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<document id="442C2B926DE6DEA71F8E6100EF10BB08" ID-DOI="10.3897/zse.100.130929" ID-ZooBank="4D6C62E4-224B-4AB5-A251-AA7ABEE66D74" ID-publisher-id="130929" URI-arpha="62A0369D-D7AA-53B4-8EA3-723D47A8742A" URI-zoobank="http://zoobank.org/4D6C62E4-224B-4AB5-A251-AA7ABEE66D74" article-type="research-article" checkinTime="1728062842149" checkinUser="pensoft" docAuthor="Jeratthitikul, Ekgachai, Sutcharit, Chirasak &amp; Prasankok, Pongpun" docDate="2024" docId="ACBEF56237745895AE444C49C4EE638E" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1333-1345" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/130929/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Isannaia occultata Jeratthitikul, Sutcharit &amp; Prasankok, 2024, sp. nov." docType="treatment" docUuid="F83569B6-8E5D-43BE-9DD4-4D44C9AD937B" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="1345" masterDocId="62A0369DD7AA53B48EA3723D47A8742A" masterDocTitle="Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin" masterLastPageNumber="1345" masterPageNumber="1333" pageNumber="1333" updateTime="1728063243823" updateUser="ExternalLinkService">
<mods:mods id="A2D96183ED7B2ABB9A3020F5E95B6919" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F998F8A7B86AF014672E3DCC998736BF">
<mods:title id="88A462495488A830883F82CB33BE48DC">Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin</mods:title>
</mods:titleInfo>
<mods:name id="64FD1EA0896AD5F64A7CECEA3E874055" type="personal">
<mods:role id="5FA9AFA3BC19E2814206D80A1992CA4C">
<mods:roleTerm id="20DF1F6DA4F8593765F34B51E59106D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AC8EA9C3B5554BB508443834D1CD7F7">Jeratthitikul, Ekgachai</mods:namePart>
<mods:nameIdentifier id="4ED10CF2F17266958AB1232A0268F5A8" type="ORCID">0000-0002-3477-9548</mods:nameIdentifier>
<mods:affiliation id="1A99BBB13FD9F294E840D1ADADF30601">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
</mods:name>
<mods:name id="03F58E64AB414E2B9FF25BAFBA608501" type="personal">
<mods:role id="05967417E477F362F9821851E22CF9C6">
<mods:roleTerm id="B03FD8375DEFBE1B217A7028A403D2BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEB1C78FB6FC79B7F1256412CAC34577">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="1C9DBB21A8D7FC6B3F17DB350688B9EF" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
<mods:affiliation id="2B52EBD3B5F4ACC19B1390C42C4F9CA5">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="D888331B6A0168991315AE23489AF49C">Prasankok, Pongpun</mods:namePart>
<mods:nameIdentifier id="B02F9E99E5658D56855A3F0F96D66172" type="ORCID">0000-0001-5967-223X</mods:nameIdentifier>
<mods:affiliation id="715FB855C02FD6ADB58619267C715DBB">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FED21FA1322CC03325DAE3EDC51D8684">text</mods:typeOfResource>
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<mods:titleInfo id="B07677CD93A24DAB9BAABB8E53E5E058">
<mods:title id="92158C41A310F85AB88A9377A77C71DC">Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part id="7CF58EDFF45F33C702F0028BE34744D5">
<mods:date id="533BD6E502A3BC3E99045B65E073C9C4">2024</mods:date>
<mods:detail id="BD241C15E9C16B37BC85212BAD7DE3E8" type="pubDate">
<mods:number id="45F53A7CD05C3C5CFB502584AEF5393A">2024-10-04</mods:number>
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<mods:detail id="441AC1CC98AC8ED6FAA14D5AC86C0691" type="volume">
<mods:number id="8B2047159CC0B36FEBD5290EE717C714">100</mods:number>
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<mods:number id="656465857ACC0A884B243FD77BB17065">4</mods:number>
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<mods:classification id="F066B8F5CCC70DC5554122E494C9CF54">journal article</mods:classification>
<mods:identifier id="BACD3852FFEF344D28F368BFF57DD97A" type="DOI">10.3897/zse.100.130929</mods:identifier>
<mods:identifier id="D238033DC78C3CA80F3BBA7FFB348CC9" type="ZooBank">4D6C62E4-224B-4AB5-A251-AA7ABEE66D74</mods:identifier>
</mods:mods>
<treatment id="ACBEF56237745895AE444C49C4EE638E" ID-DOI="http://doi.org/10.5281/zenodo.13891555" ID-Zenodo-Dep="13891555" ID-arpha="ACBEF562-3774-5895-AE44-4C49C4EE638E" ID-zoobank="https://zoobank.org/F83569B6-8E5D-43BE-9DD4-4D44C9AD937B" LSID="urn:lsid:zoobank.org:act:F83569B6-8E5D-43BE-9DD4-4D44C9AD937B" LSID-ZBK="urn:lsid:zoobank.org:act:F83569B6-8E5D-43BE-9DD4-4D44C9AD937B" httpUri="http://treatment.plazi.org/id/ACBEF56237745895AE444C49C4EE638E">
<subSubSection id="C726C456071D62251CBF782883D6CCE9" type="nomenclature">
<paragraph id="E03ACF4406958E1446ED6D13DF7F1F17">
<taxonomicName id="B4DC4CC0D29F6E10995F7D378404480B" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="occultata" status="sp. nov.">
<emphasis id="67299297093FEBBB73605B6B16CCCFFC" italics="true">Isannaia occultata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="606A5BBE293C4C6BC984A99AB78C0513" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D4DB40AB0D079396FCC3E7522A5EE306" type="description">
<paragraph id="1EF2E995D27492B0F998E15EE7D94364">
<figureCitation id="1286C2B31473A443FE44DC53A4C42036" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood (ML) tree of freshwater mussels in subfamily Gonideinae based on 1,935 - bp concatenated alignment dataset of COI and 16 S rRNA mitochondrial genes, and 28 S rRNA nuclear gene. The numbers on nodes represent the bootstrap support (BS) from the ML analysis and the bipartition posterior probability (bpp) from the BI analysis, and are shown as ML / BI. Scale bar indicates the branch length." figureDoi="10.3897/zse.100.130929.figure1" httpUri="https://binary.pensoft.net/fig/1148326">Figs 1</figureCitation>
,
<figureCitation id="8986F0A7A506DD6633A4982F60FBAEF1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing type localities of the new species: circle Lannanaia kokensis sp. nov., triangle Isannaia fortunata sp. nov., and square Isannaia occultata sp. nov. Map was generated using QGIS v 3.24. 3 with the outline of Mekong basin from the Freshwater Ecoregions of the World (Abell et al. 2008), river and lake topology from the HydroSHEDS database (https://www.hydrosheds.org), and the map raster data from the NASA EARTHDATA (https://www.earthdata.nasa.gov/)." figureDoi="10.3897/zse.100.130929.figure3" httpUri="https://binary.pensoft.net/fig/1148328">3</figureCitation>
,
<figureCitation id="663143D1A6A45AAF429FA860D71440B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shells of Isannaia gen. nov., showing the inner side of the right valve, the outer side of the left valve, and the dorsal view of both valves. A. Holotype (MUMNH - UNI 1950) and B. Paratype (MUMNH - UNI 1952) of Isannaia fortunata sp. nov.; C. Holotype (MUMNH - UNI 2895) and D. Paratype (MUMNH - UNI 2897) of Isannaia occultata sp. nov. Scale bars: 10 mm." figureDoi="10.3897/zse.100.130929.figure4" httpUri="https://binary.pensoft.net/fig/1148329">4 C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3SBG" type="type material">
<paragraph id="9C025281C52B6F78D9C2A2C9F657AA3A">
<heading id="388B99558A90C0B70E16D9254462F688" reason="title">Type material.</heading>
</paragraph>
<paragraph id="6BF064321A4CAA29FC90D14BECB3BA6D">
<materialsCitation id="F305ECBC1C6429A6386B9F22B644E775" collectionCode="MUMNH" collectorName="K. Wisittikoson" country="Thailand" county="Wang Sam Mo District" latitude="16.9886" location="Yang Stream" longLatPrecision="7" longitude="103.3638" municipality="Nong Ya Sai Subdistrict" specimenCode="UNI 2895" specimenCount="1" stateProvince="Udon Thani" typeStatus="holotype">
<emphasis id="9E4C7F95BD07D8AB4BE497F59829F009" bold="true">
<emphasis id="84E387EFB928D8FF8CAE7E13BE41F2CA" bold="true" italics="true">
<typeStatus id="B854F81E21D9622649D13E052DBD376F" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="80241AE023792EFFE01338E296C3653E" name="Thailand">Thailand</collectingCountry>
•
<collectingRegion id="7858156365AFF6AA560BC2452C20C2D1" country="Thailand" name="Udon Thani">Udon Thani Province</collectingRegion>
,
<collectingCounty id="68EA1BA3F59B13E98BFDCF4B4F820949">Wang Sam Mo District</collectingCounty>
,
<collectingMunicipality id="88DC2626B89DC50B64AB4DA98DA89760">Nong Ya Sai Subdistrict</collectingMunicipality>
,
<location id="49E49840368C79EC3BC17F507081CE44" LSID="urn:lsid:plazi:treatment:ACBEF56237745895AE444C49C4EE638E:49E49840368C79EC3BC17F507081CE44" country="Thailand" county="Wang Sam Mo District" latitude="16.9886" longLatPrecision="7" longitude="103.3638" municipality="Nong Ya Sai Subdistrict" name="Yang Stream" stateProvince="Udon Thani">Yang Stream</location>
(a tributary of
<location id="B4F9C58398F18E458D9E6BB526766924" LSID="urn:lsid:plazi:treatment:ACBEF56237745895AE444C49C4EE638E:B4F9C58398F18E458D9E6BB526766924" country="Thailand" county="Wang Sam Mo District" latitude="16.9886" longLatPrecision="7" longitude="103.3638" municipality="Nong Ya Sai Subdistrict" name="Lam Pao River" stateProvince="Udon Thani">Lam Pao River</location>
);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKTBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[103.363800,16.988600]}">
<geoCoordinate id="0D6D754A3BE73B9059A4C2DDC86D13AA" degrees="16.9886" direction="north" orientation="latitude" precision="5" value="16.9886">16.9886 ° N</geoCoordinate>
,
<geoCoordinate id="990D801FE709A7AEA4E5C17A7CBDA703" degrees="103.3638" direction="east" orientation="longitude" precision="5" value="103.3638">103.3638 ° E</geoCoordinate>
</named-content>
;
<collectorName id="694FAFDA5B9270C6EF6747AB83464C8E">K. Wisittikoson</collectorName>
leg.;
<named-content id="7645FC452C0B6F6C32927E91AC3FBB85" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">
<collectionCode id="5C88B696263464C3E3838F8C87B6BA80">MUMNH</collectionCode>
</named-content>
-
<specimenCode id="A959E553A2733671100E21AAD816755B" collectionCode="UNI" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">UNI 2895</specimenCode>
; shell length
<quantity id="ABF62BD416A574E1D32CAD2745935E14" metricMagnitude="-2" metricUnit="m" metricValue="5.384" unit="mm" value="53.84">53.84 mm</quantity>
, shell height
<quantity id="575554A3AC2BAAD730CFA336B22084C9" metricMagnitude="-2" metricUnit="m" metricValue="3.168" unit="mm" value="31.68">31.68 mm</quantity>
, and shell width
<quantity id="9F5A3D5E42B697A9E66BB143A89C0EC5" metricMagnitude="-2" metricUnit="m" metricValue="2.041" unit="mm" value="20.41">20.41 mm</quantity>
</materialsCitation>
.
<materialsCitation id="CC1DB168BFF7428258FA2F2E8023A477" collectionCode="MUMNH" collectorName="K. Wisittikoson" country="Thailand" county="Wang Sam Mo District" latitude="16.9886" location="Yang Stream" longLatPrecision="7" longitude="103.3638" municipality="Nong Ya Sai Subdistrict" specimenCode="UNI 2896, UNI 2897" specimenCount="1" stateProvince="Udon Thani" typeStatus="paratype">
<emphasis id="C971D979D0776BB3D71D99AB51224DF5" bold="true">
<emphasis id="EE5936B7CDEEE2C187DDDE267D641639" bold="true" italics="true">
<typeStatus id="027AC8541261386CFC487102F4973975" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="141251DC01A32B937003691C8CEA8C1C" name="Thailand">Thailand</collectingCountry>
• 2 shells; same collection data as for holotype;
<named-content id="271E16595E614D1D22FBFBCB27985938" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">
<collectionCode id="59368DF1DBE339DE52A70DACCCDC6CF5">MUMNH</collectionCode>
</named-content>
-
<specimenCode id="D53882925F69551366311A3E8931A9DE" collectionCode="UNI" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">UNI 2896</specimenCode>
,
<specimenCode id="C5B3181526012208C6B472E49BD8B82C" collectionCode="UNI" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">UNI 2897</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4TBG" type="diagnosis">
<paragraph id="FC7592AE52B8856E32A295FA4E0FF282">
<heading id="74EAD80425CEEA16D5ED46331686BCEA" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="7AD724860FF0EDB3C17D10747182AD1B">
This species resembles
<taxonomicName id="D9EB77168E3D2DE9B1572CEF620CC75C" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="fortunata" status="sp. nov.">
<emphasis id="8E8B3366193EF6AD6EC46BFCC56E4F79" italics="true">I. fortunata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2DA356505767319F0F3A11F3613D638" rank="species">sp. nov.</taxonomicNameLabel>
, but it can be distinguished by the overall shell shape being more rounded (vs rhomboidal), slightly higher shell (H / L ratio 0.58–0.60 vs 0.55–0.57), wider anterior margin (vs rather constricted), more truncated and rounded posterior margin (vs slightly ventrally elongate), less curved ventral margin (vs curved ventral margin). It also differs in characteristics inside the shell by having higher and more triangulate pseudocardinal tooth on the right valve (vs very shallow and hill-like pseudocardinal tooth), well-developed muscle scars (vs very shallow and sometimes almost invisible), 3 to 5 well-developed muscle scars in the umbo cavity (vs 1 to 2 tiny muscle scars in the umbo cavity), and well-marked pallial line (vs faint pallial line). It can also be distinguished from its congeners by fixed nucleotide substitution positions in the
<abbrev id="ABBRID0EOUBG" xlink_title="mitochondrial cytochrome c oxidase subunit-I gene">COI</abbrev>
gene fragment of base A on the 54
<superScript id="173B2A6618C37E7435CCD094B5A1DD2B">th</superScript>
, 114
<superScript id="E07C0F1BE551D55A8DDD81CC81AF3414">th</superScript>
, 363
<superScript id="CE8C2D7254FC601D62ED3E0D389812ED">rd</superScript>
, and 414
<superScript id="5CF4E1A6C37692A7249B7899CFC79AE4">th</superScript>
; base
<collectionCode id="188817D7E53B104AA964C3530C4385BC" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
on the 90
<superScript id="9D6B4F2E31098B5AEA3323B937C6D641">th</superScript>
, 141
<superScript id="4C43C9C2E0F6D7F4F452390D287DB6EE">st</superScript>
, 591
<superScript id="AF2C76426C9DFF6885671D3B1116D52E">st</superScript>
, and 609
<superScript id="0E2E411665B113180C7927C5354C4EAF">th</superScript>
; base C on the 57
<superScript id="1455C635BFFE4DFC220E14A3CB78A71F">th</superScript>
, and 216
<superScript id="29950E4E6202F92ACE9219A4A8A164FC">th</superScript>
; and base G on 150
<superScript id="D90D862FC264B702B9EF7132F05EC4B0">th</superScript>
, 117
<superScript id="E039DA30ADEBA2058E4C9E6FC49B77CD">th</superScript>
, 192
<superScript id="AEE9C677DA8F0612BE2A79C45B54CAFB">nd</superScript>
, 558
<superScript id="F861B7E790FAB619ADE6CF77906342D7">th</superScript>
, 480
<superScript id="D263908BA9F48778131017293CE3201D">th</superScript>
, 552
<superScript id="8B79DD821FE5E5C6843782C2705A7CB3">nd</superScript>
, 627
<superScript id="22EE8B6D395394D92869337F3527D383">th</superScript>
, 639
<superScript id="0DDA63CCFF2DD513D74FD97AE8D4B01E">th</superScript>
, 645
<superScript id="BD76888687A490B1B2D6BDD3E574A3E7">th</superScript>
, and 657
<superScript id="198FAAA37930B384869DB35212440465">th</superScript>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1VBG" type="description">
<paragraph id="13E22C4932D36BE79C66F27DE2F2729B">
<heading id="67B0A84370C9A6DF7B69E1B667B0D18C" reason="title">Description.</heading>
</paragraph>
<paragraph id="2C367F79A2E0A779F9627966F806E416">
Shell medium-sized (shell length
<quantity id="AA8F346651FB19957A258769879765F0" metricMagnitude="-2" metricUnit="m" metricValue="4.88" metricValueMax="5.38" metricValueMin="4.38" unit="mm" value="48.8" valueMax="53.8" valueMin="43.8">43.8–53.8 mm</quantity>
, shell height
<quantity id="3E061012A44DC4A5F0F7012999014CFD" metricMagnitude="-2" metricUnit="m" metricValue="2.92" metricValueMax="3.17" metricValueMin="2.67" unit="mm" value="29.2" valueMax="31.7" valueMin="26.7">26.7–31.7 mm</quantity>
, shell width
<quantity id="729CFA93AC81FC31B59FA3DDCB7AC805" metricMagnitude="-2" metricUnit="m" metricValue="1.71" metricValueMax="2.04" metricValueMin="1.38" unit="mm" value="17.1" valueMax="20.4" valueMin="13.8">13.8–20.4 mm</quantity>
; Table
<tableCitation id="120F1F806480C56F834A3684F0D2312A" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell measurements and GenBank accession numbers for the type series of the new species described in this study. Measurements in millimetres (mm)." httpUri="http://table.plazi.org/id/2EDC89F0C67A5A15CACAB0A86B456858" tableUuid="2EDC89F0C67A5A15CACAB0A86B456858">2</tableCitation>
), ovate to rectangular, elongate (H / L ratio = 0.58–0.60), rather thin, very inequilateral, moderately inflated. Anterior margin rounded, shouldered; posterior margin truncated to rounded; ventral margin straight to slightly curved. Dorsal margin slightly curved; anterior low, posterior end slightly higher. Umbo slightly elevated, heavily eroded even in young specimens. Posterior ridge wide and obtuse, not prominent; posterior slope with two faint lines running from umbo to posterior margin, invisible in old specimens. Periostracum thin, brown to brownish-black, eroded part white to coppery-brown. Shell surface with visible growth lines. Ligament long, very narrow, dark brown. One hill-like or triangulate pseudocardinal tooth on right valve; pseudocardinal tooth on left valve less developed, hill-like. Posterior end of hinge structure with small V-shaped fossette. Anterior adductor muscle scar well-developed, ovate, contiguous with anterior pedal retractor and anterior protractor muscle scars; posterior adductor muscle scar round, shallow, contiguous with posterior retractor muscle scars; posterior retractor muscle scars triangular. Pallial line well-marked, continuous. Umbo cavity moderately deep, with 3 to 5 muscle scars in cavity. Nacre blue-whitish to yellowish, more yellowish toward the umbo cavity.
</paragraph>
<paragraph id="BE4AA15F49B4614D757E71808BFFB253">Exhalant aperture almost smooth, with a row of tiny tubercle-like papillae on the edge of aperture, similar length with inhalant. Inhalant aperture with a row of very short conical papillae. Small epithelial fold divides exhalant and inhalant apertures. Gills elongated and slightly ribbed; inner gills slightly wider, and anterior margin of inner gills slightly longer than that of outer gills. Labial palps elongate, slightly curved, pointed at tip. Glochidia unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EFWBG" type="etymology">
<paragraph id="FD984FC708739BCB0842904BE9878ED2">
<heading id="82AFAFD0B666C25D8B6000A50B2D3EC9" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="20044DBD3EF7986CBE7B350D5F98F94F">
The species name “
<emphasis id="C8F9A9FA49ABB3546D52F764C826E927" italics="true">occultata</emphasis>
” is derived from the Latin adjective “
<emphasis id="CC3DD46123DA4B741D3BCA49A35A0AC9" italics="true">occultatus</emphasis>
” meaning concealed or hidden, a reference to this new freshwater mussel being hidden in a small stream far from the main river.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXWBG" type="distribution">
<paragraph id="6ABAE48CC57EFF1A85692CB047C3C44A">
<heading id="4BD2F21F9D0D35804393564991E4F48F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="963C025E00D3CA6B99AE8BFF0DA2F0AE">
This new species is known only from the
<typeStatus id="E8E9845E06C541FDBCB25BD81D7B6ECF">type</typeStatus>
locality, Yang Stream in
<collectingRegion id="F5D72440989BF9AC6B068599AE843E08" country="Thailand" name="Udon Thani">Udon Thani Province</collectingRegion>
. Yang Stream is a headwater tributary of the Lam Pao River in the Chi River Basin. The
<typeStatus id="8D63EBD204D676D78776C8998CE5D4CA">type</typeStatus>
locality is above the Lam Pao Dam.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3WBG" type="remarks">
<paragraph id="89C172CC479202A3177A64054035AC19">
<heading id="0667F13A6A4C4F7115E5AF6A2563E498" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="B1A165E37E4F5A39BAB12EAB020C122F">
This new species is rare and difficult to find despite repeated surveys. They were collected from a network of dense tree roots on a clay wall of a small stream. Nearby, there was a community of unionid mussels living in muddy or sandy substrate of the stream bottom, including
<taxonomicName id="04B20E6071A62C24A0D793445EEA6351" authority="(Lea, 1856)" baseAuthorityName="Lea" baseAuthorityYear="1856" class="Bivalvia" family="Unionidae" genus="Scabies" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="phaselus">
<emphasis id="D068FCAE817541B4E9034A6CC9047136" italics="true">Scabies phaselus</emphasis>
(
<bibRefCitation id="C7B99905C03CEC7AE2B948A3CA67F84F" author="Lea I" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pagination="92-95" refId="B32" refString="Lea I (1856) Description of twenty-five new species of exotic uniones. Proceedings of the Academy of Natural Sciences of Philadelphia 8: 92 - 95." title="Description of twenty-five new species of exotic uniones." volume="8" year="1856">Lea, 1856</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="8130F10D4091ED553F18D54E41319BF7" authority="Brandt, 1974" authorityName="Brandt" authorityYear="1974" class="Bivalvia" family="Unionidae" genus="Physunio" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="modelli">
<emphasis id="3A97373D10CE560A3558AC67D7920B22" italics="true">Physunio modelli</emphasis>
<bibRefCitation id="3F692EEA9E7A4311923F8F7D113E5EC3" author="Brandt RAM" journalOrPublisher="Archiv für Molluskenkunde" pagination="1-423" refId="B9" refString="Brandt RAM (1974) The non-marine aquatic Mollusca of Thailand. Archiv fur Molluskenkunde 105: 1 - 423." title="The non-marine aquatic Mollusca of Thailand." volume="105" year="1974">Brandt, 1974</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="18FA130F3B3B6674ED0704981BFDED6B" authority="Jeratthitikul et al., 2021" authorityName="Jeratthitikul" authorityYear="2021" class="Bivalvia" family="Unionidae" genus="Namkongnaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="inkhavilayi">
<emphasis id="ED8968328ACC6025270F6A554E18F2F0" italics="true">Namkongnaia inkhavilayi</emphasis>
Jeratthitikul et al., 2021
</taxonomicName>
,
<taxonomicName id="3B08BC34F15FF4C67D6BC9F47FC0958F" baseAuthorityName="Lea" baseAuthorityYear="1838" class="Bivalvia" family="Unionidae" genus="Pilsbryoconcha" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="exilis">
<emphasis id="EFA79418B8F79527E098E6675607E220" italics="true">P. exilis</emphasis>
</taxonomicName>
,
<taxonomicName id="7964D5D91110E7F05C680B666D1609D7" class="Bivalvia" family="Unionidae" genus="Sundadontina" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="brandti">
<emphasis id="C7DB508B89320BEF6B155A2B10E723A3" italics="true">S. brandti</emphasis>
</taxonomicName>
, and
<taxonomicName id="B3B7AEF4A4A1D892990AC88BE05BE7B6" authority="(Martens, 1860)" baseAuthorityName="Martens" baseAuthorityYear="1860" class="Bivalvia" family="Unionidae" genus="Thaiconcha" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="callifera">
<emphasis id="29CF7CC7E2E193C2755D04BA47CDE1F8" italics="true">Thaiconcha callifera</emphasis>
(
<bibRefCitation id="833CC103E66B68BF7C82CA78FE5B7E7E" author="Martens EV" journalOrPublisher="Proceedings of the Zoological Society of London" pagination="6-18" refId="B37" refString="Martens EV (1860) On the Mollusca of Siam. Proceedings of the Zoological Society of London 1860: 6 - 18." title="On the Mollusca of Siam." volume="1860" year="1860">Martens, 1860</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

227
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<mods:title id="1B68F815C8F25DF086D4D91E1D4E52DD">Four new species of Georissa W. Blanford, 1864 (Gastropoda, Hydrocenidae) from Thailand</mods:title>
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<subSubSection id="42DE5E53889209973C87A3F3207F7386" type="nomenclature">
<paragraph id="D85E1C8049830805822B866CF0F68983">
<taxonomicName id="4E379A97240374035E5E353CF652FBAD" authority="Klongkaew, Poeaim &amp; Dumrongrojwattana" authorityName="Klongkaew, Poeaim &amp; Dumrongrojwattana" authorityYear="2024" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="kohsichangensis" status="sp. nov.">
<emphasis id="0749C178C297094DB01AA224B37F8F3C" italics="true">Georissa kohsichangensis</emphasis>
Klongkaew, Poeaim &amp; Dumrongrojwattana
</taxonomicName>
<taxonomicNameLabel id="C4D93142572BC2A42262CA03983B8EF9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3579EF1424086AB410BCB01850AB3E22" type="description">
<paragraph id="9D385778F2011AF10FC3708E093E503F">
<figureCitation id="6CEBA95E01A89B9C93C0CF72F4AA7E5B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">Figs 2 B</figureCitation>
,
<figureCitation id="62DD3B78138868124111C11091484228" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 E – H</figureCitation>
,
<figureCitation id="4DE4C2437B3479D00486C2A6FA3EFDE6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of radula morphology of Georissa. A – D. G. quinquelirata sp. nov.; E – H. G. kohsichangensis sp. nov.; A, E. Radula overview; B, F. Radula segment; C, G. Lateral teeth (Lt); D, H. Marginal teeth (Mt); I – L. G. sagitta sp. nov. M – P; G. digitinota sp. nov.; I, M. Radula segment; J, M. Radula segment close up; K, O. Lateral teeth (Lt); L, P. Marginal teeth (Mt). Color highlights show radula position; blue: small central teeth (Ct), purple: lateral teeth, and orange: marginal teeth." figureDoi="10.3897/zse.100.128717.figure5" httpUri="https://binary.pensoft.net/fig/1148344">5 E – H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIMBG" type="type material">
<paragraph id="275C82E5CCF7E381B91FD571A5B82054">
<heading id="041C8D786E52448DB55402C304042B75" reason="title">Type material.</heading>
</paragraph>
<paragraph id="891319D93C2C28BB6FD687CC78497137">
<emphasis id="E4A81E1186B1705FA8C320D16839A268" bold="true" italics="true">
<typeStatus id="3332B2766DA7878F68700E9749505B74" type="holotype">Holotype</typeStatus>
</emphasis>
•
<abbrev id="ABBRID0ETMBG" xlink_title="Zoological Research Collection of Burapha University">ZRCBUU</abbrev>
0902 (Fig.
<figureCitation id="25807007E4B33AA701346D4B2201FC01" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 B</figureCitation>
).
<abbrev id="ABBRID0E2MBG" xlink_title="Shell height">SH</abbrev>
=
<quantity id="B09495CF50E6006508F1F15EE47F744B" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.10 mm</quantity>
,
<abbrev id="ABBRID0E6MBG" xlink_title="shell width">SW</abbrev>
=
<quantity id="ECEC925EB6BFF769EDE0E7906D30A720" metricMagnitude="-3" metricUnit="m" metricValue="1.43" unit="mm" value="1.43">1.43 mm</quantity>
,
<abbrev id="ABBRID0EDNBG" xlink_title="aperture height">AH</abbrev>
=
<quantity id="ECCE0C3F04DF3AB8852728B649A16194" metricMagnitude="-4" metricUnit="m" metricValue="6.3" unit="mm" value="0.63">0.63 mm</quantity>
,
<abbrev id="ABBRID0EHNBG" xlink_title="aperture width">AW</abbrev>
=
<quantity id="C5CEB8126182A35E519E6F9C38FA9654" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.90 mm</quantity>
, W = 4.25–4.5.
<emphasis id="52B86F60FC3DB7A60AE2C32BDF6ACE1E" bold="true" italics="true">
<typeStatus id="D5D1FB850B1C261E906DA58E5369268F" type="paratype">Paratype</typeStatus>
</emphasis>
•
<abbrev id="ABBRID0EONBG" xlink_title="Zoological Research Collection of Burapha University">ZRCBUU</abbrev>
0903 (Fig.
<figureCitation id="732F23443179AE995F2619CADB196C17" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 E – H</figureCitation>
) (12 shells); Shell measurements:
<abbrev id="ABBRID0EWNBG" xlink_title="Shell height">SH</abbrev>
=
<quantity id="B500C5F2CBD3557E7ED3940135A7726D" metricMagnitude="-3" metricUnit="m" metricValue="1.875" metricValueMax="2.03" metricValueMin="1.72" unit="mm" value="1.875" valueMax="2.03" valueMin="1.72">1.72–2.03 mm</quantity>
(1.86 ±
<quantity id="C06A03ED2C5B02CF842E0C1B198B1D4C" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.10 mm</quantity>
),
<abbrev id="ABBRID0E1NBG" xlink_title="shell width">SW</abbrev>
=
<quantity id="7E04D68F8535AA48CA80B0DE008F9CB7" metricMagnitude="-3" metricUnit="m" metricValue="1.315" metricValueMax="1.42" metricValueMin="1.21" unit="mm" value="1.315" valueMax="1.42" valueMin="1.21">1.21–1.42 mm</quantity>
(1.30 ±
<quantity id="CF5305084A11A97413D3BA69BB012096" metricMagnitude="-5" metricUnit="m" metricValue="7" unit="mm" value="0.07">0.07 mm</quantity>
),
<abbrev id="ABBRID0E5NBG" xlink_title="aperture height">AH</abbrev>
=
<quantity id="85D89DC0DDA2C4098A6B58C6EF6AB23D" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="7.199999999999999" metricValueMin="5.1" unit="mm" value="0.615" valueMax="0.72" valueMin="0.51">0.51–0.72 mm</quantity>
(0.62 ±
<quantity id="769B473EE3F41A437AEB832A93003A38" metricMagnitude="-5" metricUnit="m" metricValue="5.0" unit="mm" value="0.05">0.05 mm</quantity>
),
<abbrev id="ABBRID0ECOBG" xlink_title="aperture width">AW</abbrev>
=
<quantity id="FB3326E0D658FDAA614291BA5FD1848B" metricMagnitude="-4" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.4" metricValueMin="7.5" unit="mm" value="0.7949999999999999" valueMax="0.84" valueMin="0.75">0.75–0.84 mm</quantity>
(0.79 ±
<quantity id="E44AB5600CAE6755A9A67FD1742ADD84" metricMagnitude="-5" metricUnit="m" metricValue="3.0" unit="mm" value="0.03">0.03 mm</quantity>
), W = 3.75–4.5 (all type material from type locality;
<date id="AA0CD3D791FA956AD5533B3549F2B2D2" value="2021-10-30">30 October 2021</date>
, P. Dumrongrojwattana leg).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGOBG" type="type locality">
<paragraph id="8C5F61644D586561FCB7C4B65F47EDDB">
<heading id="F2A1DE59BAE6F16D4B87F20D7DD02E99" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="8E7A6B787E43B243FD4A89F22F9F1BD4">
<collectingCountry id="1A2A7969939C6F6AF60133B9C885A2DB" name="Thailand">Thailand</collectingCountry>
, Yai Man cave, Koh Sichang district,
<collectingRegion id="F58B551406EB0ADDA9F394E99C2285ED" country="Thailand" name="Chon Buri">Chonburi Province</collectingRegion>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPOBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[100.807778,13.152278]}">
<geoCoordinate id="698CBFCAD7710A5FA763FC1BAFE5AD93" degrees="13" direction="north" minutes="09" orientation="latitude" precision="1" seconds="08.2" value="13.152277">13 ° 09 ' 08.2 &quot; N</geoCoordinate>
,
<geoCoordinate id="5D415447426A29423DC62293065DBCF2" degrees="100" direction="east" minutes="48" orientation="longitude" precision="1" seconds="28.0" value="100.80778">100 ° 48 ' 28.0 &quot; E</geoCoordinate>
</named-content>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUOBG" type="etymology">
<paragraph id="8E9770729088AC93710F3409D6A320D5">
<heading id="C7E1FC711F6595CD503B1B1E9A6650E5" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="90363534511B823E05542BC6AA000B79">
This specific designation “
<emphasis id="FA6C7B77E7AE0C8F39BA0C87FC1BC8FA" italics="true">kohsichangensis</emphasis>
” is a district referring to the
<typeStatus id="EE37F01EE5F34DC1E80C5888C488151B">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPBG" type="diagnosis">
<paragraph id="8AD88C329AA16F27B9FD6B4F6C8DC9F5">
<heading id="08D9CB8CAC09740D00509AF0D1FACE4C" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4F09323B6F46354C02BBE2E1DE3167BF">Shell minute, ovately conical and the upper half part of the whorls is slightly downward, orangish to brownish, protoconch round to slightly ellipsoid and perforated sculpture. Body whorl with 6–7 spiral lirae. Aperture rhombate. Umbilicus closed. Operculum corneous with apophysis, paucispiral, opaque white, and rhombate.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJPBG" type="description">
<paragraph id="9E3237C4B396FEF948FCCF8933943CA0">
<heading id="60783CC28F85366A42979566A9774950" reason="title">Description.</heading>
</paragraph>
<paragraph id="7645EA4A3014CA13E044FFDC3F71F21F">
Shell minute, dextral, orangish to brownish, ovately conical with 4.25–4.5 whorls (Fig.
<figureCitation id="C33FAF79033FB4C4E04EDCA95FC676A4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 B</figureCitation>
). Protoconch orange, about one whorl, round to slightly ellipsoid, covered with perforated sculpture (Fig.
<figureCitation id="2AB476846CC9452D0C053A3F663DEF36" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 F, G</figureCitation>
). Teleconch orange 3.25–3.5 whorls. The upper half part of the body whorls is slightly downward, sculptured with only strong oblique growth lines, 6–7 prominent lirae, weak oblique growth lines present on the half lower part of the body whorls, more densely spaced spiral cords, ca. 6–8 cords, at the basal part of the body whorl (Fig.
<figureCitation id="5CF13A8987449F21A400E2FAF11A34E7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 B</figureCitation>
,
<figureCitation id="D038CA4B5332B774622DABAFCAE35DCE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 E</figureCitation>
). Suture incised. Aperture rhombate. Peristome thin, sharp, unexpanded or thickened, not reflected. Umbilicus closed. Operculum paucispiral, nucleus submarginal, corneous, opaque white, rhombate shape, thin, smooth outside surface with apophysis, inside surface with an arched peg arising from base (Fig.
<figureCitation id="D83F4B74ECE44E16B13FD462E1E94B07" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDQBG" type="Radula">
<paragraph id="85BB4F6D779AA6652B46865FF26BBC2B">
<heading id="3043F926AE926D393054B1021C5269ED" reason="title">Radula.</heading>
</paragraph>
<paragraph id="6E29A93B569515175C34D585A468FB71">
Ribbon-like, slender, longer and delicate towards the outside, rhipidoglossate (Fig.
<figureCitation id="24F301D1B1B35F7FB37AEA0BB9B8B91C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of radula morphology of Georissa. A – D. G. quinquelirata sp. nov.; E – H. G. kohsichangensis sp. nov.; A, E. Radula overview; B, F. Radula segment; C, G. Lateral teeth (Lt); D, H. Marginal teeth (Mt); I – L. G. sagitta sp. nov. M – P; G. digitinota sp. nov.; I, M. Radula segment; J, M. Radula segment close up; K, O. Lateral teeth (Lt); L, P. Marginal teeth (Mt). Color highlights show radula position; blue: small central teeth (Ct), purple: lateral teeth, and orange: marginal teeth." figureDoi="10.3897/zse.100.128717.figure5" httpUri="https://binary.pensoft.net/fig/1148344">5 E – H</figureCitation>
). Central tooth five small teeth, trapezoid shape. Lateral teeth five teeth, claw-like with 7–11 alternately long and short denticles decreasing in size from the first lateral tooth toward the 5
<superScript id="75FDA721AB855F69234FB8B44A77FDF0">th</superScript>
lateral tooth. Marginal teeth two teeth, claw-like with 14–18 equal size denticles. Radula formula 2: 5: 1 + 1 + 1 + 1 + 1 + 1: 5: 2.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPQBG" type="Differential diagnosis">
<paragraph id="0207FBFB944B33FDC66F3F2485FE35A0">
<heading id="8E5AD4071018DDCDA6187A480C5A80B5" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="15A727884C15670A50F5ED9931F34878">
<taxonomicName id="067F1FDEEA80FABA4F0ABFD36D0C4D9E" authorityName="Klongklaew &amp; Poeaim &amp; Dumrongrojwattana" authorityYear="2024" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="kohsichangensis" status="sp. nov.">
<emphasis id="66699F5197F2597895621B947B5E1632" italics="true">Georissa kohsichangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72262E46AABD6152E6BB7BCC5FFF5F6F" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="9C719DEF44DC7C9C8475742199846312" authority="Sutcharit &amp; Jirapatrasilp, 2020" authorityName="Sutcharit &amp; Jirapatrasilp" authorityYear="2020" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="315394B601D3B8E73C8EFF10D96301D2" italics="true">G. carinata</emphasis>
Sutcharit &amp; Jirapatrasilp, 2020
</taxonomicName>
from
<collectingCountry id="738DBE711A46673CDE4AB7442DE56633" name="Cambodia">Cambodia</collectingCountry>
.
<taxonomicName id="FD30D4A2BACC4D55D072124A3E9BE934" authorityName="Sutcharit &amp; Jirapatrasilp" authorityYear="2020" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="EFD2FBD34196E5D6434BE8378F7A9931" italics="true">Georissa carinata</emphasis>
</taxonomicName>
differs in having more convex body whorl, a smooth protoconch, and sculptured with thin and uneven growth lines.
</paragraph>
</subSubSection>
</treatment>
</document>