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<mods:title id="9D1A2FED84BC166BF73622A7E7C2B74B">Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species</mods:title>
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<mods:namePart>Qin, Yuyao</mods:namePart>
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<mods:affiliation>College of Life Sciences, Hebei University, Baoding 071002, China & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="45F08C087F8F0DDACE1ACC678E721B6D">Qin, Yuyao</mods:namePart>
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<mods:namePart>Germann, Christoph</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4051 Basel, Switzerland</mods:affiliation>
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<mods:namePart id="4A49811AC5CE5DD59CEDD045CB996C12">Germann, Christoph</mods:namePart>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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Subgenus
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<taxonomicName LSID="C1F49D6A-1F58-5AFB-9C71-D7D2124DAB99" authority="Casale, 1984" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Sphodromimus">Chlaenius Sphodromimus Casale, 1984</taxonomicName>
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<taxonomicName id="CCDE6B8E7E4EDB0E454E1A6DC018DC06" ID-CoL="8MHQY" LSID="C1F49D6A-1F58-5AFB-9C71-D7D2124DAB99" authority="Casale, 1984" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Sphodromimus">Chlaenius Sphodromimus Casale, 1984</taxonomicName>
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</paragraph>
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<paragraph pageId="0" pageNumber="61">
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<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Sphodromimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sphodromimus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Sphodromimus</taxonomicName>
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<taxonomicName id="389EC4E89B0C613E05FB4682633E4ECA" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Sphodromimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodromimus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Sphodromimus</taxonomicName>
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Casale, 1984: 372;
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<bibRefCitation author="Morvan, DM" journalOrPublisher="Elytron" pageId="0" pageNumber="61" pagination="55 - 62" refId="B22" refString="Morvan, DM, 1991. Contribụtion a la Connaissancedes Coleopteres Carabidae de Thailand. Elytron 5: 55 - 62" title="Contrib ụtion a la Connaissancedes Coleopteres Carabidae de Thailand." volume="5" year="1991">Morvan 1991</bibRefCitation>
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<bibRefCitation id="099713FEC3E451BC29148CC5881DF6D1" author="Morvan, DM" journalOrPublisher="Elytron" pageId="0" pageNumber="61" pagination="55 - 62" refId="B22" refString="Morvan, DM, 1991. Contribụtion a la Connaissancedes Coleopteres Carabidae de Thailand. Elytron 5: 55 - 62" title="Contrib ụtion a la Connaissancedes Coleopteres Carabidae de Thailand." volume="5" year="1991">Morvan 1991</bibRefCitation>
|
||||
: 60 (described new species);
|
||||
<bibRefCitation author="Morvan, DM" journalOrPublisher="Loened Aziad Amprevaned Feureskalleged C'wiledig" pageId="0" pageNumber="61" pagination="1 - 23" refId="B23" refString="Morvan, DM, 1997. Etude faunistique des coleopteres du Nepal avec extension aux provinces chinoises du Yunnan et du Sichuan. Genre Andrewesius Jedlicka et Vacinius Casale. Loened Aziad Amprevaned Feureskalleged C'wiledig 2: 1 - 23" title="Etude faunistique des coleopteres du Nepal avec extension aux provinces chinoises du Yunnan et du Sichuan. Genre Andrewesius Jedlicka et Vacinius Casale." volume="2" year="1997">Morvan 1997</bibRefCitation>
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<bibRefCitation id="79F144FCF86DBC6297C3C763E2F41B48" author="Morvan, DM" journalOrPublisher="Loened Aziad Amprevaned Feureskalleged C'wiledig" pageId="0" pageNumber="61" pagination="1 - 23" refId="B23" refString="Morvan, DM, 1997. Etude faunistique des coleopteres du Nepal avec extension aux provinces chinoises du Yunnan et du Sichuan. Genre Andrewesius Jedlicka et Vacinius Casale. Loened Aziad Amprevaned Feureskalleged C'wiledig 2: 1 - 23" title="Etude faunistique des coleopteres du Nepal avec extension aux provinces chinoises du Yunnan et du Sichuan. Genre Andrewesius Jedlicka et Vacinius Casale." volume="2" year="1997">Morvan 1997</bibRefCitation>
|
||||
: 16 (described new species);
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz 1998</bibRefCitation>
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<bibRefCitation id="A9D2001AB05421478FFFE0C10295068E" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz 1998</bibRefCitation>
|
||||
: 320 (catalogue);
|
||||
<bibRefCitation author="Lassalle, B" journalOrPublisher="Entomologiste" pageId="0" pageNumber="61" pagination="239 - 243" refId="B17" refString="Lassalle, B, 2001. Chasses en Birmanie. Entomologiste 57: 239 - 243" title="Chasses en Birmanie." volume="57" year="2001">Lassalle 2001</bibRefCitation>
|
||||
<bibRefCitation id="AD71AB9B9E6605C4EB2B1E579EDE062C" author="Lassalle, B" journalOrPublisher="Entomologiste" pageId="0" pageNumber="61" pagination="239 - 243" refId="B17" refString="Lassalle, B, 2001. Chasses en Birmanie. Entomologiste 57: 239 - 243" title="Chasses en Birmanie." volume="57" year="2001">Lassalle 2001</bibRefCitation>
|
||||
: 240 (described new species);
|
||||
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Entomofauna" pageId="0" pageNumber="61" pagination="29 - 60" refId="B12" refString="Kirschenhofer, E, 2003. Ueber neue und wenig bekannte Carabidae aus der aethiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae). Entomofauna 24: 29 - 60" title="Ueber neue und wenig bekannte Carabidae aus der aethiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae)." volume="24" year="2003">Kirschenhofer 2003</bibRefCitation>
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<bibRefCitation id="72832741843980A473D6375AC8BFA44D" author="Kirschenhofer, E" journalOrPublisher="Entomofauna" pageId="0" pageNumber="61" pagination="29 - 60" refId="B12" refString="Kirschenhofer, E, 2003. Ueber neue und wenig bekannte Carabidae aus der aethiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae). Entomofauna 24: 29 - 60" title="Ueber neue und wenig bekannte Carabidae aus der aethiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae)." volume="24" year="2003">Kirschenhofer 2003</bibRefCitation>
|
||||
: 32 (described new species);
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
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<bibRefCitation id="8CC59C1358AE426DF978F19C34A89AB2" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
|
||||
: 341 (catalogue);
|
||||
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Entomofauna" pageId="0" pageNumber="61" refId="B13" refString="Kirschenhofer, E, 2012. Neue Arten der Gattung Vachinius Casale, 1984 von Laos. Mitteilungen des Internationalen Entomologischen Vereins Frankfurt a.M. 1-2: 83-90." title="Neue Arten der Gattung Vachinius Casale, 1984 von Laos. Mitteilungen des Internationalen Entomologischen Vereins Frankfurt a. M. 1 - 2: 83 - 90." year="2012">Kirschenhofer 2012</bibRefCitation>
|
||||
<bibRefCitation id="5155B60A1E3C2E0AB8893350FD9836D3" author="Kirschenhofer, E" journalOrPublisher="Entomofauna" pageId="0" pageNumber="61" refId="B13" refString="Kirschenhofer, E, 2012. Neue Arten der Gattung Vachinius Casale, 1984 von Laos. Mitteilungen des Internationalen Entomologischen Vereins Frankfurt a.M. 1-2: 83-90." title="Neue Arten der Gattung Vachinius Casale, 1984 von Laos. Mitteilungen des Internationalen Entomologischen Vereins Frankfurt a. M. 1 - 2: 83 - 90." year="2012">Kirschenhofer 2012</bibRefCitation>
|
||||
: 84 (described new species);
|
||||
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="61" pagination="23 - 36" refId="B14" refString="Kirschenhofer, E, 2013. Ein Beitrag zur Kenntnis der Tribus Chlaeniini (= Callistini sensu auct.) der palaearktischen, afrotropischen und orientalischen Region (Coleoptera: Carabidae). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 65: 23 - 36" title="Ein Beitrag zur Kenntnis der Tribus Chlaeniini (= Callistini sensu auct.) der palaearktischen, afrotropischen und orientalischen Region (Coleoptera: Carabidae)." volume="65" year="2013">Kirschenhofer 2013</bibRefCitation>
|
||||
<bibRefCitation id="F7D3654A930FFD5112939F8D7070AFA6" author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="61" pagination="23 - 36" refId="B14" refString="Kirschenhofer, E, 2013. Ein Beitrag zur Kenntnis der Tribus Chlaeniini (= Callistini sensu auct.) der palaearktischen, afrotropischen und orientalischen Region (Coleoptera: Carabidae). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 65: 23 - 36" title="Ein Beitrag zur Kenntnis der Tribus Chlaeniini (= Callistini sensu auct.) der palaearktischen, afrotropischen und orientalischen Region (Coleoptera: Carabidae)." volume="65" year="2013">Kirschenhofer 2013</bibRefCitation>
|
||||
: 9 (new combination from
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Haplochlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Haplochlaenius</taxonomicName>
|
||||
<taxonomicName id="AB6A96C66A25A928B8E909830861ADE7" class="Insecta" family="Carabidae" genus="Haplochlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Haplochlaenius</taxonomicName>
|
||||
);
|
||||
<bibRefCitation author="Brunk, I" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="0" pageNumber="61" pagination="5 - 10" refId="B5" refString="Brunk, I, 2015. A new species of the genus Vachinius Casale, 1984 from the Philippines (Coleoptera, Carabidae, Chlaeniini). Entomologische Blatter fur Biologie und Systematik der Kafer 111: 5 - 10" title="A new species of the genus Vachinius Casale, 1984 from the Philippines (Coleoptera, Carabidae, Chlaeniini)." volume="111" year="2015">Brunk 2015</bibRefCitation>
|
||||
<bibRefCitation id="4C5EEF7D22392AAA69E63E5B7847429E" author="Brunk, I" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="0" pageNumber="61" pagination="5 - 10" refId="B5" refString="Brunk, I, 2015. A new species of the genus Vachinius Casale, 1984 from the Philippines (Coleoptera, Carabidae, Chlaeniini). Entomologische Blatter fur Biologie und Systematik der Kafer 111: 5 - 10" title="A new species of the genus Vachinius Casale, 1984 from the Philippines (Coleoptera, Carabidae, Chlaeniini)." volume="111" year="2015">Brunk 2015</bibRefCitation>
|
||||
: 5 (described new species);
|
||||
<bibRefCitation author="Kirschenhofer, E" editor="Loebl, I" journalOrPublisher="1. 2 nd Edn.). Brill, Leiden, Boston" pageId="0" pageNumber="61" pagination="491 - 497" refId="B15" refString="Kirschenhofer, E, 2017. Tribe Chlaeniini Brulle, 1834. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1. 2nd Edn.). Brill, Leiden, Boston: 491 - 497" title="Tribe Chlaeniini Brulle, 1834." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2017">Kirschenhofer 2017</bibRefCitation>
|
||||
<bibRefCitation id="34D568EBA82114D00086ABDAD057A043" author="Kirschenhofer, E" editor="Loebl, I" journalOrPublisher="1. 2 nd Edn.). Brill, Leiden, Boston" pageId="0" pageNumber="61" pagination="491 - 497" refId="B15" refString="Kirschenhofer, E, 2017. Tribe Chlaeniini Brulle, 1834. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1. 2nd Edn.). Brill, Leiden, Boston: 491 - 497" title="Tribe Chlaeniini Brulle, 1834." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2017">Kirschenhofer 2017</bibRefCitation>
|
||||
: 497 (catalogue);
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
||||
<bibRefCitation id="462D6FE4ECB1C8410A04CD55716F2870" DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
||||
: 1 (
|
||||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
|
||||
<taxonomicName id="4AA62A6A4F33418C9BB88D98289D09AD" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
|
||||
considered subgenus of
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Chlaenius</taxonomicName>
|
||||
<taxonomicName id="2A702191CAC8975F5B97D74CF0245CFF" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Chlaenius</taxonomicName>
|
||||
);
|
||||
<bibRefCitation author="Zettel, H" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="61" pagination="29 - 35" refId="B26" refString="Zettel, H, 2020. A new species of Chlaenius subgen. Sphodromimus Casale, 1984 (Coleoptera: Carabidae) from the Philippines. Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 72: 29 - 35" title="A new species of Chlaenius subgen. Sphodromimus Casale, 1984 (Coleoptera: Carabidae) from the Philippines." volume="72" year="2020">Zettel 2020</bibRefCitation>
|
||||
<bibRefCitation id="497752401CDCECF1B3C6E28D82668EF9" author="Zettel, H" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="61" pagination="29 - 35" refId="B26" refString="Zettel, H, 2020. A new species of Chlaenius subgen. Sphodromimus Casale, 1984 (Coleoptera: Carabidae) from the Philippines. Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 72: 29 - 35" title="A new species of Chlaenius subgen. Sphodromimus Casale, 1984 (Coleoptera: Carabidae) from the Philippines." volume="72" year="2020">Zettel 2020</bibRefCitation>
|
||||
: 29 (described new species).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="type species">
|
||||
<paragraph pageId="0" pageNumber="61">Type species.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius holzschuhi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="holzschuhi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Vachinius holzschuhi</emphasis>
|
||||
<subSubSection id="2C24CA336B2F0EEE8668E90E1229D6BD" pageId="0" pageNumber="61" type="type species">
|
||||
<paragraph id="510584820DB6365E4CD82E8AA2A3E00A" pageId="0" pageNumber="61">Type species.</paragraph>
|
||||
<paragraph id="E9235923469196DE8E405B2570A35065" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="07511D45B7EFCF8077A50F7DDBB48E58" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius holzschuhi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="holzschuhi">
|
||||
<emphasis id="E143DB437DF7430C45F50448B5604878" italics="true" pageId="0" pageNumber="61">Vachinius holzschuhi</emphasis>
|
||||
</taxonomicName>
|
||||
(Casale, 1984) (type locality: East Nepal, Tashigaon 2100 m; holotype in NHMB), by original designation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Sphodromimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sphodromimus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Sphodromimus</emphasis>
|
||||
<subSubSection id="08E449767901DFD41E65F22D594DFF7A" pageId="0" pageNumber="61" type="diagnosis">
|
||||
<paragraph id="EE33CBE3FC14FA4C5A9DFB183F8B0133" pageId="0" pageNumber="61">Diagnosis.</paragraph>
|
||||
<paragraph id="6DC391974CF10636AC48CF32C89F3D99" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="6B5BDA93C7D55244AAB13C2FEC515A57" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Sphodromimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodromimus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="354C5E97798C3367D7F1236EB0D62F88" italics="true" pageId="0" pageNumber="61">Sphodromimus</emphasis>
|
||||
</taxonomicName>
|
||||
can be distinguished from other subgenera in
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Chlaenius</emphasis>
|
||||
<taxonomicName id="1BB4D6EFF3663210269FC88F7B491D97" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1829651DABF2BE62ED0F0C7488CDB773" italics="true" pageId="0" pageNumber="61">Chlaenius</emphasis>
|
||||
</taxonomicName>
|
||||
by the following character combinations: body large, BL 19-26 mm; body black or metallic colored, luster matt or strongly shiny, antennae, mandibles usually dark brown, elytra black, ventral side black; head finely punctate; penultimate labial palpomere with 5-7 setae, apex truncate; antennae long, antennomere 3 longer than 4; mentum tooth stout, bifid; pronotum long, with sides usually sinuate before posterior angles, posterior lateral seta situated before posterior angles, anterior lateral seta absent; proepisterna sparsely punctate and pubescent; elytral intervals flat or convex, not ribbed, densely punctate and pubescent, basal margin reaching the scutellum; hind wings reduced in all species except
|
||||
<taxonomicName authorityName="Laporte" authorityYear="1834" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Chlaenius flavofemoratus</emphasis>
|
||||
<taxonomicName id="4F2DDBEC1D254FFA1DC84E3209590394" authorityName="Laporte" authorityYear="1834" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus">
|
||||
<emphasis id="6CBBD4AE330D08C2B4C5AB46757F496B" italics="true" pageId="0" pageNumber="61">Chlaenius flavofemoratus</emphasis>
|
||||
</taxonomicName>
|
||||
Laporte, 1834 and
|
||||
<taxonomicName baseAuthorityName="Louwerens" baseAuthorityYear="1967" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius peterseni" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="peterseni">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Chlaenius peterseni</emphasis>
|
||||
<taxonomicName id="73B4247CB1D1131F7310EFAE526EFC4A" baseAuthorityName="Louwerens" baseAuthorityYear="1967" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius peterseni" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="peterseni">
|
||||
<emphasis id="640452761250A7C50A7F318428F3AA2E" italics="true" pageId="0" pageNumber="61">Chlaenius peterseni</emphasis>
|
||||
</taxonomicName>
|
||||
(Louwerens, 1967); prosternal process unbordered at apex; metepisterna wider than long in all species except
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. flavofemoratus</emphasis>
|
||||
<taxonomicName id="9D50B4D27D74B70CCD886D5E77B996A8" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus">
|
||||
<emphasis id="E59961C7834ABE16EF9B974A1E12F59E" italics="true" pageId="0" pageNumber="61">C. flavofemoratus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. peterseni" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="peterseni">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. peterseni</emphasis>
|
||||
<taxonomicName id="C0A2E19E7B4393049CD80C2D1B85109B" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. peterseni" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="peterseni">
|
||||
<emphasis id="AB5AF08ED8696BC1FA70F5E8C68029A3" italics="true" pageId="0" pageNumber="61">C. peterseni</emphasis>
|
||||
</taxonomicName>
|
||||
, coarsely punctate, pubescent; legs sparsely pubescent, tarsi nearly smooth dorsally, claws simple, protibiae sulcate on dorsal side; abdominal sternites finely punctate laterally; apical lamella of aedeagus denticulate on dorsal side; apical gonocoxite without ensiform setae; receptaculum very short to absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="comparisons">
|
||||
<paragraph pageId="0" pageNumber="61">Comparisons.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<subSubSection id="D80DF3281231563A922EA17E3B98F2B3" pageId="0" pageNumber="61" type="comparisons">
|
||||
<paragraph id="52C9AE64B89E58450FBD49BB86D58FFC" pageId="0" pageNumber="61">Comparisons.</paragraph>
|
||||
<paragraph id="DAFDB63A8C32286A40A9A303322170F3" pageId="0" pageNumber="61">
|
||||
This subgenus is most similar to subgenera
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Haplochlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Haplochlaenius</emphasis>
|
||||
<taxonomicName id="DC8EB3448177D70CBCAB43FC8A49D3A5" class="Insecta" family="Carabidae" genus="Haplochlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="0FE1C10F16DB319CFC220196891106E7" italics="true" pageId="0" pageNumber="61">Haplochlaenius</emphasis>
|
||||
</taxonomicName>
|
||||
Lutshnik and
|
||||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Vachinius</emphasis>
|
||||
<taxonomicName id="3F5A28AB0F93C5D4918C38A118C40471" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F70BCF7CB377C46861FF74D99D7F3717" italics="true" pageId="0" pageNumber="61">Vachinius</emphasis>
|
||||
</taxonomicName>
|
||||
Casale, but differs in having elytra with intervals flat or slightly convex, densely punctate and pubescent, and with basal margin complete, connected with scutellum (intervals strongly ribbed in
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Haplochlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Haplochlaenius</emphasis>
|
||||
<taxonomicName id="EB413058EC80AA5A81AC81DE18E9E946" class="Insecta" family="Carabidae" genus="Haplochlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="59FB0FC9643DF83191D5D3B8BD398F6F" italics="true" pageId="0" pageNumber="61">Haplochlaenius</emphasis>
|
||||
</taxonomicName>
|
||||
, basal margin obsolete near scutellum; intervals smooth in
|
||||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Vachinius</emphasis>
|
||||
<taxonomicName id="D06F84B38C7386C3A041274E5ACEA86D" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A0CE45ACA5C6E591081D8389A429BE5B" italics="true" pageId="0" pageNumber="61">Vachinius</emphasis>
|
||||
</taxonomicName>
|
||||
; cfr.
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
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<bibRefCitation id="1E7C921183E8B1047EBBB25C2762BF6C" DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
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).
|
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="61">Species and distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">Subgenus is composed of 16 species distributed in the Oriental Region (China, Indonesia, Laos, Myanmar, Nepal, Philippines, Thailand, Vietnam), including the two new species described below.</paragraph>
|
||||
<subSubSection id="ED2279E4481BDD190005529D5BBCD215" pageId="0" pageNumber="61" type="distribution">
|
||||
<paragraph id="6A5A8972A00B7151CBC38775A81E4DD6" pageId="0" pageNumber="61">Species and distribution.</paragraph>
|
||||
<paragraph id="20867BCA1C15AA155994B776306AB7AA" pageId="0" pageNumber="61">Subgenus is composed of 16 species distributed in the Oriental Region (China, Indonesia, Laos, Myanmar, Nepal, Philippines, Thailand, Vietnam), including the two new species described below.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
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