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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.410.7488" ID-GBIF-Dataset="69a473d9-1152-4f3d-8b4e-a9de37e6f4c1" ID-PMC="PMC4042706" ID-Pensoft-Pub="1313-2970-410-23" ID-PubMed="24899848" ID-ZBK="E4C040C1939640F18C12D9518F59F668" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-410-23" ModsDocOrigin="ZooKeys 410" ModsDocTitle="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)" checkinTime="1451245876428" checkinUser="pensoft" docAuthor="Jochum, Adrienne, Slapnik, Rajko, Kampschulte, Marian, Martels, Gunhild, Heneka, Markus &amp; Pall-Gergely, Barna" docDate="2014" docId="0BAB027A65D6584D486CF60B35E959E1" docLanguage="en" docName="ZooKeys 410: 23-40" docOrigin="ZooKeys 410" docSource="http://dx.doi.org/10.3897/zookeys.410.7488" docTitle="Tonkinospira Jochum, Slapnik &amp; Pall-Gergely, nom. n." docType="treatment" docVersion="3" lastPageNumber="31" masterDocId="FFA7FFE8A6732834BC12004CAF57677C" masterDocTitle="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)" masterLastPageNumber="40" masterPageNumber="23" pageNumber="30" updateTime="1668158519503" updateUser="ExternalLinkService">
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<mods:title>A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)</mods:title>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:namePart>Slapnik, Rajko</mods:namePart>
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<mods:namePart>Kampschulte, Marian</mods:namePart>
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<mods:namePart>Martels, Gunhild</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Heneka, Markus</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>410</mods:number>
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Genus
<taxonomicName authority="Jochum, Slapnik &amp; Pall-Gergely" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="genus">
Tonkinospira Jochum, Slapnik &amp;
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="30">nom. n.</taxonomicNameLabel>
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<subSubSection pageId="7" pageNumber="30" type="reference_group">
<paragraph pageId="7" pageNumber="30">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenostoma" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="genus">Systenostoma</taxonomicName>
Bavay &amp; Dautzenberg, 1908: Journal de Conchyliologie, 56: 243.
</paragraph>
<paragraph pageId="7" pageNumber="30">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenostoma" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="genus">Systenostoma</taxonomicName>
-
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="10" pageNumber="33" pagination="163 - 206" title="Description de Coquilles Nouvelles de l'Indo-Chine." volume="57" year="1909">Bavay and Dautzenberg 1909</bibRefCitation>
: Journal de Conchyliologie, 57: 196. (diagnosis).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="30" type="remarks">
<paragraph pageId="7" pageNumber="30">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="30">
The name
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenostoma" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="genus">Systenostoma</taxonomicName>
Bavay &amp; Dautzenberg, 1908 is preoccupied (non
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenostoma" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="genus">Systenostoma</taxonomicName>
<bibRefCitation author="Marsson, TF" journalOrPublisher="Palaeontologische Abhandlungen" pageId="11" pageNumber="34" pagination="1 - 122" title="Die Bryozoen der weissen Schreibkreide der Insel Ruegen." volume="4" year="1887">Marsson 1887</bibRefCitation>
,
<taxonomicName class="Insecta" family="Cerambycidae" genus="Bryozoa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bryozoa" order="Coleoptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="genus">Bryozoa</taxonomicName>
). Therefore,
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="genus">Tonkinospira</taxonomicName>
Jochum, Slapnik &amp;
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
nom. n. is proposed as replacement.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="30" type="type species">
<paragraph pageId="7" pageNumber="30">Type species.</paragraph>
<paragraph pageId="7" pageNumber="30">
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Systenostoma) pauperrima" order="Stylommatophora" pageId="7" pageNumber="30" phylum="Mollusca" rank="species" species="pauperrima" subGenus="Systenostoma">Helix (Systenostoma) pauperrima</taxonomicName>
Bavay &amp; Dautzenberg, 1908 by subsequent designation (
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Conchological Department, Academy of Natural Sciences of Philadelphia, Philadelphia" pageId="11" pageNumber="34" title="Manual of Conchology, Second Series: Pulmonata" url="http://biodiversitylibrary.org/item/16727page/5/mode/1up" year="1916 - 1918">Pilsbry 1917</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="30" type="diagnosis">
<paragraph pageId="7" pageNumber="30">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="30">A genus of small, conical or depressed-conical species with regularly growing, rounded or angulated whorls. The sculpture is characterized by spiral lines on both the protoconch and the teleoconch, decussated by irregular radial lines resulting in a reticulated surface structure. The aperture is toothless, adnate or slightly adnate and shows a sharp peristome.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="30" type="etymology">
<paragraph pageId="7" pageNumber="30">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="30">The new name is established by the fusion of Tonkin (northern Vietnam, the area of distribution) and the Latin spira (a coil, twist). Gender: feminine.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="31" type="remarks">
<paragraph pageId="8" pageNumber="31">
<pageBreakToken pageId="8" pageNumber="31" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="8" pageNumber="31">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira" order="Stylommatophora" pageId="8" pageNumber="31" phylum="Mollusca" rank="genus">Tonkinospira</taxonomicName>
differs from
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Krobylos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krobylos" order="Stylommatophora" pageId="8" pageNumber="31" phylum="Mollusca" rank="genus">Krobylos</taxonomicName>
by the increased degree of angulation of the whorls and the lack of spiral lines on the teleoconch. For differences with
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="8" pageNumber="31" phylum="Mollusca" rank="genus">Angustopila</taxonomicName>
gen. n., see above.
</paragraph>
<paragraph pageId="8" pageNumber="31">
The systematic position of the genus is questionable. It most probably belongs to the family
<taxonomicName family="Hypselostomatidae" lsidName="" pageId="8" pageNumber="31" rank="family">Hypselostomatidae</taxonomicName>
, but its relationship with other families such as
<taxonomicName family="Helicodiscidae" lsidName="" pageId="8" pageNumber="31" rank="family">Helicodiscidae</taxonomicName>
(see
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica Supplement" pageId="11" pageNumber="34" pagination="129 - 261" title="Treatise on Recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidea, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachynodidae, Enidae, Sagdidae." volume="2" year="1998">Schileyko 1998</bibRefCitation>
) or
<taxonomicName family="Valloniidae" lsidName="" pageId="8" pageNumber="31" rank="family">Valloniidae</taxonomicName>
(see
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="11" pageNumber="34" pagination="1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." url="http://www.ruthenica.com/documents/vol21_Schileyko_1-68.pdf" volume="21" year="2011">Schileyko 2011</bibRefCitation>
) cannot be excluded.
</paragraph>
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<paragraph pageId="8" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="31">So far, the genus is reported from Northern Vietnam only.</paragraph>
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