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<document id="5A4D655AF624A7DE8F582C9F2B72CB66" ID-DOI="10.11646/zootaxa.5476.1.13" ID-ISSN="1175-5326" ID-ZooBank="9959FB71-BE4C-42E1-AE73-710A58C4168D" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1720374637855" checkinUser="plazi" docAuthor="Guinot, Danièle &amp; Rodríguez Moreno, Paula A." docDate="2024" docId="03B8CF7BB960FF94B1D2F8B8FD69F804" docLanguage="en" docName="zootaxa.5476.1.13.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.13" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ngan Guinot &amp; Moreno, 2024, gen. nov." docType="treatment" docUuid="A88A1642-D045-4558-8005-97A5AC781944" docUuidSource="ZooBank" docVersion="2" lastPageNumber="118" masterDocId="FF81B703B962FF97B145FF86FFB8FFD9" masterDocTitle="Ngan mayla gen. et sp. nov, a new blind potamid cave crab from Borneo, Kalimantan (Crustacea: Decapoda: Brachyura: Potamoidea, Potamidae), and three other unidentified cave crabs from the same region" masterLastPageNumber="137" masterPageNumber="115" pageNumber="117" updateTime="1720772860913" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:title id="DDF5C7D8E6658B81DEF79B7E41E0E073">Ngan mayla gen. et sp. nov, a new blind potamid cave crab from Borneo, Kalimantan (Crustacea: Decapoda: Brachyura: Potamoidea, Potamidae), and three other unidentified cave crabs from the same region</mods:title>
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<mods:namePart id="9B35AAABA5AC5B911DF3C0F5095F1DC8">Guinot, Danièle</mods:namePart>
<mods:affiliation id="BDE42DBA1CA86B80C296D74E056B8465">Institut de Systématique, Evolution, Biodiversité (ISYEB), UMR 7205 CNRS, MNHN, EPHE</mods:affiliation>
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<mods:namePart id="FFDCC9DE8A9C07D3066D52CE92652B57">Rodríguez Moreno, Paula A.</mods:namePart>
<mods:affiliation id="7009822EC808D10DF3BA596770FA275D">Direction Générale Déléguée aux Collections, Muséum national dHistoire naturelle, MNHN,</mods:affiliation>
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<mods:date id="001CB647188D397CC95D03A2067ACA86">2024</mods:date>
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<paragraph id="8BAE7E6DB960FF95B1D2F8B8FEF7F881" blockId="2.[151,953,1854,1916]" box="[151,335,1854,1880]" pageId="2" pageNumber="117">
<heading id="D0E6C901B960FF95B1D2F8B8FEF7F881" bold="true" box="[151,335,1854,1880]" fontSize="11" level="1" pageId="2" pageNumber="117" reason="1">
<emphasis id="B965A27FB960FF95B1D2F8B8FEF7F881" bold="true" box="[151,335,1854,1880]" pageId="2" pageNumber="117">
<taxonomicName id="4C1105EEB960FF95B1D2F8B8FF62F881" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,218,1854,1880]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB960FF95B1D2F8B8FF62F881" bold="true" box="[151,218,1854,1880]" italics="true" pageId="2" pageNumber="117">Ngan</emphasis>
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<taxonomicNameLabel id="A2561F04B960FF95B1A4F8B8FEF7F881" box="[225,335,1854,1880]" pageId="2" pageNumber="117" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B960FF95B1D2F82CFCA5F81D" box="[151,797,1961,1988]" pageId="2" pageNumber="117" type="type_taxon">
<paragraph id="8BAE7E6DB960FF95B1D2F82CFCA5F81D" blockId="2.[151,797,1961,1988]" box="[151,797,1961,1988]" pageId="2" pageNumber="117">
<emphasis id="B965A27FB960FF95B1D2F82CFE93F81A" bold="true" box="[151,299,1961,1988]" pageId="2" pageNumber="117">
<typeStatus id="54AAC0CFB960FF95B1D2F82CFF6AF81D" box="[151,210,1962,1988]" pageId="2" pageNumber="117">Type</typeStatus>
species
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:
<taxonomicName id="4C1105EEB960FF95B07FF82CFE7AF81D" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[314,450,1962,1988]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB960FF95B07FF82CFE7AF81D" box="[314,450,1962,1988]" italics="true" pageId="2" pageNumber="117">Ngan mayla</emphasis>
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<taxonomicNameLabel id="A2561F04B960FF95B08CF82CFD9BF81D" box="[457,547,1962,1988]" pageId="2" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(present designation).
</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B961FF94B1D2FF10FDBBFCE0" pageId="3" pageNumber="118" type="diagnosis">
<paragraph id="8BAE7E6DB961FF94B1D2FF10FDBBFCE0" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B1D2FF10FEB2FF69" bold="true" box="[151,266,150,176]" pageId="3" pageNumber="118">Diagnosis</emphasis>
. Carapace widest anteriorly, narrowing posteriorly, flat; each postorbital and epigastric ridge consist of 1 row of granules, each ridge separated from each other, epigastric ridge transverse, placed mesioposterior to postorbital ridge, postorbital ridge short, oblique, well separated from anterolateral margin; anterolateral margin long, without teeth or epibranchial incision, forming a finely granular, even finely serrated carina, joining orbital margin anteriorly, this carina posteriorly curving onto dorsal surface of carapace; posterolateral margin longer, straight, with transverse curved ridges formed by rows of granules; front flat but slightly concave medially, directed somewhat down, with very small frontal triangle (as enlarged proepistome); cervical groove not visible in median area; orbits large, orbital margin entire. Eyes strongly reduced, filling less than half the orbit, bullet-shaped, with only tiny pale brown pigment. Antenna very reduced. Mandibular palp three-articulated; last article simple, not bifurcated, oval. Third maxilipeds filling most of oral cavity; exopod without flagellum. Epistome with straight anterior margin; its concave posterior margin ending medially in triangular apex and, at either side of this triangle, two concavities separated by sharp point, one small (for reception of the palp of third maxilliped), the other rounded (forming anterior margin of efferent openings). Efferent openings very large, circular, reaching lateral margin of oral field. Chelipeds remarkably spiny, with numerous acute teeth and spines (some horn-tipped), and short simple setae or spine-like setae on all articles, may be in groups of two or three, some teeth bearing short setae at tips; fingers closing over full length and ending in sharp curved tips, and bearing short horny spines and simple setae. Pereiopods extremely long and thin, slender with short teeth and simple setae; dactyli with numerous longer simple setae on both sides. Vulvae on thoracic sternite 6, at margin with sternite 5, with wide sternal cover, and occupying two-thirds of width of sternite 6.
</paragraph>
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<subSubSection id="C30B2DE6B961FF94B182FCC5FC67FBC8" pageId="3" pageNumber="118" type="etymology">
<paragraph id="8BAE7E6DB961FF94B182FCC5FC67FBC8" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B182FCC5FEF6FC84" bold="true" box="[199,334,835,861]" pageId="3" pageNumber="118">Etymology.</emphasis>
The genus is dedicated to the memory of
<taxonomicName id="4C1105EEB961FF94B273FCC5FC63FC84" authority="Kee NG" authorityName="Kee NG" box="[822,987,835,861]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">Ngan Kee NG</taxonomicName>
(1966
<emphasis id="B965A27FB961FF94B563FCC5FB8CFC84" bold="true" box="[1062,1076,835,861]" pageId="3" pageNumber="118"></emphasis>
2022). She leaves a void in our scientific community, where she had a special place. She was more than a colleague, a very dear friend. She liked to stay in our Muséum, in Paris, and among her last works are those done here in winter over the last decade, notably the
<emphasis id="B965A27FB961FF94B185FC29FB0BFC11" box="[192,1203,943,969]" italics="true" pageId="3" pageNumber="118">
Annotated type-catalogue of
<taxonomicName id="4C1105EEB961FF94B344FC29FDC1FC10" box="[513,633,943,969]" class="Reptilia" family="Colubridae" genus="Brachyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="118" phylum="Chordata" rank="genus">Brachyura</taxonomicName>
of the Muséum national dHistoire naturelle, Paris
</emphasis>
, the first one dealing with
<taxonomicName id="4C1105EEB961FF94B197FC55FEDEFC34" authorityName="H. Milne Edwards" authorityYear="1837" box="[210,358,979,1005]" class="Malacostraca" family="Gecarcinidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Gecarcinidae</taxonomicName>
and
<taxonomicName id="4C1105EEB961FF94B0E5FC55FCADFC34" authority="(N. K. Ng et al. 2019)" baseAuthorityName="N. K. Ng" baseAuthorityYear="2019" box="[416,789,979,1005]" class="Malacostraca" family="Grapsidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">
Grapsidae (N.K.
<bibRefCitation id="EF80039CB961FF94B326FC55FCB5FC34" author="Ng, N. K. &amp; Rodriguez Moreno, P. A. &amp; Naruse, T. &amp; Guinot, D. &amp; Mollaret, N." box="[611,781,979,1005]" pageId="3" pageNumber="118" pagination="91 - 130" refId="ref15859" refString="Ng, N. K., Rodriguez Moreno, P. A., Naruse, T., Guinot, D. &amp; Mollaret, N. (2019). Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Museum national d'Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema, 41 (7), 91 - 130. https: // doi. org / 10.5252 / zoosystema 2019 v 41 a 7" type="journal article" year="2019">
Ng
<emphasis id="B965A27FB961FF94B3CBFC55FD79FC34" box="[654,705,979,1005]" italics="true" pageId="3" pageNumber="118">et al</emphasis>
. 2019
</bibRefCitation>
)
</taxonomicName>
, then
<taxonomicName id="4C1105EEB961FF94B21EFC55FC64FC34" authorityName="Dana" authorityYear="1851" box="[859,988,979,1005]" class="Malacostraca" family="Plagusiidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Plagusiidae</taxonomicName>
and
<taxonomicName id="4C1105EEB961FF94B552FC55FB30FC34" authorityName="Stevcic" authorityYear="2005" box="[1047,1160,979,1005]" class="Malacostraca" family="Percnidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Percnidae</taxonomicName>
, but the last one, on the
<taxonomicName id="4C1105EEB961FF94B1D2FC71FEB4FBC8" authorityName="H. Milne Edwards" authorityYear="1853" box="[151,268,1015,1041]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Varunidae</taxonomicName>
, her favourite group, has not been completed. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B961FF94B182FB9DFD69F804" pageId="3" pageNumber="118" type="discussion">
<paragraph id="8BAE7E6DB961FF94B182FB9DFCBCFAA0" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B182FB9DFE83FBEC" bold="true" box="[199,315,1051,1077]" pageId="3" pageNumber="118">Remarks.</emphasis>
The shape of the three-articulated mandibular palp ending by a single terminal lobe (
<figureCitation id="132A62E8B961FF94B5BAFB9DFAEBFBEC" box="[1279,1363,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="3" pageNumber="118">Fig. 3F</figureCitation>
) (
<bibRefCitation id="EF80039CB961FF94B42FFB9DFF6BFB80" author="Bott, R." pageId="3" pageNumber="118" pagination="1 - 338" refId="ref13140" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammes-geschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott 1970</bibRefCitation>
: fig. 133;
<bibRefCitation id="EF80039CB961FF94B000FBB9FE13FB80" author="Ng, P. K. L." box="[325,427,1087,1113]" pageId="3" pageNumber="118" refId="ref16006" refString="Ng, P. K. L. (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, viii + 156 pp., figs. 1 - 63, 4 colour pls." type="book" year="1988">Ng 1988</bibRefCitation>
: fig. 2A;
<bibRefCitation id="EF80039CB961FF94B350FBB9FDC3FB80" author="Ng, P. K. L." box="[533,635,1087,1113]" pageId="3" pageNumber="118" pagination="311 - 336" refId="ref16224" refString="Ng, P. K. L. (2004) Crustacea: Decapoda, Brachyura. Freshwater Invertebrates of the Malaysian Region. In: Yule, C. M. &amp; Sen, Y. H. (Eds.), Freshwater Invertebrates of the Malaysian Region. Academy of Science Malaysia, Kuala Lumpur, pp. 311 - 336." type="book chapter" year="2004">Ng 2004</bibRefCitation>
: fig. 5I, J) indicates that
<taxonomicName id="4C1105EEB961FF94B2D7FBB9FC77FB81" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[914,975,1087,1112]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B2D7FBB9FC77FB81" box="[914,975,1087,1112]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B293FBB9FBF8FB80" bold="true" box="[982,1088,1087,1113]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B293FBB9FBF8FB80" box="[982,1088,1087,1113]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C1105EEB961FF94B5A8FBB9FADFFB80" authorityName="Ortmann" authorityYear="1896" box="[1261,1383,1087,1113]" class="Malacostraca" family="Potamidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Potamidae</taxonomicName>
. The closest genus is
<taxonomicName id="4C1105EEB961FF94B009FBE5FDC8FBA4" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[332,624,1123,1149]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B009FBE5FE7AFBA4" box="[332,450,1123,1149]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
<bibRefCitation id="EF80039CB961FF94B08FFBE5FDC8FBA4" author="Holthuis, L. B." box="[458,624,1123,1149]" pageId="3" pageNumber="118" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
that consists of two species, both from the Gunung Mulu National Park, north
<collectingCountry id="F3063EFDB961FF94B19DFB01FE83FB78" box="[216,315,1159,1185]" name="Malaysia" pageId="3" pageNumber="118">Sarawak</collectingCountry>
, the Malaysian part of Borneo and close to the southern border of
<collectingCountry id="F3063EFDB961FF94B54EFB01FBE4FB78" box="[1035,1116,1159,1185]" name="Brunei" pageId="3" pageNumber="118">Brunei</collectingCountry>
:
<taxonomicName id="4C1105EEB961FF94B521FB01FAD6FB78" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[1124,1390,1159,1185]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB961FF94B521FB01FB7CFB79" box="[1124,1220,1159,1185]" italics="true" pageId="3" pageNumber="118">C. caeca</emphasis>
<bibRefCitation id="EF80039CB961FF94B58FFB01FAD6FB78" author="Holthuis, L. B." box="[1226,1390,1159,1185]" pageId="3" pageNumber="118" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1105EEB961FF94B1D2FB2DFE11FB1C" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[151,425,1195,1221]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB961FF94B1D2FB2DFF41FB1C" box="[151,249,1195,1221]" italics="true" pageId="3" pageNumber="118">C. tipula</emphasis>
<bibRefCitation id="EF80039CB961FF94B045FB2DFE11FB1C" author="Holthuis, L. B." box="[256,425,1195,1221]" pageId="3" pageNumber="118" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
. The area explored by Lips team (
<bibRefCitation id="EF80039CB961FF94B26BFB2DFC58FB1C" author="Lips, B. &amp; Lips, J. &amp; Robert, X. &amp; Senecal, P. &amp; Placido, C. &amp; Geneau, A." box="[814,992,1195,1221]" pageId="3" pageNumber="118" refId="ref15541" refString="Lips, B., Lips, J., Robert, X., Senecal, P., Placido, C. &amp; Geneau, A. (2010) Borneo 2010 Kalimantan (Indonesie) 28 juillet au 30 aout 2010. Groupe Speleologique Vulcain. Federation Francaise de Speleologie, Lyon, 66 pp." type="book" year="2010">
Lips
<emphasis id="B965A27FB961FF94B222FB2DFC20FB1C" box="[871,920,1195,1221]" italics="true" pageId="3" pageNumber="118">et al</emphasis>
. 2010
</bibRefCitation>
) in East Kalimantan, where the Lubang Gedung Cave (
<figureCitation id="132A62E8B961FF94B004FB49FE2EFB30" box="[321,406,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1567,1591]" captionTargetBox="[201,1386,179,1541]" captionTargetId="figure-23@2.[201,1386,179,1541]" captionTargetPageId="2" captionText="FIGURE 1. A, Borneo Island and Kalimantan; B, Karst of Merabu, East Kalimantan, northernmost limestone massif of the Mangkalihat Peninsula; C, D, Lubang Gedung Cave, two views. Courtesy, J. and B. Lips." pageId="3" pageNumber="118">Fig. 1B</figureCitation>
) is located, is about
<quantity id="4CE9D388B961FF94B3C5FB49FD6DFB30" box="[640,725,1231,1257]" metricMagnitude="5" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="118" unit="km" value="370.0">370 km</quantity>
southeast of the Gunung Mulu National Park. Like
<taxonomicName id="4C1105EEB961FF94B45AFB49FA2DFB30" authorityName="Holthuis" authorityYear="1979" box="[1311,1429,1231,1257]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B45AFB49FA2DFB30" box="[1311,1429,1231,1257]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1105EEB961FF94B1D2FB75FF6CFAD5" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,212,1267,1292]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B1D2FB75FF6CFAD5" box="[151,212,1267,1292]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B19BFB75FEF2FAD4" bold="true" box="[222,330,1267,1293]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B19BFB75FEF2FAD4" box="[222,330,1267,1293]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has the exopod of the third maxillipeds without a flagellum. Both species of
<taxonomicName id="4C1105EEB961FF94B594FB75FAFFFAD4" authorityName="Holthuis" authorityYear="1979" box="[1233,1351,1267,1293]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B594FB75FAFFFAD4" box="[1233,1351,1267,1293]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
have a similar arrangement of the anterior part of the carapace, a similar front and probably a similar proepistome, a similar ornamentation of the carinate and non-incised anterolateral margin, and the posterolateral portion of carapace with several striae; the rounded vulvae are similar (
<figureCitation id="132A62E8B961FF94B3D8FAD9FD4FFAA0" box="[669,759,1375,1401]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BAE7E6DB961FF94B182FA05FBC6F988" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
Differences between
<taxonomicName id="4C1105EEB961FF94B0FDFA05FE4DFA45" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[440,501,1411,1436]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B0FDFA05FE4DFA45" box="[440,501,1411,1436]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B0B8FA05FDD0FA44" bold="true" box="[509,616,1411,1437]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B0B8FA05FDD0FA44" box="[509,616,1411,1437]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1105EEB961FF94B3E7FA05FCA0FA44" authorityName="Holthuis" authorityYear="1979" box="[674,792,1411,1437]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B3E7FA05FCA0FA44" box="[674,792,1411,1437]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
concern the shape of the carapace, subquadrate in
<taxonomicName id="4C1105EEB961FF94B425FA05FA25FA45" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1376,1437,1411,1436]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B425FA05FA25FA45" box="[1376,1437,1411,1436]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2561F04B961FF94B1D2FA21FF44FA18" box="[151,252,1447,1473]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
(trapezoidal in
<taxonomicName id="4C1105EEB961FF94B0ECFA21FDA7FA18" authorityName="Holthuis" authorityYear="1979" box="[425,543,1447,1473]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B0ECFA21FDA7FA18" box="[425,543,1447,1473]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
), the postorbital and epigastric ridges marked in
<taxonomicName id="4C1105EEB961FF94B57EFA21FBC0FA19" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1083,1144,1447,1472]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B57EFA21FBC0FA19" box="[1083,1144,1447,1472]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B53BFA21FB5EFA18" bold="true" box="[1150,1254,1447,1473]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B53BFA21FB5EFA18" box="[1150,1254,1447,1473]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(inconspicuous, even absent or weak in
<taxonomicName id="4C1105EEB961FF94B0E2FA4DFDA5FA3C" authorityName="Holthuis" authorityYear="1979" box="[423,541,1483,1509]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B0E2FA4DFDA5FA3C" box="[423,541,1483,1509]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
), the chelipeds with numerous spines and setae in
<taxonomicName id="4C1105EEB961FF94B521FA4DFB19FA3D" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1124,1185,1483,1508]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B521FA4DFB19FA3D" box="[1124,1185,1483,1508]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B5ECFA4DFAACFA3C" bold="true" box="[1193,1300,1483,1509]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B5ECFA4DFAACFA3C" box="[1193,1300,1483,1509]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(practically absent in
<taxonomicName id="4C1105EEB961FF94B046FA69FEC1F9D0" authorityName="Holthuis" authorityYear="1979" box="[259,377,1519,1545]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B046FA69FEC1F9D0" box="[259,377,1519,1545]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
). In
<taxonomicName id="4C1105EEB961FF94B0F5FA69FD80F9D0" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[432,568,1519,1545]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB961FF94B0F5FA69FD80F9D0" box="[432,568,1519,1545]" italics="true" pageId="3" pageNumber="118">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B37AFA69FD21F9D0" bold="true" box="[575,665,1519,1545]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B37AFA69FD21F9D0" box="[575,665,1519,1545]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the presence on the chelipeds of horny spines accompanied by setae and of simple setae without setules (spinelike setae), which are usually chemoreceptive (
<bibRefCitation id="EF80039CB961FF94B5D4F995FA97F9F4" author="Watling, L." box="[1169,1327,1555,1581]" pageId="3" pageNumber="118" refId="ref17629" refString="Watling, L. (1989) A classification system for crustacean setae based on the homology concept. In: Felgenhauer, B., Watling, L. &amp; Thistle, A. B. (Eds.), Functional morphology of feeding and grooming in Crustacea. Crustacean Issues. Vol. 6. A. A. Balkema, Rotterdam, pp. 15 - 26. https: // doi. org / 10.1201 / 9781003079354 - 2" type="book" year="1989">Watling 1989</bibRefCitation>
;
<bibRefCitation id="EF80039CB961FF94B478F995FE8AF988" author="Garm, A. &amp; Watling, L." pageId="3" pageNumber="118" refId="ref13976" refString="Garm, A. &amp; Watling, L. (2013) The crustacean integument: setae, setules, and other ornamentation. In: Watling, L. &amp; Thiel, M. (Eds.), Functional morphology and diversity. The natural history of the Crustacea. Vol. 1. Oxford University Press, New York, New York, pp. 167 - 198. https: // doi. org / 10.1093 / acprof: osobl / 9780195398038.003.0006" type="book" year="2013">Garm &amp; Watling 2013</bibRefCitation>
), is probably the sign of a high degree of troglomorphism (see Discussion).
</paragraph>
<paragraph id="8BAE7E6DB961FF94B182F9DDFD69F804" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<taxonomicName id="4C1105EEB961FF94B182F9DDFE29F9AC" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[199,401,1626,1653]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B182F9DDFEA5F9AC" box="[199,285,1627,1653]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
<bibRefCitation id="EF80039CB961FF94B060F9DDFE29F9AC" author="Ng, P. K. L." box="[293,401,1626,1653]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
, an unmodified potamid genus from Borneo, not cavernicolous but probably semiterrestrial and known by two species,
<taxonomicName id="4C1105EEB961FF94B096F9F9FD53F940" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[467,747,1662,1689]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="aethes">
<emphasis id="B965A27FB961FF94B096F9F9FDC1F940" box="[467,633,1663,1689]" italics="true" pageId="3" pageNumber="118">Ibanum aethes</emphasis>
<bibRefCitation id="EF80039CB961FF94B3C4F9F9FD53F940" author="Ng, P. K. L." box="[641,747,1662,1689]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF80039CB961FF94B3BEF9F9FCDBF940" author="Ng, P. K. L." box="[763,867,1662,1689]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng 1995</bibRefCitation>
: figs. 4
<emphasis id="B965A27FB961FF94B2F2F9F9FC7DF940" bold="true" box="[951,965,1663,1689]" pageId="3" pageNumber="118"></emphasis>
6), the
<typeStatus id="54AAC0CFB961FF94B553F9F9FBFEF940" box="[1046,1094,1663,1689]" pageId="3" pageNumber="118">type</typeStatus>
species from Lanjak-Entimau Wildlife Sanctuary,
<collectingCountry id="F3063EFDB961FF94B03CF925FE64F964" box="[377,476,1699,1725]" name="Malaysia" pageId="3" pageNumber="118">Sarawak</collectingCountry>
, and
<taxonomicName id="4C1105EEB961FF94B35FF925FCC8F964" authority="(De Man, 1899)" baseAuthorityName="De Man" baseAuthorityYear="1899" box="[538,880,1699,1725]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="bicristatum">
<emphasis id="B965A27FB961FF94B35FF925FD0AF964" box="[538,690,1699,1725]" italics="true" pageId="3" pageNumber="118">I. bicristatum</emphasis>
(De Man, 1899)
</taxonomicName>
(
<bibRefCitation id="EF80039CB961FF94B2C5F925FC51F964" author="Ng, P. K. L." box="[896,1001,1698,1725]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng 1995</bibRefCitation>
: figs. 1
<emphasis id="B965A27FB961FF94B579F925FBF2F964" bold="true" box="[1084,1098,1699,1725]" pageId="3" pageNumber="118"></emphasis>
3), from Kapuas, Kalimantan, is close phylogenetically to
<taxonomicName id="4C1105EEB961FF94B095F941FDB5F939" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[464,525,1735,1760]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B095F941FDB5F939" box="[464,525,1735,1760]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B351F941FDC5F938" bold="true" box="[532,637,1735,1761]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B351F941FDC5F938" box="[532,637,1735,1761]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1105EEB961FF94B3F6F941FC91F938" authorityName="Holthuis" authorityYear="1979" box="[691,809,1735,1761]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B3F6F941FC91F938" box="[691,809,1735,1761]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
(P.K.L. Ng, pers. comm.). The three genera share many similarities: carapace widest anteriorly; antero-lateral margin short, arcuate, carinate, granulated to gently serrated, without teeth or epibranchial incision, with the carina curving onto the carapace dorsal surface; front flat and rather straight; and posterolateral margins long, with transverse ridges. Such as
<taxonomicName id="4C1105EEB961FF94B289F8B5FBB1F895" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[972,1033,1843,1868]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B289F8B5FBB1F895" box="[972,1033,1843,1868]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B555F8B5FBC1F894" bold="true" box="[1040,1145,1843,1869]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B555F8B5FBC1F894" box="[1040,1145,1843,1869]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C1105EEB961FF94B5CDF8B5FB66F894" authorityName="Ng" authorityYear="1995" box="[1160,1246,1843,1869]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B5CDF8B5FB66F894" box="[1160,1246,1843,1869]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
has well defined postorbital cristae.
<taxonomicName id="4C1105EEB961FF94B02AF8D1FE7DF8A8" authorityName="Ng" authorityYear="1995" box="[367,453,1879,1905]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B02AF8D1FE7DF8A8" box="[367,453,1879,1905]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
distinguishes from both
<taxonomicName id="4C1105EEB961FF94B39AF8D1FCA4F8A9" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[735,796,1879,1904]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B39AF8D1FCA4F8A9" box="[735,796,1879,1904]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B261F8D1FC36F8A8" bold="true" box="[804,910,1879,1905]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B261F8D1FC36F8A8" box="[804,910,1879,1905]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1105EEB961FF94B282F8D1FB85F8A8" authorityName="Holthuis" authorityYear="1979" box="[967,1085,1879,1905]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B282F8D1FB85F8A8" box="[967,1085,1879,1905]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
by the presence of a flagellum on the exopod of the third maxilliped (P.K.L. Ng, pers. comm.). The vulvae of
<taxonomicName id="4C1105EEB961FF94B54AF8FDFBDDF84C" authorityName="Ng" authorityYear="1995" box="[1039,1125,1915,1941]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B54AF8FDFBDDF84C" box="[1039,1125,1915,1941]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B531F8FDFB74F84C" box="[1140,1228,1915,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6D</figureCitation>
) are smaller, oval, distinctly obliquely positioned on sternite 6, instead of rounded and central in
<taxonomicName id="4C1105EEB961FF94B569F819FB0FF860" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1068,1207,1951,1977]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB961FF94B569F819FB0FF860" box="[1068,1207,1951,1977]" italics="true" pageId="3" pageNumber="118">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B587F819FAA7F860" bold="true" box="[1218,1311,1951,1977]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B587F819FAA7F860" box="[1218,1311,1951,1977]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B961FF94B477F819FA37F860" box="[1330,1423,1951,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6A</figureCitation>
),
<taxonomicName id="4C1105EEB961FF94B1D2F845FEEDF805" authorityName="Holthuis" authorityYear="1979" box="[151,341,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB961FF94B1D2F845FEEDF805" box="[151,341,1987,2013]" italics="true" pageId="3" pageNumber="118">Cerberusa caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B020F845FE03F804" box="[357,443,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6B</figureCitation>
) and
<taxonomicName id="4C1105EEB961FF94B0BFF845FDE3F804" authorityName="Holthuis" authorityYear="1979" box="[506,603,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB961FF94B0BFF845FDB5F804" box="[506,525,1987,2013]" italics="true" pageId="3" pageNumber="118">C</emphasis>
.
<emphasis id="B965A27FB961FF94B35FF845FDE3F804" box="[538,603,1987,2013]" italics="true" pageId="3" pageNumber="118">tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B32EF845FD7CF804" box="[619,708,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="3" pageNumber="118">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,743 @@
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<mods:title id="CEBD78D3C2C41712B8008C58D77C0BE1">Ngan mayla gen. et sp. nov, a new blind potamid cave crab from Borneo, Kalimantan (Crustacea: Decapoda: Brachyura: Potamoidea, Potamidae), and three other unidentified cave crabs from the same region</mods:title>
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<paragraph id="8BAE7E6DB966FF93B1D2FF1EFE28FF6B" blockId="4.[151,975,151,250]" box="[151,400,151,178]" pageId="4" pageNumber="119">
<heading id="D0E6C901B966FF93B1D2FF1EFE28FF6B" bold="true" box="[151,400,151,178]" fontSize="11" level="1" pageId="4" pageNumber="119" reason="1">
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<taxonomicName id="4C1105EEB966FF93B1D2FF1EFE92FF6B" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,298,151,178]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB966FF93B1D2FF1EFE92FF6B" bold="true" box="[151,298,151,178]" italics="true" pageId="4" pageNumber="119">Ngan mayla</emphasis>
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<taxonomicNameLabel id="A2561F04B966FF93B077FF1EFE28FF6B" box="[306,400,152,178]" pageId="4" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
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</subSubSection>
<subSubSection id="C30B2DE6B966FF93B1D2FF66FECAFF23" box="[151,370,223,250]" pageId="4" pageNumber="119" type="description">
<paragraph id="8BAE7E6DB966FF93B1D2FF66FECAFF23" blockId="4.[151,975,151,250]" box="[151,370,223,250]" pageId="4" pageNumber="119">
(
<figureCitation id="132A62E8B966FF93B1DAFF66FF57FF23" box="[159,239,224,250]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="4" pageNumber="119">Figs. 2</figureCitation>
,
<figureCitation id="132A62E8B966FF93B1BEFF66FEB2FF23" box="[251,266,224,250]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="4" pageNumber="119">3</figureCitation>
,
<figureCitation id="132A62E8B966FF93B052FF59FE84FF20" box="[279,316,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="4" pageNumber="119">6A</figureCitation>
,
<figureCitation id="132A62E8B966FF93B002FF66FED3FF23" box="[327,363,224,250]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="4" pageNumber="119">9A</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BAE7E6DB966FF93B1D2FEAEFBC7FEBF" blockId="4.[151,1436,295,610]" pageId="4" pageNumber="119">
<emphasis id="B965A27FB966FF93B1D2FEAEFEF0FE98" bold="true" box="[151,328,295,322]" pageId="4" pageNumber="119">Type material.</emphasis>
<materialsCitation id="3B797430B966FF93B017FEAEFBC7FEBF" collectingDate="2010-08-18" collectionCode="MZB" country="Indonesia" county="Borneo" elevation="850" latitude="1.6336111" location="Lubang Gedung Cave" longLatPrecision="21" longitude="117.40389" municipality="Panaan" pageId="4" pageNumber="119" specimenCount="1" stateProvince="Kalimantan Utara" typeStatus="holotype">
<emphasis id="B965A27FB966FF93B017FEAEFE01FE9B" box="[338,441,296,322]" italics="true" pageId="4" pageNumber="119">
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: female, 12.2 ×
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</date>
,
<collectionCode id="ED00E6A8B966FF93B28EFECAFBB0FEBF" box="[971,1032,332,358]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="119" type="Museum">MZB</collectionCode>
Cru 5710.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B966FF92B182FEE9FD60F8AF" lastPageId="5" lastPageNumber="120" pageId="4" pageNumber="119" type="description">
<paragraph id="8BAE7E6DB966FF93B182FEE9FD15FDBB" blockId="4.[151,1436,295,610]" pageId="4" pageNumber="119">
<emphasis id="B965A27FB966FF93B182FEE9FE5EFE53" bold="true" box="[199,486,367,394]" pageId="4" pageNumber="119">
Description of
<typeStatus id="54AAC0CFB966FF93B03DFEF6FE59FE53" box="[376,481,368,394]" pageId="4" pageNumber="119" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace subquadrate in outline, anteriorly rounded, flat, widest in anterior quarter; anterior part curved. Front rather flat, entirely visible in dorsal view, with straight margin although barely concave in the middle, smooth, slightly carinate. Surface of entire carapace appearing smooth; epigastric ridges and postorbital ridges distinct; H-shaped groove in centre of carapace indistinct. Anterolateral margin short, not bidentate or incised, about half as long as posterolateral margin, reaching anteriorly to orbital margin and merging with it, both with granules giving a serrated aspect. Antennule with large basal article. Antenna reduced, very small, with extremely short flagellum, filling entire inner gap of orbit.
</paragraph>
<caption id="DF6E2EE5B966FF93B1D2FA96FA22FA95" pageId="4" pageNumber="119" startId="4.[151,246,1296,1320]" targetBox="[173,1415,634,1272]" targetPageId="4" targetType="figure">
<paragraph id="8BAE7E6DB966FF93B1D2FA96FA22FA95" blockId="4.[151,1437,1296,1356]" pageId="4" pageNumber="119">
<emphasis id="B965A27FB966FF93B1D2FA96FEB2FAF1" bold="true" box="[151,266,1296,1320]" pageId="4" pageNumber="119">FIGURE 2</emphasis>
.
<taxonomicName id="4C1105EEB966FF93B05CFA97FE29FAF1" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[281,401,1296,1320]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB966FF93B05CFA97FE29FAF1" box="[281,401,1296,1320]" italics="true" pageId="4" pageNumber="119">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB966FF93B0DFFA96FD80FAF1" bold="true" box="[410,568,1296,1320]" pageId="4" pageNumber="119">
<taxonomicNameLabel id="A2561F04B966FF93B0DFFA96FD80FAF1" box="[410,568,1296,1320]" pageId="4" pageNumber="119" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8BAE7E6DB966FF93B182FAFCFDD1F991" blockId="4.[151,1437,1402,2040]" pageId="4" pageNumber="119">Orbit large. Eyes strongly reduced, not mobile, eyestalk with few short setae and filling less than half of orbit, bullet-shaped, and with only a tiny trace of light brown pigment, so that the species is blind. Frontal triangle very small, as enlarged proepistome. Epistome with straight upper margin; its lower margin ending in small median triangle, with rather sharp apex; on each side, lower margin with two concavities, one for reception of palp of third maxillipeds, the other forming anterior margin of efferent opening. Efferent opening large and occupying outer quarter of anterior margin of buccal field.</paragraph>
<paragraph id="8BAE7E6DB966FF93B182F9D4FB7AF9B4" blockId="4.[151,1437,1402,2040]" box="[199,1218,1618,1645]" pageId="4" pageNumber="119">Mandibular palp three-articulated; last article simple, large, pallet-shaped, with short setae.</paragraph>
<paragraph id="8BAE7E6DB966FF93B182F9F0FC10F949" blockId="4.[151,1437,1402,2040]" box="[199,936,1654,1680]" pageId="4" pageNumber="119">Third maxilliped with well-developed exopod, without flagellum.</paragraph>
<paragraph id="8BAE7E6DB966FF93B182F91CFD07F8B0" blockId="4.[151,1437,1402,2040]" pageId="4" pageNumber="119">Chelipeds equal, remarkably spiny: merus with ventral surface bordered on each margin by acute short teeth (some with corneous tip) and spines, some surounded by short simple setae; carpus inner angle with very long acute tooth bearing several smaller acessory spines, simple setae and spinelike setae; palm with sharp spines on both superior and inferior borders; fingers closing over full length and ending in sharp curved tips, with small spines and short simple setae; superior border of dactylus with some spaced spines; extensor margins of fixed finger and dactylus with corneous teeth and short stiff setae.</paragraph>
<paragraph id="8BAE7E6DB966FF93B182F8F4FE5EF821" blockId="4.[151,1437,1402,2040]" pageId="4" pageNumber="119">Ambulatory legs extremely long and slender; P4 and P5 longest. Merus with distinctly serrated aspect on superior margin, inferior side more finely, corresponding to short acute teeth mixed with simple setae, especially on P2, P3; carpus mostly toothed on inferior side; dactylus very elongated, with numerous longer simple setae on both superior and inferior margins.</paragraph>
<caption id="DF6E2EE5B967FF92B1D2F900FB60F918" pageId="5" pageNumber="120" startId="5.[151,250,1670,1694]" targetBox="[151,1435,181,1646]" targetPageId="5" targetType="figure">
<paragraph id="8BAE7E6DB967FF92B1D2F900FB60F918" blockId="5.[151,1436,1670,1730]" pageId="5" pageNumber="120">
<emphasis id="B965A27FB967FF92B1D2F900FEB5F947" bold="true" box="[151,269,1670,1694]" pageId="5" pageNumber="120">FIGURE 3</emphasis>
.
<taxonomicName id="4C1105EEB967FF92B05FF901FE2DF947" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[282,405,1670,1694]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB967FF92B05FF901FE2DF947" box="[282,405,1670,1694]" italics="true" pageId="5" pageNumber="120">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB967FF92B0D9F900FD82F947" bold="true" box="[412,570,1670,1694]" pageId="5" pageNumber="120">
<taxonomicNameLabel id="A2561F04B967FF92B0D9F900FD82F947" box="[412,570,1670,1694]" pageId="5" pageNumber="120" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm.
</paragraph>
</caption>
<paragraph id="8BAE7E6DB967FF92B182F976FE0FF88B" blockId="5.[151,1436,1776,2019]" pageId="5" pageNumber="120">Thoracic sternum anteriorly covered with small dense setae. Sternal sutures 4/5 to 6/7 interrupted. Median line at level of sternites 7 and 8. Vulvae as two large, rounded openings, close to suture 5/6 and occupying about two-thirds length of sternite 6.</paragraph>
<paragraph id="8BAE7E6DB967FF92B182F8DAFD60F8AF" blockId="5.[151,1436,1776,2019]" box="[199,728,1884,1911]" pageId="5" pageNumber="120">
<emphasis id="B965A27FB967FF92B182F8DAFEE0F8AF" bold="true" box="[199,344,1884,1910]" pageId="5" pageNumber="120">Colouration</emphasis>
. Pale on whole carapace and legs.
</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B967FF92B182F806FC38F83B" pageId="5" pageNumber="120" type="etymology">
<paragraph id="8BAE7E6DB967FF92B182F806FC38F83B" blockId="5.[151,1436,1776,2019]" pageId="5" pageNumber="120">
<emphasis id="B965A27FB967FF92B182F806FEF0F843" bold="true" box="[199,328,1920,1946]" pageId="5" pageNumber="120">Etymology</emphasis>
. Dedicated to Mayla Soubzmaigne, the young daughter of Sébastien Soubzmaigne who made all the photographs of cavernicolous crabs collected by Josiane Lips and French team in various caves of the world. Mayla, as her father, likes Nature and crabs. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B964FF9EB182FF11FC43F82A" lastPageId="9" lastPageNumber="124" pageId="6" pageNumber="121" type="discussion">
<paragraph id="8BAE7E6DB964FF91B182FF11FE6AFEBC" blockId="6.[151,1436,151,357]" pageId="6" pageNumber="121">
<emphasis id="B965A27FB964FF91B182FF11FE8CFF68" bold="true" box="[199,308,151,177]" pageId="6" pageNumber="121">Remarks</emphasis>
. In
<taxonomicName id="4C1105EEB964FF91B021FF11FE56FF68" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[356,494,151,177]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB964FF91B021FF11FE56FF68" box="[356,494,151,177]" italics="true" pageId="6" pageNumber="121">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB964FF91B0B2FF11FD13FF68" bold="true" box="[503,683,151,177]" pageId="6" pageNumber="121">
<taxonomicNameLabel id="A2561F04B964FF91B0B2FF11FD13FF68" box="[503,683,151,177]" pageId="6" pageNumber="121" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
from Lubang Gedung Cave, the pale colouration of the body, the long and slender legs (
<figureCitation id="132A62E8B964FF91B0D3FF3DFE64FF0C" box="[406,476,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="6" pageNumber="121">Fig. 2</figureCitation>
), the nearly immobile and greatly reduced, tapering eyes (
<figureCitation id="132A62E8B964FF91B52BFF3DFA8CFF0C" box="[1134,1332,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="6" pageNumber="121">Figs. 3A, C, D, F</figureCitation>
,
<figureCitation id="132A62E8B964FF91B405FF3DFADDFF0C" box="[1344,1381,187,213]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="6" pageNumber="121">9A</figureCitation>
), the length of which is less than half width of orbit, with a short bullet-shaped stalk and a vestigial unfaceted cornea, devoid of integument pigment except for a trace of pale brown pigment at tip, all are indicative of adaptation to a cavernicolous life making it a true troglobite. It is not known in which part of the cave
<taxonomicName id="4C1105EEB964FF91B525FEA1FB50FE98" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1120,1256,295,321]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB964FF91B525FEA1FB50FE98" box="[1120,1256,295,321]" italics="true" pageId="6" pageNumber="121">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB964FF91B5AAFEA1FA24FE98" bold="true" box="[1263,1436,295,321]" pageId="6" pageNumber="121">
<taxonomicNameLabel id="A2561F04B964FF91B5AAFEA1FA24FE98" box="[1263,1436,295,321]" pageId="6" pageNumber="121" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
was observed and collected.
</paragraph>
<caption id="DF6E2EE5B964FF91B1D2F8FAFC65F860" pageId="6" pageNumber="121" startId="6.[151,250,1916,1940]" targetBox="[151,1436,381,1893]" targetPageId="6" targetType="figure">
<paragraph id="8BAE7E6DB964FF91B1D2F8FAFC65F860" blockId="6.[151,1436,1916,1977]" pageId="6" pageNumber="121">
<emphasis id="B965A27FB964FF91B1D2F8FAFEB4F84D" bold="true" box="[151,268,1916,1940]" pageId="6" pageNumber="121">FIGURE 4</emphasis>
.
<taxonomicName id="4C1105EEB964FF91B05DF8FAFDD9F84D" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[280,609,1916,1940]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB964FF91B05DF8FAFE7CF84D" box="[280,452,1916,1940]" italics="true" pageId="6" pageNumber="121">Cerberusa caeca</emphasis>
<bibRefCitation id="EF80039CB964FF91B08CF8FAFDD9F84D" author="Holthuis, L. B." box="[457,609,1916,1940]" pageId="6" pageNumber="121" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng.
</paragraph>
</caption>
<caption id="DF6E2EE5B965FF90B1D2F897FB5DF897" pageId="7" pageNumber="122" startId="7.[151,250,1809,1833]" targetBox="[164,1423,181,1785]" targetPageId="7" targetType="figure">
<paragraph id="8BAE7E6DB965FF90B1D2F897FB5DF897" blockId="7.[151,1437,1809,1870]" pageId="7" pageNumber="122">
<emphasis id="B965A27FB965FF90B1D2F897FEB5F8F0" bold="true" box="[151,269,1809,1833]" pageId="7" pageNumber="122">FIGURE 5</emphasis>
.
<taxonomicName id="4C1105EEB965FF90B05FF897FDDEF8F0" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[282,614,1809,1833]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB965FF90B05FF897FE7EF8F0" box="[282,454,1809,1833]" italics="true" pageId="7" pageNumber="122">Cerberusa tipula</emphasis>
<bibRefCitation id="EF80039CB965FF90B088F897FDDEF8F0" author="Holthuis, L. B." box="[461,614,1809,1833]" pageId="7" pageNumber="122" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng.
</paragraph>
</caption>
<paragraph id="8BAE7E6DB965FF90B182F8FAFB23F804" blockId="7.[151,1436,1915,2014]" pageId="7" pageNumber="122">
<taxonomicName id="4C1105EEB965FF90B182F8FAFEF3F84C" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[199,331,1915,1941]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB965FF90B182F8FAFEF3F84C" box="[199,331,1915,1941]" italics="true" pageId="7" pageNumber="122">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB965FF90B00AF8FAFE4CF84F" bold="true" box="[335,500,1916,1942]" pageId="7" pageNumber="122">
<taxonomicNameLabel id="A2561F04B965FF90B00AF8FAFE4CF84F" box="[335,500,1916,1942]" pageId="7" pageNumber="122" rank="species">gen. et sp.nov.</taxonomicNameLabel>
</emphasis>
is immediately distinguished from all the other potamids by its multi-spined chelipeds, which additionally bear short simple setae and spinelike setae (
<figureCitation id="132A62E8B965FF90B22BF819FC78F863" box="[878,960,1951,1978]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="7" pageNumber="122">Figs. 2</figureCitation>
,
<figureCitation id="132A62E8B965FF90B28BF826FBACF863" box="[974,1044,1952,1978]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="7" pageNumber="122">3AE</figureCitation>
). The troglobitic characteristics of
<taxonomicName id="4C1105EEB965FF90B1D2F842FEA7F804" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,287,1987,2013]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB965FF90B1D2F842FEA7F804" box="[151,287,1987,2013]" italics="true" pageId="7" pageNumber="122">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB965FF90B063F842FE6DF807" bold="true" box="[294,469,1988,2014]" pageId="7" pageNumber="122">
<taxonomicNameLabel id="A2561F04B965FF90B063F842FE6DF807" box="[294,469,1988,2014]" pageId="7" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of the
<typeStatus id="54AAC0CFB965FF90B245F842FC88F807" box="[768,816,1988,2014]" pageId="7" pageNumber="122">type</typeStatus>
species of
<taxonomicName id="4C1105EEB965FF90B2E8F845FB9BF804" authorityName="Holthuis" authorityYear="1979" box="[941,1059,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB965FF90B2E8F845FB9BF804" box="[941,1059,1987,2013]" italics="true" pageId="7" pageNumber="122">Cerberusa</emphasis>
</taxonomicName>
,
<emphasis id="B965A27FB965FF90B574F842FB23F804" box="[1073,1179,1988,2014]" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="4C1105EEB965FF90B574F842FB2FF804" authorityName="Holthuis" authorityYear="1979" box="[1073,1175,1988,2014]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="caeca">C. caeca</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BAE7E6DB96AFF9FB182FF11FAE4FD5C" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="123">
<taxonomicName id="4C1105EEB96AFF9FB182FF11FE3EFF69" authorityName="Holthuis" authorityYear="1979" box="[199,390,151,177]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96AFF9FB182FF11FE3EFF69" box="[199,390,151,177]" italics="true" pageId="8" pageNumber="123">Cerberusa caeca</emphasis>
</taxonomicName>
, with all type material deposited at RMNH (
<bibRefCitation id="EF80039CB96AFF9FB23AFF11FB2DFF68" author="Fransen, C. H. J. M. &amp; Holthuis, L. B. &amp; Adema, J. P. H. M." box="[895,1173,151,177]" pageId="8" pageNumber="123" pagination="1 - 344" refId="ref13909" refString="Fransen, C. H. J. M., Holthuis, L. B. &amp; Adema, J. P. H. M. (1997) Type-Catalogue of the Decapod Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandlingen, 311, i - xvi + 1 - 344." type="journal article" year="1997">
Fransen
<emphasis id="B965A27FB96AFF9FB2A4FF11FBA2FF68" box="[993,1050,151,177]" italics="true" pageId="8" pageNumber="123">et al.</emphasis>
1997: 138
</bibRefCitation>
), was recorded in
<quantity id="4CE9D388B96AFF9FB421FF11FF15FF0C" metricMagnitude="1" metricUnit="m" metricValue="5.02412" pageId="8" pageNumber="123" unit="in" value="1978.0">1978 in</quantity>
the Gunung Mulu National Park, situated in the extreme northern part of Borneo,
<collectingCountry id="F3063EFDB96AFF9FB502FF3DFB11FF0C" box="[1095,1193,187,213]" name="Malaysia" pageId="8" pageNumber="123">Sarawak</collectingCountry>
, at about
<geoCoordinate id="EE2518AAB96AFF9FB453FF3CFAFBFF0C" box="[1302,1347,186,213]" degrees="4" direction="north" orientation="latitude" pageId="8" pageNumber="123" precision="55555" value="4.0">4°N</geoCoordinate>
<geoCoordinate id="EE2518AAB96AFF9FB416FF3CFA20FF0C" box="[1363,1432,186,213]" degrees="115" direction="east" orientation="longitude" pageId="8" pageNumber="123" precision="55555" value="115.0">115°E</geoCoordinate>
, covering more than
<quantity id="4CE9D388B96AFF9FB032FF58FE73FF20" box="[375,459,222,249]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="123" unit="km" value="500.0">500 km</quantity>
<superScript id="7C64D325B96AFF9FB08EFF58FE6BFF35" attach="left" box="[459,467,222,236]" fontSize="6" pageId="8" pageNumber="123">2</superScript>
and comprising many caves. It was collected in three caves: Green Cave (small stream at
<quantity id="4CE9D388B96AFF9FB1F2FE85FEBAFEC4" box="[183,258,259,285]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="123" unit="m" value="600.0">600 m</quantity>
from north entrance), with the
<typeStatus id="54AAC0CFB96AFF9FB332FE85FD60FEC4" box="[631,728,259,285]" pageId="8" pageNumber="123" type="holotype">holotype</typeStatus>
(RMNH.CRUS.D.31983) and
<specimenCount id="9D17B5E4B96AFF9FB57AFE85FB66FEC4" box="[1087,1246,259,285]" count="2" pageId="8" pageNumber="123" type="generic" typeStatus="paratype">two paratypes</specimenCount>
(RMNH.CRUS. D.31984); Deer Cave, known to contain the largest genuine cave passage in the world (Crab Inlet pool, small pools with gravel and guano on bottom), with a
<typeStatus id="54AAC0CFB96AFF9FB325FECDFD07FEBC" box="[608,703,331,357]" pageId="8" pageNumber="123" type="paratype">paratype</typeStatus>
(RMNH.CRUS.D.31965)); and Clearwater Cave, with the volume of Clearwater system calculated at 49 million cubic metres and the average passage diameter being
<quantity id="4CE9D388B96AFF9FB5B1FEE9FA96FE50" box="[1268,1326,367,393]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="123" unit="m" value="16.0">16 m</quantity>
along the entire
<quantity id="4CE9D388B96AFF9FB19BFE14FE8AFE74" box="[222,306,402,429]" metricMagnitude="5" metricUnit="m" metricValue="2.55" pageId="8" pageNumber="123" unit="km" value="255.0">255 km</quantity>
length (Inflation Passage, gravel floored pool in dribbly stream), with
<specimenCount id="9D17B5E4B96AFF9FB502FE15FB5AFE74" box="[1095,1250,403,429]" count="2" pageId="8" pageNumber="123" type="generic" typeStatus="paratype">two paratypes</specimenCount>
(RMNH.CRUS. D.31959) (
<bibRefCitation id="EF80039CB96AFF9FB04AFE31FE62FE08" author="Holthuis, L. B." box="[271,474,439,465]" pageId="8" pageNumber="123" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis 1979: 16</bibRefCitation>
, fig. 3, pls. 2, 8; 1986: fig. 15;
<bibRefCitation id="EF80039CB96AFF9FB26BFE31FC06FE08" author="Guinot, D." box="[814,958,439,465]" pageId="8" pageNumber="123" pagination="3 - 40" refId="ref14301" refString="Guinot, D. (1988) Les crabes cavernicoles du monde. Memoires de Biospeologie, Paris, 15, 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
: fig. 7, pl. 2, figs. 5, 6; 1990: fig. 6B; 1994: fig. 2C, pl. 1, figs. 4, 5; see
<bibRefCitation id="EF80039CB96AFF9FB08AFE5DFDD5FE2C" author="Grinang, J. A." box="[463,621,475,501]" pageId="8" pageNumber="123" refId="ref14062" refString="Grinang, J. A. (2016) Taxonomy of Freshwater Crabs (Crustacea: Decapoda: Brachyura) in Sarawak, Borneo, with Autecology of Four Species. Ph. D thesis, Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, Kota Samarahan. [unknown pagination]" type="book" year="2016">Grinang 2016</bibRefCitation>
: fig. 7).
<bibRefCitation id="EF80039CB96AFF9FB38EFE5DFC4EFE2C" author="McFarlane, D. &amp; Lundberg, J. &amp; Christenson, K." box="[715,1014,475,501]" pageId="8" pageNumber="123" pagination="1 - 18" refId="ref15641" refString="McFarlane, D., Lundberg, J. &amp; Christenson, K. (2011) New Records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of Northern Sarawak. Speleobiology Notes, 3, 1 - 18." type="journal article" year="2011">
McFarlane
<emphasis id="B965A27FB96AFF9FB20FFE5DFCC4FE2C" box="[842,892,475,501]" italics="true" pageId="8" pageNumber="123">et al</emphasis>
. (2011: 12
</bibRefCitation>
, fig. 2) found it in a pool of Lagang
<emphasis id="B965A27FB96AFF9FB4CDFE5CFA29FE2D" bold="true" box="[1416,1425,474,500]" pageId="8" pageNumber="123"></emphasis>
s Cave, connected to the Deer Cave.
<taxonomicName id="4C1105EEB96AFF9FB36AFE79FD2BFDC1" authorityName="Holthuis" authorityYear="1979" box="[559,659,511,537]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96AFF9FB36AFE79FD2BFDC1" box="[559,659,511,537]" italics="true" pageId="8" pageNumber="123">C. caeca</emphasis>
</taxonomicName>
was again collected by K. Lim
<emphasis id="B965A27FB96AFF9FB542FE79FBFAFDC0" box="[1031,1090,511,537]" italics="true" pageId="8" pageNumber="123">et al.</emphasis>
in 2006 from another cave of the Gunung Mulu National Park, the Stone Horse Cave, near the Deer Cave, and both specimens are deposited at ZRC (ZRC 2013.1220).
<taxonomicName id="4C1105EEB96AFF9FB0EFFDC1FDD0FDB9" authorityName="Holthuis" authorityYear="1979" box="[426,616,583,609]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96AFF9FB0EFFDC1FDD0FDB9" box="[426,616,583,609]" italics="true" pageId="8" pageNumber="123">Cerberusa caeca</emphasis>
</taxonomicName>
prefers to dwell in water. It is blind and long-legged; its carapace is pale yellowish or white (
<figureCitation id="132A62E8B96AFF9FB030FDEDFE01FD5C" box="[373,441,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="8" pageNumber="123">Fig. 4</figureCitation>
), thus it is fully adapted to living exclusively in the complete dark part of the caves.
</paragraph>
<paragraph id="8BAE7E6DB96AFF9FB182FD09FF67FC84" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="123">
<taxonomicName id="4C1105EEB96AFF9FB182FD09FE30FD71" authorityName="Holthuis" authorityYear="1979" box="[199,392,655,681]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96AFF9FB182FD09FE30FD71" box="[199,392,655,681]" italics="true" pageId="8" pageNumber="123">Cerberusa caeca</emphasis>
</taxonomicName>
is at least present in three massifs of Gunung Mulu National Park: Gunung Api, Southern Hills and Gunung Benerat, and is therefore a very widespread troglobitic species. Due to its extreme adaptations to a troglobitic life, it presumably has a long evolutionary history and probably diverged from its epigean ancestor before the separation of the Gunung Mulu Uplands into distinct massifs, whereas
<taxonomicName id="4C1105EEB96AFF9FB570FD7DFB2FFCCC" authorityName="Holthuis" authorityYear="1979" box="[1077,1175,763,789]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96AFF9FB570FD7DFB2FFCCC" box="[1077,1175,763,789]" italics="true" pageId="8" pageNumber="123">C. tipula</emphasis>
</taxonomicName>
, with fewer troglobitic adaptations (the eyes and the colouration), can be hypothesised to have arisen more recently (
<bibRefCitation id="EF80039CB96AFF9FB59EFC99FF6BFC84" author="McFarlane, D. &amp; Lundberg, J. &amp; Christenson, K." pageId="8" pageNumber="123" pagination="1 - 18" refId="ref15641" refString="McFarlane, D., Lundberg, J. &amp; Christenson, K. (2011) New Records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of Northern Sarawak. Speleobiology Notes, 3, 1 - 18." type="journal article" year="2011">
McFarlane
<emphasis id="B965A27FB96AFF9FB424FC99FA25FCE0" box="[1377,1437,799,825]" italics="true" pageId="8" pageNumber="123">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
<caption id="DF6E2EE5B96AFF9FB1D2F912FBC9F92C" pageId="8" pageNumber="123" startId="8.[151,250,1684,1708]" targetBox="[164,1423,885,1661]" targetPageId="8" targetType="figure">
<paragraph id="8BAE7E6DB96AFF9FB1D2F912FBC9F92C" blockId="8.[151,1437,1684,1781]" pageId="8" pageNumber="123">
<emphasis id="B965A27FB96AFF9FB1D2F912FEB6F975" bold="true" box="[151,270,1684,1708]" pageId="8" pageNumber="123">FIGURE 6</emphasis>
. Vulvae: A,
<taxonomicName id="4C1105EEB96AFF9FB0CAF913FDB3F975" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[399,523,1684,1708]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96AFF9FB0CAF913FDB3F975" box="[399,523,1684,1708]" italics="true" pageId="8" pageNumber="123">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96AFF9FB357F912FD0AF975" bold="true" box="[530,690,1684,1708]" pageId="8" pageNumber="123">
<taxonomicNameLabel id="A2561F04B96AFF9FB357F912FD0AF975" box="[530,690,1684,1708]" pageId="8" pageNumber="123" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B,
<taxonomicName id="4C1105EEB96AFF9FB5AAF912FE88F909" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96AFF9FB5AAF912FA24F975" box="[1263,1436,1684,1708]" italics="true" pageId="8" pageNumber="123">Cerberusa caeca</emphasis>
<bibRefCitation id="EF80039CB96AFF9FB1D2F93EFE88F909" author="Holthuis, L. B." box="[151,304,1720,1744]" pageId="8" pageNumber="123" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
, female, 9.9 × 13.3 mm, ZRC 2013.1220; C,
<taxonomicName id="4C1105EEB96AFF9FB3B3F93FFC54F909" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[758,1004,1720,1744]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96AFF9FB3B3F93FFCF6F909" box="[758,846,1720,1744]" italics="true" pageId="8" pageNumber="123">C. tipula</emphasis>
<bibRefCitation id="EF80039CB96AFF9FB211F93EFC54F909" author="Holthuis, L. B." box="[852,1004,1720,1744]" pageId="8" pageNumber="123" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
, female, 13.0 × 17.6 mm, ZRC 2013.1219; D,
<taxonomicName id="4C1105EEB96AFF9FB1F3F95AFE0AF92D" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[182,434,1756,1781]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="aethes">
<emphasis id="B965A27FB96AFF9FB1F3F95AFEF3F92D" box="[182,331,1756,1780]" italics="true" pageId="8" pageNumber="123">Ibanum aethes</emphasis>
<bibRefCitation id="EF80039CB96AFF9FB014F95BFE0AF92D" author="Ng, P. K. L." box="[337,434,1756,1781]" pageId="8" pageNumber="123" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng.
</paragraph>
</caption>
<paragraph id="8BAE7E6DB96AFF9EB182F8A4FA88FEC5" blockId="8.[151,1436,1826,2033]" lastBlockId="9.[151,1437,151,2036]" lastPageId="9" lastPageNumber="124" pageId="8" pageNumber="123">
The second species of
<taxonomicName id="4C1105EEB96AFF9FB0FAF8A4FD8DF8E5" authorityName="Holthuis" authorityYear="1979" box="[447,565,1826,1852]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB96AFF9FB0FAF8A4FD8DF8E5" box="[447,565,1826,1852]" italics="true" pageId="8" pageNumber="123">Cerberusa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1105EEB96AFF9FB307F8A5FD1BF8E5" authorityName="Holthuis" authorityYear="1979" box="[578,675,1826,1853]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96AFF9FB307F8A5FD1BF8E5" box="[578,675,1826,1853]" italics="true" pageId="8" pageNumber="123">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96AFF9FB3F4F8A4FD4BF8E4" box="[689,755,1826,1853]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="8" pageNumber="123">Fig. 5</figureCitation>
), was originally found in two caves in the Gunung Api massif of Gunung Mulu National Park: Clearwater Cave (
<quantity id="4CE9D388B96AFF9FB38FF8C0FCBBF8B8" box="[714,771,1862,1889]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="123" unit="m" value="15.0">15 m</quantity>
inside northwest entrance, on stone outside water), with the
<typeStatus id="54AAC0CFB96AFF9FB1D2F8ECFF41F85D" box="[151,249,1898,1924]" pageId="8" pageNumber="123" type="holotype">holotype</typeStatus>
(RMNH.CRUS.D.31968); and Wonder Cave (
<quantity id="4CE9D388B96AFF9FB242F8EDFCF9F85C" box="[775,833,1899,1925]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="123" unit="m" value="10.0">10 m</quantity>
from entrance, pool with bottom sediment of guano), with a
<typeStatus id="54AAC0CFB96AFF9FB1A4F809FEFBF870" box="[225,323,1935,1961]" pageId="8" pageNumber="123" type="paratype">Paratype</typeStatus>
(RMNH.CRUS.D.31969) (
<bibRefCitation id="EF80039CB96AFF9FB331F808FCF8F870" author="Holthuis, L. B." box="[628,832,1934,1961]" pageId="8" pageNumber="123" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis 1979: 13</bibRefCitation>
, fig. 2, pl. 1, pl. 3, fig. 1;
<bibRefCitation id="EF80039CB96AFF9FB517F808FB59F870" author="Guinot, D." box="[1106,1249,1934,1961]" pageId="8" pageNumber="123" pagination="3 - 40" refId="ref14301" refString="Guinot, D. (1988) Les crabes cavernicoles du monde. Memoires de Biospeologie, Paris, 15, 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
: fig. 6A, B, pl. 2, figs. 1
<emphasis id="B965A27FB96AFF9FB199F835FF52F814" bold="true" box="[220,234,1971,1997]" pageId="8" pageNumber="123"></emphasis>
4; 1994: fig. 2B; see
<bibRefCitation id="EF80039CB96AFF9FB08AF834FDD4F815" author="Grinang, J. A." box="[463,620,1970,1996]" pageId="8" pageNumber="123" refId="ref14062" refString="Grinang, J. A. (2016) Taxonomy of Freshwater Crabs (Crustacea: Decapoda: Brachyura) in Sarawak, Borneo, with Autecology of Four Species. Ph. D thesis, Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, Kota Samarahan. [unknown pagination]" type="book" year="2016">Grinang 2016</bibRefCitation>
: fig. 8); then in a third cave, the Cave of Winds (
<bibRefCitation id="EF80039CB96AFF9FB53BF834FAE2F815" author="Holthuis, L. B." box="[1150,1370,1970,1996]" pageId="8" pageNumber="123" pagination="589 - 615" refId="ref14830" refString="Holthuis, L. B. (1986) Decapoda. In: Botosaneanu, L. (Ed.), Stygofauna mundi. A faunistic, distributional, and ecological synthesis of the world fauna inhabiting subterranean waters (including the marine interstitial). Brill, Leiden, pp. 589 - 615. https: // doi. org / 10.1163 / 9789004631977 _ 071" type="book chapter" year="1986">Holthuis 1986: 613</bibRefCitation>
); and, later, in an additional cave located within the Southern Hills, the Fruit Bat Cave (limestone pools
<quantity id="4CE9D388B96AFF9FB5A9F850FA8CF828" box="[1260,1332,2006,2033]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="123" unit="m" value="150.0">150 m</quantity>
from the west entrance southwest massif) (
<bibRefCitation id="EF80039CB96BFF9EB34EFF11FC95FF68" author="McFarlane, D. &amp; Lundberg, J. &amp; Christenson, K." box="[523,813,151,177]" pageId="9" pageNumber="124" pagination="1 - 18" refId="ref15641" refString="McFarlane, D., Lundberg, J. &amp; Christenson, K. (2011) New Records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of Northern Sarawak. Speleobiology Notes, 3, 1 - 18." type="journal article" year="2011">
McFarlane
<emphasis id="B965A27FB96BFF9EB3C8FF11FD05FF68" box="[653,701,151,177]" italics="true" pageId="9" pageNumber="124">et al</emphasis>
. 2011: 12
</bibRefCitation>
, figs. 2, 3). As
<taxonomicName id="4C1105EEB96BFF9EB297FF11FB8BFF69" authorityName="Holthuis" authorityYear="1979" box="[978,1075,151,177]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB297FF11FB8BFF69" box="[978,1075,151,177]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1105EEB96BFF9EB505FF11FB19FF68" authorityName="Holthuis" authorityYear="1979" box="[1088,1185,151,177]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB505FF11FB19FF68" box="[1088,1185,151,177]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
was again collected in another cave of Gunung Mulu National Park, the Stone Horse Cave, near the Deer Cave, by K. Lim
<emphasis id="B965A27FB96BFF9EB445FF3DFA81FF0D" box="[1280,1337,186,212]" italics="true" pageId="9" pageNumber="124">et al.</emphasis>
in 2006, with
<specimenCount id="9D17B5E4B96BFF9EB195FF59FEF6FF21" box="[208,334,222,249]" count="2" pageId="9" pageNumber="124" type="female">two female</specimenCount>
specimens (ZRC 2013.1219). No epigean records are known.
<taxonomicName id="4C1105EEB96BFF9EB548FF58FB73FF21" authorityName="Holthuis" authorityYear="1979" box="[1037,1227,222,248]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB548FF58FB73FF21" box="[1037,1227,222,248]" italics="true" pageId="9" pageNumber="124">Cerberusa tipula</emphasis>
</taxonomicName>
can be considered a true troglobite despite its cylindrical eyes still retaining distinct pigmentation and carapace colouration.
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182FEA0FC76FD84" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
<taxonomicName id="4C1105EEB96BFF9EB182FEA0FE3EFE99" authorityName="Holthuis" authorityYear="1979" box="[199,390,294,320]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB182FEA0FE3EFE99" box="[199,390,294,320]" italics="true" pageId="9" pageNumber="124">Cerberusa tipula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1105EEB96BFF9EB0FBFEA0FD99FEE6" authorityName="Holthuis" authorityYear="1979" box="[446,545,294,320]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB0FBFEA0FD99FEE6" box="[446,545,294,320]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
cohabit in three caves: Clearwater Cave, Deer Cave and Stone Horse Cave.
<taxonomicName id="4C1105EEB96BFF9EB4C6FEA0FF60FEBA" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB4C6FEA0FF60FEBA" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
is an amphibious species, with a colourful (blue grey) dorsal surface of carapace and depigmented or lightly coloured (pale lemon-yellow or orange-yellow) legs, with practically unmodified eyes and a distinct rounded cornea containing dark pigment (
<figureCitation id="132A62E8B96BFF9EB0FBFE17FDA1FE72" box="[446,537,401,427]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">Fig. 9C</figureCitation>
). Living specimens of
<taxonomicName id="4C1105EEB96BFF9EB264FE17FC3CFE72" authorityName="Holthuis" authorityYear="1979" box="[801,900,401,427]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB264FE17FC3CFE72" box="[801,900,401,427]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
from Fruit Bat Cave and from Agong Cave in Gunung Mulu National Park (
<bibRefCitation id="EF80039CB96BFF9EB0A7FE32FD65FE16" author="McFarlane, D. &amp; Lundberg, J. &amp; Christenson, K." box="[482,733,436,463]" pageId="9" pageNumber="124" pagination="1 - 18" refId="ref15641" refString="McFarlane, D., Lundberg, J. &amp; Christenson, K. (2011) New Records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of Northern Sarawak. Speleobiology Notes, 3, 1 - 18." type="journal article" year="2011">
McFarlane
<emphasis id="B965A27FB96BFF9EB321FE33FD2CFE17" box="[612,660,436,462]" italics="true" pageId="9" pageNumber="124">et al</emphasis>
. 2011
</bibRefCitation>
: fig. 3 and
<bibRefCitation id="EF80039CB96BFF9EB213FE32FC4CFE17" author="Grinang, J. A." box="[854,1012,436,462]" pageId="9" pageNumber="124" refId="ref14062" refString="Grinang, J. A. (2016) Taxonomy of Freshwater Crabs (Crustacea: Decapoda: Brachyura) in Sarawak, Borneo, with Autecology of Four Species. Ph. D thesis, Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, Kota Samarahan. [unknown pagination]" type="book" year="2016">Grinang 2016</bibRefCitation>
: fig. 7, respectively) show spectacular colouration: carapace grey, chelipeds and legs pale yellow.
<bibRefCitation id="EF80039CB96BFF9EB274FE5EFC74FE2B" author="Guinot, D." box="[817,972,472,498]" pageId="9" pageNumber="124" pagination="3 - 40" refId="ref14301" refString="Guinot, D. (1988) Les crabes cavernicoles du monde. Memoires de Biospeologie, Paris, 15, 3 - 40." type="journal article" year="1988">Guinot (1988</bibRefCitation>
: fig. 6A, B) observed that, in the Cave of Winds, a male
<taxonomicName id="4C1105EEB96BFF9EB07FFE7AFE23FDCF" authorityName="Holthuis" authorityYear="1979" box="[314,411,508,534]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB07FFE7AFE23FDCF" box="[314,411,508,534]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
12.5 ×
<quantity id="4CE9D388B96BFF9EB0A8FE7AFD83FDCE" box="[493,571,508,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="124" unit="mm" value="16.0">16 mm</quantity>
had the eyestalk somewhat thinned distally and filling a good part of the orbit, whereas in a female of 16 ×
<quantity id="4CE9D388B96BFF9EB090FDA6FD81FDE3" box="[469,569,544,570]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="9" pageNumber="124" unit="mm" value="21.8">21.8 mm</quantity>
it was less developed, barely thinned distally and occupying a smaller part of the orbit, and with a small, pigmented cornea (
<figureCitation id="132A62E8B96BFF9EB3E4FDC5FD40FD87" box="[673,760,579,606]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">Fig. 9C</figureCitation>
, male and female).
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182FDEEFCBBFC81" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
In the 1980s, the first author had the opportunity to examine the
<typeStatus id="54AAC0CFB96BFF9EB2D9FDEEFC74FD5B" box="[924,972,616,642]" pageId="9" pageNumber="124">type</typeStatus>
material of the two species of
<taxonomicName id="4C1105EEB96BFF9EB463FDE1FA24FD58" authorityName="Holthuis" authorityYear="1979" box="[1318,1436,615,641]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB96BFF9EB463FDE1FA24FD58" box="[1318,1436,615,641]" italics="true" pageId="9" pageNumber="124">Cerberusa</emphasis>
</taxonomicName>
deposited at RMNH and published photographs of all of them together with sketches of the degenerated eyes (
<bibRefCitation id="EF80039CB96BFF9EB1DAFD29FE88FD10" author="Guinot, D." box="[159,304,687,713]" pageId="9" pageNumber="124" pagination="3 - 40" refId="ref14301" refString="Guinot, D. (1988) Les crabes cavernicoles du monde. Memoires de Biospeologie, Paris, 15, 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
, 1994). These illustrations are all deposited in the MNHN, but, as the photographs are silver, they are not reproduced in the present paper. Therefore, photographs of females specimens of
<taxonomicName id="4C1105EEB96BFF9EB529FD55FB68FD35" authorityName="Holthuis" authorityYear="1979" box="[1132,1232,723,749]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB529FD55FB68FD35" box="[1132,1232,723,749]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB5A7FD54FA8BFD34" box="[1250,1331,722,749]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Figs. 4</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB404FD54FADCFD35" box="[1345,1380,722,748]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6B</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB437FD55FA2CFD34" box="[1394,1428,723,749]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">9B</figureCitation>
) and
<taxonomicName id="4C1105EEB96BFF9EB182FD71FE92FCC9" authorityName="Holthuis" authorityYear="1979" box="[199,298,758,785]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB182FD71FE92FCC9" box="[199,298,758,785]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB07FFD70FE32FCC8" box="[314,394,758,785]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Figs. 5</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB0D2FD70FE01FCC9" box="[407,441,758,784]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6C</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB083FD70FE50FCC9" box="[454,488,758,784]" captionStart="FIGURE 9" captionStartId="15.[151,250,1709,1733]" captionTargetBox="[159,1428,281,1678]" captionTargetId="figure-73@15.[151,1436,273,1686]" captionTargetPageId="15" captionText="FIGURE 9. Ocular regression: A, Ngan mayla gen. et sp. nov., (Potamidae), holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979 (Potamidae), male, 12.5 × 17.0 mm, RMNH.CRUS.D.31904; C, C. tipula Holthuis, 1979 (Potamidae), male 12.5 × 16.0 mm, female 16.0 × 21.8 mm, RMNH.CRUS.D.33959; D, Holthuisana alba Holthuis, 1980 (Gecarcinucidae), paratype female, 22.0 × 27.0 mm, RMNH.CRUS.D.33302; E, Sendleria genuitei Guinot, 1987 (Gecarcinucidae), holotype male, 22.0 × 29.0 mm, MNHN-IU-2014-11301 (= MNHN-B17968); F, Typhlopseudothelphusa juberthiei Delamare Deboutteville, 1976 (Pseudothelphusidae Ortmann, 1893), female, 21.0 × 32.0 mm, MNHN-IU-2014- 23194 (= MNHN-B18870); G, Trogloplax joliveti Guinot, 1986 (Trogloplacinae Guinot, 1986), paratype male, 16.0 × 21.0 mm, MNHN-IU-2014-10458 (= MNHN-B12794); H, Austinograea alayseae Guinot, 1990 (Bythograeoidea Williams, 1980), allotype female, 21.0 × 32.0 mm, MNHN-IU-2008-11306 (= MNHN-B24022). Scale bar: A = 1 mm." pageId="9" pageNumber="124">9C</figureCitation>
) collected by K. Lim
<emphasis id="B965A27FB96BFF9EB3A4FD71FCA2FCC9" box="[737,794,758,784]" italics="true" pageId="9" pageNumber="124">et al.</emphasis>
in
<quantity id="4CE9D388B96BFF9EB27AFD70FC2DFCC9" box="[831,917,758,784]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="9" pageNumber="124" unit="in" value="2006.0">2006 in</quantity>
Stone Horse Cave (ZRC 2013.1220 and ZRC 2013.1219, respectively), near the Deer Cave where the
<typeStatus id="54AAC0CFB96BFF9EB257FC9CFCFAFCED" box="[786,834,794,820]" pageId="9" pageNumber="124">type</typeStatus>
series of
<taxonomicName id="4C1105EEB96BFF9EB2F5FC9CFBAAFCED" authorityName="Holthuis" authorityYear="1979" box="[944,1042,794,820]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB2F5FC9CFBAAFCED" box="[944,1042,794,820]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
was collected (
<bibRefCitation id="EF80039CB96BFF9EB584FC9CFADCFCED" author="Holthuis, L. B." box="[1217,1380,794,820]" pageId="9" pageNumber="124" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis 1979</bibRefCitation>
) and kindly made and sent by P.K.L. Ng, are published here.
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182FCE7FE9BFB8B" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
The genera
<taxonomicName id="4C1105EEB96BFF9EB00CFCE4FE3EFCA2" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[329,390,866,891]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB96BFF9EB00CFCE4FE3EFCA2" box="[329,390,866,891]" italics="true" pageId="9" pageNumber="124">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB0CEFCE7FE4BFCA2" bold="true" box="[395,499,865,891]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB0CEFCE7FE4BFCA2" box="[395,499,865,891]" pageId="9" pageNumber="124" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1105EEB96BFF9EB362FCE7FD25FCA2" authorityName="Holthuis" authorityYear="1979" box="[551,669,865,891]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB96BFF9EB362FCE7FD25FCA2" box="[551,669,865,891]" italics="true" pageId="9" pageNumber="124">Cerberusa</emphasis>
</taxonomicName>
share a similar habitus: carapace widest anteriorly; short, arcuate and carinate antero-lateral margin granulated to gently serrated, without teeth or epibranchial incision, the carina curving onto the carapace dorsal surface; front flat and rather straight; and posterolateral margins long, with tranverse ridges. Both are long-legged species. The pigmentation varies from white or pale yellow in
<taxonomicName id="4C1105EEB96BFF9EB51DFC4BFB59FC3F" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1112,1249,972,998]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB51DFC4BFB59FC3F" box="[1112,1249,972,998]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB5AFFC4BFA24FC3E" bold="true" box="[1258,1436,973,999]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB5AFFC4BFA24FC3E" box="[1258,1436,973,999]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1105EEB96BFF9EB18DFC77FE93FBD3" authorityName="Holthuis" authorityYear="1979" box="[200,299,1009,1035]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB18DFC77FE93FBD3" box="[200,299,1009,1035]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
, which, with their completely degenerated eyes, would be the most troglomorphic potamids known, to a mixed colouration (as is often the case in cave animals) in
<taxonomicName id="4C1105EEB96BFF9EB220FB92FC70FBF7" authorityName="Holthuis" authorityYear="1979" box="[869,968,1044,1070]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB220FB92FC70FBF7" box="[869,968,1044,1070]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
, which has slightly reduced eyes, but the longest legs.
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182FBDAFB46F9FB" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
<taxonomicName id="4C1105EEB96BFF9EB182FBDAFEE9FBAC" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[199,337,1115,1141]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB182FBDAFEE9FBAC" box="[199,337,1115,1141]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB019FBDAFDAAFBAF" bold="true" box="[348,530,1116,1142]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB019FBDAFDAAFBAF" box="[348,530,1116,1142]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C1105EEB96BFF9EB3EBFBDAFCAAFBAC" authorityName="Holthuis" authorityYear="1979" box="[686,786,1116,1142]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB3EBFBDAFCAAFBAC" box="[686,786,1116,1142]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
mainly by the numerous spines and simple setae on the chelipeds (
<figureCitation id="132A62E8B96BFF9EB057FBF9FEE0FB40" box="[274,344,1151,1177]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB020FBF9FE10FB40" box="[357,424,1151,1177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="9" pageNumber="124">3AE</figureCitation>
) (chelipeds unarmed, without spinelike setae in
<taxonomicName id="4C1105EEB96BFF9EB295FB06FB8BFB40" authorityName="Holthuis" authorityYear="1979" box="[976,1075,1152,1178]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB295FB06FB8BFB40" box="[976,1075,1152,1178]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
:
<figureCitation id="132A62E8B96BFF9EB501FBF9FB66FB43" box="[1092,1246,1151,1178]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4AC, E</figureCitation>
, as well as in
<taxonomicName id="4C1105EEB96BFF9EB4C6FB06FF60FB64" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB4C6FB06FF60FB64" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
:
<figureCitation id="132A62E8B96BFF9EB1ADFB25FE95FB67" box="[232,301,1187,1214]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5</figureCitation>
); by the more elongated, thinner legs (
<figureCitation id="132A62E8B96BFF9EB3A1FB25FC91FB64" box="[740,809,1187,1213]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
), in particular, the carpus, propodus and dactylus being longer and slender; by the P5 being as long as P4 (P5 shorter than P
<quantity id="4CE9D388B96BFF9EB2C1FB41FC16FB38" box="[900,942,1223,1249]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="124" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C1105EEB96BFF9EB2F1FB41FBADFB39" authorityName="Holthuis" authorityYear="1979" box="[948,1045,1223,1249]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB2F1FB41FBADFB39" box="[948,1045,1223,1249]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
:
<figureCitation id="132A62E8B96BFF9EB566FB41FB01FB38" box="[1059,1209,1223,1249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A, B, E</figureCitation>
); by certain margins of legs being finely serrated and with erect stiff, short setae (smoother, only serrated on inferior margin of propodi in
<taxonomicName id="4C1105EEB96BFF9EB1F3FA89FEA1FAF1" authorityName="Holthuis" authorityYear="1979" box="[182,281,1295,1321]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB1F3FA89FEA1FAF1" box="[182,281,1295,1321]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
). The legs of
<taxonomicName id="4C1105EEB96BFF9EB0F8FA89FD99FAF1" authorityName="Holthuis" authorityYear="1979" box="[445,545,1294,1321]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB0F8FA89FD99FAF1" box="[445,545,1294,1321]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
are very long and thin (
<figureCitation id="132A62E8B96BFF9EB27CFA88FC2CFAF0" box="[825,916,1294,1321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5A</figureCitation>
), even longer and thinner than those of
<taxonomicName id="4C1105EEB96BFF9EB425FA89FF63FA95" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB425FA89FF63FA95" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB1A6FAB4FE2BFA95" bold="true" box="[227,403,1330,1356]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB1A6FAB4FE2BFA95" box="[227,403,1330,1356]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB0E6FAB4FE5FFA95" box="[419,487,1330,1356]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
) (except that P5 is proportionally shorter in
<taxonomicName id="4C1105EEB96BFF9EB299FAB4FB86FA95" authorityName="Holthuis" authorityYear="1979" box="[988,1086,1330,1356]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB299FAB4FB86FA95" box="[988,1086,1330,1356]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
), and also longer than those of
<taxonomicName id="4C1105EEB96BFF9EB1D2FAD0FF40FAB6" authorityName="Holthuis" authorityYear="1979" box="[151,248,1366,1392]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB1D2FAD0FF40FAB6" box="[151,248,1366,1392]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
, which, of the three, is the species with the proportionally shorter legs (
<figureCitation id="132A62E8B96BFF9EB553FAD0FBC8FAA9" box="[1046,1136,1366,1392]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A</figureCitation>
). The serrated margins and numerous stiff short erect bristles of the legs of
<taxonomicName id="4C1105EEB96BFF9EB3EBFAFCFC8EFA4A" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[686,822,1401,1427]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB3EBFAFCFC8EFA4A" box="[686,822,1401,1427]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB27BFAFFFC54FA4A" bold="true" box="[830,1004,1401,1427]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB27BFAFFFC54FA4A" box="[830,1004,1401,1427]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB2BEFAFFFB87FA4A" box="[1019,1087,1401,1427]" captionStart="FIGURE 2" captionStartId="4.[151,246,1296,1320]" captionTargetBox="[173,1415,634,1272]" captionTargetId="figure-168@4.[173,1415,634,1272]" captionTargetPageId="4" captionText="FIGURE 2. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, Kalimantan (East Kalimantan), Indonesia, Borneo, Merabu Karst, Lubang Gedung Cave, 1°381”N117°2414”E, 850 m, 18 August 2010, MZB Cru 5710: habitus. Scale bar = 5 mm." pageId="9" pageNumber="124">Fig. 2</figureCitation>
) are somewhat similar to those of
<taxonomicName id="4C1105EEB96BFF9EB1F2FA1BFEA1FA6E" authorityName="Holthuis" authorityYear="1979" box="[183,281,1437,1463]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB1F2FA1BFEA1FA6E" box="[183,281,1437,1463]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB06FFA1BFE3CFA61" box="[298,388,1437,1464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5A</figureCitation>
), whereas the legs are smoother in
<taxonomicName id="4C1105EEB96BFF9EB253FA1BFCC1FA6E" authorityName="Holthuis" authorityYear="1979" box="[790,889,1437,1463]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB253FA1BFCC1FA6E" box="[790,889,1437,1463]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB2CFFA1BFC5DFA6E" box="[906,997,1437,1463]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4A</figureCitation>
), with only the posterior margin of the propodus bearing spinules. The female thoracic sternum, covered with dense setae in
<taxonomicName id="4C1105EEB96BFF9EB519FA44FB5CFA02" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1116,1252,1473,1499]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB519FA44FB5CFA02" box="[1116,1252,1473,1499]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB5A9FA47FA24FA02" bold="true" box="[1260,1436,1473,1499]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB5A9FA47FA24FA02" box="[1260,1436,1473,1499]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB1DAFA62FEB8FA26" box="[159,256,1508,1535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="9" pageNumber="124">Figs. 3E</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB049FA62FE8AFA27" box="[268,306,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6A</figureCitation>
), is bare in
<taxonomicName id="4C1105EEB96BFF9EB0F1FA63FDADFA27" authorityName="Holthuis" authorityYear="1979" box="[436,533,1509,1535]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB0F1FA63FDADFA27" box="[436,533,1509,1535]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB360FA62FD3EFA26" box="[549,646,1508,1535]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Figs. 4E</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB3D7FA62FD0BFA27" box="[658,691,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6B</figureCitation>
) and
<taxonomicName id="4C1105EEB96BFF9EB3B7FA63FCECFA27" authorityName="Holthuis" authorityYear="1979" box="[754,852,1508,1535]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB3B7FA63FCECFA27" box="[754,852,1508,1535]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB221FA62FC7DFA26" box="[868,965,1508,1535]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Figs. 5E</figureCitation>
,
<figureCitation id="132A62E8B96BFF9EB295FA62FC4DFA27" box="[976,1013,1508,1534]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. BD, courtesy P.K.L. Ng." pageId="9" pageNumber="124">6C</figureCitation>
). The efferent openings are enormous in
<taxonomicName id="4C1105EEB96BFF9EB1F1F98FFEAEF9FB" authorityName="Holthuis" authorityYear="1979" box="[180,278,1544,1571]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB1F1F98FFEAEF9FB" box="[180,278,1544,1571]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB060F98EFEC5F9FA" box="[293,381,1544,1571]" captionStart="FIGURE 5" captionStartId="7.[151,250,1809,1833]" captionTargetBox="[164,1423,181,1785]" captionTargetId="figure-31@7.[164,1423,181,1785]" captionTargetPageId="7" captionText="FIGURE 5. Cerberusa tipula Holthuis, 1979, female, 13.0 × 17.6 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1219: A, habitus; B, frontal view; C, left cheliped; D, carapace and chelipeds; E, ventral view. Courtesy P.K.L. Ng." pageId="9" pageNumber="124">Fig. 5B</figureCitation>
), normal in
<taxonomicName id="4C1105EEB96BFF9EB340F98FFDDEF9FB" authorityName="Holthuis" authorityYear="1979" box="[517,614,1545,1571]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB340F98FFDDEF9FB" box="[517,614,1545,1571]" italics="true" pageId="9" pageNumber="124">C. caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B96BFF9EB333F98EFD56F9FB" box="[630,750,1544,1570]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1940]" captionTargetBox="[151,1436,381,1893]" captionTargetId="figure-133@6.[151,1436,381,1893]" captionTargetPageId="6" captionText="FIGURE 4. Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, Borneo, Sarawak, Stone Horse Cave, ZRC 2013.1220: A, habitus; B, D, frontal views; C, left cheliped; E, ventral view. Courtesy, P.K.L. Ng." pageId="9" pageNumber="124">Fig. 4B, D</figureCitation>
) and
<taxonomicName id="4C1105EEB96BFF9EB268F98FFC0DF9FB" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[813,949,1544,1570]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB268F98FFC0DF9FB" box="[813,949,1544,1570]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB2F9F98EFBD2F9FB" bold="true" box="[956,1130,1544,1570]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB2F9F98EFBD2F9FB" box="[956,1130,1544,1570]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B96BFF9EB53FF98EFB49F9FB" box="[1146,1265,1544,1570]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: AE = 5 mm; F = 1 mm." pageId="9" pageNumber="124">Fig. 3A, F</figureCitation>
).
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182F9AAFAFDF954" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
The Lubang Gedung Cave hosting
<taxonomicName id="4C1105EEB96BFF9EB325F9ABFD65F99F" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[608,733,1580,1606]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="myla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB325F9ABFD65F99F" box="[608,733,1580,1606]" italics="true" pageId="9" pageNumber="124">Ngan myla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB3ADF9AAFC18F99F" bold="true" box="[744,928,1580,1606]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB3ADF9AAFC18F99F" box="[744,928,1580,1606]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
is around
<quantity id="4CE9D388B96BFF9EB55AF9AAFBCFF99F" box="[1055,1143,1580,1606]" metricMagnitude="5" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="124" unit="km" value="370.0">370 km</quantity>
southeast of the Gunung Mulu National Park, home to both species of
<taxonomicName id="4C1105EEB96BFF9EB3E8F9D6FC9BF9B3" authorityName="Holthuis" authorityYear="1979" box="[685,803,1616,1642]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB96BFF9EB3E8F9D6FC9BF9B3" box="[685,803,1616,1642]" italics="true" pageId="9" pageNumber="124">Cerberusa</emphasis>
</taxonomicName>
. The ranges of the two genera belong to two different hydrographic drainage systems, which implies that they are well separated geographically from each other.
</paragraph>
<paragraph id="8BAE7E6DB96BFF9EB182F91EFC43F82A" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="124">
Many cave species occur in
<collectingCountry id="F3063EFDB96BFF9EB353F911FD3BF968" box="[534,643,1687,1713]" name="Indonesia" pageId="9" pageNumber="124">Indonesia</collectingCountry>
(
<bibRefCitation id="EF80039CB96BFF9EB3D2F911FCC0F968" author="Leclerc, P. &amp; Deharveng, L. &amp; Ng, P. K. L. &amp; Juberthie, C. &amp; Decu, V." box="[663,888,1687,1714]" pageId="9" pageNumber="124" pagination="1805 - 1823" refId="ref15379" refString="Leclerc, P., Deharveng, L., Ng, P. K. L., Juberthie, C. &amp; Decu, V. (2001) Indonesie. In: Juberthie, C. &amp; Decu, V. (Eds.), Encyclopaedia Biospeologica. Vol. 3. 1805 - 1823. Societe Internationale de Biospeologie, Moulis-Bucarest. [unknwon pagination]" type="book chapter" year="2001">
Leclerc
<emphasis id="B965A27FB96BFF9EB3B0F91EFC92F968" box="[757,810,1687,1713]" italics="true" pageId="9" pageNumber="124">et al</emphasis>
. 2001
</bibRefCitation>
), and several freshwater crabs are known from the Malaysian State of
<collectingCountry id="F3063EFDB96BFF9EB0E5F93DFDBDF90C" box="[416,517,1723,1749]" name="Malaysia" pageId="9" pageNumber="124">Sarawak</collectingCountry>
: two troglobites (
<taxonomicName id="4C1105EEB96BFF9EB388F93DFC35F90C" authorityName="Holthuis" authorityYear="1979" box="[717,909,1723,1749]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB96BFF9EB388F93DFC35F90C" box="[717,909,1723,1749]" italics="true" pageId="9" pageNumber="124">Cerberusa caeca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1105EEB96BFF9EB283F93DFB92F90C" authorityName="Holthuis" authorityYear="1979" box="[966,1066,1723,1749]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB96BFF9EB283F93DFB92F90C" box="[966,1066,1723,1749]" italics="true" pageId="9" pageNumber="124">C. tipula</emphasis>
</taxonomicName>
) in Gunung Mulu; now, another troglobite,
<taxonomicName id="4C1105EEB96BFF9EB056F959FE24F920" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[275,412,1759,1785]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="mayla" status="gen. et sp. nov.">
<emphasis id="B965A27FB96BFF9EB056F959FE24F920" box="[275,412,1759,1785]" italics="true" pageId="9" pageNumber="124">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB96BFF9EB0E1F959FDEEF920" bold="true" box="[420,598,1759,1785]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="A2561F04B96BFF9EB0E1F959FDEEF920" box="[420,598,1759,1785]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, from Merabu Karst. The potamid
<taxonomicName id="4C1105EEB96BFF9EB2A1F959FAAFF920" authority="Ng, 1989" authorityName="Ng" authorityYear="1989" box="[996,1303,1759,1785]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="styx">
<emphasis id="B965A27FB96BFF9EB2A1F959FB1BF921" box="[996,1187,1759,1785]" italics="true" pageId="9" pageNumber="124">Thelphusula styx</emphasis>
<bibRefCitation id="EF80039CB96BFF9EB5EFF959FAAFF920" author="Ng, P. K. L." box="[1194,1303,1759,1785]" pageId="9" pageNumber="124" pagination="63 - 72" refId="ref16056" refString="Ng, P. K. L. (1989) The identity of the cavernicolous freshwater crab Potamon (Thelphusa) bidiense Lanchester, 1900 (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from Sarawak, Borneo, with description of a new genus. Raffles Bulletin of Zoology, 37 (1 &amp; 2), 63 - 72." type="journal article" year="1989">Ng, 1989</bibRefCitation>
</taxonomicName>
, from Long Pala Bat Cave, western corner of Gunung Mulu National Park (
<bibRefCitation id="EF80039CB96BFF9EB236F885FC64F8C4" author="Ng, P. K. L." box="[883,988,1795,1821]" pageId="9" pageNumber="124" pagination="63 - 72" refId="ref16056" refString="Ng, P. K. L. (1989) The identity of the cavernicolous freshwater crab Potamon (Thelphusa) bidiense Lanchester, 1900 (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from Sarawak, Borneo, with description of a new genus. Raffles Bulletin of Zoology, 37 (1 &amp; 2), 63 - 72." type="journal article" year="1989">Ng 1989</bibRefCitation>
), with a pale brownish carapace, dirtycream coloured walking legs, and small but relatively well-developed eyes, with distinct black pigmentation of the cornea, could be a troglobitic species (
<bibRefCitation id="EF80039CB96BFF9EB309F8CCFCF8F8BD" author="Promdam, R. &amp; Ellis, M. &amp; Jantarit, S." box="[588,832,1866,1892]" pageId="9" pageNumber="124" pagination="1 - 17" refId="ref16973" refString="Promdam, R., Ellis, M. &amp; Jantarit, S. (2022) Brachyuran crabs (Crustacea: Decapoda) in the limestone caves of Thailand, with a checklist of freshwater cave-dwelling crabs in Southeast Asia. Edaphologia, 110, 1 - 17." type="journal article" year="2022">
Promdam
<emphasis id="B965A27FB96BFF9EB386F8CDFD44F8BD" box="[707,764,1866,1892]" italics="true" pageId="9" pageNumber="124">et al.</emphasis>
2022
</bibRefCitation>
: table 2). In contrast,
<taxonomicName id="4C1105EEB96BFF9EB57DF8CCFF6AF851" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Isolapotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="collinsi">
<emphasis id="B965A27FB96BFF9EB57DF8CCFA94F8BD" box="[1080,1324,1866,1892]" italics="true" pageId="9" pageNumber="124">Isolapotamon collinsi</emphasis>
<bibRefCitation id="EF80039CB96BFF9EB470F8CCFF6AF851" author="Holthuis, L. B." pageId="9" pageNumber="124" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
, collected from the sinkhole of the Clearwater Cave River, Hidden Valley, Gunung Mulu National Park, does not show obvious adaptations to hypogean life (
<bibRefCitation id="EF80039CB96BFF9EB3E8F817FCC3F875" author="Holthuis, L. B." box="[685,891,1937,1964]" pageId="9" pageNumber="124" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis 1979: 25</bibRefCitation>
;
<bibRefCitation id="EF80039CB96BFF9EB2C2F817FB12F872" author="McFarlane, D. &amp; Lundberg, J. &amp; Christenson, K." box="[903,1194,1937,1963]" pageId="9" pageNumber="124" pagination="1 - 18" refId="ref15641" refString="McFarlane, D., Lundberg, J. &amp; Christenson, K. (2011) New Records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of Northern Sarawak. Speleobiology Notes, 3, 1 - 18." type="journal article" year="2011">
McFarlane
<emphasis id="B965A27FB96BFF9EB542F814FB87F872" box="[1031,1087,1937,1963]" italics="true" pageId="9" pageNumber="124">et al.</emphasis>
2011: 12
</bibRefCitation>
). The same is true for
<taxonomicName id="4C1105EEB96BFF9EB1D2F833FDCDF816" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[151,629,1973,1999]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="B965A27FB96BFF9EB1D2F833FE78F816" box="[151,448,1973,1999]" italics="true" pageId="9" pageNumber="124">Sundathelphusa tenebrosa</emphasis>
<bibRefCitation id="EF80039CB96BFF9EB08CF833FDCDF816" author="Holthuis, L. B." box="[457,629,1973,1999]" pageId="9" pageNumber="124" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
, known from several cave records in Gunung Mulu National Park and with an epigean record (
<bibRefCitation id="EF80039CB96BFF9EB0E3F85FFDCDF82A" author="Holthuis, L. B." box="[422,629,2009,2035]" pageId="9" pageNumber="124" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis 1979: 42</bibRefCitation>
;
<bibRefCitation id="EF80039CB96BFF9EB3C5F85FFC1BF82A" author="McFarlane, D. &amp; Lundberg, J. &amp; Christenson, K." box="[640,931,2009,2035]" pageId="9" pageNumber="124" pagination="1 - 18" refId="ref15641" refString="McFarlane, D., Lundberg, J. &amp; Christenson, K. (2011) New Records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of Northern Sarawak. Speleobiology Notes, 3, 1 - 18." type="journal article" year="2011">
McFarlane
<emphasis id="B965A27FB96BFF9EB244F85CFC81F82A" box="[769,825,2009,2035]" italics="true" pageId="9" pageNumber="124">et al.</emphasis>
2011: 12
</bibRefCitation>
, fig. 4).
</paragraph>
</subSubSection>
</treatment>
</document>