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<mods:title id="EB48B68B3B792F08B98C45577C9031CD">Review of Oweniidae Rioja, 1917 (Annelida, Palaeoannelida) from Malaysia, with a description of two new species and a key to South China Sea species</mods:title>
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<mods:title id="79E55D3848E81D05183550C3A74A6870">Review of Oweniidae Rioja, 1917 (Annelida, Palaeoannelida) from Malaysia, with a description of two new species and a key to South China Sea species</mods:title>
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<mods:namePart id="10E7E08DFF3B7F61737107AC83BB93CC">Ibrahim, Nur Fazne</mods:namePart>
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<mods:affiliation id="863B4F743794D0B8966E996528BCC4CC">Institute of Oceanography and Environment (INOS), Universiti Malaysia Terengganu, 21030, Kuala Nerus, Terengganu (Malaysia)</mods:affiliation>
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<mods:affiliation id="2727A5F1F644433B6CA6A50EA54BF6E8">Departamento Ecología Marina, Centro de Investigación Científica y de Educación Superior de Ensenada, 22860, Ensenada, Baja California (Mexico)</mods:affiliation>
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<mods:namePart id="2BD1E736FD676D90E34A9386B7677683">Villalobos-Guerrero, Tulio F.</mods:namePart>
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<mods:affiliation id="B81CB0F1A6062D13F4E5CC9B1AEAE843">South China Sea Repository and Reference Centre (RRC), Institute of Oceanography and Environment, Universiti Malaysia Terengganu (INOS), 21030, Kuala Nerus, Terengganu (Malaysia)</mods:affiliation>
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@ -68,7 +71,7 @@
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(
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<figureCitation id="132B2A0CF5338E6DFE20FF5A4CD62074" box="[403,463,247,273]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3</figureCitation>
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<figureCitation id="132B2A0CF5338E6DFE20FF5A4CD62074" box="[403,463,247,273]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3</figureCitation>
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A-J)
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@ -247,33 +250,33 @@ Complete specimen, elongated body, cylindrical in cross-section, BL:
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long, BW:
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<quantity id="4CE89B6CF5338E6DFE5EFB994F7E252A" box="[493,615,1076,1103]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="520" unit="mm" value="0.065">0.065 mm</quantity>
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, 15 segments. Creamy-white colour in alcohol, with no pigmentation. Body divided into three sections: tentacular crown, thorax and abdomen. Short, anteriorly cylindrical, posteriorly tapering and dorsoventrally flattened with two lobes (
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<figureCitation id="132B2A0CF5338E6DFD36FB194FE325AB" box="[645,762,1204,1230]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3A, D</figureCitation>
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<figureCitation id="132B2A0CF5338E6DFD36FB194FE325AB" box="[645,762,1204,1230]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3A, D</figureCitation>
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). Fragile mid and posterior body, tightly attached to tube by hooks. Short tentacular crown (C/UL ratio
|
||||
<emphasis id="B964EA9BF5338E6DFDD5FB594F692468" box="[614,624,1268,1293]" italics="true" pageId="9" pageNumber="520">c</emphasis>
|
||||
. 0.5), narrow (C/UW ratio
|
||||
<emphasis id="B964EA9BF5338E6DFEAFFAB94C3F2448" box="[284,294,1300,1325]" italics="true" pageId="9" pageNumber="520">c</emphasis>
|
||||
. 0.75), with three pairs of tentacles, slightly longer dorsal-most pair; each tentacle with short, subdistal ramification that slightly lengthens toward dorsal tentacles (
|
||||
<figureCitation id="132B2A0CF5338E6DFF3FFADE4C1B24E8" box="[140,258,1395,1421]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3B, C</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFF3FFADE4C1B24E8" box="[140,258,1395,1421]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3B, C</figureCitation>
|
||||
), except mid-ventral paired tentacle proximally with isolated and digitiform ramification (
|
||||
<figureCitation id="132B2A0CF5338E6DFDE0FA3E4FBE24C8" box="[595,679,1427,1453]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3B</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFDE0FA3E4FBE24C8" box="[595,679,1427,1453]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3B</figureCitation>
|
||||
); second dichotomy located very close to distal end, with two or three smooth lobe-like tips, with oral grooves present at the third pair of tentacles (dorsal side). Dorsally and ventrally split tentacular crown. Distinct separation between left and right paired tentacles, marked V-shaped slit at mid-ventral side (
|
||||
<figureCitation id="132B2A0CF5338E6DFF72F9FF4C232709" box="[193,314,1618,1644]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3B, C</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFF72F9FF4C232709" box="[193,314,1618,1644]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3B, C</figureCitation>
|
||||
). Two brown long elliptical eyespots present lateroventrally at base of crown (
|
||||
<figureCitation id="132B2A0CF5338E6DFDAEF9DF4F8A27E9" box="[541,659,1650,1676]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3B, C</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFDAEF9DF4F8A27E9" box="[541,659,1650,1676]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3B, C</figureCitation>
|
||||
). Junction between thorax and tentacle crown marked by short, barely membranous, straight collar, closely aligned with body wall, without covering eyespots. First and third segments half as long as third segment (RLUS): 1-2-1. Uniramous segments with three pairs of poorly developed notopodia with capillary chaetae. First and second segments present in a larger number of chaetae and shorter, at ventrolateral position; third and subsequent segments with fewer and longer chaetae located at dorsolateral side, closer to the first biramous notopodia (tori and capillary chaeta). Notochaetae with shafts composed of densely packed scales (
|
||||
<figureCitation id="132B2A0CF5338E6DFDAFF87C4F6A268E" box="[540,627,2001,2027]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3E</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFDAFF87C4F6A268E" box="[540,627,2001,2027]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3E</figureCitation>
|
||||
). Uniramous neuropodia absent. Presence of constriction on 5th segment. Width of biramous segment decreasing posteriorly, posterior segments short, compact, flattened and tapering off. Poorly developed notopodia in biramous parapodia with capillary chaeta similar to uniramous notochaetae, displaced more dorsally. Gold-coloured notochaetae bundle present from 8th segment toward posterior end (
|
||||
<figureCitation id="132B2A0CF5338E6DFB37FE3B49C720D4" box="[1156,1246,406,433]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3D</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFB37FE3B49C720D4" box="[1156,1246,406,433]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3D</figureCitation>
|
||||
). Neuropodial tori from the 4th segment, continue toward posterior end; tori wide, with several rows of neurochaetal hooks. Long, narrow and nearly rectangular tori, almost encircling the body, separated only by the notochaetal bundles on dorsal surface (
|
||||
<figureCitation id="132B2A0CF5338E6DFC87FD9B4E962335" box="[820,911,566,592]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3G</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFC87FD9B4E962335" box="[820,911,566,592]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3G</figureCitation>
|
||||
). First two biramous segments longer than total uniramous segment length. Longest segment 6th. Similar neurochaetal uncini throughout, consisting of hooks with shaft ending in two teeth, with left teeth bending toward the right teeth, concave side of teeth bearing approximately 45° angles with three bends per tip (
|
||||
<figureCitation id="132B2A0CF5338E6DFB36FD7849CF2395" box="[1157,1238,725,752]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3F</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFB36FD7849CF2395" box="[1157,1238,725,752]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3F</figureCitation>
|
||||
). Longitudinal lines present at mid ventral (two lines) position, from uniramous region to posterior part. Pygidium flattened with two lobes. Methylene blue stain concentrated at lateral side of collar and less concentrated at lateral side of uniramous region and longitudinal line (
|
||||
<figureCitation id="132B2A0CF5338E6DFC5BFCD8492222EA" box="[1000,1083,885,911]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3C</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFC5BFCD8492222EA" box="[1000,1083,885,911]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3C</figureCitation>
|
||||
). Absence of staining at mid-dorsal and ventral collar and thorax (
|
||||
<figureCitation id="132B2A0CF5338E6DFBC0FC3849D022CA" box="[1139,1225,917,943]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3C</figureCitation>
|
||||
<figureCitation id="132B2A0CF5338E6DFBC0FC3849D022CA" box="[1139,1225,917,943]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3C</figureCitation>
|
||||
). Tube short, and tapered posteriorly; made of overlapping exogenous materials of quartz grains and shell fragments (outer layer), cemented together and arranged like roof tiles (
|
||||
<figureCitation id="132B2A0CF5338E6DFB08FC59480D256A" box="[1211,1300,1012,1039]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." pageId="9" pageNumber="520">Fig. 3H</figureCitation>
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<figureCitation id="132B2A0CF5338E6DFB08FC59480D256A" box="[1211,1300,1012,1039]" captionStart="FIG" captionStartId="10.[132,143,1894,1912]" captionTargetBox="[135,1455,216,1847]" captionTargetId="figure-27@10.[700,1268,759,1304]" captionTargetPageId="10" captionText="FIG. 3. — Owenia unipinnata n. sp. holotype, UMTAnn 2186: A, complete full body of individual, with posterior part slightly bend towards the body; B, anterior end, ventral view, stained using rose bengal (microscopic image) with schematic diagram (completed with numbered arrows) of tentacular crown showing simple solitary ramification; C, anterior end, ventral (left) and dorsal (right) views, stained using methylene blue; D, flattened posterior end with broken lobes and gold coloured notochaeta; E, capillary chaeta with densely packed scales; F, hooked uncini with left teeth bend towards the right teeth; G, tori almost encircles the body; H, overlapping quartz grains and shell fragments of tube. Number 1, 2, 3, indicate the number of ramifications. Scale bars: A, 1 mm; B, C, D, G, H, 100 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13712336" httpUri="https://zenodo.org/record/13712336/files/figure.png" pageId="9" pageNumber="520">Fig. 3H</figureCitation>
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). Inner sheath consisting of a translucent successive layers, attached tightly to the neurochaetal hooks.
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</paragraph>
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(
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<figureCitation id="132B2A0CF53F8E61FBFDF8BF49972649" box="[1102,1166,1810,1836]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="5" pageNumber="516">Fig. 2</figureCitation>
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<figureCitation id="132B2A0CF53F8E61FBFDF8BF49972649" box="[1102,1166,1810,1836]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="5" pageNumber="516">Fig. 2</figureCitation>
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)
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<caption id="DF6F6601F53C8E62FF36FBD84FAE2584" ID-DOI="http://doi.org/10.5281/zenodo.13712330" ID-Zenodo-Dep="13712330" httpUri="https://zenodo.org/record/13712330/files/figure.png" pageId="6" pageNumber="517" startId="6.[133,143,1141,1159]" targetBox="[214,1335,215,1096]" targetPageId="6" targetType="figure">
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<paragraph id="8BAF3689F53C8E62FF36FBD84FAE2584" blockId="6.[132,1457,1140,1249]" pageId="6" pageNumber="517">
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<smallCapsWord id="8D49A055F53C8E62FF36FBD84D8425E0" baselines="1154,1154" box="[133,157,1141,1159]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="517">FIG</smallCapsWord>
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. 1. — Sampling positions along the coastal defence structure areas of Kuala Nerus, Terengganu:
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@ -297,23 +300,23 @@ Short, slender, thread-like body, cylindrical in cross-section, BL:
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, BW:
|
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<quantity id="4CE89B6CF53C8E62FEF4F81C4CA926A9" box="[327,432,1969,1996]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="517" unit="mm" value="0.02">0.02 mm</quantity>
|
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, 18 segments. Brownish colour in alcohol, slightly transparent, with some brownish pigmented spots encircling segment 4 immediately after tori (
|
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<figureCitation id="132B2A0CF53C8E62FC85FA9E4E922428" box="[822,907,1331,1357]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="6" pageNumber="517">Fig. 2A</figureCitation>
|
||||
<figureCitation id="132B2A0CF53C8E62FC85FA9E4E922428" box="[822,907,1331,1357]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="6" pageNumber="517">Fig. 2A</figureCitation>
|
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). Elongated head, truncated prostomium, terminal mouth extending mid-ventrally as an elongated slit (
|
||||
<figureCitation id="132B2A0CF53C8E62FAFCFAFE48BB2408" box="[1359,1442,1363,1389]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="6" pageNumber="517">Fig. 2A</figureCitation>
|
||||
<figureCitation id="132B2A0CF53C8E62FAFCFAFE48BB2408" box="[1359,1442,1363,1389]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="6" pageNumber="517">Fig. 2A</figureCitation>
|
||||
). Two ventrolateral brownish eyespots. First three anterior segments uniramous, each with notochaeta only. First and second segments twice as long as third segment (RLUS 2:2:1). White oesophageal commissures, ventral longitudinal band, Y-shaped in anterior segment, extending to segment 5. Anterior biramous segments starting from 4th segment throughout, three times longer than wide, segment 7th being longest; following ones becoming posteriorly slightly tapered and shorter, far posterior ones much compressed (
|
||||
<figureCitation id="132B2A0CF53C8E62FC85F93F4E9627C9" box="[822,911,1682,1708]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="6" pageNumber="517">Fig. 2B</figureCitation>
|
||||
<figureCitation id="132B2A0CF53C8E62FC85F93F4E9627C9" box="[822,911,1682,1708]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="6" pageNumber="517">Fig. 2B</figureCitation>
|
||||
). Capillary notochaetae on both uniramous and biramous segments decreasing in size ventrally, basally slightly striated, median and distal regions with spinules; typically, more abundant in uniramous segments (3-9 capillary chaetae) than in biramous ones (3-4 capillary chaetae), the latter with far longer capillary chaetae posteriorly (
|
||||
<figureCitation id="132B2A0CF53C8E62FA91F89C486F2629" box="[1314,1398,1841,1868]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="6" pageNumber="517">Fig. 2E</figureCitation>
|
||||
<figureCitation id="132B2A0CF53C8E62FA91F89C486F2629" box="[1314,1398,1841,1868]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="6" pageNumber="517">Fig. 2E</figureCitation>
|
||||
) but single in last three segments. Ventrolateral neurochaetae from segment 4, transverse rows of hooked uncini in long, broad tori (
|
||||
<figureCitation id="132B2A0CF53C8E62FC1FF83C493826CE" box="[940,1057,1937,1963]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="6" pageNumber="517">Fig. 2F, G</figureCitation>
|
||||
<figureCitation id="132B2A0CF53C8E62FC1FF83C493826CE" box="[940,1057,1937,1963]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="6" pageNumber="517">Fig. 2F, G</figureCitation>
|
||||
). Tori along anterior to middle segments longer than wide, with dense oblique rows of uncini (
|
||||
<figureCitation id="132B2A0CF53C8E62FC87F87C4E96268E" box="[820,911,2001,2027]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="6" pageNumber="517">Fig. 2G</figureCitation>
|
||||
<figureCitation id="132B2A0CF53C8E62FC87F87C4E96268E" box="[820,911,2001,2027]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="6" pageNumber="517">Fig. 2G</figureCitation>
|
||||
); in posterior segments (segment 14 and above) much shorter and smaller, with fewer uncini (less than 20), lying in rows forming oval fields. Bidentate, long-shafted, numerous uncini (around 100) in mid-body (6th to 7th) segments, fewer toward posterior end. Long teeth of uncini, nearly equal in size, arranged obliquely because one offset higher (
|
||||
<figureCitation id="132B2A0CF53D8E63FF69FEDA4C5420F4" box="[218,333,375,401]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="7" pageNumber="518">Fig. 2F, G</figureCitation>
|
||||
<figureCitation id="132B2A0CF53D8E63FF69FEDA4C5420F4" box="[218,333,375,401]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="7" pageNumber="518">Fig. 2F, G</figureCitation>
|
||||
). Pygidium with terminal anus and two blunt lobes (
|
||||
<figureCitation id="132B2A0CF53D8E63FEBDFE3B4C9E20D4" box="[270,391,406,433]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="7" pageNumber="518">Fig. 2B, D</figureCitation>
|
||||
<figureCitation id="132B2A0CF53D8E63FEBDFE3B4C9E20D4" box="[270,391,406,433]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="7" pageNumber="518">Fig. 2B, D</figureCitation>
|
||||
). Tube slightly longer than animal, slender, cylindrical in cross-section, composed of thin secreted layer, ornamented with sand grains and fragments of shells (
|
||||
<figureCitation id="132B2A0CF53D8E63FF5EFE5B4C702375" box="[237,361,502,528]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." pageId="7" pageNumber="518">Fig. 2C, D</figureCitation>
|
||||
<figureCitation id="132B2A0CF53D8E63FF5EFE5B4C702375" box="[237,361,502,528]" captionStart="FIG" captionStartId="8.[133,144,1962,1980]" captionTargetBox="[134,1455,216,1847]" captionTargetId="figure-19@8.[134,1455,216,1848]" captionTargetPageId="8" captionText="FIG. 2. — Galathowenia minuta n. sp. holotype, UMTAnn 2184: A, anterior end, dorsal view; B, posterior end, lateral view; C, grain sand cemented tube; midbody part; D, grain sand cemented tube; posterior end; E, posterior chaetigers with long capillary chaeta; lateral view; F, transverse rows of hooked uncini; mid body; G, hooked uncini in oblique position mid body. Abbreviation: S, segment. Scale bars: A, B, 200 µm; C, D, E, 100 µm; F, G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13712334" httpUri="https://zenodo.org/record/13712334/files/figure.png" pageId="7" pageNumber="518">Fig. 2C, D</figureCitation>
|
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).
|
||||
</paragraph>
|
||||
<paragraph id="8BAF3689F53D8E63FF37FD9A4DE5232A" blockId="7.[130,777,567,975]" box="[132,252,567,593]" pageId="7" pageNumber="518">
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