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<paragraph id="76D896A9B24E4469A81F5D348A49C8CD" pageId="0" pageNumber="113952">Distribution</paragraph>
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<paragraph id="891E1EF725672936FFB6DE1D46814710" pageId="0" pageNumber="113952">South Russian Far East to China, South Sakhalin to Japan</paragraph>
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<taxonomicName id="89B284D62563FFCFFF31FBF02D36F568" ID-CoL="Y5QH" authority="Bui, Ziegler & Bonkowski" authorityName="Bui, Ziegler & Bonkowski" authorityYear="2019" box="[151,859,1078,1104]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
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<emphasis id="7CC623472563FFCFFF31FBF02E8AF568" bold="true" box="[151,231,1078,1104]" italics="true" pageId="3" pageNumber="54">Copris</emphasis>
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(
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)
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Bui, Ziegler & Bonkowski
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,
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(
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<figureCitation id="D689E3D02563FFCFFF39FB9C2E85F54C" box="[159,232,1114,1140]" captionStart="FIGURES 1" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[215,1374,179,1936]" captionTargetId="figure@5.[206,1381,166,1945]" captionTargetPageId="5" captionText="FIGURES 1A–D. Holotype (A, B) and female paratype (C, D) of Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species. A, C, habitus, dorsal view; B, D, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3586828" httpUri="https://zenodo.org/record/3586828/files/figure.png" pageId="3" pageNumber="54">Figs. 1</figureCitation>
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A–D, 2A, C–E, 3A)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2563FFCFFF31FB642DD6F43C" pageId="3" pageNumber="54" type="materials_examined">
|
||||
<paragraph id="4E0DFF552563FFCFFF31FB642C13F584" blockId="3.[151,1437,1185,2041]" box="[151,638,1185,1213]" pageId="3" pageNumber="54">
|
||||
<materialsCitation id="FEDAF5082563FFCFFF31FB642C16F584" ID-GBIF-Occurrence="2521263207" box="[151,635,1185,1213]" country="Indonesia" location="Mt. Moeria" pageId="3" pageNumber="54" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
|
||||
<emphasis id="7CC623472563FFCFFF31FB642F5BF584" bold="true" box="[151,310,1185,1212]" pageId="3" pageNumber="54">
|
||||
<typeStatus id="910941F72563FFCFFF31FB642EBFF584" box="[151,210,1186,1212]" pageId="3" pageNumber="54">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="36A5BFC52563FFCFFE9AFB642FC1F584" box="[316,428,1186,1212]" name="Indonesia" pageId="3" pageNumber="54">Indonesia</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="8C7631B72563FFCFFE11FB652F86F585" box="[439,491,1187,1213]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="54">Java</collectingRegion>
|
||||
,
|
||||
<location id="4B6DA98E2563FFCFFE51FB652C16F584" LSID="urn:lsid:plazi:treatment:C61B4E432563FFCAFF31FBF02A5CF687:4B6DA98E2563FFCFFE51FB652C16F584" box="[503,635,1186,1213]" country="Indonesia" name="Mt. Moeria" pageId="3" pageNumber="54" stateProvince="Jawa Timur">Mt. Moeria</location>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF61FB002DD6F43C" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
|
||||
<emphasis id="7CC623472563FFCFFF61FB002FECF5D8" bold="true" box="[199,385,1221,1248]" pageId="3" pageNumber="54">Type specimen.</emphasis>
|
||||
<materialsCitation id="FEDAF5082563FFCFFE2CFB002BF4F5D8" ID-GBIF-Occurrence="2521263204" box="[394,1433,1222,1249]" collectionCode="MNHN, R" country="Indonesia" location="Oberthur" municipality="Mt. Moeria" pageId="3" pageNumber="54" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
|
||||
<emphasis id="7CC623472563FFCFFE2CFB002C6DF5D8" bold="true" box="[394,512,1222,1248]" pageId="3" pageNumber="54">
|
||||
<typeStatus id="910941F72563FFCFFE2CFB002F96F5D8" box="[394,507,1222,1248]" pageId="3" pageNumber="54" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
male (
|
||||
<collectionCode id="28A367902563FFCFFDF7FB002CC1F5D8" box="[593,684,1222,1248]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="54">MNHN</collectionCode>
|
||||
), “
|
||||
<collectingCountry id="36A5BFC52563FFCFFD6AFB002D53F5D8" box="[716,830,1222,1248]" name="Indonesia" pageId="3" pageNumber="54">Indonesia</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="8C7631B72563FFCFFCEAFB012D13F5D9" box="[844,894,1223,1249]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="54">Java</collectingRegion>
|
||||
|
|
||||
<collectingMunicipality id="AE69652F2563FFCFFC30FB012A76F5D8" box="[918,1051,1222,1249]" pageId="3" pageNumber="54">Mt. Moeria</collectingMunicipality>
|
||||
, 3–4000’ | ex Coll.
|
||||
<collectionCode id="28A367902563FFCFFAA7FB012B7AF5D9" box="[1281,1303,1223,1249]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="54">R</collectionCode>
|
||||
.
|
||||
<location id="4B6DA98E2563FFCFFA82FB002BFDF5D8" LSID="urn:lsid:plazi:treatment:C61B4E432563FFCAFF31FBF02A5CF687:4B6DA98E2563FFCFFA82FB002BFDF5D8" box="[1316,1424,1222,1248]" country="Indonesia" municipality="Mt. Moeria" name="Oberthur" pageId="3" pageNumber="54" stateProvince="Jawa Timur">Oberthür</location>
|
||||
”
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="FEDAF5082563FFCFFF31FB2C2DDAF43C" ID-GBIF-Occurrence="2521263206" box="[151,951,1258,1285]" collectionCode="MNHN" country="Indonesia" location="Oberthur" municipality="Mt. Moeria" pageId="3" pageNumber="54" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Jawa Timur" typeStatus="paratype">
|
||||
<emphasis id="7CC623472563FFCFFF31FB2C2F75F43C" bold="true" box="[151,280,1258,1284]" pageId="3" pageNumber="54">
|
||||
<typeStatus id="910941F72563FFCFFF31FB2C2F7EF43C" box="[151,275,1258,1284]" pageId="3" pageNumber="54" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCount id="58B434DC2563FFCFFEB9FB2C2F18F43C" box="[287,373,1258,1284]" pageId="3" pageNumber="54" type="male">4 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="58B434DC2563FFCFFE0AFB2D2C75F43C" box="[428,536,1258,1285]" pageId="3" pageNumber="54" type="female">7 females</specimenCount>
|
||||
(
|
||||
<collectionCode id="28A367902563FFCFFD81FB2C2CEFF43C" box="[551,642,1258,1284]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="54">MNHN</collectionCode>
|
||||
), same data as for holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2563FFC8FF61FACB2ADCF267" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54" type="description">
|
||||
<paragraph id="4E0DFF552563FFCFFF61FACB2B39F474" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
|
||||
<emphasis id="7CC623472563FFCFFF61FACB2F87F410" bold="true" box="[199,490,1293,1320]" pageId="3" pageNumber="54">
|
||||
Description of
|
||||
<typeStatus id="910941F72563FFCFFEDAFAC82F88F410" box="[380,485,1294,1320]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Total body length
|
||||
<quantity id="894A52B02563FFCFFCB7FACB2D17F411" box="[785,890,1293,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="54" unit="mm" value="15.5">15.5 mm</quantity>
|
||||
, maximum body width
|
||||
<quantity id="894A52B02563FFCFFB29FACB2A87F411" box="[1167,1258,1293,1321]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="54" unit="mm" value="7.5">7.5 mm</quantity>
|
||||
. Whole surface chocolate brown. Mouthparts and antennomeres 1–6 reddish brown. Antennal club and setae on legs yellow.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF61FA902B22F738" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
|
||||
Head
|
||||
<quantity id="894A52B02563FFCFFEACFA902F0CF449" box="[266,353,1366,1393]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="54" unit="mm" value="4.2">4.2 mm</quantity>
|
||||
long and 5.0 mm wide. Clypeal surface smooth and impunctate; clypeal apex strongly upwards protruded, and widely and shallowly emarginate. Genae rectangular, distinctly separated from clypeus by well-defined suture; most surface of genae smooth, except for small areas in middle sparsely and weakly punctate. Surface of vertex distinctly punctate; punctures denser and deeper at areas near eyes. Cephalic horn small but distinct. Antenna with 9 antennomeres; antennomere 1 longer than antennomeres 2–6 combined and also antennal club.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF61F9CD2B77F7A8" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
|
||||
Pronotum
|
||||
<quantity id="894A52B02563FFCFFE9BF9CF2FFEF71D" box="[317,403,1545,1573]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="54" unit="mm" value="5.2">5.2 mm</quantity>
|
||||
long and
|
||||
<quantity id="894A52B02563FFCFFDA4F9CC2C34F71D" box="[514,601,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="54" unit="mm" value="6.8">6.8 mm</quantity>
|
||||
wide, and without anterior declivity. Pronotal anterior margin slightly forwards curved in middle; anterolateral angles obtuse. Pronotal disc with weak median sulcus; sulcus not extending to pronotal anterior margin. Whole surface of pronotum deeply and densely punctate; most punctures equal in size, except for punctures near sulcus slightly larger. Each pronotal side with one deep and large excavation.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF61F95C2AE4F78C" blockId="3.[151,1437,1185,2041]" box="[199,1161,1690,1717]" pageId="3" pageNumber="54">Anterior angles of prothorax flat, and shallowly and sparsely punctate on ventral side.</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF61F9782C94F618" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
|
||||
Elytra (ElyL
|
||||
<quantity id="894A52B02563FFCFFEFFF97B2FDDF7E1" box="[345,432,1725,1753]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="54" unit="mm" value="8.5">8.5 mm</quantity>
|
||||
) deeply striate; striae weakly, sparsely and equally punctate. Elytral intervals convex, and deeply and densely punctate; interval punctures evenly distributed and relatively equal in size. Each elytron with one rather abrupt vertical flattening near sutural angle.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF61F8EC2EBAF6B5" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">Mesepimeron and metepisternum flat and weakly punctate. Mesosternum strongly rugose. Metasternum relatively smooth, and with median distinct groove; sides of metasternum sparsely and shallowly punctate, and without setae.</paragraph>
|
||||
<paragraph id="4E0DFF552563FFC8FF61F8512FBDF0EB" blockId="3.[151,1437,1185,2041]" lastBlockId="4.[151,1437,153,1007]" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54">Legs. Ventral surface of profemora strongly punctate; punctures unevenly distributed, and unequal in size, becoming denser and smaller towards anterior margin of profemora; a few punctures at posterior half of profemora with yellow and long setae. Anterior margin of profemora with yellow, dense and long setae. Protibia with four lateral teeth, and protibial surface strongly and unevenly punctate. Protibial spurs pointed, slightly curved outwards near apex, and equal in length to protarsomeres 2–4 combined. Protarsomere 1 and 5 equal in length, and both slightly longer than individual protarsomere 2–4. Mesofemora smooth and impunctate on ventral side. Mesotibia smooth and broadly dilated at posterior end; mesotibia with two spurs, one of which blunt, one longer and pointed. Mesotarsus with five tarsomeres, and all mesotarsomeres with yellow scanty setae; basal mesotarsomere elongated, almost equal in length to mesotibial spurs. Most ventral surface of metafemora smooth and impunctate, except for apical third sparsely punctate. Metatibia smooth and broadly dilated at posterior end, and with one spur pointed. Metatarsus with five tarsomeres covered with yellow scantly setae; basal metatarsomere apically enlarged, and shorter than metatibial spur.</paragraph>
|
||||
<paragraph id="4E0DFF552564FFC8FF61FE1B2FEAF307" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
|
||||
Abdomen and pygidium. Abdominal ventrites smooth, impunctate, and narrower at midline. Pygidium (PyL
|
||||
<quantity id="894A52B02564FFC8FADFFE1A2EABF323" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="55" unit="mm" value="1.5">1.5 mm</quantity>
|
||||
; PyW
|
||||
<quantity id="894A52B02564FFC8FEB4FDC72F04F323" box="[274,361,513,539]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="55" unit="mm" value="3.1">3.1 mm</quantity>
|
||||
) with margin complete; pygidial surface slightly convex, and deeply punctate; punctures dense but clearly separated.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552564FFC8FF61FD8F2A7FF3BF" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">Aedeagus. Phallobase slightly longer and wider than parameres. A broad angle almost reaching 180º formed by phallobase and parameres. Parameres slightly curved downwards in apical part.</paragraph>
|
||||
<paragraph id="4E0DFF552564FFC8FF61FD562A16F3CB" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FF61FD562FDAF393" bold="true" box="[199,439,656,683]" pageId="4" pageNumber="55">Sexual dimorphism.</emphasis>
|
||||
Female
|
||||
<typeStatus id="910941F72564FFC8FDBEFD572CEEF393" box="[536,643,657,683]" pageId="4" pageNumber="55" type="paratype">paratypes</typeStatus>
|
||||
differ from the male
|
||||
<typeStatus id="910941F72564FFC8FCD3FD572DBAF393" box="[885,983,657,683]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
|
||||
in the following characters: Clypeal anterior margin not protruded in middle, and without emargination. Head with one slightly curved transverse carina between clypeus and frons. Pronotal anterior margin strongly forwards curved in middle.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552564FFC8FF61FD3A2ADCF267" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FF61FD3A2F2AF22F" bold="true" box="[199,327,764,791]" pageId="4" pageNumber="55">Variability</emphasis>
|
||||
. Body length
|
||||
<quantity id="894A52B02564FFC8FE4EFD3A2CE1F22F" box="[488,652,764,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.65" metricValueMin="1.15" pageId="4" pageNumber="55" unit="mm" value="14.0" valueMax="16.5" valueMin="11.5">11.5–16.5 mm</quantity>
|
||||
, body width 6.0–
|
||||
<quantity id="894A52B02564FFC8FCF7FD3A2DCAF22F" box="[849,935,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="55" unit="mm" value="7.5">7.5 mm</quantity>
|
||||
(n=12). The two males are larger, compared to the
|
||||
<typeStatus id="910941F72564FFC8FF79FCE72F2CF203" box="[223,321,801,827]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
|
||||
possess a more strongly protruded clypeus upwards but similar in the remaining characters. There are no significant differences in external morphology among the female specimens examined.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2564FFC8FF61FCAE2C82F640" pageId="4" pageNumber="55" type="diagnosis">
|
||||
<paragraph id="4E0DFF552564FFC8FF61FCAE2C37F2D7" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FF61FCAE2FA3F2BB" bold="true" box="[199,462,872,899]" pageId="4" pageNumber="55">Differential diagnosis.</emphasis>
|
||||
The new species strongly resembles
|
||||
<taxonomicName id="89B284D62564FFC8FCC8FCAF2AB1F2BB" authorityName="Wiedemann" authorityYear="1823" box="[878,1244,873,899]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FCC8FCAF2DD7F2BB" box="[878,954,873,899]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FC6CFCAC2A25F2BB" box="[970,1096,874,899]" italics="true" pageId="4" pageNumber="55">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FBFFFCAF2AB1F2BB" box="[1113,1244,873,899]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FB44FCAF2B3FF2BB" box="[1250,1362,873,899]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
|
||||
that is widely distributed in Asia:
|
||||
<collectingCountry id="36A5BFC52564FFC8FE6DFC4B2C40F29F" box="[459,557,909,935]" name="Vietnam" pageId="4" pageNumber="55">Vietnam</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FD9FFC4B2C1FF29F" box="[569,626,909,935]" name="Laos" pageId="4" pageNumber="55">Laos</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FDD8FC4B2C99F29F" box="[638,756,909,935]" name="Cambodia" pageId="4" pageNumber="55">Cambodia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FCA6FC4B2D0BF29F" box="[768,870,909,935]" name="Thailand" pageId="4" pageNumber="55">Thailand</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FCD5FC4B2D8EF29F" box="[883,995,909,935]" name="Indonesia" pageId="4" pageNumber="55">Indonesia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FC49FC4B2A35F29F" box="[1007,1112,909,935]" name="Malaysia" pageId="4" pageNumber="55">Malaysia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FBC3FC4B2AB9F29F" box="[1125,1236,909,935]" name="Myanmar" pageId="4" pageNumber="55">Myanmar</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="36A5BFC52564FFC8FB46FC4B2B4EF29F" box="[1248,1315,909,935]" name="China" pageId="4" pageNumber="55">China</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="36A5BFC52564FFC8FAFAFC4B2BF5F29F" box="[1372,1432,909,935]" name="India" pageId="4" pageNumber="55">India</collectingCountry>
|
||||
, but has a consistently different pygidial surface and anterior pronotal margin (see detailed diagnostic characters separating these two species in
|
||||
<tableCitation id="0330CAEE2564FFC8FE51FC132C24F2D7" box="[503,585,981,1007]" captionStart="TABLE 1" captionStartId="4.[151,239,1054,1078]" captionTargetBox="[166,1420,1126,1518]" captionText="TABLE 1. Diagnostic characters separating the new species from Copris (Paracopris) punctulatus sensu lato (see also Figs. 2A–F)." httpUri="http://table.plazi.org/id/1ACDAFDD2564FFC8FF31FBD82F72F563" pageId="4" pageNumber="55" tableUuid="1ACDAFDD2564FFC8FF31FBD82F72F563">Table 1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="1ACDAFDD2564FFC8FF31FBD82F72F563" ID-Table-UUID="1ACDAFDD2564FFC8FF31FBD82F72F563" httpUri="http://table.plazi.org/id/1ACDAFDD2564FFC8FF31FBD82F72F563" pageId="4" pageNumber="55" targetBox="[166,1420,1126,1518]" targetIsTable="true" targetPageId="4">
|
||||
<paragraph id="4E0DFF552564FFC8FF31FBD82F72F563" blockId="4.[151,1436,1054,1115]" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FF31FBD82F66F50E" bold="true" box="[151,267,1054,1078]" pageId="4" pageNumber="55">TABLE 1.</emphasis>
|
||||
Diagnostic characters separating the new species from
|
||||
<taxonomicName id="89B284D62564FFC8FCC3FBD92AAEF50F" authorityName="Wiedemann" authorityYear="1823" box="[869,1219,1055,1079]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FCC3FBD92DC3F50E" box="[869,942,1055,1078]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FC19FBD92A5AF50E" box="[959,1079,1055,1078]" italics="true" pageId="4" pageNumber="55">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FBE1FBD92AAEF50F" box="[1095,1219,1055,1079]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FB6AFBD92B5AF50F" box="[1228,1335,1055,1079]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
|
||||
(see also Figs. 2A–F).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4E0DFF552564FFC8FF00FBA02A53F4D6" pageId="4" pageNumber="55">
|
||||
<table id="3CB20DF525640033FF00FBA02BE1F4D6" box="[166,1420,1126,1518]" gridcols="3" gridrows="12" pageId="4" pageNumber="55">
|
||||
<tr id="F082FD1725640033FF00FBA02BE1F547" box="[166,1420,1126,1151]" gridrow="0" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FBA02C43F547" box="[166,558,1126,1151]" gridcol="0" gridrow="0" pageId="4" pageNumber="55">Characters</th>
|
||||
<th id="B353946B25640033FDE9FBA02DAFF547" box="[591,962,1126,1151]" gridcol="1" gridrow="0" pageId="4" pageNumber="55">
|
||||
<taxonomicName id="89B284D62564FFC8FDE9FBA12D09F547" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[591,868,1126,1151]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FDE9FBA12CF9F547" bold="true" box="[591,660,1127,1151]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FD04FBA12CD8F546" bold="true" box="[674,693,1127,1150]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FD62FBA12D5FF547" bold="true" box="[708,818,1127,1151]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FC9EFBA12D09F547" box="[824,868,1127,1151]" pageId="4" pageNumber="55" rank="species">new</taxonomicNameLabel>
|
||||
</taxonomicName>
|
||||
species
|
||||
</th>
|
||||
<th id="B353946B25640033FC42FBA02BE1F547" box="[996,1420,1126,1151]" gridcol="2" gridrow="0" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FC42FBA12B75F547" bold="true" box="[996,1304,1126,1151]" pageId="4" pageNumber="55">
|
||||
<taxonomicName id="89B284D62564FFC8FC42FBA12AC5F547" authorityName="Wiedemann" authorityYear="1823" box="[996,1192,1126,1151]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FC42FBA12D96F547" bold="true" box="[996,1019,1127,1151]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FBACFBA12A70F546" bold="true" box="[1034,1053,1127,1150]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FB8AFBA02AC5F547" bold="true" box="[1068,1192,1126,1151]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FB08FBA12B75F547" box="[1198,1304,1127,1151]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FB402BE1F5A7" box="[166,1420,1158,1183]" gridrow="1" pageId="4" pageNumber="55" rowspan-0="1">
|
||||
<td id="B353946B25640033FDE9FB402DAFF5A7" box="[591,962,1158,1183]" gridcol="1" gridrow="1" pageId="4" pageNumber="55">
|
||||
(
|
||||
<specimenCount id="58B434DC2564FFC8FDF0FB402C88F5A7" box="[598,741,1158,1183]" pageId="4" pageNumber="55" type="generic">12 specimens</specimenCount>
|
||||
examined)
|
||||
</td>
|
||||
<td id="B353946B25640033FC42FB402BE1F5A7" box="[996,1420,1158,1183]" gridcol="2" gridrow="1" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FC42FB402A88F5A7" bold="true" box="[996,1253,1158,1183]" pageId="4" pageNumber="55">
|
||||
(
|
||||
<specimenCount id="58B434DC2564FFC8FC4AFB402A03F5A7" box="[1004,1134,1158,1183]" pageId="4" pageNumber="55" type="generic">9 specimens</specimenCount>
|
||||
examined)
|
||||
</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FB612BE1F5F8" box="[166,1420,1191,1216]" gridrow="2" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FB612C43F5F8" box="[166,558,1191,1216]" gridcol="0" gridrow="2" pageId="4" pageNumber="55">Coloration</th>
|
||||
<td id="B353946B25640033FDE9FB612DAFF5F8" box="[591,962,1191,1216]" gridcol="1" gridrow="2" pageId="4" pageNumber="55">Chocolate brown</td>
|
||||
<td id="B353946B25640033FC42FB612BE1F5F8" box="[996,1420,1191,1216]" gridcol="2" gridrow="2" pageId="4" pageNumber="55">Reddish brown</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FB0A2BE1F5DD" box="[166,1420,1228,1253]" gridrow="3" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FB0A2C43F5DD" box="[166,558,1228,1253]" gridcol="0" gridrow="3" pageId="4" pageNumber="55">Anterior margin of pronotum in males</th>
|
||||
<td id="B353946B25640033FDE9FB0A2DAFF5DD" box="[591,962,1228,1253]" gridcol="1" gridrow="3" pageId="4" pageNumber="55">Slightly forwards curved in middle</td>
|
||||
<td id="B353946B25640033FC42FB0A2BE1F5DD" box="[996,1420,1228,1253]" gridcol="2" gridrow="3" pageId="4" pageNumber="55">Slightly backwards curved in middle</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FB2A2BE1F43D" box="[166,1420,1260,1285]" gridrow="4" pageId="4" pageNumber="55" rowspan-0="1">
|
||||
<td id="B353946B25640033FDE9FB2A2DAFF43D" box="[591,962,1260,1285]" gridcol="1" gridrow="4" pageId="4" pageNumber="55">
|
||||
(
|
||||
<figureCitation id="D689E3D02564FFC8FDF0FB2A2CCBF43D" box="[598,678,1260,1285]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2A</figureCitation>
|
||||
)
|
||||
</td>
|
||||
<td id="B353946B25640033FC42FB2A2BE1F43D" box="[996,1420,1260,1285]" gridcol="2" gridrow="4" pageId="4" pageNumber="55">
|
||||
(
|
||||
<figureCitation id="D689E3D02564FFC8FC4AFB2A2A57F43D" box="[1004,1082,1260,1285]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2B</figureCitation>
|
||||
)
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FACB2BE1F41E" box="[166,1420,1293,1318]" gridrow="5" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FACB2C43F41E" box="[166,558,1293,1318]" gridcol="0" gridrow="5" pageId="4" pageNumber="55">Anterior margin of pronotum in</th>
|
||||
<td id="B353946B25640033FDE9FACB2DAFF41E" box="[591,962,1293,1318]" gridcol="1" gridrow="5" pageId="4" pageNumber="55">Strongly forwards curved in middle</td>
|
||||
<td id="B353946B25640033FC42FACB2BE1F41E" box="[996,1420,1293,1318]" gridcol="2" gridrow="5" pageId="4" pageNumber="55">Slightly forwards curved in middle (Fig.</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FAEB2BE1F47E" box="[166,1420,1325,1350]" gridrow="6" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FAEB2C43F47E" box="[166,558,1325,1350]" gridcol="0" gridrow="6" pageId="4" pageNumber="55">females</th>
|
||||
<td id="B353946B25640033FDE9FAEB2DAFF47E" box="[591,962,1325,1350]" gridcol="1" gridrow="6" pageId="4" pageNumber="55">
|
||||
(
|
||||
<figureCitation id="D689E3D02564FFC8FDF0FAEB2CCEF47E" box="[598,675,1325,1350]" captionStart="FIGURES 1" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[215,1374,179,1936]" captionTargetId="figure@5.[206,1381,166,1945]" captionTargetPageId="5" captionText="FIGURES 1A–D. Holotype (A, B) and female paratype (C, D) of Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species. A, C, habitus, dorsal view; B, D, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3586828" httpUri="https://zenodo.org/record/3586828/files/figure.png" pageId="4" pageNumber="55">Fig. 1C</figureCitation>
|
||||
)
|
||||
</td>
|
||||
<td id="B353946B25640033FC42FAEB2BE1F47E" box="[996,1420,1325,1350]" gridcol="2" gridrow="6" pageId="4" pageNumber="55">4C)</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FA892BE1F450" box="[166,1420,1359,1384]" gridrow="7" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FA892C43F450" box="[166,558,1359,1384]" gridcol="0" gridrow="7" pageId="4" pageNumber="55">Punctures on pygidium surface</th>
|
||||
<td id="B353946B25640033FDE9FA892DAFF450" box="[591,962,1359,1384]" gridcol="1" gridrow="7" pageId="4" pageNumber="55">
|
||||
Clearly separated (
|
||||
<figureCitation id="D689E3D02564FFC8FCABFA892D36F45F" box="[781,859,1359,1384]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2C</figureCitation>
|
||||
)
|
||||
</td>
|
||||
<td id="B353946B25640033FC42FA892BE1F450" box="[996,1420,1359,1384]" gridcol="2" gridrow="7" pageId="4" pageNumber="55">
|
||||
Contiguous (
|
||||
<figureCitation id="D689E3D02564FFC8FBC1FA892ADBF45F" box="[1127,1206,1359,1384]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2D</figureCitation>
|
||||
)
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FAB22BE1F4B5" box="[166,1420,1396,1421]" gridrow="8" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FAB22C43F4B5" box="[166,558,1396,1421]" gridcol="0" gridrow="8" pageId="4" pageNumber="55">Phallobase width relative to parameres</th>
|
||||
<td id="B353946B25640033FDE9FAB22DAFF4B5" box="[591,962,1396,1421]" gridcol="1" gridrow="8" pageId="4" pageNumber="55">PhaW slightly wider than PaW</td>
|
||||
<td id="B353946B25640033FC42FAB22BE1F4B5" box="[996,1420,1396,1421]" gridcol="2" gridrow="8" pageId="4" pageNumber="55">
|
||||
PhaW greatly wider than PaW (
|
||||
<figureCitation id="D689E3D02564FFC8FA83FAB22B1CF4B5" box="[1317,1393,1396,1421]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2F</figureCitation>
|
||||
)
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FA522BE1F495" box="[166,1420,1428,1453]" gridrow="9" pageId="4" pageNumber="55" rowspan-2="1">
|
||||
<th id="B353946B25640033FF00FA522C43F495" box="[166,558,1428,1453]" gridcol="0" gridrow="9" pageId="4" pageNumber="55">width</th>
|
||||
<td id="B353946B25640033FDE9FA522DAFF495" box="[591,962,1428,1453]" gridcol="1" gridrow="9" pageId="4" pageNumber="55">
|
||||
(
|
||||
<figureCitation id="D689E3D02564FFC8FDF0FA522CCEF495" box="[598,675,1428,1453]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2E</figureCitation>
|
||||
)
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FA732BE1F4F6" box="[166,1420,1461,1486]" gridrow="10" pageId="4" pageNumber="55">
|
||||
<th id="B353946B25640033FF00FA732C43F4F6" box="[166,558,1461,1486]" gridcol="0" gridrow="10" pageId="4" pageNumber="55">Shape of parameres</th>
|
||||
<td id="B353946B25640033FDE9FA732DAFF4F6" box="[591,962,1461,1486]" gridcol="1" gridrow="10" pageId="4" pageNumber="55">Slightly curved downwards in apical</td>
|
||||
<td id="B353946B25640033FC42FA732BE1F4F6" box="[996,1420,1461,1486]" gridcol="2" gridrow="10" pageId="4" pageNumber="55">Strongly curved downwards in apical part</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725640033FF00FA132BE1F4D6" box="[166,1420,1493,1518]" gridrow="11" pageId="4" pageNumber="55" rowspan-0="1">
|
||||
<td id="B353946B25640033FDE9FA132DAFF4D6" box="[591,962,1493,1518]" gridcol="1" gridrow="11" pageId="4" pageNumber="55">
|
||||
part (
|
||||
<figureCitation id="D689E3D02564FFC8FD22FA132CBCF4D6" box="[644,721,1493,1518]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2E</figureCitation>
|
||||
)
|
||||
</td>
|
||||
<td id="B353946B25640033FC42FA132BE1F4D6" box="[996,1420,1493,1518]" gridcol="2" gridrow="11" pageId="4" pageNumber="55">
|
||||
(
|
||||
<figureCitation id="D689E3D02564FFC8FC4AFA132A5AF4D6" box="[1004,1079,1493,1518]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2F</figureCitation>
|
||||
)
|
||||
</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552564FFC8FF61F9DC2C82F640" blockId="4.[151,1437,1562,1949]" pageId="4" pageNumber="55">
|
||||
The males of both
|
||||
<taxonomicName id="89B284D62564FFC8FE3FF9DC2CABF70C" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[409,710,1562,1588]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FE3FF9DC2F88F70C" box="[409,485,1562,1588]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FE53F9DD2C67F70C" box="[501,522,1563,1588]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FDBCF9DD2CFDF70C" box="[538,656,1563,1588]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FD31F9DC2CABF70C" box="[663,710,1562,1588]" pageId="4" pageNumber="55" rank="species">new</taxonomicNameLabel>
|
||||
</taxonomicName>
|
||||
species and
|
||||
<taxonomicName id="89B284D62564FFC8FCF5F9DC2A48F70C" authorityName="Wiedemann" authorityYear="1823" box="[851,1061,1562,1588]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FCF5F9DC2D00F70C" box="[851,877,1562,1588]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FCDBF9DD2DFEF70C" box="[893,915,1563,1588]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FC04F9DC2A48F70C" box="[930,1061,1562,1588]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FB8DF9DC2AF6F70C" box="[1067,1179,1562,1588]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
|
||||
possess a clypeal apex strongly upwards protruded and shallowly and widely emarginated, making these two species distinct from most Asian members of the subgenus
|
||||
<taxonomicName id="89B284D62564FFC8FDA1F9A52CE8F744" authorityName="Balthasar" authorityYear="1939" box="[519,645,1635,1660]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FDA1F9A52CE8F744" box="[519,645,1635,1660]" italics="true" pageId="4" pageNumber="55">Paracopris</emphasis>
|
||||
</taxonomicName>
|
||||
(for details see
|
||||
<tableCitation id="0330CAEE2564FFC8FC9DF9A42DFDF744" box="[827,912,1634,1660]" captionStart="TABLE 2" captionStartId="6.[151,239,154,178]" captionTargetBox="[166,1421,261,1584]" captionText="TABLE 2. Diagnostic characters of clypeus separating the new species from the other Copris (Paracopris) species (based on the specimens examined by us in MNHN and NMPC, and data published by Wiedemann 1823; Arrow 1931; Balthasar 1939, 1963; Ochi & Kon 1996; Hanboonsong et al. 2003; Ochi et al. 2019a,b)" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="4" pageNumber="55" tableUuid="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2">Table 2</tableCitation>
|
||||
), except for
|
||||
<taxonomicName id="89B284D62564FFC8FBB9F9A42A9BF744" authorityName="Lansberge" authorityYear="1886" box="[1055,1270,1634,1660]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pedarioides" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FBB9F9A42A54F744" box="[1055,1081,1634,1660]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FBECF9A52A32F744" box="[1098,1119,1635,1660]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FBC9F9A42A9BF744" box="[1135,1270,1634,1660]" italics="true" pageId="4" pageNumber="55">pedarioides</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7CC623472564FFC8FB50F9A42A97F744" box="[1270,1274,1634,1660]" italics="true" pageId="4" pageNumber="55">,</emphasis>
|
||||
<taxonomicName id="89B284D62564FFC8FAA4F9A42BF0F744" authorityName="Arrow" authorityYear="1931" box="[1282,1437,1634,1660]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="surdus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FAA4F9A42B71F744" box="[1282,1308,1634,1660]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FA8BF9A52B2FF744" box="[1325,1346,1635,1660]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FAF5F9A42BF0F744" box="[1363,1437,1634,1660]" italics="true" pageId="4" pageNumber="55">surdus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="89B284D62564FFC8FF60F9402FE7F798" authorityName="Felsche" authorityYear="1910" box="[198,394,1670,1696]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="cariniceps" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FF60F9402E8DF798" box="[198,224,1670,1696]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FF56F9412F68F798" box="[240,261,1671,1696]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FEB3F9412FE7F798" box="[277,394,1671,1696]" italics="true" pageId="4" pageNumber="55">cariniceps</emphasis>
|
||||
</taxonomicName>
|
||||
. The three latter can be clearly distinguished from the two former due to having very densely to contiguously punctate elytra and very densely and deeply punctate genae (
|
||||
<figureCitation id="D689E3D02564FFC8FBA2F96C2A3DF7FC" box="[1028,1104,1706,1732]" captionStart="FIGURES 3" captionStartId="9.[151,264,1806,1831]" captionTargetBox="[163,1427,949,1787]" captionTargetId="figure@9.[151,1435,949,1787]" captionTargetPageId="9" captionText="FIGURES 3A–C. Surface of elytral intervals of Copris (Paracopris) species. A, Copris (P.) javanensis Bui, Ziegler & Bonkowski, new species; B, Copris (P.) punctulatus punctulatus; C, Copris (P.) cariniceps." figureDoi="http://doi.org/10.5281/zenodo.3586832" httpUri="https://zenodo.org/record/3586832/files/figure.png" pageId="4" pageNumber="55">Figs. 3</figureCitation>
|
||||
A–C). In addition, characters on the male cephalic horn clearly differentiate
|
||||
<taxonomicName id="89B284D62564FFC8FD0AF9082DECF7D0" authorityName="Lansberge" authorityYear="1886" box="[684,897,1742,1768]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pedarioides" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FD0AF9082CABF7D0" box="[684,710,1742,1768]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FD71F9092C81F7D0" box="[727,748,1743,1768]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FD5BF9082DECF7D0" box="[765,897,1742,1768]" italics="true" pageId="4" pageNumber="55">pedarioides</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="89B284D62564FFC8FC1DF9082A3BF7D0" authorityName="Arrow" authorityYear="1931" box="[955,1110,1742,1768]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="surdus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FC1DF9082DB8F7D0" box="[955,981,1742,1768]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FC40F9092D91F7D0" box="[998,1020,1743,1768]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FBAAF9082A3BF7D0" box="[1036,1110,1742,1768]" italics="true" pageId="4" pageNumber="55">surdus</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<taxonomicName id="89B284D62564FFC8FB3BF9082B08F7D0" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="Bui & Ziegler & Bonkowski" authorityYear="2019" box="[1181,1381,1742,1768]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FB3BF9082ADAF7D0" box="[1181,1207,1742,1768]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FB6FF9092AB3F7D0" box="[1225,1246,1743,1768]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FB49F9092B08F7D0" box="[1263,1381,1743,1768]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FAC8F9082E85F634" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="89B284D62564FFC8FE86F9342F9EF634" authorityName="Wiedemann" authorityYear="1823" box="[288,499,1778,1804]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FE86F9342F57F634" box="[288,314,1778,1804]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FEEDF9352F0DF634" box="[331,352,1779,1804]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FED6F9342F9EF634" box="[368,499,1778,1804]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FE5DF9342C03F634" box="[507,622,1778,1804]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
|
||||
. In the two former, the male cephalic horn is relatively long, whereas in the two latter, the male cephalic horn is small. The males of
|
||||
<taxonomicName id="89B284D62564FFC8FC96F8D02D9EF608" authorityName="Felsche" authorityYear="1910" box="[816,1011,1814,1840]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="cariniceps" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FC96F8D02D27F608" box="[816,842,1814,1840]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FCFCF8D12D02F608" box="[858,879,1815,1840]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FCD8F8D12D9EF608" box="[894,1011,1815,1840]" italics="true" pageId="4" pageNumber="55">cariniceps</emphasis>
|
||||
</taxonomicName>
|
||||
bear a clypeus strongly longitudinally carinate in middle, distinctly separating this species from
|
||||
<taxonomicName id="89B284D62564FFC8FC96F8FC2D94F66C" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="Bui & Ziegler & Bonkowski" authorityYear="2019" box="[816,1017,1850,1876]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FC96F8FC2D24F66C" box="[816,841,1850,1876]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FCFAF8FD2D1CF66C" box="[860,881,1851,1876]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FC25F8FD2D94F66C" box="[899,1017,1851,1876]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FBA4F8FC2AE6F66C" box="[1026,1163,1850,1876]" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="89B284D62564FFC8FB60F8FC2BF0F66C" authorityName="Wiedemann" authorityYear="1823" box="[1222,1437,1850,1876]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472564FFC8FB60F8FC2A8DF66C" box="[1222,1248,1850,1876]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472564FFC8FB54F8FD2B65F66C" box="[1266,1288,1851,1876]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472564FFC8FABCF8FC2BF0F66C" box="[1306,1437,1850,1876]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="67F59E3C2564FFC8FF31F8982F6BF640" box="[151,262,1886,1912]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
|
||||
whose clypeus has no longitudinal carinae.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2564FFCAFF61F8442A5CF687" lastPageId="6" lastPageNumber="57" pageId="4" pageNumber="55" type="etymology">
|
||||
<paragraph id="4E0DFF552564FFC8FF61F8442B51F6A5" blockId="4.[151,1437,1562,1949]" box="[199,1340,1922,1949]" pageId="4" pageNumber="55">
|
||||
<emphasis id="7CC623472564FFC8FF61F8442F23F6A4" bold="true" box="[199,334,1922,1948]" pageId="4" pageNumber="55">Etymology.</emphasis>
|
||||
The specific epithet “javanensis” refers to the name of the
|
||||
<typeStatus id="910941F72564FFC8FC42F8442A79F6A4" box="[996,1044,1922,1948]" pageId="4" pageNumber="55">type</typeStatus>
|
||||
locality,
|
||||
<collectingRegion id="8C7631B72564FFC8FBDAF8442AC3F6A4" box="[1148,1198,1922,1948]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="55">Java</collectingRegion>
|
||||
(
|
||||
<collectingCountry id="36A5BFC52564FFC8FB1AF8442B5DF6A5" box="[1212,1328,1922,1949]" name="Indonesia" pageId="4" pageNumber="55">Indonesia</collectingCountry>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="1ACDAFDD2565FFC9FF31F86B2D2AF6D2" ID-DOI="http://doi.org/10.5281/zenodo.3586828" ID-Zenodo-Dep="3586828" httpUri="https://zenodo.org/record/3586828/files/figure.png" pageId="5" pageNumber="56" startId="5.[151,264,1965,1990]" targetBox="[215,1374,179,1936]" targetPageId="5">
|
||||
<paragraph id="4E0DFF552565FFC9FF31F86B2D2AF6D2" blockId="5.[151,1437,1965,2026]" pageId="5" pageNumber="56">
|
||||
<emphasis id="7CC623472565FFC9FF31F86B2F3EF6FE" bold="true" box="[151,339,1965,1990]" pageId="5" pageNumber="56">FIGURES 1A–D.</emphasis>
|
||||
Holotype (A, B) and female paratype (C, D) of
|
||||
<taxonomicName id="89B284D62565FFC9FC99F8692BF5F6FE" authority="Bui, Ziegler & Bonkowski" authorityName="Bui, Ziegler & Bonkowski" authorityYear="2019" box="[831,1432,1965,1990]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
|
||||
<emphasis id="7CC623472565FFC9FC99F8692DE9F6FE" box="[831,900,1967,1990]" italics="true" pageId="5" pageNumber="56">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472565FFC9FC35F8692A68F6FE" box="[915,1029,1967,1990]" italics="true" pageId="5" pageNumber="56">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472565FFC9FBB3F8692AEDF6FE" box="[1045,1152,1967,1990]" italics="true" pageId="5" pageNumber="56">javanensis</emphasis>
|
||||
Bui, Ziegler & Bonkowski
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="67F59E3C2565FFC9FF31F8152F7CF6D2" box="[151,273,2001,2026]" pageId="5" pageNumber="56" rank="species">new species</taxonomicNameLabel>
|
||||
. A, C, habitus, dorsal view; B, D, habitus, ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" ID-Table-UUID="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="6" pageNumber="57" targetBox="[166,1421,261,1584]" targetIsTable="true" targetPageId="6">
|
||||
<paragraph id="4E0DFF552566FFCAFF31FF5C2DB7F1C2" blockId="6.[151,1436,154,250]" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF31FF5C2F64F18A" bold="true" box="[151,265,154,178]" pageId="6" pageNumber="57">
|
||||
<tableCitation id="0330CAEE2566FFCAFF31FF5C2F69F18A" box="[151,260,154,178]" captionStart="TABLE 2" captionStartId="6.[151,239,154,178]" captionTargetBox="[166,1421,261,1584]" captionText="TABLE 2. Diagnostic characters of clypeus separating the new species from the other Copris (Paracopris) species (based on the specimens examined by us in MNHN and NMPC, and data published by Wiedemann 1823; Arrow 1931; Balthasar 1939, 1963; Ochi & Kon 1996; Hanboonsong et al. 2003; Ochi et al. 2019a,b)" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="6" pageNumber="57" tableUuid="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2">TABLE 2</tableCitation>
|
||||
.
|
||||
</emphasis>
|
||||
Diagnostic characters of clypeus separating the new species from the other
|
||||
<taxonomicName id="89B284D62566FFCAFB80FF5C2A92F18A" authority="" authorityName="Balthasar" authorityYear="1939" box="[1062,1279,154,178]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFB80FF5C2A02F189" box="[1062,1135,154,177]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFBD8FF5C2A9BF189" box="[1150,1270,154,177]" italics="true" pageId="6" pageNumber="57">Paracopris</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species (based on the specimens examined by us in MNHN and NMPC, and data published by
|
||||
<bibRefCitation id="2A2382A42566FFCAFC44FF782ACDF1EE" author="Wiedemann, C. R. W." box="[994,1184,190,214]" pageId="6" pageNumber="57" pagination="1 - 164" refId="ref9330" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoolo- gisches Magazin, 2, 1 - 164." type="journal article" year="1823">Wiedemann 1823</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42566FFCAFB0FFF782B40F1EE" author="Arrow, G. J." box="[1193,1325,190,214]" pageId="6" pageNumber="57" refId="ref7267" refString="Arrow, G. J. (1931) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42566FFCAFA91FF782BF1F1EE" author="Balthasar, V." box="[1335,1436,190,214]" pageId="6" pageNumber="57" pagination="1 - 36" refId="ref7382" refString="Balthasar, V. (1939) Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico delle R. Universita di Firenze. Redia, 25, 1 - 36." type="journal article" year="1939">Balthasar</bibRefCitation>
|
||||
1939,
|
||||
<bibRefCitation id="2A2382A42566FFCAFF7CFF242F7FF1C2" author="Balthasar, V." box="[218,274,226,250]" pageId="6" pageNumber="57" refId="ref7444" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region. Coleo- ptera: Lamellicornia. Band 1. Allgemeiner Teil, Systematischer Teil: 1. Scarabaeinae, 2. Coprinae (Pinotini, Coprini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 391 pp." type="book" year="1963">1963</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42566FFCAFEBBFF242F8CF1C2" author="Ochi, T. & Kon, M." box="[285,481,226,250]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi & Kon 1996</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42566FFCAFE4DFF242C96F1C2" author="Hanboonsong, Y. & Masumoto, K. & Ochi, T." box="[491,763,226,250]" pageId="6" pageNumber="57" pagination="103 - 124" refId="ref7902" refString="Hanboonsong, Y., Masumoto, K. & Ochi, T. (2003) Dung beetles (Coleoptera, Scarabaeidae) of Thailand. Part 5. Genera Copris and Microcopris (Coprini). Elytra, 31, 103 - 124." type="journal article" year="2003">
|
||||
Hanboonsong
|
||||
<emphasis id="7CC623472566FFCAFD20FF252CD8F1C2" box="[646,693,226,250]" italics="true" pageId="6" pageNumber="57">et al</emphasis>
|
||||
. 2003
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42566FFCAFCA3FF242DACF1C2" author="Ochi, T. & Kon, M. & Kawahara, M." box="[773,961,226,250]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. & Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
|
||||
Ochi
|
||||
<emphasis id="7CC623472566FFCAFCE6FF252D02F1C2" box="[832,879,226,250]" italics="true" pageId="6" pageNumber="57">et al</emphasis>
|
||||
. 2019a
|
||||
</bibRefCitation>
|
||||
,b)
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4E0DFF552566FFCAFF00FEC32A1CF717" pageId="6" pageNumber="57">
|
||||
<table id="3CB20DF525660033FF00FEC32BE0F708" box="[166,1421,261,1584]" gridcols="2" gridrows="26" pageId="6" pageNumber="57">
|
||||
<tr id="F082FD1725660033FF00FEC32BE0F026" box="[166,1421,261,286]" gridrow="0" pageId="6" pageNumber="57">
|
||||
<th id="B353946B25660033FF00FEC32FD1F026" box="[166,444,261,286]" gridcol="0" gridrow="0" pageId="6" pageNumber="57">Species</th>
|
||||
<th id="B353946B25660033FDAEFEC32BE0F026" box="[520,1421,261,286]" gridcol="1" gridrow="0" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFDAEFEC32D53F026" bold="true" box="[520,830,261,286]" pageId="6" pageNumber="57">Character of clypeus (males)</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FEED2BE0F07C" box="[166,1421,299,324]" gridrow="1" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FEED2FD1F07C" box="[166,444,299,324]" gridcol="0" gridrow="1" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FEEA2FD1F07B" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[166,444,300,324]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FEEA2E86F07B" box="[166,235,300,323]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFEA6FEEB2F7EF07C" box="[256,275,301,324]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFE87FEEB2FE1F07C" box="[289,396,301,324]" italics="true" pageId="6" pageNumber="57">javanensis</emphasis>
|
||||
new
|
||||
</taxonomicName>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFEED2BE0F07C" box="[520,1421,299,324]" gridcol="1" gridrow="1" pageId="6" pageNumber="57">Clypeal apex distinctly upwards protruded, and widely and shallowly emarginated.</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FE892BE0F050" box="[166,1421,335,360]" gridrow="2" pageId="6" pageNumber="57" rowspan-1="1">
|
||||
<td id="B353946B25660033FF00FE892FD1F050" box="[166,444,335,360]" gridcol="0" gridrow="2" pageId="6" pageNumber="57">species</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FEB62BE0F0B1" box="[166,1421,368,393]" gridrow="3" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FEB62FD1F0B1" box="[166,444,368,393]" gridcol="0" gridrow="3" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FEB72ED0F0B0" box="[166,189,369,392]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFEB72EB2F0B0" box="[204,223,369,392]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFEB72FFDF0B1" box="[237,400,369,393]" italics="true" pageId="6" pageNumber="57">anomiopseoides</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFEB62BE0F0B1" box="[520,1421,368,393]" gridcol="1" gridrow="3" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFEB62C3AF0B1" authority="Clypeal" authorityName="Clypeal" box="[520,599,368,393]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="anomiopseoides" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex deeply and almost semicircularly emarginated; middle part of
|
||||
<tr id="F082FD1725660033FF00FE522BE0F0F6" box="[166,1421,404,462]" gridrow="4" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FE522FD1F0F6" box="[166,444,404,462]" gridcol="0" gridrow="4" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFAAFFEB62ED0F0F5" italics="true" pageId="6" pageNumber="57">emarginaC.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFE702EB2F0F5" box="[204,223,438,461]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFE732F39F0F5" box="[237,340,437,461]" italics="true" pageId="6" pageNumber="57">andrewesi</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFE522BE0F0F6" box="[520,1421,404,462]" gridcol="1" gridrow="4" pageId="6" pageNumber="57">
|
||||
tion strongly protruded upwards, and bifurcated.
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFE732C3AF0F6" authority="Clypeal" authorityName="Clypeal" box="[520,599,437,462]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="andrewesi" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex strongly reflexed and emarginated; middle part of emargination protruded,
|
||||
</td>
|
||||
</tr>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FE1F2BE0F32A" box="[166,1421,473,530]" gridrow="5" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FE1F2FD1F32A" box="[166,444,473,530]" gridcol="0" gridrow="5" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FE3C2F34F32A" authorityName="Ochi, Kon & Pham" authorityYear="2019" box="[166,345,506,530]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bartolozzii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FE3C2ED0F329" box="[166,189,506,529]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFE3D2EB2F32A" box="[204,223,507,530]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFE3C2F34F32A" box="[237,345,506,530]" italics="true" pageId="6" pageNumber="57">bartolozzii</emphasis>
|
||||
</taxonomicName>
|
||||
(**)
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFE1F2BE0F32A" box="[520,1421,473,530]" gridcol="1" gridrow="5" pageId="6" pageNumber="57">
|
||||
forming a rounded lobe. “Clypeal margin rather broadly but shallowly emarginate at apex, each side of
|
||||
<tr id="F082FD1725660033FF00FDDB2BE0F30E" box="[166,1421,541,566]" gridrow="6" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FDDB2BE0F30E" box="[166,1421,541,566]" colspan="2" colspanRight="1" gridcol="0" gridrow="6" pageId="6" pageNumber="57">
|
||||
emargination weakly produced forward, forming small, rounded and reflexed tooth” (
|
||||
<bibRefCitation id="2A2382A42566FFCAFAA8FDD82B14F30E" author="Ochi, T. & Kon, M. & Pham, H. T." box="[1294,1401,542,566]" pageId="6" pageNumber="57" pagination="435 - 446" refId="ref8783" refString="Ochi, T., Kon, M. & Pham, H. T. (2019 b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435 - 446." type="journal article" year="2019">
|
||||
<emphasis id="7CC623472566FFCAFAA8FDD82B14F30E" box="[1294,1401,542,566]" italics="true" pageId="6" pageNumber="57">Ochi et al.</emphasis>
|
||||
</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FD872BE0F343" box="[166,1421,577,635]" gridrow="7" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FD872FD1F343" box="[166,444,577,635]" gridcol="0" gridrow="7" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FDA52F16F343" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[166,379,611,635]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chinmontanus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FDA52ED0F342" box="[166,189,611,634]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFDA52EB2F342" box="[204,223,611,634]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFDA52F16F343" box="[237,379,611,635]" italics="true" pageId="6" pageNumber="57">chinmontanus</emphasis>
|
||||
</taxonomicName>
|
||||
(**)
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFD872BE0F343" box="[520,1421,577,635]" gridcol="1" gridrow="7" pageId="6" pageNumber="57">2019b) “Clypeus with anterior margin suddenly produced upward as a clypeal horn at middle,</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FD402BE0F3A7" box="[166,1421,646,671]" gridrow="8" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FD402BE0F3A7" box="[166,1421,646,671]" colspan="2" colspanRight="1" gridcol="0" gridrow="8" pageId="6" pageNumber="57">
|
||||
which is curved backward near the pointed apex, ca.
|
||||
<quantity id="894A52B02566FFCAFB87FD412A03F3A6" box="[1057,1134,647,671]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="57" unit="mm" value="1.2">1.2 mm</quantity>
|
||||
in length in lateral view…”
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FD6C2BE0F3DC" box="[166,1421,682,740]" gridrow="9" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FD6C2FD1F3DC" box="[166,444,682,740]" gridcol="0" gridrow="9" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FD0A2ED0F3DB" box="[166,189,716,739]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFD0A2EB2F3DB" box="[204,223,716,739]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFD0A2FF8F3DB" box="[237,405,716,739]" italics="true" pageId="6" pageNumber="57">compressipennis</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFD6C2BE0F3DC" box="[520,1421,682,740]" gridcol="1" gridrow="9" pageId="6" pageNumber="57">
|
||||
(
|
||||
<bibRefCitation id="2A2382A42566FFCAFDA9FD6C2CD2F3FB" author="Ochi, T. & Kon, M. & Kawahara, M." box="[527,703,682,707]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. & Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
|
||||
Ochi
|
||||
<emphasis id="7CC623472566FFCAFDEEFD6D2C16F3FB" box="[584,635,683,707]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
|
||||
2019a
|
||||
</bibRefCitation>
|
||||
)
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFD0D2C3AF3DC" authority="Clypeal" authorityName="Clypeal" box="[520,599,715,740]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="compressipennis" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex widely emarginated, and not protruded.
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FD362BE0F231" box="[166,1421,752,777]" gridrow="10" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FD362FD1F231" box="[166,444,752,777]" gridcol="0" gridrow="10" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FD342ED0F231" box="[166,189,754,777]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFD342EB2F231" box="[204,223,754,777]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFD372F27F231" box="[237,330,753,777]" italics="true" pageId="6" pageNumber="57">cribratus</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFD362BE0F231" box="[520,1421,752,777]" gridcol="1" gridrow="10" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFD362C3AF231" authority="Clypeal" authorityName="Clypeal" box="[520,599,752,777]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="cribratus" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex distinctly emarginated, and not protruded.
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FCD02BE0F217" box="[166,1421,790,815]" gridrow="11" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FCD02FD1F217" box="[166,444,790,815]" gridcol="0" gridrow="11" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FCD12ED0F216" box="[166,189,791,814]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFCDE2EB2F217" box="[204,223,792,815]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFCD12F2EF217" box="[237,323,791,815]" italics="true" pageId="6" pageNumber="57">davisoni</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFCD02BE0F217" box="[520,1421,790,815]" gridcol="1" gridrow="11" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFCD02C3AF217" authority="Clypeal" authorityName="Clypeal" box="[520,599,790,815]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="davisoni" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex upwards protruded, forming one backwardly curved and pointed horn.
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FCFA2BE0F26D" box="[166,1421,828,853]" gridrow="12" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FCFA2FD1F26D" box="[166,444,828,853]" gridcol="0" gridrow="12" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FCFB2ED0F26C" box="[166,189,829,852]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFCFB2EB2F26C" box="[204,223,829,852]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFCFB2F5BF26C" box="[237,310,829,852]" italics="true" pageId="6" pageNumber="57">excisus</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFCFA2BE0F26D" box="[520,1421,828,853]" gridcol="1" gridrow="12" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFCFA2C3AF26D" authority="Clypeal" authorityName="Clypeal" box="[520,599,828,853]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="excisus" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex widely and deeply emarginated, forming a pair of slender lateral processes
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FCA62BE0F2A2" box="[166,1421,864,922]" gridrow="13" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FCA62FD1F2A2" box="[166,444,864,922]" gridcol="0" gridrow="13" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FC442ED0F2A1" box="[166,189,898,921]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFC442EB2F2A1" box="[204,223,898,921]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFC442F24F2A2" box="[237,329,898,922]" italics="true" pageId="6" pageNumber="57">furciceps</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFCA62BE0F2A2" box="[520,1421,864,922]" gridcol="1" gridrow="13" pageId="6" pageNumber="57">
|
||||
forwards directed.
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFC472C3AF2A2" authority="Clypeal" authorityName="Clypeal" box="[520,599,897,922]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="furciceps" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex deeply and widely emarginated, and with one backwards and pointed horn
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FC632BE0F2E6" box="[166,1421,933,990]" gridrow="14" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FC632FD1F2E6" box="[166,444,933,990]" gridcol="0" gridrow="14" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FC012ED0F2E6" box="[166,189,967,990]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFC012EB2F2E6" box="[204,223,967,990]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFC012F55F2E6" box="[237,312,967,990]" italics="true" pageId="6" pageNumber="57">imitans</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFC632BE0F2E6" box="[520,1421,933,990]" gridcol="1" gridrow="14" pageId="6" pageNumber="57">
|
||||
in middle.
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFC032C3AF2E6" authority="Clypeal" authorityName="Clypeal" box="[520,599,965,990]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="imitans" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex reflexed, and weakly emarginated; middle part of emargination upwards
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FC2F2BE0F51B" box="[166,1421,1001,1059]" gridrow="15" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FC2F2FD1F51B" box="[166,444,1001,1059]" gridcol="0" gridrow="15" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FBCD2F5BF51B" authorityName="Ochi & Kon" authorityYear="1996" box="[166,310,1035,1059]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FBCD2ED0F51A" box="[166,189,1035,1058]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFBCA2EB2F51B" box="[204,223,1036,1059]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFBCD2F5BF51B" box="[237,310,1035,1059]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
|
||||
</taxonomicName>
|
||||
(*)
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFC2F2BE0F51B" box="[520,1421,1001,1059]" gridcol="1" gridrow="15" pageId="6" pageNumber="57">protruded, forming one short erect process. Clypeal anterior margin distinctly bidentate in middle, and not protruded.</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FBF62BE0F571" box="[166,1421,1072,1097]" gridrow="16" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FBF62FD1F571" box="[166,444,1072,1097]" gridcol="0" gridrow="16" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FBF72F3BF571" authorityName="Ochi, Kon & Pham" authorityYear="2019" box="[166,342,1073,1097]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="montreuili" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FBF72ED0F570" box="[166,189,1073,1096]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFBF72EB2F570" box="[204,223,1073,1096]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFBF72F3BF571" box="[237,342,1073,1097]" italics="true" pageId="6" pageNumber="57">montreuili</emphasis>
|
||||
</taxonomicName>
|
||||
(**)
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFBF62BE0F571" box="[520,1421,1072,1097]" gridcol="1" gridrow="16" pageId="6" pageNumber="57">“Clypeus with antero-median portion noticeably broadly incised, posterior portion of the</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FB922BE0F555" box="[166,1421,1108,1133]" gridrow="17" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FB922BE0F555" box="[166,1421,1108,1133]" colspan="2" colspanRight="1" gridcol="0" gridrow="17" pageId="6" pageNumber="57">incision developed as a short straight prominence which is strongly inclined backward”</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FBBE2BE0F58A" box="[166,1421,1144,1202]" gridrow="18" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FBBE2FD1F58A" box="[166,444,1144,1202]" gridcol="0" gridrow="18" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FB5C2F5BF589" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[166,310,1177,1201]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="olivieri" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FB5C2ED0F589" box="[166,189,1178,1201]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFB5C2EB2F589" box="[204,223,1178,1201]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFB5F2F5BF589" box="[237,310,1177,1201]" italics="true" pageId="6" pageNumber="57">olivieri</emphasis>
|
||||
</taxonomicName>
|
||||
(**)
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFBBE2BE0F58A" box="[520,1421,1144,1202]" gridcol="1" gridrow="18" pageId="6" pageNumber="57">
|
||||
(
|
||||
<bibRefCitation id="2A2382A42566FFCAFDA9FBBE2CA8F5A9" author="Ochi, T. & Kon, M. & Pham, H. T." box="[527,709,1144,1169]" pageId="6" pageNumber="57" pagination="435 - 446" refId="ref8783" refString="Ochi, T., Kon, M. & Pham, H. T. (2019 b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435 - 446." type="journal article" year="2019">
|
||||
Ochi
|
||||
<emphasis id="7CC623472566FFCAFDEEFBBF2C16F5A9" box="[584,635,1145,1169]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
|
||||
2019b
|
||||
</bibRefCitation>
|
||||
). “Clypeus with a slender horn situated at middle just behind clypeal margin, which is ca.
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FB7B2BE0F5EE" box="[166,1421,1213,1238]" gridrow="19" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FB7B2BE0F5EE" box="[166,1421,1213,1238]" colspan="2" colspanRight="1" gridcol="0" gridrow="19" pageId="6" pageNumber="57">
|
||||
<quantity id="894A52B02566FFCAFDAEFB7B2C38F5ED" box="[520,597,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="57" unit="mm" value="1.7">1.7 mm</quantity>
|
||||
long in lateral view, slightly curved backward, another short point a little behind
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FB272BE0F5C2" box="[166,1421,1249,1274]" gridrow="20" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FB272BE0F5C2" box="[166,1421,1249,1274]" colspan="2" colspanRight="1" gridcol="0" gridrow="20" pageId="6" pageNumber="57">
|
||||
the cephalic horn is present, which is ca.
|
||||
<quantity id="894A52B02566FFCAFC0FFB272D9BF5C1" box="[937,1014,1249,1273]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="57" unit="mm" value="0.3">0.3 mm</quantity>
|
||||
in height and slightly inclined forward”
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FAC32BE0F406" box="[166,1421,1285,1342]" gridrow="21" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FAC32FD1F406" box="[166,444,1285,1342]" gridcol="0" gridrow="21" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFF00FAE02F3FF406" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[166,338,1318,1342]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="popaensis" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF00FAE02ED0F405" box="[166,189,1318,1341]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFAE12EB2F406" box="[204,223,1319,1342]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFAE12F3FF406" box="[237,338,1319,1342]" italics="true" pageId="6" pageNumber="57">popaensis</emphasis>
|
||||
</taxonomicName>
|
||||
(**)
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFAC32BE0F406" box="[520,1421,1285,1342]" gridcol="1" gridrow="21" pageId="6" pageNumber="57">
|
||||
(
|
||||
<bibRefCitation id="2A2382A42566FFCAFDA9FAC32CAEF425" author="Ochi, T. & Kon, M. & Kawahara, M." box="[527,707,1285,1310]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. & Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
|
||||
Ochi
|
||||
<emphasis id="7CC623472566FFCAFDEEFAC02C16F425" box="[584,635,1285,1309]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
|
||||
2019a
|
||||
</bibRefCitation>
|
||||
). “Clypeus also with another stout subtriangular horn which is situated at median portion
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FA8F2BE0F45A" box="[166,1421,1353,1378]" gridrow="22" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FA8F2BE0F45A" box="[166,1421,1353,1378]" colspan="2" colspanRight="1" gridcol="0" gridrow="22" pageId="6" pageNumber="57">
|
||||
of clypeus, ca.
|
||||
<quantity id="894A52B02566FFCAFD07FA8C2C83F459" box="[673,750,1354,1378]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="57" unit="mm" value="2.1">2.1 mm</quantity>
|
||||
in length in lateral view, flat on anterior face, strongly
|
||||
<tr id="F082FD1725660033FF00FAAB2BE0F49F" box="[166,1421,1389,1447]" gridrow="23" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FAAB2FD1F49F" box="[166,444,1389,1447]" gridcol="0" gridrow="23" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFABBFA8C2ED0F49E" italics="true" pageId="6" pageNumber="57">comC.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFA492EB2F49E" box="[204,223,1423,1446]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFA492F0BF49F" box="[237,358,1423,1447]" italics="true" pageId="6" pageNumber="57">punjabensis</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFAAB2BE0F49F" box="[520,1421,1389,1447]" gridcol="1" gridrow="23" pageId="6" pageNumber="57">
|
||||
pressed on both sides and bidentate at apex” (
|
||||
<bibRefCitation id="2A2382A42566FFCAFC73FAAB2AE4F4BE" author="Ochi, T. & Kon, M. & Kawahara, M." box="[981,1161,1389,1414]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. & Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
|
||||
Ochi
|
||||
<emphasis id="7CC623472566FFCAFBA9FAA92A2FF4BE" box="[1039,1090,1390,1414]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
|
||||
2019a
|
||||
</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFA482C3AF49F" authority="Clypeal" authorityName="Clypeal" box="[520,599,1422,1447]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="punjabensis" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex shallowly emarginated; middle part of emargination protruded, forming a
|
||||
</td>
|
||||
</tr>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FA742BE0F4D4" box="[166,1421,1458,1516]" gridrow="24" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FA742FD1F4D4" box="[166,444,1458,1516]" gridcol="0" gridrow="24" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00FA122ED0F4D3" box="[166,189,1492,1515]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AFA122EB2F4D3" box="[204,223,1492,1515]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BFA122F0DF4D3" box="[237,352,1492,1515]" italics="true" pageId="6" pageNumber="57">ramosiceps</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFA742BE0F4D4" box="[520,1421,1458,1516]" gridcol="1" gridrow="24" pageId="6" pageNumber="57">
|
||||
rounded tongue-like lobe.
|
||||
<taxonomicName id="89B284D62566FFCAFDAEFA152C3AF4D4" authority="Clypeal" authorityName="Clypeal" box="[520,599,1491,1516]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="ramosiceps" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex protruded, forming one upwards protruded horn which widely bifurcated;
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="F082FD1725660033FF00FA312BE0F708" box="[166,1421,1527,1584]" gridrow="25" pageId="6" pageNumber="57">
|
||||
<td id="B353946B25660033FF00FA312FD1F708" box="[166,444,1527,1584]" gridcol="0" gridrow="25" pageId="6" pageNumber="57">
|
||||
<emphasis id="7CC623472566FFCAFF00F9DE2ED0F717" box="[166,189,1560,1583]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF6AF9DF2EB2F708" box="[204,223,1561,1584]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF4BF9DF2F2CF708" box="[237,321,1561,1584]" italics="true" pageId="6" pageNumber="57">signatus</emphasis>
|
||||
</td>
|
||||
<td id="B353946B25660033FDAEFA312BE0F708" box="[520,1421,1527,1584]" gridcol="1" gridrow="25" pageId="6" pageNumber="57">
|
||||
two branches upwards and backwards curved.
|
||||
<taxonomicName id="89B284D62566FFCAFDAEF9D12C3AF708" authority="Clypeal" authorityName="Clypeal" box="[520,599,1559,1584]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="signatus" subGenus="Paracopris">Clypeal</taxonomicName>
|
||||
apex with two narrow and almost parallel processes.
|
||||
</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552566FFCAFF31F9F92A5CF687" blockId="6.[151,1437,1599,1983]" pageId="6" pageNumber="57">
|
||||
(*)
|
||||
<bibRefCitation id="2A2382A42566FFCAFF66F9F92FABF76F" author="Ochi, T. & Kon, M." box="[192,454,1599,1623]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi & Kon (1996: 25)</bibRefCitation>
|
||||
originally showed the clypeus of
|
||||
<taxonomicName id="89B284D62566FFCAFCE7F9F92A1CF76F" authorityName="Ochi & Kon" authorityYear="1996" box="[833,1137,1599,1623]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFCE7F9F92DE7F76E" box="[833,906,1599,1622]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFC3FF9F92A7CF76E" box="[921,1041,1599,1622]" italics="true" pageId="6" pageNumber="57">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFB82F9F92A1CF76F" box="[1060,1137,1599,1623]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
|
||||
</taxonomicName>
|
||||
with distinct emargination: “Clypeus produced as a reflexed point in the middle, which is medially notched as obtuse angle and distinctly bidentate, from anterior aspect”, though the illustrated photos (
|
||||
<bibRefCitation id="2A2382A42566FFCAFD6AF9412DD1F7A7" author="Ochi, T. & Kon, M." box="[716,956,1671,1695]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi & Kon 1996: 26</bibRefCitation>
|
||||
) showed an indistinctly emarginated clypeal anterior margin. However, both the description and photos of
|
||||
<taxonomicName id="89B284D62566FFCAFCE5F96D2DB3F7FB" authorityName="Ochi & Kon" authorityYear="1996" box="[835,990,1707,1731]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFCE5F96D2D31F7FA" box="[835,860,1707,1730]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFCCDF96A2D12F7FB" box="[875,895,1708,1731]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFC37F96D2DB3F7FB" box="[913,990,1707,1731]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="2A2382A42566FFCAFBAAF96D2A8BF7FB" author="Ochi, T. & Kon, M." box="[1036,1254,1707,1731]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi & Kon (1996)</bibRefCitation>
|
||||
showed that the clypeal anterior margin was not protruded, clearly distinguishing
|
||||
<taxonomicName id="89B284D62566FFCAFCF7F9092D84F7DF" authorityName="Ochi & Kon" authorityYear="1996" box="[849,1001,1743,1767]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFCF7F9092D07F7DE" box="[849,874,1743,1766]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFCDFF9162DE3F7DF" box="[889,910,1744,1767]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFC3AF9092D84F7DF" box="[924,1001,1743,1767]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
|
||||
</taxonomicName>
|
||||
from the new species. (**) We have not been able to examine the following five newly discovered species:
|
||||
<taxonomicName id="89B284D62566FFCAFB99F9352BF8F633" authorityName="Ochi, Kon & Pham" authorityYear="2019" box="[1087,1429,1779,1803]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bartolozzii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFB99F9352AE5F632" box="[1087,1160,1779,1802]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFB3CF9352B7FF632" box="[1178,1298,1779,1802]" italics="true" pageId="6" pageNumber="57">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFA85F9352BF8F633" box="[1315,1429,1779,1803]" italics="true" pageId="6" pageNumber="57">bartolozzii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="89B284D62566FFCAFF31F8D12F14F617" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[151,377,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chinmontanus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFF31F8D12EDDF616" box="[151,176,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF66F8DE2EB9F617" box="[192,212,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF45F8D12F14F617" box="[227,377,1815,1839]" italics="true" pageId="6" pageNumber="57">chinmontanus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="89B284D62566FFCAFE20F8D12C2DF617" authorityName="Ochi, Kon & Pham" authorityYear="2019" box="[390,576,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="montreuili" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFE20F8D12FF2F616" box="[390,415,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFE09F8DE2FAEF617" box="[431,451,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFE74F8D12C2DF617" box="[466,576,1815,1839]" italics="true" pageId="6" pageNumber="57">montreuili</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="89B284D62566FFCAFDE8F8D12C8BF617" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[590,742,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="olivieri" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFDE8F8D12C0BF616" box="[590,614,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFDD0F8DE2CE6F617" box="[630,651,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFD3FF8D12C8BF617" box="[665,742,1815,1839]" italics="true" pageId="6" pageNumber="57">olivieri</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="89B284D62566FFCAFCBDF8D12DBFF616" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[795,978,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="popaensis" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFCBDF8D12D59F616" box="[795,820,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFCE5F8DE2D35F617" box="[835,856,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFCC1F8D12DBFF616" box="[871,978,1815,1838]" italics="true" pageId="6" pageNumber="57">popaensis</emphasis>
|
||||
</taxonomicName>
|
||||
. The original descriptions and illustrations showed clear separation in clypeal morphological characters between these five species and
|
||||
<taxonomicName id="89B284D62566FFCAFBDEF8FC2BF1F66A" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[1144,1436,1850,1876]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFBDEF8FC2AA9F66C" box="[1144,1220,1850,1876]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFB72F8FA2A84F66B" box="[1236,1257,1852,1875]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFB5EF8FD2B04F66A" box="[1272,1385,1851,1874]" italics="true" pageId="6" pageNumber="57">javanensis</emphasis>
|
||||
new
|
||||
</taxonomicName>
|
||||
species (
|
||||
<tableCitation id="0330CAEE2566FFCAFF55F8992F2AF64F" box="[243,327,1887,1911]" captionStart="TABLE 2" captionStartId="6.[151,239,154,178]" captionTargetBox="[166,1421,261,1584]" captionText="TABLE 2. Diagnostic characters of clypeus separating the new species from the other Copris (Paracopris) species (based on the specimens examined by us in MNHN and NMPC, and data published by Wiedemann 1823; Arrow 1931; Balthasar 1939, 1963; Ochi & Kon 1996; Hanboonsong et al. 2003; Ochi et al. 2019a,b)" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="6" pageNumber="57" tableUuid="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2">Table 2</tableCitation>
|
||||
). In addition, the densely to contiguously punctate elytra of
|
||||
<taxonomicName id="89B284D62566FFCAFC69F8992AE0F64F" authorityName="Ochi, Kon & Pham" authorityYear="2019" box="[975,1165,1887,1911]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bartolozzii" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFC69F8992D8AF64E" box="[975,999,1887,1910]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFC51F8A62A61F64F" box="[1015,1036,1888,1911]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFBBDF8992AE0F64F" box="[1051,1165,1887,1911]" italics="true" pageId="6" pageNumber="57">bartolozzii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="7CC623472566FFCAFB3DF8992BF0F64E" box="[1179,1437,1887,1911]" italics="true" pageId="6" pageNumber="57">
|
||||
<taxonomicName id="89B284D62566FFCAFB3DF8992B15F64F" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[1179,1400,1887,1911]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chinmontanus" subGenus="Paracopris">C.(P.) chinmontanus</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="89B284D62566FFCAFA22F8992BF0F64E" authorityName="Ochi, Kon & Pham" authorityYear="2019" box="[1412,1437,1887,1910]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="montreuili" subGenus="Paracopris">C.</taxonomicName>
|
||||
</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFF06F8422ED8F6A3" box="[160,181,1924,1947]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFF62F8452F5FF6A3" box="[196,306,1923,1947]" italics="true" pageId="6" pageNumber="57">montreuili</emphasis>
|
||||
and
|
||||
<taxonomicName id="89B284D62566FFCAFEC0F8452C70F6A2" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[358,541,1923,1947]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="popaensis" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFEC0F8452F12F6A2" box="[358,383,1923,1946]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFE29F8422FCEF6A3" box="[399,419,1924,1947]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFE14F8452C70F6A2" box="[434,541,1923,1946]" italics="true" pageId="6" pageNumber="57">popaensis</emphasis>
|
||||
</taxonomicName>
|
||||
differentiate these species from
|
||||
<taxonomicName id="89B284D62566FFCAFCDCF8442AF3F6A2" authority="Bui & Ziegler & Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[890,1182,1922,1948]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFCDCF8442DABF6A4" box="[890,966,1922,1948]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFC70F8422D86F6A3" box="[982,1003,1924,1947]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFC5CF8452A06F6A2" box="[1018,1131,1923,1946]" italics="true" pageId="6" pageNumber="57">javanensis</emphasis>
|
||||
<taxonomicNameLabel id="67F59E3C2566FFCAFBD4F8452AF3F6A2" box="[1138,1182,1923,1946]" pageId="6" pageNumber="57" rank="species">new</taxonomicNameLabel>
|
||||
</taxonomicName>
|
||||
species, while the deep median sulcus of pronotal disc of
|
||||
<taxonomicName id="89B284D62566FFCAFDA7F8612CF4F687" authorityName="Ochi, Kon & Kawahara" authorityYear="2019" box="[513,665,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="olivieri" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472566FFCAFDA7F8612C74F686" box="[513,537,1959,1982]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472566FFCAFD8FF86E2C50F687" box="[553,573,1960,1983]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472566FFCAFDEAF8612CF4F687" box="[588,665,1959,1983]" italics="true" pageId="6" pageNumber="57">olivieri</emphasis>
|
||||
</taxonomicName>
|
||||
differentiates it from the new species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
120
data/C6/1B/4E/C61B4E432563FFCFFF31FDD92A97F2DC.xml
Normal file
120
data/C6/1B/4E/C61B4E432563FFCFFF31FDD92A97F2DC.xml
Normal file
|
|
@ -0,0 +1,120 @@
|
|||
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<mods:title id="1737D3550FAE0A345EA9B5B6E72B0B49">Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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</mods:mods>
|
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<treatment id="C61B4E432563FFCFFF31FDD92A97F2DC" ID-GBIF-Taxon="160797966" LSID="urn:lsid:plazi:treatment:C61B4E432563FFCFFF31FDD92A97F2DC" httpUri="http://treatment.plazi.org/id/C61B4E432563FFCFFF31FDD92A97F2DC" lastPageNumber="54" pageId="3" pageNumber="54">
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<subSubSection id="06A8ACDE2563FFCFFF31FDD92C09F302" box="[151,612,543,570]" pageId="3" pageNumber="54" type="nomenclature">
|
||||
<paragraph id="4E0DFF552563FFCFFF31FDD92C09F302" blockId="3.[151,612,543,570]" box="[151,612,543,570]" pageId="3" pageNumber="54">
|
||||
<heading id="154548392563FFCFFF31FDD92C09F302" bold="true" box="[151,612,543,570]" fontSize="11" level="1" pageId="3" pageNumber="54" reason="1">
|
||||
<emphasis id="7CC623472563FFCFFF31FDD92C09F302" bold="true" box="[151,612,543,570]" pageId="3" pageNumber="54">
|
||||
Subgenus
|
||||
<taxonomicName id="89B284D62563FFCFFEB1FDE62C09F302" ID-CoL="8CP9" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" box="[279,612,544,570]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472563FFCFFEB1FDE62FF6F302" bold="true" box="[279,411,544,570]" italics="true" pageId="3" pageNumber="54">Paracopris</emphasis>
|
||||
<bibRefCitation id="2A2382A42563FFCFFE05FDE62C09F302" author="Balthasar, V." box="[419,612,544,570]" pageId="3" pageNumber="54" pagination="1 - 36" refId="ref7382" refString="Balthasar, V. (1939) Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico delle R. Universita di Firenze. Redia, 25, 1 - 36." type="journal article" year="1939">Balthasar, 1939</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2563FFCFFF31FDA22A44F208" pageId="3" pageNumber="54" type="materials_examined">
|
||||
<paragraph id="4E0DFF552563FFCFFF31FDA22DCCF344" blockId="3.[151,929,611,636]" box="[151,929,611,636]" pageId="3" pageNumber="54">
|
||||
<typeStatus id="910941F72563FFCFFF31FDA22EA4F344" box="[151,201,612,636]" pageId="3" pageNumber="54">Type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="89B284D62563FFCFFEB9FDA22CCDF344" authority="Wiedemann, 1823" authorityName="Wiedemann" authorityYear="1823" box="[287,672,611,636]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="punctulatus">
|
||||
<emphasis id="7CC623472563FFCFFEB9FDA22F8CF344" box="[287,481,612,636]" italics="true" pageId="3" pageNumber="54">Copris punctulatus</emphasis>
|
||||
<bibRefCitation id="2A2382A42563FFCFFE41FDA52CCDF344" author="Wiedemann, C. R. W." box="[487,672,611,636]" pageId="3" pageNumber="54" pagination="1 - 164" refId="ref9330" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoolo- gisches Magazin, 2, 1 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(by original designation)
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552563FFCFFF31FD6C2A44F208" blockId="3.[151,1437,681,997]" pageId="3" pageNumber="54">
|
||||
<emphasis id="7CC623472563FFCFFF31FD6C2FC4F3FC" bold="true" box="[151,425,681,708]" pageId="3" pageNumber="54">Nomenclatural history.</emphasis>
|
||||
Recently, the subgenus
|
||||
<taxonomicName id="89B284D62563FFCFFD15FD6D2D5CF3FC" authorityName="Balthasar" authorityYear="1939" box="[691,817,683,708]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472563FFCFFD15FD6D2D5CF3FC" box="[691,817,683,708]" italics="true" pageId="3" pageNumber="54">Paracopris</emphasis>
|
||||
</taxonomicName>
|
||||
was considered as a genus by
|
||||
<bibRefCitation id="2A2382A42563FFCFFBD9FD6C2B28F3FC" author="Sabu, T. K. & Nithya, S. & Vinod, K. V." box="[1151,1349,682,709]" pageId="3" pageNumber="54" pagination="29 - 38" refId="ref8935" refString="Sabu, T. K., Nithya, S. & Vinod, K. V. (2011) Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa, 2830 (1), 29 - 38. https: // doi. org / 10.11646 / zootaxa. 2830.1.3" type="journal article" year="2011">
|
||||
Sabu
|
||||
<emphasis id="7CC623472563FFCFFB1AFD6D2A81F3FC" box="[1212,1260,682,708]" italics="true" pageId="3" pageNumber="54">et al</emphasis>
|
||||
. (2011)
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="2A2382A42563FFCFFAE9FD6C2F50F3D1" author="Kral, D. & Bezdek, A." pageId="3" pageNumber="54" pagination="167 - 171" refId="ref8237" refString="Kral, D. & Bezdek, A. (2016) Tribe Coprini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scara- baeoidea-Scirtoidea-Dasciloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 167 - 171." type="book chapter" year="2016">Král & Bezděk (2016)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="2A2382A42563FFCFFED4FD082C36F3D0" author="Cheung, K. & Bai, M. & Leung, M. & Lee, Y. & Chan, K. & Yang, X." box="[370,603,718,745]" pageId="3" pageNumber="54" pagination="233 - 267" refId="ref7699" refString="Cheung, K., Bai, M., Leung, M., Lee, Y., Chan, K. & Yang, X. (2018) Scarabaeinae (Coleoptera: Scarabaeidae) from Hong Kong. Zoological Systematics, 43 (3), 233 - 267. https: // doi. org / 10.11865 / zs. 201830" type="journal article" year="2018">
|
||||
Cheung
|
||||
<emphasis id="7CC623472563FFCFFE76FD092C6CF3D0" box="[464,513,718,744]" italics="true" pageId="3" pageNumber="54">et al</emphasis>
|
||||
. (2018)
|
||||
</bibRefCitation>
|
||||
, but a lack of the genetic analysis makes the generic placement of
|
||||
<emphasis id="7CC623472563FFCFFAE6FD092EAEF234" italics="true" pageId="3" pageNumber="54">Paraco- pris</emphasis>
|
||||
unclarified. In this study, we regarded
|
||||
<taxonomicName id="89B284D62563FFCFFDD7FD352C82F234" authorityName="Balthasar" authorityYear="1939" box="[625,751,755,780]" class="Insecta" family="Scarabaeidae" genus="Paracopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7CC623472563FFCFFDD7FD352C82F234" box="[625,751,755,780]" italics="true" pageId="3" pageNumber="54">Paracopris</emphasis>
|
||||
</taxonomicName>
|
||||
as a subgenus of the genus
|
||||
<taxonomicName id="89B284D62563FFCFFBB9FD352A06F235" authorityName="Geoffroy" authorityYear="1762" box="[1055,1131,755,781]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7CC623472563FFCFFBB9FD352A06F235" box="[1055,1131,755,781]" italics="true" pageId="3" pageNumber="54">Copris</emphasis>
|
||||
</taxonomicName>
|
||||
according to the traditional classification of
|
||||
<bibRefCitation id="2A2382A42563FFCFFEF6FCD02C61F208" author="Balthasar, V." box="[336,524,790,816]" pageId="3" pageNumber="54" pagination="1 - 36" refId="ref7382" refString="Balthasar, V. (1939) Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico delle R. Universita di Firenze. Redia, 25, 1 - 36." type="journal article" year="1939">Balthasar (1939)</bibRefCitation>
|
||||
that has been widely used in taxonomic studies.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2563FFCFFF61FCFF2A97F2DC" pageId="3" pageNumber="54" type="diagnosis">
|
||||
<paragraph id="4E0DFF552563FFCFFF61FCFF2A97F2DC" blockId="3.[151,1437,681,997]" pageId="3" pageNumber="54">
|
||||
<emphasis id="7CC623472563FFCFFF61FCFF2F2FF26C" bold="true" box="[199,322,825,852]" pageId="3" pageNumber="54">Diagnosis.</emphasis>
|
||||
Color: grayish-black, reddish brown or chocolate-brown. Whole surface slightly convex to fairly flat. Head broad. Clypeal surface smooth (i.e. minutely punctate). Clypeus and frons typically armed with a horn and/or carina. Pronotum broad, anteriorly untruncated and distinctly punctate. Elytra distinctly striate; elytral intervals strongly punctate. Tibia, compared with the other subgenera slender. Metatibia bear a weak transverse outer carina. Metatarsus slender, and with five tarsomeres; metatarsomeres approximately equal in length.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
80
data/C6/1B/4E/C61B4E432563FFCFFF31FEB72A28F0F5.xml
Normal file
80
data/C6/1B/4E/C61B4E432563FFCFFF31FEB72A28F0F5.xml
Normal file
|
|
@ -0,0 +1,80 @@
|
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<mods:mods id="352B38B7389F207D8918FF947BDD94AD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2BB5FA0D9CA1998769C6D3646AB00907">
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<mods:title id="C5B77E5F292046B66265DB5DB5625377">Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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|
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<mods:namePart id="B48D5CEA85B40C7CEF52CC937F66905A">Bui, Van Bac</mods:namePart>
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<mods:namePart id="194640031CF943501B0DA0C8832931F8">Ziegler, Thomas</mods:namePart>
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<mods:namePart id="013A9E439DF47167E090D9218C2841F1">Bonkowski, Michael</mods:namePart>
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<subSubSection id="06A8ACDE2563FFCFFF31FEB72F93F0B3" box="[151,510,369,395]" pageId="3" pageNumber="54" type="nomenclature">
|
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<paragraph id="4E0DFF552563FFCFFF31FEB72F93F0B3" blockId="3.[151,510,369,395]" box="[151,510,369,395]" pageId="3" pageNumber="54">
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<heading id="154548392563FFCFFF31FEB72F93F0B3" bold="true" box="[151,510,369,395]" fontSize="11" level="1" pageId="3" pageNumber="54" reason="1">
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<emphasis id="7CC623472563FFCFFF31FEB72F93F0B3" bold="true" box="[151,510,369,395]" pageId="3" pageNumber="54">
|
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Genus
|
||||
<taxonomicName id="89B284D62563FFCFFF48FEB72F93F0B3" ID-CoL="3TQY" authority="Geoffroy, 1762" authorityName="Geoffroy" authorityYear="1762" box="[238,510,369,395]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7CC623472563FFCFFF48FEB72F53F0B3" bold="true" box="[238,318,369,395]" italics="true" pageId="3" pageNumber="54">Copris</emphasis>
|
||||
Geoffroy, 1762
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2563FFCFFF31FE702A28F0F5" box="[151,1093,437,462]" pageId="3" pageNumber="54" type="materials_examined">
|
||||
<paragraph id="4E0DFF552563FFCFFF31FE702A28F0F5" blockId="3.[151,1093,437,462]" box="[151,1093,437,462]" pageId="3" pageNumber="54">
|
||||
<typeStatus id="910941F72563FFCFFF31FE702EA4F0F6" box="[151,201,438,462]" pageId="3" pageNumber="54">Type</typeStatus>
|
||||
species:
|
||||
<taxonomicName id="89B284D62563FFCFFE80FE732CE3F0F6" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[294,654,437,462]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="lunaris">
|
||||
<emphasis id="7CC623472563FFCFFE80FE732F86F0F5" box="[294,491,437,461]" italics="true" pageId="3" pageNumber="54">Scarabaeus lunaris</emphasis>
|
||||
Linnaeus, 1758
|
||||
</taxonomicName>
|
||||
(subsequent designation by Latreille 1810)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
535
data/C6/1B/4E/C61B4E432567FFC7FF31FF5F2AD6F088.xml
Normal file
535
data/C6/1B/4E/C61B4E432567FFC7FF31FF5F2AD6F088.xml
Normal file
|
|
@ -0,0 +1,535 @@
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<mods:title id="AC2B0D1ED81AE45082E1830C4B2C8EC3">Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<taxonomicName id="89B284D62567FFCBFF31FF5F2DF1F18B" ID-CoL="7ZBDP" authority="punctulatus Wiedemann, 1823" authorityName="punctulatus Wiedemann" authorityYear="1823" box="[151,924,153,180]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="punctulatus" subGenus="Paracopris" subSpecies="punctulatus">
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<emphasis id="7CC623472567FFCBFF31FF5F2DF1F18B" bold="true" box="[151,924,153,180]" pageId="7" pageNumber="58">
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<emphasis id="7CC623472567FFCBFF31FF5F2E8AF18B" bold="true" box="[151,231,153,179]" italics="true" pageId="7" pageNumber="58">Copris</emphasis>
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(
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<emphasis id="7CC623472567FFCBFF5EFF5F2F11F18B" bold="true" box="[248,380,153,179]" italics="true" pageId="7" pageNumber="58">Paracopris</emphasis>
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)
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<emphasis id="7CC623472567FFCBFE2BFF5F2CDEF18C" bold="true" box="[397,691,153,180]" italics="true" pageId="7" pageNumber="58">punctulatus punctulatus</emphasis>
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<bibRefCitation id="2A2382A42567FFCBFD1DFF5F2DF1F18B" author="Wiedemann, C. R. W." box="[699,924,153,179]" pageId="7" pageNumber="58" pagination="1 - 164" refId="ref9330" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoolo- gisches Magazin, 2, 1 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
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</emphasis>
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<paragraph id="4E0DFF552567FFCBFF31FF7B2FA5F1EF" blockId="7.[151,924,153,216]" box="[151,456,189,216]" pageId="7" pageNumber="58">
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(
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<figureCitation id="D689E3D02567FFCBFF39FF7B2F2DF1E0" box="[159,320,189,216]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="7" pageNumber="58">Figs. 2B, D, F</figureCitation>
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,
|
||||
<figureCitation id="D689E3D02567FFCBFEEDFF7B2F03F1EF" box="[331,366,189,215]" captionStart="FIGURES 3" captionStartId="9.[151,264,1806,1831]" captionTargetBox="[163,1427,949,1787]" captionTargetId="figure@9.[151,1435,949,1787]" captionTargetPageId="9" captionText="FIGURES 3A–C. Surface of elytral intervals of Copris (Paracopris) species. A, Copris (P.) javanensis Bui, Ziegler & Bonkowski, new species; B, Copris (P.) punctulatus punctulatus; C, Copris (P.) cariniceps." figureDoi="http://doi.org/10.5281/zenodo.3586832" httpUri="https://zenodo.org/record/3586832/files/figure.png" pageId="7" pageNumber="58">3B</figureCitation>
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,
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<figureCitation id="D689E3D02567FFCBFEDCFF7B2FE5F1EF" box="[378,392,189,215]" captionStart="FIGURES 4" captionStartId="10.[151,264,1962,1987]" captionTargetBox="[218,1366,169,1941]" captionTargetId="figure@10.[213,1375,166,1943]" captionTargetPageId="10" captionText="FIGURES 4A–D. Male specimen (A, B) and female specimen (C, D) of Copris (Paracopris) punctulatus punctulatus. A, C, habitus, dorsal view; B, D, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3586834" httpUri="https://zenodo.org/record/3586834/files/figure.png" pageId="7" pageNumber="58">4</figureCitation>
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A–D)
|
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<subSubSection id="06A8ACDE2567FFCBFF31FEC42DEFF08B" pageId="7" pageNumber="58" type="reference_group">
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<paragraph id="4E0DFF552567FFCBFF31FEC42D75F022" blockId="7.[151,1437,257,435]" box="[151,792,257,282]" pageId="7" pageNumber="58">
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<emphasis id="7CC623472567FFCBFF31FEC42F23F022" box="[151,334,258,282]" italics="true" pageId="7" pageNumber="58">Copris punctulata</emphasis>
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<treatmentCitation id="CF13D9442567FFCBFEF3FEC72C5EF022" author="Wiedemann, C. R. W." box="[341,563,257,282]" page="11" pageId="7" pageNumber="58" year="1823">
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<bibRefCitation id="2A2382A42567FFCBFEF3FEC72C5EF022" author="Wiedemann, C. R. W." box="[341,563,257,282]" pageId="7" pageNumber="58" pagination="1 - 164" refId="ref9330" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoolo- gisches Magazin, 2, 1 - 164." type="journal article" year="1823">Wiedemann, 1823: 11</bibRefCitation>
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(original description).
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</paragraph>
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<paragraph id="4E0DFF552567FFCBFF31FEE72DC2F000" blockId="7.[151,1437,257,435]" box="[151,943,288,313]" pageId="7" pageNumber="58">
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<taxonomicName id="89B284D62567FFCBFF31FEE72C7BF001" ID-CoL="Y5QP" authority=": Arrow 1931: 126" authorityName="Arrow" authorityPageNumber="126" authorityYear="1931" box="[151,534,288,313]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="punctulatus">
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<emphasis id="7CC623472567FFCBFF31FEE72F35F000" box="[151,344,288,312]" italics="true" pageId="7" pageNumber="58">Copris punctulatus</emphasis>
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:
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<treatmentCitation id="CF13D9442567FFCBFEC3FEE62C7BF001" author="Arrow, G. J." box="[357,534,288,313]" page="126" pageId="7" pageNumber="58" year="1931">
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<bibRefCitation id="2A2382A42567FFCBFEC3FEE62C7BF001" author="Arrow, G. J." box="[357,534,288,313]" pageId="7" pageNumber="58" refId="ref7267" refString="Arrow, G. J. (1931) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, 428 pp." type="book" year="1931">Arrow 1931: 126</bibRefCitation>
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</taxonomicName>
|
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;
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<treatmentCitation id="CF13D9442567FFCBFD87FEE62CD4F000" author="Sewak, R." box="[545,697,288,313]" page="3" pageId="7" pageNumber="58" year="2009">
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<bibRefCitation id="2A2382A42567FFCBFD87FEE62CD4F000" author="Sewak, R." box="[545,697,288,313]" pageId="7" pageNumber="58" pagination="1 - 106" refId="ref9154" refString="Sewak, R. (2009) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the zoological survey of India, 296, 1 - 106." type="journal article" year="2009">Sewak 2009: 3</bibRefCitation>
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</treatmentCitation>
|
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;
|
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<treatmentCitation id="CF13D9442567FFCBFD63FEE62DC6F000" author="Shahabuddin" box="[709,939,288,313]" page="31" pageId="7" pageNumber="58" year="2010">
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<bibRefCitation id="2A2382A42567FFCBFD63FEE62DC6F000" author="Shahabuddin" box="[709,939,288,313]" pageId="7" pageNumber="58" pagination="29 - 33" refId="ref9187" refString="Shahabuddin (2010) Diversity and community structure of dung beetles (Coleoptera: Scarabaeidae) across a habitat disturbance gradient in Lore Lindu National Park, Central Sulawesi. Biodiversitas, 11, 29 - 33. https: // doi. org / 10.13057 / biodiv / d 110107" type="journal article" year="2010">Shahabuddin 2010: 31</bibRefCitation>
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</treatmentCitation>
|
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="4E0DFF552567FFCBFF31FEF92AA1F04E" blockId="7.[151,1437,257,435]" pageId="7" pageNumber="58">
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<treatmentCitationGroup id="6EA2D87B2567FFCBFF31FEF92AA1F04E" pageId="7" pageNumber="58">
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<taxonomicName id="89B284D62567FFCBFF31FEF92CD1F06F" ID-CoL="Y5QP" authority=": Balthasar 1963: 370" authorityName=": Balthasar" authorityPageNumber="370" authorityYear="1963" box="[151,700,318,343]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="punctulatus" subGenus="Paracopris">
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<emphasis id="7CC623472567FFCBFF31FEF92EB1F06E" box="[151,220,319,342]" italics="true" pageId="7" pageNumber="58">Copris</emphasis>
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(
|
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<emphasis id="7CC623472567FFCBFF4DFE862F30F06F" box="[235,349,320,343]" italics="true" pageId="7" pageNumber="58">Paracopris</emphasis>
|
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)
|
||||
<emphasis id="7CC623472567FFCBFECDFEF92F8CF06F" box="[363,481,319,343]" italics="true" pageId="7" pageNumber="58">punctulatus</emphasis>
|
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:
|
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<treatmentCitation id="CF13D9442567FFCBFE49FEF82CD1F06F" author="Balthasar, V." box="[495,700,318,343]" page="370" pageId="7" pageNumber="58" year="1963">
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<bibRefCitation id="2A2382A42567FFCBFE49FEF82CD1F06F" author="Balthasar, V." box="[495,700,318,343]" pageId="7" pageNumber="58" refId="ref7444" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region. Coleo- ptera: Lamellicornia. Band 1. Allgemeiner Teil, Systematischer Teil: 1. Scarabaeinae, 2. Coprinae (Pinotini, Coprini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 391 pp." type="book" year="1963">Balthasar 1963: 370</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
, fig. 133;
|
||||
<treatmentCitation id="CF13D9442567FFCBFC84FEF82A6CF06F" author="Ochi, T. & Kon, M." box="[802,1025,318,343]" page="27" pageId="7" pageNumber="58" year="1996">
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<bibRefCitation id="2A2382A42567FFCBFC84FEF82A6CF06F" author="Ochi, T. & Kon, M." box="[802,1025,318,343]" pageId="7" pageNumber="58" pagination="29 - 35" refId="ref8462" refString="Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi & Kon 1996: 27</bibRefCitation>
|
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</treatmentCitation>
|
||||
, figs. 25–26;
|
||||
<treatmentCitation id="CF13D9442567FFCBFB2CFEF82E8EF04E" author="Kabakov, O. N. & Napolov, A." page="68" pageId="7" pageNumber="58" year="1999">
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<bibRefCitation id="2A2382A42567FFCBFB2CFEF82E8EF04E" author="Kabakov, O. N. & Napolov, A." pageId="7" pageNumber="58" pagination="58 - 96" refId="ref8124" refString="Kabakov, O. N. & Napolov, A. (1999) Fauna and ecology of Lamellicornia of subfamily Scarabaeinae of Vietnam and some parts of adjacent countries: South China, Laos, and Thailand. Latvijas Entomologs, 37, 58 - 96." type="journal article" year="1999">Kabakov & Napolov 1999: 68</bibRefCitation>
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</treatmentCitation>
|
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;
|
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<treatmentCitation id="CF13D9442567FFCBFF48FE9B2C4CF04E" author="Hanboonsong, Y. & Masumoto, K. & Ochi, T." box="[238,545,349,374]" page="116" pageId="7" pageNumber="58" year="2003">
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<bibRefCitation id="2A2382A42567FFCBFF48FE9B2C4CF04E" author="Hanboonsong, Y. & Masumoto, K. & Ochi, T." box="[238,545,349,374]" pageId="7" pageNumber="58" pagination="103 - 124" refId="ref7902" refString="Hanboonsong, Y., Masumoto, K. & Ochi, T. (2003) Dung beetles (Coleoptera, Scarabaeidae) of Thailand. Part 5. Genera Copris and Microcopris (Coprini). Elytra, 31, 103 - 124." type="journal article" year="2003">
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Hanboonsong
|
||||
<emphasis id="7CC623472567FFCBFE27FE982FC3F04E" box="[385,430,350,374]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
|
||||
. 2003: 116
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, figs. 65–68;
|
||||
<treatmentCitation id="CF13D9442567FFCBFD0DFE9B2DCFF04E" author="Mittal, I. C. & Jain, R." box="[683,930,349,374]" page="390" pageId="7" pageNumber="58" year="2015">
|
||||
<bibRefCitation id="2A2382A42567FFCBFD0DFE9B2DCFF04E" author="Mittal, I. C. & Jain, R." box="[683,930,349,374]" pageId="7" pageNumber="58" pagination="383 - 404" refId="ref8362" refString="Mittal, I. C. & Jain, R. (2015) A checklist of Indian dung beetles (Coleoptera: Scarabaeidae). Indian Journal of Entomology, 77 (4), 383 - 404. https: // doi. org / 10.5958 / 0974 - 8172.2015.00076.0" type="journal article" year="2015">Mittal & Jain 2015: 390</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="CF13D9442567FFCBFC0AFE9B2A15F04E" author="Bui, V. B. & Dumack, K. & Bonkowski, M." box="[940,1144,349,374]" page="180" pageId="7" pageNumber="58" year="2018">
|
||||
<bibRefCitation id="2A2382A42567FFCBFC0AFE9B2A15F04E" author="Bui, V. B. & Dumack, K. & Bonkowski, M." box="[940,1144,349,374]" pageId="7" pageNumber="58" pagination="167 - 191" refId="ref7585" refString="Bui, V. B., Dumack, K. & Bonkowski, M. (2018) Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species. European Journal of Entomology, 115, 167 - 191. https: // doi. org / 10.14411 / eje. 2018.016" type="journal article" year="2018">
|
||||
Bui
|
||||
<emphasis id="7CC623472567FFCBFC71FE982A6EF04E" box="[983,1027,350,374]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
|
||||
. 2018: 180
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, fig. 45.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF31FEBB2DEFF08B" blockId="7.[151,1437,257,435]" pageId="7" pageNumber="58">
|
||||
<treatmentCitationGroup id="6EA2D87B2567FFCBFF31FEBB2DEFF08B" pageId="7" pageNumber="58">
|
||||
<taxonomicName id="89B284D62567FFCBFF31FEBB2C06F0AD" ID-CoL="Y5QP" authority=": Lobl et al. 2006: 153" authorityName="Lobl" authorityPageNumber="153" authorityYear="2006" box="[151,619,380,405]" class="Insecta" family="Scarabaeidae" genus="Paracopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="punctulatus">
|
||||
<emphasis id="7CC623472567FFCBFF31FEBB2FE8F0AC" box="[151,389,380,404]" italics="true" pageId="7" pageNumber="58">Paracopris punctulatus</emphasis>
|
||||
:
|
||||
<treatmentCitation id="CF13D9442567FFCBFE35FEBA2C06F0AD" author="Lobl, I. & Krell, F. - T. & Kral, D." box="[403,619,380,405]" page="153" pageId="7" pageNumber="58" year="2006">
|
||||
<bibRefCitation id="2A2382A42567FFCBFE35FEBA2C06F0AD" author="Lobl, I. & Krell, F. - T. & Kral, D." box="[403,619,380,405]" pageId="7" pageNumber="58" pagination="151 - 154" refId="ref8299" refString="Lobl, I., Krell, F. - T. & Kral, D. (2006) Scarabaeidae: Scarabaeinae: Coprini. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 151 - 154." type="book chapter" year="2006">
|
||||
Löbl
|
||||
<emphasis id="7CC623472567FFCBFE6FFEBB2F91F0AC" box="[457,508,380,404]" italics="true" pageId="7" pageNumber="58">et al.</emphasis>
|
||||
2006: 153
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
;
|
||||
<treatmentCitation id="CF13D9442567FFCBFDD3FEBA2D4BF0AC" author="Bai, M. & Mccullough, E. & Song, K. - Q. & Liu, W. - G. & Yang, X. - K." box="[629,806,380,405]" page="4" pageId="7" pageNumber="58" year="2011">
|
||||
<bibRefCitation id="2A2382A42567FFCBFDD3FEBA2D4BF0AC" author="Bai, M. & Mccullough, E. & Song, K. - Q. & Liu, W. - G. & Yang, X. - K." box="[629,806,380,405]" pageId="7" pageNumber="58" pagination="21600" refId="ref7305" refString="Bai, M., Mccullough, E., Song, K. - Q., Liu, W. - G. & Yang, X. - K. (2011) Evolutionary constraints in hind wing shape in Chinese dung beetles (Coleoptera: Scarabaeinae). PLoS ONE, 6 (6), e 21600. https: // doi. org / 10.1371 / journal. pone. 0021600" type="journal article" year="2011">
|
||||
Bai
|
||||
<emphasis id="7CC623472567FFCBFD39FEBB2CA6F0AC" box="[671,715,380,404]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
|
||||
. 2011: 4
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, 6;
|
||||
<treatmentCitation id="CF13D9442567FFCBFCECFEBA2A7AF0AC" author="Sabu, T. K. & Nithya, S. & Vinod, K. V." box="[842,1047,380,405]" page="32" pageId="7" pageNumber="58" year="2011">
|
||||
<bibRefCitation id="2A2382A42567FFCBFCECFEBA2A7AF0AC" author="Sabu, T. K. & Nithya, S. & Vinod, K. V." box="[842,1047,380,405]" pageId="7" pageNumber="58" pagination="29 - 38" refId="ref8935" refString="Sabu, T. K., Nithya, S. & Vinod, K. V. (2011) Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa, 2830 (1), 29 - 38. https: // doi. org / 10.11646 / zootaxa. 2830.1.3" type="journal article" year="2011">
|
||||
Sabu
|
||||
<emphasis id="7CC623472567FFCBFC25FEBB2DC2F0AC" box="[899,943,380,404]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
|
||||
. 2011: 32
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="CF13D9442567FFCBFB87FEBA2B39F0AC" author="Kabakov, O. N. & Shokhin, I. V." box="[1057,1364,380,405]" page="49" pageId="7" pageNumber="58" year="2014">
|
||||
<bibRefCitation id="2A2382A42567FFCBFB87FEBA2B39F0AC" author="Kabakov, O. N. & Shokhin, I. V." box="[1057,1364,380,405]" pageId="7" pageNumber="58" pagination="47 - 59" refId="ref8172" refString="Kabakov, O. N. & Shokhin, I. V. (2014) Contribution to the knowledge of the subfamily Scarabaeinae (Coleoptera) from China with nomenclatural notes. Caucasian Entomological Bulletin, 10, 47 - 59. https: // doi. org / 10.23885 / 1814 - 3326 - 2014 - 10 - 1 - 47 - 59" type="journal article" year="2014">Kabakov & Shokhin 2014: 49</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="CF13D9442567FFCBFAF8FEBA2F04F08B" author="Gupta, D. & Chandra, K. & Khan, S." page="229" pageId="7" pageNumber="58" year="2014">
|
||||
<bibRefCitation id="2A2382A42567FFCBFAF8FEBA2F04F08B" author="Gupta, D. & Chandra, K. & Khan, S." pageId="7" pageNumber="58" pagination="225 - 240" refId="ref7848" refString="Gupta, D., Chandra, K. & Khan, S. (2014) An updated checklist of scarabaeoid beetles (Coleoptera: Scarabaeoidea) of Pench Tiger Reserve, Madhya Pradesh, India. Journal of Entomology and Zoology Studies, 2 (5), 225 - 240." type="journal article" year="2014">
|
||||
Gupta
|
||||
<emphasis id="7CC623472567FFCBFF61FE5A2E99F08B" box="[199,244,411,435]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
|
||||
. 2014: 229
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
; Bezděk &
|
||||
<treatmentCitation id="CF13D9442567FFCBFE46FE5C2C11F08B" author="Kral, D. & Bezdek, A." box="[480,636,410,435]" page="171" pageId="7" pageNumber="58" year="2016">
|
||||
<bibRefCitation id="2A2382A42567FFCBFE46FE5C2C11F08B" author="Kral, D. & Bezdek, A." box="[480,636,410,435]" pageId="7" pageNumber="58" pagination="167 - 171" refId="ref8237" refString="Kral, D. & Bezdek, A. (2016) Tribe Coprini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scara- baeoidea-Scirtoidea-Dasciloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 167 - 171." type="book chapter" year="2016">Král 2016: 171</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="CF13D9442567FFCBFD21FE5C2D13F08B" author="Cheung, K. & Bai, M. & Leung, M. & Lee, Y. & Chan, K. & Yang, X." box="[647,894,410,435]" page="242" pageId="7" pageNumber="58" year="2018">
|
||||
<bibRefCitation id="2A2382A42567FFCBFD21FE5C2D13F08B" author="Cheung, K. & Bai, M. & Leung, M. & Lee, Y. & Chan, K. & Yang, X." box="[647,894,410,435]" pageId="7" pageNumber="58" pagination="233 - 267" refId="ref7699" refString="Cheung, K., Bai, M., Leung, M., Lee, Y., Chan, K. & Yang, X. (2018) Scarabaeinae (Coleoptera: Scarabaeidae) from Hong Kong. Zoological Systematics, 43 (3), 233 - 267. https: // doi. org / 10.11865 / zs. 201830" type="journal article" year="2018">
|
||||
Cheung
|
||||
<emphasis id="7CC623472567FFCBFD7AFE5A2D65F08B" box="[732,776,411,435]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
|
||||
. 2018: 242
|
||||
</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2567FFC5FF31FE242FA0F026" lastPageId="9" lastPageNumber="60" pageId="7" pageNumber="58" type="materials_examined">
|
||||
<paragraph id="4E0DFF552567FFCBFF31FE242B0DF327" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
<emphasis id="7CC623472567FFCBFF31FE242F23F0C4" bold="true" box="[151,334,481,508]" pageId="7" pageNumber="58">Type specimen.</emphasis>
|
||||
<materialsCitation id="FEDAF5082567FFCBFEF2FE242B0DF327" ID-GBIF-Occurrence="2521263208" collectionCode="MNHN, R" country="Indonesia" location="Mt. Moeria" pageId="7" pageNumber="58" specimenCount="2" stateProvince="Jawa Timur" typeStatus="neotype">
|
||||
<emphasis id="7CC623472567FFCBFEF2FE242CD8F0C4" bold="true" box="[340,693,481,508]" pageId="7" pageNumber="58">
|
||||
<typeStatus id="910941F72567FFCBFEF2FE242FD5F0C4" box="[340,440,482,508]" pageId="7" pageNumber="58" type="neotype">Neotype</typeStatus>
|
||||
, present designation:
|
||||
</emphasis>
|
||||
male (
|
||||
<collectionCode id="28A367902567FFCBFCA6FE272D36F0C3" box="[768,859,481,507]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="58">MNHN</collectionCode>
|
||||
), “
|
||||
<collectingCountry id="36A5BFC52567FFCBFCDEFE272A7FF0C3" box="[888,1042,481,507]" name="Indonesia" pageId="7" pageNumber="58">INDONESIA</collectingCountry>
|
||||
|
|
||||
<collectingRegion id="8C7631B72567FFCBFB83FE242A3AF0C4" box="[1061,1111,482,508]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="58">Java</collectingRegion>
|
||||
Island |
|
||||
<location id="4B6DA98E2567FFCBFB12FE242B5BF0C3" LSID="urn:lsid:plazi:treatment:C61B4E432567FFC7FF31FF5F2AD6F088:4B6DA98E2567FFCBFB12FE242B5BF0C3" box="[1204,1334,481,508]" country="Indonesia" name="Mt. Moeria" pageId="7" pageNumber="58" stateProvince="Jawa Timur">Mt. Moeria</location>
|
||||
, 3–4000’ | ex Coll.
|
||||
<collectionCode id="28A367902567FFCBFEA5FDC02F74F318" box="[259,281,518,544]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="58">R</collectionCode>
|
||||
. Oberthür”, “
|
||||
<taxonomicName id="89B284D62567FFCBFE0BFDC32CEEF327" authorityName=": Balthasar" authorityYear="1963" box="[429,643,517,543]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="punctulatus">Copris punctulatus</taxonomicName>
|
||||
|
|
||||
<bibRefCitation id="2A2382A42567FFCBFD30FDC32D0FF327" author="Wiedemann, C. R. W." box="[662,866,517,543]" pageId="7" pageNumber="58" pagination="1 - 164" refId="ref9330" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoolo- gisches Magazin, 2, 1 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
|
||||
|
|
||||
<typeStatus id="910941F72567FFCBFCD0FDC32D94F327" box="[886,1017,517,543]" pageId="7" pageNumber="58" type="neotype">NEOTYPE</typeStatus>
|
||||
, male | Van Bac Bui des. 2019”.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FDEF2F22F35F" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
<emphasis id="7CC623472567FFCBFF61FDEF2C26F37C" bold="true" box="[199,587,553,580]" pageId="7" pageNumber="58">Additional specimens examined.</emphasis>
|
||||
<materialsCitation id="FEDAF5082567FFCBFDF2FDEF2AE6F37C" ID-GBIF-Occurrence="2521263205" box="[596,1163,552,580]" collectionCode="R" country="Indonesia" location="Mt Moeria" pageId="7" pageNumber="58" specimenCount="1" stateProvince="Jawa Timur">
|
||||
<emphasis id="7CC623472567FFCBFDF2FDEF2D7DF37C" bold="true" box="[596,784,553,580]" pageId="7" pageNumber="58">
|
||||
<collectingCountry id="36A5BFC52567FFCBFDF2FDEF2CA7F37C" box="[596,714,553,580]" name="Indonesia" pageId="7" pageNumber="58">Indonesia</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="8C7631B72567FFCBFD7EFDEC2D7DF37C" box="[728,784,554,580]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="58">Java</collectingRegion>
|
||||
</emphasis>
|
||||
:
|
||||
<location id="4B6DA98E2567FFCBFC86FDEC2DF7F37B" LSID="urn:lsid:plazi:treatment:C61B4E432567FFC7FF31FF5F2AD6F088:4B6DA98E2567FFCBFC86FDEC2DF7F37B" box="[800,922,553,580]" country="Indonesia" name="Mt Moeria" pageId="7" pageNumber="58" stateProvince="Jawa Timur">Mt Moeria</location>
|
||||
3–4000
|
||||
<superScript id="B9C7521D2567FFCBFC50FDEE2D96F30E" attach="none" box="[1014,1019,552,566]" fontSize="6" pageId="7" pageNumber="58">’</superScript>
|
||||
, ex. Coll.
|
||||
<collectionCode id="28A367902567FFCBFBD3FDEC2AE6F37C" box="[1141,1163,554,580]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="58">R</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="FEDAF5082567FFCBFB31FDEF2F26F35F" ID-GBIF-Occurrence="2521263203" collectionCode="MNHN" country="Indonesia" location="Oberthur" pageId="7" pageNumber="58" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Jawa Timur">
|
||||
<location id="4B6DA98E2567FFCBFB31FDEF2A93F37B" LSID="urn:lsid:plazi:treatment:C61B4E432567FFC7FF31FF5F2AD6F088:4B6DA98E2567FFCBFB31FDEF2A93F37B" box="[1175,1278,553,579]" country="Indonesia" name="Oberthur" pageId="7" pageNumber="58" stateProvince="Jawa Timur">Oberthür</location>
|
||||
,
|
||||
<specimenCount id="58B434DC2567FFCBFAACFDEF2B37F37B" box="[1290,1370,553,579]" pageId="7" pageNumber="58" type="male">1 male</specimenCount>
|
||||
,
|
||||
<specimenCount id="58B434DC2567FFCBFAC1FDEF2EB5F35F" pageId="7" pageNumber="58" type="female">3 females</specimenCount>
|
||||
(
|
||||
<collectionCode id="28A367902567FFCBFF41FD8B2F2EF35F" box="[231,323,589,615]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="58">MNHN</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FDB72A7AF397" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
Description of
|
||||
<typeStatus id="910941F72567FFCBFED6FDB42FA1F3B4" box="[368,460,626,652]" pageId="7" pageNumber="58" type="neotype">neotype</typeStatus>
|
||||
. Male. Total body length
|
||||
<quantity id="894A52B02567FFCBFD4AFDB72D3EF3B4" box="[748,851,625,652]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="7" pageNumber="58" unit="mm" value="17.5">17.5 mm</quantity>
|
||||
, body width 9.0 mm. Whole surface reddish brown. Mouthparts and antennomeres 1–6 reddish. Antennal club and setae on legs red.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FD7F2D85F25B" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
Head almost semicircular (HeadL
|
||||
<quantity id="894A52B02567FFCBFDECFD7F2CCEF3EC" box="[586,675,697,724]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="58" unit="mm" value="4.5">4.5 mm</quantity>
|
||||
, HeadW 7.0 mm). Clypeus smooth and impunctate; clypeal surface near anterior margin very shiny; clypeal apex distinctly upwards protruded, and widely and shallowly emarginate. Genae rectangular, separated from clypeus by well-defined clypeo-genal suture; genal surface minutely and sparsely punctate. Vertex strongly and densely punctate; punctures of vertex unevenly distributed and unequal in size, denser and coarser at areas near eyes. Cephalic horn small but distinct.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FCA82CECF520" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
Pronotum (PronL
|
||||
<quantity id="894A52B02567FFCBFE3FFCAB2F98F2B0" box="[409,501,877,904]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="58" unit="mm" value="6.2">6.2 mm</quantity>
|
||||
, PronW
|
||||
<quantity id="894A52B02567FFCBFDF8FCAB2CD5F2B0" box="[606,696,877,904]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="58" unit="mm" value="7.5">7.5 mm</quantity>
|
||||
) without anterior declivity; anterior margin slightly downwards curved in middle; anterolateral angles obtuse. Pronotal median sulcus weak; sulcus not extending to pronotal anterior margin. Whole surface of pronotum deeply and densely punctate; punctures slightly larger in middle than in sides and also in anterior part. Each side of pronotum with one deep and large excavation in middle; surface of excavation coarsely but shallowly punctate.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FBE72ACEF504" blockId="7.[151,1437,481,1984]" box="[199,1187,1057,1084]" pageId="7" pageNumber="58">Ventral surface of anterior angles of prothorax flat, and shallowly and sparsely punctate.</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FB832AD5F59F" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
Elytra (ElyL
|
||||
<quantity id="894A52B02567FFCBFEFCFB832FDDF558" box="[346,432,1093,1120]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="7" pageNumber="58" unit="mm" value="9.3">9.3 mm</quantity>
|
||||
) deeply striate; striae sparsely and equally punctate. Elytral intervals convex and strongly punctate; interval punctures dense, evenly distributed, and relatively equal in size; interval punctures slightly deeper than strial punctures. Each elytron with one rather abrupt vertical flattening near sutural angle.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FB772C54F5C8" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">Mesepimeron flat; surface weakly punctate. Metepisternum flat; surface sparsely but distinctly punctate. Mesosternum flat; surface strongly rugose.</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FB3F2FA4F400" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">Metasternal plate smooth, and with median deep groove at basal half; metasternal sides sparsely but distinctly punctate and without setae.</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61FA842C7EF737" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">Legs. Ventral surface of profemora strongly punctate; punctures unevenly distributed, and unequal in size, becoming denser and coarser at posterior half of femora; a few punctures near profemoral posterior margin with yellow setae. Profemoral margin with yellow, dense and long setae. Protibia with four lateral teeth, and protibial surface smooth and impunctate. Protibial spurs broadly dilated in middle, relatively pointed and slightly curved outwards near apex. Protarsus with five tarsomeres; protarsomere 1 and 5 equal in length, and both slightly longer than individual protarsomere 2–4.</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61F9DC2F48F798" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">Most ventral surface of mesofemora impunctate, except for small area behind anterior margin with a few strong but sparse punctures. Mesotibia smooth and broadly dilated at posterior end. Two mesotibial spurs almost equal in length and pointed. Mesotarsus with five tarsomeres; tarsomeres elongated, almost equal in length and with yellow scanty setae.</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61F96F2D58F66B" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">Almost whole ventral surface of metafemora smooth and impunctate, except for apical areas near anterior margin distinctly punctate; punctures with relatively long and red setae. Metatibia smooth and broadly dilated at posterior end, with one spur long and pointed. Metatarsus with five tarsomeres, all tarsomeres bear red and scantly setae; basal metatarsomere slender, and slightly longer than individual metatarsomeres 2–4; basal metatarsomere, metatarsomere 5 and metatibial spur almost equal in length.</paragraph>
|
||||
<paragraph id="4E0DFF552567FFCBFF61F89B2AEBF687" blockId="7.[151,1437,481,1984]" pageId="7" pageNumber="58">
|
||||
Abdomen and pygidium. Abdominal ventrites slightly narrower at midline; most surface of abdomen smooth, except for small areas at sides distinctly punctate. Pygidium (PyL
|
||||
<quantity id="894A52B02567FFCBFC2DF8472D8BF6A4" box="[907,998,1921,1948]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="58" unit="mm" value="1.6">1.6 mm</quantity>
|
||||
; PyW
|
||||
<quantity id="894A52B02567FFCBFB95F8472AE6F6A4" box="[1075,1163,1921,1948]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="58" unit="mm" value="3.3">3.3 mm</quantity>
|
||||
) with complete margin; surface slightly convex and deeply punctate; pygidial punctures very dense to contiguous.
|
||||
</paragraph>
|
||||
<paragraph id="4E0DFF552568FFC4FF61FF5F2A77F1EF" blockId="8.[151,1437,153,323]" pageId="8" pageNumber="59">Aedeagus. Phallobase much longer and wider than parameres. A broad angle almost reaching 180º formed by phallobase and parameres. Parameres strongly curved downwards in apical part.</paragraph>
|
||||
<paragraph id="4E0DFF552568FFC4FF61FF262D3DF07B" blockId="8.[151,1437,153,323]" pageId="8" pageNumber="59">
|
||||
<emphasis id="7CC623472568FFC4FF61FF262FD5F1C3" bold="true" box="[199,440,224,251]" pageId="8" pageNumber="59">Sexual dimorphism.</emphasis>
|
||||
Females differ from males in the following characters: Clypeal anterior margin not protruded, and without emargination. Head with one strongly curved transverse carina between clypeus and frons. Pronotal anterior margin distinctly forwards curved in middle.
|
||||
</paragraph>
|
||||
<caption id="1ACDAFDD2568FFC4FF31F8A72EA3F6FA" ID-DOI="http://doi.org/10.5281/zenodo.3586830" ID-Zenodo-Dep="3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="8" pageNumber="59" startId="8.[151,264,1889,1914]" targetBox="[164,1427,351,1859]" targetPageId="8">
|
||||
<paragraph id="4E0DFF552568FFC4FF31F8A72EA3F6FA" blockId="8.[151,1436,1889,1986]" pageId="8" pageNumber="59">
|
||||
<emphasis id="7CC623472568FFC4FF31F8A72F21F642" bold="true" box="[151,332,1889,1914]" pageId="8" pageNumber="59">FIGURES 2A–F.</emphasis>
|
||||
Diagnostic characters separating
|
||||
<taxonomicName id="89B284D62568FFC4FD06F8A42A9EF642" authority="Bui, Ziegler & Bonkowski" authorityName="Bui, Ziegler & Bonkowski" authorityYear="2019" box="[672,1267,1889,1914]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
|
||||
<emphasis id="7CC623472568FFC4FD06F8A42C88F641" box="[672,741,1890,1913]" italics="true" pageId="8" pageNumber="59">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472568FFC4FD55F8A42D08F641" box="[755,869,1890,1913]" italics="true" pageId="8" pageNumber="59">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472568FFC4FCD5F8A42DB3F641" box="[883,990,1890,1913]" italics="true" pageId="8" pageNumber="59">javanensis</emphasis>
|
||||
Bui, Ziegler & Bonkowski
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="67F59E3C2568FFC4FB5AF8A42B18F642" box="[1276,1397,1889,1914]" pageId="8" pageNumber="59" rank="species">new species</taxonomicNameLabel>
|
||||
(A, C, E) from
|
||||
<taxonomicName id="89B284D62568FFC4FEAAF8402C25F6A5" authority="punctulatus" authorityName="punctulatus Wiedemann" authorityYear="1823" box="[268,584,1925,1950]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="subSpecies" species="punctulatus" subGenus="Paracopris" subSpecies="punctulatus">
|
||||
<emphasis id="7CC623472568FFC4FEAAF8402F49F6A5" box="[268,292,1926,1949]" italics="true" pageId="8" pageNumber="59">C.</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472568FFC4FE95F8402F2AF6A5" box="[307,327,1926,1949]" italics="true" pageId="8" pageNumber="59">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472568FFC4FEF3F8432C25F6A5" box="[341,584,1925,1949]" italics="true" pageId="8" pageNumber="59">punctulatus punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
(B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4E0DFF552569FFC5FF61FF5E2FA0F026" blockId="9.[151,1437,152,934]" pageId="9" pageNumber="60">
|
||||
<emphasis id="7CC623472569FFC5FF61FF5E2F2AF18B" bold="true" box="[199,327,152,179]" pageId="9" pageNumber="60">Variability</emphasis>
|
||||
. Total body length
|
||||
<quantity id="894A52B02569FFC5FD85FF5E2CA4F18B" box="[547,713,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.85" metricValueMin="1.25" pageId="9" pageNumber="60" unit="mm" value="15.5" valueMax="18.5" valueMin="12.5">12.5–18.5 mm</quantity>
|
||||
, body width
|
||||
<quantity id="894A52B02569FFC5FCFDFF5E2D8EF18B" box="[859,995,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="9.5" metricValueMin="6.2" pageId="9" pageNumber="60" unit="mm" value="7.85" valueMax="9.5" valueMin="6.2">6.2–9.5 mm</quantity>
|
||||
(n=5). There is a significant difference in size of the protruding part of clypeal apex between small and large-bodied males with the latter having a more strongly upwards protruded clypeal apex. Females show no significant differences in external morphology between large and small individuals.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="06A8ACDE2569FFC7FF61FEEF2AD6F088" lastPageId="11" lastPageNumber="62" pageId="9" pageNumber="60" type="discussion">
|
||||
<paragraph id="4E0DFF552569FFC5FF61FEEF2F18F29E" blockId="9.[151,1437,152,934]" pageId="9" pageNumber="60">
|
||||
<emphasis id="7CC623472569FFC5FF61FEEF2F56F07B" bold="true" box="[199,315,297,323]" pageId="9" pageNumber="60">Remarks.</emphasis>
|
||||
<taxonomicName id="89B284D62569FFC5FEE7FEEF2CC1F07A" authorityName=": Balthasar" authorityYear="1963" box="[321,684,296,323]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472569FFC5FEE7FEEF2FE0F07B" box="[321,397,297,323]" italics="true" pageId="9" pageNumber="60">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472569FFC5FE3AFEEF2C77F07A" box="[412,538,297,322]" italics="true" pageId="9" pageNumber="60">Paracopris</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472569FFC5FD8FFEEE2CC1F07A" box="[553,684,296,322]" italics="true" pageId="9" pageNumber="60">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
was originally described with only 20 text lines and lacking illustrations. Although this species was redescribed by
|
||||
<bibRefCitation id="2A2382A42569FFC5FD0CFE8A2D09F05F" author="Balthasar, V." box="[682,868,332,359]" pageId="9" pageNumber="60" refId="ref7444" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region. Coleo- ptera: Lamellicornia. Band 1. Allgemeiner Teil, Systematischer Teil: 1. Scarabaeinae, 2. Coprinae (Pinotini, Coprini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 391 pp." type="book" year="1963">Balthasar (1963)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="2A2382A42569FFC5FC3FFE8A2A18F05F" author="Ochi, T. & Kon, M." box="[921,1141,332,359]" pageId="9" pageNumber="60" pagination="29 - 35" refId="ref8462" refString="Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi & Kon (1996)</bibRefCitation>
|
||||
, details on its intraspecific morphological variation are still lacking. Additionally the lack of illustrations of this species poses a great challenge for the reliable identification of this species. Therefore we provide detailed measurements of numerous morphological traits of this species based on a series of examined individuals along with the photographs. Although Christian Rudolf Wilhelm Wiedemann (1770–1840) worked mainly on Diptera, he also published descriptions of
|
||||
<taxonomicName id="89B284D62569FFC5FA87FE1A2BF1F0CE" authorityName="Linnaeus" authorityYear="1758" box="[1313,1436,476,502]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="60" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="2A2382A42569FFC5FF39FDC62F24F322" author="Fairchild, G. B." box="[159,329,512,538]" pageId="9" pageNumber="60" pagination="73 - 104" refId="ref7805" refString="Fairchild, G. B. (1967) Notes on neotropical Tabanidae (Diptera), VII. The species described by C. R. W. Wiedemann. Pacific Insects, 9 (1), 73 - 104." type="journal article" year="1967">Fairchild 1967</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42569FFC5FEF1FDC72FA2F323" author="Pont, A. C." box="[343,463,513,539]" pageId="9" pageNumber="60" pagination="87 - 122" refId="ref8832" refString="Pont, A. C. (1997) The Muscidae and Fanniidae (Insecta, Diptera) described by C. R. W. Wiedemann. Steenstrupia, 23, 87 - 122." type="journal article" year="1997">Pont 1997</bibRefCitation>
|
||||
). Most of the
|
||||
<taxonomicName id="89B284D62569FFC5FDD6FDC62C86F322" authorityName="Linnaeus" authorityYear="1758" box="[624,747,512,538]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="60" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
|
||||
types of Wiedemann were deposited in the Natural History Museum of
|
||||
<collectingCountry id="36A5BFC52569FFC5FE84FDE22FE3F306" box="[290,398,548,574]" name="Denmark" pageId="9" pageNumber="60">Denmark</collectingCountry>
|
||||
and the Zoological Museum Hamburg (ZMH) (
|
||||
<bibRefCitation id="2A2382A42569FFC5FC06FDE22A5AF307" author="Zimsen, E." box="[928,1079,548,575]" pageId="9" pageNumber="60" pagination="1 - 43" refId="ref9365" refString="Zimsen, E. (1954) The insect types of C. R. W. Wiedemann in the Zoological Museum in Copenhagen. Spolia Zoologica Musei Hauniensis, 14, 1 - 43." type="journal article" year="1954">Zimsen 1954</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="2A2382A42569FFC5FBE7FDE32A97F307" author="Horn, W. & Kahle, I. & Friese, G. & Gaedike, R." box="[1089,1274,548,575]" pageId="9" pageNumber="60" pagination="223 - 573" refId="ref8021" refString="Horn, W., Kahle, I., Friese, G. & Gaedike, R. (1990) Collectiones Entomologicae. Ein Kompendium uber den Verbleib ento- mologischer Sammlungen der Welt bis 1960. Teil II. L bis Z. Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin, 351 pp. [pp. 223 - 573]" type="book chapter" year="1990">
|
||||
Horn
|
||||
<emphasis id="7CC623472569FFC5FB27FDE32AD5F306" box="[1153,1208,548,574]" italics="true" pageId="9" pageNumber="60">et al.</emphasis>
|
||||
1990
|
||||
</bibRefCitation>
|
||||
). According to
|
||||
<bibRefCitation id="2A2382A42569FFC5FF31FD8F2F58F35B" author="Arrow, G. J." box="[151,309,585,611]" pageId="9" pageNumber="60" refId="ref7267" refString="Arrow, G. J. (1931) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, 428 pp." type="book" year="1931">Arrow (1931)</bibRefCitation>
|
||||
, the type of
|
||||
<taxonomicName id="89B284D62569FFC5FE66FD8F2CFBF35A" authorityName=": Balthasar" authorityYear="1963" box="[448,662,584,611]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="punctulatus">
|
||||
<emphasis id="7CC623472569FFC5FE66FD8F2CFBF35A" box="[448,662,584,611]" italics="true" pageId="9" pageNumber="60">Copris punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
was deposited in ZMH. However, the major parts of the entomology collection of ZMH were lost in 1943 including an estimated 10.000 insect types (
|
||||
<bibRefCitation id="2A2382A42569FFC5FB93FDAA2AB4F3BF" author="Weidner, H." box="[1077,1241,620,647]" pageId="9" pageNumber="60" pagination="87 - 264" refId="ref9288" refString="Weidner, H. (1976) Die Entomologischen Sammlungen des Zoologischen Instituts und Zoologischen Museums der Universitat Hamburg, IX. Teil, Insecta VI. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut, 73, 87 - 264." type="journal article" year="1976">Weidner 1976</bibRefCitation>
|
||||
). It is very likely that the type of
|
||||
<taxonomicName id="89B284D62569FFC5FEE3FD572F85F392" authorityName=": Balthasar" authorityYear="1963" box="[325,488,656,683]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="punctulatus">
|
||||
<emphasis id="7CC623472569FFC5FEE3FD572F85F392" box="[325,488,656,683]" italics="true" pageId="9" pageNumber="60">C. punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
was also lost during this time, as the entomology collection of ZMH (curator: Martin Husemann) currently contains only the specimens of
|
||||
<taxonomicName id="89B284D62569FFC5FD57FD732DF8F3F6" authorityName=": Balthasar" authorityYear="1963" box="[753,917,692,719]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="punctulatus">
|
||||
<emphasis id="7CC623472569FFC5FD57FD732DF8F3F6" box="[753,917,692,719]" italics="true" pageId="9" pageNumber="60">C. punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
collected by E. Ross in
|
||||
<date id="3A0CD9952569FFC5FB0EFD732B3FF3F7" box="[1192,1362,692,719]" pageId="9" pageNumber="60" value="1911" valueMax="1914">1911 and 1914</date>
|
||||
, Klara Herre in 1949 and Le Moult in 1957. As the real type specimen was lost, we here designate a
|
||||
<typeStatus id="910941F72569FFC5FB69FD1F2B45F3CB" box="[1231,1320,729,755]" pageId="9" pageNumber="60" type="neotype">neotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="89B284D62569FFC5FAF6FD1F2F77F22E" authorityName=": Balthasar" authorityYear="1963" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="punctulatus">
|
||||
<emphasis id="7CC623472569FFC5FAF6FD1F2F77F22E" italics="true" pageId="9" pageNumber="60">Copris punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
using a male specimen from
|
||||
<collectingRegion id="8C7631B72569FFC5FDC5FD3B2CF8F22F" box="[611,661,765,791]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="60">Java</collectingRegion>
|
||||
(
|
||||
<collectingCountry id="36A5BFC52569FFC5FD05FD3A2D7BF22F" box="[675,790,764,791]" name="Indonesia" pageId="9" pageNumber="60">Indonesia</collectingCountry>
|
||||
), being from the same locality mentioned by Wiedemann. The
|
||||
<typeStatus id="910941F72569FFC5FF6DFCE72F49F203" box="[203,292,801,827]" pageId="9" pageNumber="60" type="neotype">neotype</typeStatus>
|
||||
was designated to maintain nomenclatural stability as Articles 75.3.4 and 75.3.6 of ICZN (1999) required. These detailed morphological description and
|
||||
<typeStatus id="910941F72569FFC5FD5EFC832D3CF267" box="[760,849,837,863]" pageId="9" pageNumber="60" type="neotype">neotype</typeStatus>
|
||||
designation intent to enable a reliable identification of species boundaries within the subgenus
|
||||
<taxonomicName id="89B284D62569FFC5FDD8FCAF2C91F2BA" authorityName="Balthasar" authorityYear="1939" box="[638,764,873,898]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472569FFC5FDD8FCAF2C91F2BA" box="[638,764,873,898]" italics="true" pageId="9" pageNumber="60">Paracopris</emphasis>
|
||||
</taxonomicName>
|
||||
, and provide a guideline for discovery of potentially novel
|
||||
<taxonomicName id="89B284D62569FFC5FF31FC4B2F78F29E" authorityName="Balthasar" authorityYear="1939" box="[151,277,909,934]" class="Insecta" family="Scarabaeidae" genus="Paracopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7CC623472569FFC5FF31FC4B2F78F29E" box="[151,277,909,934]" italics="true" pageId="9" pageNumber="60">Paracopris</emphasis>
|
||||
</taxonomicName>
|
||||
species.
|
||||
</paragraph>
|
||||
<caption id="1ACDAFDD2569FFC5FF31F8C82A5EF672" ID-DOI="http://doi.org/10.5281/zenodo.3586832" ID-Zenodo-Dep="3586832" httpUri="https://zenodo.org/record/3586832/files/figure.png" pageId="9" pageNumber="60" startId="9.[151,264,1806,1831]" targetBox="[163,1427,949,1787]" targetPageId="9">
|
||||
<paragraph id="4E0DFF552569FFC5FF31F8C82A5EF672" blockId="9.[151,1437,1806,1867]" pageId="9" pageNumber="60">
|
||||
<emphasis id="7CC623472569FFC5FF31F8C82F35F61F" bold="true" box="[151,344,1806,1831]" pageId="9" pageNumber="60">FIGURES 3A–C.</emphasis>
|
||||
Surface of elytral intervals of
|
||||
<taxonomicName id="89B284D62569FFC5FD0CF8C92D10F61F" authority="" authorityName="Balthasar" authorityYear="1939" box="[682,893,1807,1831]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472569FFC5FD0CF8C92C82F61E" box="[682,751,1807,1830]" italics="true" pageId="9" pageNumber="60">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472569FFC5FCA5F8D62D18F61F" box="[771,885,1808,1831]" italics="true" pageId="9" pageNumber="60">Paracopris</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species. A,
|
||||
<taxonomicName id="89B284D62569FFC5FBA0F8C92A26F61E" authorityName="Geoffroy" authorityYear="1762" box="[1030,1099,1807,1830]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7CC623472569FFC5FBA0F8C92A26F61E" box="[1030,1099,1807,1830]" italics="true" pageId="9" pageNumber="60">Copris</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicName id="89B284D62569FFC5FBF0F8C92F60F673" authority="Bui, Ziegler & Bonkowski" authorityName="Bui, Ziegler & Bonkowski" authorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
|
||||
(
|
||||
<emphasis id="7CC623472569FFC5FBF8F8D62A00F61F" box="[1118,1133,1808,1831]" italics="true" pageId="9" pageNumber="60">P</emphasis>
|
||||
.)
|
||||
<emphasis id="7CC623472569FFC5FB2EF8D62A9EF61F" box="[1160,1267,1808,1831]" italics="true" pageId="9" pageNumber="60">javanensis</emphasis>
|
||||
Bui, Ziegler & Bonkowski
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="67F59E3C2569FFC5FEB1F8F52FF9F673" box="[279,404,1842,1867]" pageId="9" pageNumber="60" rank="species">new species</taxonomicNameLabel>
|
||||
; B,
|
||||
<taxonomicName id="89B284D62569FFC5FE1AF8F52D49F673" authority="punctulatus" authorityName="punctulatus Wiedemann" authorityYear="1823" box="[444,804,1843,1867]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="punctulatus" subGenus="Paracopris" subSpecies="punctulatus">
|
||||
<emphasis id="7CC623472569FFC5FE1AF8F52C6CF672" box="[444,513,1843,1866]" italics="true" pageId="9" pageNumber="60">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472569FFC5FDB6F8F22C4EF673" box="[528,547,1844,1867]" italics="true" pageId="9" pageNumber="60">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472569FFC5FD97F8F52D49F673" box="[561,804,1843,1867]" italics="true" pageId="9" pageNumber="60">punctulatus punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
; C,
|
||||
<taxonomicName id="89B284D62569FFC5FCE9F8F52A43F673" authorityName="Felsche" authorityYear="1910" box="[847,1070,1843,1867]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="cariniceps" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472569FFC5FCE9F8F52DF9F672" box="[847,916,1843,1866]" italics="true" pageId="9" pageNumber="60">Copris</emphasis>
|
||||
(
|
||||
<emphasis id="7CC623472569FFC5FC04F8F22DDBF673" box="[930,950,1844,1867]" italics="true" pageId="9" pageNumber="60">P.</emphasis>
|
||||
)
|
||||
<emphasis id="7CC623472569FFC5FC62F8F22A43F673" box="[964,1070,1844,1867]" italics="true" pageId="9" pageNumber="60">cariniceps</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4E0DFF552569FFC7FF61F8BF2AD6F088" blockId="9.[151,1436,1913,2012]" lastBlockId="11.[151,1437,153,432]" lastPageId="11" lastPageNumber="62" pageId="9" pageNumber="60">
|
||||
Recently, Ochi
|
||||
<emphasis id="7CC623472569FFC5FED3F8BC2FC3F6AB" box="[373,430,1913,1939]" italics="true" pageId="9" pageNumber="60">et al.</emphasis>
|
||||
(2019) described a new subspecies of
|
||||
<taxonomicName id="89B284D62569FFC5FCC7F8BC2A55F6AB" authorityName=": Balthasar" authorityYear="1963" box="[865,1080,1913,1940]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC623472569FFC5FCC7F8BC2D19F6AC" box="[865,884,1914,1940]" italics="true" pageId="9" pageNumber="60">C</emphasis>
|
||||
. (
|
||||
<emphasis id="7CC623472569FFC5FC2AF8BC2DF0F6AB" box="[908,925,1914,1939]" italics="true" pageId="9" pageNumber="60">P</emphasis>
|
||||
.)
|
||||
<emphasis id="7CC623472569FFC5FC13F8BF2A55F6AB" box="[949,1080,1913,1939]" italics="true" pageId="9" pageNumber="60">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
under the name
|
||||
<emphasis id="7CC623472569FFC5FB55F8BC2B6BF6AC" box="[1267,1286,1914,1940]" italics="true" pageId="9" pageNumber="60">C</emphasis>
|
||||
. (
|
||||
<emphasis id="7CC623472569FFC5FAB8F8BC2B42F6AB" box="[1310,1327,1914,1939]" italics="true" pageId="9" pageNumber="60">P</emphasis>
|
||||
.)
|
||||
<emphasis id="7CC623472569FFC5FAE1F8BC2F11F68F" italics="true" pageId="9" pageNumber="60">punctu- latus septentrionalis</emphasis>
|
||||
<bibRefCitation id="2A2382A42569FFC5FE23F85B2CC7F68F" author="Ochi, T. & Kon, M. & Pham, H. T." box="[389,682,1949,1976]" pageId="9" pageNumber="60" pagination="435 - 446" refId="ref8783" refString="Ochi, T., Kon, M. & Pham, H. T. (2019 b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435 - 446." type="journal article" year="2019">Ochi, Kon & Pham, 2019</bibRefCitation>
|
||||
based on eight specimens from northern
|
||||
<collectingCountry id="36A5BFC52569FFC5FB22F85B2A8EF68F" box="[1156,1251,1949,1975]" name="Vietnam" pageId="9" pageNumber="60">Vietnam</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="8C7631B72569FFC5FB55F85B2EA7F6E4" country="Vietnam" name="Phu Tho" pageId="9" pageNumber="60">Phu Tho Province</collectingRegion>
|
||||
). According to the original description and illustration (
|
||||
<bibRefCitation id="2A2382A42569FFC5FC9CF8072A1FF6E4" author="Ochi, T. & Kon, M. & Pham, H. T." box="[826,1138,1985,2012]" pageId="9" pageNumber="60" pagination="435 - 446" refId="ref8783" refString="Ochi, T., Kon, M. & Pham, H. T. (2019 b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435 - 446." type="journal article" year="2019">
|
||||
Ochi
|
||||
<emphasis id="7CC623472569FFC5FCDFF8042DDFF6E3" box="[889,946,1985,2011]" italics="true" pageId="9" pageNumber="60">et al.</emphasis>
|
||||
2019b: 444–445
|
||||
</bibRefCitation>
|
||||
, fig. 5) and four examined specimens (
|
||||
<specimenCount id="58B434DC256BFFC7FEBBFF5F2F05F18B" box="[285,360,153,179]" pageId="11" pageNumber="62" type="male">1 male</specimenCount>
|
||||
,
|
||||
<specimenCount id="58B434DC256BFFC7FED4FF5F2FB2F18C" box="[370,479,153,180]" pageId="11" pageNumber="62" type="female">3 females</specimenCount>
|
||||
) in MNHN (labelled:
|
||||
<collectingCountry id="36A5BFC5256BFFC7FD75FF5F2D58F18B" box="[723,821,153,179]" name="Vietnam" pageId="11" pageNumber="62">Vietnam</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="8C7631B7256BFFC7FC98FF5F2A2EF18B" box="[830,1091,153,179]" country="Vietnam" name="Tuyen Quang" pageId="11" pageNumber="62">Tuyen Quang Province</collectingRegion>
|
||||
: Chiem Hoa District, Phuc Son Village, November-December, Fruhstorfer lgt.), this subspecies possesses similar morphological characters to the nominotypical subspecies. There are only minute morphological differences in the surface of the pronotum between the two subspecies. While
|
||||
<taxonomicName id="89B284D6256BFFC7FE6EFEC32CCDF027" authorityName=": Balthasar" authorityYear="1963" box="[456,672,261,287]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC62347256BFFC7FE6EFEC32FB6F027" box="[456,475,261,287]" italics="true" pageId="11" pageNumber="62">C</emphasis>
|
||||
. (
|
||||
<emphasis id="7CC62347256BFFC7FE55FEC02C69F027" box="[499,516,262,287]" italics="true" pageId="11" pageNumber="62">P</emphasis>
|
||||
.)
|
||||
<emphasis id="7CC62347256BFFC7FDBBFEC32CCDF027" box="[541,672,261,287]" italics="true" pageId="11" pageNumber="62">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7CC62347256BFFC7FD0EFEC32D23F027" box="[680,846,261,287]" italics="true" pageId="11" pageNumber="62">septentrionalis</emphasis>
|
||||
possesses a shallowly and sparsely punctate pronotum, the nominotypical subspecies bears a pronotum with deep and dense punctures. The anterior marginal line is thicker in
|
||||
<taxonomicName id="89B284D6256BFFC7FEABFE8B2F8BF05F" authorityName=": Balthasar" authorityYear="1963" box="[269,486,333,359]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC62347256BFFC7FEABFE8B2F4DF05F" box="[269,288,333,359]" italics="true" pageId="11" pageNumber="62">C</emphasis>
|
||||
. (
|
||||
<emphasis id="7CC62347256BFFC7FE9EFE882F24F05F" box="[312,329,334,359]" italics="true" pageId="11" pageNumber="62">P</emphasis>
|
||||
.)
|
||||
<emphasis id="7CC62347256BFFC7FEC5FE8B2F8BF05F" box="[355,486,333,359]" italics="true" pageId="11" pageNumber="62">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7CC62347256BFFC7FE48FE8B2CF9F05F" box="[494,660,333,359]" italics="true" pageId="11" pageNumber="62">septentrionalis</emphasis>
|
||||
, compared to the nominotypical subspecies. In terms of geographic distribution, the nominotypical subspecies has a wide distribution throughout Asia, while
|
||||
<taxonomicName id="89B284D6256BFFC7FB28FEB72B0BF0B3" authorityName=": Balthasar" authorityYear="1963" box="[1166,1382,369,395]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="punctulatus" subGenus="Paracopris">
|
||||
<emphasis id="7CC62347256BFFC7FB28FEB72ACCF0B3" box="[1166,1185,369,395]" italics="true" pageId="11" pageNumber="62">C</emphasis>
|
||||
. (
|
||||
<emphasis id="7CC62347256BFFC7FB1FFEB42AA7F0B3" box="[1209,1226,370,395]" italics="true" pageId="11" pageNumber="62">P</emphasis>
|
||||
.)
|
||||
<emphasis id="7CC62347256BFFC7FB45FEB72B0BF0B3" box="[1251,1382,369,395]" italics="true" pageId="11" pageNumber="62">punctulatus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7CC62347256BFFC7FAC8FEB42F75F097" italics="true" pageId="11" pageNumber="62">sep- tentrionalis</emphasis>
|
||||
has so far been found in northern
|
||||
<collectingCountry id="36A5BFC5256BFFC7FD3EFE532C9AF097" box="[664,759,405,431]" name="Vietnam" pageId="11" pageNumber="62">Vietnam</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="8C7631B7256BFFC7FCA3FE532D0BF097" box="[773,870,405,431]" country="Vietnam" name="Phu Tho" pageId="11" pageNumber="62">Phu Tho</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="8C7631B7256BFFC7FC3AFE532A58F097" box="[924,1077,405,431]" country="Vietnam" name="Tuyen Quang" pageId="11" pageNumber="62">Tuyen Quang</collectingRegion>
|
||||
Provinces).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
70
data/C6/1B/5F/C61B5FEEF79AF14C0133C929269FCB4E.xml
Normal file
70
data/C6/1B/5F/C61B5FEEF79AF14C0133C929269FCB4E.xml
Normal file
|
|
@ -0,0 +1,70 @@
|
|||
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.81.833" ID-GBIF-Dataset="29a6ff09-6c38-42e2-9681-31637b91cc81" ID-PMC="PMC3088066" ID-Pensoft-Pub="1313-2970-81-39" ID-PubMed="21594160" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-81-39" ModsDocOrigin="ZooKeys 81" ModsDocTitle="Review of the genus Fibuloides Kuznetsov in China (Lepidoptera, Tortricidae, Olethreutinae)" checkinTime="1451250790816" checkinUser="pensoft" docAuthor="Zhang, Aihuan & Li, Houhun" docDate="2011" docId="C61B5FEEF79AF14C0133C929269FCB4E" docLanguage="en" docName="ZooKeys 81: 39-50" docOrigin="ZooKeys 81" docSource="http://dx.doi.org/10.3897/zookeys.81.833" docTitle="Fibuloides modificana Kuznetsov 1997" docType="treatment" docVersion="4" lastPageNumber="45" masterDocId="D55DFFA9FFB3FFF4FFFC963F1D53FFA9" masterDocTitle="Review of the genus Fibuloides Kuznetsov in China (Lepidoptera, Tortricidae, Olethreutinae)" masterLastPageNumber="50" masterPageNumber="39" pageNumber="45" updateTime="1668166707950" updateUser="ExternalLinkService">
|
||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Review of the genus Fibuloides Kuznetsov in China (Lepidoptera, Tortricidae, Olethreutinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Zhang, Aihuan</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Li, Houhun</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>ZooKeys</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2011</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>81</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>39</mods:start>
|
||||
<mods:end>50</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://dx.doi.org/10.3897/zookeys.81.833</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.81.833</mods:identifier>
|
||||
<mods:identifier type="Pensoft-Pub">1313-2970-81-39</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-GBIF-Taxon="159361319" LSID="urn:lsid:plazi:treatment:C61B5FEEF79AF14C0133C929269FCB4E" httpUri="http://treatment.plazi.org/id/C61B5FEEF79AF14C0133C929269FCB4E" lastPageNumber="45" pageId="6" pageNumber="45">
|
||||
<subSubSection pageId="6" pageNumber="45" type="nomenclature">
|
||||
<paragraph pageId="6" pageNumber="45">
|
||||
<taxonomicName authority="Kuznetsov, 1997" authorityName="Kuznetsov" authorityYear="1997" class="Insecta" family="Tortricidae" genus="Fibuloides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fibuloides modificana" order="Lepidoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="modificana">
|
||||
<pageBreakToken pageId="6" pageNumber="45" start="start">Fibuloides</pageBreakToken>
|
||||
modificana Kuznetsov, 1997
|
||||
</taxonomicName>
|
||||
Fig. 817
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="6" pageNumber="45" type="reference_group">
|
||||
<paragraph pageId="6" pageNumber="45">
|
||||
<taxonomicName class="Insecta" family="Tortricidae" genus="Fibuloides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fibuloides modificana" order="Lepidoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="modificana">Fibuloides modificana</taxonomicName>
|
||||
Kuznetsov, 1997: 810. Holotype ♂, TL: Vietnam, deposited in ZMAS.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="6" pageNumber="45" type="material examined">
|
||||
<paragraph pageId="6" pageNumber="45">Material examined.</paragraph>
|
||||
<paragraph pageId="6" pageNumber="45">China, Guangxi Zhuangzu Autonomous Region: 1 ♂, Leye County, 665 m, 24.VII.2004, coll. Jiasheng Xu.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="6" pageNumber="45" type="distribution">
|
||||
<paragraph pageId="6" pageNumber="45">Distribution.</paragraph>
|
||||
<paragraph pageId="6" pageNumber="45">China (Guangxi), Vietnam.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="6" pageNumber="45" type="remarks">
|
||||
<paragraph pageId="6" pageNumber="45">Remarks.</paragraph>
|
||||
<paragraph pageId="6" pageNumber="45">The uncus of this species is broad and distally bifurcate, the valva has a long, sinuate, flattened bristle on the neck, and the cucullus is elongatesubrectangular and distally downcurved. In figure 17 the long flattened bristle is off the inserted hole which is located on the ventral side of the neck instead of on the angle of the sacculus (see arrow in fig. 17).It is new for China.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
54
data/C6/1B/DB/C61BDB6F81A8304F34607BDDED1A0318.xml
Normal file
54
data/C6/1B/DB/C61BDB6F81A8304F34607BDDED1A0318.xml
Normal file
|
|
@ -0,0 +1,54 @@
|
|||
<document id="D1C74AAB1A3509185A933CC71A061245" ID-DOI="http://dx.doi.org/10.3897/BDJ.3.e4567" ID-PMC="PMC4355676" ID-Pensoft-Pub="1314-2828-3-4567" ID-PubMed="25829860" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1314-2828-3-e4567" ModsDocOrigin="Biodiversity Data Journal 3" ModsDocTitle="Updated list of the mosquitoes of Colombia (Diptera: Culicidae)" checkinTime="1451253157155" checkinUser="pensoft" docAuthor="Rozo-Lopez, Paula & Mengual, Ximo" docDate="2015" docId="C61BDB6F81A8304F34607BDDED1A0318" docLanguage="en" docName="BiodivDatJour 3: e4567" docOrigin="Biodiversity Data Journal 3" docSource="http://dx.doi.org/10.3897/BDJ.3.e4567" docTitle="Culex (Culex) inflictus Theobald 1901" docType="treatment" docVersion="4" lastPageNumber="4567" masterDocId="656A44447C0AFFEDFFE0E467FFFEFFD0" masterDocTitle="Updated list of the mosquitoes of Colombia (Diptera: Culicidae)" masterLastPageNumber="4567" masterPageNumber="4567" pageNumber="4567" updateTime="1668122931701" updateUser="ExternalLinkService">
|
||||
<mods:mods id="E60C8BA9CEB12BCFE5F349DD87A7FD9F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="0B6909BF28E84CC935A5AF62CFCAE20F">
|
||||
<mods:title id="C4BD47650E3E1EB57989407DB4CD5F3D">Updated list of the mosquitoes of Colombia (Diptera: Culicidae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="042853884D1A16016C9DC8178D8A0F78" type="personal">
|
||||
<mods:role id="A1DB557FDAC84DBACC7AE37D37F132C7">
|
||||
<mods:roleTerm id="F098E4F921DD8754F725FC02C05630EA">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="379954129043E3E2F8AD76F95752B6D6">Rozo-Lopez, Paula</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="56E1E5F3739D837263190B97C3686697" type="personal">
|
||||
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<mods:namePart id="A6DC874B03CBE9FC1C0C34253FEAD7A0">Mengual, Ximo</mods:namePart>
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<mods:title id="153240476C4FE42217E6B979FF0D95ED">Biodiversity Data Journal</mods:title>
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<mods:part id="BC79D3B54EA1D7ACCB298798E9028C6B">
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<mods:date id="0FAFABECF3FE0EDF95B25E325D649BA3">2015</mods:date>
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<mods:number id="1411BB58A11FFA57D45CA0CE2CE44ACA">3</mods:number>
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<mods:start id="15D684D1F075D32C826E001B5B6CA827">4567</mods:start>
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<mods:url id="81E554A76B577A1A3EF84BA4E7E6EC8A">http://dx.doi.org/10.3897/BDJ.3.e4567</mods:url>
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<mods:identifier id="79725076481A509469490CB2324FD7DD" type="DOI">http://dx.doi.org/10.3897/BDJ.3.e4567</mods:identifier>
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<mods:identifier id="77F2145E264662F9499EFF2839BC8076" type="Pensoft-Pub">1314-2828-3-4567</mods:identifier>
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<treatment id="C61BDB6F81A8304F34607BDDED1A0318" LSID="urn:lsid:plazi:treatment:C61BDB6F81A8304F34607BDDED1A0318" httpUri="http://treatment.plazi.org/id/C61BDB6F81A8304F34607BDDED1A0318" lastPageNumber="4567" pageId="0" pageNumber="4567">
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<subSubSection id="5CDF8EAAD9C0CCDFA787FF7ECB226355" pageId="0" pageNumber="4567" type="nomenclature">
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<paragraph id="26C1AFF29F9A284CF303AAB743F7963E" pageId="0" pageNumber="4567">
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<taxonomicName id="BCDB8D37A774078EF7A2A049C715CAB7" ID-CoL="8BMKK" authority="Theobald, 1901" authorityName="Theobald" authorityYear="1901" class="Insecta" family="Culicidae" genus="Culex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Culex (Culex) inflictus" order="Diptera" pageId="0" pageNumber="4567" phylum="Arthropoda" rank="species" species="inflictus" subGenus="Culex">Culex (Culex) inflictus Theobald, 1901</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="71D5482CDC584A629AD7287064157D85" pageId="0" pageNumber="4567" type="notes">
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<paragraph id="D6F411512403D39B5AB1464C66B984CA" pageId="0" pageNumber="4567">Notes</paragraph>
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<paragraph id="F82C10B10FA1B6A791B96C3334A90FBB" pageId="0" pageNumber="4567">
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<bibRefCitation id="F0BEB00D3DBB0F843AF4BA2913346480" author="Barreto-Reyes, P." journalOrPublisher="Anales de la Sociedad de Biologia de Bogota" pageId="0" pageNumber="4567" pagination="46 - 94" title="Lista de mosquitos de Colombia (Diptera: Culicidae)." volume="7" year="1955">Barreto-Reyes 1955</bibRefCitation>
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</paragraph>
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</subSubSection>
|
||||
</treatment>
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||||
</document>
|
||||
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