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<document id="C4F3A3B18BC95EB5DD6250E96564735B" ID-DOI="10.1093/zoolinnean/zlae118" ID-ISSN="0024-4082" ID-Zenodo-Dep="14542407" ID-ZooBank="1C0D753-0F6F-4D0C-BD1D-8D1C6D588F30" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1732850095598" checkinUser="plazi" docAuthor="Pacheco, Leonel I., Teso, Valeria &amp; Pastorino, Guido" docDate="2024" docId="03857E58A10FFF99FC63FAEDFED8FE9A" docLanguage="en" docName="zlae118.pdf" docOrigin="Zoological Journal of the Linnean Society 202" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Lyonsiella tentaculata Pacheco &amp; Teso &amp; Pastorino 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="18" masterDocId="FFBC0620A100FF8BFFA7FFF2FFD9FF9C" masterDocTitle="Use of traditional tools and micro-computed tomography for the taxonomy of carnivorous bivalves from the deep waters of Southwestern Atlantic" masterLastPageNumber="23" masterPageNumber="1" pageNumber="16" updateTime="1734965400749" updateUser="karina">
<mods:mods id="94C3CF52A93DD714FEDC44DBF5135C2B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DF1A2439225CFFD034934DD711EE7DAA">
<mods:title id="B3F0E114D57ACF5F7456C5F4A567750C">Use of traditional tools and micro-computed tomography for the taxonomy of carnivorous bivalves from the deep waters of Southwestern Atlantic</mods:title>
</mods:titleInfo>
<mods:name id="D74B6B0C9235BB7D499A177B15FDCE52" type="personal">
<mods:role id="90605D0CBABD08EF1A50B5DE3F86C603">
<mods:roleTerm id="3597AAD1AF6C93DD0F6F376E0B65B93D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F34A47EB294F39814131F2EF4EC3A96">Pacheco, Leonel I.</mods:namePart>
</mods:name>
<mods:name id="6E91B6BBF0DF9DE2263BA977683C5780" type="personal">
<mods:role id="A41A7AB9E0BD571AC488D95694469742">
<mods:roleTerm id="528508923995CDDA9C7117C5FAD6A19C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19E685DA59C9B9F4BCE2F13BC3C348AF">Teso, Valeria</mods:namePart>
<mods:nameIdentifier id="F9B1996F10CF37CE7937E1249AA7E6D7" type="email">valeriateso@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B4AFC4C168D56290D900A643A8C69E6D" type="personal">
<mods:role id="7689A22E39BC60AEE90C4E1D0F010EB0">
<mods:roleTerm id="8670AF3905702FB4829900D01DF87E52">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6F79891DBCD7D5B25B79A146CBE515B">Pastorino, Guido</mods:namePart>
</mods:name>
<mods:typeOfResource id="8F31F55BCAA7C55B542C4736D8C1DD63">text</mods:typeOfResource>
<mods:relatedItem id="3DA4FE18ACADBC7D7276B5546BC95459" type="host">
<mods:titleInfo id="41AB131E0029CAD294AFD6231F298B53">
<mods:title id="1C44D58DCCE3F82EC572E50F7591D820">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="C57B2D1E86A1372F45092177EF151D98">
<mods:date id="D7E4D44748136C80F77AA6556D5863DE">2024</mods:date>
<mods:detail id="5FD1EB93B821E91EB84AE298B7EE0715" type="volume">
<mods:number id="603FA41B69E24FCD92FD78FC736B9C44">202</mods:number>
</mods:detail>
<mods:extent id="ABA448868DC006FFEFADB93007AA2952" unit="page">
<mods:start id="8314CB07588369E29B4B77802440326F">1</mods:start>
<mods:end id="8A6F862C9B53E4583356C36ACC376C4C">23</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="04850D8AA5E206FDE51BE0CE147097E5">journal article</mods:classification>
<mods:identifier id="F66DC282B24F699CA0DBF2980A5029DD" type="DOI">10.1093/zoolinnean/zlae118</mods:identifier>
<mods:identifier id="C57360BCC879E39AD953F9128A9FE49B" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="1F59DD133CCC3B47A89121552AAF79AE" type="Zenodo-Dep">14542407</mods:identifier>
<mods:identifier id="80E0C109ACC4EE1BBB14E66AB83F9C20" type="ZooBank">1C0D753-0F6F-4D0C-BD1D-8D1C6D588F30</mods:identifier>
</mods:mods>
<treatment id="03857E58A10FFF99FC63FAEDFED8FE9A" LSID="urn:lsid:plazi:treatment:03857E58A10FFF99FC63FAEDFED8FE9A" httpUri="http://treatment.plazi.org/id/03857E58A10FFF99FC63FAEDFED8FE9A" lastPageId="18" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="C3369CC5A10FFF84FC63FAEDFAC0FAA6" box="[964,1305,1311,1338]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B93CF4EA10FFF84FC63FAEDFAC0FAA6" blockId="15.[964,1305,1311,1338]" box="[964,1305,1311,1338]" pageId="15" pageNumber="16">
<heading id="D0DB7822A10FFF84FC63FAEDFAC0FAA6" bold="true" box="[964,1305,1311,1338]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<emphasis id="B958135CA10FFF84FC63FAEDFAC0FAA6" bold="true" box="[964,1305,1311,1338]" pageId="15" pageNumber="16">
<taxonomicName id="4C2CB4CDA10FFF84FC63FAEDFB64FAA6" authority="Pacheco &amp; Teso &amp; Pastorino, 2024" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[964,1213,1311,1338]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="tentaculata" status="sp. nov.">
<emphasis id="B958135CA10FFF84FC63FAEDFB64FAA6" bold="true" box="[964,1213,1311,1338]" italics="true" pageId="15" pageNumber="16">Lyonsiella tentaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BAE27A10FFF84FB65FAD2FAC0FAA6" box="[1218,1305,1312,1338]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A10FFF84FC4AFABDFC0CFA31" pageId="15" pageNumber="16" type="description">
<paragraph id="8B93CF4EA10FFF84FC4AFABDFB29FAF5" blockId="15.[810,1459,1359,1453]" box="[1005,1264,1359,1385]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FC4AFABDFB29FAF5" bold="true" box="[1005,1264,1359,1385]" pageId="15" pageNumber="16">
(
<figureCitation id="1317D3CBA10FFF84FC5FFABDFB19FAF5" box="[1016,1216,1359,1385]" captionStart="Figure 11" captionStartId="16.[129,194,1719,1743]" captionTargetBox="[129,1473,144,1691]" captionTargetId="figure-56@16.[129,1473,144,1691]" captionTargetPageId="16" captionText="Figure 11. Lyonsiella spp. A, B, Lyonsiella tentaculata paratype MACN-IN 44326:right valve (A) and less valve (B). C, Lyonsiella abyssicola lectotype NHMO D 31115: less valve.D, Lyonsiella tentaculata MACN-In 44328:right valve. E, Lyonsiella subquadrata syntype USNM 63238:right valve. F, G, Lyonsiella tentaculata holotype MACN-In 44327 soss parts: ventral view of septum (F) and frontal view of siphons (G). Scale bars:5 mm in A, B; 1 mm in C–G." figureDoi="http://doi.org/10.5281/zenodo.14542429" httpUri="https://zenodo.org/record/14542429/files/figure.png" pageId="15" pageNumber="16">Figs 11A, B, D, F, G</figureCitation>
,
<figureCitation id="1317D3CBA10FFF84FB6DFABDFB3FFAF5" box="[1226,1254,1359,1385]" captionStart="Figure 12" captionStartId="17.[111,176,1116,1140]" captionTargetBox="[144,1424,143,1088]" captionTargetId="figure-438@17.[144,1424,143,1088]" captionTargetPageId="17" captionText="Figure 12. Lyonsiella tentaculata ornamentation through scanning electron microscopy.A, radial lines and spines. B, well-preserved group of spines. C, D, lateral (C) and frontal (D) view of spine. Scale bars: 200 µm in A; 50 µm in B; 10 µm in C; 5 µm in D." figureDoi="http://doi.org/10.5281/zenodo.14542431" httpUri="https://zenodo.org/record/14542431/files/figure.png" pageId="15" pageNumber="16">12</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B93CF4EA10FFF84FC8DFA84FC0CFA31" blockId="15.[810,1459,1359,1453]" pageId="15" pageNumber="16">urn:lsid:zoobank.org:act:1443D27C-D033-4721-B12CE1B7BA98F3E8</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A10FFF84FC8DFA3DFC29F9F5" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B93CF4EA10FFF84FC8DFA3DFB1FF99A" blockId="15.[810,1460,1486,1542]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FC8DFA3DFC18FA7A" box="[810,961,1486,1510]" italics="true" pageId="15" pageNumber="16">Type material:</emphasis>
<materialsCitation id="3B44C513A10FFF84FC77FA3CFA69FA7A" box="[976,1456,1486,1511]" elevation="2212" latitude="-38.023888" location="Mar del Plata" longLatPrecision="19" longitude="-53.850277" municipality="Canyon" pageId="15" pageNumber="16" specimenCode="MACN-In 44327" specimenCount="1" typeStatus="holotype">
<typeStatus id="549771ECA10FFF84FC77FA3CFBEEFA7A" box="[976,1079,1486,1510]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
:
<emphasis id="B958135CA10FFF84FBEFFA3DFB24FA7B" box="[1096,1277,1487,1511]" italics="true" pageId="15" pageNumber="16">
<specimenCode id="DB8A6735A10FFF84FBEFFA3DFB24FA7B" box="[1096,1277,1487,1511]" pageId="15" pageNumber="16">MACN-In 44327</specimenCode>
</emphasis>
,
<specimenCount id="9D2A04C7A10FFF84FAB6FA3DFAE1FA7A" box="[1297,1336,1487,1510]" pageId="15" pageNumber="16">one</specimenCount>
individual
</materialsCitation>
.
<materialsCitation id="3B44C513A10FFF84FC8DFA1CFB1BF99A" box="[810,1218,1518,1542]" elevation="2212" latitude="-38.023888" location="Mar del Plata" longLatPrecision="19" longitude="-53.850277" municipality="Canyon" pageId="15" pageNumber="16" specimenCode="MACN-In 44326" specimenCount="1" typeStatus="paratype">
<typeStatus id="549771ECA10FFF84FC8DFA1CFC4BF99A" box="[810,914,1518,1542]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="DB8A6735A10FFF84FC3DFA1CFB97F99A" box="[922,1102,1518,1542]" pageId="15" pageNumber="16">MACN-In 44326</specimenCode>
,
<specimenCount id="9D2A04C7A10FFF84FBF1FA1CFBA4F999" box="[1110,1149,1518,1541]" pageId="15" pageNumber="16">two</specimenCount>
valves
</materialsCitation>
.
</paragraph>
<paragraph id="8B93CF4EA10FFF84FC8DF9C1FC29F9F5" blockId="15.[810,1459,1585,1642]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FC8DF9C1FC77F9D6" box="[810,942,1586,1610]" italics="true" pageId="15" pageNumber="16">Type locality:</emphasis>
<location id="8EF39995A10FFF84FC14F9C1FBE5F9D6" LSID="urn:lsid:plazi:treatment:03857E58A10FFF99FC63FAEDFED8FE9A:8EF39995A10FFF84FC14F9C1FBE5F9D6" box="[947,1084,1586,1610]" latitude="-38.023888" longLatPrecision="19" longitude="-53.850277" municipality="Canyon" name="Mar del Plata" pageId="15" pageNumber="16">Mar del Plata</location>
submarine Canyon (
<geoCoordinate id="EE18A989A10FFF84FAB5F9C1FA5EF9D6" box="[1298,1415,1585,1611]" degrees="38" direction="south" minutes="01" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="26" value="-38.023888">
38°01
<emphasis id="B958135CA10FFF84FAF6F9C3FA8EF9D6" box="[1361,1367,1585,1610]" italics="true" pageId="15" pageNumber="16">ʹ</emphasis>
26″S
</geoCoordinate>
,
<geoCoordinate id="EE18A989A10FFF84FA36F9C0FC55F9F5" degrees="53" direction="west" minutes="51" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="01" value="-53.850277">
53° 51
<emphasis id="B958135CA10FFF84FCE3F9A2FC93F9F5" box="[836,842,1616,1641]" italics="true" pageId="15" pageNumber="16">ʹ</emphasis>
01″W
</geoCoordinate>
),
<quantity id="4CD462ABA10FFF84FC3BF9A0FC35F9F5" box="[924,1004,1618,1642]" metricMagnitude="3" metricUnit="m" metricValue="2.2119999999999997" pageId="15" pageNumber="16" unit="m" value="2212.0">
<elevation id="0001287DA10FFF84FC3BF9A0FC35F9F5" box="[924,1004,1618,1642]" metricMagnitude="3" metricUnit="m" metricValue="2.2119999999999997" pageId="15" pageNumber="16" unit="m" value="2212.0">2212 m</elevation>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A10FFF84FC8DF979FC23F97E" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8B93CF4EA10FFF84FC8DF979FC23F97E" blockId="15.[810,1459,1675,1762]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FC8DF979FC43F93F" box="[810,922,1675,1699]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
The name
<taxonomicName id="4C2CB4CDA10FFF84FBABF979FBA1F93F" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[1036,1144,1675,1699]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA10FFF84FBABF979FBA1F93F" box="[1036,1144,1675,1699]" italics="true" pageId="15" pageNumber="16">tentaculata</emphasis>
</taxonomicName>
, from the Latin
<emphasis id="B958135CA10FFF84FAB8F979FA52F93F" box="[1311,1419,1675,1699]" italics="true" pageId="15" pageNumber="16">tentaculum</emphasis>
, refers to the numerous siphonal tentacles in comparison to other species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A10FFF9BFB92F8F5FD9DF836" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8B93CF4EA10FFF84FB92F8F5FB71F8BD" blockId="15.[810,1459,1799,1893]" box="[1077,1192,1799,1825]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FB92F8F5FB71F8BD" box="[1077,1192,1799,1825]" italics="true" pageId="15" pageNumber="16">Distribution</emphasis>
</paragraph>
<paragraph id="8B93CF4EA10FFF84FC8DF8DDFB85F8F9" blockId="15.[810,1459,1799,1893]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FC8DF8DDFC70F8DA" box="[810,937,1838,1862]" italics="true" pageId="15" pageNumber="16">New records:</emphasis>
Mar del Plata Submarine Canyon (
<quantity id="4CD462ABA10FFF84FAB2F8DCFA72F8DA" box="[1301,1451,1838,1862]" metricMagnitude="3" metricUnit="m" metricValue="1.8035" metricValueMax="2.2119999999999997" metricValueMin="1.395" pageId="15" pageNumber="16" unit="m" value="1803.5" valueMax="2212.0" valueMin="1395.0">1395–2212 m</quantity>
) and Burdwood Bank (
<quantity id="4CD462ABA10FFF84FBAEF8BFFB89F8F9" box="[1033,1104,1869,1893]" metricMagnitude="2" metricUnit="m" metricValue="6.47" pageId="15" pageNumber="16" unit="m" value="647.0">647 m</quantity>
).
</paragraph>
<paragraph id="8B93CF4EA10FFF84FC8DF87EFBEBF838" blockId="15.[810,1074,1932,1956]" box="[810,1074,1932,1956]" pageId="15" pageNumber="16">
<emphasis id="B958135CA10FFF84FC8DF87EFC7DF838" box="[810,932,1932,1956]" italics="true" pageId="15" pageNumber="16">Bathymetry:</emphasis>
<quantity id="4CD462ABA10FFF84FC0EF87EFBF7F838" box="[937,1070,1932,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.4294999999999998" metricValueMax="2.2119999999999997" metricValueMin="0.647" pageId="15" pageNumber="16" unit="m" value="1429.5" valueMax="2212.0" valueMin="647.0">647–2212 m</quantity>
.
</paragraph>
<caption id="DF539FC6A110FF9BFF26F945FDF3F8BF" ID-DOI="http://doi.org/10.5281/zenodo.14542429" ID-Zenodo-Dep="14542429" httpUri="https://zenodo.org/record/14542429/files/figure.png" pageId="16" pageNumber="17" startId="16.[129,194,1719,1743]" targetBox="[129,1473,144,1691]" targetPageId="16" targetType="figure">
<paragraph id="8B93CF4EA110FF9BFF26F945FDF3F8BF" blockId="16.[129,1452,1719,1827]" pageId="16" pageNumber="17">
<emphasis id="B958135CA110FF9BFF26F945FF3CF953" bold="true" box="[129,229,1719,1743]" pageId="16" pageNumber="17">Figure 11.</emphasis>
<taxonomicName id="4C2CB4CDA110FF9BFF4DF945FE4BF953" box="[234,402,1719,1743]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="undefined-A">
<emphasis id="B958135CA110FF9BFF4DF945FEA2F953" bold="true" box="[234,379,1719,1743]" pageId="16" pageNumber="17">
<emphasis id="B958135CA110FF9BFF4DF945FE94F953" bold="true" box="[234,333,1719,1743]" italics="true" pageId="16" pageNumber="17">Lyonsiella</emphasis>
spp.
</emphasis>
A
</taxonomicName>
, B,
<taxonomicName id="4C2CB4CDA110FF9BFE14F945FD53F953" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[435,650,1719,1743]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA110FF9BFE14F945FD53F953" bold="true" box="[435,650,1719,1743]" italics="true" pageId="16" pageNumber="17">Lyonsiella tentaculata</emphasis>
</taxonomicName>
paratype MACN-IN 44326: right valve (A) and less valve (B). C,
<taxonomicName id="4C2CB4CDA110FF9BFB54F945FA75F953" baseAuthorityName="GO Sars" baseAuthorityYear="1872" box="[1267,1452,1719,1743]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="abyssicola">
<emphasis id="B958135CA110FF9BFB54F945FA75F953" box="[1267,1452,1719,1743]" italics="true" pageId="16" pageNumber="17">Lyonsiella abyssicola</emphasis>
</taxonomicName>
lectotype NHMO D 31115: less valve. D,
<taxonomicName id="4C2CB4CDA110FF9BFDA0F921FD07F977" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[519,734,1747,1771]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA110FF9BFDA0F921FD07F977" bold="true" box="[519,734,1747,1771]" italics="true" pageId="16" pageNumber="17">Lyonsiella tentaculata</emphasis>
</taxonomicName>
MACN-In 44328: right valve. E,
<taxonomicName id="4C2CB4CDA110FF9BFBB0F921FB3EF977" baseAuthorityName="Jeffreys" baseAuthorityYear="1882" box="[1047,1255,1747,1771]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="subquadrata">
<emphasis id="B958135CA110FF9BFBB0F921FB3EF977" box="[1047,1255,1747,1771]" italics="true" pageId="16" pageNumber="17">Lyonsiella subquadrata</emphasis>
</taxonomicName>
syntype USNM 63238: right valve. F, G,
<taxonomicName id="4C2CB4CDA110FF9BFEC2F91DFDE5F89B" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[357,572,1775,1799]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA110FF9BFEC2F91DFDE5F89B" bold="true" box="[357,572,1775,1799]" italics="true" pageId="16" pageNumber="17">Lyonsiella tentaculata</emphasis>
</taxonomicName>
holotype
<emphasis id="B958135CA110FF9BFD3BF91DFCE5F89B" box="[668,828,1775,1799]" italics="true" pageId="16" pageNumber="17">MACN-In 44327</emphasis>
soss parts: ventral view of septum (F) and frontal view of siphons (G). Scale bars: 5 mm in A, B; 1 mm in C–G.
</paragraph>
</caption>
<paragraph id="8B93CF4EA110FF9BFF26F8A1FD9DF836" blockId="16.[129,778,1874,1962]" pageId="16" pageNumber="17">
<emphasis id="B958135CA110FF9BFF26F8A1FE98F8F7" box="[129,321,1875,1899]" italics="true" pageId="16" pageNumber="17">Material examined:</emphasis>
MACN-In 44328 (
<geoCoordinate id="EE18A989A110FF9BFDAFF8A6FDA7F8F7" box="[520,638,1874,1900]" degrees="37" direction="south" minutes="49" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="40" value="-37.827778">
37°49
<emphasis id="B958135CA110FF9BFDE0F8A0FD94F8F7" box="[583,589,1874,1899]" italics="true" pageId="16" pageNumber="17">ʹ</emphasis>
40″S
</geoCoordinate>
,
<geoCoordinate id="EE18A989A110FF9BFD2FF8A1FCDFF8F7" box="[648,774,1874,1899]" degrees="54" direction="west" minutes="07" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="56" value="-54.13222">
54°07
<emphasis id="B958135CA110FF9BFD63F8A0FD13F8F7" box="[708,714,1874,1899]" italics="true" pageId="16" pageNumber="17">ʹ</emphasis>
56″W
</geoCoordinate>
,
<quantity id="4CD462ABA110FF9BFF26F881FF07F817" box="[129,222,1907,1931]" metricMagnitude="3" metricUnit="m" metricValue="1.395" pageId="16" pageNumber="17" unit="m" value="1395.0">1395 m</quantity>
) one individual; MACN-In 44326 (
<geoCoordinate id="EE18A989A110FF9BFD37F881FCDCF816" box="[656,773,1905,1931]" degrees="54" direction="south" minutes="36" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="14" value="-54.603886">
54°36
<emphasis id="B958135CA110FF9BFD68F883FD0CF816" box="[719,725,1905,1930]" italics="true" pageId="16" pageNumber="17">ʹ</emphasis>
14″S
</geoCoordinate>
,
<geoCoordinate id="EE18A989A110FF9BFF26F860FF27F836" box="[129,254,1937,1962]" degrees="62" direction="west" minutes="50" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="55" value="-62.84861">
62°50
<emphasis id="B958135CA110FF9BFF1AF863FF1AF836" box="[189,195,1937,1962]" italics="true" pageId="16" pageNumber="17">ʹ</emphasis>
55″W
</geoCoordinate>
,
<quantity id="4CD462ABA110FF9BFEA0F860FE90F836" box="[263,329,1938,1962]" metricMagnitude="2" metricUnit="m" metricValue="6.47" pageId="16" pageNumber="17" unit="m" value="647.0">647 m</quantity>
) two valves (
<typeStatus id="549771ECA110FF9BFE6AF860FDE0F836" box="[461,569,1938,1962]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A110FF9AFBE8F8A6FEF8FB6B" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B93CF4EA110FF9BFBE8F8A6FB75F8F1" blockId="16.[825,1474,1876,1969]" box="[1103,1196,1876,1901]" pageId="16" pageNumber="17">
<emphasis id="B958135CA110FF9BFBE8F8A6FB75F8F1" box="[1103,1196,1876,1901]" italics="true" pageId="16" pageNumber="17">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B93CF4EA110FF9AFC9EF88BFEF8FB6B" blockId="16.[825,1474,1876,1969]" lastBlockId="17.[113,762,1216,1271]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Shell ≤
<quantity id="4CD462ABA110FF9BFC27F88BFC1FF80D" box="[896,966,1913,1937]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="17" unit="mm" value="14.0">14 mm</quantity>
long, ornamented by ≤50 radial lines spaced by radially oriented groups of pyramidal spines. Siphons surrounded by 25 siphonal tentacles, 12 associated with inhalant siphon and 13 with exhalant.
</paragraph>
</subSubSection>
<caption id="DF539FC6A111FF9AFFC8FBAEFB46FB0C" ID-DOI="http://doi.org/10.5281/zenodo.14542431" ID-Zenodo-Dep="14542431" httpUri="https://zenodo.org/record/14542431/files/figure.png" pageId="17" pageNumber="18" startId="17.[111,176,1116,1140]" targetBox="[144,1424,143,1088]" targetPageId="17" targetType="figure">
<paragraph id="8B93CF4EA111FF9AFFC8FBAEFB46FB0C" blockId="17.[111,1443,1116,1169]" pageId="17" pageNumber="18">
<emphasis id="B958135CA111FF9AFFC8FBAEFE76FBE8" bold="true" box="[111,431,1116,1140]" pageId="17" pageNumber="18">
Figure 12.
<taxonomicName id="4C2CB4CDA111FF9AFF7FFBAEFE76FBE8" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[216,431,1116,1140]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA111FF9AFF7FFBAEFE76FBE8" bold="true" box="[216,431,1116,1140]" italics="true" pageId="17" pageNumber="18">Lyonsiella tentaculata</emphasis>
</taxonomicName>
</emphasis>
ornamentation through scanning electron microscopy. A, radial lines and spines. B, well-preserved group of spines. C, D, lateral (C) and frontal (D) view of spine. Scale bars: 200 µm in A; 50 µm in B; 10 µm in C; 5 µm in D.
</paragraph>
</caption>
<subSubSection id="C3369CC5A111FF9AFE27FAEAFB27FB6B" pageId="17" pageNumber="18" type="description">
<paragraph id="8B93CF4EA111FF9AFE27FAEAFE35FAAD" blockId="17.[113,763,1304,1992]" box="[384,492,1304,1329]" pageId="17" pageNumber="18">
<emphasis id="B958135CA111FF9AFE27FAEAFE35FAAD" box="[384,492,1304,1329]" italics="true" pageId="17" pageNumber="18">Description</emphasis>
</paragraph>
<paragraph id="8B93CF4EA111FF9AFFD6FACFFB27FB6B" blockId="17.[113,763,1304,1992]" lastBlockId="17.[810,1458,1216,1272]" pageId="17" pageNumber="18">
Shell medium in size (≤
<quantity id="4CD462ABA111FF9AFE26FACFFE17FAC9" box="[385,462,1341,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="18" unit="mm" value="14.0">14 mm</quantity>
L,
<quantity id="4CD462ABA111FF9AFE5EFACFFD9DFAC9" box="[505,580,1341,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="mm" value="11.0">11 mm</quantity>
H,
<quantity id="4CD462ABA111FF9AFDD3FACFFD1CFAC9" box="[628,709,1341,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="17" pageNumber="18" unit="mm" value="4.6">4.6 mm</quantity>
W), fragile, somewhat translucid, oval-trapezoidal, inequilateral. Inequivalve; left valve more rectangular, right valve slightly larger and with dorsal and ventral margins projecting beyond the left valve. Surface covered by ≤50 radial lines spaced by radially oriented groups of four to eight pyramidal spines, usually covered by debris. Umbo prosogyrus, displaced anteriorly. Anterodorsal margin diagonally inclined, lunule weak, anterior to resilium. Anterior margin rounded, slightly extended beyond umbo. Posterodorsal margin somewhat straight, almost parallel to ventral margin; both margins connect with nearly vertical posterior margin through rounded corners. Hinge edentulous, with elongated lithodesma. Interior iridescent, with radial lines corresponding to external striae. Adductor muscle scars slightly marked, anterior more than posterior. Palial line noticeable. Ctenidium composed of a reduced outer demibranch and an inner demibranch with ~50 and ~60 branchial filaments, respectively. Inhalant siphon with 12 inner arborescent tentacles on the edge of the aperture and 10 outer small tentacles surrounding it, plus two small dorsal tentacles. Exhalant siphon surrounded by 13 outer small tentacles (seven larger intercalated by six smaller). Byssus present.
</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A111FF9AFBE3FAE5FB41FAAD" box="[1092,1176,1303,1329]" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8B93CF4EA111FF9AFBE3FAE5FB41FAAD" blockId="17.[810,1459,1303,1992]" box="[1092,1176,1303,1329]" pageId="17" pageNumber="18">
<emphasis id="B958135CA111FF9AFBE3FAE5FB41FAAD" box="[1092,1176,1303,1329]" italics="true" pageId="17" pageNumber="18">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3369CC5A111FF99FC8DFACFFED8FE9A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8B93CF4EA111FF9AFC8DFACFFBCAF890" blockId="17.[810,1459,1303,1992]" pageId="17" pageNumber="18">
<taxonomicName id="4C2CB4CDA111FF9AFC8DFACFFC26FAC9" ID-CoL="9G5B5" baseAuthorityName="GO Sars" baseAuthorityYear="1872" box="[810,1023,1341,1365]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="abyssicola">
<emphasis id="B958135CA111FF9AFC8DFACFFC26FAC9" box="[810,1023,1341,1365]" italics="true" pageId="17" pageNumber="18">Lyonsiella abyssicola</emphasis>
</taxonomicName>
is a comparable species; however, it has 16 tentacles surrounding both siphons, whereas
<taxonomicName id="4C2CB4CDA111FF9AFA80FAAFFA6AFAE9" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[1319,1459,1373,1397]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA111FF9AFA80FAAFFA6AFAE9" box="[1319,1459,1373,1397]" italics="true" pageId="17" pageNumber="18">L. tentaculata</emphasis>
</taxonomicName>
has 25. The number of tentacles is the same both in the largest individual (
<emphasis id="B958135CA111FF9AFC0CFA6EFBB9FA2F" box="[939,1120,1435,1459]" italics="true" pageId="17" pageNumber="18">MACN-In 44327</emphasis>
,
<quantity id="4CD462ABA111FF9AFBC8FA69FB63FA2F" box="[1135,1210,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="18" unit="mm" value="14.0">14 mm</quantity>
L) and in the smallest individual (MACN-In 44328,
<quantity id="4CD462ABA111FF9AFBDFFA49FB6FFA4E" box="[1144,1206,1467,1490]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="mm" value="7.0">7 mm</quantity>
L), with a similar size to
<taxonomicName id="4C2CB4CDA111FF9AFCEEFA29FC14FA6E" ID-CoL="9G5B5" baseAuthorityName="GO Sars" baseAuthorityYear="1872" box="[841,973,1498,1522]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="abyssicola">
<emphasis id="B958135CA111FF9AFCEEFA29FC14FA6E" box="[841,973,1498,1522]" italics="true" pageId="17" pageNumber="18">L. abyssicola</emphasis>
</taxonomicName>
, hence it can be discarded as a variation attributable to size. Additionally, the new species reaches a larger size and exhibits more radial lines than
<taxonomicName id="4C2CB4CDA111FF9AFA8EF9EBFA74F9AD" ID-CoL="9G5B5" baseAuthorityName="GO Sars" baseAuthorityYear="1872" box="[1321,1453,1561,1585]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="abyssicola">
<emphasis id="B958135CA111FF9AFA8EF9EBFA74F9AD" box="[1321,1453,1561,1585]" italics="true" pageId="17" pageNumber="18">L. abyssicola</emphasis>
</taxonomicName>
, (≤
<quantity id="4CD462ABA111FF9AFCE4F9CAFC49F9D3" box="[835,912,1592,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="18" unit="mm" value="14.0">14 mm</quantity>
and 50 lines vs.
<quantity id="4CD462ABA111FF9AFB9CF9CAFBA1F9D3" box="[1083,1144,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
and 20 lines) (
<figureCitation id="1317D3CBA111FF9AFAB0F9CAFA55F9D3" box="[1303,1420,1592,1616]" captionStart="Figure 11" captionStartId="16.[129,194,1719,1743]" captionTargetBox="[129,1473,144,1691]" captionTargetId="figure-56@16.[129,1473,144,1691]" captionTargetPageId="16" captionText="Figure 11. Lyonsiella spp. A, B, Lyonsiella tentaculata paratype MACN-IN 44326:right valve (A) and less valve (B). C, Lyonsiella abyssicola lectotype NHMO D 31115: less valve.D, Lyonsiella tentaculata MACN-In 44328:right valve. E, Lyonsiella subquadrata syntype USNM 63238:right valve. F, G, Lyonsiella tentaculata holotype MACN-In 44327 soss parts: ventral view of septum (F) and frontal view of siphons (G). Scale bars:5 mm in A, B; 1 mm in C–G." figureDoi="http://doi.org/10.5281/zenodo.14542429" httpUri="https://zenodo.org/record/14542429/files/figure.png" pageId="17" pageNumber="18">Fig. 11A, B</figureCitation>
). A clear character visible by SEM is the shape of the microscopic spines.
<taxonomicName id="4C2CB4CDA111FF9AFCDBF985FB84F913" authorityName="Pacheco &amp; Teso &amp; Pastorino" authorityYear="2024" box="[892,1117,1655,1679]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="B958135CA111FF9AFCDBF985FB84F913" box="[892,1117,1655,1679]" italics="true" pageId="17" pageNumber="18">Lyonsiella tentaculata</emphasis>
</taxonomicName>
has pyramidal and acuminated spines, whereas
<taxonomicName id="4C2CB4CDA111FF9AFC7CF965FB8CF932" ID-CoL="9G5B5" baseAuthorityName="GO Sars" baseAuthorityYear="1872" box="[987,1109,1686,1710]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="abyssicola">
<emphasis id="B958135CA111FF9AFC7CF965FB8CF932" box="[987,1109,1686,1710]" italics="true" pageId="17" pageNumber="18">L abyssicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2CB4CDA111FF9AFBC5F964FCB9F951" authority="Allen &amp; Turner, 1974" authorityName="Allen &amp; Turner" authorityYear="1974" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="frieli">
<emphasis id="B958135CA111FF9AFBC5F964FB26F932" box="[1122,1279,1686,1710]" italics="true" pageId="17" pageNumber="18">Lyonsiella frieli</emphasis>
<bibRefCitation id="EFBDB2BFA111FF9AFAA0F964FCB9F951" author="Allen JA &amp; Turner JF" pageId="17" pageNumber="18" pagination="401 - 532" refId="ref15118" refString="Allen JA, Turner JF. On the functional morphology of the family Verticordiidae (Bivalvia) with descriptions of new species from the abyssal Atlantic. Philosophical Transactions of the Royal Society of London B, Biological Sciences 1974; 268: 401 - 532." type="journal article" year="1974">Allen &amp; Turner, 1974</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2CB4CDA111FF9AFC3EF947FAC5F951" ID-CoL="5TWNB" authority="(Jeffreys, 1882)" authorityName="Machado et al." authorityYear="2019" baseAuthorityName="Jeffreys" baseAuthorityYear="1882" box="[921,1308,1717,1741]" class="Bivalvia" family="Lyonsiellidae" genus="Allogramma" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="formosa">
<emphasis id="B958135CA111FF9AFC3EF947FAC5F951" box="[921,1308,1717,1741]" italics="true" pageId="17" pageNumber="18">Allogramma formosa (Jeffreys, 1882)</emphasis>
</taxonomicName>
, have thinner spines with stellated termination (
<bibRefCitation id="EFBDB2BFA111FF9AFB67F927FCB2F890" author="Oliveira CDdC &amp; Absalao RS" pageId="17" pageNumber="18" pagination="305 - 16" refId="ref17314" refString="Oliveira CDdC, Absalao RS. Review of the Septibranchia (Mollusca: Pelecypoda) fromthedeepseaofCamposBasin, Brazil: familyLyonsiellidae, with description of a new species. Scientia Marina 2010 a; 74: 305 - 16." type="journal article" year="2010">Oliveira and Absalão 2010a</bibRefCitation>
) (
<figureCitation id="1317D3CBA111FF9AFC2FF906FBDBF897" box="[904,1026,1780,1804]" captionStart="Figure 12" captionStartId="17.[111,176,1116,1140]" captionTargetBox="[144,1424,143,1088]" captionTargetId="figure-438@17.[144,1424,143,1088]" captionTargetPageId="17" captionText="Figure 12. Lyonsiella tentaculata ornamentation through scanning electron microscopy.A, radial lines and spines. B, well-preserved group of spines. C, D, lateral (C) and frontal (D) view of spine. Scale bars: 200 µm in A; 50 µm in B; 10 µm in C; 5 µm in D." figureDoi="http://doi.org/10.5281/zenodo.14542431" httpUri="https://zenodo.org/record/14542431/files/figure.png" pageId="17" pageNumber="18">Fig. 12C, D</figureCitation>
).
</paragraph>
<paragraph id="8B93CF4EA111FF99FCE2F8E1FED8FE9A" blockId="17.[810,1459,1303,1992]" lastBlockId="18.[129,778,144,262]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Other comparable
<taxonomicName id="4C2CB4CDA111FF9AFB86F8E1FB5AF8B7" box="[1057,1155,1811,1835]" pageId="17" pageNumber="18">
<emphasis id="B958135CA111FF9AFB86F8E1FB5AF8B7" box="[1057,1155,1811,1835]" italics="true" pageId="17" pageNumber="18">Lyonsiella</emphasis>
</taxonomicName>
species include
<taxonomicName id="4C2CB4CDA111FF9AFAF6F8E1FB81F8D7" ID-CoL="3WSXL" authority="(Jeffreys, 1882)" baseAuthorityName="Jeffreys" baseAuthorityYear="1882" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="subquadrata">
<emphasis id="B958135CA111FF9AFAF6F8E1FC7DF8D7" italics="true" pageId="17" pageNumber="18">Lyonsiella subquadrata</emphasis>
(Jeffreys, 1882)
</taxonomicName>
, distinguished by a wider, more prolonged, and ovate anterior margin (
<figureCitation id="1317D3CBA111FF9AFB7FF8A0FAEBF8F6" box="[1240,1330,1874,1898]" captionStart="Figure 11" captionStartId="16.[129,194,1719,1743]" captionTargetBox="[129,1473,144,1691]" captionTargetId="figure-56@16.[129,1473,144,1691]" captionTargetPageId="16" captionText="Figure 11. Lyonsiella spp. A, B, Lyonsiella tentaculata paratype MACN-IN 44326:right valve (A) and less valve (B). C, Lyonsiella abyssicola lectotype NHMO D 31115: less valve.D, Lyonsiella tentaculata MACN-In 44328:right valve. E, Lyonsiella subquadrata syntype USNM 63238:right valve. F, G, Lyonsiella tentaculata holotype MACN-In 44327 soss parts: ventral view of septum (F) and frontal view of siphons (G). Scale bars:5 mm in A, B; 1 mm in C–G." figureDoi="http://doi.org/10.5281/zenodo.14542429" httpUri="https://zenodo.org/record/14542429/files/figure.png" pageId="17" pageNumber="18">Fig. 11E</figureCitation>
);
<taxonomicName id="4C2CB4CDA111FF9AFAF6F8A0FBBDF815" ID-CoL="3WSX8" authority="Allen &amp; Turner, 1974" authorityName="Allen &amp; Turner" authorityYear="1974" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="perplexa">
<emphasis id="B958135CA111FF9AFAF6F8A0FCA7F815" italics="true" pageId="17" pageNumber="18">Lyonsiella perplexa</emphasis>
<bibRefCitation id="EFBDB2BFA111FF9AFC23F883FBBDF815" author="Allen JA &amp; Turner JF" box="[900,1124,1905,1929]" pageId="17" pageNumber="18" pagination="401 - 532" refId="ref15118" refString="Allen JA, Turner JF. On the functional morphology of the family Verticordiidae (Bivalvia) with descriptions of new species from the abyssal Atlantic. Philosophical Transactions of the Royal Society of London B, Biological Sciences 1974; 268: 401 - 532." type="journal article" year="1974">Allen &amp; Turner, 1974</bibRefCitation>
</taxonomicName>
, with an angular anterior margin and a lower number of tentacles (~15);
<taxonomicName id="4C2CB4CDA111FF9AFB65F863FAB0F835" ID-CoL="3WSWN" baseAuthorityName="Allen and Turner" baseAuthorityYear="1974" box="[1218,1385,1937,1961]" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="fragilis">
<emphasis id="B958135CA111FF9AFB65F863FAB0F835" box="[1218,1385,1937,1961]" italics="true" pageId="17" pageNumber="18">Lyonsiella fragilis</emphasis>
</taxonomicName>
, with a much more horizontal anterodorsal margin, a more prolonged anterior margin, and fewer radial lines (8–18); and
<taxonomicName id="4C2CB4CDA112FF99FD00FF62FE0CFF5B" ID-CoL="3WSXB" authority="Oliveira &amp; Absalao, 2010" authorityName="Oliveira &amp; Absalao" authorityYear="2010" class="Bivalvia" family="Lyonsiellidae" genus="Lyonsiella" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="pipoca">
<emphasis id="B958135CA112FF99FD00FF62FF18FF5B" italics="true" pageId="18" pageNumber="19">Lyonsiella pipoca</emphasis>
<bibRefCitation id="EFBDB2BFA112FF99FF6EFF5DFE0CFF5B" author="Oliveira CDdC &amp; Absalao RS" box="[201,469,175,199]" pageId="18" pageNumber="19" pagination="305 - 16" refId="ref17314" refString="Oliveira CDdC, Absalao RS. Review of the Septibranchia (Mollusca: Pelecypoda) fromthedeepseaofCamposBasin, Brazil: familyLyonsiellidae, with description of a new species. Scientia Marina 2010 a; 74: 305 - 16." type="journal article" year="2010">Oliveira &amp; Absalão, 2010</bibRefCitation>
</taxonomicName>
, with short and wider spines oriented irregularly, not radially, and with a few isolated delicate ‘projections’.
</paragraph>
</subSubSection>
</treatment>
</document>