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<document id="33DA3B4862D8BD21B98C7A6DEE15F24C" ID-CLB-Dataset="8361" ID-DOI="10.5281/zenodo.4526089" ID-GBIF-Dataset="ceb677fa-1aec-4d3c-bd12-ad41c762c044" ID-ISSN="0253-116X" ID-Zenodo-Dep="4526089" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612223888760" checkinUser="felipe" docAuthor="Gusenleitner, J., Fallahzadeh, M., Haghighi, A. &amp; Dousti, A. F." docDate="2013" docId="0B130540271AFFE1FF0BFBC1FC7BAD95" docLanguage="en" docName="LinzerbiolBeitr.45.1.0109-0116.pdf" docOrigin="Linzer biologische Beiträge 45 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Eustenancistrocerus (Parastenancistrocerus) iranicus Gusenleitner &amp; Fallahzadeh &amp; Haghighi &amp; Dousti 2013, nov.sp." docType="treatment" docVersion="8" lastPageNumber="111" masterDocId="F72A7D38271BFFE3FFA8FFF4FFF9AE5D" masterDocTitle="Two new species of Eumeninae from Iran (Hymenoptera: Vespidae)" masterLastPageNumber="116" masterPageNumber="109" pageNumber="110" updateTime="1698926294665" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4700C0A2A4929E9726D96A538FA3012B">Two new species of Eumeninae from Iran (Hymenoptera: Vespidae)</mods:title>
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<heading id="D84D033A271AFFE2FF0BFBC1FBD9AA16" box="[163,1056,1077,1099]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="110" reason="2">
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<taxonomicName id="44BACFD5271AFFE2FF0BFBC1FC90AA17" authority="GUSENLEITNER" authorityName="Gusenleitner &amp; Fallahzadeh &amp; Haghighi &amp; Dousti" authorityYear="2013" box="[163,873,1077,1099]" class="Insecta" family="Eumenidae" genus="Eustenancistrocerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov." subGenus="Parastenancistrocerus">
<emphasis id="B1CE6844271AFFE2FF0BFBC1FE8BAA17" bold="true" box="[163,370,1077,1098]" italics="true" pageId="1" pageNumber="110">Eustenancistrocerus</emphasis>
(
<emphasis id="B1CE6844271AFFE2FE28FBC1FD9AAA17" bold="true" box="[384,611,1077,1098]" italics="true" pageId="1" pageNumber="110">Parastenancistrocerus</emphasis>
)
<emphasis id="B1CE6844271AFFE2FDD8FBC1FD3AAA17" bold="true" box="[624,707,1077,1098]" italics="true" pageId="1" pageNumber="110">iranicus</emphasis>
GUSENLEITNER
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<taxonomicNameLabel id="AAFDD53F271AFFE2FCC5FBC1FC4EAA16" box="[877,951,1077,1099]" pageId="1" pageNumber="110" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1B81A8D3271AFFE2FC6BFBC1FBE1AA16" box="[963,1048,1077,1099]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[163,196,1560,1580]" captionStartId-1="7.[163,196,909,929]" captionTargetBox-0="[163,1032,922,1543]" captionTargetBox-1="[163,1070,204,892]" captionTargetId-0="figure-45@6.[163,1032,922,1543]" captionTargetId-1="figure-89@7.[163,1070,204,892]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 6: Eustenancistrocerus iranicus nov.sp. 3, habitus." captionText-1="Fig. 7-10: (7) Eustenancistrocerus iranicus nov.sp. 3, clypeus; (8) Eustenancistrocerus iranicus nov.sp. 3, last segment of flagellum; (9) Eustenancistrocerus iranicus nov.sp. 3, second and third tergites; (10) Eustenancistrocerus khuzestanicus GIORDANI SOIKA, 3, second and third tergites. Scale bar: figs. 7, 9, 10: 0,1 mm, fig. 8: 0,5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4526103" figureDoi-1="http://doi.org/10.5281/zenodo.4526105" httpUri-0="https://zenodo.org/record/4526103/files/figure.png" httpUri-1="https://zenodo.org/record/4526105/files/figure.png" pageId="1" pageNumber="110">Figs 6-9</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8305B456271AFFE2FF0BFBA9FE69AAE4" blockId="1.[163,1070,1115,1209]" pageId="1" pageNumber="110">
<materialsCitation id="33D2BE0B271AFFE2FF0BFBA9FD23AAD5" ID-GBIF-Occurrence="3033377303" collectorName="A. Falahatpishe &amp; Same" country="Iran" elevation="870" latitude="27.533333" location="Larestan" longLatPrecision="1236" longitude="54.366665" pageId="1" pageNumber="110" specimenCount="1" stateProvince="Fars">
H o l o t y p e:
<collectingCountry id="FBADF4C6271AFFE2FE87FBA9FEAFAA2C" box="[303,342,1117,1137]" name="Iran" pageId="1" pageNumber="110">Iran</collectingCountry>
,
<collectingRegion id="417E7AB4271AFFE2FEC8FBA9FE70AA2C" box="[352,393,1117,1137]" country="Iran" name="Fars" pageId="1" pageNumber="110">Fars</collectingRegion>
,
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,
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,
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, 3, 30.11.201 2, leg.
<collectorName id="2E4FD180271AFFE2FC30FBA9FBD3AA2C" box="[920,1066,1117,1137]" pageId="1" pageNumber="110">A. Falahatpishe</collectorName>
, coll. OLM. P a r a t y p e s:
<collectorName id="2E4FD180271AFFE2FE74FB81FDF7AAD4" box="[476,526,1141,1161]" pageId="1" pageNumber="110">Same</collectorName>
data as holotype, 13
</materialsCitation>
;
<materialsCitation id="33D2BE0B271AFFE2FD4FFB81FDEAAAFC" ID-GBIF-Occurrence="3033377301" collectorName="A. Haghighi" country="Iran" elevation="1124" latitude="28.7" location="Darab" longLatPrecision="1230" longitude="54.566666" pageId="1" pageNumber="110" specimenCount="1" stateProvince="Fars">
<collectingCountry id="FBADF4C6271AFFE2FD4FFB81FCF7AAD4" box="[743,782,1141,1161]" name="Iran" pageId="1" pageNumber="110">Iran</collectingCountry>
,
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,
<location id="8665E28D271AFFE2FCE6FB81FC71AAD4" LSID="urn:lsid:plazi:treatment:0B130540271AFFE1FF0BFBC1FC7BAD95:8665E28D271AFFE2FCE6FB81FC71AAD4" box="[846,904,1141,1161]" country="Iran" latitude="28.7" longLatPrecision="1230" longitude="54.566666" name="Darab" pageId="1" pageNumber="110" stateProvince="Fars">Darab</location>
,
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<elevation id="08975365271AFFE2FC3BFB81FC2DAAD4" box="[915,980,1141,1161]" metricMagnitude="3" metricUnit="m" metricValue="1.124" pageId="1" pageNumber="110" unit="m" value="1124.0">1124m</elevation>
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,
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<geoCoordinate id="E68ED291271AFFE2FF70FB79FEDFAAFC" box="[216,294,1165,1185]" degrees="54" direction="east" minutes="34" orientation="longitude" pageId="1" pageNumber="110" precision="925" value="54.566666">54°34E</geoCoordinate>
, 233, leg.
<collectorName id="2E4FD180271AFFE2FE35FB79FDEAAAFC" box="[413,531,1165,1185]" pageId="1" pageNumber="110">A. Haghighi</collectorName>
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;
<materialsCitation id="33D2BE0B271AFFE2FD88FB79FE75AAE4" ID-GBIF-Occurrence="3033377305" collectorName="Dept. Entom." country="Iran" location="Iran" pageId="1" pageNumber="110" specimenCount="1">
all in coll.
<collectorName id="2E4FD180271AFFE2FD23FB79FCF2AAFC" box="[651,779,1165,1185]" pageId="1" pageNumber="110">Dept. Entom.</collectorName>
, Jahrom Branch, Islamic Azad Univ., Jahrom,
<collectingCountry id="FBADF4C6271AFFE2FECDFB51FE75AAE4" box="[357,396,1189,1209]" name="Iran" pageId="1" pageNumber="110">Iran</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection id="CBA0E7DD271AFFE1FF0BFB3EFC7BAD95" lastPageId="2" lastPageNumber="111" pageId="1" pageNumber="110" type="description">
<paragraph id="8305B456271AFFE2FF0BFB3EFEEAABD2" blockId="1.[163,1070,1225,1423]" pageId="1" pageNumber="110">
D i a g n o s i s: This species belongs to the group of
<taxonomicName id="44BACFD5271AFFE2FCC8FB3EFCD8AAA0" authority="(DE SAUSSURE 1863)" baseAuthorityName="DE SAUSSURE" baseAuthorityYear="1863" class="Insecta" family="Eumenidae" genus="Eustenancistrocerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="amadanensis" subGenus="Parastenancistrocerus">
<emphasis id="B1CE6844271AFFE2FCC8FB3EFBD7AA82" box="[864,1070,1226,1247]" italics="true" pageId="1" pageNumber="110">Eustenancistrocerus</emphasis>
(
<emphasis id="B1CE6844271AFFE2FF03FB13FE69AAA1" box="[171,400,1255,1276]" italics="true" pageId="1" pageNumber="110">Parastenancistrocerus</emphasis>
)
<emphasis id="B1CE6844271AFFE2FE00FB13FDD5AAA1" box="[424,556,1255,1276]" italics="true" pageId="1" pageNumber="110">amadanensis</emphasis>
(DE SAUSSURE 1863)
</taxonomicName>
and is very similar to
<taxonomicName id="44BACFD5271AFFE2FF0BFAF1FC04AB47" authority="GIORDANI SOIKA 1970" authorityName="GIORDANI SOIKA" authorityYear="1970" box="[163,1021,1284,1306]" class="Insecta" family="Eumenidae" genus="Eustenancistrocerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="khuzestanicus" subGenus="Parastenancistrocerus">
<emphasis id="B1CE6844271AFFE2FF0BFAF1FE88AB47" box="[163,369,1285,1306]" italics="true" pageId="1" pageNumber="110">Eustenancistrocerus</emphasis>
(
<emphasis id="B1CE6844271AFFE2FE2BFAF1FD91AB47" box="[387,616,1285,1306]" italics="true" pageId="1" pageNumber="110">Parastenancistrocerus</emphasis>
)
<emphasis id="B1CE6844271AFFE2FDD1FAF0FCFCAB44" box="[633,773,1284,1305]" italics="true" pageId="1" pageNumber="110">khuzestanicus</emphasis>
<bibRefCitation id="E72BC9A7271AFFE2FCA7FAF0FC04AB47" author="GIORDANI SOIKA A" box="[783,1021,1284,1306]" pageId="1" pageNumber="110" pagination="27 - 183" refId="ref1648" refString="GIORDANI SOIKA A. (1970): Contributo alla conoscenza degli Eumenidi del Medio Oriente (Missione Giordani Soika in Iran 1965 III). - Boll. Mus. Civ. Stor. Nat. Venezia 20 - 21: 27 - 183." type="journal article" year="1970">GIORDANI SOIKA 1970</bibRefCitation>
</taxonomicName>
. But the species described here differs in the form of the last segments of the antennae (
<bibRefCitation id="E72BC9A7271AFFE2FF02FACAFE78AB09" author="GIORDANI SOIKA A" box="[170,385,1342,1364]" pageId="1" pageNumber="110" pagination="27 - 183" refId="ref1648" refString="GIORDANI SOIKA A. (1970): Contributo alla conoscenza degli Eumenidi del Medio Oriente (Missione Giordani Soika in Iran 1965 III). - Boll. Mus. Civ. Stor. Nat. Venezia 20 - 21: 27 - 183." type="journal article" year="1970">Giordani Soika 1970</bibRefCitation>
, p. 98) and the second tergite is simply truncate and not extendedly rounded (fig. 10). The occipital carina narrowly lamellated but in
<emphasis id="B1CE6844271AFFE2FCF1FAA9FBFDAB2C" box="[857,1028,1372,1394]" italics="true" pageId="1" pageNumber="110">P. kuzhestanicus</emphasis>
not lamellated.
</paragraph>
<paragraph id="8305B456271AFFE1FF0BFA6BFE8BAF0B" blockId="1.[163,1070,1438,1577]" lastBlockId="2.[163,1070,203,343]" lastPageId="2" lastPageNumber="111" pageId="1" pageNumber="110">
D e s c r i p t i o n:
<taxonomicName id="44BACFD5271AFFE2FEC9FA6BFE28ABE9" authorityName="Gusenleitner &amp; Fallahzadeh &amp; Haghighi &amp; Dousti" authorityYear="2013" box="[353,465,1439,1460]" class="Insecta" family="Eumenidae" genus="Eustenancistrocerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="B1CE6844271AFFE2FEC9FA6BFE28ABE9" box="[353,465,1439,1460]" italics="true" pageId="1" pageNumber="110">E. iranicus</emphasis>
</taxonomicName>
(fig. 6) is black with nearly white markings, distributed as follows: mandibles except for black teeth; labrum; clypeus; top part of face up to the ocelli; antennal scape; first and last segments of the flagellum; temples and pronotum completely; a dot on mesonotum in front of scutellum; mesopleurae nearly completely; tegulae except for two translucent dots and a translucent margin; parategulae; scutellum and postscutellum completely; two large and two small patches on each side of propodeum; legs; first tergite except for the front; an apical band on second tergite, attached to big patches at the sides; bisinuate apical bands on tergites 3 to 6; first and second sternites completely, and sternites 3 to 6 with bisinuate apical bands. Wings pale, without dusky parts.
</paragraph>
<paragraph id="8305B4562719FFE1FF0BFE92FE57ACDE" blockId="2.[163,1071,358,643]" pageId="2" pageNumber="111">Clypeus (fig. 7) wider than long (2,2: 1,7), emargination flat (length: depth = 1,5: 0,3), and broader than distance between antennal grooves (length: depth = 1,5: 1,2). Clypeus coarse and densely punctured, except for a sparsely punctured, narrow band at the base. Puncturation of frons, vertex and temples similar to that of clypeus. Antennal scapes glossy, with only a few microscopical punctures. Last segment of flagellum long and narrow (fig. 1). Pronotum, mesonotum, scutellum and mesopleurae much more coarsely punctured than face. Frontal border between horizontal and vertical part of pronotum lamellated like occipital carina. Postscutellum very finely sculptured. Propodeum with coarse punctures, only the concavity shows microscopical striation. Tegulae and legs glossy, without punctures.</paragraph>
<paragraph id="8305B4562719FFE1FF0BFD67FD74AD05" blockId="2.[163,1070,659,856]" pageId="2" pageNumber="111">Puncturation of first tergite very similar to that of scutellum; transverse carina with a central depression. Second tergite uniform and densely punctured. Third tergite densely punctured and indented in the middle (fig. 8, 10); other tergites more finely but not as densely punctured. In profile, second sternite flat, sloping steeply to the basal furrow, with dense, deep puncturation similar to that on second tergite. Tergites 3 to 7 more finely punctured than second tergite, with glossier intervals. The whole insect has a microscopical pubescense that is difficult to see.</paragraph>
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Body length:
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.
</paragraph>
<paragraph id="8305B4562719FFE1FF0BFC7AFE69ADFE" blockId="2.[163,400,909,931]" box="[163,400,909,931]" pageId="2" pageNumber="111">F e m a l e: unknown.</paragraph>
<paragraph id="8305B4562719FFE1FF0BFC47FC7BAD95" blockId="2.[163,898,946,968]" box="[163,898,946,968]" pageId="2" pageNumber="111">
E t y m o l o g y: The name is derived from the country of origin,
<collectingCountry id="FBADF4C62719FFE1FCFAFC47FC87AD95" box="[850,894,947,968]" name="Iran" pageId="2" pageNumber="111">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,176 @@
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<mods:title id="75B34B71E8E61BEACC3AA393268BB8EA">Two new species of Eumeninae from Iran (Hymenoptera: Vespidae)</mods:title>
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<heading id="D84D033A271BFFE3FF0BFB59FC5EAA99" bold="true" box="[163,935,1197,1220]" fontSize="9" level="2" pageId="0" pageNumber="109" reason="3">
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<taxonomicName id="44BACFD5271BFFE3FF0BFB59FD1EAA9F" authority="GUSENLEITNER &amp; FALLAHZADEH" authorityName="Gusenleitner &amp; Fallahzadeh &amp; Haghighi &amp; Dousti" authorityYear="2013" box="[163,743,1197,1220]" class="Insecta" family="Eumenidae" genus="Pterocheilus" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B1CE6844271BFFE3FF0BFB59FE7BAA9E" bold="true" box="[163,386,1197,1219]" italics="true" pageId="0" pageNumber="109">Pterocheilus persicus</emphasis>
GUSENLEITNER &amp; FALLAHZADEH
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(
<figureCitation id="1B81A8D3271BFFE3FCE0FB5AFC67AA99" box="[840,926,1198,1220]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[163,196,788,808]" captionStartId-1="4.[163,196,1438,1458]" captionStartId-2="5.[163,196,848,868]" captionStartId-3="5.[163,196,1538,1558]" captionTargetBox-0="[163,1070,204,768]" captionTargetBox-1="[163,1070,823,1419]" captionTargetBox-2="[163,994,204,828]" captionTargetBox-3="[163,994,896,1520]" captionTargetId-0="figure-1@4.[163,1070,204,768]" captionTargetId-1="figure-11@4.[163,1070,823,1419]" captionTargetId-2="figure-44@5.[163,994,204,828]" captionTargetId-3="figure-45@5.[163,994,896,1520]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionText-0="Fig. 1: Pterocheilus persicus nov.sp. ♀, habitus." captionText-1="Fig. 2: Pterocheilus persicus nov.sp. ♀, clypeus." captionText-2="Fig. 3: Pterocheilus persicus nov.sp., first tergite. ♀" captionText-3="Fig. 4: Pterocheilus persicus nov.sp. ♀, sternites." figureDoi-0="http://doi.org/10.5281/zenodo.4526091" figureDoi-1="http://doi.org/10.5281/zenodo.4526095" figureDoi-2="http://doi.org/10.5281/zenodo.4526097" figureDoi-3="http://doi.org/10.5281/zenodo.4526099" httpUri-0="https://zenodo.org/record/4526091/files/figure.png" httpUri-1="https://zenodo.org/record/4526095/files/figure.png" httpUri-2="https://zenodo.org/record/4526097/files/figure.png" httpUri-3="https://zenodo.org/record/4526099/files/figure.png" pageId="0" pageNumber="109">Figs 1-4</figureCitation>
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H o l o t y p e:
<collectingCountry id="FBADF4C6271BFFE3FE99FB21FEA1AAB4" box="[305,344,1237,1257]" name="Iran" pageId="0" pageNumber="109">Iran</collectingCountry>
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, 5.7.201 2, leg.
<collectorName id="2E4FD180271BFFE3FC3EFB21FBD3AAB4" box="[918,1066,1237,1257]" pageId="0" pageNumber="109">A. Falahatpishe</collectorName>
, coll. OLM.
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P a r a t y p e s:
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, Niriz,
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, 8.7.291 2, leg.
<collectorName id="2E4FD180271BFFE3FBB0FAF1FED6AB6C" pageId="0" pageNumber="109">A. Haghighi</collectorName>
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;
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both coll.
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<collectingCountry id="FBADF4C6271BFFE3FC12FAE9FC18AB6C" box="[954,993,1309,1329]" name="Iran" pageId="0" pageNumber="109">Iran</collectingCountry>
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.
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<paragraph id="8305B456271BFFE3FF0BFAB6FDA2ABF2" blockId="0.[163,1070,1346,1456]" pageId="0" pageNumber="109">
D i a g n o s i s:
<emphasis id="B1CE6844271BFFE3FEFBFAB6FDB8AB05" box="[339,577,1346,1368]" italics="true" pageId="0" pageNumber="109">Pterocheilius persicus</emphasis>
(fig. 1) is similar to
<taxonomicName id="44BACFD5271BFFE3FCCEFAB6FE79AB28" authority="RADOSZKOWSKI 1876" authorityName="RADOSZKOWSKI" authorityYear="1876" class="Insecta" family="Eumenidae" genus="Pterocheilus" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="dives">
<emphasis id="B1CE6844271BFFE3FCCEFAB6FBD7AB0A" box="[870,1070,1346,1367]" italics="true" pageId="0" pageNumber="109">Pterocheilus dives</emphasis>
RADOSZKOWSKI 1876
</taxonomicName>
(fig. 5), but differs in colouration; the pronotum is longer, the first tergite is stronger and the second sternite is more densely punctured than in
<taxonomicName id="44BACFD5271BFFE3FC01FA89FC03ABCF" authorityName="RADOSZKOWSKI" authorityYear="1876" box="[937,1018,1405,1426]" class="Insecta" family="Eumenidae" genus="Pterocheilus" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="dives">
<emphasis id="B1CE6844271BFFE3FC01FA89FC03ABCF" box="[937,1018,1405,1426]" italics="true" pageId="0" pageNumber="109">P. dives</emphasis>
</taxonomicName>
. The occipital carina is lamellated in
<taxonomicName id="44BACFD5271BFFE3FE4DFA6FFDAFABF2" authorityName="Gusenleitner &amp; Fallahzadeh &amp; Haghighi &amp; Dousti" authorityYear="2013" box="[485,598,1434,1456]" class="Insecta" family="Eumenidae" genus="Pterocheilus" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B1CE6844271BFFE3FE4DFA6FFDAFABF2" box="[485,598,1434,1456]" italics="true" pageId="0" pageNumber="109">P. persicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8305B456271BFFE2FF0BFA34FC9EAFF3" blockId="0.[163,1071,1471,1581]" lastBlockId="1.[163,1070,203,430]" lastPageId="1" lastPageNumber="110" pageId="0" pageNumber="109">D e s c r i p t i o n: Primary colour black, with yellow colouring distributed as follows: mandibles nearly completely, labrum, clypeus (except for translucent anterior margin), and orbital bands from sinus to clypeus; one small dot above the antennae; antennal scape except for apical end; big dots present on the temples and the dorsal surface of the pronotum; patches on the upper part of the mesopleurae; tegulae except for the translucent margins and the disk; two dots on scutellum; a transverse band on postscutellum; claps on the sides of the scutellum and postscutellum present to tegulae; two patches on the metanotum; legs (the base of femora III and the trochanters black except for small yellow dots); broad, laterally enlarged apical bands on first and second tergites; a bisinuate narrow band on third tergite; tergites 4 to 6 totally, and the first sternite and other sternites almost completely yellow. Three red spots present in the middle of second sternite (fig. 4). Wings pale, with dusky radial cells.</paragraph>
<paragraph id="8305B456271AFFE2FF0BFE4AFC78AC86" blockId="1.[163,1071,446,731]" pageId="1" pageNumber="110">Clypeus (fig. 2) much broader than long (5,0: 3,0) and the straight „emargination“ is longer than distance between the antennal grooves. Disc of clypeus strongly punctured; spaces between the dots are glossy and similar in size to the punctures. Clypeus covered with microscopical white pubescence. Frons very densely punctured with small spots. Vertex and temples slightly punctured. On pronotum, mesonotum, mesopleurae and scutellum the dots are coarser and fewer than on the face. The posterior part of postscutellum has no punctures. Lateral edges of pronotum with spikes. Mesonotum has a prescutal furrow (extending from scutellum to close to middle of mesonotum). Propodeum sides rounded and uniformly punctured; only the lower part of each side is finely cross-striped. Discs of tegulae and legs glossy, without punctures.</paragraph>
<paragraph id="8305B456271AFFE2FF0BFD1FFDC3AD05" blockId="1.[163,1070,746,856]" pageId="1" pageNumber="110">Dots on the first tergite (fig. 3) not as coarse as those on mesonotum; the intervals between the dots have microscopical dots. Second tergite has a few, not very coarse, microscopical dots. Tergites 3 to 6 also have fine microsopical dots. Sternites 2 to 6 glossy, with scattered punctures (fig. 4).</paragraph>
<paragraph id="8305B456271AFFE2FF0BFC9CFEE6ADC6" blockId="1.[163,1070,872,923]" pageId="1" pageNumber="110">Head and thorax with long, wavy, white hairs; abdomen with only microscopical pubescence.</paragraph>
<paragraph id="8305B456271AFFE2FF0BFC5EFE97AD9D" blockId="1.[163,366,938,960]" box="[163,366,938,960]" pageId="1" pageNumber="110">
Body length:
<quantity id="444219B3271AFFE2FE85FC5EFE93AD9D" box="[301,362,938,960]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="110" unit="mm" value="9.0">9 mm</quantity>
.
</paragraph>
<paragraph id="8305B456271AFFE2FF0BFC3BFEBAADB8" blockId="1.[163,323,975,997]" box="[163,323,975,997]" pageId="1" pageNumber="110">Male unknown.</paragraph>
<paragraph id="8305B456271AFFE2FF0BFC01FCD5AA56" blockId="1.[163,812,1013,1035]" box="[163,812,1013,1035]" pageId="1" pageNumber="110">
Etymology: The name is derived from
<collectingCountry id="FBADF4C6271AFFE2FD84FC01FD95AA56" box="[556,620,1013,1035]" name="Iran" pageId="1" pageNumber="110">Persia</collectingCountry>
, the area of origin.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,159 @@
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<mods:title id="25857965991C2847F6A1D382A4A32746">Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae)</mods:title>
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<subSubSection id="CBA06506FFFBFFB7AA7AFB2DCEC5FABF" pageId="1" pageNumber="247" type="nomenclature">
<paragraph id="8305368DFFFBFFB7AA7AFB2DCEC5FABF" blockId="1.[810,1451,1274,1372]" pageId="1" pageNumber="247">
<heading id="D84D81E1FFFBFFB7AA7AFB2DCEC5FABF" bold="true" centered="true" fontSize="10" level="2" pageId="1" pageNumber="247" reason="3">
<emphasis id="B1CEEA9FFFFBFFB7AA7AFB2DCEC5FABF" bold="true" pageId="1" pageNumber="247">
<taxonomicName id="44BA4D0EFFFBFFB7AA7AFB2DCE25FABF" authority="Li, Hsiao, Yoshitomi &amp; Cai" authorityName="Li, Hsiao, Yoshitomi &amp; Cai" authorityYear="2022" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="species" species="jarzembowskii" status="sp. nov.">
<emphasis id="B1CEEA9FFFFBFFB7AA7AFB2DC9E0FA92" bold="true" box="[810,1169,1274,1301]" italics="true" pageId="1" pageNumber="247">Homalenchodes jarzembowskii</emphasis>
Li, Hsiao, Yoshitomi &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="AAFD57E4FFFBFFB7AA0BFAC9CEC5FABF" box="[859,948,1310,1336]" pageId="1" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="CBA06506FFFBFFB7AA7AFA95CEECFADB" box="[810,925,1346,1372]" pageId="1" pageNumber="247" type="description">
<paragraph id="8305368DFFFBFFB7AA7AFA95CEECFADB" blockId="1.[810,1451,1274,1372]" box="[810,925,1346,1372]" pageId="1" pageNumber="247">
(
<figureCitation id="1B812A08FFFBFFB7AA62FA95CEE5FADB" box="[818,916,1346,1372]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[136,235,1383,1407]" captionStartId-1="3.[136,235,1699,1723]" captionStartId-2="4.[136,235,551,575]" captionTargetBox-0="[151,1436,196,1359]" captionTargetBox-1="[165,1423,168,1675]" captionTargetBox-2="[151,1436,185,527]" captionTargetId-0="figure-215@2.[151,1436,196,1359]" captionTargetId-1="figure-59@3.[165,1423,168,1675]" captionTargetId-2="figure-191@4.[151,1436,185,527]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. General habitus of Homalenchodes jarzembowskii Li, Hsiao,Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 1 mm." captionText-1="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." captionText-2="FIGURE 3. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Mesotarsus, lateral view. B, Metatarsus, dorsal view. C, Metatarsus, ventral view. Abbreviations: mst15, mesotarsomeres 15; mtt14, metatarsomeres 14. Scale bars = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.6820855" figureDoi-1="http://doi.org/10.5281/zenodo.6820858" figureDoi-2="http://doi.org/10.5281/zenodo.6820860" httpUri-0="https://zenodo.org/record/6820855/files/figure.png" httpUri-1="https://zenodo.org/record/6820858/files/figure.png" httpUri-2="https://zenodo.org/record/6820860/files/figure.png" pageId="1" pageNumber="247">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB7AA7AFA5DC9C2FA22" box="[810,1203,1418,1445]" pageId="1" pageNumber="247" type="materials_examined">
<paragraph id="8305368DFFFBFFB7AA7AFA5DC9C2FA22" blockId="1.[810,1452,1418,2093]" box="[810,1203,1418,1445]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7AA7AFA5DCEE8FA23" bold="true" box="[810,921,1418,1444]" pageId="1" pageNumber="247">Material.</emphasis>
<materialsCitation id="33D23CD0FFFBFFB7AAF0FA5DC9C2FA22" ID-GBIF-Occurrence="3862612301" box="[928,1203,1418,1445]" collectionCode="NIGP" pageId="1" pageNumber="247" specimenCode="NIGP180475" specimenCount="1" typeStatus="holotype">
<typeStatus id="5C01882FFFFBFFB7AAF0FA5DC97AFA23" box="[928,1035,1418,1444]" pageId="1" pageNumber="247" type="holotype">Holotype</typeStatus>
,
<specimenCode id="D31C9EF6FFFBFFB7AD46FA5DC9DEFA22" box="[1046,1199,1418,1445]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="247">NIGP180475</specimenCode>
.
</materialsCitation>
</paragraph>
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<subSubSection id="CBA06506FFFBFFB7AA0FFA79C82EFA6B" pageId="1" pageNumber="247" type="etymology">
<paragraph id="8305368DFFFBFFB7AA0FFA79C82EFA6B" blockId="1.[810,1452,1418,2093]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7AA0FFA79CE97FA4F" bold="true" box="[863,998,1454,1480]" pageId="1" pageNumber="247">Etymology.</emphasis>
The species is named in honor of the palaeoentomologist Dr Edmund A. Jarzembowski.
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB7AA0FFA21C824F927" pageId="1" pageNumber="247" type="diagnosis">
<paragraph id="8305368DFFFBFFB7AA0FFA21C9EFF996" blockId="1.[810,1452,1418,2093]" box="[863,1182,1526,1553]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7AA0FFA21CEA8F997" bold="true" box="[863,985,1526,1552]" pageId="1" pageNumber="247">Diagnosis.</emphasis>
As for the genus.
</paragraph>
<paragraph id="8305368DFFFBFFB7AA0FF9CDC824F927" blockId="1.[810,1452,1418,2093]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7AA0FF9CDC91CF9B3" bold="true" box="[863,1133,1562,1588]" pageId="1" pageNumber="247">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="417EF86FFFFBFFB7AA7AF9B5CEA1F9FB" box="[810,976,1634,1660]" country="Myanmar" name="Kachin" pageId="1" pageNumber="247">Kachin State</collectingRegion>
,
<collectingCountry id="FBAD761DFFFBFFB7AABFF9B4C911F9FA" box="[1007,1120,1635,1661]" name="Myanmar" pageId="1" pageNumber="247">Myanmar</collectingCountry>
; unnamed horizon, midCretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB4AA0FF97DC99AF7A2" lastPageId="2" lastPageNumber="248" pageId="1" pageNumber="247" type="description">
<paragraph id="8305368DFFFBFFB7AA0FF97DCE19F88B" blockId="1.[810,1452,1418,2093]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7AA0FF97DCE80F943" bold="true" box="[863,1009,1706,1732]" pageId="1" pageNumber="247">Description.</emphasis>
Body strongly elongate, about
<quantity id="44429B68FFFBFFB7AC05F97DC8DDF942" box="[1365,1452,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="247" unit="mm" value="2.7">2.7 mm</quantity>
long,
<quantity id="44429B68FFFBFFB7AA3AF919CEB1F96E" box="[874,960,1742,1769]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="247" unit="mm" value="0.6">0.6 mm</quantity>
wide. Surface finely punctate and with fine hairs.
</paragraph>
<paragraph id="8305368DFFFBFFB4AA0FF8C1CFE8F913" blockId="1.[810,1452,1418,2093]" lastBlockId="2.[136,778,1514,2080]" lastPageId="2" lastPageNumber="248" pageId="1" pageNumber="247">
Head well exposed from above. Compound eyes lateral, large, emarginate in front of antennal insertions, coarsely facetted on the surface, without interfacetal setae. Antennal insertions fully concealed from above, widely separated. Subantennal grooves absent. Antennae 11-segmented, long, filiform; antennomere 1 elongate, almost parallel-sided; antennomere 2 short, subquadrate; antennomeres 311 elongate, subequal. Clypeus with anterior margin convex. Mandibles bilobed apically. Maxillary palps 4-segmented, simple, not clearly serrate; palpomere 1 small; palpomere 4 possibly securiform.
<collectingRegion id="417EF86FFFF8FFB4A9D8F981CD96F9F7" box="[136,231,1622,1648]" country="Myanmar" name="Chin" pageId="2" pageNumber="248">Mentum</collectingRegion>
subtrapezoidal. Submentum relatively smooth, with hairs somewhat denser than other regions.
</paragraph>
<caption id="D7C56605FFF8FFB4A9D8FAB0CE1CFA25" ID-DOI="http://doi.org/10.5281/zenodo.6820855" ID-Zenodo-Dep="6820855" httpUri="https://zenodo.org/record/6820855/files/figure.png" pageId="2" pageNumber="248" startId="2.[136,235,1383,1407]" targetBox="[151,1436,196,1359]" targetPageId="2">
<paragraph id="8305368DFFF8FFB4A9D8FAB0CE1CFA25" blockId="2.[136,1452,1383,1443]" pageId="2" pageNumber="248">
<emphasis id="B1CEEA9FFFF8FFB4A9D8FAB0CC73FAF8" bold="true" box="[136,258,1383,1407]" pageId="2" pageNumber="248">FIGURE 1.</emphasis>
General habitus of
<taxonomicName id="44BA4D0EFFF8FFB4A893FAB0C963FAF8" authority="Li, Hsiao, Yoshitomi &amp; Cai" authorityName="Li, Hsiao, Yoshitomi &amp; Cai" authorityYear="2022" box="[451,1042,1383,1407]" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="248" phylum="Arthropoda" rank="species" species="jarzembowskii" status="gen. et sp. nov.">
<emphasis id="B1CEEA9FFFF8FFB4A893FAB0CF8CFAF8" box="[451,765,1383,1407]" italics="true" pageId="2" pageNumber="248">Homalenchodes jarzembowskii</emphasis>
Li, Hsiao, Yoshitomi &amp; Cai
</taxonomicName>
<emphasis id="B1CEEA9FFFF8FFB4AD46FAB0C9DEFAF8" bold="true" box="[1046,1199,1383,1407]" pageId="2" pageNumber="248">
<taxonomicNameLabel id="AAFD57E4FFF8FFB4AD46FAB0C9DEFAF8" box="[1046,1199,1383,1407]" pageId="2" pageNumber="248" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP180475, under incident light.
<emphasis id="B1CEEA9FFFF8FFB4A80BFA5CCC1CFA24" bold="true" box="[347,365,1419,1443]" pageId="2" pageNumber="248">A</emphasis>
, Dorsal view.
<emphasis id="B1CEEA9FFFF8FFB4AB50FA5CCF60FA24" bold="true" box="[512,529,1419,1443]" pageId="2" pageNumber="248">B</emphasis>
, Ventral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8305368DFFF8FFB4A9EDF949CD9DF85F" blockId="2.[136,778,1514,2080]" pageId="2" pageNumber="248">Pronotum widest posteriorly, with slightly produced posterior pronotal angles; disc without depression medially; posterior edge margined. Notosternal sutures distinct. Prosternum about as long as longitudinal procoxal diameter; prosternal process complete,gradually narrowed posteriorly, slightly extending beyond posterior procoxal margin, apically acute. Procoxal cavities narrowly separated, externally broadly open. Protrochantins well exposed.</paragraph>
<paragraph id="8305368DFFF8FFB4A9EDF835C991F918" blockId="2.[136,778,1514,2080]" lastBlockId="2.[810,1452,1514,2085]" pageId="2" pageNumber="248">Scutellar shield transverse, subtrapezoidal, widest posteriorly; posterior edge slightly rounded, without notch. Elytra long, about 3.1 times as long as width combined, at base almost as wide as pronotum, apically separately rounded; surface without longitudinal striae. Mesocoxal cavities narrowly separated. Metacoxae almost contiguous; coxal plates absent.</paragraph>
<paragraph id="8305368DFFF8FFB4AA0FF97AC8E1F837" blockId="2.[810,1452,1514,2085]" pageId="2" pageNumber="248">Legs slender. Protrochanter small, obliquely attached to profemur, with profemur abutting procoxa. All tibiae with 2 spurs, without transverse rows of bristles or spines. Metatibial spurs normal-sized. Tarsi 5-5-4; penultimate tarsomeres ventrally lobed in all three pairs of legs; metatarsomere 1 long, distinctly longer than metatarsomeres 24 combined. Pretarsal claws simple.</paragraph>
<paragraph id="8305368DFFF8FFB4AA0FF86AC99AF7A2" blockId="2.[810,1452,1514,2085]" pageId="2" pageNumber="248">Abdomen with five ventrites. Ratio of ventrite lengths along middle: 3.2:2.4:2.1:1.7:2.4. Ventrite 5 apically rounded, without emargination.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,237 @@
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<mods:title id="12D0226823E3C16343CD0697A698E914">Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae)</mods:title>
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<treatment id="0B13879BFFFBFFB7A9D8FBC1C96CFB4B" ID-DOI="http://doi.org/10.5281/zenodo.6828737" ID-GBIF-Taxon="197516838" ID-Zenodo-Dep="6828737" LSID="urn:lsid:plazi:treatment:0B13879BFFFBFFB7A9D8FBC1C96CFB4B" httpUri="http://treatment.plazi.org/id/0B13879BFFFBFFB7A9D8FBC1C96CFB4B" lastPageNumber="247" pageId="1" pageNumber="247">
<subSubSection id="CBA06506FFFBFFB7A9D8FBC1CD80FBD2" pageId="1" pageNumber="247" type="nomenclature">
<paragraph id="8305368DFFFBFFB7A9D8FBC1CD80FBD2" blockId="1.[136,778,1046,1109]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7A9D8FBC1CD80FBD2" bold="true" pageId="1" pageNumber="247">
Genus
<taxonomicName id="44BA4D0EFFFBFFB7A9B3FBC1CE7BFBB7" authority="Li, Hsiao, Yoshitomi &amp; Cai" authorityName="Li, Hsiao, Yoshitomi &amp; Cai" authorityYear="2022" box="[227,778,1046,1073]" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B1CEEA9FFFFBFFB7A9B3FBC1CCEFFBB6" bold="true" box="[227,414,1046,1073]" italics="true" pageId="1" pageNumber="247">Homalenchodes</emphasis>
Li, Hsiao, Yoshitomi &amp; Cai
</taxonomicName>
<heading id="D84D81E1FFFBFFB7A9D8FBECCD80FBD2" bold="true" box="[136,241,1083,1109]" fontSize="10" level="2" pageId="1" pageNumber="247" reason="3">
<taxonomicNameLabel id="AAFD57E4FFFBFFB7A9D8FBECCD80FBD2" box="[136,241,1083,1109]" pageId="1" pageNumber="247" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</emphasis>
</paragraph>
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<subSubSection id="CBA06506FFFBFFB7A9D8FB53CF99FB19" box="[136,744,1155,1182]" pageId="1" pageNumber="247" type="type_taxon">
<paragraph id="8305368DFFFBFFB7A9D8FB53CF99FB19" blockId="1.[136,778,1155,2092]" box="[136,744,1155,1182]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7A9D8FB53CC52FB1A" bold="true" box="[136,291,1155,1182]" pageId="1" pageNumber="247">
<typeStatus id="5C01882FFFFBFFB7A9D8FB53CDB2FB19" box="[136,195,1156,1182]" pageId="1" pageNumber="247">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="44BA4D0EFFFBFFB7A87AFB53CFF6FB19" authority="LI &amp; HSIAO &amp; YOSHITOMI &amp; HUANG &amp; CAI, 2022" authorityName="LI &amp; HSIAO &amp; YOSHITOMI &amp; HUANG &amp; CAI" authorityYear="2022" box="[298,647,1156,1182]" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="species" species="jarzembowskii" status="sp. nov.">
<emphasis id="B1CEEA9FFFFBFFB7A87AFB53CFF6FB19" box="[298,647,1156,1182]" italics="true" pageId="1" pageNumber="247">Homalenchodes jarzembowskii</emphasis>
</taxonomicName>
<emphasis id="B1CEEA9FFFFBFFB7ABDEFB53CF99FB19" bold="true" box="[654,744,1156,1182]" pageId="1" pageNumber="247">
<taxonomicNameLabel id="AAFD57E4FFFBFFB7ABDEFB53CF99FB19" box="[654,744,1156,1182]" pageId="1" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB7A9EDFB7FCD86FAD3" pageId="1" pageNumber="247" type="etymology">
<paragraph id="8305368DFFFBFFB7A9EDFB7FCD86FAD3" blockId="1.[136,778,1155,2092]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7A9EDFB7FCC35FB45" bold="true" box="[189,324,1192,1218]" pageId="1" pageNumber="247">Etymology.</emphasis>
The generic name is a combination of the Greek “
<emphasis id="B1CEEA9FFFFBFFB7A842FB1BCC00FB61" box="[274,369,1228,1254]" italics="true" pageId="1" pageNumber="247">homalós</emphasis>
”, smooth, referring to the absence of longitudinal striae on its elytra, and “
<taxonomicName id="44BA4D0EFFFBFFB7AB20FB26CFACFA8C" authorityName=", LeConte" authorityYear="1866" box="[624,733,1265,1291]" class="Insecta" family="Melandryidae" genus="Enchodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7AB20FB26CFACFA8C" box="[624,733,1265,1291]" italics="true" pageId="1" pageNumber="247">Enchodes</emphasis>
</taxonomicName>
”, a presumably closely related genus. The name is feminine in gender.
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB7A9EDFA89CC13F88E" pageId="1" pageNumber="247" type="discussion">
<paragraph id="8305368DFFFBFFB7A9EDFA89CC13F88E" blockId="1.[136,778,1155,2092]" pageId="1" pageNumber="247">
The generic name
<taxonomicName id="44BA4D0EFFFBFFB7A8FDFA89CFFDFAFF" authority="LeConte" authorityName=", LeConte" authorityYear="1866" box="[429,652,1374,1400]" class="Insecta" family="Melandryidae" genus="Enchodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7A8FDFA89CF6BFAFF" box="[429,538,1374,1400]" italics="true" pageId="1" pageNumber="247">Enchodes</emphasis>
LeConte
</taxonomicName>
has been sometimes mistakenly treated as masculine (
<emphasis id="B1CEEA9FFFFBFFB7ABC3FA54CFEEFA1B" box="[659,671,1411,1436]" italics="true" pageId="1" pageNumber="247">e</emphasis>
.
<emphasis id="B1CEEA9FFFFBFFB7ABF6FA54CFC5FA1B" box="[678,692,1411,1436]" italics="true" pageId="1" pageNumber="247">g</emphasis>
.,
<bibRefCitation id="E72B4B7CFFFBFFB7AB9EFA54CDB4FA46" author="Jung, B. H." pageId="1" pageNumber="247" refId="ref3517" refString="Jung, B. H. (2019) Insect fauna of Korea, Volume 12, Number 27: Tenebrionoidea - Arthropoda: Insecta: Coleoptera: Tenebrionoidea: Melandryidae &amp; Tetratomidae. National Institute of Biological Resources, Incheon, 85 pp." type="book" year="2019">Jung, 2019</bibRefCitation>
). According to Article 30.1.4.4 of ICZN (1999), “a compound genus-group name ending in the suffix -
<emphasis id="B1CEEA9FFFFBFFB7AB8CFA1BCE72FA62" box="[732,771,1484,1509]" italics="true" pageId="1" pageNumber="247">ites</emphasis>
, -
<emphasis id="B1CEEA9FFFFBFFB7A9C1FA27CDBDF98D" box="[145,204,1520,1546]" italics="true" pageId="1" pageNumber="247">oides</emphasis>
, -
<emphasis id="B1CEEA9FFFFBFFB7A9B4FA27CC60F98D" box="[228,273,1520,1546]" italics="true" pageId="1" pageNumber="247">ides</emphasis>
, -
<emphasis id="B1CEEA9FFFFBFFB7A879FA27CC2DF98D" box="[297,348,1520,1546]" italics="true" pageId="1" pageNumber="247">odes</emphasis>
, or -
<emphasis id="B1CEEA9FFFFBFFB7A8C3FA26CCB4F98D" box="[403,453,1521,1546]" italics="true" pageId="1" pageNumber="247">istes</emphasis>
is to be treated as masculine unless its author, when establishing the name, … treated it as such by combining it with an adjectival species-group name in another gender form”. When establishing genus
<taxonomicName id="44BA4D0EFFFBFFB7A9D8F955CCB4F91C" authority=", LeConte (1866)" authorityName=", LeConte" authorityYear="1866" box="[136,453,1665,1692]" class="Insecta" family="Melandryidae" genus="Enchodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7A9D8F955CD84F91B" box="[136,245,1666,1692]" italics="true" pageId="1" pageNumber="247">Enchodes</emphasis>
,
<bibRefCitation id="E72B4B7CFFFBFFB7A85BF955CCB4F91C" author="LeConte, J. L." box="[267,453,1665,1692]" pageId="1" pageNumber="247" pagination="87 - 168" refId="ref3562" refString="LeConte, J. L. (1866) New species of North American Coleoptera. Prepared for the Smithsonian Institution. Part I. Smithsonian Miscellaneous Collections, 6 (167), 87 - 168. https: // doi. org / 10.5962 / bhl. title. 17758" type="journal article" year="1866">LeConte (1866)</bibRefCitation>
</taxonomicName>
added a single previously described species,
<taxonomicName id="44BA4D0EFFFBFFB7A80AF970CFF8F947" authority="Haldeman" authorityName="Haldeman" box="[346,649,1702,1728]" class="Insecta" family="Melandryidae" genus="Dircaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="species" species="sericea">
<emphasis id="B1CEEA9FFFFBFFB7A80AF970CF7DF947" box="[346,524,1703,1728]" italics="true" pageId="1" pageNumber="247">Dircaea sericea</emphasis>
Haldeman
</taxonomicName>
, citing it as
<taxonomicName id="44BA4D0EFFFBFFB7A9D8F91DCCAFF963" authority="(Haldeman)" baseAuthorityName="Haldeman" baseAuthorityYear="1848" box="[136,478,1738,1764]" class="Insecta" family="Melandryidae" genus="Enchodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="species" species="sericea">
<emphasis id="B1CEEA9FFFFBFFB7A9D8F91DCC3CF963" box="[136,333,1738,1764]" italics="true" pageId="1" pageNumber="247">Enchodes sericea</emphasis>
(Haldeman)
</taxonomicName>
. Thus, the name should be feminine in gender.
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB7A9EDF8C4C978FEEB" pageId="1" pageNumber="247" type="diagnosis">
<paragraph id="8305368DFFFBFFB7A9EDF8C4C978FEEB" blockId="1.[136,778,1155,2092]" lastBlockId="1.[810,1452,158,1228]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7A9EDF8C4CC49F8AA" bold="true" box="[189,312,1811,1837]" pageId="1" pageNumber="247">Diagnosis.</emphasis>
Body relatively small, strongly elongate. Compound eyes coarsely facetted (
<figureCitation id="1B812A08FFFBFFB7AB41F8EFCFFAF8D5" box="[529,651,1848,1874]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2A, B</figureCitation>
). Antennae long, filiform. Maxillary palps simple (
<figureCitation id="1B812A08FFFBFFB7AB62F88BCFFBF8F1" box="[562,650,1884,1910]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2A</figureCitation>
). Pronotum widest posteriorly, with somewhat produced posterior pronotal angles (
<figureCitation id="1B812A08FFFBFFB7A81CF872CCD9F838" box="[332,424,1957,1983]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2B</figureCitation>
). Prosternal process complete, gradually narrowed posteriorly, slightly extending beyond posterior procoxal margin (
<figureCitation id="1B812A08FFFBFFB7A891F839CF6CF78F" box="[449,541,2030,2056]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2A</figureCitation>
). Procoxae narrowly separated (
<figureCitation id="1B812A08FFFBFFB7A857F7C5CC17F7AB" box="[263,358,2066,2092]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2A</figureCitation>
). Protrochantins well exposed (
<figureCitation id="1B812A08FFFBFFB7AB8CF7C5CE21FF3F" box="[732,848,158,2092]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2A</figureCitation>
). Elytra without longitudinal striae (
<figureCitation id="1B812A08FFFBFFB7ADA3FF49C83FFF3F" box="[1267,1358,158,184]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2E</figureCitation>
). Meso- and metatibiae without transverse rows of bristles or spines (
<figureCitation id="1B812A08FFFBFFB7AA2EFF31CEA6FE87" box="[894,983,230,256]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2D</figureCitation>
). Metatibial spurs normal-sized (distinctly shorter than half length of metatarsomere 1) (
<figureCitation id="1B812A08FFFBFFB7AC11FEDDC8EEFEA3" box="[1345,1439,266,292]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Fig. 2D</figureCitation>
). Penultimate tarsomeres ventrally lobed in all three pairs of legs (
<figureCitation id="1B812A08FFFBFFB7AAD4FE85CE91FEEB" box="[900,992,338,364]" captionStart="FIGURE 2" captionStartId="3.[136,235,1699,1723]" captionTargetBox="[165,1423,168,1675]" captionTargetId="figure-59@3.[165,1423,168,1675]" captionTargetPageId="3" captionText="FIGURE 2. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Head and prothorax, ventral view. B, Head and prothorax, dorsal view. C, Abdominal base and fore leg. D, Abdominal apex and metatibia, ventral view. E, Elytral apex, dorsal view. Abbreviations: an19, antennomeres 19; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mtc, metacoxa; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; ptcn, protrochantin; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6820858" httpUri="https://zenodo.org/record/6820858/files/figure.png" pageId="1" pageNumber="247">Figs 2C</figureCitation>
,
<figureCitation id="1B812A08FFFBFFB7AABBFE85CE8AFEEB" box="[1003,1019,338,364]" captionStart="FIGURE 3" captionStartId="4.[136,235,551,575]" captionTargetBox="[151,1436,185,527]" captionTargetId="figure-191@4.[151,1436,185,527]" captionTargetPageId="4" captionText="FIGURE 3. Details of Homalenchodes jarzembowskii Li, Hsiao, Yoshitomi &amp; Cai gen. et sp. nov., holotype, NIGP180475, under confocal microscopy. A, Mesotarsus, lateral view. B, Metatarsus, dorsal view. C, Metatarsus, ventral view. Abbreviations: mst15, mesotarsomeres 15; mtt14, metatarsomeres 14. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6820860" httpUri="https://zenodo.org/record/6820860/files/figure.png" pageId="1" pageNumber="247">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CBA06506FFFBFFB7AA0FFEA1C96CFB4B" pageId="1" pageNumber="247" type="discussion">
<paragraph id="8305368DFFFBFFB7AA0FFEA1C96CFB4B" blockId="1.[810,1452,158,1228]" pageId="1" pageNumber="247">
<emphasis id="B1CEEA9FFFFBFFB7AA0FFEA1CEA2FE17" bold="true" box="[863,979,374,400]" pageId="1" pageNumber="247">Remarks.</emphasis>
<taxonomicName id="44BA4D0EFFFBFFB7AA8BFEA1C9FEFE17" authorityName="Li, Hsiao, Yoshitomi &amp; Cai" authorityYear="2022" box="[987,1167,374,400]" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7AA8BFEA1C9FEFE17" box="[987,1167,374,400]" italics="true" pageId="1" pageNumber="247">Homalenchodes</emphasis>
</taxonomicName>
shares with
<taxonomicName id="44BA4D0EFFFBFFB7AC4FFEA1C8DAFE17" authorityName="Lacordaire" authorityYear="1859" box="[1311,1451,374,400]" class="Insecta" family="Synchroidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="family">Synchroidae</taxonomicName>
a similar elongate habitus, lateral and oval eyes, and transverse head and pronotum. In fact, the adults of
<taxonomicName id="44BA4D0EFFFBFFB7AA7AFE35CEC7FE7B" authorityName="Lacordaire" authorityYear="1859" box="[810,950,482,508]" class="Insecta" family="Synchroidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="family">Synchroidae</taxonomicName>
have an appearance similar to
<taxonomicName id="44BA4D0EFFFBFFB7AC59FE35C8D6FE7B" authorityName="Leach" authorityYear="1815" box="[1289,1447,482,508]" class="Insecta" family="Melandryidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="family">Melandryidae</taxonomicName>
, but these two families are otherwise readily separable based on larval characters (
<bibRefCitation id="E72B4B7CFFFBFFB7AD03FDFDC873FDC3" author="Crowson, R. A." box="[1107,1282,554,580]" pageId="1" pageNumber="247" pagination="507 - 513" refId="ref3075" refString="Crowson, R. A. (1966) Observations on the constitution and subfamilies of the family Melandryidae (Coleoptera). Eos: Revista Espanola de Entomologia, 41, 507 - 513." type="journal article" year="1966">Crowson, 1966</bibRefCitation>
). Nevertheless, synchroids are generally characterised by the long interfacetal setae (absent in
<taxonomicName id="44BA4D0EFFFBFFB7AD29FDA5C849FD0A" authority="Horn" authorityName="Horn" authorityYear="1888" box="[1145,1336,626,653]" class="Insecta" family="Synchroidae" genus="Mallodrya" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7AD29FDA5C99EFD0B" box="[1145,1263,626,652]" italics="true" pageId="1" pageNumber="247">Mallodrya</emphasis>
Horn
</taxonomicName>
, although this genus might have been inappropriately placed in
<taxonomicName id="44BA4D0EFFFBFFB7AA7AFD6DCEC7FD53" authorityName="Lacordaire" authorityYear="1859" box="[810,950,698,724]" class="Insecta" family="Synchroidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="family">Synchroidae</taxonomicName>
and needs to be further evaluated;
<bibRefCitation id="E72B4B7CFFFBFFB7AC3AFD6DCEC8FD7F" author="Hsiao, Y. &amp; Konvicka, O. &amp; Ko, C. C." pageId="1" pageNumber="247" pagination="1 - 33" refId="ref3459" refString="Hsiao, Y., Konvicka, O. &amp; Ko, C. C. (2018) The world fauna of Synchroidae Lacordaire, 1859 (Coleoptera, Tenebrionoidea, Synchroidae). European Journal of Taxonomy, 407, 1 - 33. https: // doi. org / 10.5852 / ejt. 2018.407" type="journal article" year="2018">
Hsiao
<emphasis id="B1CEEA9FFFFBFFB7AA7AFD08CE13FD7F" box="[810,866,734,760]" italics="true" pageId="1" pageNumber="247">et al</emphasis>
., 2018
</bibRefCitation>
), unmargined posterior edge of pronotum (distinctly margined in
<taxonomicName id="44BA4D0EFFFBFFB7AD15FCD5C9CAFC9B" authorityName="Horn" authorityYear="1888" box="[1093,1211,770,796]" class="Insecta" family="Synchroidae" genus="Mallodrya" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7AD15FCD5C9CAFC9B" box="[1093,1211,770,796]" italics="true" pageId="1" pageNumber="247">Mallodrya</emphasis>
</taxonomicName>
), prosternal process markedly extending beyond procoxae with long oval apex (prosternal process laminate, ending just before the posterior edges of procoxae in
<taxonomicName id="44BA4D0EFFFBFFB7ADBBFCB9C810FC0F" authorityName="Horn" authorityYear="1888" box="[1259,1377,878,904]" class="Insecta" family="Synchroidae" genus="Mallodrya" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7ADBBFCB9C810FC0F" box="[1259,1377,878,904]" italics="true" pageId="1" pageNumber="247">Mallodrya</emphasis>
</taxonomicName>
), and unlobed penultimate tarsomeres (
<bibRefCitation id="E72B4B7CFFFBFFB7ADF7FC43CE17FC57" author="Slipinski, A. &amp; Lawrence, J. F." pageId="1" pageNumber="247" pagination="667 - 669" refId="ref4050" refString="Slipinski, A. &amp; Lawrence, J. F. (2010) Synchroidae Lacordaire, 1859. In: Leschen, R. A. B., Beutel, R. G., Lawrence, J. F. (Eds), Handbook of zoology, Arthropoda: Insecta, Coleoptera, beetles, Vol. 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Walter de Gruyter, Berlin, pp. 667 - 669. https: // doi. org / 10.1515 / 9783110911213.667" type="book chapter" year="2010">Ślipiński &amp; Lawrence, 2010</bibRefCitation>
;
<bibRefCitation id="E72B4B7CFFFBFFB7AA23FC61C934FC57" author="Hsiao, Y. &amp; Konvicka, O. &amp; Ko, C. C." box="[883,1093,950,977]" pageId="1" pageNumber="247" pagination="1 - 33" refId="ref3459" refString="Hsiao, Y., Konvicka, O. &amp; Ko, C. C. (2018) The world fauna of Synchroidae Lacordaire, 1859 (Coleoptera, Tenebrionoidea, Synchroidae). European Journal of Taxonomy, 407, 1 - 33. https: // doi. org / 10.5852 / ejt. 2018.407" type="journal article" year="2018">
Hsiao
<emphasis id="B1CEEA9FFFFBFFB7AAEEFC60CE83FC57" box="[958,1010,950,976]" italics="true" pageId="1" pageNumber="247">et al</emphasis>
., 2018
</bibRefCitation>
), while
<taxonomicName id="44BA4D0EFFFBFFB7ADF3FC61C826FC57" authorityName="Li, Hsiao, Yoshitomi &amp; Cai" authorityYear="2022" box="[1187,1367,950,976]" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7ADF3FC61C826FC57" box="[1187,1367,950,976]" italics="true" pageId="1" pageNumber="247">Homalenchodes</emphasis>
</taxonomicName>
has no interfacetal setae, a margined posterior edge of pronotum, a posteriorly gradually narrowed prosternal process with acute apex, slightly extending beyond posterior edges of procoxae, and lobed penultimate tarsomeres. Therefore, here we believe that it is appropriate to place
<taxonomicName id="44BA4D0EFFFBFFB7AA7AFB59CEAFFB2F" authorityName="Li, Hsiao, Yoshitomi &amp; Cai" authorityYear="2022" box="[810,990,1166,1192]" class="Insecta" family="Melandryidae" genus="Homalenchodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B1CEEA9FFFFBFFB7AA7AFB59CEAFFB2F" box="[810,990,1166,1192]" italics="true" pageId="1" pageNumber="247">Homalenchodes</emphasis>
</taxonomicName>
in the (currently non-monophyletic) family
<taxonomicName id="44BA4D0EFFFBFFB7AA2AFB65C968FB4B" authorityName="Leach" authorityYear="1815" box="[890,1049,1202,1228]" class="Insecta" family="Melandryidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="247" phylum="Arthropoda" rank="family">Melandryidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,208 @@
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Borko, Spela</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fuchs, Andreas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Altermatt, Florian</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>760</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>113</mods:start>
<mods:end>141</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.760.24978</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.760.24978</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-760-113</mods:identifier>
<mods:identifier type="ZBK">5C2A6C152AE3499DB71DED5C8685B05F</mods:identifier>
<mods:identifier type="ZooBank">5C2A6C152AE3499DB71DED5C8685B05F</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="144148287" LSID="urn:lsid:zoobank.org:act:E1C7C812-1494-40A8-8844-C6DC45C7AF07" httpUri="http://treatment.plazi.org/id/0B13BDF0D53AEE29AB2D50CC492E39E0" lastPageNumber="113" pageId="0" pageNumber="113">
<subSubSection pageId="0" pageNumber="113" type="nomenclature">
<paragraph pageId="0" pageNumber="113">
<taxonomicName LSID="http://zoobank.org/E1C7C812-1494-40A8-8844-C6DC45C7AF07" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Niphargus luchoffmanni" order="Amphipoda" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="luchoffmanni">Niphargus luchoffmanni</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="113">sp. n.</taxonomicNameLabel>
Figs 3, 4, 5, 6, 7, 8
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="holotype">
<paragraph pageId="0" pageNumber="113">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="113">
Male, 6.7 mm. The specimen is mounted on two slides and deposited in the collection of the the
<normalizedToken originalValue="Musée">Musee</normalizedToken>
de Zoologie, Lausanne, Switzerland under voucher numbers GBIFCH00585716 and GBIFCH00585717. Sampled on May 29, 2014 by Verena
<taxonomicName lsidName="" pageId="0" pageNumber="113" rank="tribe" tribe="Lubini">Lubini</taxonomicName>
. Additional paratypes include 9.15 mm long and partially dissected female deposited under voucher number GBIFCH00587519, a male 6.8 mm long deposited under voucher number GBIFCH00587518 and several un-dissected specimens deposited in vials under GBIFCH00329353 and GBIFCH00329354.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="materials_examined">
<paragraph pageId="0" pageNumber="113">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="113">Two males of lengths 6.7 and 6.8 mm and a female 9.15 mm long; specimens are partially dissected and mounted on slides with voucher numbers GBIFCH00585716, GBIFCH00585717, GBIFCH00587518 and GBIFCH00587519; seven other specimens were sequenced.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="type locality">
<paragraph pageId="0" pageNumber="113">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="113">Marchbachquelle, Wolfenschiessen, Switzerland (CH1903: 672,490/190,300).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="diagnosis">
<paragraph pageId="0" pageNumber="113">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="113">
Mid-sized species, in general appearance similar to
<taxonomicName lsidName="N. forelii" pageId="0" pageNumber="113" rank="species" species="forelii">N. forelii</taxonomicName>
. Epimeral plates angular. Telson with three long apical spines, one lateral, and one dorsal spine per lobe. Propods of gnathopods I and II of rectangular shape, propodus of gnathopod II large when compared to body length and propodus I. Maxilla outer lobe with seven spiniform setae, the inner four comb-like with long subapical denticles, the remaining three spines with one such denticle. Uropod I inner ramus slightly shorter than outer ramus; uropod II inner ramus slightly longer than outer ramus. Uropod III distal article elongated in males, as long as 0.5 times proximal article.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="description">
<paragraph pageId="0" pageNumber="113">Description</paragraph>
<paragraph pageId="0" pageNumber="113">
(based on dissected specimens).Head and trunk (Fig. 3). Body length up to 9.2 mm. Head length approximately 10% of body length; rostrum absent. Pereonites
<normalizedToken originalValue="IVI">I-VI</normalizedToken>
without setae, pereonite VII with one seta ventro-posteriorly.
</paragraph>
<paragraph pageId="0" pageNumber="113">
Pleonites
<normalizedToken originalValue="IIII">I-III</normalizedToken>
with up to four setae along the entire respective dorso-posterior margins. Epimeral plate II only slightly inclined, posterior and ventral margins slightly convex; ventro-postero-distal corner distinct; two spines along ventral margin; three to six setae along posterior margin. Epimeral plate III inclined, posterior and ventral margin concave and convex, respectively; ventro-postero-distal corner distinct but not produced; two to three spiniform setae along ventral margin; four to five thin setae along posterior margin.
</paragraph>
<paragraph pageId="0" pageNumber="113">Urosomite I postero-dorso-laterally with one slender and flexible seta; urosomite II postero-dorso-laterally with one strong spiniform setae accompanied with one slender and flexible seta; urosomite III without setae. At the base of uropod I, a single strong spiniform seta.</paragraph>
<paragraph pageId="0" pageNumber="113">Telson length: width ratio is 1: 0.81-0.91; cleft is 0.69-0.72 telson length; telson margins straight and narrowing apically. Telson spiniform setae (per lobe, left-right lobe asymmetry commonly observed): three apical, one dorsal and one lateral spiniform; mesial setae were not observed. Apical spiniform setae 0.44-0.5 telson length. Pairs of plumose setae laterally.</paragraph>
<paragraph pageId="0" pageNumber="113">Antennae (Fig. 4). Antenna I 0.41-0.56 times body length. Flagellum with 17-20 articles; each article with one aesthetasc. Peduncle articles in ratio 1: 0.79-0.87: 0.37-0.47. Proximal article of peduncle dorso-distally slightly produced. Accessory flagellum biarticulated; distal article shorter than one quarter of proximal article length.</paragraph>
<paragraph pageId="0" pageNumber="113">Length ratio antenna I: antenna II is 1: 0.48-0.57. Flagellum of antenna II with nine to ten articles; each article with setae and elongated sensillae of unknown function. Peduncle articles lengths 4: 5 is 1: 0.93-0.95; flagellum 0.69-0.77 of length of peduncle articles 4 and 5.</paragraph>
<paragraph pageId="0" pageNumber="113">Mouthparts (Fig. 5). Labrum typical; inner lobes of labium hardly visible.</paragraph>
<paragraph pageId="0" pageNumber="113">Left mandible: incisor with five teeth, lacinia mobilis with four teeth; between lacinia and molar a row of serrated setae, few spatulate setae and a long seta at the base of molar. Right mandible: incisor processus with four teeth, lacinia mobilis with several small teeth, between lacinia and molar a row of thick serrated setae. Ratio of mandibular palp article 2: article 3 (distal) is 1: 1.01-1.11. Proximal palp article without setae; the second article with seven to eleven setae; distal article with a group of two to four A setae; two to three groups of B setae; 15-20 D setae and three E setae.</paragraph>
<paragraph pageId="0" pageNumber="113">Maxilla I distal palp article with five to six apical setae. Outer lobe of maxilla I with a row of seven stout setae, inner four comb-like, with many long subapical denticles, the remaining three setae with one denticle; inner lobe with two to three apical setae.</paragraph>
<paragraph pageId="0" pageNumber="113">Maxilla II inner lobe slightly smaller than outer lobe; both lobes setose apically.</paragraph>
<paragraph pageId="0" pageNumber="113">Maxilliped palp article 2 with seven to eight rows of setae along inner margin; distal article with a dorsal seta, and setae at the base of nail. Maxilliped outer lobe with nine to eleven stout setae mesially to subapically, and three to five setae apically; inner lobe apically with three to four stout setae and seven serrated setae.</paragraph>
<paragraph pageId="0" pageNumber="113">
Coxal plates, and gills (Figs 3, 6, 7). Coxal plate I of parallelogram shape, with rounded antero-ventral corner and armed with four to six setae. Coxal plates II-IV width: depth ratios as 0.87-1.07: 1, 1.03-1.12: 1 and 0.96-1.13: 1; anterior and ventral margins with four to seven, five to six and five to seven setae. Coxal plate IV posteriorly distinctly concave. Coxal plates
<normalizedToken originalValue="VVI">V-VI</normalizedToken>
anteriorly with large lobes; posterior margins with one seta. Coxal plate VII half-rounded shaped with one posterior seta. Gills
<normalizedToken originalValue="IIVI">II-VI</normalizedToken>
ovoid.
</paragraph>
<paragraph pageId="0" pageNumber="113">Gnathopod I (Fig. 6). Ischium with one group of four to six postero-distal setae. Carpus 0.58-0.61 of basis length and 0.84-0.97 of propodus length; broadened distally. Carpus with single distal group of setae anteriorly, rarely accompanied by an additional seta in the mid of article; transverse rows of setae along posterior margin and a row of setae postero-laterally. Propodus rectangular. Along posterior margin, three to five rows of setae. Anterior margin with two to three groups of total four to eleven setae in addition to antero-distal group of eight setae. Several groups of short setae on the inner surface present. Palmar corner armed with a long spiniform palmar seta, two to three serrated spiniform seta, a single supporting spiniform seta on inner surface and three to five long setae below palmar spine. Palm setose. Nail length 0.31-0.32 of total dactylus length; four setae along anterior margin; a row of short setae along inner margin.</paragraph>
<paragraph pageId="0" pageNumber="113">Gnathopod II (Figs 6). Basis width: length is 0.28-0.30: 1. Ischium with three to four postero-distal setae. Carpus 0.52-0.57 of basis length and 0.83-0.91 of propodus length, distally broadened. Carpus with distal group of setae anteriorly, rarely accompanied by an additional seta in the middle of the article; few transverse rows of setae along posterior margin and a row of setae postero-laterally. Propodus rectangular, large (circumference measures up to 0.20-0.23 of body length), much larger than propodus of gnathopod I (I: II as 0.75-0.76: 1). Posterior margin with six to eight rows of setae. Anterior margin with two to three groups of total four to six setae in addition to seven to ten antero-distal setae. Individual surface setae present. Palmar corner with one strong palmar spiniform seta, a single supporting spiniform seta on inner surface and two denticulated thick-spiniform setae on outer side. Palm setose, below spiniform palmar seta, a group of three to four long setae. Nail length 0.30-0.32 of total dactylus length; three to six setae along anterior margin; a few short setae along inner margin.</paragraph>
<paragraph pageId="0" pageNumber="113">
Pereopods
<normalizedToken originalValue="IIIIV">III-IV</normalizedToken>
(Fig. 7): Lengths of pereopods III and IV subequal. Dactylus IV 0.46-0.52 of propodus IV; nail length 0.52-0.59 of total dactylus length. Dactyli III-IV with a dorsal plumose seta; one spiniform seta at the base of nail, sometimes accompanied by a tiny seta.
</paragraph>
<paragraph pageId="0" pageNumber="113">
Pereopods
<normalizedToken originalValue="VVII">V-VII</normalizedToken>
(Fig. 7): Lengths of pereopods V: VI: VII is 1: 1.33-1.34: 1.39-1.42; pereopod VII measures 0.55-0.59 of body length.
</paragraph>
<paragraph pageId="0" pageNumber="113">
Bases V-VII broad, respective length: width ratios as 1: 0.64-0.67, 1: 0.60-0.65 and 1: 0.60-0.63; posterior margins straight to convex; bases V-VII with moderate posterior lobes; posteriorly eight to eleven, nine to twelve and seven to ten setae, respectively; anteriorly six to seven, six and five to seven groups of spines, respectively. Dactyli
<normalizedToken originalValue="VVII">V-VII</normalizedToken>
with dorsal plumose seta; spiniform seta at the base of nail, in most cases accompanied by one tiny seta.
</paragraph>
<paragraph pageId="0" pageNumber="113">
Pleopods and uropods (Fig. 8): Pleopods
<normalizedToken originalValue="IIII">I-III</normalizedToken>
with two hooked retinacles. Pleopod II rami with seven to nine and nine to ten articles.
</paragraph>
<paragraph pageId="0" pageNumber="113">Uropod I protopodite with three to six dorso-lateral spiniform setae and three to four dorso-medial spiniform setae. Exopodite: endopodite lengths is 1: 0.82-0.99; rami straight. Endopodite with two individual spiniform setae laterally and five spiniform setae apically. Exopodite with two to four groups totalling three to eight setae; mesial groups comprise individual spiniform setae, whereas lateral groups comprise spiniform and flexible setae groups; apically five spiniform setae.</paragraph>
<paragraph pageId="0" pageNumber="113">Uropod II exopodite: endopodite lengths is 1: 1.02-1.12.</paragraph>
<paragraph pageId="0" pageNumber="113">Uropod III rod-shaped, 0.22-0.41 of body length. Protopodite with one to two lateral setae and six to seven apical spiniform setae. Endopodite 0.45-0.50 of protopodite length, laterally with 0-1 seta, apically with two setae, at least one spiniform. Exopodite of uropod III distal article 0.28-0.48 of the proximal article length. Proximal article with four to six groups of thin-flexible, spiniform and plumose setae along inner margin and four groups of thin-flexible and spiniform setae along outer margin. Distal article with one to four groups of thin-flexible setae along each margin, and five to six of setae apically.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="etymology">
<paragraph pageId="0" pageNumber="113">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="113">
The species is named in honour of Hans Lukas
<normalizedToken originalValue="“Luc”">&quot;Luc&quot;</normalizedToken>
Hoffmann (1923-2016), naturalist and ecologists, who importantly influenced nature conservation worldwide. Among others, he was the founder of the MAVA foundation and co-founder of the World Wide Fund for Nature (WWF).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="habitat">
<paragraph pageId="0" pageNumber="113">Habitat and distribution.</paragraph>
<paragraph pageId="0" pageNumber="113">The species has been hitherto reported from springs, and seems to be endemic to central Switzerland (Fig. 1).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="variability">
<paragraph pageId="0" pageNumber="113">Variability.</paragraph>
<paragraph pageId="0" pageNumber="113">
The variability of the species is poorly understood, as we could analyse relatively little material, with numerous sub-adult and damaged specimens. Males and females differ in length of distal article of uropod III, which is remarkably longer in males. Larger specimens tend to have narrower bases of pereopods
<normalizedToken originalValue="VVII">V-VII</normalizedToken>
. The pattern of denticulation on spines on outer lobe of maxilla I is, however, stable and the most important diagnostic trait.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="remarks">
<paragraph pageId="0" pageNumber="113">Remarks and affiliation.</paragraph>
<paragraph pageId="0" pageNumber="113">
In a morphological sense,
<taxonomicName lsidName="N. luchoffmanni" pageId="0" pageNumber="113" rank="species" species="luchoffmanni">N. luchoffmanni</taxonomicName>
sp. n. shows some similarities to
<taxonomicName lsidName="N. forelii" pageId="0" pageNumber="113" rank="species" species="forelii">N. forelii</taxonomicName>
Humbert, 1876. We compared
<taxonomicName lsidName="N. luchoffmanni" pageId="0" pageNumber="113" rank="species" species="luchoffmanni">N. luchoffmanni</taxonomicName>
sp. n. with neotypes from Bodensee from Berlin Museum and species descriptions (
<bibRefCitation author="Karaman, GS" journalOrPublisher="Bolletino Museo Civico di Storia naturale Verona" pageId="0" pageNumber="113" pagination="57 - 68" title="Niphargusforelii Humbert, 1876 and its taxonomic position." volume="17" year="1993">Karaman and Ruffo 1993</bibRefCitation>
,
<bibRefCitation pageId="0" pageNumber="113">Ginet 1996</bibRefCitation>
). Both species have long dactyls, long telson spines, multiple setae along gnathopod dactyls, an elongated distal article of uropod III in males, and the endopodite of uropod I shorter than the exopodite. Yet, there are few distinct traits separating both species. The differences in sizes of propods of gnathopods I and II is more pronounced in
<taxonomicName lsidName="N. luchoffmanni" pageId="0" pageNumber="113" rank="species" species="luchoffmanni">N. luchoffmanni</taxonomicName>
sp. n. than in
<taxonomicName lsidName="N. forelii" pageId="0" pageNumber="113" rank="species" species="forelii">N. forelii</taxonomicName>
. In addition, endopodite of the uropod II is longer and shorter than the exopodite in
<taxonomicName lsidName="N. luchoffmanni" pageId="0" pageNumber="113" rank="species" species="luchoffmanni">N. luchoffmanni</taxonomicName>
sp. n. and
<taxonomicName lsidName="N. forelii" pageId="0" pageNumber="113" rank="species" species="forelii">N. forelii</taxonomicName>
, respectively.
<taxonomicName class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Niphargus luchoffmanni" order="Amphipoda" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="luchoffmanni">Niphargus luchoffmanni</taxonomicName>
sp. n. has one dorsal spine on telson, while
<taxonomicName lsidName="N. forelii" pageId="0" pageNumber="113" rank="species" species="forelii">N. forelii</taxonomicName>
is lacking dorsal telson spines. Finally, the spines on the outer lobe of maxilla I are different: while
<taxonomicName lsidName="N. luchoffmanni" pageId="0" pageNumber="113" rank="species" species="luchoffmanni">N. luchoffmanni</taxonomicName>
sp. n. has at least four spines multidenticulate,
<taxonomicName lsidName="N. forelii" pageId="0" pageNumber="113" rank="species" species="forelii">N. forelii</taxonomicName>
has, at most, one such spine.
</paragraph>
</subSubSection>
</treatment>
</document>