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@ -78,7 +80,7 @@ locality.
(
<figureCitation id="618DCB8EFF90FF95FE7DFDF8FE27DF6A" box="[395,463,581,602]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="2" pageNumber="583">Figs. 2</figureCitation>
,
<figureCitation id="618DCB8EFF90FF95FE2EFDF8FE0CDF6A" box="[472,484,581,602]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." pageId="2" pageNumber="583">5</figureCitation>
<figureCitation id="618DCB8EFF90FF95FE2EFDF8FE0CDF6A" box="[472,484,581,602]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="2" pageNumber="583">5</figureCitation>
7), a nearly complete crushed skull, lacking the anterior part of the rostrum. Originally preserved in a phosphatic block, it has been mechanically prepared on both sides. It is very crushed and dorsoventrally flattened from the anterior level of the palatine to its posterior end. Some sutures are hardly visible, but almost all the bones can be reconstituted. All the measurements are taken for the
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.
@ -118,7 +120,7 @@ there are:
</paragraph>
<paragraph id="F909D70BFF90FF95FCA4FCCCFAB0D93A" blockId="2.[812,1485,849,1099]" pageId="2" pageNumber="583">
OCP DEKGE 18 (
<figureCitation id="618DCB8EFF90FF95FBCEFCCCFB68DEBA" box="[1080,1152,881,906]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="2" pageNumber="583">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF90FF95FBCEFCCCFB68DEBA" box="[1080,1152,881,906]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="2" pageNumber="583">Fig. 3</figureCitation>
). An almost complete skull, with mandible. The skull is crushed, and the occipital part is strongly damaged. In the left ramus of the mandible the posterior part is missing a little behind the end of the mandibular symphysis; the right mandible is badly preserved.
</paragraph>
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@ -142,7 +144,7 @@ Fig. 2.
</paragraph>
</caption>
<paragraph id="F909D70BFF91FF94FF2FFB82FBACD963" blockId="3.[217,663,1087,1108]" box="[217,1092,1087,1108]" lastBlockId="3.[875,1092,1087,1108]" pageId="3" pageNumber="584">quadratojugal postorbital jugal orbit tooth splenial</paragraph>
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Fig. 3.
<taxonomicName id="3EB6AC88FF91FF94FF53FBD3FE08D9B2" authorityName="Jouve &amp; Iarochène &amp; Bouya &amp; Amaghzaz" authorityYear="2005" box="[165,480,1134,1154]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
@ -173,7 +175,7 @@ The supratemporal fenestrae are longer (about
), separated medially by a very narrow interfenestral bar constituted by the frontal (for about one fifth) and in a major part by the parietal (
<figureCitation id="618DCB8EFF91FF94FDC4F9CCFD7DDBBA" box="[562,661,1649,1674]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF91FF94FD54F9CCFD20DBBA" box="[674,712,1649,1674]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">4A</figureCitation>
<figureCitation id="618DCB8EFF91FF94FD54F9CCFD20DBBA" box="[674,712,1649,1674]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4A</figureCitation>
). The fenestra appears to be bordered laterally more by the postorbital than by the squamosal. The posterior margin is constituted for about the same proportion by squamosal and parietal.
</paragraph>
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@ -186,7 +188,7 @@ The infratemporal fenestra is not preserved, but can be reconstructed after the
;
<figureCitation id="618DCB8EFF91FF94FD56F8ACFCE9DA1A" box="[672,769,1809,1834]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2A</figureCitation>
,
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<figureCitation id="618DCB8EFF91FF94FF90F88CFF64DA7A" box="[102,140,1841,1866]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4A</figureCitation>
). It is anteriorly limited by the postorbital bar, constituted half by the postorbital and jugal, this latter constituting the major part of the ventral margin. The quadratojugal borders the posterior margin, constitutes the posterior part of the dorsal edge, excluding the quadrate from the dorsal margin. It participates for a small length to the ventral edge
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.
@ -196,24 +198,24 @@ The infratemporal fenestra is not preserved, but can be reconstructed after the
The suborbital fenestra is formed medially by the palatine and the pterygoid, posterolaterally by the ectopterygoid, and anterolaterally by the maxilla (
<figureCitation id="618DCB8EFF91FF94FB89FAC5FB08D8A1" box="[1151,1248,1400,1425]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2B</figureCitation>
,
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). The medial and lateral edges are curved, and the most posterior part is acute and bears the ectopterygoidpterygoid suture.
</paragraph>
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The choanae are deep, ventrally oriented (slightly caudally), and separated by a pterygoidian septum (
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,
<figureCitation id="618DCB8EFF91FF94FA6BFA43FA29DB27" box="[1437,1473,1534,1559]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">4B</figureCitation>
<figureCitation id="618DCB8EFF91FF94FA6BFA43FA29DB27" box="[1437,1473,1534,1559]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4B</figureCitation>
). They are largely surrounded by the pterygoid, only the most anterior border being formed by the palatine. The choanae are not very abruptly pierced within the pterygoids: the choanae lie within a depression on the ventral pterygoidal surface. The depression extends posterolaterally on to the lateral branch of the pterygoid, tapering before the pterygoid contacts the ectopterygoid. Thus, the choanal margin is not abrupt.
</paragraph>
<paragraph id="F909D70BFF91FF93FCA4F8B7FE27D9E7" blockId="3.[812,1486,1232,2026]" lastBlockId="4.[102,774,1182,1239]" lastPageId="4" lastPageNumber="585" pageId="3" pageNumber="584">
On the occipital face, the parietal forms the dorsomedial quarter of the posttemporal fenestra (
<figureCitation id="618DCB8EFF91FF94FB3DF896FAE7DA74" box="[1227,1295,1835,1860]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." pageId="3" pageNumber="584">Fig. 5</figureCitation>
<figureCitation id="618DCB8EFF91FF94FB3DF896FAE7DA74" box="[1227,1295,1835,1860]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="3" pageNumber="584">Fig. 5</figureCitation>
). The squamosal contributes to it dorsolaterally (less than the lateral half), and very slightly to the ventrolateral margin. The supraoccipital constitutes a small part of the ventral margin of the posttemporal fenestra (the rest formed by exoccipital), but constitutes the major part of this same margin to contact the squamosal more deeply within the posttemporal fenestra (excluding the exoccipital) (
<figureCitation id="618DCB8EFF96FF93FE8BFB03FE29D9E7" box="[381,449,1214,1239]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." pageId="4" pageNumber="585">Fig. 5</figureCitation>
<figureCitation id="618DCB8EFF96FF93FE8BFB03FE29D9E7" box="[381,449,1214,1239]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="4" pageNumber="585">Fig. 5</figureCitation>
).
</paragraph>
<caption id="ADC98783FF96FF93FF90FBE0FB70D941" box="[102,1176,1116,1137]" pageId="4" pageNumber="585" startId="4.[102,135,1117,1137]" targetBox="[171,1417,225,1090]" targetPageId="4" targetType="figure">
<caption id="ADC98783FF96FF93FF90FBE0FB70D941" ID-DOI="http://doi.org/10.5281/zenodo.13620805" ID-Zenodo-Dep="13620805" box="[102,1176,1116,1137]" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="4" pageNumber="585" startId="4.[102,135,1117,1137]" targetBox="[171,1417,225,1090]" targetPageId="4" targetType="figure">
<paragraph id="F909D70BFF96FF93FF90FBE0FB70D941" blockId="4.[102,1176,1116,1137]" box="[102,1176,1116,1137]" pageId="4" pageNumber="585">
Fig. 4.
<taxonomicName id="3EB6AC88FF96FF93FF5AFBE0FE00D941" authorityName="Jouve &amp; Iarochène &amp; Bouya &amp; Amaghzaz" authorityYear="2005" box="[172,488,1117,1137]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
@ -230,22 +232,22 @@ Fig. 4.
<paragraph id="F909D70BFF96FF93FF90FB5AFE33DBEF" blockId="4.[102,775,1255,1857]" pageId="4" pageNumber="585">
<emphasis id="CBC20B19FF96FF93FF90FB5AFF08D9CF" box="[102,224,1255,1279]" italics="true" pageId="4" pageNumber="585">Premaxilla</emphasis>
.—The premaxilla is only preserved on OCP DEKGE 18 (
<figureCitation id="618DCB8EFF96FF93FF0DFABAFED4D810" box="[251,316,1287,1312]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="4" pageNumber="585">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF96FF93FF0DFABAFED4D810" box="[251,316,1287,1312]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
). It is crushed, but its general shape can be reconstructed. Only the three first right teeth are preserved, but four teeth seem to have existed on each premaxilla. The first tooth is smaller than the two other, and a deep concavity is present between the first and the second. The second tooth is larger, but less than the third tooth which seems to be the largest (apparently confirmed by the large space between the second and third mandibular tooth). The fourth premaxillary tooth is not preserved on the skull of OCP DEKGE 18, but the space between its third and fourth mandibular teeth is short (as in OCP DEKGE 1200), implying the fourth tooth would have been the smallest. The space between the fourth and the first maxillary tooth is large, dues to the large size of the fourth dentary tooth, and the premaxillamaxilla suture is at this level on the lateral surface of the snout.
</paragraph>
<paragraph id="F909D70BFF96FF93FF7AF955FF28DA71" blockId="4.[102,775,1255,1857]" pageId="4" pageNumber="585">
The dorsal posterior process of the premaxilla is long and ends posteriorly at the level of the third maxillary tooth (
<figureCitation id="618DCB8EFF96FF93FF98F895FF5ADA71" box="[110,178,1832,1857]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="4" pageNumber="585">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF96FF93FF98F895FF5ADA71" box="[110,178,1832,1857]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF96FF93FF90F8EFFD11DADA" blockId="4.[102,775,1873,2026]" pageId="4" pageNumber="585">
<emphasis id="CBC20B19FF96FF93FF90F8EFFF53DA5A" box="[102,187,1874,1898]" italics="true" pageId="4" pageNumber="585">Maxilla</emphasis>
.—The number of teeth on each maxilla can be estimated to about 17 (
<figureCitation id="618DCB8EFF96FF93FEBFF8CCFE67DABA" box="[329,399,1905,1930]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="4" pageNumber="585">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF96FF93FEBFF8CCFE67DABA" box="[329,399,1905,1930]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
). The snout is relatively narrow, slender, and exhibits a longirostrine
<typeStatus id="260D69A9FF96FF93FE09F82FFDC6DA9A" box="[511,558,1938,1962]" pageId="4" pageNumber="585">type</typeStatus>
morphology (
<figureCitation id="618DCB8EFF96FF93FD26F82CFF7BDAFA" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[102,135,1976,1996]" captionStartId-1="3.[102,135,1134,1154]" captionStartId-2="4.[102,135,1117,1137]" captionTargetBox-1="[200,1388,233,1080]" captionTargetBox-2="[171,1417,225,1090]" captionTargetId-1="figure-433@3.[298,1372,225,1080]" captionTargetId-2="figure-520@4.[278,1288,249,1063]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." captionText-1="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." captionText-2="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="4" pageNumber="585">Figs. 24</figureCitation>
<figureCitation id="618DCB8EFF96FF93FD26F82CFF7BDAFA" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[102,135,1976,1996]" captionStartId-1="3.[102,135,1134,1154]" captionStartId-2="4.[102,135,1117,1137]" captionTargetBox-1="[200,1388,233,1080]" captionTargetBox-2="[171,1417,225,1090]" captionTargetId-1="figure-433@3.[298,1372,225,1080]" captionTargetId-2="figure-520@4.[278,1288,249,1063]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." captionText-1="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." captionText-2="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi-1="http://doi.org/10.5281/zenodo.13620803" figureDoi-2="http://doi.org/10.5281/zenodo.13620805" httpUri-1="https://zenodo.org/record/13620803/files/figure.png" httpUri-2="https://zenodo.org/record/13620805/files/figure.png" pageId="4" pageNumber="585">Figs. 24</figureCitation>
). The maxilla is relatively sculptured laterally compared to other skull area, with longitudinal deep ridges and furrows.
</paragraph>
<paragraph id="F909D70BFF96FF93FCA4FB23FA62D807" blockId="4.[812,1485,1182,1756]" pageId="4" pageNumber="585">
@ -256,7 +258,7 @@ The alveoli are well developed, circular, with their base in relief and very mar
<paragraph id="F909D70BFF96FF93FCACFAFDFB38DB09" blockId="4.[812,1485,1182,1756]" pageId="4" pageNumber="585">The maxilla is laterally straight, and the tooth row does not display a festooned outline in dorsal view. Occlusal pits are present from the space between the twelfth and thirteenth right maxillary alveoli, and between the thirteenth and fourteenth left alveoli. These pits, lined with the alveoli tooth row, increase in depth posteriorly. Anteriorly, the space between the right and left alveoli is large (twice the diameter of the alveoli) and increases posteriorly.</paragraph>
<paragraph id="F909D70BFF96FF93FCA4F9FEFB52DBEC" blockId="4.[812,1485,1182,1756]" pageId="4" pageNumber="585">
The contact with the palatines is expected between the level of the eleventh and twelfth maxillary alveoli (OCP DEKGE 300;
<figureCitation id="618DCB8EFF96FF93FC23F93EFBC5DBAC" box="[981,1069,1667,1692]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="4" pageNumber="585">Fig. 4B</figureCitation>
<figureCitation id="618DCB8EFF96FF93FC23F93EFBC5DBAC" box="[981,1069,1667,1692]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="4" pageNumber="585">Fig. 4B</figureCitation>
). The maxilla contacts the ectopterygoid medially, and ends ventrally on the jugal. Dorsally, the maxillae are separated by the nasal.
</paragraph>
<paragraph id="F909D70BFF96FF92FCDAF94FFD13DF89" blockId="4.[812,1485,1777,2026]" lastBlockId="5.[102,774,640,697]" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
@ -264,12 +266,12 @@ The contact with the palatines is expected between the level of the eleventh and
.—The nasal is a single bone (the two nasals are fused), ornamented with only discrete and sparse furrows. It is narrow between the maxillae, and its anterior process penetrates deeply between the posterior premaxillae processes, but terminates
<quantity id="3E4E7AEEFF96FF93FC9DF8CFFC49DAB9" box="[875,929,1906,1929]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="585" unit="cm" value="7.0">7 cm</quantity>
(in OCP DEKGE 18) posterior to the external nares, which it does not reach (
<figureCitation id="618DCB8EFF96FF93FBADF82CFB73DA9A" box="[1115,1179,1937,1962]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="4" pageNumber="585">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF96FF93FBADF82CFB73DA9A" box="[1115,1179,1937,1962]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
). The nasal is narrow anteriorly between the maxillae, with a constant width from the posterior contact with the premaxillae to the anterior contact with the lacrimals. Then, it widens posteriorly and finally sends a long posterior process between frontal and prefrontal (
<figureCitation id="618DCB8EFF97FF92FD5AFD1DFD06DF89" box="[684,750,672,697]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Fig. 4</figureCitation>
<figureCitation id="618DCB8EFF97FF92FD5AFD1DFD06DF89" box="[684,750,672,697]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">Fig. 4</figureCitation>
).
</paragraph>
<caption id="ADC98783FF97FF92FF90FD99FC8CDF64" pageId="5" pageNumber="586" startId="5.[102,135,548,568]" targetBox="[108,1485,235,515]" targetPageId="5" targetType="figure">
<caption id="ADC98783FF97FF92FF90FD99FC8CDF64" ID-DOI="http://doi.org/10.5281/zenodo.13620807" ID-Zenodo-Dep="13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="5" pageNumber="586" startId="5.[102,135,548,568]" targetBox="[108,1485,235,515]" targetPageId="5" targetType="figure">
<paragraph id="F909D70BFF97FF92FF90FD99FC8CDF64" blockId="5.[102,1485,548,596]" pageId="5" pageNumber="586">
Fig. 5.
<taxonomicName id="3EB6AC88FF97FF92FF51FD99FE0BDF08" authorityName="Jouve &amp; Iarochène &amp; Bouya &amp; Amaghzaz" authorityYear="2005" box="[167,483,548,568]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="586" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
@ -290,13 +292,13 @@ Fig. 5.
), almost complete (the most anterior part is absent). It is short, narrow and longer than wide (
<figureCitation id="618DCB8EFF97FF92FD38FCB1FF62DE75" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FF62FC91FF53DE75" box="[148,187,812,837]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FF62FC91FF53DE75" box="[148,187,812,837]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
). Dorsally, the contact with the frontal is as long as its contact with the nasal.
</paragraph>
<paragraph id="F909D70BFF97FF92FF90FCC5FDE7D920" blockId="5.[102,775,887,1040]" pageId="5" pageNumber="586">
<emphasis id="CBC20B19FF97FF92FF90FCC5FF25DEA0" box="[102,205,888,912]" italics="true" pageId="5" pageNumber="586">Lacrimal</emphasis>
.—The right lacrimal of specimen OCP DEKGE 18 is well preserved (
<figureCitation id="618DCB8EFF97FF92FEB9FC2AFE79DE80" box="[335,401,919,944]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="5" pageNumber="586">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF97FF92FEB9FC2AFE79DE80" box="[335,401,919,944]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="5" pageNumber="586">Fig. 3</figureCitation>
), and is well anteriorly expanded, reaching the level of the tenth maxillary tooth. Its ornamentation is very light, with some shallow pits. It is large, and forms the anterior margin of the orbit.
</paragraph>
<paragraph id="F909D70BFF97FF92FF90FB9EFEB5D8AC" blockId="5.[102,775,1059,1501]" pageId="5" pageNumber="586">
@ -304,13 +306,13 @@ Fig. 5.
.—It extends anteriorly within the nasal as far as the prefrontal, and its width, at the orbital level appears moderate (
<figureCitation id="618DCB8EFF97FF92FF98FBDEFF39D94C" box="[110,209,1123,1148]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FF2AFBDEFEEAD94C" box="[220,258,1123,1148]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FF2AFBDEFEEAD94C" box="[220,258,1123,1148]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
). Its lateral extension is short, and contacts the postorbital to constitute the postorbital bar. Posteromedially, at the angle between the postorbital bar and the interfenestral bar, the frontal forms a very light dorsal overhang within the supratemporal fenestra. Ventrally, and below this overhang, the frontal contacts an extremely laterally elongated laterosphenoid in a flat lateroventral extension. Posteromedially, the frontal takes part in the narrow interfenestral bar (
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width), since its total proportion is about onefifth of the total length (
<figureCitation id="618DCB8EFF97FF92FF4DFA3EFEF7D8AC" box="[187,287,1411,1436]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FEDCFA3EFEB9D8AC" box="[298,337,1411,1436]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FEDCFA3EFEB9D8AC" box="[298,337,1411,1436]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF97FF92FF7AFA19FDC4D8ED" blockId="5.[102,775,1059,1501]" pageId="5" pageNumber="586">The frontal is smooth, without ornamentation, with only three medial grooves between the orbits.</paragraph>
@ -321,7 +323,7 @@ width), since its total proportion is about onefifth of the total length (
anteriorly, is only 0.5 cm in width posterior to the suture (OCP DEKGE 300;
<figureCitation id="618DCB8EFF97FF92FEE1F9CDFE96DBB9" box="[279,382,1648,1673]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FE78F9CDFE5CDBB9" box="[398,436,1648,1673]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FE78F9CDFE5CDBB9" box="[398,436,1648,1673]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
). Anteroventrally, its contact with the laterosphenoid is visible, with a long anterior process between the frontal and the laterosphenoid, reducing the contact between these two bones.
</paragraph>
<paragraph id="F909D70BFF97FF92FF7AF94CFAD8D909" blockId="5.[102,775,1520,2026]" lastBlockId="5.[812,1485,640,1305]" pageId="5" pageNumber="586">
@ -330,20 +332,20 @@ Ventrally, the parietallaterosphenoid suture appears parallel to the skull ro
medial to the squamosalparietal suture, and appears to end rapidly in the interfenestral bar. The posterior wall of the supratemporal fenestra is very inclined posteriorly, and is largely visible in dorsal view (
<figureCitation id="618DCB8EFF97FF92FA63FD7DFCB9DFC9" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FCAAFD5DFC6BDFC9" box="[860,899,736,761]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FCAAFD5DFC6BDFC9" box="[860,899,736,761]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
). The parietalquadrate suture, visible in dorsal view on the posterior wall of the supratemporal fenestra, continues in the same direction as the parietallaterosphenoid suture. It extends dorsally to join the squamosalquadrate suture at the same level (in the right supratemporal fenestra) or slightly below the temporal canal (in the left supratemporal fenestra), about
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laterally to the temporal canal. The ventral squamosalparietal suture joins the temporal canal in its most lateral part, continues dorsally to this one (crosses the temporal canal medially, in its onethird dorsomedial part), and continues in a vertical direction (with zigzag).
</paragraph>
<paragraph id="F909D70BFF97FF92FCA4FBFDFA9ED829" blockId="5.[812,1485,640,1305]" pageId="5" pageNumber="586">
The parietal is not ornamented on either the interfenestral bar or on the posterior skull roof. It contributes to half of the posterior wall of the supratemporal fenestra, and tapers posteromedially with a strong and acute process in the occipital face between the occipital tuberosities. The dorsal surface of this process is posteroventrally inclined, and the supraoccipital forms its ventrooccipital part (
<figureCitation id="618DCB8EFF97FF92FAD5FABDFA80D829" box="[1315,1384,1280,1305]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="5" pageNumber="586">Fig. 6</figureCitation>
<figureCitation id="618DCB8EFF97FF92FAD5FABDFA80D829" box="[1315,1384,1280,1305]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="5" pageNumber="586">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF97FF92FCDAFA94FBF7D8D2" blockId="5.[812,1486,1321,2026]" pageId="5" pageNumber="586">
<emphasis id="CBC20B19FF97FF92FCDAFA94FC42D871" box="[812,938,1321,1345]" italics="true" pageId="5" pageNumber="586">Postorbital</emphasis>
.—It is badly preserved on OCP DEKGE 18 (
<figureCitation id="618DCB8EFF97FF92FCC3FAF4FC9FD852" box="[821,887,1353,1378]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="5" pageNumber="586">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF97FF92FCC3FAF4FC9FD852" box="[821,887,1353,1378]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="5" pageNumber="586">Fig. 3</figureCitation>
), and extremely fractured in OCP DEKGE 300 (
<figureCitation id="618DCB8EFF97FF92FA56FAF4FCBAD8B2" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Fig. 2A</figureCitation>
). The postorbital bar, comprised of the postorbital and the jugal, seems to be gently concave medially. It was probably inclined ventrolaterally, not ornamented, mediolaterally flat and longer than wide.
@ -352,11 +354,11 @@ The parietal is not ornamented on either the interfenestral bar or on the poster
The postorbital is the most important part of the lateral arcade of the supratemporal fenestra (
<figureCitation id="618DCB8EFF97FF92FB33F9B4FAC2DB12" box="[1221,1322,1545,1570]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FAC1F9B4FAB4DB12" box="[1335,1372,1545,1570]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FAC1F9B4FAB4DB12" box="[1335,1372,1545,1570]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
, 7). It appears dorsally and laterally ornamented with spaced pits, contacts the squamosal posteriorly, and the quadratojugal posteroventrally. It seems to participate largely to the dorsal margin of the infratemporal fenestra (Fig. 7). Anteriorly, it bears a robust lateral process, directed anteroventrally, and ornamented with deep furrows, which seems to contact the ventral margin of the orbit (jugal) (
<figureCitation id="618DCB8EFF97FF92FB42F954FAEADA32" box="[1204,1282,1769,1794]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="5" pageNumber="586">Figs. 3</figureCitation>
<figureCitation id="618DCB8EFF97FF92FB42F954FAEADA32" box="[1204,1282,1769,1794]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="5" pageNumber="586">Figs. 3</figureCitation>
,
<figureCitation id="618DCB8EFF97FF92FAF9F954FADEDA32" box="[1295,1334,1769,1794]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">4A</figureCitation>
<figureCitation id="618DCB8EFF97FF92FAF9F954FADEDA32" box="[1295,1334,1769,1794]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF97FF91FCDAF8AFFDC1D9D2" blockId="5.[812,1486,1321,2026]" lastBlockId="6.[102,775,1033,2026]" lastPageId="6" lastPageNumber="587" pageId="5" pageNumber="586">
@ -365,7 +367,7 @@ The postorbital is the most important part of the lateral arcade of the supratem
<figureCitation id="618DCB8EFF97FF92FC04F8CCFBA3DABA" box="[1010,1099,1905,1930]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Fig. 2A</figureCitation>
). Weakly high in the posterior wall of the supratemporal fenestra, it constitutes the major part of the dorsal border of the temporal canal. It contacts the postorbital anterolaterally, but is separated from the quadratojugal by the quadrate. It takes part in the dorsal part of the external ear, and sends an important squamosal wing roofing the external otic aperture (Fig. 7). Posterior to the ear, the squamosal constitutes a long and high blade sinking deeply beneath the skull roof and forms the anterior wall of the paroccipital process. The paroccipital process extends more ventrally than the level of the skull roof.
</paragraph>
<caption id="ADC98783FF94FF91FF90FCC0FD28DEF9" pageId="6" pageNumber="587" startId="6.[102,136,893,913]" targetBox="[102,774,231,864]" targetPageId="6" targetType="figure">
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<paragraph id="F909D70BFF94FF91FF90FCC0FD28DEF9" blockId="6.[102,775,893,969]" pageId="6" pageNumber="587">
Fig. 6.
<taxonomicName id="3EB6AC88FF94FF91FF5CFCC0FE1DDEA1" authorityName="Jouve &amp; Iarochène &amp; Bouya &amp; Amaghzaz" authorityYear="2005" box="[170,501,893,913]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
@ -381,7 +383,7 @@ Fig. 6.
</caption>
<paragraph id="F909D70BFF94FF91FF7AFB57FDB0D892" blockId="6.[102,775,1033,2026]" pageId="6" pageNumber="587">
In occipital view, the squamosal contributes dorsolaterally to the posttemporal fenestra (less than the lateral half), and to a small lateral part of the occipital tuberosity. It contributes to a small part to the occipital face, and more posteriorly, it participates in the dorsal edge of the paroccipital process, ending before the extremity one of the process (
<figureCitation id="618DCB8EFF94FF91FDF1FA34FDA3D892" box="[519,587,1417,1442]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">Fig. 6</figureCitation>
<figureCitation id="618DCB8EFF94FF91FDF1FA34FDA3D892" box="[519,587,1417,1442]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF94FF91FF90FA0FFEA9DA7A" blockId="6.[102,775,1033,2026]" pageId="6" pageNumber="587">
@ -389,16 +391,16 @@ In occipital view, the squamosal contributes dorsolaterally to the posttemporal
.—Very fragmentary (
<figureCitation id="618DCB8EFF94FF91FE5CFA0CFDE1D8FA" box="[426,521,1457,1482]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Fig. 2A</figureCitation>
), the jugal forms the lateroventral edge of the orbit, and the ventral part of the postorbital pillar (
<figureCitation id="618DCB8EFF94FF91FEC1FA4CFE7DDB3A" box="[311,405,1521,1546]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Fig. 4A</figureCitation>
<figureCitation id="618DCB8EFF94FF91FEC1FA4CFE7DDB3A" box="[311,405,1521,1546]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">Fig. 4A</figureCitation>
). There is no lateral jugal edge raised bordering this pillar, and the postorbital pillar is not laterally in continuity with the lateral edge of the jugal. The postorbital bar is displaced medially and is not laterally in alignment with the lateral jugal edge. The base of the postorbital pillar is completely pierced by a foramen, anteroposteriorly directed. Apparently, the posterior aperture of this foramen is situated on the external face of the postorbital pillar. Posteriorly, the internal part of the jugal is exposed and exhibits a deep and long groove, including two distinct foramina (
<figureCitation id="618DCB8EFF94FF91FF2CF88CFEDCDA7A" box="[218,308,1841,1866]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF94FF91FF7AF8ECFE0FDADA" blockId="6.[102,775,1033,2026]" pageId="6" pageNumber="587">
The jugal is laterally ornamented with spaced deep pits. Posteriorly, it is high, slightly convex dorsally, lateromedially narrow and ends just before the quadratojugal lateral notch (see below). Anteriorly, it reaches the level of the anterior process of the prefrontal (
<figureCitation id="618DCB8EFF94FF91FEADF86CFE40DADA" box="[347,424,2001,2026]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">Figs. 3</figureCitation>
<figureCitation id="618DCB8EFF94FF91FEADF86CFE40DADA" box="[347,424,2001,2026]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="6" pageNumber="587">Figs. 3</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FE43F86CFE33DADA" box="[437,475,2001,2026]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">4A</figureCitation>
<figureCitation id="618DCB8EFF94FF91FE43F86CFE33DADA" box="[437,475,2001,2026]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4A</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF94FF91FCDAFF55FC95DCD1" blockId="6.[812,1485,232,611]" pageId="6" pageNumber="587">
@ -415,45 +417,45 @@ The jugal is laterally ornamented with spaced deep pits. Posteriorly, it is high
.—It is a small bone, “V” shaped in posterior view, which contributes to the posteromedial occipital process with the parietal (
<figureCitation id="618DCB8EFF94FF91FBDCFD0BFB66DFFF" box="[1066,1166,694,719]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FB6DFD0BFB28DFFF" box="[1179,1216,694,719]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">4A</figureCitation>
<figureCitation id="618DCB8EFF94FF91FB6DFD0BFB28DFFF" box="[1179,1216,694,719]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4A</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FB3AFD0BFB34DFFF" box="[1228,1244,694,719]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">6</figureCitation>
<figureCitation id="618DCB8EFF94FF91FB3AFD0BFB34DFFF" box="[1228,1244,694,719]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">6</figureCitation>
). It forms the medioventral mid part of the posttemporal fenestra, posteriorly on the occipital tuberosity, but contacts the squamosal one centimeter anteriorly, within the posttemporal fenestra (
<figureCitation id="618DCB8EFF94FF91FA8CFCABFA57DE1F" box="[1402,1471,790,815]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">Fig. 5</figureCitation>
<figureCitation id="618DCB8EFF94FF91FA8CFCABFA57DE1F" box="[1402,1471,790,815]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="6" pageNumber="587">Fig. 5</figureCitation>
). It does not seem to contribute to the occipital tuberosities.
</paragraph>
<paragraph id="F909D70BFF94FF91FCDAFCDFFBBBD9EB" blockId="6.[812,1485,866,1534]" pageId="6" pageNumber="587">
<emphasis id="CBC20B19FF94FF91FCDAFCDFFC43DE4A" box="[812,939,866,890]" italics="true" pageId="6" pageNumber="587">Exoccipital</emphasis>
.—They form the main part of the occipital face, contributing laterally to the occipital condyle (each exoccipital constitutes to one third of the half width), and almost completely surround the foramen magnum (threequarter) (
<figureCitation id="618DCB8EFF94FF91FC93FC5FFC44DECB" box="[869,940,994,1019]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">Fig. 6</figureCitation>
<figureCitation id="618DCB8EFF94FF91FC93FC5FFC44DECB" box="[869,940,994,1019]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
). Dorsally, they form the occipital tuberosities, which are well developed, dorsoventrally flattened, and posteriorly directed under the posttemporal fenestra. Laterally, they constitute the main part of the robust paroccipital process, and surround dorsally, medially and ventrally to the cranioquadrate canal. The paroccipital process is flat, composed of the exoccipital and squamosal, and its posterior extremity is quadrangular.
</paragraph>
<paragraph id="F909D70BFF94FF91FCA4FB59FAF1D84D" blockId="6.[812,1485,866,1534]" pageId="6" pageNumber="587">
Ventrally, the exoccipitals contribute posterolaterally, onethird to each basioccipital tuberosity in a very broad ventral process (
<figureCitation id="618DCB8EFF94FF91FC3EFA99FBD9D80D" box="[968,1073,1316,1341]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2B</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FBB4FA99FB8DD80D" box="[1090,1125,1316,1341]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">4B</figureCitation>
<figureCitation id="618DCB8EFF94FF91FBB4FA99FB8DD80D" box="[1090,1125,1316,1341]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4B</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FB81FA99FB73D80D" box="[1143,1179,1316,1341]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">6B</figureCitation>
<figureCitation id="618DCB8EFF94FF91FB81FA99FB73D80D" box="[1143,1179,1316,1341]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">6B</figureCitation>
). The suture between the basioccipital and the exoccipital, posteriorly to the basioccipital tuberosities, lay deeply in a cavity.
</paragraph>
<paragraph id="F909D70BFF94FF91FCA4FA38FB37D8CE" blockId="6.[812,1485,866,1534]" pageId="6" pageNumber="587">
The foramen for nerve XII is small and laterally directed on the exoccipital. The vagus foramen and posterior carotid foramen are situated anterior to the foramen for nerve XII, and directed ventroposteriorly (
<figureCitation id="618DCB8EFF94FF91FB7BFA58FB39D8CE" box="[1165,1233,1509,1534]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">Fig. 6</figureCitation>
<figureCitation id="618DCB8EFF94FF91FB7BFA58FB39D8CE" box="[1165,1233,1509,1534]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF94FF91FCDAF9AFFB8EDADA" blockId="6.[812,1485,1553,2026]" pageId="6" pageNumber="587">
<emphasis id="CBC20B19FF94FF91FCDAF9AFFC28DB1A" box="[812,960,1554,1578]" italics="true" pageId="6" pageNumber="587">Basioccipital</emphasis>
.—It constitutes the main part of the occipital condyle, which is very wide. Its anterior part (basioccipital tuberosities) is not strongly projected ventrally and therefore does not proceed ventrally to the occipital condyle in occipital view (
<figureCitation id="618DCB8EFF94FF91FC60F92CFC32DB9A" box="[918,986,1681,1706]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." pageId="6" pageNumber="587">Fig. 6</figureCitation>
<figureCitation id="618DCB8EFF94FF91FC60F92CFC32DB9A" box="[918,986,1681,1706]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
). The area between the basioccipital tuberosities and the occipital condyle is gently curved dorsally, almost horizontally oriented. In ventral view, the posterior margin of the basioccipital tuberosities is concave anteriorly, relatively thin in its medial part (just behind the medial eustachian foramen), with a small medial ridge posteriorly to the medial eustachian foramen (
<figureCitation id="618DCB8EFF94FF91FB9CF8ECFB24DA5A" box="[1130,1228,1873,1898]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2B</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FB2EF8ECFB14DA5A" box="[1240,1276,1873,1898]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">4B</figureCitation>
<figureCitation id="618DCB8EFF94FF91FB2EF8ECFB14DA5A" box="[1240,1276,1873,1898]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4B</figureCitation>
). The anteroposterior thinness of the medial part of the basioccipital, posterior to the medial eustachian foramen, separates the two basioccipital tuberosities, each one having a “drop” shape in ventral view (
<figureCitation id="618DCB8EFF94FF91FC31F86CFBC1DADA" box="[967,1065,2001,2026]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2B</figureCitation>
,
<figureCitation id="618DCB8EFF94FF91FBC3F86CFBB1DADA" box="[1077,1113,2001,2026]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">4B</figureCitation>
<figureCitation id="618DCB8EFF94FF91FBC3F86CFBB1DADA" box="[1077,1113,2001,2026]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4B</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF95FF90FF90FDDDFF07DEF1" blockId="7.[102,241,608,628]" lastBlockId="7.[112,239,941,961]" pageId="7" pageNumber="588">laterosphenoid parietal frontal postorbital postorbital bar jugal ectopterygoid</paragraph>
@ -462,7 +464,7 @@ The foramen for nerve XII is small and laterally directed on the exoccipital. Th
.—It seems to enclose completely the medial eustachian foramen, but the limit between basioccipital and basisphenoid cannot be seen. The bone is narrow laterally to the medial eustachian foramen, with a lateral “pinching” (
<figureCitation id="618DCB8EFF95FF90FF98FB26FF38D984" box="[110,208,1179,1204]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Figs. 2B</figureCitation>
,
<figureCitation id="618DCB8EFF95FF90FF2AFB26FEE8D984" box="[220,256,1179,1204]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">4B</figureCitation>
<figureCitation id="618DCB8EFF95FF90FF2AFB26FEE8D984" box="[220,256,1179,1204]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">4B</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF95FF90FF90FB77FD15D8F3" blockId="7.[102,775,1226,1475]" pageId="7" pageNumber="588">
@ -470,7 +472,7 @@ The foramen for nerve XII is small and laterally directed on the exoccipital. Th
.—It is long, posteroventrally directed, and forms the jaw joint with the quadratojugal (
<figureCitation id="618DCB8EFF95FF90FDE8FB57FD6ED833" box="[542,646,1258,1283]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF95FF90FD61FB57FD40D833" box="[663,680,1258,1283]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">4</figureCitation>
<figureCitation id="618DCB8EFF95FF90FD61FB57FD40D833" box="[663,680,1258,1283]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">4</figureCitation>
). In the supratemporal fenestra, it contacts the squamosal lateral to the temporal canal. Anterolaterally, it slips in between the squamosal and the quadratojugal by a thin anterior process that contacts the postorbital. Anteriorly, on its ventral face, the quadrate bears a small crest at its mid width, which probably corresponds to the “crest B” of
<bibRefCitation id="9D27AAFAFF95FF90FE42FA17FD82D8F3" author="Iordansky, N. N." box="[436,618,1450,1475]" pageId="7" pageNumber="588" pagination="256 - 280" refId="ref10559" refString="Iordansky, N. N. 1964. The jaw muscles of the crocodiles and some relating structures of the crocodilian skull. Anatomischer Anzeiger 115: 256 - 280." type="journal article" year="1964">Iordansky (1964</bibRefCitation>
,
@ -484,7 +486,7 @@ The foramen for nerve XII is small and laterally directed on the exoccipital. Th
.—The palatines are very crushed (
<figureCitation id="618DCB8EFF95FF90FDADFA65FD5ED8C1" box="[603,694,1496,1521]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
). They seem enlarged and curved laterally before contacting the maxillae anteriorly. This contact, dues to the bad preservation, is not available. Its posterior border seems flattened, enlarged posteriorly and deviates laterally before joining the pterygoid (lateral border of the palatine not parallel, but laterally curved anteriorly and posteriorly, medial border of suborbital fenestra medially curved). The contact with the pterygoid is not clearly visible, but it seems to form only the anterior part of the choanae (
<figureCitation id="618DCB8EFF95FF90FE5EF942FDE9DA28" box="[424,513,1791,1816]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 4B</figureCitation>
<figureCitation id="618DCB8EFF95FF90FE5EF942FDE9DA28" box="[424,513,1791,1816]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF95FF90FF90F893FBFBDEFF" blockId="7.[102,774,1838,2026]" lastBlockId="7.[895,1486,834,991]" pageId="7" pageNumber="588">
@ -492,7 +494,7 @@ The foramen for nerve XII is small and laterally directed on the exoccipital. Th
.—The left ectopterygoid is well preserved (
<figureCitation id="618DCB8EFF95FF90FF98F8F3FF2DDA57" box="[110,197,1870,1895]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
). It is wide and twisted between its contacts with the jugal and the pterygoid. It curves gently anteriorly to form the posterolateral part of the suborbital fenestra (
<figureCitation id="618DCB8EFF95FF90FD68F82DFD11DA99" box="[670,761,1936,1961]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 4B</figureCitation>
<figureCitation id="618DCB8EFF95FF90FD68F82DFD11DA99" box="[670,761,1936,1961]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
). The posterior process is wide, decreasing posteriorly, and covers the pterygoid almost as far as its posterior extremity occipital tuberosities exoccipital squamosal quadrate quadratojugal Fig. 7.
<taxonomicName id="3EB6AC88FF95FF90FB67FCFFFA25DE66" authorityName="Jouve &amp; Iarochène &amp; Bouya &amp; Amaghzaz" authorityYear="2005" box="[1169,1485,834,854]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
<emphasis id="CBC20B19FF95FF90FB67FCFFFA25DE66" box="[1169,1485,834,854]" italics="true" pageId="7" pageNumber="588">Arambourgisuchus khouribgaensis</emphasis>
@ -519,12 +521,12 @@ tory drawing of the same (
.—The pterygoids are damaged (
<figureCitation id="618DCB8EFF95FF90FAF1FBDDFAB5D949" box="[1287,1373,1120,1145]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
), but their shape can be trace out. Anteriorly, they almost completely surround the choanae. A septum, formed by the pterygoids separates the choanae in two openings (
<figureCitation id="618DCB8EFF95FF90FB1EFB7DFAA8D9E9" box="[1256,1344,1216,1241]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 4B</figureCitation>
<figureCitation id="618DCB8EFF95FF90FB1EFB7DFAA8D9E9" box="[1256,1344,1216,1241]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF95FF90FCA4FB5CFACBDB0A" blockId="7.[812,1485,1048,2026]" pageId="7" pageNumber="588">
The pterygoids diverge posterolateraly to form two lateral wings in contact with the ectopterygoid. In front and anterolaterally to the choanae, they seem flattened and enlarged laterally; they are anteroposteriorly narrow between the choanae and their contact with the ectopterygoids (
<figureCitation id="618DCB8EFF95FF90FA56FADCFCB8D8AA" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 4B</figureCitation>
<figureCitation id="618DCB8EFF95FF90FA56FADCFCB8D8AA" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
). Contact with the ectopterygoid is small, and increases rapidly lateroposteriorly with the increase of the anteroposterior length of the pterygoid wing. The lateral part of this wing is extremely thickened dorsoventrally to form a strong
<emphasis id="CBC20B19FF95FF90FCDAF9BFFC0CDB29" box="[812,996,1537,1561]" italics="true" pageId="7" pageNumber="588">torus transiliens</emphasis>
, with a very important anterior thickening, decreasing progressively posteriorly (Fig. 7).
@ -534,7 +536,7 @@ The pterygoids diverge posterolateraly to form two lateral wings in contact with
.—Anteriorly, it is laterally expanded ventrally to the frontal, and its anterior margin is lateromedially directed (
<figureCitation id="618DCB8EFF95FF90FC60F934FC12DB92" box="[918,1018,1673,1698]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Figs. 2A</figureCitation>
,
<figureCitation id="618DCB8EFF95FF90FBF1F934FBC5DB92" box="[1031,1069,1673,1698]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">4A</figureCitation>
<figureCitation id="618DCB8EFF95FF90FBF1F934FBC5DB92" box="[1031,1069,1673,1698]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">4A</figureCitation>
). The posterior part is dorsoventrally crushed, and its shape is hardly distinguished (
<figureCitation id="618DCB8EFF95FF90FADBF914FA99DBF2" box="[1325,1393,1705,1730]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2</figureCitation>
). Suture with the frontal and parietal (in continuity) seems horizontal, parallel to the skull roof (dorsal limit of the interfenestral bar).
@ -542,11 +544,11 @@ The pterygoids diverge posterolateraly to form two lateral wings in contact with
<paragraph id="F909D70BFF95FF9FFCDAF8AFFD91DC91" blockId="7.[812,1485,1048,2026]" lastBlockId="8.[102,775,232,613]" lastPageId="8" lastPageNumber="589" pageId="7" pageNumber="588">
<emphasis id="CBC20B19FF95FF90FCDAF8AFFC7FDA1A" box="[812,919,1810,1834]" italics="true" pageId="7" pageNumber="588">Mandible</emphasis>
.—Two isolated mandibular fragments have been found, one anterior, preserved from the first to the twelfth tooth (OCP DEKGE 1200;
<figureCitation id="618DCB8EFF95FF90FB82F8ECFB20DA5A" box="[1140,1224,1873,1898]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." pageId="7" pageNumber="588">Fig 8A</figureCitation>
<figureCitation id="618DCB8EFF95FF90FB82F8ECFB20DA5A" box="[1140,1224,1873,1898]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="7" pageNumber="588">Fig 8A</figureCitation>
), and a more posterior part, including the eleventh to the sixteenth teeth (OCP DEKGE 269;
<figureCitation id="618DCB8EFF95FF90FC20F82CFBA7DA9A" box="[982,1103,1937,1962]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." pageId="7" pageNumber="588">Fig. 8B, C</figureCitation>
<figureCitation id="618DCB8EFF95FF90FC20F82CFBA7DA9A" box="[982,1103,1937,1962]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="7" pageNumber="588">Fig. 8B, C</figureCitation>
). Unfortunately, only a small part of its posterior branch is preserved on the specimen OCP DEKGE 300, but too poorly preserved to be interpreted. The first tooth is relatively robust (
<figureCitation id="618DCB8EFF9AFF9FFE4DFF55FDFDDC31" box="[443,533,232,257]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." pageId="8" pageNumber="589">Fig. 8A</figureCitation>
<figureCitation id="618DCB8EFF9AFF9FFE4DFF55FDFDDC31" box="[443,533,232,257]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="8" pageNumber="589">Fig. 8A</figureCitation>
), the second alveolus smaller, and the space between it and the third one is greater than between the other. The third tooth is about the same size as the second, and it is near the very enlarged fourth. This one has its alveolar edge higher dorsally. The next alveoli seem relatively regular in their spacing and diameter.
</paragraph>
<caption id="ADC98783FF9AFF9FFCDAFF57FBBEDC09" pageId="8" pageNumber="589" startId="8.[812,867,234,255]" targetType="table">
@ -623,16 +625,16 @@ The pterygoids diverge posterolateraly to form two lateral wings in contact with
</paragraph>
<paragraph id="F909D70BFF9AFF9FFF7AFE17FDB4DF33" blockId="8.[102,775,232,613]" pageId="8" pageNumber="589">
The mandible is wider than high (
<figureCitation id="618DCB8EFF9AFF9FFDE7FE17FD89DCF3" box="[529,609,426,451]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="8" pageNumber="589">Figs. 3</figureCitation>
<figureCitation id="618DCB8EFF9AFF9FFDE7FE17FD89DCF3" box="[529,609,426,451]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="8" pageNumber="589">Figs. 3</figureCitation>
,
<figureCitation id="618DCB8EFF9AFF9FFD86FE17FD5EDCF3" box="[624,694,426,451]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." pageId="8" pageNumber="589">8B, C</figureCitation>
<figureCitation id="618DCB8EFF9AFF9FFD86FE17FD5EDCF3" box="[624,694,426,451]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="8" pageNumber="589">8B, C</figureCitation>
), lacks festooned outlines, and there are occlusal pits posterior to the fifteenth teeth (OCP DEKGE 269;
<figureCitation id="618DCB8EFF9AFF9FFE0EFE57FDA6DF33" box="[504,590,490,515]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." pageId="8" pageNumber="589">Fig. 8B</figureCitation>
<figureCitation id="618DCB8EFF9AFF9FFE0EFE57FDA6DF33" box="[504,590,490,515]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEKGE 1200 in ventral view. B. OCP DEKGE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="8" pageNumber="589">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF9AFF9FFF7AFDB1FE55DF55" blockId="8.[102,775,232,613]" pageId="8" pageNumber="589">
The symphysis ends posteriorly at the level of the sixteenth tooth, and the splenial ends anteriorly between the tenth and the eleventh (
<figureCitation id="618DCB8EFF9AFF9FFE9DFDF1FE58DF55" box="[363,432,588,613]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." pageId="8" pageNumber="589">Fig. 3</figureCitation>
<figureCitation id="618DCB8EFF9AFF9FFE9DFDF1FE58DF55" box="[363,432,588,613]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEKGE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="8" pageNumber="589">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="F909D70BFF9AFF9FFF90FDC4FEFDDE21" blockId="8.[102,775,632,785]" pageId="8" pageNumber="589">