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<mods:title id="EBDD912F7B0FB4540F2345CE78167FCB">Ngan mayla gen. et sp. nov, a new blind potamid cave crab from Borneo, Kalimantan (Crustacea: Decapoda: Brachyura: Potamoidea, Potamidae), and three other unidentified cave crabs from the same region</mods:title>
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<mods:namePart id="9B35AAABA5AC5B911DF3C0F5095F1DC8">Guinot, Danièle</mods:namePart>
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<mods:namePart id="FFDCC9DE8A9C07D3066D52CE92652B57">Rodríguez Moreno, Paula A.</mods:namePart>
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@ -126,7 +129,7 @@ and
<paragraph id="8BAE7E6DB961FF94B182FB9DFCBCFAA0" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B182FB9DFE83FBEC" bold="true" box="[199,315,1051,1077]" pageId="3" pageNumber="118">Remarks.</emphasis>
The shape of the three-articulated mandibular palp ending by a single terminal lobe (
<figureCitation id="132A62E8B961FF94B5BAFB9DFAEBFBEC" box="[1279,1363,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="3" pageNumber="118">Fig. 3F</figureCitation>
<figureCitation id="132A62E8B961FF94B5BAFB9DFAEBFBEC" box="[1279,1363,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="3" pageNumber="118">Fig. 3F</figureCitation>
) (
<bibRefCitation id="EF80039CB961FF94B42FFB9DFF6BFB80" author="Bott, R." pageId="3" pageNumber="118" pagination="1 - 338" refId="ref13140" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammes-geschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott 1970</bibRefCitation>
: fig. 133;
@ -168,7 +171,7 @@ Lips
. 2010
</bibRefCitation>
) in East Kalimantan, where the Lubang Gedung Cave (
<figureCitation id="132A62E8B961FF94B004FB49FE2EFB30" box="[321,406,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1567,1591]" captionTargetBox="[201,1386,179,1541]" captionTargetId="figure-23@2.[201,1386,179,1541]" captionTargetPageId="2" captionText="FIGURE 1. A, Borneo Island and Kalimantan; B, Karst of Merabu, East Kalimantan, northernmost limestone massif of the Mangkalihat Peninsula; C, D, Lubang Gedung Cave, two views. Courtesy, J. and B. Lips." pageId="3" pageNumber="118">Fig. 1B</figureCitation>
<figureCitation id="132A62E8B961FF94B004FB49FE2EFB30" box="[321,406,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1567,1591]" captionTargetBox="[201,1386,179,1541]" captionTargetId="figure-23@2.[201,1386,179,1541]" captionTargetPageId="2" captionText="FIGURE 1. A, Borneo Island and Kalimantan; B, Karst of Merabu, East Kalimantan, northernmost limestone massif of the Mangkalihat Peninsula; C, D, Lubang Gedung Cave, two views. Courtesy, J. and B. Lips." figureDoi="http://doi.org/10.5281/zenodo.12731231" httpUri="https://zenodo.org/record/12731231/files/figure.png" pageId="3" pageNumber="118">Fig. 1B</figureCitation>
) is located, is about
<quantity id="4CE9D388B961FF94B3C5FB49FD6DFB30" box="[640,725,1231,1257]" metricMagnitude="5" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="118" unit="km" value="370.0">370 km</quantity>
southeast of the Gunung Mulu National Park. Like
@ -187,7 +190,7 @@ has the exopod of the third maxillipeds without a flagellum. Both species of
<emphasis id="B965A27FB961FF94B594FB75FAFFFAD4" box="[1233,1351,1267,1293]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
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have a similar arrangement of the anterior part of the carapace, a similar front and probably a similar proepistome, a similar ornamentation of the carinate and non-incised anterolateral margin, and the posterolateral portion of carapace with several striae; the rounded vulvae are similar (
<figureCitation id="132A62E8B961FF94B3D8FAD9FD4FFAA0" box="[669,759,1375,1401]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6A</figureCitation>
<figureCitation id="132A62E8B961FF94B3D8FAD9FD4FFAA0" box="[669,759,1375,1401]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BAE7E6DB961FF94B182FA05FBC6F988" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
@ -315,7 +318,7 @@ by the presence of a flagellum on the exopod of the third maxilliped (P.K.L. Ng,
<emphasis id="B965A27FB961FF94B54AF8FDFBDDF84C" box="[1039,1125,1915,1941]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B531F8FDFB74F84C" box="[1140,1228,1915,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6D</figureCitation>
<figureCitation id="132A62E8B961FF94B531F8FDFB74F84C" box="[1140,1228,1915,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6D</figureCitation>
) are smaller, oval, distinctly obliquely positioned on sternite 6, instead of rounded and central in
<taxonomicName id="4C1105EEB961FF94B569F819FB0FF860" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1068,1207,1951,1977]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB961FF94B569F819FB0FF860" box="[1068,1207,1951,1977]" italics="true" pageId="3" pageNumber="118">Ngan mayla</emphasis>
@ -324,13 +327,13 @@ by the presence of a flagellum on the exopod of the third maxilliped (P.K.L. Ng,
<taxonomicNameLabel id="A2561F04B961FF94B587F819FAA7F860" box="[1218,1311,1951,1977]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B961FF94B477F819FA37F860" box="[1330,1423,1951,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6A</figureCitation>
<figureCitation id="132A62E8B961FF94B477F819FA37F860" box="[1330,1423,1951,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6A</figureCitation>
),
<taxonomicName id="4C1105EEB961FF94B1D2F845FEEDF805" authorityName="Holthuis" authorityYear="1979" box="[151,341,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB961FF94B1D2F845FEEDF805" box="[151,341,1987,2013]" italics="true" pageId="3" pageNumber="118">Cerberusa caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B020F845FE03F804" box="[357,443,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6B</figureCitation>
<figureCitation id="132A62E8B961FF94B020F845FE03F804" box="[357,443,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6B</figureCitation>
) and
<taxonomicName id="4C1105EEB961FF94B0BFF845FDE3F804" authorityName="Holthuis" authorityYear="1979" box="[506,603,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB961FF94B0BFF845FDB5F804" box="[506,525,1987,2013]" italics="true" pageId="3" pageNumber="118">C</emphasis>
@ -338,7 +341,7 @@ by the presence of a flagellum on the exopod of the third maxilliped (P.K.L. Ng,
<emphasis id="B965A27FB961FF94B35FF845FDE3F804" box="[538,603,1987,2013]" italics="true" pageId="3" pageNumber="118">tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B32EF845FD7CF804" box="[619,708,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6C</figureCitation>
<figureCitation id="132A62E8B961FF94B32EF845FD7CF804" box="[619,708,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6C</figureCitation>
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</paragraph>
</subSubSection>

165
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<figureCitation id="132A62E8B966FF93B1DAFF66FF57FF23" box="[159,239,224,250]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="4" pageNumber="119">Figs. 2</figureCitation>
,
<figureCitation id="132A62E8B966FF93B1BEFF66FEB2FF23" box="[251,266,224,250]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="4" pageNumber="119">3</figureCitation>
<figureCitation id="132A62E8B966FF93B1BEFF66FEB2FF23" box="[251,266,224,250]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="4" pageNumber="119">3</figureCitation>
,
<figureCitation id="132A62E8B966FF93B052FF59FE84FF20" box="[279,316,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="4" pageNumber="119">6A</figureCitation>
<figureCitation id="132A62E8B966FF93B052FF59FE84FF20" box="[279,316,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="4" pageNumber="119">6A</figureCitation>
,
<figureCitation id="132A62E8B966FF93B002FF66FED3FF23" box="[327,363,224,250]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="4" pageNumber="119">9A</figureCitation>
<figureCitation id="132A62E8B966FF93B002FF66FED3FF23" box="[327,363,224,250]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731258" httpUri="https://zenodo.org/record/12731258/files/figure.png" pageId="4" pageNumber="119">9A</figureCitation>
)
</paragraph>
</subSubSection>
@ -125,7 +128,7 @@ Description of
</emphasis>
Carapace subquadrate in outline, anteriorly rounded, flat, widest in anterior quarter; anterior part curved. Front rather flat, entirely visible in dorsal view, with straight margin although barely concave in the middle, smooth, slightly carinate. Surface of entire carapace appearing smooth; epigastric ridges and postorbital ridges distinct; H-shaped groove in centre of carapace indistinct. Anterolateral margin short, not bidentate or incised, about half as long as posterolateral margin, reaching anteriorly to orbital margin and merging with it, both with granules giving a serrated aspect. Antennule with large basal article. Antenna reduced, very small, with extremely short flagellum, filling entire inner gap of orbit.
</paragraph>
<caption id="DF6E2EE5B966FF93B1D2FA96FA22FA95" pageId="4" pageNumber="119" startId="4.[151,246,1296,1320]" targetBox="[173,1415,634,1272]" targetPageId="4" targetType="figure">
<caption id="DF6E2EE5B966FF93B1D2FA96FA22FA95" ID-DOI="http://doi.org/10.5281/zenodo.12731233" ID-Zenodo-Dep="12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="4" pageNumber="119" startId="4.[151,246,1296,1320]" targetBox="[173,1415,634,1272]" targetPageId="4" targetType="figure">
<paragraph id="8BAE7E6DB966FF93B1D2FA96FA22FA95" blockId="4.[151,1437,1296,1356]" pageId="4" pageNumber="119">
<emphasis id="B965A27FB966FF93B1D2FA96FEB2FAF1" bold="true" box="[151,266,1296,1320]" pageId="4" pageNumber="119">FIGURE 2</emphasis>
.
@ -143,7 +146,7 @@ Carapace subquadrate in outline, anteriorly rounded, flat, widest in anterior qu
<paragraph id="8BAE7E6DB966FF93B182F9F0FC10F949" blockId="4.[151,1437,1402,2040]" box="[199,936,1654,1680]" pageId="4" pageNumber="119">Third maxilliped with well-developed exopod, without flagellum.</paragraph>
<paragraph id="8BAE7E6DB966FF93B182F91CFD07F8B0" blockId="4.[151,1437,1402,2040]" pageId="4" pageNumber="119">Chelipeds equal, remarkably spiny: merus with ventral surface bordered on each margin by acute short teeth (some with corneous tip) and spines, some surounded by short simple setae; carpus inner angle with very long acute tooth bearing several smaller acessory spines, simple setae and spinelike setae; palm with sharp spines on both superior and inferior borders; fingers closing over full length and ending in sharp curved tips, with small spines and short simple setae; superior border of dactylus with some spaced spines; extensor margins of fixed finger and dactylus with corneous teeth and short stiff setae.</paragraph>
<paragraph id="8BAE7E6DB966FF93B182F8F4FE5EF821" blockId="4.[151,1437,1402,2040]" pageId="4" pageNumber="119">Ambulatory legs extremely long and slender; P4 and P5 longest. Merus with distinctly serrated aspect on superior margin, inferior side more finely, corresponding to short acute teeth mixed with simple setae, especially on P2, P3; carpus mostly toothed on inferior side; dactylus very elongated, with numerous longer simple setae on both superior and inferior margins.</paragraph>
<caption id="DF6E2EE5B967FF92B1D2F900FB60F918" pageId="5" pageNumber="120" startId="5.[151,250,1670,1694]" targetBox="[151,1435,181,1646]" targetPageId="5" targetType="figure">
<caption id="DF6E2EE5B967FF92B1D2F900FB60F918" ID-DOI="http://doi.org/10.5281/zenodo.12731237" ID-Zenodo-Dep="12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="5" pageNumber="120" startId="5.[151,250,1670,1694]" targetBox="[151,1435,181,1646]" targetPageId="5" targetType="figure">
<paragraph id="8BAE7E6DB967FF92B1D2F900FB60F918" blockId="5.[151,1436,1670,1730]" pageId="5" pageNumber="120">
<emphasis id="B965A27FB967FF92B1D2F900FEB5F947" bold="true" box="[151,269,1670,1694]" pageId="5" pageNumber="120">FIGURE 3</emphasis>
.
@ -179,11 +182,11 @@ Carapace subquadrate in outline, anteriorly rounded, flat, widest in anterior qu
<taxonomicNameLabel id="A2561F04B964FF91B0B2FF11FD13FF68" box="[503,683,151,177]" pageId="6" pageNumber="121" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
from Lubang Gedung Cave, the pale colouration of the body, the long and slender legs (
<figureCitation id="132A62E8B964FF91B0D3FF3DFE64FF0C" box="[406,476,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="6" pageNumber="121">Fig. 2</figureCitation>
<figureCitation id="132A62E8B964FF91B0D3FF3DFE64FF0C" box="[406,476,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="6" pageNumber="121">Fig. 2</figureCitation>
), the nearly immobile and greatly reduced, tapering eyes (
<figureCitation id="132A62E8B964FF91B52BFF3DFA8CFF0C" box="[1134,1332,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="6" pageNumber="121">Figs. 3A, C, D, F</figureCitation>
<figureCitation id="132A62E8B964FF91B52BFF3DFA8CFF0C" box="[1134,1332,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="6" pageNumber="121">Figs. 3A, C, D, F</figureCitation>
,
<figureCitation id="132A62E8B964FF91B405FF3DFADDFF0C" box="[1344,1381,187,213]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="6" pageNumber="121">9A</figureCitation>
<figureCitation id="132A62E8B964FF91B405FF3DFADDFF0C" box="[1344,1381,187,213]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731258" httpUri="https://zenodo.org/record/12731258/files/figure.png" pageId="6" pageNumber="121">9A</figureCitation>
), the length of which is less than half width of orbit, with a short bullet-shaped stalk and a vestigial unfaceted cornea, devoid of integument pigment except for a trace of pale brown pigment at tip, all are indicative of adaptation to a cavernicolous life making it a true troglobite. It is not known in which part of the cave
<taxonomicName id="4C1105EEB964FF91B525FEA1FB50FE98" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1120,1256,295,321]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB964FF91B525FEA1FB50FE98" box="[1120,1256,295,321]" italics="true" pageId="6" pageNumber="121">Ngan mayla</emphasis>
@ -193,7 +196,7 @@ from Lubang Gedung Cave, the pale colouration of the body, the long and slender
</emphasis>
was observed and collected.
</paragraph>
<caption id="DF6E2EE5B964FF91B1D2F8FAFC65F860" pageId="6" pageNumber="121" startId="6.[151,250,1916,1940]" targetBox="[151,1436,381,1893]" targetPageId="6" targetType="figure">
<caption id="DF6E2EE5B964FF91B1D2F8FAFC65F860" ID-DOI="http://doi.org/10.5281/zenodo.12731243" ID-Zenodo-Dep="12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="6" pageNumber="121" startId="6.[151,250,1916,1940]" targetBox="[151,1436,381,1893]" targetPageId="6" targetType="figure">
<paragraph id="8BAE7E6DB964FF91B1D2F8FAFC65F860" blockId="6.[151,1436,1916,1977]" pageId="6" pageNumber="121">
<emphasis id="B965A27FB964FF91B1D2F8FAFEB4F84D" bold="true" box="[151,268,1916,1940]" pageId="6" pageNumber="121">FIGURE 4</emphasis>
.
@ -204,7 +207,7 @@ was observed and collected.
, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng.
</paragraph>
</caption>
<caption id="DF6E2EE5B965FF90B1D2F897FB5DF897" pageId="7" pageNumber="122" startId="7.[151,250,1809,1833]" targetBox="[164,1423,181,1785]" targetPageId="7" targetType="figure">
<caption id="DF6E2EE5B965FF90B1D2F897FB5DF897" ID-DOI="http://doi.org/10.5281/zenodo.12731250" ID-Zenodo-Dep="12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="7" pageNumber="122" startId="7.[151,250,1809,1833]" targetBox="[164,1423,181,1785]" targetPageId="7" targetType="figure">
<paragraph id="8BAE7E6DB965FF90B1D2F897FB5DF897" blockId="7.[151,1437,1809,1870]" pageId="7" pageNumber="122">
<emphasis id="B965A27FB965FF90B1D2F897FEB5F8F0" bold="true" box="[151,269,1809,1833]" pageId="7" pageNumber="122">FIGURE 5</emphasis>
.
@ -223,9 +226,9 @@ was observed and collected.
<taxonomicNameLabel id="A2561F04B965FF90B00AF8FAFE4CF84F" box="[335,500,1916,1942]" pageId="7" pageNumber="122" rank="species">gen. et sp.nov.</taxonomicNameLabel>
</emphasis>
is immediately distinguished from all the other potamids by its multi-spined chelipeds, which additionally bear short simple setae and spinelike setae (
<figureCitation id="132A62E8B965FF90B22BF819FC78F863" box="[878,960,1951,1978]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="7" pageNumber="122">Figs. 2</figureCitation>
<figureCitation id="132A62E8B965FF90B22BF819FC78F863" box="[878,960,1951,1978]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="7" pageNumber="122">Figs. 2</figureCitation>
,
<figureCitation id="132A62E8B965FF90B28BF826FBACF863" box="[974,1044,1952,1978]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="7" pageNumber="122">3A–E</figureCitation>
<figureCitation id="132A62E8B965FF90B28BF826FBACF863" box="[974,1044,1952,1978]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="7" pageNumber="122">3A–E</figureCitation>
). The troglobitic characteristics of
<taxonomicName id="4C1105EEB965FF90B1D2F842FEA7F804" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,287,1987,2013]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB965FF90B1D2F842FEA7F804" box="[151,287,1987,2013]" italics="true" pageId="7" pageNumber="122">Ngan mayla</emphasis>
@ -305,7 +308,7 @@ in 2006 from another cave of the Gunung Mulu National Park, the Stone Horse Cave
<emphasis id="B965A27FB96AFF9FB0EFFDC1FDD0FDB9" box="[426,616,583,609]" italics="true" pageId="8" pageNumber="123">Cerberusa caeca</emphasis>
</taxonomicName>
prefers to dwell in water. It is blind and long-legged; its carapace is pale yellowish or white (
<figureCitation id="132A62E8B96AFF9FB030FDEDFE01FD5C" box="[373,441,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="8" pageNumber="123">Fig. 4</figureCitation>
<figureCitation id="132A62E8B96AFF9FB030FDEDFE01FD5C" box="[373,441,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="8" pageNumber="123">Fig. 4</figureCitation>
), thus it is fully adapted to living exclusively in the complete dark part of the caves.
</paragraph>
<paragraph id="8BAE7E6DB96AFF9FB182FD09FF67FC84" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="123">
@ -324,7 +327,7 @@ McFarlane
</bibRefCitation>
).
</paragraph>
<caption id="DF6E2EE5B96AFF9FB1D2F912FBC9F92C" pageId="8" pageNumber="123" startId="8.[151,250,1684,1708]" targetBox="[164,1423,885,1661]" targetPageId="8" targetType="figure">
<caption id="DF6E2EE5B96AFF9FB1D2F912FBC9F92C" ID-DOI="http://doi.org/10.5281/zenodo.12731252" ID-Zenodo-Dep="12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="8" pageNumber="123" startId="8.[151,250,1684,1708]" targetBox="[164,1423,885,1661]" targetPageId="8" targetType="figure">
<paragraph id="8BAE7E6DB96AFF9FB1D2F912FBC9F92C" blockId="8.[151,1437,1684,1781]" pageId="8" pageNumber="123">
<emphasis id="B965A27FB96AFF9FB1D2F912FEB6F975" bold="true" box="[151,270,1684,1708]" pageId="8" pageNumber="123">FIGURE 6</emphasis>
. Vulvae: A,
@ -362,7 +365,7 @@ The second species of
<emphasis id="B965A27FB96AFF9FB307F8A5FD1BF8E5" box="[578,675,1826,1853]" italics="true" pageId="8" pageNumber="123">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96AFF9FB3F4F8A4FD4BF8E4" box="[689,755,1826,1853]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="8" pageNumber="123">Fig. 5</figureCitation>
<figureCitation id="132A62E8B96AFF9FB3F4F8A4FD4BF8E4" box="[689,755,1826,1853]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="8" pageNumber="123">Fig. 5</figureCitation>
), was originally found in two caves in the Gunung Api massif of Gunung Mulu National Park: Clearwater Cave (
<quantity id="4CE9D388B96AFF9FB38FF8C0FCBBF8B8" box="[714,771,1862,1889]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="123" unit="m" value="15.0">15 m</quantity>
inside northwest entrance, on stone outside water), with the
@ -420,7 +423,7 @@ cohabit in three caves: Clearwater Cave, Deer Cave and Stone Horse Cave.
<emphasis id="B965A27FB96BFF9EB4C6FEA0FF60FEBA" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
is an amphibious species, with a colourful (blue grey) dorsal surface of carapace and depigmented or lightly coloured (pale lemon-yellow or orange-yellow) legs, with practically unmodified eyes and a distinct rounded cornea containing dark pigment (
<figureCitation id="132A62E8B96BFF9EB0FBFE17FDA1FE72" box="[446,537,401,427]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">Fig. 9C</figureCitation>
<figureCitation id="132A62E8B96BFF9EB0FBFE17FDA1FE72" box="[446,537,401,427]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731258" httpUri="https://zenodo.org/record/12731258/files/figure.png" pageId="9" pageNumber="124">Fig. 9C</figureCitation>
). Living specimens of
<taxonomicName id="4C1105EEB96BFF9EB264FE17FC3CFE72" authorityName="Holthuis" authorityYear="1979" box="[801,900,401,427]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB264FE17FC3CFE72" box="[801,900,401,427]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
@ -444,7 +447,7 @@ McFarlane
had the eyestalk somewhat thinned distally and filling a good part of the orbit, whereas in a female of 16 ×
<quantity id="4CE9D388B96BFF9EB090FDA6FD81FDE3" box="[469,569,544,570]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="9" pageNumber="124" unit="mm" value="21.8">21.8 mm</quantity>
it was less developed, barely thinned distally and occupying a smaller part of the orbit, and with a small, pigmented cornea (
<figureCitation id="132A62E8B96BFF9EB3E4FDC5FD40FD87" box="[673,760,579,606]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">Fig. 9C</figureCitation>
<figureCitation id="132A62E8B96BFF9EB3E4FDC5FD40FD87" box="[673,760,579,606]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731258" httpUri="https://zenodo.org/record/12731258/files/figure.png" pageId="9" pageNumber="124">Fig. 9C</figureCitation>
, male and female).
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182FDEEFCBBFC81" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
@ -461,21 +464,21 @@ deposited at RMNH and published photographs of all of them together with sketche
<emphasis id="B965A27FB96BFF9EB529FD55FB68FD35" box="[1132,1232,723,749]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB5A7FD54FA8BFD34" box="[1250,1331,722,749]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Figs. 4</figureCitation>
<figureCitation id="132A62E8B96BFF9EB5A7FD54FA8BFD34" box="[1250,1331,722,749]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Figs. 4</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB404FD54FADCFD35" box="[1345,1380,722,748]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6B</figureCitation>
<figureCitation id="132A62E8B96BFF9EB404FD54FADCFD35" box="[1345,1380,722,748]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="9" pageNumber="124">6B</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB437FD55FA2CFD34" box="[1394,1428,723,749]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">9B</figureCitation>
<figureCitation id="132A62E8B96BFF9EB437FD55FA2CFD34" box="[1394,1428,723,749]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731258" httpUri="https://zenodo.org/record/12731258/files/figure.png" pageId="9" pageNumber="124">9B</figureCitation>
) and
<taxonomicName id="4C1105EEB96BFF9EB182FD71FE92FCC9" authorityName="Holthuis" authorityYear="1979" box="[199,298,758,785]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB182FD71FE92FCC9" box="[199,298,758,785]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB07FFD70FE32FCC8" box="[314,394,758,785]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Figs. 5</figureCitation>
<figureCitation id="132A62E8B96BFF9EB07FFD70FE32FCC8" box="[314,394,758,785]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="9" pageNumber="124">Figs. 5</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB0D2FD70FE01FCC9" box="[407,441,758,784]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6C</figureCitation>
<figureCitation id="132A62E8B96BFF9EB0D2FD70FE01FCC9" box="[407,441,758,784]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="9" pageNumber="124">6C</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB083FD70FE50FCC9" box="[454,488,758,784]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">9C</figureCitation>
<figureCitation id="132A62E8B96BFF9EB083FD70FE50FCC9" box="[454,488,758,784]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731258" httpUri="https://zenodo.org/record/12731258/files/figure.png" pageId="9" pageNumber="124">9C</figureCitation>
) collected by K. Lim
<emphasis id="B965A27FB96BFF9EB3A4FD71FCA2FCC9" box="[737,794,758,784]" italics="true" pageId="9" pageNumber="124">et al.</emphasis>
in
@ -531,30 +534,30 @@ differs from
<emphasis id="B965A27FB96BFF9EB3EBFBDAFCAAFBAC" box="[686,786,1116,1142]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
mainly by the numerous spines and simple setae on the chelipeds (
<figureCitation id="132A62E8B96BFF9EB057FBF9FEE0FB40" box="[274,344,1151,1177]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
<figureCitation id="132A62E8B96BFF9EB057FBF9FEE0FB40" box="[274,344,1151,1177]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="9" pageNumber="124">Fig. 2</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB020FBF9FE10FB40" box="[357,424,1151,1177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="9" pageNumber="124">3A–E</figureCitation>
<figureCitation id="132A62E8B96BFF9EB020FBF9FE10FB40" box="[357,424,1151,1177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="9" pageNumber="124">3A–E</figureCitation>
) (chelipeds unarmed, without spinelike setae in
<taxonomicName id="4C1105EEB96BFF9EB295FB06FB8BFB40" authorityName="Holthuis" authorityYear="1979" box="[976,1075,1152,1178]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB295FB06FB8BFB40" box="[976,1075,1152,1178]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
:
<figureCitation id="132A62E8B96BFF9EB501FBF9FB66FB43" box="[1092,1246,1151,1178]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A–C, E</figureCitation>
<figureCitation id="132A62E8B96BFF9EB501FBF9FB66FB43" box="[1092,1246,1151,1178]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Fig. 4A–C, E</figureCitation>
, as well as in
<taxonomicName id="4C1105EEB96BFF9EB4C6FB06FF60FB64" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB4C6FB06FF60FB64" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
:
<figureCitation id="132A62E8B96BFF9EB1ADFB25FE95FB67" box="[232,301,1187,1214]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5</figureCitation>
<figureCitation id="132A62E8B96BFF9EB1ADFB25FE95FB67" box="[232,301,1187,1214]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="9" pageNumber="124">Fig. 5</figureCitation>
); by the more elongated, thinner legs (
<figureCitation id="132A62E8B96BFF9EB3A1FB25FC91FB64" box="[740,809,1187,1213]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
<figureCitation id="132A62E8B96BFF9EB3A1FB25FC91FB64" box="[740,809,1187,1213]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="9" pageNumber="124">Fig. 2</figureCitation>
), in particular, the carpus, propodus and dactylus being longer and slender; by the P5 being as long as P4 (P5 shorter than P
<quantity id="4CE9D388B96BFF9EB2C1FB41FC16FB38" box="[900,942,1223,1249]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="124" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C1105EEB96BFF9EB2F1FB41FBADFB39" authorityName="Holthuis" authorityYear="1979" box="[948,1045,1223,1249]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB2F1FB41FBADFB39" box="[948,1045,1223,1249]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
:
<figureCitation id="132A62E8B96BFF9EB566FB41FB01FB38" box="[1059,1209,1223,1249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A, B, E</figureCitation>
<figureCitation id="132A62E8B96BFF9EB566FB41FB01FB38" box="[1059,1209,1223,1249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Fig. 4A, B, E</figureCitation>
); by certain margins of legs being finely serrated and with erect stiff, short setae (smoother, only serrated on inferior margin of propodi in
<taxonomicName id="4C1105EEB96BFF9EB1F3FA89FEA1FAF1" authorityName="Holthuis" authorityYear="1979" box="[182,281,1295,1321]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB1F3FA89FEA1FAF1" box="[182,281,1295,1321]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
@ -564,7 +567,7 @@ mainly by the numerous spines and simple setae on the chelipeds (
<emphasis id="B965A27FB96BFF9EB0F8FA89FD99FAF1" box="[445,545,1294,1321]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
are very long and thin (
<figureCitation id="132A62E8B96BFF9EB27CFA88FC2CFAF0" box="[825,916,1294,1321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5A</figureCitation>
<figureCitation id="132A62E8B96BFF9EB27CFA88FC2CFAF0" box="[825,916,1294,1321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="9" pageNumber="124">Fig. 5A</figureCitation>
), even longer and thinner than those of
<taxonomicName id="4C1105EEB96BFF9EB425FA89FF63FA95" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB425FA89FF63FA95" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
@ -573,7 +576,7 @@ are very long and thin (
<taxonomicNameLabel id="A2561F04B96BFF9EB1A6FAB4FE2BFA95" box="[227,403,1330,1356]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB0E6FAB4FE5FFA95" box="[419,487,1330,1356]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
<figureCitation id="132A62E8B96BFF9EB0E6FAB4FE5FFA95" box="[419,487,1330,1356]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="9" pageNumber="124">Fig. 2</figureCitation>
) (except that P5 is proportionally shorter in
<taxonomicName id="4C1105EEB96BFF9EB299FAB4FB86FA95" authorityName="Holthuis" authorityYear="1979" box="[988,1086,1330,1356]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB299FAB4FB86FA95" box="[988,1086,1330,1356]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
@ -583,7 +586,7 @@ are very long and thin (
<emphasis id="B965A27FB96BFF9EB1D2FAD0FF40FAB6" box="[151,248,1366,1392]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
, which, of the three, is the species with the proportionally shorter legs (
<figureCitation id="132A62E8B96BFF9EB553FAD0FBC8FAA9" box="[1046,1136,1366,1392]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A</figureCitation>
<figureCitation id="132A62E8B96BFF9EB553FAD0FBC8FAA9" box="[1046,1136,1366,1392]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Fig. 4A</figureCitation>
). The serrated margins and numerous stiff short erect bristles of the legs of
<taxonomicName id="4C1105EEB96BFF9EB3EBFAFCFC8EFA4A" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[686,822,1401,1427]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB3EBFAFCFC8EFA4A" box="[686,822,1401,1427]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
@ -592,19 +595,19 @@ are very long and thin (
<taxonomicNameLabel id="A2561F04B96BFF9EB27BFAFFFC54FA4A" box="[830,1004,1401,1427]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB2BEFAFFFB87FA4A" box="[1019,1087,1401,1427]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
<figureCitation id="132A62E8B96BFF9EB2BEFAFFFB87FA4A" box="[1019,1087,1401,1427]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°38’1”N–117°24’14”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731233" httpUri="https://zenodo.org/record/12731233/files/figure.png" pageId="9" pageNumber="124">Fig. 2</figureCitation>
) are somewhat similar to those of
<taxonomicName id="4C1105EEB96BFF9EB1F2FA1BFEA1FA6E" authorityName="Holthuis" authorityYear="1979" box="[183,281,1437,1463]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB1F2FA1BFEA1FA6E" box="[183,281,1437,1463]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB06FFA1BFE3CFA61" box="[298,388,1437,1464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5A</figureCitation>
<figureCitation id="132A62E8B96BFF9EB06FFA1BFE3CFA61" box="[298,388,1437,1464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="9" pageNumber="124">Fig. 5A</figureCitation>
), whereas the legs are smoother in
<taxonomicName id="4C1105EEB96BFF9EB253FA1BFCC1FA6E" authorityName="Holthuis" authorityYear="1979" box="[790,889,1437,1463]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB253FA1BFCC1FA6E" box="[790,889,1437,1463]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB2CFFA1BFC5DFA6E" box="[906,997,1437,1463]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A</figureCitation>
<figureCitation id="132A62E8B96BFF9EB2CFFA1BFC5DFA6E" box="[906,997,1437,1463]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Fig. 4A</figureCitation>
), with only the posterior margin of the propodus bearing spinules. The female thoracic sternum, covered with dense setae in
<taxonomicName id="4C1105EEB96BFF9EB519FA44FB5CFA02" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1116,1252,1473,1499]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB519FA44FB5CFA02" box="[1116,1252,1473,1499]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
@ -613,37 +616,37 @@ are very long and thin (
<taxonomicNameLabel id="A2561F04B96BFF9EB5A9FA47FA24FA02" box="[1260,1436,1473,1499]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB1DAFA62FEB8FA26" box="[159,256,1508,1535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="9" pageNumber="124">Figs. 3E</figureCitation>
<figureCitation id="132A62E8B96BFF9EB1DAFA62FEB8FA26" box="[159,256,1508,1535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="9" pageNumber="124">Figs. 3E</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB049FA62FE8AFA27" box="[268,306,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6A</figureCitation>
<figureCitation id="132A62E8B96BFF9EB049FA62FE8AFA27" box="[268,306,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="9" pageNumber="124">6A</figureCitation>
), is bare in
<taxonomicName id="4C1105EEB96BFF9EB0F1FA63FDADFA27" authorityName="Holthuis" authorityYear="1979" box="[436,533,1509,1535]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB0F1FA63FDADFA27" box="[436,533,1509,1535]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB360FA62FD3EFA26" box="[549,646,1508,1535]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Figs. 4E</figureCitation>
<figureCitation id="132A62E8B96BFF9EB360FA62FD3EFA26" box="[549,646,1508,1535]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Figs. 4E</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB3D7FA62FD0BFA27" box="[658,691,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6B</figureCitation>
<figureCitation id="132A62E8B96BFF9EB3D7FA62FD0BFA27" box="[658,691,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="9" pageNumber="124">6B</figureCitation>
) and
<taxonomicName id="4C1105EEB96BFF9EB3B7FA63FCECFA27" authorityName="Holthuis" authorityYear="1979" box="[754,852,1508,1535]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB3B7FA63FCECFA27" box="[754,852,1508,1535]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB221FA62FC7DFA26" box="[868,965,1508,1535]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Figs. 5E</figureCitation>
<figureCitation id="132A62E8B96BFF9EB221FA62FC7DFA26" box="[868,965,1508,1535]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="9" pageNumber="124">Figs. 5E</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB295FA62FC4DFA27" box="[976,1013,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6C</figureCitation>
<figureCitation id="132A62E8B96BFF9EB295FA62FC4DFA27" box="[976,1013,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="9" pageNumber="124">6C</figureCitation>
). The efferent openings are enormous in
<taxonomicName id="4C1105EEB96BFF9EB1F1F98FFEAEF9FB" authorityName="Holthuis" authorityYear="1979" box="[180,278,1544,1571]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB1F1F98FFEAEF9FB" box="[180,278,1544,1571]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB060F98EFEC5F9FA" box="[293,381,1544,1571]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5B</figureCitation>
<figureCitation id="132A62E8B96BFF9EB060F98EFEC5F9FA" box="[293,381,1544,1571]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731250" httpUri="https://zenodo.org/record/12731250/files/figure.png" pageId="9" pageNumber="124">Fig. 5B</figureCitation>
), normal in
<taxonomicName id="4C1105EEB96BFF9EB340F98FFDDEF9FB" authorityName="Holthuis" authorityYear="1979" box="[517,614,1545,1571]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB340F98FFDDEF9FB" box="[517,614,1545,1571]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB333F98EFD56F9FB" box="[630,750,1544,1570]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4B, D</figureCitation>
<figureCitation id="132A62E8B96BFF9EB333F98EFD56F9FB" box="[630,750,1544,1570]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731243" httpUri="https://zenodo.org/record/12731243/files/figure.png" pageId="9" pageNumber="124">Fig. 4B, D</figureCitation>
) and
<taxonomicName id="4C1105EEB96BFF9EB268F98FFC0DF9FB" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[813,949,1544,1570]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB268F98FFC0DF9FB" box="[813,949,1544,1570]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
@ -652,7 +655,7 @@ are very long and thin (
<taxonomicNameLabel id="A2561F04B96BFF9EB2F9F98EFBD2F9FB" box="[956,1130,1544,1570]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB53FF98EFB49F9FB" box="[1146,1265,1544,1570]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." pageId="9" pageNumber="124">Fig. 3A, F</figureCitation>
<figureCitation id="132A62E8B96BFF9EB53FF98EFB49F9FB" box="[1146,1265,1544,1570]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="9" pageNumber="124">Fig. 3A, F</figureCitation>
).
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182F9AAFAFDF954" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">

89
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@ -66,7 +69,7 @@ Family
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<paragraph id="8BAE7E6DB969FF9CB1D2FC4EFF53FC3B" blockId="11.[151,361,932,994]" box="[151,235,968,994]" pageId="11" pageNumber="126">
(
<figureCitation id="132A62E8B969FF9CB1DAFC4EFF5AFC3B" box="[159,226,968,994]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." pageId="11" pageNumber="126">Fig. 8</figureCitation>
<figureCitation id="132A62E8B969FF9CB1DAFC4EFF5AFC3B" box="[159,226,968,994]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731256" httpUri="https://zenodo.org/record/12731256/files/figure.png" pageId="11" pageNumber="126">Fig. 8</figureCitation>
)
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@ -113,20 +116,20 @@ Cru 5712
sp.
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mentioned before (
<figureCitation id="132A62E8B969FF9CB293FBDEFBE1FBAB" box="[982,1113,1112,1138]" captionStart="FIGURE 7" captionStartId="10.[151,250,1898,1922]" captionTargetBox="[189,1398,381,1875]" captionTargetId="figure-42@10.[189,1398,381,1875]" captionTargetPageId="10" captionText="FIGURE 7. A, Isolapotamon sp., Lubang Bayan Cave, Merabu Karst, East Kalimantan, Borneo, 01°28’56”N, 117°11’32”E, August 2010, photographed in situ in the river preying on barb; crab not collected, probably a trogloxene; B, Isolapotamon sp., juvenile female, 7.8 × 8.9 mm, Lubang Tebot Cave, Kalimantan (East Kalimantan), Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5711. Scale bar: B = 5 mm." pageId="11" pageNumber="126">Fig. 7A, B</figureCitation>
<figureCitation id="132A62E8B969FF9CB293FBDEFBE1FBAB" box="[982,1113,1112,1138]" captionStart="FIGURE 7" captionStartId="10.[151,250,1898,1922]" captionTargetBox="[189,1398,381,1875]" captionTargetId="figure-42@10.[189,1398,381,1875]" captionTargetPageId="10" captionText="FIGURE 7. A, Isolapotamon sp., Lubang Bayan Cave, Merabu Karst, East Kalimantan, Borneo, 01°28’56”N, 117°11’32”E, August 2010, photographed in situ in the river preying on barb; crab not collected, probably a trogloxene; B, Isolapotamon sp., juvenile female, 7.8 × 8.9 mm, Lubang Tebot Cave, Kalimantan (East Kalimantan), Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5711. Scale bar: B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12731254" httpUri="https://zenodo.org/record/12731254/files/figure.png" pageId="11" pageNumber="126">Fig. 7A, B</figureCitation>
), the present specimen is a gecarcinucid as shown by the bilobed last article of the mandibular palp. It is depigmented (only brownish pale coloured body; white pereiopods); the eyes barely fill one-third of the orbit and tend to be ogive-shaped; the cornea is black-pigmented (
<figureCitation id="132A62E8B969FF9CB039FB42FE6CFB07" box="[380,468,1220,1246]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." pageId="11" pageNumber="126">Fig. 8B</figureCitation>
<figureCitation id="132A62E8B969FF9CB039FB42FE6CFB07" box="[380,468,1220,1246]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731256" httpUri="https://zenodo.org/record/12731256/files/figure.png" pageId="11" pageNumber="126">Fig. 8B</figureCitation>
).
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A short diagnosis of the juvenile female follows: carapace dorsal surface gently convex (
<figureCitation id="132A62E8B969FF9CB5E6FB6EFB44FADB" box="[1187,1276,1256,1282]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." pageId="11" pageNumber="126">Fig. 8A</figureCitation>
<figureCitation id="132A62E8B969FF9CB5E6FB6EFB44FADB" box="[1187,1276,1256,1282]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731256" httpUri="https://zenodo.org/record/12731256/files/figure.png" pageId="11" pageNumber="126">Fig. 8A</figureCitation>
); anterolateral margin distinctly convex, with marked indentation, lined with small rounded granules; areas adjacent to antero- and posterolateral margins covered with numerous distinct oblique striae, anteriorly mixed with very short setae; rugose epigastric cristae on two prominences (
<figureCitation id="132A62E8B969FF9CB3E2FAD2FC9BFAB7" box="[679,803,1364,1390]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." pageId="11" pageNumber="126">Fig. 8A, B</figureCitation>
<figureCitation id="132A62E8B969FF9CB3E2FAD2FC9BFAB7" box="[679,803,1364,1390]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731256" httpUri="https://zenodo.org/record/12731256/files/figure.png" pageId="11" pageNumber="126">Fig. 8A, B</figureCitation>
); postorbital cristae well discernible; frontal margin not gradually confluent with supraorbital margin, conversely the both being almost at right angle; a frontal triangle not developed and, instead, a narrow proepistome; posterior margin of epistome undulated; chelipeds (
<figureCitation id="132A62E8B969FF9CB47EFA1AFA2BFA6F" box="[1339,1427,1436,1462]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." pageId="11" pageNumber="126">Fig. 8C</figureCitation>
<figureCitation id="132A62E8B969FF9CB47EFA1AFA2BFA6F" box="[1339,1427,1436,1462]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731256" httpUri="https://zenodo.org/record/12731256/files/figure.png" pageId="11" pageNumber="126">Fig. 8C</figureCitation>
) elongate; chela slender, outer surface covered with small sharp granules; fingers slightly longer than palm, curved distally to corneous tip; ambulatory legs very long and thin, second pair longest (
<figureCitation id="132A62E8B969FF9CB564FA62FB24FA27" box="[1057,1180,1508,1534]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." pageId="11" pageNumber="126">Fig. 8A, D</figureCitation>
<figureCitation id="132A62E8B969FF9CB564FA62FB24FA27" box="[1057,1180,1508,1534]" captionStart="FIGURE 8" captionStartId="12.[151,250,1929,1953]" captionTargetBox="[177,1411,309,1905]" captionTargetId="figure-95@12.[177,1411,309,1905]" captionTargetPageId="12" captionText="FIGURE 8. Gecarcinucid sp., juvenile female, 7.7 × 9.9 mm, Lubang Tebot Cave, Merabu Karst, East Kalimantan, Borneo, 1°34’58”N, 117°21’07”E, 360 m, 09 August 2010, MZB Cru 5712:A, habitus; B, frontal view; C, cheliped; D, ventral view; E, thoracic sternum. Scale bars: A, B, D, E = 5 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731256" httpUri="https://zenodo.org/record/12731256/files/figure.png" pageId="11" pageNumber="126">Fig. 8A, D</figureCitation>
); outer surface rugose; dorsal, ventral margins of meri subcristate, gently serrated; propodus with small sharp granules on margins; dactylus very long, slightly downcurved, with numerous strong, anteriorly directed spines on dorsal, ventral margins, and corneous tip.
</paragraph>
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