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<mods:title id="AE09B045CBBFB1D6B4FCB901FBD71E88">Three new species of the long-tailed wasp genus † Megacoxa Brazidec et al., 2024 (Hymenoptera: Megalyridae) from Kachin amber</mods:title>
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<mods:namePart id="8894DCCD082ED45BB2C36DDC68FDC7FD">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="999C0A144D6E4F362BF5E9D989291C56">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark.</mods:affiliation>
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<mods:namePart id="D7F6D212837729611CA1AF85D5FE441B">Wu, Qiong</mods:namePart>
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<mods:affiliation id="912F55278DD0979D0228006328733A5A">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark. & College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:nameIdentifier id="4CB4625CDB341545696E3E9E64EECE72" type="email">narsleslie@163.com</mods:nameIdentifier>
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<mods:namePart id="4394DB908EA6657E303E818F2FA89495">Wang, Zhen</mods:namePart>
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<mods:affiliation id="EC97C54D27D3C40FE834AB13822B79C5">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark. & College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:namePart id="5CC76E56E7ADCB91E2389E7126F848DB">Gao, Tai-Ping</mods:namePart>
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<mods:affiliation id="AF9F3B2B5D31F00BBB37D9E91E05893B">College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:namePart id="1969C0043F7760E28E2ED2A29FE9631D">Ren, Dong</mods:namePart>
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<mods:date id="1EACD8DD244623D05841AA73AD034187">2025</mods:date>
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†
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<taxonomicName id="72DB4D4DFFD4AD6CFDBCC070FBFBA552" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[536,1067,1294,1321]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="248" phylum="Arthropoda" rank="species" species="synchrotron">
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<emphasis id="87AFEADCFFD4AD6CFDBCC070FCE0A552" bold="true" box="[536,816,1294,1321]" italics="true" pageId="10" pageNumber="248">Megacoxa synchrotron</emphasis>
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<bibRefCitation id="D14A4B3FFFD4AD6CFC93C070FBFBA552" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[823,1067,1295,1321]" pageId="10" pageNumber="248" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
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Brazidec
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<emphasis id="87AFEADCFFD4AD6CFC02C06FFC31A552" box="[934,993,1295,1321]" italics="true" pageId="10" pageNumber="248">et al.</emphasis>
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, 2024
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</taxonomicName>
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</paragraph>
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<figureCitation id="2DE02A4BFFD4AD6CFD53C04DFCECA536" box="[759,828,1330,1357]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="10" pageNumber="248">Fig. 5</figureCitation>
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<emphasis id="87AFEADCFFD4AD6CFF19C00CFEE6A5F6" bold="true" box="[189,310,1395,1421]" pageId="10" pageNumber="248">Diagnosis</emphasis>
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</heading>
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<paragraph id="B56436CEFFD4AD6CFF19C0DDFE26A5C7" blockId="10.[189,502,1442,1468]" box="[189,502,1442,1468]" pageId="10" pageNumber="248">
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See
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<bibRefCitation id="D14A4B3FFFD4AD6CFF4BC0DDFE23A5C7" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[239,499,1442,1468]" pageId="10" pageNumber="248" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
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Brazidec
|
||||
<emphasis id="87AFEADCFFD4AD6CFEFAC0DCFE49A5C7" box="[350,409,1442,1468]" italics="true" pageId="10" pageNumber="248">et al.</emphasis>
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(2024)
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</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="FDC16545FFD4AD6CFF19C09CFCD9A609" pageId="10" pageNumber="248" type="materials_examined">
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<emphasis id="87AFEADCFFD4AD6CFF19C09CFE7AA586" bold="true" box="[189,426,1507,1533]" pageId="10" pageNumber="248">Material examined</emphasis>
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<materialsCitation id="05B33C93FFD4AD6CFF19C36DFCD9A609" country="Myanmar" latitude="26.333334" location="Hukawng Valley" longLatPrecision="1242" longitude="96.6" municipality="Myitkyina District" pageId="10" pageNumber="248" specimenCode="CNU-HYM-MA-2016208" specimenCount="1" specimenCount-female="1" stateProvince="Kachin State" typeStatus="holotype">
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•
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;
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<collectingRegion id="771FF82CFFD4AD6CFE3DC36DFDE5A657" box="[409,565,1554,1580]" country="Myanmar" name="Kachin" pageId="10" pageNumber="248">Kachin State</collectingRegion>
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,
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,
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; [
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,
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<geologicalTimeScale id="C7FBB665FFD4AD6CFAAFC36DFBA9A62B" pageId="10" pageNumber="248">unnamed horizon, mid-Cretaceous, upper Albian to lower Cenomanian, 98.79± 0.62 Ma</geologicalTimeScale>
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(
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<bibRefCitation id="D14A4B3FFFD4AD6CFB29C34AFF2BA608" author="Cruickshank R. D. & Ko K." pageId="10" pageNumber="248" pagination="441 - 455" refId="ref7759" refString="Cruickshank R. D. & Ko K. 2003. Geology of an amber locality in the Hukawng Valley, northern Myanmar. Journal of Asian Earth Sciences 21: 441-455. https://doi.org/10.1016/S1367-9120(02)00044-5" type="journal article" year="2003">Cruickshank & Ko 2003</bibRefCitation>
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;
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<bibRefCitation id="D14A4B3FFFD4AD6CFEA3C327FE68A608" author="Shi G. H. & Grimaldi D. A. & Harlow G. E. & Wang J. & Wang J. & Yang M. C. & Lei W. Y. & Li Q. L. & Li X. H." box="[263,440,1624,1651]" pageId="10" pageNumber="248" pagination="155 - 163" refId="ref8697" refString="Shi G. H., Grimaldi D. A., Harlow G. E., Wang J., Wang J., Yang M. C., Lei W. Y., Li Q. L. & Li X. H. 2012. Age constraint on Burmese amber based on U-Pb dating of zircons. Cretaceous Research 37: 155-163. https://doi.org/10.1016/j.cretres.2012.03.014" type="journal article" year="2012">
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Shi
|
||||
<emphasis id="87AFEADCFFD4AD6CFE92C326FEB9A609" box="[310,361,1624,1650]" italics="true" pageId="10" pageNumber="248">et al</emphasis>
|
||||
. 2012
|
||||
</bibRefCitation>
|
||||
);
|
||||
<specimenCode id="E57D9EB5FFD4AD6CFE6FC327FCD6A609" box="[459,774,1624,1650]" pageId="10" pageNumber="248">CNU-HYM-MA-2016208</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD4AD6AFF19C3E5FE33A50A" lastPageId="12" lastPageNumber="250" pageId="10" pageNumber="248" type="description">
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C3E5FE9EA6CF" blockId="10.[189,334,1690,1716]" box="[189,334,1690,1716]" pageId="10" pageNumber="248">
|
||||
<heading id="EE2C81A2FFD4AD6CFF19C3E5FE9EA6CF" bold="true" box="[189,334,1690,1716]" fontSize="11" level="3" pageId="10" pageNumber="248" reason="3">
|
||||
<emphasis id="87AFEADCFFD4AD6CFF19C3E5FE9EA6CF" bold="true" box="[189,334,1690,1716]" pageId="10" pageNumber="248">Description</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6CFF4EC3B6FE95A698" blockId="10.[188,1399,1737,1869]" box="[234,325,1737,1763]" pageId="10" pageNumber="248">
|
||||
<emphasis id="87AFEADCFFD4AD6CFF4EC3B6FE95A698" bold="true" box="[234,325,1737,1763]" pageId="10" pageNumber="248">Female</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C393FA80A737" blockId="10.[188,1399,1737,1869]" pageId="10" pageNumber="248">
|
||||
BODY. Length (except ovipositor)
|
||||
<quantity id="72239B2BFFD4AD6CFDEAC394FD7CA77D" box="[590,684,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="10" pageNumber="248" unit="mm" value="5.4">5.4 mm</quantity>
|
||||
, fore wing length
|
||||
<quantity id="72239B2BFFD4AD6CFC26C393FC30A77D" box="[898,992,1772,1798]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="248" unit="mm" value="3.4">3.4 mm</quantity>
|
||||
. Specimen in slightly gritty amber piece together with small beetle (Coleoptera) and midge (Diptera); wasp well preserved except for right foreleg cut off at base of tibia and ovipositor (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFD71C24DFCE1A736" box="[725,817,1842,1869]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="10" pageNumber="248">Fig. 5A</figureCitation>
|
||||
) cut off approx.
|
||||
<quantity id="72239B2BFFD4AD6CFC51C24DFB9FA736" box="[1013,1103,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="248" unit="mm" value="0.5">0.5 mm</quantity>
|
||||
from tip of abdomen.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6AFF19C20BFECCA1A8" blockId="10.[189,1400,1907,2005]" lastBlockId="12.[189,1399,265,467]" lastPageId="12" lastPageNumber="250" pageId="10" pageNumber="248">
|
||||
HEAD. Wider than long, head capsule without prominent sculpture or setation; eyes oval (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFAACC20CFAB7A7F5" box="[1288,1383,1907,1934]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="10" pageNumber="248">Fig. 5B</figureCitation>
|
||||
), higher than long, without postocular carina; antennal insertions in depressions, frons overhangs them above; occipital carina well developed (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFD0BC2C5FCC3A7AE" box="[687,787,1978,2005]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="10" pageNumber="248">Fig. 5A</figureCitation>
|
||||
), crenulate, area around occipital foramen deeply concave; clypeus short and wide, triangular with tip between antennal insertions; antenna (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFAB4C476FABDA15F" box="[1296,1389,265,292]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5B</figureCitation>
|
||||
) slightly shorter than head + mesosoma combined, with short globular scapus and short flat pedicellus, antennomeres 3–14 flattened (probably an artefact), antennomeres 3–6 longer than 7–13, antennomere 14 approx. twice as long as wide, strongly tapered apically; labrum a small narrow flap below clypeus, concealed when mandibles are closed; mandibles broad, with three teeth (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFB91C4E9FB41A1CA" box="[1077,1169,406,433]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5B</figureCitation>
|
||||
) each, apical tooth longest.
|
||||
</paragraph>
|
||||
<caption id="E1A46646FFD5AD6DFF19C274FC29A7AD" ID-DOI="http://doi.org/10.5281/zenodo.14981663" ID-Zenodo-Dep="14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="11" pageNumber="249" startId="11.[189,232,1803,1829]" targetBox="[189,1398,266,1777]" targetPageId="11">
|
||||
<paragraph id="B56436CEFFD5AD6DFF19C274FC29A7AD" blockId="11.[187,1400,1803,2006]" pageId="11" pageNumber="249">
|
||||
<emphasis id="87AFEADCFFD5AD6DFF19C274FED9A75E" bold="true" box="[189,265,1803,1829]" pageId="11" pageNumber="249">Fig. 5.</emphasis>
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD5AD6DFEBBC273FCF5A75E" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[287,805,1803,1830]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="249" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFD5AD6DFEBBC273FDFBA75E" box="[287,555,1803,1829]" italics="true" pageId="11" pageNumber="249">Megacoxa synchrotron</emphasis>
|
||||
<bibRefCitation id="D14A4B3FFFD5AD6DFD94C274FCF5A75E" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[560,805,1803,1830]" pageId="11" pageNumber="249" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD5AD6DFD04C273FD0AA75E" box="[672,730,1803,1829]" italics="true" pageId="11" pageNumber="249">et al.</emphasis>
|
||||
, 2024
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, ♀ (CNU-HYM-MA-2016208).
|
||||
<emphasis id="87AFEADCFFD5AD6DFB07C274FB68A75E" bold="true" box="[1187,1208,1803,1829]" pageId="11" pageNumber="249">A</emphasis>
|
||||
. Habitus, dorsal view.
|
||||
<emphasis id="87AFEADCFFD5AD6DFEA0C250FEC8A732" bold="true" box="[260,280,1839,1865]" pageId="11" pageNumber="249">B</emphasis>
|
||||
. Head and mesosoma, ventral view.
|
||||
<emphasis id="87AFEADCFFD5AD6DFD68C251FD30A733" bold="true" box="[716,736,1838,1864]" pageId="11" pageNumber="249">C</emphasis>
|
||||
. Hind leg, ventral view.
|
||||
<emphasis id="87AFEADCFFD5AD6DFBA3C250FBCBA732" bold="true" box="[1031,1051,1839,1865]" pageId="11" pageNumber="249">D</emphasis>
|
||||
. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C77DFC4FA3E4" blockId="12.[186,1401,514,928]" pageId="12" pageNumber="250">
|
||||
MESOSOMA. With elongate, extended propectus (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFCA8C77DFC43A266" box="[780,915,514,541]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5A–B</figureCitation>
|
||||
); pronotum narrow medially, triangular laterally; fore femur spindle-shaped, fore tibia slender, with elongate, curved calcar; mesoscutum strongly arched in lateral view, angled between vertical anterior surface and horizontal surface; horizontal surface flat, level with mesoscutellum, not prominently sculptured; median mesoscutal sulcus (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFB09C713FAD8A2FD" box="[1197,1288,619,646]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5A</figureCitation>
|
||||
) distinct, crenulate, extends to transscutal articulation; axillae distinct, triangular, abut medially, separated from mesoscutellum by oblique foveate scutoscutellar sulcus; mesoscutellum flat, smooth, pointed anteriorly, rounded posteriorly; mesosoma difficult to observe laterally because of position of legs and curvature of amber piece; mid femur spindle-shaped, mid tibia slender; hind leg much larger than fore and mid legs, hind femur strongly swollen, hind tibia (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFD49C664FC9AA34D" box="[749,842,795,822]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5C</figureCitation>
|
||||
) strongly expanded distally, widest at approx. two-thirds from proximal end, with distinct fringe of setae distally on median ventral margin, two apical tibial spurs inserted just distally of fringe, inner spur longer than outer; hind basitarsomere approx. equal in length to remaining tarsomeres; tarsal claws slender, simple.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C6B1FE82A429" blockId="12.[189,1400,974,1106]" pageId="12" pageNumber="250">
|
||||
WINGS. Fore wing (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFE0FC6B1FDDAA392" box="[427,522,974,1001]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5D</figureCitation>
|
||||
) hyaline with veins C, R1, Sc+R, M+Cu, A, Rs+M, Rs, M and Cu fully pigmented; Rs+M aligned with M+Cu; medial cell wide distally; Rs present between Rs+M and r-rs, closing first submarginal cell; marginal cell narrow, closed in straight line by Rs; hind wing could not be observed.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C1FEFAA9A57F" blockId="12.[189,1401,1153,1285]" pageId="12" pageNumber="250">
|
||||
METASOMA. 1.68 × as long as mesosoma, fusiform, with weak sculpture and setation in ventral view; sternum 2 tapering anteriorly, broadening posteriorly to broadest point of metasoma, approx. twice as long as sternum 3; sternum 7 strongly tapered apically, folded over medially along longitudinal axis; base of ovipositor (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFE3AC195FE28A57E" box="[414,504,1258,1285]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5A</figureCitation>
|
||||
) extends straight from dorsal part of metasomal tip and beyond tips of wings.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD2AD6AFF4EC04CFEFAA536" blockId="12.[189,483,1331,1393]" box="[234,298,1331,1357]" pageId="12" pageNumber="250">
|
||||
<emphasis id="87AFEADCFFD2AD6AFF4EC04CFEFAA536" bold="true" box="[234,298,1331,1357]" pageId="12" pageNumber="250">Male</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C028FE33A50A" blockId="12.[189,483,1331,1393]" box="[189,483,1366,1393]" pageId="12" pageNumber="250">
|
||||
See
|
||||
<bibRefCitation id="D14A4B3FFFD2AD6AFF4BC029FE0EA50A" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[239,478,1366,1393]" pageId="12" pageNumber="250" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD2AD6AFEFAC028FE49A50B" box="[350,409,1366,1392]" italics="true" pageId="12" pageNumber="250">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD2AD6AFF19C0DFFC56A75E" pageId="12" pageNumber="250" type="discussion">
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C0DFFEFFA5C1" blockId="12.[189,303,1440,1466]" box="[189,303,1440,1466]" pageId="12" pageNumber="250">
|
||||
<heading id="EE2C81A2FFD2AD6AFF19C0DFFEFFA5C1" bold="true" box="[189,303,1440,1466]" fontSize="11" level="3" pageId="12" pageNumber="250" reason="3">
|
||||
<emphasis id="87AFEADCFFD2AD6AFF19C0DFFEFFA5C1" bold="true" box="[189,303,1440,1466]" pageId="12" pageNumber="250">Remarks</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C0B1FC56A75E" blockId="12.[188,1399,1486,1829]" pageId="12" pageNumber="250">
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD2AD6AFF68C0B0FE0DA593" box="[204,477,1486,1512]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="250" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFD2AD6AFF68C0B0FE0DA593" box="[204,477,1486,1512]" italics="true" pageId="12" pageNumber="250">Megacoxa synchrotron</emphasis>
|
||||
</taxonomicName>
|
||||
was described from a male and the
|
||||
<typeStatus id="6A60886CFFD2AD6AFC39C0B1FBD4A593" box="[925,1028,1486,1512]" pageId="12" pageNumber="250" type="holotype">holotype</typeStatus>
|
||||
was deposited in Phyletisches Museum Jena,
|
||||
<collectingCountry id="CDCC765EFFD2AD6AFED4C08EFE0DA670" box="[368,477,1521,1547]" name="Germany" pageId="12" pageNumber="250">Germany</collectingCountry>
|
||||
(CASENT0753237;
|
||||
<bibRefCitation id="D14A4B3FFFD2AD6AFD7DC08EFC1DA670" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[729,973,1521,1547]" pageId="12" pageNumber="250" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD2AD6AFCEFC08DFC56A670" box="[843,902,1521,1547]" italics="true" pageId="12" pageNumber="250">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: figs 2f, 8). The female we describe here resembles the
|
||||
<typeStatus id="6A60886CFFD2AD6AFE07C36BFDDAA655" box="[419,522,1556,1582]" pageId="12" pageNumber="250" type="holotype">holotype</typeStatus>
|
||||
in a number of features: the antennal configuration, the crenulate median mesoscutal sulcus, the medially abutting axillae, the uniquely enlarged hind tibia (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFB04C347FB2FA629" box="[1184,1279,1591,1618]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5C</figureCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD2AD6AFAAAC347FE94A60E" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." pageId="12" pageNumber="250" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD2AD6AFF19C323FF2BA60E" box="[189,251,1627,1653]" italics="true" pageId="12" pageNumber="250">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 8a). The wing venation is virtually identical in the
|
||||
<typeStatus id="6A60886CFFD2AD6AFC57C324FB8AA60E" box="[1011,1114,1627,1653]" pageId="12" pageNumber="250" type="holotype">holotype</typeStatus>
|
||||
and the present female (compare
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFE96C300FE5AA6E2" box="[306,394,1662,1689]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="12" pageNumber="250">Fig. 5D</figureCitation>
|
||||
with fig. 2f in
|
||||
<bibRefCitation id="D14A4B3FFFD2AD6AFD95C301FCCFA6E3" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[561,799,1662,1688]" pageId="12" pageNumber="250" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD2AD6AFD04C300FD09A6E3" box="[672,729,1662,1688]" italics="true" pageId="12" pageNumber="250">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
). The main difference between the two specimens is in their size, the female being almost twice as long as the
|
||||
<typeStatus id="6A60886CFFD2AD6AFCF1C3DEFC10A6C0" box="[853,960,1697,1723]" pageId="12" pageNumber="250" type="holotype">holotype</typeStatus>
|
||||
. However, such sexual dimorphism is not unusual in wood-living wasps (L. Vilhelmsen, pers. obs.). In most of the phylogenetic analyses (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFAE2C3BBFF19A779" captionStart="Fig" captionStartId="3.[189,232,1311,1337]" captionTargetBox="[189,1397,269,1294]" captionTargetId="figure-187@3.[189,1398,265,1299]" captionTargetPageId="3" captionText="Fig. 1. Phylogeny of Megalyridae. Strict consensus tree of six trees obtained by implied weights analyses with k = 5. All genera except †Megacoxa represented by only one terminal, number in brackets indicates total number of recognized species." figureDoi="http://doi.org/10.5281/zenodo.14981653" httpUri="https://zenodo.org/record/14981653/files/figure.png" pageId="12" pageNumber="250">Fig. 1</figureCitation>
|
||||
), the female specimen CNU-HYM-MA-2016208 comes out in a polytomy with the three previously described species of †
|
||||
<taxonomicName id="72DB4D4DFFD2AD6AFE1BC273FDE9A75E" box="[447,569,1804,1829]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="250" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="87AFEADCFFD2AD6AFE1BC273FDE9A75E" box="[447,569,1804,1829]" italics="true" pageId="12" pageNumber="250">Megacoxa</emphasis>
|
||||
</taxonomicName>
|
||||
, including †
|
||||
<taxonomicName id="72DB4D4DFFD2AD6AFD69C273FC50A75E" box="[717,896,1803,1829]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="250" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFD2AD6AFD69C273FD35A75E" box="[717,741,1804,1829]" italics="true" pageId="12" pageNumber="250">M</emphasis>
|
||||
.
|
||||
<emphasis id="87AFEADCFFD2AD6AFD50C274FC50A75E" box="[756,896,1803,1829]" italics="true" pageId="12" pageNumber="250">synchrotron</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
305
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305
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<mods:namePart id="5B5CAFDE195AD8C84F21CA3B625CA2DF">Wang, Zhen</mods:namePart>
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<mods:namePart id="C35B18B28FBB8A1FA20FA4A9048280C9">Gao, Tai-Ping</mods:namePart>
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†
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFDDCC6CBFCB9A3B5" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[632,873,947,974]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD6AD6EFDDCC6CBFCB9A3B5" bold="true" box="[632,873,947,974]" italics="true" pageId="8" pageNumber="246">Megacoxa mjoelner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD6AD6EFCD4C6CBFC1AA3B5" box="[880,970,948,974]" pageId="8" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD6AD6EFED2C6A8FCECA46F" pageId="8" pageNumber="246" type="description">
|
||||
<paragraph id="B56436CEFFD6AD6EFED2C6A8FB6EA389" blockId="8.[374,1214,947,1044]" box="[374,1214,983,1010]" pageId="8" pageNumber="246">
|
||||
<uri id="C14A3ACCFFD6AD6EFED2C6A8FB6EA389" box="[374,1214,983,1010]" pageId="8" pageNumber="246">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="C17D0C1BFFD6AD6EFD32C6A8FB6EA389" box="[662,1214,983,1010]" pageId="8" pageNumber="246">215840BF-F89F-4A1B-98F1-A738E89B3299</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFD53C685FCECA46F" blockId="8.[374,1214,947,1044]" box="[759,828,1018,1044]" pageId="8" pageNumber="246">
|
||||
<figureCitation id="2DE02A4BFFD6AD6EFD53C685FCECA46F" box="[759,828,1018,1044]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="8" pageNumber="246">Fig. 4</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD6AD6EFF19C144FD83A56A" pageId="8" pageNumber="246" type="diagnosis">
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C144FEE6A42E" blockId="8.[189,310,1083,1109]" box="[189,310,1083,1109]" pageId="8" pageNumber="246">
|
||||
<heading id="EE2C81A2FFD6AD6EFF19C144FEE6A42E" bold="true" box="[189,310,1083,1109]" fontSize="11" level="3" pageId="8" pageNumber="246" reason="3">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF19C144FEE6A42E" bold="true" box="[189,310,1083,1109]" pageId="8" pageNumber="246">Diagnosis</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C115FD83A56A" blockId="8.[188,1399,1130,1297]" pageId="8" pageNumber="246">
|
||||
Body comparatively slender (
|
||||
<figureCitation id="2DE02A4BFFD6AD6EFDBCC115FDA7A4FF" box="[536,631,1130,1156]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="8" pageNumber="246">Fig. 4A</figureCitation>
|
||||
), metasoma longer than mesosoma (shorter in †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFB0EC114FAA7A4FF" box="[1194,1399,1130,1156]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
||||
<emphasis id="87AFEADCFFD6AD6EFB0EC114FAA7A4FF" box="[1194,1399,1130,1156]" italics="true" pageId="8" pageNumber="246">M. chandrahrasa</emphasis>
|
||||
</taxonomicName>
|
||||
and †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFF5AC1F1FE57A4DC" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[254,391,1166,1191]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="gungner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF5AC1F1FE57A4DC" box="[254,391,1166,1191]" italics="true" pageId="8" pageNumber="246">M. gungner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD6AD6EFE2AC1F1FE3DA4D3" box="[398,493,1166,1192]" pageId="8" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
). Antennae shorter than head + mesosoma (longer in †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFBCBC1F1FAD3A4DC" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[1135,1283,1165,1191]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="brazideci" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD6AD6EFBCBC1F1FAD3A4DC" box="[1135,1283,1165,1191]" italics="true" pageId="8" pageNumber="246">M. brazideci</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD6AD6EFAAFC1F1FABAA4D3" box="[1291,1386,1166,1192]" pageId="8" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
). Median mesoscutal sulcus smooth (crenulate in †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFD58C1CEFC7EA4B0" box="[764,942,1201,1227]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFD6AD6EFD58C1CEFC7EA4B0" box="[764,942,1201,1227]" italics="true" pageId="8" pageNumber="246">M. synchrotron</emphasis>
|
||||
</taxonomicName>
|
||||
), axillae not contiguous (contiguous in †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFF68C1AAFE8FA495" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[204,351,1236,1262]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="brazideci">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF68C1AAFE8FA495" box="[204,351,1236,1262]" italics="true" pageId="8" pageNumber="246">M. brazideci</emphasis>
|
||||
</taxonomicName>
|
||||
). Hind coxa without carina or foveae along posterior margin (coxa carinate and foveate in †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFF68C187FE98A56A" box="[204,328,1272,1297]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="janzeni">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF68C187FE98A56A" box="[204,328,1272,1297]" italics="true" pageId="8" pageNumber="246">M. janzeni</emphasis>
|
||||
</taxonomicName>
|
||||
and †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFE2BC187FD92A56A" box="[399,578,1271,1297]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFD6AD6EFE2BC187FD92A56A" box="[399,578,1271,1297]" italics="true" pageId="8" pageNumber="246">M. synchrotron</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD6AD6EFF19C046FB72A675" pageId="8" pageNumber="246" type="etymology">
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C046FE94A528" blockId="8.[189,324,1337,1363]" box="[189,324,1337,1363]" pageId="8" pageNumber="246">
|
||||
<heading id="EE2C81A2FFD6AD6EFF19C046FE94A528" bold="true" box="[189,324,1337,1363]" fontSize="11" level="3" pageId="8" pageNumber="246" reason="3">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF19C046FE94A528" bold="true" box="[189,324,1337,1363]" pageId="8" pageNumber="246">Etymology</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C018FB72A675" blockId="8.[188,1399,1383,1550]" pageId="8" pageNumber="246">
|
||||
The new species takes its name after Mjølner (or Mjolnir), the magic hammer that was the chief weapon and primary attribute of Thor, the thunder god in Norse mythology. Like the ovipositor in †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFB43C0F4FAA6A5DF" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[1255,1398,1418,1444]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD6AD6EFB43C0F4FAA6A5DF" box="[1255,1398,1418,1444]" italics="true" pageId="8" pageNumber="246">M. mjoelner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD6AD6EFF19C0D1FEC6A5B3" box="[189,278,1454,1480]" pageId="8" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(see below), the shaft of the hammer was shorter than it was originally supposed to be, due to the interference of the trickster god Loke in the form of a fly; he viciously stung the dwarven smith Brokk as Mjølner was being finalized, thus confirming the timeless malevolence of Diptera.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD6AD6EFF19C34AFCD9A69C" pageId="8" pageNumber="246" type="materials_examined">
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C34AFEBCA634" blockId="8.[189,364,1589,1615]" box="[189,364,1589,1615]" pageId="8" pageNumber="246">
|
||||
<heading id="EE2C81A2FFD6AD6EFF19C34AFEBCA634" bold="true" box="[189,364,1589,1615]" fontSize="11" level="3" pageId="8" pageNumber="246" reason="3">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF19C34AFEBCA634" bold="true" box="[189,364,1589,1615]" pageId="8" pageNumber="246">Type material</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF4EC31BFE8CA605" blockId="8.[189,1399,1636,1768]" box="[234,348,1636,1662]" pageId="8" pageNumber="246">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF4EC31BFE8CA605" bold="true" box="[234,348,1636,1662]" pageId="8" pageNumber="246">
|
||||
<typeStatus id="6A60886CFFD6AD6EFF4EC31BFE8CA605" box="[234,348,1636,1662]" pageId="8" pageNumber="246" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C3F8FCD9A69C" blockId="8.[189,1399,1636,1768]" pageId="8" pageNumber="246">
|
||||
<materialsCitation id="05B33C93FFD6AD6EFF19C3F8FCD9A69C" country="Myanmar" latitude="26.333334" location="Hukawng Valley" longLatPrecision="1242" longitude="96.6" municipality="Myitkyina District" pageId="8" pageNumber="246" specimenCode="CNU-HYM-MA-2016209" specimenCount="1" specimenCount-female="1" stateProvince="Kachin State" typeStatus="holotype">
|
||||
<collectingCountry id="CDCC765EFFD6AD6EFF19C3F8FE86A6DA" box="[189,342,1671,1697]" name="Myanmar" pageId="8" pageNumber="246">MYANMAR</collectingCountry>
|
||||
•
|
||||
<specimenCount id="A3DDFD47FFD6AD6EFED6C3F7FE59A6D9" box="[370,393,1672,1698]" count="1" pageId="8" pageNumber="246" type="female">♀</specimenCount>
|
||||
;
|
||||
<collectingRegion id="771FF82CFFD6AD6EFE3DC3F8FDE5A6DA" box="[409,565,1671,1697]" country="Myanmar" name="Kachin" pageId="8" pageNumber="246">Kachin State</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="5500ACB4FFD6AD6EFDE6C3F8FCF0A6D9" box="[578,800,1671,1698]" pageId="8" pageNumber="246">Myitkyina District</collectingMunicipality>
|
||||
,
|
||||
<location id="B0046015FFD6AD6EFC89C3F8FC27A6DA" LSID="urn:lsid:plazi:treatment:3D7287D8FFD6AD6CFDCDC6CBFC0EA4AA:B0046015FFD6AD6EFC89C3F8FC27A6DA" box="[813,1015,1671,1697]" country="Myanmar" latitude="26.333334" longLatPrecision="1242" longitude="96.6" municipality="Myitkyina District" name="Hukawng Valley" pageId="8" pageNumber="246" stateProvince="Kachin State">Hukawng Valley</location>
|
||||
; [
|
||||
<geoCoordinate id="D0EF5009FFD6AD6EFBAFC3FAFBABA6D9" box="[1035,1147,1669,1698]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="8" pageNumber="246" precision="925" value="26.333334">26°20′ N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D0EF5009FFD6AD6EFB2CC3FAFB24A6D9" box="[1160,1268,1669,1698]" degrees="96" direction="east" minutes="36" orientation="longitude" pageId="8" pageNumber="246" precision="925" value="96.6">96°36′ E</geoCoordinate>
|
||||
];
|
||||
<geologicalTimeScale id="C7FBB665FFD6AD6EFAAFC3F8FBA9A6BE" pageId="8" pageNumber="246">unnamed horizon, mid-Cretaceous, upper Albian to lower Cenomanian, 98.79± 0.62 Ma</geologicalTimeScale>
|
||||
(
|
||||
<bibRefCitation id="D14A4B3FFFD6AD6EFB29C3D5FF2BA693" author="Cruickshank R. D. & Ko K." pageId="8" pageNumber="246" pagination="441 - 455" refId="ref7759" refString="Cruickshank R. D. & Ko K. 2003. Geology of an amber locality in the Hukawng Valley, northern Myanmar. Journal of Asian Earth Sciences 21: 441-455. https://doi.org/10.1016/S1367-9120(02)00044-5" type="journal article" year="2003">Cruickshank & Ko 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD6AD6EFEA3C3B2FE68A693" author="Shi G. H. & Grimaldi D. A. & Harlow G. E. & Wang J. & Wang J. & Yang M. C. & Lei W. Y. & Li Q. L. & Li X. H." box="[263,440,1741,1768]" pageId="8" pageNumber="246" pagination="155 - 163" refId="ref8697" refString="Shi G. H., Grimaldi D. A., Harlow G. E., Wang J., Wang J., Yang M. C., Lei W. Y., Li Q. L. & Li X. H. 2012. Age constraint on Burmese amber based on U-Pb dating of zircons. Cretaceous Research 37: 155-163. https://doi.org/10.1016/j.cretres.2012.03.014" type="journal article" year="2012">
|
||||
Shi
|
||||
<emphasis id="87AFEADCFFD6AD6EFE92C3B1FEB9A69C" box="[310,361,1741,1767]" italics="true" pageId="8" pageNumber="246">et al</emphasis>
|
||||
. 2012
|
||||
</bibRefCitation>
|
||||
);
|
||||
<specimenCode id="E57D9EB5FFD6AD6EFE6FC3B2FCD5A69C" box="[459,773,1741,1767]" pageId="8" pageNumber="246">CNU-HYM-MA-2016209</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD6AD6CFF19C270FEE9A445" lastPageId="10" lastPageNumber="248" pageId="8" pageNumber="246" type="description">
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C270FE9EA752" blockId="8.[189,334,1807,1833]" box="[189,334,1807,1833]" pageId="8" pageNumber="246">
|
||||
<heading id="EE2C81A2FFD6AD6EFF19C270FE9EA752" bold="true" box="[189,334,1807,1833]" fontSize="11" level="3" pageId="8" pageNumber="246" reason="3">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF19C270FE9EA752" bold="true" box="[189,334,1807,1833]" pageId="8" pageNumber="246">Description</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF4EC241FDF5A79E" blockId="8.[189,1400,1854,2021]" pageId="8" pageNumber="246">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF4EC241FE95A723" bold="true" box="[234,325,1854,1880]" pageId="8" pageNumber="246">Female</emphasis>
|
||||
BODY. Length (except ovipositor)
|
||||
<quantity id="72239B2BFFD6AD6EFDC1C21EFD18A700" box="[613,712,1889,1915]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="246" unit="mm" value="4.3">4.3 mm</quantity>
|
||||
, fore wing length
|
||||
<quantity id="72239B2BFFD6AD6EFC12C21EFBC9A700" box="[950,1049,1889,1915]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="246" unit="mm" value="3.3">3.3 mm</quantity>
|
||||
, ovipositor length
|
||||
<quantity id="72239B2BFFD6AD6EFAA2C21EFA81A700" box="[1286,1361,1889,1915]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="246" unit="mm" value="1.0">1 mm</quantity>
|
||||
as preserved (
|
||||
<figureCitation id="2DE02A4BFFD6AD6EFEE5C2FBFE72A7E5" box="[321,418,1924,1950]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="8" pageNumber="246">Fig. 4A</figureCitation>
|
||||
), probably broken (see below). Specimen in mostly clear amber piece, with small midge (Diptera) syninclusion. Exudate covers lower part of head and a few diffraction planes extend from dorsal part of mesosoma.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD7AD6FFF19C476FB57A18D" blockId="9.[189,1399,265,503]" pageId="9" pageNumber="247">
|
||||
HEAD (
|
||||
<figureCitation id="2DE02A4BFFD7AD6FFEB5C476FEBFA158" box="[273,367,265,291]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="9" pageNumber="247">Fig. 4D</figureCitation>
|
||||
). Globular, slightly wider than long, without prominent sculpture or setation; eyes oval, cover most of lateral surface of head; ocelli situated on top of head in approx. equilateral triangle; postocular carina absent; occipital carina well developed (
|
||||
<figureCitation id="2DE02A4BFFD7AD6FFC29C430FC20A112" box="[909,1008,335,361]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="9" pageNumber="247">Fig. 4D</figureCitation>
|
||||
), smooth, area around occipital foramen deeply concave; antenna clearly shorter than head + mesosoma combined, scapus curved, pedicellus short, antennomeres 3–13 cylindrical (
|
||||
<figureCitation id="2DE02A4BFFD7AD6FFCB1C4E9FCA7A1CB" box="[789,887,406,432]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="9" pageNumber="247">Fig. 4D</figureCitation>
|
||||
), slightly longer than wide, decreasing in length distally, antennomere 14 approx. twice as long as wide, tapering distally; mouthparts difficult to observe, mandible apparently broad, teeth not visible; short palps can be observed.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD7AD6CFF19C769FD42A2F8" blockId="9.[189,1398,534,595]" lastBlockId="10.[188,1400,265,643]" lastPageId="10" lastPageNumber="248" pageId="9" pageNumber="247">
|
||||
MESOSOMA. With elongate, extended propectus; pronotum narrow medially, triangular laterally, with anterior thoracic spiracle in notch in lateral margin (
|
||||
<figureCitation id="2DE02A4BFFD7AD6FFC9BC746FC71A228" box="[831,929,569,595]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="9" pageNumber="247">Fig. 4C</figureCitation>
|
||||
); fore femur spindle-shaped, fore tibia slightly expanded distally (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFDAFC476FDBEA158" box="[523,622,265,291]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4D</figureCitation>
|
||||
), with elongate, curved calcar; mesoscutum strongly arched in lateral view, angled between vertical anterior surface and horizontal surface; horizontal surface flat, level with mesoscutellum, not prominently sculptured; median mesoscutal sulcus distinct (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFB4DC430FAB9A112" box="[1257,1385,335,361]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4B–C</figureCitation>
|
||||
), smooth, extends to transscutal articulation; axillae distinct, triangular, not abutting medially, separated from mesoscutellum by oblique foveate scutoscutellar sulcus; mesoscutellum flat, smooth with small punctures, diamond-shaped; mesopleuron smooth, with depression medially, mesopleuro-metapleural sulcus distinct, crenulate (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFE55C4A3FD9FA18D" box="[497,591,476,502]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4C</figureCitation>
|
||||
); mid femur spindle-shaped, mid tibia slender, with two short apical spurs; hind coxa broad, with carina along posterior margin but without foveae; hind femur spindle-shaped; hind tibia (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFE3EC75CFE25A246" box="[410,501,547,573]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4A</figureCitation>
|
||||
) flattened, expanded distally, without fringe of setae, with two apical spurs of subequal length, inner spur shorter than distal tibial width; hind basitarsus slightly longer than remaining tarsomeres, tarsal claws simple, slender.
|
||||
</paragraph>
|
||||
<caption id="E1A46646FFD7AD6FFF19C385FD7AA793" ID-DOI="http://doi.org/10.5281/zenodo.14981661" ID-Zenodo-Dep="14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="9" pageNumber="247" startId="9.[189,232,1786,1812]" targetBox="[189,1398,636,1766]" targetPageId="9">
|
||||
<paragraph id="B56436CEFFD7AD6FFF19C385FD7AA793" blockId="9.[188,1399,1786,2024]" pageId="9" pageNumber="247">
|
||||
<emphasis id="87AFEADCFFD7AD6FFF19C385FED9A76F" bold="true" box="[189,265,1786,1812]" pageId="9" pageNumber="247">Fig. 4.</emphasis>
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD7AD6FFEBBC384FDD7A76F" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[287,519,1786,1812]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="247" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD7AD6FFEBBC384FDD7A76F" box="[287,519,1786,1812]" italics="true" pageId="9" pageNumber="247">Megacoxa mjoelner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD7AD6FFDA9C384FDB9A76E" box="[525,617,1787,1813]" pageId="9" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, ♀, holotype (CNU-HYM-MA-2016209).
|
||||
<emphasis id="87AFEADCFFD7AD6FFBF8C385FBA1A76F" bold="true" box="[1116,1137,1786,1812]" pageId="9" pageNumber="247">A</emphasis>
|
||||
. Habitus, lateral view.
|
||||
<emphasis id="87AFEADCFFD7AD6FFF18C261FF00A743" bold="true" box="[188,208,1822,1848]" pageId="9" pageNumber="247">B</emphasis>
|
||||
. Fore wings.
|
||||
<emphasis id="87AFEADCFFD7AD6FFECBC262FE54A74C" bold="true" box="[367,388,1821,1847]" pageId="9" pageNumber="247">C</emphasis>
|
||||
. Mesosoma, lateral view.
|
||||
<emphasis id="87AFEADCFFD7AD6FFD10C261FD19A743" bold="true" box="[692,713,1822,1848]" pageId="9" pageNumber="247">D</emphasis>
|
||||
. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C7D4FE82A355" blockId="10.[189,1400,682,814]" pageId="10" pageNumber="248">
|
||||
WINGS. Fore wing (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFE08C7D5FDD9A2BE" box="[428,521,682,709]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4B</figureCitation>
|
||||
) hyaline with veins C, R1, Sc+R, M+Cu, A, Rs+M, Rs, M and Cu fully pigmented; Rs+M aligned with M+Cu; medial cell wide distally; Rs present between Rs+M and r-rs, closing first submarginal cell; marginal cell narrow, closed in straight line by Rs; hind wing could not be observed.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C629FC08A3A2" blockId="10.[189,1398,853,986]" pageId="10" pageNumber="248">
|
||||
METASOMA (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFEF4C62AFE60A30B" box="[336,432,853,880]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4A</figureCitation>
|
||||
). 1.27 × as long as mesosoma, slender, fusiform, without prominent sculpture or pilosity; metasomal segment 2 largest, metasoma widest there; sternum 7 tapered apically, folded over medially along longitudinal axis; ovipositor (
|
||||
<figureCitation id="2DE02A4BFFD4AD6CFD40C6E3FC94A3CD" box="[740,836,924,950]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="10" pageNumber="248">Fig. 4A</figureCitation>
|
||||
) bent ventrally, apparently broken, 1
|
||||
<superScript id="42AE9B86FFD4AD6CFAAEC6E4FAC5A3D2" attach="left" box="[1290,1301,923,937]" fontSize="6" pageId="10" pageNumber="248">st</superScript>
|
||||
and 2
|
||||
<superScript id="42AE9B86FFD4AD6CFAC1C6E4FAA6A3D2" attach="none" box="[1381,1398,923,937]" fontSize="6" pageId="10" pageNumber="248">nd</superScript>
|
||||
valvulae separated at tip and no teeth observed on tip of 2
|
||||
<superScript id="42AE9B86FFD4AD6CFCC6C6C1FCA3A3B7" attach="left" box="[866,883,958,972]" fontSize="6" pageId="10" pageNumber="248">nd</superScript>
|
||||
valvula.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6CFF4EC17EFEFAA460" blockId="10.[189,313,1025,1086]" box="[234,298,1025,1051]" pageId="10" pageNumber="248">
|
||||
<emphasis id="87AFEADCFFD4AD6CFF4EC17EFEFAA460" bold="true" box="[234,298,1025,1051]" pageId="10" pageNumber="248">Male</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C15BFEE9A445" blockId="10.[189,313,1025,1086]" box="[189,313,1060,1086]" pageId="10" pageNumber="248">Unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD4AD6CFF19C11AFC0EA4AA" pageId="10" pageNumber="248" type="discussion">
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C11AFEFFA404" blockId="10.[189,303,1125,1151]" box="[189,303,1125,1151]" pageId="10" pageNumber="248">
|
||||
<heading id="EE2C81A2FFD4AD6CFF19C11AFEFFA404" bold="true" box="[189,303,1125,1151]" fontSize="11" level="3" pageId="10" pageNumber="248" reason="3">
|
||||
<emphasis id="87AFEADCFFD4AD6CFF19C11AFEFFA404" bold="true" box="[189,303,1125,1151]" pageId="10" pageNumber="248">Remarks</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD4AD6CFF19C1EBFC0EA4AA" blockId="10.[189,1398,1172,1233]" pageId="10" pageNumber="248">
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD4AD6CFF68C1EBFE65A4D5" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[204,437,1172,1198]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="248" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD4AD6CFF68C1EBFE65A4D5" box="[204,437,1172,1198]" italics="true" pageId="10" pageNumber="248">Megacoxa mjoelner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD4AD6CFE1FC1EBFDC5A4D5" box="[443,533,1172,1198]" pageId="10" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
mostly resembles †
|
||||
<taxonomicName id="72DB4D4DFFD4AD6CFD59C1EBFC40A4D5" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[765,912,1172,1198]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="248" phylum="Arthropoda" rank="species" species="brazideci" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD4AD6CFD59C1EBFC40A4D5" box="[765,912,1172,1198]" italics="true" pageId="10" pageNumber="248">M. brazideci</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD4AD6CFC33C1EBFC25A4D5" box="[919,1013,1172,1198]" pageId="10" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, from which it differs in having a smooth frons, shorter antennae and the axillae not being contiguous.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
269
data/3D/72/87/3D7287D8FFD8AD6EFDC8C740FC13A30D.xml
Normal file
269
data/3D/72/87/3D7287D8FFD8AD6EFDC8C740FC13A30D.xml
Normal file
|
|
@ -0,0 +1,269 @@
|
|||
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<mods:title id="978AB31BC004347EEBA5D901C8AA9C3E">Three new species of the long-tailed wasp genus † Megacoxa Brazidec et al., 2024 (Hymenoptera: Megalyridae) from Kachin amber</mods:title>
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<mods:namePart id="A0222170CDC73F7F1C296589866AC757">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="1BB1B3C59C692A465AC3335CCC1C933A">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark.</mods:affiliation>
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<mods:namePart id="4A7403FECD67343D27FD5B6A0873A123">Wu, Qiong</mods:namePart>
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<mods:affiliation id="6FC6C7950F632DB9FEB6A472C7A2E3E0">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark. & College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:nameIdentifier id="8614892D782E406330742AF868294B51" type="email">narsleslie@163.com</mods:nameIdentifier>
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<mods:namePart id="EC58CE6B2ADF356382949C34CC952CB6">Wang, Zhen</mods:namePart>
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<mods:affiliation id="7828FEFB6C873F2807EBBD9B2365D049">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark. & College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:nameIdentifier id="4F4E98A7425921A8D608D240CC8D29DA" type="email">wangzhen@cnu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="90C6005D2CB08F774F15479C7F0B38BD">Gao, Tai-Ping</mods:namePart>
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<mods:affiliation id="CE0FE472D9EE83E354D9F31CCB34A4A0">College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:namePart id="B668CEF768B2AFF22175DC8B07B37F30">Ren, Dong</mods:namePart>
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<mods:date id="CAA188E32AF80D5FA6F75CFE7C555234">2025</mods:date>
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†
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<taxonomicName id="72DB4D4DFFD8AD60FDDFC740FCB6A222" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[635,870,575,601]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="gungner" status="sp. nov.">
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<emphasis id="87AFEADCFFD8AD60FDDFC740FCB6A222" bold="true" box="[635,870,575,601]" italics="true" pageId="6" pageNumber="244">Megacoxa gungner</emphasis>
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<taxonomicNameLabel id="9C9C57A7FFD8AD60FCC9C73FFC18A221" box="[877,968,576,602]" pageId="6" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="B56436CEFFD8AD60FD53C7FAFCECA2DB" blockId="6.[359,1228,575,672]" box="[759,828,645,672]" pageId="6" pageNumber="244">
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<figureCitation id="2DE02A4BFFD8AD60FD53C7FAFCECA2DB" box="[759,828,645,672]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="6" pageNumber="244">Fig. 3</figureCitation>
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</paragraph>
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<subSubSection id="FDC16545FFD8AD60FF19C7B8FDD8A398" pageId="6" pageNumber="244" type="diagnosis">
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<paragraph id="B56436CEFFD8AD60FF19C7B8FEE6A29A" blockId="6.[189,310,711,737]" box="[189,310,711,737]" pageId="6" pageNumber="244">
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<heading id="EE2C81A2FFD8AD60FF19C7B8FEE6A29A" bold="true" box="[189,310,711,737]" fontSize="11" level="3" pageId="6" pageNumber="244" reason="3">
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<emphasis id="87AFEADCFFD8AD60FF19C7B8FEE6A29A" bold="true" box="[189,310,711,737]" pageId="6" pageNumber="244">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B56436CEFFD8AD60FF19C789FDD8A398" blockId="6.[188,1400,757,995]" pageId="6" pageNumber="244">
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Compact body, metasoma of approx. same length as mesosoma (
|
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<figureCitation id="2DE02A4BFFD8AD60FC76C78AFB87A36B" box="[978,1111,757,784]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="6" pageNumber="244">Fig. 3A–B</figureCitation>
|
||||
). Antennae shorter than head + mesosoma. Median mesoscutal sulcus smooth, axillae contiguous. Hind coxa without carina and foveae along posterior margin (coxa carinate and foveate in †
|
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<taxonomicName id="72DB4D4DFFD8AD60FC33C642FBB5A32D" box="[919,1125,828,854]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
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<emphasis id="87AFEADCFFD8AD60FC33C642FBB5A32D" box="[919,1125,828,854]" italics="true" pageId="6" pageNumber="244">M. chandrahrasa</emphasis>
|
||||
</taxonomicName>
|
||||
). Hind femur and tibia swollen, tibial apical spurs very short, less than half max. tibial width (spurs longer than tibial width in †
|
||||
<taxonomicName id="72DB4D4DFFD8AD60FF4EC6FCFE67A3E7" box="[234,439,898,924]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
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<emphasis id="87AFEADCFFD8AD60FF4EC6FCFE67A3E7" box="[234,439,898,924]" italics="true" pageId="6" pageNumber="244">M. chandrahrasa</emphasis>
|
||||
</taxonomicName>
|
||||
), hind basitarsomere slightly shorter than remaining tarsomeres combined (much longer in †
|
||||
<taxonomicName id="72DB4D4DFFD8AD60FE9BC6D9FDDDA3C4" box="[319,525,933,959]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
||||
<emphasis id="87AFEADCFFD8AD60FE9BC6D9FDDDA3C4" box="[319,525,933,959]" italics="true" pageId="6" pageNumber="244">M. chandrahrasa</emphasis>
|
||||
</taxonomicName>
|
||||
). Ovipositor elongate, longer than body (
|
||||
<figureCitation id="2DE02A4BFFD8AD60FC58C6DAFB8CA3BB" box="[1020,1116,933,960]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="6" pageNumber="244">Fig. 3A</figureCitation>
|
||||
; ovipositor shorter than body in †
|
||||
<taxonomicName id="72DB4D4DFFD8AD60FE88C6B6FE28A398" box="[300,504,969,995]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
||||
<emphasis id="87AFEADCFFD8AD60FE88C6B6FE28A398" box="[300,504,969,995]" italics="true" pageId="6" pageNumber="244">M. chandrahrasa</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection id="FDC16545FFD8AD60FF19C175FDBFA40D" pageId="6" pageNumber="244" type="etymology">
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<paragraph id="B56436CEFFD8AD60FF19C175FE94A45F" blockId="6.[189,324,1034,1060]" box="[189,324,1034,1060]" pageId="6" pageNumber="244">
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<heading id="EE2C81A2FFD8AD60FF19C175FE94A45F" bold="true" box="[189,324,1034,1060]" fontSize="11" level="3" pageId="6" pageNumber="244" reason="3">
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<emphasis id="87AFEADCFFD8AD60FF19C175FE94A45F" bold="true" box="[189,324,1034,1060]" pageId="6" pageNumber="244">Etymology</emphasis>
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</heading>
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<paragraph id="B56436CEFFD8AD60FF19C146FDBFA40D" blockId="6.[189,1399,1081,1142]" pageId="6" pageNumber="244">The species epithet is after Gungner, the never-missing spear of Odin, king of the Norse gods. It refers to the long ovipositor of this species.</paragraph>
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</subSubSection>
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<emphasis id="87AFEADCFFD8AD60FF19C1E2FEBCA4CC" bold="true" box="[189,364,1181,1207]" pageId="6" pageNumber="244">Type material</emphasis>
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|
||||
<paragraph id="B56436CEFFD8AD60FF4EC1B3FE8CA49D" blockId="6.[189,1399,1228,1360]" box="[234,348,1228,1254]" pageId="6" pageNumber="244">
|
||||
<emphasis id="87AFEADCFFD8AD60FF4EC1B3FE8CA49D" bold="true" box="[234,348,1228,1254]" pageId="6" pageNumber="244">
|
||||
<typeStatus id="6A60886CFFD8AD60FF4EC1B3FE8CA49D" box="[234,348,1228,1254]" pageId="6" pageNumber="244" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD8AD60FF19C190FCD8A534" blockId="6.[189,1399,1228,1360]" pageId="6" pageNumber="244">
|
||||
<materialsCitation id="05B33C93FFD8AD60FF19C190FCD8A534" country="Myanmar" latitude="26.333334" location="Hukawng Valley" longLatPrecision="1242" longitude="96.6" municipality="Myitkyina District" pageId="6" pageNumber="244" specimenCode="CNU-HYM-MA-2016211" specimenCount="1" specimenCount-female="1" stateProvince="Kachin State" typeStatus="holotype">
|
||||
<collectingCountry id="CDCC765EFFD8AD60FF19C190FE86A572" box="[189,342,1263,1289]" name="Myanmar" pageId="6" pageNumber="244">MYANMAR</collectingCountry>
|
||||
•
|
||||
<specimenCount id="A3DDFD47FFD8AD60FED6C18FFE59A571" box="[370,393,1264,1290]" count="1" pageId="6" pageNumber="244" type="female">♀</specimenCount>
|
||||
;
|
||||
<collectingRegion id="771FF82CFFD8AD60FE3DC190FDE5A572" box="[409,565,1263,1289]" country="Myanmar" name="Kachin" pageId="6" pageNumber="244">Kachin State</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="5500ACB4FFD8AD60FDE6C190FCF0A571" box="[578,800,1263,1290]" pageId="6" pageNumber="244">Myitkyina District</collectingMunicipality>
|
||||
,
|
||||
<location id="B0046015FFD8AD60FC89C190FC27A572" LSID="urn:lsid:plazi:treatment:3D7287D8FFD8AD6EFDC8C740FC13A30D:B0046015FFD8AD60FC89C190FC27A572" box="[813,1015,1263,1289]" country="Myanmar" latitude="26.333334" longLatPrecision="1242" longitude="96.6" municipality="Myitkyina District" name="Hukawng Valley" pageId="6" pageNumber="244" stateProvince="Kachin State">Hukawng Valley</location>
|
||||
; [
|
||||
<geoCoordinate id="D0EF5009FFD8AD60FBAFC192FBABA571" box="[1035,1147,1261,1290]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="6" pageNumber="244" precision="925" value="26.333334">26°20′ N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D0EF5009FFD8AD60FB2CC192FB24A571" box="[1160,1268,1261,1290]" degrees="96" direction="east" minutes="36" orientation="longitude" pageId="6" pageNumber="244" precision="925" value="96.6">96°36′ E</geoCoordinate>
|
||||
];
|
||||
<geologicalTimeScale id="C7FBB665FFD8AD60FAAFC190FBA9A556" pageId="6" pageNumber="244">unnamed horizon, mid-Cretaceous, upper Albian to lower Cenomanian, 98.79± 0.62 Ma</geologicalTimeScale>
|
||||
(
|
||||
<bibRefCitation id="D14A4B3FFFD8AD60FB29C06DFF2BA52B" author="Cruickshank R. D. & Ko K." pageId="6" pageNumber="244" pagination="441 - 455" refId="ref7759" refString="Cruickshank R. D. & Ko K. 2003. Geology of an amber locality in the Hukawng Valley, northern Myanmar. Journal of Asian Earth Sciences 21: 441-455. https://doi.org/10.1016/S1367-9120(02)00044-5" type="journal article" year="2003">Cruickshank & Ko 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD8AD60FEA3C04AFE68A52B" author="Shi G. H. & Grimaldi D. A. & Harlow G. E. & Wang J. & Wang J. & Yang M. C. & Lei W. Y. & Li Q. L. & Li X. H." box="[263,440,1333,1360]" pageId="6" pageNumber="244" pagination="155 - 163" refId="ref8697" refString="Shi G. H., Grimaldi D. A., Harlow G. E., Wang J., Wang J., Yang M. C., Lei W. Y., Li Q. L. & Li X. H. 2012. Age constraint on Burmese amber based on U-Pb dating of zircons. Cretaceous Research 37: 155-163. https://doi.org/10.1016/j.cretres.2012.03.014" type="journal article" year="2012">
|
||||
Shi
|
||||
<emphasis id="87AFEADCFFD8AD60FE92C049FEB9A534" box="[310,361,1333,1359]" italics="true" pageId="6" pageNumber="244">et al</emphasis>
|
||||
. 2012
|
||||
</bibRefCitation>
|
||||
);
|
||||
<specimenCode id="E57D9EB5FFD8AD60FE6FC04AFCD4A534" box="[459,772,1333,1359]" pageId="6" pageNumber="244">CNU-HYM-MA-2016211</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD8AD6EFF19C008FEE9A2C4" lastPageId="8" lastPageNumber="246" pageId="6" pageNumber="244" type="description">
|
||||
<paragraph id="B56436CEFFD8AD60FF19C008FE9EA5EA" blockId="6.[189,334,1399,1425]" box="[189,334,1399,1425]" pageId="6" pageNumber="244">
|
||||
<heading id="EE2C81A2FFD8AD60FF19C008FE9EA5EA" bold="true" box="[189,334,1399,1425]" fontSize="11" level="3" pageId="6" pageNumber="244" reason="3">
|
||||
<emphasis id="87AFEADCFFD8AD60FF19C008FE9EA5EA" bold="true" box="[189,334,1399,1425]" pageId="6" pageNumber="244">Description</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD8AD60FF4EC0D9FE95A5BB" blockId="6.[187,1399,1446,1683]" box="[234,325,1446,1472]" pageId="6" pageNumber="244">
|
||||
<emphasis id="87AFEADCFFD8AD60FF4EC0D9FE95A5BB" bold="true" box="[234,325,1446,1472]" pageId="6" pageNumber="244">Female</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD8AD60FF19C0B6FE2EA6E8" blockId="6.[187,1399,1446,1683]" pageId="6" pageNumber="244">
|
||||
BODY. Length (except ovipositor)
|
||||
<quantity id="72239B2BFFD8AD60FDE1C0B7FD70A598" box="[581,672,1480,1507]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="244" unit="mm" value="5.1">5.1 mm</quantity>
|
||||
, fore wing length
|
||||
<quantity id="72239B2BFFD8AD60FCCAC0B6FC19A598" box="[878,969,1481,1507]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="244" unit="mm" value="3.2">3.2 mm</quantity>
|
||||
, ovipositor length
|
||||
<quantity id="72239B2BFFD8AD60FB3AC0B6FB29A598" box="[1182,1273,1481,1507]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="6" pageNumber="244" unit="mm" value="6.9">6.9 mm</quantity>
|
||||
. Specimen complete, generally well preserved in clear piece of amber with moderate amount of grit, no major occlusions around specimen but several diffraction planes, particularly in lower part of specimen; lower part of head obscured by frothy mass of bubbles with hair-like extensions. Small to minute air bubbles on surface of parts of specimen, especially around antennae, making observations difficult. Syninclusion: remains of spider exuvium.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD8AD60FF19C3C5FD73A7FF" blockId="6.[188,1400,1722,1924]" pageId="6" pageNumber="244">
|
||||
HEAD (
|
||||
<figureCitation id="2DE02A4BFFD8AD60FEABC3C5FEB9A6AF" box="[271,361,1722,1748]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="6" pageNumber="244">Fig. 3B</figureCitation>
|
||||
). Held closely appressed to mesosoma, globular, without prominent sculpture or setation, eyes oval; ocelli situated on anterodorsal part of head in approx. equilateral triangle; postocular carina absent; occipital carina well developed, finely crenulate, slightly incurved dorsally; antennae shorter than head and mesosoma combined, basal antennomeres difficult to observe, distal antennomeres not much longer than wide except apical antennomere, approx. twice as long as wide and tapering; mouthparts difficult to observe, short palps observed.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD9AD61FF19C476FB39A36B" blockId="7.[189,1400,265,784]" pageId="7" pageNumber="245">
|
||||
MESOSOMA. With propectus concealed; pronotum narrow medially, triangular laterally, with anterior thoracic spiracle in notch in upper lateral margin; fore femur spindle-shaped, thickened, fore tibia slender, slightly widening distally, calcar not observed; mesoscutum strongly arched in lateral view, vertical anterior surface curves smoothly into horizontal surface; horizontal surface flat, level with mesoscutellum, not prominently sculptured, with short, evenly spaced hairs; median mesoscutal sulcus distinct (
|
||||
<figureCitation id="2DE02A4BFFD9AD61FE8CC4C6FE57A1A8" box="[296,391,441,467]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="7" pageNumber="245">Fig. 3B</figureCitation>
|
||||
), apparently smooth, extends to transscutal articulation; axillae narrow, continuous medially, separated from mesoscutellum by angled foveate scutoscutellar sulcus; mesoscutellum flat, smooth with small hairs, rounded posteriorly; mesopleuron smooth with small, evenly spaced hairs, with slight depression in middle, mesopleuro-metapleural sulcus straight, distinct, crenulate; mid femur thickened, spindle-shaped, mid tibia slender, slightly expanded distally, with two short apical spurs of approx. same length as distal tibial width; hind coxa broad basally, cone-shaped, without carina or foveae along posterior margin; hind femur very thickened, broadened in middle (
|
||||
<figureCitation id="2DE02A4BFFD9AD61FB2BC7F3FB3DA2DD" box="[1167,1261,652,678]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="7" pageNumber="245">Fig. 3B</figureCitation>
|
||||
); hind tibia very thickened, expanded distally, with dense mat of setae on inner side, two minute apical spurs of subequal length present, spur length less than half distal tibial width; hind basitarsus with short erect pegs on inner side, slightly shorter than remaining tarsomeres, tarsal claws simple, slender.
|
||||
</paragraph>
|
||||
<caption id="E1A46646FFD9AD61FF19C267FDDDA798" ID-DOI="http://doi.org/10.5281/zenodo.14981659" ID-Zenodo-Dep="14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="7" pageNumber="245" startId="7.[189,232,1816,1842]" targetBox="[189,1398,840,1789]" targetPageId="7">
|
||||
<paragraph id="B56436CEFFD9AD61FF19C267FDDDA798" blockId="7.[188,1400,1816,2019]" pageId="7" pageNumber="245">
|
||||
<emphasis id="87AFEADCFFD9AD61FF19C267FEDAA749" bold="true" box="[189,266,1816,1842]" pageId="7" pageNumber="245">Fig. 3.</emphasis>
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD9AD61FE84C266FDD3A749" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[288,515,1817,1842]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="245" phylum="Arthropoda" rank="species" species="gungner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD9AD61FE84C266FDD3A749" box="[288,515,1817,1842]" italics="true" pageId="7" pageNumber="245">Megacoxa gungner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD9AD61FDAEC266FDB7A748" box="[522,615,1817,1843]" pageId="7" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, ♀, holotype (CNU-HYM-MA-2016211).
|
||||
<emphasis id="87AFEADCFFD9AD61FBFFC267FBA0A749" bold="true" box="[1115,1136,1816,1842]" pageId="7" pageNumber="245">A</emphasis>
|
||||
. Habitus, lateral view.
|
||||
<emphasis id="87AFEADCFFD9AD61FF19C243FF01A72D" bold="true" box="[189,209,1852,1878]" pageId="7" pageNumber="245">B</emphasis>
|
||||
. Body, lateral view.
|
||||
<emphasis id="87AFEADCFFD9AD61FE6AC244FE33A72E" bold="true" box="[462,483,1851,1877]" pageId="7" pageNumber="245">C</emphasis>
|
||||
. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C476FE1AA1F7" blockId="8.[188,1399,265,396]" pageId="8" pageNumber="246">
|
||||
WINGS. Fore wing (
|
||||
<figureCitation id="2DE02A4BFFD6AD6EFE08C476FDD8A158" box="[428,520,265,291]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="8" pageNumber="246">Fig. 3C</figureCitation>
|
||||
) hyaline with veins C, R1, Sc+R, M+Cu, A, Rs+M, Rs, M and Cu fully pigmented; Rs+M aligned with M+Cu; medial cell wide distally, rectangular; Rs present between Rs+M and r-rs, closing first submarginal cell; marginal cell narrow, closed in straight line by Rs; hind wings covered by fore wings.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C4CBFCADA220" blockId="8.[189,1399,436,603]" pageId="8" pageNumber="246">
|
||||
METASOMA (
|
||||
<figureCitation id="2DE02A4BFFD6AD6EFEF7C4CBFE63A1B5" box="[339,435,436,462]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="8" pageNumber="246">Fig. 3B</figureCitation>
|
||||
). 1.09 × as long as mesosoma, compact, turnip-shaped, hardly separated from mesosoma, without prominent sculpture but with short, evenly spaced hairs; metasomal segment 2 largest, metasoma widest there; sternum 7 tapered apically, folded over medially along longitudinal axis; ovipositor (
|
||||
<figureCitation id="2DE02A4BFFD6AD6EFE20C761FE31A243" box="[388,481,542,568]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="8" pageNumber="246">Fig. 3A</figureCitation>
|
||||
) extending posteriorly, elongate, length 1.35 × body length; ovipositor proper and sheaths slender, teeth on tip of 2
|
||||
<superScript id="42AE9B86FFD6AD6EFDCCC73FFDA9A235" attach="left" box="[616,633,576,590]" fontSize="6" pageId="8" pageNumber="246">nd</superScript>
|
||||
valvula not observed.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF4EC7FDFEFAA2E7" blockId="8.[189,313,642,703]" box="[234,298,642,668]" pageId="8" pageNumber="246">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF4EC7FDFEFAA2E7" bold="true" box="[234,298,642,668]" pageId="8" pageNumber="246">Male</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C7DAFEE9A2C4" blockId="8.[189,313,642,703]" box="[189,313,677,703]" pageId="8" pageNumber="246">Unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD6AD6EFF19C798FC13A30D" pageId="8" pageNumber="246" type="discussion">
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C798FEFFA37A" blockId="8.[189,303,743,769]" box="[189,303,743,769]" pageId="8" pageNumber="246">
|
||||
<heading id="EE2C81A2FFD6AD6EFF19C798FEFFA37A" bold="true" box="[189,303,743,769]" fontSize="11" level="3" pageId="8" pageNumber="246" reason="3">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF19C798FEFFA37A" bold="true" box="[189,303,743,769]" pageId="8" pageNumber="246">Remarks</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD6AD6EFF19C669FC13A30D" blockId="8.[188,1398,789,886]" pageId="8" pageNumber="246">
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFF68C669FE7EA354" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[204,430,790,815]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="gungner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD6AD6EFF68C669FE7EA354" box="[204,430,790,815]" italics="true" pageId="8" pageNumber="246">Megacoxa gungner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD6AD6EFE10C669FDDEA34B" box="[436,526,790,816]" pageId="8" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
mostly resembles †
|
||||
<taxonomicName id="72DB4D4DFFD6AD6EFD50C669FC6FA354" box="[756,959,789,815]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="246" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
||||
<emphasis id="87AFEADCFFD6AD6EFD50C669FC6FA354" box="[756,959,789,815]" italics="true" pageId="8" pageNumber="246">M. chandrahrasa</emphasis>
|
||||
</taxonomicName>
|
||||
among the other species in the genus, but differs from the latter in having the hind coxa without prominent carina or foveae, the apical hind tibial spurs and hind barsitarsus shorter, and the ovipositor longer.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
349
data/3D/72/87/3D7287D8FFDCAD65FD89C225FD18A6BF.xml
Normal file
349
data/3D/72/87/3D7287D8FFDCAD65FD89C225FD18A6BF.xml
Normal file
|
|
@ -0,0 +1,349 @@
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<mods:title id="D8F240B6D4A7BCCCD4037D435AE1CB11">Three new species of the long-tailed wasp genus † Megacoxa Brazidec et al., 2024 (Hymenoptera: Megalyridae) from Kachin amber</mods:title>
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<mods:namePart id="EB56B427162E16F9A5B6C4ABB677B5A4">Wu, Qiong</mods:namePart>
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<mods:affiliation id="C770A50A10A06B8480DB52CB3D931D5A">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark. & College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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Genus †
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<taxonomicName id="72DB4D4DFFDCAD64FD2AC224FBD7A70F" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[654,1031,1882,1909]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="240" phylum="Arthropoda" rank="genus">
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<emphasis id="87AFEADCFFDCAD64FD2AC224FCDBA70E" bold="true" box="[654,779,1883,1909]" italics="true" pageId="2" pageNumber="240">Megacoxa</emphasis>
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<bibRefCitation id="D14A4B3FFFDCAD64FCB7C225FBD7A70F" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[787,1031,1882,1909]" pageId="2" pageNumber="240" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
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Brazidec
|
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<emphasis id="87AFEADCFFDCAD64FC26C224FC6DA70F" box="[898,957,1882,1908]" italics="true" pageId="2" pageNumber="240">et al.</emphasis>
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, 2024
|
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</bibRefCitation>
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</taxonomicName>
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</heading>
|
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</paragraph>
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</subSubSection>
|
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<paragraph id="B56436CEFFDCAD64FF19C2E3FE88A7CE" blockId="2.[189,344,1947,1974]" box="[189,344,1947,1974]" pageId="2" pageNumber="240">
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<heading id="EE2C81A2FFDCAD64FF19C2E3FE88A7CE" bold="true" box="[189,344,1947,1974]" fontSize="11" level="3" pageId="2" pageNumber="240" reason="3">
|
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<emphasis id="87AFEADCFFDCAD64FF19C2E3FE88A7CE" bold="true" box="[189,344,1947,1974]" pageId="2" pageNumber="240">Type species</emphasis>
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</heading>
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</paragraph>
|
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</subSubSection>
|
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<subSubSection id="FDC16545FFDCAD64FF19C2B5FD75A79F" box="[189,677,1994,2021]" pageId="2" pageNumber="240" type="reference_group">
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<paragraph id="B56436CEFFDCAD64FF19C2B5FD75A79F" blockId="2.[189,677,1994,2021]" box="[189,677,1994,2021]" pageId="2" pageNumber="240">
|
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†
|
||||
<taxonomicName id="72DB4D4DFFDCAD64FF68C2B4FD70A79F" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[204,672,1994,2021]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="240" phylum="Arthropoda" rank="species" species="janzeni">
|
||||
<emphasis id="87AFEADCFFDCAD64FF68C2B4FE72A79F" box="[204,418,1995,2020]" italics="true" pageId="2" pageNumber="240">Megacoxa janzeni</emphasis>
|
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<bibRefCitation id="D14A4B3FFFDCAD64FE0DC2B5FD70A79F" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[425,672,1994,2021]" pageId="2" pageNumber="240" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
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Brazidec
|
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<emphasis id="87AFEADCFFDCAD64FDBDC2B4FD84A79F" box="[537,596,1994,2020]" italics="true" pageId="2" pageNumber="240">et al.</emphasis>
|
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, 2024
|
||||
</bibRefCitation>
|
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</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="E1A46646FFDDAD65FF19C060FD8DA5FB" ID-DOI="http://doi.org/10.5281/zenodo.14981653" ID-Zenodo-Dep="14981653" httpUri="https://zenodo.org/record/14981653/files/figure.png" pageId="3" pageNumber="241" startId="3.[189,232,1311,1337]" targetBox="[189,1397,269,1294]" targetPageId="3">
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<paragraph id="B56436CEFFDDAD65FF19C060FD8DA5FB" blockId="3.[187,1398,1311,1408]" pageId="3" pageNumber="241">
|
||||
<emphasis id="87AFEADCFFDDAD65FF19C060FED9A542" bold="true" box="[189,265,1311,1337]" pageId="3" pageNumber="241">Fig. 1.</emphasis>
|
||||
Phylogeny of
|
||||
<taxonomicName id="72DB4D4DFFDDAD65FE0BC060FD97A542" authorityName="Schletterer" authorityYear="1889" box="[431,583,1311,1337]" class="Insecta" family="Megalyridae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="241" phylum="Arthropoda" rank="family">Megalyridae</taxonomicName>
|
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. Strict consensus tree of six trees obtained by implied weights analyses with k = 5. All genera except †
|
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<taxonomicName id="72DB4D4DFFDDAD65FDBBC03CFD49A527" box="[543,665,1347,1372]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="241" phylum="Arthropoda" rank="genus">
|
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<emphasis id="87AFEADCFFDDAD65FDBBC03CFD49A527" box="[543,665,1347,1372]" italics="true" pageId="3" pageNumber="241">Megacoxa</emphasis>
|
||||
</taxonomicName>
|
||||
represented by only one terminal, number in brackets indicates total number of recognized species.
|
||||
</paragraph>
|
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</caption>
|
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<paragraph id="B56436CEFFDDAD65FF19C0D7FE7FA5B9" blockId="3.[189,431,1448,1474]" box="[189,431,1448,1474]" pageId="3" pageNumber="241">
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<heading id="EE2C81A2FFDDAD65FF19C0D7FE7FA5B9" bold="true" box="[189,431,1448,1474]" fontSize="11" level="3" pageId="3" pageNumber="241" reason="3">
|
||||
<emphasis id="87AFEADCFFDDAD65FF19C0D7FE7FA5B9" bold="true" box="[189,431,1448,1474]" pageId="3" pageNumber="241">Emended diagnosis</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDDAD65FF19C0A8FD18A6BF" blockId="3.[189,1400,1495,1732]" pageId="3" pageNumber="241">Compound eye oval; scape wider than long; low occipital carina slightly crenulate. Mesoscutoscutellar sulcus crenulate. Fore wing venation with veins R1, M+Cu, Sc+R, A and Cu fully pigmented; vein A connected to medial cell with vein cu-a; vein Rs closing marginal cell without bend. Metacoxa much enlarged, with dorsal surface rounded to angular, outer metacoxal surface posteriorly foveate, inner metacoxal surface almost entirely accommodated in anterolateral, shallow concavity of metasoma. Metatibia and metabasitarsus sometimes with ventral row of comb-like setae; two mesotibial and two metatibial spurs. Metasomal length variable.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD2AD6AFF19C22BFD8AA715" box="[189,602,1876,1902]" pageId="12" pageNumber="250" type="nomenclature">
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<paragraph id="B56436CEFFD2AD6AFF19C22BFD8AA715" blockId="12.[189,602,1876,1902]" box="[189,602,1876,1902]" pageId="12" pageNumber="250">
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<heading id="EE2C81A2FFD2AD6AFF19C22BFD8AA715" bold="true" box="[189,602,1876,1902]" fontSize="11" level="3" pageId="12" pageNumber="250" reason="3">
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<emphasis id="87AFEADCFFD2AD6AFF19C22BFD8AA715" bold="true" box="[189,602,1876,1902]" pageId="12" pageNumber="250">
|
||||
Key to species of †
|
||||
<taxonomicName id="72DB4D4DFFD2AD6AFE07C22BFDF0A715" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[419,544,1876,1902]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="250" phylum="Arthropoda" rank="genus">
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<emphasis id="87AFEADCFFD2AD6AFE07C22BFDF0A715" bold="true" box="[419,544,1876,1902]" italics="true" pageId="12" pageNumber="250">Megacoxa</emphasis>
|
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</taxonomicName>
|
||||
spp.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFD2AD6BFF19C2FCFAA7A4E0" lastPageId="13" lastPageNumber="251" pageId="12" pageNumber="250" type="key">
|
||||
<key id="C14A61FFFFD2AD6BFF19C2FCFAA7A4E0" lastPageId="13" lastPageNumber="251" pageId="12" pageNumber="250">
|
||||
<keyStep id="0E2F2E6BFFD2AD6AFF19C2FCFAA7A7BB" pageId="12" pageNumber="250">
|
||||
<paragraph id="B56436CEFFD2AD6AFF19C2FCFAA7A7E6" blockId="12.[189,1399,1922,1984]" box="[189,1399,1922,1949]" pageId="12" pageNumber="250">
|
||||
<keyLead id="0E2A95FBFFD2AD6AFF19C2FCFAA7A7E6" box="[189,1399,1922,1949]" pageId="12" pageNumber="250">
|
||||
1. Metasoma at most as long as mesosoma (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFD74C2FDFC80A7E6" box="[720,848,1922,1949]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="12" pageNumber="250">Fig. 3A–B</figureCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD2AD6AFCF8C2FDFB9DA7E6" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[860,1101,1922,1949]" pageId="12" pageNumber="250" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD2AD6AFC68C2FCFBD6A7E7" box="[972,1030,1922,1948]" italics="true" pageId="12" pageNumber="250">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 6b, e) ................... 2
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD2AD6AFF65C2D9FAA7A7BB" blockId="12.[189,1399,1922,1984]" box="[193,1399,1958,1984]" pageId="12" pageNumber="250">
|
||||
<keyLead id="0E2A95FBFFD2AD6AFF65C2D9FAA7A7BB" box="[193,1399,1958,1984]" pageId="12" pageNumber="250">
|
||||
– Metasoma clearly longer than mesosoma (
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFD7EC2D9FCECA7BB" box="[730,828,1958,1984]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="12" pageNumber="250">Figs 2A</figureCitation>
|
||||
,
|
||||
<figureCitation id="2DE02A4BFFD2AD6AFCECC2D9FCBDA7BB" box="[840,877,1958,1984]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="12" pageNumber="250">4A</figureCitation>
|
||||
) ................................................................. 3
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0E2F2E6BFFD3AD6BFF19C476FAA7A246" pageId="13" pageNumber="251">
|
||||
<paragraph id="B56436CEFFD3AD6BFF19C476FAA7A1CB" blockId="13.[189,1399,265,573]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF19C476FAA7A1CB" pageId="13" pageNumber="251">
|
||||
2. Hind coxal posterior margin strongly carinate and with row of foveae (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFB15C476FEF9A13D" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFA8BC475FAA7A158" box="[1327,1399,265,291]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 6c); hind tibial apical spurs longer than max. width of hind tibia (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFB19C453FEF9A111" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFA91C452FAA7A13D" box="[1333,1399,300,326]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 6b, g); hind basitarsus approx. 1.5 × as long as remaining tarsomeres combined (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFF56C40DFE3FA1F6" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[242,495,370,397]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFEC3C40CFE76A1F7" box="[359,422,370,396]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 6b, g); ovipositor shorter than body (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFC57C40DFB20A1F6" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[1011,1264,370,397]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFBCCC40CFB76A1F7" box="[1128,1190,370,396]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 6b, e) ................................................................................. †
|
||||
<taxonomicName id="72DB4D4DFFD3AD6BFCF1C4E8FAA7A1CB" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[853,1399,406,432]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="251" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
||||
<emphasis id="87AFEADCFFD3AD6BFCF1C4E8FBACA1CB" box="[853,1148,406,432]" italics="true" pageId="13" pageNumber="251">Megacoxa chandrahrasa</emphasis>
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFB27C4E9FAA7A1CB" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[1155,1399,406,432]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFB56C4E8FAFDA1CB" box="[1266,1325,406,432]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
, 2024
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD3AD6BFF65C4C5FAA7A246" blockId="13.[189,1399,265,573]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF65C4C5FAA7A246" pageId="13" pageNumber="251">
|
||||
– Hind coxal posterior margin without carina or row of foveae (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFBBFC4C6FBAFA1A8" box="[1051,1151,441,467]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="13" pageNumber="251">Fig. 3B</figureCitation>
|
||||
); hind tibial apical spurs much shorter than max. width of hind tibia (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFCDCC4A3FC0BA18D" box="[888,987,476,502]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="13" pageNumber="251">Fig. 3B</figureCitation>
|
||||
); hind basitarsus slightly shorter than combined length of remaining tarsomeres (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFCD0C480FC31A261" box="[884,993,511,538]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="13" pageNumber="251">Fig. 3A</figureCitation>
|
||||
); ovipositor longer than body (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFF57C75DFE9FA246" box="[243,335,546,573]" captionStart="Fig" captionStartId="7.[189,232,1816,1842]" captionTargetBox="[189,1398,840,1789]" captionTargetId="figure-309@7.[189,1398,840,1789]" captionTargetPageId="7" captionText="Fig. 3. †Megacoxa gungner sp. nov., ♀, holotype (CNU-HYM-MA-2016211). A. Habitus, lateral view. B. Body, lateral view. C. Fore wing. Yellow arrow = median mesoscutal sulcus; blue arrows = hind tibial apical spurs; white arrow = medial cell. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesonotum; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, M+Cu, Rs) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981659" httpUri="https://zenodo.org/record/14981659/files/figure.png" pageId="13" pageNumber="251">Fig. 3A</figureCitation>
|
||||
) ............................................................................................... †
|
||||
<taxonomicName id="72DB4D4DFFD3AD6BFB96C75CFAC5A247" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[1074,1301,547,572]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="251" phylum="Arthropoda" rank="species" species="gungner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD3AD6BFB96C75CFAC5A247" box="[1074,1301,547,572]" italics="true" pageId="13" pageNumber="251">Megacoxa gungner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD3AD6BFAB8C75CFAA7A246" box="[1308,1399,547,573]" pageId="13" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0E2F2E6BFFD3AD6BFF19C713FAA7A28B" pageId="13" pageNumber="251">
|
||||
<paragraph id="B56436CEFFD3AD6BFF19C713FAA7A2D2" blockId="13.[189,1399,619,752]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF19C713FAA7A2D2" pageId="13" pageNumber="251">
|
||||
3. Hind coxal posterior margin with row of large foveae (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFCCBC714FBB3A2FD" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[879,1123,619,646]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFC46C713FBCDA2FE" box="[994,1053,619,645]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: figs 7b, 8c, g); median mesoscutal sulcus smooth or crenulate ............................................................................................. 4
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD3AD6BFF65C7CCFAA7A28B" blockId="13.[189,1399,619,752]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF65C7CCFAA7A28B" pageId="13" pageNumber="251">
|
||||
– Hind coxal posterior margin without foveae (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFCB9C7CDFC54A2B7" box="[797,900,690,716]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="13" pageNumber="251">Figs 2D</figureCitation>
|
||||
,
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFC32C7CDFC6DA2B7" box="[918,957,690,716]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="13" pageNumber="251">4C</figureCitation>
|
||||
); median mesoscutal sulcus smooth (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFF57C7AAFE9EA294" box="[243,334,725,751]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="13" pageNumber="251">Fig. 4B</figureCitation>
|
||||
) ............................................................................................................................................ 5
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0E2F2E6BFFD3AD6BFF19C661FAA7A393" pageId="13" pageNumber="251">
|
||||
<paragraph id="B56436CEFFD3AD6BFF19C661FAA7A304" blockId="13.[189,1399,798,1001]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF19C661FAA7A304" pageId="13" pageNumber="251">
|
||||
4. Median mesosocutal sulcus smooth (e.g.,
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFD6BC661FCFAA343" box="[719,810,798,824]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="13" pageNumber="251">Fig. 4B</figureCitation>
|
||||
); axillae not abutting medially (e.g.,
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFB77C661FAFEA343" box="[1235,1326,798,824]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="13" pageNumber="251">Fig. 4B</figureCitation>
|
||||
); hind tibia triangular, broadest close to apex (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFD13C63EFC7BA320" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[695,939,833,859]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFC8EC63DFCB5A320" box="[810,869,833,859]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 7d) ............................................... ............................................................................................ †
|
||||
<taxonomicName id="72DB4D4DFFD3AD6BFC01C61AFAA7A304" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[933,1399,868,895]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="251" phylum="Arthropoda" rank="species" species="janzeni">
|
||||
<emphasis id="87AFEADCFFD3AD6BFC01C61AFBACA305" box="[933,1148,869,894]" italics="true" pageId="13" pageNumber="251">Megacoxa janzeni</emphasis>
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFB27C61BFAA7A304" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[1155,1399,868,895]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFB56C61AFAFDA305" box="[1266,1325,868,894]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
, 2024
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD3AD6BFF65C6F7FAA7A393" blockId="13.[189,1399,798,1001]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF65C6F7FAA7A393" pageId="13" pageNumber="251">
|
||||
– Median mesoscutal sulcus crenulate (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFD38C6F7FD28A3D8" box="[668,760,904,931]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="13" pageNumber="251">Fig. 5A</figureCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFCA7C6F7FC3FA3D9" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[771,1007,904,930]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFCD4C6F6FC79A3D9" box="[880,937,904,930]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 8f); axillae abutting medially (
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFF56C6D4FE36A3BE" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[242,486,939,965]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFEC0C6D3FE4FA3BE" box="[356,415,939,965]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 8f); hind tibia spindle-shaped, broadest at two-thirds from base (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFAB1C6D4FAA1A3BD" box="[1301,1393,939,966]" captionStart="Fig" captionStartId="11.[189,232,1803,1829]" captionTargetBox="[189,1398,266,1777]" captionTargetId="figure-118@11.[189,1398,266,1777]" captionTargetPageId="11" captionText="Fig. 5. †Megacoxa synchrotron Brazidec et al., 2024, ♀ (CNU-HYM-MA-2016208). A. Habitus, dorsal view. B. Head and mesosoma, ventral view. C. Hind leg, ventral view. D. Fore wings. Yellow arrow = median mesoscutal sulcus; red arrow = occipital carina; green arrows = mandibular teeth; blue arrow = hind tibial apical spur; white arrow = medial cell. Abbreviations: ax = axilla; fe3 = hind femur; mc = marginal cell; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti3 = hind tibia. Remaining abbreviations (A, Cu, M, 1cu-a etc.) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981663" httpUri="https://zenodo.org/record/14981663/files/figure.png" pageId="13" pageNumber="251">Fig. 5C</figureCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFF4EC6B1FE0BA393" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[234,475,974,1000]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFEFEC6B0FE45A393" box="[346,405,974,1000]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
: fig. 8a) .......................................†
|
||||
<taxonomicName id="72DB4D4DFFD3AD6BFCCAC6B0FAA7A393" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[878,1399,974,1001]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="251" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFD3AD6BFCCAC6B0FBACA393" box="[878,1148,974,1000]" italics="true" pageId="13" pageNumber="251">Megacoxa synchrotron</emphasis>
|
||||
<bibRefCitation id="D14A4B3FFFD3AD6BFB27C6B1FAA7A393" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[1155,1399,974,1001]" pageId="13" pageNumber="251" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFD3AD6BFB56C6B0FAFDA393" box="[1266,1325,974,1000]" italics="true" pageId="13" pageNumber="251">et al.</emphasis>
|
||||
, 2024
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0E2F2E6BFFD3AD6BFF19C168FAA7A4E0" pageId="13" pageNumber="251">
|
||||
<paragraph id="B56436CEFFD3AD6BFF19C168FAA7A42E" blockId="13.[189,1399,1047,1179]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF19C168FAA7A42E" pageId="13" pageNumber="251">
|
||||
5. Head with transverse striae on frons (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFD08C168FCDAA44A" box="[684,778,1047,1073]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="13" pageNumber="251">Fig. 2B</figureCitation>
|
||||
); antennae longer than head + mesosoma combined (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFF51C145FE83A42F" box="[245,339,1082,1108]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="13" pageNumber="251">Fig. 2A</figureCitation>
|
||||
); axillae abutting medially .....................................................†
|
||||
<taxonomicName id="72DB4D4DFFD3AD6BFB82C144FAC5A42F" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[1062,1301,1082,1108]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="251" phylum="Arthropoda" rank="species" species="brazideci" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD3AD6BFB82C144FAC5A42F" box="[1062,1301,1082,1108]" italics="true" pageId="13" pageNumber="251">Megacoxa brazideci</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD3AD6BFAB8C144FAA7A42E" box="[1308,1399,1083,1109]" pageId="13" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD3AD6BFF65C121FAA7A4E0" blockId="13.[189,1399,1047,1179]" pageId="13" pageNumber="251">
|
||||
<keyLead id="0E2A95FBFFD3AD6BFF65C121FAA7A4E0" pageId="13" pageNumber="251">
|
||||
– Frons smooth; antennae shorter than head + mesosoma combined (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFBB0C122FBA6A403" box="[1044,1142,1117,1144]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="13" pageNumber="251">Fig. 4A</figureCitation>
|
||||
); axillae not abutting medially (
|
||||
<figureCitation id="2DE02A4BFFD3AD6BFEC5C1FEFE6CA4E0" box="[353,444,1153,1179]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[189,1398,636,1766]" captionTargetId="figure-256@9.[189,1398,636,1770]" captionTargetPageId="9" captionText="Fig. 4. †Megacoxa mjoelner sp. nov., ♀, holotype (CNU-HYM-MA-2016209). A. Habitus, lateral view. B. Fore wings. C. Mesosoma, lateral view. D. Head, lateral view. Blue arrows = hind tibial apical spurs; black arrow = median mesoscutal sulcus; white arrow = medial cell; green arrow = spiracular notch; red arrow = occipital carina. Abbreviations: ax = axilla; cx3 = hind coxa; fe3 = hind femur; mc = marginal cell; n1 = pronotum; n2 = mesoscutum; pl2 = mesopleuron; pl3 = metapleuron; pt = pterostigma; sc2 = mesoscutellum; smc = submarginal cell; ti1 = fore tibia; ti3 = hind tibia. Remaining abbreviations (Rs, M+Cu) refer to wing veins; see main text." figureDoi="http://doi.org/10.5281/zenodo.14981661" httpUri="https://zenodo.org/record/14981661/files/figure.png" pageId="13" pageNumber="251">Fig. 4B</figureCitation>
|
||||
) ................................................................................ †
|
||||
<taxonomicName id="72DB4D4DFFD3AD6BFB8FC1FDFAC5A4E0" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[1067,1301,1153,1179]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="251" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
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<emphasis id="87AFEADCFFD3AD6BFB8FC1FDFAC5A4E0" box="[1067,1301,1153,1179]" italics="true" pageId="13" pageNumber="251">Megacoxa mjoelner</emphasis>
|
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<taxonomicNameLabel id="9C9C57A7FFD3AD6BFAB8C1FEFAA7A4E0" box="[1308,1399,1153,1179]" pageId="13" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
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|
||||
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|
||||
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||||
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311
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<mods:title id="C907B9765BE3B5024748F6E0E6EC42BC">Three new species of the long-tailed wasp genus † Megacoxa Brazidec et al., 2024 (Hymenoptera: Megalyridae) from Kachin amber</mods:title>
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<mods:namePart id="A1969A1FCBD50506122287FD2827663C">Wu, Qiong</mods:namePart>
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<mods:affiliation id="506D2C8F6DE16F441A74E525FCDA7B27">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark. & College of Life Sciences, Capital Normal University, Beijing 100048, China.</mods:affiliation>
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<mods:namePart id="F00726C3D940F7826ED2E6999A9F1B32">Wang, Zhen</mods:namePart>
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†
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<taxonomicName id="72DB4D4DFFDDAD65FDD3C384FCB9A76E" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[631,873,1786,1813]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="241" phylum="Arthropoda" rank="species" species="brazideci" status="sp. nov.">
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<emphasis id="87AFEADCFFDDAD65FDD3C384FCB9A76E" bold="true" box="[631,873,1786,1813]" italics="true" pageId="3" pageNumber="241">Megacoxa brazideci</emphasis>
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<emphasis id="87AFEADCFFDDAD65FF19C205FEE6A7EF" bold="true" box="[189,310,1914,1940]" pageId="3" pageNumber="241">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B56436CEFFDDAD62FF19C2D6FEE2A1CB" blockId="3.[189,1399,1961,2023]" lastBlockId="4.[189,1399,265,432]" lastPageId="4" lastPageNumber="242" pageId="3" pageNumber="241">
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Body comparatively slender (
|
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<figureCitation id="2DE02A4BFFDDAD65FDBCC2D6FDA7A7B8" box="[536,631,1961,1987]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="3" pageNumber="241">Fig. 2A</figureCitation>
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), metasoma longer than mesosoma (shorter in †
|
||||
<taxonomicName id="72DB4D4DFFDDAD65FB0EC2D5FE12A79C" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="241" phylum="Arthropoda" rank="species" species="chandrahrasa">
|
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<emphasis id="87AFEADCFFDDAD65FB0EC2D5FAA7A7B8" box="[1194,1399,1961,1987]" italics="true" pageId="3" pageNumber="241">M. chandrahrasa</emphasis>
|
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<bibRefCitation id="D14A4B3FFFDDAD65FF19C2B3FE12A79C" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[189,450,1996,2023]" pageId="3" pageNumber="241" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
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Brazidec
|
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<emphasis id="87AFEADCFFDDAD65FE96C2B2FEA3A79D" box="[306,371,1996,2022]" italics="true" pageId="3" pageNumber="241">et al.</emphasis>
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, 2024
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</bibRefCitation>
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</taxonomicName>
|
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and †
|
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<taxonomicName id="72DB4D4DFFDDAD65FDB0C2B2FD73A79D" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[532,675,1997,2022]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="241" phylum="Arthropoda" rank="species" species="gungner" status="sp. nov.">
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<emphasis id="87AFEADCFFDDAD65FDB0C2B2FD73A79D" box="[532,675,1997,2022]" italics="true" pageId="3" pageNumber="241">M. gungner</emphasis>
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</taxonomicName>
|
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<taxonomicNameLabel id="9C9C57A7FFDDAD65FD14C2B2FCC4A79C" box="[688,788,1997,2023]" pageId="3" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
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). Head with transverse striae on frons (
|
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<figureCitation id="2DE02A4BFFDDAD65FAA3C2B3FAB9A79C" box="[1287,1385,1996,2023]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="3" pageNumber="241">Fig. 2B</figureCitation>
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||||
). Antennae longer than head + mesosoma combined. Median mesoscutal sulcus smooth (crenulate in †
|
||||
<taxonomicName id="72DB4D4DFFDAAD62FF68C452FD40A13D" authority="Brazidec et al., 2024" authorityName="Brazidec" authorityYear="2024" box="[204,656,300,327]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="242" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFDAAD62FF68C452FE52A13D" box="[204,386,300,326]" italics="true" pageId="4" pageNumber="242">M. synchrotron</emphasis>
|
||||
<bibRefCitation id="D14A4B3FFFDAAD62FE29C453FD40A13D" author="Brazidec M. & Vilhelmsen L. & Boudinot B. & Richter A. K. F. & Hammel J. & Perkovsky E. E. & Fan Y. & Wang B. & Wang Z. & Wu Q. & Perrichot V." box="[397,656,300,327]" pageId="4" pageNumber="242" pagination="151 - 181" refId="ref7498" refString="Brazidec M., Vilhelmsen L., Boudinot B., Richter A. K. F., Hammel J., Perkovsky E. E., Fan Y., Wang B., Wang Z., Wu Q. & Perrichot V. 2024. Unveiling ancient diversity of long-tailed wasps (Hymenoptera: MegalyrIdae): new taxa frOm CretaceOus KachIn and TaImyr ambers and theIr phylOgenetIc affinItIes. Arthropod Systematics & Phylogeny 82: 151-181. https://doi.org/10.3897/asp.82.e111148" type="book chapter" year="2024">
|
||||
Brazidec
|
||||
<emphasis id="87AFEADCFFDAAD62FDA4C452FDEFA13D" box="[512,575,300,326]" italics="true" pageId="4" pageNumber="242">et al.</emphasis>
|
||||
, 2024
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), axillae contiguous (not contiguous in †
|
||||
<taxonomicName id="72DB4D4DFFDAAD62FB20C452FE96A112" authority="Bracidec et al., 2024" authorityName="Bracidec" authorityYear="2024" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="242" phylum="Arthropoda" rank="species" species="janzeni">
|
||||
<emphasis id="87AFEADCFFDAAD62FB20C452FAD3A13D" box="[1156,1283,301,326]" italics="true" pageId="4" pageNumber="242">M. janzeni</emphasis>
|
||||
Bracidec
|
||||
<emphasis id="87AFEADCFFDAAD62FF19C42FFF2AA112" box="[189,250,335,361]" italics="true" pageId="4" pageNumber="242">et al.</emphasis>
|
||||
, 2024
|
||||
</taxonomicName>
|
||||
and †
|
||||
<taxonomicName id="72DB4D4DFFDAAD62FE35C42FFDF2A112" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[401,546,335,361]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="242" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFDAAD62FE35C42FFDF2A112" box="[401,546,335,361]" italics="true" pageId="4" pageNumber="242">M. mjoelner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFDAAD62FD88C42FFD5DA112" box="[556,653,335,362]" pageId="4" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
). Hind coxa without carina and foveae along posterior margin (coxa carinate and foveate in †
|
||||
<taxonomicName id="72DB4D4DFFDAAD62FD83C40CFD74A1F7" box="[551,676,371,396]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="242" phylum="Arthropoda" rank="species" species="janzeni">
|
||||
<emphasis id="87AFEADCFFDAAD62FD83C40CFD74A1F7" box="[551,676,371,396]" italics="true" pageId="4" pageNumber="242">M. janzeni</emphasis>
|
||||
</taxonomicName>
|
||||
and †
|
||||
<taxonomicName id="72DB4D4DFFDAAD62FD48C40CFC4FA1F7" box="[748,927,370,396]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="242" phylum="Arthropoda" rank="species" species="synchrotron">
|
||||
<emphasis id="87AFEADCFFDAAD62FD48C40CFC4FA1F7" box="[748,927,370,396]" italics="true" pageId="4" pageNumber="242">M. synchrotron</emphasis>
|
||||
</taxonomicName>
|
||||
). Ovipositor elongate, longer than body (
|
||||
<figureCitation id="2DE02A4BFFDAAD62FF62C4E9FEF4A1CB" box="[198,292,406,432]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="4" pageNumber="242">Fig. 2A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFDAAD62FF19C4A8FCA3A238" pageId="4" pageNumber="242" type="etymology">
|
||||
<paragraph id="B56436CEFFDAAD62FF19C4A8FE94A18A" blockId="4.[189,324,471,497]" box="[189,324,471,497]" pageId="4" pageNumber="242">
|
||||
<heading id="EE2C81A2FFDAAD62FF19C4A8FE94A18A" bold="true" box="[189,324,471,497]" fontSize="11" level="3" pageId="4" pageNumber="242" reason="3">
|
||||
<emphasis id="87AFEADCFFDAAD62FF19C4A8FE94A18A" bold="true" box="[189,324,471,497]" pageId="4" pageNumber="242">Etymology</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF19C779FCA3A238" blockId="4.[189,1398,518,579]" pageId="4" pageNumber="242">
|
||||
The new species is named after Manuel Brazidec, the French palaeoentomologist who described †
|
||||
<taxonomicName id="72DB4D4DFFDAAD62FF68C755FE96A238" box="[204,326,554,579]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="242" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="87AFEADCFFDAAD62FF68C755FE96A238" box="[204,326,554,579]" italics="true" pageId="4" pageNumber="242">Megacoxa</emphasis>
|
||||
</taxonomicName>
|
||||
as well as a number of other extinct wasp taxa.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFDAAD62FF19C715FCD9A367" pageId="4" pageNumber="242" type="materials_examined">
|
||||
<paragraph id="B56436CEFFDAAD62FF19C715FEBCA2FF" blockId="4.[189,364,618,644]" box="[189,364,618,644]" pageId="4" pageNumber="242">
|
||||
<heading id="EE2C81A2FFDAAD62FF19C715FEBCA2FF" bold="true" box="[189,364,618,644]" fontSize="11" level="3" pageId="4" pageNumber="242" reason="3">
|
||||
<emphasis id="87AFEADCFFDAAD62FF19C715FEBCA2FF" bold="true" box="[189,364,618,644]" pageId="4" pageNumber="242">Type material</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF4EC7E6FE8CA2C8" blockId="4.[189,1399,665,797]" box="[234,348,665,691]" pageId="4" pageNumber="242">
|
||||
<emphasis id="87AFEADCFFDAAD62FF4EC7E6FE8CA2C8" bold="true" box="[234,348,665,691]" pageId="4" pageNumber="242">
|
||||
<typeStatus id="6A60886CFFDAAD62FF4EC7E6FE8CA2C8" box="[234,348,665,691]" pageId="4" pageNumber="242" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF19C7C3FCD9A367" blockId="4.[189,1399,665,797]" pageId="4" pageNumber="242">
|
||||
<materialsCitation id="05B33C93FFDAAD62FF19C7C3FCD9A367" country="Myanmar" latitude="26.333334" location="Hukawng Valley" longLatPrecision="1242" longitude="96.6" municipality="MyItkyIna DIstrIct" pageId="4" pageNumber="242" specimenCode="CNU-HYM-MA-2016210" specimenCount="1" specimenCount-female="1" stateProvince="KachIn State" typeStatus="holotype">
|
||||
<collectingCountry id="CDCC765EFFDAAD62FF19C7C3FE86A2AD" box="[189,342,700,726]" name="Myanmar" pageId="4" pageNumber="242">MYANMAR</collectingCountry>
|
||||
•
|
||||
<specimenCount id="A3DDFD47FFDAAD62FED6C7C2FE59A2AC" box="[370,393,701,727]" count="1" pageId="4" pageNumber="242" type="female">♀</specimenCount>
|
||||
;
|
||||
<collectingRegion id="771FF82CFFDAAD62FE3DC7C3FDE5A2AD" box="[409,565,700,726]" country="Myanmar" name="Kachin" pageId="4" pageNumber="242">Kachin State</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="5500ACB4FFDAAD62FDE6C7C3FCF0A2AC" box="[578,800,700,727]" pageId="4" pageNumber="242">Myitkyina District</collectingMunicipality>
|
||||
,
|
||||
<location id="B0046015FFDAAD62FC89C7C3FC27A2AD" LSID="urn:lsid:plazi:treatment:3D7287D8FFDDAD60FDCDC384FB28A27A:B0046015FFDAAD62FC89C7C3FC27A2AD" box="[813,1015,700,726]" country="Myanmar" latitude="26.333334" longLatPrecision="1242" longitude="96.6" municipality="MyItkyIna DIstrIct" name="Hukawng Valley" pageId="4" pageNumber="242" stateProvince="KachIn State">Hukawng Valley</location>
|
||||
; [
|
||||
<geoCoordinate id="D0EF5009FFDAAD62FBAFC7C4FBABA2AC" box="[1035,1147,699,727]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="4" pageNumber="242" precision="925" value="26.333334">26°20′ N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D0EF5009FFDAAD62FB2CC7C4FB24A2AC" box="[1160,1268,699,727]" degrees="96" direction="east" minutes="36" orientation="longitude" pageId="4" pageNumber="242" precision="925" value="96.6">96°36′ E</geoCoordinate>
|
||||
];
|
||||
<geologicalTimeScale id="C7FBB665FFDAAD62FAAFC7C3FBA9A281" pageId="4" pageNumber="242">unnamed horizon, mid-Cretaceous, upper Albian to lower Cenomanian, 98.79± 0.62 Ma</geologicalTimeScale>
|
||||
(
|
||||
<bibRefCitation id="D14A4B3FFFDAAD62FB29C7A0FF2BA366" author="Cruickshank R. D. & Ko K." pageId="4" pageNumber="242" pagination="441 - 455" refId="ref7759" refString="Cruickshank R. D. & Ko K. 2003. Geology of an amber locality in the Hukawng Valley, northern Myanmar. Journal of Asian Earth Sciences 21: 441-455. https://doi.org/10.1016/S1367-9120(02)00044-5" type="journal article" year="2003">Cruickshank & Ko 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFDAAD62FEA3C67DFE68A366" author="Shi G. H. & Grimaldi D. A. & Harlow G. E. & Wang J. & Wang J. & Yang M. C. & Lei W. Y. & Li Q. L. & Li X. H." box="[263,440,770,797]" pageId="4" pageNumber="242" pagination="155 - 163" refId="ref8697" refString="Shi G. H., Grimaldi D. A., Harlow G. E., Wang J., Wang J., Yang M. C., Lei W. Y., Li Q. L. & Li X. H. 2012. Age constraint on Burmese amber based on U-Pb dating of zircons. Cretaceous Research 37: 155-163. https://doi.org/10.1016/j.cretres.2012.03.014" type="journal article" year="2012">
|
||||
Shi
|
||||
<emphasis id="87AFEADCFFDAAD62FE92C67CFEA0A367" box="[310,368,770,796]" italics="true" pageId="4" pageNumber="242">et al.</emphasis>
|
||||
2012
|
||||
</bibRefCitation>
|
||||
);
|
||||
<specimenCode id="E57D9EB5FFDAAD62FE6FC67DFCD5A367" box="[459,773,770,796]" pageId="4" pageNumber="242">CNU-HYM-MA-2016210</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FDC16545FFDAAD63FF19C63BFEE9A2F6" lastPageId="5" lastPageNumber="243" pageId="4" pageNumber="242" type="description">
|
||||
<paragraph id="B56436CEFFDAAD62FF19C63BFE9EA325" blockId="4.[189,334,836,862]" box="[189,334,836,862]" pageId="4" pageNumber="242">
|
||||
<heading id="EE2C81A2FFDAAD62FF19C63BFE9EA325" bold="true" box="[189,334,836,862]" fontSize="11" level="3" pageId="4" pageNumber="242" reason="3">
|
||||
<emphasis id="87AFEADCFFDAAD62FF19C63BFE9EA325" bold="true" box="[189,334,836,862]" pageId="4" pageNumber="242">Description</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF4EC60CFE95A3F6" blockId="4.[187,1399,883,1049]" box="[234,325,883,909]" pageId="4" pageNumber="242">
|
||||
<emphasis id="87AFEADCFFDAAD62FF4EC60CFE95A3F6" bold="true" box="[234,325,883,909]" pageId="4" pageNumber="242">Female</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF19C6E9FDC7A462" blockId="4.[187,1399,883,1049]" pageId="4" pageNumber="242">
|
||||
BODY. Length (except ovipositor)
|
||||
<quantity id="72239B2BFFDAAD62FDE1C6E9FD70A3CB" box="[581,672,918,944]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="242" unit="mm" value="3.7">3.7 mm</quantity>
|
||||
, fore wing length
|
||||
<quantity id="72239B2BFFDAAD62FCCAC6E9FC19A3CB" box="[878,969,918,944]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="242" unit="mm" value="2.6">2.6 mm</quantity>
|
||||
, ovipositor length
|
||||
<quantity id="72239B2BFFDAAD62FB3AC6E9FB29A3CB" box="[1182,1273,918,944]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="242" unit="mm" value="4.3">4.3 mm</quantity>
|
||||
. Specimen complete, generally well preserved (left hind femur and tibia somewhat compressed), in clear amber piece with few and small air bubbles, no major occlusions obscuring specimen, anterior part of head covered with thin milky film.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF19C13EFEA3A52A" blockId="4.[188,1400,1089,1361]" pageId="4" pageNumber="242">
|
||||
HEAD. Oval in lateral view, wider than long, eyes oval, occupy most of lateral surface of head (
|
||||
<figureCitation id="2DE02A4BFFDAAD62FF62C11BFEF5A405" box="[198,293,1124,1150]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="4" pageNumber="242">Fig. 2B</figureCitation>
|
||||
); ocelli situated on top of head in approx. equilateral triangle; frons with transverse, wavy striation; postocular carina absent; occipital carina well developed, smooth, slightly incurved dorsally, area around occipital foramen deeply concave; antennae slender, elongate, longer than combined length of head + mesosoma, scapus short, expanded distally, short pedicellus inserted subapically on scapus, antennomeres 4 and 5 longest (
|
||||
<figureCitation id="2DE02A4BFFDAAD62FD95C18EFD5EA570" box="[561,654,1265,1291]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="4" pageNumber="242">Fig. 2B</figureCitation>
|
||||
), antennomeres gradually shortening until apical antennomere, which is approx. 2× as long as penultimate antennomere; mandibles with three teeth each; short, slender palps observed.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD62FF19C006FEC8A727" blockId="4.[187,1401,1400,1884]" pageId="4" pageNumber="242">
|
||||
MESOSOMA. With elongate, extended propectus (
|
||||
<figureCitation id="2DE02A4BFFDAAD62FD4CC007FC94A5E8" box="[744,836,1400,1427]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="4" pageNumber="242">Fig. 2D</figureCitation>
|
||||
); pronotum narrow medially, triangular laterally, with anterior thoracic spiracle in notch in lateral margin; fore femur slender, fore tibia slightly expanded distally, with elongate, curved calcar; mesoscutum strongly arched in lateral view, vertical anterior surface curves smoothly into horizontal surface; horizontal surface flat, level with mesoscutellum, not prominently sculptured; median mesoscutal sulcus distinct, apparently smooth, extends to transscutal articulation; axillae distinct, triangular, continuous medially, separated from mesoscutellum by curved foveate scutoscutellar sulcus (
|
||||
<figureCitation id="2DE02A4BFFDAAD62FD8DC333FD5AA61D" box="[553,650,1612,1638]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="4" pageNumber="242">Fig. 2C</figureCitation>
|
||||
); mesoscutellum flat, smooth with small punctures, triangular posteriorly; mesopleuron smooth, with depression medially, mesopleuro-metapleural sulcus distinct, crenulate; mid femur slightly spindle-shaped, mid tibia slender, with two short apical spurs; hind coxa broad basally, cone-shaped, without carina or foveae along posterior margin; hind femur spindle-shaped, broadened in middle; hind tibia flattened, expanded distally, without fringe of setae, with two apical spurs of subequal length (
|
||||
<figureCitation id="2DE02A4BFFDAAD62FE4EC383FD97A76D" box="[490,583,1788,1814]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="4" pageNumber="242">Fig. 2A</figureCitation>
|
||||
), inner spur approx. to distal tibial width of tibia; hind basitarsus with short erect pegs on inner side, of approx. same length as remaining tarsomeres, tarsal claws simple, slender.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDAAD63FF19C2FBFC6AA13D" blockId="4.[189,1399,1923,1985]" lastBlockId="5.[189,1399,265,326]" lastPageId="5" lastPageNumber="243" pageId="4" pageNumber="242">WINGS. Fore wing hyaline with veins C, R1, Sc+R, M+Cu, A, Rs+M, Rs, M and Cu fully pigmented; Rs+M aligned with M+Cu; medial cell wide distally, rectangular; Rs present between Rs+M and r-rs, closing first submarginal cell; marginal cell narrow, closed in straight line by Rs; hind wings mostly covered by fore wings, apparently with little venation developed.</paragraph>
|
||||
<paragraph id="B56436CEFFDBAD63FF19C419FE9FA24B" blockId="5.[188,1400,358,560]" pageId="5" pageNumber="243">
|
||||
METASOMA. 1.12 × as long as mesosoma, slender, fusiform, without prominent sculpture or pilosity; metasomal segment 2 largest, metasoma widest there, sternum 2 with concavities for accommodating hind coxae; slender, slightly swollen cerci extend from tip of metasoma (
|
||||
<figureCitation id="2DE02A4BFFDBAD63FB88C4D3FB5BA1BC" box="[1068,1163,428,455]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="5" pageNumber="243">Fig. 2C</figureCitation>
|
||||
); sternum 7 tapered apically, folded over medially along longitudinal axis; ovipositor extending posteriorly, elongate, length 1.16 × body length (
|
||||
<figureCitation id="2DE02A4BFFDBAD63FE09C48CFDDDA276" box="[429,525,499,525]" captionStart="Fig" captionStartId="5.[189,232,1819,1845]" captionTargetBox="[189,1398,698,1798]" captionTargetId="figure-211@5.[189,1398,698,1801]" captionTargetPageId="5" captionText="Fig. 2. †Megacoxa brazideci sp. nov., ♀, holotype (CNU-HYM-MA-2016210). A. Habitus, lateral view. B. Head, lateral view, with antennomeres enumerated. C. Metasoma, lateral view. D. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="5" pageNumber="243">Fig. 2A</figureCitation>
|
||||
), ovipositor proper and sheaths slender, tip of 2
|
||||
<superScript id="42AE9B86FFDBAD63FBEEC48DFB8BA27B" attach="left" box="[1098,1115,498,512]" fontSize="6" pageId="5" pageNumber="243">nd</superScript>
|
||||
valvula with 6–7 small dorsal teeth.
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDBAD63FF4EC72FFEFAA211" blockId="5.[189,313,592,653]" box="[234,298,592,618]" pageId="5" pageNumber="243">
|
||||
<emphasis id="87AFEADCFFDBAD63FF4EC72FFEFAA211" bold="true" box="[234,298,592,618]" pageId="5" pageNumber="243">Male</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFDBAD63FF19C70CFEE9A2F6" blockId="5.[189,313,592,653]" box="[189,313,627,653]" pageId="5" pageNumber="243">Unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="E1A46646FFDBAD63FF19C264FC9AA79D" ID-DOI="http://doi.org/10.5281/zenodo.14981657" ID-Zenodo-Dep="14981657" httpUri="https://zenodo.org/record/14981657/files/figure.png" pageId="5" pageNumber="243" startId="5.[189,232,1819,1845]" targetBox="[189,1398,698,1798]" targetPageId="5">
|
||||
<paragraph id="B56436CEFFDBAD63FF19C264FC9AA79D" blockId="5.[188,1399,1819,2022]" pageId="5" pageNumber="243">
|
||||
<emphasis id="87AFEADCFFDBAD63FF19C264FED9A74E" bold="true" box="[189,265,1819,1845]" pageId="5" pageNumber="243">Fig. 2.</emphasis>
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFDBAD63FEBAC263FDDBA74D" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[286,523,1820,1846]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="243" phylum="Arthropoda" rank="species" species="brazideci" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFDBAD63FEBAC263FDDBA74D" box="[286,523,1820,1846]" italics="true" pageId="5" pageNumber="243">Megacoxa brazideci</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFDBAD63FDB4C263FDBCA74D" box="[528,620,1820,1846]" pageId="5" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, ♀, holotype (CNU-HYM-MA-2016210).
|
||||
<emphasis id="87AFEADCFFDBAD63FBFAC264FBA3A74E" bold="true" box="[1118,1139,1819,1845]" pageId="5" pageNumber="243">A</emphasis>
|
||||
. Habitus, lateral view.
|
||||
<emphasis id="87AFEADCFFDBAD63FF18C240FF00A722" bold="true" box="[188,208,1855,1881]" pageId="5" pageNumber="243">B</emphasis>
|
||||
. Head, lateral view, with antennomeres enumerated.
|
||||
<emphasis id="87AFEADCFFDBAD63FC9FC240FC80A722" bold="true" box="[827,848,1855,1881]" pageId="5" pageNumber="243">C</emphasis>
|
||||
. Metasoma, lateral view.
|
||||
<emphasis id="87AFEADCFFDBAD63FBD8C240FB41A722" bold="true" box="[1148,1169,1855,1881]" pageId="5" pageNumber="243">D</emphasis>
|
||||
. Mesosoma, lateral view. Red arrows = transverse striae on frons; blue arrows = hind tibial apical spurs; white arrow = cercus; green arrow = spiracular notch. Abbreviations: ax = axilla; cx3 = hind coxa; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pl3 = metapleuron; S2 = second abdominal sternum; S7 = seventh abdominal sternum; sc2 = mesoscutellum.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="FDC16545FFD8AD60FF19C476FB28A27A" pageId="6" pageNumber="244" type="discussion">
|
||||
<paragraph id="B56436CEFFD8AD60FF19C476FEFFA158" blockId="6.[189,303,265,291]" box="[189,303,265,291]" pageId="6" pageNumber="244">
|
||||
<heading id="EE2C81A2FFD8AD60FF19C476FEFFA158" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="6" pageNumber="244" reason="3">
|
||||
<emphasis id="87AFEADCFFD8AD60FF19C476FEFFA158" bold="true" box="[189,303,265,291]" pageId="6" pageNumber="244">Remarks</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B56436CEFFD8AD60FF19C448FB28A27A" blockId="6.[185,1400,311,514]" pageId="6" pageNumber="244">
|
||||
†
|
||||
<taxonomicName id="72DB4D4DFFD8AD60FF68C447FE6CA12A" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[204,444,311,337]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="brazideci" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD8AD60FF68C447FE6CA12A" box="[204,444,311,337]" italics="true" pageId="6" pageNumber="244">Megacoxa brazideci</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD8AD60FE61C447FDF1A129" box="[453,545,312,338]" pageId="6" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
can be most easily recognized by the transverse striae on the frons and the elongate antennae; the ovipositor is also longer than in most other species of the genus. The frontal sculpture might help facilitate eclosion from a pupal chamber in wood; many other wood-living wasps have prominent sculpture on the head or mesosoma for this purpose (
|
||||
<bibRefCitation id="D14A4B3FFFD8AD60FC2AC4DEFB18A1C0" author="Turrisi G. F. & Vilhelmsen L." box="[910,1224,417,443]" pageId="6" pageNumber="244" pagination="244 - 258" refId="ref8910" refString="Turrisi G. F. & Vilhelmsen L. 2010. Into the wood and back: morphological adaptations to the wood-boring parasitoid lifestyle in adult aulacid wasps (Insecta, Hymenoptera: Aulacidae). Journal of Hymenoptera Research 19 (2): 244-258. Available from https://www.biodiversitylibrary.org/page/35235262#page/46 [accessed 5 Feb. 2025]." type="journal article" year="2010">Turrisi & Vilhelmsen 2010</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="D14A4B3FFFD8AD60FB75C4DEFE81A1A5" author="Vilhelmsen L. & Turrisi G. F." pageId="6" pageNumber="244" pagination="2 - 20" refId="ref8985" refString="Vilhelmsen L. & Turrisi G. F. 2011. Per arborem ad astra: morphological adaptations to exploiting the woody habitat in the early evolution of Hymenoptera. Arthropod Structure & Development 40: 2-20. https://doi.org/10.1016/j.asd.2010.10.001" type="journal article" year="2011">Vilhelmsen & Turrisi 2011</bibRefCitation>
|
||||
). The species resembles †
|
||||
<taxonomicName id="72DB4D4DFFD8AD60FDDBC4BAFCDEA1A5" authorityName="Vilhelmsen & Wu & Wang & Gao & Ren" authorityYear="2025" box="[639,782,452,478]" class="Malacostraca" family="Australomicroprotopidae" genus="Megacoxa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="244" phylum="Arthropoda" rank="species" species="mjoelner" status="sp. nov.">
|
||||
<emphasis id="87AFEADCFFD8AD60FDDBC4BAFCDEA1A5" box="[639,782,452,478]" italics="true" pageId="6" pageNumber="244">M. mjoelner</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9C9C57A7FFD8AD60FCB2C4BBFCA0A1A5" box="[790,880,452,478]" pageId="6" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in having the median sulcus smooth and the hind coxa without carina and foveae posteriorly, but differs in having the axillae contiguous.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue