182 lines
21 KiB
XML
182 lines
21 KiB
XML
<document id="1CA85378DCCC224D86998E6831D10F1A" ID-DOI="10.11646/zootaxa.5492.2.6" ID-ISSN="1175-5326" ID-ZooBank="4C877712-B5C4-451A-87BE-B0C08B2BEC24" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722841366327" checkinUser="plazi" docAuthor="Zamani, Alireza, Khudhur, Farhad A. & Marusik, Yuri M." docDate="2024" docId="03E7FE0B5152FFEEFF6514BAFAC8775B" docLanguage="en" docName="zootaxa.5492.1.6.pdf" docOrigin="Zootaxa 5492 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zelotes hazarmerdensis Zamani & Khudhur & Marusik 2024, sp. n." docType="treatment" docVersion="1" lastPageNumber="269" masterDocId="FFDE8673515BFFE7FFF21641FFA37152" masterDocTitle="New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records" masterLastPageNumber="278" masterPageNumber="260" pageNumber="269" updateTime="1722841495305" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="61F9DF7CD19BFD88789633E58262CD41">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:name id="B5DBCFFA03D99AD40FEB6D221208DF70" type="personal">
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<mods:namePart id="0F17780E6B6CECD10AC5097748729058">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="5BC1341A6EBE9BAE98E1D4A22052B98B">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku 20500, Finland</mods:affiliation>
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<mods:nameIdentifier id="250E85473B868873CACE21CB82CDC787" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="15CB0AACFC1278BAD2E50F677C2CB56B">Khudhur, Farhad A.</mods:namePart>
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<mods:nameIdentifier id="FA004EC11276FD55F7FB647A3EF72C39" type="ORCID">0000-0001-5267-6334</mods:nameIdentifier>
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<mods:affiliation id="F997BDC003E407101BC0F8B902D07A83">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq & University of Mendel, Brno, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="5A209B54C1CDF2A6E1DDEED164186F9E" type="email">qi.ude.lusvinu@ruhduhk.dahraf</mods:nameIdentifier>
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<mods:namePart id="C808E600AC8D8101E365087FF130AE2F">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="AE30B3021EA4E707AA96A777AF33E48D" type="ORCID">0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation id="5278F25EA6D288E147E365195B43E712">Institute for Biological Problems of the North FEB RAS, Magadan, Russia & Altai State University, Barnaul, Russia & Department of Zoology & Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<mods:nameIdentifier id="1EDAB2B0A425F682998474E29802240F" type="email">yurmar@mail.ru</mods:nameIdentifier>
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<mods:title id="5CD122C976270774FFC5D7C436D06C81">Zootaxa</mods:title>
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<mods:part id="347F492D99BCBFA3AABB5EC391BDE180">
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<mods:date id="07BA8857F3DFCD4EAD751CA98047A1B0">2024</mods:date>
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<mods:number id="307AE5A0BA598AD3FC376CD4C44CF6A1">2024-08-05</mods:number>
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<mods:number id="02D888EEE78429F39ED09AB4D06D6B67">5492</mods:number>
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<mods:url id="DDA72DE452915F76A2D32D0CD24F9F42">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:classification id="7921153E396B4E050EE54723A8E8CA3C">journal article</mods:classification>
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<treatment id="03E7FE0B5152FFEEFF6514BAFAC8775B" LSID="urn:lsid:plazi:treatment:03E7FE0B5152FFEEFF6514BAFAC8775B" httpUri="http://treatment.plazi.org/id/03E7FE0B5152FFEEFF6514BAFAC8775B" lastPageNumber="269" pageId="9" pageNumber="269">
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<subSubSection id="C3541C965152FFEEFF6514BAFCA37247" box="[151,768,763,790]" pageId="9" pageNumber="269" type="nomenclature">
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<paragraph id="8BF14F1D5152FFEEFF6514BAFCA37247" blockId="9.[151,768,763,826]" box="[151,768,763,790]" pageId="9" pageNumber="269">
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<heading id="D0B9F8715152FFEEFF6514BAFCA37247" bold="true" box="[151,768,763,790]" fontSize="11" level="1" pageId="9" pageNumber="269" reason="1">
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<emphasis id="B93A930F5152FFEEFF6514BAFCA37247" bold="true" box="[151,768,763,790]" pageId="9" pageNumber="269">
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<taxonomicName id="4C4E349E5152FFEEFF6514BAFD127247" authority="Zamani & Marusik" authorityName="Zamani & Khudhur & Marusik" authorityYear="2024" box="[151,689,763,790]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="269" phylum="Arthropoda" rank="species" species="hazarmerdensis" status="sp. nov.">
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<emphasis id="B93A930F5152FFEEFF6514BAFE177244" bold="true" box="[151,436,763,790]" italics="true" pageId="9" pageNumber="269">Zelotes hazarmerdensis</emphasis>
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Zamani & Marusik
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2092E745152FFEEFD4C14BAFCA37247" box="[702,768,763,789]" pageId="9" pageNumber="269" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3541C965152FFEEFF65155EFEE3726B" box="[151,320,799,826]" pageId="9" pageNumber="269" type="description">
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<paragraph id="8BF14F1D5152FFEEFF65155EFEE3726B" blockId="9.[151,768,763,826]" box="[151,320,799,826]" pageId="9" pageNumber="269">
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<figureCitation id="137553985152FFEEFF65155EFF537268" box="[151,240,799,826]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–F, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (A–F, H, I) and frontal (G). A–C, F–G, I males; D, E, H females." pageId="9" pageNumber="269">Figs 1D</figureCitation>
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,
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<figureCitation id="137553985152FFEEFF0D155EFEE3726B" box="[255,320,799,825]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="9" pageNumber="269">6B–C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541C965152FFEEFF651526FCB772F7" pageId="9" pageNumber="269" type="materials_examined">
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<paragraph id="8BF14F1D5152FFEEFF651526FCB772F7" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
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<emphasis id="B93A930F5152FFEEFF651526FEE972D3" bold="true" box="[151,330,871,897]" pageId="9" pageNumber="269">Type material.</emphasis>
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<materialsCitation id="3B2645405152FFEEFEA41526FCB772F7" collectingDate="2022-03-28" collectionCode="ZMUT" collectorName="F. A. Khudhur" country="Iraq" latitude="35.49789" location="Hazar Merd" longLatPrecision="1" longitude="45.31561" municipality="Bakrajo" pageId="9" pageNumber="269" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah" typeStatus="holotype">
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<typeStatus id="54F5F1BF5152FFEEFEA41526FE1D72D3" box="[342,446,871,897]" pageId="9" pageNumber="269" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D4884945152FFEEFE381529FE7C72D0" box="[458,479,872,898]" count="1" pageId="9" pageNumber="269" type="female">♀</specimenCount>
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(
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<collectionCode id="ED5FD7D85152FFEEFE001526FDE472D3" box="[498,583,871,897]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="9" pageNumber="269">ZMUT</collectionCode>
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),
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<collectingCountry id="F3590F8D5152FFEEFDAD1526FD0B72D3" box="[607,680,871,897]" name="Iraq" pageId="9" pageNumber="269">IRAQ</collectingCountry>
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:
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<emphasis id="B93A930F5152FFEEFD451526FC5272D0" box="[695,1009,871,898]" italics="true" pageId="9" pageNumber="269">
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<collectingRegion id="498A81FF5152FFEEFD451526FC5272D0" box="[695,1009,871,898]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
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</emphasis>
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||
:
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<collectingMunicipality id="6B95D5675152FFEEFBF71526FBC272D3" box="[1029,1121,871,897]" pageId="9" pageNumber="269">Bakrajo</collectingMunicipality>
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,
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<location id="8E9119C65152FFEEFB821529FB5C72D3" LSID="urn:lsid:plazi:treatment:03E7FE0B5152FFEEFF6514BAFAC8775B:8E9119C65152FFEEFB821529FB5C72D3" box="[1136,1279,871,898]" country="Iraq" latitude="35.49789" longLatPrecision="1" longitude="45.31561" municipality="Bakrajo" name="Hazar Merd" pageId="9" pageNumber="269" stateProvince="As Sulaymaniyah">Hazar Merd</location>
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, Dekon Vill.,
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<geoCoordinate id="EE7A29DA5152FFEEFF6515CAFE9372F4" box="[151,304,907,934]" degrees="35" direction="north" minutes="29" orientation="latitude" pageId="9" pageNumber="269" precision="1" seconds="52.4" value="35.49789">35°29'52.4"N</geoCoordinate>
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<geoCoordinate id="EE7A29DA5152FFEEFEC515CAFE6C72F4" box="[311,463,907,934]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="9" pageNumber="269" precision="1" seconds="56.2" value="45.31561">45°18'56.2"E</geoCoordinate>
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,
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<date id="FFF069DD5152FFEEFE2915CAFDFA72F7" box="[475,601,907,933]" pageId="9" pageNumber="269" value="2022-03-28">
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<collectingDate id="EFB490355152FFEEFE2915CAFDFA72F7" box="[475,601,907,933]" pageId="9" pageNumber="269" value="2022-03-28">28.03.2022</collectingDate>
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</date>
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(
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<collectorName id="26BB2ACB5152FFEEFD9A15CAFCAA72F7" box="[616,777,907,933]" pageId="9" pageNumber="269">F.A. Khudhur</collectorName>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541C965152FFEEFF3515EEFB3672BF" pageId="9" pageNumber="269" type="etymology">
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<paragraph id="8BF14F1D5152FFEEFF3515EEFB3672BF" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
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<emphasis id="B93A930F5152FFEEFF3515EEFEED729B" bold="true" box="[199,334,943,969]" pageId="9" pageNumber="269">Etymology.</emphasis>
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The specific epithet refers to Hazar Merd ancient archeological cave within Baranan mountain chain in
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<collectingRegion id="498A81FF5152FFEEFF0A1592FD8D72BF" box="[248,558,979,1005]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
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, near which the
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<typeStatus id="54F5F1BF5152FFEEFD171595FCB672BC" box="[741,789,980,1006]" pageId="9" pageNumber="269">type</typeStatus>
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locality of this species is situated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541C965152FFEEFF3515B6FDD8752F" pageId="9" pageNumber="269" type="diagnosis">
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<paragraph id="8BF14F1D5152FFEEFF3515B6FDD8752F" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
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<emphasis id="B93A930F5152FFEEFF3515B6FEE17543" bold="true" box="[199,322,1015,1041]" pageId="9" pageNumber="269">Diagnosis.</emphasis>
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The new species is similar to
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<taxonomicName id="4C4E349E5152FFEEFD5115B9FC4A7543" authority="Chatzaki, 2010" authorityName="Chatzaki" authorityYear="2010" box="[675,1001,1015,1042]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="269" phylum="Arthropoda" rank="species" species="helicoides">
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||
<emphasis id="B93A930F5152FFEEFD5115B9FD107543" box="[675,691,1016,1041]" italics="true" pageId="9" pageNumber="269">Z</emphasis>
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.
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<emphasis id="B93A930F5152FFEEFD3015B6FC977543" box="[706,820,1015,1041]" italics="true" pageId="9" pageNumber="269">helicoides</emphasis>
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<bibRefCitation id="EFDF32EC5152FFEEFCCF15B6FC4A7543" author="Chatzaki, M." box="[829,1001,1015,1041]" pageId="9" pageNumber="269" pagination="43 - 61" refId="ref8257" refString="Chatzaki, M. (2010) New data on the least known zelotines (Araneae, Gnaphosidae) of Greece and adjacent regions. Zootaxa, 2564 (1), 43 - 61. https: // doi. org / 10.11646 / zootaxa. 2564.1.3" type="journal article" year="2010">Chatzaki, 2010</bibRefCitation>
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</taxonomicName>
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from
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<collectingCountry id="F3590F8D5152FFEEFBC015B6FB227543" box="[1074,1153,1015,1041]" name="Greece" pageId="9" pageNumber="269">Greece</collectingCountry>
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by having large anterior hoods, but can be distinguished by the relatively larger receptacles and converging lateral margins of the epigynal fovea (vs. subparallel), as well as by the receptacle span being wider than the anterior pockets (vs. subequal) (cf.
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<figureCitation id="137553985152FFEEFF651222FEAF752F" box="[151,268,1123,1149]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="9" pageNumber="269">Fig. 6B–C</figureCitation>
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||
and
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<bibRefCitation id="EFDF32EC5152FFEEFEB01222FE4A752F" author="Chatzaki, M." box="[322,489,1123,1149]" pageId="9" pageNumber="269" pagination="43 - 61" refId="ref8257" refString="Chatzaki, M. (2010) New data on the least known zelotines (Araneae, Gnaphosidae) of Greece and adjacent regions. Zootaxa, 2564 (1), 43 - 61. https: // doi. org / 10.11646 / zootaxa. 2564.1.3" type="journal article" year="2010">Chatzaki 2010</bibRefCitation>
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||
: figs 22, 23).
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C3541C965152FFEEFF3512C6FE2674B7" pageId="9" pageNumber="269" type="description">
|
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<paragraph id="8BF14F1D5152FFEEFF3512C6FAD47463" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
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<emphasis id="B93A930F5152FFEEFF3512C6FEFA75F3" bold="true" box="[199,345,1159,1185]" pageId="9" pageNumber="269">Description.</emphasis>
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||
Female. Habitus as in
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<figureCitation id="137553985152FFEEFDA412C6FD0975F0" box="[598,682,1159,1186]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A–F, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (A–F, H, I) and frontal (G). A–C, F–G, I males; D, E, H females." pageId="9" pageNumber="269">Fig. 1D</figureCitation>
|
||
. Total length 5.70. Carapace 2.44 long, 1.95 wide. Eye sizes: AME: 0.08, ALE: 0.12, PME: 0.09, PLE: 0.08. Carapace, chelicerae, maxillae, labium and sternum black. Legs proximally black and distally light brown, without annulations. Abdomen dark gray, without any pattern. Spinnerets slightly darker than abdomen, uniform in color. Measurements of legs: I: Fe: 1.63, Pa: 1.04 (rest of segments missing), II: 4.65 (1.37, 0.86, 0.95, 0.80, 0.67), III: 4.44 (1.26, 0.76, 0.79, 0.99, 0.64), IV: 6.09 (1.73, 0.94, 1.22, 1.47, 0.73).
|
||
</paragraph>
|
||
<paragraph id="8BF14F1D5152FFEEFF35137AFF587493" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
|
||
Epigyne as in
|
||
<figureCitation id="137553985152FFEEFE9A137AFE427407" box="[360,481,1339,1365]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="9" pageNumber="269">Fig. 6B–C</figureCitation>
|
||
; epigynal plate ca. 1.3 times longer than wide; fovea (
|
||
<emphasis id="B93A930F5152FFEEFBB6137DFBC07407" box="[1092,1123,1340,1365]" italics="true" pageId="9" pageNumber="269">Fo</emphasis>
|
||
) ovate, ca. 1.4 times longer than wide; anterior hoods (
|
||
<emphasis id="B93A930F5152FFEEFE35131EFE45742B" box="[455,486,1375,1401]" italics="true" pageId="9" pageNumber="269">Ah</emphasis>
|
||
) broad, subequal to half width of fovea, spaced by less than their width; receptacles (
|
||
<emphasis id="B93A930F5152FFEEFF5313C5FF1C74CF" box="[161,191,1412,1437]" italics="true" pageId="9" pageNumber="269">Re</emphasis>
|
||
) large, globular, their diameters equal to width of fovea, spaced by less than one radius; accessorial glands (
|
||
<emphasis id="B93A930F5152FFEEFA8613C2FA3074CF" box="[1396,1427,1411,1437]" italics="true" pageId="9" pageNumber="269">Ag</emphasis>
|
||
) globular.
|
||
</paragraph>
|
||
<paragraph id="8BF14F1D5152FFEEFF35138AFE2674B7" blockId="9.[151,1437,871,1546]" box="[199,389,1483,1509]" pageId="9" pageNumber="269">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3541C965152FFEEFF3513AEFAC8775B" box="[199,1387,1519,1546]" pageId="9" pageNumber="269" type="distribution">
|
||
<paragraph id="8BF14F1D5152FFEEFF3513AEFAC8775B" blockId="9.[151,1437,871,1546]" box="[199,1387,1519,1546]" pageId="9" pageNumber="269">
|
||
<emphasis id="B93A930F5152FFEEFF3513AEFEC2775B" bold="true" box="[199,353,1519,1545]" pageId="9" pageNumber="269">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54F5F1BF5152FFEEFD9213B1FD337758" box="[608,656,1520,1546]" pageId="9" pageNumber="269">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="498A81FF5152FFEEFCFD13AEFBE5775B" box="[783,1094,1519,1545]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
|
||
, Iraqi
|
||
<collectingRegion id="498A81FF5152FFEEFB7E13AEFB59775B" box="[1164,1274,1519,1545]" country="Iran" name="Kordestan" pageId="9" pageNumber="269">Kurdistan</collectingRegion>
|
||
(
|
||
<figureCitation id="137553985152FFEEFAF813AEFAFE775B" box="[1290,1373,1519,1545]" captionStart="FIGURE 10" captionStartId="16.[152,255,1294,1318]" captionTargetBox="[151,1436,381,1271]" captionTargetId="figure-58@16.[151,1436,381,1271]" captionTargetPageId="16" captionText="FIGURE 10. Collection localities of the new species described in this paper in Iraqi Kurdistan: Tegenaria sorani sp. n. (circle), Dysdera goyzha sp. n. (triangle), Dysdera kurdistanica sp. n. and Zelotes hazarmerdensis sp. n. (square), and Palpimanus garmiyanus sp. n. (hexagon)." pageId="9" pageNumber="269">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |