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data/27/C7/3C/27C73C8902D0594AB24F5F4699BA0D99.xml
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, left mandible with p4-m1 and alveoli of c1-p3 (
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).
</materialsCitation>
</paragraph>
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. — Hasnot,
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,
<collectingCountry id="0DB2E295FFBCA916FB62FEF6AAB92116" box="[1169,1250,346,370]" name="Pakistan" pageId="7" pageNumber="306">Pakistan</collectingCountry>
(
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).
</paragraph>
</subSubSection>
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<paragraph id="751AA205FFBCA916FCDEFE26AA2721B2" blockId="7.[812,1457,394,471]" pageId="7" pageNumber="306">
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. — Large species of the genus
<taxonomicName id="B2A5D986FFBCA916FB2BFE26AB7721C5" authorityName="Pilgrim" authorityYear="1931" box="[1240,1324,394,417]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FB2BFE26AB7721C5" box="[1240,1324,394,417]" italics="true" pageId="7" pageNumber="306">Sivaonyx</emphasis>
</taxonomicName>
with slightly sectorial aspect to P4 and m1, but showing a talonid basin broader than the trigonid (
<bibRefCitation id="1134DFF4FFBCA916FC11FE13AA3421B3" author="PICKFORD M." box="[994,1135,447,471]" pageId="7" pageNumber="306" pagination="83 - 127" refId="ref9573" refString="PICKFORD M. 2007. - Revision of the Mio-Pliocene otter-like mammals of the Indian Subcontinent. Estudios Geologicos 63: 83-127. https://doi.org/10.3989/egeol.07631192" type="journal article" year="2007">Pickford 2007</bibRefCitation>
).
</paragraph>
<paragraph id="751AA205FFBCA916FCDEFE43AA1E2231" blockId="7.[811,1455,494,598]" pageId="7" pageNumber="306">
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. — This species well-known from the Late Miocene, Dhok Pathan Formation of Siwaliks (
<bibRefCitation id="1134DFF4FFBCA916FC3FFD8FAA10225F" author="PILGRIM G. E." box="[972,1099,547,571]" pageId="7" pageNumber="306" pagination="1 - 232" refId="ref9719" refString="PILGRIM G. E. 1932. - The fossil Carnivora of India. Memoirs of the Geological Survey of India. Palaeontologia indica NS 18: 1-232." type="journal article" year="1932">Pilgrim 1932</bibRefCitation>
;
<bibRefCitation id="1134DFF4FFBCA916FBA5FD8FAABA225F" author="PICKFORD M." box="[1110,1249,547,571]" pageId="7" pageNumber="306" pagination="83 - 127" refId="ref9573" refString="PICKFORD M. 2007. - Revision of the Mio-Pliocene otter-like mammals of the Indian Subcontinent. Estudios Geologicos 63: 83-127. https://doi.org/10.3989/egeol.07631192" type="journal article" year="2007">Pickford 2007</bibRefCitation>
) and Southern
<collectingCountry id="0DB2E295FFBCA916FA80FD8FABF4225F" box="[1395,1455,547,571]" name="China" pageId="7" pageNumber="306">China</collectingCountry>
(
<bibRefCitation id="1134DFF4FFBCA916FCC6FD92ADF82232" author="ZONG G." box="[821,931,574,598]" pageId="7" pageNumber="306" pagination="69 - 89" refId="ref10685" refString="ZONG G. 1997. - Carnivora, in HE Z. (ed.), Yunamou Hominoid Fauna. Yunnan Science and Technology Press: 69-89." type="book chapter" year="1997">Zong 1997</bibRefCitation>
;
<bibRefCitation id="1134DFF4FFBCA916FC5CFD92AA622232" author="QI G. &amp; DONG W. &amp; ZHENG L. &amp; ZHAO L. &amp; GAO F. &amp; YUE L. &amp; ZHANG Y." box="[943,1081,574,598]" pageId="7" pageNumber="306" pagination="704 - 712" refId="ref9866" refString="QI G., DONG W., ZHENG L., ZHAO L., GAO F., YUE L. &amp; ZHANG Y. 2006. - Taxonomy, age and environment status of the Yuanmou hominoids. Chinese Science Bulletin 51 (6): 704-712. https://doi.org/10.1007/s11434-006-0704-5" type="journal article" year="2006">
Qi
<emphasis id="47D17E17FFBCA916FC23FD93ADA42232" box="[976,1023,574,598]" italics="true" pageId="7" pageNumber="306">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DBFF18EFFBCA916FCDEFDC2AAB92669" pageId="7" pageNumber="306" type="description">
<paragraph id="751AA205FFBCA916FCDEFDC2AA8E22E1" blockId="7.[813,1237,621,646]" box="[813,1237,621,646]" pageId="7" pageNumber="306">
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<smallCapsWord id="73FC34D9FFBCA916FC65FDDFADAF22E0" baselines="640" box="[918,1012,627,644]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="7" pageNumber="306">MATERIAL</smallCapsWord>
. — PUPC 16/29, lp4.
</paragraph>
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<paragraph id="751AA205FFBCA916FCDEFD7AAA1C246B" blockId="7.[813,1457,694,1039]" pageId="7" pageNumber="306">PUPC 16/29 is a left p4. The base of tooth is partially broken postero-lingually. It is monocuspid, slightly worn and subtriangular in shape. It has a thick and strong cingulum at the base labio-lingually. The protoconid is large, pyramidal, slightly compressed buccally and highest. The distal cuspid is small, slightly buccally positioned, and separated from the protoconid by a small notch. A wide basal platform is present in the front of the distal cuspid lingually. It is double-rooted, and the distal root is more preserved (Fig. 2C1-C3).</paragraph>
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</paragraph>
<paragraph id="751AA205FFBCA916FCDEFBF8AAB92669" blockId="7.[811,1457,1076,1549]" pageId="7" pageNumber="306">
The described p4, PUPC 16/29, is broad distally, with a strong, and a broad cingulid, no mesial accessory cusp, outwardly situated distal cusp; and these are the characters of p4 given in the diagnosis of
<taxonomicName id="B2A5D986FFBCA916FB89FB18AB3524AA" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[1146,1390,1204,1230]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBCA916FB89FB18AB3524AA" box="[1146,1390,1204,1230]" italics="true" pageId="7" pageNumber="306">Sivaonyx bathygnathus</emphasis>
</taxonomicName>
given by
<bibRefCitation id="1134DFF4FFBCA916FCA2FB78ADA1248A" author="PILGRIM G. E." box="[849,1018,1236,1262]" pageId="7" pageNumber="306" pagination="1 - 232" refId="ref9719" refString="PILGRIM G. E. 1932. - The fossil Carnivora of India. Memoirs of the Geological Survey of India. Palaeontologia indica NS 18: 1-232." type="journal article" year="1932">Pilgrim (1932)</bibRefCitation>
. Further, while describing p4 (GSI-D 244),
<bibRefCitation id="1134DFF4FFBCA916FC9EFB5FAA4B256A" author="PILGRIM G. E." box="[877,1040,1267,1294]" pageId="7" pageNumber="306" pagination="1 - 232" refId="ref9719" refString="PILGRIM G. E. 1932. - The fossil Carnivora of India. Memoirs of the Geological Survey of India. Palaeontologia indica NS 18: 1-232." type="journal article" year="1932">Pilgrim (1932)</bibRefCitation>
writes: “distal accessory cusp is almost as strong and probably little inferior in height to the main cusp and it lies well on the outside of the median axis” and the same morphology is described by
<bibRefCitation id="1134DFF4FFBCA916FB35FAFFAB2C2509" author="PICKFORD M." box="[1222,1399,1363,1389]" pageId="7" pageNumber="306" pagination="83 - 127" refId="ref9573" refString="PICKFORD M. 2007. - Revision of the Mio-Pliocene otter-like mammals of the Indian Subcontinent. Estudios Geologicos 63: 83-127. https://doi.org/10.3989/egeol.07631192" type="journal article" year="2007">Pickford (2007)</bibRefCitation>
, i.e., presence of a distal accessory cusp, a cingulid that covers the base of whole tooth, and a broadly expanded distal platform. In this regard, morphology and dimensions of the specimen (Fig. 2C1-C3;
<tableCitation id="382797BEFFBCA916FC3DFA7FAA7B2589" box="[974,1056,1491,1517]" captionStart="TABLE" captionStartId="10.[133,143,219,236]" captionText="TABLE 2. — Comparative measurements of Enhydriodon cf. falconeri, Sivaonyx sarwari n. sp. and Sivaonyx bathygnathus (Lydekker, 1884) (in mm). Comparative data taken from Pilgrim (1932), Colbert (1935), Chow (1961), Qi (1983), Zong (1997), Werdelin (2003), Morales &amp; Pickford (2005), Morales et al. (2005), Pickford (2007), Peigné et al. (2008), Geraads et al. (2011), Werdelin &amp; Manthi (2012), Grohé et al. (2013, 2022), Werdelin &amp; Lewis (2013), Peigné (2016), Koufos et al. (2018) and Valenciano &amp; Govender (2020). *, studied specimens; **, measurements as preserved; est., estimated measurements." pageId="7" pageNumber="306">Table 2</tableCitation>
) are close to
<taxonomicName id="B2A5D986FFBCA916FB43FA7FAB022589" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[1200,1369,1491,1517]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBCA916FB43FA7FAAE62589" box="[1200,1213,1491,1517]" italics="true" pageId="7" pageNumber="306">S</emphasis>
.
<emphasis id="47D17E17FFBCA916FB3FFA7FAB022589" box="[1228,1369,1491,1517]" italics="true" pageId="7" pageNumber="306">bathygnathus</emphasis>
</taxonomicName>
, hence, we referred it to
<taxonomicName id="B2A5D986FFBCA916FC14FA5FAA802669" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[999,1243,1523,1549]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBCA916FC14FA5FAA802669" box="[999,1243,1523,1549]" italics="true" pageId="7" pageNumber="306">Sivaonyx bathygnathus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

107
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@ -0,0 +1,107 @@
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. — After Dr Muhammad Sarwar, the second vertebrate palaeontologist of the
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University and who have done extensive field work during the 1963-1977 throughout
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, that resulted in the identification and exploration of new localities and became a source of guidance for the field work for the future palaeontologists of
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.
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. — Hasnot, Jhelum,
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,
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,
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cf.
<emphasis id="47D17E17FFBDA917FE55FA67AFB525B8" box="[422,494,1483,1500]" italics="true" pageId="6" pageNumber="305">falconeri</emphasis>
</taxonomicName>
:
<emphasis id="47D17E17FFBDA917FE0AFA67AC4A25B8" bold="true" box="[505,529,1483,1500]" pageId="6" pageNumber="305">A4</emphasis>
, PUPC 83/01, right mandibular fragment with m1 and root of m2;
<emphasis id="47D17E17FFBDA917FBBFFA67AA3F25B8" bold="true" box="[1100,1124,1483,1500]" pageId="6" pageNumber="305">A5</emphasis>
, GSI-D 161 from
<bibRefCitation id="1134DFF4FFBDA917FB0EFA67AB3325B8" author="PILGRIM G. E." box="[1277,1384,1483,1500]" pageId="6" pageNumber="305" pagination="1 - 232" refId="ref9719" refString="PILGRIM G. E. 1932. - The fossil Carnivora of India. Memoirs of the Geological Survey of India. Palaeontologia indica NS 18: 1-232." type="journal article" year="1932">Pilgrim 1932</bibRefCitation>
;
<emphasis id="47D17E17FFBDA917FA80FA67ABF125B8" bold="true" box="[1395,1450,1483,1500]" pageId="6" pageNumber="305">B1-B3</emphasis>
,
<taxonomicName id="B2A5D986FFBDA917FF77FA4EAF542597" authorityName="Mahmood &amp; Morlo &amp; Abbas &amp; Babar &amp; Khan" authorityYear="2025" box="[132,271,1506,1523]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="sarwari" status="sp. nov.">
<emphasis id="47D17E17FFBDA917FF77FA4EAF542597" box="[132,271,1506,1523]" italics="true" pageId="6" pageNumber="305">Sivaonyx sarwari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5CE2C36CFFBDA917FEE6FA4EAF1C2597" box="[277,327,1506,1523]" pageId="6" pageNumber="305" rank="species">n. sp.</taxonomicNameLabel>
; PUPC 17/64, lP4;
<emphasis id="47D17E17FFBDA917FE14FA4EAC452597" bold="true" box="[487,542,1506,1523]" pageId="6" pageNumber="305">C1-C3</emphasis>
,
<taxonomicName id="B2A5D986FFBDA917FDDAFA4EAD1F2597" authority="(Lydekker, 1884)" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[553,836,1506,1523]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBDA917FDDAFA4EAC6E2597" box="[553,565,1506,1523]" italics="true" pageId="6" pageNumber="305">S</emphasis>
.
<emphasis id="47D17E17FFBDA917FDB3FA4EACEB2597" box="[576,688,1506,1523]" italics="true" pageId="6" pageNumber="305">bathygnathus</emphasis>
(
<bibRefCitation id="1134DFF4FFBDA917FD49FA4EAD642597" author="LYDEKKER R." box="[698,831,1506,1523]" pageId="6" pageNumber="305" pagination="1 - 258" refId="ref9028" refString="LYDEKKER R. 1884. - Indian Tertiary and post-Tertiary Vertebrata. Memoirs of the Geological Survey of India. Palaeontologia indica (3): 1-258." type="journal article" year="1884">Lydekker, 1884</bibRefCitation>
)
</taxonomicName>
, PUPC 16/29, lp4.
<emphasis id="47D17E17FFBDA917FC10FA4EADA12597" bold="true" box="[995,1018,1506,1523]" pageId="6" pageNumber="305">A1</emphasis>
,
<emphasis id="47D17E17FFBDA917FBF6FA4EAA472597" bold="true" box="[1029,1052,1506,1523]" pageId="6" pageNumber="305">A4</emphasis>
,
<emphasis id="47D17E17FFBDA917FBD4FA4EAA652597" bold="true" box="[1063,1086,1506,1523]" pageId="6" pageNumber="305">A5</emphasis>
,
<emphasis id="47D17E17FFBDA917FBBAFA4EAA3B2597" bold="true" box="[1097,1120,1506,1523]" pageId="6" pageNumber="305">B1</emphasis>
,
<emphasis id="47D17E17FFBDA917FB98FA4EAAD82597" bold="true" box="[1131,1155,1506,1523]" pageId="6" pageNumber="305">C1</emphasis>
, occlusal views;
<emphasis id="47D17E17FFBDA917FAE1FA4EAB722597" bold="true" box="[1298,1321,1506,1523]" pageId="6" pageNumber="305">A2</emphasis>
,
<emphasis id="47D17E17FFBDA917FAC7FA4EAB172597" bold="true" box="[1332,1356,1506,1523]" pageId="6" pageNumber="305">B2</emphasis>
,
<emphasis id="47D17E17FFBDA917FAA5FA4EAB352597" bold="true" box="[1366,1390,1506,1523]" pageId="6" pageNumber="305">C2</emphasis>
, lingual views;
<emphasis id="47D17E17FFBDA917FF4DFA54AE8E266D" bold="true" box="[190,213,1528,1545]" pageId="6" pageNumber="305">A3</emphasis>
,
<emphasis id="47D17E17FFBDA917FF2CFA54AEAC266D" bold="true" box="[223,247,1528,1545]" pageId="6" pageNumber="305">B3</emphasis>
,
<emphasis id="47D17E17FFBDA917FEF2FA54AF42266D" bold="true" box="[257,281,1528,1545]" pageId="6" pageNumber="305">C3</emphasis>
, buccal views. Scale bars: A1-A3, B, C, 10 mm; A4, A5, not to scale.
</paragraph>
</caption>
<subSubSection id="3DBFF18EFFBDA917FF77F9FDAA45278F" pageId="6" pageNumber="305" type="diagnosis">
<paragraph id="751AA205FFBDA917FF77F9FDAC1C2691" blockId="6.[132,776,1617,1781]" pageId="6" pageNumber="305">
DIAGNOSIS. — A small
<taxonomicName id="B2A5D986FFBDA917FE88F9FDAC47260D" box="[379,540,1617,1641]" pageId="6" pageNumber="305">
<emphasis id="47D17E17FFBDA917FE88F9FDAF8A260C" box="[379,465,1617,1640]" italics="true" pageId="6" pageNumber="305">Sivaonyx</emphasis>
species
</taxonomicName>
having P4 with round borders/external walls, basal lingual cingulum is straight at mesial border, postparacrista short and having less buccal concavity, lack a cusplet between proto- and hypocone, small mesial valley hence, less pinched mesial border, and very long posthypocrista, the crest that joins hypocone with metastyle/metacone.
</paragraph>
<paragraph id="751AA205FFBDA917FF77F8A3AA45278F" blockId="6.[131,777,1807,2027]" lastBlockId="6.[811,1456,1616,2027]" pageId="6" pageNumber="305">
<smallCapsWord id="73FC34D9FFBDA917FF77F8A3AF4F2741" baselines="1824,1825" box="[132,276,1807,1830]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Differential" pageId="6" pageNumber="305">DIFFERENTIAL</smallCapsWord>
<smallCapsWord id="73FC34D9FFBDA917FEEFF8B8AFDD2741" baselines="1825" box="[284,390,1812,1829]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="6" pageNumber="305">DIAGNOSIS</smallCapsWord>
. — Differs from all
<taxonomicName id="B2A5D986FFBDA917FD97F8A3AD5C2743" box="[612,775,1807,1831]" pageId="6" pageNumber="305">
<emphasis id="47D17E17FFBDA917FD97F8A3ACE02742" box="[612,699,1807,1830]" italics="true" pageId="6" pageNumber="305">Sivaonyx</emphasis>
species
</taxonomicName>
being small in size, having round outline, basal lingual cingulum is straight at mesial border, and small mesial valley hence, less pinched mesial border. It differs from
<taxonomicName id="B2A5D986FFBDA917FDFCF8CFACB7271F" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[527,748,1891,1915]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBDA917FDFCF8CFACB7271F" box="[527,748,1891,1915]" italics="true" pageId="6" pageNumber="305">Sivaonyx bathygnathus</emphasis>
</taxonomicName>
in having short postparacrista, less spaced proto- and hypocone, and having no cusplet between proto- and hypocone; from
<taxonomicName id="B2A5D986FFBDA917FD40F830AF5127AB" authorityName="Pickford" authorityYear="2007" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="gandakasensis">
<emphasis id="47D17E17FFBDA917FD40F830AF5127AB" italics="true" pageId="6" pageNumber="305">Sivaonyx gandakasensis</emphasis>
</taxonomicName>
in short postparacrista, long posthypocrista, and more buccal concavity of postparacrista; from
<taxonomicName id="B2A5D986FFBDA917FD90F878AD9F260C" authority="Lydekker, 1890" authorityName="Lydekker" authorityYear="1890" box="[611,964,1616,2027]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="hessicus">
<emphasis id="47D17E17FFBDA917FD90F878AD53278F" box="[611,776,2003,2027]" italics="true" pageId="6" pageNumber="305">Sivaonyx hessicus</emphasis>
<bibRefCitation id="1134DFF4FFBDA917FCDEF9FCAD9F260C" author="LYDEKKER R." box="[813,964,1616,1640]" pageId="6" pageNumber="305" pagination="3 - 5" refId="ref9058" refString="LYDEKKER R. 1890. - On a new species of otter from the lower Pliocene of Eppelsheim. Proceedings of the Zoological Society of London 1890: 3-5. https://www.biodiversitylibrary.org/page/31014104" type="book chapter" year="1890">
Lydekker,
<quantity id="B25D0FE0FFBDA917FC62F9FCAD9F260C" box="[913,964,1616,1640]" metricMagnitude="1" metricUnit="m" metricValue="4.8006" pageId="6" pageNumber="305" unit="in" value="1890.0">1890</quantity>
</bibRefCitation>
</taxonomicName>
in having weak buccal concavity of postparacrista, no cusplet between proto- and hypocone, very long posthypocrista; from
<taxonomicName id="B2A5D986FFBDA917FC6DF926ABC826C5" authority="Morales, Pickford &amp; Soria, 2005" authorityName="Morales, Pickford &amp; Soria" authorityYear="2005" box="[926,1427,1673,1698]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="hendeyi">
<emphasis id="47D17E17FFBDA917FC6DF926AA1826C6" box="[926,1091,1674,1698]" italics="true" pageId="6" pageNumber="305">Sivaonyx hendeyi</emphasis>
<bibRefCitation id="1134DFF4FFBDA917FBBAF926ABC826C5" author="MORALES J. &amp; PICKFORD M. &amp; SORIA D." box="[1097,1427,1673,1698]" pageId="6" pageNumber="305" pagination="39 - 61" refId="ref9288" refString="MORALES J., PICKFORD M. &amp; SORIA D. 2005. - Carnivores from the late Miocene and Basal Pliocene of the Tugen Hills, Kenya. Revista de la Societad Geologica de Espana 18: 39-61." type="journal article" year="2005">
Morales, Pickford &amp; Soria,
<quantity id="B25D0FE0FFBDA917FA93F925ABC826C5" box="[1376,1427,1673,1697]" metricMagnitude="1" metricUnit="m" metricValue="5.0927" pageId="6" pageNumber="305" unit="in" value="2005.0">2005</quantity>
</bibRefCitation>
</taxonomicName>
in lacking notch between paracone and metastyle, having very long posthypocrista, and weak buccal cingulum; from
<taxonomicName id="B2A5D986FFBDA917FB0EF96FAD932693" authority="Werdelin, 2003" authorityName="Werdelin" authorityYear="2003" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="ekecaman">
<emphasis id="47D17E17FFBDA917FB0EF96FABF526BF" box="[1277,1454,1731,1755]" italics="true" pageId="6" pageNumber="305">Sivaonyx ekecaman</emphasis>
<bibRefCitation id="1134DFF4FFBDA917FCDEF973AD932693" author="WERDELIN L." box="[813,968,1759,1783]" pageId="6" pageNumber="305" pagination="115 - 132" refId="ref10443" refString="WERDELIN L. 2003. - Carnivores from the Kanapoi Hominid site, Turkana Basin, northern Kenya. Contributions in Science 498: 115-132. https://doi.org/10.5962/p.214388" type="journal article" year="2003">
Werdelin,
<quantity id="B25D0FE0FFBDA917FC66F973AD932693" box="[917,968,1759,1783]" metricMagnitude="1" metricUnit="m" metricValue="5.087619999999999" pageId="6" pageNumber="305" unit="in" value="2003.0">2003</quantity>
</bibRefCitation>
</taxonomicName>
in having longer pre- and postparacrista, slightly distally positioned protocone, lack of notch between paracone and metastyle, weak buccal concavity of postparacrista, no cusplet between proto- and hypocone, large median valley, and very long posthypocrista; and from
<taxonomicName id="B2A5D986FFBDA917FBDDF8FEAA9B270E" authorityName="Morales &amp; Pickford, 2005 BAR" authorityYear="1720" box="[1070,1216,1874,1898]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="305" phylum="Chordata" rank="species" species="soriae">
<emphasis id="47D17E17FFBDA917FBDDF8FEAA9B270E" box="[1070,1216,1874,1898]" italics="true" pageId="6" pageNumber="305">Sivaonyx soriae</emphasis>
</taxonomicName>
in having slightly distal position of protocone, weak buccal concavity of postparacrista, no cusplet between proto- and hypocone, very long posthypocrista.
<smallCapsWord id="73FC34D9FFBDA917FCDEF81BAD9F27A9" baselines="1993,1993" box="[813,964,1975,1999]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Stratigraphic" pageId="6" pageNumber="305">STRATIGRAPHIC</smallCapsWord>
<smallCapsWord id="73FC34D9FFBDA917FC39F810AA5127A9" baselines="1993" box="[970,1034,1980,1997]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="range" pageId="6" pageNumber="305">RANGE</smallCapsWord>
. — Dhok Pathan Formation of the Middle Siwaliks (Late Miocene).
</paragraph>
</subSubSection>
<subSubSection id="3DBFF18EFFBCA916FF77FF74AFB8278F" pageId="7" pageNumber="306" type="description">
<paragraph id="751AA205FFBCA916FF77FF74AF4E2094" blockId="7.[131,777,216,911]" box="[132,277,216,242]" pageId="7" pageNumber="306">
<smallCapsWord id="73FC34D9FFBCA916FF77FF74AF4E2094" baselines="235,235" box="[132,277,216,242]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="306">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="751AA205FFBCA916FF77FF5BAF3A23EB" blockId="7.[131,777,216,911]" pageId="7" pageNumber="306">PUPC 17/64 is nearly rectangular with round outline, well-preserved and slightly worn (Fig. 2B1-B3). All the cusps are well-developed and prominent. The paracone is the highest, massive, with a rounded external wall and a prominent parastyle. The parastyle is low in height and it is separated from paracone by a small notch. Three crests originate from the paracone, mesial crest (preparacrista) connects the paracone to the parastyle, distal (postparacrista) to the metacone and the metacrista, and the third (lingual crest) connects to the protocone through crista obliqua. The metacrista is small and the metastyle/metacone is bulbous. The paracone is connected to the metacone. A prominent slanting cingulum is also present between the paracone and metacone and is connected to metastyle. The coupled protocone and hypocone are well-developed and distinct, separated by only a small notch. The protocone is slightly higher than the hypocone. The hypocrista is extremely long and becomes slightly round distally, rendering tooth round outline. The median fossette is large, oval in shape and separates trigon and talon. A thick cingulum fully covers the base of the tooth that has three roots (Fig. 2B1-B3).</paragraph>
<paragraph id="751AA205FFBCA916FF77FC19AF4E23A9" blockId="7.[130,777,949,2028]" box="[132,277,949,975]" pageId="7" pageNumber="306">
<smallCapsWord id="73FC34D9FFBCA916FF77FC19AF4E23A9" baselines="968,968" box="[132,277,949,975]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparison" pageId="7" pageNumber="306">COMPARISON</smallCapsWord>
</paragraph>
<paragraph id="751AA205FFBCA916FF77FC78AFB8278F" blockId="7.[130,777,949,2028]" pageId="7" pageNumber="306">
The presence of a protocone far from the paracone and a large cingulum that covers the base of the tooth in PUPC 17/64 differentiate it from
<taxonomicName id="B2A5D986FFBCA916FEA9FBB8AF8F244A" authorityName="Hürzeler" authorityYear="1987" box="[346,468,1044,1070]" class="Mammalia" family="Mustelidae" genus="Paludolutra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FEA9FBB8AF8F244A" box="[346,468,1044,1070]" italics="true" pageId="7" pageNumber="306">Paludolutra</emphasis>
</taxonomicName>
, and presence of a less developed hypocone, a protocone far from the paracone, a more sectorial paracone-metastyle, and that it is longer than wide in addition to its small size differentiate it from
<taxonomicName id="B2A5D986FFBCA916FD8AFBD8ACA524EA" authorityName="Falconer" authorityYear="1868" box="[633,766,1140,1166]" class="Mammalia" family="Mustelidae" genus="Enhydriodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FD8AFBD8ACA524EA" box="[633,766,1140,1166]" italics="true" pageId="7" pageNumber="306">Enhydriodon</emphasis>
</taxonomicName>
.
<taxonomicName id="B2A5D986FFBCA916FF77FB38AEA024CA" authorityName="Pilgrim" authorityYear="1932" box="[132,251,1172,1198]" class="Mammalia" family="Mustelidae" genus="Vishnuonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FF77FB38AEA024CA" box="[132,251,1172,1198]" italics="true" pageId="7" pageNumber="306">Vishnuonyx</emphasis>
</taxonomicName>
and
<taxonomicName id="B2A5D986FFBCA916FED8FB38AFDD24CA" authorityName="Pilgrim" authorityYear="1931" box="[299,390,1172,1198]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FED8FB38AFDD24CA" box="[299,390,1172,1198]" italics="true" pageId="7" pageNumber="306">Sivaonyx</emphasis>
</taxonomicName>
are the other two “enhydriodontine” genera present in the Siwaliks. PUPC 17/64 differs from the P4 of all
<taxonomicName id="B2A5D986FFBCA916FF17FB78AFF6248A" box="[228,429,1236,1262]" pageId="7" pageNumber="306">
<emphasis id="47D17E17FFBCA916FF17FB78AF06248A" box="[228,349,1236,1262]" italics="true" pageId="7" pageNumber="306">Vishnuonyx</emphasis>
species
</taxonomicName>
in having round outline, robust parastyle, short metacrista, very long posthypocrista, and wider central basin in addition to incomplete lingual shelf. Even the space between the protocone and hypocone is smaller in PUPC 17/64 than in
<taxonomicName id="B2A5D986FFBCA916FE88FAFFAFAE2509" authorityName="Pilgrim" authorityYear="1932" box="[379,501,1363,1389]" class="Mammalia" family="Mustelidae" genus="Vishnuonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FE88FAFFAFAE2509" box="[379,501,1363,1389]" italics="true" pageId="7" pageNumber="306">Vishnuonyx</emphasis>
</taxonomicName>
while the space between the parastyle and paracone is wider in
<taxonomicName id="B2A5D986FFBCA916FDD7FADFACC625E9" authorityName="Pilgrim" authorityYear="1932" box="[548,669,1395,1421]" class="Mammalia" family="Mustelidae" genus="Vishnuonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FDD7FADFACC625E9" box="[548,669,1395,1421]" italics="true" pageId="7" pageNumber="306">Vishnuonyx</emphasis>
</taxonomicName>
.
<taxonomicName id="B2A5D986FFBCA916FD58FADFAD5C25E9" authorityName="Pilgrim" authorityYear="1931" box="[683,775,1395,1421]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FD58FADFAD5C25E9" box="[683,775,1395,1421]" italics="true" pageId="7" pageNumber="306">Sivaonyx</emphasis>
</taxonomicName>
is represented by two species, a small,
<taxonomicName id="B2A5D986FFBCA916FDD7FA3FAEC125A9" authorityName="Pickford" authorityYear="2007" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="gandakasensis">
<emphasis id="47D17E17FFBCA916FDD7FA3FAEC125A9" italics="true" pageId="7" pageNumber="306">Sivaonyx gandakasensis</emphasis>
</taxonomicName>
and a large,
<taxonomicName id="B2A5D986FFBCA916FEDBFA1FAF9725A9" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[296,460,1459,1485]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBCA916FEDBFA1FAF6F25A9" box="[296,308,1459,1485]" italics="true" pageId="7" pageNumber="306">S</emphasis>
.
<emphasis id="47D17E17FFBCA916FEB0FA1FAF9725A9" box="[323,460,1459,1485]" italics="true" pageId="7" pageNumber="306">bathygnathus</emphasis>
</taxonomicName>
(
<bibRefCitation id="1134DFF4FFBCA916FE2EFA1FAC2325A9" author="PICKFORD M." box="[477,632,1459,1485]" pageId="7" pageNumber="306" pagination="83 - 127" refId="ref9573" refString="PICKFORD M. 2007. - Revision of the Mio-Pliocene otter-like mammals of the Indian Subcontinent. Estudios Geologicos 63: 83-127. https://doi.org/10.3989/egeol.07631192" type="journal article" year="2007">Pickford 2007</bibRefCitation>
;
<bibRefCitation id="1134DFF4FFBCA916FD74FA1FAEE72589" author="GROHE C. &amp; DE BONIS L. &amp; CHAIMANEE Y. &amp; BLONDEL C. &amp; JAEGER J. J." pageId="7" pageNumber="306" refId="ref8158" refString="GROHE C., DE BONIS L., CHAIMANEE Y., BLONDEL C. &amp; JAEGER J. J. 2013. - The oldest Asian Sivaonyx (Lutrinae, Mustelidae): A contribution to the evolutionary history of bunodont otters. Palaeontologia Electronica 16: 29 A. https://doi.org/10.26879/404" type="journal volume" year="2013">
Grohé
<emphasis id="47D17E17FFBCA916FD20FA18AD5C25A9" box="[723,775,1459,1485]" italics="true" pageId="7" pageNumber="306">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="1134DFF4FFBCA916FF3AFA7FAF9D2589" author="GHAFFAR A. &amp; AKHTAR M." box="[201,454,1491,1517]" pageId="7" pageNumber="306" pagination="347 - 356" refId="ref8104" refString="GHAFFAR A. &amp; AKHTAR M. 2016. - New material of Sivaonyx bathygnathus (Lutrinae, Mustelidae) from the early Pliocene of Siwaliks, Pakistan. Revista Brasileira de Paleontologia 19 (2): 347-356. https://doi.org/10.4072/rbp.2016.2.16" type="journal article" year="2016">Ghaffar &amp; Akhtar 2016</bibRefCitation>
). PUPC 17/64 is smaller than any known P4 of both of these species (
<tableCitation id="382797BEFFBCA916FDC7FA5FACD22669" box="[564,649,1523,1549]" captionStart="TABLE" captionStartId="10.[133,143,219,236]" captionText="TABLE 2. — Comparative measurements of Enhydriodon cf. falconeri, Sivaonyx sarwari n. sp. and Sivaonyx bathygnathus (Lydekker, 1884) (in mm). Comparative data taken from Pilgrim (1932), Colbert (1935), Chow (1961), Qi (1983), Zong (1997), Werdelin (2003), Morales &amp; Pickford (2005), Morales et al. (2005), Pickford (2007), Peigné et al. (2008), Geraads et al. (2011), Werdelin &amp; Manthi (2012), Grohé et al. (2013, 2022), Werdelin &amp; Lewis (2013), Peigné (2016), Koufos et al. (2018) and Valenciano &amp; Govender (2020). *, studied specimens; **, measurements as preserved; est., estimated measurements." pageId="7" pageNumber="306">Table 2</tableCitation>
). However, it shares more morphological similarity with the P4, GSI-D 157, of
<taxonomicName id="B2A5D986FFBCA916FF29F99FAF9D2628" authorityName="GSI K" baseAuthorityName="Lydekker" baseAuthorityYear="1884" box="[218,454,1586,1613]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="bathygnathus">
<emphasis id="47D17E17FFBCA916FF29F99FAF9D2628" box="[218,454,1586,1613]" italics="true" pageId="7" pageNumber="306">Sivaonyx bathygnathus</emphasis>
</taxonomicName>
first described and figured by
<bibRefCitation id="1134DFF4FFBCA916FF76F9FEAF442608" author="PILGRIM G. E." box="[133,287,1618,1644]" pageId="7" pageNumber="306" pagination="1 - 232" refId="ref9719" refString="PILGRIM G. E. 1932. - The fossil Carnivora of India. Memoirs of the Geological Survey of India. Palaeontologia indica NS 18: 1-232." type="journal article" year="1932">Pilgrim (1932</bibRefCitation>
: pl. 2, fig. 16) and later figured by
<bibRefCitation id="1134DFF4FFBCA916FD5EF9FEAE9E26E8" author="PICKFORD M." pageId="7" pageNumber="306" pagination="83 - 127" refId="ref9573" refString="PICKFORD M. 2007. - Revision of the Mio-Pliocene otter-like mammals of the Indian Subcontinent. Estudios Geologicos 63: 83-127. https://doi.org/10.3989/egeol.07631192" type="journal article" year="2007">Pickford (2007</bibRefCitation>
: fig. 9A). These similarities include the large paracone with a round lingual wall, low but heavy parastyle, a large and bulbous metastyle/metacone, a large and wide median fossette, and a very long posthypocrista (Fig. 2B1-B3). Based on these similarities, we are allocating PUPC 17/64 to
<taxonomicName id="B2A5D986FFBCA916FD51F95EACA52768" authorityName="Pilgrim" authorityYear="1931" box="[674,766,1778,1804]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="47D17E17FFBCA916FD51F95EACA52768" box="[674,766,1778,1804]" italics="true" pageId="7" pageNumber="306">Sivaonyx</emphasis>
</taxonomicName>
. However, due to its minute size, its rounded outline, the short metacrista, small space between protocone and hypocone, the more developed cingulum, less pinched preprotocrista or anterotransverse wall of the tooth, we do not assign it to one of the known species but designate it as a new species,
<taxonomicName id="B2A5D986FFBCA916FF75F81DAF6C27AF" authorityName="Mahmood &amp; Morlo &amp; Abbas &amp; Babar &amp; Khan" authorityYear="2025" box="[134,311,1969,1995]" class="Mammalia" family="Mustelidae" genus="Sivaonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="306" phylum="Chordata" rank="species" species="sarwari" status="sp. nov.">
<emphasis id="47D17E17FFBCA916FF75F81DAF6C27AF" box="[134,311,1969,1995]" italics="true" pageId="7" pageNumber="306">Sivaonyx sarwari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5CE2C36CFFBCA916FECFF81EAF2C27A8" box="[316,375,1970,1996]" pageId="7" pageNumber="306" rank="species">n. sp.</taxonomicNameLabel>
It differs from other
<taxonomicName id="B2A5D986FFBCA916FDA8F81DAD5C27AF" box="[603,775,1969,1995]" pageId="7" pageNumber="306">
<emphasis id="47D17E17FFBCA916FDA8F81DACEC27AF" box="[603,695,1969,1995]" italics="true" pageId="7" pageNumber="306">Sivaonyx</emphasis>
species
</taxonomicName>
as given in differential diagnosis.
</paragraph>
</subSubSection>
</treatment>
</document>