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Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Lebia +( +Metalebia +) +ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 34 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.34A–B +), a male, in NMNH, labeled: “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +1100m + + +22-31 October 1988 + +W. E. Steiner +”/ “ +Malaise trap +in small clearing, montane rain forest”/ “ +HOLOTYPE + +Lebia +( +Metalebia +) +ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(only 1): a female (in +CAS +) labeled “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +900m + + +23-28 February 1990 + +W. E. Steiner +”/ “ +Malaise trap +in small clearing, montane rain forest”/ “ +PARATYPE + +Lebia +( +Metalebia +) +ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size slightly small for genus, SBL of male = +4.3 mm +, of female = +4.2 mm +. Members of this species ( +Fig. 34A +) key to subgenus + +Metalebia +Jeannel (1949) + +and the + +madagascariensis + +group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody (head plus pronotum) has no metallic reflection and the elytra are uniform in color (but not necessarily in reflection); and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the + +madagascariensis + +group, + +L. ranomafanae + +members can be distinguished from those of all species except + +L. alluaudana + +, + +L. apicoviolacea + +sp. nov. +, + +L. mirana + +and + +L. tanala + +by the presence of elytral metallic reflection (metallic reflection absent from the elytra of members of the remaining seven species). They are distinguished from members of + +L. mirana + +by their reddish labrum (black in + +L. mirana + +members) and from those of + +L. alluaudana + +by their small size and relatively short and broad elytra, with ratio of elytra length to width = 1.3 (size larger, SBL of males = 4.6–5.0 mm, of females 4.6– +5.3 mm +, and elytra relatively longer, with ratio EL/EW = 1.4, in + +L. alluaudana + +members). They are distinguished from members of + +L. tanala + +in having very dark and relatively faint elytral metallic reflection, pronotum with anteri- or angles indistinct, smoothly rounded from apical to lateral margin and not at all projected anteriorly beyond apical margin, lateral margins nearly parallel in posterior two-thirds and not or only very slightly sinuate anterior to obtusely angulate hind angles (elytral metallic reflection more distinct, pronotum with anterior angles broadly round but slightly projected anteriorly beyond apical margin, lateral margins distinctly sinuate anterior to sharp, rectangular hind angles slightly projected laterally in members of + +L. tanala + +). Finally, they can be distinguished from members of + +L. apicoviolacea + +by slightly larger size and the faint, concolorous dark blue-green elytral metallic reflection and (size slightly smaller, SBL of male = +3.8 mm +, of female = 4.0 mm, and elytra with distinct tricolored metallic reflection, green basally and on medial two-thirds, brassy on lateral one-third and violet in apical one-fifth, in + +L. apicoviolacea + +members. Shape of the median lobe of the male aedeagus ( +Fig. 34C–D +) is also diagnostic, with the shaft broadest at basal one-third, slightly and evenly arcuate on ventral margin in lateral view ( +Fig. 34C +) and apex parallel-side basally and broadly rounded apically in dorsal view ( +Fig. 34D +). + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Both specimens of the +type +series were collected in malaise traps in secondary montane rainforest in the Talatakely area at elevations ranging from +900 to 1100 m +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573042FFADFF8EFA33BF1E0823.xml b/data/0A/3E/26/0A3E26573042FFADFF8EFA33BF1E0823.xml new file mode 100644 index 00000000000..d5d162ff94a --- /dev/null +++ b/data/0A/3E/26/0A3E26573042FFADFF8EFA33BF1E0823.xml @@ -0,0 +1,284 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Eunostus minimus +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 36–37 + + + +Eunostus +n. sp. +1 + +; Rainio 2009: 31, Rainio 2012: 72 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 36A–B +), a male, in CAS, labeled: “CASENT 8069195”/ “ +MADAGASCAR +: +Fianarantsoa +Parc National de Ranomafana +, +Vatoharanana River +, + +4.1 km +231° SW Ranomafana + +elev + +1100m + + +27-31 Mar 2003 + +”/ “ +21°17’24”S +047°26’00”E +California Acad. of Sciences +coll. +Fisher +, Griswold et al. sifted litter, montane rainforest collection code: BLF8400”/ “ +HOLOTYPE + +Eunostus minimus +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 2): +1 male +(in +MNHN +) labeled “CASENT 8069129”/ “ +MADAGASCAR +: +Fianarantsoa +, +Ranomafana +JIRAMA water works, +21°14.91’S +, +047°27.13’E +”/ “ + +21-24 December 2001 + +collector: + +R. +Harin’Hala California Acad + +of Sciences malaise trap near river elev. + +690 m + +, MA-02-09D-08”; +1 female +(in +CAS +) labeled “CASENT 1051146”/ “ +MADAGASCAR +Ranomafana +NP, Vatoharana trail S 600 + +24.8.2004 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”. Both +paratypes +also bear the following label: “ +PARATYPE + +Eunostus minimus +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + +FIGURE +36. Digital images of holotype male of + +Eunostus minimus +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +minimus + +, is the Latin adjective meaning smallest, a reference to fact that the known specimens of this species are smaller than those of any of the other known species that we have examined. + + + +RECOGNITION + +.— Small for genus, SBL of male = +5.5 mm +, of female 5.0 +– +5. +1 mm +. Members of this species ( +Fig. 36A +) do not key out in Jeannel’s (1949) key for + +Eunostus +species. + +They are much smaller and shinier than members of any other described species. Their heads are slightly narrower, with the tempora more evenly rounded to the back of the head, their antennal scapes shorter and thicker than in other species, thickest slightly distal to mid-length. Lateral margins of the pronotum are smoothly rounded in anterior half with a moderately short and deep subbasal sinuation anterior to slightly obtuse to rectangular hind angles. The elytral setae are generally sparser than in other species and arranged in 2 more or less distinct longitudinal rows on each interval. Male ( +Fig. 37A +) and female ( +Fig. 37B +) hind femora are sexual dimorphic, each unique among Malagasy species for each sex. Median lobe of male genitalia +Figs. 37C–D +markedly different in shape from those of the other species for which males are known (see Jeannel 1949, Fig. 513d–e). + + + +FIGURE +37. Digital images of + +Eunostus minimus +Kavanaugh & Rainio + +sp. nov. +A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Specimens of the +type +series were collected in both primary montane rainforest in the Vatoharanana area (at elevations ranging from of +1100 to 1200 m +) and in secondary montane rainforest +1 km +north of Ranomafana village at an elevation of + +690 m +. + +The specimens were collected in a malaise trap and sifted from forest litter, respectively, at the primary forest site and by hand at the secondary forest site. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573043FFABFF8EFF49BE9D098D.xml b/data/0A/3E/26/0A3E26573043FFABFF8EFF49BE9D098D.xml new file mode 100644 index 00000000000..50fd8bfcbd0 --- /dev/null +++ b/data/0A/3E/26/0A3E26573043FFABFF8EFF49BE9D098D.xml @@ -0,0 +1,312 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Lebia +( +Metalebia +) +apicoviolacea +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 35 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 35A–B +), a male, in CAS, labeled: “ +MADAGASCAR +: + +Fianaran. Pr. +Ranomafana +N.P. + +HQ. area +21°15’24”S +47°25’15E + +25NOV1994 + +, at night +M.A. Ivie +& +D.A. Pollack +”/ “ +HOLOTYPE + +Lebia +( +Metalebia +) +apicoviolacea +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(only 1): a female (in +NMHN +) labeled “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +1100 m + + +6-21 October 1988 + +W. E. Steiner +”/ “At black light in montane rain forest”/ “ +PARATYPE + +Lebia +( +Metalebia +) +apicoviolacea +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +apicoviolacea +, + +is an adjective derived from the Latin words, +apex +, meaning tip, and +violaceus +, meaning violet-colored. The name refers to the violet or purple metallic reflection of elytral apices, which contrasts distinctly with the green or bronze metallic reflection of the remainder of the elytra in members of this species. + + + +FIGURE +35. Digital images of holotype male of + +Lebia +( +Metalebia +) +apicoviolacea +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm. + + + + +RECOGNITION + +.— Size moderately small for genus, SBL of male = +3.8 mm +, of female = 4.0 mm. Members of this species ( +Fig. 35A +) key to subgenus + +Metalebia +Jeannel (1949) + +and the + +madagascariensis + +group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody has no metallic reflection and the elytra are uniform in color (but not necessarily in reflection); and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the + +madagascariensis + +group, + +L. apicoviolacea + +members can be distinguished from those of all species except + +L. alluaudana + +, + +L. mirana +Alluaud + +, + +L. ranomafanae + +and + +L. tanala + +by the presence of elytral metallic reflection (metallic reflection absent from the elytra of members of the remaining seven species). They are distinguished from members of + +L. mirana + +by their reddish labrum (black in + +L. mirana + +members) and from those of + +L. alluaudana + +by their small size and relatively short and broad elytra, with ratio of elytra length to width = 1.2–1.3 (size larger, SBL of males = 4.6–5.0 mm, of females 4.6– +5.3 mm +, and elytra relatively longer, with ratio of elytra length to width = 1.4, in + +L. alluaudana + +members). They are distinguished from members of + +L. tanala + +in having the pronotum with lateral margins slightly sinuate anterior to slightly obtuse hind angles (lateral margins distinctly sinuate anterior to sharp, rectangular hind angles slightly projected laterally in members of + +L. tanala + +) and from members of both + +L. ranomafanae + +and + +L. tanala + +in having elytra with distinct tricolored metallic reflection, green basally and on medial two-thirds, brassy on lateral one-third and violet in apical one-fifth (elytral metallic reflection uniformly faint or distinct metallic green or blue in members of + +L. ranomafanae + +and + +L. tanala + +). Shape of the median lobe of the male aedeagus ( +Fig. 35C–D +) is also diagnostic, with the shaft very slightly arcuate, dorsal and ventral margins more or less parallel in basal two-thirds in lateral view ( +Fig. 35C +) and apex moderate in length and relatively broader in both lateral and dorsal ( +Fig. 35D +) views than in males of other species of this species group (compare with Jeannel 1949, Fig. 440a-l). + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Both specimens of the +type +series were collected in secondary montane rainforest in the Talatakely area at elevations ranging from +900 to 1100 m +. Both were collect- ed at night, one by hand from vegetation, and the other attracted to ultraviolet light. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573044FFADFE49FA1EBE7C08E2.xml b/data/0A/3E/26/0A3E26573044FFADFE49FA1EBE7C08E2.xml new file mode 100644 index 00000000000..250f2f2d334 --- /dev/null +++ b/data/0A/3E/26/0A3E26573044FFADFE49FA1EBE7C08E2.xml @@ -0,0 +1,72 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + +Genus + +Erephognathus +Alluaud 1932 + + + + + +This genus is endemic to +Madagascar +and includes only two previously described species + + + + +( +Lorenz 2005 +). Jeannel (1949) provided a key to those species. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573044FFAFFF8EF9B3BE7F0E68.xml b/data/0A/3E/26/0A3E26573044FFAFFF8EF9B3BE7F0E68.xml new file mode 100644 index 00000000000..c89fc41266d --- /dev/null +++ b/data/0A/3E/26/0A3E26573044FFAFFF8EF9B3BE7F0E68.xml @@ -0,0 +1,288 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Erephognathus ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 38 + + + + + +TYPE +MATERIAL + +.— +Holotype +( +Figs.38A–B +), a female, in CAS, labeled: “CASENT 8068531”/ + + +“ + +MADAGASCAR +: +Province +Fianarantsoa +, +Parc National Ranomafana +, +Vohiparara +, at broken bridge, el + +1110 m + + +8-15 November 2001 + +”/ “ +21°13.57’S +, +047°22.19’E +collector: + +R. +Harin’Hala California Acad + +of Sciences malaise trap in high altitude rainforest, MA-02-09A-02”/ “ +HOLOTYPE + +Erephognathus ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label]. +Paratypes +(total of 2): +1 female +(in +CAS +) labeled same as +holotype +, except “CASENT 8068532”; +1 female +(in +CAS +) labeled “CASENT 8068672”/; “ +MADAGASCAR +: +Province d’Antananarivo + +3 km +41° NE Andranomay + +, + +11.5 km +147° SSE Anjozorobe + +elev + +1300 m + + +5-13 December 2000 + +”/ “ +18°28’24”S +47°57’38”E +coll. +Fisher +, Griswold et al. +California Acad. of Sciences +montane rainforest, +malaise trap +, coll. code: BLF2375”. +Both +paratypes +also bear the following label: “ +PARATYPE + +Erephognathus ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + +FIGURE +38. Digital images of holotype female of + +Erephognathus ranomafanae +Kavanaugh and Rainio + +sp. nov. + +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size moderate for genus, SBL of females = 7.5 +- +8. +9 mm +. In Jeannel’s (1949) key to species of + +Erephognathus + +, members of + +E. ranomafanae + +key to + +E. coerulescens +( +Fairmaire 1903 +) + +based on their short antennae (extended posteriorly only to the basal one-fourth of the elytra or less) and the absence of the tuft of setae from the apex of antennomere 11 (terminal antennomere) found in + +E. margarithrix +Alluaud (1936a) + +members. The pronotum ( +Fig. 38A +) is distinctly longer, narrower and less convex and with anterior angles more projected, narrowed and angulate than in members of + +E. coerulescens + +. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— + +We have examined specimens from only two areas. +Two female +specimens were collected in +Ranomafana National Park +, the +type +locality, on the eastern slope off the central plateau in the northeastern part of the southern half of the island of +Madagascar +. A third specimen, also a female, was collected near +Anjozorobe +, in +Antananarivo Province +, slightly north of the center of the island and just over +300 km +north of +Ranomafana +along the eastern slope off the central plateau. +No +specimens have been recorded from intervening areas + +. + + + +HABITAT +DISTRIBUTION + +.— Specimens of the +type +series were all collected in malaise traps in montane rainforest at elevations ranging from +1110 to 1300 m +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573049FFA3FF8EFA93BE8D09FD.xml b/data/0A/3E/26/0A3E26573049FFA3FF8EFA93BE8D09FD.xml new file mode 100644 index 00000000000..523d3aa8c09 --- /dev/null +++ b/data/0A/3E/26/0A3E26573049FFA3FF8EFA93BE8D09FD.xml @@ -0,0 +1,338 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Pseudomasoreus ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 29 +, +30A, 30C + + + +Deuveilla +n. sp. +1 + +; Rainio 2009: 31(informal designation). + + +Lebiinae +(n. sp. 1); Rainio 2012: 72 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.29A–B +), a male, in CAS, labeled: “CASENT 1004471”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, +D.H. & K.M Kavanugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, +R. Ranaivosolo +,”/ “ +E.F. Randrianirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime from large accumulation of dead leaf, twig, and branch debris on and extended up to + +2 m + +above forest floor”/ “ +HOLOTYPE + +Pseudomasoreus ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 8): +1 male +(in +CAS +) labeled same as +holotype +except “CASENT 1004472” + +; + +2 males +(in +CAS +) labeled “CASENT 1004003” and “CASENT 1004004”, respectively + +/ + + + +FIGURE +29. Digital images of holotype male of + +Pseudomasoreus ranomafanae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm. + + + + +FIGURE +30. Digital images of + +Pseudomasoreus + + + +spp. A–C. Holotype male of + +P. ranomafanae + + + +Kavanaugh & Rainio +sp. nov. +; D, E. + +P. catalai +Jeannel + +(from Ranomafana National Park). A. Right labial palpus, ventral aspect; B–E. Aedeagus of male genitalia; B, D. Left lateral aspect; C, E. Ventral aspect. Scale lines = 0.5 mm. + + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size large for genus, SBL of males = 9.7–10. +6 mm +, of females = +10.3 mm +. In +Mateu’s (1980) +key to the Malagasy species of this genus, members of + +P. ranomafanae + +( +Fig. 29A +) key to + +Pseudomasoreus decorsei +Jeannel (1941b) + +based on the following combination of features: size large, SBL = +8.5 mm +or more; elytra concolorous brown or piceous, without pale areas. However, they differ from the unique +holotype +of + +P. decorsei + +in having even larger body size (SBL = +8.5 mm +in + +P. decorsei + +type), pronotum narrower overall and especially basally and with less rounded, more evident hind angles (pronotum broad, rounded, with hind angles effaced by smoothly rounded transition from lateral to basal margin in + +P. decorsei + +type), and elytral lateral margins nearly parallel (more evenly rounded in + +P. decorsei + +type). Members of + +P. ranomafanae + +are actually more similar to those of + +P. deuvei + +, which was not included in Mateu’s key. They share large size, general body form with long, parallel-sided elytra, and antennomere 3 glabrous except for the apical whorl of fixed setae (additional pubescence seen in apical part of antennomere +3 in +most other species of genus). However, their body size is slightly smaller on average (SBL = 10.2–11. +5 in + +P. deuvei + +members), the pronotum is narrower overall and more narrowed basally, with the hind angles more distinct than in + +P. deuvei + +members and the lateral margins either straight or with slight sinuation anterior to the hind angles (lateral margins not at all sinuate in the latter). Also, the penultimate labial palpomeres in all specimens of the type series have three or four setae ( +Fig. 29C +), whereas members of + +P. deuvei + +have only two such setae, a feature particularly noted by Casale in his original description. Finally, the median lobe of male genitalia ( +Figs. 30A,C +) has the shaft slightly less arcuate and the apex slightly longer on more rounded apically than in + +P. deuvei + +males + + +(see +Casale 1998 +, +Figs.4–5 +). For future comparative purposes, we also provide digital photographs of the median lobe of the genitalia of a male + +Pseudomasoreus catalai +Jeannel (1949) + +( +Figs. 30B,D +) [male previously unknown] to complement the illustrations of other + +Pseudomasoreus +species + +provided by Jeannel (1949), +Mateu (1980) +and +Casale (1998) +. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Specimens of this species were are collected in secondary montane rainforest in the Talatakely and Vohiparara areas at elevations ranging from +900–1170 m +. Adults were collected by hand, by beating live vegetation and suspended dead leaves and twigs, particularly dead leaves suspended is leaf whirls of + +Pandanus +sp. + +Two specimens +were found in daytime in masses of dead leaves, twigs and branch debris accumulated on the forest floor. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657304AFFA3FF8EFA35B9F60BA6.xml b/data/0A/3E/26/0A3E2657304AFFA3FF8EFA35B9F60BA6.xml new file mode 100644 index 00000000000..0ead91ab131 --- /dev/null +++ b/data/0A/3E/26/0A3E2657304AFFA3FF8EFA35B9F60BA6.xml @@ -0,0 +1,143 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Assadecma basilewskyi +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 31A–B +, 32 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 31A–B +), a male, in NMNH, labeled: “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +940m + +21°16’S +, +47°25’E + +18 April 1994 + +”/ “ +pitfall traps +in montane rainforest, +E. Rajeriarison +& +R. Malenky +”/ “ +HOLOTYPE + +Assadecma basilewskyi +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 2): +1 male +(in +NMNH +) labeled same as +holotype + +; + +1 male +(in +CAS +) labeled “CASENT 1051147”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, 26 +April + + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657304FFFA7FF8EFD0BBFBD0B61.xml b/data/0A/3E/26/0A3E2657304FFFA7FF8EFD0BBFBD0B61.xml new file mode 100644 index 00000000000..4b8e723738a --- /dev/null +++ b/data/0A/3E/26/0A3E2657304FFFA7FF8EFD0BBFBD0B61.xml @@ -0,0 +1,320 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Lebia +( +Metalebia +) +laterolucida +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 33 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 33A–B +), a female, in CAS, labeled: “CASENT 1006164”/ +MADAGASCAR +Fianarantsoa Prov. + +Parc +Nacional Ranomafana Bell Vue + +Trail, tropical forest +21°15.5’S +47°25.6’E + +1000 m + +M.E.Irwin +and +E.I. Schlinger +MEI 99-MA-7 + +21-XII-1999 + +”/ “ +HOLOTYPE + +Lebia +( +Metalebia +) +laterolucida +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + +Paratypes +(only 1): a female (in +CAS +) labelled “CASENT 1049030”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +850 m + +, + +13 April 1998 + +,”/ “ +21°14’S +47°22’E +Malaise trap +in mixed tropical forest, +M.E.Irwin +and +E.I. Schlinger +collectors, Stop 98- MAD-1”/ “ +PARATYPE + +Lebia +( +Metalebia +) +laterolucida +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + +TYPE +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, Ranomafana National Park. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +laterolucida + +, is an adjective derived from the Latin words, +lateris +, meaning side, and +lucidus +, meaning bright or full of light. The name refers to the pale, translucent lateral pronotal margins, which contrast markedly with the dark pronotal disc in members of this species. + + + +RECOGNITION + +.— Size moderate for genus, SBL of females = 4.5–5.0 mm. Members of this species ( +Fig. 33A +) key to subgenus + +Metalebia +Jeannel (1949) + +and the + +madagascariensis + +group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody (head plus pronotum) has no metallic reflection and the elytra are uniform in color; and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the + +madagascariensis + +group, + +L. laterolucida + +members are distinguished from those of + +L. alluaudana +Jeannel (1949) + +, + +L. apicoviolacea + +sp. nov. +, + +L. mirana +Alluaud (1936a) + +, + +L. ranomafanae + +sp. nov. +, and + +L. tanala +Jeannel (1949) + +by the absence of metallic reflection on the elytra and punctation of the anterior dorsum of the head (both features present in members of the other five species). They differ from members of + +L. sulcipennis +(Fairmaire) (1889) + +, + +L. nana +Jeannel (1949) + +, and + +L. perrieri +Jeannel (1949) + +in having the pronotum broad, ratio PW/PL = at least 1.5, and with lateral margins distinctly rounded at least in the anterior half pronotum (narrower and less rounded anteriorly in members of the three other species). From adults of + +L. rufa +Jeannel (1949) + +, they differ in having the pronotal basal lobe distinctly margined (absent from the former). + +Lebia laterolucida + +adults are most similar to those of the two remaining species in Jeannel’s key, + +L. brunneipennis +Jeannel (1949) + +and + +L. madagascariensis +Chaudoir (1850) + +, but differ from them in having the dorsum of the head, pronotum and elytra dark piceous, with markedly contrasting pale yellow lateral pronotal margins (dorsum uniformly reddish brown and the lateral pronotal margins not or less markedly contrasting in color in the together two species). In addition, they are distinctly smaller than + +L. madagascariensis + +adults and have distinct elytral microsculpture (absent or at least very faint in + +L. brunneipennis + +adults). + + + +FIGURE +33. Digital images of holotype female of + +Lebia +( +Metalebia +) +laterolucida +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— + +Both specimens of the +type +series were collected in mixed secondary tropical forest in the +Talatakely area +in malaise traps + +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573051FFBAFF8EF999BC920B24.xml b/data/0A/3E/26/0A3E26573051FFBAFF8EF999BC920B24.xml new file mode 100644 index 00000000000..5b353eafcb0 --- /dev/null +++ b/data/0A/3E/26/0A3E26573051FFBAFF8EF999BC920B24.xml @@ -0,0 +1,281 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Eurydera ocellata +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 23–24 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 23A–B +), a male, in CAS, labeled: “CASENT 8068800”/ “ +MADAGASCAR +: +Province +Fianarantsoa +, + +Parc National +Ranomafana + +, +Bellevue +at +Telatakely +, elev + +1020 m + + +12-19 February 2002 + +”/ “ +21°15.99’S +, +47°25.21’E +collector: + +R. +Harin’Hala California Academy + +of Sciences malaise, secondary tropical forest MA-02-09C-16”/ “ +HOLOTYPE + +Eurydera ocellata +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(only 1): a female (in +NMNH +) labeled “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +900 m + + +1-9 February 1990 + +W. E. Steiner +”/ “at black light in montane rainforest near river and stream”/ “ +PARATYPE + +Eurydera ocellata +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + +FIGURE +23. Digital images of holotype male of + +Eurydera ocellata +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Label. Scale line = 1.0 mm. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ocellata + +, is the Latin adjective meaning having little eyes or marked with spots. The name refers to the small, faint dark spot on interval 3 within the subapical pale area of each elytron at the insertion point of the subapical discal setiferous pore. + + + +RECOGNITION + +.— Size small for genus, SBL of male and female = +9.2 mm +. Members of this species ( +Fig. 23A +) key to + +Eurydera mormolycoides +Coquerel (1851) + +in Jeannel’s (1949) key on the basis of the following combination of features: size small; head ( +Fig. 24A +) with frontal furrows deeply impressed, smooth, divergent posteriorly and without a transverse impression uniting them, posterior part of frons smooth, narrow and triangular, without lateral grooves prolonged posteriorly beyond the posterior border of the eyes; mandibles short and broad, lateral border not broadly explanate, lateral margin only slightly convex; pronotum cordate, distinctly narrowed basally, lateral margins angulate at insertions of midlateral setae; elytra short, round and black, each with a reddish subapical spot tangential to the medial suture as well as subhumeral and subapicolateral reddish areas and reddish apical spines, elytral intervals moderately convex; median lobe of male genitalia without a large, angulate projection on the ventral margin of the shaft near the base. They differ from + +E. mormolycoides + +members in having: dorsal surface slightly shiny (dull in + +E. mormolycoides + +members); pronotum with lateral explanations pale testaceous (darker in + +E. mormolycoides + +members), anterior angles broadly rounded (more narrowly rounded in + +E. mormolycoides + +members) and lateral margins with shallow but distinct sinuation anterior to hind angles (sinuation absent from or less distinct in + +E. mormolycoides + +members); elytra with larger but less distinct (less contrasting with rest of elytra) pale subapical macula with black eye spot on interval three (macula extended further laterally and lateroanteriorly than in + +E. mormolycoides + +members) and apical spine about half as long as spine in + +E. mormolycoides + +members. Finally, the median lobe of the male genitalia ( +Figs. 24B–C +) has a thicker shaft throughout and an apex shorter and thicker in lateral view ( +Fig. 24B +) and broader and straighter in dorsal view ( +Fig. 24C +) than in + +E. mormolycoides + +males (see Jeannel 1949, Fig. 492b). + + + +FIGURE +24. Digital images of holotype male of + +Eurydera ocellata +Kavanaugh & Rainio + +sp. nov. +A. Head, dorsal aspect; + +B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The +holotype +specimen was collected in a malaise trap in secondary montane rainforest in the Talatakely area at an elevation of +1030 m +. The +paratype +specimen was collected at ultraviolet light in secondary montane rainforest near the junction of the Namorona River and a small tributary stream. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573057FFA0FF8EFF49BC7D0FEE.xml b/data/0A/3E/26/0A3E26573057FFA0FF8EFF49BC7D0FEE.xml new file mode 100644 index 00000000000..ef0ef37bda2 --- /dev/null +++ b/data/0A/3E/26/0A3E26573057FFA0FF8EFF49BC7D0FEE.xml @@ -0,0 +1,346 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Eurydera simplica +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 27–28 + + + +Eurydera sp. + +; Rainio and Niemelä 2006: 227 (informal designation). + + + +Eurydera +n. sp. +3 + +; Rainio 2009: 31, Rainio 2012: 72 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.27A–B +), a male, in CAS, labeled: “ +MADAGASCAR +, Ranomafana NP., +Talatakely +, + +900 m + +, Trail SP at + +50 m + +mark, + +08.03.2004 + +”/ “collected by hand in daytime in secondary montane rainforest, +F. Ratalata +& +J. Rainio +collectors”/ “106” [handwritten label]/ “ + + +HOLOTYPE + +Eurydera simplica +Kavanaugh & Rainio + +sp. n. +2015” [red label]. +Paratypes +(only 1): a female (in +MNHN +) labeled ““ +MADAGASCAR +, Ranomafana NP., +Talatakely +, + +900 m + +, Trail B at + +1000 m + +mark, + +15.06.2000 + +”/ “collected by hand in daytime in secondary montane rainforest, +F. Ratalata +& +J. Rainio +collectors”/ “7/13” [handwritten label]/ “ + +PARATYPE + +Eurydera simplica +Kavanaugh & Rainio + +sp. n. +2015” [yellow label]. + + + +FIGURE +27. Digital images of holotype male of + +Eurydera simplica +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + +FIGURE +28. Digital images of holotype male of + +Eurydera simplica +Kavanaugh & Rainio + +sp. nov. +A. Head, dorsal aspect; + +B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +simplica + +, is an adjective derived from the Latin word, +simplex +, meaning simple. The name refers to the relative simplicity of form and surface macrosculpture of member of this species compared with those of other species of + +Eurydera + +. + + + +RECOGNITION + +.— Size average for genus, SBL of male = +13.1 mm +, of female = 12.4. Members of this species ( +Figs. 27A +) are unlike those of any other species in the genus. In Jeannel’s (1949) key species of + +Eurydera + +, they key to his + +unicolor + +species group, based on the following features: size medium; head ( +Fig. 28A +) with posterior part of frons formed as a broad, flat, smooth, elevat- ed plateau, laterally delimited by deep furrows extended posteriorly beyond the level of the posterior margin of the eye then slightly and arcuately convergent posteriorly and terminated abruptly at the level of the posterior margin of the ocular swelling and just less than halfway from the midline to the lateral margin of the head, area between the furrow smooth and only slightly elevated; mandibles moderately short and broad; median lobe of male genitalia without a large, angulate projection on the ventral margin of the shaft near the base. Jeannel included only + +Eurydera unicolor +( +Klug 1833 +) + +in this species group, but +Mateu (1973) +added another species, + +Eurydera fossulata +Mateu + +to the group, in part based on the shared lack of sexual dimorphism in the front tarsi in members of these two species. Unlike males of most + +Eurydera +species + +, those of + +E. unicolor + +and + +E. fossulata + +have tarsomeres 1 to 3 no wider than in females and without ventral adhesive setae. Members of + +E. simplica + +are easily distinguished from those of these two species, as well as from all other described + +Eurydera +species + +, by the uniquely shaped pronotum ( +Fig. 27A +), which is relatively small and short for the genus, with the disc smooth, lateral explanation very narrow, lateral margins with very short and shallow sinuation anterior to markedly obtuse hind angles, and basal margin obliquely angulate laterally. Unlike males of + +E. unicolor + +and + +E. fossulata + +, those of + +E. simplica + +have front tarsi with tarsomeres 1 to 3 wider than in females and with at least a few adhesive setae ventrally, which may call into question a close relationship with these two these species. Members of + +E. simplica + +also differ from those of + +E. unicolor + +in having the pronotum with anterior angles much less projected anteriorly. They differ from members of + +E. fossulata + +in having the posterior part of the head, between the lateral furrows, smooth and slightly convex, whereas + +E. fossulata + +members have a deep, transverse medial depression, the bottom of which is flat and covered with deep transverse creases, on that part of the head. They also have the elytral intervals without the roughened surfaces typical of members of several species in genus, including + +E. fossulata + +(at least subapically), and the sutural spines are very long and straight (similar to those in + +E. unicolor + +and + +E. fossulata + +members). Finally, the median lobe of male genitalia of the +holotype +of + +E. simplica + +( +Fig. 28B–C +) has the shaft much narrower basally and the apical orifice shorter than in +E. unicol- or +males (see Jeannel 1949, Fig. 495). No male of + +E. fossulata + +has been available for comparison. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The two known specimens of this species both were collected by hand in daytime in secondary montane rainforest in the Talatakely area at elevations ranging from +900 to 1000 m +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573058FFB3FF8EFCB7B95E09C5.xml b/data/0A/3E/26/0A3E26573058FFB3FF8EFCB7B95E09C5.xml new file mode 100644 index 00000000000..22e0f87be36 --- /dev/null +++ b/data/0A/3E/26/0A3E26573058FFB3FF8EFCB7B95E09C5.xml @@ -0,0 +1,563 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Madecassina bimaculata +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 19 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.19A–B +), a male, in CAS, labeled: “CASENT 1049022”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +13 April 1998 + +, +21°14.9’S +47°25.6’E +,”/ “Stop # 98-29, +D. H. Kavanaugh +collector collected by beating suspended clusters of dead leaves and twigs”/ “ +HOLOTYPE + +Madecassina bimaculata +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 16): +1 female +(in +CAS +) labeled “CASENT 1004001”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, +D.H. & K.M Kavanaugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, +R. Ranaivosolo +,”/ “ +E.F. Randrianirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime by beating live understory vegetation and/or dead leaves and other suspended debris” + +; + +1 female +(in +CAS +) labeled “CASENT 1049023”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, Talatakely area, + +900 m + +, +Trail +FF, +21°14.9’S +47° 25.6’E +,”/ “ + +10-16 November 1998 + +, beaten from vines, +V. F. Lee +& +K. J. Ribardo +collector” + +; + +1 male +(in +CAS +) labeled “CASENT 1049024”/ “ +MADAGASCAR +, +Fianarantsoa Province +, Parc National +Ranomafana +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +24 April 1998 + +,”/ “ +21°15.3’S +47°25.9’E +,”/ “Stop # 98-76, +D. H. Kavanaugh +collector collected at night on standing tree trunk” + +; + +1 male +(in +CAS +) labeled “CASENT 8068817”/ “ +MADAGASCAR +: +Province +Fianarantsoa +, +Ranomafana National Park +, radio tower at forest edge, elev + +1130 m + + +9-20 March 2003 + +”/ “ +21°15.05’S +, +47°24.43’E +collector: +R. Harin’Hala California Academy +of Sciences malaise, mixed tropical forest MA-02-09B-55” + +; + +1 male +(in +CAS +) labeled “CASENT 1049026”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Vohiparara area +, + +1150 m + +, mixed tropical forest, + +18 April 1998 + +, +21°12.8’S +47°23.0’E +,”/ “Stop# 98-52, +D. H. Kavanaugh +collector collected by beating live vegetation and suspended dead leaves and twigs” + +; + +1 female +(in +CAS +) labeled “CASENT 1049025”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, Vatoharanana area, + +1050 m + +, +Ambatovory +stream, + +29 April 1998 + +, +21°16.7’S +47° 26.1’E +,”/ “Stop# 98-101, +D. H. Kavanaugh +collector collected by beating suspended clusters of dead leaves and twigs” + +; + +2 males +and +1 female +(in +CAS +and EM) labeled “CASENT 8072308” or “CASENT 8072309” or “CASENT 8072307”, respectively/ “ +MADAGASCAR +: +Fianarantsoa +Parc National de Ranomafana +, +Vatoharanana River +, + +4.1 km +231° SW Ranomafana + +elev + +1100 m + + +27-31 March 2003 + +”/ “ +21°15’24”S +, +47°26’00”E +California Academy of Sciences +coll. +Fisher +, Griswold et al. beating low veg., montane rainforest code: BLF8401” + +; + +1 male +(in +MNHN +) labeled “ +MADAGASCAR +, +Ranomafana +NP., +Vatoharanana +, Trail K at + +150 m + +mark, + +25.7.2005 + +”/ “collected by hand in daytime from dead leaves of bushes and trees, +F. Ratalata +& +J. Rainio +collectors” + +; + +2 females +(in +MNHN +and +MZF +) labeled “ +MADAGASCAR +, +Ranomafana +NP., +Talatakely +, Trail SP at + +50 m + +mark, + +28.07.2005 + +”/ “collected by hand in daytime from dead leaves of bushes and trees, +F. Ratalata +& +J. Rainio +collectors” + +; + +2 females +(in +MNHN +and +MZF +) labeled “ +MADAGASCAR +, +Ranomafana +NP., +Valohoaka +, Trail B at + +555 m + +mark, + +28.01.2005 + +”/ “collected by hand in daytime from dead leaves of bushes and trees, +F. Ratalata +& +J. Rainio +collectors”; and + + +2 females +(in +MNHN +and +MNHN +) labeled “ +MADAGASCAR +, +Ranomafana +NP., +Valohoaka +, Trail B at + +555 m + +mark, + +16.11.2005 + +”/ “collected by hand in daytime from dead leaves of bushes and trees, +F. Ratalata +& +J. Rainio +collector”. +All +paratypes +also bear the following label: “ +PARATYPE + +Madecassina bimaculata +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + +FIGURE +19. Digital images of holotype male of + +Madecassina bimaculata +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +bimaculata + +, is an adjective derived from the Latin words, +bis +, meaning two, and +macula +, meaning spot or spotted. The name refers to the pair of subapical elytral spots seen in members of this species. + + + +RECOGNITION + +.— Size average for genus, SBL of males = 4.7– +5.5 mm +, of females = 5.3–5. +6 mm +. Members of this species ( +Fig. 19A +) key most closely, to + +Madecassina picta +Alluaud (1897) + +in Jeannel’s (1949) key based on the following combination of features: pronotum impunctate, elytra with two subapical pale spots and striae deeply impressed and legs testaceous in color. The only feature used in the key with which + +T. bimaculata + +members differ is the presence of is metallic reflection on the forebody—members of + +T. picta + +have evident aeneous or greenish metallic reflection on the head and pronotum, whereas + +T. bimaculata + +members are without metallic reflection. These two species are otherwise very similar externally; however the pronotum is proportionately slightly longer and more distinctly narrowed basally and the hind angles more sharply defined and rectangular in + +T. bimaculata + +members than in + +T. picta + +members (which have the hind angles slightly obtuse). Also, the apex of the median lobe of the male genitalia is shorter, thicker and straighter in lateral aspect ( +Fig. 19C +) and more bluntly triangular in dorsal aspect ( +Fig. 19D +) in + +T. bimaculata + +males than in those of + +T. picta + +males, which have the apex longer, narrower and distinctly hooked dorsally in lateral aspect and more narrowly rounded in dorsal aspect (see Jeannel 1949, Figs. +474g +and 474i). + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Specimens of this species have been collected in primary montane rainforest in the Vatoharanana and Valohoaka areas and in secondary montane rainforest in the Talatakely and Vohiparara areas, at elevations ranging from +900 to 1150 m +. They have been collected mainly by beating live vegetation and suspended clusters of dead leaves and twigs. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657305AFFB5FF8EFA95B9710F61.xml b/data/0A/3E/26/0A3E2657305AFFB5FF8EFA95B9710F61.xml new file mode 100644 index 00000000000..7206917fc77 --- /dev/null +++ b/data/0A/3E/26/0A3E2657305AFFB5FF8EFA95B9710F61.xml @@ -0,0 +1,222 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Madecassina quadrimaculata +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 20 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.20A–B +), a male, in CAS, labeled: “ +MADAGASCAR +, Ranomafana NP., montane rainforest near +Tsinjorano village +, + +100 m + +in from forest edge,”/ “ +21.09637°S +47.52085°E +, + +1050 m + +, + +19.05.2005 + +, collected by hand in daytime, +F.J. Ratelolahy +collector”/ “ +HOLOTYPE + +Madecassina quadrimaculata +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +quadrimaculata + +, is an adjective derived from the Latin words, +quattuor +, meaning four, and +macula +, meaning spot or spotted. The name refers to the two (subbasal and subapical) pairs of elytral spots seen in members of this species. + + + +RECOGNITION + +.— Size average for genus, SBL = +5.2 mm +. The +holotype +male of this species ( +Fig. 20A +) keys to + +Madecassina maculata +Alluaud (1899b) + +in Jeannel’s (1949) key based on the following combination of features: body black, forebody without metallic reflection, pronotum punctate laterally, and elytra shiny with four pale spots. It differs from members of + +M. maculata + +in having a smaller body size (SBL = 5.5–6.0 mm in + +M. maculata + +), the pronotum punctate laterally and laterobasally only, with conspicuous transverse grooves and ridges on disc (in + +M. maculata + +the pronotum is punctate on the disc also, between the ridges, in contrast to what Jeannel said in his key), elytral intervals shinier due to more shallowly impressed isodiametric microsculpture (elytral intervals duller due to more deeply impressed isodiametric microsculpture in + +M. maculata + +), the elytral spots much more distinct (much less so in + +M. maculata + +) and median lobe of male genitalia with apex hooked dorsally and thicker in lateral aspect ( +Fig. 20C +) and slightly swollen and bent right in dorsal aspect ( +Fig. 20D +) (apex hooked but thinner in lateral aspect and more parallel-sided and straight in dorsal aspect in + +M. maculata + +males) (see Jeannel 1949, Fig. 474f). + + + +FIGURE +20. Digital images of holotype male of + +Madecassina quadrimaculata +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The unique +holotype +was collected in montane rainforest near Tsinjorano village at a distance of +100 m +inside the forest edge at an elevation of +1050 m +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657305CFFB8FF8EFB4BBCA909E0.xml b/data/0A/3E/26/0A3E2657305CFFB8FF8EFB4BBCA909E0.xml new file mode 100644 index 00000000000..148d250e0e6 --- /dev/null +++ b/data/0A/3E/26/0A3E2657305CFFB8FF8EFB4BBCA909E0.xml @@ -0,0 +1,689 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Pristacrus ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 21 +–22 + + + +Pristacrus +n. sp. +1 + +; Rainio 2009: 33, Rainio 2012: 73 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.21A–B +), a male, in CAS, labeled: “CASENT 1003833”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, +D.H. & K.M Kavanugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, +R. Ranaivosolo +,”/ “ +E.F. Randrianirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected at night on surface of rotting logs on forest floor or suspended just above it”/ “ +HOLOTYPE + +Pristacrus ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 19): +1 male +and +1 female +(in +CAS +) + + +labelled “CASENT 1003832” or “CASENT 1003831”, respectively, otherwise labeled same as +holotype +; +1 female +(in +CAS +) + + +labeled “CASENT 1004470”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, +D.H. & K.M Kavanugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, +R. Ranaivosolo +,”/ “ +E.F. Randrianirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime from large accumulation of dead leaf, twig, and branch debris on and extended up to + +2 m + +above forest floor”; +1 male +(in +CAS +) + + +labeled “CASENT 1002838”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, +D.H. & K.M Kavanugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, +R. Ranaivosolo +,”/ “ +E.F. Randrianirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime under loss bark of rotting log logs on forest floor or suspended just above it”; +1 male +and +two females + + + + +FIGURE +21. Digital images of holotype male of + +Pristacrus ranomafanae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm. + + + +(in +CAS +and +MZF +) labeled “CASENT + +1003752”, “CASENT 1003751” or “CASENT + +1003753”, respectively/ “ +MADAGASCAR +, + + + +Fianarantsoa Province +, +Ranomafana +National + + + + +Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S + +/ + + +47.41945°E +, Stop# DHK-01-001, D.H. & K.M + +Kavanugh, R.L. Brett, E. Elsom, F. Vargas, +R. Ranaivosolo,”/ “E.F. Randrianirina, +N. Rasoamananana, T.J. Ravelomanana, and + +H.C. Raveloson collectors”/ “collected in daytime on underside of rotting log logs on forest floor or suspended just above it”; +1 male +(in + + + +CAS +) labeled “CASENT 1004313”/ “MADA- +FIGURE +22. +Digital +images of aedeagus of genitalia of GASCAR, +Fianarantsoa Province +, Ranoma- male +holotype +of + +Pristacrus ranomafanae +Kavanaugh + +& fana +National Park +, +Talatakely area +, + +900 m + +, +Rainio +sp. nov. +A. Left +lateral aspect; +B. Dorsal +aspect. +Scale +line = +0.5 mm + +. + + +mixed tropical forest, +2-22 January 2001 +,”/ + + +“ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, D.H. & K.M Kavanugh, R.L. Brett, E. Elsom, F. Vargas, R. Ranaivosolo,”/ “E.F. Randrianirina, N. Rasoamananana, T.J. Ravelomanana, and H.C. Raveloson collectors”/ “collected in daytime from leaf litter on forest floor”; + +1 female +(in +CAS +) labeled “CASENT 1051144”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +27 April 1998 + +”/ “ +21°15.3’S +47°25.9’E +, Stop # 98-85 +A D.H. Kavanaugh +collector collected in and under rotten logs on forest floor” + +; + +1 female +(in +CAS +) labeled “CASENT 8068264”/ “ +MADAGASCAR +: +Province +Fianarantsoa +, Parc National +Ranomafana +, +Belle Vue +at +Talatakely +, elev + +1020 m + + +28 April – 5 May 2002 + +”/ “ +21°15.99’S +47°25.21’E +collector: +R. Harin’Hala California Acad +of Sciences malaise secondary tropical forest MA-02-09C-27” + +; + +1 female +(in +CAS +) labeled “CASENT 8068264”/ “ +MADAGASCAR +: +Province +Fianarantsoa +, Parc National +Ranomafana +, +Belle Vue +at +Talatakely +, elev + +1020 m + + +15-22 November 2001 + +”/ “ +21°15.99’S +47°25.21’E +collector: +R. Harin’Hala California Acad +of Sciences malaise secondary tropical forest MA-02-09C-03” + +; + +1 male +(in +NMNH +) labeled “ +MADAGASCAR +: +Prov. +Fianarantsoa +, + +7 km +W Ranomafana + +, + +900 m + + +1-9 February 1990 + +W. E. Steiner +”/ “at black light in montane rainforest near river and stream” + +; + +1 male +(in +CAS +) labeled “CASENT 8005762”/ “ +21.25041°S +/ +47.41945°E +MADAGASCAR +, +Fianarantsoa +Prov Ranomafana National Park, Talatakely area, + +900m + +, 10:1:2001 +Col. K. Will +headlamp search logs” + +; + +5 males +and +1 female +(in +EMEC +and +MNHN +) labeled “CASENT 8005762”/ “ +21.25041°S +/ +47.41945°E +MADAGASCAR +, +Fianarantsoa +Prov Ranomafana National Park, Talatakely area, + +900m + +, 10:1:2001 +Col. K. Will +headlamp search logs”. +All +paratypes +also bear the following label: “ +PARATYPE + +Pristacrus ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park +. + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected + +. + + + +RECOGNITION + +.— Size small for genus, SBL of males = 8.3–8. +5 mm +, of female = +8.6 mm +. Members of this species ( +Fig. 21A +) are distinguished from those of all other species in the genus except + +Pristacrus binotatus +Klug (1833) + +by their small body size (SBL greater than 10.0 mm in members of the three other species). They differ from members of + +Pristacrus semipiceus +( +Fairmaire 1887 +) + +, in having much smaller body size (SBL = 13.6 to 14. +5 mm +in + +P. semipiceus + +members), a shinier dorsal surface, shorter and broadly rounded mandibles and an impunctate (but longitudinally rugulose) frons (dorsal surface duller, mandibles distinctly longer and more tapered, and frons punctate as well as rugulose in + +P. semipiceus + +members). They differ from members of + +P. laticollis + +in having slightly smaller body size (SBL = 10.2–11. +1 mm +), slightly shinier dorsal surface, relatively longer and narrower pronotum and elytra that are broadly ovoid with lateral margin arcuate and lateral explanation broad throughout, especially at middle, and elytral apices distinctly angulate and toothed (duller dorsal surface, relatively broader and shorter pronotum and elytra that are nearly parallel-sided at middle with lateral explanation narrower throughout and elytral apices broadly rounded in + +P. laticollis + +members). They differ from the unique +type +of + +Pristacrus rotundatus +( +Fairmaire 1892 +) + +in having smaller body size (SBL = +11.9 mm +in + +P. rotundatus + +type +), shinier dorsal surface, elytra less broadly rounded and humeri less anteriorly projected (dorsal surface dull and elytra markedly broad and rounded and humeri more projected anteriorly in + +P. rotundatus + +type +). Members of + +P. ranomafanae + +are very similar to those of + +P. binotatus + +in size and overall form. Specimens of the former have the dorsal surface slightly shinier, the pronotum relatively broader basally, with subbasal sinuation of the lateral margins absent or only very faint and hind angles less sharply defined, and elytra with striae slightly more deeply impressed and intervals more convex and more roughened at the raised center than in most members of + +P. binotatus + +. However, at least a few specimens of + +P. binotatus + +share one or more of these features with + +P. ranomofanae + +members. They only features that most reliably distinguish members of + +P. ranomafanae + +from those of + +P. binotatus + +, as well as from members of the other species, are the male genitalia. In + +P. ranomafanae + +males (Fig. 22A–B), the shaft of the median lobe is inflated subbasally but lacks the large, well-defined dorsal bulge seen in males of + +P. binotatus + +(see Jeannel 1949, Fig. 486) and, in lateral aspect ( +Fig. 21A +), the apex is distinctly hooked dorsally (apex not hooked in + +P. binotatus + +males. Males of the each of the other species also have the shaft of the median lobe uniquely shaped. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Specimens of the +type +series were all collected in secondary montane rainforest in the Talatakely area at elevations of +900 to 1020 m +. The area is replete with invasive guava ( + +Psidium cattleianum + +). Specimens were found in daytime in masses of dead leaves, twigs and branch debris accumulated on the forest floor, in leaf litter, and under loose bark or on the underside of rotting logs on the forest floor. Several were found at night running on the surfaces of rotting logs on the forest floor; two were collected in malaise traps and one was attracted to and collected at an ultraviolet light. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573060FF8BFF8EFF49BEB20CA8.xml b/data/0A/3E/26/0A3E26573060FF8BFF8EFF49BEB20CA8.xml new file mode 100644 index 00000000000..ca4d0197137 --- /dev/null +++ b/data/0A/3E/26/0A3E26573060FF8BFF8EFF49BEB20CA8.xml @@ -0,0 +1,409 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Perigona +( +Ripogena + +) + +deuvei +Kavanaugh and Rainio + + + +, +sp. nov. + + + + + + +Figure 14 + + + +Ripogena +n. sp. +1 + +; Rainio 2009: 33 (informal designation). + + + +Perigona +( +Ripogena +) + +n. sp. 1 +; Rainio 2012: 73, Rainio 2013: 96 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.14A–B +), a male, in CAS, labeled: “ +MADAGASCAR +, Ranomafana NP., +Vatoharanana +, Trail K at + +350 m + +mark, + +24.07.2005 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “ +HOLOTYPE + +Perigona +( +Ripogena +) +deuvei +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 4): +1 female +(in +MNHN +) labeled “ +MADAGASCAR +, Ranomafana NP., +Vatoharanana +, +Trail +J at + +50 m + +mark, + +6.10.2005 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “239.” [handwritten label] + +; + +1 female +(in +CAS +) labeled “ +MADAGASCAR +, Ranomafana NP., +Vatoharanana +, +Trail +S at + +600 m + +mark, + +9.9.2005 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “239.” [handwritten label] + +; + +1 male +(in +MZF +) labeled “ +MADAGASCAR +, Ranomafana NP., +Vatoharanana +, +Trail +S at + +600 m + +mark, + +24.7.2005 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “253.” [handwritten label] + +; + +1 female +(in +MZF +) labeled “ +MADAGASCAR +, Ranomafana NP., +Vatoharanana +, +Trail +S at + +600 m + +mark, + +6.10.2005 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “245.” [handwritten label]. All +paratypes +also bear the following label: “ +PARATYPE + +Perigona +( +Ripogena +) +deuvei +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +deuvei + +, is a noun in apposition, derived from the surname of Dr. Thierry Deuve of the Laboratoire d’Entomologie at the Muséum National d’Histoire Naturelle in Paris. We are pleased to name this new species in honor of Dr. Deuve in thanks for his friendship and kind hospitality during our several visits to MNHN in the course of this study. + + + +RECOGNITION + +.— Size average for subgenus, SBL of males = 3.5– +3.8 mm +, of females 3.8– +3.9 mm +. Members of this species ( +Fig. 14A +) share with those of + +Perigona heterodera +Alluaud (1936) + +and + +Perigona prasinus +Alluaud (1936) + +the following features: pronotum with lateral margins without sinuation anterior to hind angles, posterior margin slightly and smoothly convex, and hind angles either obtuse or broadly rounded. These features distinguish members of these three species from those of + +Perigona bembidioides +Alluaud (1936) + +and + +Perigona viridimicans +Jeannel (1948) + +, which have distinct sinuation of the lateral margins anterior to rectangular hind angles and a straight basal margin. Members of + +P. deuvei + +differ from those of + +P. prasinus + +in having smaller body size (SBL = +4.7 mm +in + +P. prasinus + +), narrower body form, and dorsum without metallic reflection (dorsum with metallic green reflection in + +P. prasinus + +) but elytra with moderate but distinct iridescence generated by microsculpture comprised of transverse microlines and marjkedly transverse sculpticells. They are most similar to members of + +P. heterodera + +in body form and size but differ from them in having the pronotum proportionately longer and male genitalia ( +Figs. 14C–D +) with shaft markedly narrowed basally, markedly inflated in apical two-thirds, and apex longer, slender and ventrally bent in lateral view ( +Fig. 14C +), slightly deflected left and slightly longer in dorsal view ( +Fig. 14D +) than in + +P. heterodera + +(see Figs. 351b-c in Jeannel 1948). They differ from members of + +Perigona descarpentriesi +( +Deuve 1998 +) + +in having moderately large eyes (eyes markedly reduced eyes in size in + +P. descarpentriesi + +), pronotum distinctly narrower and with less rounded lateral margins, and elytra with only stria 1 deeply impressed, stria 2 and 3 shallowly impressed but evident on disk, striae 4 to 7 effaced, and stria 8 impressed only in apical one-third (striae 1–7 shallowly impressed but evident at least on disc and stria 8 deeply impressed throughout in + +P. descarpentriesi + +members). Also, the apex of the median lobe of the male genitalia is narrower in lateral aspect in + +P. deuvei + +males (Fig. 143C) than in those of + +P. descarpentriesi + +(see +Deuve 1998 +, +Fig. 2 +.). Finally, + +P. deuvei + +members differ from those of the new species described below in having small- er body size, pronotum with lateral margination very narrow throughout, fewer elytral striae evident, and median lobe of the male genitalia with a longer and ventrally bent apex. + + + +FIGURE +14. Digital images of holotype male of + +Perigona +( +Ripogena +) +deuvei +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— All specimens of this species were collected by hand in daytime in primary montane rainforest in the Vatoharanana area at an elevation of +1200 m +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573062FF8CFF8EFE4EBC080B0A.xml b/data/0A/3E/26/0A3E26573062FF8CFF8EFE4EBC080B0A.xml new file mode 100644 index 00000000000..44e5239062d --- /dev/null +++ b/data/0A/3E/26/0A3E26573062FF8CFF8EFE4EBC080B0A.xml @@ -0,0 +1,320 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Perigona +( +Ripogena + +) + +ranomafanae +Kavanaugh and Rainio + + + +, +sp. nov. + + + + + + +Figure 15 + + + +Ripogena +n. sp. +2 + +; Rainio 2009: 33 (informal designation). + + + +Perigona +( +Ripogena +) + +n. sp. 2 +; Rainio 2012: 73, Rainio 2013: 96 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.15A–B +), a male, in CAS, labeled: ““CASENT 1049017”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Vatoharanana area +, + +1050 m + +, +Abotovory +stream, + +29 April 1998 + +”/ “ +21°16.7’S +47° 26.1’E +, Stop # 98-100, +D.H. Kavanaugh +collector collected in large rotting log just under loose bark with cavities”/ “ +HOLOTYPE + +Perigona +( +Ripogena +) +ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size larger than average for subgenus, SBL = +4.4 mm +. The +holotype +male of this species ( +Fig. 15A +) shares with members of + +Perigona heterodera +Alluaud (1936) + +, + +Perigona prasinus +Alluaud (1936) + +and + +P. deuvei + +sp. nov. +the following features: pronotum with lateral margins without sinuation anterior to hind angles, posterior margin slightly and smoothly convex, and hind angles either obtuse or broadly rounded. These features distinguish members of these four species from those of + +Perigona bembidioides +Alluaud (1936) + +and + +Perigona viridimicans +Jeannel (1948) + +, which have distinct sinuation of the lateral margins anterior to rectangular hind angles and a straight basal margin. Members of + +P. ranomafanae + +differ from those of + +P. prasinus + +in having slightly smaller body size and dorsum without metallic reflection (dorsum with metallic green reflection in + +P. prasinus + +). They differ from members of + +P. heterodera + +and + +P. deuvei + +in having larg- er body size (SBL = +3.4 mm +in + +P. heterodera + +, 3.5 to +3.9 mm +in + +P. deuvei + +), pronotal hind angles more sharply angulate, lateral explanation of pronotum markedly widened and flattened in region of hind angles (pronotal hind angles rounded or at least less sharply angulate and lateral explanation only slightly wider near hind angles than anteriorly in + +P. heterodera + +and + +P. deuvei + +). The unique male +holotype +of + +P. ranomafanae + +is slightly teneral and the genitalia are very soft and only lightly sclerotized. However, the apex is sclerotized sufficiently to allow comparisons with those of + +P. heterodera + +and + +P. deuvei + +males. In lateral view ( +Fig. 15C +), the apex is slightly longer than in + +P. heterodera + +males (see Jeannel 1948, Fig. 351b) and distinctly shorter and straighter than in + +P. deuvei + +males. Finally, they differ from members of + +P. descarpentriesi + +in having moderately large eyes (eyes markedly reduced eyes in size in + +P. descarpentriesi + +), pronotum widest anterior to middle and lateral margins nearly straight in posterior half (pronotum widest at middle and lateral margins evenly arcuate throughout + +P. descarpentriesi + +). Also, the apex of the median lobe of the male genitalia is narrower and more pointed in lateral aspect in + +P. ranomafanae + +males ( +Fig. 15C +) than in those of + +P. descarpentriesi + +(see +Deuve 1998 +, +Fig. 2 +.). + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The unique +holotype +was collected by hand in primary montane rainforest in the Vatoharanana area at an elevation of +1050 m +from under the loose bark of a large log on the forest floor. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573064FF8EFF8EFD99BEDB0B4C.xml b/data/0A/3E/26/0A3E26573064FF8EFF8EFD99BEDB0B4C.xml new file mode 100644 index 00000000000..b1f772efcf7 --- /dev/null +++ b/data/0A/3E/26/0A3E26573064FF8EFF8EFD99BEDB0B4C.xml @@ -0,0 +1,285 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Archicolliuris ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 16 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.16A–B +), a female, in NMNH, labeled: “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +1100m + + +22-31 October 1988 + +W. E. Steiner +”/ “ +Flight intercept-yellow pan trap +, island in stream, montane rainforest”/ “ +HOLOTYPE + +Archicolliuris ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size large for genus, SBL = +8.2 mm +. The +holotype +female of this species ( +Fig. 16A +) is easily distinguished from members of all other Malagasy odacanthine species based on numerous features. The following features used by Jeannel (1948) in his key to genera are shared with the three species that he included in “ + +Casnonia + +”: integument glabrous (without pubescence), pronotum long, narrow and tubular anteriorly, impunctate, with one or more setae present along each lateral margin. The +holotype +of + +A. ranomafanae + +differs from members of + +Casnonia fairmairei +Gestro (1895) + +[currently included in genus + +Erectocolliuris +Liebke (1931) + +] in having a pronotum black in color and with only a single pair of midlateral setae (pronotum rufotestaceous in color and with 5 or 6 pairs of lateral setae present in + +E. fairmairei + +members). It differs from members + +Casnonia coerulans +Künckel d’Herculais (1887) + +[currently included in genus + +Protocolliuris +Liebke (1931) + +in having the pronotum only about twice as long as wide (three times as long as wide in + +P. coerulans + +), the elytra without metallic reflection (elytra with a dark metallic blue reflection), and legs dark black to piceous, except trochanthers rufous and basal parts of all femora pale (legs pale in + +P. coerulans + +, except apical parts of femora dark). It is similar to members of the last of Jeannel’s “ + +Casnonia + +” species, + +Colliuris olsoufieffi +Alluaud (1935) + +[currently included in + +Archicolliuris + +] in having a shiny black dorsum (with the only pale areas present as elytral pale spots), the pronotum with transverse grooves and ridges in the basal half and anterior angles projected laterally, and the elytra with a deep transverse depression at the basal one-fourth and with three or four discal setiferous pores on elytral interval 3. However, it differs with members of + +A. olsoufieffi + +in several features: body size larger (SBL = +6.8 mm +in + +A. olsoufieffi + +); pronotum impunctate (coarsely punctate laterally and basally in + +A. olsoufieffi + +), slightly less than twice as long as wide (2.5 times as long as wide in + +A. olsoufieffi + +), with lateral borders vaguely present but only as very faintly impressed lines (lateral borders absent from + +A. olsoufieffi + +members); elytra with only one pair of small pale spots, located at apical one-third on interval 4 (both subapical and subbasal pairs of spots present in + +A. olsoufieffi + +) and with three or four discal setiferous pores also on interval 5 (absent from + +A. olsoufieffi + +members); and legs dark, except pale at base of femora and on trochanters (legs pale thoughout in + +A. olsoufieffi + +). + + +In addition, the +holotype +of + +A. ranomafanae + +is unique among members of all Malagasy odcanthine species in having the elytra distinctly convex throughout, except in the area of the transverse depression, elytral intervals flat, and elytral striae 3 to 7 effaced except in the transverse depression and apically, where they are evident. Striae 1 and 2 are very shallowly impressed but evident (stria 2 less so) in and posterior to the transverse depression. The form and location of the transverse depression of the elytra and the nearly effaced striae are similar to these features of members and genus + +Mimocolliuris +Liebke (1933) + +of the eastern Palearctic and Oriental Regions. + + + +FIGURE +16. Digital images of holotype female of + +Archicolliuris ranomafanae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The unique +holotype +was collected in a flight intercept-yellow pan trap on an island in a stream in montane rainforest at an elevation of +1100 m +. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573066FFB1FF8EFE48BDA50C48.xml b/data/0A/3E/26/0A3E26573066FFB1FF8EFE48BDA50C48.xml new file mode 100644 index 00000000000..7951731b7f6 --- /dev/null +++ b/data/0A/3E/26/0A3E26573066FFB1FF8EFE48BDA50C48.xml @@ -0,0 +1,346 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Thysanotus bimaculatoides +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 17–18 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.16A–B +), a male, in CAS, labeled: “ +MADAGASCAR + + + +Ranomafana NP. Mangevo plot +1600m +20.4.2005 +”/ “240” [handwritten label]/ “ +HOLOTYPE + + + +FIGURE +17. A-D. Digital images of holotype male of + +Thysanotus bimaculatoides +Kavanaugh & Rainio + +sp. nov. + +A–D. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + +FIGURE +18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. A–B. Holotype male of + +Thysanotus bimaculatoides +Kavanaugh & Rainio + +sp. nov. +; B, D. + +Thysanotus spinosus +Alluaud + +(from Ranomafana National Park). Scale lines = 0.5 mm. + + + + +Thysanotus bimaculatoides +Kavanaugh & Rainio + +sp. n. +2015” [red label]. +Paratypes +(total of 2): +1 female +(in +MNHN +) labeled “ +MADAGASCAR +Ranomafana NP. +Mangevo +plot + +1600m + + +22.4.2005 + +”/ “240” [handwritten label]; +1 female +(in +CAS +) labeled “CASENT 8069538”/ “ +MADAGASCAR +: +Province +Fianarantsoa +, +Forêt d’Ambalagoavy Nord +, +Ikonga +, +Abatombe + +Nov-2000 + +”/ “ +21°49’39”S +, +47°20’20”E +Calif. Acad. of Sciences +colls: +R. Harin’Hala +& +M.E. Irwin +, +malaise trap + +625m + +MA-01-12-01”. Both +paratypes +also bear the following label: “ +PARATYPE + +Thysanotus bimaculatoides +Kavanaugh & Rainio + +sp. n. +2015” [yellow label]. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +bimaculatoides + +, is an adjective derived from the Latin words, +bis +, meaning two, and +macula +, meaning spot or spotted, and the Greek word +eidos +, meaning like. The name refers to the similarity between members of this species and those of + +Thysanotus bimaculatus +Deuve (2010) + +. + + + +RECOGNITION + +.— Size slightly less than average for genus, SBL of male = +5.7 mm +, of females = +5.3 mm +. Members of this species ( +Fig. 17A +) key to + +Thysanotus bimaculatus +Deuve (2010) + +in Deuve’s key based on the following combination of features: elytra shiny, intervals smooth, elytral microsculpture indistinct (comprised of faintly impressed microlines and markedly transverse sculpticells, each elytron with a large, pale macula near the base on intervals 2 to 4, also extended partially onto interval 1 and to scutellum at basomedial part of macula, elytral apex bluntly round- ed, not toothed. Members of + +T. bimaculatoides + +differ from the +holotype +of + +T. bimaculatus + +in having slightly larger body size (SBL of +holotype +female of + +T. bimaculatus + += +4.9 mm +), pronotum slightly narrower, ratio of PW/PL = 1.1 (ratio of PW/PL = +1.2 in + +T. bimaculatus + +), narrower basally with subbasal sinuation of the lateral margin slightly longer and deeper and hind angles sharp- er, rectangular to distinctly acute (compare with +Deuve 2010 +, +Fig. 17 +). Deuve did not have a male specimen of + +T. bimaculatus + +, so we cannot compare male genitalia of these two species. However, for future comparative purposes, we provide digital photographs of the genitalia ( +Figs. 18A,C +) of the +holotype +of + +T. bimaculatoides + +as well as those of a male of + +Thysanotus spinosus +( +Alluaud 1936c +) + +( +Figs. 18B,D +) [males of this species previously unknown], to complement the illustrations of other + +Thysanotus +species + +provided by Jeannel (1949) and +Deuve (2010) +. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from two localities, +52 km +apart, in central +Fianarantsoa Province +. Both localities are on the eastern slope off the central plateau. + + + +HABITAT +DISTRIBUTION + +.— + +The +holotype +and +one paratype + + +were collected by hand in the +Mangevo area +, about + +1600 m + +distance in from the edge of montane rainforest, at an elevation of + +1600 m + +. +The +other +paratype +was collected in a +malaise trap +in montane rainforest at an elevation of + + +650 m + +. + + + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573069FF82FF8EFD07BCD40E48.xml b/data/0A/3E/26/0A3E26573069FF82FF8EFD07BCD40E48.xml new file mode 100644 index 00000000000..ee3b4de6b46 --- /dev/null +++ b/data/0A/3E/26/0A3E26573069FF82FF8EFD07BCD40E48.xml @@ -0,0 +1,313 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Trichinillus +( +Mallopelmus +) +ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 10 + + + +Mallopelmus +n. sp. +1 + +; Rainio 2009: 32, Rainio 2012: 74 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 10A–B +), a male, in CAS, labeled: “ +MADAGASCAR +Ranomafana NP. +Buffer +zone, degraded forest + +14.10.2002 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “ +HOLOTYPE + +Trichinillus +( +Mallopelmus +) +ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(only 1): a female (in +MNHN +) labelled “ +MADAGASCAR +Ranomafana NP. +Talatakely +, trail B, + + +600 m + +. + + +05.03.2004 + +”/ “collected by hand in daytime from dead leaves in bushes, +F. Ratalata +and +J. Rainio +collectors”/ “ +PARATYPE + +Trichinillus +( +Mallopelmus +) +ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Slightly smaller than average for subgenus, SBL male = +7.2 mm +, female = +7.5 mm +. The following combination of features identify members of this species as coelostomine pterostichines of the genus + +Trichinillus + +, subgenus + +Mallopelmus + +: antennae with antennomere 3 attached symmetrically or nearly symmetrically to antennomere 2; mentum with deep emargination, apex of median mental tooth distinctly posterior to apices of epilobes; elytron with epipleuron interrupted by internal plica subapically; parascutellar stria short but evident; striae smooth, impunctate; basal setiferious puncture situated at base of stria 2 + parascutellar stria; tarsomeres 1 to 4 of all legs with dense pads of setae ventrally; front and middle tarsomeres 1 to 3 with more or less faintly-defined longitudinal groove medially and depressed areas paralaterally on dorsal surface; male front tarsomeres 1 to 4 wide, dilated, not latero- or medioapically toothed; aedeagus of male with right face dorsal in repose, right paramere conchoid, left paramere reduced, short, apically digitiform. Of the three previously described species represented in Jeannel’s (1948) key to + +Mallopelmus +species + +, members of + +T. ranomafanae + +( +Fig. 10A +) are most similar to + +Trichinillus abacetoides +(Alluaud 1936) + +members based on the following shared features: head with temporal area long, nearly as long as the diameter of the eye, and joined to the neck region at a very obtuse (about 165°) angle; eyes moderately convex and projected; pronotum with basal area distinctly and abruptly depressed relative to disc; and elytral base without margination medial to base of stria 6. Members of the other two species, + +Trichinillus dactyleuryoides +(Alluaud 1936) + +and + +Trichinillus perrieri +(Jeannel 1948) + +, differ in having the head with the temporal area shorter, about one-third as long as the diameter of the eye, and joined to the neck region at a moderately obtuse (about 135°) angle, eyes more convex and projected, pronotum not or less depressed, especially medially, and margination of the elytra base more varied, extended medially to the base of stria +3 in +most individuals). The apex of the median lobe of the male genitalia is also differently shaped in each of these species (compare with Figs. 191c-d and 191f-g, respectively, in Jeannel 1948). Members of + +T. ranomafanae + +differ from those of + +T. abacetoides + +in having: slightly larger size (SBL = +6.4 mm +in + +T. abacetoides + +); pronotum with shallow but distinct subbasal sinuation of the lateral margin (absent from + +T. abacetoides + +members); elytra relatively shorter and wider, especially across the base, and humeri broadly rounded and not at all sloped (humeri more slope in + +T. abacetoides + +, and median lobe of male genitalia with apex straighter, slightly bent ventrally toward apex in lateral aspect ( +Fig. 10C +) and slightly deflect right in dorsal view ( +Fig. 10D +) (not recurved dorsally and then ventrally in lateral aspect and straight in dorsal aspect as in + +T. abacetoides + +males; compare with Fig. 191e in Jeannel 1948) + + + +FIGURE +10. Digital images of holotype male of + +Trichinillus +( +Mallopelmus +) +ranomafanae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The male +holotype +of + +T. ranomafanae + +was collected in degraded secondary montane rainforest at an elevation of +800 m +, near a village in the peripheral zone of Ranomafana National Park about1000 meters from the main gate. The female +paratype +was collected in secondary montane rainforest in the Talatakely area, at an elevation of +900 m +, at the +600 m +mark along Trail B. Vegetation in this area was dominated by young trees (DBH < +10 cm +) and guava ( + +Psidium cattleianum + +). + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657306BFF84FF8EFBE3BF8A0985.xml b/data/0A/3E/26/0A3E2657306BFF84FF8EFBE3BF8A0985.xml new file mode 100644 index 00000000000..9404029a322 --- /dev/null +++ b/data/0A/3E/26/0A3E2657306BFF84FF8EFBE3BF8A0985.xml @@ -0,0 +1,328 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Chlaenius kathrynae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 11 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.11A–B +), a male, in CAS, labeled: “CASENT 1002782”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Vohiparara area +, + +1150 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.24032°S +/ +47.39399°E +, Stop# DHK-01-002, +D.H. & K.M. Kavanaugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, Ranaivosolo,”/ “ +E.F. Randrianifirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime by treading marsh vegetation and shore”/ “ +HOLOTYPE + +Chlaenius kathrynae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(total of 8): +4 males +(in +CAS +, +MNHN +and +MZF +) and + + +1 female +(in +CAS +) labeled same as +holotype +except “CASENT 1002783”, “CASENT 1002784”, “CASENT 1002786”, “CASENT 1002787”, and “CASENT 1002785”, respectively + +; + +1 male +( +CAS +) labeled “CASENT 1049008”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Vohiparara area +, + +1050 m + +, mixed tropical forest, + +23 April 1998 + +,”/ “ +21°13.6’S +/ +47°23.0’E +,”/ “Stop # 98-71, +D.H. Kavanaugh +collector collected in soil cracks, under dirt clods, and under root clods in abandoned rice paddy area” + +; + +2 females +(in +MTEC +) labeled “ +MADAGASCAR +: +Fianaran. Pr. Ranomafana +N.P. HQ. area +21°15’24”S +47°25’15”E + +25NOV1994 + +, at night +M.A. Ivie +& +D.A. Pollack +”. All +paratypes +also bear the following label: “ +PARATYPE + +Chlaenius kathrynae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + +FIGURE +11. Digital images of holotype male of + +Chlaenius kathrynae +Kavanaugh & Rainio + +sp. nov. +A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +kathrynae + +, is a noun in the genitive case, derived from the first given name of Kathryn May Kavanaugh, daughter of DHK and one of the collectors of the +type +series. + + + +RECOGNITION + +.— Size slightly below average for genus, SBL of males = 10.2–10. +8 mm +, of females 9.8–11. +1 mm +. Members of this species ( +Fig. 11A +) share the following features, which place them in genus + +Chlaenites + +sense Jeannel (1948): antennomere 3 longer than antennomere 4; maxillary palpi setose, penultimate labial palpomeres 4 or 5 setose; pronotum with basolateral setae inserted distinctly anterior to the hind angles; elytra with pubescence restricted to near the striae on disk ( +i.e. +, not present on the centers of intervals), except more generally and densely present on lateral intervals and near apices; and tarsi asetose dorsally, tarsomeres 5 with two rows of stout setae ventrally. In Jeannel’s (1949) key to subgenera of + +Chlaenites + +, members of this species key best, although not fully, to subgenus + +Chlaeniostenus +Kuntzen (1919) + +, which is now considered a subgenus of + +Chlaenius +( +Lorenz 2005 +) + +. Shared with other Malagasy members of this subgenus are the following features: pronotum relatively narrow, more or less cordiform, narrowed basally, with the anterior angles not projected anterior of the basal margin and the anterolateral areas of the pronotum distinctly curved ventrally; and the setose punctures along the lateral margins of the elytral intervals not or only faintly evident, most clearly evident in the apical half only. Jeannel included three other species in this subgenus; namely, + +Chlaenius attenuatus +Klug (1833) + +, + +Chlaenius subovatus +Chaudoir (1876) + +, and + +Chlaenius sellatus +Dejean (1831) + +. Members of + +C. kathrynae + +differ from those of the other species in lacking any trace of the pale lateral and apical elytral border distinct in members of the other three species. They are also smaller than members of the other species (SBL in the latter ranges from 11.9 to 15. +3 mm +), have slightly broader and less cordiform pronota and males have genitalia ( +Figs. 11C–D +) that are very different in form from those of any of the other species (compare with Jeannel’s (1949) Figs. 388c-i) + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Most specimens of the +type +series were collected in open habitats at the edges of mixed tropical forest in the Vohiparara area at an elevation of +1050 m +. Several were collected by treading down vegetation at the edge of a marsh, and +one specimen +was found under loose dirt and root clods in an abandoned rice paddy area. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657306DFF86FF8EFB54BE4608B9.xml b/data/0A/3E/26/0A3E2657306DFF86FF8EFB54BE4608B9.xml new file mode 100644 index 00000000000..f76c5bd624b --- /dev/null +++ b/data/0A/3E/26/0A3E2657306DFF86FF8EFB54BE4608B9.xml @@ -0,0 +1,341 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Chlaenius robertae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 12 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.12A–B +), a male, in CAS, labeled: “CASENT 1002781”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Vohiparara area +, + +1150 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.24032°S +/ +47.39399°E +, Stop# DHK-01-002, +D.H. & K.M. Kavanaugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, Ranaivosolo,”/ “ +E.F. Randrianifirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime by treading marsh vegetation and shore”/ “ +HOLOTYPE + +Chlaenius robertae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(only 1): a female (in +CAS +) labeled “CASENT1049009”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Vohiparara area +, + +1050 m + +, mixed tropical forest, + +27 April 1998 + +,”/ “ +21°13.6’S +/ +47°23.0’E +,”/ “Stop # 98-91, +D.H. Kavanaugh +collector collected on bare soil at night in abandoned rice paddy area”/ “ +PARATYPE + +Chlaenius robertae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +robertae + +, is a noun in the genitive case, + + + +FIGURE +12. Digital images of holotype male of + +Chlaenius robertae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm. + + + +derived from the given name of Roberta L. Brett, former graduate student and technical assistant of DHK and one of the collectors of the +type +series. + + + +RECOGNITION + +.— Sized moderate for genus, SBL of male = +12.9 mm +, of female = +13.3 mm +. Members of this species ( +Fig. 12A +) exhibit the following features, which place them in genus + +Chlaenites +sensu Jeannel (1949) + +: antennomere 3 longer than antennomere 4; maxillary palpi setose; penultimate labial palpomeres 4 or 5 setose; pronotum with basolateral setae inserted distinctly anterior to the hind angles; elytra with pubescence restricted to near the striae on disk (i.e., not present on the centers of intervals), except more generally and densely present on lateral intervals and near apices; and tarsi asetose dorsally, tarsomeres 5 with two rows of stout setae ventrally. In Jeannel’s (1949) key to subgenera of + +Chlaenites + +, members of this species key to his new subgenus + +Chlaenitidius + +, which is now considered a junior synonym of subgenus + +Amblygenius +Laferté-Sénectère (1851) + +( +Lorenz 2005 +), in having the pronotum relatively broad and the setose punctures along the lateral margins of the elytral intervals slightly foveate, especially in the apical half apically. Member s of + +C. robertae + +differ distinctly from those of the each of the four species Jeannel included in this subgenus. They differ from members of + +Chlaenius cupreolus +Faimaire (1901) + +in having striae deeply impressed and intervals moderately convex (striae superficial only and intervals flat in + +C. cupreolus + +members), head with metallic green reflection but pronotum black without a trace of metallic reflection (both head and pronotum with green metallic reflection in + +C. cupreolus + +members), and elytra without a pale lateral and apical margin (a thin pale margin present in + +C. cupreolus + +members). They differ from members of + +Chlaenius allacteus +Alluaud (1919) + +, which is current classified in subgenus + +Oochlaenius +Alluaud (1933) + +( +Lorenz 2005 +), in having all elytral interval similarly and moderately convex (intervals 1, 3, 5 and 7 more convex and intervals 2, 4, 6 and 8 flat in + +C. allacteus + +members) and, again, elytra without a pale lateral and apical margin (a wide pale margin present in + +C. allacteus + +members). They differ from members of + +Chlaenius inaequalis +Faimaire (1901) + +in having the pronotum black without metallic reflection (metallic green reflection present on pronotum in + +C. inaequalis + +members), a longer and narrower elytral form (shorter and broader elytra in + +C. inaequalis + +members) and elytral epipleurae black (pale in + +C. inaequalis + +members). Overall, members of + +C. robertae + +are most similar to those the fourth species, + +Chlaenius lyperus +Jeannel (1949) + +, with which they share similar elytral shape, depth of striae and convexity of intervals. However, they differ from + +C. lyperus + +members in having a pronotum broader basally than the latter and legs that are pale in color (dark in + +C. lyperus + +members). Males of + +C. robertae + +also have genitalia ( +Figs. 12C–D +) that are markedly different in form those of the other four species (compare with Jeannel’s (1949) Fig. 386a-d). + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Both specimens of the +type +series were collected in open habitats at the edges of mixed tropical forest in the Vohiparara area at elevations ranging from +1050 to 1150 m +. One was collected in daytime by treading down vegetation at the edge of a marsh, the other was found at night running on bare soil in an abandoned rice paddy. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657306EFF87FF8EFF49BEB4082C.xml b/data/0A/3E/26/0A3E2657306EFF87FF8EFF49BEB4082C.xml new file mode 100644 index 00000000000..694d8bcd0ef --- /dev/null +++ b/data/0A/3E/26/0A3E2657306EFF87FF8EFF49BEB4082C.xml @@ -0,0 +1,239 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Omphreoides ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 13 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs.13A–B +), a female, in NMNH, labeled: “ +MADAGASCAR +: Prov. +Fianarantsoa +, +7 km +W +Ranomafana +, + +900 m + + +17-22 February 1990 + +W. E. Steiner +”/ “at black light in montane rainforest near river and stream”/ “ +HOLOTYPE + +Omphreoides ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size larger than average for genus, SBL = +13.4 mm +. The unique +holotype +female of + +O. ranomafanae + +( +Fig. 13A +) is clearly unlike members of any described species of + +Omphreoides + +, with several features intermediate between the four typical species and + +O. quodi + +. The head of the +holotype +( +Fig. 13C +) is longer and more slender than that of females of any other species except + +O. quodi + +, which has an even longer and narrower head (see Jeannel 1949, Fig. 364). In + +O. ranomafanae + +, the tempora are straight, parallel in anterior their two-thirds, slightly more than twice as long as the diameter of eye, distinctly delimited posteriorly by narrowly round angles. The dorsal longitudinal grooves typical of all members of this genus, except those of + +Omphreoides bispinus +Fairmaire (1896) + +, are extended as sharply-defined grooves only about halfway from the point of insertion of the anterior supraorbital setae to that of posterior supraorbital setae and not continued on to base of the head as they are in + +Omphreoides bucculentus +Alluaud (1899a) + +and + +Omphreoides distinctus +Alluaud (1936) + +. Posterior to the sharply-defined segments, broad and shallow depressions extend posteriorly, arcuately convergent and nearly joined in the midline near the back of the head. The pronotum ( +Fig. 13C +) is subquadrate, longer than wide (ratio PW/PL = 0.75), with a wavy diagonal row of sparse, coarse punctures (seen also in + +O. distinctus + +and + +O. quodi + +members but absent from those of + +O. bucculentus + +and + +O. bispinus + +) extended from near the anterior transverse impression paramedially to the basal foveae posteriorly; lateral pronotal margins with short but distinct sinuations anterior to slightly obtuse hind angles; median longitudinal impression deep, wider than in all other + +Omphreoides + +except + +O. quodi + +(in which the impression is even wider) and sparsely but coarsely punctate; lateral explanation narrow (narrower at middle than that in + +O. bucculentus + +members and similar to that in + +O. distinctus + +member. The elytra are relatively wider than in all other species; each elytron with a straight apical spine in line with interval 3. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The unique +holotype +was collected at ultraviolet light in montane rainforest near the junction of the Namorona River and a small tributary stream. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573071FF9AFF8EFD4FB9350CCD.xml b/data/0A/3E/26/0A3E26573071FF9AFF8EFD4FB9350CCD.xml new file mode 100644 index 00000000000..b262a262839 --- /dev/null +++ b/data/0A/3E/26/0A3E26573071FF9AFF8EFD4FB9350CCD.xml @@ -0,0 +1,272 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Caelostomus rotundiformis +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 5 + + + +Caelostomus +n. sp. +1 + +; Rainio 2009: 31, Rainio 2012: 62 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 5A–B +), a female, in CAS, labeled: “ +MADAGASCAR +Ranomafana NP., +Vatoharanana +, Trail K at + +50 m + +mark, + +07.06.2000 + +”/ “collected by hand in daytime, +F. Ratalata +& +J. Rainio +collectors”/ “13.” [handwritten label]/ “7.6.00 K 50” [handwritten label]/ “ + +HOLOTYPE + +Caelostomus rotundiformis +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + +Paratypes +(only 1): a female (in +MNHN +) labeled “ +MADAGASCAR +Ranomafana NP. Talatakely, C1450 + +11.3.2004 + +Johanna Rainio +leg.”/ “112.” [handwritten label]/ “ + +PARATYPE + +Caelostomus rotundiformis +Kavanaugh & Rainio + +sp. n. +2015” [yellow label]. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +rotundiformis + +, is an adjective derived from the Latin words, +rotundus +, meaning round, and +forma +, meaning form or shape. The name refers to the very short, broad, rounded form of members of this species. + + + +RECOGNITION + +.— Size moderately small for genus, SBL = 5.4–5. +5 mm +(females only), body form ( +Fig. 5A +) short and broad, ratio SBL/EW = 1.97–2.01. This species is undoubtedly a member of genus + +Caelostomus + +as conceived by Jeannel (1948), although we cannot confirm this based on his key because he used male features only in two couplets to distinguish + +Caelostomus + +members from those of several other genera and we have only female specimens. However, we examined specimens of all species in all the other Malagasy caelostomine genera while at MNHN and confirmed that this species represents none of them. Observable female features, including body form short and broad, frontal furrows simple, not doubled, parascutellar striae of elytra indistinct, basal setiferous pore situated at base of elytral stria 3, tarsomeres not densely setose ventrally, and anterior tarsomere 4 not bilobed apically ( +Fig. 5C +), are consistent with + +Caelostomus + +females. Within this genus, members of + +C. rotundiformis + +share the following features with those of other species of the nominate subgenus: head with frons and frontal furrows smooth, impunctate; elytral base distinctly margined from humerus medially to base of stria 3 or 2; elytral striae 6 and 7 deeply impressed throughout, not effaced apically; lateral elytral intervals narrow, moderately convex; apical part of umbilicate series of setiferous pores not inserted in a deep groove; and female apical abdominal ventrite with (two or) three pairs of setae subapically. However, members of this species confound Jeannel’s (1948) key to species of + +Caelostomus + +s. str. +Their short and broad elytra (ratio EL/EW = 1.13–1.15) are shared with members of the + +convexiusculus + +species group, but their elytral striae are impuncate, whereas those of other species group members are markedly punctate throughout their length. Within the group, their small size (less than 6.0 mm) is shared with + +Caelostomus humilis +( +Tschitschérine 1903 +) + +, + +Caelostomus minisculus +Straneo (1940) + +, and + +Caelostomus minutissimus +Jeannel (1948) + +; however, members of these three species are even smaller (SBL = 5.0 mm or less) and, again, have markedly punctate elytral striae. The striae in + +C. rotundiformis + +members are smooth, without evident punctures, and the shape of the pronotum ( +Fig. 5A +) is markedly different, with lateral margins evenly arcuate throughout their length and hind angles markedly obtuse. This species probably represents a distinct species group of its own within subgenus + +Caelostomus + +in +Madagascar +. + + + +FIGURE +5. Digital images of holotype female of + +Caelostomus rotundiformis +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The +holotype +of this species was collected by hand in daytime in primary montane rain forest in the Vatoharanana area at an elevation of +1200 m +, along Trail K at the +50 meter +mark. The +paratype +specimen was collected in a pitfall trap in secondary montane rain forest in the Talatakely area at an elevation of +900 m +, along Trail C at the +1450 meter +mark. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573073FF9BFF8EFD4FBCF10BA1.xml b/data/0A/3E/26/0A3E26573073FF9BFF8EFD4FBCF10BA1.xml new file mode 100644 index 00000000000..c14da814ba1 --- /dev/null +++ b/data/0A/3E/26/0A3E26573073FF9BFF8EFD4FBCF10BA1.xml @@ -0,0 +1,266 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Dactyleurys minimus +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 6 +, +8A +, +9A, 9D + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 6A–B +), a male, in CAS, labeled: “ +MADAGASCAR +, Ranomafana NP, Talatakely, X at + +130 m + +mark + +12.4.2005 + +”/ “collected by hand in daytime, +F. Ratalata +& Johanna Rainio collectors”/ “ +HOLOTYPE + +Dactyleurys minimus +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +minimus + +, is the Latin adjective meaning smallest, a reference to fact that the known adult male of this species is smaller than males of any of the other known species that we have examined. + + + +RECOGNITION + +.— Smallest adult known for the genus, SBL = +6.9 mm +. The following combination of features identify members of this species as coelostomine pterostichines of the genus + +Dactyleurys + +: antennae with antennomere 3 attached symmetrically or nearly symmetrically to antennomere 2; mentum with deep emargination, apex of median mental tooth distinctly posterior to apices of epilobes; elytron with epipleuron interrupted by internal plica subapically; parascutellar stria not evident; basal setiferious puncture situated at base of stria 3; tarsomeres 1 to 4 of all legs with dense pads of setae ventrally; front and middle tarsomeres 1 to 3 smoothly convex dorsally; male front tarsomeres 1 to 4 wide, dilated, not latero- or medioapically toothed; aedeagus of male with right face dorsal in repose, right paramere conchoid, left paramere reduced, short, apically digitiform. Members of + +D. minimus + +( +Fig. 6A +) differ from those of + +D. anomalus + +in the following features: smaller size (SBL = +6.9 mm +compared with 7.7–9. +4 mm +in + +D. anomalus + +); pronotum ( +Fig. 8A +) relatively narrower, widest at middle (relatively wider ( +Fig. 8C +) and clearly widest anterior to middle in + +D. anomalus + +); elytra faintly but evidently iridescence and elytral microsculpture markedly transverse with sculpticells great than 2.5 times as wide as long (elytra not iridescent and elytral microsculpture moderately transverse with sculpticells less than 2.5 times as wide as long in + +D. anomalus + +); and median lobe of male genitalia ( +Fig. 9A +) with shaft abruptly bent ventrally in apical fourth with apex nearly straight in lateral aspect (shaft nearly straight ventrally ( +Fig. 8C +) in apical fourth with apex slightly recurved dorsally in + +D. anomalus + +), apex narrowly rounded in dorsal aspect ( +Fig. 9D +) (more broadly rounded ( +Fig. 9F +) in + +D. anomalus + +). Members of + +D. minimus + +differ from those of the other new species, + +Dactyleurys ranomafanae + +, in the following features: smaller size (SBL = 7.7–8. +3 mm +in + +D. ranomafanae + +); pronotum ( +Fig. 8A +) relatively narrower and with evenly rounded lateral margin (slightly wider ( +Fig. 8B +) and with lateral margin slightly straighter in basal one-third in + +D. ranomafanae + +); and median lobe of male genitalia ( +Fig. 9A +) with apex nearly straight in lateral aspect (apex slightly recurved dorsally ( +Fig. 9C +) in + +D. ranomafanae + +), apical part of shaft ( +Fig. 9D +) slightly deflected right in dorsal aspect (apical part of shaft ( +Fig. 9E +) straight in + +D. ranomafanae + +). + + + +FIGURE +6. Digital images of holotype male of + +Dactyleurys minimus +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The unique +holotype +was collected by hand in daytime in secondary montane rain forest in the Talatakely area, at an elevation of +900 m +, at the +130 meter +mark along Trail “X”. Vegetation in that area consisted of bamboo, small young trees and some larger trees with diameters at breast-height (DBH) of +20–30 cm +. Tree trunks and branches were heavily laden with epiphytes and lianas. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E26573075FF9FFF8EFF49BCE50B42.xml b/data/0A/3E/26/0A3E26573075FF9FFF8EFF49BCE50B42.xml new file mode 100644 index 00000000000..47634aeefc6 --- /dev/null +++ b/data/0A/3E/26/0A3E26573075FF9FFF8EFF49BCE50B42.xml @@ -0,0 +1,764 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +0068-547X + + + + + + + +Dactyleurys ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 7 +, +8B +, +9B, 9E + + + +Dactyleurys +n. sp. +1 + +; Rainio 2009: 31, Rainio 2012: 62, Rainio 2013: 95 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 7A–B +), a male, in CAS, labeled: “CASENT 1009744”/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +26 April 1998 + +”/ “ +21°15.3’S +/ +47°25.9’E +, Stop #98-83 +D.H. Kavanaugh +collector collected by beating + +Pandanus + +tree boluses with debris”/ “ +HOLOTYPE + +Dactyleurys ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label] + +. + +Paratypes +(a total of 35): +1 male +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Mangevo +, plot 0 m., + +21.4.2005 + +” + +; + +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Mangevo +, plot + + +100 m + +. + +, + +21.4.2005 + +” + +; + +2 females +(in +CAS +) labeled same as +holotype +except “CASENT 1009742” and “CASENT 1009743”, respectively + +; + +3 males +and +2 females +(in +CAS +) labeled “CASENT 1009737”, “CASENT 1009738”, “CASENT 1009739”, “CASENT 1009740” and “CASENT 1009741”, respectively/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +25 April 1998 + +”/ “ +21°15.3’S +/ +47°25.9’E +, Stop #98-78 +D.H. Kavanaugh +collector beaten from vegetation and suspended dead leaves” + +; + +3 males +and +2 females +(in +CAS +and +MNHN +) labeled “CASENT 1009734”, “CASENT 1009735”, “CASENT 1009736”, “CASENT 1009732” and “CASENT 1009733”, respectively/ “ +MADAGASCAR +, +Fianarantsoa Province +, +Ranomafana National Park +, +Talatakely area +, + +900 m + +, mixed tropical forest, + +2-22 January 2001 + +,”/ “ +21.25041°S +/ +47.41945°E +, Stop# DHK-01-001, +D.H. & K.M Kavanaugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +, +R. Ranaivosolo +,”/ “ +E.F. Randrianirina +, +N. Rasoamananana +, +T.J. Ravelomanana +, and +H.C. Raveloson +collectors”/ “collected in daytime by beating live understory vegetation and/or dead leaves and other suspended debris” + +; + +1 male +and +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, + +26.2.2001 + +” + +; + +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail B + +1150 m + +., in + +Pandanus +sp. +8 + +.6.2000” + +; + +1 male +and +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail BF + + +380 m + +. + +, in + +Pandanus +sp. + + +16.6.2000 + +” + +; + +1 male +and +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail BM + + +200 m + +. + +, in + +Pandanus +sp. + + +16.6.2000 + +” + +; + +1 male +and +2 females +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail FBF + + +50 m + +. + +, in + +Pandanus +sp. +6 + +.6.2000” + +; + +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail FBF + + +380 m + +. + +, in + +Pandanus +sp. +6 + +.6.2000” + +; + +2 females +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail SP + + +50 m + +. + +, + +22.3.2001 + +” + +; + +1 female +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail SP + + +50 m + +. + +, + +9.2.2005 + +” + +; + +1 male +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Talatakely +, trail X + + +130 m + +. + +, + +12.4.2005 + +” + +; + +1 male +(in +MNHN +) labeled “ +MADAGASCAR +Ranomafana +NP. +Valohoaka +, trail F + + +900 m + +. + +, + +28.2.2001 + +” + +; + +1 male +(in +MZF +) labeled “ +MADAGASCAR +Ranomafana +NP. +Valohoaka +, trail F + + +850 m + +. + +, + +16.10.2002 + +” + +; + +1 male +(in +MNHN +) labeled “ +MADAGASCAR +Ranomafana +NP. +Valohoaka +, trail F + +850 m + +down., + +18.12.2004 + +” + +; + +1 female +(in +MNHN +) labeled “ +MADAGASCAR +Ranomafana +NP. +Valohoaka +, trail F + + +850 m + +. + +, + +25.1.2005 + +” + +; + +2 males +and +1 female +(in +MNHN +and +MZF +) labeled “ +MADAGASCAR +Ranomafana NP. +Valohoaka +, trail F + + +850 m + +. + +, + +26.1.2005 + +”. +All +paratypes +also bear the following label: “ +PARATYPE + +Dactyleurys ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [yellow label] + +. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Size average for genus, SBL of males = 7.7–8. +2 mm +, of females = 7.8– +8.4 mm +. The following combination of features identify members of this species as coelostomine ( +Fig. 8B +) relatively broader and with lateral margins slightly straighter in basal one-third (slightly narrower and with lateral margins more evenly rounded ( +Fig. 8A +) in + +D. minimus + +); and median lobe of male genitalia ( +Fig. 9B +) with apex slightly recurved dorsally in lateral aspect (apex ( +Fig. 9A +) nearly straight in + +D. minimus + +), apical part of shaft ( +Fig. 9E +) straight (apical part of shaft slightly deflected right ( +Fig. 9D +) in + +D. minimus + +). + + + +FIGURE +7. Digital images of holotype male of + +Dactyleurys ranomafanae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + +FIGURE +8. Digital images of pronota of + +Dactyleurys +spp. + +, dorsal aspect. A. Holotype of + +D. minimus +Kavanaugh & Rainio + +sp. nov. +; + + +B. Holotype of + +D. ranomafanae +Kavanaugh & Rainio + +sp. nov. +; + + +C. + +D. anomalus +Tschitschérine. Scale + +lines = 1.0 mm + + + + +FIGURE +9. Digital images of aedeagus of male genitalia of + +Dactyleurys +spp. A + +–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of + +D. minimus +Kavanaugh & Rainio + +sp. nov. +; B, E. Holotype of + +D. ranomafanae +Kavanaugh & Rainio + +sp. nov. +; C, F. + +D. anomalus +Tschitschérine. Scale + +lines = 0.5 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— Specimens of this species have been collected in both primary and secondary montane rainforest throughout RNP, at elevations ranging from of +900 to 1180 m +. Most of the specimens were collected by beating vegetation, especially plants of + +Pandanus +sp. + +, and suspended accumulations of dead vegetative debris in shrubs and understory trees. One was collected in a malaise trap, another sifted from forest litter at the primary forest site, and another by hand at the secondary forest site. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657307DFF95FF8EFCD9BC8D0B62.xml b/data/0A/3E/26/0A3E2657307DFF95FF8EFCD9BC8D0B62.xml new file mode 100644 index 00000000000..bc16719846f --- /dev/null +++ b/data/0A/3E/26/0A3E2657307DFF95FF8EFCD9BC8D0B62.xml @@ -0,0 +1,309 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +10.5281/zenodo.13799439 +0068-547X +13799439 + + + + + + + +Brachypelus ranomafanae +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figures 2–3 + + + +Brachypelus +n. sp. +1 + +; Rainio 2009: 31 (informal designation). + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 2A–B +), a female, in CAS, labeled: “ +MADAGASCAR +Ranomafana NP. Talatakely, X, + + +130 m + +. + + +15.1.2001 + +”/ “ +collected in pitfall trap +on forest floor, +F. Ratalata +& +J. Rainio +collectors”/ “32.” [handwritten label]/ “ + +HOLOTYPE + +Brachypelus ranomafanae +Kavanaugh & Rainio + +sp. n. +2015” [red label]. + + + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +ranomafanae + +, is a noun in apposition, derived from the name of the national park in which the +type +was collected. + + + +RECOGNITION + +.— Large for genus, SBL = +6.8 mm +. Members of this species ( +Fig. 2A +) are most similar to those of + +Brachypelus obesus +Putzeys (1866) + +, with which they share large size, only six setae on the labrum, and a single discal setiferous pore on elytral interval 3 (at apical one-fifth). They differ from + +B. obesus + +members in the following: transverse fronto-clypeal suture present and deeply impressed ( +Fig. 3A +) (absent from + +B. obesus + +members); frons with longitudinal frontal furrows narrower and shallower than in + +B. obesus + +; posterior lateral margins of the eyes broadly round- ed (more narrowly and angularly rounded in + +B. obesus + +members); elytral base ( +Fig. 3B +) straight between intervals 3 and obliquely sloped laterally from interval 4 (elytral base straight between intervals 4 and gradually rounded laterally from interval +3 in + +B. obesus + +members); elytra with lateral margin distinctly toothed at humerus (smoothly rounded at humerus, without a tooth in + +B. obesus + +members); and elytral intervals slightly convex apically (nearly flat in + +B. obesus + +members). Others species with members of comparably large size include + +Brachypelus rolandi +Bulirsch et al. (2005) + +, + +Brachypelus betsileo +Bulirsch et al. (2005) + +, + +Brachypelas fischeri +Bulirsch et al. (2005) + +and + +Brachypelus janaki +Bulirsch and Moravec (2009) + +. However, + +B. ranomafanae + +members differ from those of + +B. rolandi + +in having the labrum with only six setae (seven in + +B. rolandi + +members), elytra widest at middle (anterior to middle in + +B. rolandi + +members), and elytral intervals moderately convex throughout (flattened apical in + +B. rolandi + +members). Members of both + +B. betsileo + +and + +B. fischeri + +have seven labral setae (whereas those of + +B. ranomafanae + +have only six) and lack discal setiferous pores (those of + +B. ranomafanae + +have a single setiferous pore in the apical one-fifth on interval 3. Finally, + +B. ranomafanae + +members differ from those of + +B. janaki + +in having only six setae on the labrum (seven labral setae in + +B. janaki + +members), the elytral margin with a distinct humeral tooth (tooth absent from + +B. janaki + +members), and only a single discal setiferous pore on elytral interval 3, with intervals 5 and 7 without such punctures (five or more such punctures on intervals 3, 5 and +7 in + +B. janaki + +members). + + + +FIGURE +2. Digital images of holotype female of + +Brachypelus ranomafanae +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm. + + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— The unique +holotype +was collected in a pitfall trap placed in secondary montane rain forest in the Talatakely area, at an elevation of +900 m +, at the +130 meter +mark along Trail “X”. Vegetation in that area consisted of bamboo, small young trees and some larger trees with diameters at breast-height (DBH) of +20–30 cm +. Tree trunks and branches were mostly covered with epiphytes and lianas. + + + + \ No newline at end of file diff --git a/data/0A/3E/26/0A3E2657307FFF98FF8EFA4EBF6C0FAE.xml b/data/0A/3E/26/0A3E2657307FFF98FF8EFA4EBF6C0FAE.xml new file mode 100644 index 00000000000..867670d18b5 --- /dev/null +++ b/data/0A/3E/26/0A3E2657307FFF98FF8EFA4EBF6C0FAE.xml @@ -0,0 +1,234 @@ + + + +Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar + + + +Author + +Kavanaugh, David H. +Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) + + + +Author + +Rainio, Johanna +Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com + +text + + +Proceedings of the California Academy of Sciences + + +2016 + +2016-04-29 + + +63 + + +7 + + +201 +268 + + + +journal article +10.5281/zenodo.13799439 +0068-547X +13799439 + + + + + + + +Caelostomus latus +Kavanaugh and Rainio + +, +sp. nov. + + + + + + +Figure 4 + + + + + +TYPE +MATERIAL + +.— + +Holotype +( +Figs. 4A–B +), a female, in CAS, labeled: “ +MADAGASCAR + + + +Ranomafana NP. Talatakely, Trail C, + +350 m +. + +10.5.2005 +”/ “ +HOLOTYPE + +Caelostomus latus + + + +Kavanaugh & Rainio +sp. n. +2015” [red label]. + + + +TYPE + +LOCALITY + +.— +Madagascar +, +Fianarantsoa Province +, +Ranomafana National Park + +. + + + +FIGURE +4. Digital images of holotype female of + +Caelostomus latus +Kavanaugh & Rainio + +sp. nov. +A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm. + + + + +DERIVATION +OF +SPECIES +NAME + +.— The species epithet, + +latus + +, is the Latin adjective meaning broad or wide, a reference to relatively broad pronotum of the +holotype +specimen and, presumably, other members of this species. + + + +RECOGNITION + +.— Size moderate for genus, SBL = +6.3 mm +. Based on Jeannel’s (1948) key, this species is a member of subgenus + +Caelostomus + +s. str. +It’s members ( +Fig. 4A +) share with those of other members of the subgenus the following features: head with frons and frontal furrows smooth, impunctate; elytral base distinctly margined from humerus medially to base of stria 3 or 2; elytral striae 6 and 7 deeply impressed throughout, not effaced apically; lateral elytral intervals narrow, moderately convex; apical part of umbilicate series of setiferous pores not inserted in a deep groove; and female apical abdominal ventrite with (two or) three pairs of setae subapically. Within the subgenus, the following features are shared only with members of two species of Jeannel’s + +convexiusculus + +group, namely + +Caelostomus convexiusculus +Tschitschérine (1899) + +and + +Caelostomus alluaudi +Jeannel (1948) + +: SBL greater than 6.0 mm, elytra short and broad, ratio EL/EW = 1.19, elytral disc convex; and elytral intervals distinctly punctate throughout, including in apical part. The +holotype +female of + +C. latus + +is slightly larger than members of these other two species, has a distinct and broad transverse depression between the eyes seen best in lateral view ( +Fig. 4C +) (depression absent from members of the other two species). The pronotum in the +holotype +of + +C. latus + +is much wider than in + +C. convexiusculus + +members, slightly wider than in + +C. alluaudi + +members, and less depressed anterolaterally than in members of either of the other species; and the pronotal basal foveae are straighter and more sharply defined laterally than in members of the other two species. The elytra are less abruptly rounded laterally and apically than in + +C. alluaudi + +members but similar to shape in to those in + +C. convexiusculus + +members. Finally, anterior tarsomeres 1 to 3 of the female +holotype +of + +C. latus + +are distinctly toothed medioapically ( +Fig. 4D +), a feature not seen in females of the other two species. + + + +GEOGRAPHICAL +DISTRIBUTION + +.— At present, known only from the +type +locality. + + + +HABITAT +DISTRIBUTION + +.— + +The unique +holotype +female was collected by hand in secondary montane forest in the +Talatakely area + +. + + + + \ No newline at end of file