes Latreille, 1804 in Kazakhstan (Hymenoptera: Halictidae). . Far Eastern Entomologist 369: 1 - 47" title="The cuckoo bee genus Sphecodes Latreille, 1804 in Kazakhstan (Hymenoptera: Halictidae)." url="https://doi.org/10.25221/fee.369.1" volume="369" year="2018 a">Astafurova et al. 2018a</bibRefCitation>
: 39.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="materials_examined">
<paragraph pageId="20" pageNumber="33">Material examined.</paragraph>
<paragraph pageId="20" pageNumber="33">ISRAEL: 1 ♀, 8 ♂♂, Jerusalem, 10-25.VIII.1960, Bytinski (MNHB).</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="distribution">
<paragraph pageId="20" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="20" pageNumber="33">*Israel; Turkey, Iran, Central Asia, Kazakhstan.</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="DA684B0EDBD343BFFE6E73AB7A5D93FB" italics="true" pageNumber="135">Saguinus melanoleucus</emphasis>
Miranda Ribeiro 1912
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<taxonomicName id="90257D4519FE46A3692101557E89D188" authority="Miranda Ribeiro 1912" authorityName="Miranda Ribeiro" authorityYear="1912" class="Mammalia" family="Cebidae" genus="Saguinus" kingdom="Animalia" order="Primates" pageNumber="135" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="E4DEA2BDC14E261B2C670E77814CC027" italics="true" pageNumber="135">Saguinus melanoleucus</emphasis>
Miranda Ribeiro 1912
</taxonomicName>
,
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<collectingCountry id="9E5E476341442E530A904DBC4944ED85" name="Brazil">Brasil</collectingCountry>
. Rundsch.: 22
</bibCitation>
</treatmentCitation>
.
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<subSubSection id="261D3F149F640DE6EC2E56D24FDDE9C9" pageNumber="135" type="materials_examined">
<paragraph id="F9E8CD271CD55F21E6FD064D72B6BD7A" pageNumber="135">
<emphasis id="7522D3E06F3F36C3CD7A17CA6F7F076B" inLineHeading="true" pageNumber="135">Type Locality:</emphasis>
<materialsCitation id="0C30E6C91F43BF3BDDA6AD88C3467C74" ID-GBIF-Occurrence="4867331665" country="Brazil" pageNumber="135" stateProvince="Amazonas">
<collectingCountry id="DF2F0817B905C933638C55C788115F6C" name="Brazil">Brazil</collectingCountry>
,
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, Santo Antonio, Rio Eiru, upper Rio Jurua.
</materialsCitation>
</paragraph>
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<paragraph id="5FFA4F27B2770CA3AE1678DFC764098E" pageNumber="135">
<emphasis id="28BCDA0DEAB37BC1C71AB18C1119EDD5" inLineHeading="true" pageNumber="135">Vernacular Names:</emphasis>
<vernacularName id="9CC5F42A88ED4035E59BB7E59AA1C458" language="eng" pageNumber="135">White-mantled Tamarin</vernacularName>
.
</paragraph>
</subSubSection>
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<paragraph id="218B545E91BB5B07D1E43F647BD0B738" pageNumber="135">
<emphasis id="E2F706AA0100228DE72A2D1B0AAB3638" inLineHeading="true" pageNumber="135">Synonyms:</emphasis>
<taxonomicName id="5A070B6E4EF854B70B6157860F3C1050" authority="(Carvalho 1957)" baseAuthorityName="Carvalho" baseAuthorityYear="1957" class="Mammalia" family="Cebidae" genus="Saguinus" kingdom="Animalia" order="Primates" pageNumber="135" phylum="Chordata" rank="species" species="acrensis">
<emphasis id="016199B92CD3869D8BA6E917A56ED6F6" italics="true" pageNumber="135">Saguinus acrensis</emphasis>
(Carvalho 1957)
</taxonomicName>
;
<taxonomicName id="29E4E94AC6A06DF4C19C64D70484A13E" authority="Hershkovitz 1966" authorityName="Hershkovitz" authorityYear="1966" class="Mammalia" family="Cebidae" genus="Saguinus" kingdom="Animalia" order="Primates" pageNumber="135" phylum="Chordata" rank="species" species="crandalli">
<emphasis id="791AB571226D4F97A1D42C1E917CF540" italics="true" pageNumber="135">Saguinus crandalli</emphasis>
Hershkovitz 1966
</taxonomicName>
;
<taxonomicName id="974EC2BF2646C03EE2775614ED03B1E4" authority="(Pinto 1937)" baseAuthorityName="Pinto" baseAuthorityYear="1937" class="Mammalia" family="Cebidae" genus="Saguinus" kingdom="Animalia" order="Primates" pageNumber="135" phylum="Chordata" rank="species" species="hololeucus">
<emphasis id="60B87E17CC5194F58F46A732A96453CB" italics="true" pageNumber="135">Saguinus hololeucus</emphasis>
(Pinto 1937)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C1A5B12805A1B32C34DB5672D1686B1C" pageNumber="135" type="distribution">
<paragraph id="293B8FD55253E8B9E3807091FBC94992" pageNumber="135">
<emphasis id="4E9348B127BAF625641C4FC9BCFCA455" inLineHeading="true" pageNumber="135">Distribution:</emphasis>
<collectingCountry id="53AFD242C3E137CDCADD994EBA1D0A00" name="Brazil">Brazil</collectingCountry>
, between Rios Jurua and Tarauca.
</paragraph>
</subSubSection>
<subSubSection id="652F62B33538C280E2D4F36A67DC32BF" pageNumber="135" type="conservation">
<paragraph id="104675AC8C1A30DC0A73151EF7FB2E87" pageNumber="135">
<emphasis id="E213CD75D03A24B7C3C279CB9A0928E4" inLineHeading="true" pageNumber="135">Conservation:</emphasis>
<collectionCode id="6C50D939384D3956232E882337A8B413">CITES</collectionCode>
Appendix II;
<collectionCode id="3A9E5491DA719F73176A7F9D4EA8378D">IUCN</collectionCode>
Data Deficient as
<taxonomicName id="8CDC629F11F2CBE216FDEB399335C310" authorityName="Hershkovitz" authorityYear="1966" class="Mammalia" family="Callitrichidae" genus="Saguinus" kingdom="Animalia" order="Primates" phylum="Chordata" rank="subSpecies" species="fuscicolis" subSpecies="crandalli">
<emphasis id="940DB95E07A22DC580BEAC1268977AC7" italics="true" pageNumber="135">S. fuscicolis crandalli</emphasis>
</taxonomicName>
, otherwise not listed.
</paragraph>
</subSubSection>
<subSubSection id="2D0B4C76816B28D447E345BCB9B37F8A" pageNumber="135" type="discussion">
<paragraph id="1AB77E9B36A560BF48048C94A8219A9A" pageNumber="135">
<emphasis id="DBCCF17BFA035FB6994D6E8F592DC216" inLineHeading="true" pageNumber="135">Discussion:</emphasis>
<taxonomicName id="1E124F046B4E2097084FAE3336FB6188" authorityName="Spix" authorityYear="1823" class="Mammalia" family="Callitrichidae" genus="Saguinus" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="nigricollis">
<emphasis id="390717F1C588AE4BF481D82CDE68E95D" italics="true" pageNumber="135">S. nigricollis</emphasis>
</taxonomicName>
species group.
<bibRefCitation id="82F613A44D95FCBD37206C228534533F" author="Coimbra-Filho, A. F." refId="ref63910" refString="Coimbra-Filho, A. F. 1990. Sistematica, distribucao geografica e situacao atual dos simios Brasileiros (Platyrrhini-Primates). Revista Brasileira de Biologia, 50: 1063 - 1079." year="1990">Coimbra-Filho (1990)</bibRefCitation>
regarded
<taxonomicName id="7AC5285E31E807104466AE77EFE7EA47" authorityName="Miranda Ribeiro" authorityYear="1912" class="Mammalia" family="Callitrichidae" genus="Saguinus" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="32D0216C778E33059583F76F78C02FFC" italics="true" pageNumber="135">melanoleucus</emphasis>
</taxonomicName>
as a species distinct from
<taxonomicName id="3EF9B4113486F446E63E7296C58370C1" authorityName="Spix" authorityYear="1823" class="Mammalia" family="Callitrichidae" genus="Saguinus" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="fuscicollis">
<emphasis id="B9ECD63F5AC1C26732B93CFC70BC2A8A" italics="true" pageNumber="135">S. fuscicollis</emphasis>
</taxonomicName>
, and this was followed by
<bibRefCitation id="FDCC3DA2F93167EC45B88A9ED2E801B2" author="Groves, C. P." refId="ref132585" refString="Groves, C. P. 2001 c. Primate taxonomy. Smithsonian Institution Press, Washington, DC, 350 pp." year="2001">
Groves (2001
<emphasis id="01D4850FE701B4B7EEE7D410ADE4F44E" italics="true" pageNumber="135">c</emphasis>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:namePart id="7F49456665CB206A5118AEBF0BFA4902">Deyrup, Mark</mods:namePart>
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<mods:namePart id="453FBFD93451DC4C6BBA9B0B7E694CF6">Sorger, D. Magdalena</mods:namePart>
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<mods:part id="76A3C468B16DDDC4DC44188F43F42A92">
<mods:date id="1065F8C655D5A619417DA52E0192776F">2014</mods:date>
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<mods:identifier id="0F04751055D6957DEAD9F61934F8E0AD" type="DOI">10.11646/zootaxa.3802.4.6</mods:identifier>
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<treatment id="C66787A55B22FF9F9AE0FCE4FB63F9CC" ID-DOI="http://doi.org/10.5281/zenodo.6129408" ID-GBIF-Taxon="119429108" ID-Zenodo-Dep="6129408" LSID="urn:lsid:plazi:treatment:C66787A55B22FF9F9AE0FCE4FB63F9CC" httpUri="http://treatment.plazi.org/id/C66787A55B22FF9F9AE0FCE4FB63F9CC" lastPageId="14" lastPageNumber="529" pageId="11" pageNumber="526">
<subSubSection id="06D465385B22FF9A9AE0FCE4FDEDFC5E" pageId="11" pageNumber="526" type="nomenclature">
<paragraph id="4E7136B35B22FF9A9AE0FCE4FD6DFCBC" blockId="11.[151,640,862,922]" box="[151,640,862,888]" pageId="11" pageNumber="526">
<heading id="153981DF5B22FF9A9AE0FCE4FD6DFCBC" bold="true" box="[151,640,862,888]" fontSize="11" level="1" pageId="11" pageNumber="526" reason="1">
<taxonomicName id="89CE4D305B22FF9A9AE0FCE4FD6DFCBC" ID-CoL="6SH5N" authority="Patton, 1894" authorityName="Patton" authorityYear="1894" box="[151,640,862,888]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9AE0FCE4FD6DFCBC" bold="true" box="[151,640,862,888]" pageId="11" pageNumber="526">
<emphasis id="7CBAEAA15B22FF9A9AE0FCE4FE27FCBC" bold="true" box="[151,458,862,888]" italics="true" pageId="11" pageNumber="526">Odontomachus brunneus</emphasis>
(
<bibRefCitation id="2A5F4B425B22FF9A9BAEFCE4FD95FCBC" author="Patton" box="[473,632,862,888]" pageId="11" pageNumber="526" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton, 1894</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4E7136B35B22FF9A9AE0FC3BFDEDFC5E" blockId="11.[151,640,862,922]" box="[151,512,897,922]" pageId="11" pageNumber="526">
(
<figureCitation id="D6F52A365B22FF9A9AE8FC3BFF03FC5D" box="[159,238,897,922]" captionStart="FIGURE 4" captionStartId="7.[151,249,1425,1447]" captionTargetBox="[151,1435,486,1403]" captionTargetId="figure@7.[151,1435,486,1403]" captionTargetPageId="7" captionText="FIGURE 4. First gastral tergites of workers, dorsal view: (A) O. brunneus and (B) O. clarus." httpUri="https://zenodo.org/record/249900/files/figure.png" pageId="11" pageNumber="526">Figs. 4</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9A8DFC38FEE7FC5D" box="[250,266,898,921]" captionStart="FIGURE 7" captionStartId="9.[151,249,1080,1102]" captionTargetBox="[151,1435,410,1059]" captionTargetId="figure@9.[151,1435,410,1059]" captionTargetPageId="9" captionText="FIGURE 7. Male heads, full-face view: (A) O. brunneus and (B) O. haematodus. LOC = lateral ocelli, MOC = median ocellus." httpUri="https://zenodo.org/record/249903/files/figure.png" pageId="11" pageNumber="526">7</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9B61FC3BFEC8FC5D" box="[278,293,897,921]" captionStart="FIGURE 8" captionStartId="9.[151,249,1900,1922]" captionTargetBox="[151,1435,1331,1878]" captionTargetId="figure@9.[151,1435,1331,1879]" captionTargetPageId="9" captionText="FIGURE 8. Male mesoscuta, dorsal view: (A) O. brunneus (inset shows transverse striae) and (B) O. relictus (inset shows longitudinal striae)." httpUri="https://zenodo.org/record/249904/files/figure.png" pageId="11" pageNumber="526">8</figureCitation>
, 12, 13, 24, 25, 26)
</paragraph>
</subSubSection>
<subSubSection id="06D465385B22FF9A9AE0FC70FED1FB9E" pageId="11" pageNumber="526" type="reference_group">
<paragraph id="4E7136B35B22FF9A9AE0FC70FD21FC1B" blockId="11.[151,1436,969,1114]" box="[151,716,969,991]" pageId="11" pageNumber="526">
<treatmentCitationGroup id="6EDE119D5B22FF9A9AE0FC70FD21FC1B" box="[151,716,969,991]" pageId="11" pageNumber="526">
<taxonomicName id="89CE4D305B22FF9A9AE0FC70FE22FC1B" authority="Patton 1894: 618" authorityName="Patton" authorityPageNumber="618" authorityYear="1894" box="[151,463,969,991]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="7CBAEAA15B22FF9A9AE0FC70FEF6FC1B" box="[151,283,969,991]" italics="true" pageId="11" pageNumber="526">Atta brunnea</emphasis>
<treatmentCitation id="CF6F10A25B22FF9A9B55FC70FE22FC1B" author="Patton" box="[290,463,969,991]" page="618" pageId="11" pageNumber="526" year="1894">
<bibRefCitation id="2A5F4B425B22FF9A9B55FC70FE72FC1B" author="Patton" box="[290,415,969,991]" pageId="11" pageNumber="526" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton 1894</bibRefCitation>
: 618
</treatmentCitation>
</taxonomicName>
(worker)
<collectingCountry id="36D976235B22FF9A9843FC73FD65FC1B" box="[564,648,969,991]" name="Georgia" pageId="11" pageNumber="526">Georgia</collectingCountry>
,
<collectingCountry id="36D976235B22FF9A98E5FC73FD25FC1B" box="[658,712,969,991]" name="United States of America" pageId="11" pageNumber="526">USA</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B22FF9A9AE0FC52FED1FB9E" blockId="11.[151,1436,969,1114]" pageId="11" pageNumber="526">
<treatmentCitationGroup id="6EDE119D5B22FF9A9AE0FC52FED1FB9E" pageId="11" pageNumber="526">
<taxonomicName id="89CE4D305B22FF9A9AE0FC52FDDDFC3A" ID-CoL="6SH5N" authority="Patton, 1894" authorityName="Patton" authorityYear="1894" box="[151,560,1000,1022]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9AE0FC52FE78FC3A" box="[151,405,1000,1022]" italics="true" pageId="11" pageNumber="526">Odontomachus brunneus</emphasis>
(
<bibRefCitation id="2A5F4B425B22FF9A9BD4FC52FDC4FC3A" author="Patton" box="[419,553,1000,1022]" pageId="11" pageNumber="526" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton, 1894</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B22FF9A984CFC53FD1CFC3A" author="Emery" box="[571,753,1000,1022]" page="268" pageId="11" pageNumber="526" year="1895">
<bibRefCitation id="2A5F4B425B22FF9A984CFC53FD2CFC3A" author="Emery" box="[571,705,1000,1022]" pageId="11" pageNumber="526" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery, 1895</bibRefCitation>
: 268
</treatmentCitation>
(as junior synonym of
<taxonomicName id="89CE4D305B22FF9A9996FC52FF13FBD8" ID-CoL="48QBW" authority="Guerin-Meneville, 1844" authorityName="Guerin-Meneville" authorityYear="1844" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="7CBAEAA15B22FF9A9996FC52FB37FC3A" box="[993,1242,1000,1022]" italics="true" pageId="11" pageNumber="526">Odontomachus insularis</emphasis>
<bibRefCitation id="2A5F4B425B22FF9A9E97FC52FF13FBD8" author="Guerin-Meneville" pageId="11" pageNumber="526" refString="Guerin-Meneville, F. E. (1844) Iconographie du regne animal de G. Cuvier, ou representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees, de chaque genre d'animaux. Insectes. J. B. Bailliere, Paris, 576 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10331" type="book" year="1844">Guérin-Méneville, 1844</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="CF6F10A25B22FF9A9B78FBBDFE24FBD8" author="Brown" box="[271,457,1030,1052]" page="103" pageId="11" pageNumber="526" year="1976">
<bibRefCitation id="2A5F4B425B22FF9A9B78FBBDFE75FBD8" author="Brown" box="[271,408,1030,1052]" pageId="11" pageNumber="526" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
, 139 (as senior synonym of
<taxonomicName id="89CE4D305B22FF9A9891FBBDFCA9FBD8" box="[742,836,1030,1052]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="7CBAEAA15B22FF9A9891FBBDFCA9FBD8" box="[742,836,1030,1052]" italics="true" pageId="11" pageNumber="526">O. fuscus</emphasis>
</taxonomicName>
);
<treatmentCitation id="CF6F10A25B22FF9A992DFBBDFB2AFBD8" author="Deyrup" box="[858,1223,1030,1052]" page="191" pageId="11" pageNumber="526" year="1985">
<bibRefCitation id="2A5F4B425B22FF9A992DFBBDFB7AFBD8" author="Deyrup" box="[858,1175,1030,1052]" pageId="11" pageNumber="526" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article" year="1985">Deyrup, Trager &amp; Carlin, 1985</bibRefCitation>
: 191
</treatmentCitation>
(male).
<taxonomicName id="89CE4D305B22FF9A9AE0FB9FFDCDFBFF" authority="Stitz, 1925:115" authorityName="Stitz" authorityPageNumber="115" authorityYear="1925" box="[151,544,1061,1083]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="7CBAEAA15B22FF9A9AE0FB9FFE95FBFF" box="[151,376,1061,1083]" italics="true" pageId="11" pageNumber="526">Odontomachus fuscus</emphasis>
<treatmentCitation id="CF6F10A25B22FF9A9BF6FB9FFDCDFBFF" author="Stitz" box="[385,544,1061,1083]" page="115" pageId="11" pageNumber="526" year="1925">
<bibRefCitation id="2A5F4B425B22FF9A9BF6FB9FFE18FBFF" author="Stitz" box="[385,501,1061,1083]" pageId="11" pageNumber="526" refString="Stitz, H. (1925) Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1923, 110 - 136." type="journal article" year="1925">Stitz, 1925</bibRefCitation>
:115
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B22FF9A985AFB9CFD01FBFF" author="Brown" box="[557,748,1061,1083]" page="103" pageId="11" pageNumber="526" year="1976">
<bibRefCitation id="2A5F4B425B22FF9A985AFB9CFD55FBFF" author="Brown" box="[557,696,1061,1083]" pageId="11" pageNumber="526" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
, 139 (unresolved junior homonym of
<taxonomicName id="89CE4D305B22FF9A9E0AFB9FFEDCFB9E" authority="Stitz, 1916: 372" authorityName="Stitz" authorityPageNumber="372" authorityYear="1916" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="7CBAEAA15B22FF9A9E0AFB9FFAB3FBFF" box="[1149,1374,1061,1083]" italics="true" pageId="11" pageNumber="526">Odontomachus fuscus</emphasis>
<treatmentCitation id="CF6F10A25B22FF9A9F1FFB9FFEDCFB9E" author="Stitz" page="372" pageId="11" pageNumber="526" year="1916">
<bibRefCitation id="2A5F4B425B22FF9A9F1FFB9FFF10FB9E" author="Stitz" pageId="11" pageNumber="526" refString="Stitz, H. (1916) Formiciden. Ergebnisse der Zweiten Deutschen Zentral-Afrika Expedition, 1, 369 - 405." type="journal article" year="1916">Stitz, 1916</bibRefCitation>
: 372
</treatmentCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="06D465385B22FF9A9AE0FB3CFF02F98B" pageId="11" pageNumber="526" type="diagnosis">
<paragraph id="4E7136B35B22FF9A9AE0FB3CFF02F98B" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="526">
<emphasis id="7CBAEAA15B22FF9A9AE0FB3CFEFCFB5B" bold="true" box="[151,273,1158,1183]" pageId="11" pageNumber="526">Diagnosis.</emphasis>
Workers of this species are distinguished from others in our region by the much finer and denser pubescence on the first gastral tergite. Males are yellow-colored and most easily confused with those of
<taxonomicName id="89CE4D305B22FF9A9FF6FB11FEF3FB22" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B22FF9A9FF6FB11FEF3FB22" italics="true" pageId="11" pageNumber="526">O. haematodus</emphasis>
</taxonomicName>
; those of
<taxonomicName id="89CE4D305B22FF9A9BE1FB75FDC9FB22" box="[406,548,1230,1254]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9BE1FB75FDC9FB22" box="[406,548,1230,1254]" italics="true" pageId="11" pageNumber="526">O. brunneus</emphasis>
</taxonomicName>
are clearly separated by the following characters: 1) ocelli larger, projecting beyond the posterior border of the head; 2) metasternal processes short, not elongate and spine-like; 3) posterior face of propodeum not offset from dorsal and lateral faces by distinct carina, and by numerous genitalic characters (see
<figureCitation id="D6F52A365B22FF9A9AB9FA80FEC0FA97" box="[206,301,1338,1363]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="11" pageNumber="526">Figs. 24</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9B4BFA80FEB6FA97" box="[316,347,1338,1363]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="11" pageNumber="526">25</figureCitation>
). Male
<taxonomicName id="89CE4D305B22FF9A9BC1FA81FDA9FA97" box="[438,580,1339,1363]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9BC1FA81FDA9FA97" box="[438,580,1339,1363]" italics="true" pageId="11" pageNumber="526">O. brunneus</emphasis>
</taxonomicName>
are further separated from other
<collectingCountry id="36D976235B22FF9A99BEFA81FC00FA97" box="[969,1005,1339,1363]" name="United States of America" pageId="11" pageNumber="526">US</collectingCountry>
species by the transverse mesoscutal striae, and by the anteroposteriorly slender mesosoma. Genitalia of
<taxonomicName id="89CE4D305B22FF9A99D5FAE5FBC2FAB2" box="[930,1071,1374,1398]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A99D5FAE5FBC2FAB2" box="[930,1071,1374,1398]" italics="true" pageId="11" pageNumber="526">O. brunneus</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="89CE4D305B22FF9A9FF6FAE5FEEEFA5F" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9FF6FAE5FEEEFA5F" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B22FF9A9B40FA39FE73FA5F" box="[311,414,1411,1435]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9B40FA39FE73FA5F" box="[311,414,1411,1435]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
, but are distinguished from these two species by the following characters (see
<figureCitation id="D6F52A365B22FF9A9F7FFA38FA89FA5F" box="[1288,1380,1410,1435]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="11" pageNumber="526">Figs. 24</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9F18FA38FA62FA5F" box="[1391,1423,1410,1435]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="11" pageNumber="526">25</figureCitation>
): abdominal sternum IX narrower and longer than
<taxonomicName id="89CE4D305B22FF9A98B5FA1DFCC1FA7A" box="[706,812,1446,1470]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A98B5FA1DFCC1FA7A" box="[706,812,1446,1470]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
, and broader than
<taxonomicName id="89CE4D305B22FF9A9E76FA1DFB62FA7A" box="[1025,1167,1446,1470]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9E76FA1DFB62FA7A" box="[1025,1167,1446,1470]" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
; telomere apex broadly rounded, posterodorsal margin nearly linear (apex narrower and posterodorsal margin sinuate in
<taxonomicName id="89CE4D305B22FF9A9EA9FA71FAAAFA27" box="[1246,1351,1483,1507]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9EA9FA71FAAAFA27" box="[1246,1351,1483,1507]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B22FF9A9FF6FA71FEEFF9C2" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9FF6FA71FEEFF9C2" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
); valviceps apicodorsal lobe broad (narrow in
<taxonomicName id="89CE4D305B22FF9A9963FA55FC91F9C2" box="[788,892,1518,1542]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9963FA55FC91F9C2" box="[788,892,1518,1542]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
), subapical lamina narrow (broad
<taxonomicName id="89CE4D305B22FF9A9F74FA55FA86F9C2" box="[1283,1387,1518,1542]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9F74FA55FA86F9C2" box="[1283,1387,1518,1542]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B22FF9A9AE0F9A9FECAF9EF" box="[151,295,1555,1579]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9AE0F9A9FECAF9EF" box="[151,295,1555,1579]" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
).
<taxonomicName id="89CE4D305B22FF9A9B34F9A9FDB3F9EF" box="[323,606,1555,1579]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9B34F9A9FDB3F9EF" box="[323,606,1555,1579]" italics="true" pageId="11" pageNumber="526">Odontomachus brunneus</emphasis>
</taxonomicName>
has the proportionally shortest (dorsoventral) valviceps of the Nearctic species.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B22FF9F9AB0F9E0FDD4FD6C" lastPageId="14" lastPageNumber="529" pageId="11" pageNumber="526" type="description">
<paragraph id="4E7136B35B22FF9D9AB0F9E0FB43FED8" blockId="11.[151,1437,1158,2011]" lastBlockId="12.[151,1436,151,285]" lastPageId="12" lastPageNumber="527" pageId="11" pageNumber="526">
<emphasis id="7CBAEAA15B22FF9A9AB0F9E0FEB4F9B7" bold="true" box="[199,345,1626,1651]" pageId="11" pageNumber="526">Description.</emphasis>
Worker: HL 2.102.44, HW 1.581.92, SL 1.842.10, EL 0.340.42, ML 1.161.36, WL 2.443.00, PTH 0.961.20, PTL 0.480.52 (n=10). Entire body generally shiny except where obscured by dense pubescence; head, mesosoma, and petiole dark reddish-brown to dark brown; gaster darker brown; scape and legs brown. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; dorsally with numerous, fine, appressed pubescence and scattered elongate, erect setae present. Pronotum with sub-circular, concentric striae that become longitudinal near rear margin; pubescence appressed, abundant; 58 elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; propleuron, mesopleuron, and basalar lobe lacking sculpture; pubescence abundant dorsally, appressed. Metasternum lacking paired, elongate, spiniform processes between hind coxae. Petiole widest at base, gradually tapering apically to a short spine directed rearward; mostly lacking striae with only faint striae present near base; subpetiolar process rounded triangular; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but with fine coriaceous sculpture (seen at high magnification); fine, appressed pubescence dense, spaces between hairs less than 1/3 the length of a hair, often overlapping one another; scattered erect, elongate setae present.
</paragraph>
<paragraph id="4E7136B35B25FF9D9AE0F8A4FE82F8B6" pageId="12" pageNumber="527">
<table id="3CCEC4135B25006E9AE0F8A4FA71F8B6" box="[151,1436,1822,1906]" gridcols="4" gridrows="3" pageId="12" pageNumber="527">
<tr id="F0FE34F15B25006E9AE0F8A4FA71F8F0" box="[151,1436,1822,1844]" gridrow="0" pageId="12" pageNumber="527">
<th id="B32F5D8D5B25006E9AE0F8A4FDD7F8F0" box="[151,570,1822,1844]" gridcol="0" gridrow="0" pageId="12" pageNumber="527">
<emphasis id="7CBAEAA15B25FF9D9AE0F8A4FEC8F8F0" bold="true" box="[151,293,1822,1844]" pageId="12" pageNumber="527">FIGURE 12.</emphasis>
<taxonomicName id="89CE4D305B25FF9D9B46F8A4FDD9F8F0" box="[305,564,1822,1844]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="527" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B25FF9D9B46F8A4FDD9F8F0" box="[305,564,1822,1844]" italics="true" pageId="12" pageNumber="527">Odontomachus brunneus</emphasis>
</taxonomicName>
,
</th>
<th id="B32F5D8D5B25006E9836F8A4FD76F8F0" box="[577,667,1822,1844]" gridcol="1" gridrow="0" pageId="12" pageNumber="527">worker:</th>
<th id="B32F5D8D5B25006E98D5F8A4FBADF8F0" box="[674,1088,1822,1844]" gridcol="2" gridrow="0" pageId="12" pageNumber="527">(A) head in full-face view, (B) lateral</th>
<th id="B32F5D8D5B25006E9E30F8A4FA71F8F0" box="[1095,1436,1822,1844]" gridcol="3" gridrow="0" pageId="12" pageNumber="527">habitus, (C) dorsal habitus, (D)</th>
</tr>
<tr id="F0FE34F15B25006E9AE0F887FA71F897" box="[151,1436,1853,1875]" gridrow="1" pageId="12" pageNumber="527">
<th id="B32F5D8D5B25006E9AE0F887FDD7F897" box="[151,570,1853,1875]" gridcol="0" gridrow="1" pageId="12" pageNumber="527">mesosoma in dorsal view, (E) mesosoma</th>
<td id="B32F5D8D5B25006E9836F887FD76F897" box="[577,667,1853,1875]" gridcol="1" gridrow="1" pageId="12" pageNumber="527">in lateral</td>
<td id="B32F5D8D5B25006E98D5F887FBADF897" box="[674,1088,1853,1875]" gridcol="2" gridrow="1" pageId="12" pageNumber="527">view, (F) petiole in lateral view, and (G)</td>
<td id="B32F5D8D5B25006E9E30F887FA71F897" box="[1095,1436,1853,1875]" gridcol="3" gridrow="1" pageId="12" pageNumber="527">first gastral tergite in dorsal view</td>
</tr>
<tr id="F0FE34F15B25006E9AE0F8E6FA71F8B6" box="[151,1436,1884,1906]" gridrow="2" pageId="12" pageNumber="527" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="B32F5D8D5B25006E9AE0F8E6FDD7F8B6" box="[151,570,1884,1906]" gridcol="0" gridrow="2" pageId="12" pageNumber="527">showing pubescence.</th>
</tr>
</table>
</paragraph>
<caption id="1AB1663B5B24FF9C9AE0F892FB92F899" httpUri="https://zenodo.org/record/249908/files/figure.png" pageId="13" pageNumber="528" targetBox="[151,1436,193,1810]" targetPageId="13">
<paragraph id="4E7136B35B24FF9C9AE0F892FB92F899" blockId="13.[151,1436,1832,1885]" pageId="13" pageNumber="528">
<emphasis id="7CBAEAA15B24FF9C9AE0F892FEF2F8FA" bold="true" box="[151,287,1832,1854]" pageId="13" pageNumber="528">FIGURE 13.</emphasis>
<taxonomicName id="89CE4D305B24FF9C9B51F892FDCEF8FA" box="[294,547,1832,1854]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B24FF9C9B51F892FDCEF8FA" box="[294,547,1832,1854]" italics="true" pageId="13" pageNumber="528">Odontomachus brunneus</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<paragraph id="4E7136B35B24FF9F9AB0F83EFA82FD9B" blockId="13.[151,1437,1924,2021]" lastBlockId="14.[151,1437,151,1544]" lastPageId="14" lastPageNumber="529" pageId="13" pageNumber="528">Male: HL 1.001.08, HW 1.241.34, SL 0.180.22, EL 0.680.76, EW 0.400.44, OL 0.24, OES 0.16, WL 2.522.76, PTH 0.720.88, PTL 0.460.54, FWL 5.105.65 (n=5). Head, mesosoma, and petiole generally shiny except where obscured by dense pubescence; head, meso- and metasoma, legs, scape and pedicel yellow to yellowish-brown, remainder of funiculus brown. Head and mesosoma with abundant fine, white pubescence except on anepisternum where pubescence is mostly absent. Eyes extremely large, maximum diameter of each eye at least 70% of the length of the head in full-face view. Ocelli large, the length of each ocellus wider than distance between lateral ocellus and eye margin; in full-face view, lateral ocelli protrude beyond posterior border of head. Mesosoma: pronotum lacking sculpture; mesoscutum with fine transversely arcuate striae anteriorly, striae becoming longitudinal posteriorly; mesoscutellum raised and convex, lacking sculpture; propodeum with weak to moderately strong transverse striae laterally, and especially posterodorsally; mesopleuron mostly lacking striae. Petiole bluntly rounded apically, with rounded triangular subpetiolar process anteriorly; densely pubescent anteriorly and laterally, but reduced pubescence posteriorly. Disc of abdominal sternum IX trapezoidal, breadth slightly greater than length; posterior lobe length approximately twice maximum lobe width, basal half narrowed. Telomeral apex broadly rounded; telomere length distinctly greater than height; valviceps ventral apex strongly produced, narrow; valviceps apical margin broadly concave; vertical portion of dorsolateral carina and lateral margin of subapical lamina meeting at an angle; apicodorsal lobe of valviceps broad; subapical lamina narrow.</paragraph>
<paragraph id="4E7136B35B27FF9F9AB0FDD6FDD4FD6C" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">Queen: No specimens measured, but similar to workers in color and general appearance except slightly larger, with mesosoma developed for wings.</paragraph>
</subSubSection>
<subSubSection id="06D465385B27FF9F9AB0FD0EFED8FBD4" pageId="14" pageNumber="529" type="distribution">
<paragraph id="4E7136B35B27FF9F9AB0FD0EFED8FBD4" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">
<emphasis id="7CBAEAA15B27FF9F9AB0FD0EFDF2FD09" bold="true" box="[199,543,692,717]" pageId="14" pageNumber="529">
Distribution (
<figureCitation id="D6F52A365B27FF9F9B18FD0EFE26FD09" box="[367,459,692,717]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="14" pageNumber="529">Fig. 26</figureCitation>
).
<collectingCountry id="36D976235B27FF9F9B90FD0EFDF2FD09" box="[487,543,692,717]" name="United States of America" pageId="14" pageNumber="529">USA</collectingCountry>
</emphasis>
. Alabama: Baldwin and Houston Counties. Florida: Alachua,
<collectingCountry id="36D976235B27FF9F9F65FD0EFABAFD09" box="[1298,1367,692,717]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="529">Baker</collectingCountry>
, Bay, Bradford, Broward, Citrus, Clay, Collier,
<collectingCountry id="36D976235B27FF9F9818FD6DFD0DFD34" box="[623,736,727,752]" name="United States of America" pageId="14" pageNumber="529">Columbia</collectingCountry>
, Dade, De Soto, Duval, Franklin, Gadsden, Gilchrist, Glades, Hamilton, Hendry, Hernando, Highlands, Hillsborough, Indian River, Jackson, Jefferson, Lake, Lee, Leon, Levy, Liberty, Madison, Marion, Martin, Monroe, Nassau, Okeechobee, Orange, Osceola, Palm Beach, Pasco, Polk, Putnam, Sarasota, St. Lucie, Sumter, Taylor, Volusia, Wakulla, and Walton Counties.
<collectingCountry id="36D976235B27FF9F9E1DFCFEFB2EFC99" box="[1130,1219,836,861]" name="Georgia" pageId="14" pageNumber="529">Georgia</collectingCountry>
: Chatham, Clinch, Dougherty, Glynn, Lowndes, and Ware Counties. Mississippi: Harrison County. According to
<bibRefCitation id="2A5F4B425B27FF9F9EBEFCD2FF09FC60" author="Deyrup" pageId="14" pageNumber="529" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
, the occurrence of this species in the Caribbean region and Central and South
<collectingCountry id="36D976235B27FF9F9E05FC36FB3FFC61" box="[1138,1234,908,933]" name="United States of America" pageId="14" pageNumber="529">America</collectingCountry>
is unclear due to historical misidentifications and nomenclatural changes. Distributional information from
<bibRefCitation id="2A5F4B425B27FF9F9E0AFC0AFA71FC0C" author="Deyrup" box="[1149,1436,943,968]" pageId="14" pageNumber="529" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
and examination of specimens from the ABS, MEM, UGA, USNM, and personal collections of Dan Suitor and Doug Booher.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B27FF9F9AB0FBA6FB63F9CC" pageId="14" pageNumber="529" type="discussion">
<paragraph id="4E7136B35B27FF9F9AB0FBA6FD29FB2C" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">
<emphasis id="7CBAEAA15B27FF9F9AB0FBA6FEA1FBF1" bold="true" box="[199,332,1052,1077]" pageId="14" pageNumber="529">Discussion.</emphasis>
<taxonomicName id="89CE4D305B27FF9F9B23FBA6FD82FBF0" box="[340,623,1052,1076]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9B23FBA6FD82FBF0" box="[340,623,1052,1076]" italics="true" pageId="14" pageNumber="529">Odontomachus brunneus</emphasis>
</taxonomicName>
appears to be restricted to the southeastern
<collectingCountry id="36D976235B27FF9F9E10FBA6FB63FBF0" box="[1127,1166,1052,1076]" name="United States of America" pageId="14" pageNumber="529">US</collectingCountry>
. Previous records of
<taxonomicName id="89CE4D305B27FF9F9FF6FBA6FF12FB9C" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9FF6FBA6FF12FB9C" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
from the Caribbean, and Central and South
<collectingCountry id="36D976235B27FF9F9885FB85FCBEFB9C" box="[754,851,1087,1112]" name="United States of America" pageId="14" pageNumber="529">America</collectingCountry>
(
<bibRefCitation id="2A5F4B425B27FF9F9915FBFAFC1DFB9C" author="Brown" box="[866,1008,1087,1112]" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown 1976</bibRefCitation>
) all appear to be of
<taxonomicName id="89CE4D305B27FF9F9EA3FBFAFA8CFB9C" box="[1236,1377,1088,1112]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B27FF9F9EA3FBFAFA8CFB9C" box="[1236,1377,1088,1112]" italics="true" pageId="14" pageNumber="529">O. ruginodis</emphasis>
</taxonomicName>
. The situation was clarified by
<bibRefCitation id="2A5F4B425B27FF9F9BBAFBDFFD2FFBB9" box="[461,706,1124,1149]" pageId="14" pageNumber="529" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article">
Deyrup
<emphasis id="7CBAEAA15B27FF9F9859FBDFFD86FBB8" box="[558,619,1124,1148]" italics="true" pageId="14" pageNumber="529">et al.</emphasis>
(1985)
</bibRefCitation>
who recognized the distinction between
<taxonomicName id="89CE4D305B27FF9F9EDAFBDEFAADFBB8" box="[1197,1344,1124,1148]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B27FF9F9EDAFBDEFAADFBB8" box="[1197,1344,1124,1148]" italics="true" pageId="14" pageNumber="529">O. ruginodis</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B27FF9F9FF6FBDEFF12FB64" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9FF6FBDEFF12FB64" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
and revived the former from synonymy with the latter. Browns statement (1976) that
<taxonomicName id="89CE4D305B27FF9F9EB5FB32FAA1FB64" box="[1218,1356,1160,1184]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9EB5FB32FAA1FB64" box="[1218,1356,1160,1184]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
is well adapted to “marginal habitats is consistent with the ecology of
<emphasis id="7CBAEAA15B27FF9F992FFB16FC00FB00" box="[856,1005,1196,1220]" italics="true" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F992FFB16FC04FB00" box="[856,1001,1196,1220]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="ruginodis">O. ruginodis</taxonomicName>
,
</emphasis>
whereas
<taxonomicName id="89CE4D305B27FF9F9E20FB16FB0CFB00" box="[1111,1249,1196,1220]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9E20FB16FB0CFB00" box="[1111,1249,1196,1220]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
in the Southeast is generally found in undisturbed natural habitats.
</paragraph>
<paragraph id="4E7136B35B27FF9F9AB0FB4FFB63F9CC" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">
In the
<collectingCountry id="36D976235B27FF9F9B64FB4EFED4FAC8" box="[275,313,1268,1292]" name="United States of America" pageId="14" pageNumber="529">US</collectingCountry>
,
<taxonomicName id="89CE4D305B27FF9F9B30FB4EFE39FAC8" box="[327,468,1268,1292]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9B30FB4EFE39FAC8" box="[327,468,1268,1292]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
occurs in a wide variety of natural habitats including flatwoods, mesic forests, pine savannas, swamp forests, oak-pine scrub, upland scrub, sandhills, bayheads, edges of seasonal ponds, and elevated tussocks. Nests of
<taxonomicName id="89CE4D305B27FF9F9B1AFA86FE17FA90" box="[365,506,1340,1364]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9B1AFA86FE17FA90" box="[365,506,1340,1364]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
have been found in leaf litter, rotting logs, at tree bases, and in open to partially covered sandy areas. Nest architecture has been explored and discussed by
<bibRefCitation id="2A5F4B425B27FF9F9E50FADAFA75FABC" author="Cerquera" box="[1063,1432,1375,1400]" pageId="14" pageNumber="529" refString="Cerquera, L. M &amp; Tschinkel, W. R. (2010) The nest architecture of the ant Odontomachus brunneus. Journal of Insect Science, 10, 1 - 12. http: // dx. doi. org / 10.1673 / 031.010.6401" type="journal article" year="2010">Cerquera and Tschinkel (2010)</bibRefCitation>
. Workers occasionally forage during the day, but are more active at night. Upon colony disturbance, workers are not aggressive, but instead quickly retreat or vacate the nest. This is in sharp contrast to the aggressive defensive stinging behavior of
<emphasis id="7CBAEAA15B27FF9F9BFFFA76FDD1FA20" box="[392,572,1484,1508]" italics="true" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9BFFFA76FDD5FA20" box="[392,568,1484,1508]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">O. haematodus</taxonomicName>
.
</emphasis>
Alates have been collected from May through December. This broad time frame for alate activity contrasts sharply with the early summer activity of
<emphasis id="7CBAEAA15B27FF9F99AAFA4AFB63F9CC" box="[989,1166,1520,1544]" italics="true" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F99AAFA4AFB67F9CC" box="[989,1162,1520,1544]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">O. haematodus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,474 @@
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<mods:title id="20AB20FD7C46FC2A346BC234912DA27C">A review of the Nearctic Odontomachus (Hymenoptera: Formicidae: Ponerinae) with a treatment of the males</mods:title>
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<treatment id="C66787A55B27FF839AE0F9E6FEF0FBF0" ID-DOI="http://doi.org/10.5281/zenodo.6129410" ID-GBIF-Taxon="119429107" ID-Zenodo-Dep="6129410" LSID="urn:lsid:plazi:treatment:C66787A55B27FF839AE0F9E6FEF0FBF0" httpUri="http://treatment.plazi.org/id/C66787A55B27FF839AE0F9E6FEF0FBF0" lastPageId="18" lastPageNumber="533" pageId="14" pageNumber="529">
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<taxonomicName id="89CE4D305B27FF9F9AE0F9E6FDD1F9B2" ID-CoL="48QBK" authority="Roger, 1861" authorityName="Roger" authorityYear="1861" box="[151,572,1628,1654]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9AE0F9E6FDD1F9B2" bold="true" box="[151,572,1628,1654]" pageId="14" pageNumber="529">
<emphasis id="7CBAEAA15B27FF9F9AE0F9E6FE4CF9B2" bold="true" box="[151,417,1628,1654]" italics="true" pageId="14" pageNumber="529">Odontomachus clarus</emphasis>
<bibRefCitation id="2A5F4B425B27FF9F9BDFF9E6FDD1F9B2" author="Roger" box="[424,572,1628,1654]" pageId="14" pageNumber="529" refString="Roger, J. (1861) Die Ponera - artigen Ameisen (Schluss). Berliner Entomologische Zeitschrift, 5, 1 - 54." type="journal article" year="1861">Roger, 1861</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F9C4FD8FF953" blockId="14.[151,610,1628,1687]" box="[151,610,1662,1687]" pageId="14" pageNumber="529">
(
<figureCitation id="D6F52A365B27FF9F9AE8F9C4FF03F953" box="[159,238,1662,1687]" captionStart="FIGURE 2" captionStartId="6.[151,249,1213,1235]" captionTargetBox="[152,1434,306,1191]" captionTargetId="figure@6.[152,1435,300,1192]" captionTargetPageId="6" captionText="FIGURE 2. Worker petioles, profile view: (A) O. ruginodis and (B) O. clarus." httpUri="https://zenodo.org/record/249898/files/figure.png" pageId="14" pageNumber="529">Figs. 2</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9A8DF9C5FEE7F953" box="[250,266,1663,1687]" captionStart="FIGURE 4" captionStartId="7.[151,249,1425,1447]" captionTargetBox="[151,1435,486,1403]" captionTargetId="figure@7.[151,1435,486,1403]" captionTargetPageId="7" captionText="FIGURE 4. First gastral tergites of workers, dorsal view: (A) O. brunneus and (B) O. clarus." httpUri="https://zenodo.org/record/249900/files/figure.png" pageId="14" pageNumber="529">4</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B61F9C4FEC8F953" box="[278,293,1662,1687]" captionStart="FIGURE 5" captionStartId="8.[151,249,1008,1030]" captionTargetBox="[211,1380,183,992]" captionTargetId="figure@8.[211,1380,183,993]" captionTargetPageId="8" captionText="FIGURE 5. Worker basalar lobes: (A) O. relictus, basalar lobe ovate, with striae (inset at bottom right with simple diagram showing strial pattern of basalar lobe) and (B) O. clarus, basalar lobe spheroid, mostly smooth (inset with simple diagram showing general lack of sculpture of basalar lobe)." httpUri="https://zenodo.org/record/249901/files/figure.png" pageId="14" pageNumber="529">5</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B45F9C4FEAFF953" box="[306,322,1662,1687]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="14" pageNumber="529">6</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B39F9C5FEB3F953" box="[334,350,1663,1687]" captionStart="FIGURE 9" captionStartId="10.[151,249,823,845]" captionTargetBox="[151,1435,324,784]" captionTargetId="figure@10.[151,1435,324,784]" captionTargetPageId="10" captionText="FIGURE 9. Male propodea (Pp), oblique view: (A) O. clarus (rectangular inset shows striate sculpture, arrow indicates lack of obvious propodeal carina) and (B) O. ruginodis (rectangular inset shows reticulated sculpture, arrows point to propodeal carina)." httpUri="https://zenodo.org/record/249905/files/figure.png" pageId="14" pageNumber="529">9</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B1DF9C5FE65F953" box="[362,392,1663,1687]" captionStart="FIGURE 10" captionStartId="10.[151,250,1657,1679]" captionTargetBox="[152,1435,1182,1627]" captionTargetId="figure@10.[152,1435,1182,1627]" captionTargetPageId="10" captionText="FIGURE 10. Male petiolar sterna (ps), profile view: (A) O. desertorum, left arrow indicates subrectangular subpetiolar process (spp), middle arrow indicates the angular posterior process of the petiolar sternum, and right arrow points to the depressed region behind the posterior angled process; and (B) O. clarus, left arrow indicates triangular subpetiolar process, middle arrow points to tubercles with setae, and right arrow indicates the lack of angular posterior process and accompanying shallow depression." httpUri="https://zenodo.org/record/249906/files/figure.png" pageId="14" pageNumber="529">10</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9BE3F9C5FE5FF953" box="[404,434,1663,1687]" captionStart="FIGURE 14" captionStartId="15.[151,249,1651,1673]" captionTargetBox="[151,1436,195,1629]" captionTargetId="figure@15.[151,1436,195,1629]" captionTargetPageId="15" captionText="FIGURE 14. Odontomachus clarus, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in lateral view, (E) mesosoma in dorsal view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence." httpUri="https://zenodo.org/record/249909/files/figure.png" pageId="14" pageNumber="529">14</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9BC9F9C4FE31F953" box="[446,476,1662,1687]" captionStart="FIGURE 15" captionStartId="16.[151,249,1908,1930]" captionTargetBox="[151,1436,409,1886]" captionTargetId="figure@16.[151,1436,409,1886]" captionTargetPageId="16" captionText="FIGURE 15. Odontomachus clarus, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view." httpUri="https://zenodo.org/record/249910/files/figure.png" pageId="14" pageNumber="529">15</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B9FF9C5FDEBF953" box="[488,518,1663,1687]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="14" pageNumber="529">24</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9865F9C4FDDCF953" box="[530,561,1662,1687]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="14" pageNumber="529">25</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F984BF9C4FDB4F953" box="[572,601,1662,1687]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="14" pageNumber="529">26</figureCitation>
)
</paragraph>
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<emphasis id="7CBAEAA15B27FF9F9AE0F97CFE98F918" box="[151,373,1734,1756]" italics="true" pageId="14" pageNumber="529">Odontomachus clarus</emphasis>
<treatmentCitation id="CF6F10A25B27FF9F9B0CF97DFDF1F918" author="Roger" box="[379,540,1734,1756]" page="26" pageId="14" pageNumber="529" year="1861">
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: 26
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(worker) Texas,
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;
<treatmentCitation id="CF6F10A25B27FF9F9970F97DFC57F918" author="Patton" box="[775,954,1734,1756]" page="618" pageId="14" pageNumber="529" year="1894">
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: 618
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(male);
<treatmentCitation id="CF6F10A25B27FF9F9E78F97CFB38F918" author="Wheeler" box="[1039,1237,1734,1756]" page="407" pageId="14" pageNumber="529" year="1908">
<bibRefCitation id="2A5F4B425B27FF9F9E78F97CFB4BF918" author="Wheeler" box="[1039,1190,1734,1756]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1908) The ants of Texas, New Mexico and Arizona. (Part 1.). Bulletin of the American Museum of Natural History, 24, 399 - 485." type="journal article" year="1908">Wheeler, 1908</bibRefCitation>
: 407
</treatmentCitation>
(queen);
<treatmentCitation id="CF6F10A25B27FF9F9F42F97CFE66F93F" author="Petralia" page="381" pageId="14" pageNumber="529" year="1980">
<bibRefCitation id="2A5F4B425B27FF9F9F42F97CFEBBF93F" author="Petralia" pageId="14" pageNumber="529" refString="Petralia, R. S. &amp; Vinson, S. B. (1980 [&quot; 1979 &quot;]) Comparative anatomy of the ventral region of ant larvae, and its relation to feeding behavior. Psyche, 86, 375 - 394. [Cambridge]" type="journal article" year="1980">Petralia &amp; Vinson, 1980</bibRefCitation>
: 381
</treatmentCitation>
(larva). Subspecies of
<taxonomicName id="89CE4D305B27FF9F98F0F95CFC1AF93F" ID-CoL="48QBS" authority="Emery, 1911: 115" authorityName="Emery" authorityPageNumber="115" authorityYear="1911" box="[647,1015,1765,1787]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B27FF9F98F0F95CFCC8F93F" box="[647,805,1765,1787]" italics="true" pageId="14" pageNumber="529">O. haematodus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9940F95CFC1AF93F" author="Emery" box="[823,1015,1765,1787]" page="115" pageId="14" pageNumber="529" year="1911">
<bibRefCitation id="2A5F4B425B27FF9F9940F95CFC2DF93F" author="Emery" box="[823,960,1765,1787]" pageId="14" pageNumber="529" refString="Emery, C. (1911) Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118, 1 - 125." type="journal article" year="1911">Emery, 1911</bibRefCitation>
: 115
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B27FF9F9E71F95FFB23F93F" author="Wheeler" box="[1030,1230,1765,1787]" page="40" pageId="14" pageNumber="529" year="1914">
<bibRefCitation id="2A5F4B425B27FF9F9E71F95FFB4FF93F" author="Wheeler" box="[1030,1186,1765,1787]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1914) Ants collected by W. M. Mann in the state of Hidalgo, Mexico. Journal of the New York Entomological Society, 22, 37 - 61." type="journal article" year="1914">Wheeler, 1914</bibRefCitation>
: 40
</treatmentCitation>
;
<treatmentCitation id="CF6F10A25B27FF9F9EAAF95FFA74F93F" author="Smith" box="[1245,1433,1765,1787]" page="129" pageId="14" pageNumber="529" year="1939">
<bibRefCitation id="2A5F4B425B27FF9F9EAAF95FFA8CF93F" author="Smith" box="[1245,1377,1765,1787]" pageId="14" pageNumber="529" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">Smith, 1939</bibRefCitation>
: 129
</treatmentCitation>
. Revived status as species:
<treatmentCitation id="CF6F10A25B27FF9F9BA4F8BEFD1EF8DE" author="Taylor" box="[467,755,1796,1818]" page="142" pageId="14" pageNumber="529" year="1962">
<bibRefCitation id="2A5F4B425B27FF9F9BA4F8BEFD2DF8DE" author="Taylor" box="[467,704,1796,1818]" pageId="14" pageNumber="529" refString="Taylor, R. W. &amp; Wilson, E. O. (1962 [&quot; 1961 &quot;]) Ants from three remote oceanic islands. Psyche 68, 137 - 144. http: // dx. doi. org / 10.1155 / 1961 / 87982" type="journal article" year="1962">Taylor &amp; Wilson, 1962</bibRefCitation>
: 142
</treatmentCitation>
;
<treatmentCitation id="CF6F10A25B27FF9F988AF8BFFC55F8DE" author="Brown" box="[765,952,1796,1818]" page="136" pageId="14" pageNumber="529" year="1976">
<bibRefCitation id="2A5F4B425B27FF9F988AF8BFFC68F8DE" author="Brown" box="[765,901,1796,1818]" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 136
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F898FA71F8FC" blockId="14.[151,1436,1734,1848]" box="[151,1436,1826,1848]" pageId="14" pageNumber="529">
<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F898FA71F8FC" box="[151,1436,1826,1848]" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F898FDABF8FC" authority="Buckley, 1867: 355" authorityName="Buckley" authorityPageNumber="355" authorityYear="1867" box="[151,582,1826,1848]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="7CBAEAA15B27FF9F9AE0F898FE97F8FC" box="[151,378,1826,1848]" italics="true" pageId="14" pageNumber="529">Odontomachus texana</emphasis>
Buckley, 1867: 355
</taxonomicName>
. Junior synonym of
<taxonomicName id="89CE4D305B27FF9F9965F899FBA1F8FC" ID-CoL="48QBW" authority="Emery, 1895: 268" authorityName="Emery" authorityPageNumber="268" authorityYear="1895" box="[786,1100,1826,1848]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="7CBAEAA15B27FF9F9965F899FC67F8FC" box="[786,906,1826,1848]" italics="true" pageId="14" pageNumber="529">O. insularis</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F99E1F899FBA1F8FC" author="Emery" box="[918,1100,1826,1848]" page="268" pageId="14" pageNumber="529" year="1895">
<bibRefCitation id="2A5F4B425B27FF9F99E1F899FBF7F8FC" author="Emery" box="[918,1050,1826,1848]" pageId="14" pageNumber="529" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery, 1895</bibRefCitation>
: 268
</treatmentCitation>
</taxonomicName>
; of
<taxonomicName id="89CE4D305B27FF9F9E06F899FA75F8FC" ID-CoL="48QBK" authority="Wheeler, 1902: 26" authorityName="Wheeler" authorityPageNumber="26" authorityYear="1902" box="[1137,1432,1826,1848]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9E06F899FB22F8FC" box="[1137,1231,1826,1848]" italics="true" pageId="14" pageNumber="529">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9EABF898FA75F8FC" author="Wheeler" box="[1244,1432,1826,1848]" page="26" pageId="14" pageNumber="529" year="1902">
<bibRefCitation id="2A5F4B425B27FF9F9EABF898FA9EF8FC" author="Wheeler" box="[1244,1395,1826,1848]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1902) A consideration of S. B. Buckley's &quot; North American Formicidae. &quot; Transactions of the Texas Academy of Sciences, 4, 17 - 31." type="journal article" year="1902">Wheeler, 1902</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F8DAFED9F850" blockId="14.[151,1437,1888,2002]" pageId="14" pageNumber="529">
<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F8DAFED9F850" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F8DAFE5EF8B2" ID-CoL="48QBS" box="[151,435,1888,1910]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B27FF9F9AE0F8DAFE5EF8B2" box="[151,435,1888,1910]" italics="true" pageId="14" pageNumber="529">Odontomachus haematodus</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="89CE4D305B27FF9F9873F8DAFCD3F8B2" authority="Wheeler, 1915: 391" authorityName="Wheeler" authorityPageNumber="391" authorityYear="1915" box="[516,830,1888,1910]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="coninodis">
<emphasis id="7CBAEAA15B27FF9F9873F8DAFD8AF8B2" box="[516,615,1888,1910]" italics="true" pageId="14" pageNumber="529">coninodis</emphasis>
<treatmentCitation id="CF6F10A25B27FF9F9818F8DAFCD3F8B2" author="Wheeler" box="[623,830,1888,1910]" page="391" pageId="14" pageNumber="529" year="1915">
<bibRefCitation id="2A5F4B425B27FF9F9818F8DAFCE4F8B2" author="Wheeler" box="[623,777,1888,1910]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
: 391
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B27FF9F993CF8DBFBE7F8B2" author="Brown" box="[843,1034,1888,1910]" page="103" pageId="14" pageNumber="529" year="1976">
<bibRefCitation id="2A5F4B425B27FF9F993CF8DBFC39F8B2" author="Brown" box="[843,980,1888,1910]" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
. Junior synonym of
<taxonomicName id="89CE4D305B27FF9F9E96F8DBFEDDF850" ID-CoL="48QBK" authority="Brown, 1976: 103" authorityName="Brown" authorityPageNumber="103" authorityYear="1976" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9E96F8DBFAACF8B2" box="[1249,1345,1888,1910]" italics="true" pageId="14" pageNumber="529">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9F26F8DBFEDDF850" author="Brown" page="103" pageId="14" pageNumber="529" year="1976">
<bibRefCitation id="2A5F4B425B27FF9F9F26F8DBFF10F850" author="Brown" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F827FF19F816" blockId="14.[151,1437,1888,2002]" pageId="14" pageNumber="529">
<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F827FF19F816" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F827FE5BF877" box="[151,438,1949,1971]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haemataoda">
<emphasis id="7CBAEAA15B27FF9F9AE0F827FE5BF877" box="[151,438,1949,1971]" italics="true" pageId="14" pageNumber="529">Odontomachus haemataoda</emphasis>
</taxonomicName>
subsp.
<emphasis id="7CBAEAA15B27FF9F987FF827FD69F877" box="[520,644,1949,1971]" italics="true" pageId="14" pageNumber="529">clarionensis</emphasis>
<treatmentCitation id="CF6F10A25B27FF9F98FBF827FCB6F877" author="Wheeler" box="[652,859,1949,1971]" page="141" pageId="14" pageNumber="529" year="1934">
<bibRefCitation id="2A5F4B425B27FF9F98FBF827FCCBF877" author="Wheeler" box="[652,806,1949,1971]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1934) Ants from the islands off the west coast of Lower California and Mexico. Pan-Pacific Entomologist, 10, 132 - 144." type="journal article" year="1934">Wheeler, 1934</bibRefCitation>
: 141
</treatmentCitation>
. Junior synonym of
<taxonomicName id="89CE4D305B27FF9F9E44F824FF1DF816" ID-CoL="48QBK" authority="Taylor &amp; Wilson, 1962: 142" authorityName="Taylor &amp; Wilson" authorityPageNumber="142" authorityYear="1962" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9E44F824FB79F877" box="[1075,1172,1949,1971]" italics="true" pageId="14" pageNumber="529">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9ED3F827FF1DF816" author="Taylor" page="142" pageId="14" pageNumber="529" year="1962">
<bibRefCitation id="2A5F4B425B27FF9F9ED3F827FA74F877" author="Taylor" box="[1188,1433,1949,1971]" pageId="14" pageNumber="529" refString="Taylor, R. W. &amp; Wilson, E. O. (1962 [&quot; 1961 &quot;]) Ants from three remote oceanic islands. Psyche 68, 137 - 144. http: // dx. doi. org / 10.1155 / 1961 / 87982" type="journal article" year="1962">Taylor &amp; Wilson, 1962</bibRefCitation>
: 142
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="1AB1663B5B26FF9E9AE0F9C9FEFEF902" httpUri="https://zenodo.org/record/249909/files/figure.png" pageId="15" pageNumber="530" targetBox="[151,1436,195,1629]" targetPageId="15">
<paragraph id="4E7136B35B26FF9E9AE0F9C9FEFEF902" blockId="15.[151,1436,1651,1734]" pageId="15" pageNumber="530">
<emphasis id="7CBAEAA15B26FF9E9AE0F9C9FEF2F94D" bold="true" box="[151,287,1651,1673]" pageId="15" pageNumber="530">FIGURE 14.</emphasis>
<taxonomicName id="89CE4D305B26FF9E9B51F9C9FDE8F94D" box="[294,517,1651,1673]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E9B51F9C9FDE8F94D" box="[294,517,1651,1673]" italics="true" pageId="15" pageNumber="530">Odontomachus clarus</emphasis>
</taxonomicName>
, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in lateral view, (E) mesosoma in dorsal view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence.
</paragraph>
</caption>
<subSubSection id="06D465385B26FF819AE0F957FB0BFEA1" lastPageId="16" lastPageNumber="531" pageId="15" pageNumber="530" type="diagnosis">
<paragraph id="4E7136B35B26FF819AE0F957FB0BFEA1" blockId="15.[151,1436,1773,2014]" lastBlockId="16.[151,1436,151,357]" lastPageId="16" lastPageNumber="531" pageId="15" pageNumber="530">
<emphasis id="7CBAEAA15B26FF9E9AE0F957FEFCF8C2" bold="true" box="[151,273,1773,1798]" pageId="15" pageNumber="530">Diagnosis.</emphasis>
Workers of
<taxonomicName id="89CE4D305B26FF9E9BE8F954FDE5F8C1" box="[415,520,1773,1797]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E9BE8F954FDE5F8C1" box="[415,520,1773,1797]" italics="true" pageId="15" pageNumber="530">O. clarus</emphasis>
</taxonomicName>
are most similar to
<taxonomicName id="89CE4D305B26FF9E989AF954FC89F8C1" box="[749,868,1773,1797]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E989AF954FC89F8C1" box="[749,868,1773,1797]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B26FF9E99EBF954FBD2F8C1" box="[924,1087,1773,1797]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B26FF9E99EBF954FBD2F8C1" box="[924,1087,1773,1797]" italics="true" pageId="15" pageNumber="530">O. desertorum</emphasis>
</taxonomicName>
, but are distinguished from
<taxonomicName id="89CE4D305B26FF9E9FF6F954FF07F8ED" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E9FF6F954FF07F8ED" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
by their smooth basilar lobes (striate in
<taxonomicName id="89CE4D305B26FF9E98DCF8A8FCCDF8ED" box="[683,800,1809,1833]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E98DCF8A8FCCDF8ED" box="[683,800,1809,1833]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
), and from
<taxonomicName id="89CE4D305B26FF9E99D4F8A8FBA9F8ED" box="[931,1092,1809,1833]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B26FF9E99D4F8A8FBA9F8ED" box="[931,1092,1809,1833]" italics="true" pageId="15" pageNumber="530">O. desertorum</emphasis>
</taxonomicName>
by the following: Size smaller (HL &lt;
<quantity id="89369B565B26FF9E9A9FF88CFEAFF889" box="[232,322,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="530" unit="mm" value="2.2">2.2 mm</quantity>
, WL &lt;
<quantity id="89369B565B26FF9E9BE8F88CFE16F88A" box="[415,507,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="530" unit="mm" value="2.8">2.8 mm</quantity>
); dorsomedian petiolar process nub-like or well-differentiated; propleurae almost entirely smooth and polished-looking; pronotal cervical lobe longer than broad; and antennal scapes relatively shorter (SL/HW = 0.991.07 vs. 1.111.15).
<taxonomicName id="89CE4D305B26FF9E98D4F8C7FC70F851" box="[675,925,1917,1941]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E98D4F8C7FC70F851" box="[675,925,1917,1941]" italics="true" pageId="15" pageNumber="530">Odontomachus clarus</emphasis>
</taxonomicName>
is further differentiated from
<taxonomicName id="89CE4D305B26FF9E9E8AF8C4FA9BF851" box="[1277,1398,1917,1941]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E9E8AF8C4FA9BF851" box="[1277,1398,1917,1941]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
by geography:
<taxonomicName id="89CE4D305B26FF9E9B56F818FE62F87D" box="[289,399,1953,1977]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E9B56F818FE62F87D" box="[289,399,1953,1977]" italics="true" pageId="15" pageNumber="530">O. clarus</emphasis>
</taxonomicName>
only occurs west of the Mississippi river; whereas,
<taxonomicName id="89CE4D305B26FF9E9E73F818FB92F87D" box="[1028,1151,1953,1977]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E9E73F818FB92F87D" box="[1028,1151,1953,1977]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
only occurs east of the Mississippi river. Males are uniquely identifiable among the Nearctic fauna by, at minimum, the following combination of characters: 1) dark brown coloration; 2) ocelli small, weakly bulging past posterior head margin; 3) mesoscutum longitudinally striate (striae may be posterolaterally divergent); 4) posterior margin of petiolar sternum lacking an angular process, but with paired setae on tubercles; and 5) propodeum finely striate. The ninth abdominal sternum of
<taxonomicName id="89CE4D305B39FF819BEAFEBEFDE4FED8" box="[413,521,260,284]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B39FF819BEAFEBEFDE4FED8" box="[413,521,260,284]" italics="true" pageId="16" pageNumber="531">O. clarus</emphasis>
</taxonomicName>
is unique, having a short, broad apical portion of abdominal sternum IX (
<figureCitation id="D6F52A365B39FF819F18FEBEFF5AFE84" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="16" pageNumber="531">Fig. 24</figureCitation>
B). The genitalia are otherwise most similar to those of
<taxonomicName id="89CE4D305B39FF819935FE92FC0BFE84" box="[834,998,296,320]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B39FF819935FE92FC0BFE84" box="[834,998,296,320]" italics="true" pageId="16" pageNumber="531">O. desertorum</emphasis>
</taxonomicName>
, differing primarily in having a more strongly sclerotized digitus and anteroventral process shorter, sharply triangular, and not upturned.
</paragraph>
</subSubSection>
<caption id="1AB1663B5B39FF819AE0F8CEFB92F86C" httpUri="https://zenodo.org/record/249910/files/figure.png" pageId="16" pageNumber="531" targetBox="[151,1436,409,1886]" targetPageId="16">
<paragraph id="4E7136B35B39FF819AE0F8CEFB92F86C" blockId="16.[151,1436,1908,1960]" pageId="16" pageNumber="531">
<emphasis id="7CBAEAA15B39FF819AE0F8CEFECCF84E" bold="true" box="[151,289,1908,1930]" pageId="16" pageNumber="531">FIGURE 15.</emphasis>
<taxonomicName id="89CE4D305B39FF819B5EF8CEFDE4F84E" box="[297,521,1908,1930]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B39FF819B5EF8CEFDE4F84E" box="[297,521,1908,1930]" italics="true" pageId="16" pageNumber="531">Odontomachus clarus</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<subSubSection id="06D465385B38FF809AB0FF2DFDD4FABC" pageId="17" pageNumber="532" type="description">
<paragraph id="4E7136B35B38FF809AB0FF2DFAE2FD63" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">
<emphasis id="7CBAEAA15B38FF809AB0FF2DFEB4FF74" bold="true" box="[199,345,151,176]" pageId="17" pageNumber="532">Description.</emphasis>
Worker: HL 1.922.32, HW 1.571.90, SL 1.681.90, EL 0.310.36, ML 1.061.30, WL 2.432.83, PTH 0.811.10, PTL 0.420.48 (n=7). Entire body generally shiny except where obscured by dense pubescence; head, mesosoma, and petiole reddish-brown, legs yellowish-brown to orangish-brown, and gaster blackish. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; with numerous, fine, appressed pubescence and scattered elongate, erect setae present dorsally. Pronotum with circular, slightly concentric striae that become transverse near rear margin; appressed pubescence abundant; 56 elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; propleuron, mesopleuron, and basalar lobe lacking sculpture; abundant pubescence present dorsally. Metasternum lacking paired elongate, spiniform processes between hind coxae. Petiole widest at base, either gradually tapering apically to a short spiniform spine or stoutly conical (see
<figureCitation id="D6F52A365B38FF809964FE45FCB2FDDC" box="[787,863,511,536]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="17" pageNumber="532">Figs. 6</figureCitation>
D, E and 14F); mostly lacking striae with only faint striae present near base; subpetiolar process rounded triangular; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but with fine coriaceous sculpture (seen at high magnification); slightly coarse, appressed pubescence somewhat sparse, hairs separated by at least 1/2 of their length; scattered erect, elongate setae present.
</paragraph>
<paragraph id="4E7136B35B38FF809AB0FD0EFA8AFAF4" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">Male: HL 0.910.97, HW 1.181.30, SL 0.170.25, EL 0.620.64, EW 0.350.37, OL 0.120.14, OES 0.250.26, PTH 0.660.73, WL 2.392.65, PTL 0.400.59, FWL 4.564.97 (n=3). Body generally shiny except where obscured by dense pubescence; head and mesosoma brownish-black, petiole and gaster dark reddish-brown to brownish-black, legs and antennae reddish-brown. Head and body with abundant semi erect, whitish pubescence. Eyes extremely large, maximum diameter of each eye at least 70% of the length of the head in fullface view. Ocelli small, the length of each ocellus about half the distance between lateral ocellus and eye margin; in full-face view, lateral ocelli do not noticeably protrude beyond posterior border of head. Mesosoma: pronotum lacking sculpture; mesoscutum with fine transversely arcuate striae anteriorly, striae becoming longitudinal posteriorly; mesoscutellum raised and convex, with longitudinal striae; propodeum and mesopleuron with weak to slightly strong longitudinal striae. Petiole bluntly rounded apically, with apically acute to apically rounded triangular subpetiolar process present anteriorly; densely pubescent anteriorly and laterally, with reduced pubescence posteriorly; petiolar sternum lacking angular process near posterior margin, but with paired setae on small tubercles. Abdominal sternum IX disc subrectangular, breadth almost twice length; posterior lobe length less than twice maximum width, basal half barely narrowed. Telomeral apex narrowly rounded; telomere length distinctly greater than height; valviceps ventral apex strongly produced and very narrow; apical margin of valviceps linear until abruptly curving posteriorly at apicoventral process; anteroventral process of valviceps sharply triangular, and not upturned; vertical portion of dorsolateral carina and lateral margin of subapical lamina curving into one another; apicodorsal lobe of valviceps narrow and dorsally elongate; subapical lamina broad.</paragraph>
<paragraph id="4E7136B35B38FF809AB0FA86FDD4FABC" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">Queen: No specimens measured, but similar to workers in color and general appearance except slightly larger, with mesosoma developed for wings.</paragraph>
</subSubSection>
<subSubSection id="06D465385B38FF809AB0FA3EFF0CF98B" pageId="17" pageNumber="532" type="distribution">
<paragraph id="4E7136B35B38FF809AB0FA3EFF0CF98B" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">
<emphasis id="7CBAEAA15B38FF809AB0FA3EFDD1FA59" bold="true" box="[199,572,1412,1437]" pageId="17" pageNumber="532">
Distribution (
<figureCitation id="D6F52A365B38FF809B19FA3EFE27FA59" box="[366,458,1412,1437]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="17" pageNumber="532">Fig. 26</figureCitation>
).
<collectingCountry id="36D976235B38FF809B92FA3EFDD1FA59" box="[485,572,1412,1437]" name="Mexico" pageId="17" pageNumber="532">Mexico</collectingCountry>
</emphasis>
. Chihuahua, Coahuila, Hidalgo, Nuevo Leon, and Clarión Island.
<emphasis id="7CBAEAA15B38FF809F2AFA3EFA71FA59" bold="true" box="[1373,1436,1412,1437]" pageId="17" pageNumber="532">
<collectingCountry id="36D976235B38FF809F2AFA3EFA7AFA59" box="[1373,1431,1412,1437]" name="United States of America" pageId="17" pageNumber="532">USA</collectingCountry>
.
</emphasis>
Arizona: Cochise, Pima, Santa Cruz, and Yavapai Counties. New
<collectingCountry id="36D976235B38FF8099D6FA1DFC1AFA04" box="[929,1015,1447,1472]" name="Mexico" pageId="17" pageNumber="532">Mexico</collectingCountry>
. McKinley County. Texas: Bastrop, Bexar, Bosque, Brewster, Brown, Denton, Jeff Davis, and Travis Counties. Louisiana: Natchitoches Parish. Distributional information from AntWeb (2013), Anonymous (2013),
<bibRefCitation id="2A5F4B425B38FF8099DCFA4AFBA6F9CC" author="Brown" box="[939,1099,1519,1544]" pageId="17" pageNumber="532" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
,
<bibRefCitation id="2A5F4B425B38FF809E2FFA55FAD1F9CC" box="[1112,1340,1519,1544]" pageId="17" pageNumber="532" refString="Adams, B. J., Chen, X. &amp; Hooper-Bui, L. M. (2010) Odontomachus clarus Roger (Hymenoptera: Formicidae) reported in Kisatchie National Forest, Louisiana. Midsouth Entomologist, 3, 104 - 105." type="journal article">
Adams
<emphasis id="7CBAEAA15B38FF809ED9FA4BFB32F9CC" box="[1198,1247,1520,1544]" italics="true" pageId="17" pageNumber="532">et al</emphasis>
. (2010)
</bibRefCitation>
,
<bibRefCitation id="2A5F4B425B38FF809F3FFA4AFE88F9E8" author="Deyrup" pageId="17" pageNumber="532" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
, and examination of specimens from ABS, JTLC, LACM, LSAM, MEM, UCDC, UGA, and UTEP.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B38FF839AB0F9E6FEF0FBF0" lastPageId="18" lastPageNumber="533" pageId="17" pageNumber="532" type="discussion">
<paragraph id="4E7136B35B38FF839AB0F9E6FA84FE84" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,1077]" lastPageId="18" lastPageNumber="533" pageId="17" pageNumber="532">
<emphasis id="7CBAEAA15B38FF809AB0F9E6FEA1F9B1" bold="true" box="[199,332,1628,1653]" pageId="17" pageNumber="532">Discussion.</emphasis>
<taxonomicName id="89CE4D305B38FF809B23F9E6FDA6F9B0" box="[340,587,1628,1652]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B38FF809B23F9E6FDA6F9B0" box="[340,587,1628,1652]" italics="true" pageId="17" pageNumber="532">Odontomachus clarus</emphasis>
</taxonomicName>
has had four specific synonyms until the present paper. While the Buckley name may be safely ignored as superfluous and
<taxonomicName id="89CE4D305B38FF8098CAF93AFC8CF95C" box="[701,865,1664,1688]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B38FF8098CAF93AFC8CF95C" box="[701,865,1664,1688]" italics="true" pageId="17" pageNumber="532">O. desertorum</emphasis>
</taxonomicName>
is here recognized as a valid species, the speciesstatus of the other two names stands to be reassessed.
<taxonomicName id="89CE4D305B38FF809979F91EFBEAF978" box="[782,1031,1700,1724]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B38FF809979F91EFBEAF978" box="[782,1031,1700,1724]" italics="true" pageId="17" pageNumber="532">Odontomachus clarus</emphasis>
</taxonomicName>
is variable across its range, with a broad longitudinal gradient from Texas and Louisiana South to Hidalgo,
<collectingCountry id="36D976235B38FF809992F97DFBD0F924" box="[997,1085,1735,1760]" name="Mexico" pageId="17" pageNumber="532">Mexico</collectingCountry>
, and a very narrow shift from Texas into the mountains of Arizona. The US-Mexico cline is manifested in a North-to-South trend of increasing intensity of gastral sculpturation, erection of gastral pubescence, and anteroposterior compression of the petiolar node. The Mexican High Plateau population is otherwise identical to those of the Great and South Central Plains. The montane Arizonan populations, located in the Chiricahua, Huachua, and Santa Rita Mountains, as well as the Colorado Plateau Transition Zone, have previously been provided with the subspecific name
<taxonomicName id="89CE4D305B38FF809ECBF8C6FEA6F87D" authority="Wheeler 1915" authorityName="Wheeler" authorityYear="1915" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="subSpecies" species="clarus" subSpecies="coninodis">
<emphasis id="7CBAEAA15B38FF809ECBF8C6FA71F850" box="[1212,1436,1916,1940]" italics="true" pageId="17" pageNumber="532">O. clarus coninodis</emphasis>
(
<bibRefCitation id="2A5F4B425B38FF809AE9F81AFEAFF87D" author="Wheeler" box="[158,322,1952,1977]" pageId="17" pageNumber="532" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler 1915</bibRefCitation>
)
</taxonomicName>
due to their nub-like dorsomedian petiolar process (see
<figureCitation id="D6F52A365B38FF8099A8F81AFBC9F87D" box="[991,1060,1952,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="17" pageNumber="532">Fig. 6</figureCitation>
E). Upon careful examination of material across the range of
<taxonomicName id="89CE4D305B38FF809BA1F87EFDD2F818" box="[470,575,1988,2012]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B38FF809BA1F87EFDD2F818" box="[470,575,1988,2012]" italics="true" pageId="17" pageNumber="532">O. clarus</emphasis>
</taxonomicName>
it is clear that this is the only distinguishing character for these populations, and that the conical dorsomedian petiolar process merely represents a foreshortening of the process as the rest of the petiole is otherwise unmodified. Moreover, males attributable to “
<taxonomicName id="89CE4D305B3BFF8399C5FF06FBCCFF11" box="[946,1057,188,213]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="coninodis">coninodis</taxonomicName>
are inseparable from those of
<taxonomicName id="89CE4D305B3BFF839FF6FF06FF30FF3C" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839FF6FF06FF30FF3C" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
except for reduced size. These observations strongly suggest that the name
<taxonomicName id="89CE4D305B3BFF839E16FF5AFAABFF3C" box="[1121,1350,224,248]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="clarus" subSpecies="coninodis">
<emphasis id="7CBAEAA15B3BFF839E16FF5AFAABFF3C" box="[1121,1350,224,248]" italics="true" pageId="18" pageNumber="533">O. clarus coninodis</emphasis>
</taxonomicName>
should remain in synonymy with
<taxonomicName id="89CE4D305B3BFF839BB6FEBEFDC6FED8" box="[449,555,260,284]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BB6FEBEFDC6FED8" box="[449,555,260,284]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
. It would still be valuable to conduct a sampling transect from Texas through the southeastern Arizona mountains to detect intergradation of petiolar node form as
<bibRefCitation id="2A5F4B425B3BFF839E39FE92FB01FE84" author="Brown" box="[1102,1260,295,320]" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
suggested.
</paragraph>
<paragraph id="4E7136B35B3BFF839AB0FEF6FB4AFD41" blockId="18.[151,1437,151,1077]" pageId="18" pageNumber="533">
Finally, the status of the name
<taxonomicName id="89CE4D305B3BFF839844FEF6FCDEFEA0" box="[563,819,332,356]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="clarus" subSpecies="clarionensis">
<emphasis id="7CBAEAA15B3BFF839844FEF6FCDEFEA0" box="[563,819,332,356]" italics="true" pageId="18" pageNumber="533">O. clarus clarionensis</emphasis>
</taxonomicName>
remains to be absolutely determined. The name was provided by
<bibRefCitation id="2A5F4B425B3BFF839B53FED5FE38FE4C" author="Wheeler" box="[292,469,367,392]" pageId="18" pageNumber="533" refString="Wheeler, W. M. (1934) Ants from the islands off the west coast of Lower California and Mexico. Pan-Pacific Entomologist, 10, 132 - 144." type="journal article" year="1934">Wheeler (1934)</bibRefCitation>
for an insular population on Clarión Island (about
<quantity id="89369B565B3BFF839E78FECAFB89FE4C" box="[1039,1124,367,392]" metricMagnitude="5" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="533" unit="km" value="700.0">700 km</quantity>
WSW from the tip of Baja) of what he considered to be
<taxonomicName id="89CE4D305B3BFF839BA5FE2EFD96FE68" box="[466,635,404,428]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3BFF839BA5FE2EFD96FE68" box="[466,635,404,428]" italics="true" pageId="18" pageNumber="533">O. haematodus</emphasis>
</taxonomicName>
. Wheeler erected the subspecies based on the larger size, broader head, and smoother posterolateral head corners than compared to
<taxonomicName id="89CE4D305B3BFF839944FE02FC3FFE14" box="[819,978,440,464]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="haematoda">
<emphasis id="7CBAEAA15B3BFF839944FE02FC3FFE14" box="[819,978,440,464]" italics="true" pageId="18" pageNumber="533">O. haematoda</emphasis>
</taxonomicName>
(an artificial assembly not recognized as such at the time).
<bibRefCitation id="2A5F4B425B3BFF839B2AFE66FD91FE31" author="Taylor" box="[349,636,476,501]" pageId="18" pageNumber="533" refString="Taylor, R. W. &amp; Wilson, E. O. (1962 [&quot; 1961 &quot;]) Ants from three remote oceanic islands. Psyche 68, 137 - 144. http: // dx. doi. org / 10.1155 / 1961 / 87982" type="journal article" year="1962">Taylor and Wilson (1962)</bibRefCitation>
synonymized the subspecies with
<taxonomicName id="89CE4D305B3BFF839989FE66FB8BFE30" box="[1022,1126,476,500]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839989FE66FB8BFE30" box="[1022,1126,476,500]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
without discussion, and the population has been noted as “typical for
<taxonomicName id="89CE4D305B3BFF83980DFDBAFD08FDDC" box="[634,741,512,536]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF83980DFDBAFD08FDDC" box="[634,741,512,536]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
by
<bibRefCitation id="2A5F4B425B3BFF839963FDBAFC5AFDDC" author="Brown" box="[788,951,511,536]" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
. Examination of one of the cotypes at the MCZ does not reveal any salient distinctions of the Clarión Island population from mainland
<taxonomicName id="89CE4D305B3BFF839E87FD9EFAB0FDF8" box="[1264,1373,548,572]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839E87FD9EFAB0FDF8" box="[1264,1373,548,572]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
. The synonymy is thus tentatively confirmed here; however, it would be valuable to sequence genetic material of the Clarión Island population of
<taxonomicName id="89CE4D305B3BFF839BADFDD6FDAFFD40" box="[474,578,620,644]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BADFDD6FDAFFD40" box="[474,578,620,644]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
to compare to material from across the species range.
</paragraph>
<paragraph id="4E7136B35B3BFF839AB0FD35FEF0FBF0" blockId="18.[151,1437,151,1077]" pageId="18" pageNumber="533">
The first report of
<taxonomicName id="89CE4D305B3BFF839BD7FD2AFD77FD6C" box="[416,666,656,680]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BD7FD2AFD77FD6C" box="[416,666,656,680]" italics="true" pageId="18" pageNumber="533">Odontomachus clarus</emphasis>
</taxonomicName>
from Louisiana was based on specimens collected in Provencal in Natchitoches Parish (
<bibRefCitation id="2A5F4B425B3BFF839BF3FD0EFDE6FD08" author="Smith" box="[388,523,692,717]" pageId="18" pageNumber="533" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">Smith 1939</bibRefCitation>
). Unfortunately, we were unable to locate Smiths specimens to verify this record. Subsequent authorities implicitly questioned this disjunct record:
<bibRefCitation id="2A5F4B425B3BFF839905FD62FBE2FD34" author="Brown" box="[882,1039,727,752]" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
omitted it entirely from his lengthy monograph, and this record was later omitted by
<bibRefCitation id="2A5F4B425B3BFF8398B0FD47FC02FCD0" author="Deyrup" box="[711,1007,764,789]" pageId="18" pageNumber="533" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
. Smith's efforts were later vindicated, however, as specimens of
<taxonomicName id="89CE4D305B3BFF839BCCFC9AFDCEFCFC" box="[443,547,800,824]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BCCFC9AFDCEFCFC" box="[443,547,800,824]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
were collected nesting under large rocks in sandy soil in the Kisatchie National Forest in Natchitoches Parish just
<quantity id="89369B565B3BFF83986DFCFEFD8FFC99" box="[538,610,836,861]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="533" unit="km" value="20.0">20 km</quantity>
SW of where Smith reported this species (
<bibRefCitation id="2A5F4B425B3BFF839E32FCFEFAF6FC98" author="Adams" box="[1093,1307,836,861]" pageId="18" pageNumber="533" refString="Adams, B. J., Chen, X. &amp; Hooper-Bui, L. M. (2010) Odontomachus clarus Roger (Hymenoptera: Formicidae) reported in Kisatchie National Forest, Louisiana. Midsouth Entomologist, 3, 104 - 105." type="journal article" year="2010">
Adams
<emphasis id="7CBAEAA15B3BFF839EE9FCFFFB3DFC98" box="[1182,1232,836,860]" italics="true" pageId="18" pageNumber="533">et al</emphasis>
. 2010
</bibRefCitation>
). Similarly, in Texas, nesting sites are typically in dry sites. In Arizona, its nesting habits appear more flexible, as it has been collected under rocks and grass tussocks in dry to mesic habitats (
<bibRefCitation id="2A5F4B425B3BFF8399FEFC36FB4FFC60" author="Deyrup" box="[905,1186,908,933]" pageId="18" pageNumber="533" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover 2004</bibRefCitation>
). These differences in nest site selection may yet reveal whether the “
<taxonomicName id="89CE4D305B3BFF8398CBFC15FCC6FC0C" box="[700,811,943,968]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="coninodis">coninodis</taxonomicName>
populations represent a distinct species, although this seems unlikely. Regardless of the results of increased sampling and molecular work on the Arizona populations, the Louisiana populations of
<taxonomicName id="89CE4D305B3BFF839B95FC42FDA6FBD4" box="[482,587,1016,1040]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839B95FC42FDA6FBD4" box="[482,587,1016,1040]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
would certainly retain that specific epithet as the
<typeStatus id="917588115B3BFF839EF4FC42FB5EFBD4" box="[1155,1203,1016,1040]" pageId="18" pageNumber="533">type</typeStatus>
series was collected from Texas.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,323 @@
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<heading bold="true" box="[151,743,1375,1402]" fontSize="11" level="1" pageId="25" pageNumber="540" reason="1">
<taxonomicName authority="Deyrup and Cover, 2004" authorityName="Deyrup and Cover" authorityYear="2004" box="[151,743,1375,1402]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="relictus">
<emphasis bold="true" box="[151,743,1375,1402]" pageId="25" pageNumber="540">
<emphasis bold="true" box="[151,432,1375,1401]" italics="true" pageId="25" pageNumber="540">Odontomachus relictus</emphasis>
<bibRefCitation author="Deyrup" box="[439,743,1376,1402]" pageId="25" pageNumber="540" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">
Deyrup and Cover,
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</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="25.[151,743,1375,1435]" box="[151,484,1410,1435]" pageId="25" pageNumber="540">
(
<figureCitation box="[159,237,1410,1435]" captionStart="FIGURE 5" captionStartId="8.[151,249,1008,1030]" captionTargetBox="[211,1380,183,992]" captionTargetId="figure@8.[211,1380,183,993]" captionTargetPageId="8" captionText="FIGURE 5. Worker basalar lobes: (A) O. relictus, basalar lobe ovate, with striae (inset at bottom right with simple diagram showing strial pattern of basalar lobe) and (B) O. clarus, basalar lobe spheroid, mostly smooth (inset with simple diagram showing general lack of sculpture of basalar lobe)." httpUri="https://zenodo.org/record/249901/files/figure.png" pageId="25" pageNumber="540">Figs. 5</figureCitation>
,
<figureCitation box="[250,266,1411,1435]" captionStart="FIGURE 8" captionStartId="9.[151,249,1900,1922]" captionTargetBox="[151,1435,1331,1878]" captionTargetId="figure@9.[151,1435,1331,1879]" captionTargetPageId="9" captionText="FIGURE 8. Male mesoscuta, dorsal view: (A) O. brunneus (inset shows transverse striae) and (B) O. relictus (inset shows longitudinal striae)." httpUri="https://zenodo.org/record/249904/files/figure.png" pageId="25" pageNumber="540">8</figureCitation>
,
<figureCitation box="[278,308,1411,1435]" captionStart="FIGURE 20" captionStartId="26.[151,249,1848,1870]" captionTargetBox="[153,1433,193,1826]" captionTargetId="figure@26.[153,1433,193,1826]" captionTargetPageId="26" captionText="FIGURE 20. Odontomachus relictus, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in dorsal view, (E) mesosoma in lateral view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence." httpUri="https://zenodo.org/record/249915/files/figure.png" pageId="25" pageNumber="540">20</figureCitation>
,
<figureCitation box="[320,350,1411,1435]" captionStart="FIGURE 21" captionStartId="27.[151,249,1709,1731]" captionTargetBox="[153,1433,193,1687]" captionTargetId="figure@27.[153,1433,193,1688]" captionTargetPageId="27" captionText="FIGURE 21. Odontomachus relictus, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view." httpUri="https://zenodo.org/record/249916/files/figure.png" pageId="25" pageNumber="540">21</figureCitation>
,
<figureCitation box="[362,393,1411,1435]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="25" pageNumber="540">24</figureCitation>
,
<figureCitation box="[404,435,1410,1435]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="25" pageNumber="540">25</figureCitation>
,
<figureCitation box="[446,475,1410,1435]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="25" pageNumber="540">
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)
</paragraph>
</subSubSection>
<subSubSection box="[151,1097,1482,1504]" pageId="25" pageNumber="540" type="reference_group">
<paragraph blockId="25.[151,1097,1482,1504]" box="[151,1097,1482,1504]" pageId="25" pageNumber="540">
<treatmentCitationGroup box="[151,1097,1482,1504]" pageId="25" pageNumber="540">
<taxonomicName authority="Deyrup &amp; Cover, 2004: 137" authorityName="Deyrup &amp; Cover" authorityPageNumber="137" authorityYear="2004" box="[151,675,1482,1504]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[151,385,1482,1504]" italics="true" pageId="25" pageNumber="540">Odontomachus relictus</emphasis>
<treatmentCitation author="Deyrup" box="[392,675,1482,1504]" page="137" pageId="25" pageNumber="540" year="2004">
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Deyrup &amp; Cover,
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:
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(worker, queen, and male) Florida,
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.
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</subSubSection>
<subSubSection pageId="25" pageNumber="540" type="diagnosis">
<paragraph blockId="25.[151,1436,1549,1970]" pageId="25" pageNumber="540">
<emphasis bold="true" box="[151,273,1549,1574]" pageId="25" pageNumber="540">Diagnosis.</emphasis>
Very similar to
<taxonomicName box="[456,561,1549,1573]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="clarus">
<emphasis box="[456,561,1549,1573]" italics="true" pageId="25" pageNumber="540">O. clarus</emphasis>
</taxonomicName>
, but endemic to Florida;
<taxonomicName box="[847,965,1549,1573]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[847,965,1549,1573]" italics="true" pageId="25" pageNumber="540">O. relictus</emphasis>
</taxonomicName>
has never been reported from west of the Mississippi river. Workers are most similar to
<taxonomicName box="[664,768,1585,1609]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="clarus">
<emphasis box="[664,768,1585,1609]" italics="true" pageId="25" pageNumber="540">O. clarus</emphasis>
</taxonomicName>
, but may be separated by the striate basalar lobes (
<figureCitation box="[1334,1399,1585,1610]" captionStart="FIGURE 5" captionStartId="8.[151,249,1008,1030]" captionTargetBox="[211,1380,183,992]" captionTargetId="figure@8.[211,1380,183,993]" captionTargetPageId="8" captionText="FIGURE 5. Worker basalar lobes: (A) O. relictus, basalar lobe ovate, with striae (inset at bottom right with simple diagram showing strial pattern of basalar lobe) and (B) O. clarus, basalar lobe spheroid, mostly smooth (inset with simple diagram showing general lack of sculpture of basalar lobe)." httpUri="https://zenodo.org/record/249901/files/figure.png" pageId="25" pageNumber="540">
Fig.
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A). Workers are further separated from other Nearctic species by the following combination:
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) gastric pubescence sparse;
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) basalar lobes striate; and
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) petiolar node lacking conspicuous, transverse striae. Males are uniquely identified by the following combination:
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) body and head dark brown, antennae yellowish-brown;
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) very large, strongly bulging ocelli, and
<number box="[465,479,1730,1754]" pageId="25" pageNumber="540" value="3.0">3</number>
) longitudinally striate mesoscutum. The ninth abdominal sternum is unique, having a very short disc relative to length. Genitalia of
<taxonomicName box="[724,849,1765,1789]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[724,849,1765,1789]" italics="true" pageId="25" pageNumber="540">O. relictus</emphasis>
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are most similar to those of
<taxonomicName box="[1219,1330,1765,1789]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="clarus">
<emphasis box="[1219,1330,1765,1789]" italics="true" pageId="25" pageNumber="540">O. clarus</emphasis>
</taxonomicName>
, but are distinguished by the following character combination:
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) telomere apex very narrowly rounded;
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) penisvalvar apicodorsal lobe broader;
<number box="[442,456,1838,1862]" pageId="25" pageNumber="540" value="3.0">3</number>
) apicoventral penisvalvar process shorter, broader (see
<figureCitation box="[1080,1164,1837,1862]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="25" pageNumber="540">
Fig.
<number box="[1132,1164,1837,1862]" pageId="25" pageNumber="540" value="25.0">25</number>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="543" pageId="25" pageNumber="540" type="description">
<paragraph blockId="25.[151,1436,1549,1970]" lastBlockId="28.[151,1437,151,1616]" lastPageId="28" lastPageNumber="543" pageId="25" pageNumber="540">
<emphasis bold="true" box="[199,345,1873,1898]" pageId="25" pageNumber="540">Description.</emphasis>
Worker: HL 2.00
<number box="[564,614,1873,1898]" pageId="25" pageNumber="540" value="2.06">2.06</number>
, HW
<number box="[682,797,1873,1898]" pageId="25" pageNumber="540" value="1.74" valueMax="1.8" valueMin="1.68">1.681.80</number>
, SL
<number box="[852,904,1874,1898]" pageId="25" pageNumber="540" value="1.94">1.94</number>
, EL
<number box="[959,1074,1874,1898]" pageId="25" pageNumber="540" value="0.35" valueMax="0.4" valueMin="0.3">0.300.40</number>
, ML
<number box="[1138,1190,1874,1898]" pageId="25" pageNumber="540" value="1.2">1.20</number>
, WL
<number box="[1255,1370,1873,1898]" pageId="25" pageNumber="540" value="2.675" valueMax="2.68" valueMin="2.67">2.672.68</number>
, PTH
<number box="[151,203,1910,1934]" pageId="25" pageNumber="540" value="0.9">0.90</number>
, PTL
<number box="[275,387,1909,1934]" pageId="25" pageNumber="540" value="0.48" valueMax="0.52" valueMin="0.44">0.440.52</number>
(one
<collectionCode box="[454,521,1910,1933]" pageId="25" pageNumber="540">MEM</collectionCode>
specimen measured, other measurements from
<bibRefCitation author="Deyrup" box="[1067,1347,1909,1934]" pageId="25" pageNumber="540" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">
Deyrup and Cover
<number box="[1286,1347,1910,1934]" pageId="25" pageNumber="540" value="2004.0">2004</number>
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). Entire body generally shiny except where dulled by dense pubescence; head, mesosoma, and petiole reddish-brown, legs yellowish-brown, and gaster blackish. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; with numerous, fine, appressed pubescence and scattered elongate erect setae present dorsally. Pronotum with somewhat circular concentric striae that become longitudinal posteriorly, then transverse at posterior margin; appressed pubescence abundant;
<number box="[1064,1106,1945,1970]" pageId="27" pageNumber="542" value="5.5" valueMax="6.0" valueMin="5.0">56</number>
elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; with abundant pubescence present dorsally. Propleuron mostly lacking sculpture, smooth and shiny; mesopleuron with longitudinal striae at dorsal and ventral margins, otherwise smooth and shiny; pubescence mostly lacking. Basalar lobe with coarse striate sculpture. Metasternum lacking paired elongate, spiniform processes between hind coxae. Petiole widest at base, gradually tapering apically to a short spine directed rearward; mostly lacking striae with only faint striae present near base; subpetiolar process rounded triangular; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but with fine coriaceous sculpture (seen at high magnification); fine, appressed pubescence sparse, spaces between hairs more than the length of a hair; scattered erect, elongate setae present.
</paragraph>
<caption httpUri="https://zenodo.org/record/249915/files/figure.png" pageId="26" pageNumber="541" targetBox="[153,1433,193,1826]" targetPageId="26">
<paragraph blockId="26.[151,1436,1848,1931]" pageId="26" pageNumber="541">
<emphasis bold="true" box="[151,287,1848,1870]" pageId="26" pageNumber="541">FIGURE 20.</emphasis>
<taxonomicName box="[293,527,1848,1870]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[293,527,1848,1870]" italics="true" pageId="26" pageNumber="541">Odontomachus relictus</emphasis>
</taxonomicName>
, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in dorsal view, (E) mesosoma in lateral view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/249916/files/figure.png" pageId="27" pageNumber="542" targetBox="[153,1433,193,1687]" targetPageId="27">
<paragraph blockId="27.[151,1436,1709,1762]" pageId="27" pageNumber="542">
<emphasis bold="true" box="[151,288,1709,1731]" pageId="27" pageNumber="542">FIGURE 21.</emphasis>
<taxonomicName box="[296,531,1709,1731]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="542" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[296,531,1709,1731]" italics="true" pageId="27" pageNumber="542">Odontomachus relictus</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<paragraph blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="543">
Male: HL
<number box="[328,442,440,464]" pageId="28" pageNumber="543" value="1.025" valueMax="1.07" valueMin="0.98">0.981.07</number>
, HW
<number box="[520,635,439,464]" pageId="28" pageNumber="543" value="1.375" valueMax="1.45" valueMin="1.3">1.301.45</number>
, SL
<number box="[699,814,440,464]" pageId="28" pageNumber="543" value="0.22999999999999998" valueMax="0.24" valueMin="0.22">0.220.24</number>
, EL
<number box="[878,930,439,464]" pageId="28" pageNumber="543" value="0.76">0.76</number>
, EW
<number box="[1005,1057,439,464]" pageId="28" pageNumber="543" value="0.5">0.50</number>
, OL
<number box="[1125,1239,440,464]" pageId="28" pageNumber="543" value="0.225" valueMax="0.23" valueMin="0.22">0.220.23</number>
, OES
<number box="[1323,1375,440,464]" pageId="28" pageNumber="543" value="0.13">0.13</number>
, WL
<number box="[151,266,476,501]" pageId="28" pageNumber="543" value="2.465" valueMax="2.5" valueMin="2.43">2.432.50</number>
, PTH
<number box="[337,452,476,501]" pageId="28" pageNumber="543" value="0.6699999999999999" valueMax="0.7" valueMin="0.64">0.640.70</number>
, PTL
<number box="[521,636,476,500]" pageId="28" pageNumber="543" value="0.45499999999999996" valueMax="0.47" valueMin="0.44">0.440.47</number>
, FWL
<number box="[713,761,476,501]" pageId="28" pageNumber="543" value="4.95">4.95</number>
5.00 (two
<collectionCode box="[891,958,477,500]" pageId="28" pageNumber="543">MEM</collectionCode>
specimens measured, other measurements from
<bibRefCitation author="Deyrup" box="[217,506,511,536]" pageId="28" pageNumber="543" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">
Deyrup and Cover
<number box="[445,506,512,536]" pageId="28" pageNumber="543" value="2004.0">2004</number>
</bibRefCitation>
). Body generally shiny except where obscured by dense pubescence; head and mesosoma brownish-black, petiole and gaster dark reddish-brown to brownish-black, femora and tibiae dark brown, tarsi pale yellowish-brown, and antennae yellowish-brown and contrasting with the dark head. Head, meso- and metasoma with abundant whitish subdecumbent pubescence. Eyes extremely large, maximum diameter of each eye at least
<number box="[281,311,656,680]" pageId="28" pageNumber="543" value="70.0">70</number>
% of the length of the head in full-face view. Ocelli large, length of each ocellus greater than distance between lateral ocellus and eye margin; in full-face view, lateral ocelli protrude beyond posterior border of head. Mesosoma: pronotum lacking sculpture; mesoscutum with fine transversely arcuate striae anteriorly, striae becoming longitudinal posteriorly; mesoscutellum convex, shiny, with longitudinal striae; propodeum long and low without a declivious posterior face, with fine striae in concentric ovals medially, transversely bisinuate posteriorly, and longitudinal laterally. Mesopleuron with weak to slightly strong transverse striae. Petiole bluntly rounded apically, with rounded triangular subpetiolar process anteriorly; densely pubescent anteriorly and laterally, with reduced pubescence posteriorly. Abdominal sternum IX disc very short, breadth almost three times length; posterior lobe length about twice width. Telomeral apex very narrowly rounded, almost angular; telomere length distinctly greater than height; valviceps ventral apex weakly produced, narrow; valviceps apical margin linear to weakly concave; apicodorsal lobe of valviceps somewhat narrow and short; vertical portion of dorsolateral carina and lateral margin of subapical lamina curving into one another.
</paragraph>
<paragraph blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="543">
Queen: HL
<number box="[335,387,1087,1112]" pageId="28" pageNumber="543" value="2.15">2.15</number>
, HW
<number box="[457,508,1088,1112]" pageId="28" pageNumber="543" value="1.77">1.77</number>
, EL
<number box="[565,617,1088,1112]" pageId="28" pageNumber="543" value="0.27">0.27</number>
, ML
<number box="[682,734,1088,1112]" pageId="28" pageNumber="543" value="1.2">1.20</number>
, PTL
<number box="[806,858,1087,1112]" pageId="28" pageNumber="543" value="0.52">0.52</number>
, WL
<number box="[924,973,1087,1112]" pageId="28" pageNumber="543" value="2.75">2.75</number>
(measurements from
<bibRefCitation author="Deyrup" pageId="28" pageNumber="543" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">
Deyrup and Cover
<number box="[151,210,1124,1148]" pageId="28" pageNumber="543" value="2004.0">2004</number>
</bibRefCitation>
, one individual measured). Structure and color similar to worker, but with presence of ocelli and expanded mesosoma (for flight); pronotum with transverse striae; and mesonotum with longitudinal striae.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="543" type="distribution">
<paragraph blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="543">
<emphasis bold="true" box="[199,476,1196,1221]" pageId="28" pageNumber="543">
Distribution (
<figureCitation box="[368,461,1196,1221]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="28" pageNumber="543">
Fig.
<number box="[429,461,1196,1221]" pageId="28" pageNumber="543" value="26.0">26</number>
</figureCitation>
):
</emphasis>
Described from Highlands, Polk, Orange, and Citrus Counties in south-central peninsular Florida (
<bibRefCitation author="Deyrup" box="[370,646,1231,1256]" pageId="28" pageNumber="543" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">
Deyrup and Cover
<number box="[585,646,1232,1256]" pageId="28" pageNumber="543" value="2004.0">2004</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="543" type="discussion">
<paragraph blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="543">
<emphasis bold="true" box="[199,332,1268,1293]" pageId="28" pageNumber="543">Discussion.</emphasis>
<taxonomicName box="[342,605,1268,1292]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[342,605,1268,1292]" italics="true" pageId="28" pageNumber="543">Odontomachus relictus</emphasis>
</taxonomicName>
is endemic to sand ridges in central Florida. It was described from the Lake
<collectingCountry box="[215,283,1303,1328]" name="United Kingdom" pageId="28" pageNumber="543">Wales</collectingCountry>
Ridge, including an outlier site previously thought to be part of the Orlando Ridge, and the Southern Brooksville Ridge, although the latter may be a distinctive allopatric population that deserves taxonomic recognition (Sorger, unpublished genetic data).
</paragraph>
<paragraph blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="543">
Little is known about the habitat requirements and nest structure of
<taxonomicName box="[956,1073,1412,1436]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[956,1073,1412,1436]" italics="true" pageId="28" pageNumber="543">O. relictus</emphasis>
</taxonomicName>
. This species has been observed to nest in the ground at the base of oak bushes and is often found in places with at least some leaf litter cover. Due to specific habitat preferences or microclimatological needs,
<taxonomicName box="[835,952,1484,1508]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[835,952,1484,1508]" italics="true" pageId="28" pageNumber="543">O. relictus</emphasis>
</taxonomicName>
may be threatened by human development and disturbance. This species is predominately nocturnal. Male flight activity peaks conspicuously during July through October, apparently correlated with moonlight (see
<bibRefCitation author="Deyrup" box="[820,1031,1556,1581]" pageId="28" pageNumber="543" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article" year="1985">
Deyrup
<emphasis box="[910,966,1556,1580]" italics="true" pageId="28" pageNumber="543">et al.</emphasis>
<number box="[972,1031,1556,1581]" pageId="28" pageNumber="543" value="1985.0">1985</number>
</bibRefCitation>
, where
<taxonomicName box="[1116,1233,1556,1580]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="relictus">
<emphasis box="[1116,1233,1556,1580]" italics="true" pageId="28" pageNumber="543">O. relictus</emphasis>
</taxonomicName>
is identified as
<taxonomicName class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="clarus">
<emphasis italics="true" pageId="28" pageNumber="543">O. clarus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,313 @@
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<treatment id="C66787A55B35FFB09AE0F91EFF3FFC28" ID-DOI="http://doi.org/10.5281/zenodo.6129416" ID-GBIF-Taxon="119429105" ID-Zenodo-Dep="6129416" LSID="urn:lsid:plazi:treatment:C66787A55B35FFB09AE0F91EFF3FFC28" httpUri="http://treatment.plazi.org/id/C66787A55B35FFB09AE0F91EFF3FFC28" lastPageId="33" lastPageNumber="548" pageId="28" pageNumber="543">
<subSubSection id="06D465385B35FF8D9AE0F91EFE2AF91B" pageId="28" pageNumber="543" type="nomenclature">
<paragraph id="4E7136B35B35FF8D9AE0F91EFD59F97A" blockId="28.[151,692,1700,1759]" box="[151,692,1700,1726]" pageId="28" pageNumber="543">
<heading id="153981DF5B35FF8D9AE0F91EFD59F97A" bold="true" box="[151,692,1700,1726]" fontSize="11" level="1" pageId="28" pageNumber="543" reason="1">
<taxonomicName id="89CE4D305B35FF8D9AE0F91EFD59F97A" ID-CoL="48QCK" authority="M. R. Smith, 1937" authorityName="M. R. Smith" authorityYear="1937" box="[151,692,1700,1726]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B35FF8D9AE0F91EFD59F97A" bold="true" box="[151,692,1700,1726]" pageId="28" pageNumber="543">
<emphasis id="7CBAEAA15B35FF8D9AE0F91EFE27F97A" bold="true" box="[151,458,1700,1726]" italics="true" pageId="28" pageNumber="543">Odontomachus ruginodis</emphasis>
<bibRefCitation id="2A5F4B425B35FF8D9BA6F91EFD59F97A" author="Smith" box="[465,692,1700,1726]" pageId="28" pageNumber="543" refString="Smith, M. R. (1937) (&quot; 1936 &quot;) The ants of Puerto Rico. Journal of Agriculture of the University of Puerto Rico, 20, 819 - 875." type="journal article" year="1937">M. R. Smith, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4E7136B35B35FF8D9AE0F97CFE2AF91B" blockId="28.[151,692,1700,1759]" box="[151,455,1734,1759]" pageId="28" pageNumber="543">
(
<figureCitation id="D6F52A365B35FF8D9AE8F97CFF03F91B" box="[159,238,1734,1759]" captionStart="FIGURE 2" captionStartId="6.[151,249,1213,1235]" captionTargetBox="[152,1434,306,1191]" captionTargetId="figure@6.[152,1435,300,1192]" captionTargetPageId="6" captionText="FIGURE 2. Worker petioles, profile view: (A) O. ruginodis and (B) O. clarus." httpUri="https://zenodo.org/record/249898/files/figure.png" pageId="28" pageNumber="543">Figs. 2</figureCitation>
,
<figureCitation id="D6F52A365B35FF8D9A8DF97DFEE7F91B" box="[250,266,1735,1759]" captionStart="FIGURE 3" captionStartId="6.[151,249,1979,2001]" captionTargetBox="[151,1435,1277,1957]" captionTargetId="figure@6.[151,1435,1277,1958]" captionTargetPageId="6" captionText="FIGURE 3. Worker metasterna, anteroventral oblique view: (A) O. haematodus with both hindcoxae removed, arrow indicating elongate metasternal processes and (B) O. ruginodis with one hindcoxa removed, arrow indicating cleft ridge." httpUri="https://zenodo.org/record/249899/files/figure.png" pageId="28" pageNumber="543">3</figureCitation>
,
<figureCitation id="D6F52A365B35FF8D9B61F97DFECBF91A" box="[278,294,1735,1758]" captionStart="FIGURE 7" captionStartId="9.[151,249,1080,1102]" captionTargetBox="[151,1435,410,1059]" captionTargetId="figure@9.[151,1435,410,1059]" captionTargetPageId="9" captionText="FIGURE 7. Male heads, full-face view: (A) O. brunneus and (B) O. haematodus. LOC = lateral ocelli, MOC = median ocellus." httpUri="https://zenodo.org/record/249903/files/figure.png" pageId="28" pageNumber="543">7</figureCitation>
,
<figureCitation id="D6F52A365B35FF8D9B45F97DFEAFF91B" box="[306,322,1735,1759]" captionStart="FIGURE 9" captionStartId="10.[151,249,823,845]" captionTargetBox="[151,1435,324,784]" captionTargetId="figure@10.[151,1435,324,784]" captionTargetPageId="10" captionText="FIGURE 9. Male propodea (Pp), oblique view: (A) O. clarus (rectangular inset shows striate sculpture, arrow indicates lack of obvious propodeal carina) and (B) O. ruginodis (rectangular inset shows reticulated sculpture, arrows point to propodeal carina)." httpUri="https://zenodo.org/record/249905/files/figure.png" pageId="28" pageNumber="543">9</figureCitation>
,
<figureCitation id="D6F52A365B35FF8D9B39F97DFE86F91B" box="[334,363,1735,1759]" captionStart="FIGURE 11" captionStartId="11.[151,249,658,680]" captionTargetBox="[151,1435,193,636]" captionTargetId="figure@11.[151,1435,193,637]" captionTargetPageId="11" captionText="FIGURE 11. Male metasterna, anteroventral oblique view, with arrow indicating sternal processes: (A) O. ruginodis and (B) O. haematodus." httpUri="https://zenodo.org/record/249907/files/figure.png" pageId="28" pageNumber="543">11</figureCitation>
, 2226)
</paragraph>
</subSubSection>
<subSubSection id="06D465385B35FF8D9AE0F8B4FD5FF844" pageId="28" pageNumber="543" type="reference_group">
<paragraph id="4E7136B35B35FF8D9AE0F8B4FD5FF844" blockId="28.[151,1436,1806,1920]" pageId="28" pageNumber="543">
<treatmentCitationGroup id="6EDE119D5B35FF8D9AE0F8B4FD5FF844" pageId="28" pageNumber="543">
<taxonomicName id="89CE4D305B35FF8D9AE0F8B4FE56F8E0" ID-CoL="48QBS" box="[151,443,1806,1828]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B35FF8D9AE0F8B4FE56F8E0" box="[151,443,1806,1828]" italics="true" pageId="28" pageNumber="543">Odontomachus haematodus</emphasis>
</taxonomicName>
<taxonomicName id="89CE4D305B35FF8D9BBCF8B5FCC1F8E0" authority="Smith, 1937: 828" authorityName="Smith" authorityPageNumber="828" authorityYear="1937" box="[459,812,1806,1828]" pageId="28" pageNumber="543" rank="variety" variety="ruginodis">
var.
<emphasis id="7CBAEAA15B35FF8D9876F8B4FD8FF8E0" box="[513,610,1806,1828]" italics="true" pageId="28" pageNumber="543">ruginodis</emphasis>
<treatmentCitation id="CF6F10A25B35FF8D981FF8B4FCC1F8E0" author="Smith" box="[616,812,1806,1828]" page="828" pageId="28" pageNumber="543" year="1937">
<bibRefCitation id="2A5F4B425B35FF8D981FF8B4FD1FF8E0" author="Smith" box="[616,754,1806,1828]" pageId="28" pageNumber="543" refString="Smith, M. R. (1937) (&quot; 1936 &quot;) The ants of Puerto Rico. Journal of Agriculture of the University of Puerto Rico, 20, 819 - 875." type="journal article" year="1937">Smith, 1937</bibRefCitation>
: 828
</treatmentCitation>
</taxonomicName>
(worker and queen),
<collectingCountry id="36D976235B35FF8D9E59F8B4FB63F8E0" box="[1070,1166,1806,1828]" name="Bahamas" pageId="28" pageNumber="543">Bahamas</collectingCountry>
. [First available use of
<taxonomicName id="89CE4D305B35FF8D9AB0F897FDA9F887" ID-CoL="48QBW" box="[199,580,1837,1859]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="haematodes" subSpecies="insularis">
<emphasis id="7CBAEAA15B35FF8D9AB0F897FDA9F887" box="[199,580,1837,1859]" italics="true" pageId="28" pageNumber="543">Odontomachus haematodes insularis</emphasis>
</taxonomicName>
<taxonomicName id="89CE4D305B35FF8D983BF897FC9AF887" ID-CoL="48QCK" authority="Wheeler, 1905: 82" authorityName="Wheeler" authorityPageNumber="82" authorityYear="1905" box="[588,887,1837,1859]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B35FF8D983BF897FD40F887" box="[588,685,1837,1859]" italics="true" pageId="28" pageNumber="543">ruginodis</emphasis>
<treatmentCitation id="CF6F10A25B35FF8D98C1F897FC9AF887" author="Wheeler" box="[694,887,1837,1859]" page="82" pageId="28" pageNumber="543" year="1905">
<bibRefCitation id="2A5F4B425B35FF8D98C1F897FCBDF887" author="Wheeler" box="[694,848,1837,1859]" pageId="28" pageNumber="543" refString="Wheeler, W. M. (1905) The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History, 21, 79 - 135." type="journal article" year="1905">Wheeler, 1905</bibRefCitation>
: 82
</treatmentCitation>
</taxonomicName>
, unavailable name.]
<treatmentCitation id="CF6F10A25B35FF8D9E26F894FF00F8A6" author="Deyrup" page="192" pageId="28" pageNumber="543" year="1985">
<bibRefCitation id="2A5F4B425B35FF8D9E26F894FA75F887" author="Deyrup" box="[1105,1432,1837,1859]" pageId="28" pageNumber="543" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article" year="1985">Deyrup, Trager &amp; Carlin, 1985</bibRefCitation>
: 192
</treatmentCitation>
(male). Raised to species:
<treatmentCitation id="CF6F10A25B35FF8D9862F8F6FD2EF8A6" author="Wilson" box="[533,707,1868,1890]" page="4" pageId="28" pageNumber="543" year="1964">
<bibRefCitation id="2A5F4B425B35FF8D9862F8F6FD48F8A6" author="Wilson" box="[533,677,1868,1890]" pageId="28" pageNumber="543" refString="Wilson, E. O. (1964) The ants of the Florida Keys. Breviora, 210, 1 - 14." type="journal article" year="1964">Wilson, 1964</bibRefCitation>
: 4
</treatmentCitation>
. Junior synonym of
<taxonomicName id="89CE4D305B35FF8D99D3F8F6FB10F8A6" ID-CoL="6SH5N" authority="Brown, 1976: 103" authorityName="Brown" authorityPageNumber="103" authorityYear="1976" box="[932,1277,1868,1890]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="543" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B35FF8D99D3F8F6FBCBF8A6" box="[932,1062,1868,1890]" italics="true" pageId="28" pageNumber="543">O. brunneus</emphasis>
:
<treatmentCitation id="CF6F10A25B35FF8D9E4EF8F7FB10F8A6" author="Brown" box="[1081,1277,1868,1890]" page="103" pageId="28" pageNumber="543" year="1976">
<bibRefCitation id="2A5F4B425B35FF8D9E4EF8F7FB28F8A6" author="Brown" box="[1081,1221,1868,1890]" pageId="28" pageNumber="543" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
. Revived from synonymy:
<treatmentCitation id="CF6F10A25B35FF8D9B49F8D1FD43F844" author="Deyrup" box="[318,686,1898,1920]" page="192" pageId="28" pageNumber="543" year="1985">
<bibRefCitation id="2A5F4B425B35FF8D9B49F8D1FD96F844" author="Deyrup" box="[318,635,1898,1920]" pageId="28" pageNumber="543" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article" year="1985">Deyrup, Trager &amp; Carlin, 1985</bibRefCitation>
: 192
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="06D465385B35FF8C9AE0F817FAD5FE4C" lastPageId="29" lastPageNumber="544" pageId="28" pageNumber="543" type="diagnosis">
<paragraph id="4E7136B35B35FF8C9AE0F817FAD5FE4C" blockId="28.[151,1436,1965,2026]" lastBlockId="29.[151,1436,151,392]" lastPageId="29" lastPageNumber="544" pageId="28" pageNumber="543">
<emphasis id="7CBAEAA15B35FF8D9AE0F817FEFCF802" bold="true" box="[151,273,1965,1990]" pageId="28" pageNumber="543">Diagnosis.</emphasis>
Workers may be identified by the unique following character combination: 1) body small, somewhat tricolored orangish, reddish-brown, and blackish; 2) petiolar node with conspicuous transverse striae; and 3) metasternal ridge notched, with short, rounded triangular teeth (not elongate as in
<taxonomicName id="89CE4D305B34FF8C9E16FF22FAE3FF74" box="[1121,1294,152,176]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="544" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B34FF8C9E16FF22FAE3FF74" box="[1121,1294,152,176]" italics="true" pageId="29" pageNumber="544">O. haematodus</emphasis>
</taxonomicName>
). Males are uniquely identified by the following character combination: 1) ocelli small, weakly projecting beyond posterior head margin; 2) metasternal processes short; 3) mesoscutum smooth to weakly longitudinally striate; 4) propodeum coarsely rugose; and 5) petiolar node conspicuously striate. Abdominal sternum IX is most similar to that of
<taxonomicName id="89CE4D305B34FF8C9FF6FEBEFEF3FE84" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="544" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B34FF8C9FF6FEBEFEF3FE84" italics="true" pageId="29" pageNumber="544">O. haematodus</emphasis>
</taxonomicName>
, but the apex of the posterior lobe is narrower (see
<figureCitation id="D6F52A365B34FF8C992AFE9DFC5CFE84" box="[861,945,295,320]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="29" pageNumber="544">Fig. 24</figureCitation>
). Genitalia are most similar to
<taxonomicName id="89CE4D305B34FF8C9F7EFE92FA9CFE84" box="[1289,1393,296,320]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="544" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B34FF8C9F7EFE92FA9CFE84" box="[1289,1393,296,320]" italics="true" pageId="29" pageNumber="544">O. clarus</emphasis>
</taxonomicName>
but are differentiated by the following characters: telomeral apex broader, more evenly rounded; telomeral posterodorsal margin less concave; valviceps apicodorsal lobe and ventroapical process broader (
<figureCitation id="D6F52A365B34FF8C9EA0FED5FAC6FE4C" box="[1239,1323,367,392]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="29" pageNumber="544">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<paragraph id="4E7136B35B34FF8C9AE0F820FE82F829" pageId="29" pageNumber="544">
<table id="3CCEC4135B34006E9AE0F820FA71F829" box="[151,1436,1946,2029]" gridcols="3" gridrows="3" pageId="29" pageNumber="544">
<tr id="F0FE34F15B34006E9AE0F820FA71F874" box="[151,1436,1946,1968]" gridrow="0" pageId="29" pageNumber="544">
<th id="B32F5D8D5B34006E9AE0F820FD76F874" box="[151,667,1946,1968]" gridcol="0" gridrow="0" pageId="29" pageNumber="544">
<emphasis id="7CBAEAA15B34FF8C9AE0F820FEC8F874" bold="true" box="[151,293,1946,1968]" pageId="29" pageNumber="544">FIGURE 22.</emphasis>
<taxonomicName id="89CE4D305B34FF8C9B47F820FDDBF874" box="[304,566,1946,1968]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="544" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B34FF8C9B47F820FDDBF874" box="[304,566,1946,1968]" italics="true" pageId="29" pageNumber="544">Odontomachus ruginodis</emphasis>
</taxonomicName>
, worker:
</th>
<th id="B32F5D8D5B34006E98D5F820FBADF874" box="[674,1088,1946,1968]" gridcol="1" gridrow="0" pageId="29" pageNumber="544">(A) head in full-face view, (B) lateral</th>
<th id="B32F5D8D5B34006E9E30F820FA71F874" box="[1095,1436,1946,1968]" gridcol="2" gridrow="0" pageId="29" pageNumber="544">habitus, (C) dorsal habitus, (D)</th>
</tr>
<tr id="F0FE34F15B34006E9AE0F802FA71F80A" box="[151,1436,1976,1998]" gridrow="1" pageId="29" pageNumber="544">
<th id="B32F5D8D5B34006E9AE0F802FD76F80A" box="[151,667,1976,1998]" gridcol="0" gridrow="1" pageId="29" pageNumber="544">mesosoma in dorsal view, (E) mesosoma in lateral</th>
<td id="B32F5D8D5B34006E98D5F802FBADF80A" box="[674,1088,1976,1998]" gridcol="1" gridrow="1" pageId="29" pageNumber="544">view, (F) petiole in lateral view, and (G)</td>
<td id="B32F5D8D5B34006E9E30F802FA71F80A" box="[1095,1436,1976,1998]" gridcol="2" gridrow="1" pageId="29" pageNumber="544">first gastral tergite in dorsal view</td>
</tr>
<tr id="F0FE34F15B34006E9AE0F86DFA71F829" box="[151,1436,2007,2029]" gridrow="2" pageId="29" pageNumber="544" rowspan-1="1" rowspan-2="1">
<th id="B32F5D8D5B34006E9AE0F86DFD76F829" box="[151,667,2007,2029]" gridcol="0" gridrow="2" pageId="29" pageNumber="544">showing pubescence.</th>
</tr>
</table>
</paragraph>
<caption id="1AB1663B5B37FF8F9AE0F971FB92F8C4" httpUri="https://zenodo.org/record/249917/files/figure.png" pageId="30" pageNumber="545" targetBox="[153,1433,185,1716]" targetPageId="30">
<paragraph id="4E7136B35B37FF8F9AE0F971FB92F8C4" blockId="30.[151,1436,1739,1792]" pageId="30" pageNumber="545">
<emphasis id="7CBAEAA15B37FF8F9AE0F971FEF2F925" bold="true" box="[151,287,1739,1761]" pageId="30" pageNumber="545">FIGURE 23.</emphasis>
<taxonomicName id="89CE4D305B37FF8F9B52F971FDCBF925" box="[293,550,1739,1761]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="545" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B37FF8F9B52F971FDCBF925" box="[293,550,1739,1761]" italics="true" pageId="30" pageNumber="545">Odontomachus ruginodis</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<subSubSection id="06D465385B37FFB09AB0F89DFCB9FDDC" lastPageId="33" lastPageNumber="548" pageId="30" pageNumber="545" type="description">
<paragraph id="4E7136B35B37FF8E9AB0F89DFCABFE30" blockId="30.[151,1436,1831,2036]" lastBlockId="31.[151,1436,151,501]" lastPageId="31" lastPageNumber="546" pageId="30" pageNumber="545">
<emphasis id="7CBAEAA15B37FF8F9AB0F89DFEB4F884" bold="true" box="[199,345,1831,1856]" pageId="30" pageNumber="545">Description.</emphasis>
Worker: HL 2.002.18, HW 1.581.76, SL 1.781.90, EL 0.330.38, ML 1.161.20, WL 2.502.60, PTH 0.881.04, PTL 0.440.46 (n=5). Entire body generally shiny except where dulled by dense pubescence; head including mandibles and antennae and legs orangish-brown to reddish-brown, mesosoma and waist darker reddish-brown, and gaster brownish-black. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; with numerous, fine, appressed pubescence and scattered elongate, erect setae present dorsally. Pronotum with somewhat circular concentric striae that become transverse near rear margin; appressed pubescence abundant; 610 elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; propleuron, mesopleuron, and basalar lobe lacking sculpture; abundant pubescence present dorsally. Metasternum with ridge that can be bilobed or cleft, but lobes not extending out as elongate, spiniform processes (
<figureCitation id="D6F52A365B36FF8E9B85FE9DFDD9FE84" box="[498,564,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,249,1979,2001]" captionTargetBox="[151,1435,1277,1957]" captionTargetId="figure@6.[151,1435,1277,1958]" captionTargetPageId="6" captionText="FIGURE 3. Worker metasterna, anteroventral oblique view: (A) O. haematodus with both hindcoxae removed, arrow indicating elongate metasternal processes and (B) O. ruginodis with one hindcoxa removed, arrow indicating cleft ridge." httpUri="https://zenodo.org/record/249899/files/figure.png" pageId="31" pageNumber="546">Fig. 3</figureCitation>
B). Petiole widest at base, gradually tapering apically to a short spine directed rearward; with transverse striae present and obvious of all surfaces; subpetiolar process somewhat anvil shaped; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but with fine coriaceous sculpture (seen at high magnification); slightly coarse, appressed pubescence somewhat sparse, spaces between hairs more than 1/2 the length of a single hair; scattered erect, elongate setae present.
</paragraph>
<caption id="1AB1663B5B36FF8E9AE0F813FBE6F819" httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="31" pageNumber="546" targetBox="[151,1435,553,1937]" targetPageId="31">
<paragraph id="4E7136B35B36FF8E9AE0F813FBE6F819" blockId="31.[151,1436,1961,2013]" pageId="31" pageNumber="546">
<emphasis id="7CBAEAA15B36FF8E9AE0F813FECCF87B" bold="true" box="[151,289,1961,1983]" pageId="31" pageNumber="546">FIGURE 24.</emphasis>
Male
<taxonomicName id="89CE4D305B36FF8E9B12F813FE13F87B" box="[357,510,1961,1983]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="genus">
<emphasis id="7CBAEAA15B36FF8E9B12F813FE13F87B" box="[357,510,1961,1983]" italics="true" pageId="31" pageNumber="546">Odontomachus</emphasis>
</taxonomicName>
abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A)
<taxonomicName id="89CE4D305B36FF8E9EE2F813FA78F87B" box="[1173,1429,1961,1983]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B36FF8E9EE2F813FA78F87B" box="[1173,1429,1961,1983]" italics="true" pageId="31" pageNumber="546">Odontomachus brunneus</emphasis>
</taxonomicName>
; (B)
<taxonomicName id="89CE4D305B36FF8E9AC8F872FEF0F819" box="[191,285,1991,2013]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B36FF8E9AC8F872FEF0F819" box="[191,285,1991,2013]" italics="true" pageId="31" pageNumber="546">O. clarus</emphasis>
</taxonomicName>
; (C)
<taxonomicName id="89CE4D305B36FF8E9B24F872FE08F819" box="[339,485,1991,2013]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B36FF8E9B24F872FE08F819" box="[339,485,1991,2013]" italics="true" pageId="31" pageNumber="546">O. desertorum</emphasis>
</taxonomicName>
; (D)
<taxonomicName id="89CE4D305B36FF8E986BF872FD58F819" box="[540,693,1991,2013]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B36FF8E986BF872FD58F819" box="[540,693,1991,2013]" italics="true" pageId="31" pageNumber="546">O. haematodus</emphasis>
</taxonomicName>
; (E)
<taxonomicName id="89CE4D305B36FF8E989EF872FCBEF819" box="[745,851,1991,2013]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B36FF8E989EF872FCBEF819" box="[745,851,1991,2013]" italics="true" pageId="31" pageNumber="546">O. relictus</emphasis>
</taxonomicName>
; (F)
<taxonomicName id="89CE4D305B36FF8E99F2F872FBE8F819" box="[901,1029,1991,2013]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="546" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B36FF8E99F2F872FBE8F819" box="[901,1029,1991,2013]" italics="true" pageId="31" pageNumber="546">O. ruginodis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="1AB1663B5B09FFB19AE0FA52FACCF9BD" httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="32" pageNumber="547" targetBox="[151,1435,193,1489]" targetPageId="32">
<paragraph id="4E7136B35B09FFB19AE0FA52FACCF9BD" blockId="32.[151,1436,1512,1657]" pageId="32" pageNumber="547">
<emphasis id="7CBAEAA15B09FFB19AE0FA52FECFFA3A" bold="true" box="[151,290,1512,1534]" pageId="32" pageNumber="547">FIGURE 25.</emphasis>
Male
<taxonomicName id="89CE4D305B09FFB19B10FA52FDEDFA3A" box="[359,512,1512,1534]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="7CBAEAA15B09FFB19B10FA52FDEDFA3A" box="[359,512,1512,1534]" italics="true" pageId="32" pageNumber="547">Odontomachus</emphasis>
</taxonomicName>
genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (AF) Paramere and volsella, medial view; (GL) penisvalva, lateral view. (A, G)
<taxonomicName id="89CE4D305B09FFB199D7F9BDFB72F9D9" box="[928,1183,1543,1565]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B09FFB199D7F9BDFB72F9D9" box="[928,1183,1543,1565]" italics="true" pageId="32" pageNumber="547">Odontomachus brunneus</emphasis>
</taxonomicName>
; (B, H)
<taxonomicName id="89CE4D305B09FFB19E81F9B2FAB8F9D9" box="[1270,1365,1543,1565]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B09FFB19E81F9B2FAB8F9D9" box="[1270,1365,1543,1565]" italics="true" pageId="32" pageNumber="547">O. clarus</emphasis>
</taxonomicName>
; (C, I)
<taxonomicName id="89CE4D305B09FFB19AE0F99DFEC7F9F8" box="[151,298,1574,1596]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B09FFB19AE0F99DFEC7F9F8" box="[151,298,1574,1596]" italics="true" pageId="32" pageNumber="547">O. desertorum</emphasis>
</taxonomicName>
; (D, J)
<taxonomicName id="89CE4D305B09FFB19B0AF99DFDF5F9F8" box="[381,536,1574,1596]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B09FFB19B0AF99DFDF5F9F8" box="[381,536,1574,1596]" italics="true" pageId="32" pageNumber="547">O. haematodus</emphasis>
</taxonomicName>
; (E, K)
<taxonomicName id="89CE4D305B09FFB19807F99DFD31F9F8" box="[624,732,1574,1596]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B09FFB19807F99DFD31F9F8" box="[624,732,1574,1596]" italics="true" pageId="32" pageNumber="547">O. relictus</emphasis>
</taxonomicName>
; (F, L)
<taxonomicName id="89CE4D305B09FFB1995AF99DFC42F9F8" box="[813,943,1574,1596]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="547" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B09FFB1995AF99DFC42F9F8" box="[813,943,1574,1596]" italics="true" pageId="32" pageNumber="547">O. ruginodis</emphasis>
</taxonomicName>
. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura.
</paragraph>
</caption>
<paragraph id="4E7136B35B09FFB09AB0F91AFCD2FE69" blockId="32.[151,1436,1696,2009]" lastBlockId="33.[151,1437,151,1004]" lastPageId="33" lastPageNumber="548" pageId="32" pageNumber="547">Male: HL 0.880.96, HW 1.101.20, SL 0.150.18, EL 0.580.64, EW 0.320.38, OL 0.150.18, OES 0.220.23, WL 2.282.50, PTH 0.720.76, PTL 0.420.48;, FWL 4.084.60 (n=5). Head, mesosoma, and petiole generally shiny except where obscured by dense pubescence; head and mesosoma yellowish-brown with a darker brown infuscation on pronotum, propodeum dark brown, gaster yellowish-brown to darker brown, antennae and legs pale yellowish-brown. Head, meso- and metasoma with abundant fine, white pubescence except on anepisternum where pubescence is mostly absent. Eyes relatively large, maximum diameter of each eye 5060% of the length of the head in full-face view. Ocelli relatively small, length of each ocellus less than distance between lateral ocellus and eye margin; in full-face view, lateral ocelli only protrude slightly beyond posterior border of head. Mesosoma: pronotum lacking sculpture; mesoscutum feebly shining, lacking obvious sculpture; mesoscutellum raised and rounded, with longitudinal striae; propodeum with distinct declivious face in profile, strongly rugoreticulate; mesopleuron mostly lacking striae. Petiole bluntly rounded apically, with triangular subpetiolar process anteriorly; densely pubescent anteriorly and laterally, but reduced pubescence posteriorly. Abdominal sternum IX disc breadth about twice length; posterior lobe length about twice maximum lobe width, basal half strongly narrowed. Telomeral apex narrowly to somewhat broadly rounded; telomere length distinctly greater than height; valviceps ventral apex weakly produced, broad; valviceps apical margin broadly concave; vertical portion of dorsolateral carina and lateral margin of subapical lamina meeting at an angle; apicodorsal lobe of valviceps somewhat broad, short; subapical lamina broad.</paragraph>
<paragraph id="4E7136B35B08FFB09AB0FE02FCB9FDDC" blockId="33.[151,1437,151,1004]" pageId="33" pageNumber="548">Queen: HL 2.182.20, HW 1.821.88, SL 1.902.02, EL 0.400.46, ML 1.101.30, OL 0.12, WL, 2.943.06, PTH 1.141.24, PTL 0.520.56, FWL 6.006.25 (n=2). Similar to workers in color and general appearance except slightly larger, alate, and with mesosoma developed for wings.</paragraph>
</subSubSection>
<subSubSection id="06D465385B08FFB09AB0FD9EFC66FD34" pageId="33" pageNumber="548" type="distribution">
<paragraph id="4E7136B35B08FFB09AB0FD9EFC66FD34" blockId="33.[151,1437,151,1004]" pageId="33" pageNumber="548">
<emphasis id="7CBAEAA15B08FFB09AB0FD9EFE23FDF9" bold="true" box="[199,462,548,573]" pageId="33" pageNumber="548">
Distribution (
<figureCitation id="D6F52A365B08FFB09B1EFD9EFE52FDF9" box="[361,447,548,573]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="33" pageNumber="548">Fig. 26</figureCitation>
):
</emphasis>
In the
<collectingCountry id="36D976235B08FFB09869FD9EFDA9FDF8" box="[542,580,548,572]" name="United States of America" pageId="33" pageNumber="548">US</collectingCountry>
, this species is known only from peninsular Florida, where it is abundant in the Keys and the southern Peninsula, found sporadically in the central and northern Peninsula: Alachua, Brevard, Broward, Charlotte, Collier, Dade, Glades, Hendrey, Highlands, Hillsborough, Indian River, Lee, Manatee, Martin, Monroe, Orange, Polk, Sarasota, St. Lucie, and Volusia Counties (
<bibRefCitation id="2A5F4B425B08FFB099F4FD2AFB76FD6C" author="Deyrup" box="[899,1179,655,680]" pageId="33" pageNumber="548" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover 2004</bibRefCitation>
). The species has been reported from the West Indies (
<bibRefCitation id="2A5F4B425B08FFB09B86FD0EFD1CFD08" author="Deyrup" box="[497,753,692,716]" pageId="33" pageNumber="548" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup &amp; Cover 2004</bibRefCitation>
), and is apparently widespread in Central and South
<collectingCountry id="36D976235B08FFB09F4BFD0EFA71FD09" box="[1340,1436,692,717]" name="United States of America" pageId="33" pageNumber="548">America</collectingCountry>
(based on specimens examined from UMNM, UCDC, and UGCA).
</paragraph>
</subSubSection>
<subSubSection id="06D465385B08FFB09AB0FD46FF3FFC28" pageId="33" pageNumber="548" type="discussion">
<paragraph id="4E7136B35B08FFB09AB0FD46FF3FFC28" blockId="33.[151,1437,151,1004]" pageId="33" pageNumber="548">
<emphasis id="7CBAEAA15B08FFB09AB0FD46FEA1FCD1" bold="true" box="[199,332,764,789]" pageId="33" pageNumber="548">Discussion.</emphasis>
The status of
<taxonomicName id="89CE4D305B08FFB09B82FD46FCF9FCD0" box="[501,788,764,788]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="548" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B08FFB09B82FD46FCF9FCD0" box="[501,788,764,788]" italics="true" pageId="33" pageNumber="548">Odontomachus ruginodis</emphasis>
</taxonomicName>
as an exotic species is unclear. The Florida populations may be recent arrivals from Antillean populations, as evidenced by the frequent collection in disturbed areas, such as near homes, rather than in natural habitats. Indeed, Puerto Rican populations of
<taxonomicName id="89CE4D305B08FFB09E4DFCFEFB25FC98" box="[1082,1224,836,860]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="548" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B08FFB09E4DFCFEFB25FC98" box="[1082,1224,836,860]" italics="true" pageId="33" pageNumber="548">O. ruginodis</emphasis>
</taxonomicName>
show a preference for forests that are at least 25 years old (
<bibRefCitation id="2A5F4B425B08FFB09824FCDDFC84FC44" author="Osorio-Perez" box="[595,873,871,896]" pageId="33" pageNumber="548" refString="Osorio-Perez, K., Barberena-Arias, M. F. &amp; Aide, T. M. (2007) Changes in ant species richness and composition during plant secondary succession in Puerto Rico. Caribbean Journal of Science, 43, 244 - 253." type="journal article" year="2007">
Osorio-Pérez
<emphasis id="7CBAEAA15B08FFB09898FCD3FCCAFC44" box="[751,807,872,896]" italics="true" pageId="33" pageNumber="548">et al.</emphasis>
2007
</bibRefCitation>
), and males of Nearctic
<taxonomicName id="89CE4D305B08FFB09E01FCD2FAE9FC44" box="[1142,1284,872,896]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="548" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B08FFB09E01FCD2FAE9FC44" box="[1142,1284,872,896]" italics="true" pageId="33" pageNumber="548">O. ruginodis</emphasis>
</taxonomicName>
more-closely resemble those of the Antilles rather than of the mainland Neotropics. During the past 20 years, this species appears to have been steadily expanding its range northward in Florida. Males have been collected from early May through June.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="C66787A55B3BFF849AE0FB3DFB4CF979" ID-DOI="http://doi.org/10.5281/zenodo.6129412" ID-GBIF-Taxon="119429104" ID-Zenodo-Dep="6129412" LSID="urn:lsid:plazi:treatment:C66787A55B3BFF849AE0FB3DFB4CF979" httpUri="http://treatment.plazi.org/id/C66787A55B3BFF849AE0FB3DFB4CF979" lastPageId="21" lastPageNumber="536" pageId="18" pageNumber="533">
<subSubSection id="06D465385B3BFF839AE0FB3DFE1FFB07" pageId="18" pageNumber="533" type="nomenclature">
<paragraph id="4E7136B35B3BFF839AE0FB3DFCDAFB66" blockId="18.[151,823,1159,1219]" box="[151,823,1159,1186]" pageId="18" pageNumber="533">
<heading id="153981DF5B3BFF839AE0FB3DFCDAFB66" bold="true" box="[151,823,1159,1186]" fontSize="11" level="1" pageId="18" pageNumber="533" reason="1">
<emphasis id="7CBAEAA15B3BFF839AE0FB3DFCDAFB66" bold="true" box="[151,823,1159,1186]" pageId="18" pageNumber="533">
<taxonomicName id="89CE4D305B3BFF839AE0FB3DFD7AFB66" authority="Wheeler, 1915" authorityName="Wheeler" authorityYear="1915" box="[151,663,1159,1186]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="desertorum" status="stat. nov.">
<emphasis id="7CBAEAA15B3BFF839AE0FB3DFE32FB65" bold="true" box="[151,479,1159,1185]" italics="true" pageId="18" pageNumber="533">Odontomachus desertorum</emphasis>
<bibRefCitation id="2A5F4B425B3BFF839B91FB32FD7AFB66" author="Wheeler" box="[486,663,1160,1186]" pageId="18" pageNumber="533" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="678957DA5B3BFF8398E9FB32FCDAFB66" box="[670,823,1160,1186]" pageId="18" pageNumber="533">STAT. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4E7136B35B3BFF839AE0FB10FE1FFB07" blockId="18.[151,823,1159,1219]" box="[151,498,1194,1219]" pageId="18" pageNumber="533">
(
<figureCitation id="D6F52A365B3BFF839AE8FB10FF03FB07" box="[159,238,1194,1219]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="18" pageNumber="533">Figs. 6</figureCitation>
,
<figureCitation id="D6F52A365B3BFF839A8DFB11FEF5FB07" box="[250,280,1195,1219]" captionStart="FIGURE 10" captionStartId="10.[151,250,1657,1679]" captionTargetBox="[152,1435,1182,1627]" captionTargetId="figure@10.[152,1435,1182,1627]" captionTargetPageId="10" captionText="FIGURE 10. Male petiolar sterna (ps), profile view: (A) O. desertorum, left arrow indicates subrectangular subpetiolar process (spp), middle arrow indicates the angular posterior process of the petiolar sternum, and right arrow points to the depressed region behind the posterior angled process; and (B) O. clarus, left arrow indicates triangular subpetiolar process, middle arrow points to tubercles with setae, and right arrow indicates the lack of angular posterior process and accompanying shallow depression." httpUri="https://zenodo.org/record/249906/files/figure.png" pageId="18" pageNumber="533">10</figureCitation>
,
<figureCitation id="D6F52A365B3BFF839B52FB10FEAEFB07" box="[293,323,1194,1219]" captionStart="FIGURE 16" captionStartId="19.[151,250,1600,1622]" captionTargetBox="[155,1432,181,1578]" captionTargetId="figure@19.[155,1432,181,1579]" captionTargetPageId="19" captionText="FIGURE 16. Odontomachus desertorum, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in dorsal view, (E) mesosoma in lateral view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence." httpUri="https://zenodo.org/record/249911/files/figure.png" pageId="18" pageNumber="533">16</figureCitation>
,
<figureCitation id="D6F52A365B3BFF839B39FB11FE81FB07" box="[334,364,1195,1219]" captionStart="FIGURE 17" captionStartId="20.[151,250,1944,1966]" captionTargetBox="[153,1433,471,1922]" captionTargetId="figure@20.[153,1433,471,1923]" captionTargetPageId="20" captionText="FIGURE 17. Odontomachus desertorum, male (note: prothorax damaged and genitalia dissected out): (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view." httpUri="https://zenodo.org/record/249912/files/figure.png" pageId="18" pageNumber="533">17</figureCitation>
,
<figureCitation id="D6F52A365B3BFF839B0FFB11FE7BFB07" box="[376,406,1195,1219]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="18" pageNumber="533">24</figureCitation>
,
<figureCitation id="D6F52A365B3BFF839BD5FB10FE2CFB07" box="[418,449,1194,1219]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="18" pageNumber="533">25</figureCitation>
,
<figureCitation id="D6F52A365B3BFF839BBBFB10FE04FB07" box="[460,489,1194,1219]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="18" pageNumber="533">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="06D465385B3BFF839AE0FB48FD4CFA82" pageId="18" pageNumber="533" type="reference_group">
<paragraph id="4E7136B35B3BFF839AE0FB48FD4CFA82" blockId="18.[151,1437,1266,1350]" pageId="18" pageNumber="533">
<treatmentCitationGroup id="6EDE119D5B3BFF839AE0FB48FD4CFA82" infered="true" pageId="18" pageNumber="533">
<taxonomicName id="89CE4D305B3BFF839AE0FB48FCE6FACC" authority="Wheeler, 1915: 391" authorityName="Wheeler" authorityPageNumber="391" authorityYear="1915" box="[151,779,1266,1288]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="haematoda" subSpecies="desertorum">
<emphasis id="7CBAEAA15B3BFF839AE0FB48FDC1FACC" box="[151,556,1266,1288]" italics="true" pageId="18" pageNumber="533">Odontomachus haematoda desertorum</emphasis>
<treatmentCitation id="CF6F10A25B3BFF83984FFB48FCE6FACC" author="Wheeler" box="[568,779,1266,1288]" page="391" pageId="18" pageNumber="533" year="1915">
<bibRefCitation id="2A5F4B425B3BFF83984FFB48FD3BFACC" author="Wheeler" box="[568,726,1266,1288]" pageId="18" pageNumber="533" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
: 391
</treatmentCitation>
</taxonomicName>
<typeStatus id="917588115B3BFF839960FB49FC91FACC" box="[791,892,1267,1288]" pageId="18" pageNumber="533" type="lectotype">Lectotype</typeStatus>
worker
<emphasis id="7CBAEAA15B3BFF8399ABFB48FC09FACC" bold="true" box="[988,996,1266,1288]" pageId="18" pageNumber="533">(</emphasis>
here designated):
<collectingCountry id="36D976235B3BFF839ED3FB48FAD9FACC" box="[1188,1332,1266,1288]" name="United States of America" pageId="18" pageNumber="533">United States</collectingCountry>
, Arizona: Tucson,
<date id="3A7010735B3BFF839B69FAABFE09FAE3" box="[286,484,1297,1319]" pageId="18" pageNumber="533" value="1910-11-24">24 November 1910</date>
(W.M. Wheeler) [MCZC, MCZT_20523] (examined). Junior synonym of
<taxonomicName id="89CE4D305B3BFF839E94FAA8FEDDFA82" ID-CoL="48QBK" authority="Brown, 1976: 103" authorityName="Brown" authorityPageNumber="103" authorityYear="1976" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839E94FAA8FAAFFAE3" box="[1251,1346,1297,1319]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B3BFF839F27FAA8FEDDFA82" author="Brown" page="103" pageId="18" pageNumber="533" year="1976">
<bibRefCitation id="2A5F4B425B3BFF839F27FAA8FF10FA82" author="Brown" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
. See also:
<bibRefCitation id="2A5F4B425B3BFF839BECFA8AFDF4FA82" author="Smith" box="[411,537,1328,1350]" pageId="18" pageNumber="533" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">Smith, 1939</bibRefCitation>
.
<emphasis id="7CBAEAA15B3BFF839854FA8AFD4CFA82" bold="true" box="[547,673,1328,1350]" pageId="18" pageNumber="533">New Status.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="06D465385B3BFF839AE0FAC8FCAAF82B" pageId="18" pageNumber="533" type="diagnosis">
<paragraph id="4E7136B35B3BFF839AE0FAC8FCAAF82B" blockId="18.[151,1437,1394,2031]" pageId="18" pageNumber="533">
<emphasis id="7CBAEAA15B3BFF839AE0FAC8FEFCFA4F" bold="true" box="[151,273,1394,1419]" pageId="18" pageNumber="533">Diagnosis.</emphasis>
Workers of
<taxonomicName id="89CE4D305B3BFF839BD4FAC9FD38FA4F" box="[419,725,1395,1419]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3BFF839BD4FAC9FD38FA4F" box="[419,725,1395,1419]" italics="true" pageId="18" pageNumber="533">Odontomachus desertorum</emphasis>
</taxonomicName>
can be separated from the introduced species
<taxonomicName id="89CE4D305B3BFF839E87FAC9FA76FA4F" box="[1264,1435,1395,1419]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3BFF839E87FAC9FA76FA4F" box="[1264,1435,1395,1419]" italics="true" pageId="18" pageNumber="533">O. haematodus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B3BFF839AB1FA2DFEB9FA6A" box="[198,340,1430,1454]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B3BFF839AB1FA2DFEB9FA6A" box="[198,340,1430,1454]" italics="true" pageId="18" pageNumber="533">O. ruginodis</emphasis>
</taxonomicName>
by the smooth, mostly unsculptured petiole (striate in
<taxonomicName id="89CE4D305B3BFF8399CCFA2DFB88FA6A" box="[955,1125,1430,1454]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3BFF8399CCFA2DFB88FA6A" box="[955,1125,1430,1454]" italics="true" pageId="18" pageNumber="533">O. haematodus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B3BFF839EEBFA2DFAC7FA6A" box="[1180,1322,1430,1454]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B3BFF839EEBFA2DFAC7FA6A" box="[1180,1322,1430,1454]" italics="true" pageId="18" pageNumber="533">O. ruginodis</emphasis>
</taxonomicName>
); from
<taxonomicName id="89CE4D305B3BFF839FF6FA2DFF12FA17" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B3BFF839FF6FA2DFF12FA17" italics="true" pageId="18" pageNumber="533">O. brunneus</emphasis>
</taxonomicName>
by coarse and somewhat sparse gastral pubescence (densely pubescent in
<taxonomicName id="89CE4D305B3BFF839E13FA01FB1FFA17" box="[1124,1266,1467,1491]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B3BFF839E13FA01FB1FFA17" box="[1124,1266,1467,1491]" italics="true" pageId="18" pageNumber="533">O. brunneus</emphasis>
</taxonomicName>
); and from
<taxonomicName id="89CE4D305B3BFF839FF6FA01FF07FA32" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B3BFF839FF6FA01FF07FA32" italics="true" pageId="18" pageNumber="533">O. relictus</emphasis>
</taxonomicName>
by the smooth basilar lobes (striate in
<taxonomicName id="89CE4D305B3BFF8398EEFA65FCE3FA32" box="[665,782,1502,1526]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B3BFF8398EEFA65FCE3FA32" box="[665,782,1502,1526]" italics="true" pageId="18" pageNumber="533">O. relictus</emphasis>
</taxonomicName>
). Additionally,
<taxonomicName id="89CE4D305B3BFF8399C9FA65FB8DFA32" box="[958,1120,1502,1526]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3BFF8399C9FA65FB8DFA32" box="[958,1120,1502,1526]" italics="true" pageId="18" pageNumber="533">O. desertorum</emphasis>
</taxonomicName>
can be separated from these species by geography as it is restricted to the Sonoran Desert region of the southwestern
<collectingCountry id="36D976235B3BFF839EDEF9B9FB20F9DF" box="[1193,1229,1539,1563]" name="United States of America" pageId="18" pageNumber="533">US</collectingCountry>
and northwestern
<collectingCountry id="36D976235B3BFF839AE0F99CFF1DF9FB" box="[151,240,1574,1599]" name="Mexico" pageId="18" pageNumber="533">Mexico</collectingCountry>
; whereas, all of the aforementioned species are only known to occur east of the Mississippi River in the
<collectingCountry id="36D976235B3BFF839AE0F9F1FF53F9A7" box="[151,190,1611,1635]" name="United States of America" pageId="18" pageNumber="533">US</collectingCountry>
. Workers of
<taxonomicName id="89CE4D305B3BFF839B3BF9F1FE03F9A7" box="[332,494,1611,1635]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3BFF839B3BF9F1FE03F9A7" box="[332,494,1611,1635]" italics="true" pageId="18" pageNumber="533">O. desertorum</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="89CE4D305B3BFF83995BF9F1FC79F9A7" box="[812,916,1611,1635]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF83995BF9F1FC79F9A7" box="[812,916,1611,1635]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
(also a western species), with workers of both species being similar in coloration, having coarse gastric pubescence and a smooth petiole and basalar lobe, but may be differentiated from
<taxonomicName id="89CE4D305B3BFF839BA6F929FDD7F96F" box="[465,570,1683,1707]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BA6F929FDD7F96F" box="[465,570,1683,1707]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
by the much larger size (WL 3.41 3.57 compared to WL 2.432.83 for
<taxonomicName id="89CE4D305B3BFF839FF6F929FF30F90A" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839FF6F929FF30F90A" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
), stout, poorly differentiated dorsomedian petiolar spine (
<figureCitation id="D6F52A365B3BFF83991DF90CFC40F90B" box="[874,941,1718,1743]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="18" pageNumber="533">Fig. 6</figureCitation>
C) (may be nublike or well differentiated in
<taxonomicName id="89CE4D305B3BFF839AE0F961FEEDF937" box="[151,256,1755,1779]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839AE0F961FEEDF937" box="[151,256,1755,1779]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
), the entirely striate propleurae (
<figureCitation id="D6F52A365B3BFF839802F960FD55F937" box="[629,696,1754,1779]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="18" pageNumber="533">Fig. 6</figureCitation>
A), pronotal cervical lobes which are wider than long, and by the relatively longer scapes (SL/HW = 1.111.15 vs. 0.991.07). Males are uniquely identifiable among
<collectingCountry id="36D976235B3BFF839E89F945FACFF8D3" box="[1278,1314,1791,1815]" name="United States of America" pageId="18" pageNumber="533">US</collectingCountry>
species by the following character combination: 1) generally large size (WL 2.613.06); 2) ocelli relatively large and bulging beyond posterior border of head; 3) body light golden brown and appendages honey yellow; 4) propodeum finely striate; and 5) petiolar sternum posteriorly glabrous and with a distinct angular process near the posterior margin. The genitalia of
<taxonomicName id="89CE4D305B3BFF839B2AF835FDEEF862" box="[349,515,1934,1958]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3BFF839B2AF835FDEEF862" box="[349,515,1934,1958]" italics="true" pageId="18" pageNumber="533">O. desertorum</emphasis>
</taxonomicName>
are most similar to
<taxonomicName id="89CE4D305B3BFF83988EF835FC8BF862" box="[761,870,1934,1958]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF83988EF835FC8BF862" box="[761,870,1934,1958]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
from which they differ mainly by the narrower posterior ninth sternal lobe (
<figureCitation id="D6F52A365B3BFF839BACF808FDDCF80F" box="[475,561,1970,1995]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="18" pageNumber="533">Fig. 24</figureCitation>
C), the less strongly-sclerotized digitus, and by the longer, slightly upturned, anteroventral valviceps process with rounded apex (
<figureCitation id="D6F52A365B3BFF8398AAF86CFCDDF82B" box="[733,816,2006,2031]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="18" pageNumber="533">
Fig.
<date id="3A7010735B3BFF839966F86CFCDDF82B" box="[785,816,2006,2031]" pageId="18" pageNumber="533" value="1925-01">25</date>
</figureCitation>
I).
</paragraph>
</subSubSection>
<caption id="1AB1663B5B3AFF829AE0F9FAFE82F957" httpUri="https://zenodo.org/record/249911/files/figure.png" pageId="19" pageNumber="534" targetBox="[155,1432,181,1578]" targetPageId="19">
<paragraph id="4E7136B35B3AFF829AE0F9FAFE82F957" blockId="19.[151,1436,1600,1683]" pageId="19" pageNumber="534">
<emphasis id="7CBAEAA15B3AFF829AE0F9FAFEC9F992" bold="true" box="[151,292,1600,1622]" pageId="19" pageNumber="534">FIGURE 16.</emphasis>
<taxonomicName id="89CE4D305B3AFF829B59F9FAFDABF992" box="[302,582,1600,1622]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="534" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3AFF829B59F9FAFDABF992" box="[302,582,1600,1622]" italics="true" pageId="19" pageNumber="534">Odontomachus desertorum</emphasis>
</taxonomicName>
, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in dorsal view, (E) mesosoma in lateral view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence.
</paragraph>
</caption>
<subSubSection id="06D465385B3AFF849AB0F901FD49FD34" lastPageId="21" lastPageNumber="536" pageId="19" pageNumber="534" type="description">
<paragraph id="4E7136B35B3AFF859AB0F901FEC3FE68" blockId="19.[151,1436,1723,2036]" lastBlockId="20.[151,1437,151,428]" lastPageId="20" lastPageNumber="535" pageId="19" pageNumber="534">
<emphasis id="7CBAEAA15B3AFF829AB0F901FEB4F910" bold="true" box="[199,345,1723,1748]" pageId="19" pageNumber="534">Description.</emphasis>
Worker: HL 2.592.74, HW 2.002.17, SL 2.302.40, EL 0.460.49, ML 1.451.57, WL 3.41 3.57, PTH 1.331.35, PTL 0.590.60 (n=2). Entire body generally shiny except where obscured by dense pubescence; head, mesosoma, and petiole deep red, legs and antennae ferrugineous, nearly orange, and gaster piceous black. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning on frontal lobes and clypeus and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head smooth with fine piligerous punctures; head with fine appressed pubescence and with sparse, variable setae present on postgenal bridge and in ocellar triangle. Pronotum with circular, concentric striae which are transverse along rear margin; appressed pubescence abundant; 04 elongate erect setae present. Mesonotum and propodeum with transverse striae, striae coarser on propodeum; propleuron entirely striate; mesopleuron striate in dorsal fifth and near mesocoxae; basalar lobe smooth; pubescence abundant on mesonotum and propodeum. Metasternum lacking paired elongate, spiniform processes between hind coxae. Petiole in profile view widest at base and tapering gradually toward posterodorsally-directed spine; petiole, in anterior view, with dorsomedian process thick at base, long, and poorly differentiated; node mostly smooth and shining, with weak striae present near base; subpetiolar process rounded, triangular; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly smooth and shining, although faintly coriacious dorsomedially; fine appressed pubescence somewhat sparse, hairs separated by somewhat less than one hair length; scattered standing setae present.
</paragraph>
<caption id="1AB1663B5B3DFF859AE0F822FE2BF82F" httpUri="https://zenodo.org/record/249912/files/figure.png" pageId="20" pageNumber="535" targetBox="[153,1433,471,1922]" targetPageId="20">
<paragraph id="4E7136B35B3DFF859AE0F822FE2BF82F" blockId="20.[151,1436,1944,2027]" pageId="20" pageNumber="535">
<emphasis id="7CBAEAA15B3DFF859AE0F822FECFF86A" bold="true" box="[151,290,1944,1966]" pageId="20" pageNumber="535">FIGURE 17.</emphasis>
<taxonomicName id="89CE4D305B3DFF859B5DF822FDD2F86A" box="[298,575,1944,1966]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="535" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3DFF859B5DF822FDD2F86A" box="[298,575,1944,1966]" italics="true" pageId="20" pageNumber="535">Odontomachus desertorum</emphasis>
</taxonomicName>
, male (note: prothorax damaged and genitalia dissected out): (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<paragraph id="4E7136B35B3CFF849AB0FF2DFD49FD34" blockId="21.[151,1437,151,1725]" pageId="21" pageNumber="536">Male: HL 0.971.06, HW 1.341.37, SL 0.230.25, EL 0.690.75, EW 0.430.44, OL 0.18; OES 0.210.26, WL 2.613.06, PTH 0.870.94, PTL 0.540.82, FWL 5.325.64 (n=2). Body generally shiny except where obscured by dense pubescence; head and appendages honey yellow, mesosoma and metasoma light golden brown. Head and body with dense pubescence of variable stature. Eyes extremely large, maximum diameter of each eye at least 70% of head length in full-face view. Ocelli relatively large but not hypertrophied, the length of each ocellus slightly more than half the distance between lateral ocellus and eye margin; in full-face view, lateral ocelli protrude beyond posterior margin of head. Mesosoma: pronotum coriaceous to granulose; mesoscutum covered with weakly rugose striae which are transversely arcuate anteriorly and becoming longitudinal posteriorly; mesoscutellum raised and convex, with longitudinal striae; mesopleuron roughened by dense, weak sculpture; propodeum finely striate. Petiole bluntly rounded apically, with digitate subpetiolar process; densely pubescent anteriorly and laterally, with reduced pubescence posteriorly. Abdominal sternum IX disc trapezoidal, breadth about twice length; posterior lobe length almost three times maximum width, basal half barely narrowed. Telomeral apex narrowly rounded; telomere length distinctly greater than height; valviceps ventral apex strongly produced and very narrow; apical margin of valviceps linear before weakly and abruptly curving posteriorly at apicoventral process; anteroventral process of valviceps slightly elongated, with a rounded, slightly upturned apex; vertical portion of dorsolateral carina and lateral margin of subapical lamina curving into one another; apicodorsal lobe of valviceps narrow and elongated; subapical lamina broad.</paragraph>
</subSubSection>
<subSubSection id="06D465385B3CFF849AB0FD46FB20FCFC" pageId="21" pageNumber="536" type="distribution">
<paragraph id="4E7136B35B3CFF849AB0FD46FB20FCFC" blockId="21.[151,1437,151,1725]" pageId="21" pageNumber="536">
<emphasis id="7CBAEAA15B3CFF849AB0FD46FD84FCD1" bold="true" box="[199,617,764,789]" pageId="21" pageNumber="536">
Distribution (
<figureCitation id="D6F52A365B3CFF849B1DFD46FE2DFCD1" box="[362,448,764,789]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="21" pageNumber="536">Fig. 26</figureCitation>
, Appendix 1).
</emphasis>
Sonoran Desert.
<emphasis id="7CBAEAA15B3CFF849946FD46FC84FCD1" bold="true" box="[817,873,764,789]" pageId="21" pageNumber="536">
<collectingCountry id="36D976235B3CFF849946FD46FC84FCD1" box="[817,873,764,789]" name="United States of America" pageId="21" pageNumber="536">USA</collectingCountry>
</emphasis>
. Arizona: Maricopa and Pima Counties.
<emphasis id="7CBAEAA15B3CFF849F49FD46FA71FCD1" bold="true" box="[1342,1436,764,789]" pageId="21" pageNumber="536">
<collectingCountry id="36D976235B3CFF849F49FD46FA7AFCD1" box="[1342,1431,764,789]" name="Mexico" pageId="21" pageNumber="536">Mexico</collectingCountry>
.
</emphasis>
Sonora. Distributional information from specimens examined from LACM, MCZC, and UCDC.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B3CFF849AB0FCFEFB4CF979" pageId="21" pageNumber="536" type="discussion">
<paragraph id="4E7136B35B3CFF849AB0FCFEFBF2FB01" blockId="21.[151,1437,151,1725]" pageId="21" pageNumber="536">
<emphasis id="7CBAEAA15B3CFF849AB0FCFEFEA1FC99" bold="true" box="[199,332,836,861]" pageId="21" pageNumber="536">Discussion.</emphasis>
<taxonomicName id="89CE4D305B3CFF849B23FCFEFD68FC98" box="[340,645,836,860]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849B23FCFEFD68FC98" box="[340,645,836,860]" italics="true" pageId="21" pageNumber="536">Odontomachus desertorum</emphasis>
</taxonomicName>
is restricted to the Sonoran Desert; its range in Arizona is surrounded by that of
<taxonomicName id="89CE4D305B3CFF849B6BFCD2FE67FC44" box="[284,394,872,896]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849B6BFCD2FE67FC44" box="[284,394,872,896]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
to the north, east, and south. When first described as a subspecies of
<taxonomicName id="89CE4D305B3CFF849E9BFCD2FA76FC44" box="[1260,1435,872,896]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3CFF849E9BFCD2FA76FC44" box="[1260,1435,872,896]" italics="true" pageId="21" pageNumber="536">O. haematodus</emphasis>
</taxonomicName>
(“
<taxonomicName id="89CE4D305B3CFF849ADFFC36FECAFC61" box="[168,295,908,933]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="haematoda">haematoda</taxonomicName>
) by
<bibRefCitation id="2A5F4B425B3CFF849B17FC36FDFBFC61" author="Wheeler" box="[352,534,908,933]" pageId="21" pageNumber="536" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler (1915)</bibRefCitation>
, the coloration and petiolar node form were used to characterize the taxon, a character set scarcely improved by
<bibRefCitation id="2A5F4B425B3CFF849854FC0AFD15FC0C" author="Smith" box="[547,760,943,968]" pageId="21" pageNumber="536" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">M.R. Smith (1939)</bibRefCitation>
who noted the larger size and well-developed lateral ocellar pits of
<taxonomicName id="89CE4D305B3CFF849A98FC6EFE78FC28" box="[239,405,980,1004]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849A98FC6EFE78FC28" box="[239,405,980,1004]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
.
<bibRefCitation id="2A5F4B425B3CFF849BDEFC6EFD82FC29" author="Creighton" box="[425,623,980,1005]" pageId="21" pageNumber="536" refString="Creighton, W. S. (1950) The Ants of North America. Bulletin of the Museum of Comparative Zoology, 104, 1 - 585." type="journal article" year="1950">Creighton (1950)</bibRefCitation>
repeated this diagnosis in his derivative key to the North American
<taxonomicName id="89CE4D305B3CFF849AE0FC42FEACFBD4" box="[151,321,1016,1040]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="7CBAEAA15B3CFF849AE0FC42FEACFBD4" box="[151,321,1016,1040]" italics="true" pageId="21" pageNumber="536">Odontomachus</emphasis>
</taxonomicName>
.
<bibRefCitation id="2A5F4B425B3CFF849B38FC42FE03FBD4" author="Brown" box="[335,494,1015,1040]" pageId="21" pageNumber="536" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
synonymized
<taxonomicName id="89CE4D305B3CFF8498E3FC42FC2BFBD4" box="[660,966,1016,1040]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="subSpecies" species="haematodus" subSpecies="desertorum">
<emphasis id="7CBAEAA15B3CFF8498E3FC42FC2BFBD4" box="[660,966,1016,1040]" italics="true" pageId="21" pageNumber="536">O. haematodus desertorum</emphasis>
</taxonomicName>
with
<taxonomicName id="89CE4D305B3CFF849E7EFC42FB9CFBD4" box="[1033,1137,1016,1040]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849E7EFC42FB9CFBD4" box="[1033,1137,1016,1040]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
, but unfortunately did not provide support for this action. Based on the discovery and examination of the male of
<taxonomicName id="89CE4D305B3CFF849EDDFBA6FABDFBF0" box="[1194,1360,1052,1076]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849EDDFBA6FABDFBF0" box="[1194,1360,1052,1076]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
and a reassessment of worker variation for names attributable to
<taxonomicName id="89CE4D305B3CFF849930FBFAFC5EFB9C" box="[839,947,1088,1112]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849930FBFAFC5EFB9C" box="[839,947,1088,1112]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
, it is here determined that
<taxonomicName id="89CE4D305B3CFF849E81FBFAFA76FB9C" box="[1270,1435,1088,1112]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849E81FBFAFA76FB9C" box="[1270,1435,1088,1112]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
should be revived from synonymy and elevated to species. While no sympatric material of
<taxonomicName id="89CE4D305B3CFF849ED3FBDEFAA5FBB8" box="[1188,1352,1124,1148]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849ED3FBDEFAA5FBB8" box="[1188,1352,1124,1148]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B3CFF849FF6FBDEFF30FB64" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849FF6FBDEFF30FB64" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
were examined during this study,
<bibRefCitation id="2A5F4B425B3CFF849816FB32FCDAFB64" author="Smith" box="[609,823,1159,1184]" pageId="21" pageNumber="536" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">M.R. Smith (1939)</bibRefCitation>
indicated that the two species may occur within a few miles of one another, supporting recognition of
<taxonomicName id="89CE4D305B3CFF8498DBFB16FCA3FB00" box="[684,846,1196,1220]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF8498DBFB16FCA3FB00" box="[684,846,1196,1220]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
as a valid species.
</paragraph>
<paragraph id="4E7136B35B3CFF849AB0FB6AFB4CF979" blockId="21.[151,1437,151,1725]" pageId="21" pageNumber="536">
New worker-based characters for separating
<taxonomicName id="89CE4D305B3CFF8498BEFB6AFC81FB2C" box="[713,876,1232,1256]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF8498BEFB6AFC81FB2C" box="[713,876,1232,1256]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
from
<taxonomicName id="89CE4D305B3CFF8499C1FB6AFBCCFB2C" box="[950,1057,1232,1256]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF8499C1FB6AFBCCFB2C" box="[950,1057,1232,1256]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
are the formers longer antennal scapes (see diagnosis), shorter cervical pronotal lobe (which contacts the occiput when the head is reflexed), and entirely or almost entirely striate propleurae (which forms the longitudinally divided area beneath the pronotum and above the procoxae, and is technically the proepisterna) (
<figureCitation id="D6F52A365B3CFF849936FA86FC69FA91" box="[833,900,1340,1365]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="21" pageNumber="536">Fig. 6</figureCitation>
A). The male of
<taxonomicName id="89CE4D305B3CFF849E4CFA86FB36FA90" box="[1083,1243,1340,1364]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849E4CFA86FB36FA90" box="[1083,1243,1340,1364]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
is conspicuously different from that of
<taxonomicName id="89CE4D305B3CFF849BE6FADAFE16FABC" box="[401,507,1376,1400]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849BE6FADAFE16FABC" box="[401,507,1376,1400]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
in terms of coloration and with respect to the petiolar sternum, which is strongly angled in
<taxonomicName id="89CE4D305B3CFF849B70FA3EFE47FA58" box="[263,426,1412,1436]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849B70FA3EFE47FA58" box="[263,426,1412,1436]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
(
<figureCitation id="D6F52A365B3CFF849BCDFA3EFDE6FA58" box="[442,523,1412,1437]" captionStart="FIGURE 10" captionStartId="10.[151,250,1657,1679]" captionTargetBox="[152,1435,1182,1627]" captionTargetId="figure@10.[152,1435,1182,1627]" captionTargetPageId="10" captionText="FIGURE 10. Male petiolar sterna (ps), profile view: (A) O. desertorum, left arrow indicates subrectangular subpetiolar process (spp), middle arrow indicates the angular posterior process of the petiolar sternum, and right arrow points to the depressed region behind the posterior angled process; and (B) O. clarus, left arrow indicates triangular subpetiolar process, middle arrow points to tubercles with setae, and right arrow indicates the lack of angular posterior process and accompanying shallow depression." httpUri="https://zenodo.org/record/249906/files/figure.png" pageId="21" pageNumber="536">Fig. 10</figureCitation>
A) and nearly flat in
<taxonomicName id="89CE4D305B3CFF84988DFA3EFC8EFA58" box="[762,867,1412,1436]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF84988DFA3EFC8EFA58" box="[762,867,1412,1436]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
(
<figureCitation id="D6F52A365B3CFF849903FA3EFC2AFA58" box="[884,967,1412,1437]" captionStart="FIGURE 10" captionStartId="10.[151,250,1657,1679]" captionTargetBox="[152,1435,1182,1627]" captionTargetId="figure@10.[152,1435,1182,1627]" captionTargetPageId="10" captionText="FIGURE 10. Male petiolar sterna (ps), profile view: (A) O. desertorum, left arrow indicates subrectangular subpetiolar process (spp), middle arrow indicates the angular posterior process of the petiolar sternum, and right arrow points to the depressed region behind the posterior angled process; and (B) O. clarus, left arrow indicates triangular subpetiolar process, middle arrow points to tubercles with setae, and right arrow indicates the lack of angular posterior process and accompanying shallow depression." httpUri="https://zenodo.org/record/249906/files/figure.png" pageId="21" pageNumber="536">Fig. 10</figureCitation>
B). Genitalic differences between the two species are indicated in the diagnosis above. The ocelli of male
<taxonomicName id="89CE4D305B3CFF8499FFFA12FBC1FA04" box="[904,1068,1448,1472]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF8499FFFA12FBC1FA04" box="[904,1068,1448,1472]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
are also slightly larger than
<taxonomicName id="89CE4D305B3CFF849FF6FA12FF30FA20" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849FF6FA12FF30FA20" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
; this may reflect a more light-restricted flight timing or perhaps size-based allometry. Until further males are collected and phenological data are recorded and collated, it is too early to infer temporal isolation of diel flight times between the two species. Beyond worker and male morphology, little is known about the biology of
<taxonomicName id="89CE4D305B3CFF849FF6F9AEFEFAF994" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849FF6F9AEFEFAF994" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
. Whereas
<taxonomicName id="89CE4D305B3CFF849BE2F982FDECF994" box="[405,513,1592,1616]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3CFF849BE2F982FDECF994" box="[405,513,1592,1616]" italics="true" pageId="21" pageNumber="536">O. clarus</emphasis>
</taxonomicName>
is usually collected in woodland habitats,
<taxonomicName id="89CE4D305B3CFF84998FF982FB70F994" box="[1016,1181,1592,1616]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF84998FF982FB70F994" box="[1016,1181,1592,1616]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
seems to prefer open desert/desert-scrub habitats.
<bibRefCitation id="2A5F4B425B3CFF849BA1F9E6FD87F9B0" author="Smith" box="[470,618,1628,1653]" pageId="21" pageNumber="536" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">Smith (1939)</bibRefCitation>
noted that
<taxonomicName id="89CE4D305B3CFF849892F9E6FC6AF9B0" box="[741,903,1628,1652]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B3CFF849892F9E6FC6AF9B0" box="[741,903,1628,1652]" italics="true" pageId="21" pageNumber="536">O. desertorum</emphasis>
</taxonomicName>
may be common but infrequently collected due to nocturnal or matinal foraging times. Material provided by Stefan Cover (MCZ) suggest that this may indeed be true, as the label data indicate that the workers were collected while “nocturnal[ly] foraging.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="4E7136B35B3CFF849AE0F8B5FD2DF8EE" blockId="21.[151,704,1807,1867]" box="[151,704,1807,1834]" pageId="21" pageNumber="536">
<heading id="153981DF5B3CFF849AE0F8B5FD2DF8EE" bold="true" box="[151,704,1807,1834]" fontSize="11" level="1" pageId="21" pageNumber="536" reason="1">
<taxonomicName id="89CE4D305B3CFF849AE0F8B5FD2DF8EE" ID-CoL="48QBS" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,704,1807,1834]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3CFF849AE0F8B5FD2DF8EE" bold="true" box="[151,704,1807,1834]" pageId="21" pageNumber="536">
<emphasis id="7CBAEAA15B3CFF849AE0F8B5FE04F8ED" bold="true" box="[151,489,1807,1833]" italics="true" pageId="21" pageNumber="536">Odontomachus haematodus</emphasis>
(
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)
</emphasis>
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</paragraph>
<paragraph id="4E7136B35B3CFF849AE0F888FE1CF88F" blockId="21.[151,704,1807,1867]" box="[151,497,1842,1867]" pageId="21" pageNumber="536">
(
<figureCitation id="D6F52A365B3CFF849AE8F888FF00F88F" box="[159,237,1842,1867]" captionStart="FIGURE 3" captionStartId="6.[151,249,1979,2001]" captionTargetBox="[151,1435,1277,1957]" captionTargetId="figure@6.[151,1435,1277,1958]" captionTargetPageId="6" captionText="FIGURE 3. Worker metasterna, anteroventral oblique view: (A) O. haematodus with both hindcoxae removed, arrow indicating elongate metasternal processes and (B) O. ruginodis with one hindcoxa removed, arrow indicating cleft ridge." httpUri="https://zenodo.org/record/249899/files/figure.png" pageId="21" pageNumber="536">Figs. 3</figureCitation>
,
<figureCitation id="D6F52A365B3CFF849A8DF889FEFAF88F" box="[250,279,1843,1867]" captionStart="FIGURE 11" captionStartId="11.[151,249,658,680]" captionTargetBox="[151,1435,193,636]" captionTargetId="figure@11.[151,1435,193,637]" captionTargetPageId="11" captionText="FIGURE 11. Male metasterna, anteroventral oblique view, with arrow indicating sternal processes: (A) O. ruginodis and (B) O. haematodus." httpUri="https://zenodo.org/record/249907/files/figure.png" pageId="21" pageNumber="536">11</figureCitation>
,
<figureCitation id="D6F52A365B3CFF849B54F889FEACF88F" box="[291,321,1843,1867]" captionStart="FIGURE 18" captionStartId="22.[151,249,1943,1965]" captionTargetBox="[153,1433,318,1920]" captionTargetId="figure@22.[153,1433,318,1921]" captionTargetPageId="22" captionText="FIGURE 18. Odontomachus haematodus, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in dorsal view, (E) mesosoma in lateral view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence." httpUri="https://zenodo.org/record/249913/files/figure.png" pageId="21" pageNumber="536">18</figureCitation>
,
<figureCitation id="D6F52A365B3CFF849B3AF889FE86F88F" box="[333,363,1843,1867]" captionStart="FIGURE 19" captionStartId="23.[151,249,1810,1832]" captionTargetBox="[153,1433,185,1796]" captionTargetId="figure@23.[153,1433,185,1797]" captionTargetPageId="23" captionText="FIGURE 19. Odontomachus haematodus, male: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) body in lateral view, (E) mesosoma in lateral view, (F) body in dorsal view, and (G) mesosoma in dorsal view." httpUri="https://zenodo.org/record/249914/files/figure.png" pageId="21" pageNumber="536">19</figureCitation>
,
<figureCitation id="D6F52A365B3CFF849B00F889FE78F88F" box="[375,405,1843,1867]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="21" pageNumber="536">24</figureCitation>
,
<figureCitation id="D6F52A365B3CFF849BD6F888FE52F88F" box="[417,447,1842,1867]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="21" pageNumber="536">25</figureCitation>
,
<figureCitation id="D6F52A365B3CFF849BBCF888FE05F88F" box="[459,488,1842,1867]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="21" pageNumber="536">26</figureCitation>
)
</paragraph>
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<taxonomicName id="89CE4D305B3CFF849AE0F8C1FDADF854" authority="Linnaeus, 1758: 582" authorityName="Linnaeus" authorityPageNumber="582" authorityYear="1758" box="[151,576,1914,1936]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="haematoda">
<emphasis id="7CBAEAA15B3CFF849AE0F8C1FE8AF854" box="[151,359,1914,1936]" italics="true" pageId="21" pageNumber="536">Formica haematoda</emphasis>
<treatmentCitation id="CF6F10A25B3CFF849B19F8C0FDADF854" ID-DOI="http://doi.org/10.5281/zenodo.3509159" ID-Zenodo-Dep="3509159" LSID="urn:lsid:zoobank.org:act:EBBA3176-6D80-490B-9F90-3A47BC468BDA" author="Linnaeus" box="[366,576,1914,1936]" httpUri="http://treatment.plazi.org/id/5FF6EF675F585A25D52772B20D71F3F8" page="582" pageId="21" pageNumber="536" year="1758">
<bibRefCitation id="2A5F4B425B3CFF849B19F8C0FDE2F854" author="Linnaeus" box="[366,527,1914,1936]" pageId="21" pageNumber="536" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. L. Salvii, Holmiae [= Stockholm], 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
: 582
</treatmentCitation>
</taxonomicName>
(queen) “
<collectingCountry id="36D976235B3CFF8498D1F8C0FD12F854" box="[678,767,1914,1936]" name="United States of America" pageId="21" pageNumber="536">America</collectingCountry>
Meridionali;
<treatmentCitation id="CF6F10A25B3CFF8499F9F8C1FBC1F854" author="Emery" box="[910,1068,1914,1936]" page="5" pageId="21" pageNumber="536" year="1899">
<bibRefCitation id="2A5F4B425B3CFF8499F9F8C1FBF9F854" author="Emery" box="[910,1044,1914,1936]" pageId="21" pageNumber="536" refString="Emery, C. (1899) Intorno alle larve di alcune formiche. Rendiconti delle Sessioni della Reale Accademia delle Scienze dell'Istituto di Bologna (n. s.), 3, 93." type="journal article" year="1899">Emery, 1899</bibRefCitation>
: 5
</treatmentCitation>
(larva);
<treatmentCitation id="CF6F10A25B3CFF849EF1F8C0FF00F86B" author="Wheeler" page="646" pageId="21" pageNumber="536" year="1952">
<bibRefCitation id="2A5F4B425B3CFF849EF1F8C0FA74F854" author="Wheeler" box="[1158,1433,1914,1936]" pageId="21" pageNumber="536" refString="Wheeler, W. M. &amp; Wheeler, J. (1952) The ant larvae of the subfamily Ponerinae-Part II. American Midland Naturalist, 48, 604 - 672. http: // dx. doi. org / 10.2307 / 2422200" type="journal article" year="1952">Wheeler &amp; Wheeler, 1952</bibRefCitation>
: 646
</treatmentCitation>
(larva). Combination in
<taxonomicName id="89CE4D305B3CFF849B9FF823FCB7F86B" ID-CoL="66Y3" authority="Latreille, 1804: 179" authorityName="Latreille" authorityPageNumber="179" authorityYear="1804" box="[488,858,1945,1967]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="7CBAEAA15B3CFF849B9FF823FD6CF86B" box="[488,641,1945,1967]" italics="true" pageId="21" pageNumber="536">Odontomachus</emphasis>
:
<treatmentCitation id="CF6F10A25B3CFF8498F9F823FCB7F86B" author="Latreille" box="[654,858,1945,1967]" page="179" pageId="21" pageNumber="536" year="1804">
<bibRefCitation id="2A5F4B425B3CFF8498F9F823FCC5F86B" author="Latreille" box="[654,808,1945,1967]" pageId="21" pageNumber="536" refString="Latreille, P. A. (1804) Tableau methodique des insectes. In: Societ de Naturalistes et d'Agriculteurs 1804. Nouveau dictionnaire d'histoire naturelle. Tome 24. Deterville, Paris, pp. 129 - 200." type="book chapter" year="1804">Latreille, 1804</bibRefCitation>
: 179
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B3CFF849AE0F803FED9F828" blockId="21.[151,1436,1914,2028]" pageId="21" pageNumber="536">
<treatmentCitationGroup id="6EDE119D5B3CFF849AE0F803FED9F828" pageId="21" pageNumber="536">
<taxonomicName id="89CE4D305B3CFF849AE0F803FDC7F80A" ID-CoL="6JGGW" authority="DeGeer, 1773: 601" authorityName="DeGeer" authorityPageNumber="601" authorityYear="1773" box="[151,554,1976,1998]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="maxillosa">
<emphasis id="7CBAEAA15B3CFF849AE0F803FEB5F80A" box="[151,344,1976,1998]" italics="true" pageId="21" pageNumber="536">Formica maxillosa</emphasis>
DeGeer, 1773: 601
</taxonomicName>
. Junior synonym of
<taxonomicName id="89CE4D305B3CFF849976F803FB88F80A" ID-CoL="48QBS" authority="Retzius, 1783: 75" authorityName="Retzius" authorityPageNumber="75" authorityYear="1783" box="[769,1125,1976,1998]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3CFF849976F803FC71F80A" box="[769,924,1976,1998]" italics="true" pageId="21" pageNumber="536">O. haematodus</emphasis>
:
<treatmentCitation id="CF6F10A25B3CFF8499DBF802FB88F80A" author="Retzius" box="[940,1125,1976,1998]" page="75" pageId="21" pageNumber="536" year="1783">
<bibRefCitation id="2A5F4B425B3CFF8499DBF802FBD3F80A" author="Retzius" box="[940,1086,1976,1998]" pageId="21" pageNumber="536" refString="Retzius, A. J. (1783) Caroli de Geer. Genera et species insectorum e generosissimi auctoris scriptis extraxit, digessit, Latine quoad partem reddidit, et terminologiam insectorum Linneanam addidit. Cruse, Lipsiae [= Leipzig], 220 pp. http: // dx. doi. org / 10.5962 / bhl. title. 35661" type="book" year="1783">Retzius, 1783</bibRefCitation>
: 75
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B3CFF849E05F802FADEF80A" author="Olivier" box="[1138,1331,1976,1998]" page="502" pageId="21" pageNumber="536" year="1792">
<bibRefCitation id="2A5F4B425B3CFF849E05F802FB13F80A" author="Olivier" box="[1138,1278,1976,1998]" pageId="21" pageNumber="536" refString="Olivier, A. G. (1792) Encyclopedie methodique. Histoire naturelle. Insectes. Tome 6. (pt. 2). Panckoucke, Paris, pp. 369 - 704." type="journal article" year="1792">Olivier, 1792</bibRefCitation>
: 502
</treatmentCitation>
;
<treatmentCitation id="CF6F10A25B3CFF849F37F802FEDDF828" author="Latreille" page="192" pageId="21" pageNumber="536" year="1802">
<bibRefCitation id="2A5F4B425B3CFF849F37F802FF10F828" author="Latreille" pageId="21" pageNumber="536" refString="Latreille, P. A. (1802) Histoire naturelle des fourmis, et recueil de memoires et d'observations sur les abeilles, les araignees, les faucheurs, et autres insectes. Impr. Crapelet (chez T. Barrois), Paris, xvi + 445 pp." type="book" year="1802">Latreille, 1802</bibRefCitation>
: 192
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B3FFF879AE0FF22FA71FF2F" blockId="22.[151,1436,152,266]" pageId="22" pageNumber="537">
<treatmentCitationGroup id="6EDE119D5B3FFF879AE0FF22FA71FF2F" pageId="22" pageNumber="537">
<taxonomicName id="89CE4D305B3FFF879AE0FF22FD62FF6A" authority="Smith, F., 1858: 78" authorityName="Smith, F." authorityPageNumber="78" authorityYear="1858" box="[151,655,152,174]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="hirsutiusculus">
<emphasis id="7CBAEAA15B3FFF879AE0FF22FE29FF6A" box="[151,452,152,174]" italics="true" pageId="22" pageNumber="537">Odontomachus hirsutiusculus</emphasis>
<treatmentCitation id="CF6F10A25B3FFF879BBCFF22FD62FF6A" ID-DOI="http://doi.org/10.5281/zenodo.3507426" ID-Zenodo-Dep="3507426" author="Smith" box="[459,655,152,174]" httpUri="http://treatment.plazi.org/id/F4EEFCA44D6952971AC8077B0B40B9E4" page="78" pageId="22" pageNumber="537" year="1858">
<bibRefCitation id="2A5F4B425B3FFF879BBCFF22FD84FF6A" author="Smith" box="[459,617,152,174]" pageId="22" pageNumber="537" refString="Smith, F. (1858) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part VI. Formicidae. British Museum, London, 216 pp." type="book" year="1858">Smith, F., 1858</bibRefCitation>
: 78
</treatmentCitation>
</taxonomicName>
. Junior synonym of
<taxonomicName id="89CE4D305B3FFF87992AFF23FB44FF6A" ID-CoL="48QBS" authority="Roger, 1861: 24" authorityName="Roger" authorityPageNumber="24" authorityYear="1861" box="[861,1193,152,174]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3FFF87992AFF23FC1BFF6A" box="[861,1014,152,174]" italics="true" pageId="22" pageNumber="537">O. haematodus</emphasis>
:
<treatmentCitation id="CF6F10A25B3FFF879E74FF23FB44FF6A" author="Roger" box="[1027,1193,152,174]" page="24" pageId="22" pageNumber="537" year="1861">
<bibRefCitation id="2A5F4B425B3FFF879E74FF23FB6EFF6A" author="Roger" box="[1027,1155,152,174]" pageId="22" pageNumber="537" refString="Roger, J. (1861) Die Ponera - artigen Ameisen (Schluss). Berliner Entomologische Zeitschrift, 5, 1 - 54." type="journal article" year="1861">Roger, 1861</bibRefCitation>
: 24
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B3FFF879EC4FF23FA83FF6A" author="Brown" box="[1203,1390,152,174]" page="104" pageId="22" pageNumber="537" year="1976">
<bibRefCitation id="2A5F4B425B3FFF879EC4FF23FAD6FF6A" author="Brown" box="[1203,1339,152,174]" pageId="22" pageNumber="537" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 104
</treatmentCitation>
.
<taxonomicName id="89CE4D305B3FFF879AE0FF0CFE4BFF08" box="[151,422,182,204]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="haematoda">
<emphasis id="7CBAEAA15B3FFF879AE0FF0CFE4BFF08" box="[151,422,182,204]" italics="true" pageId="22" pageNumber="537">Odontomachus haematoda</emphasis>
</taxonomicName>
<taxonomicName id="89CE4D305B3FFF879BDAFF0DFD14FF08" authority="Crawley, 1916: 368" authorityName="Crawley" authorityPageNumber="368" authorityYear="1916" box="[429,761,182,204]" pageId="22" pageNumber="537" rank="variety" variety="pallipes">
var.
<emphasis id="7CBAEAA15B3FFF879BAEFF0CFDC4FF08" box="[473,553,182,204]" italics="true" pageId="22" pageNumber="537">pallipes</emphasis>
<treatmentCitation id="CF6F10A25B3FFF879858FF0CFD14FF08" author="Crawley" box="[559,761,182,204]" page="368" pageId="22" pageNumber="537" year="1916">
<bibRefCitation id="2A5F4B425B3FFF879858FF0CFD2BFF08" author="Crawley" box="[559,710,182,204]" pageId="22" pageNumber="537" refString="Crawley, W. C. (1916) Ants from British Guiana. Annals and Magazine of Natural History, 17, 366 - 378. http: // dx. doi. org / 10.1080 / 00222931608693799" type="journal article" year="1916">Crawley, 1916</bibRefCitation>
: 368
</treatmentCitation>
</taxonomicName>
. Junior synonym of
<taxonomicName id="89CE4D305B3FFF8799B0FF0DFAC5FF08" ID-CoL="48QBS" authority="Brown, 1976: 104" authorityName="Brown" authorityPageNumber="104" authorityYear="1976" box="[967,1320,182,204]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3FFF8799B0FF0DFB8DFF08" box="[967,1120,182,204]" italics="true" pageId="22" pageNumber="537">O. haematodus</emphasis>
:
<treatmentCitation id="CF6F10A25B3FFF879E1AFF0DFAC5FF08" author="Brown" box="[1133,1320,182,204]" page="104" pageId="22" pageNumber="537" year="1976">
<bibRefCitation id="2A5F4B425B3FFF879E1AFF0DFB18FF08" author="Brown" box="[1133,1269,182,204]" pageId="22" pageNumber="537" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="89CE4D305B3FFF879AE0FF6FFE55FF2F" ID-CoL="48QBS" box="[151,440,213,235]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3FFF879AE0FF6FFE55FF2F" box="[151,440,213,235]" italics="true" pageId="22" pageNumber="537">Odontomachus haematodus</emphasis>
</taxonomicName>
<taxonomicName id="89CE4D305B3FFF879BB0FF6CFC56FF2F" authority="Emery, 1893: 91" authorityName="Emery" authorityPageNumber="91" authorityYear="1893" box="[455,955,213,235]" pageId="22" pageNumber="537" rank="variety" subSpecies="pubescens" variety="beuneipes">
r.
<emphasis id="7CBAEAA15B3FFF879B94FF6FFDA6FF2F" box="[483,587,213,235]" italics="true" pageId="22" pageNumber="537">pubescens</emphasis>
var.
<emphasis id="7CBAEAA15B3FFF8798FAFF6FFD1FFF2F" box="[653,754,213,235]" italics="true" pageId="22" pageNumber="537">beuneipes</emphasis>
<treatmentCitation id="CF6F10A25B3FFF879977FF6CFC56FF2F" author="Emery" box="[768,955,213,235]" page="91" pageId="22" pageNumber="537" year="1893">
<bibRefCitation id="2A5F4B425B3FFF879977FF6CFC60FF2F" author="Emery" box="[768,909,213,235]" pageId="22" pageNumber="537" refString="Emery, C. (1893) Notice sur quelques fourmis des iles Galapagos. Annales de la Societe Entomologique de France, 62, 89 - 92." type="journal article" year="1893">Emery, 1893</bibRefCitation>
: 91
</treatmentCitation>
</taxonomicName>
. Unavailable name; material referred to
<emphasis id="7CBAEAA15B3FFF879FF4FF6CFA71FF2F" box="[1411,1436,214,235]" italics="true" pageId="22" pageNumber="537">O.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B3FFF879AB0FF4EFDE0FECE" blockId="22.[151,1436,152,266]" box="[199,525,244,266]" pageId="22" pageNumber="537">
<treatmentCitationGroup id="6EDE119D5B3FFF879AB0FF4EFDE0FECE" box="[199,525,244,266]" pageId="22" pageNumber="537">
<taxonomicName id="89CE4D305B3FFF879AB0FF4EFDE4FECE" ID-CoL="48QBS" authority="Brown, 1976: 104" authorityName="Brown" authorityPageNumber="104" authorityYear="1976" box="[199,521,244,266]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3FFF879AB0FF4EFEACFECE" box="[199,321,244,266]" italics="true" pageId="22" pageNumber="537">haematodus</emphasis>
:
<treatmentCitation id="CF6F10A25B3FFF879B39FF4FFDE4FECE" author="Brown" box="[334,521,244,266]" page="104" pageId="22" pageNumber="537" year="1976">
<bibRefCitation id="2A5F4B425B3FFF879B39FF4FFE3BFECE" author="Brown" box="[334,470,244,266]" pageId="22" pageNumber="537" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="1AB1663B5B3FFF879AE0F82DFE82F82E" httpUri="https://zenodo.org/record/249913/files/figure.png" pageId="22" pageNumber="537" targetBox="[153,1433,318,1920]" targetPageId="22">
<paragraph id="4E7136B35B3FFF879AE0F82DFE82F82E" blockId="22.[151,1436,1943,2026]" pageId="22" pageNumber="537">
<emphasis id="7CBAEAA15B3FFF879AE0F82DFECEF869" bold="true" box="[151,291,1943,1965]" pageId="22" pageNumber="537">FIGURE 18.</emphasis>
<taxonomicName id="89CE4D305B3FFF879B5AF82DFDA1F869" box="[301,588,1943,1965]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3FFF879B5AF82DFDA1F869" box="[301,588,1943,1965]" italics="true" pageId="22" pageNumber="537">Odontomachus haematodus</emphasis>
</taxonomicName>
, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in dorsal view, (E) mesosoma in lateral view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence.
</paragraph>
</caption>
<caption id="1AB1663B5B3EFF869AE0F8A8FB72F883" httpUri="https://zenodo.org/record/249914/files/figure.png" pageId="23" pageNumber="538" targetBox="[153,1433,185,1796]" targetPageId="23">
<paragraph id="4E7136B35B3EFF869AE0F8A8FB72F883" blockId="23.[151,1436,1810,1863]" pageId="23" pageNumber="538">
<emphasis id="7CBAEAA15B3EFF869AE0F8A8FEF2F8EC" bold="true" box="[151,287,1810,1832]" pageId="23" pageNumber="538">FIGURE 19.</emphasis>
<taxonomicName id="89CE4D305B3EFF869B52F8A8FDD2F8EC" box="[293,575,1810,1832]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="538" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3EFF869B52F8A8FDD2F8EC" box="[293,575,1810,1832]" italics="true" pageId="23" pageNumber="538">Odontomachus haematodus</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) body in lateral view, (E) mesosoma in lateral view, (F) body in dorsal view, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<subSubSection id="06D465385B3EFF899AE0F8D4FBCFFED8" lastPageId="24" lastPageNumber="539" pageId="23" pageNumber="538" type="diagnosis">
<paragraph id="4E7136B35B3EFF899AE0F8D4FBCFFED8" blockId="23.[151,1436,1902,2035]" lastBlockId="24.[151,1437,151,2013]" lastPageId="24" lastPageNumber="539" pageId="23" pageNumber="538">
<emphasis id="7CBAEAA15B3EFF869AE0F8D4FEFCF843" bold="true" box="[151,273,1902,1927]" pageId="23" pageNumber="538">Diagnosis.</emphasis>
Workers in the Nearctic region are identified by their unique large size, dark color, lightly shagreened gastral dorsum, and presence of elongate metasternal processes between the hind coxae (
<figureCitation id="D6F52A365B3EFF869E01F828FB54F86E" box="[1142,1209,1938,1963]" captionStart="FIGURE 3" captionStartId="6.[151,249,1979,2001]" captionTargetBox="[151,1435,1277,1957]" captionTargetId="figure@6.[151,1435,1277,1958]" captionTargetPageId="6" captionText="FIGURE 3. Worker metasterna, anteroventral oblique view: (A) O. haematodus with both hindcoxae removed, arrow indicating elongate metasternal processes and (B) O. ruginodis with one hindcoxa removed, arrow indicating cleft ridge." httpUri="https://zenodo.org/record/249899/files/figure.png" pageId="23" pageNumber="538">Fig. 3</figureCitation>
), a character unique for the Nearctic fauna. The yellow colored males are most similar to
<emphasis id="7CBAEAA15B3EFF8699C8F80DFBBEF80A" box="[959,1107,1974,1998]" italics="true" pageId="23" pageNumber="538">
<taxonomicName id="89CE4D305B3EFF8699C8F80DFBA2F80A" box="[959,1103,1974,1998]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="538" phylum="Arthropoda" rank="species" species="brunneus">O. brunneus</taxonomicName>
,
</emphasis>
but can be separated by the following: 1) ocelli small, weakly bulging past posterior head margin; 2) posterior propodeal face offset dorsally and laterally by distinct carina; 3) metasternal processes long and narrow; and by numerous genitalic characters. Genitalia of
<taxonomicName id="89CE4D305B31FF899B52FF06FE3DFF10" box="[293,464,188,212]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B31FF899B52FF06FE3DFF10" box="[293,464,188,212]" italics="true" pageId="24" pageNumber="539">O. haematodus</emphasis>
</taxonomicName>
are unique among Nearctic species, with the telomere distinctly taller than long. The ninth abdominal sternum is most similar to that of
<taxonomicName id="89CE4D305B31FF8998AFFF5AFC8AFF3C" box="[728,871,224,248]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B31FF8998AFFF5AFC8AFF3C" box="[728,871,224,248]" italics="true" pageId="24" pageNumber="539">O. ruginodis</emphasis>
</taxonomicName>
, having similarly proportioned discs and strongly constricted posterior lobes, but
<taxonomicName id="89CE4D305B31FF899B80FEBEFD4DFED8" box="[503,672,260,284]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B31FF899B80FEBEFD4DFED8" box="[503,672,260,284]" italics="true" pageId="24" pageNumber="539">O. haematodus</emphasis>
</taxonomicName>
has a broader posterior lobe apex.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B31FF899AB0FE9DFE29F9E9" pageId="24" pageNumber="539" type="description">
<paragraph id="4E7136B35B31FF899AB0FE9DFBFDFCF3" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="539">
<emphasis id="7CBAEAA15B31FF899AB0FE9DFEB4FE84" bold="true" box="[199,345,295,320]" pageId="24" pageNumber="539">Description.</emphasis>
Worker: HL 2.322.75, HW 1.882.16, SL 2.162.48, EL 0.440.54, ML 1.381.56, WL 3.043.48, PTH 1.221.38, PTL 0.540.60 (n=10). Entire body generally shiny except where dulled by dense pubescence; concolorous dark reddish-brown to blackish. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; with numerous, fine, appressed pubescence and scattered elongate, erect setae present (dorsally). Pronotum with somewhat circular concentric striae that become longitudinal near rear margin; appressed pubescence abundant; 5-8 elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; propleuron, mesopleuron, and basalar lobe lacking sculpture; abundant pubescence present dorsally. Metasternum with paired narrowed, elongate, spiniform processes between hind coxae (
<figureCitation id="D6F52A365B31FF899A9DFDD6FEC3FD40" box="[234,302,620,645]" captionStart="FIGURE 3" captionStartId="6.[151,249,1979,2001]" captionTargetBox="[151,1435,1277,1957]" captionTargetId="figure@6.[151,1435,1277,1958]" captionTargetPageId="6" captionText="FIGURE 3. Worker metasterna, anteroventral oblique view: (A) O. haematodus with both hindcoxae removed, arrow indicating elongate metasternal processes and (B) O. ruginodis with one hindcoxa removed, arrow indicating cleft ridge." httpUri="https://zenodo.org/record/249899/files/figure.png" pageId="24" pageNumber="539">Fig. 3</figureCitation>
A). Petiole widest at base, gradually tapering apically to a long spine directed rearward; transverse striae completely surrounding petiole except upper dorso-posteriorly where striae are faint or lacking; subpetiolar process somewhat anvil shaped; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but weakly shagreened (seen at high magnification); fine, appressed pubescence moderately dense, spaces between hairs usually less than 1/2 the length of a hair; scattered erect, elongate setae present.
</paragraph>
<paragraph id="4E7136B35B31FF899AB0FCFEFAA5FA59" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="539">Male: HL 1.001.12, HW 1.201.30, SL 0.180.20, EL 0.620.68, EW 0.360.40, OL 0.180.20, OES 0.200.22, WL 2.682.83, PTH 0.900.94, PTL 0.500.56, FWL 4.705.45 (n=5). Body generally shiny except where obscured by dense pubescence; head, meso- and metasoma yellowish-orange, antenna and legs pale yellowish-orange. Head and body with abundant fine, white pubescence except on anepisternum, where it is mostly absent. Eyes extremely large, maximum diameter of each eye at least 70% of the length of the head in full-face view. Ocelli small to average in size, the length of each ocellus slightly less to approximately the same as the distance between lateral ocellus and eye margin; in full-face view, lateral ocelli do not protrude beyond posterior border of head. Mesosoma: pronotum with faint transverse striae, especially laterally, but mostly appearing to lack sculpture; mesoscutum with faint transversely arcuate striae anteriorly, striae becoming transverse posteriorly; mesoscutellum raised and convex, lacking sculpture; propodeum evenly rounded without obvious declivious face, with strong rugoreticulation; mesopleuron mostly lacking striae or with very faint longitudinal striae. Petiole bluntly rounded apically, with rounded triangular subpetiolar process anteriorly; densely pubescent anteriorly and laterally, but reduced pubescence posteriorly. Abdominal sternum IX disc subrectangular, breadth about twice length; posterior lobe length about twice maximum lobe width, basal half strongly narrowed. Telomeral apex very narrowly rounded, nearly angular; telomere length distinctly shorter than height; valviceps apex weakly produced, broad; apical margin of valviceps linear; vertical portion of dorsolateral carina and lateral margin of subapical lamina meeting at an angle; apicodorsal lobe of valviceps narrow, somewhat short; subapical lamina broad.</paragraph>
<paragraph id="4E7136B35B31FF899AB0FA12FE29F9E9" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="539">Queen: HL 2.482.55, HW 2.142.00, SL 2.352.36, EL 0.520.55, ML 1.561.58, OL 0.120.13, WL 3.56- 3.60, PTH 1.281.62, PTL 0.580.66, FWL 6.5 (n=2 except for FWL for which only one individual had wings). Similar to workers in color and general appearance except with mesosoma developed for wings and with wings present in alate specimens.</paragraph>
</subSubSection>
<subSubSection id="06D465385B31FF899AB0F98DFBECF818" pageId="24" pageNumber="539" type="distribution">
<paragraph id="4E7136B35B31FF899AB0F98DFBECF818" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="539">
<emphasis id="7CBAEAA15B31FF899AB0F98DFD80F994" bold="true" box="[199,621,1591,1616]" pageId="24" pageNumber="539">
Distribution (
<figureCitation id="D6F52A365B31FF899B1DF98DFE2CF994" box="[362,449,1591,1616]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="24" pageNumber="539">Fig. 26</figureCitation>
, Appendix 1):
</emphasis>
Due to misidentifications, the worldwide distribution of this species is not clearly understood at this time. However, based on recent examination of numerous specimens from the
<collectingCountry id="36D976235B31FF899F3EF9E6FA82F9B0" box="[1353,1391,1628,1652]" name="United States of America" pageId="24" pageNumber="539">US</collectingCountry>
, the distributional records for this species from this country given here (Appendix 1) can be considered reliable. Additionally, specimens from
<collectingCountry id="36D976235B31FF899B84F91EFDD7F979" box="[499,570,1700,1725]" name="Brazil" pageId="24" pageNumber="539">Brazil</collectingCountry>
,
<collectingCountry id="36D976235B31FF89983EF91EFD1EF979" box="[585,755,1700,1725]" name="Guyana" pageId="24" pageNumber="539">British Guiana</collectingCountry>
, and
<collectingCountry id="36D976235B31FF899943F91FFC85F978" box="[820,872,1701,1724]" name="Peru" pageId="24" pageNumber="539">Peru</collectingCountry>
were identified as
<taxonomicName id="89CE4D305B31FF899E3EF91EFB18F978" box="[1097,1269,1700,1724]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B31FF899E3EF91EFB18F978" box="[1097,1269,1700,1724]" italics="true" pageId="24" pageNumber="539">O. haematodus</emphasis>
</taxonomicName>
by MacGown [LSAM, UGCA, and UMMZ].
<bibRefCitation id="2A5F4B425B31FF899B80F972FD78F924" author="Brown" box="[503,661,1735,1760]" pageId="24" pageNumber="539" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
reported this species' distribution as continental South
<collectingCountry id="36D976235B31FF899E89F97DFAB3F924" box="[1278,1374,1735,1760]" name="United States of America" pageId="24" pageNumber="539">America</collectingCountry>
from Orinoco Delta to Tucuma,
<collectingCountry id="36D976235B31FF899BB8F956FDAFF8C1" box="[463,578,1772,1797]" name="Argentina" pageId="24" pageNumber="539">Argentina</collectingCountry>
.
<bibRefCitation id="2A5F4B425B31FF899827F956FCFDF8C0" author="McGlynn" box="[592,784,1772,1797]" pageId="24" pageNumber="539" refString="McGlynn, T. P. (1999) The worldwide transfer of ants: Geographical distribution and ecological invasions. Journal of Biogeography, 26, 535 - 548. http: // dx. doi. org / 10.1046 / j. 1365 - 2699.1999.00310. x" type="journal article" year="1999">McGlynn (1999)</bibRefCitation>
reported
<taxonomicName id="89CE4D305B31FF8999F6F956FBC0F8C0" box="[897,1069,1772,1796]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B31FF8999F6F956FBC0F8C0" box="[897,1069,1772,1796]" italics="true" pageId="24" pageNumber="539">O. haematodus</emphasis>
</taxonomicName>
as being introduced to Hawaii; however, these records have not been verified, nor is this species thought to have been established in that state.
<bibRefCitation id="2A5F4B425B31FF899AE0F88EFE29F889" box="[151,452,1844,1869]" pageId="24" pageNumber="539" refString="Krushelnycky, P. D., Loope, L. L. &amp; Reimer, N. J. (2005) The ecology, policy, and management of ants in Hawaii. Proceedings of the Hawaiian Entomological Society, 37, 1 - 25." type="journal article">
Krushelnycky
<emphasis id="7CBAEAA15B31FF899B4CF88FFE87F888" box="[315,362,1844,1868]" italics="true" pageId="24" pageNumber="539">et al</emphasis>
. (2005)
</bibRefCitation>
did not include
<taxonomicName id="89CE4D305B31FF89980DF88EFCCEF888" box="[634,803,1844,1868]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B31FF89980DF88EFCCEF888" box="[634,803,1844,1868]" italics="true" pageId="24" pageNumber="539">O. haematodus</emphasis>
</taxonomicName>
in their recent list of ant species from Hawaii. In the
<collectingCountry id="36D976235B31FF899F0FF88EFA71F888" box="[1400,1436,1844,1868]" name="United States of America" pageId="24" pageNumber="539">US</collectingCountry>
we have verified records of established populations of this species only from the Gulf Coast region, specifically from: Alabama: Baldwin, Escambia, and Mobile Counties. Florida: Escambia County. Louisiana: Orleans Parish. Mississippi: Greene and Jackson Counties. Distributional information for
<collectingCountry id="36D976235B31FF89998FF81AFBF1F87C" box="[1016,1052,1952,1976]" name="United States of America" pageId="24" pageNumber="539">US</collectingCountry>
specimens from AntWeb (2013) (Florida record), and examination of specimens in AUEM, MEM and USNM.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B30FF889AB0FF2DFEC5FAC9" pageId="25" pageNumber="540" type="discussion">
<paragraph id="4E7136B35B30FF889AB0FF2DFAF8FEA0" blockId="25.[151,1437,151,1293]" pageId="25" pageNumber="540">
<emphasis id="7CBAEAA15B30FF889AB0FF2DFEA1FF74" bold="true" box="[199,332,151,176]" pageId="25" pageNumber="540">Discussion.</emphasis>
<taxonomicName id="89CE4D305B30FF889B22FF22FD63FF74" box="[341,654,152,176]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF889B22FF22FD63FF74" box="[341,654,152,176]" italics="true" pageId="25" pageNumber="540">Odontomachus haematodus</emphasis>
</taxonomicName>
is native to South
<collectingCountry id="36D976235B30FF88991DFF2DFC23FF74" box="[874,974,151,176]" name="United States of America" pageId="25" pageNumber="540">America</collectingCountry>
. The earliest record in the
<collectingCountry id="36D976235B30FF889F73FF22FAC5FF74" box="[1284,1320,152,176]" name="United States of America" pageId="25" pageNumber="540">US</collectingCountry>
we found was of three specimens collected on
<date id="3A7010735B30FF889858FF06FD58FF11" box="[559,693,188,213]" pageId="25" pageNumber="540" value="1956-06-01">1 June 1956</date>
from Mobile, Alabama. These specimens were borrowed from the USNM and examined by MacGown. Earliest MEM records are from 2000 from Baldwin County, Alabama, by which time this species had become locally abundant. Until recently, specimens of this species from the Gulf Coast were identified as
<emphasis id="7CBAEAA15B30FF889B06FE92FD74FE84" box="[369,665,296,320]" italics="true" pageId="25" pageNumber="540">
<taxonomicName id="89CE4D305B30FF889B06FE92FE11FE84" box="[369,508,296,320]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="insularis">O. insularis</taxonomicName>
,
<taxonomicName id="89CE4D305B30FF88987CFE92FD74FE84" box="[523,665,296,320]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="brunneus">O. brunneus</taxonomicName>
</emphasis>
, and/or
<emphasis id="7CBAEAA15B30FF88988AFE92FC7AFE84" box="[765,919,296,320]" italics="true" pageId="25" pageNumber="540">
<taxonomicName id="89CE4D305B30FF88988AFE92FC7EFE84" box="[765,915,296,320]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="ruginodis">O. ruginodis</taxonomicName>
.
</emphasis>
However, after examination of workers and males of all three species, it became clear that this now-common Gulf Coast species is
<taxonomicName id="89CE4D305B30FF889E13FEF6FAE0FEA0" box="[1124,1293,332,356]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF889E13FEF6FAE0FEA0" box="[1124,1293,332,356]" italics="true" pageId="25" pageNumber="540">O. haematodus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4E7136B35B30FF889AB0FED5FD4DFC0C" blockId="25.[151,1437,151,1293]" pageId="25" pageNumber="540">
This species has been collected in a variety of habitats along the Gulf Coast including bottomland forests, pine forests, in a mixed forest at the edge of estuary, at the edge of a pitcher plant bog, fore dunes, shell midden, pine savanna, sandhill, and in residential areas. In non-urban areas, nests have been discovered in the soil, in leaf litter, in rotting logs and stumps, at tree bases, in dead parts of standing live trees (colonies observed in tree cavities at least 2.0 m from ground level); in urban areas nests have been discovered under mulch, debris, under roots of ornamental grass bushes, in building foundations, and under PVC pipes. Colonies range in size from less than a hundred workers to several thousand with some colonies polygynous. Similar to other members of the genus, workers are typically nocturnal, but may also forage during the day. In areas in the Gulf Coast where this species occurs, populations may be extremely abundant, especially in parts of Baldwin and Mobile Counties, Alabama; and Jackson County, Mississippi. Large populations have also been observed at the Audubon Zoo in New Orleans, Louisiana (
<collectingCountry id="36D976235B30FF889B60FD62FEA1FD34" box="[279,332,728,752]" name="Iran" pageId="25" pageNumber="540">Pers</collectingCountry>
. Comm. Fred Larabee and Andy Saurez). Compared to other
<collectingCountry id="36D976235B30FF889E69FD62FBAFFD34" box="[1054,1090,728,752]" name="United States of America" pageId="25" pageNumber="540">US</collectingCountry>
species,
<taxonomicName id="89CE4D305B30FF889ED9FD62FAB7FD34" box="[1198,1370,728,752]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF889ED9FD62FAB7FD34" box="[1198,1370,728,752]" italics="true" pageId="25" pageNumber="540">O. haematodus</emphasis>
</taxonomicName>
is an aggressive stinger. Upon placing one's hand in leaf litter where a nest is located, workers may immediately sting. The sting is painful, but not long lasting (MacGown, personal observation). Homeowner's in southern Mississippi have complained of bites and stings received from this species (MacGown, unpublished data). One homeowner stated, &quot;[the] ants would latch on with their mandibles and sting repeatedly, lashing out with their abdomens. One ant can sting 45 times before you realize it. They are very aggressive.&quot; Alate males and queens have been collected from late June through early August.
</paragraph>
<paragraph id="4E7136B35B30FF889AB0FC6FFEC5FAC9" blockId="25.[151,1437,151,1293]" pageId="25" pageNumber="540">
It is clear from collecting records that the abundance of
<taxonomicName id="89CE4D305B30FF889911FC6EFBF9FC28" box="[870,1044,980,1004]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF889911FC6EFBF9FC28" box="[870,1044,980,1004]" italics="true" pageId="25" pageNumber="540">O. haematodus</emphasis>
</taxonomicName>
along the Gulf Coast is a recent phenomenon, and it is also clear that this species is continuing to spread. This suggests that
<taxonomicName id="89CE4D305B30FF889ED9FC42FAB5FBD4" box="[1198,1368,1016,1040]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF889ED9FC42FAB5FBD4" box="[1198,1368,1016,1040]" italics="true" pageId="25" pageNumber="540">O. haematodus</emphasis>
</taxonomicName>
spent around 50 years in North
<collectingCountry id="36D976235B30FF889BCCFBA6FDF6FBF1" box="[443,539,1052,1077]" name="United States of America" pageId="25" pageNumber="540">America</collectingCountry>
before some threshold condition allowed it to form such large populations that they became conspicuous. This kind of history would not be unique to
<taxonomicName id="89CE4D305B30FF8899CAFBFAFB8AFB9C" box="[957,1127,1088,1112]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF8899CAFBFAFB8AFB9C" box="[957,1127,1088,1112]" italics="true" pageId="25" pageNumber="540">O. haematodus</emphasis>
</taxonomicName>
.
<taxonomicName id="89CE4D305B30FF889E01FBFAFA81FB9C" box="[1142,1388,1088,1112]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="7CBAEAA15B30FF889E01FBFAFA81FB9C" box="[1142,1388,1088,1112]" italics="true" pageId="25" pageNumber="540">Camponotus planatus</emphasis>
</taxonomicName>
, for example was restricted to a small area of tropical Florida for many decades, arriving some time before 1910 (
<bibRefCitation id="2A5F4B425B30FF889AE9FB32FE69FB64" author="Deyrup" box="[158,388,1159,1184]" pageId="25" pageNumber="540" refString="Deyrup, M., Cover, S. &amp; Davis, L. (2000) Exotic ants in Florida. Transactions of the American Entomological Society, 126, 293 - 325." type="journal article" year="2000">Deyrup et al. 2000</bibRefCitation>
). This species is now common in some sites (including some natural habitats) in central peninsular Florida (Deyrup 2014, unpublished data) and has even been recorded in Mississippi (
<bibRefCitation id="2A5F4B425B30FF889EA3FB16FA7CFB00" author="MacGown" box="[1236,1425,1196,1220]" pageId="25" pageNumber="540" refString="MacGown, J. A. (2010) Camponotus planatus (Hymenoptera: Formicidae), an exotic carpenter ant found in Mississippi. Journal of the Mississippi Academy of Sciences, 55, 187 - 188." type="journal article" year="2010">MacGown 2010</bibRefCitation>
). It is always possible, however, that the 1956 collection of
<taxonomicName id="89CE4D305B30FF889915FB6AFBFCFB2C" box="[866,1041,1232,1256]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B30FF889915FB6AFBFCFB2C" box="[866,1041,1232,1256]" italics="true" pageId="25" pageNumber="540">O. haematodus</emphasis>
</taxonomicName>
represents an independent failed introduction.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="58AE2320815ABE657679544A0C676CEA">A checklist of vascular plants of the W National Park in Burkina Faso, including the adjacent hunting zones of Tapoa-Djerma and Kondio</mods:title>
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<mods:namePart id="B68CCF0A802611AA7E7AB6992654CB5A">Nacoulma, Blandine M. I.</mods:namePart>
<mods:affiliation id="435FE4C4857268E0FD62B8BEF6EE2D4E">Universite Joseph Ki-Zerbo, Ouagadougou, Burkina Faso</mods:affiliation>
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Willd.
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<paragraph id="E684C61F43BB14E9431714A77C68268C" pageId="0" pageNumber="54205">Pantropical</paragraph>
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<paragraph id="F8F7236EE9469E1FBA7D05F51358FF75" pageId="0" pageNumber="54205">Life Form: phanerophyte; Voucher: Nacoulma 4538 (OUA-17107)</paragraph>
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<bibRefCitation id="3858D625686477A66A23AD3539B21C07" author="Papp, J." journalOrPublisher="Entomologische Abhandlungen" pageId="0" pageNumber="8151" pagination="95 - 140" title="Palaearctic species of Microgaster Latreille (= Microplitis Foerster) with description of seven new species (Hymenoptera, Braconidae, Microgastrinae)" volume="47" year="1984">Papp (1984)</bibRefCitation>
suggests that semicircularis (Ratzeburg, 1844,
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) (type destroyed) may be a senior synonym.
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