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<mods:title id="70077A5A97B8DD1B506779D8EE88513E">Chariodema villosa Blanchard, 1850 is also a Blepharotoma Blanchard, 1850 (Coleoptera: Scarabaeidae: Sericoidinae)</mods:title>
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<mods:namePart id="8250164A25BC6DE655B60779600B2705">Clavijo-Bustos, Julián</mods:namePart>
<mods:affiliation id="8D095D1D1011126AB4D5BBD0B4FC0F0F">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Instituto de Investigación de Recursos Biológicos Alexander von Humboldt. Claustro San Agustín, Villa de Leyva, Boyacá, Colombia.</mods:affiliation>
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<mods:namePart id="CA99F26738218DEC1B5951BC7A819451">Salazar, Karen</mods:namePart>
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<mods:affiliation id="E47ECB6E25304CD1562443BB50540F70">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, Paris cedex 05 75231, France.</mods:affiliation>
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<mods:namePart id="BEC127082AAA36C690DC72C2B7B887C1">Moreno, Jhon César Neita</mods:namePart>
<mods:affiliation id="ED1682C28B0B59B60A94E906D053FD19">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Instituto de Investigación de Recursos Biológicos Alexander von Humboldt. Claustro San Agustín, Villa de Leyva, Boyacá, Colombia.</mods:affiliation>
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<mods:date id="86782C2A9695D377FE909039FB9E59E2">2025</mods:date>
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<taxonomicName id="4C35F1BD3639481D7B8032C4FD36FA25" authority="(Blanchard, 1850)" authorityName="Clavijo-Bustos &amp; Salazar &amp; Moreno" authorityYear="2025" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[151,643,1497,1524]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="villosa" status="comb. nov.">
<emphasis id="B941562C3639481D7B8032C4FE2CFA25" bold="true" box="[151,409,1497,1524]" italics="true" pageId="0" pageNumber="193">Blepharotoma villosa</emphasis>
(
<bibRefCitation id="EFA4F7CF3639481D7ABE32C7FDCFFA25" author="Blanchard, C. E." box="[425,634,1498,1524]" pageId="0" pageNumber="193" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A272EB573639481D798732C7FCD3FA25" box="[656,870,1498,1524]" pageId="0" pageNumber="193" rank="species">new combination</taxonomicNameLabel>
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<figureCitation id="130E96BB3639481D7B80311DFF45F9C9" box="[151,240,1536,1560]" captionStart="FIGURE 1" captionStartId="1.[151,250,1943,1967]" captionTargetBox="[188,1412,188,1918]" captionTargetId="figure-21@1.[176,1412,180,1918]" captionTargetPageId="1" captionText="FIGURE 1. Blepharotoma villosa (Blanchard, 1850), new combination (female lectotype). A, Dorsal habitus, B, ventral habitus; C, lateral habitus; D, dorsal head and pronotum; E, ventral head and pronotum; F. ventral metatibiae and abdomen; G. labels. Scale bars = 1 mm." pageId="0" pageNumber="193">Figure 1</figureCitation>
</paragraph>
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<taxonomicName id="4C35F1BD3639481D7B803103FDF6F9E6" ID-CoL="69PNF" authority="Blanchard, 1850: 118" authorityName="Blanchard" authorityPageNumber="118" authorityYear="1850" box="[151,579,1566,1591]" class="Insecta" family="Scarabaeidae" genus="Chariodema" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C3639481D7B803103FED5F9E7" box="[151,352,1566,1590]" italics="true" pageId="0" pageNumber="193">Chariodema villosa</emphasis>
<bibRefCitation id="EFA4F7CF3639481D7A713103FDF6F9E6" author="Blanchard, C. E." box="[358,579,1566,1591]" pageId="0" pageNumber="193" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard, 1850: 118</bibRefCitation>
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.
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<paragraph id="8B8A8A3E3639481D7B80315EFC03F93B" blockId="0.[151,1437,1497,2023]" pageId="0" pageNumber="193">
<emphasis id="B941562C3639481D7B80315EFD9CF98A" bold="true" box="[151,553,1602,1627]" pageId="0" pageNumber="193">
<emphasis id="B941562C3639481D7B80315EFE07F98B" box="[151,434,1602,1627]" inLineHeading="true" pageId="0" pageNumber="193" reason="1">Type material examined.</emphasis>
<typeStatus id="548E349C3639481D7AAF315EFD9CF98A" box="[440,553,1603,1627]" pageId="0" pageNumber="193" type="lectotype">Lectotype</typeStatus>
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of
<taxonomicName id="4C35F1BD3639481D795A315FFC95F98B" authorityName="Blanchard" authorityYear="1850" box="[589,800,1602,1626]" class="Insecta" family="Scarabaeidae" genus="Chariodema" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C3639481D795A315FFC95F98B" box="[589,800,1602,1626]" italics="true" pageId="0" pageNumber="193">Chariodema villosa</emphasis>
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,
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, deposited in
<collectionCode id="ED2412FB3639481D78CF315EFB9BF98B" box="[984,1070,1603,1626]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="193">MNHN</collectionCode>
: “2113 [handwritten label] // 7112 | 34 [circular, handwritten label] //
<collectionCode id="ED2412FB3639481D7900317AFD6BF9AE" box="[535,734,1639,1665]" pageId="0" pageNumber="193">MUSÉUM PARIS</collectionCode>
| Chiquitos | dOrbigny [label with the first line printed and the remainder handwritten] //
<typeStatus id="548E349C3639481D7ABB3196FDF1F972" box="[428,580,1675,1699]" pageId="0" pageNumber="193" type="lectotype">LECTOTYPE</typeStatus>
[printed label] //
<typeStatus id="548E349C3639481D79EC3196FC26F972" box="[763,915,1675,1699]" pageId="0" pageNumber="193" type="lectotype">LECTOTYPE</typeStatus>
|
<taxonomicName id="4C35F1BD3639481D78B33197FB3DF973" authorityName="Clavijo-Bustos &amp; Salazar &amp; Moreno" authorityYear="2025" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[932,1160,1674,1698]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C3639481D78B33197FB3DF973" box="[932,1160,1674,1698]" italics="true" pageId="0" pageNumber="193">Blepharotoma villosa</emphasis>
</taxonomicName>
| (
<bibRefCitation id="EFA4F7CF3639481D7FB63197FAE6F973" author="Blanchard, C. E." box="[1185,1363,1674,1698]" pageId="0" pageNumber="193" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard, 1850</bibRefCitation>
) | Des: Clavijo-Bustos, Salazar | &amp; Neita Moreno, 2024 [printed label] //
<typeStatus id="548E349C3639481D784731B2FC7DF916" box="[848,968,1711,1735]" pageId="0" pageNumber="193" type="syntype">SYNTYPE</typeStatus>
[printed label] //
<typeStatus id="548E349C3639481D7F9631B2FB4CF916" box="[1153,1273,1711,1735]" pageId="0" pageNumber="193" type="syntype">SYNTYPE</typeStatus>
|
<taxonomicName id="4C35F1BD3639481D7E1C31B3FA25F917" authorityName="Blanchard" authorityYear="1850" box="[1291,1424,1710,1734]" class="Insecta" family="Scarabaeidae" genus="Chariodema" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="B941562C3639481D7E1C31B3FA25F917" box="[1291,1424,1710,1734]" italics="true" pageId="0" pageNumber="193">Chariodema</emphasis>
</taxonomicName>
|
<taxonomicName id="4C35F1BD3639481D7B8031CFFE3BF93B" authority="Blanchard 1850" authorityName="Blanchard" authorityYear="1850" box="[151,398,1746,1770]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C3639481D7B8031CFFF6BF93B" box="[151,222,1746,1770]" italics="true" pageId="0" pageNumber="193">villosa</emphasis>
<bibRefCitation id="EFA4F7CF3639481D7BF231CFFE3BF93B" author="Blanchard, C. E." box="[229,398,1746,1770]" pageId="0" pageNumber="193" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard 1850</bibRefCitation>
</taxonomicName>
[printed label] //
<collectionCode id="ED2412FB3639481D795E31CEFD2FF93B" box="[585,666,1747,1770]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="193">MNHN</collectionCode>
| EC9782 [printed label]”.
</paragraph>
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<subSubSection id="C32FD9B53639481F7BD031EAFB85FC8A" lastPageId="2" lastPageNumber="195" pageId="0" pageNumber="193" type="discussion">
<paragraph id="8B8A8A3E3639481F7BD031EAFC9DFE7A" blockId="0.[151,1437,1497,2023]" lastBlockId="2.[151,1437,151,860]" lastPageId="2" lastPageNumber="195" pageId="0" pageNumber="193">
<emphasis id="B941562C3639481D7BD031EAFE83F8DE" bold="true" box="[199,310,1783,1807]" inLineHeading="true" pageId="0" pageNumber="193" reason="1">Remarks.</emphasis>
<bibRefCitation id="EFA4F7CF3639481D7A5731EBFDA5F8DF" author="Blanchard, C. E." box="[320,528,1782,1806]" pageId="0" pageNumber="193" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchards (1850)</bibRefCitation>
publication and specimen labels state that the specimen was collected in
<collectingCountry id="F322CAAE3639481D7E5031EBFA2DF8DF" box="[1351,1432,1782,1806]" name="Bolivia" pageId="0" pageNumber="193">Bolivia</collectingCountry>
, Department of
<collectingRegion id="49F144DC3639481D7A573006FE09F8E2" box="[320,444,1819,1843]" country="Bolivia" name="Santa Cruz" pageId="0" pageNumber="193">Santa Cruz</collectingRegion>
, Chiquitos Province, during dOrbignys exploration trip. DOrbignys expeditions across South America occurred between 1826 and 1833. According to the reference numbers on the specimen labels and the corresponding registration information in the MNHN Entry Register, this specimen was incorporated into the museums collection in 1834. There is an annotation indicating “(
<emphasis id="B941562C3639481D7816309BFBA5F84F" box="[769,1040,1926,1950]" italics="true" pageId="0" pageNumber="193">du voyage de dOrbigny</emphasis>
) (
<emphasis id="B941562C3639481D7F3A309AFEB7F813" italics="true" pageId="0" pageNumber="193">voire le catalogue manuscript de dOrbigny</emphasis>
)” “(from dOrbignys voyage) (see dOrbignys manuscript catalogue)”. The handwritten catalog,
<emphasis id="B941562C3639481D7E0D30B7FDE6F837" italics="true" pageId="0" pageNumber="193">DOrbigny - Catalogue des Coléoptères de sa collection</emphasis>
, (DOrbigny - Catalogue of
<taxonomicName id="4C35F1BD3639481D786930D3FC5BF837" box="[894,1006,1998,2022]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="0" pageNumber="193" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
in his collection) authored by DOrbigny and housed in the entomology library of the MNHN, indicates that his stay in Chiquitos occurred between July and November of 1831. Therefore, following the catalog number of the specimen, this was likely collected during that period and captured by a light trap. Additionally, associated with the specimen reference number in the DOrbigny catalog, there is a word that is difficult to decipher. However, the context of the text that mentions Chiquitos Province allows us to deduce that this word refers to the locality of Concepción. This locality, effectively located in the province of Chiquitos (
<geoCoordinate id="EE01ECF9363B481F7B883657FEA8FEB2" box="[159,285,330,355]" degrees="16" direction="south" minutes="07" orientation="latitude" pageId="2" pageNumber="195" precision="15" seconds="55" value="-16.131945">16°0755″S</geoCoordinate>
,
<geoCoordinate id="EE01ECF9363B481F7A303657FE05FEB2" box="[295,432,330,355]" degrees="62" direction="west" minutes="01" orientation="longitude" pageId="2" pageNumber="195" precision="15" seconds="34" value="-62.02611">62°0134″W</geoCoordinate>
), is at an elevation of approximately
<quantity id="4CCD27DB363B481F78213656FC1BFEB2" box="[822,942,331,355]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="195" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500600 m</quantity>
and is part of the Chiquitano dry forest biome, and is close to the locality of Santa Ana de Chiquitos, where dOrbigny also collected. Furthermore, DOrbigny attributed the identification of this specimen to the genus
<taxonomicName id="4C35F1BD363B481F7987368EFC9DFE7A" authority="Illiger." authorityName="Illiger." box="[656,808,403,427]" class="Insecta" family="Scarabaeidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B941562C363B481F7987368EFD6FFE7A" box="[656,730,403,427]" italics="true" pageId="2" pageNumber="195">Hoplia</emphasis>
Illiger.
</taxonomicName>
</paragraph>
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<paragraph id="8B8A8A3E3638481C7B80308AFE16F826" blockId="1.[151,1436,1943,2040]" pageId="1" pageNumber="194">
<emphasis id="B941562C3638481C7B80308AFEA2F87E" bold="true" box="[151,279,1943,1967]" pageId="1" pageNumber="194">FIGURE 1.</emphasis>
<taxonomicName id="4C35F1BD3638481C7A363085FD72F87E" authority="(Blanchard, 1850)" authorityName="Clavijo-Bustos &amp; Salazar &amp; Moreno" authorityYear="2025" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[289,711,1943,1968]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="villosa" status="comb. nov.">
<emphasis id="B941562C3638481C7A363085FE4AF861" box="[289,511,1944,1968]" italics="true" pageId="1" pageNumber="194">Blepharotoma villosa</emphasis>
(
<bibRefCitation id="EFA4F7CF3638481C79073085FD75F87E" author="Blanchard, C. E." box="[528,704,1943,1968]" pageId="1" pageNumber="194" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B941562C3638481C79C13085FC26F87E" bold="true" box="[726,915,1943,1968]" pageId="1" pageNumber="194">
<taxonomicNameLabel id="A272EB573638481C79C13085FC26F87E" box="[726,915,1943,1968]" pageId="1" pageNumber="194" rank="species">new combination</taxonomicNameLabel>
</emphasis>
(female lectotype).
<emphasis id="B941562C3638481C7F7D308AFB36F87E" bold="true" box="[1130,1155,1943,1967]" pageId="1" pageNumber="194">A,</emphasis>
Dorsal habitus,
<emphasis id="B941562C3638481C7E233085FAFEF861" bold="true" box="[1332,1355,1944,1968]" pageId="1" pageNumber="194">B,</emphasis>
ventral habitus;
<emphasis id="B941562C3638481C7BF930A6FEB5F802" bold="true" box="[238,256,1979,2003]" pageId="1" pageNumber="194">C</emphasis>
, lateral habitus;
<emphasis id="B941562C3638481C7ABE30A1FE0EF805" bold="true" box="[425,443,1980,2004]" pageId="1" pageNumber="194">D</emphasis>
, dorsal head and pronotum;
<emphasis id="B941562C3638481C79CB30A1FD58F805" bold="true" box="[732,749,1980,2004]" pageId="1" pageNumber="194">E</emphasis>
, ventral head and pronotum;
<emphasis id="B941562C3638481C7F0030A1FB9FF805" bold="true" box="[1047,1066,1980,2004]" pageId="1" pageNumber="194">F.</emphasis>
ventral metatibiae and abdomen;
<emphasis id="B941562C3638481C7E9530A6FA29F802" bold="true" box="[1410,1436,1979,2003]" pageId="1" pageNumber="194">G.</emphasis>
labels. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B8A8A3E363B481F7BD036AAFAB8FD1A" blockId="2.[151,1437,151,860]" pageId="2" pageNumber="195">
This
<typeStatus id="548E349C363B481F7BE836AAFED4FE1E" box="[255,353,439,463]" pageId="2" pageNumber="195" type="lectotype">lectotype</typeStatus>
fits in the genus
<taxonomicName id="4C35F1BD363B481F793636AAFD0CFE1E" authorityName="Blanchard" authorityYear="1850" box="[545,697,439,463]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B941562C363B481F793636AAFD0CFE1E" box="[545,697,439,463]" italics="true" pageId="2" pageNumber="195">Blepharotoma</emphasis>
</taxonomicName>
as diagnosed by
<bibRefCitation id="EFA4F7CF363B481F786036AAFBBCFE01" author="Smith, A. B. T." box="[887,1033,439,464]" pageId="2" pageNumber="195" pagination="1 - 28" refId="ref1638" refString="Smith, A. B. T. (2008) South American Melolonthinae (Coleoptera: Scarabaeidae) classification and nomenclature: some problems and solutions. Insecta Mundi, 60, 1-28." type="journal article" year="2008">Smith (2008)</bibRefCitation>
, sharing the clypeal margin reflexed, labrum hidden beneath the clypeus in dorsal view, labrum, and clypeus separated by a suture, meso-meta ventral process absent, abdomen with six ventrites, metatibia apically with two spurs and spurs set below and above tarsal articulation, and tarsal claws symmetric and apically toothed (
<figureCitation id="130E96BB363B481F796E353EFD0DFDED" box="[633,696,547,572]" captionStart="FIGURE 1" captionStartId="1.[151,250,1943,1967]" captionTargetBox="[188,1412,188,1918]" captionTargetId="figure-21@1.[176,1412,180,1918]" captionTargetPageId="1" captionText="FIGURE 1. Blepharotoma villosa (Blanchard, 1850), new combination (female lectotype). A, Dorsal habitus, B, ventral habitus; C, lateral habitus; D, dorsal head and pronotum; E, ventral head and pronotum; F. ventral metatibiae and abdomen; G. labels. Scale bars = 1 mm." pageId="2" pageNumber="195">Fig. 1</figureCitation>
). The specimen (Total length ≈
<quantity id="4CCD27DB363B481F7F1A3539FBEBFDEA" box="[1037,1118,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="195" unit="mm" value="8.8">8.8 mm</quantity>
from clypeus to the abdomen apex; width ≈
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at the base of the elytra) is overall in a good state of preservation but is missing the antennae and some tibiae and tarsi.
<bibRefCitation id="EFA4F7CF363B481F7A283576FE4FFD52" author="Blanchard, C. E." box="[319,506,619,643]" pageId="2" pageNumber="195" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard (1850)</bibRefCitation>
mentioned the species within “Division III” (
<taxonomicName id="4C35F1BD363B481F78CC3571FB49FD52" authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[987,1276,619,643]" class="Insecta" family="Scarabaeidae" genus="Chariodema" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B941562C363B481F78CC3571FBF2FD52" box="[987,1095,619,643]" italics="true" pageId="2" pageNumber="195">C. pallens</emphasis>
<bibRefCitation id="EFA4F7CF363B481F7F5B3576FB49FD52" author="Blanchard, C. E." box="[1100,1276,619,643]" pageId="2" pageNumber="195" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard, 1850</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C35F1BD363B481F7E393571FA26FD52" authorityName="Clavijo-Bustos &amp; Salazar &amp; Moreno" authorityYear="2025" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[1326,1427,619,643]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C363B481F7E393571FA26FD52" box="[1326,1427,619,643]" italics="true" pageId="2" pageNumber="195">C. villosa</emphasis>
</taxonomicName>
) have eight antennomeres with the ovate club. This number of antennomeres is a diagnostic character of
<taxonomicName id="4C35F1BD363B481F7E123592FA28FD76" authorityName="Blanchard" authorityYear="1850" box="[1285,1437,655,679]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B941562C363B481F7E123592FA28FD76" box="[1285,1437,655,679]" italics="true" pageId="2" pageNumber="195">Blepharotoma</emphasis>
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according to
<bibRefCitation id="EFA4F7CF363B481F7A3535AEFE06FD1D" author="Smith, A. B. T." box="[290,435,691,716]" pageId="2" pageNumber="195" pagination="1 - 28" refId="ref1638" refString="Smith, A. B. T. (2008) South American Melolonthinae (Coleoptera: Scarabaeidae) classification and nomenclature: some problems and solutions. Insecta Mundi, 60, 1-28." type="journal article" year="2008">Smith (2008)</bibRefCitation>
, and one of the characters
<bibRefCitation id="EFA4F7CF363B481F79D935AEFC3FFD1A" author="Blanchard, C. E." box="[718,906,691,715]" pageId="2" pageNumber="195" pagination="1 - 128" refId="ref1384" refString="Blanchard, C. E. (1850) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E., &amp; Lucas, H. (Eds.), Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Part I. Gide and Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard (1850)</bibRefCitation>
mentioned in the genus description.
</paragraph>
<paragraph id="8B8A8A3E363B481F7BD035CAFB85FC8A" blockId="2.[151,1437,151,860]" pageId="2" pageNumber="195">
Similar to the situation with
<taxonomicName id="4C35F1BD363B481F791235C5FDD1FD3E" baseAuthorityName="Mannerheim" baseAuthorityYear="1828" box="[517,612,727,751]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B941562C363B481F791235C5FDD1FD3E" box="[517,612,727,751]" italics="true" pageId="2" pageNumber="195">B. nitida</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFA4F7CF363B481F79B535CAFC05FD21" author="Pacheco, T. L. &amp; Ahrens, D." box="[674,944,727,752]" pageId="2" pageNumber="195" pagination="298 - 300" refId="ref1580" refString="Pacheco, T. L. &amp; Ahrens, D. (2024) Omaloplia nitida Mannerheim, 1829 is a species of Blepharotoma Blanchard, 1850 (Coleoptera: Scarabaeidae: Sericoidinae). Zootaxa, 5447 (2), 298-300. https://doi.org/10.11646/zootaxa.5447.2.12" type="journal article" year="2024">Pacheco &amp; Ahrens 2024</bibRefCitation>
), the only known specimen (
<typeStatus id="548E349C363B481F7FE735CAFAE0FD3E" box="[1264,1365,727,751]" pageId="2" pageNumber="195" type="lectotype">lectotype</typeStatus>
) of
<taxonomicName id="4C35F1BD363B481F7E9235C5FF6BFCC2" authorityName="Clavijo-Bustos &amp; Salazar &amp; Moreno" authorityYear="2025" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C363B481F7E9235C5FF6BFCC2" italics="true" pageId="2" pageNumber="195">B. villosa</emphasis>
</taxonomicName>
is a female, which complicates identification at the species level. Therefore, examining the male genitalia from close specimens and doing molecular analyses is a priority to determine whether
<taxonomicName id="4C35F1BD363B481F78EE343DFBE8FCE6" authorityName="Clavijo-Bustos &amp; Salazar &amp; Moreno" authorityYear="2025" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[1017,1117,799,823]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="B941562C363B481F78EE343DFBE8FCE6" box="[1017,1117,799,823]" italics="true" pageId="2" pageNumber="195">B. villosa</emphasis>
</taxonomicName>
is a junior or senior synonym of other species currently placed in the genus
<taxonomicName id="4C35F1BD363B481F7997345EFCADFC8A" authorityName="Blanchard" authorityYear="1850" box="[640,792,835,859]" class="Insecta" family="Scarabaeidae" genus="Blepharotoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B941562C363B481F7997345EFCADFC8A" box="[640,792,835,859]" italics="true" pageId="2" pageNumber="195">Blepharotoma</emphasis>
</taxonomicName>
, as this could be possible.
</paragraph>
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Pesarini, 1975
</taxonomicName>
(
<figureCitation id="60C02A0B7079FFD7FD505761310ECA99" box="[659,798,870,897]" captionStart="FIGURE 18" captionStartId="2.[151,250,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-41@2.[215,1372,181,1885]" captionTargetPageId="2" captionText="FIGURE 18. Fig. 1. Lectotype male and Fig. 2. Paralectotype female of Phyllobius brenskei Schilsky, 1911, designated herein, with their labels (Photos: Ivo Jurisch). Fig. 3. Holotype male of Ph. nigrofasciatus Pesarini, 1975 (Photo: Michele Zilioli). Fig. 4. Male of Ph. nigrofasciatus from Afyonkarahisar. Fig. 5. Ditto female. Fig. 6. Detail of long hairs and scales of apical part of left elytron of Ph. brenskei. Fig. 7. Ditto Ph. nigrofasciatus. Fig. 8. Ditto Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="1" pageNumber="182">Figs 35; 7</figureCitation>
;
<figureCitation id="60C02A0B7079FFD7FCEE5761315ECA98" box="[813,846,870,896]" captionStart="FIGURE 916" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[152,1436,181,1723]" captionTargetId="figure-41@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 916. Fig. 9. Holotype male of Phyllobius occultatus sp. nov. Fig. 10. Ditto, paratype female. Fig. 11. Elytral apex of the female of Ph. occultatus sp. nov. Fig. 12. Ditto of the female of Ph. nigrofasciatus. Figs 1314. Male and female of Ph. brenskei, freshly collected specimens showing variability in colour of scales and body shape. Fig. 15. Apical sternites of male and 16. Female of Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="1" pageNumber="182">12</figureCitation>
;
<figureCitation id="60C02A0B7079FFD7FC98576131B6CA98" box="[859,934,870,896]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="1" pageNumber="182">2930</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B0E165057079FFD7FF5457A831F3CD2D" pageId="1" pageNumber="182" type="materials_examined">
<paragraph id="F844368E7079FFD7FF5457A831F3CD2D" blockId="1.[151,1436,942,1077]" pageId="1" pageNumber="182">
<materialsCitation id="48933CD37079FFD7FF5457A831F3CD2D" collectingDate="1998-05-26" collectingDateMax="2004-05-22" collectingDateMin="1998-05-26" collectionCode="MNMI, BKMP, BMNH" collectorName="L. Magnano &amp; J. Vorisek &amp; Skoupy &amp; Basoren Skoupy" country="Turkey" location="Basoren" municipality="Basoren" pageId="1" pageNumber="182" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Afyonkarahisar" typeStatus="holotype">
<typeStatus id="2740882C7079FFD7FF5457A832EFCAD1" box="[151,255,943,969]" pageId="1" pageNumber="182" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="60C02A0B7079FFD7FE8B57A83398CAD1" box="[328,392,943,969]" captionStart="FIGURE 18" captionStartId="2.[151,250,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-41@2.[215,1372,181,1885]" captionTargetPageId="2" captionText="FIGURE 18. Fig. 1. Lectotype male and Fig. 2. Paralectotype female of Phyllobius brenskei Schilsky, 1911, designated herein, with their labels (Photos: Ivo Jurisch). Fig. 3. Holotype male of Ph. nigrofasciatus Pesarini, 1975 (Photo: Michele Zilioli). Fig. 4. Male of Ph. nigrofasciatus from Afyonkarahisar. Fig. 5. Ditto female. Fig. 6. Detail of long hairs and scales of apical part of left elytron of Ph. brenskei. Fig. 7. Ditto Ph. nigrofasciatus. Fig. 8. Ditto Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="1" pageNumber="182">Fig. 3</figureCitation>
) of
<taxonomicName id="3FFB4D0D7079FFD7FE7057A83126CAD1" authority="&quot; Bozdag" authorityName="Bozdag" box="[435,822,943,969]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="nigrofasciatus">
<emphasis id="CA8FEA9C7079FFD7FE7057A830DFCAD1" box="[435,719,943,969]" italics="true" pageId="1" pageNumber="182">Phyllobius nigrofasciatus</emphasis>
“Bozdag
</taxonomicName>
(
<collectingCountry id="80EC761E7079FFD7FC8157A8318ECAD1" box="[834,926,943,969]" name="Turkey" pageId="1" pageNumber="182">Turchia</collectingCountry>
)” (
<collectionCode id="9EEAAE4B7079FFD7FC7E57A83604CAD1" box="[957,1044,943,969]" pageId="1" pageNumber="182">MNMI</collectionCode>
).—
<specimenCount id="EEFDFD077079FFD7FBFC57A83689CAD1" box="[1087,1177,943,969]" count="1" pageId="1" pageNumber="182" type="female">1 female</specimenCount>
<date id="8C45104E7079FFD7FB6B57A93739CAD1" box="[1192,1321,942,969]" pageId="1" pageNumber="182" value="2004-05-22">
<collectingDate id="9C01E9A67079FFD7FB6B57A93739CAD1" box="[1192,1321,942,969]" pageId="1" pageNumber="182" value="2004-05-22">22.05.2004</collectingDate>
</date>
/ Çıngırak Yolu // Gölbaşı
<collectionCode id="9EEAAE4B7079FFD7FE8A57D43387CAF5" box="[329,407,979,1005]" pageId="1" pageNumber="182">BKMP</collectionCode>
/
<collectingRegion id="3A3FF86C7079FFD7FE6F57D433E4CAF5" box="[428,500,979,1005]" country="Turkey" name="Afyonkarahisar" pageId="1" pageNumber="182">Afyon</collectingRegion>
92 / O.
<collectingRegion id="3A3FF86C7079FFD7FD8C57D4306CCAF5" box="[591,636,979,1005]" country="Turkey" name="Siirt" pageId="1" pageNumber="182">Sert</collectingRegion>
// collezione
<collectorName id="550E53587079FFD7FCD757D431B0CAF5" box="[788,928,979,1005]" pageId="1" pageNumber="182">L. Magnano</collectorName>
” (
<collectionCode id="9EEAAE4B7079FFD7FC7A57D43605CAF5" box="[953,1045,979,1005]" pageId="1" pageNumber="182">BMNH</collectionCode>
).—
<specimenCount id="EEFDFD077079FFD7FB8357D43696CAF5" box="[1088,1158,979,1005]" count="1" pageId="1" pageNumber="182" type="male">1 male</specimenCount>
“TR-prov.
<collectingRegion id="3A3FF86C7079FFD7FAC657D4375DCAF5" box="[1285,1357,979,1005]" country="Turkey" name="Afyonkarahisar" pageId="1" pageNumber="182">Afyon</collectingRegion>
/
<locationDeviation id="E9F62FAC7079FFD7FAA057D43329CD09" location="Basoren" pageId="1" pageNumber="182">
<quantity id="3F039B6B7079FFD7FAA057D4378CCAF5" box="[1379,1436,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="182" unit="km" value="3.0">3 km</quantity>
W of Başören
</locationDeviation>
/
<date id="8C45104E7079FFD7FE9757F033D7CD09" box="[340,455,1015,1041]" pageId="1" pageNumber="182" value="1998-05-26">
<collectingDate id="9C01E9A67079FFD7FE9757F033D7CD09" box="[340,455,1015,1041]" pageId="1" pageNumber="182" value="1998-05-26">26.V.1998</collectingDate>
</date>
/
<collectorName id="550E53587079FFD7FE2057F03044CD09" box="[483,596,1015,1041]" pageId="1" pageNumber="182">J. Voříšek</collectorName>
lgt.” (
<collectionCode id="9EEAAE4B7079FFD7FD6257F030EDCD09" box="[673,765,1015,1041]" pageId="1" pageNumber="182">BMNH</collectionCode>
).—
<specimenCount id="EEFDFD077079FFD7FCEB57F03197CD09" box="[808,903,1015,1041]" count="1" pageId="1" pageNumber="182" type="female">1 female</specimenCount>
“TR. (
<collectingRegion id="3A3FF86C7079FFD7FC1957F03635CD09" box="[986,1061,1015,1041]" country="Turkey" name="Afyonkarahisar" pageId="1" pageNumber="182">Afyon</collectingRegion>
)
<date id="8C45104E7079FFD7FBF557F13688CD09" box="[1078,1176,1014,1041]" pageId="1" pageNumber="182" value="2002-06-05">
<collectingDate id="9C01E9A67079FFD7FBF557F13688CD09" box="[1078,1176,1014,1041]" pageId="1" pageNumber="182" value="2002-06-05">5.6.2002</collectingDate>
</date>
/
<collectingMunicipality id="1820ACF47079FFD7FB6257F136ECCD09" box="[1185,1276,1014,1041]" pageId="1" pageNumber="182">Basören</collectingMunicipality>
/
<collectorName id="550E53587079FFD7FAC657F1374ACD09" box="[1285,1370,1014,1041]" pageId="1" pageNumber="182">Skoupý</collectorName>
leg.// (cHW).”—
<specimenCount id="EEFDFD077079FFD7FEDA501C334FCD2D" box="[281,351,1051,1077]" count="1" pageId="1" pageNumber="182" type="male">1 male</specimenCount>
“Tr. (
<collectingRegion id="3A3FF86C7079FFD7FE62501C33FCCD2D" box="[417,492,1051,1077]" country="Turkey" name="Afyonkarahisar" pageId="1" pageNumber="182">Afyon</collectingRegion>
)
<date id="8C45104E7079FFD7FE39501D304CCD2D" box="[506,604,1050,1077]" pageId="1" pageNumber="182" value="2002-06-05">
<collectingDate id="9C01E9A67079FFD7FE39501D304CCD2D" box="[506,604,1050,1077]" pageId="1" pageNumber="182" value="2002-06-05">5.6.2002</collectingDate>
</date>
/
<collectorName id="550E53587079FFD7FDB1501C313BCD2D" box="[626,811,1051,1077]" pageId="1" pageNumber="182">
<location id="FD2460557079FFD7FDB1501C30DECD2D" LSID="urn:lsid:plazi:treatment:705287987079FFD7FF54576131F3CD2D:FD2460557079FFD7FDB1501C30DECD2D" box="[626,718,1051,1077]" country="Turkey" municipality="Basoren" name="Basoren" pageId="1" pageNumber="182" stateProvince="Afyonkarahisar">Basören</location>
Skoupý
</collectorName>
leg. ” (
<collectionCode id="9EEAAE4B7079FFD7FCBE501C31C7CD2D" box="[893,983,1051,1077]" pageId="1" pageNumber="182">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,540 @@
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<heading id="A30C81E2707BFFD5FF545490373CC9A9" bold="true" box="[151,1324,151,178]" fontSize="11" level="1" pageId="3" pageNumber="184" reason="1">
<emphasis id="CA8FEA9C707BFFD5FF545490373CC9A9" bold="true" box="[151,1324,151,178]" pageId="3" pageNumber="184">
<taxonomicName id="3FFB4D0D707BFFD5FF545490316DC9A9" authority="Germann, Wartmann &amp; Winkelmann" authorityName="Germann, Wartmann &amp; Winkelmann" authorityYear="2025" box="[151,893,151,178]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707BFFD5FF545490338BC9AA" bold="true" box="[151,411,151,178]" italics="true" pageId="3" pageNumber="184">Phyllobius occultatus</emphasis>
Germann, Wartmann &amp; Winkelmann
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,
<taxonomicNameLabel id="D1BC57E7707BFFD5FC49549031F7C9A9" box="[906,999,151,177]" pageId="3" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="60C02A0B707BFFD5FC34549036DBC9A9" box="[1015,1227,151,177]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 916" captionStartId-0="2.[151,250,1909,1933]" captionStartId-1="4.[151,250,1746,1770]" captionTargetBox-0="[215,1372,181,1885]" captionTargetBox-1="[152,1436,181,1723]" captionTargetId-0="figure-41@2.[215,1372,181,1885]" captionTargetId-1="figure-41@4.[151,1436,181,1723]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 18. Fig. 1. Lectotype male and Fig. 2. Paralectotype female of Phyllobius brenskei Schilsky, 1911, designated herein, with their labels (Photos: Ivo Jurisch). Fig. 3. Holotype male of Ph. nigrofasciatus Pesarini, 1975 (Photo: Michele Zilioli). Fig. 4. Male of Ph. nigrofasciatus from Afyonkarahisar. Fig. 5. Ditto female. Fig. 6. Detail of long hairs and scales of apical part of left elytron of Ph. brenskei. Fig. 7. Ditto Ph. nigrofasciatus. Fig. 8. Ditto Ph. occultatus sp. nov. (Photos: C. Germann)." captionText-1="FIGURE 916. Fig. 9. Holotype male of Phyllobius occultatus sp. nov. Fig. 10. Ditto, paratype female. Fig. 11. Elytral apex of the female of Ph. occultatus sp. nov. Fig. 12. Ditto of the female of Ph. nigrofasciatus. Figs 1314. Male and female of Ph. brenskei, freshly collected specimens showing variability in colour of scales and body shape. Fig. 15. Apical sternites of male and 16. Female of Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="3" pageNumber="184">Figs 811; 1516</figureCitation>
;
<figureCitation id="60C02A0B707BFFD5FB1A54903733C9A9" box="[1241,1315,151,177]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">1726</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B0E16505707BFFD0FF5455043099CCB0" lastPageId="6" lastPageNumber="187" pageId="3" pageNumber="184" type="materials_examined">
<paragraph id="F844368E707BFFD5FF545504361FC87D" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
<materialsCitation id="48933CD3707BFFD5FF545504361FC87D" collectingDate="2019-05-09" collectionCode="R, JACOB, NMB" elevation="805" latitude="36.883247" location="Taygetos Mts." longLatPrecision="1" longitude="22.291082" municipality="Mani" pageId="3" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Peloponnisos" typeStatus="holotype">
<typeStatus id="2740882C707BFFD5FF5455043314C805" box="[151,260,259,285]" pageId="3" pageNumber="184" type="holotype">Holotype</typeStatus>
:
<specimenCount id="EEFDFD07707BFFD5FECD55033335C806" box="[270,293,260,286]" count="1" pageId="3" pageNumber="184" type="male"></specimenCount>
, “GR
<collectingRegion id="3A3FF86C707BFFD5FEA9550433FBC805" box="[362,491,259,285]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnes</collectingRegion>
<collectingMunicipality id="1820ACF4707BFFD5FE325504303CC805" box="[497,556,259,285]" pageId="3" pageNumber="184">Mani</collectingMunicipality>
/ SE Kalamata,
<location id="FD246055707BFFD5FD225503316FC806" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FD225503316FC806" box="[737,895,260,286]" latitude="36.883247" longLatPrecision="1" longitude="22.291082" municipality="Mani" name="Taygetos Mts." pageId="3" pageNumber="184" stateProvince="Peloponnisos">Taygetos Mts.</location>
/ 0,
<quantity id="3F039B6B707BFFD5FC64550431F1C805" box="[935,993,259,285]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="184" unit="km" value="7.0">7 km</quantity>
NE SAIDONA /
<date id="8C45104E707BFFD5FB6755043707C805" box="[1188,1303,259,285]" pageId="3" pageNumber="184" value="2019-05-09">
<collectingDate id="9C01E9A6707BFFD5FB6755043707C805" box="[1188,1303,259,285]" pageId="3" pageNumber="184" value="2019-05-09">09.V.2019</collectingDate>
</date>
<quantity id="3F039B6B707BFFD5FADE55043773C806" box="[1309,1379,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="3" pageNumber="184" unit="m" value="805.0">
<elevation id="73D6D1BD707BFFD5FADE55043773C806" box="[1309,1379,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="3" pageNumber="184" unit="m" value="805.0">805 m</elevation>
</quantity>
/ lgt.
<collectionCode id="9EEAAE4B707BFFD5FF54552F32BDC85A" box="[151,173,296,322]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="184">R</collectionCode>
.
<collectionCode id="9EEAAE4B707BFFD5FF7A55203302C859" box="[185,274,295,321]" pageId="3" pageNumber="184">JACOB</collectionCode>
//
<geoCoordinate id="9DCF5049707BFFD5FEF1552E33C7C859" box="[306,471,295,324]" degrees="36" direction="north" minutes="52" orientation="latitude" pageId="3" pageNumber="184" precision="1" seconds="59,7" value="36.883247">36°5259,7″ N</geoCoordinate>
/
<geoCoordinate id="9DCF5049707BFFD5FE24552E3088C85A" box="[487,664,296,324]" degrees="022" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="184" precision="1" seconds="27,9" value="22.291082">022°1727,9″ E</geoCoordinate>
// von Griech. Tanne/
<taxonomicName id="3FFB4D0D707BFFD5FC5A55203679C859" box="[921,1129,295,321]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="184" phylum="Tracheophyta" rank="species" species="cephalonica">
<emphasis id="CA8FEA9C707BFFD5FC5A55203679C859" box="[921,1129,295,321]" italics="true" pageId="3" pageNumber="184">Abies cephalonica</emphasis>
</taxonomicName>
//
<typeStatus id="2740882C707BFFD5FB4B5520370CC859" box="[1160,1308,295,321]" pageId="3" pageNumber="184" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="3FFB4D0D707BFFD5FAE75520331AC87D" authorityName="Germann, Wartmann &amp; Winkelmann" authorityYear="2025" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="occultatus">
<emphasis id="CA8FEA9C707BFFD5FAE75520331AC87D" italics="true" pageId="3" pageNumber="184">Phyllobius occultatus</emphasis>
</taxonomicName>
des. Germann, Wartmann &amp; Winkelmann, 2025” [red label] (
<collectionCode id="9EEAAE4B707BFFD5FC7E554C3613C87D" box="[957,1027,331,357]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="184">NMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F844368E707BFFD5FF0455773008CBEA" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
<materialsCitation id="48933CD3707BFFD5FF0455773751C891" box="[199,1345,367,394]" collectionCode="KITA" collectorName="Fam. Dillge" elevation="90" location="Mani" pageId="3" pageNumber="184" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="2740882C707BFFD5FF0455773328C892" box="[199,312,368,394]" pageId="3" pageNumber="184" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="EEFDFD07707BFFD5FE8555683365C892" box="[326,373,367,394]" count="1" pageId="3" pageNumber="184" type="male">1 ♂</specimenCount>
,
<specimenCount id="EEFDFD07707BFFD5FE47556833A1C892" box="[388,433,367,394]" count="1" pageId="3" pageNumber="184" type="female">1 ♀</specimenCount>
“S-GR.
<location id="FD246055707BFFD5FDDB55683043C891" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FDDB55683043C891" box="[536,595,367,393]" name="Mani" pageId="3" pageNumber="184">Mani</location>
4.[19]93 /
<quantity id="3F039B6B707BFFD5FD1B55683132C891" box="[728,802,367,393]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="184" unit="km" value="10.0">10 km</quantity>
nördl.
<collectionCode id="9EEAAE4B707BFFD5FCBB556831ACC891" box="[888,956,367,393]" pageId="3" pageNumber="184">KITA</collectionCode>
; /
<quantity id="3F039B6B707BFFD5FC1F55683602C891" box="[988,1042,367,393]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="184" unit="m" value="90.0">
<elevation id="73D6D1BD707BFFD5FC1F55683602C891" box="[988,1042,367,393]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="184" unit="m" value="90.0">90m</elevation>
</quantity>
; leg.:
<collectorName id="550E5358707BFFD5FB98557736F5C891" box="[1115,1253,367,394]" pageId="3" pageNumber="184">Fam. Dillge</collectorName>
” (cCG)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FAA055683388C8B5" collectionCode="KITA" collectorName="Fam. Dillge" elevation="90" location="Mani" pageId="3" pageNumber="184" specimenCount="6" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FAA05568378DC892" box="[1379,1437,367,394]" count="6" pageId="3" pageNumber="184" type="generic">6 ex.</specimenCount>
same data as HT (cRJ)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FE795594369CC8C9" collectingDate="2019-05-08" collectionCode="V, R, JACOB, NMB" county="Mani" elevation="500" latitude="36.854942" location="Taygetos Mts." longLatPrecision="1" longitude="22.343555" municipality="Kalamata" pageId="3" pageNumber="184" specimenCount="12" specimenCount-female="1" specimenCount-male="2" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FE79559433F7C8B6" box="[442,487,403,430]" count="2" pageId="3" pageNumber="184" type="male">2 ♂</specimenCount>
,
<specimenCount id="EEFDFD07707BFFD5FE305594300EC8B6" box="[499,542,403,430]" count="1" pageId="3" pageNumber="184" type="female">1 ♀</specimenCount>
,
<specimenCount id="EEFDFD07707BFFD5FDEE55943074C8B6" box="[557,612,403,430]" count="9" pageId="3" pageNumber="184" type="generic">9 ex.</specimenCount>
“GR
<collectingRegion id="3A3FF86C707BFFD5FD6455943138C8B5" box="[679,808,403,429]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnes</collectingRegion>
<collectingCounty id="11254E02707BFFD5FCEC5594317AC8B5" box="[815,874,403,429]" pageId="3" pageNumber="184">Mani</collectingCounty>
/ SE
<collectingMunicipality id="1820ACF4707BFFD5FC6955943608C8B5" box="[938,1048,403,429]" pageId="3" pageNumber="184">Kalamata</collectingMunicipality>
,
<location id="FD246055707BFFD5FBE7559336D3C8B6" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FBE7559336D3C8B6" box="[1060,1219,404,430]" county="Mani" latitude="36.854942" longLatPrecision="1" longitude="22.343555" municipality="Kalamata" name="Taygetos Mts." pageId="3" pageNumber="184" stateProvince="Peloponnisos">Taygetos Mts.</location>
/ KARIOVOUNI /
<date id="8C45104E707BFFD5FF5455B0331AC8C9" box="[151,266,439,465]" pageId="3" pageNumber="184" value="2019-05-08">
<collectingDate id="9C01E9A6707BFFD5FF5455B0331AC8C9" box="[151,266,439,465]" pageId="3" pageNumber="184" value="2019-05-08">
08.
<collectionCode id="9EEAAE4B707BFFD5FF7455B032DCC8C9" box="[183,204,439,465]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="184" type="Museum">V</collectionCode>
.2019
</collectingDate>
</date>
<quantity id="3F039B6B707BFFD5FED155B0334BC8CA" box="[274,347,439,466]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="184" unit="m" value="500.0">
<elevation id="73D6D1BD707BFFD5FED155B0334BC8CA" box="[274,347,439,466]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="184" unit="m" value="500.0">500 m</elevation>
</quantity>
/ lgt.
<collectionCode id="9EEAAE4B707BFFD5FE6255BF33A7C8CA" box="[417,439,440,466]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="184">R</collectionCode>
.
<collectionCode id="9EEAAE4B707BFFD5FE0055B0300CC8C9" box="[451,540,439,465]" pageId="3" pageNumber="184">JACOB</collectionCode>
//
<geoCoordinate id="9DCF5049707BFFD5FDFF55BE30F2C8C9" box="[572,738,439,468]" degrees="36" direction="north" minutes="51" orientation="latitude" pageId="3" pageNumber="184" precision="1" seconds="17,8" value="36.854942">36°5117,8″ N</geoCoordinate>
/
<geoCoordinate id="9DCF5049707BFFD5FD3055BE31B6C8C9" box="[755,934,439,468]" degrees="022" direction="east" minutes="20" orientation="longitude" pageId="3" pageNumber="184" precision="1" seconds="36,8" value="22.343555">022°2036,8″ E</geoCoordinate>
” (cRJ, cSW,
<collectionCode id="9EEAAE4B707BFFD5FBFC55B03695C8C9" box="[1087,1157,439,465]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="184">NMB</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FB7355B032EDCB01" collectingDate="2019-05-08" collectionCode="NW, MILA, V, R, JACOB" county="Mani" elevation="534" latitude="36.83836" location="Taygetos Mts." longLatPrecision="1" longitude="22.341251" municipality="Kalamata" pageId="3" pageNumber="184" specimenCount="1" specimenCount-female="1" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FB7355B036C7C8CA" box="[1200,1239,439,466]" count="1" pageId="3" pageNumber="184" type="female">1 ♀</specimenCount>
“GR
<collectingRegion id="3A3FF86C707BFFD5FAD855B0378CC8C9" box="[1307,1436,439,465]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnes</collectingRegion>
<collectingCounty id="11254E02707BFFD5FF5455DC32C2C8ED" box="[151,210,475,501]" pageId="3" pageNumber="184">Mani</collectingCounty>
/ SE
<collectingMunicipality id="1820ACF4707BFFD5FECE55DC336BC8ED" box="[269,379,475,501]" pageId="3" pageNumber="184">Kalamata</collectingMunicipality>
,
<location id="FD246055707BFFD5FE4655DB3033C8EE" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FE4655DB3033C8EE" box="[389,547,476,502]" county="Mani" latitude="36.83836" longLatPrecision="1" longitude="22.341251" municipality="Kalamata" name="Taygetos Mts." pageId="3" pageNumber="184" stateProvince="Peloponnisos">Taygetos Mts.</location>
/
<collectionCode id="9EEAAE4B707BFFD5FDF455DB30A2C8ED" box="[567,690,475,502]" pageId="3" pageNumber="184">NW MILA</collectionCode>
/
<date id="8C45104E707BFFD5FD0255DC3124C8ED" box="[705,820,475,501]" pageId="3" pageNumber="184" value="2019-05-08">
<collectingDate id="9C01E9A6707BFFD5FD0255DC3124C8ED" box="[705,820,475,501]" pageId="3" pageNumber="184" value="2019-05-08">
08.
<collectionCode id="9EEAAE4B707BFFD5FD2355DC30E5C8ED" box="[736,757,475,501]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="184" type="Museum">V</collectionCode>
.2019
</collectingDate>
</date>
<quantity id="3F039B6B707BFFD5FCF955DC3190C8EE" box="[826,896,475,502]" metricMagnitude="2" metricUnit="m" metricValue="5.34" pageId="3" pageNumber="184" unit="m" value="534.0">
<elevation id="73D6D1BD707BFFD5FCF955DC3190C8EE" box="[826,896,475,502]" metricMagnitude="2" metricUnit="m" metricValue="5.34" pageId="3" pageNumber="184" unit="m" value="534.0">534 m</elevation>
</quantity>
/ lgt.
<collectionCode id="9EEAAE4B707BFFD5FC7C55DB31C5C8EE" box="[959,981,476,502]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="184">R</collectionCode>
.
<collectionCode id="9EEAAE4B707BFFD5FC1C55DC3628C8ED" box="[991,1080,475,501]" pageId="3" pageNumber="184">JACOB</collectionCode>
//
<geoCoordinate id="9DCF5049707BFFD5FB9055DA36E7C8ED" box="[1107,1271,475,504]" degrees="36" direction="north" minutes="50" orientation="latitude" pageId="3" pageNumber="184" precision="1" seconds="18,1" value="36.83836">36°5018,1″ N</geoCoordinate>
/
<geoCoordinate id="9DCF5049707BFFD5FAC655DA32BACB01" degrees="022" direction="east" minutes="20" orientation="longitude" pageId="3" pageNumber="184" precision="1" seconds="28,5" value="22.341251">022°2028,5″ E</geoCoordinate>
” (cRJ)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FEE355F83016CB25" collectingDate="2019-05-08" collectionCode="V, R, JACOB, MNFB, BMNH" county="Mani" elevation="380" location="Kalamata" municipality="Taygetos Mts." pageId="3" pageNumber="184" specimenCount="8" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FEE355F83349CB02" box="[288,345,511,538]" count="8" pageId="3" pageNumber="184" type="generic">8 ex.</specimenCount>
“GR
<collectingRegion id="3A3FF86C707BFFD5FE6355F83031CB01" box="[416,545,511,537]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnes</collectingRegion>
<collectingCounty id="11254E02707BFFD5FDEF55F83077CB01" box="[556,615,511,537]" pageId="3" pageNumber="184">Mani</collectingCounty>
/
<collectingMunicipality id="1820ACF4707BFFD5FD4056073135CB02" box="[643,805,512,538]" pageId="3" pageNumber="184">Taygetos Mts.</collectingMunicipality>
SE
<location id="FD246055707BFFD5FC9955F831D5CB01" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FC9955F831D5CB01" box="[858,965,511,537]" county="Mani" municipality="Taygetos Mts." name="Kalamata" pageId="3" pageNumber="184" stateProvince="Peloponnisos">Kalamata</location>
/ NE Platsa /
<date id="8C45104E707BFFD5FBB355F836F3CB01" box="[1136,1251,511,537]" pageId="3" pageNumber="184" value="2019-05-08">
<collectingDate id="9C01E9A6707BFFD5FBB355F836F3CB01" box="[1136,1251,511,537]" pageId="3" pageNumber="184" value="2019-05-08">
08.
<collectionCode id="9EEAAE4B707BFFD5FB5355F836B5CB01" box="[1168,1189,511,537]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="184" type="Museum">V</collectionCode>
.2019
</collectingDate>
</date>
<quantity id="3F039B6B707BFFD5FB2E55F83727CB02" box="[1261,1335,511,538]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="184" unit="m" value="380.0">
<elevation id="73D6D1BD707BFFD5FB2E55F83727CB02" box="[1261,1335,511,538]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="184" unit="m" value="380.0">380 m</elevation>
</quantity>
/ lgt.
<collectionCode id="9EEAAE4B707BFFD5FA4056073786CB02" box="[1411,1430,512,538]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="184">R</collectionCode>
.
<collectionCode id="9EEAAE4B707BFFD5FF54562432E2CB25" box="[151,242,547,573]" pageId="3" pageNumber="184">JACOB</collectionCode>
” (cRJ,
<collectionCode id="9EEAAE4B707BFFD5FE895623338CCB26" box="[330,412,548,574]" pageId="3" pageNumber="184">MNFB</collectionCode>
,
<collectionCode id="9EEAAE4B707BFFD5FE69562333EECB26" box="[426,510,548,574]" pageId="3" pageNumber="184">BMNH</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FDE95624318CCB79" collectingDate="2011-05-19" collectionCode="FO" collectorName="Winkelmann" county="Messinia" elevation="800" latitude="36.883053" location="Saidona" longLatPrecision="19" longitude="22.290277" municipality="Mt. Taygetos" pageId="3" pageNumber="184" specimenCode="FO5" specimenCount="7" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FDE956243074CB26" box="[554,612,547,574]" count="7" pageId="3" pageNumber="184" type="generic">7 ex.</specimenCount>
“GR:
<collectingRegion id="3A3FF86C707BFFD5FD7156243154CB25" box="[690,836,547,573]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnese</collectingRegion>
:
<collectingCounty id="11254E02707BFFD5FC92562431A9CB25" box="[849,953,547,573]" pageId="3" pageNumber="184">Messinia</collectingCounty>
,
<collectingMunicipality id="1820ACF4707BFFD5FC055623364CCB26" box="[966,1116,548,574]" pageId="3" pageNumber="184">Mt. Taygetos</collectingMunicipality>
W,
<location id="FD246055707BFFD5FB4E562436F7CB25" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FB4E562436F7CB25" box="[1165,1255,547,573]" county="Messinia" latitude="36.883053" longLatPrecision="19" longitude="22.290277" municipality="Mt. Taygetos" name="Saidona" pageId="3" pageNumber="184" stateProvince="Peloponnisos">Saidona</location>
E,
<geoCoordinate id="9DCF5049707BFFD5FAD05624378CCB26" box="[1299,1436,547,574]" degrees="36" direction="north" minutes="52" orientation="latitude" pageId="3" pageNumber="184" precision="15" seconds="59" value="36.883053">36°5259”N</geoCoordinate>
<geoCoordinate id="9DCF5049707BFFD5FF5456403330CB7A" box="[151,288,583,610]" degrees="22" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="184" precision="15" seconds="25" value="22.290277">22°1725”E</geoCoordinate>
,
<quantity id="3F039B6B707BFFD5FEF35640336FCB7A" box="[304,383,583,610]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="184" unit="m" value="800.0">
<elevation id="73D6D1BD707BFFD5FEF35640336FCB7A" box="[304,383,583,610]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="184" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="8C45104E707BFFD5FE4C5640301FCB7A" box="[399,527,583,610]" pageId="3" pageNumber="184" value="2011-05-19">
<collectingDate id="9C01E9A6707BFFD5FE4C5640301FCB7A" box="[399,527,583,610]" pageId="3" pageNumber="184" value="2011-05-19">19.05.2011</collectingDate>
</date>
, leg.
<collectorName id="550E5358707BFFD5FD97564030F4CB79" box="[596,740,583,609]" pageId="3" pageNumber="184">Winkelmann</collectorName>
(
<specimenCode id="A85D9EF5707BFFD5FD3B5640313CCB7A" box="[760,812,583,610]" collectionCode="FO" pageId="3" pageNumber="184">FO5</specimenCode>
)” (cHW)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FC7E56403716CB9E" collectingDate="2011-05-21" collectionCode="FO" collectorName="Winkelmann" county="Messinia" elevation="1300" latitude="36.896114" location="Marathos" longLatPrecision="19" longitude="22.319168" municipality="Mt. Taygetos" pageId="3" pageNumber="184" specimenCode="FO8" specimenCount="22" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FC7E56403618CB7A" box="[957,1032,583,610]" count="22" pageId="3" pageNumber="184" type="generic">22 ex.</specimenCount>
“GR:
<collectingRegion id="3A3FF86C707BFFD5FB98564036FDCB79" box="[1115,1261,583,609]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnese</collectingRegion>
:
<collectingCounty id="11254E02707BFFD5FB3F56403774CB79" box="[1276,1380,583,609]" pageId="3" pageNumber="184">Messinia</collectingCounty>
,
<collectingMunicipality id="1820ACF4707BFFD5FAB6564F32EFCB9E" pageId="3" pageNumber="184">Mt. Taygetos</collectingMunicipality>
,
<location id="FD246055707BFFD5FECB566C3362CB9D" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FECB566C3362CB9D" box="[264,370,619,645]" county="Messinia" latitude="36.896114" longLatPrecision="19" longitude="22.319168" municipality="Mt. Taygetos" name="Marathos" pageId="3" pageNumber="184" stateProvince="Peloponnisos">Marathos</location>
S,
<geoCoordinate id="9DCF5049707BFFD5FE50566C300DCB9E" box="[403,541,619,646]" degrees="36" direction="north" minutes="53" orientation="latitude" pageId="3" pageNumber="184" precision="15" seconds="46" value="36.896114">36°5346”N</geoCoordinate>
<geoCoordinate id="9DCF5049707BFFD5FDE1566C30BBCB9D" box="[546,683,619,645]" degrees="22" direction="east" minutes="19" orientation="longitude" pageId="3" pageNumber="184" precision="15" seconds="09" value="22.319168">22°1909”E</geoCoordinate>
,
<quantity id="3F039B6B707BFFD5FD77566C311ACB9E" box="[692,778,619,646]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="184" unit="m" value="1300.0">
<elevation id="73D6D1BD707BFFD5FD77566C311ACB9E" box="[692,778,619,646]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="184" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<date id="8C45104E707BFFD5FCD7566C3184CB9E" box="[788,916,619,646]" pageId="3" pageNumber="184" value="2011-05-21">
<collectingDate id="9C01E9A6707BFFD5FCD7566C3184CB9E" box="[788,916,619,646]" pageId="3" pageNumber="184" value="2011-05-21">21.05.2011</collectingDate>
</date>
, leg.
<collectorName id="550E5358707BFFD5FC08566C364BCB9D" box="[971,1115,619,645]" pageId="3" pageNumber="184">Winkelmann</collectorName>
(
<specimenCode id="A85D9EF5707BFFD5FBAB566C368BCB9D" box="[1128,1179,619,645]" collectionCode="FO" pageId="3" pageNumber="184">FO8</specimenCode>
)” (cHW)
</materialsCitation>
.—
<materialsCitation id="48933CD3707BFFD5FAEB566C3004CBEA" collectingDate="2011-05-23" collectionCode="FO" collectorName="Winkelmann &amp; All" county="Messinia" elevation="680" latitude="36.883614" location="Saidona" longLatPrecision="19" longitude="22.285276" municipality="Mt. Taygetos" pageId="3" pageNumber="184" specimenCode="FO11" specimenCount="5" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="EEFDFD07707BFFD5FAEB566C374CCB9E" box="[1320,1372,619,646]" count="5" pageId="3" pageNumber="184" type="generic">5 ex.</specimenCount>
“GR:
<collectingRegion id="3A3FF86C707BFFD5FF5456883339CBB1" box="[151,297,655,681]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="184">Peloponnese</collectingRegion>
:
<collectingCounty id="11254E02707BFFD5FEF75688338CCBB1" box="[308,412,655,681]" pageId="3" pageNumber="184">Messinia</collectingCounty>
,
<collectingMunicipality id="1820ACF4707BFFD5FE6B5697302DCBB2" box="[424,573,656,682]" pageId="3" pageNumber="184">Mt. Taygetos</collectingMunicipality>
W,
<location id="FD246055707BFFD5FDA9568830D7CBB1" LSID="urn:lsid:plazi:treatment:70528798707BFFD0FF5454903099CCB0:FD246055707BFFD5FDA9568830D7CBB1" box="[618,711,655,681]" county="Messinia" latitude="36.883614" longLatPrecision="19" longitude="22.285276" municipality="Mt. Taygetos" name="Saidona" pageId="3" pageNumber="184" stateProvince="Peloponnisos">Saidona</location>
,
<quantity id="3F039B6B707BFFD5FD175688310DCBB2" box="[724,797,655,682]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="184" unit="m" value="680.0">
<elevation id="73D6D1BD707BFFD5FD175688310DCBB2" box="[724,797,655,682]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="184" unit="m" value="680.0">680 m</elevation>
</quantity>
,
<geoCoordinate id="9DCF5049707BFFD5FCE9568831A3CBB2" box="[810,947,655,682]" degrees="36" direction="north" minutes="53" orientation="latitude" pageId="3" pageNumber="184" precision="15" seconds="01" value="36.883614">36°5301”N</geoCoordinate>
<geoCoordinate id="9DCF5049707BFFD5FC7856883654CBB1" box="[955,1092,655,681]" degrees="22" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="184" precision="15" seconds="07" value="22.285276">22°1707”E</geoCoordinate>
,
<date id="8C45104E707BFFD5FB93568836C0CBB2" box="[1104,1232,655,682]" pageId="3" pageNumber="184" value="2011-05-23">
<collectingDate id="9C01E9A6707BFFD5FB93568836C0CBB2" box="[1104,1232,655,682]" pageId="3" pageNumber="184" value="2011-05-23">23.05.2011</collectingDate>
</date>
, leg.
<collectorName id="550E5358707BFFD5FACE5688378DCBB1" box="[1293,1437,655,681]" pageId="3" pageNumber="184">Winkelmann</collectorName>
(
<specimenCode id="A85D9EF5707BFFD5FF5C56B432F0CBD5" box="[159,224,691,717]" collectionCode="FO" pageId="3" pageNumber="184">FO11</specimenCode>
)” (cHW).
<collectorName id="550E5358707BFFD5FE9E56B43391CBD5" box="[349,385,691,717]" pageId="3" pageNumber="184">All</collectorName>
<typeStatus id="2740882C707BFFD5FE4F56B333E7CBD6" box="[396,503,692,718]" pageId="3" pageNumber="184" type="paratype">paratypes</typeStatus>
with an additional red label: “
<typeStatus id="2740882C707BFFD5FC9E56B431FBCBD5" box="[861,1003,691,717]" pageId="3" pageNumber="184" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="3FFB4D0D707BFFD5FC3556B436FCCBD5" authorityName="Germann, Wartmann &amp; Winkelmann" authorityYear="2025" box="[1014,1260,691,717]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="occultatus">
<emphasis id="CA8FEA9C707BFFD5FC3556B436FCCBD5" box="[1014,1260,691,717]" italics="true" pageId="3" pageNumber="184">Phyllobius occultatus</emphasis>
</taxonomicName>
des. Germann, Wartmann &amp; Winkelmann, 2025”
</materialsCitation>
.
</paragraph>
<paragraph id="F844368E707BFFD5FF0456FC31B1CA0E" blockId="3.[151,1437,259,2013]" box="[199,929,763,790]" pageId="3" pageNumber="184">
Size. (Without rostrum) males 3.65.0 mm, females
<quantity id="3F039B6B707BFFD5FCD056FC318DCA0E" box="[787,925,763,790]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="6.1" metricValueMin="4.2" pageId="3" pageNumber="184" unit="mm" value="5.15" valueMax="6.1" valueMin="4.2">4.26.1 mm</quantity>
.
</paragraph>
<paragraph id="F844368E707BFFD5FF0457183064CA45" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
Colour. Body and head black; antennae and legs blackish, dark brown or reddish-brown, often apex of scape and club of antennae blackish (
<figureCitation id="60C02A0B707BFFD5FE3057443078CA45" box="[499,616,835,861]" captionStart="FIGURE 916" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[152,1436,181,1723]" captionTargetId="figure-41@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 916. Fig. 9. Holotype male of Phyllobius occultatus sp. nov. Fig. 10. Ditto, paratype female. Fig. 11. Elytral apex of the female of Ph. occultatus sp. nov. Fig. 12. Ditto of the female of Ph. nigrofasciatus. Figs 1314. Male and female of Ph. brenskei, freshly collected specimens showing variability in colour of scales and body shape. Fig. 15. Apical sternites of male and 16. Female of Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="3" pageNumber="184">Figs 910</figureCitation>
).
</paragraph>
<paragraph id="F844368E707BFFD5FF0457603037CD66" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
Head. Eyes convex, protruding from outline of head, oval in section. Rostrum elongate, slightly longer than wide (L/W: 1.12), parallel-sided, epistome with glossy triangular area. Epifrons elevated, minimal width at first third, abruptly widening forward, flattened towards frons. Frons impressed in middle with short bar-shaped sulcus. Scrobes deep, oval, visible from above. Temples twice as long as eyes diameter. Head and rostrum with thin brownish hairs, without scales. Antennal scape long slightly bowed and thickened towards apex, reaching fore margin of prothorax in repose. Funiculus slender, 1
<superScript id="0F8E9BC6707BFFD5FD21501C30FDCD31" attach="left" box="[738,749,1051,1065]" fontSize="6" pageId="3" pageNumber="184">st</superScript>
and 2
<superScript id="0F8E9BC6707BFFD5FCF6501C3155CD31" attach="left" box="[821,837,1051,1065]" fontSize="6" pageId="3" pageNumber="184">nd</superScript>
funicular antennomeres two and a half times longer than wide, 3rd one and a half longer than wide, 4
<superScript id="0F8E9BC6707BFFD5FD00503830C0CD55" attach="left" box="[707,720,1087,1101]" fontSize="6" pageId="3" pageNumber="184">th</superScript>
to 7
<superScript id="0F8E9BC6707BFFD5FCCB50383105CD55" attach="right" box="[776,789,1087,1101]" fontSize="6" pageId="3" pageNumber="184">th</superScript>
as long as wide, club fusiform, long and slender, slightly wider as 7
<superScript id="0F8E9BC6707BFFD5FECD5064330BCD69" attach="left" box="[270,283,1123,1137]" fontSize="6" pageId="3" pageNumber="184">th</superScript>
funicular antennomere.
</paragraph>
<paragraph id="F844368E707BFFD5FF04508030CBCC15" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">Prothorax. (L/W): 0.6, transverse, widest in middle, laterally strongly rounded, densely and irregularly punctuate on disc, vestiture consisting of oval, light green metallic recumbent scales, often with golden shimmer and adjacent long brownish hairs. Scales arranged in 2 spots at each side of prothorax from base to middle, separated in middle. Metathorax. Hind wings present, longer than elytra.</paragraph>
<paragraph id="F844368E707BFFD5FF045110331ACCD9" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">Elytra. (L/W): 1.92.0, parallel-sided. Base somewhat wider than prothorax. Shoulders well-pronounced. Striae thin, linear, interstriae four times wider, with thin and short, bright bowed hairs and strong, raised pointed blackish hairs, longer than width of 2 interstriae. Vestiture consisting of light green-metallic, ovoid, drop-shaped to broad oval scales with golden shimmer, loosely spread over elytra with gaps in between. Scutellum bare, glossy and triangular.</paragraph>
<paragraph id="F844368E707BFFD5FF0451CB3656CF49" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
Legs. Strong, femora thickened, with strong denticle, profemora of males strongly swollen, about 3 times as thick as meso- and metafemora, denticle shaped as equilateral triangle, tibiae straight along outer margin, inner margin bisinuate. Four visible tarsal segments, 1
<superScript id="0F8E9BC6707BFFD5FD00521430DECF39" attach="left" box="[707,718,1555,1569]" fontSize="6" pageId="3" pageNumber="184">st</superScript>
tarsomere 1.5 times longer than 2
<superScript id="0F8E9BC6707BFFD5FB9A52143679CF39" attach="none" box="[1113,1129,1555,1569]" fontSize="6" pageId="3" pageNumber="184">nd</superScript>
, 3
<superScript id="0F8E9BC6707BFFD5FB4B52143686CF39" attach="left" box="[1160,1174,1555,1569]" fontSize="6" pageId="3" pageNumber="184">rd</superScript>
shorter, bilobed, twice wider than 2
<superScript id="0F8E9BC6707BFFD5FEE052303323CF5D" attach="none" box="[291,307,1591,1605]" fontSize="6" pageId="3" pageNumber="184">nd</superScript>
, 4
<superScript id="0F8E9BC6707BFFD5FE8C5230334CCF5D" attach="left" box="[335,348,1591,1605]" fontSize="6" pageId="3" pageNumber="184">th</superScript>
tiny, onychium as long as segment 1. Claws simple, fused at base.
</paragraph>
<paragraph id="F844368E707BFFD5FF04525C3353CF81" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
Abdomen. Ventrite
<quantity id="3F039B6B707BFFD5FE68525C33CACF6D" box="[427,474,1627,1654]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="184" unit="in" value="5.0">5 in</quantity>
males apically flat, apical margin blunt, in females ventrite 5 vaulted and weakly pointed apicad.
</paragraph>
<paragraph id="F844368E707BFFD5FF0452A4334DCE31" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
Male terminalia. (
<figureCitation id="60C02A0B707BFFD5FE5052A43052CFA5" box="[403,578,1699,1725]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Figs 1719; 23</figureCitation>
) Penis elongate, parallel sided, apically triangular with prolonged tip (
<figureCitation id="60C02A0B707BFFD5FAA952A432F0CFF9" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Figs 1718</figureCitation>
), in lateral view regularly bowed, prolonged tip reflexed (
<figureCitation id="60C02A0B707BFFD5FCB752C031D8CFF9" box="[884,968,1735,1761]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Fig. 18</figureCitation>
). Tegmen with apodeme twice as long as parameres, those as long as width of tegminal ring. Sternite VIII (with hemisternite
<collectionCode id="9EEAAE4B707BFFD5FB9A52EC3668CE1D" box="[1113,1144,1771,1797]" pageId="3" pageNumber="184">IX</collectionCode>
) weakly S-shaped, 2/3 as long as aedeagus.
</paragraph>
<paragraph id="F844368E707BFFD5FF04533432D4CEC5" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="184">
Female terminalia. (
<figureCitation id="60C02A0B707BFFD5FE79533430B1CE55" box="[442,673,1843,1869]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Figs 2022; 2426</figureCitation>
) Gonocoxites dorso-ventrally flattened, without styli, proximal gonocoxites tube-like, distal gonocoxites heavier sclerotized towards tip and set with long light sensillae (
<figureCitation id="60C02A0B707BFFD5FAF853503780CE6A" box="[1339,1424,1879,1906]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Fig. 25</figureCitation>
). Sternite VIII with very long and slender apodeme, sclerotized part of plate strongly rounded and set with long hairs (
<figureCitation id="60C02A0B707BFFD5FF5C539832E3CEA1" box="[159,243,1951,1977]" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Fig. 20</figureCitation>
). Spermatheca shaped like number 5, with short recumbent cornu, and long tube-like and bent collum (
<figureCitation id="60C02A0B707BFFD5FAAC539832A7CEC5" captionStart="FIGURES 1730" captionStartId="5.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1837]" captionTargetId="figure-21@5.[151,1436,181,1837]" captionTargetPageId="5" captionText="FIGURES 1730. Figs 1718. Penis in ventral and lateral view of Phyllobius occultatus sp. nov. 19. Male sternite VIII and hemisternite IX, ditto. Fig. 20. Female sternite VIII, ditto. Fig. 21. Ovipositor in full length with gonocoxites, ditto. Fig. 22. Spermatheca, ditto. Figs 2324. Dorsal view on tergites VII and VIII of male and female, ditto. Figs 2526. Details of gonocoxites, lateral and dorso-ventral views, ditto. Fig. 2728. Penis of Ph. brenskei in ventral and lateral view. Figs 2930. Ditto, Ph. nigrofasciatus (Photos: C. Germann)." pageId="3" pageNumber="184">Fig. 22</figureCitation>
).
</paragraph>
<caption id="AC846606707CFFD2FF5452D53136CE4F" pageId="4" pageNumber="185" startId="4.[151,250,1746,1770]" targetBox="[152,1436,181,1723]" targetPageId="4" targetType="figure">
<paragraph id="F844368E707CFFD2FF5452D53136CE4F" blockId="4.[151,1437,1746,1879]" pageId="4" pageNumber="185">
<emphasis id="CA8FEA9C707CFFD2FF5452D53395CFF3" bold="true" box="[151,389,1746,1771]" pageId="4" pageNumber="185">FIGURE 916. Fig. 9.</emphasis>
Holotype male of
<taxonomicName id="3FFB4D0D707CFFD2FD8452D43132CFF3" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[583,802,1747,1771]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707CFFD2FD8452D43132CFF3" box="[583,802,1747,1771]" italics="true" pageId="4" pageNumber="185">Phyllobius occultatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BC57E7707CFFD2FCE952D43169CFF2" box="[810,889,1747,1770]" pageId="4" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="CA8FEA9C707CFFD2FC4352D531C0CFF3" bold="true" box="[896,976,1746,1771]" pageId="4" pageNumber="185">Fig. 10.</emphasis>
Ditto, paratype female.
<emphasis id="CA8FEA9C707CFFD2FB0952D53709CFF3" bold="true" box="[1226,1305,1746,1771]" pageId="4" pageNumber="185">Fig. 11.</emphasis>
Elytral apex of the female of
<taxonomicName id="3FFB4D0D707CFFD2FE8352F033C1CE17" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[320,465,1783,1807]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707CFFD2FE8352F033C1CE17" box="[320,465,1783,1807]" italics="true" pageId="4" pageNumber="185">Ph. occultatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BC57E7707CFFD2FE1B52F03037CE16" box="[472,551,1783,1806]" pageId="4" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="CA8FEA9C707CFFD2FDEE52F1306CCE17" bold="true" box="[557,636,1782,1807]" pageId="4" pageNumber="185">Fig. 12.</emphasis>
Ditto of the female of
<taxonomicName id="3FFB4D0D707CFFD2FCA552F03630CE17" authorityName="Pesarini. Thanks" authorityYear="1975" box="[870,1056,1783,1807]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="nigrofasciatus">
<emphasis id="CA8FEA9C707CFFD2FCA552F03630CE17" box="[870,1056,1783,1807]" italics="true" pageId="4" pageNumber="185">Ph. nigrofasciatus</emphasis>
</taxonomicName>
.
<emphasis id="CA8FEA9C707CFFD2FBEE52F136B6CE17" bold="true" box="[1069,1190,1782,1807]" pageId="4" pageNumber="185">Figs 1314.</emphasis>
Male and female of
<taxonomicName id="3FFB4D0D707CFFD2FAB952F032FCCE2B" authorityName="Schilsky" authorityYear="1911" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="brenskei">
<emphasis id="CA8FEA9C707CFFD2FAB952F032FCCE2B" italics="true" pageId="4" pageNumber="185">Ph. brenskei</emphasis>
</taxonomicName>
, freshly collected specimens showing variability in colour of scales and body shape.
<emphasis id="CA8FEA9C707CFFD2FB91531D36B2CE2B" bold="true" box="[1106,1186,1818,1843]" pageId="4" pageNumber="185">Fig. 15.</emphasis>
Apical sternites of male and
<emphasis id="CA8FEA9C707CFFD2FF01533832F2CE4F" bold="true" box="[194,226,1855,1879]" pageId="4" pageNumber="185">16.</emphasis>
Female of
<taxonomicName id="3FFB4D0D707CFFD2FE97533833F5CE4F" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[340,485,1855,1879]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707CFFD2FE97533833F5CE4F" box="[340,485,1855,1879]" italics="true" pageId="4" pageNumber="185">Ph. occultatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BC57E7707CFFD2FE2F5338302BCE4E" box="[492,571,1855,1878]" pageId="4" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
(Photos: C. Germann).
</paragraph>
</caption>
<caption id="AC846606707DFFD3FF545343306DCEF5" pageId="5" pageNumber="186" startId="5.[151,264,1860,1884]" targetBox="[151,1436,181,1837]" targetPageId="5" targetType="figure">
<paragraph id="F844368E707DFFD3FF545343306DCEF5" blockId="5.[151,1437,1860,2029]" pageId="5" pageNumber="186">
<emphasis id="CA8FEA9C707DFFD3FF54534333C8CE44" bold="true" box="[151,472,1860,1884]" pageId="5" pageNumber="186">FIGURES 1730. Figs 1718.</emphasis>
Penis in ventral and lateral view of
<taxonomicName id="3FFB4D0D707DFFD3FC905342363FCE45" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[851,1071,1861,1885]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707DFFD3FC905342363FCE45" box="[851,1071,1861,1885]" italics="true" pageId="5" pageNumber="186">Phyllobius occultatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BC57E7707DFFD3FBF453423697CE44" box="[1079,1159,1861,1884]" pageId="5" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="CA8FEA9C707DFFD3FB4D534336BECE44" bold="true" box="[1166,1198,1860,1884]" pageId="5" pageNumber="186">19.</emphasis>
Male sternite VIII and hemisternite IX, ditto.
<emphasis id="CA8FEA9C707DFFD3FE4A536F33CCCE98" bold="true" box="[393,476,1896,1920]" pageId="5" pageNumber="186">Fig. 20.</emphasis>
Female sternite VIII, ditto.
<emphasis id="CA8FEA9C707DFFD3FCC9536F314DCE98" bold="true" box="[778,861,1896,1920]" pageId="5" pageNumber="186">Fig. 21.</emphasis>
Ovipositor in full length with gonocoxites, ditto.
<emphasis id="CA8FEA9C707DFFD3FAB0536F32A7CEBC" bold="true" pageId="5" pageNumber="186">Fig. 22.</emphasis>
Spermatheca, ditto.
<emphasis id="CA8FEA9C707DFFD3FE4C538B301BCEBC" bold="true" box="[399,523,1932,1956]" pageId="5" pageNumber="186">Figs 2324.</emphasis>
Dorsal view on tergites VII and VIII of male and female, ditto.
<emphasis id="CA8FEA9C707DFFD3FB71538B373ECEBC" bold="true" box="[1202,1326,1932,1956]" pageId="5" pageNumber="186">Figs 2526.</emphasis>
Details of gonocoxites, lateral and dorso-ventral views, ditto.
<emphasis id="CA8FEA9C707DFFD3FD6453B7310ECED0" bold="true" box="[679,798,1968,1992]" pageId="5" pageNumber="186">Fig. 2728.</emphasis>
Penis of
<taxonomicName id="3FFB4D0D707DFFD3FC4253B63610CED1" authorityName="Schilsky" authorityYear="1911" box="[897,1024,1969,1993]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="brenskei">
<emphasis id="CA8FEA9C707DFFD3FC4253B63610CED1" box="[897,1024,1969,1993]" italics="true" pageId="5" pageNumber="186">Ph. brenskei</emphasis>
</taxonomicName>
in ventral and lateral view.
<emphasis id="CA8FEA9C707DFFD3FAE153B7378DCED0" bold="true" box="[1314,1437,1968,1992]" pageId="5" pageNumber="186">Figs 2930.</emphasis>
Ditto,
<taxonomicName id="3FFB4D0D707DFFD3FF1B53D23381CEF5" authorityName="Pesarini. Thanks" authorityYear="1975" box="[216,401,2005,2029]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="nigrofasciatus">
<emphasis id="CA8FEA9C707DFFD3FF1B53D23381CEF5" box="[216,401,2005,2029]" italics="true" pageId="5" pageNumber="186">Ph. nigrofasciatus</emphasis>
</taxonomicName>
(Photos: C. Germann).
</paragraph>
</caption>
<caption id="AC846606707EFFD0FF545004300ACD27" pageId="6" pageNumber="187" startId="6.[151,250,1027,1051]" targetBox="[151,1435,234,1003]" targetPageId="6" targetType="figure">
<paragraph id="F844368E707EFFD0FF545004300ACD27" blockId="6.[151,1436,1027,1087]" pageId="6" pageNumber="187">
<emphasis id="CA8FEA9C707EFFD0FF5450043330CD03" bold="true" box="[151,288,1027,1051]" pageId="6" pageNumber="187">FIGURE 31.</emphasis>
Habitat of
<taxonomicName id="3FFB4D0D707EFFD0FE515004307DCD03" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[402,621,1027,1051]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="187" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707EFFD0FE515004307DCD03" box="[402,621,1027,1051]" italics="true" pageId="6" pageNumber="187">Phyllobius occultatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BC57E7707EFFD0FDB0500330D2CD03" box="[627,706,1028,1051]" pageId="6" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
on Peloponnese (Marathos) on about 1300 m a.s.l. with forests of
<taxonomicName id="3FFB4D0D707EFFD0FAA750043303CD27" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="187" phylum="Tracheophyta" rank="species" species="cephalonica">
<emphasis id="CA8FEA9C707EFFD0FAA750043303CD27" italics="true" pageId="6" pageNumber="187">Abies cephalonica</emphasis>
</taxonomicName>
(Photo: H. Winkelmann).
</paragraph>
</caption>
<paragraph id="F844368E707EFFD0FF0450953649CDEC" blockId="6.[151,1436,1170,1448]" pageId="6" pageNumber="187">
Sexual dimorphism and variability. Males in dorsal view with elytra more parallel (in females more rounded in apical third), profemora more strongly swollen (
<figureCitation id="60C02A0B707EFFD0FD7150B13135CDC8" box="[690,805,1206,1232]" captionStart="FIGURE 916" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[152,1436,181,1723]" captionTargetId="figure-41@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 916. Fig. 9. Holotype male of Phyllobius occultatus sp. nov. Fig. 10. Ditto, paratype female. Fig. 11. Elytral apex of the female of Ph. occultatus sp. nov. Fig. 12. Ditto of the female of Ph. nigrofasciatus. Figs 1314. Male and female of Ph. brenskei, freshly collected specimens showing variability in colour of scales and body shape. Fig. 15. Apical sternites of male and 16. Female of Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="6" pageNumber="187">Figs 910</figureCitation>
) in males, less so in females. Head with rostrum slender (a bit more inflated in females), apex of elytra in females weakly mucronate (
<figureCitation id="60C02A0B707EFFD0FC3950DD365BCDEC" box="[1018,1099,1242,1268]" captionStart="FIGURE 916" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[152,1436,181,1723]" captionTargetId="figure-41@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 916. Fig. 9. Holotype male of Phyllobius occultatus sp. nov. Fig. 10. Ditto, paratype female. Fig. 11. Elytral apex of the female of Ph. occultatus sp. nov. Fig. 12. Ditto of the female of Ph. nigrofasciatus. Figs 1314. Male and female of Ph. brenskei, freshly collected specimens showing variability in colour of scales and body shape. Fig. 15. Apical sternites of male and 16. Female of Ph. occultatus sp. nov. (Photos: C. Germann)." pageId="6" pageNumber="187">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="F844368E707EFFD0FF0450F9313ACC24" blockId="6.[151,1436,1170,1448]" pageId="6" pageNumber="187">
Derivation of name: The name
<taxonomicName id="3FFB4D0D707EFFD0FDE950F9308DCC00" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[554,669,1278,1304]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="187" phylum="Arthropoda" rank="species" species="occultatus">
<emphasis id="CA8FEA9C707EFFD0FDE950F9308DCC00" box="[554,669,1278,1304]" italics="true" pageId="6" pageNumber="187">occultatus</emphasis>
</taxonomicName>
derives from the Latin verb “occultare” in English “to hide”, as the new species was hidden in the catalogue of Greek weevils.
</paragraph>
<paragraph id="F844368E707EFFD0FF0451413099CCB0" blockId="6.[151,1436,1170,1448]" pageId="6" pageNumber="187">
Ecological note: Most specimens were collected on higher elevations (
<quantity id="3F039B6B707EFFD0FC1651423673CC78" box="[981,1123,1349,1376]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="6" pageNumber="187" unit="m" value="900.0" valueMax="1300.0" valueMin="500.0">5001300 m</quantity>
a.s.l.) on
<taxonomicName id="3FFB4D0D707EFFD0FB0C514132FECC9C" authority="Loudon" authorityName="Loudon" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="187" phylum="Tracheophyta" rank="species" species="cephalonica">
<emphasis id="CA8FEA9C707EFFD0FB0C5141378CCC78" box="[1231,1436,1350,1376]" italics="true" pageId="6" pageNumber="187">Abies cephalonica</emphasis>
Loudon
</taxonomicName>
(
<figureCitation id="60C02A0B707EFFD0FF3E516D335FCC9C" box="[253,335,1386,1412]" captionStart="FIGURE 31" captionStartId="6.[151,250,1027,1051]" captionTargetBox="[151,1435,234,1003]" captionTargetId="figure-41@6.[151,1435,234,1003]" captionTargetPageId="6" captionText="FIGURE 31. Habitat of Phyllobius occultatus sp. nov. on Peloponnese (Marathos) on about 1300 m a.s.l. with forests of Abies cephalonica (Photo: H. Winkelmann)." pageId="6" pageNumber="187">Fig. 31</figureCitation>
), which seems to be the preferred host of
<taxonomicName id="3FFB4D0D707EFFD0FCE1516D31D2CC9C" authorityName="Germann &amp; Wartmann &amp; Winkelmann" authorityYear="2025" box="[802,962,1386,1412]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="187" phylum="Arthropoda" rank="species" species="occultatus" status="sp. nov.">
<emphasis id="CA8FEA9C707EFFD0FCE1516D31D2CC9C" box="[802,962,1386,1412]" italics="true" pageId="6" pageNumber="187">Ph. occultatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BC57E7707EFFD0FC0A516D360FCC9C" box="[969,1055,1386,1412]" pageId="6" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
In contrast to that observation, all
<taxonomicName id="3FFB4D0D707EFFD0FF5451893332CCB0" authorityName="Schilsky" authorityYear="1911" box="[151,290,1422,1448]" class="Insecta" family="Curculionidae" genus="Phyllobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="187" phylum="Arthropoda" rank="species" species="brenskei">
<emphasis id="CA8FEA9C707EFFD0FF5451893332CCB0" box="[151,290,1422,1448]" italics="true" pageId="6" pageNumber="187">Ph. brenskei</emphasis>
</taxonomicName>
were collected on
<taxonomicName id="3FFB4D0D707EFFD0FE3551893099CCB0" box="[502,649,1422,1448]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="187" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="CA8FEA9C707EFFD0FE3551893044CCB0" box="[502,596,1422,1448]" italics="true" pageId="6" pageNumber="187">Quercus</emphasis>
spp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,79 +1,81 @@
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<mods:namePart id="6A92260028E83360D50A459FAB4A64FF">Mahmood, Khalid</mods:namePart>
<mods:affiliation id="F11E0A1E732E3CB45A4C7F9EC5181D5B">Dr. Abu Bakr Fossil Display &amp; Research Centre, Department of Zoology, University of the Punjab, Quid-e-Azam Campus, Lahore 54590 (Pakistan) and Department of Zoology, Govt. Islamia Graduate College Kasur, Punjab (Pakistan) khalidkasuri 1 @ gmail. com (corresponding author)</mods:affiliation>
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<mods:affiliation id="1A08F3829BDC9C0C54FA8AF950AEBA24">Dr. Abu Bakr Fossil Display &amp; Research Centre, Department of Zoology, University of the Punjab, Quid-e-Azam Campus, Lahore 54590 (Pakistan) and Department of Zoology, Govt. Islamia Graduate College Kasur, Punjab (Pakistan) khalidkasuri 1 @ gmail. com (corresponding author)</mods:affiliation>
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