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<mods:title id="1AC92D37652244CA191235B09AE3AAF1">Species plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</mods:title>
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1. Kaempferia foliis ovatis sessilibus.
<emphasis id="A1348BA9806862FB6DBD6B1AFB6A5A95" italics="true">Fl. zeyl.8. Mat. med.3.</emphasis>
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Kaempferia.
<emphasis id="90ADC6C7CA44EC4C32A43F7286F6E71B" italics="true">Hort. cliff.2. t.3. Roy. lugdb.12.</emphasis>
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<pageBreakToken id="F3A06B5D66274E2D40CBAB4B03965BEC" pageNumber="3">Wanhom</pageBreakToken>
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<emphasis id="F61EC7E59483802CAEE148DBDB527B66" italics="true">Habitat in</emphasis>
India
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Beier
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Paulian
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Roy
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;
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Ehrmann
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:
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;
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Otte &amp; Spearman
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<emphasis box="[807,929,1625,1651]" italics="true" pageId="12" pageNumber="221">Nilomantis</emphasis>
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<emphasis bold="true" box="[98,160,951,973]" pageId="13" pageNumber="222">Fig. 9.</emphasis>
Male genitalic complex of
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<emphasis box="[419,596,951,972]" italics="true" pageId="13" pageNumber="222">Ilomantis thalassina</emphasis>
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. Abbreviations: an, anterior apodeme; ap, anterior process; bm, right arm; L1, L2, L4B, a sub-sclerite of the dorsal sclerotization of the left phallomeric complex; L4A, the ventral sclerotization of the left phallomeric complex; loa, lobo membranoso; paa, apical process; pda, processo distale; pia, piastra ventrale; pva, processo ventrale sclerificato; R1, a sub-sclerite of the right phallomere; R3, a sub-sclerite of the right phallomere. Red lettering indicates structures; black lettering indicates characters. This figure is published in colour in the online edition of this journal, which can be accessed via http://booksandjournals.brillonline.com/content/journals/1876312x.
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<typeStatus box="[98,147,1179,1204]" pageId="13" pageNumber="222">Type</typeStatus>
species
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<paragraph blockId="13.[98,672,1230,1257]" box="[98,672,1230,1257]" pageId="13" pageNumber="222">
<taxonomicName authority="Saussure, 1899: 593" authorityName="Saussure" authorityPageNumber="593" authorityYear="1899" box="[98,606,1230,1257]" class="Insecta" family="Iridopterygidae" genus="Tropidomantis" kingdom="Animalia" order="Mantodea" pageId="13" pageNumber="222" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[98,370,1230,1256]" italics="true" pageId="13" pageNumber="222">Tropidomantis thalassina</emphasis>
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Saussure,
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.
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<emphasis box="[98,202,1292,1317]" italics="true" pageId="13" pageNumber="222">Diagnosis</emphasis>
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Small, slender mantises (body length
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), green to ochre, fully developed wings. Body surface and anterior margin of forewings lightly ciliated. Pronotal medial keel present. Head capsule dorsoventrally compressed. Compound eyes dorsoventrally compressed, laterally conical, blunted. Cranial vertex with a cosinusoidal margin, juxtaocular bulges present; elevation on the vertex in the shape of a truncated pyramid. Lower frons featuring a medially bifurcated carina.
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<emphasis box="[98,241,1578,1604]" italics="true" pageId="13" pageNumber="222">Redescription</emphasis>
</heading>
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<emphasis box="[98,157,1630,1656]" italics="true" pageId="13" pageNumber="222">Head</emphasis>
(
<figureCitation box="[174,242,1630,1656]" captionStart="Fig. 8" captionStartId="12.[106,139,1140,1162]" captionTargetBox="[155,1023,131,1114]" captionTargetId="figure@12.[155,1024,131,1115]" captionTargetPageId="12" captionText="Fig. 8. Illustrations of anterior perspective of the Ilomantis and Nilomantis head (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male; (E) Nilomantis floweri, female; (F) Nilomantis floweri, male. (B, D, E, F) Ilomantis species feature a cosinusoidal vertical margin, whereas Nilomantis species feature a relatively straight vertical margin. Gray markings on the ocellar tubercle and within the corridors of the frontal sulci represent the presence of reddish-brown markings. LF, lower frons." httpUri="https://zenodo.org/record/269847/files/figure.png" pageId="13" pageNumber="222">
Fig.
<number box="[226,242,1630,1656]" pageId="13" pageNumber="222" value="8.0">8</number>
</figureCitation>
). Prognathous (based on observed resting position); head capsule wider than long. Juxtaocular bulges present, highlighted by parietal sulci. Head capsule dorsoventrally compressed with likewise compressed, laterally conical compound eyes. Four gently sloping carinal ridges on the vertex (two of which originate from the mid-vertex, the other two originate from the mid-ocular region) converge into an elevation on the postero-medial vertex in the shape of a truncated pyramid, which is bisected apically by the coronal suture. Vertex slightly concave posterior to lateral ocelli. Ocelli of both sexes are situated atop an elevated region; ocelli amber to yellow. Lower frons transverse, featuring a subtle, medially bifurcated sulcus. Clypeus broad, with elevated margins. Labrum rounded anteriorly. Maxillary and labial palpi pale, light green to ochre. Compound eye pigmentation darker than cuticle of the head capsule. Antenna long and filiform, lightly ciliated, tapering to a point, with darkened terminal antennomeres.
</paragraph>
<caption httpUri="https://zenodo.org/record/269849/files/figure.png" pageId="14" pageNumber="223" targetBox="[155,1024,131,1220]" targetPageId="14">
<paragraph blockId="14.[106,1074,1247,1387]" pageId="14" pageNumber="223">
<emphasis bold="true" box="[106,180,1247,1269]" pageId="14" pageNumber="223">Fig. 10.</emphasis>
Male genitalic complex of
<taxonomicName box="[435,618,1248,1269]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="14" pageNumber="223" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[435,618,1248,1269]" italics="true" pageId="14" pageNumber="223">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[624,668,1248,1269]" pageId="14" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
(A) Right phallomere, ventral sclerotization of the left phallomeric complex, and dorsal (i.e., “left”) sclerotization of the left phallomeric complex (from left to right, respectively); (B, C, D) examples of shape and sclerotization variability present within the anterior process (ap) of L4B. This figure is published in colour in the online edition of this journal, which can be accessed via http://booksandjournals.brillonline.com/content/journals/1876312x.
</paragraph>
</caption>
<paragraph blockId="15.[98,1067,273,924]" pageId="15" pageNumber="224">
<emphasis box="[98,182,273,299]" italics="true" pageId="15" pageNumber="224">Thorax.</emphasis>
Pronotum roughly
<number box="[413,448,273,300]" pageId="15" pageNumber="224" value="2.5">2.5</number>
-times longer than wide at supracoxal sulcus dilation, featuring a pronounced medial keel that bisects the anteriorly positioned supracoxal sulcus and terminates at the posterior margin of the metazone (
<figureCitation box="[844,930,342,369]" captionStart="Fig. 11" captionStartId="16.[106,139,1517,1539]" captionTargetBox="[243,935,131,1491]" captionTargetId="figure@16.[243,937,131,1492]" captionTargetPageId="16" captionText="Fig. 11. Illustrations of dorsal perspective of the Ilomantis pronotum (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. Ilomantis thalassina exhibits a medial keel that fully traverses the pronotum (A and B); Ilomantis ginsburgae sp. n. exhibits a medial keel that originates in the mid-prozone and concludes at the posterior pronotal margin (C and D). MK, medial keel." httpUri="https://zenodo.org/record/269850/files/figure.png" pageId="15" pageNumber="224">
Fig.
<number box="[899,930,343,369]" pageId="15" pageNumber="224" value="11.0">11</number>
</figureCitation>
). Metazone posterior margin elevated into a shelf extending slightly over anterior margin of the mesothorax. Margin of the lateral lamellar expansion of the pronotum is relatively smooth with socketed setae projecting around the perimeter. Region containing pronotal medial keel elevated, sloping down to lateral margins. Thorax medially traversed by a yellow stripe (
<figureCitation box="[306,389,516,542]" captionStart="Fig. 12" captionStartId="17.[98,131,1460,1482]" captionTargetBox="[147,1015,131,1433]" captionTargetId="figure@17.[147,1016,131,1434]" captionTargetPageId="17" captionText="Fig. 12. Example of yellow stripe that originates in the posterior half of the ocellar tubercle and traverses the thorax of both Nilomantis and Ilomantis species. Species shown: Ilomantis ginsburgae sp. n., male. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269851/files/figure.png" pageId="15" pageNumber="224">
Fig.
<number box="[357,389,516,542]" pageId="15" pageNumber="224" value="12.0">12</number>
</figureCitation>
). Prozone with bilaterally symmetric sculpting, which tapers to the edge just prior to the lateral margins. Anterior metazone features a slight indentation on either side of the medial keel. Cervix bears lateral cervical sclerites and intercervical sclerites; ventral cervical sclerites absent. Lateral cervical sclerite anterior margin extends just past the anterior margin of the prozone and is lightly ciliated. Postcervical plate and exposed thoracic membrane is contained within the boundaries of the intercervical sclerites and the episternum, with either a broad or narrow hexagonal shape (
<figureCitation box="[216,298,758,785]" captionStart="Fig. 13" captionStartId="18.[106,139,1269,1291]" captionTargetBox="[155,1024,131,1244]" captionTargetId="figure@18.[155,1024,131,1244]" captionTargetPageId="18" captionText="Fig. 13. Illustrations of ventral perspective of the Ilomantis cervix (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. The postcervical plate and exposed thoracic membrane are contained within the boundaries of the intercervical sclerites and the episternum, with either a broad (Ilomantis thalassina, A and B) or narrow (Ilomantis ginsburgae sp. n., C and D) hexagonal shape. Abbreviations: eps, episternum; etm, exposed thoracic membrane; ics, intercervical sclerite; lcs, lateral cervical sclerite; pcpl, postcervical plate; tss, T-shaped sclerite." httpUri="https://zenodo.org/record/269852/files/figure.png" pageId="15" pageNumber="224">
Fig.
<number box="[266,298,759,785]" pageId="15" pageNumber="224" value="13.0">13</number>
</figureCitation>
). A furcasternal tubercle projects medially at the base of the T-shaped sclerite, posterior to the prothoracic coxae, surfaced with setation (
<figureCitation box="[852,936,793,819]" captionStart="Fig. 14" captionStartId="19.[98,131,607,629]" captionTargetBox="[365,799,131,582]" captionTargetId="figure@19.[365,800,131,582]" captionTargetPageId="19" captionText="Fig. 14. Oblique ventral view of the furcasternal tubercle (FT) located medially at the base of the T-shaped sclerite, posterior to the prothoracic coxae. Arrow indicates FT. Species shown: Nilomantis floweri, male. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269853/files/figure.png" pageId="15" pageNumber="224">
Fig.
<number box="[904,936,793,819]" pageId="15" pageNumber="224" value="14.0">14</number>
</figureCitation>
). DK hearing organ present on metathoracic ventral surface (see
<bibRefCitation author="Yager, D. D. &amp; Svenson, G. J." box="[737,1023,828,854]" journalOrPublisher="Biological Journal of the Linnean Society" pageId="15" pageNumber="224" pagination="541 - 568" part="94" refString="Yager, D. D. &amp; Svenson, G. J. (2008) Patterns of praying mantis auditory system evolution based on morphological, molecular, neurophysiological, and behavioural data. Biological Journal of the Linnean Society 94: 541 - 568." title="Patterns of praying mantis auditory system evolution based on morphological, molecular, neurophysiological, and behavioural data" type="journal article" year="2008">
Yager &amp; Svenson (
<number box="[954,1015,828,854]" pageId="15" pageNumber="224" value="2008.0">2008</number>
)
</bibRefCitation>
for hearing organ description). Wings of both sexes well developed, extending beyond base of terminalia, hyaline to opaque; variably ciliated.
</paragraph>
<paragraph blockId="15.[98,1067,966,1374]" pageId="15" pageNumber="224">
<emphasis box="[98,268,966,992]" italics="true" pageId="15" pageNumber="224">Prothoracic legs</emphasis>
. Lightly ciliated. Forecoxae long, extending past the base of the pronotum; postero- and anteroventral margins with socketed setae; apical lobes squared, convergent. Forefemora feature relatively straight dorsal margins that narrow distally. Posteroventral femoral spines robust and darkened apically, interspersed with cilia and a row of crenulation. Genicular lobes of the femora each with a slightly curved spine. Tibial spur groove lies between first discoidal spine and the first anteroventral femoral spine. Anteroventral femoral spines alternate between short and long, darkened apically. Second discoidal spine significantly longer than the first and third. Posteroventral and anteroventral tibial spines gradually elongating towards the tibial spur, darkening apically, interspersed with cilia. Tarsi
<number box="[717,730,1278,1305]" pageId="15" pageNumber="224" value="5.0">5</number>
-segmented, ciliated, with an enlarged penultimate euplantulae; darkened ungues.
<emphasis box="[777,793,1314,1339]" italics="true" pageId="15" pageNumber="224">F</emphasis>
=
<number box="[809,821,1313,1339]" pageId="15" pageNumber="224" value="3.0">3</number>
DS/
<number box="[869,937,1313,1339]" pageId="15" pageNumber="224" value="11.5" valueMax="12.0" valueMin="11.0">1112</number>
AvS/
<number box="[999,1014,1313,1339]" pageId="15" pageNumber="224" value="4.0">4</number>
PvS;
<emphasis box="[98,117,1349,1374]" italics="true" pageId="15" pageNumber="224">T</emphasis>
=
<number box="[133,203,1348,1374]" pageId="15" pageNumber="224" value="12.0" valueMax="14.0" valueMin="10.0">1014</number>
AvS/
<number box="[265,321,1348,1374]" pageId="15" pageNumber="224" value="11.5" valueMax="14.0" valueMin="9.0">914</number>
PvS.
</paragraph>
<paragraph blockId="15.[98,1066,1417,1548]" pageId="15" pageNumber="224">
<emphasis box="[98,424,1417,1443]" italics="true" pageId="15" pageNumber="224">Meso- and metathoracic legs.</emphasis>
Lightly ciliated across surface with increased ciliation density distally. Femoral carina on the posteroventral margin; femoral genicular lobes lacking spines. Tibiae tubular with an apical lobe and two apical spurs. Tarsi
<number box="[98,111,1521,1548]" pageId="15" pageNumber="224" value="5.0">5</number>
-segmented, ciliated, with an enlarged penultimate euplantulae; darkened ungues.
</paragraph>
<paragraph blockId="15.[98,1066,1590,1686]" lastBlockId="19.[98,1066,810,1011]" lastPageId="19" lastPageNumber="228" pageId="15" pageNumber="224">
<emphasis box="[98,198,1590,1616]" italics="true" pageId="15" pageNumber="224">Abdomen</emphasis>
. Smooth, surface ciliated. Segmentation of tergites straightly traversing the abdomen. Female supra-anal plate triangular, ciliated; narrower than the subgenital plate. Female subgenital plate posterior margin bifid, ciliated, narrowing towards the terminus; cerci long, filiform, tapering. Male supra-anal plate triangular, ciliated; narrower than subgenital plate. Male subgenital plate approximately rounded at the terminus; cerci long, compressed, tapering; styli short. Female genitalia: gonoplac (GL) features a rectangular dilation on the anteroventral margin; gonapophysis IX (GP) apex approximately mammiform to rounded. Male genitalia: lobo membranoso (loa) lightly sclerotized and without apical crenulation; processo distale (pda) narrow and rounded apically.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/269850/files/figure.png" pageId="16" pageNumber="225" targetBox="[243,935,131,1491]" targetPageId="16">
<paragraph blockId="16.[106,1074,1517,1657]" pageId="16" pageNumber="225">
<emphasis bold="true" box="[106,182,1517,1539]" pageId="16" pageNumber="225">Fig. 11.</emphasis>
Illustrations of dorsal perspective of the
<taxonomicName box="[570,652,1517,1538]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis box="[570,652,1517,1538]" italics="true" pageId="16" pageNumber="225">Ilomantis</emphasis>
</taxonomicName>
pronotum (scale bar = 1 mm). (A)
<taxonomicName class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis italics="true" pageId="16" pageNumber="225">Ilomantis thalassina</emphasis>
</taxonomicName>
, female; (B)
<taxonomicName box="[311,484,1547,1568]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[311,484,1547,1568]" italics="true" pageId="16" pageNumber="225">Ilomantis thalassina</emphasis>
</taxonomicName>
, male; (C)
<taxonomicName box="[591,771,1547,1568]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[591,771,1547,1568]" italics="true" pageId="16" pageNumber="225">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[777,822,1547,1568]" pageId="16" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
, female; (D)
<taxonomicName class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis italics="true" pageId="16" pageNumber="225">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[171,216,1577,1598]" pageId="16" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
, male.
<taxonomicName box="[283,456,1576,1597]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[283,456,1576,1597]" italics="true" pageId="16" pageNumber="225">Ilomantis thalassina</emphasis>
</taxonomicName>
exhibits a medial keel that fully traverses the pronotum (A and B);
<taxonomicName box="[106,287,1605,1626]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="16" pageNumber="225" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[106,287,1605,1626]" italics="true" pageId="16" pageNumber="225">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[295,339,1606,1627]" pageId="16" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
exhibits a medial keel that originates in the mid-prozone and concludes at the posterior pronotal margin (C and D). MK, medial keel.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/269851/files/figure.png" pageId="17" pageNumber="226" targetBox="[147,1015,131,1433]" targetPageId="17">
<paragraph blockId="17.[98,1066,1460,1570]" pageId="17" pageNumber="226">
<emphasis bold="true" box="[98,173,1460,1482]" pageId="17" pageNumber="226">Fig. 12.</emphasis>
Example of yellow stripe that originates in the posterior half of the ocellar tubercle and traverses the thorax of both
<taxonomicName box="[342,446,1489,1510]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="17" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis box="[342,446,1489,1510]" italics="true" pageId="17" pageNumber="226">Nilomantis</emphasis>
</taxonomicName>
and
<taxonomicName box="[494,581,1489,1510]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="17" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis box="[494,581,1489,1510]" italics="true" pageId="17" pageNumber="226">Ilomantis</emphasis>
</taxonomicName>
species. Species shown:
<taxonomicName box="[816,1007,1489,1510]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="17" pageNumber="226" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[816,1007,1489,1510]" italics="true" pageId="17" pageNumber="226">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1014,1062,1490,1511]" pageId="17" pageNumber="226" rank="species">sp.n.</taxonomicNameLabel>
, male. This figure is published in colour in the online edition of this journal, which can be accessed via http://booksandjournals.brillonline.com/content/journals/1876312x.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/269852/files/figure.png" pageId="18" pageNumber="227" targetBox="[155,1024,131,1244]" targetPageId="18">
<paragraph blockId="18.[106,1074,1269,1468]" pageId="18" pageNumber="227">
<emphasis bold="true" box="[106,179,1269,1291]" pageId="18" pageNumber="227">Fig. 13.</emphasis>
Illustrations of ventral perspective of the
<taxonomicName box="[563,646,1270,1291]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis box="[563,646,1270,1291]" italics="true" pageId="18" pageNumber="227">Ilomantis</emphasis>
</taxonomicName>
cervix (scale bar = 1 mm). (A)
<taxonomicName class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis italics="true" pageId="18" pageNumber="227">Ilomantis thalassina</emphasis>
</taxonomicName>
, female; (B)
<taxonomicName box="[259,435,1299,1320]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[259,435,1299,1320]" italics="true" pageId="18" pageNumber="227">Ilomantis thalassina</emphasis>
</taxonomicName>
, male; (C)
<taxonomicName box="[538,721,1299,1320]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[538,721,1299,1320]" italics="true" pageId="18" pageNumber="227">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[725,772,1300,1321]" pageId="18" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
, female; (D)
<taxonomicName box="[891,1074,1299,1320]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[891,1074,1299,1320]" italics="true" pageId="18" pageNumber="227">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[106,152,1329,1350]" pageId="18" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
, male. The postcervical plate and exposed thoracic membrane are contained within the boundaries of the intercervical sclerites and the episternum, with either a broad (
<taxonomicName box="[765,944,1358,1379]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[765,944,1358,1379]" italics="true" pageId="18" pageNumber="227">Ilomantis thalassina</emphasis>
</taxonomicName>
, A and B) or narrow (
<taxonomicName box="[189,376,1387,1408]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="227" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[189,376,1387,1408]" italics="true" pageId="18" pageNumber="227">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[385,432,1388,1409]" pageId="18" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
, C and D) hexagonal shape. Abbreviations: eps, episternum; etm, exposed thoracic membrane; ics, intercervical sclerite; lcs, lateral cervical sclerite; pcpl, postcervical plate; tss, T-shaped sclerite.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/269853/files/figure.png" pageId="19" pageNumber="228" targetBox="[365,799,131,582]" targetPageId="19">
<paragraph blockId="19.[98,1066,607,718]" pageId="19" pageNumber="228">
<emphasis bold="true" box="[98,177,607,629]" pageId="19" pageNumber="228">Fig. 14.</emphasis>
Oblique ventral view of the furcasternal tubercle (FT) located medially at the base of the T-shaped sclerite, posterior to the prothoracic coxae. Arrow indicates FT. Species shown:
<taxonomicName class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="species" species="floweri">
<emphasis italics="true" pageId="19" pageNumber="228">Nilomantis floweri</emphasis>
</taxonomicName>
, male. This figure is published in colour in the online edition of this journal, which can be accessed via http://booksandjournals.brillonline.com/content/journals/1876312x.
</paragraph>
</caption>
<subSubSection pageId="19" pageNumber="228" type="key">
<paragraph blockId="19.[98,595,1087,1114]" box="[98,595,1087,1114]" pageId="19" pageNumber="228">
<heading bold="true" box="[98,595,1087,1114]" fontSize="11" level="1" pageId="19" pageNumber="228" reason="0">
<emphasis bold="true" box="[98,595,1087,1114]" pageId="19" pageNumber="228">
Key to
<taxonomicName authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[184,506,1087,1114]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[184,298,1087,1114]" italics="true" pageId="19" pageNumber="228">Ilomantis</emphasis>
<bibRefCitation author="Giglio-Tos, E." box="[305,506,1087,1113]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino" pageId="19" pageNumber="228" pagination="1 - 16" part="30" refString="Giglio-Tos, E. (1915) Mantidi esotici. VIII. Acromantinae. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino 30: 1 - 16." title="Mantidi esotici. VIII. Acromantinae" type="journal article" year="1915">
Giglio-Tos,
<number box="[445,506,1087,1113]" pageId="19" pageNumber="228" value="1915.0">1915</number>
</bibRefCitation>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[98,1066,1140,1270]" pageId="19" pageNumber="228">
<number box="[98,114,1140,1166]" pageId="19" pageNumber="228" value="1.0">1</number>
. Medial keel of pronotum originates from the anterior margin of prozone. The lower frons features a latitudinal, medially bifurcated sulcus that traverses the anterior margin of the sclerite. Postcervical plate and exposed thoracic membrane broadly framed by the intercervical sclerites and episternum.
<taxonomicName box="[957,1064,1244,1270]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[957,1064,1244,1270]" italics="true" pageId="19" pageNumber="228">thalassina</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="19.[98,1066,1313,1444]" pageId="19" pageNumber="228">
<number box="[98,113,1313,1339]" pageId="19" pageNumber="228" value="1.0">1</number>
. Medial keel of pronotum originates from the mid-prozone. The lower frons features a latitudinal, medially bifid sulcus that traverses the anterior region of the sclerite, but not across the anterior margin. Postcervical plate and exposed thoracic membrane narrowly framed by the intercervical sclerites and episternum.
<taxonomicName box="[896,1007,1417,1443]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[896,1007,1417,1443]" italics="true" pageId="19" pageNumber="228">ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1013,1066,1418,1444]" pageId="19" pageNumber="228" rank="species">sp.n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,610 @@
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<mods:title>Leveraging female genitalic characters for generic and species delimitation in Nilomantis Werner, 1907 and Ilomantis Giglio-Tos, 1915 (Mantodea, Nilomantinae)</mods:title>
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<taxonomicName authority="Saussure, 1899" authorityName="Saussure" authorityYear="1899" box="[98,546,1520,1547]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="species" species="thalassina">
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<emphasis bold="true" box="[98,340,1520,1547]" italics="true" pageId="19" pageNumber="228">Ilomantis thalassina</emphasis>
(
<bibRefCitation author="Saussure, H." box="[354,537,1520,1546]" editor="Voeltzkow, A." journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1" pageId="19" pageNumber="228" pagination="567 - 664" part="24" refString="Saussure, H. (1899). In: Voeltzkow, A.: Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1 (24): 567 - 664." type="journal article" volumeTitle="Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae" year="1899">
Saussure,
<number box="[475,537,1520,1546]" pageId="19" pageNumber="228" value="1899.0">1899</number>
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)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection lastPageId="20" lastPageNumber="229" pageId="19" pageNumber="228" type="reference_group">
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<taxonomicName authority="Saussure, 1899: 593" authorityName="Saussure" authorityPageNumber="593" authorityYear="1899" box="[98,608,1573,1600]" class="Insecta" family="Iridopterygidae" genus="Tropidomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[98,370,1573,1599]" italics="true" pageId="19" pageNumber="228">Tropidomantis thalassina</emphasis>
<treatmentCitation author="Saussure, H." box="[377,608,1573,1600]" page="593" pageId="19" pageNumber="228" year="1899">
<bibRefCitation author="Saussure, H." box="[377,550,1573,1599]" editor="Voeltzkow, A." journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1" pageId="19" pageNumber="228" pagination="567 - 664" part="24" refString="Saussure, H. (1899). In: Voeltzkow, A.: Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1 (24): 567 - 664." type="journal article" volumeTitle="Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae" year="1899">
Saussure,
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</bibRefCitation>
:
<number box="[561,608,1573,1600]" pageId="19" pageNumber="228" value="593.0">593</number>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation author="Kirby, W. F." box="[620,814,1573,1599]" page="227" pageId="19" pageNumber="228" year="1904">
<bibRefCitation author="Kirby, W. F." box="[620,758,1573,1599]" journalOrPublisher="Bulletin of the British Museum (Natural History), London" pageId="19" pageNumber="228" pagination="207 - 316" part="1" refString="Kirby, W. F. (1904) A synonymic Catalogue of Orthoptera. (Forficilidae, Hemimeridae, Blattidae, Mantidae and Phasmidae). Bulletin of the British Museum (Natural History), London 1: 207 - 316." title="A synonymic Catalogue of Orthoptera. (Forficilidae, Hemimeridae, Blattidae, Mantidae and Phasmidae)" type="journal article" year="1904">
Kirby,
<number box="[696,758,1573,1599]" pageId="19" pageNumber="228" value="1904.0">1904</number>
</bibRefCitation>
:
<number box="[770,814,1573,1599]" pageId="19" pageNumber="228" value="227.0">227</number>
</treatmentCitation>
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</paragraph>
<paragraph blockId="19.[98,1066,1573,1669]" pageId="19" pageNumber="228">
<treatmentCitationGroup pageId="19" pageNumber="228">
<emphasis box="[98,495,1608,1634]" italics="true" pageId="19" pageNumber="228">
<taxonomicName box="[98,200,1608,1634]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="genus">Ilomantis</taxonomicName>
[
<taxonomicName box="[217,373,1608,1634]" class="Insecta" family="Iridopterygidae" genus="Tropidomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="genus">Tropidomantis</taxonomicName>
]
<taxonomicName authority="(Saussure, 1899) Giglio-Tos, 1915: 46" authorityName="(Saussure, 1899) Giglio-Tos" authorityPageNumber="46" authorityYear="1915" box="[388,932,1608,1635]" class="Insecta" family="Iridopterygidae" genus="Tropidomantis" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="228" phylum="Arthropoda" rank="species" species="thalassina">
thalassina (
<bibRefCitation author="Saussure, H." box="[509,680,1608,1634]" editor="Voeltzkow, A." journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1" pageId="19" pageNumber="228" pagination="567 - 664" part="24" refString="Saussure, H. (1899). In: Voeltzkow, A.: Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1 (24): 567 - 664." type="journal article" volumeTitle="Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae" year="1899">
Saussure,
<number box="[620,680,1608,1634]" pageId="19" pageNumber="228" value="1899.0">1899</number>
</bibRefCitation>
)
<treatmentCitation author="Giglio-Tos, E." box="[694,932,1608,1635]" page="46" pageId="19" pageNumber="228" year="1915">
<bibRefCitation author="Giglio-Tos, E." box="[694,890,1608,1635]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino" pageId="19" pageNumber="228" pagination="1 - 16" part="30" refString="Giglio-Tos, E. (1915) Mantidi esotici. VIII. Acromantinae. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino 30: 1 - 16." title="Mantidi esotici. VIII. Acromantinae" type="journal article" year="1915">
Giglio-Tos,
<number box="[828,890,1608,1635]" pageId="19" pageNumber="228" value="1915.0">1915</number>
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: 46
</treatmentCitation>
</taxonomicName>
</emphasis>
, 1927:
<number box="[1014,1060,1608,1634]" pageId="19" pageNumber="228" value="137.0">137</number>
;
<treatmentCitation author="Paulian, R." box="[98,295,1642,1669]" page="30" pageId="19" pageNumber="228" year="1957">
<bibRefCitation author="Paulian, R." box="[98,251,1642,1669]" journalOrPublisher="Insectes Mantodea. Publications de l'Institut de Recherche Scientifique Tananarive - Tsimbazaza" pageId="19" pageNumber="228" pagination="1 - 102" part="5" refString="Paulian, R. (1957) Faune de Madagascar. Insectes Mantodea. Publications de l'Institut de Recherche Scientifique Tananarive - Tsimbazaza 5: 1 - 102." title="Faune de Madagascar" type="journal article" year="1957">
Paulian
<number box="[189,251,1642,1669]" pageId="19" pageNumber="228" value="1957.0">1957</number>
</bibRefCitation>
:
<number box="[263,295,1643,1669]" pageId="19" pageNumber="228" value="30.0">30</number>
</treatmentCitation>
;
<treatmentCitation author="Roy, R." box="[307,479,1643,1669]" page="121" pageId="19" pageNumber="228" year="1987">
<bibRefCitation author="Roy, R." box="[307,421,1643,1669]" journalOrPublisher="Bulletin de la Societe zoologique de France" pageId="19" pageNumber="228" pagination="117 - 125" part="112" refString="Roy, R. (1987) Le peuplement en Mantes des iles situees au large de l'Afrique. Bulletin de la Societe zoologique de France 112: 117 - 125." title="Le peuplement en Mantes des iles situees au large de l'Afrique" type="journal article" year="1987">
Roy
<number box="[359,421,1643,1669]" pageId="19" pageNumber="228" value="1987.0">1987</number>
</bibRefCitation>
:
<number box="[433,479,1643,1669]" pageId="19" pageNumber="228" value="121.0">121</number>
</treatmentCitation>
;
<treatmentCitation author="Ehrmann, R." box="[491,731,1642,1669]" page="193" pageId="19" pageNumber="228" year="2002">
<bibRefCitation author="Ehrmann, R." box="[491,675,1642,1669]" journalOrPublisher="Munster, Natur und Tier Verlag Munster" pageId="19" pageNumber="228" refString="Ehrmann, R. (2002) Mantodea: Gottesanberinnen der Welt. Munster, Natur und Tier Verlag Munster." title="Mantodea: Gottesanberinnen der Welt" type="book" year="2002">
Ehrmann,
<number box="[613,675,1643,1669]" pageId="19" pageNumber="228" value="2002.0">2002</number>
</bibRefCitation>
:
<number box="[687,731,1643,1669]" pageId="19" pageNumber="228" value="193.0">193</number>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="20.[106,1074,134,196]" pageId="20" pageNumber="229">
<treatmentCitationGroup pageId="20" pageNumber="229">
<emphasis box="[106,463,134,160]" italics="true" pageId="20" pageNumber="229">
<taxonomicName box="[106,227,134,160]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="20" pageNumber="229" phylum="Arthropoda" rank="genus">Nilomantis</taxonomicName>
[
<taxonomicName box="[242,341,134,160]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="20" pageNumber="229" phylum="Arthropoda" rank="genus">Ilomantis</taxonomicName>
]
<taxonomicName authority="(Saussure, 1899) Beier 1935: 54" authorityName="(Saussure, 1899) Beier" authorityPageNumber="54" authorityYear="1935" box="[357,820,134,161]" class="Insecta" family="Iridopterygidae" genus="Tropidomantis" kingdom="Animalia" order="Mantodea" pageId="20" pageNumber="229" phylum="Arthropoda" rank="species" species="thalassina">
thalassina (
<bibRefCitation author="Saussure, H." box="[475,644,135,161]" editor="Voeltzkow, A." journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1" pageId="20" pageNumber="229" pagination="567 - 664" part="24" refString="Saussure, H. (1899). In: Voeltzkow, A.: Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 1 (24): 567 - 664." type="journal article" volumeTitle="Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889 - 1895. Orthoptera: Mantidae" year="1899">
Saussure,
<number box="[584,644,135,161]" pageId="20" pageNumber="229" value="1899.0">1899</number>
</bibRefCitation>
)
<treatmentCitation author="Beier, M." box="[657,820,134,161]" page="54" pageId="20" pageNumber="229" year="1935">
<bibRefCitation author="Beier, M." box="[657,782,134,161]" journalOrPublisher="Wytsman, Brussels" pageId="20" pageNumber="229" refString="Beier, M. (1935) Genera Insectorum, Vol. 201. Mantodea. Subfamilie: Orthoderinae, Choeradodinae, Deroplatinae. Wytsman, Brussels." title="Genera Insectorum, Vol. 201. Mantodea. Subfamilie: Orthoderinae, Choeradodinae, Deroplatinae" type="book" year="1935">
Beier
<number box="[720,782,134,161]" pageId="20" pageNumber="229" value="1935.0">1935</number>
</bibRefCitation>
:
<number box="[790,866,134,161]" pageId="20" pageNumber="229" value="54.5" valueMax="55.0" valueMin="54.0">5455</number>
</treatmentCitation>
</taxonomicName>
</emphasis>
;
<treatmentCitation author="Otte, D. &amp; Spearman, L. A." page="120" pageId="20" pageNumber="229" year="2005">
<bibRefCitation author="Otte, D. &amp; Spearman, L. A." journalOrPublisher="Catalog of the mantids of the world" pageId="20" pageNumber="229" part="1" refString="Otte, D. &amp; Spearman, L. A. (2005) Mantida species file. Catalog of the mantids of the world, volume 1. Association of Insect Diversity, Philadelphia, PA." title="Mantida species file" type="journal volume" year="2005">
Otte &amp; Spearman
<number box="[106,168,169,196]" pageId="20" pageNumber="229" value="2005.0">2005</number>
</bibRefCitation>
:
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</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="229" type="materials_examined">
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<heading box="[106,230,239,264]" fontSize="11" level="2" pageId="20" pageNumber="229" reason="1">
<emphasis box="[106,230,239,264]" italics="true" pageId="20" pageNumber="229">
<typeStatus box="[106,155,239,264]" pageId="20" pageNumber="229">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[106,1074,290,352]" box="[106,1073,290,317]" pageId="20" pageNumber="229">
<typeStatus box="[106,213,290,316]" pageId="20" pageNumber="229">Holotype</typeStatus>
♂. Senckenberg Forschungsinstitut und Naturmuseum, Frankfurt,
<collectingCountry box="[960,1069,291,317]" name="Germany" pageId="20" pageNumber="229">Germany</collectingCountry>
;
</paragraph>
<paragraph blockId="20.[106,1074,290,352]" box="[106,1074,325,352]" pageId="20" pageNumber="229">
Allotype ♀. Senckenberg Forschungsinstitut und Naturmuseum, Frankfurt,
<collectingCountry box="[962,1070,325,351]" name="Germany" pageId="20" pageNumber="229">Germany</collectingCountry>
.
</paragraph>
<paragraph blockId="20.[106,248,394,420]" box="[106,248,394,420]" pageId="20" pageNumber="229">
<heading box="[106,248,394,420]" fontSize="11" level="2" pageId="20" pageNumber="229" reason="1">
<emphasis box="[106,248,394,420]" italics="true" pageId="20" pageNumber="229">
<typeStatus box="[106,155,395,420]" pageId="20" pageNumber="229">Type</typeStatus>
location
</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[106,355,446,473]" box="[106,355,446,473]" pageId="20" pageNumber="229">
<heading box="[106,355,446,473]" fontSize="11" level="2" pageId="20" pageNumber="229" reason="1">
<collectingCountry box="[106,165,447,473]" name="Iceland" pageId="20" pageNumber="229">Nosy</collectingCountry>
Be,
<collectingCountry box="[216,351,446,472]" name="Madagascar" pageId="20" pageNumber="229">Madagascar</collectingCountry>
.
</heading>
</paragraph>
<paragraph blockId="20.[106,326,516,542]" box="[106,326,516,542]" pageId="20" pageNumber="229">
<heading box="[106,326,516,542]" fontSize="11" level="2" pageId="20" pageNumber="229" reason="1">
<emphasis box="[106,326,516,542]" italics="true" pageId="20" pageNumber="229">Examined specimens</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[106,1074,568,907]" pageId="20" pageNumber="229">
The list consists of
<specimenCount box="[308,443,568,594]" pageId="20" pageNumber="229" type="generic">
<number box="[308,322,568,594]" pageId="20" pageNumber="229" value="8.0">8</number>
specimens
</specimenCount>
.
<collectingCountry box="[452,584,568,594]" name="Madagascar" pageId="20" pageNumber="229">Madagascar</collectingCountry>
:
<emphasis box="[593,703,568,594]" italics="true" pageId="20" pageNumber="229">Ampijoroa</emphasis>
,
<specimenCount box="[716,752,568,595]" pageId="20" pageNumber="229" type="generic">
<number box="[716,730,568,594]" pageId="20" pageNumber="229" value="1.0">1</number>
</specimenCount>
, Ankarafantsika,
<quantity box="[941,1008,568,594]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="20" pageNumber="229" unit="m" value="170.0">170m</quantity>
,
<date box="[1019,1069,568,595]" pageId="20" pageNumber="229" value="1957-01">
I-
<number box="[1039,1069,568,595]" pageId="20" pageNumber="229" value="57.0">57</number>
</date>
, R.E., Insitut Scientifique de
<collectingCountry box="[412,542,602,628]" name="Madagascar" pageId="20" pageNumber="229">Madagascar</collectingCountry>
, Genitalia No. 0 0 0
<geoCoordinate box="[748,799,603,629]" direction="south" orientation="latitude" pageId="20" pageNumber="229" precision="925" value="-0.11666667">
0
<number box="[762,776,603,629]" pageId="20" pageNumber="229" value="7.0">7</number>
S
</geoCoordinate>
. K. Brannoch (
<collectionCode box="[962,1061,603,629]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="229">MNHN</collectionCode>
);
<emphasis box="[106,288,637,663]" italics="true" pageId="20" pageNumber="229">Distr. dAmbanja</emphasis>
,
<number box="[303,317,637,663]" pageId="20" pageNumber="229" value="1.0">1</number>
♀,
<collectingCountry box="[353,482,637,663]" name="Madagascar" pageId="20" pageNumber="229">Madagascar</collectingCountry>
Nord, N. de Beangona-Ambevy, Vallée dAntremabe,
<quantity box="[106,174,672,698]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="229" unit="m" value="400.0">400m</quantity>
,
<date box="[187,275,672,698]" pageId="20" pageNumber="229" value="1964-02">
II-
<number box="[216,275,672,698]" pageId="20" pageNumber="229" value="1964.0">1964</number>
</date>
, P. Soga, Genitalia No. 0 0 0
<geoCoordinate box="[582,635,672,698]" direction="south" orientation="latitude" pageId="20" pageNumber="229" precision="925" value="-0.15">
0
<number box="[596,610,672,698]" pageId="20" pageNumber="229" value="9.0">9</number>
S
</geoCoordinate>
. K. Brannoch (
<collectionCode box="[807,905,672,698]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="229">MNHN</collectionCode>
);
<emphasis box="[926,1022,672,698]" italics="true" pageId="20" pageNumber="229">Nossi Bé,</emphasis>
<specimenCount box="[1030,1066,672,699]" pageId="20" pageNumber="229" type="generic">
<number box="[1030,1044,672,698]" pageId="20" pageNumber="229" value="1.0">1</number>
</specimenCount>
, Voeltzkow, genitalia R. Roy No.
<number box="[479,536,707,733]" pageId="20" pageNumber="229" value="4072.0">4072</number>
(
<collectionCode box="[555,653,707,733]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="229">MNHN</collectionCode>
);
<number box="[676,690,707,733]" pageId="20" pageNumber="229" value="2.0">2</number>
♀ (
<collectionCode box="[731,829,707,733]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="229">MNHN</collectionCode>
);
<number box="[852,866,707,733]" pageId="20" pageNumber="229" value="1.0">1</number>
♀,
<number box="[905,965,707,733]" pageId="20" pageNumber="229" value="1900.0">1900</number>
, Dr. Joly, Genitalia No. 0 0 0
<geoCoordinate box="[300,349,741,767]" direction="south" orientation="latitude" pageId="20" pageNumber="229" precision="925" value="-0.13333334">
0
<number box="[315,328,741,767]" pageId="20" pageNumber="229" value="8.0">8</number>
S
</geoCoordinate>
. K. Brannoch (
<collectionCode box="[507,605,741,767]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="229">MNHN</collectionCode>
);
<emphasis box="[623,833,741,768]" italics="true" pageId="20" pageNumber="229">Nossi Bé Sambirano,</emphasis>
<number box="[837,851,741,767]" pageId="20" pageNumber="229" value="2.0">2</number>
♀, J. de Coudenfiore (
<collectionCode box="[114,212,776,802]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="229">MNHN</collectionCode>
);
<emphasis box="[240,383,776,802]" italics="true" pageId="20" pageNumber="229">Toliara Prov.</emphasis>
, Réserve Spéciale de Cap Sainte Marie,
<quantity box="[872,975,776,802]" metricMagnitude="4" metricUnit="m" metricValue="1.23" pageId="20" pageNumber="229" unit="km" value="12.3">12.3 km</quantity>
,
<number box="[994,1037,776,802]" pageId="20" pageNumber="229" value="262.0">262</number>
°W Marovato, elev.
<quantity box="[279,352,811,837]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="229" unit="m" value="200.0">200 m</quantity>
<date box="[358,543,810,837]" pageId="20" pageNumber="229" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">
<number box="[358,430,810,837]" pageId="20" pageNumber="229" value="13.0" valueMax="15.0" valueMin="11.0">1115</number>
Feb 2002
</date>
,
<geoCoordinate box="[554,680,810,837]" direction="south" orientation="latitude" pageId="20" pageNumber="229" precision="15" value="-23.581667">
23°
<number box="[592,621,810,837]" pageId="20" pageNumber="229" value="34.0">34</number>
<number box="[626,656,810,837]" pageId="20" pageNumber="229" value="54.0">54</number>
”S
</geoCoordinate>
<geoCoordinate box="[687,803,810,837]" direction="east" orientation="longitude" pageId="20" pageNumber="229" precision="15" value="45.168335">
<number box="[687,717,810,837]" pageId="20" pageNumber="229" value="45.0">45</number>
°
<number box="[727,755,810,837]" pageId="20" pageNumber="229" value="10.0">10</number>
<number box="[760,774,810,837]" pageId="20" pageNumber="229" value="6.0">6</number>
”E
</geoCoordinate>
, coll: Fisher, Griswold et al., California Acad. of Sciences, at light in spiny forest thicket, collection code: BLF
<number box="[1013,1069,845,872]" pageId="20" pageNumber="229" value="5501.0">5501</number>
, CASENT 3006172, 5501, MN
<number box="[455,498,880,906]" pageId="20" pageNumber="229" value="413.0">413</number>
Primary Voucher,
<collectionCode box="[704,786,880,906]" pageId="20" pageNumber="229">GSMC</collectionCode>
000141 (CASENT).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="229" type="biology_ecology">
<paragraph blockId="20.[106,270,949,975]" box="[106,270,949,975]" pageId="20" pageNumber="229">
<heading box="[106,270,949,975]" fontSize="11" level="2" pageId="20" pageNumber="229" reason="1">
<emphasis box="[106,270,949,975]" italics="true" pageId="20" pageNumber="229">Natural history</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[106,1074,1001,1062]" pageId="20" pageNumber="229">
Specimens studied were collected between January and February in Northern
<collectingCountry box="[938,1070,1001,1027]" name="Madagascar" pageId="20" pageNumber="229">Madagascar</collectingCountry>
. No specific behavioral or ecological information is known from the literature.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="229" type="diagnosis">
<paragraph blockId="20.[106,210,1106,1131]" box="[106,210,1106,1131]" pageId="20" pageNumber="229">
<heading box="[106,210,1106,1131]" fontSize="11" level="2" pageId="20" pageNumber="229" reason="1">
<emphasis box="[106,210,1106,1131]" italics="true" pageId="20" pageNumber="229">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[106,1074,1157,1530]" pageId="20" pageNumber="229">
Pronotum with a medial keel that runs from the anterior margin of the prozone to the posterior margin of the metazone. Postcervical plate and exposed thoracic membrane of the cervix broad with an approximately hexagonal shape. Females (ca.
<quantity box="[916,1066,1226,1253]" metricMagnitude="-2" metricUnit="m" metricValue="2.675" metricValueMax="2.7" metricValueMin="2.65" pageId="20" pageNumber="229" unit="mm" value="26.75" valueMax="27.0" valueMin="26.5">26.527 mm</quantity>
) relatively longer than males (ca.
<quantity box="[473,582,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="229" unit="mm" value="22.5">22.5 mm</quantity>
); both females and males of this species are relatively longer than those within
<taxonomicName box="[495,718,1296,1322]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="20" pageNumber="229" phylum="Arthropoda" rank="species" species="ginsburgae" status="n.sp.">
<emphasis box="[495,718,1296,1322]" italics="true" pageId="20" pageNumber="229">Ilomantis ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[724,778,1297,1323]" pageId="20" pageNumber="229" rank="species">n.sp.</taxonomicNameLabel>
Female genitalia: gonapophysis VIII (GA) with pronounced medial outgrowth (MO); gonoplac (GL) with divergent apical cleft (AC). Male genitalia: processo distale (pda) small, adjacent to a relatively rounded posterodextral margin; processo ventrale sclerificato (pva) slightly arched along the anterior margin with a relatively straight posterior margin. Distributed within the Southern region of the
<collectingCountry box="[501,777,1469,1495]" name="Madagascar" pageId="20" pageNumber="229">Republic of Madagascar</collectingCountry>
.
<emphasis box="[789,805,1470,1495]" italics="true" pageId="20" pageNumber="229">F</emphasis>
=
<number box="[821,833,1469,1495]" pageId="20" pageNumber="229" value="3.0">3</number>
DS/
<number box="[879,946,1469,1495]" pageId="20" pageNumber="229" value="11.5" valueMax="12.0" valueMin="11.0">1112</number>
AvS/
<number box="[1007,1021,1469,1495]" pageId="20" pageNumber="229" value="4.0">4</number>
PvS;
<emphasis box="[106,125,1505,1530]" italics="true" pageId="20" pageNumber="229">T</emphasis>
=
<number box="[140,208,1504,1530]" pageId="20" pageNumber="229" value="12.5" valueMax="14.0" valueMin="11.0">1114</number>
AvS/
<number box="[269,339,1504,1530]" pageId="20" pageNumber="229" value="13.0" valueMax="14.0" valueMin="12.0">1214</number>
PvS.
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="232" pageId="20" pageNumber="229" type="description">
<paragraph blockId="20.[106,1074,1573,1669]" box="[106,823,1573,1600]" pageId="20" pageNumber="229">
<emphasis box="[106,249,1573,1599]" italics="true" pageId="20" pageNumber="229">Redescription</emphasis>
(all measurements in mm, unless noted otherwise)
</paragraph>
<paragraph blockId="20.[106,1074,1573,1669]" lastBlockId="21.[98,1066,134,300]" lastPageId="21" lastPageNumber="230" pageId="20" pageNumber="229">
<emphasis box="[106,181,1608,1634]" italics="true" pageId="20" pageNumber="229">Female</emphasis>
(
<figureCitation box="[197,258,1608,1634]" captionStart="Fig. 1" captionStartId="2.[106,138,1569,1591]" captionTargetBox="[155,1024,396,1542]" captionTargetId="figure@2.[155,1024,396,1544]" captionTargetPageId="2" captionText="Fig. 1. Ilomantis thalassina. (A) Female, dorsal habitus; (B) female, ventral habitus; (C) male, dorsal habitus; (D) male, ventral habitus. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269840/files/figure.png" pageId="20" pageNumber="229">
Fig.
<number box="[248,258,1608,1634]" pageId="20" pageNumber="229" value="1.0">1</number>
</figureCitation>
A,B). Length
<number box="[416,548,1608,1635]" pageId="20" pageNumber="229" value="26.775" valueMax="27.05" valueMin="26.5">26.527.05</number>
; forewing length
<number box="[744,876,1608,1635]" pageId="20" pageNumber="229" value="19.09" valueMax="19.6" valueMin="18.58">18.5819.6</number>
; hindwing length
<number box="[106,238,1642,1669]" pageId="20" pageNumber="229" value="19.88" valueMax="20.56" valueMin="19.2">19.220.56</number>
; pronotum length
<number box="[443,560,1642,1669]" pageId="20" pageNumber="229" value="5.425000000000001" valueMax="5.53" valueMin="5.32">5.325.53</number>
; prozone length 2.0
<number box="[793,830,1643,1669]" pageId="20" pageNumber="229" value="2.2">2.2</number>
; pronotum maximum width
<number box="[171,289,134,161]" pageId="21" pageNumber="230" value="2.04" valueMax="2.13" valueMin="1.95">1.952.13</number>
; pronotum minimum width
<number box="[618,735,134,161]" pageId="21" pageNumber="230" value="1.455" valueMax="1.52" valueMin="1.39">1.391.52</number>
; head width
<number box="[883,1000,134,161]" pageId="21" pageNumber="230" value="4.57" valueMax="4.58" valueMin="4.56">4.564.58</number>
; head vertex to clypeus
<number box="[286,403,169,195]" pageId="21" pageNumber="230" value="1.7999999999999998" valueMax="1.97" valueMin="1.63">1.631.97</number>
; frons width
<number box="[546,663,169,196]" pageId="21" pageNumber="230" value="1.8050000000000002" valueMax="1.85" valueMin="1.76">1.761.85</number>
; frons height
<number box="[811,929,169,196]" pageId="21" pageNumber="230" value="0.5" valueMax="0.53" valueMin="0.47">0.470.53</number>
; prothoracic femur length
<number box="[250,354,204,231]" pageId="21" pageNumber="230" value="6.37" valueMax="6.54" valueMin="6.2">6.26.54</number>
; mesothoracic femur length
<number box="[670,774,204,231]" pageId="21" pageNumber="230" value="6.01" valueMax="6.12" valueMin="5.9">5.96.12</number>
; mesothoracic tibia length
<number box="[98,218,239,265]" pageId="21" pageNumber="230" value="4.135" valueMax="4.17" valueMin="4.1">4.104.17</number>
; mesothoracic tarsus length
<number box="[553,607,238,264]" pageId="21" pageNumber="230" value="3.06">3.06</number>
; metathoracic femur length
<number box="[942,1062,238,265]" pageId="21" pageNumber="230" value="6.475" valueMax="6.59" valueMin="6.36">6.366.59</number>
; metathoracic tibia length
<number box="[382,500,273,299]" pageId="21" pageNumber="230" value="6.92" valueMax="7.01" valueMin="6.83">6.837.01</number>
; metathoracic tarsus length
<number box="[807,926,273,299]" pageId="21" pageNumber="230" value="4.505" valueMax="4.74" valueMin="4.27">4.274.74</number>
.
</paragraph>
<paragraph blockId="21.[98,1066,342,542]" pageId="21" pageNumber="230">
<emphasis box="[98,157,342,368]" italics="true" pageId="21" pageNumber="230">Head</emphasis>
(
<figureCitation box="[172,237,342,369]" captionStart="Fig. 7" captionStartId="11.[98,131,884,906]" captionTargetBox="[147,1016,131,858]" captionTargetId="figure@11.[147,1016,131,859]" captionTargetPageId="11" captionText="Fig. 7. Species distribution map for Nilomantis and Ilomantis species. Examined Nilomantis and Ilomantis specimens, as well as additional Nilomantis taxa listed in Roy &amp; Leston (1975), have been incorporated. Collecting localities without coordinates were georeferenced using Google Earth. Ilomantis species are distributed within Madagascar; Nilomantis species are distributed across the Sahara-Sahel region of Africa, including Ghana, the Central African Republic, and Saudi Arabia. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269846/files/figure.png" pageId="21" pageNumber="230">
Fig.
<number box="[223,237,343,369]" pageId="21" pageNumber="230" value="7.0">7</number>
</figureCitation>
A). Cranial vertex elevation yellow. Area posterior to the lateral ocelli concave. Ocelli reduced with relatively larger, lateral ocelli sitting within a slightly raised cylindrical ocellar tubercle; the smaller, unpaired median ocellus positioned anterior to a sulcus that lies medially between the unpaired ocellus and the antennal sclerites, spanning the distance between the antennae; area surrounding the median ocellus is very slightly raised. Antennae extend to mid-body.
</paragraph>
<paragraph blockId="21.[98,1066,585,716]" pageId="21" pageNumber="230">
<emphasis box="[98,182,585,611]" italics="true" pageId="21" pageNumber="230">Thorax.</emphasis>
Pronotal medial keel reaching anterior margin (
<figureCitation box="[741,822,585,611]" captionStart="Fig. 11" captionStartId="16.[106,139,1517,1539]" captionTargetBox="[243,935,131,1491]" captionTargetId="figure@16.[243,937,131,1492]" captionTargetPageId="16" captionText="Fig. 11. Illustrations of dorsal perspective of the Ilomantis pronotum (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. Ilomantis thalassina exhibits a medial keel that fully traverses the pronotum (A and B); Ilomantis ginsburgae sp. n. exhibits a medial keel that originates in the mid-prozone and concludes at the posterior pronotal margin (C and D). MK, medial keel." httpUri="https://zenodo.org/record/269850/files/figure.png" pageId="21" pageNumber="230">
Fig.
<number box="[795,822,585,611]" pageId="21" pageNumber="230" value="11.0">11</number>
</figureCitation>
A). Postcervical plate and exposed thoracic membrane of the cervix broad with an approximately hexagonal shape (
<figureCitation box="[177,257,654,681]" captionStart="Fig. 13" captionStartId="18.[106,139,1269,1291]" captionTargetBox="[155,1024,131,1244]" captionTargetId="figure@18.[155,1024,131,1244]" captionTargetPageId="18" captionText="Fig. 13. Illustrations of ventral perspective of the Ilomantis cervix (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. The postcervical plate and exposed thoracic membrane are contained within the boundaries of the intercervical sclerites and the episternum, with either a broad (Ilomantis thalassina, A and B) or narrow (Ilomantis ginsburgae sp. n., C and D) hexagonal shape. Abbreviations: eps, episternum; etm, exposed thoracic membrane; ics, intercervical sclerite; lcs, lateral cervical sclerite; pcpl, postcervical plate; tss, T-shaped sclerite." httpUri="https://zenodo.org/record/269852/files/figure.png" pageId="21" pageNumber="230">
Fig.
<number box="[229,257,655,681]" pageId="21" pageNumber="230" value="13.0">13</number>
</figureCitation>
A). Forewings opaque with light green costal region, hindwings opaque to hyaline; costal vein of both fore- and hindwings with light ciliation.
</paragraph>
<paragraph blockId="21.[98,1066,758,819]" pageId="21" pageNumber="230">
<emphasis box="[98,271,758,784]" italics="true" pageId="21" pageNumber="230">Prothoracic legs.</emphasis>
Femur maximum width ca.
<number box="[593,682,758,785]" pageId="21" pageNumber="230" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5</number>
times the value of the male femur maximum width.
<emphasis box="[303,319,794,819]" italics="true" pageId="21" pageNumber="230">F</emphasis>
=
<number box="[334,346,793,819]" pageId="21" pageNumber="230" value="3.0">3</number>
DS/
<number box="[392,460,793,819]" pageId="21" pageNumber="230" value="11.5" valueMax="12.0" valueMin="11.0">1112</number>
AvS/
<number box="[521,535,793,819]" pageId="21" pageNumber="230" value="4.0">4</number>
PvS;
<emphasis box="[594,613,794,819]" italics="true" pageId="21" pageNumber="230">T</emphasis>
=
<number box="[628,697,793,819]" pageId="21" pageNumber="230" value="12.5" valueMax="13.0" valueMin="12.0">1213</number>
AvS/
<number box="[758,827,793,819]" pageId="21" pageNumber="230" value="13.0" valueMax="14.0" valueMin="12.0">1214</number>
PvS.
</paragraph>
<paragraph blockId="21.[98,1066,862,993]" pageId="21" pageNumber="230">
<emphasis box="[98,403,862,888]" italics="true" pageId="21" pageNumber="230">Meso- and metathoracic legs.</emphasis>
Lightly ciliated across surface with ciliation density increasing distally. Mesothoracic basitarsus approximately the same length as the remaining tarsal segments combined; metathoracic basitarsus
<number box="[665,702,932,959]" pageId="21" pageNumber="230" value="1.5">1.5</number>
times longer than the remaining tarsal segments combined.
</paragraph>
<paragraph blockId="21.[98,1066,1036,1096]" pageId="21" pageNumber="230">
<emphasis box="[98,209,1036,1062]" italics="true" pageId="21" pageNumber="230">Abdomen.</emphasis>
Supra-anal plate triangular, ciliated across the surface, narrower than the subgenital plate. Cerci filiform, tapering.
</paragraph>
<paragraph blockId="21.[98,1067,1140,1652]" lastBlockId="22.[106,1074,134,196]" lastPageId="22" lastPageNumber="231" pageId="21" pageNumber="230">
<emphasis box="[98,199,1140,1166]" italics="true" pageId="21" pageNumber="230">Genitalia</emphasis>
(
<figureCitation box="[216,285,1140,1166]" captionStart="Fig. 3" captionStartId="7.[98,131,1511,1533]" captionTargetBox="[147,1016,360,1484]" captionTargetId="figure@7.[147,1016,360,1485]" captionTargetPageId="7" captionText="Fig. 3. Female genitalic complex of Ilomantis shown in the lateral perspective. (A) Ilomantis thalassina; (B) Ilomantis ginsburgae sp. n. Abbreviations: AC, apical cleft; AL, apical lobe; GA, gonapophysis VIII; GL, gonoplac; MO, medial outgrowth; RTD, rectangular dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269842/files/figure.png" pageId="21" pageNumber="230">
Figs
<number box="[272,285,1140,1166]" pageId="21" pageNumber="230" value="3.0">3</number>
</figureCitation>
A,
<number box="[321,335,1140,1167]" pageId="21" pageNumber="230" value="5.0">5</number>
A,B). Gonapophysis VIII (GA) moderately slender, variably ciliated, and visibly curved from both the ventral (VH) and lateral (LH) perspectives; GA is heavily surfaced with fossettes. From VH, GA angles inward; a rugose, sclerotized tubercle projects posterolaterad from this region; apical lobe (AL) ovoid; heavily ciliated. AL with a membrane that projects from the internal dorsal margin of the structure, forming a shallow pocket as the membrane merges with the dorsal margin of GA, lending AL the appearance of being bilobed or mitten-shaped. From LH, GA is curved ventrad; GA ventral margin features a tumescent medial outgrowth (MO), which projects posterior to the crest of the curve. GA dorsal margin is invaginated anterior to the curve towards the ventral habitus. Gonapophysis IX (GP) is relatively short, apically mammiform, and invaginated along the ventral margin; GP ventral margin features a subulate overhang, the medial tine (MT). Gonoplac (GL) is shorter than GA, broad, tumescent, variably ciliated, and obtusely arched ventrad; from LH, GL base has a rectangular dilation on the anteroventral margin. GL features a prominent apical cleft (AC) with divergent, blunted tines. From LH, GL possesses a pronounced sclerotization that traverses the length of the structure, terminating just before the AC. GL ventral and dorsal margins are sculpturate, rugose.
</paragraph>
<paragraph blockId="22.[106,1074,238,507]" pageId="22" pageNumber="231">
<emphasis box="[106,162,238,264]" italics="true" pageId="22" pageNumber="231">Male</emphasis>
(
<figureCitation box="[176,238,238,265]" captionStart="Fig. 1" captionStartId="2.[106,138,1569,1591]" captionTargetBox="[155,1024,396,1542]" captionTargetId="figure@2.[155,1024,396,1544]" captionTargetPageId="2" captionText="Fig. 1. Ilomantis thalassina. (A) Female, dorsal habitus; (B) female, ventral habitus; (C) male, dorsal habitus; (D) male, ventral habitus. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269840/files/figure.png" pageId="22" pageNumber="231">
Fig.
<number box="[227,238,239,265]" pageId="22" pageNumber="231" value="1.0">1</number>
</figureCitation>
C,D). Body length
<number box="[460,610,238,265]" pageId="22" pageNumber="231" value="22.5" valueMax="22.51" valueMin="22.49">22.4922.51</number>
; forewing length
<number box="[804,954,238,264]" pageId="22" pageNumber="231" value="16.69" valueMax="16.92" valueMin="16.46">16.4616.92</number>
; hindwing length
<number box="[185,335,273,299]" pageId="22" pageNumber="231" value="17.36" valueMax="17.64" valueMin="17.08">17.0817.64</number>
; pronotum length
<number box="[551,671,273,299]" pageId="22" pageNumber="231" value="4.505000000000001" valueMax="4.69" valueMin="4.32">4.324.69</number>
; prozone length
<number box="[863,983,273,299]" pageId="22" pageNumber="231" value="1.75" valueMax="1.81" valueMin="1.69">1.691.81</number>
; pronotum maximum width
<number box="[360,480,308,334]" pageId="22" pageNumber="231" value="1.605" valueMax="1.72" valueMin="1.49">1.491.72</number>
; pronotum minimum width
<number box="[813,933,308,334]" pageId="22" pageNumber="231" value="1.2200000000000002" valueMax="1.36" valueMin="1.08">1.081.36</number>
; head width
<number box="[106,226,342,368]" pageId="22" pageNumber="231" value="3.92" valueMax="4.08" valueMin="3.76">3.764.08</number>
; head vertex to clypeus
<number box="[522,627,342,369]" pageId="22" pageNumber="231" value="1.58" valueMax="1.6" valueMin="1.56">1.561.6</number>
; frons width
<number box="[794,915,342,368]" pageId="22" pageNumber="231" value="1.47" valueMax="1.48" valueMin="1.46">1.461.48</number>
; frons height
<number box="[106,226,377,404]" pageId="22" pageNumber="231" value="0.5" valueMax="0.53" valueMin="0.47">0.470.53</number>
; prothoracic femur length
<number box="[525,645,377,404]" pageId="22" pageNumber="231" value="5.35" valueMax="5.48" valueMin="5.22">5.225.48</number>
; mesothoracic femur length
<number box="[963,1069,377,404]" pageId="22" pageNumber="231" value="5.525" valueMax="5.7" valueMin="5.35">5.355.7</number>
; mesothoracic tibia length
<number box="[405,510,412,438]" pageId="22" pageNumber="231" value="3.7199999999999998" valueMax="3.84" valueMin="3.6">3.63.84</number>
; mesothoracic tarsus length
<number box="[836,956,412,439]" pageId="22" pageNumber="231" value="2.875" valueMax="3.17" valueMin="2.58">2.583.17</number>
; metathoracic femur length
<number box="[335,455,446,473]" pageId="22" pageNumber="231" value="5.62" valueMax="5.71" valueMin="5.53">5.535.71</number>
; metathoracic tibia length
<number box="[779,833,446,472]" pageId="22" pageNumber="231" value="6.02">6.02</number>
; metathoracic tarsus length
<number box="[185,255,481,507]" pageId="22" pageNumber="231" value="2.903">2.903</number>
.
</paragraph>
<paragraph blockId="22.[106,1074,550,716]" pageId="22" pageNumber="231">
<emphasis box="[106,165,550,576]" italics="true" pageId="22" pageNumber="231">Head</emphasis>
(
<figureCitation box="[180,246,550,577]" captionStart="Fig. 8" captionStartId="12.[106,139,1140,1162]" captionTargetBox="[155,1023,131,1114]" captionTargetId="figure@12.[155,1024,131,1115]" captionTargetPageId="12" captionText="Fig. 8. Illustrations of anterior perspective of the Ilomantis and Nilomantis head (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male; (E) Nilomantis floweri, female; (F) Nilomantis floweri, male. (B, D, E, F) Ilomantis species feature a cosinusoidal vertical margin, whereas Nilomantis species feature a relatively straight vertical margin. Gray markings on the ocellar tubercle and within the corridors of the frontal sulci represent the presence of reddish-brown markings. LF, lower frons." httpUri="https://zenodo.org/record/269847/files/figure.png" pageId="22" pageNumber="231">
Fig.
<number box="[232,246,551,577]" pageId="22" pageNumber="231" value="8.0">8</number>
</figureCitation>
B). Cranial vertical margin and posterior half of lateral ocelli yellow. Area posterior to the lateral ocelli slightly concave. Ocelli well developed and sit atop a raised, ocellar tubercle; lateral ocelli relatively larger than unpaired, median ocellus. Reddish-brown pigmentation may or may not be present adjacent to the compound eyes, near the frontal sulci. Antennae extend past mid-abdomen.
</paragraph>
<paragraph blockId="22.[106,1073,758,854]" pageId="22" pageNumber="231">
<emphasis box="[106,190,758,784]" italics="true" pageId="22" pageNumber="231">Thorax.</emphasis>
Pronotum as in females (
<figureCitation box="[493,577,758,785]" captionStart="Fig. 11" captionStartId="16.[106,139,1517,1539]" captionTargetBox="[243,935,131,1491]" captionTargetId="figure@16.[243,937,131,1492]" captionTargetPageId="16" captionText="Fig. 11. Illustrations of dorsal perspective of the Ilomantis pronotum (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. Ilomantis thalassina exhibits a medial keel that fully traverses the pronotum (A and B); Ilomantis ginsburgae sp. n. exhibits a medial keel that originates in the mid-prozone and concludes at the posterior pronotal margin (C and D). MK, medial keel." httpUri="https://zenodo.org/record/269850/files/figure.png" pageId="22" pageNumber="231">
Fig.
<number box="[548,577,759,785]" pageId="22" pageNumber="231" value="11.0">11</number>
</figureCitation>
B). Postcervical plate and exposed thoracic membrane of the cervix as in females (
<figureCitation box="[547,629,793,819]" captionStart="Fig. 13" captionStartId="18.[106,139,1269,1291]" captionTargetBox="[155,1024,131,1244]" captionTargetId="figure@18.[155,1024,131,1244]" captionTargetPageId="18" captionText="Fig. 13. Illustrations of ventral perspective of the Ilomantis cervix (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. The postcervical plate and exposed thoracic membrane are contained within the boundaries of the intercervical sclerites and the episternum, with either a broad (Ilomantis thalassina, A and B) or narrow (Ilomantis ginsburgae sp. n., C and D) hexagonal shape. Abbreviations: eps, episternum; etm, exposed thoracic membrane; ics, intercervical sclerite; lcs, lateral cervical sclerite; pcpl, postcervical plate; tss, T-shaped sclerite." httpUri="https://zenodo.org/record/269852/files/figure.png" pageId="22" pageNumber="231">
Fig.
<number box="[599,629,793,819]" pageId="22" pageNumber="231" value="13.0">13</number>
</figureCitation>
B). Wings generally hyaline and iridescent, light ciliation across surface of both fore- and hindwings.
</paragraph>
<paragraph blockId="22.[106,1073,897,958]" pageId="22" pageNumber="231">
<emphasis box="[106,282,897,923]" italics="true" pageId="22" pageNumber="231">Prothoracic legs.</emphasis>
Femur maximum width ca.
<number box="[621,723,897,924]" pageId="22" pageNumber="231" value="0.725" valueMax="0.75" valueMin="0.7">0.70.75</number>
times the value of the female femur maximum width.
<emphasis box="[385,401,933,958]" italics="true" pageId="22" pageNumber="231">F</emphasis>
=
<number box="[417,429,932,958]" pageId="22" pageNumber="231" value="3.0">3</number>
DS/
<number box="[475,542,932,958]" pageId="22" pageNumber="231" value="11.5" valueMax="12.0" valueMin="11.0">1112</number>
AvS/
<number box="[603,617,932,958]" pageId="22" pageNumber="231" value="4.0">4</number>
PvS;
<emphasis box="[677,696,933,958]" italics="true" pageId="22" pageNumber="231">T</emphasis>
=
<number box="[711,779,932,958]" pageId="22" pageNumber="231" value="12.5" valueMax="14.0" valueMin="11.0">1114</number>
AvS/
<number box="[841,909,932,958]" pageId="22" pageNumber="231" value="13.5" valueMax="14.0" valueMin="13.0">1314</number>
PvS.
</paragraph>
<paragraph blockId="22.[106,1074,1001,1097]" pageId="22" pageNumber="231">
<emphasis box="[106,411,1001,1027]" italics="true" pageId="22" pageNumber="231">Meso- and metathoracic legs.</emphasis>
Lightly ciliated across surface with ciliation density increasing distally. Meso- and metathoracic basitarsus approximately the same length as the remaining tarsal segments combined.
</paragraph>
<paragraph blockId="22.[106,1073,1140,1235]" pageId="22" pageNumber="231">
<emphasis box="[106,216,1140,1166]" italics="true" pageId="22" pageNumber="231">Abdomen.</emphasis>
Supra-anal plate triangular, ciliated across the surface; much narrower than the subgenital plate. Subgenital plate approximately rounded at the terminus. Cerci long, compressed, tapering. Styli short.
</paragraph>
<paragraph blockId="22.[106,1074,1278,1652]" lastBlockId="23.[98,1066,134,299]" lastPageId="23" lastPageNumber="232" pageId="22" pageNumber="231">
<emphasis box="[106,208,1278,1304]" italics="true" pageId="22" pageNumber="231">Genitalia</emphasis>
(
<figureCitation box="[224,292,1278,1305]" captionStart="Fig. 9" captionStartId="13.[98,131,951,973]" captionTargetBox="[147,1015,131,925]" captionTargetId="figure@13.[147,1016,131,925]" captionTargetPageId="13" captionText="Fig. 9. Male genitalic complex of Ilomantis thalassina. Abbreviations: an, anterior apodeme; ap, anterior process; bm, right arm; L 1, L 2, L 4 B, a sub-sclerite of the dorsal sclerotization of the left phallomeric complex; L 4 A, the ventral sclerotization of the left phallomeric complex; loa, lobo membranoso; paa, apical process; pda, processo distale; pia, piastra ventrale; pva, processo ventrale sclerificato; R 1, a sub-sclerite of the right phallomere; R 3, a sub-sclerite of the right phallomere. Red lettering indicates structures; black lettering indicates characters. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269848/files/figure.png" pageId="22" pageNumber="231">
Fig.
<number box="[275,292,1279,1305]" pageId="22" pageNumber="231" value="9.0">9</number>
</figureCitation>
). Dorsal sclerotization of the left phallomeric complex elongate anteriorly, broadening towards the posterior margin; anterior process (ap) moderately narrow, recurved, and lightly sclerotized; L
<number box="[551,565,1348,1374]" pageId="22" pageNumber="231" value="4.0">4</number>
B dextral posterolateral margin rugose; apical process (paa) tumescent, recurved distally with a rounded and moderately sclerotized apical margin. Lobo membranoso (loa) relatively short, apex moderately sclerotized. Ventral sclerotization of the left phallomere complex (i.e., the ventral phallomere) (L
<number box="[131,145,1487,1513]" pageId="22" pageNumber="231" value="4.0">4</number>
A) longer than wide with a slightly undulate and sclerotized antero-sinistral margin; L
<number box="[122,135,1521,1547]" pageId="22" pageNumber="231" value="4.0">4</number>
A posterior region broad, lightly ciliated; processo distale (pda) pouch-shaped, rounded apically. Anterior apodeme (an) rounded anteriorly with a slightly sclerotized sinistral margin; processo ventrale sclerificato (pva) obtusely arched, tumescent, and heavily sclerotized along the posterior and dextral margins; piastra ventrale (pia) heavily sclerotized and linearly shaped with a triangular projection along the anterior margin; pia situated on the anterior margin of a membranous fold that features a generally straight, moderately sclerotized lateral margin. R
<number box="[676,686,204,230]" pageId="23" pageNumber="232" value="1.0">1</number>
posterior region ciliated with a slightly tapered, rounded posterior margin; right arm (bm) moderately narrow, pointed apically.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,671 @@
<document ID-DOI="http://doi.org/10.5281/zenodo.6066869" ID-GBIF-Dataset="09101595-7414-42fa-b53f-5a1e9baadd98" ID-GBIF-Taxon="121067103" ID-Zenodo-Dep="6066869" checkinTime="1467004166608" checkinUser="plazi" docAuthor="Sydney K. Brannoch &amp; Gavin J. Svenson" docDate="2016" docId="4A09B0023436FFA7FF88FE83653BFA24" docLanguage="en" docName="InsectSystematicsEvoluition.47.4.209.pdf" docOrigin="Insect Systematics &amp; Evolution 47" docStyle="DocumentStyle{}" docTitle="Ilomantis ginsburgae Brannoch &amp; Svenson, sp.n." docType="treatment" docVersion="5" lastPageId="26" lastPageNumber="235" masterDocId="B630C87A3421FFBDFFEAFFFB666EFF94" masterDocTitle="Leveraging female genitalic characters for generic and species delimitation in Nilomantis Werner, 1907 and Ilomantis Giglio-Tos, 1915 (Mantodea, Nilomantinae)" masterLastPageNumber="244" masterPageNumber="209" pageId="23" pageNumber="232" updateTime="1644787795423" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Leveraging female genitalic characters for generic and species delimitation in Nilomantis Werner, 1907 and Ilomantis Giglio-Tos, 1915 (Mantodea, Nilomantinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sydney K. Brannoch</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gavin J. Svenson</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Insect Systematics &amp; Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>209</mods:start>
<mods:end>244</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1163/1876312X-47032141</mods:identifier>
<mods:identifier type="GBIF-Dataset">09101595-7414-42fa-b53f-5a1e9baadd98</mods:identifier>
<mods:identifier type="Zenodo-Dep">269839</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6066869" ID-GBIF-Taxon="121067103" ID-Zenodo-Dep="6066869" LSID="urn:lsid:plazi:treatment:4A09B0023436FFA7FF88FE83653BFA24" httpUri="http://treatment.plazi.org/id/4A09B0023436FFA7FF88FE83653BFA24" lastPageId="26" lastPageNumber="235" pageId="23" pageNumber="232">
<subSubSection box="[98,681,376,403]" pageId="23" pageNumber="232" type="nomenclature">
<paragraph blockId="23.[98,681,376,403]" box="[98,681,376,403]" pageId="23" pageNumber="232">
<heading bold="true" box="[98,681,376,403]" fontSize="11" level="1" pageId="23" pageNumber="232" reason="0">
<emphasis bold="true" box="[98,681,376,403]" pageId="23" pageNumber="232">
<taxonomicName authority="Brannoch &amp; Svenson" authorityName="Brannoch &amp; Svenson" box="[98,611,376,403]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="23" pageNumber="232" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis bold="true" box="[98,347,376,403]" italics="true" pageId="23" pageNumber="232">Ilomantis ginsburgae</emphasis>
Brannoch &amp; Svenson
</taxonomicName>
,
<taxonomicNameLabel box="[623,681,377,403]" pageId="23" pageNumber="232" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="232" type="materials_examined">
<paragraph blockId="23.[98,222,430,455]" box="[98,222,430,455]" pageId="23" pageNumber="232">
<heading box="[98,222,430,455]" fontSize="11" level="2" pageId="23" pageNumber="232" reason="1">
<emphasis box="[98,222,430,455]" italics="true" pageId="23" pageNumber="232">
<typeStatus box="[98,147,430,455]" pageId="23" pageNumber="232">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph blockId="23.[98,1066,481,646]" box="[98,1066,481,508]" pageId="23" pageNumber="232">
<typeStatus box="[98,205,481,507]" pageId="23" pageNumber="232">Holotype</typeStatus>
♀. Ihosy,
<date box="[319,456,481,507]" pageId="23" pageNumber="232" value="1967-02-14">
<number box="[319,346,481,507]" pageId="23" pageNumber="232" value="14.0">14</number>
II
<number box="[398,456,481,507]" pageId="23" pageNumber="232" value="1967.0">1967</number>
</date>
(Muséum national dHistoire naturelle, Paris,
<collectingCountry box="[974,1055,482,508]" name="France" pageId="23" pageNumber="232">France</collectingCountry>
).
</paragraph>
<paragraph blockId="23.[98,1066,481,646]" pageId="23" pageNumber="232">
Allotype ♂. Bekily, Rég. Sud de lIle, A. Seyrig (Muséum national dHistoire naturelle, Paris,
<collectingCountry box="[166,247,551,577]" name="France" pageId="23" pageNumber="232">France</collectingCountry>
).
<typeStatus box="[265,362,551,577]" pageId="23" pageNumber="232">Paratype</typeStatus>
♀. Ihosy,
<date box="[479,616,550,576]" pageId="23" pageNumber="232" value="1967-02-14">
<number box="[479,506,550,576]" pageId="23" pageNumber="232" value="14.0">14</number>
II
<number box="[558,616,550,576]" pageId="23" pageNumber="232" value="1967.0">1967</number>
</date>
(Muséum national dHistoire naturelle, Paris,
<collectingCountry box="[171,251,586,612]" name="France" pageId="23" pageNumber="232">France</collectingCountry>
);
<typeStatus box="[275,372,586,612]" pageId="23" pageNumber="232">Paratype</typeStatus>
♂. Bekily, Rég. Sud de lIle, A. Seyrig (Muséum national dHistoire naturelle, Paris,
<collectingCountry box="[400,481,620,646]" name="France" pageId="23" pageNumber="232">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="232" type="etymology">
<paragraph blockId="23.[98,209,689,715]" box="[98,209,689,715]" pageId="23" pageNumber="232">
<heading box="[98,209,689,715]" fontSize="11" level="2" pageId="23" pageNumber="232" reason="1">
<emphasis box="[98,209,689,715]" italics="true" pageId="23" pageNumber="232">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="23.[98,1066,741,906]" pageId="23" pageNumber="232">
A noun in the genitive case, we named the species in honor of Ruth Joan Bader Ginsburg, Associate Justice of the Supreme Court of the
<collectingCountry box="[742,897,776,802]" name="United States of America" pageId="23" pageNumber="232">United States</collectingCountry>
, for her relentless fight for gender equality, as well as for her sartorial appreciation of the jabot, which is reminiscent of the postcervical plate of
<emphasis box="[562,671,845,871]" italics="true" pageId="23" pageNumber="232">
<taxonomicName box="[562,667,845,871]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="23" pageNumber="232" phylum="Arthropoda" rank="genus">Ilomantis</taxonomicName>
,
</emphasis>
a diagnostic character that embodies this judicial accessory.
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="233" pageId="23" pageNumber="232" type="materials_examined">
<paragraph blockId="23.[98,319,949,975]" box="[98,319,949,975]" pageId="23" pageNumber="232">
<heading box="[98,319,949,975]" fontSize="11" level="2" pageId="23" pageNumber="232" reason="1">
<emphasis box="[98,319,949,975]" italics="true" pageId="23" pageNumber="232">Examined specimens</emphasis>
</heading>
</paragraph>
<paragraph blockId="23.[98,1067,1001,1652]" lastBlockId="24.[106,1073,134,196]" lastPageId="24" lastPageNumber="233" pageId="23" pageNumber="232">
The list consists of
<number box="[328,343,1001,1027]" pageId="23" pageNumber="232" value="4.0">4</number>
<typeStatus box="[354,412,1002,1028]" pageId="23" pageNumber="232">types</typeStatus>
and
<number box="[477,506,1001,1027]" pageId="23" pageNumber="232" value="13.0">13</number>
additional specimens.
<collectingCountry box="[776,913,1001,1027]" name="Madagascar" pageId="23" pageNumber="232">Madagascar</collectingCountry>
:
<emphasis box="[928,1011,1001,1027]" italics="true" pageId="23" pageNumber="232">Androy,</emphasis>
<specimenCount box="[1022,1058,1001,1028]" pageId="23" pageNumber="232" type="generic">
<number box="[1022,1037,1001,1027]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
, (Pays Androy (Nord)),
<collectingCountry box="[363,496,1036,1062]" name="Madagascar" pageId="23" pageNumber="232">Madagascar</collectingCountry>
Sud,
<number box="[564,622,1036,1062]" pageId="23" pageNumber="232" value="1901.0">1901</number>
/
<number box="[631,691,1036,1062]" pageId="23" pageNumber="232" value="1926.0">1926</number>
(?)(this specimen has conflicting labels: one label indicates the specimen was collected in
<number box="[737,798,1071,1097]" pageId="23" pageNumber="232" value="1901.0">1901</number>
, the other indicates the collection year was
<number box="[321,383,1105,1132]" pageId="23" pageNumber="232" value="1926.0">1926</number>
), Alluaud, Genitalia No. 0 0 0
<number box="[720,748,1105,1131]" pageId="23" pageNumber="232" value="14.0">14</number>
S. K. Brannoch (
<collectionCode box="[952,1053,1105,1131]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[98,222,1140,1166]" italics="true" pageId="23" pageNumber="232">Ankiliarivo</emphasis>
,
<specimenCount box="[239,275,1140,1167]" pageId="23" pageNumber="232" type="generic">
<number box="[239,254,1140,1166]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
,
<date box="[292,442,1140,1166]" pageId="23" pageNumber="232" value="1962-03-26">
<number box="[292,322,1140,1166]" pageId="23" pageNumber="232" value="26.0">26</number>
III
<number box="[383,442,1140,1166]" pageId="23" pageNumber="232" value="1962.0">1962</number>
</date>
, D. Wintrebert, Genitalia No. 0 0 0
<number box="[849,878,1140,1166]" pageId="23" pageNumber="232" value="12.0">12</number>
S. K. Brannoch (
<collectionCode box="[107,208,1175,1201]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[227,298,1174,1200]" italics="true" pageId="23" pageNumber="232">Bekily,</emphasis>
<number box="[305,320,1175,1201]" pageId="23" pageNumber="232" value="1.0">1</number>
♀, Reg. Sud.
<collectingCountry box="[469,503,1175,1201]" name="South Georgia and the South Sandwich Islands" pageId="23" pageNumber="232">De</collectingCountry>
LIle, A. Seyrig (
<collectionCode box="[686,787,1175,1201]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<specimenCount box="[806,843,1175,1201]" pageId="23" pageNumber="232" type="generic">
<number box="[806,821,1175,1201]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
, Reg. Sud.
<collectingCountry box="[970,1004,1175,1201]" name="South Georgia and the South Sandwich Islands" pageId="23" pageNumber="232">De</collectingCountry>
LIle, A. Seyrig (
<collectionCode box="[216,317,1209,1235]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<number box="[337,352,1209,1235]" pageId="23" pageNumber="232" value="1.0">1</number>
♀, Reg. Sud.
<collectingCountry box="[504,538,1210,1236]" name="South Georgia and the South Sandwich Islands" pageId="23" pageNumber="232">De</collectingCountry>
LIle,
<date box="[608,674,1209,1235]" bridgedPair="-" pageId="23" pageNumber="232" value="1934-02">
II
<number box="[644,674,1209,1235]" pageId="23" pageNumber="232" value="34.0">34</number>
</date>
, A. Seyrig, Genitalia No. 0 0 0
<number box="[1008,1037,1209,1235]" pageId="23" pageNumber="232" value="10.0">10</number>
S. K. Brannoch (
<collectionCode box="[262,363,1244,1270]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<specimenCount box="[385,422,1244,1271]" pageId="23" pageNumber="232" type="generic">
<number box="[385,400,1244,1270]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
, Reg. Sud.
<collectingCountry box="[557,591,1244,1270]" name="South Georgia and the South Sandwich Islands" pageId="23" pageNumber="232">De</collectingCountry>
LIle,
<date box="[664,741,1244,1270]" bridgedPair="-" pageId="23" pageNumber="232" value="1932-03">
III
<number box="[711,741,1244,1270]" pageId="23" pageNumber="232" value="32.0">32</number>
</date>
, A. Seyrig, Genitalia R. Roy No.
<number box="[154,213,1279,1305]" pageId="23" pageNumber="232" value="4071.0">4071</number>
(
<collectionCode box="[234,335,1279,1305]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[361,440,1278,1304]" italics="true" pageId="23" pageNumber="232">Betioky</emphasis>
(Sud),
<number box="[535,550,1279,1305]" pageId="23" pageNumber="232" value="1.0">1</number>
♀,
<collectingCountry box="[592,725,1279,1305]" name="Madagascar" pageId="23" pageNumber="232">Madagascar</collectingCountry>
Sud-Ouest,
<date box="[878,1019,1279,1305]" pageId="23" pageNumber="232" value="1968-05-26">
<number box="[878,908,1279,1305]" pageId="23" pageNumber="232" value="26.0">26</number>
V
<number box="[961,1019,1279,1305]" pageId="23" pageNumber="232" value="1968.0">1968</number>
</date>
, D. Wintrebert Rec. (
<collectionCode box="[298,399,1313,1339]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[420,623,1313,1339]" italics="true" pageId="23" pageNumber="232">Clairière Erinkady</emphasis>
(?),
<number box="[674,689,1313,1339]" pageId="23" pageNumber="232" value="1.0">1</number>
♀,
<date box="[726,832,1313,1339]" pageId="23" pageNumber="232" value="1963-03-07">
<number box="[726,740,1313,1339]" pageId="23" pageNumber="232" value="7.0">7</number>
III
<number box="[803,832,1313,1339]" pageId="23" pageNumber="232" value="63.0">63</number>
</date>
, D. Wintrebert Reç. (
<collectionCode box="[107,208,1348,1374]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[233,394,1348,1374]" italics="true" pageId="23" pageNumber="232">Fort-Dauphin,</emphasis>
<number box="[406,421,1348,1374]" pageId="23" pageNumber="232" value="1.0">1</number>
♀,
<date box="[462,558,1348,1374]" bridgedPair="-" pageId="23" pageNumber="232" value="1937-05">
V
<number box="[500,558,1348,1374]" pageId="23" pageNumber="232" value="1937.0">1937</number>
</date>
, A. Seyrig (
<collectionCode box="[702,803,1348,1374]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[829,883,1348,1374]" italics="true" pageId="23" pageNumber="232">Ifaty,</emphasis>
<specimenCount box="[895,932,1348,1375]" pageId="23" pageNumber="232" type="generic">
<number box="[895,910,1348,1374]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
,
<quantity box="[952,1030,1348,1374]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="232" unit="km" value="20.0">20 km</quantity>
N Tulear
<quantity box="[177,241,1383,1409]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="232" unit="m" value="30.0">30 m</quantity>
,
<date box="[254,442,1383,1409]" pageId="23" pageNumber="232" value="2003-12-10" valueMax="2003-12-27" valueMin="2003-12-10">
<number box="[254,328,1383,1409]" pageId="23" pageNumber="232" value="18.5" valueMax="27.0" valueMin="10.0">1027</number>
.XII.
<number box="[380,442,1383,1409]" pageId="23" pageNumber="232" value="2003.0">2003</number>
</date>
, S. Murzin &amp; A. Shamaev leg, Genitalia No. 0 0 0
<number box="[1008,1036,1383,1409]" pageId="23" pageNumber="232" value="13.0">13</number>
S. K. Brannoch (
<collectionCode box="[258,359,1417,1443]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<specimenCount box="[379,416,1417,1444]" pageId="23" pageNumber="232" type="generic">
<number box="[379,394,1417,1443]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
,
<quantity box="[430,503,1417,1443]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="232" unit="km" value="20.0">20 km</quantity>
N Tulear
<quantity box="[617,679,1417,1444]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="232" unit="m" value="30.0">30 m</quantity>
,
<date box="[690,878,1417,1443]" pageId="23" pageNumber="232" value="2003-12-10" valueMax="2003-12-27" valueMin="2003-12-10">
<number box="[690,765,1417,1443]" pageId="23" pageNumber="232" value="18.5" valueMax="27.0" valueMin="10.0">1027</number>
.XII.
<number box="[816,878,1417,1443]" pageId="23" pageNumber="232" value="2003.0">2003</number>
</date>
, S. Murzin &amp; A. Shamaev leg, Genitalia No. 0 0 0
<number box="[458,486,1452,1479]" pageId="23" pageNumber="232" value="15.0">15</number>
S. K. Brannoch (
<collectionCode box="[682,783,1452,1478]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[802,856,1452,1478]" italics="true" pageId="23" pageNumber="232">Ihosy</emphasis>
,
<number box="[870,885,1452,1478]" pageId="23" pageNumber="232" value="1.0">1</number>
♀,
<date box="[921,1062,1452,1478]" pageId="23" pageNumber="232" value="1967-02-14">
<number box="[921,949,1452,1478]" pageId="23" pageNumber="232" value="14.0">14</number>
II
<number box="[1002,1062,1452,1478]" pageId="23" pageNumber="232" value="1967.0">1967</number>
</date>
, Genitalia No. 0 0 0
<number box="[315,343,1487,1513]" pageId="23" pageNumber="232" value="18.0">18</number>
S. K. Brannoch (
<collectionCode box="[560,661,1487,1513]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<number box="[687,702,1487,1513]" pageId="23" pageNumber="232" value="1.0">1</number>
♀,
<date box="[743,880,1487,1513]" pageId="23" pageNumber="232" value="1967-02-14">
<number box="[743,770,1487,1513]" pageId="23" pageNumber="232" value="14.0">14</number>
II
<number box="[822,880,1487,1513]" pageId="23" pageNumber="232" value="1967.0">1967</number>
</date>
(
<collectionCode box="[901,1002,1487,1513]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis italics="true" pageId="23" pageNumber="232">Lac Tsimanampetsotsa,</emphasis>
<specimenCount box="[309,345,1521,1548]" pageId="23" pageNumber="232" type="generic">
<number box="[309,324,1521,1547]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
,
<collectingCountry box="[361,494,1521,1547]" name="Madagascar" pageId="23" pageNumber="232">Madagascar</collectingCountry>
Sud, Rés. Nat. Int. 10, 7/
<date box="[790,931,1521,1547]" pageId="23" pageNumber="232" value="1969-02-10">
<number box="[790,818,1521,1547]" pageId="23" pageNumber="232" value="10.0">10</number>
II
<number box="[871,931,1521,1547]" pageId="23" pageNumber="232" value="1969.0">1969</number>
</date>
, P. Viette et P. Griveaud, Genitalia No. 0 0 0
<number box="[457,484,1556,1582]" pageId="23" pageNumber="232" value="11.0">11</number>
S. K. Brannoch (
<collectionCode box="[688,789,1556,1582]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[810,1006,1556,1582]" italics="true" pageId="23" pageNumber="232">Plateau Mahafaly</emphasis>
,
<specimenCount box="[1022,1058,1556,1583]" pageId="23" pageNumber="232" type="generic">
<number box="[1022,1037,1556,1582]" pageId="23" pageNumber="232" value="1.0">1</number>
</specimenCount>
,
<date box="[98,184,1591,1617]" bridgedPair="-" pageId="23" pageNumber="232" value="1966-01">
I
<number box="[124,184,1591,1617]" pageId="23" pageNumber="232" value="1966.0">1966</number>
</date>
, Paul Grivaud,
<collectionCode box="[374,456,1591,1617]" pageId="23" pageNumber="232">GSMC</collectionCode>
0001444 (
<collectionCode box="[581,682,1591,1617]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="232">MNHN</collectionCode>
);
<emphasis box="[707,865,1591,1617]" italics="true" pageId="23" pageNumber="232">
<taxonomicName box="[707,808,1591,1617]" class="Insecta" family="Tineidae" genus="Ranohira" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="232" phylum="Arthropoda" rank="genus">Ranohira</taxonomicName>
(W)
</emphasis>
,
<number box="[884,899,1591,1617]" pageId="23" pageNumber="232" value="1.0">1</number>
<collectingCountry box="[933,1066,1591,1617]" name="Madagascar" pageId="23" pageNumber="232">Madagascar</collectingCountry>
Sud,
<date box="[156,266,1625,1651]" pageId="23" pageNumber="232" value="1963-06-12">
<number box="[156,185,1625,1651]" pageId="23" pageNumber="232" value="12.0">12</number>
-VI-
<number box="[236,266,1625,1651]" pageId="23" pageNumber="232" value="63.0">63</number>
</date>
, Sité à Tapia, Q. Wintrebert Reç., Genitalia No. 0 0 0
<number box="[858,886,1625,1651]" pageId="23" pageNumber="232" value="16.0">16</number>
S. K. Brannoch (
<collectionCode box="[114,215,135,161]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="233">MNHN</collectionCode>
);
<emphasis box="[237,307,134,160]" italics="true" pageId="24" pageNumber="233">Tuléar</emphasis>
(
<quantity box="[324,400,134,161]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="233" unit="km" value="20.0">20 km</quantity>
sud de Tuléar),
<number box="[587,602,135,161]" pageId="24" pageNumber="233" value="1.0">1</number>
<collectingCountry box="[633,766,134,160]" name="Madagascar" pageId="24" pageNumber="233">Madagascar</collectingCountry>
S. S. Ouest,
<date box="[919,1069,135,161]" pageId="24" pageNumber="233" value="1973-03-30">
<number box="[919,949,135,161]" pageId="24" pageNumber="233" value="30.0">30</number>
III
<number box="[1011,1069,135,161]" pageId="24" pageNumber="233" value="1973.0">1973</number>
</date>
, Descamps &amp; Wintrebert Reç., Genitalia No. 0 0 0
<number box="[663,691,169,195]" pageId="24" pageNumber="233" value="17.0">17</number>
S. K. Brannoch (
<collectionCode box="[889,990,169,195]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="233">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="233" type="biology_ecology">
<paragraph blockId="24.[106,270,244,270]" box="[106,270,244,270]" pageId="24" pageNumber="233">
<heading box="[106,270,244,270]" fontSize="11" level="2" pageId="24" pageNumber="233" reason="1">
<emphasis box="[106,270,244,270]" italics="true" pageId="24" pageNumber="233">Natural history</emphasis>
</heading>
</paragraph>
<paragraph blockId="24.[106,1074,296,357]" pageId="24" pageNumber="233">
Specimens examined were collected January through June in Southern
<collectingCountry box="[935,1070,296,322]" name="Madagascar" pageId="24" pageNumber="233">Madagascar</collectingCountry>
. No specific behavioral or ecological information is known.
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="233" type="diagnosis">
<paragraph blockId="24.[106,210,406,431]" box="[106,210,406,431]" pageId="24" pageNumber="233">
<heading box="[106,210,406,431]" fontSize="11" level="2" pageId="24" pageNumber="233" reason="1">
<emphasis box="[106,210,406,431]" italics="true" pageId="24" pageNumber="233">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="24.[106,1074,457,726]" pageId="24" pageNumber="233">
Pronotum with a medial keel that runs from the mid-prozone to the posterior margin of the metazone. Postcervical plate and exposed thoracic membrane of the cervix transverse. Females (ca.
<quantity box="[317,467,526,553]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" metricValueMax="2.06" metricValueMin="1.5" pageId="24" pageNumber="233" unit="mm" value="17.8" valueMax="20.6" valueMin="15.0">1520.6 mm</quantity>
) only marginally longer than males (ca.
<quantity box="[916,1066,527,553]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" metricValueMax="2.04" metricValueMin="1.4" pageId="24" pageNumber="233" unit="mm" value="17.2" valueMax="20.4" valueMin="14.0">1420.4 mm</quantity>
) on average. Female genitalia: GA with reduced MO; GL with convergent AC. Male genitalia: pda small, adjacent to a moderately pronounced dextral lobe on the posterodextral margin; pva slightly arched along the anterior margin with a curved posterior margin. Distributed within the Northern region of the
<collectingCountry box="[783,1070,665,691]" name="Madagascar" pageId="24" pageNumber="233">Republic of Madagascar</collectingCountry>
.
<emphasis box="[106,122,701,726]" italics="true" pageId="24" pageNumber="233">F</emphasis>
=
<number box="[137,149,700,726]" pageId="24" pageNumber="233" value="3.0">3</number>
DS/
<number box="[194,221,700,726]" pageId="24" pageNumber="233" value="12.0">12</number>
AvS/
<number box="[281,295,700,726]" pageId="24" pageNumber="233" value="4.0">4</number>
PvS;
<emphasis box="[353,372,701,726]" italics="true" pageId="24" pageNumber="233">T</emphasis>
=
<number box="[388,455,700,726]" pageId="24" pageNumber="233" value="10.5" valueMax="11.0" valueMin="10.0">1011</number>
AvS/
<number box="[516,572,700,726]" pageId="24" pageNumber="233" value="11.0" valueMax="13.0" valueMin="9.0">913</number>
PvS.
</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="235" pageId="24" pageNumber="233" type="description">
<paragraph blockId="24.[106,230,775,800]" box="[106,230,775,800]" pageId="24" pageNumber="233">
<heading box="[106,230,775,800]" fontSize="11" level="2" pageId="24" pageNumber="233" reason="1">
<emphasis box="[106,230,775,800]" italics="true" pageId="24" pageNumber="233">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="24.[106,1074,826,1477]" pageId="24" pageNumber="233">
<emphasis box="[106,191,826,852]" italics="true" pageId="24" pageNumber="233">Female.</emphasis>
<typeStatus box="[202,310,826,852]" pageId="24" pageNumber="233">Holotype</typeStatus>
(
<figureCitation box="[329,400,826,853]" captionStart="Fig. 2" captionStartId="3.[98,131,1339,1361]" captionTargetBox="[147,1016,131,1312]" captionTargetId="figure@3.[147,1016,131,1313]" captionTargetPageId="3" captionText="Fig. 2. Ilomantis ginsburgae sp. n. (A) Holotype female, dorsal habitus; (B) holotype female, ventral habitus; (C) allotype male, dorsal habitus; (D) allotype male, ventral habitus. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269841/files/figure.png" pageId="24" pageNumber="233">
Fig.
<number box="[385,400,827,853]" pageId="24" pageNumber="233" value="2.0">2</number>
</figureCitation>
A,B). While male mantodean specimens have historically been assigned as
<typeStatus box="[303,416,861,887]" pageId="24" pageNumber="233">holotypes</typeStatus>
, we have assigned a female specimen as the
<typeStatus box="[931,1031,861,887]" pageId="24" pageNumber="233">holotype</typeStatus>
for this new species. In so doing, we hope to underscore the point that when detailed character descriptions of both external and genitalic morphology are presented, a female
<typeStatus box="[190,290,965,991]" pageId="24" pageNumber="233">holotype</typeStatus>
can be as informative as a male
<typeStatus box="[673,777,965,991]" pageId="24" pageNumber="233">holotype</typeStatus>
. Traditionally, male
<typeStatus pageId="24" pageNumber="233">holotypes</typeStatus>
were preferred because the male genitalia could be used to perform species-level determinations, while making species-level determinations in females was less certain due to ambiguous boundaries in external morphological characters. However, in this study we have demonstrated that female genitalia can be informative for species identifications, eliminating a potential problem with choosing females as the
<typeStatus box="[933,1033,1138,1164]" pageId="24" pageNumber="233">holotype</typeStatus>
for new species.
</paragraph>
<paragraph blockId="24.[106,1074,826,1477]" pageId="24" pageNumber="233">
Body length
<number box="[282,434,1208,1234]" pageId="24" pageNumber="233" value="20.799999999999997" valueMax="21.61" valueMin="19.99">19.9921.61</number>
; forewing length
<number box="[632,784,1208,1235]" pageId="24" pageNumber="233" value="14.98" valueMax="15.48" valueMin="14.48">14.4815.48</number>
; hindwing length
<number pageId="24" pageNumber="233" value="15.295" valueMax="15.95" valueMin="14.64">14.64 15.95</number>
; pronotum maximum length
<number box="[521,643,1243,1269]" pageId="24" pageNumber="233" value="4.0600000000000005" valueMax="4.29" valueMin="3.83">3.834.29</number>
; prozone length
<number box="[837,944,1242,1269]" pageId="24" pageNumber="233" value="1.675" valueMax="1.8" valueMin="1.55">1.551.8</number>
; pronotum width
<number box="[185,307,1277,1304]" pageId="24" pageNumber="233" value="1.73" valueMax="1.85" valueMin="1.61">1.611.85</number>
; pronotum minimum width
<number box="[662,784,1277,1303]" pageId="24" pageNumber="233" value="1.3900000000000001" valueMax="1.44" valueMin="1.34">1.341.44</number>
; head width
<number box="[947,1069,1277,1304]" pageId="24" pageNumber="233" value="3.985" valueMax="4.22" valueMin="3.75">3.754.22</number>
; head vertex to clypeus
<number box="[361,430,1312,1338]" pageId="24" pageNumber="233" value="1.1" valueMax="1.2" valueMin="1.0">11.2</number>
; frons width
<number box="[578,700,1312,1338]" pageId="24" pageNumber="233" value="1.52" valueMax="1.63" valueMin="1.41">1.411.63</number>
; frons height
<number box="[853,975,1312,1338]" pageId="24" pageNumber="233" value="0.42" valueMax="0.47" valueMin="0.37">0.370.47</number>
; prothoracic femur length
<number box="[329,451,1346,1373]" pageId="24" pageNumber="233" value="5.075" valueMax="5.23" valueMin="4.92">4.925.23</number>
; mesothoracic femur length
<number box="[786,909,1346,1372]" pageId="24" pageNumber="233" value="4.595" valueMax="4.93" valueMin="4.26">4.264.93</number>
; mesothoracic tibia length
<number box="[251,372,1381,1408]" pageId="24" pageNumber="233" value="3.2350000000000003" valueMax="3.45" valueMin="3.02">3.023.45</number>
; mesothoracic tarsus length
<number box="[709,831,1381,1408]" pageId="24" pageNumber="233" value="2.37" valueMax="2.58" valueMin="2.16">2.162.58</number>
; metathoracic femur length
<number box="[189,310,1416,1443]" pageId="24" pageNumber="233" value="5.324999999999999" valueMax="5.63" valueMin="5.02">5.025.63</number>
; metathoracic tibia length
<number box="[629,751,1416,1443]" pageId="24" pageNumber="233" value="5.535" valueMax="5.85" valueMin="5.22">5.225.85</number>
; metathoracic tarsus length
<number box="[106,229,1451,1477]" pageId="24" pageNumber="233" value="3.425" valueMax="3.71" valueMin="3.14">3.143.71</number>
.
</paragraph>
<paragraph blockId="24.[106,1074,1520,1650]" pageId="24" pageNumber="233">
<emphasis box="[106,165,1520,1546]" italics="true" pageId="24" pageNumber="233">Head</emphasis>
(
<figureCitation box="[179,247,1520,1546]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="24" pageNumber="233">
Fig.
<number box="[230,247,1520,1546]" pageId="24" pageNumber="233" value="4.0">4</number>
</figureCitation>
C). Juxtaocular bulges present, slight. Vertex elevation yellow. Region posterior to the lateral ocelli slightly concave. Ocelli relatively small; lateral ocelli marginally larger than the median ocellus; area containing the ocelli slightly convex. Antennae extend to at least mid-body.
</paragraph>
<paragraph blockId="25.[98,1066,134,265]" pageId="25" pageNumber="234">
<emphasis box="[98,182,134,160]" italics="true" pageId="25" pageNumber="234">Thorax.</emphasis>
Pronotal medial keel reaching mid-prozone (
<figureCitation box="[695,777,134,161]" captionStart="Fig. 11" captionStartId="16.[106,139,1517,1539]" captionTargetBox="[243,935,131,1491]" captionTargetId="figure@16.[243,937,131,1492]" captionTargetPageId="16" captionText="Fig. 11. Illustrations of dorsal perspective of the Ilomantis pronotum (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. Ilomantis thalassina exhibits a medial keel that fully traverses the pronotum (A and B); Ilomantis ginsburgae sp. n. exhibits a medial keel that originates in the mid-prozone and concludes at the posterior pronotal margin (C and D). MK, medial keel." httpUri="https://zenodo.org/record/269850/files/figure.png" pageId="25" pageNumber="234">
Fig.
<number box="[747,777,135,161]" pageId="25" pageNumber="234" value="11.0">11</number>
</figureCitation>
C). Postcervical plate and exposed thoracic membrane of the cervix transverse with an approximately hexagonal shape (
<figureCitation box="[180,265,204,230]" captionStart="Fig. 13" captionStartId="18.[106,139,1269,1291]" captionTargetBox="[155,1024,131,1244]" captionTargetId="figure@18.[155,1024,131,1244]" captionTargetPageId="18" captionText="Fig. 13. Illustrations of ventral perspective of the Ilomantis cervix (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. The postcervical plate and exposed thoracic membrane are contained within the boundaries of the intercervical sclerites and the episternum, with either a broad (Ilomantis thalassina, A and B) or narrow (Ilomantis ginsburgae sp. n., C and D) hexagonal shape. Abbreviations: eps, episternum; etm, exposed thoracic membrane; ics, intercervical sclerite; lcs, lateral cervical sclerite; pcpl, postcervical plate; tss, T-shaped sclerite." httpUri="https://zenodo.org/record/269852/files/figure.png" pageId="25" pageNumber="234">
Fig.
<number box="[235,265,204,230]" pageId="25" pageNumber="234" value="13.0">13</number>
</figureCitation>
C). Wings opaque with margins sparsely ciliated; anterior margin of forewing costal area with light green cells.
</paragraph>
<paragraph blockId="25.[98,1066,308,369]" pageId="25" pageNumber="234">
<emphasis box="[98,275,308,334]" italics="true" pageId="25" pageNumber="234">Prothoracic legs.</emphasis>
Femur maximum width ca.
<number box="[617,654,308,334]" pageId="25" pageNumber="234" value="1.3">1.3</number>
times the value of the male femur maximum width.
<emphasis box="[303,319,343,368]" italics="true" pageId="25" pageNumber="234">F</emphasis>
=
<number box="[334,346,343,369]" pageId="25" pageNumber="234" value="3.0">3</number>
DS/
<number box="[391,418,343,369]" pageId="25" pageNumber="234" value="12.0">12</number>
AvS/
<number box="[478,492,343,369]" pageId="25" pageNumber="234" value="4.0">4</number>
PvS;
<emphasis box="[550,569,343,368]" italics="true" pageId="25" pageNumber="234">T</emphasis>
=
<number box="[584,608,343,369]" pageId="25" pageNumber="234" value="11.0">11</number>
AvS/
<number box="[670,739,343,369]" pageId="25" pageNumber="234" value="11.0" valueMax="12.0" valueMin="10.0">1012</number>
PvS.
</paragraph>
<paragraph blockId="25.[98,1066,412,542]" pageId="25" pageNumber="234">
<emphasis box="[98,413,412,438]" italics="true" pageId="25" pageNumber="234">Meso- and metathoracic legs.</emphasis>
Ciliated across surface. Mesothoracic basitarsus approximately
<number box="[186,224,446,473]" pageId="25" pageNumber="234" value="1.5">1.5</number>
times the length of the remaining tarsal segments combined; metathoracic basitarsus approximately the same length as the remaining tarsal segments combined.
</paragraph>
<paragraph blockId="25.[98,1066,585,646]" pageId="25" pageNumber="234">
<emphasis box="[98,209,585,611]" italics="true" pageId="25" pageNumber="234">Abdomen.</emphasis>
Supra-anal plate triangular, ciliated across surface, narrower than the subgenital plate. Cerci filiform, tapering.
</paragraph>
<paragraph blockId="25.[98,1066,689,1339]" pageId="25" pageNumber="234">
<emphasis box="[98,202,689,715]" italics="true" pageId="25" pageNumber="234">Genitalia</emphasis>
(
<figureCitation box="[220,292,689,715]" captionStart="Fig. 3" captionStartId="7.[98,131,1511,1533]" captionTargetBox="[147,1016,360,1484]" captionTargetId="figure@7.[147,1016,360,1485]" captionTargetPageId="7" captionText="Fig. 3. Female genitalic complex of Ilomantis shown in the lateral perspective. (A) Ilomantis thalassina; (B) Ilomantis ginsburgae sp. n. Abbreviations: AC, apical cleft; AL, apical lobe; GA, gonapophysis VIII; GL, gonoplac; MO, medial outgrowth; RTD, rectangular dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269842/files/figure.png" pageId="25" pageNumber="234">
Figs
<number box="[278,292,689,715]" pageId="25" pageNumber="234" value="3.0">3</number>
</figureCitation>
B,
<number box="[329,344,689,716]" pageId="25" pageNumber="234" value="5.0">5</number>
C,D). Gonapophysis VIII (GA) slender, variably ciliated, and visibly curved from both the ventral (VH) and lateral (LH) perspectives; GA lightly surfaced with fossettes and sculpturing. From VH, GA angles inward; apical lobe (AL) ovoid, heavily ciliated. From LH, AL ranges from ovoid, to blunt, to thin and arched. AL has a membrane that projects from the internal dorsal margin of the structure, forming a shallow pocket as the membrane merges with the dorsal margin of GA, lending AL the appearance of being bilobed or mitten-shaped. From LH, GA is obtusely curved ventrad; GA ventral margin features a reduced medial outgrowth (MO), which projects posterior to the crest of the curve. GA dorsal margin is invaginated anterior to the curve towards the ventral habitus. Gonapophysis IX (GP) is relatively short, terminating in a weakly rounded apex; GP ventral margin features a subulate overhang, the medial tine (MT). Gonoplac (GL) is shorter than GA, broad, variably ciliated, and arched ventrad; from LH, GL base has a rectangular dilation on the anteroventral margin. GL narrows toward the apex; from LH, GL apex slants ventrad. GL features a significantly reduced apical cleft (AC), with the lobe of the dorsal margin appearing to converge with the ventral lobe, resulting in an approximately subulate apex. From LH, GL possesses a slight sclerotization that traverses the length of the structure, terminating just before AC. GL ventral and dorsal margins are relatively smooth.
</paragraph>
<paragraph blockId="25.[98,1066,1382,1652]" pageId="25" pageNumber="234">
<emphasis box="[98,162,1382,1408]" italics="true" pageId="25" pageNumber="234">Male.</emphasis>
Allotype (
<figureCitation box="[278,344,1382,1409]" captionStart="Fig. 2" captionStartId="3.[98,131,1339,1361]" captionTargetBox="[147,1016,131,1312]" captionTargetId="figure@3.[147,1016,131,1313]" captionTargetPageId="3" captionText="Fig. 2. Ilomantis ginsburgae sp. n. (A) Holotype female, dorsal habitus; (B) holotype female, ventral habitus; (C) allotype male, dorsal habitus; (D) allotype male, ventral habitus. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269841/files/figure.png" pageId="25" pageNumber="234">
Fig.
<number box="[328,344,1383,1409]" pageId="25" pageNumber="234" value="2.0">2</number>
</figureCitation>
C, D). Body length
<number box="[568,717,1383,1409]" pageId="25" pageNumber="234" value="19.89" valueMax="20.44" valueMin="19.34">19.3420.44</number>
; forewing length
<number box="[912,1061,1382,1409]" pageId="25" pageNumber="234" value="14.805" valueMax="15.37" valueMin="14.24">14.2415.37</number>
; hindwing length
<number box="[289,439,1417,1444]" pageId="25" pageNumber="234" value="15.515" valueMax="16.18" valueMin="14.85">14.8516.18</number>
; pronotum maximum length
<number box="[768,888,1417,1443]" pageId="25" pageNumber="234" value="3.65" valueMax="3.84" valueMin="3.46">3.463.84</number>
; prozone length
<number box="[98,219,1452,1478]" pageId="25" pageNumber="234" value="1.405" valueMax="1.49" valueMin="1.32">1.321.49</number>
; pronotum width
<number box="[433,554,1452,1479]" pageId="25" pageNumber="234" value="1.4" valueMax="1.45" valueMin="1.35">1.351.45</number>
; pronotum minimum width
<number box="[893,998,1452,1478]" pageId="25" pageNumber="234" value="1.21" valueMax="1.3" valueMin="1.12">1.121.3</number>
; head width
<number box="[175,280,1486,1512]" pageId="25" pageNumber="234" value="3.52" valueMax="3.6" valueMin="3.44">3.443.6</number>
; head vertex to clypeus
<number box="[564,647,1487,1513]" pageId="25" pageNumber="234" value="1.0899999999999999" valueMax="1.18" valueMin="1.0">11.18</number>
; frons width
<number box="[806,911,1486,1512]" pageId="25" pageNumber="234" value="1.33" valueMax="1.36" valueMin="1.3">1.31.36</number>
; frons height
<number box="[98,219,1521,1547]" pageId="25" pageNumber="234" value="0.37" valueMax="0.4" valueMin="0.34">0.340.40</number>
; prothoracic femur length
<number box="[539,659,1521,1547]" pageId="25" pageNumber="234" value="4.34" valueMax="4.49" valueMin="4.19">4.194.49</number>
; mesothoracic femur length
<number pageId="25" pageNumber="234" value="3.9850000000000003" valueMax="4.19" valueMin="3.78">3.78 4.19</number>
; mesothoracic tibia length
<number box="[480,600,1556,1582]" pageId="25" pageNumber="234" value="2.71" valueMax="2.94" valueMin="2.48">2.482.94</number>
; mesothoracic tarsus length
<number box="[941,1061,1556,1582]" pageId="25" pageNumber="234" value="1.94" valueMax="2.17" valueMin="1.71">1.712.17</number>
; metathoracic femur length
<number box="[407,527,1591,1617]" pageId="25" pageNumber="234" value="4.695" valueMax="4.92" valueMin="4.47">4.474.92</number>
; metathoracic tibia length
<number box="[830,951,1590,1617]" pageId="25" pageNumber="234" value="4.914999999999999" valueMax="5.27" valueMin="4.56">4.565.27</number>
; metathoracic tarsus length
<number box="[309,430,1625,1651]" pageId="25" pageNumber="234" value="3.2" valueMax="3.66" valueMin="2.74">2.743.66</number>
.
</paragraph>
<paragraph blockId="26.[106,1074,134,300]" pageId="26" pageNumber="235">
<emphasis box="[106,165,134,160]" italics="true" pageId="26" pageNumber="235">Head</emphasis>
(
<figureCitation box="[180,246,134,161]" captionStart="Fig. 8" captionStartId="12.[106,139,1140,1162]" captionTargetBox="[155,1023,131,1114]" captionTargetId="figure@12.[155,1024,131,1115]" captionTargetPageId="12" captionText="Fig. 8. Illustrations of anterior perspective of the Ilomantis and Nilomantis head (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male; (E) Nilomantis floweri, female; (F) Nilomantis floweri, male. (B, D, E, F) Ilomantis species feature a cosinusoidal vertical margin, whereas Nilomantis species feature a relatively straight vertical margin. Gray markings on the ocellar tubercle and within the corridors of the frontal sulci represent the presence of reddish-brown markings. LF, lower frons." httpUri="https://zenodo.org/record/269847/files/figure.png" pageId="26" pageNumber="235">
Fig.
<number box="[231,246,135,161]" pageId="26" pageNumber="235" value="8.0">8</number>
</figureCitation>
D). Juxtaocular bulges present, highlighted by parietal sulci. Vertex elevation yellow. Region posterior to the lateral ocelli concave. Ocelli larger than females; lateral ocelli slightly larger than unpaired median ocellus. Ocelli sit within an elevated ocellar tubercle that has three brown markings on the cuticle between the ocelli. Antennae as in females.
</paragraph>
<paragraph blockId="26.[106,1073,345,440]" pageId="26" pageNumber="235">
<emphasis box="[106,190,345,371]" italics="true" pageId="26" pageNumber="235">Thorax.</emphasis>
Pronotum as in females (
<figureCitation box="[491,575,345,371]" captionStart="Fig. 11" captionStartId="16.[106,139,1517,1539]" captionTargetBox="[243,935,131,1491]" captionTargetId="figure@16.[243,937,131,1492]" captionTargetPageId="16" captionText="Fig. 11. Illustrations of dorsal perspective of the Ilomantis pronotum (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. Ilomantis thalassina exhibits a medial keel that fully traverses the pronotum (A and B); Ilomantis ginsburgae sp. n. exhibits a medial keel that originates in the mid-prozone and concludes at the posterior pronotal margin (C and D). MK, medial keel." httpUri="https://zenodo.org/record/269850/files/figure.png" pageId="26" pageNumber="235">
Fig.
<number box="[545,575,345,371]" pageId="26" pageNumber="235" value="11.0">11</number>
</figureCitation>
D). Postcervical plate and exposed thoracic membrane of the cervix as in females (
<figureCitation box="[545,627,380,406]" captionStart="Fig. 13" captionStartId="18.[106,139,1269,1291]" captionTargetBox="[155,1024,131,1244]" captionTargetId="figure@18.[155,1024,131,1244]" captionTargetPageId="18" captionText="Fig. 13. Illustrations of ventral perspective of the Ilomantis cervix (scale bar = 1 mm). (A) Ilomantis thalassina, female; (B) Ilomantis thalassina, male; (C) Ilomantis ginsburgae sp. n., female; (D) Ilomantis ginsburgae sp. n., male. The postcervical plate and exposed thoracic membrane are contained within the boundaries of the intercervical sclerites and the episternum, with either a broad (Ilomantis thalassina, A and B) or narrow (Ilomantis ginsburgae sp. n., C and D) hexagonal shape. Abbreviations: eps, episternum; etm, exposed thoracic membrane; ics, intercervical sclerite; lcs, lateral cervical sclerite; pcpl, postcervical plate; tss, T-shaped sclerite." httpUri="https://zenodo.org/record/269852/files/figure.png" pageId="26" pageNumber="235">
Fig.
<number box="[597,627,380,406]" pageId="26" pageNumber="235" value="13.0">13</number>
</figureCitation>
D). Wings hyaline and iridescent; fore- and hindwing margins and surface with light ciliation.
</paragraph>
<paragraph blockId="26.[106,1074,486,547]" pageId="26" pageNumber="235">
<emphasis box="[106,282,486,512]" italics="true" pageId="26" pageNumber="235">Prothoracic legs.</emphasis>
Femur maximum width ca.
<number box="[618,669,487,513]" pageId="26" pageNumber="235" value="0.74">0.74</number>
times the value of the male femur maximum width.
<emphasis box="[310,326,522,547]" italics="true" pageId="26" pageNumber="235">F</emphasis>
=
<number box="[341,353,521,547]" pageId="26" pageNumber="235" value="3.0">3</number>
DS/
<number box="[399,426,521,547]" pageId="26" pageNumber="235" value="12.0">12</number>
AvS/
<number box="[485,499,521,547]" pageId="26" pageNumber="235" value="4.0">4</number>
PvS;
<emphasis box="[557,576,522,547]" italics="true" pageId="26" pageNumber="235">T</emphasis>
=
<number box="[592,660,521,547]" pageId="26" pageNumber="235" value="10.5" valueMax="11.0" valueMin="10.0">1011</number>
AvS/
<number box="[722,790,521,547]" pageId="26" pageNumber="235" value="11.5" valueMax="12.0" valueMin="11.0">1112</number>
PvS.
</paragraph>
<paragraph blockId="26.[106,1074,593,723]" pageId="26" pageNumber="235">
<emphasis box="[106,425,593,619]" italics="true" pageId="26" pageNumber="235">Meso- and metathoracic legs.</emphasis>
Densely ciliated across surface. Mesothoracic basitarsus approximately
<number box="[274,311,628,655]" pageId="26" pageNumber="235" value="1.5">1.5</number>
times the length of the remaining tarsal segments combined; metathoracic basitarsus approximately the same length as the remaining tarsal segments combined.
</paragraph>
<paragraph blockId="26.[106,1074,769,864]" pageId="26" pageNumber="235">
<emphasis box="[106,216,769,795]" italics="true" pageId="26" pageNumber="235">Abdomen.</emphasis>
Supra-anal plate triangular, ciliated across surface; much narrower than the subgenital plate. Subgenital plate approximately rounded at the terminus, ciliated. Cerci long, compressed, tapering. Styli short.
</paragraph>
<paragraph blockId="26.[106,1074,910,1457]" pageId="26" pageNumber="235">
<emphasis box="[106,209,910,936]" italics="true" pageId="26" pageNumber="235">Genitalia</emphasis>
(
<figureCitation box="[225,309,910,937]" captionStart="Fig. 10" captionStartId="14.[106,139,1247,1269]" captionTargetBox="[155,1024,131,1220]" captionTargetId="figure@14.[155,1024,131,1222]" captionTargetPageId="14" captionText="Fig. 10. Male genitalic complex of Ilomantis ginsburgae sp. n. (A) Right phallomere, ventral sclerotization of the left phallomeric complex, and dorsal (i. e., “ left ”) sclerotization of the left phallomeric complex (from left to right, respectively); (B, C, D) examples of shape and sclerotization variability present within the anterior process (ap) of L 4 B. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269849/files/figure.png" pageId="26" pageNumber="235">
Fig.
<number box="[277,309,911,937]" pageId="26" pageNumber="235" value="10.0">10</number>
</figureCitation>
). Dorsal sclerotization of the left phallomeric complex is anteriorly narrow and elongate, broadening towards the posterior margin; anterior process (ap) ranges from narrow and elongate to compact, tapering anteriorly; ap anterior margin variably sclerotized; Apical process (paa) dilated on the anterior margin of its visible “base,” recurved distally, narrow; paa with a rounded apical margin. Lobo membranoso (loa) relatively short, apex moderately sclerotized. Ventral sclerotization of the left phallomere complex (i.e., the ventral phallomere) (L
<number box="[720,734,1119,1145]" pageId="26" pageNumber="235" value="4.0">4</number>
A) longer than wide with an undulate, lightly sclerotized antero-sinistral margin; L
<number box="[704,717,1153,1179]" pageId="26" pageNumber="235" value="4.0">4</number>
A anterior margin varies slightly in width; L
<number box="[233,246,1188,1214]" pageId="26" pageNumber="235" value="4.0">4</number>
A posterior region lightly ciliated and bilobed; processo distale (pda) narrow, apically rounded, and well separated from a projecting dextral lobe that is broad and approximately rounded along the posterior margin. Anterior apodeme (an) anteriorly rounded with a moderately sclerotized sinistral margin; processo ventrale sclerificato (pva) obtusely curved, tumescent, with a heavily sclerotized posterior margin; piastra ventrale (pia) variably shaped, ranging from relatively straight to arcuate with a rugose surface. R
<number box="[310,320,1396,1422]" pageId="26" pageNumber="235" value="1.0">1</number>
posterior region lightly ciliated with a slightly tapered, rounded posterior margin; right arm (bm) varies in width, pointed apically.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,700 @@
<document ID-DOI="http://doi.org/10.5281/zenodo.6066871" ID-GBIF-Dataset="09101595-7414-42fa-b53f-5a1e9baadd98" ID-GBIF-Taxon="121067094" ID-Zenodo-Dep="6066871" checkinTime="1467004166608" checkinUser="plazi" docAuthor="Sydney K. Brannoch &amp; Gavin J. Svenson" docDate="2016" docId="4A09B002343BFFA6FF80F9FB6246F9E7" docLanguage="en" docName="InsectSystematicsEvoluition.47.4.209.pdf" docOrigin="Insect Systematics &amp; Evolution 47" docStyle="DocumentStyle{}" docTitle="Nilomantis Werner 1907" docType="key" docVersion="5" lastPageId="27" lastPageNumber="237" masterDocId="B630C87A3421FFBDFFEAFFFB666EFF94" masterDocTitle="Leveraging female genitalic characters for generic and species delimitation in Nilomantis Werner, 1907 and Ilomantis Giglio-Tos, 1915 (Mantodea, Nilomantinae)" masterLastPageNumber="244" masterPageNumber="209" pageId="26" pageNumber="235" updateTime="1644787795423" updateUser="ExternalLinkService">
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<mods:title>Leveraging female genitalic characters for generic and species delimitation in Nilomantis Werner, 1907 and Ilomantis Giglio-Tos, 1915 (Mantodea, Nilomantinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sydney K. Brannoch</mods:namePart>
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<mods:name type="personal">
<mods:role>
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<mods:namePart>Gavin J. Svenson</mods:namePart>
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<mods:number>47</mods:number>
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<taxonomicName authority="Werner, 1907" authorityName="Werner" authorityYear="1907" box="[106,410,1536,1563]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="235" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[106,410,1536,1563]" pageId="26" pageNumber="235">
<emphasis bold="true" box="[106,335,1536,1563]" italics="true" pageId="26" pageNumber="235">
Nilomantis
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Werner,
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</emphasis>
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<emphasis box="[106,223,1589,1615]" italics="true" pageId="26" pageNumber="235">Nilomantis</emphasis>
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Beier
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Uvarov
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7, 34;
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Ehrmann
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Otte &amp; Spearman
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<taxonomicName authority="Giglio-Tos, 1915: 45" authorityName="Giglio-Tos" authorityPageNumber="45" authorityYear="1915" box="[128,518,204,231]" class="Insecta" family="Iridopterygidae" genus="Cryptomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="27" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis box="[128,274,204,230]" italics="true" pageId="27" pageNumber="236">Cryptomantis</emphasis>
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Giglio-Tos,
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Giglio-Tos
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Ehrmann
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</paragraph>
</subSubSection>
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<paragraph blockId="27.[98,222,309,334]" box="[98,222,309,334]" pageId="27" pageNumber="236">
<heading box="[98,222,309,334]" fontSize="11" level="2" pageId="27" pageNumber="236" reason="1">
<emphasis box="[98,222,309,334]" italics="true" pageId="27" pageNumber="236">
<typeStatus box="[98,147,309,334]" pageId="27" pageNumber="236">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph blockId="27.[98,470,360,386]" box="[98,470,360,386]" pageId="27" pageNumber="236">
<heading box="[98,470,360,386]" fontSize="11" level="2" pageId="27" pageNumber="236" reason="1">
<taxonomicName authority="Werner, 1907" authorityName="Werner" authorityYear="1907" box="[98,466,360,386]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="27" pageNumber="236" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[98,300,360,386]" italics="true" pageId="27" pageNumber="236">Nilomantis floweri</emphasis>
<bibRefCitation author="Werner, F." box="[307,466,360,386]" journalOrPublisher="In Kommission bei Alfred Holder, Vienna" pageId="27" pageNumber="236" refString="Werner, F. (1907) Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. Franz Werner's in den agyptischen Sudan und nach Nord-Uganda: Orthoptera Blattaeformia. VII. aus der Kaiserlich-Koniglichen Hof-und Staatsdruckerei. In Kommission bei Alfred Holder, Vienna." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. Franz Werner's in den agyptischen Sudan und nach Nord-Uganda: Orthoptera Blattaeformia. VII. aus der Kaiserlich-Koniglichen Hof-und Staatsdruckerei" type="book" year="1907">
Werner,
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.
</heading>
</paragraph>
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<subSubSection pageId="27" pageNumber="236" type="diagnosis">
<paragraph blockId="27.[98,202,430,455]" box="[98,202,430,455]" pageId="27" pageNumber="236">
<heading box="[98,202,430,455]" fontSize="11" level="2" pageId="27" pageNumber="236" reason="1">
<emphasis box="[98,202,430,455]" italics="true" pageId="27" pageNumber="236">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="27.[98,1067,481,785]" pageId="27" pageNumber="236">
Small, slender mantises (body length
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), green; fully developed wings. Body and anterior margin of forewings lightly ciliated. Pronotal medial keel present. Head capsule dorsoventrally compressed; compound eyes dorsoventrally compressed, laterally conical, blunted. Head capsule with relatively straight vertical margin; cranial vertex with no projections. Lower frons featuring an arcuate carina. Female genitalia: gonoplac (GL) features a rounded dilation on the anteroventral margin; gonapophysis IX (GP) apex acuminate. Male genitalia: lobo membranoso (loa) with varying levels of sclerotization and distal crenulation; ventral sclerotization of the left phallomere (L
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A) terminates in two lobes.
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<subSubSection pageId="27" pageNumber="236" type="discussion">
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<heading box="[98,190,828,854]" fontSize="11" level="2" pageId="27" pageNumber="236" reason="1">
<emphasis box="[98,190,828,854]" italics="true" pageId="27" pageNumber="236">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="27.[98,1066,880,976]" pageId="27" pageNumber="236">
As evidenced by external and internal morphological characters, as well as by the geographical distribution of the genera, we contend that
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<emphasis box="[733,855,914,940]" italics="true" pageId="27" pageNumber="236">Nilomantis</emphasis>
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and
<taxonomicName box="[920,1022,914,940]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="27" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis box="[920,1022,914,940]" italics="true" pageId="27" pageNumber="236">Ilomantis</emphasis>
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are separate, valid genera.
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<subSubSection pageId="27" pageNumber="236" type="key">
<paragraph blockId="27.[98,575,1052,1079]" box="[98,575,1052,1079]" pageId="27" pageNumber="236">
<heading bold="true" box="[98,575,1052,1079]" fontSize="11" level="1" pageId="27" pageNumber="236" reason="0">
<emphasis bold="true" box="[98,575,1052,1079]" pageId="27" pageNumber="236">
Key to
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<emphasis bold="true" box="[184,317,1052,1079]" italics="true" pageId="27" pageNumber="236">Nilomantis</emphasis>
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Werner,
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species
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</heading>
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<number box="[98,114,1105,1131]" pageId="27" pageNumber="236" value="1.0">1</number>
. Female genitalia: Gonocoxa VIII (VF) internal margin rounded; from LH, VF without projection. Apical lobe (AL) ovoid, ciliated. Gonoplac (GL) ventral margin rugose, dorsal margin relatively smooth. Male genitalia: lobo membranoso (loa) heavily sclerotized and strongly crenulated distally; processo apical (paa) narrow and recurved apically. The ventral sclerotization of the left phallomere (L
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A) relatively narrow, terminating into two triangular lobes; processo distale (pda) terminating in a lightly sclerotized point. Processo ventrale sclerificato (pva) and piastra ventrale (pia) reduced and closely adjoined. Distributed within the Sahara-Sahel region of Africa and
<collectingCountry box="[98,247,1382,1408]" name="Saudi Arabia" pageId="27" pageNumber="236">Saudi Arabia</collectingCountry>
.
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. Female genitalia: VF internal margin approximately angular in appearance; from LH, VF possesses a distinct knob-like projection that is directed dorsad. AL ovoid, pitted; GL dorsal and ventral margins relatively smooth. Male genitalia: loa invaginated and very lightly crenulated distally; paa broad and recurved. L
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A broad, terminating into two well separated, apically rounded lobes; pda broad, largely rounded. Pva and pia sclerotized, finger-like. Distributed within
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Nilomantis floweri
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<emphasis box="[106,333,290,317]" italics="true" pageId="28" pageNumber="237">Cryptomantis tenella</emphasis>
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<emphasis box="[106,313,360,386]" italics="true" pageId="28" pageNumber="237">Nilomantis arabica</emphasis>
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[
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tenella (
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The list consists of
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.
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:
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♀, Formation arborée à Ficus et Olea,
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N -
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E,
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-VI-
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, M. Donskoff reç., Genitalia No. 0 0 0
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E,
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, M. Donskoff reç., Genitalia No. 0 0 0
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, M. Donskoff reç. (
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);
<emphasis box="[106,262,764,790]" italics="true" pageId="28" pageNumber="237">Wadi Turabah</emphasis>
,
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, milieu de vallée,
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°
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<number box="[592,620,764,790]" pageId="28" pageNumber="237" value="12.0">12</number>
N -
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°
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E,
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,
<date box="[913,1068,764,790]" pageId="28" pageNumber="237" value="1992-07-03">
03-VII-
<number box="[1007,1068,764,790]" pageId="28" pageNumber="237" value="1992.0">1992</number>
</date>
, M., Donskoff reç., Genitalia No. 0 0 0
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0
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S
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. K. Brannoch (
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)
</paragraph>
</subSubSection>
<subSubSection lastPageId="29" lastPageNumber="238" pageId="28" pageNumber="237" type="description">
<paragraph blockId="28.[106,386,873,899]" box="[106,386,873,899]" pageId="28" pageNumber="237">
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<emphasis box="[106,386,873,899]" italics="true" pageId="28" pageNumber="237">Male genitalia description</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[106,1074,925,1195]" pageId="28" pageNumber="237">
Based on an examination of the genital complex of
<taxonomicName box="[708,820,925,951]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[708,820,925,951]" italics="true" pageId="28" pageNumber="237">N. floweri</emphasis>
</taxonomicName>
, as well as plates presented by
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Roy &amp; Leston (
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, the anterior process (ap) is lightly sclerotized, undulate. Apical process (paa) features a broad base with an anterior dilation; paa narrow and recurved distally. Apophysis phalloide (L
<number box="[634,644,1029,1055]" pageId="28" pageNumber="237" value="1.0">1</number>
) does not retain the apophysis; lobo membranoso (loa) heavily sclerotized and strongly crenulated distally. Processo distale (pda) narrows apically, terminating in a moderately sclerotized, blunted point. There is a second, triangular lobe to the right of pda. Processo ventrale sclerificato (pva) and piastra ventrale (pia) reduced, heavily sclerotized, and closely adjoined.
</paragraph>
<paragraph blockId="28.[106,1074,1242,1650]" lastBlockId="29.[98,1066,134,404]" lastPageId="29" lastPageNumber="238" pageId="28" pageNumber="237">
<emphasis box="[106,406,1242,1268]" italics="true" pageId="28" pageNumber="237">Female genitalia description</emphasis>
(
<figureCitation box="[420,488,1242,1269]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="28" pageNumber="237">
Figs
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A,
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A,B). Gonapophysis VIII (GA) slender, variably ciliated, and visibly curved from both the ventral (VH) and lateral (LH) perspectives; GA lightly surfaced with fossettes. From VH, gonocoxa VIII (VF) internal margin rounded; from LH, VF does not retain a projection. From VH, GA curves strongly inward; apical lobe (AL) ovoid, ciliated. AL has a small membrane which projects from the internal dorsal margin of the structure, forming a shallow pocket as the membrane merges with the dorsal margin of GA. From LH, GA is gently curved ventrad; GA ventral margin features a pronounced medial outgrowth (MO), which projects posterior to the crest of the curve, with a row of cilia that runs from the crest of MO towards AL. GA external margin possesses a moderately sclerotized tubercle anterior to MO. From LH, AL narrows posteriorly with a bluntly rounded apical margin. GA dorsal margin is invaginated anterior to the curve towards the ventral habitus. GA very lightly sclerotized. Gonapophysis IX (GP) relatively short, broadening at the site of the medial tine (MT) before narrowing into a tapered, acuminate apex; GP ventral margin invaginated, covered anteriorly by MT. Gonoplac (GL) shorter than GA, broad, variably ciliated, and arched ventrad; from LH, GL base has a rounded dilation on the anteroventral margin. GL narrows toward the apex; from LH, GL apex slants ventrad. GL features a reduced apical cleft (AC) resulting in a slight apical notch. From LH, GL possesses a slight sclerotization that traverses the length of the structure, terminating just before the AC. GL ventral margin rugose, dorsal margin relatively smooth.
</paragraph>
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</treatment>
</emphasis>
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,451 @@
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<taxonomicName authority="Roy, 1975" authorityName="Roy" authorityYear="1975" box="[98,482,480,507]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="29" pageNumber="238" phylum="Arthropoda" rank="species" species="edmundsi">
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<emphasis bold="true" box="[98,354,480,507]" italics="true" pageId="29" pageNumber="238">Nilomantis edmundsi</emphasis>
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Edmunds
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7, 34;
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Edmunds
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Ehrmann
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:
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Otte &amp; Spearman
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The list consists of
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specimen
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.
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;
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♀,
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001500, MN
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Secondary voucher, Genitalia No. 0 0 0
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0
<number box="[423,436,724,751]" pageId="29" pageNumber="238" value="5.0">5</number>
S
</geoCoordinate>
. K. Brannoch.
</paragraph>
</subSubSection>
<subSubSection lastPageId="30" lastPageNumber="239" pageId="29" pageNumber="238" type="description">
<paragraph blockId="29.[98,379,793,819]" box="[98,379,793,819]" pageId="29" pageNumber="238">
<heading box="[98,379,793,819]" fontSize="11" level="2" pageId="29" pageNumber="238" reason="1">
<emphasis box="[98,379,793,819]" italics="true" pageId="29" pageNumber="238">Male genitalia description</emphasis>
</heading>
</paragraph>
<paragraph blockId="29.[98,1066,845,1115]" pageId="29" pageNumber="238">
An examination of the plates presented by
<bibRefCitation author="Roy, R. &amp; Leston, D." box="[602,855,845,872]" journalOrPublisher="Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A" pageId="29" pageNumber="238" pagination="297 - 344" part="37" refString="Roy, R. &amp; Leston, D. (1975) Mantodea of Ghana: new species, further records and habitats. Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A 37: 297 - 344." title="Mantodea of Ghana: new species, further records and habitats" type="journal article" year="1975">
Roy &amp; Leston (
<number box="[786,846,845,872]" pageId="29" pageNumber="238" value="1975.0">1975</number>
)
</bibRefCitation>
indicates that the anterior process (ap) is narrow. Apical process (paa) broad and recurved. Apophysis phalloide (L
<number box="[239,249,915,941]" pageId="29" pageNumber="238" value="1.0">1</number>
) does not retain the apophysis; lobo membranoso (loa) lightly sclerotized and crenulated on the distal margin. Ventral sclerotization of the left phallomeric process (L
<number box="[176,190,984,1010]" pageId="29" pageNumber="238" value="4.0">4</number>
A) terminates in two lobes; processo distale (pda) largely rounded and well separated from a secondary lobe that is rounded and relatively smaller. Processo ventrale sclerificato (pva) and piastra ventrale (pia) sclerotized and more well defined than in
<taxonomicName box="[129,238,1088,1114]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="29" pageNumber="238" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[129,238,1088,1114]" italics="true" pageId="29" pageNumber="238">N. floweri</emphasis>
</taxonomicName>
; right arm (bm) pointed distally and lightly sclerotized.
</paragraph>
<paragraph blockId="29.[98,584,1157,1183]" box="[98,584,1157,1183]" pageId="29" pageNumber="238">
<heading box="[98,584,1157,1183]" fontSize="11" level="2" pageId="29" pageNumber="238" reason="1">
<emphasis box="[98,584,1157,1183]" italics="true" pageId="29" pageNumber="238">
Female genitalia description (
<figureCitation box="[414,481,1157,1183]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="29" pageNumber="238">
Figs
<number box="[466,481,1157,1183]" pageId="29" pageNumber="238" value="4.0">4</number>
</figureCitation>
B,
<number box="[512,528,1157,1183]" pageId="29" pageNumber="238" value="6.0">6</number>
C,D)
</emphasis>
</heading>
</paragraph>
<paragraph blockId="29.[98,1067,1209,1652]" lastBlockId="30.[106,1074,134,403]" lastPageId="30" lastPageNumber="239" pageId="29" pageNumber="238">Gonapophysis VIII (GA) slender, variably ciliated, slightly arcuate, and visibly curved from both the ventral (VH) and lateral (LH) perspectives; GA surfaced with fossettes. From VH, gonocoxa VIII (VF) internal margin narrow, rounded, lending VF an angular appearance; from LH, VF possesses a distinct knob-like projection that is directed dorsad. From VH, GA gently curves inward. From LH, GA obtusely curved ventrad; GA ventral margin features a rugose, sparsely ciliated medial outgrowth (MO), which projects posterior to the crest of the curve. GA external margin possesses a moderately sclerotized tubercle anterior to MO. From VH, apical lobe (AL) ovoid, pitted, distinctly lacking cilia across surface majority; from LH, AL thin, arched, pitted, and approximately pointed; no visible cilia observed. AL has a pronounced membrane that projects from the internal dorsal margin of the structure, forming a shallow pocket as the membrane merges with the dorsal margin of GA, lending AL the appearance of being bilobed or mitten-shaped. GA dorsal margin invaginated at the region that bends towards the ventral habitus. Gonapophysis IX (GP) relatively short, broadening at the site of the medial tine (MT) before narrowing into a tapered, acuminate apex; GP ventral margin invaginated, covered anteriorly by MT. Gonoplac (GL) shorter than GA, broad, and obtusely arched ventrad; from LH, GL base has a rounded dilation on the anteroventral margin. GL narrows toward the apex; from LH, GL apex slants ventrad; apical cleft (AC) reduced, resulting in a slight apical notch. From LH, GL possesses a pronounced sclerotization that traverses the length of the structure, terminating just before the AC. GL dorsal and ventral margins relatively smooth.</paragraph>
</subSubSection>
<subSubSection lastPageId="31" lastPageNumber="240" pageId="30" pageNumber="239" type="discussion">
<paragraph blockId="30.[106,237,465,491]" box="[106,237,465,491]" pageId="30" pageNumber="239">
<heading bold="true" box="[106,237,465,491]" fontSize="11" level="1" pageId="30" pageNumber="239" reason="0">
<emphasis bold="true" box="[106,237,465,491]" pageId="30" pageNumber="239">Discussion</emphasis>
</heading>
</paragraph>
<paragraph blockId="30.[106,1074,517,1653]" pageId="30" pageNumber="239">
As a means to consider the validity of
<taxonomicName box="[535,638,517,543]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[535,638,517,543]" italics="true" pageId="30" pageNumber="239">Ilomantis</emphasis>
</taxonomicName>
, we examined the female genital complexes of the species included in
<taxonomicName box="[488,611,552,578]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[488,611,552,578]" italics="true" pageId="30" pageNumber="239">Nilomantis</emphasis>
</taxonomicName>
and
<taxonomicName box="[672,775,552,578]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[672,775,552,578]" italics="true" pageId="30" pageNumber="239">Ilomantis</emphasis>
</taxonomicName>
for potential intergeneric diagnostic characters. The shape of the dilation on the anteroventral margin of the gonoplac (GL) varies between the genera, with
<taxonomicName box="[645,768,621,647]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[645,768,621,647]" italics="true" pageId="30" pageNumber="239">Nilomantis</emphasis>
</taxonomicName>
possessing a rounded dilation, whereas
<taxonomicName box="[266,369,656,682]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[266,369,656,682]" italics="true" pageId="30" pageNumber="239">Ilomantis</emphasis>
</taxonomicName>
features a dilation that is approximately rectangular in shape (compare
<figureCitation box="[224,296,690,717]" captionStart="Fig. 3" captionStartId="7.[98,131,1511,1533]" captionTargetBox="[147,1016,360,1484]" captionTargetId="figure@7.[147,1016,360,1485]" captionTargetPageId="7" captionText="Fig. 3. Female genitalic complex of Ilomantis shown in the lateral perspective. (A) Ilomantis thalassina; (B) Ilomantis ginsburgae sp. n. Abbreviations: AC, apical cleft; AL, apical lobe; GA, gonapophysis VIII; GL, gonoplac; MO, medial outgrowth; RTD, rectangular dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269842/files/figure.png" pageId="30" pageNumber="239">
Figs
<number box="[281,296,691,717]" pageId="30" pageNumber="239" value="3.0">3</number>
</figureCitation>
and
<figureCitation box="[360,378,691,717]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="30" pageNumber="239">
<number box="[360,378,691,717]" pageId="30" pageNumber="239" value="4.0">4</number>
</figureCitation>
). Furthermore, in
<taxonomicName box="[601,724,690,716]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[601,724,690,716]" italics="true" pageId="30" pageNumber="239">Nilomantis</emphasis>
</taxonomicName>
the apex of gonapophysis IX (GP) is acuminate, whereas in
<taxonomicName box="[465,568,725,751]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[465,568,725,751]" italics="true" pageId="30" pageNumber="239">Ilomantis</emphasis>
</taxonomicName>
the apex of GP is mammiform to approximately rounded in shape (compare
<figureCitation box="[514,583,760,787]" captionStart="Fig. 5" captionStartId="9.[98,131,1328,1350]" captionTargetBox="[147,1016,131,1303]" captionTargetId="figure@9.[147,1016,131,1303]" captionTargetPageId="9" captionText="Fig. 5. Female genitalic complex of Ilomantis shown in the ventral perspective with natural and spread positioning. (A) Ilomantis thalassina, natural; (B) Ilomantis thalassina, spread; (C) Ilomantis ginsburgae sp. n., natural; (D) Ilomantis ginsburgae sp. n., spread. Abbreviations: AC, apical cleft; AL, apical lobe; MO, medial outgrowth; MT, medial tine; VF, gonocoxa VIII; GA, gonapophysis VIII; GP, gonapophysis IX II; GL, gonoplac. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269844/files/figure.png" pageId="30" pageNumber="239">
Figs
<number box="[568,583,760,787]" pageId="30" pageNumber="239" value="5.0">5</number>
</figureCitation>
and
<figureCitation box="[641,658,760,787]" captionStart="Fig. 6" captionStartId="10.[106,139,1438,1460]" captionTargetBox="[155,1024,131,1412]" captionTargetId="figure@10.[155,1024,131,1412]" captionTargetPageId="10" captionText="Fig. 6. Female genitalic complex of Nilomantis shown in the ventral perspective with natural and spread positioning. (A) Nilomantis floweri, natural; (B) Nilomantis floweri, spread; (C) Nilomantis edmundsi, natural; (D) Nilomantis edmundsi, spread. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269845/files/figure.png" pageId="30" pageNumber="239">
<number box="[641,658,760,787]" pageId="30" pageNumber="239" value="6.0">6</number>
</figureCitation>
). These character states were able to consistently and unambiguously distinguish the genera, which, when compared against external morphological characters, male genitalic characters, and geographic distribution, enabled us to resurrect
<taxonomicName box="[521,624,864,890]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis box="[521,624,864,890]" italics="true" pageId="30" pageNumber="239">Ilomantis</emphasis>
</taxonomicName>
as a valid genus with confidence.
</paragraph>
<paragraph blockId="30.[106,1074,517,1653]" pageId="30" pageNumber="239">
We likewise examined the female genital complexes of
<taxonomicName box="[748,880,898,924]" class="Insecta" family="Iridopterygidae" genus="Tropidomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[748,880,898,924]" italics="true" pageId="30" pageNumber="239">I. thalassina</emphasis>
</taxonomicName>
and
<taxonomicName box="[935,1073,898,924]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[935,1073,898,924]" italics="true" pageId="30" pageNumber="239">I. ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[106,160,934,960]" pageId="30" pageNumber="239" rank="species">sp.n.</taxonomicNameLabel>
as a site for potential interspecific diagnostic characters. We found that gonapophysis VIII (GA) and GL feature compelling interspecific character states that proved successful in delimiting the species. Generally,
<taxonomicName box="[654,873,1002,1028]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[654,873,1002,1028]" italics="true" pageId="30" pageNumber="239">Ilomantis thalassina</emphasis>
</taxonomicName>
features broader, more robust gonapophyses and gonoplacs than
<taxonomicName box="[620,752,1037,1063]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[620,752,1037,1063]" italics="true" pageId="30" pageNumber="239">I. ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[758,810,1038,1064]" pageId="30" pageNumber="239" rank="species">sp.n.</taxonomicNameLabel>
(compare
<figureCitation box="[925,989,1037,1063]" captionStart="Fig. 3" captionStartId="7.[98,131,1511,1533]" captionTargetBox="[147,1016,360,1484]" captionTargetId="figure@7.[147,1016,360,1485]" captionTargetPageId="7" captionText="Fig. 3. Female genitalic complex of Ilomantis shown in the lateral perspective. (A) Ilomantis thalassina; (B) Ilomantis ginsburgae sp. n. Abbreviations: AC, apical cleft; AL, apical lobe; GA, gonapophysis VIII; GL, gonoplac; MO, medial outgrowth; RTD, rectangular dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269842/files/figure.png" pageId="30" pageNumber="239">
Figs
<number box="[975,989,1037,1063]" pageId="30" pageNumber="239" value="3.0">3</number>
</figureCitation>
and
<figureCitation box="[1043,1059,1037,1064]" captionStart="Fig. 5" captionStartId="9.[98,131,1328,1350]" captionTargetBox="[147,1016,131,1303]" captionTargetId="figure@9.[147,1016,131,1303]" captionTargetPageId="9" captionText="Fig. 5. Female genitalic complex of Ilomantis shown in the ventral perspective with natural and spread positioning. (A) Ilomantis thalassina, natural; (B) Ilomantis thalassina, spread; (C) Ilomantis ginsburgae sp. n., natural; (D) Ilomantis ginsburgae sp. n., spread. Abbreviations: AC, apical cleft; AL, apical lobe; MO, medial outgrowth; MT, medial tine; VF, gonocoxa VIII; GA, gonapophysis VIII; GP, gonapophysis IX II; GL, gonoplac. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269844/files/figure.png" pageId="30" pageNumber="239">
<number box="[1043,1059,1037,1064]" pageId="30" pageNumber="239" value="5.0">5</number>
</figureCitation>
). Within
<taxonomicName box="[196,328,1072,1098]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[196,328,1072,1098]" italics="true" pageId="30" pageNumber="239">I. thalassina</emphasis>
</taxonomicName>
, GA features a pronounced medial outgrowth (MO); GA anterodorsal margin is moderately sclerotized, with the sclerotized region concluding just posterior to the “heel” of GA. The apical cleft (AC) of GL is distinctly divergent, the resultant lobes blunt apically; the ventral and dorsal margins of GL are rugose, sculpturate (see
<figureCitation box="[232,297,1210,1237]" captionStart="Fig. 3" captionStartId="7.[98,131,1511,1533]" captionTargetBox="[147,1016,360,1484]" captionTargetId="figure@7.[147,1016,360,1485]" captionTargetPageId="7" captionText="Fig. 3. Female genitalic complex of Ilomantis shown in the lateral perspective. (A) Ilomantis thalassina; (B) Ilomantis ginsburgae sp. n. Abbreviations: AC, apical cleft; AL, apical lobe; GA, gonapophysis VIII; GL, gonoplac; MO, medial outgrowth; RTD, rectangular dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269842/files/figure.png" pageId="30" pageNumber="239">
Fig.
<number box="[284,297,1211,1237]" pageId="30" pageNumber="239" value="3.0">3</number>
</figureCitation>
A). However, in
<taxonomicName box="[491,631,1210,1236]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[491,631,1210,1236]" italics="true" pageId="30" pageNumber="239">I. ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[640,696,1211,1237]" pageId="30" pageNumber="239" rank="species">sp.n.</taxonomicNameLabel>
, MO of GA is reduced; and GA anterodorsal margin is less sclerotized than in
<taxonomicName box="[626,758,1245,1271]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[626,758,1245,1271]" italics="true" pageId="30" pageNumber="239">I. thalassina</emphasis>
</taxonomicName>
, with the region of sclerotization terminating just prior to the apical lobes (AL). The AC of GL is significantly reduced, with the lobe of the dorsal margin appearing to converge with the ventral lobe, resulting in a slightly notched, acuminate apex; GL ventral and dorsal margins are smooth (see
<figureCitation box="[288,352,1384,1410]" captionStart="Fig. 3" captionStartId="7.[98,131,1511,1533]" captionTargetBox="[147,1016,360,1484]" captionTargetId="figure@7.[147,1016,360,1485]" captionTargetPageId="7" captionText="Fig. 3. Female genitalic complex of Ilomantis shown in the lateral perspective. (A) Ilomantis thalassina; (B) Ilomantis ginsburgae sp. n. Abbreviations: AC, apical cleft; AL, apical lobe; GA, gonapophysis VIII; GL, gonoplac; MO, medial outgrowth; RTD, rectangular dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269842/files/figure.png" pageId="30" pageNumber="239">
Fig.
<number box="[338,352,1384,1410]" pageId="30" pageNumber="239" value="3.0">3</number>
</figureCitation>
B).
</paragraph>
<paragraph blockId="30.[106,1074,517,1653]" lastBlockId="31.[98,1067,134,1652]" lastPageId="31" lastPageNumber="240" pageId="30" pageNumber="239">
Although the number of specimens were limited, we examined intraspecific variation within
<specimenCount box="[240,345,1453,1479]" pageId="30" pageNumber="239" type="female">
<number box="[240,255,1453,1479]" pageId="30" pageNumber="239" value="2.0">2</number>
females
</specimenCount>
of
<taxonomicName box="[382,514,1453,1479]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[382,514,1453,1479]" italics="true" pageId="30" pageNumber="239">I. thalassina</emphasis>
</taxonomicName>
and
<specimenCount box="[570,674,1453,1479]" pageId="30" pageNumber="239" type="female">
<number box="[570,585,1453,1479]" pageId="30" pageNumber="239" value="3.0">3</number>
females
</specimenCount>
of
<taxonomicName box="[712,850,1453,1479]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="30" pageNumber="239" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[712,850,1453,1479]" italics="true" pageId="30" pageNumber="239">I. ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[858,913,1454,1480]" pageId="30" pageNumber="239" rank="species">sp.n.</taxonomicNameLabel>
; variation was most frequently observed in the apical lobe (AL) when viewed from the lateral perspective, ranging from rounded, to blunted, to angled in shape. However, this might be due to AL being a membranous structure, which can be actively or passively moved by an insect, and which might also slightly change shape when submerged within ethanol. Furthermore, AL is a lobe-like apical dilation of GA that is folded toward the external surface along the ventral margin, which could lend AL a varied appearance. We found that the overall shape of the female genitalia was more uniform in comparison to the intraspecific variation of character states observed in male genitalia, specifically in the overall length and width of the apical process (ap) of L
<number box="[741,756,239,265]" pageId="31" pageNumber="240" value="4.0">4</number>
B, as well as the shape and sclerotization of the anterior margin of ap (see
<figureCitation box="[616,695,273,299]" captionStart="Fig. 10" captionStartId="14.[106,139,1247,1269]" captionTargetBox="[155,1024,131,1220]" captionTargetId="figure@14.[155,1024,131,1222]" captionTargetPageId="14" captionText="Fig. 10. Male genitalic complex of Ilomantis ginsburgae sp. n. (A) Right phallomere, ventral sclerotization of the left phallomeric complex, and dorsal (i. e., “ left ”) sclerotization of the left phallomeric complex (from left to right, respectively); (B, C, D) examples of shape and sclerotization variability present within the anterior process (ap) of L 4 B. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269849/files/figure.png" pageId="31" pageNumber="240">
Fig.
<number box="[665,695,273,299]" pageId="31" pageNumber="240" value="10.0">10</number>
</figureCitation>
) of the one
<taxonomicName box="[828,958,273,299]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="thalassina">
<emphasis box="[828,958,273,299]" italics="true" pageId="31" pageNumber="240">I. thalassina</emphasis>
</taxonomicName>
male and four
<taxonomicName box="[153,292,308,334]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[153,292,308,334]" italics="true" pageId="31" pageNumber="240">I. ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[299,353,308,334]" pageId="31" pageNumber="240" rank="species">sp.n.</taxonomicNameLabel>
males examined.
</paragraph>
<paragraph blockId="31.[98,1067,134,1652]" pageId="31" pageNumber="240">
An examination of the female genital complexes of
<taxonomicName box="[708,910,342,368]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[708,910,342,368]" italics="true" pageId="31" pageNumber="240">Nilomantis floweri</emphasis>
</taxonomicName>
(
<emphasis box="[927,942,343,368]" italics="true" pageId="31" pageNumber="240">n</emphasis>
=
<number box="[960,970,343,369]" pageId="31" pageNumber="240" value="1.0">1</number>
) and
<taxonomicName class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="edmundsi">
<emphasis italics="true" pageId="31" pageNumber="240">N. edmundsi</emphasis>
</taxonomicName>
(
<emphasis box="[218,233,378,403]" italics="true" pageId="31" pageNumber="240">n</emphasis>
=
<number box="[252,262,377,403]" pageId="31" pageNumber="240" value="1.0">1</number>
) led to the addition of new characters with which to delimit the species. In particular,
<taxonomicName box="[250,359,412,438]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[250,359,412,438]" italics="true" pageId="31" pageNumber="240">N. floweri</emphasis>
</taxonomicName>
features a gonocoxa VIII (VF) with a rounded internal margin; which, in
<taxonomicName box="[212,352,446,472]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="edmundsi">
<emphasis box="[212,352,446,472]" italics="true" pageId="31" pageNumber="240">N. edmundsi</emphasis>
</taxonomicName>
, is approximately angular (see
<figureCitation box="[699,766,446,473]" captionStart="Fig. 6" captionStartId="10.[106,139,1438,1460]" captionTargetBox="[155,1024,131,1412]" captionTargetId="figure@10.[155,1024,131,1412]" captionTargetPageId="10" captionText="Fig. 6. Female genitalic complex of Nilomantis shown in the ventral perspective with natural and spread positioning. (A) Nilomantis floweri, natural; (B) Nilomantis floweri, spread; (C) Nilomantis edmundsi, natural; (D) Nilomantis edmundsi, spread. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269845/files/figure.png" pageId="31" pageNumber="240">
Fig.
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</figureCitation>
). From LH, VF is without projection in
<taxonomicName box="[258,371,481,507]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[258,371,481,507]" italics="true" pageId="31" pageNumber="240">N. floweri</emphasis>
</taxonomicName>
, whereas VF features a distinct knob-like projection that is directed dorsad (see
<figureCitation box="[328,394,516,542]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="31" pageNumber="240">
Fig.
<number box="[377,394,516,542]" pageId="31" pageNumber="240" value="4.0">4</number>
</figureCitation>
). The apical lobe (AL) of
<taxonomicName box="[683,792,516,542]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[683,792,516,542]" italics="true" pageId="31" pageNumber="240">N. floweri</emphasis>
</taxonomicName>
is heavily ciliated, while in
<taxonomicName box="[128,269,550,576]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="edmundsi">
<emphasis box="[128,269,550,576]" italics="true" pageId="31" pageNumber="240">N. edmundsi</emphasis>
</taxonomicName>
AL cilia were not observed across the majority of its surface (see
<figureCitation box="[999,1066,550,577]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="31" pageNumber="240">
Figs
<number box="[1051,1066,551,577]" pageId="31" pageNumber="240" value="4.0">4</number>
</figureCitation>
and
<figureCitation box="[147,164,585,612]" captionStart="Fig. 6" captionStartId="10.[106,139,1438,1460]" captionTargetBox="[155,1024,131,1412]" captionTargetId="figure@10.[155,1024,131,1412]" captionTargetPageId="10" captionText="Fig. 6. Female genitalic complex of Nilomantis shown in the ventral perspective with natural and spread positioning. (A) Nilomantis floweri, natural; (B) Nilomantis floweri, spread; (C) Nilomantis edmundsi, natural; (D) Nilomantis edmundsi, spread. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksandjournals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269845/files/figure.png" pageId="31" pageNumber="240">
<number box="[147,164,585,612]" pageId="31" pageNumber="240" value="6.0">6</number>
</figureCitation>
). Lastly, the ventral margin of the gonoplac (GL) is rugose and the dorsal margin is relatively smooth in
<taxonomicName box="[358,468,620,646]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[358,468,620,646]" italics="true" pageId="31" pageNumber="240">N. floweri</emphasis>
</taxonomicName>
, whereas both the ventral and dorsal margins of GL in
<taxonomicName box="[129,269,654,680]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="edmundsi">
<emphasis box="[129,269,654,680]" italics="true" pageId="31" pageNumber="240">N. edmundsi</emphasis>
</taxonomicName>
are relatively smooth (see
<figureCitation box="[567,634,654,681]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="31" pageNumber="240">
Fig.
<number box="[617,634,655,681]" pageId="31" pageNumber="240" value="4.0">4</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="31.[98,1067,134,1652]" pageId="31" pageNumber="240">
The examined specimens were collected between
<number box="[709,753,689,715]" pageId="31" pageNumber="240" value="116.0">116</number>
to
<number box="[800,829,689,715]" pageId="31" pageNumber="240" value="13.0">13</number>
years ago, with the majority collected around
<number box="[408,437,724,751]" pageId="31" pageNumber="240" value="50.0">50</number>
years ago. With material of this age, desiccation and damage from drawer movement, museum environmental conditions (e.g., fluctuations in temperature or humidity), and specimen handling may occur. While we took great pains to ensure that the entire female and male genital complex of each specimen was removed intact, we were unable to isolate the spermatheca, vestibular floor, and the laterosternal shelf of the female genitalia. This might be due to the fact that the genitalia of these species are membranous, and when coupled with specimen age, more susceptible to being destroyed over time or while submerged in potassium hydroxide, a solution frequently used to clear insect genitalia. While we were not able to access these structures within the female genital complex of
<taxonomicName box="[635,737,1036,1062]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis box="[635,737,1036,1062]" italics="true" pageId="31" pageNumber="240">Ilomantis</emphasis>
</taxonomicName>
and
<taxonomicName box="[793,915,1036,1062]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis box="[793,915,1036,1062]" italics="true" pageId="31" pageNumber="240">Nilomantis</emphasis>
</taxonomicName>
, we were still able to find characters on the gonapophyses, gonoplacs, and gonocoxae, which unambiguously delimited the genera and the species.
</paragraph>
<paragraph blockId="31.[98,1067,134,1652]" pageId="31" pageNumber="240">
Our investigation of mantodean female genitalia revealed a novel character system that successfully resurrected
<taxonomicName box="[417,518,1174,1200]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis box="[417,518,1174,1200]" italics="true" pageId="31" pageNumber="240">Ilomantis</emphasis>
</taxonomicName>
as a valid genus, delimited the species contained within both
<taxonomicName box="[237,338,1209,1235]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis box="[237,338,1209,1235]" italics="true" pageId="31" pageNumber="240">Ilomantis</emphasis>
</taxonomicName>
and
<taxonomicName box="[393,514,1209,1235]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis box="[393,514,1209,1235]" italics="true" pageId="31" pageNumber="240">Nilomantis</emphasis>
</taxonomicName>
(i.e.,
<emphasis box="[577,828,1209,1235]" italics="true" pageId="31" pageNumber="240">
<taxonomicName box="[577,710,1209,1235]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="thalassina">I. thalassina</taxonomicName>
,
<taxonomicName box="[720,828,1209,1235]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="floweri">N. floweri</taxonomicName>
</emphasis>
and
<taxonomicName box="[883,1022,1209,1235]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="edmundsi">
<emphasis box="[883,1022,1209,1235]" italics="true" pageId="31" pageNumber="240">N. edmundsi</emphasis>
</taxonomicName>
), as well as described a new
<collectingCountry box="[360,464,1244,1270]" name="Madagascar" pageId="31" pageNumber="240">Malagasy</collectingCountry>
praying mantis species (i.e.,
<taxonomicName box="[782,920,1244,1270]" class="Insecta" family="Iridopterygidae" genus="Ilomantis" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="species" species="ginsburgae" status="sp.n.">
<emphasis box="[782,920,1244,1270]" italics="true" pageId="31" pageNumber="240">I. ginsburgae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[926,982,1244,1270]" pageId="31" pageNumber="240" rank="species">sp.n.</taxonomicNameLabel>
). These results underscore the utility of female genitalia in
<taxonomicName box="[691,806,1278,1304]" class="Insecta" family="Mantodea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="order">Mantodea</taxonomicName>
taxonomy, and subsequently demonstrate the need for thorough examination of female genital complexes as sites of taxonomic characters. Furthermore, by describing both male and female genitalic characters (when available), the ability to diagnose females in externally homogenous genera might be resolved within
<taxonomicName box="[638,757,1417,1443]" class="Insecta" family="Mantodea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="31" pageNumber="240" phylum="Arthropoda" rank="order">Mantodea</taxonomicName>
. For this study, traditional taxonomic data, such as external morphology, male genitalic characters, and geographic distribution, supported the results of the investigation of the female genitalic character system. As such, it is our hope that future taxonomic descriptions of praying mantises assess female genitalic characters in addition to traditional taxonomic treatments. A more thorough investigation of female genitalia across a wide taxon sampling might provide a new motif from which to build a novel taxonomic character system.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,440 @@
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<taxonomicName authority="Werner, 1907" authorityName="Werner" authorityYear="1907" box="[106,498,134,161]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="species" species="floweri">
<emphasis bold="true" box="[106,498,134,161]" pageId="28" pageNumber="237">
Nilomantis floweri
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Werner,
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</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<emphasis box="[106,307,186,212]" italics="true" pageId="28" pageNumber="237">Nilomantis floweri</emphasis>
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Werner,
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:
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</taxonomicName>
;
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Beier
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Uvarov
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Kaltenbach
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;
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<bibRefCitation author="Ehrmann, R." box="[634,809,221,247]" journalOrPublisher="Munster, Natur und Tier Verlag Munster" pageId="28" pageNumber="237" refString="Ehrmann, R. (2002) Mantodea: Gottesanberinnen der Welt. Munster, Natur und Tier Verlag Munster." title="Mantodea: Gottesanberinnen der Welt" type="book" year="2002">
Ehrmann
<number box="[747,809,221,247]" pageId="28" pageNumber="237" value="2002.0">2002</number>
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:
<number box="[818,865,221,247]" pageId="28" pageNumber="237" value="242.0">242</number>
</treatmentCitation>
;
<treatmentCitation author="Otte, D. &amp; Spearman, L. A." page="120" pageId="28" pageNumber="237" year="2005">
<bibRefCitation author="Otte, D. &amp; Spearman, L. A." journalOrPublisher="Catalog of the mantids of the world" pageId="28" pageNumber="237" part="1" refString="Otte, D. &amp; Spearman, L. A. (2005) Mantida species file. Catalog of the mantids of the world, volume 1. Association of Insect Diversity, Philadelphia, PA." title="Mantida species file" type="journal volume" year="2005">
Otte &amp; Spearman
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:
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</treatmentCitation>
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</paragraph>
<paragraph blockId="28.[106,1074,186,456]" pageId="28" pageNumber="237">
<treatmentCitationGroup pageId="28" pageNumber="237">
<taxonomicName authority="Giglio-Tos, 1915: 45" authorityName="Giglio-Tos" authorityPageNumber="45" authorityYear="1915" box="[106,584,290,317]" class="Insecta" family="Iridopterygidae" genus="Cryptomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="species" species="tenella">
<emphasis box="[106,333,290,317]" italics="true" pageId="28" pageNumber="237">Cryptomantis tenella</emphasis>
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<bibRefCitation author="Giglio-Tos, E." box="[342,541,290,317]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino" pageId="28" pageNumber="237" pagination="1 - 16" part="30" refString="Giglio-Tos, E. (1915) Mantidi esotici. VIII. Acromantinae. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino 30: 1 - 16." title="Mantidi esotici. VIII. Acromantinae" type="journal article" year="1915">
Giglio-Tos,
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:
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;
<treatmentCitation author="Giglio-Tos, E." box="[644,896,290,317]" page="136" pageId="28" pageNumber="237" year="1927">
<bibRefCitation author="Giglio-Tos, E." box="[644,835,290,317]" journalOrPublisher="Walter de Gruyter, Berlin" pageId="28" pageNumber="237" refString="Giglio-Tos, E. (1927) Das Tierreich. Orthoptera-Mantidae. Walter de Gruyter, Berlin." title="Das Tierreich. Orthoptera-Mantidae" type="book" year="1927">
Giglio-Tos
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:
<number box="[849,896,290,316]" pageId="28" pageNumber="237" value="136.0">136</number>
</treatmentCitation>
;
<treatmentCitation author="Roy, R. &amp; Leston, D." page="311" pageId="28" pageNumber="237" year="1975">
<bibRefCitation author="Roy, R. &amp; Leston, D." journalOrPublisher="Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A" pageId="28" pageNumber="237" pagination="297 - 344" part="37" refString="Roy, R. &amp; Leston, D. (1975) Mantodea of Ghana: new species, further records and habitats. Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A 37: 297 - 344." title="Mantodea of Ghana: new species, further records and habitats" type="journal article" year="1975">
Roy &amp; Leston
<number box="[106,168,325,352]" pageId="28" pageNumber="237" value="1975.0">1975</number>
</bibRefCitation>
:
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</treatmentCitation>
;
<bibRefCitation author="Ehrmann, R." box="[296,473,325,351]" journalOrPublisher="Munster, Natur und Tier Verlag Munster" pageId="28" pageNumber="237" refString="Ehrmann, R. (2002) Mantodea: Gottesanberinnen der Welt. Munster, Natur und Tier Verlag Munster." title="Mantodea: Gottesanberinnen der Welt" type="book" year="2002">
Ehrmann
<number box="[411,473,325,351]" pageId="28" pageNumber="237" value="2002.0">2002</number>
</bibRefCitation>
;
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</treatmentCitationGroup>
</paragraph>
<paragraph blockId="28.[106,1074,186,456]" pageId="28" pageNumber="237">
<treatmentCitationGroup pageId="28" pageNumber="237">
<taxonomicName authority="Beier, 1930: 438" authorityName="Beier" authorityPageNumber="438" authorityYear="1930" box="[106,506,360,386]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="species" species="arabica">
<emphasis box="[106,313,360,386]" italics="true" pageId="28" pageNumber="237">Nilomantis arabica</emphasis>
<treatmentCitation author="Beier, M." box="[319,506,360,386]" page="438" pageId="28" pageNumber="237" year="1930">
<bibRefCitation author="Beier, M." box="[319,451,360,386]" journalOrPublisher="The Annals and Magazine of Natural History" pageId="28" pageNumber="237" pagination="432 - 460" part="10" refString="Beier, M. (1930) New and rare Mantodea (Orthoptera) in the British Museum. The Annals and Magazine of Natural History 10: 432 - 460." title="New and rare Mantodea (Orthoptera) in the British Museum" type="journal article" year="1930">
Beier,
<number box="[389,451,360,386]" pageId="28" pageNumber="237" value="1930.0">1930</number>
</bibRefCitation>
:
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</treatmentCitation>
</taxonomicName>
;
<treatmentCitation author="Beier, M." box="[577,745,360,387]" page="55" pageId="28" pageNumber="237" year="1935">
<bibRefCitation author="Beier, M." box="[577,703,360,387]" journalOrPublisher="Wytsman, Brussels" pageId="28" pageNumber="237" refString="Beier, M. (1935) Genera Insectorum, Vol. 201. Mantodea. Subfamilie: Orthoderinae, Choeradodinae, Deroplatinae. Wytsman, Brussels." title="Genera Insectorum, Vol. 201. Mantodea. Subfamilie: Orthoderinae, Choeradodinae, Deroplatinae" type="book" year="1935">
Beier
<number box="[641,703,360,387]" pageId="28" pageNumber="237" value="1935.0">1935</number>
</bibRefCitation>
:
<number box="[713,745,360,387]" pageId="28" pageNumber="237" value="55.0">55</number>
</treatmentCitation>
;
<treatmentCitation author="Uvarov, B. P." box="[756,962,360,387]" page="551" pageId="28" pageNumber="237" year="1939">
<bibRefCitation author="Uvarov, B. P." box="[756,905,360,386]" journalOrPublisher="Journal of the Linnean Society of London, Zoology" pageId="28" pageNumber="237" pagination="547 - 559" part="40" refString="Uvarov, B. P. (1939). Studies in the Arabian Orthoptera. II. New and little known Mantidae and Phasmidae. Journal of the Linnean Society of London, Zoology 40: 547 - 559." title="Studies in the Arabian Orthoptera. II. New and little known Mantidae and Phasmidae" type="journal article" year="1939">
Uvarov
<number box="[843,905,360,386]" pageId="28" pageNumber="237" value="1939.0">1939</number>
</bibRefCitation>
:
<number box="[915,962,360,387]" pageId="28" pageNumber="237" value="551.0">551</number>
</treatmentCitation>
;
<treatmentCitation author="Chopard, L." page="132" pageId="28" pageNumber="237" year="1950">
<bibRefCitation author="Chopard, L." journalOrPublisher="Memoire de l'Institut Francais d'Afrique Noire" pageId="28" pageNumber="237" pagination="127 - 146" part="10" refString="Chopard, L. (1950) Contribution a l'etude de l'Air. Orthoptera: Mantodea. Memoire de l'Institut Francais d'Afrique Noire 10: 127 - 146." title="Contribution a l'etude de l'Air. Orthoptera: Mantodea" type="journal article" year="1950">
Chopard
<number box="[106,168,394,421]" pageId="28" pageNumber="237" value="1950.0">1950</number>
</bibRefCitation>
:
<number box="[179,226,395,421]" pageId="28" pageNumber="237" value="132.0">132</number>
</treatmentCitation>
;
<treatmentCitation author="Roy, R. &amp; Leston, D." box="[238,522,394,421]" page="311" pageId="28" pageNumber="237" year="1975">
<bibRefCitation author="Roy, R. &amp; Leston, D." box="[238,467,394,421]" journalOrPublisher="Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A" pageId="28" pageNumber="237" pagination="297 - 344" part="37" refString="Roy, R. &amp; Leston, D. (1975) Mantodea of Ghana: new species, further records and habitats. Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A 37: 297 - 344." title="Mantodea of Ghana: new species, further records and habitats" type="journal article" year="1975">
Roy &amp; Leston
<number box="[405,467,394,421]" pageId="28" pageNumber="237" value="1975.0">1975</number>
</bibRefCitation>
:
<number box="[478,584,394,420]" pageId="28" pageNumber="237" value="313.5" valueMax="316.0" valueMin="311.0">311316</number>
</treatmentCitation>
;
<treatmentCitation author="Ehrmann, R." box="[596,828,394,421]" page="242" pageId="28" pageNumber="237" year="2002">
<bibRefCitation author="Ehrmann, R." box="[596,772,394,421]" journalOrPublisher="Munster, Natur und Tier Verlag Munster" pageId="28" pageNumber="237" refString="Ehrmann, R. (2002) Mantodea: Gottesanberinnen der Welt. Munster, Natur und Tier Verlag Munster." title="Mantodea: Gottesanberinnen der Welt" type="book" year="2002">
Ehrmann
<number box="[710,772,395,421]" pageId="28" pageNumber="237" value="2002.0">2002</number>
</bibRefCitation>
:
<number box="[784,828,395,421]" pageId="28" pageNumber="237" value="242.0">242</number>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="28.[106,1074,186,456]" box="[106,1072,429,456]" pageId="28" pageNumber="237">
<treatmentCitationGroup box="[106,1072,429,456]" pageId="28" pageNumber="237">
<emphasis box="[106,459,429,456]" italics="true" pageId="28" pageNumber="237">
<taxonomicName box="[106,222,429,455]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="genus">Nilomantis</taxonomicName>
[
<taxonomicName box="[238,376,430,456]" class="Insecta" family="Iridopterygidae" genus="Cryptomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="genus">Cryptomantis</taxonomicName>
]
<taxonomicName authority="(Giglio-Tos, 1915) Beier, 1935: 55" authorityName="(Giglio-Tos, 1915) Beier" authorityPageNumber="55" authorityYear="1935" box="[392,840,429,456]" class="Insecta" family="Iridopterygidae" genus="Cryptomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="species" species="tenella">
tenella (
<bibRefCitation author="Giglio-Tos, E." box="[473,659,429,456]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino" pageId="28" pageNumber="237" pagination="1 - 16" part="30" refString="Giglio-Tos, E. (1915) Mantidi esotici. VIII. Acromantinae. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino 30: 1 - 16." title="Mantidi esotici. VIII. Acromantinae" type="journal article" year="1915">
Giglio-Tos,
<number box="[602,659,429,456]" pageId="28" pageNumber="237" value="1915.0">1915</number>
</bibRefCitation>
)
<treatmentCitation author="Beier, M." box="[674,840,429,456]" page="55" pageId="28" pageNumber="237" year="1935">
<bibRefCitation author="Beier, M." box="[674,800,429,456]" journalOrPublisher="Wytsman, Brussels" pageId="28" pageNumber="237" refString="Beier, M. (1935) Genera Insectorum, Vol. 201. Mantodea. Subfamilie: Orthoderinae, Choeradodinae, Deroplatinae. Wytsman, Brussels." title="Genera Insectorum, Vol. 201. Mantodea. Subfamilie: Orthoderinae, Choeradodinae, Deroplatinae" type="book" year="1935">
Beier,
<number box="[740,800,429,456]" pageId="28" pageNumber="237" value="1935.0">1935</number>
</bibRefCitation>
:
<number box="[810,840,429,456]" pageId="28" pageNumber="237" value="55.0">55</number>
</treatmentCitation>
</taxonomicName>
</emphasis>
;
<treatmentCitation author="Ehrmann, R." box="[851,1072,429,455]" page="242" pageId="28" pageNumber="237" year="2002">
<bibRefCitation author="Ehrmann, R." box="[851,1021,429,455]" journalOrPublisher="Munster, Natur und Tier Verlag Munster" pageId="28" pageNumber="237" refString="Ehrmann, R. (2002) Mantodea: Gottesanberinnen der Welt. Munster, Natur und Tier Verlag Munster." title="Mantodea: Gottesanberinnen der Welt" type="book" year="2002">
Ehrmann
<number box="[961,1021,429,455]" pageId="28" pageNumber="237" value="2002.0">2002</number>
</bibRefCitation>
:
<number box="[1030,1072,429,455]" pageId="28" pageNumber="237" value="242.0">242</number>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="237" type="materials_examined">
<paragraph blockId="28.[106,326,504,530]" box="[106,326,504,530]" pageId="28" pageNumber="237">
<heading box="[106,326,504,530]" fontSize="11" level="2" pageId="28" pageNumber="237" reason="1">
<emphasis box="[106,326,504,530]" italics="true" pageId="28" pageNumber="237">Examined specimens</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[106,1074,556,825]" pageId="28" pageNumber="237">
The list consists of
<specimenCount box="[321,462,556,582]" pageId="28" pageNumber="237" type="generic">
<number box="[321,336,556,582]" pageId="28" pageNumber="237" value="4.0">4</number>
specimens
</specimenCount>
.
<collectingCountry box="[473,621,556,582]" name="Saudi Arabia" pageId="28" pageNumber="237">Saudi Arabia</collectingCountry>
:
<emphasis box="[632,895,556,582]" italics="true" pageId="28" pageNumber="237">Escarpement de Raydah,</emphasis>
<number box="[902,917,556,582]" pageId="28" pageNumber="237" value="1.0">1</number>
♀, Formation arborée à Ficus et Olea,
<number box="[407,436,590,616]" pageId="28" pageNumber="237" value="18.0">18</number>
°
<number box="[447,474,590,616]" pageId="28" pageNumber="237" value="11.0">11</number>
<number box="[479,511,590,616]" pageId="28" pageNumber="237" value="67.0">67</number>
N -
<number box="[583,612,591,617]" pageId="28" pageNumber="237" value="42.0">42</number>
<emphasis box="[613,625,591,617]" italics="true" pageId="28" pageNumber="237">°</emphasis>
<number box="[624,655,591,617]" pageId="28" pageNumber="237" value="23.0">23</number>
<number box="[660,690,591,617]" pageId="28" pageNumber="237" value="33.0">33</number>
E,
<quantity box="[741,825,591,617]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="28" pageNumber="237" unit="m" value="1920.0">1920m</quantity>
,
<number box="[843,873,591,617]" pageId="28" pageNumber="237" value="29.0">29</number>
/
<date box="[882,1022,591,617]" pageId="28" pageNumber="237" value="1992-06-30">
<number box="[882,912,591,617]" pageId="28" pageNumber="237" value="30.0">30</number>
-VI-
<number box="[962,1022,591,617]" pageId="28" pageNumber="237" value="1992.0">1992</number>
</date>
, M. Donskoff reç., Genitalia No. 0 0 0
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0
<number box="[493,503,625,651]" pageId="28" pageNumber="237" value="1.0">1</number>
S
</geoCoordinate>
. K. Brannoch (
<collectionCode box="[695,796,625,651]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="237">MNHN</collectionCode>
);
<number box="[815,830,625,651]" pageId="28" pageNumber="237" value="1.0">1</number>
♀, Formation arborée à Ficus et Olea,
<number box="[283,312,660,686]" pageId="28" pageNumber="237" value="18.0">18</number>
°
<number box="[322,349,660,686]" pageId="28" pageNumber="237" value="11.0">11</number>
<number box="[355,387,660,686]" pageId="28" pageNumber="237" value="67.0">67</number>
N -
<number box="[438,467,660,686]" pageId="28" pageNumber="237" value="42.0">42</number>
<emphasis box="[468,480,660,686]" italics="true" pageId="28" pageNumber="237">°</emphasis>
<number box="[479,510,660,686]" pageId="28" pageNumber="237" value="23.0">23</number>
<number box="[515,545,660,686]" pageId="28" pageNumber="237" value="33.0">33</number>
E,
<quantity box="[582,673,660,686]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="28" pageNumber="237" unit="m" value="1920.0">1920 m</quantity>
,
<number box="[684,714,660,686]" pageId="28" pageNumber="237" value="29.0">29</number>
/
<date box="[723,862,660,686]" pageId="28" pageNumber="237" value="1992-06-30">
<number box="[723,753,660,686]" pageId="28" pageNumber="237" value="30.0">30</number>
-VI-
<number box="[802,862,660,686]" pageId="28" pageNumber="237" value="1992.0">1992</number>
</date>
, M. Donskoff reç., Genitalia No. 0 0 0
<geoCoordinate box="[318,375,695,721]" direction="south" orientation="latitude" pageId="28" pageNumber="237" precision="925" value="-0.033333335">
0
<number box="[333,347,695,721]" pageId="28" pageNumber="237" value="2.0">2</number>
S
</geoCoordinate>
. K. Brannoch (
<collectionCode box="[558,659,695,721]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="237">MNHN</collectionCode>
);
<specimenCount box="[682,719,695,721]" pageId="28" pageNumber="237" type="generic">
<number box="[682,697,695,721]" pageId="28" pageNumber="237" value="1.0">1</number>
</specimenCount>
, Formation arborée à Ficus et Olea,
<number box="[173,202,729,755]" pageId="28" pageNumber="237" value="18.0">18</number>
°
<number box="[212,239,729,755]" pageId="28" pageNumber="237" value="11.0">11</number>
<number box="[244,276,729,755]" pageId="28" pageNumber="237" value="67.0">67</number>
N -
<number box="[326,355,729,755]" pageId="28" pageNumber="237" value="42.0">42</number>
<emphasis box="[356,368,729,755]" italics="true" pageId="28" pageNumber="237">°</emphasis>
<number box="[367,398,729,755]" pageId="28" pageNumber="237" value="23.0">23</number>
<number box="[403,433,729,755]" pageId="28" pageNumber="237" value="33.0">33</number>
E,
<quantity box="[469,553,729,755]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="28" pageNumber="237" unit="m" value="1920.0">1920m</quantity>
,
<number box="[564,594,729,755]" pageId="28" pageNumber="237" value="29.0">29</number>
/
<date box="[603,743,729,755]" pageId="28" pageNumber="237" value="1992-06-30">
<number box="[603,633,729,755]" pageId="28" pageNumber="237" value="30.0">30</number>
-VI-
<number box="[683,743,729,755]" pageId="28" pageNumber="237" value="1992.0">1992</number>
</date>
, M. Donskoff reç. (
<collectionCode box="[959,1060,729,755]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="237">MNHN</collectionCode>
);
<emphasis box="[106,262,764,790]" italics="true" pageId="28" pageNumber="237">Wadi Turabah</emphasis>
,
<specimenCount box="[276,313,764,791]" pageId="28" pageNumber="237" type="generic">
<number box="[276,291,764,790]" pageId="28" pageNumber="237" value="1.0">1</number>
</specimenCount>
, milieu de vallée,
<number box="[517,548,764,790]" pageId="28" pageNumber="237" value="20.0">20</number>
°
<number box="[557,587,764,790]" pageId="28" pageNumber="237" value="27.0">27</number>
<number box="[592,620,764,790]" pageId="28" pageNumber="237" value="12.0">12</number>
N -
<number box="[667,697,764,790]" pageId="28" pageNumber="237" value="41.0">41</number>
°
<number box="[707,737,764,790]" pageId="28" pageNumber="237" value="10.0">10</number>
<number box="[741,770,764,790]" pageId="28" pageNumber="237" value="61.0">61</number>
E,
<quantity box="[809,901,764,790]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="28" pageNumber="237" unit="m" value="1330.0">1330 m</quantity>
,
<date box="[913,1068,764,790]" pageId="28" pageNumber="237" value="1992-07-03">
03-VII-
<number box="[1007,1068,764,790]" pageId="28" pageNumber="237" value="1992.0">1992</number>
</date>
, M., Donskoff reç., Genitalia No. 0 0 0
<geoCoordinate box="[530,582,799,825]" direction="south" orientation="latitude" pageId="28" pageNumber="237" precision="925" value="-0.05">
0
<number box="[545,558,799,825]" pageId="28" pageNumber="237" value="3.0">3</number>
S
</geoCoordinate>
. K. Brannoch (
<collectionCode box="[756,854,799,825]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="237">MNHN</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="29" lastPageNumber="238" pageId="28" pageNumber="237" type="description">
<paragraph blockId="28.[106,386,873,899]" box="[106,386,873,899]" pageId="28" pageNumber="237">
<heading box="[106,386,873,899]" fontSize="11" level="2" pageId="28" pageNumber="237" reason="1">
<emphasis box="[106,386,873,899]" italics="true" pageId="28" pageNumber="237">Male genitalia description</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[106,1074,925,1195]" pageId="28" pageNumber="237">
Based on an examination of the genital complex of
<taxonomicName box="[708,820,925,951]" class="Insecta" family="Iridopterygidae" genus="Nilomantis" kingdom="Animalia" order="Mantodea" pageId="28" pageNumber="237" phylum="Arthropoda" rank="species" species="floweri">
<emphasis box="[708,820,925,951]" italics="true" pageId="28" pageNumber="237">N. floweri</emphasis>
</taxonomicName>
, as well as plates presented by
<bibRefCitation author="Roy, R. &amp; Leston, D." box="[221,470,960,987]" journalOrPublisher="Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A" pageId="28" pageNumber="237" pagination="297 - 344" part="37" refString="Roy, R. &amp; Leston, D. (1975) Mantodea of Ghana: new species, further records and habitats. Bulletin de l'Institut Fondamental d'Afrique Noire (IFAN) Serie A 37: 297 - 344." title="Mantodea of Ghana: new species, further records and habitats" type="journal article" year="1975">
Roy &amp; Leston (
<number box="[399,461,960,987]" pageId="28" pageNumber="237" value="1975.0">1975</number>
)
</bibRefCitation>
, the anterior process (ap) is lightly sclerotized, undulate. Apical process (paa) features a broad base with an anterior dilation; paa narrow and recurved distally. Apophysis phalloide (L
<number box="[634,644,1029,1055]" pageId="28" pageNumber="237" value="1.0">1</number>
) does not retain the apophysis; lobo membranoso (loa) heavily sclerotized and strongly crenulated distally. Processo distale (pda) narrows apically, terminating in a moderately sclerotized, blunted point. There is a second, triangular lobe to the right of pda. Processo ventrale sclerificato (pva) and piastra ventrale (pia) reduced, heavily sclerotized, and closely adjoined.
</paragraph>
<paragraph blockId="28.[106,1074,1242,1650]" lastBlockId="29.[98,1066,134,404]" lastPageId="29" lastPageNumber="238" pageId="28" pageNumber="237">
<emphasis box="[106,406,1242,1268]" italics="true" pageId="28" pageNumber="237">Female genitalia description</emphasis>
(
<figureCitation box="[420,488,1242,1269]" captionStart="Fig. 4" captionStartId="8.[106,139,1540,1562]" captionTargetBox="[155,1024,390,1513]" captionTargetId="figure@8.[155,1024,390,1515]" captionTargetPageId="8" captionText="Fig. 4. Female genitalic complex of Nilomantis shown in the lateral perspective. (A) Nilomantis floweri; (B) Nilomantis edmundsi. Abbreviations: KLP, knob-like projection; RD, round dilation. This figure is published in colour in the online edition of this journal, which can be accessed via http: // booksand journals. brillonline. com / content / journals / 1876312 x." httpUri="https://zenodo.org/record/269843/files/figure.png" pageId="28" pageNumber="237">
Figs
<number box="[474,488,1243,1269]" pageId="28" pageNumber="237" value="4.0">4</number>
</figureCitation>
A,
<number box="[521,536,1242,1269]" pageId="28" pageNumber="237" value="6.0">6</number>
A,B). Gonapophysis VIII (GA) slender, variably ciliated, and visibly curved from both the ventral (VH) and lateral (LH) perspectives; GA lightly surfaced with fossettes. From VH, gonocoxa VIII (VF) internal margin rounded; from LH, VF does not retain a projection. From VH, GA curves strongly inward; apical lobe (AL) ovoid, ciliated. AL has a small membrane which projects from the internal dorsal margin of the structure, forming a shallow pocket as the membrane merges with the dorsal margin of GA. From LH, GA is gently curved ventrad; GA ventral margin features a pronounced medial outgrowth (MO), which projects posterior to the crest of the curve, with a row of cilia that runs from the crest of MO towards AL. GA external margin possesses a moderately sclerotized tubercle anterior to MO. From LH, AL narrows posteriorly with a bluntly rounded apical margin. GA dorsal margin is invaginated anterior to the curve towards the ventral habitus. GA very lightly sclerotized. Gonapophysis IX (GP) relatively short, broadening at the site of the medial tine (MT) before narrowing into a tapered, acuminate apex; GP ventral margin invaginated, covered anteriorly by MT. Gonoplac (GL) shorter than GA, broad, variably ciliated, and arched ventrad; from LH, GL base has a rounded dilation on the anteroventral margin. GL narrows toward the apex; from LH, GL apex slants ventrad. GL features a reduced apical cleft (AC) resulting in a slight apical notch. From LH, GL possesses a slight sclerotization that traverses the length of the structure, terminating just before the AC. GL ventral margin rugose, dorsal margin relatively smooth.
</paragraph>
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<mods:title>A revision and key to the genera of Afrotropical Mantispidae (Neuropterida, Neuroptera), with the description of a new genus</mods:title>
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<mods:namePart>Ohl, Michael</mods:namePart>
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<mods:namePart>Mansell, Mervyn W.</mods:namePart>
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<treatment ID-GBIF-Taxon="152034737" LSID="urn:lsid:zoobank.org:act:23AFF0D1-4D05-4E15-8B91-79F32AE4A9D0" httpUri="http://treatment.plazi.org/id/4A09CDEB7E2F3ECABBC9F9F8D42E6CAB" lastPageId="7" lastPageNumber="74" pageId="5" pageNumber="72">
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Genus
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<taxonomicNameLabel pageId="5" pageNumber="72">gen. n.</taxonomicNameLabel>
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<paragraph pageId="5" pageNumber="72">
<taxonomicName class="Insecta" family="Mantispidae" genus="Afromantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afromantispa" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Afromantispa</taxonomicName>
Snyman and Ohl gen. n. Type species:
<taxonomicName class="Insecta" family="Mantispidae" genus="Afromantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afromantispa tenella" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="tenella">Afromantispa tenella</taxonomicName>
(Erichson, 1839: 169) comb. n., designated here.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="72" type="remarks">
<paragraph pageId="5" pageNumber="72">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="72">
Both
<bibRefCitation author="Handschin, E" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="16" pageNumber="83" pagination="185 - 227" title="Beitraege zu einer Revision der Mantispiden (Neuroptera). I Teil. Mantispiden des Musee Royal du Congo Belge, Tervuren." volume="59" year="1959">Handschin (1959</bibRefCitation>
,
<bibRefCitation author="Handschin, E" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="16" pageNumber="83" pagination="181 - 245" title="Beitraege zu einer Revision der Mantispiden (Neuroptera). II Teil. Mantispiden des &quot; Musee Royal du Congo Belge &quot;, Tervuren." volume="62" year="1960 a">1960a</bibRefCitation>
) and
<bibRefCitation author="Poivre, C" journalOrPublisher="Neuroptera International" pageId="17" pageNumber="84" pagination="77 - 89" title="Nouvelle description et morphologie externe comparee de Mantispa nana Erichson (Planipennia, Mantispidae)." volume="1" year="1980">Poivre (1980</bibRefCitation>
,
<bibRefCitation author="Poivre, C" journalOrPublisher="Neuroptera International" pageId="17" pageNumber="84" pagination="3 - 25" title="Mantispides nouveaux d'Afrique et d'Europe (Neuroptera, Planipennia) (seconde partie)." volume="2" year="1982 b">1982b</bibRefCitation>
) considered the Afrotropical
<taxonomicName family="Mantispidae" lsidName="" pageId="5" pageNumber="72" rank="family">Mantispidae</taxonomicName>
to be part of
<taxonomicName class="Insecta" family="Mantispidae" genus="Mantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mantispa" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Mantispa</taxonomicName>
without comparing the Afrotropical species to the type species of
<taxonomicName class="Insecta" family="Mantispidae" genus="Mantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mantispa" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Mantispa</taxonomicName>
(
<taxonomicName class="Insecta" family="Mantidae" genus="Mantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mantis pagana" order="Mantodea" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="pagana">Mantis pagana</taxonomicName>
Fabricius, 1775 (=
<taxonomicName class="Insecta" family="Mantispidae" genus="Mantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mantispa styriaca" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="styriaca">Mantispa styriaca</taxonomicName>
Poda, 1761)). The new genus
<taxonomicName class="Insecta" family="Mantispidae" genus="Afromantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afromantispa" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Afromantispa</taxonomicName>
can easily be identified and clearly differs morphologically from the Palaearctic
<taxonomicName class="Insecta" family="Mantispidae" genus="Mantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mantispa" order="Neuroptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Mantispa</taxonomicName>
although the two genera are possibly closely related. (see discussion below). Species complexes within this genus must be investigated, with at least one species with a distribution from South Africa to the Arabian Peninsula lacking morphological variation. Sexual dimorphism has not been formally investigated, but it appears to be absent in the genus.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="72" type="distribution">
<paragraph pageId="5" pageNumber="72">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="72">Widespread throughout Africa. A few species have been collected in the Palaearctic Region sharing borders with Africa such as Spain and the Arabian Peninsula.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="73" pageId="5" pageNumber="72" type="diagnosis">
<paragraph pageId="5" pageNumber="72">Diagnosis.</paragraph>
<paragraph lastPageId="6" lastPageNumber="73" pageId="5" pageNumber="72">
Prothorax granulated; granules dark (Fig. 3e). Antennae with distinct yellowish white band in the apical third (Fig. 4a). Even species with pale antennae have a few darker flagellomeres two-thirds apically from the base of the antennae and on the apex to form a yellowish-white band. The crossvein between Cua and Cup+Aa in hind
<pageBreakToken pageId="6" pageNumber="73" start="start">wing</pageBreakToken>
attenuated or absent (Fig. 2b). These characteristics combined are unique to this genus and can be used to distinguish
<taxonomicName class="Insecta" family="Mantispidae" genus="Afromantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afromantispa" order="Neuroptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Afromantispa</taxonomicName>
species from all other genera.
</paragraph>
<caption pageId="6" pageNumber="73">
<paragraph pageId="6" pageNumber="73">
Figure 2. Wing variation in the Afrotropical
<taxonomicName lsidName="" pageId="6" pageNumber="73" rank="subFamily" subFamily="Mantispinae">Mantispinae</taxonomicName>
. a
<taxonomicName class="Insecta" family="Mantispidae" genus="Pseudoclimaciella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoclimaciella apicipennis" order="Neuroptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="apicipennis">Pseudoclimaciella apicipennis</taxonomicName>
(Kolbe) b
<taxonomicName class="Insecta" family="Mantispidae" genus="Afromantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afromantispa" order="Neuroptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Afromantispa</taxonomicName>
sp. c
<taxonomicName class="Insecta" family="Mantispidae" genus="Sagittalata" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sagittalata" order="Neuroptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Sagittalata</taxonomicName>
sp. d
<taxonomicName class="Insecta" family="Mantispidae" genus="Cercomantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercomantispa" order="Neuroptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Cercomantispa</taxonomicName>
sp. e
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(
<normalizedToken originalValue="Navás">Navas</normalizedToken>
) f
<taxonomicName class="Insecta" family="Mantispidae" genus="Rectinerva" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rectinerva braconidiformis" order="Neuroptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="braconidiformis">Rectinerva braconidiformis</taxonomicName>
Handschin.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73" type="description">
<paragraph pageId="6" pageNumber="73">Description.</paragraph>
<paragraph pageId="6" pageNumber="73">Head: Antennae moniliform; colour variable but all with conspicuous yellowish-white band in apical third; scape and pedicel yellow. Posterior vertex concave except for slight convex elevation directly posterior to and between antennal bases; median tubercle projection at posterior margin of vertex, vertex not visible in lateral view. Compound eyes large, each eye slightly broader medially at epistomal suture. Labrum circular. Mandible with dark apices; inner margins dark.</paragraph>
<paragraph lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
Thorax: Pronotum narrow and elongated; prothorax longer than pterothorax; granulated; granules dark; pronotum transversely slightly wrinkled or rugulose; setae
<pageBreakToken pageId="7" pageNumber="74" start="start">present</pageBreakToken>
. Maculae slightly raised and inconspicuous; not pigmented in lighter coloured species, pigmented and shiny in darker species. Prozona slightly broader than base. Meso- and metathorax of similar size and distinctly separated by a deep cleft.
</paragraph>
<paragraph pageId="7" pageNumber="74">Wings (Fig. 2b): Wings always hyaline, lacking pigment except for the pterostigma. Pterostigma slightly concave in dorsal view; semi-circular and truncate appearance; pterostigma of most species with reddish appearance. Radial cell 1 and 2 of similar size with radial cell 3 smaller and narrower; a single crossvein from third radial cell to anterior margin (C). Hind wings: Crossvein between Cua and Cup+Aa attenuated or absent; Cua with sharp angle to and from attenuated crossvein to form inverted triangle.</paragraph>
<paragraph pageId="7" pageNumber="74">Legs: Median line on the anterior surface of the forecoxae never continuous from thorax to femur (Fig. 4a). Mid- and hind legs differ considerably between species. Fore tarsal claw reduced to a single claw lacking arolium; Mid- and hind pretarsal claws pectinate (5 -6 teeth) with the middle tooth elongated giving the claw a sharp triangular appearance; arolium present on mid and hind tarsi.</paragraph>
<paragraph pageId="7" pageNumber="74">Abdomen/Genitalia: Males with ectoprocts slightly enlarged (Fig. 4j). Pseudopenis visible in lateral and dorsal view. EEG present. No morphological significance regarding the female genitalia.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="74" type="etymology">
<paragraph pageId="7" pageNumber="74">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="74">The new genus name is a combination of Afro- and Mantispa, which emphasises the African distribution of this Mantispa-like taxon.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="74" type="included species">
<paragraph pageId="7" pageNumber="74">Included species.</paragraph>
<paragraph pageId="7" pageNumber="74">
Besides the type species, 18 confirmed and 7 unconfirmed species names will be added in the future. These numbers are, however, certain to change. Synonyms need to be identified and new species described. A subsequent full revision of
<taxonomicName class="Insecta" family="Mantispidae" genus="Afromantispa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afromantispa" order="Neuroptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="genus">Afromantispa</taxonomicName>
is currently in progress. The genus for the time being will therefore be based on the type species only.
</paragraph>
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<paragraph pageId="0" pageNumber="133">
<taxonomicName ID-CoL="78YL6" LSID="4A09F31A-CC4C-5B70-BE14-139EDB4D472D" authority="(Saalmueller, 1884)" baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Metarbelidae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
Saalmulleria stumpffi (
<normalizedToken originalValue="Saalmüller">Saalmueller</normalizedToken>
, 1884)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a. Morondavania mineti sp. nov., holotype, male, Madagascar, Western Region, north of Morondava, Marofandilia Forest; b. M. mineti sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest; c. Eberhardfischeria husemanni sp. nov., holotype, female, Madagascar, Western Region, Diego Suarez (today Antsira ṅana) or areas nearby; d. E. husemanni sp. nov., paratype, female, Madagascar, Western Region, Diego Suarez (today Antsira ṅana) or areas nearby; e. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female, Madagascar, Sambirano Region, Nosy Be Island, Lokobe; f. S. analameranaensis sp. nov., holotype, female, Madagascar, Western Region, Analamerana Forest, ca. 10 km to 40 km west of the Indian Ocean coastline; g. S. ampandrandavaensis sp. nov., holotype, female, Madagascar, Central Region, Ampandrandava, ca. 50 km northeast of Bekily." httpUri="https://binary.pensoft.net/fig/845634" pageId="0" pageNumber="133">Figs 4e</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing venation: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Morondavania mineti sp. nov., holotype, male; c. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female." httpUri="https://binary.pensoft.net/fig/845635" pageId="0" pageNumber="133">, 5c</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female postabdominal structures and male genitalia in a not pressed condition: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female; c. Shimbania kerstinhempae sp. nov., holotype, female; d. Morondavania mineti sp. nov., paratype, male, with aedeagus below; e. S. wichgrafi (Gruenberg, 1910), &quot; Type &quot;, male, with aedeagus below (all drawings by I. L.)." httpUri="https://binary.pensoft.net/fig/845637" pageId="0" pageNumber="133">, 7b</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female postabdominal structures of a. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female, Madagascar, Sambirano Region, Nosy Be Island, Lokobe; b. S. analameranaensis sp. nov., holotype, female, Madagascar, Western Region, Analamerana Forest, ca. 10 km to 40 km west of the Indian Ocean coastline." httpUri="https://binary.pensoft.net/fig/845643" pageId="0" pageNumber="133">, 14a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="reference_group">
<paragraph pageId="0" pageNumber="133">
<taxonomicName authorityName="Saalmuller" authorityYear="1884" class="Insecta" family="Metarbelidae" genus="Cossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cossus stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">Cossus stumpffi</taxonomicName>
<normalizedToken originalValue="Saalmüller">Saalmueller</normalizedToken>
, 1884: In:
<normalizedToken originalValue="Saalmüller">Saalmueller</normalizedToken>
, M. (Ed.) Lepidopteren von Madagaskar. Neue und wenig bekannte Arten. Zumeist aus der Sammlung der
<normalizedToken originalValue="Senckenbergschen">Senckenberg'schen</normalizedToken>
naturforschenden Gesellschaft zu Frankfurt am Main. Erste Abtheilung:
<taxonomicName higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" rank="subOrder" subOrder="Rhopalocera">Rhopalocera</taxonomicName>
.
<taxonomicName higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" rank="subOrder" subOrder="Heterocera">Heterocera</taxonomicName>
. Shinges et Bombyces.
<taxonomicName family="Cossidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" rank="family">Cossidae</taxonomicName>
: 210-211. Original combination.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="materials_examined">
<paragraph pageId="0" pageNumber="133">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="133">
<materialsCitation collectingDate="1882-02-10" collectorName="I. Lehmann" country="Madagascar" location="Female" specimenCount="one" typeStatus="Type">
<location LSID="urn:lsid:plazi:treatment:4A09F31ACC4C5B70BE14139EDB4D472D:7B2307C6F938D05AB93A58CB0579AA76" country="Madagascar" name="Female">Female</location>
,
<normalizedToken originalValue="“Type”">
&quot;
<typeStatus>Type</typeStatus>
&quot;
</normalizedToken>
,
<normalizedToken originalValue="“Madag.”">&quot;Madag.&quot;</normalizedToken>
[
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
],
<normalizedToken originalValue="“Loucoubé”">&quot;Loucoube&quot;</normalizedToken>
[Lokobe, Nosy Be Island, Sambirano Region], &quot;Stumpff 82&quot; [1882],
<normalizedToken originalValue="“X-II-82”">&quot;X-II-82&quot;</normalizedToken>
[on another tiny label,
<collectingDate value="1882-02-10">10th February 1882</collectingDate>
] Stumpff [Anton Stumpff from
<normalizedToken originalValue="“Nossi-Bé”">&quot;Nossi-Be&quot;</normalizedToken>
leg.;
<specimenCount type="generic">one specimen</specimenCount>
only,
<emphasis italics="true" pageId="0" pageNumber="133">cf.</emphasis>
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8759" author="Saalmueller, M" editor="Saalmueller, M" journalOrPublisher="Neue und wenig bekannte Arten. Zumeist aus der Sammlung der Senckenberg'schen naturforschenden Gesellschaft zu Frankfurt am Main. Erste Abtheilung: Rhopalocera. Heterocera. Shinges et Bombyces. Cossidae" pageId="0" pageNumber="133" pagination="210 - 211" refId="B91" refString="Saalmueller, M, 1884. Cossus stumpffi n. sp. In: Saalmueller, M, Ed., Lepidopteren von Madagaskar. Neue und wenig bekannte Arten. Zumeist aus der Sammlung der Senckenberg'schen naturforschenden Gesellschaft zu Frankfurt am Main. Erste Abtheilung: Rhopalocera. Heterocera. Shinges et Bombyces. Cossidae: 210 - 211, DOI: https://doi.org/10.5962/bhl.title.8759" title="Cossus stumpffi n. sp." url="https://doi.org/10.5962/bhl.title.8759" volumeTitle="Lepidopteren von Madagaskar." year="1884">
<normalizedToken originalValue="Saalmüller">Saalmueller</normalizedToken>
1884
</bibRefCitation>
, pp. 210-211]; genitalia slide number 20/122008
<collectorName>I. Lehmann</collectorName>
(SNMF)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">Re-description.</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" pageId="0" pageNumber="133">
Female
<emphasis italics="true" pageId="0" pageNumber="133">Head</emphasis>
</emphasis>
: Unusually small, 3.0 mm in diameter; rough-scaled; long hair-like scales of light brownish-olive mixed with sepia scales on fronto-clypeus, some with a light lilac and golden glint; eyes brownish-olive with small dark olive spots; a pair of pits is entirely absent on lower fronto-clypeus, a pair of conical projections absent, pits behind labial palpi absent; lower fronto-clypeus is broad, as broad as half of eye-diameter (viewed anteriorly) and smooth, without any structures; labial palpi dark olive-buff, very short, less than half of eye diameter, narrow, consisting of three segments, all segments are of almost equal length, 1st (basal) segment is very broad, at least 1.5
<normalizedToken originalValue="×">x</normalizedToken>
broader than central segment, rectangular. Antennae bipectinate, narrow and long branches 3.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of shaft, branches are widely separated at base with 2.0
<normalizedToken originalValue="×">x</normalizedToken>
width of branch, dorsal and lateral sides of branches not scaled, but with many setae in pairs ventrally and laterally, dorsal and lateral sides of flagellum scaled deep olive-buff mixed with brownish-olive.
</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Thorax</emphasis>
: Densely covered with hair-like scales and broader scales of olive-brown with dark olive or sepia tips on patagia and on tegulae, sepia scales on tegulae with a light lilac glint; scale crest on metathorax is pronounced with long olive-brown hair-like scales with a slightly broader sepia tip. Fore and mid legs olive-brown, dorsally sepia, with long dense hair-like structures and a light lilac glint. Epiphyses present, 2.5 mm long, medium broad and flat. Hind legs with two pairs of tibial spurs, upper pair narrower and longer, up to 2.1 mm long, spurs in lower pair broader, up to 1.4 mm long, all spurs with a claw-like tip. Wingspan is 70.5 mm. Forewing length is 32.0 mm (as long as abdomen), large, broad with a rounded apex, upperside without any geometric design, deep olive-buff and towards termen with a light golden glint, a simple scale pattern is present, with a lunule-like sepia patch at center of forewing from base of M2 to base of CuA1, the patch is edged inwards by a small transparent spot, various terminal, sub-terminal and post-medial patches and bands of
<normalizedToken originalValue="Saccardos">Saccardo's</normalizedToken>
umber, with a prominent broad V-shaped band of
<normalizedToken originalValue="Saccardos">Saccardo's</normalizedToken>
umber from apex and costa to near end of CuA2, the latter vein is not marked in a different colour, termen with weak triangular dark olive lunules, a sepia patch is present below 30% of length of 1A+2A. Hindwing rounded but with a prominent pointed apex, termen not bent inwards, largely with short scales of deep olive-buff with a light golden glint, with a sepia patch at center of hindwing from base of M2 to base of CuA1, but the patch is not edged inwards by a small transparent spot as in forewing. Underside with scales of deep olive-buff with a light golden glint. Cilia very short for such a large species, 1.1 mm long, deep olive-buff with a glint. Forewing venation with strongly sclerotized and broad veins, 1A+2A deeply forked at base, fork is 25% the length of 1A+2A; CuP absent, but represented by a fold, slightly sclerotized on first 2/3 of its length; CuA2 originating from near hind margin of posterior cell; CuA1, M3 and M2 separated and initiating from apical angle of posterior cell; M1 originating from distal margin of median cell and is broadly separated from its anterior angle; areole absent; R1+R2 originating from a long stalk (the stalk has the length of
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
40% of R3) and initiating from near anterior angle of median cell; R3+R4+R5 are long stalked and originating from anterior angle of median cell, the basal point of this stalk is exactly opposite of the basal point of the stalk of R1+R2; Sc more or less parallel to R1. Hindwing venation with three anal veins, 3A present, 1A+2A present with a well sclerotized fork at base, CuP present and well sclerotized; CuA2 originating from near hind margin of posterior cell; CuA1, M3 and M2 originating from apical angle of posterior cell, separated, M2 has a tiny fork at base; M1 and Rs originating from anterior cell, broadly separated, with M1 at center of distal margin of anterior cell; a short bar from Rs to Sc+R1 is weak, a strongly sclerotized vein in discocellular cell on both fore- and hindwing is present and long forked distally in forewing (a very rare character in
<taxonomicName authorityName="Strand" authorityYear="1909" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
). The discocellular cell on forewing and hindwing is small, only
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
15% of wing size. The discocellular cell on the hindwing is similar in shape like a fish-tail, with the upper and lower tip in opposite position, and the upper tip is strongly pointed and bent. Fringe scales short if compared to the large hindwing size, 2.0 mm long, deep olive-buff with a glint. Retinaculum and frenulum absent.
</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Abdomen</emphasis>
: Very long with 32.0 mm in length and very broad at center with 9.0 mm, with hair-like scales of deep olive-buff, some with chestnut tips, and a strong light golden glint, upper part of abdomen broadly sepia, end of abdomen broadly sepia, abdominal tuft short with 5.0 mm length, sepia.
</paragraph>
<paragraph pageId="0" pageNumber="133">
Female postabdominal structure with very large, pear-shaped lobes of papillae anales in horizontal position, one lobe 50% the size of papillae anales, lobes ventrally with long setae and few long setae along the edge, each lobe with a long and deep oblique graben-like structure that has no setae at center; papillae anales covered with many short and many long setae. Segment 8 represents a medium broad rectangular sclerotized band, more narrow ventrally, setose along its posterior margin with long setae but without any setae on dorsal part of posterior margin and on segment 8, a very narrow band is attached ventrally extending to the base of anterior apophysis, where the band is weakly attached and has a very broad rectangular-shaped base that fits like a segment into segment 8 (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female postabdominal structures and male genitalia in a not pressed condition: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female; c. Shimbania kerstinhempae sp. nov., holotype, female; d. Morondavania mineti sp. nov., paratype, male, with aedeagus below; e. S. wichgrafi (Gruenberg, 1910), &quot; Type &quot;, male, with aedeagus below (all drawings by I. L.)." httpUri="https://binary.pensoft.net/fig/845637" pageId="0" pageNumber="133">7b</figureCitation>
, only visible in fresh preparations); anterior apophysis slightly bent downwards, 3.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as segment 8 dorsally, on basal half of length 2
<normalizedToken originalValue="×">x</normalizedToken>
as broad as at tip, within the first 30% of their length strongly knee-like shaped, on almost the whole length with a deep horizontal graben-like structure; posterior apophysis narrow but with four times broader base, 50% the length of anterior apophysis, with large sclerotized base that is 40% the size of papillae anales in lateral view, posterior apophysis equal in length of dorsal part of segment 8; ductus bursae and corpus bursae are unknown.
</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" pageId="0" pageNumber="133">Male</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="diagnosis">
<paragraph pageId="0" pageNumber="133">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="133">
<taxonomicName baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
has by far the largest wing size in
<taxonomicName authorityName="Strand" authorityYear="1909" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
worldwide and its small transparent spot in the forewing near base of M2 to base of CuA1 distinguishes it from all other species of
<taxonomicName authorityName="Mabille" authorityYear="1891" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Saalmulleria</emphasis>
</taxonomicName>
. The size of one lobe of the papillae anales is 50% of the size of the whole papillae anales and is at present the largest among
<taxonomicName authorityName="Strand" authorityYear="1909" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
. Remarkable differences to the other species of
<taxonomicName authorityName="Mabille" authorityYear="1891" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Saalmulleria</emphasis>
</taxonomicName>
comprise also the very long setae along the posterior margin of segment 8 that are absent from its dorsal part, the strongly knee-like shape of the anterior apophyses present on 30% of their length and the very large sclerotized base of posterior apophysis that is 40% the size of the papillae anales.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="distribution">
<paragraph pageId="0" pageNumber="133">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="133">
<taxonomicName baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
is only known from Lokobe, including the
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
Naturelle de Lokobe (740 hectare in size), located on the southeastern part of the small island of Nosy Be (320 km2 in size)
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
14 km off the northwestern coast of the main island of Madagascar. The whole area belongs to the &quot;Sambirano Region&quot; and to the &quot;Madagascar Subhumid Forests&quot; ecoregion.
<taxonomicName baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
is classified herein as an endemic species to the &quot;Sambirano Region&quot; and might be restricted today to Nosy Be Island and scattered forest areas nearby located on the &quot;main island&quot; of northern Madagascar. Nosy Be Island has an altitude range of 5-419 m, average annual rainfall is among the highest along the west coast of Madagascar ranging from 2000-2356 mm at Lokobe, with highest rainfall in November-April (
<bibRefCitation author="White, F" journalOrPublisher="Unesco, Paris" pageId="0" pageNumber="133" refId="B104" refString="White, F, 1983. The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco/AETFAT/UNSO Vegetation Map of Africa. Natural Resources Research no. XX. Unesco, Paris" title="The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco / AETFAT / UNSO Vegetation Map of Africa. Natural Resources Research no. XX." year="1983">White 1983</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1002/joc.1276" author="Hijmans, R J" journalOrPublisher="International Journal for Climatology" pageId="0" pageNumber="133" pagination="1965 - 1978" refId="B32" refString="Hijmans, R J, Cameron, SE, Parra, JL, Jones, PG, Jarvis, A, 2005. Very high resolution interpolated climate surfaces for global land areas. International Journal for Climatology 25 (15): 1965 - 1978, DOI: https://doi.org/10.1002/joc.1276" title="Very high resolution interpolated climate surfaces for global land areas." url="https://doi.org/10.1002/joc.1276" volume="25" year="2005">Hijmans et al. 2005</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3389/fevo.2021.783770" author="Reinhardt, AL" journalOrPublisher="Primate Report, 46 - 1, Special Issue, Berne" pageId="0" pageNumber="133" refId="B87" refString="Reinhardt, AL, Kasper, T, Lochner, M, Bliedtner, M, Krahn, KJ, Haberzettl, T, Shumilovskikh, L, Rahobisoa, J-J, Zech, R, Favier, C, Behling, H, Bremond, L, Daut, G, Montade, V, 2022. Rain forest fragmentation and environmental dynamics on Nosy Be Island (NW Madagascar) at 1300 cal BP is attributable to intensified human impact. Frontiers in Ecology and Evolution 9: 783770. https://doi.org/10.3389/fevo.2021.783770" title="Rain forest fragmentation and environmental dynamics on Nosy Be Island (NW Madagascar) at 1300 cal BP is attributable to intensified human impact. Frontiers in Ecology and Evolution 9: 783770." url="https://doi.org/10.3389/fevo.2021.783770" year="2022">Reinhardt et al. 2022</bibRefCitation>
). Prior to 1300 BP, Nosy Be Island was largely covered by lowland rain forest comprising also genera such as
<taxonomicName class="Dicotyledoneae" family="Leguminosae" genus="Podocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Podocarpus" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Podocarpus</emphasis>
</taxonomicName>
<normalizedToken originalValue="LHér">L'Her</normalizedToken>
. ex Pers., but became increasingly a mosaic of grassland, bushland and forest until present due to human activities including the use of fire to destroy forests for agriculture (
<bibRefCitation DOI="https://doi.org/10.3389/fevo.2021.783770" author="Reinhardt, AL" journalOrPublisher="Primate Report, 46 - 1, Special Issue, Berne" pageId="0" pageNumber="133" refId="B87" refString="Reinhardt, AL, Kasper, T, Lochner, M, Bliedtner, M, Krahn, KJ, Haberzettl, T, Shumilovskikh, L, Rahobisoa, J-J, Zech, R, Favier, C, Behling, H, Bremond, L, Daut, G, Montade, V, 2022. Rain forest fragmentation and environmental dynamics on Nosy Be Island (NW Madagascar) at 1300 cal BP is attributable to intensified human impact. Frontiers in Ecology and Evolution 9: 783770. https://doi.org/10.3389/fevo.2021.783770" title="Rain forest fragmentation and environmental dynamics on Nosy Be Island (NW Madagascar) at 1300 cal BP is attributable to intensified human impact. Frontiers in Ecology and Evolution 9: 783770." url="https://doi.org/10.3389/fevo.2021.783770" year="2022">Reinhardt et al. 2022</bibRefCitation>
). The Lokobe forest comprises
<taxonomicName class="Dicotyledoneae" family="Leguminosae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="family">Leguminosae</taxonomicName>
tree species such as
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Parkia" kingdom="Plantae" lsidName="Parkia madagascariensis" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="133">Parkia madagascariensis</emphasis>
</taxonomicName>
Viguier (
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="subFamily" subFamily="Mimosoideae">Mimosoideae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Adenanthera" kingdom="Plantae" lsidName="Adenanthera pavonina" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="pavonina">
<emphasis italics="true" pageId="0" pageNumber="133">Adenanthera pavonina</emphasis>
</taxonomicName>
L. (
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="subFamily" subFamily="Mimosoideae">Mimosoideae</taxonomicName>
) and
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Cordyla" kingdom="Plantae" lsidName="Cordyla madagascariensis" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="133">Cordyla madagascariensis</emphasis>
</taxonomicName>
R. Vig. (
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="subFamily" subFamily="Papilionoideae">Papilionoideae</taxonomicName>
); other tree species include,
<emphasis italics="true" pageId="0" pageNumber="133">
e.g.
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" lsidName="Croton" order="Malpighiales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">Croton</taxonomicName>
loucoubensis
</emphasis>
Baill. (
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" lsidName="" order="Malpighiales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Sapindaceae" genus="Macphersonia" kingdom="Plantae" lsidName="Macphersonia madagascariensis" order="Sapindales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="133">Macphersonia madagascariensis</emphasis>
</taxonomicName>
Blume (
<taxonomicName class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
), the extremely rare
<taxonomicName class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" lsidName="Pandanus androcephalanthos" order="Pandanales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="androcephalanthos">
<emphasis italics="true" pageId="0" pageNumber="133">Pandanus androcephalanthos</emphasis>
</taxonomicName>
Martelli (
<taxonomicName class="Liliopsida" family="Pandanaceae" kingdom="Plantae" lsidName="" order="Pandanales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="family">Pandanaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Anacardiaceae" genus="Sorindeia" kingdom="Plantae" lsidName="Sorindeia madagascariensis" order="Sapindales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="133">Sorindeia madagascariensis</emphasis>
</taxonomicName>
Thouars (
<taxonomicName authorityName="Du Petit-Thouars" authorityYear="1806" class="Magnoliopsida" family="Anacardiaceae" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="family">Anacardiaceae</taxonomicName>
), one species of
<taxonomicName class="Magnoliopsida" family="Monimiaceae" genus="Tambourissa" kingdom="Plantae" lsidName="Tambourissa" order="Laurales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Tambourissa</emphasis>
</taxonomicName>
Sonn. (
<taxonomicName class="Magnoliopsida" family="Monimiaceae" kingdom="Plantae" lsidName="" order="Laurales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="family">Monimiaceae</taxonomicName>
) and
<taxonomicName class="Magnoliopsida" family="Moraceae" genus="Trilepisium" kingdom="Plantae" lsidName="Trilepisium madagascariense" order="Rosales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="madagascariense">
<emphasis italics="true" pageId="0" pageNumber="133">Trilepisium madagascariense</emphasis>
</taxonomicName>
DC. (
<taxonomicName authorityName="Du Petit-Thouars" authorityYear="1806" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
) (
<bibRefCitation DOI="https://doi.org/10.1111/j.1744-7429.2001.tb00201.x" author="Birkinshaw, C" journalOrPublisher="Biotropica" pageId="0" pageNumber="133" pagination="478 - 486" refId="B5" refString="Birkinshaw, C, 2001. Fruit characteristics of species dispersed by the Black Lemur (Eulemur macaco) in the Lokobe Forest, Madagascar. Biotropica 33 (3): 478 - 486, DOI: https://doi.org/10.1111/j.1744-7429.2001.tb00201.x" title="Fruit characteristics of species dispersed by the Black Lemur (Eulemur macaco) in the Lokobe Forest, Madagascar." url="https://doi.org/10.1111/j.1744-7429.2001.tb00201.x" volume="33" year="2001">Birkinshaw 2001</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.90.20586" author="Berry, PE" journalOrPublisher="Phytokeys" pageId="0" pageNumber="133" pagination="1 - 87" refId="B4" refString="Berry, PE, Kainulainen, K, van Ee, BW, 2017. A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands. Phytokeys 90: 1 - 87, DOI: https://doi.org/10.3897/phytokeys.90.20586" title="A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands." url="https://doi.org/10.3897/phytokeys.90.20586" volume="90" year="2017">Berry et al. 2017</bibRefCitation>
). Pierre Viette took one of the possibly oldest pictures published of this forest in November 1958 (
<emphasis italics="true" pageId="0" pageNumber="133">cf.</emphasis>
<bibRefCitation author="Viette, P" journalOrPublisher="1, annee 1962" pageId="0" pageNumber="133" refId="B99" refString="Viette, P, 1962. Noctuelles Trifides de Madagascar, Ecologie, Biogeographie, Morphologie et Taxonomie (Lep.). Theses presentees a la Faculte des Sciences de L'Universite de Nancy. Annales de la Societe Entomologique de France, tome 131, fasc. 1, annee 1962" title="Noctuelles Trifides de Madagascar, Ecologie, Biogeographie, Morphologie et Taxonomie (Lep.). Theses presentees a la Faculte des Sciences de L'Universite de Nancy. Annales de la Societe Entomologique de France, tome 131, fasc." year="1962">Viette 1962</bibRefCitation>
, plate III, fig. 6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="biological traits">
<paragraph pageId="0" pageNumber="133">Biological traits.</paragraph>
<paragraph pageId="0" pageNumber="133">
The biology of
<taxonomicName baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>