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<mods:title id="71385398438240647EAC933132D1C438">A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy</mods:title>
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<mods:namePart>Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation>Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany & Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:namePart id="F4047905F8AF6380366D4857857BC11B">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="2BA8FBD6CA4A55721478371C076BD36F">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany & Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:namePart>Velde, Sabrina van de</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:namePart>Yanina, Tamara</mods:namePart>
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<mods:affiliation>Moscow State University, Faculty of Geography, Leninskie Gory, 1, 119991 Moscow, Russia</mods:affiliation>
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<mods:namePart id="57ED559685E36906D90207409E597DF0">Yanina, Tamara</mods:namePart>
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<mods:affiliation id="B9D4D8E1BE4A26475E1D6CF764804D43">Moscow State University, Faculty of Geography, Leninskie Gory, 1, 119991 Moscow, Russia</mods:affiliation>
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<mods:namePart>Wesselingh, Frank P.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:namePart id="2AD8367A95002EC222502262C49EA8F7">Wesselingh, Frank P.</mods:namePart>
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<treatment ID-GBIF-Taxon="182229616" LSID="urn:lsid:plazi:treatment:5B47BC86DFFFDDA3DF763C68664B1073" httpUri="http://treatment.plazi.org/id/5B47BC86DFFFDDA3DF763C68664B1073" lastPageNumber="43" pageId="0" pageNumber="43">
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<treatment id="5B47BC86DFFFDDA3DF763C68664B1073" ID-GBIF-Taxon="182229616" LSID="urn:lsid:plazi:treatment:5B47BC86DFFFDDA3DF763C68664B1073" httpUri="http://treatment.plazi.org/id/5B47BC86DFFFDDA3DF763C68664B1073" lastPageNumber="43" pageId="0" pageNumber="43" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_subClass="Caenogastropoda" scope_superFamily="Truncatelloidea">
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Subfamily
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<taxonomicName authority="B. Dybowski, 1913" authorityName="B. Dybowski" authorityYear="1913" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Caspiinae">Caspiinae B. Dybowski, 1913</taxonomicName>
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<taxonomicName id="EA267337F00233E67B521064422C22AC" ID-CoL="7NXHX" authority="B. Dybowski, 1913" authorityName="B. Dybowski" authorityYear="1913" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="subFamily" subFamily="Caspiinae">Caspiinae B. Dybowski, 1913</taxonomicName>
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</paragraph>
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<subSubSection pageId="0" pageNumber="43" type="discussion">
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<paragraph pageId="0" pageNumber="43">Discussion.</paragraph>
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<paragraph id="D6F9C869AE8CB90CCF556066CDE64410" pageId="0" pageNumber="43">Discussion.</paragraph>
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<paragraph id="03E1399F337833BE610CCACEAB127652" pageId="0" pageNumber="43">
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The genus
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<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
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<taxonomicName id="63D4DEC784742EC34E602A0CC580F035" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
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<emphasis id="630B1978DCE7CB13F23DD34D3B95F550" italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
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</taxonomicName>
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has been widely used for species with small ovoid shells, occasionally with spiral or reticulate teleoconch sculpture. Based on the expression of sculpture, some authors have divided the species among the (sub)genera
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<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
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||||
<taxonomicName id="6BFC7044AC40CBEE9C6293AF8F82E16E" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
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<emphasis id="DCCC93F89CEB968CAF2FBDE430D6DAF1" italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
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</taxonomicName>
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s.s., with a single spiral line below the suture, and
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<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
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<emphasis italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
<taxonomicName id="491AC2AB22DA6083F7C1158DFA20411C" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="6F8BEF8CE931781F0754DBC39D8A19DE" italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
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||||
</taxonomicName>
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Lindholm, 1930, exposing a reticulate pattern (e.g.,
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<bibRefCitation author="Anistratenko, VV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="43" pagination="15 - 21" refId="B10" refString="Anistratenko, VV, Prisyazhniuk, VA, 1992. Novyye dannyye o mollyuskakh golotsenovykh otlozheniy Chernogo morya na Ukraine. Vestnik Zoologii 5: 15 - 21" title="Novyye dannyye o mollyuskakh golotsenovykh otlozheniy Chernogo morya na Ukraine." volume="5" year="1992">Anistratenko and Prisyazhniuk 1992</bibRefCitation>
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<bibRefCitation id="66E48C4249F57CBE75513709397E3386" author="Anistratenko, VV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="43" pagination="15 - 21" refId="B10" refString="Anistratenko, VV, Prisyazhniuk, VA, 1992. Novyye dannyye o mollyuskakh golotsenovykh otlozheniy Chernogo morya na Ukraine. Vestnik Zoologii 5: 15 - 21" title="Novyye dannyye o mollyuskakh golotsenovykh otlozheniy Chernogo morya na Ukraine." volume="5" year="1992">Anistratenko and Prisyazhniuk 1992</bibRefCitation>
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||||
,
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2012.742934" author="Anistratenko, VV" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="43" pagination="51 - 64" refId="B8" refString="Anistratenko, VV, 2013. On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae). Journal of Natural History 47 (1-2): 51 - 64, DOI: https://doi.org/10.1080/00222933.2012.742934" title="On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae)." url="https://doi.org/10.1080/00222933.2012.742934" volume="47" year="2013">Anistratenko 2013</bibRefCitation>
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<bibRefCitation id="6F142A8EBFDE61A31F2462A71303A74C" DOI="https://doi.org/10.1080/00222933.2012.742934" author="Anistratenko, VV" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="43" pagination="51 - 64" refId="B8" refString="Anistratenko, VV, 2013. On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae). Journal of Natural History 47 (1-2): 51 - 64, DOI: https://doi.org/10.1080/00222933.2012.742934" title="On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae)." url="https://doi.org/10.1080/00222933.2012.742934" volume="47" year="2013">Anistratenko 2013</bibRefCitation>
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,
|
||||
<bibRefCitation DOI="https://doi.org/10.12657/folmal.023.014" author="Boeters, HD" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="43" pagination="177 - 186" refId="B14" refString="Boeters, HD, Gloeer, P, Georgiev, D, Dedov, I, 2015. A new species of Caspia Clessin et W. Dybowski, 1887 (Gastropoda: Truncatelloidea: Hydrobiidae) in the Danube of Bulgaria. Folia Malacologica 23: 177 - 186, DOI: https://doi.org/10.12657/folmal.023.014" title="A new species of Caspia Clessin et W. Dybowski, 1887 (Gastropoda: Truncatelloidea: Hydrobiidae) in the Danube of Bulgaria." url="https://doi.org/10.12657/folmal.023.014" volume="23" year="2015">Boeters et al. 2015</bibRefCitation>
|
||||
<bibRefCitation id="763354208C127832E3DD1B79407A2B13" DOI="https://doi.org/10.12657/folmal.023.014" author="Boeters, HD" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="43" pagination="177 - 186" refId="B14" refString="Boeters, HD, Gloeer, P, Georgiev, D, Dedov, I, 2015. A new species of Caspia Clessin et W. Dybowski, 1887 (Gastropoda: Truncatelloidea: Hydrobiidae) in the Danube of Bulgaria. Folia Malacologica 23: 177 - 186, DOI: https://doi.org/10.12657/folmal.023.014" title="A new species of Caspia Clessin et W. Dybowski, 1887 (Gastropoda: Truncatelloidea: Hydrobiidae) in the Danube of Bulgaria." url="https://doi.org/10.12657/folmal.023.014" volume="23" year="2015">Boeters et al. 2015</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation DOI="https://doi.org/10.1016/j.quaint.2016.03.018" author="Bueyuekmeric, Y" journalOrPublisher="Actienbuchdruckerei, Agram" pageId="0" pageNumber="43" refId="B20" refString="Bueyuekmeric, Y, Wesselingh, FP, 2018. New cockles (Bivalvia: Cardiidae: Lymnocardiinae) from Late Pleistocene Lake Karapinar (Turkey): Discovery of a Pontocaspian refuge? Quaternary International 465 (A): 37-45. https://doi.org/10.1016/j.quaint.2016.03.018" title="New cockles (Bivalvia: Cardiidae: Lymnocardiinae) from Late Pleistocene Lake Karapinar (Turkey): Discovery of a Pontocaspian refuge? Quaternary International 465 (A): 37 - 45." url="https://doi.org/10.1016/j.quaint.2016.03.018" year="2018">
|
||||
<normalizedToken originalValue="Büyükmeriç">Bueyuekmeric</normalizedToken>
|
||||
<bibRefCitation id="0392DE722271E39319E86450BAAAF603" DOI="https://doi.org/10.1016/j.quaint.2016.03.018" author="Bueyuekmeric, Y" journalOrPublisher="Actienbuchdruckerei, Agram" pageId="0" pageNumber="43" refId="B20" refString="Bueyuekmeric, Y, Wesselingh, FP, 2018. New cockles (Bivalvia: Cardiidae: Lymnocardiinae) from Late Pleistocene Lake Karapinar (Turkey): Discovery of a Pontocaspian refuge? Quaternary International 465 (A): 37-45. https://doi.org/10.1016/j.quaint.2016.03.018" title="New cockles (Bivalvia: Cardiidae: Lymnocardiinae) from Late Pleistocene Lake Karapinar (Turkey): Discovery of a Pontocaspian refuge? Quaternary International 465 (A): 37 - 45." url="https://doi.org/10.1016/j.quaint.2016.03.018" year="2018">
|
||||
<normalizedToken id="D97ADA9DFEF88C2CC2A212A04639F865" originalValue="Büyükmeriç">Bueyuekmeric</normalizedToken>
|
||||
and Wesselingh 2018
|
||||
</bibRefCitation>
|
||||
). Species lacking teleoconch sculpture were grouped under the new taxon
|
||||
<taxonomicName authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
<taxonomicName id="31845DF17E00A70373DD468683597F43" authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="95C93F1981678443B838DED8276437A7" italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation author="Logvinenko, BM" editor="Birshtein, YaA" journalOrPublisher="Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva" pageId="0" pageNumber="43" pagination="308 - 385" refId="B70" refString="Logvinenko, BM, Starobogatov, YaI, 1969. Mollusca. In: Birshtein, YaA, Vinogradov, LG, Kondakov, NN, Kuhn, MS, Astakhova, TV, Romanova, NN, Eds., Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva: 308 - 385" title="Mollusca." volumeTitle="Atlas bespozvonochnykh Kaspiyskogo morya." year="1969">Logvinenko and Starobogatov (1969)</bibRefCitation>
|
||||
<bibRefCitation id="AAADF6ECE16D98D4AB4A731C535B2420" author="Logvinenko, BM" editor="Birshtein, YaA" journalOrPublisher="Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva" pageId="0" pageNumber="43" pagination="308 - 385" refId="B70" refString="Logvinenko, BM, Starobogatov, YaI, 1969. Mollusca. In: Birshtein, YaA, Vinogradov, LG, Kondakov, NN, Kuhn, MS, Astakhova, TV, Romanova, NN, Eds., Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva: 308 - 385" title="Mollusca." volumeTitle="Atlas bespozvonochnykh Kaspiyskogo morya." year="1969">Logvinenko and Starobogatov (1969)</bibRefCitation>
|
||||
. While those authors considered it a subgenus of
|
||||
<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
|
||||
<taxonomicName id="D71F4EE8C6BAB345F11FD3ED25A50233" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="01B1A2F73080F12D981DCD36E612DF06" italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
|
||||
</taxonomicName>
|
||||
, W. Dybowski (1887) originally treated its
|
||||
<typeStatus>type</typeStatus>
|
||||
<typeStatus id="701DADB5EF52C60955222084CFDA0EBC">type</typeStatus>
|
||||
species (
|
||||
<taxonomicName authorityName="Clessin & W.Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia ulskii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="ulskii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia ulskii</emphasis>
|
||||
<taxonomicName id="CB16EEC2532A87ED4EBBE0F20119D8B2" authorityName="Clessin & W.Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia ulskii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="ulskii">
|
||||
<emphasis id="D1F8DBFD3943FB9FC825C88CE712A98E" italics="true" pageId="0" pageNumber="43">Caspia ulskii</emphasis>
|
||||
</taxonomicName>
|
||||
Clessin & W. Dybowski in W. Dybowski, 1887, see below) as a sculpture-less form of
|
||||
<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
<taxonomicName id="2E83B2F880383BD04D0E3F7FA840ACB4" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="5D3F744574C8D26260CD8A75F53122DA" italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="43">
|
||||
<taxonomicName baseAuthorityName="Clessin & W. Dybowski in W. Dybowski" baseAuthorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia ulskii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="ulskii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Ulskia ulskii</emphasis>
|
||||
<paragraph id="A776845298015B40D77EA44623359ED3" pageId="0" pageNumber="43">
|
||||
<taxonomicName id="B18368BACCAB3E1905628A9E44E762CA" baseAuthorityName="Clessin & W. Dybowski in W. Dybowski" baseAuthorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia ulskii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="ulskii">
|
||||
<emphasis id="A1F7F50EBE5530C2BECB2BD52601FE1A" italics="true" pageId="0" pageNumber="43">Ulskia ulskii</emphasis>
|
||||
</taxonomicName>
|
||||
is available in the present material, and we have investigated the
|
||||
<typeStatus>type</typeStatus>
|
||||
<typeStatus id="3B095727C7DA02FE82F4E99ADE3E3217">type</typeStatus>
|
||||
species of
|
||||
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
<taxonomicName id="B0964DD477F0A21A2667330568640E19" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="30F5AF12CDC95A73F7ADFC4196D4716B" italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName authorityName="Clessin & W. Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia pallasii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="pallasii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia pallasii</emphasis>
|
||||
<taxonomicName id="61200847BF8B1B1630DEB44ADF27366C" authorityName="Clessin & W. Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia pallasii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="pallasii">
|
||||
<emphasis id="AD343ECE3F9B54E7E31D85F7CE1A512E" italics="true" pageId="0" pageNumber="43">Caspia pallasii</emphasis>
|
||||
</taxonomicName>
|
||||
Clessin & W. Dybowski in W. Dybowski, 1887) obtained from Holocene deposits of the northern and southern Caspian Sea. However, the
|
||||
<typeStatus>type</typeStatus>
|
||||
<typeStatus id="8A9C3B385E8D6033C20952940B7BE19C">type</typeStatus>
|
||||
species of
|
||||
<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
<taxonomicName id="1B2122A8948E130F346F7EEB4EC38AB2" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="2F4ABDD588481445E472A92F36BF3DE1" italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName authorityName="Clessin & W. Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia baerii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="baerii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia baerii</emphasis>
|
||||
<taxonomicName id="3B610D7235AA798F6C2F689C2D703A7E" authorityName="Clessin & W. Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia baerii" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="baerii">
|
||||
<emphasis id="7DBD97744EC1EFD71BFE010028563D64" italics="true" pageId="0" pageNumber="43">Caspia baerii</emphasis>
|
||||
</taxonomicName>
|
||||
Clessin & W. Dybowski in W. Dybowski, 1887, is unknown to us. The original description suggests that it is similar to
|
||||
<taxonomicName authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
<taxonomicName id="178E4F4C692DC1CAEFFF7E03E4A007B2" authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="2C327441F33B6734261D86CDA5343396" italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
<taxonomicName id="229D84E33DB3A8533141586660B683AB" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="03AED9553D8F375DA8AE2CC4660A1338" italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
</taxonomicName>
|
||||
in terms of size and shape, yet to differ in the presence of a single line below to suture, demarcating a narrow subsutural ramp. All three genera are probably closely related, which is also suggested by the similar protoconchs of
|
||||
<taxonomicName authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
<taxonomicName id="CBF73E5E4FADC2E017FE6E9B23B022AB" authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="F6C2EB8CD0D31C4E99692203C86D9CB4" italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
<taxonomicName id="6340B47B8F61EEB3995376804FAB0E2A" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="7250166C6AD858A8D1F8ED17EC862019" italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
</taxonomicName>
|
||||
(pers. obs. T.A.N.). Since
|
||||
<taxonomicName authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
<taxonomicName id="F23BECC2D0B6F6AE25A35919210590E4" authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="1B8BC9E9ABE830316A98F390AD323F44" italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
<taxonomicName id="6843975A50AE6FA9BEEFE775B1C0E7A3" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="2FF76151807CC81ED2C09F7E0C24ED3A" italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
</taxonomicName>
|
||||
can be easily distinguished based on the presence of sculpture, we propose to treat them as distinct genera. The status of
|
||||
<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
<taxonomicName id="E56C7F66D70A69DDD5E756F435B9E480" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="39B3F816E7EC0837D0EF55F95693B127" italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
</taxonomicName>
|
||||
remains doubtful until the
|
||||
<typeStatus>type</typeStatus>
|
||||
<typeStatus id="C72DB55431B0C423403B38803FFE922D">type</typeStatus>
|
||||
species is properly re-investigated.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="43">
|
||||
<paragraph id="60EADA5AFA761F6E9AF21D86F4239B62" pageId="0" pageNumber="43">
|
||||
The
|
||||
<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
<taxonomicName id="AC25C0AE6A5C379715069E2B452CD952" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="64F0CBA90B360FD5C1A92F87E237E4E6" italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
|
||||
</taxonomicName>
|
||||
-
|
||||
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
<taxonomicName id="241E11B817B715C021DD15EE98244EF4" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clathrocaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="7AD84C53C1C77C841E8D09AF85819FD4" italics="true" pageId="0" pageNumber="43">Clathrocaspia</emphasis>
|
||||
</taxonomicName>
|
||||
-
|
||||
<taxonomicName authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
<taxonomicName id="208F58754CB2DA419C10BA8E51F0C624" authorityName="Logvinenko & Starobogatov" authorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulskia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="24F80A30CE2AFB4B3068A554BD788F15" italics="true" pageId="0" pageNumber="43">Ulskia</emphasis>
|
||||
</taxonomicName>
|
||||
species group can be well delimited from the larger, elongate-conical or -ovoid
|
||||
<taxonomicName authorityName="B. Dybowski & Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
|
||||
<taxonomicName id="23B534B4CCC9D12434FDC04D06E096F2" authorityName="B. Dybowski & Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="D3B53A99425886248D1F85B4BE5E2B6C" italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
|
||||
</taxonomicName>
|
||||
auct. and
|
||||
<taxonomicName authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
|
||||
<taxonomicName id="5F84E1E814F61D95B48AC14AF34EC060" authorityName="Neubauer & Velde & Yanina & Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer & Velde & Yanina & Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="CB2FA2EAFCA0AF44D4915A02B54EE101" italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
|
||||
</taxonomicName>
|
||||
auct. Moreover, unpublished molecular data suggest that the group is unrelated to
|
||||
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
|
||||
<taxonomicName id="D1B5554FF1E634CE130A0D8504148676" baseAuthorityName="Brusina" baseAuthorityYear="1881" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
|
||||
(T. Wilke, pers. comm. 04/2018). We follow
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2012.742934" author="Anistratenko, VV" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="43" pagination="51 - 64" refId="B8" refString="Anistratenko, VV, 2013. On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae). Journal of Natural History 47 (1-2): 51 - 64, DOI: https://doi.org/10.1080/00222933.2012.742934" title="On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae)." url="https://doi.org/10.1080/00222933.2012.742934" volume="47" year="2013">Anistratenko (2013)</bibRefCitation>
|
||||
<bibRefCitation id="676E46D4B9C4636FF3F3BD7DB242B6AB" DOI="https://doi.org/10.1080/00222933.2012.742934" author="Anistratenko, VV" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="43" pagination="51 - 64" refId="B8" refString="Anistratenko, VV, 2013. On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae). Journal of Natural History 47 (1-2): 51 - 64, DOI: https://doi.org/10.1080/00222933.2012.742934" title="On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae)." url="https://doi.org/10.1080/00222933.2012.742934" volume="47" year="2013">Anistratenko (2013)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation DOI="https://doi.org/10.4002/040.061.0201" author="Bouchet, P" journalOrPublisher="Malacologia" pageId="0" pageNumber="43" pagination="1 - 526" refId="B15" refString="Bouchet, P, Rocroi, J-P, Hausdorf, B, Kaim, A, Kano, Y, Nuetzel, A, Parkhaev, P, Schroedl, M, Strong, EE, 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1-2): 1 - 526, DOI: https://doi.org/10.4002/040.061.0201" title="Revised classification, nomenclator and typification of gastropod and monoplacophoran families." url="https://doi.org/10.4002/040.061.0201" volume="61" year="2017">Bouchet et al. (2017)</bibRefCitation>
|
||||
<bibRefCitation id="CCB68835C8E846849BDD6537DE593E75" DOI="https://doi.org/10.4002/040.061.0201" author="Bouchet, P" journalOrPublisher="Malacologia" pageId="0" pageNumber="43" pagination="1 - 526" refId="B15" refString="Bouchet, P, Rocroi, J-P, Hausdorf, B, Kaim, A, Kano, Y, Nuetzel, A, Parkhaev, P, Schroedl, M, Strong, EE, 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1-2): 1 - 526, DOI: https://doi.org/10.4002/040.061.0201" title="Revised classification, nomenclator and typification of gastropod and monoplacophoran families." url="https://doi.org/10.4002/040.061.0201" volume="61" year="2017">Bouchet et al. (2017)</bibRefCitation>
|
||||
, who listed the
|
||||
<taxonomicName authorityName="B.Dybowski" authorityYear="1913" lsidName="" pageId="0" pageNumber="43" rank="subFamily" subFamily="Caspiinae">Caspiinae</taxonomicName>
|
||||
<taxonomicName id="27E0DAB6B7CE76CC410C812D04881672" authorityName="B.Dybowski" authorityYear="1913" lsidName="" pageId="0" pageNumber="43" rank="subFamily" subFamily="Caspiinae">Caspiinae</taxonomicName>
|
||||
as separate subfamily.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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