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<mods:title id="652777026A2B297B8D263F71916181C4">Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species</mods:title>
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<mods:namePart>Qin, Yuyao</mods:namePart>
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<mods:affiliation>College of Life Sciences, Hebei University, Baoding 071002, China & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart>Germann, Christoph</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4051 Basel, Switzerland</mods:affiliation>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<taxonomicName LSID="33485B29-28AD-5248-9B49-E1065845E20A" authority="Mandl, 1992" authorityName="Mandl" authorityYear="1992" baseAuthorityName="Qin & Germann & Liang" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis" status="stat. nov." subGenus="Sphodromimus">Chlaenius (Sphodromimus) enleensis Mandl, 1992</taxonomicName>
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<taxonomicNameLabel id="1127D41D5CCB189BFB6315F6B76EDC70" pageId="0" pageNumber="61">stat. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A, B Chlaenius (Sphodromimus) flavofemoratus Laporte, 1834 (male, Yunnan, Menglun) C C. (Sphodromimus) flavofemoratus Laporte, 1834 (female, Yunnan, Menglun) 8 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, Indo Chine) Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures7-8" httpUri="https://binary.pensoft.net/fig/780979" pageId="0" pageNumber="61">Figs 8</figureCitation>
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<figureCitation captionStart="Figures 9, 10" captionStartId="F6" captionText="Figures 9, 10. 9 A, B Chlaenius (Sphodromimus) davidi nom. nov. (male, Guangdong, Xinyi) 10 A, B C. (Sphodromimus) enleensis (male, Vietnam, Tam Dao). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures9-10" httpUri="https://binary.pensoft.net/fig/780980" pageId="0" pageNumber="61">, 10A, B</figureCitation>
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<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">, 16</figureCitation>
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<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">, 24</figureCitation>
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<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">, 32A-F</figureCitation>
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<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">, 33A-E</figureCitation>
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<figureCitation captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">, 40</figureCitation>
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<figureCitation id="CAFAE4E7E659DA0589A0C32A27C5D7DC" captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A, B Chlaenius (Sphodromimus) flavofemoratus Laporte, 1834 (male, Yunnan, Menglun) C C. (Sphodromimus) flavofemoratus Laporte, 1834 (female, Yunnan, Menglun) 8 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, Indo Chine) Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures7-8" httpUri="https://binary.pensoft.net/fig/780979" pageId="0" pageNumber="61">Figs 8</figureCitation>
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<figureCitation id="E9EC2F8037AB9389CC7FBA5F2484FEE8" captionStart="Figures 9, 10" captionStartId="F6" captionText="Figures 9, 10. 9 A, B Chlaenius (Sphodromimus) davidi nom. nov. (male, Guangdong, Xinyi) 10 A, B C. (Sphodromimus) enleensis (male, Vietnam, Tam Dao). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures9-10" httpUri="https://binary.pensoft.net/fig/780980" pageId="0" pageNumber="61">, 10A, B</figureCitation>
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<figureCitation id="DCF6F34BAC06E76969868E9A35EB823C" captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">, 16</figureCitation>
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<figureCitation id="A9E50A1BB16805D89F5996D76E5DC548" captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">, 24</figureCitation>
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<figureCitation id="68CA8B59198F8BC7D44681A6CDC28409" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">, 32A-F</figureCitation>
|
||||
<figureCitation id="0681C855682C72DC18161D7FC644A892" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">, 33A-E</figureCitation>
|
||||
<figureCitation id="92980E7762EDBC1644647F27A25EDE1E" captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">, 40</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" baseAuthorityName="Mandl" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="flavofemoratus" status="stat. nov." subSpecies="enleensis">Chlaenius flavofemoratus enleensis</taxonomicName>
|
||||
<subSubSection id="249F192984AA96D52031EC0D0713CC65" pageId="0" pageNumber="61" type="reference_group">
|
||||
<paragraph id="0068B7A522168A31FACFECF98E77AE5F" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="63D3CD01A3C032D7E96873861FC387B9" authorityName="Qin & Germann & Liang" authorityYear="2022" baseAuthorityName="Mandl" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="flavofemoratus" status="stat. nov." subSpecies="enleensis">Chlaenius flavofemoratus enleensis</taxonomicName>
|
||||
Mandl, 1992: 100;
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz 1998</bibRefCitation>
|
||||
<bibRefCitation id="21494DEF7531AB07B5C3AB3DDD0A7DA6" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz 1998</bibRefCitation>
|
||||
: 318 (synonymized with
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus" status="stat. nov.">C. flavofemoratus</taxonomicName>
|
||||
<taxonomicName id="3B584625E575F1FFBF8B2DDD8E76726C" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus" status="stat. nov.">C. flavofemoratus</taxonomicName>
|
||||
Laporte,1834, catalogue);
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
|
||||
<bibRefCitation id="804AE76FAC8E77BB8BF8A7673E65C007" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
|
||||
: 338 (catalogue);
|
||||
<bibRefCitation author="Kirschenhofer, E" editor="Loebl, I" journalOrPublisher="1. 2 nd Edn.). Brill, Leiden, Boston" pageId="0" pageNumber="61" pagination="491 - 497" refId="B15" refString="Kirschenhofer, E, 2017. Tribe Chlaeniini Brulle, 1834. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1. 2nd Edn.). Brill, Leiden, Boston: 491 - 497" title="Tribe Chlaeniini Brulle, 1834." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2017">Kirschenhofer 2017</bibRefCitation>
|
||||
<bibRefCitation id="1CD49B8180F43FA7DFD27E80B58DE5C2" author="Kirschenhofer, E" editor="Loebl, I" journalOrPublisher="1. 2 nd Edn.). Brill, Leiden, Boston" pageId="0" pageNumber="61" pagination="491 - 497" refId="B15" refString="Kirschenhofer, E, 2017. Tribe Chlaeniini Brulle, 1834. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1. 2nd Edn.). Brill, Leiden, Boston: 491 - 497" title="Tribe Chlaeniini Brulle, 1834." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2017">Kirschenhofer 2017</bibRefCitation>
|
||||
: 491 (as a subspecies of
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus" status="stat. nov.">C. flavofemoratus</taxonomicName>
|
||||
<taxonomicName id="BC57D24E3B58BE59576E9C14BBF5A4C9" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus" status="stat. nov.">C. flavofemoratus</taxonomicName>
|
||||
; catalogue);
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
||||
<bibRefCitation id="A634C758D7510C658F631D04103047A2" DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
||||
: 1 (transferred to subgenus
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
|
||||
<taxonomicName id="ED4E946AFDFF3FADCBA95BE401AB1360" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
|
||||
from subgenus
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Haplochlaenius)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Haplochlaenius">Chlaenius Haplochlaenius</taxonomicName>
|
||||
<taxonomicName id="5ABF4FBE8604F8002B2F483AECB776F9" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Haplochlaenius)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Haplochlaenius">Chlaenius Haplochlaenius</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis" status="stat. nov.">Chlaenius tamdaoensis</taxonomicName>
|
||||
<paragraph id="576897195B4A501522E3FAC4790BDCC3" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="EC7BA7A52FC2CD1EF1EE1CCBA8714458" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis" status="stat. nov.">Chlaenius tamdaoensis</taxonomicName>
|
||||
Kirschenhofer, 2003: 32 (type locality: Vietnam, Tam Dao; genus
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="stat. nov.">Chlaenius</taxonomicName>
|
||||
<taxonomicName id="542808ED51772A1AF6C94D660478F5CF" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="stat. nov.">Chlaenius</taxonomicName>
|
||||
, subgenus
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Haplochlaenius)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Haplochlaenius">Chlaenius Haplochlaenius</taxonomicName>
|
||||
<taxonomicName id="8B14443A5F3955318A9078E8FC425173" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Haplochlaenius)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Haplochlaenius">Chlaenius Haplochlaenius</taxonomicName>
|
||||
);
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
|
||||
<bibRefCitation id="FB40924461A2AC10A9E622EB6FC5F1B7" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
|
||||
: 342 (catalogue);
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
||||
<bibRefCitation id="792D1BE4B1D45FFCCD3D5BC5546984D7" DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
|
||||
: 1 (genus
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="stat. nov.">Chlaenius</taxonomicName>
|
||||
<taxonomicName id="6E25751E2675421002F2FB282058903C" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="stat. nov.">Chlaenius</taxonomicName>
|
||||
, subgenus
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
|
||||
<taxonomicName id="FE7A5311375277D787A6D818058F638C" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status="stat. nov." subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
|
||||
) syn. nov.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="61">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">Indo Chine.</paragraph>
|
||||
<subSubSection id="30369B268A54F821F47CE448D0EABAB9" pageId="0" pageNumber="61" type="type locality">
|
||||
<paragraph id="289AD44E129D5D6CA7FDDAE101A225B8" pageId="0" pageNumber="61">Type locality.</paragraph>
|
||||
<paragraph id="E4D285ECC5D487F616702E99A7484511" pageId="0" pageNumber="61">Indo Chine.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="61">Material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<subSubSection id="4A9187107847CDB51302C3A7FF853ED9" pageId="0" pageNumber="61" type="materials_examined">
|
||||
<paragraph id="AD50C1F32FA60A440C3CD8B6DED001B8" pageId="0" pageNumber="61">Material examined.</paragraph>
|
||||
<paragraph id="05C78F6AECBA594874736948E8FAA56A" pageId="0" pageNumber="61">
|
||||
Total
|
||||
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Male, Indo Chine, Dr. K." country="Vietnam" location="Chl" specimenCount="5" typeStatus="Holotype">
|
||||
<specimenCount type="generic">5 specimens</specimenCount>
|
||||
<materialsCitation id="7D2FBDAF42A0012A3857C4BADC7A7228" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Male, Indo Chine, Dr. K." country="Vietnam" location="Chl" specimenCount="5" typeStatus="Holotype">
|
||||
<specimenCount id="5118FAE5669187B1AC3C2CFAF98767E8" type="generic">5 specimens</specimenCount>
|
||||
.
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||||
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
|
||||
<emphasis id="69ACE85CBBB47757A9BA87F608EB2723" bold="true" pageId="0" pageNumber="61">
|
||||
<collectingCountry id="19A6468ADA9673D1EDE85E1EA8BAFCBA" name="Vietnam">Vietnam</collectingCountry>
|
||||
</emphasis>
|
||||
:
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
<emphasis id="C9FB647AD0747A48205DB7D38A970FF4" bold="true" italics="true" pageId="0" pageNumber="61">
|
||||
<typeStatus id="600480F45638F0DD25416343380C68A8">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
,
|
||||
<collectorName>Male</collectorName>
|
||||
<collectorName id="19BC5FA6BBB0998BBE7CABFD3DBE7865">Male</collectorName>
|
||||
(NHMB),
|
||||
<collectorName>Indo Chine</collectorName>
|
||||
<collectorName id="DFE9F0E8D0C7D1C6943ED06A096C5C51">Indo Chine</collectorName>
|
||||
coll. Dussault/ En-Le 1908/
|
||||
<location LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:54A005A5E051D6D1E1EF865C86E2053E" country="Vietnam" name="Chl">Chl</location>
|
||||
<location id="54A005A5E051D6D1E1EF865C86E2053E" LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:54A005A5E051D6D1E1EF865C86E2053E" country="Vietnam" name="Chl">Chl</location>
|
||||
(
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Macrochlaeniles)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Macrochlaeniles">Chlaenius Macrochlaeniles</taxonomicName>
|
||||
<taxonomicName id="08E2A8FD127697F0F4197631A7A9ED64" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Macrochlaeniles)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Macrochlaeniles">Chlaenius Macrochlaeniles</taxonomicName>
|
||||
)
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" baseAuthorityName="Mandl" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Macrochlaeniles) flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="flavofemoratus" subGenus="Macrochlaeniles" subSpecies="enleensis">Chlaenius Macrochlaeniles flavofemoratus ssp. enleensis</taxonomicName>
|
||||
<collectorName>Dr. K.</collectorName>
|
||||
<taxonomicName id="427188F6B5BC092FFAD90C9B17E60508" authorityName="Qin & Germann & Liang" authorityYear="2022" baseAuthorityName="Mandl" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Macrochlaeniles) flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="flavofemoratus" subGenus="Macrochlaeniles" subSpecies="enleensis">Chlaenius Macrochlaeniles flavofemoratus ssp. enleensis</taxonomicName>
|
||||
<collectorName id="6CADE0391BD88BACE01EC37328C9BD8E">Dr. K.</collectorName>
|
||||
MANDL det. 1978/
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
<typeStatus id="FE30C298DFB18181CEF9649145D9CD64">Holotype</typeStatus>
|
||||
[red label].
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">
|
||||
<typeStatus>Paratype</typeStatus>
|
||||
<emphasis id="3E18BE91D07A3E3B3FA3E1C8CBA2BF24" bold="true" italics="true" pageId="0" pageNumber="61">
|
||||
<typeStatus id="A3429FEC108F7F8227A225FC4B0FE5A0">Paratype</typeStatus>
|
||||
</emphasis>
|
||||
(DWC, photo)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation collectingDate="1990-06-20" collectingDateMax="1990-06-28" collectingDateMin="1990-06-20" collectorName="Dr. Blazicek" country="Vietnam" determinerName="Kirschenhofer" location="Tam Dao" municipality="Vietnam" specimenCount="1" specimenCount-male="1" typeStatus="Paratypus">
|
||||
<specimenCount type="male">1 ♂</specimenCount>
|
||||
<materialsCitation id="10453F0FB88D6AF82EE00979596862C1" collectingDate="1990-06-20" collectingDateMax="1990-06-28" collectingDateMin="1990-06-20" collectorName="Dr. Blazicek" country="Vietnam" determinerName="Kirschenhofer" location="Tam Dao" municipality="Vietnam" specimenCount="1" specimenCount-male="1" typeStatus="Paratypus">
|
||||
<specimenCount id="3698B943936D86A598BA4ED49F2430A9" type="male">1 ♂</specimenCount>
|
||||
(IZAS),
|
||||
<collectingMunicipality>Vietnam</collectingMunicipality>
|
||||
<collectingMunicipality id="B481A662A249EBCE9F902EB441B20FC6">Vietnam</collectingMunicipality>
|
||||
,
|
||||
<location LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:23606970FB0FD1139D09CA59241E0566" country="Vietnam" municipality="Vietnam" name="Tam Dao">Tam Dao</location>
|
||||
<location id="23606970FB0FD1139D09CA59241E0566" LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:23606970FB0FD1139D09CA59241E0566" country="Vietnam" municipality="Vietnam" name="Tam Dao">Tam Dao</location>
|
||||
,
|
||||
<collectingDate value="1990-06-20" valueMax="1990-06-28" valueMin="1990-06-20">20-28.VI.1990</collectingDate>
|
||||
<collectingDate id="56F0D20CE0F7F3803E391FEE31ABED87" value="1990-06-20" valueMax="1990-06-28" valueMin="1990-06-20">20-28.VI.1990</collectingDate>
|
||||
,
|
||||
<collectorName>Dr. Blazicek</collectorName>
|
||||
<collectorName id="752C52B27D84171C342F2AB76525143F">Dr. Blazicek</collectorName>
|
||||
lgt./
|
||||
<typeStatus>Paratypus</typeStatus>
|
||||
<typeStatus id="3D49CE8B4FCC446CF4A40710623AE24F">Paratypus</typeStatus>
|
||||
,
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Haplochlaenius) tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis" subGenus="Haplochlaenius">Chlaenius (Haplochlaenius) tamdaoensis</taxonomicName>
|
||||
<taxonomicName id="CB3D701FF12C6A1E0A917B918F9597C2" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Haplochlaenius) tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis" subGenus="Haplochlaenius">Chlaenius (Haplochlaenius) tamdaoensis</taxonomicName>
|
||||
mihi det.
|
||||
<determinerName>Kirschenhofer</determinerName>
|
||||
<determinerName id="F1F540DF9D76B29EEB42FF3C832BEC5A">Kirschenhofer</determinerName>
|
||||
2001[red label]/COLL WRASE, BERLIN
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="May-June, S, Ryabov" country="Vietnam" elevation="900" location="Tam Dao" municipality="Vietnam" specimenCount="1" specimenCount-male="1" stateProvince="Hanoi" typeStatus="paratype">
|
||||
<specimenCount type="male">1 ♂</specimenCount>
|
||||
<materialsCitation id="AC4ADC6D096A90F0FEBCF98B0D6ECF64" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="May-June, S, Ryabov" country="Vietnam" elevation="900" location="Tam Dao" municipality="Vietnam" specimenCount="1" specimenCount-male="1" stateProvince="Hanoi" typeStatus="paratype">
|
||||
<specimenCount id="89085694BF2390CDAAC0BA8192DC52FB" type="male">1 ♂</specimenCount>
|
||||
(IZAS),
|
||||
<collectingMunicipality>Vietnam</collectingMunicipality>
|
||||
<collectingMunicipality id="C6B9ED57A41F2398FA4792CBB15FA424">Vietnam</collectingMunicipality>
|
||||
,
|
||||
<location LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:A8CEDEFD0F38CE176695B42CA7A1E10F" country="Vietnam" municipality="Vietnam" name="Tam Dao" stateProvince="Hanoi">Tam Dao</location>
|
||||
<location id="A8CEDEFD0F38CE176695B42CA7A1E10F" LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:A8CEDEFD0F38CE176695B42CA7A1E10F" country="Vietnam" municipality="Vietnam" name="Tam Dao" stateProvince="Hanoi">Tam Dao</location>
|
||||
,
|
||||
<locationDeviation location="Hanoi">
|
||||
<quantity metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
|
||||
<locationDeviation id="DF687309863EFB9CCA3FEDEAFAD01FE2" location="Hanoi">
|
||||
<quantity id="D4830FF881701BBBFC0AAEF18EBD5F54" metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
|
||||
NW
|
||||
<collectingRegion country="Vietnam" name="Ha Noi">Hanoi</collectingRegion>
|
||||
<collectingRegion id="3CC2CE22D6C8FC8BF78B9B23CF527993" country="Vietnam" name="Ha Noi">Hanoi</collectingRegion>
|
||||
</locationDeviation>
|
||||
,
|
||||
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
|
||||
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
|
||||
<quantity id="F9BB325E4135C8D7D3243ACBE24E60BF" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
|
||||
<elevation id="FA545E397AB087F6F8EB8B4D46FFC714" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
|
||||
</quantity>
|
||||
, 1997
|
||||
<collectorName>May-June, S</collectorName>
|
||||
<collectorName id="080DD78A8CEF93F5FE3D675704C54010">May-June, S</collectorName>
|
||||
.
|
||||
<collectorName>Ryabov</collectorName>
|
||||
<collectorName id="A564F0817B6232728E11BC562FA54D47">Ryabov</collectorName>
|
||||
[internal sac fully everted]
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation collectingDate="2003-05-13" collectorName="S. Ryabov" country="Vietnam" elevation="800" location="Nguen Binh" municipality="Vietnam" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
|
||||
<specimenCount type="male">1 ♂</specimenCount>
|
||||
<materialsCitation id="18E303FA2F9EA20F1B52A620778824A6" collectingDate="2003-05-13" collectorName="S. Ryabov" country="Vietnam" elevation="800" location="Nguen Binh" municipality="Vietnam" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
|
||||
<specimenCount id="F8DDC2484CC5EA85822C66CCFFDB70EF" type="male">1 ♂</specimenCount>
|
||||
(IZAS),
|
||||
<collectingMunicipality>Vietnam</collectingMunicipality>
|
||||
<collectingMunicipality id="0ED928C4661219D862DDBB2C60B959BE">Vietnam</collectingMunicipality>
|
||||
,
|
||||
<collectingRegion country="Vietnam" name="Cao Bang">Cao Bang</collectingRegion>
|
||||
<collectingRegion id="DEE2EC2E4EA101B0819931E0469FAE49" country="Vietnam" name="Cao Bang">Cao Bang</collectingRegion>
|
||||
,
|
||||
<location LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:F4B7FB9FA345742B684B2FC327F0FE82" country="Vietnam" municipality="Vietnam" name="Nguen Binh" stateProvince="Cao Bang">Nguen Binh</location>
|
||||
<location id="F4B7FB9FA345742B684B2FC327F0FE82" LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:F4B7FB9FA345742B684B2FC327F0FE82" country="Vietnam" municipality="Vietnam" name="Nguen Binh" stateProvince="Cao Bang">Nguen Binh</location>
|
||||
,
|
||||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
|
||||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
|
||||
<quantity id="B202DC2FA3DA3B49E93D9CD63DA85706" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
|
||||
<elevation id="921B0B6D320692731B6B2EF0FF0F4C62" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectingDate value="2003-05-13">2003.V.13</collectingDate>
|
||||
<collectingDate id="F5D9202D1298C72F4DABC1BFA48D552A" value="2003-05-13">2003.V.13</collectingDate>
|
||||
,
|
||||
<collectorName>S. Ryabov</collectorName>
|
||||
<collectorName id="F870ED33329DCAD03714B0BE2CD30DCA">S. Ryabov</collectorName>
|
||||
[internal sac partially everted]
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. de Cooman" country="Vietnam" location="Hoa-Binh" municipality="Tonkin" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
|
||||
<specimenCount type="male">1 ♂</specimenCount>
|
||||
<materialsCitation id="A9D1200EA056C96CADF7213B88EAC21A" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. de Cooman" country="Vietnam" location="Hoa-Binh" municipality="Tonkin" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
|
||||
<specimenCount id="3B5567B2CBEEDC9950A40277B556E443" type="male">1 ♂</specimenCount>
|
||||
(IZAS),
|
||||
<collectingMunicipality>Tonkin</collectingMunicipality>
|
||||
<collectingMunicipality id="21691DD6BFAE54C259780EE2D9A346F8">Tonkin</collectingMunicipality>
|
||||
,
|
||||
<location LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:ED1DD5D56279457D2E9B751B09A898F6" country="Vietnam" municipality="Tonkin" name="Hoa-Binh" stateProvince="Cao Bang">Hoa-Binh</location>
|
||||
<location id="ED1DD5D56279457D2E9B751B09A898F6" LSID="urn:lsid:plazi:treatment:33485B2928AD52489B49E1065845E20A:ED1DD5D56279457D2E9B751B09A898F6" country="Vietnam" municipality="Tonkin" name="Hoa-Binh" stateProvince="Cao Bang">Hoa-Binh</location>
|
||||
, leg.
|
||||
<collectorName>A. de Cooman</collectorName>
|
||||
<collectorName id="B90BFA43F5FCF48D1186F9ADE0013443">A. de Cooman</collectorName>
|
||||
[genitalia damaged by dermestes beetle]
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<subSubSection id="86FF12AB5160393AA2200956C1356DB0" pageId="0" pageNumber="61" type="diagnosis">
|
||||
<paragraph id="2A7227699BB7F5A0419C5C8D80FCBCB5" pageId="0" pageNumber="61">Diagnosis.</paragraph>
|
||||
<paragraph id="CD1CE1C4C2469863198CA2FF2A1084AE" pageId="0" pageNumber="61">
|
||||
Pronotum green to coppery. PW/PL = 1.06-1.12; PAW/PBW = 0.82-0.95 (Fig.
|
||||
<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">24</figureCitation>
|
||||
<figureCitation id="76E4AEC32756BF7EAA0F18D234019563" captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">24</figureCitation>
|
||||
); pronotum cordate with anterior angles rounded, not projected forward; disc sparsely punctate and pubescent, with shallow transverse rugosities. Elytral intervals gently convex throughout; densely punctate and pubescent. Hind wings reduced. Metepisterna short; MW/ML = 1.11-1.17 (Fig.
|
||||
<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">16</figureCitation>
|
||||
<figureCitation id="23E9F8E3B068F3FAA393981AC545691A" captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">16</figureCitation>
|
||||
). Apex of femora dark brown or yellow-brown, the rest of legs black.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="description">
|
||||
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">BL = 21.7-24.1 mm, BW = 8.2-8.7 mm. PL = 4.5-4.9 mm, PW = 5.0-5.3 mm, MW = 2.0-2.1 mm, ML = 1.7-1.8 mm. Head, elytra, venter dark and black; pronotum green to coppery; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown; apex of femora dark brown or yellow-brown, the rest of legs black.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Head</emphasis>
|
||||
<subSubSection id="ECFBE02FE6D42E7FABF1853EA1225857" pageId="0" pageNumber="61" type="description">
|
||||
<paragraph id="4B31A0978B5F8E133CFBDE6FD82FC031" pageId="0" pageNumber="61">Description.</paragraph>
|
||||
<paragraph id="EB0B73B78047B7F6AD81AB729C82CF5B" pageId="0" pageNumber="61">BL = 21.7-24.1 mm, BW = 8.2-8.7 mm. PL = 4.5-4.9 mm, PW = 5.0-5.3 mm, MW = 2.0-2.1 mm, ML = 1.7-1.8 mm. Head, elytra, venter dark and black; pronotum green to coppery; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown; apex of femora dark brown or yellow-brown, the rest of legs black.</paragraph>
|
||||
<paragraph id="E6D07D5D228983BFCD16B135E3DF5E22" pageId="0" pageNumber="61">
|
||||
<emphasis id="CAFE1C88C84D0B457A277AFFB397ADFD" bold="true" pageId="0" pageNumber="61">
|
||||
<emphasis id="7B16CCA41FB9F1EC412581360AD12D80" italics="true" pageId="0" pageNumber="61">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Vertex finely punctate, pubescent, without a distinct glabrous area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="2B7CE56DAD68F276AF61441AEB57F4B3" originalValue="×">x</normalizedToken>
|
||||
longer than antennomere 4.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Pronotum</emphasis>
|
||||
<paragraph id="E80D9D4896A6BB5E62D373F8617465BD" pageId="0" pageNumber="61">
|
||||
<emphasis id="8D2228B3C8F155CE96FB76189AFF066E" bold="true" italics="true" pageId="0" pageNumber="61">Pronotum</emphasis>
|
||||
cordiform, PW/PL = 1.06-1.12 (Fig.
|
||||
<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">24</figureCitation>
|
||||
<figureCitation id="CC315474F64011C20BCF11CA7946EE8F" captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">24</figureCitation>
|
||||
), widest at apical third; anterior margin slightly concave, PAW/PBW = 0.82-0.95; lateral margins distinctly narrowed from middle to base, slightly sinuate before posterior angles; anterior angles rounded, not projected forward; posterior angles nearly right angled, rounded at tips; disc gently convex, sparsely punctate and pubescent, with shallow, transverse rugosities, without glabrous area; median line distinct, fine, not reaching anterior margin and base; basal foveae deeply arcuate, punctate and pubescent.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Elytra</emphasis>
|
||||
<paragraph id="39B9953FD48889B6EB04E10BA449EC20" pageId="0" pageNumber="61">
|
||||
<emphasis id="68D8D02468A93FC3E440C7116B187AC0" bold="true" italics="true" pageId="0" pageNumber="61">Elytra</emphasis>
|
||||
elongate, EL/BW = 1.58-2.22, gently convex near anterior third, widest near posterior third, rounded at apex in males; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals gently convex throughout, densely punctate and pubescent; sutural angles sharp at tips; hind wings reduced.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Venter</emphasis>
|
||||
<paragraph id="FF4237E5EF1E3A588291943F7B894B90" pageId="0" pageNumber="61">
|
||||
<emphasis id="825C856D01656D520BAFF825A725A98A" bold="true" italics="true" pageId="0" pageNumber="61">Venter</emphasis>
|
||||
densely punctate, pubescent, metepisterna (Fig.
|
||||
<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">16</figureCitation>
|
||||
<figureCitation id="6B41D97A8558A62DD6B26B097C7C4EAA" captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">16</figureCitation>
|
||||
) short, MW/ML = 1.11-1.17; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males; all abdominal sternites with distinct impressions laterally.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Legs</emphasis>
|
||||
<paragraph id="8DCECC51E22EFD57BEDEAE89C651260A" pageId="0" pageNumber="61">
|
||||
<emphasis id="656C3E0B65D18D0F1A58001B32379BBE" bold="true" italics="true" pageId="0" pageNumber="61">Legs</emphasis>
|
||||
long and slender; tarsi nearly smooth dorsally.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
|
||||
<paragraph id="AA5CA62CCB9D878FBE6A3B3D8DC31D01" pageId="0" pageNumber="61">
|
||||
<emphasis id="0E89B9A8F280583B0F1115BB3C2F5CEA" bold="true" pageId="0" pageNumber="61">
|
||||
<emphasis id="4F2EF51C54373F8254B09498AE1B8B93" italics="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Median lobe (Figs
|
||||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32B-E</figureCitation>
|
||||
<figureCitation id="19E340955AB05BBEE5A31E4FAF5F7EC2" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32B-E</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">33B-E</figureCitation>
|
||||
<figureCitation id="A06FB125FC98789F5CA4A5F00588EC0A" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">33B-E</figureCitation>
|
||||
) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella linear, longer than basal width (Figs
|
||||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32A</figureCitation>
|
||||
<figureCitation id="16BF309A26C93C494472C44C53A6A194" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32A</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">33A</figureCitation>
|
||||
<figureCitation id="59635ED49692DF2E25E5A2A5732AB244" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">33A</figureCitation>
|
||||
), distinctly bent to right side, with a denticulation laterally in the middle, left denticulation distinctly larger than the right one; in left lateral view, apical portion slightly bent dorsally at apex, basal orifice ~ 90° relative to preapical shaft; left paramere larger than right paramere, both helically curved; endophallus (Fig.
|
||||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32F</figureCitation>
|
||||
<figureCitation id="C229D2061BAB97F36EAC9058AA46DF8D" captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32F</figureCitation>
|
||||
) with flagellum helically thick and straight; basal part of flagellum with irregular bursa; apical part of flagellum with triangular sclerite.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61" start="Figures 31–33" startId="F11">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">Figures 31-33.</emphasis>
|
||||
<caption id="18BFD9B5C6E6F77DCE46795892483DF4" doi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61" start="Figures 31–33" startId="F11">
|
||||
<paragraph id="7BC665A195C418B666530B561959C381" pageId="0" pageNumber="61">
|
||||
<emphasis id="CCF39EF241269FB2D8E8CE26CDDB6FC1" bold="true" pageId="0" pageNumber="61">Figures 31-33.</emphasis>
|
||||
Aedeagus of
|
||||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Sphodromimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sphodromimus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Sphodromimus</emphasis>
|
||||
<taxonomicName id="3D8BF07F962033477A3944A540ECBB61" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Sphodromimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodromimus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="6C714DAA4B6976E6DB09E1629C7C1F85" italics="true" pageId="0" pageNumber="61">Sphodromimus</emphasis>
|
||||
</taxonomicName>
|
||||
spp.
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">31</emphasis>
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="davidi" subGenus="Sphodromimus">Chlaenius (Sphodromimus) davidi</taxonomicName>
|
||||
<emphasis id="C09193286A3EE3FE37BAC060381A9203" bold="true" pageId="0" pageNumber="61">31</emphasis>
|
||||
<taxonomicName id="88044FEE00AF359EE11590CC75E542F8" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="davidi" subGenus="Sphodromimus">Chlaenius (Sphodromimus) davidi</taxonomicName>
|
||||
nom. nov. (Guangdong, Xinyi)
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">32</emphasis>
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis" subGenus="Sphodromimus">C. (Sphodromimus) enleensis</taxonomicName>
|
||||
<emphasis id="8819D450A2F7549DE1A3E81F0D155C48" bold="true" pageId="0" pageNumber="61">32</emphasis>
|
||||
<taxonomicName id="2341073E926BCB349C514FD481138D6C" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis" subGenus="Sphodromimus">C. (Sphodromimus) enleensis</taxonomicName>
|
||||
Mandl, 1992 (Vietnam, Tam Dao)
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">33</emphasis>
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis" subGenus="Sphodromimus">C. (Sphodromimus) enleensis</taxonomicName>
|
||||
<emphasis id="BF408FECB908FD288E0B89267F88BC28" bold="true" pageId="0" pageNumber="61">33</emphasis>
|
||||
<taxonomicName id="B36105AB194CD9EBBAB32977644625E7" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis" subGenus="Sphodromimus">C. (Sphodromimus) enleensis</taxonomicName>
|
||||
Mandl, 1992 (holotype, "Indo Chine")
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
<emphasis id="D7505E9C77A2783F53A1CE1E48216F7D" bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
apical lamella
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
<emphasis id="B252A43D58C17C862C3FE25A29CD5B34" bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
left lateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
<emphasis id="1B770D912AC1B87C657FEA3C8EE5EEA4" bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
|
||||
<emphasis id="35BAA1BFF852E37EB596A4916D5D2EB4" bold="true" pageId="0" pageNumber="61">D</emphasis>
|
||||
ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
|
||||
<emphasis id="F14C0FBBB18D0FCDA249D9DDF44F4193" bold="true" pageId="0" pageNumber="61">E</emphasis>
|
||||
right lateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">F</emphasis>
|
||||
<emphasis id="46AFAB1A822BA7534F5182CD02517065" bold="true" pageId="0" pageNumber="61">F</emphasis>
|
||||
endophallus. Left scale bars: 0.5 mm (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
<emphasis id="86105010DF2A2B5E550A6D52EE9A6D0F" bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
); right scale bars: 1.0 mm (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B-F</emphasis>
|
||||
<emphasis id="B6CFAD46C2F8FBC52FF6E747518EB5B6" bold="true" pageId="0" pageNumber="61">B-F</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption doi="10.3897/zookeys.1135.93843.figures34-36" httpUri="https://binary.pensoft.net/fig/780986" pageId="0" pageNumber="61" start="Figures 34–36" startId="F12">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">Figures 34-36.</emphasis>
|
||||
<caption id="C08AEC73C0DBE1FFC4922CD7223FBCC8" doi="10.3897/zookeys.1135.93843.figures34-36" httpUri="https://binary.pensoft.net/fig/780986" pageId="0" pageNumber="61" start="Figures 34–36" startId="F12">
|
||||
<paragraph id="D650E464B43EC61DA0DB4B89B294CFD0" pageId="0" pageNumber="61">
|
||||
<emphasis id="B004A3768BC0CB87C1C13FF6D1377A93" bold="true" pageId="0" pageNumber="61">Figures 34-36.</emphasis>
|
||||
Internal reproductive system of females
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">34A-C</emphasis>
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) caperatus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="caperatus" subGenus="Sphodromimus">Chlaenius (Sphodromimus) caperatus</taxonomicName>
|
||||
<emphasis id="8E1406E6E9A32183AA1914D2A5729AC6" bold="true" pageId="0" pageNumber="61">34A-C</emphasis>
|
||||
<taxonomicName id="BFA79CE34DD408FD93D224ADA892FAF6" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) caperatus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="caperatus" subGenus="Sphodromimus">Chlaenius (Sphodromimus) caperatus</taxonomicName>
|
||||
sp. nov., paratype
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">35A-C</emphasis>
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) hunanus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="hunanus" subGenus="Sphodromimus">C. (Sphodromimus) hunanus</taxonomicName>
|
||||
<emphasis id="8E5BDE93F7E0607AD9FEF9CB8FE20FF3" bold="true" pageId="0" pageNumber="61">35A-C</emphasis>
|
||||
<taxonomicName id="93BDD79CF4E44A2B40F23F852C89CC5B" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. (Sphodromimus) hunanus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="hunanus" subGenus="Sphodromimus">C. (Sphodromimus) hunanus</taxonomicName>
|
||||
(Morvan, 1997)
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">36A-C</emphasis>
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) deuvei" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="deuvei" subGenus="Sphodromimus">C. (Sphodromimus) deuvei</taxonomicName>
|
||||
<emphasis id="C20DB3D7CBF4C4A90B721229B607F7F8" bold="true" pageId="0" pageNumber="61">36A-C</emphasis>
|
||||
<taxonomicName id="01C9F7A9757FA53810DD759BF07639A7" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. (Sphodromimus) deuvei" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="deuvei" subGenus="Sphodromimus">C. (Sphodromimus) deuvei</taxonomicName>
|
||||
(Morvan, 1997)
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
<emphasis id="1B34D7565F5680B76B77EF3B0785E92D" bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
spermatheca
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
<emphasis id="90A1492998252F3627624E05D91B4E7C" bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
<emphasis id="2CC6AB1C4BD391555B929BF13F4D2A36" bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
ventral view. Vertical scale bars: 0.5 mm (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">A)</emphasis>
|
||||
<emphasis id="B663CFD02FF8B79A42ACF82A6C4BC5AB" bold="true" pageId="0" pageNumber="61">A)</emphasis>
|
||||
; horizontal scale bars: 1.0 mm (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B, C</emphasis>
|
||||
<emphasis id="C0F2238761A940D760FE74D25CDB65C8" bold="true" pageId="0" pageNumber="61">B, C</emphasis>
|
||||
)
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption doi="10.3897/zookeys.1135.93843.figures37-39" httpUri="https://binary.pensoft.net/fig/780987" pageId="0" pageNumber="61" start="Figures 37–39" startId="F13">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">Figures 37-39.</emphasis>
|
||||
<caption id="0589B91422948464FA965A567EB532AB" doi="10.3897/zookeys.1135.93843.figures37-39" httpUri="https://binary.pensoft.net/fig/780987" pageId="0" pageNumber="61" start="Figures 37–39" startId="F13">
|
||||
<paragraph id="382171A42B78A26D3BB55AA7B938A8E0" pageId="0" pageNumber="61">
|
||||
<emphasis id="091350BEE2843C3809539CC3771B2E8B" bold="true" pageId="0" pageNumber="61">Figures 37-39.</emphasis>
|
||||
Internal reproductive system of females
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">37</emphasis>
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) yinggelingensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="yinggelingensis" subGenus="Sphodromimus">Chlaenius (Sphodromimus) yinggelingensis</taxonomicName>
|
||||
<emphasis id="7FB0DF58642D1D8FA4C7C9B94C396076" bold="true" pageId="0" pageNumber="61">37</emphasis>
|
||||
<taxonomicName id="FF57F4540A2D4C52BF9CCAD9B7357508" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) yinggelingensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="yinggelingensis" subGenus="Sphodromimus">Chlaenius (Sphodromimus) yinggelingensis</taxonomicName>
|
||||
sp. nov., paratype
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">38</emphasis>
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus" subGenus="Sphodromimus">C. (Sphodromimus) flavofemoratus</taxonomicName>
|
||||
<emphasis id="3092F689CD75679875B4A08CB06AD283" bold="true" pageId="0" pageNumber="61">38</emphasis>
|
||||
<taxonomicName id="F281D35A1B5E81EFC4D234D4540AD16E" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. (Sphodromimus) flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus" subGenus="Sphodromimus">C. (Sphodromimus) flavofemoratus</taxonomicName>
|
||||
Laporte, 1834
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">39</emphasis>
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="davidi" subGenus="Sphodromimus">C. (Sphodromimus) davidi</taxonomicName>
|
||||
<emphasis id="90A7EEBA62742CD5F1D7791C99ADDE3F" bold="true" pageId="0" pageNumber="61">39</emphasis>
|
||||
<taxonomicName id="0A6E87A3AB878A7C15138FC7840D8D19" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="davidi" subGenus="Sphodromimus">C. (Sphodromimus) davidi</taxonomicName>
|
||||
nom. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
<emphasis id="A9C6783B8DD005BA8B54B259B9E286D8" bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
bursa copulatrix
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
<emphasis id="5E39F6BB3C86405C4D8F3A601D55E689" bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
<emphasis id="8D470FC279E70E9A2564C5C1A8DABAA4" bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
ventral view. Scale bars: 1.0 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Female genitalia</emphasis>
|
||||
<paragraph id="3DE5285E7C3A4D7596EB3482B8E042E8" pageId="0" pageNumber="61">
|
||||
<emphasis id="368A68CA8E1D0D94DF226E46E9DE080A" bold="true" italics="true" pageId="0" pageNumber="61">Female genitalia</emphasis>
|
||||
unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<subSubSection id="7883A0CF96518425E0B88F2EB3C5704E" pageId="0" pageNumber="61" type="distribution">
|
||||
<paragraph id="D47A17DFD7D9A933D1BB5FBF001ECFDC" pageId="0" pageNumber="61">Distribution.</paragraph>
|
||||
<paragraph id="DF90CD6CE445DFFA13F0525F934E7300" pageId="0" pageNumber="61">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">40</figureCitation>
|
||||
<figureCitation id="9B8390816B18AE5F20FE2A3150BC8362" captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">40</figureCitation>
|
||||
) Vietnam (Indo Chine). We mark En-le, Yunnan on the map with a question mark
|
||||
<normalizedToken originalValue="‘?’">'?'</normalizedToken>
|
||||
<normalizedToken id="16E9632D112329EE5B3DC77CDB228D1A" originalValue="‘?’">'?'</normalizedToken>
|
||||
.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61" start="Figure 40" startId="F14">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">Figure 40.</emphasis>
|
||||
<caption id="CFE560C430B50AA83C7766EB21FD472B" doi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61" start="Figure 40" startId="F14">
|
||||
<paragraph id="9F580CA26FB1B03C1273219FA6A2DEBE" pageId="0" pageNumber="61">
|
||||
<emphasis id="EE1A677662EE0DE564BCB479FAA58126" bold="true" pageId="0" pageNumber="61">Figure 40.</emphasis>
|
||||
Distribution of
|
||||
<taxonomicName authority="" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Sphodromimus">Chlaenius (Sphodromimus)</taxonomicName>
|
||||
<taxonomicName id="61BA22B57F52C4397CDDFB2C58E58EB3" authority="" authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Sphodromimus">Chlaenius (Sphodromimus)</taxonomicName>
|
||||
species in China and adjacent areas (a possible distribution of
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||||
<taxonomicName id="751080AEA8AE3ED86DF5C92FBE0C9B88" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis">
|
||||
<emphasis id="A88183D3134DE582675C33EF86D3DE5A" italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||||
</taxonomicName>
|
||||
, En-le, Yunnan, represented by a question mark).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="61">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz (1998)</bibRefCitation>
|
||||
<subSubSection id="D1E0126C35DDDD6A9CC5A527D39DD539" pageId="0" pageNumber="61" type="remarks">
|
||||
<paragraph id="37176655BFDCCB7DD146C62C1CB231CB" pageId="0" pageNumber="61">Remarks.</paragraph>
|
||||
<paragraph id="E4A8B91DA7C0D14261861502C6087792" pageId="0" pageNumber="61">
|
||||
<bibRefCitation id="F1FC17E03F262B6B955822C31DDF2831" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz (1998)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz (2005)</bibRefCitation>
|
||||
<bibRefCitation id="46BE1685281780D2EE57E614F8FE8C78" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz (2005)</bibRefCitation>
|
||||
proposed
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="subSpecies" species="flavofemoratus" subSpecies="enleensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. flavofemoratus enleensis</emphasis>
|
||||
<taxonomicName id="C14105BF707AD242EB47ED51196D5E9D" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="flavofemoratus" subSpecies="enleensis">
|
||||
<emphasis id="69811BF6B2C1D239DB95662942A93A77" italics="true" pageId="0" pageNumber="61">C. flavofemoratus enleensis</emphasis>
|
||||
</taxonomicName>
|
||||
Mandl, 1992 as a synonym of
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. flavofemoratus</emphasis>
|
||||
<taxonomicName id="21B9620AB0545F2C39C3FCDD1653BA28" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus">
|
||||
<emphasis id="F85D22736E0CF0D712D922E5FD3D2232" italics="true" pageId="0" pageNumber="61">C. flavofemoratus</emphasis>
|
||||
</taxonomicName>
|
||||
, but
|
||||
<bibRefCitation author="Kirschenhofer, E" editor="Loebl, I" journalOrPublisher="1. 2 nd Edn.). Brill, Leiden, Boston" pageId="0" pageNumber="61" pagination="491 - 497" refId="B15" refString="Kirschenhofer, E, 2017. Tribe Chlaeniini Brulle, 1834. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1. 2nd Edn.). Brill, Leiden, Boston: 491 - 497" title="Tribe Chlaeniini Brulle, 1834." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2017">Kirschenhofer (2017)</bibRefCitation>
|
||||
<bibRefCitation id="30C5E9A559502EA6AD194955CAE69E0E" author="Kirschenhofer, E" editor="Loebl, I" journalOrPublisher="1. 2 nd Edn.). Brill, Leiden, Boston" pageId="0" pageNumber="61" pagination="491 - 497" refId="B15" refString="Kirschenhofer, E, 2017. Tribe Chlaeniini Brulle, 1834. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1. 2nd Edn.). Brill, Leiden, Boston: 491 - 497" title="Tribe Chlaeniini Brulle, 1834." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2017">Kirschenhofer (2017)</bibRefCitation>
|
||||
treated it as a distinct subspecies. Based on its original description, its pronotum longer than
|
||||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. flavofemoratus</emphasis>
|
||||
<taxonomicName id="8AC11B021AD99149FB14AC5159A08052" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus">
|
||||
<emphasis id="F99EA81633ABEBE2BE466DAF8796956F" italics="true" pageId="0" pageNumber="61">C. flavofemoratus</emphasis>
|
||||
</taxonomicName>
|
||||
, and black femora indicated it probably represented a different species. After the examination of the holotype and its dissected genitalia, we find that it has the same apical lamella as
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="tamdaoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. tamdaoensis</emphasis>
|
||||
<taxonomicName id="8A716E7AB6FF71B9FC7A9C22DE3B07D1" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis">
|
||||
<emphasis id="259E0A46A2364E37702B056C15B8B170" italics="true" pageId="0" pageNumber="61">C. tamdaoensis</emphasis>
|
||||
</taxonomicName>
|
||||
. As a consequence of this surprising discovery, the locality should also be critically revised. The labels of the holotype contain two localities (
|
||||
<normalizedToken originalValue="“En-le”">"En-le"</normalizedToken>
|
||||
<normalizedToken id="4A5CECB8E69C58C060BF84EEEEE430E6" originalValue="“En-le”">"En-le"</normalizedToken>
|
||||
and "Indo Chine"), the latter including
|
||||
<normalizedToken originalValue="today’s">today's</normalizedToken>
|
||||
<normalizedToken id="C8733FD6AC219D0C8FA032E0EC991FBB" originalValue="today’s">today's</normalizedToken>
|
||||
Vietnam. Based on the type locality of
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="tamdaoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. tamdaoensis</emphasis>
|
||||
<taxonomicName id="FAC21DA6AFA87BA5B645044E397CFE91" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis">
|
||||
<emphasis id="EC6F096F6F35443793E813E3B2F6C3B0" italics="true" pageId="0" pageNumber="61">C. tamdaoensis</emphasis>
|
||||
</taxonomicName>
|
||||
, it is very unlikely that
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="subSpecies" species="flavofemoratus" subSpecies="enleensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. flavofemoratus enleensis</emphasis>
|
||||
<taxonomicName id="82B1500400934E00A36562B3DEE77375" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. flavofemoratus subsp. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="flavofemoratus" subSpecies="enleensis">
|
||||
<emphasis id="0ACC901C24E39F13F30FA30481888D46" italics="true" pageId="0" pageNumber="61">C. flavofemoratus enleensis</emphasis>
|
||||
</taxonomicName>
|
||||
also occurs in
|
||||
<normalizedToken originalValue="Yunnan’s">Yunnan's</normalizedToken>
|
||||
<normalizedToken id="63A02BF594EFA42ADBC1A9F3F380CC87" originalValue="Yunnan’s">Yunnan's</normalizedToken>
|
||||
En-le. Hence, we think that the label of En-le is likely to be the wrong one and may have been erroneously added, as it rarely happened in the historical collections of the NHMB (e.g.,
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5169.4.6" author="Caldara, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="61" pagination="371 - 380" refId="B7" refString="Caldara, R, Tosevski, I, Mendel, H, Germann, C, 2022. In search of some type-specimens of Rhamphus [Clairville], 1798 (Coleoptera: Curculionidae). Zootaxa 5169 (4): 371 - 380, DOI: https://doi.org/10.11646/zootaxa.5169.4.6" title="In search of some type-specimens of Rhamphus [Clairville], 1798 (Coleoptera: Curculionidae)." url="https://doi.org/10.11646/zootaxa.5169.4.6" volume="5169" year="2022">Caldara et al. 2022</bibRefCitation>
|
||||
<bibRefCitation id="574A0F1C16952DBA003B8A4C7DA8DAB7" DOI="https://doi.org/10.11646/zootaxa.5169.4.6" author="Caldara, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="61" pagination="371 - 380" refId="B7" refString="Caldara, R, Tosevski, I, Mendel, H, Germann, C, 2022. In search of some type-specimens of Rhamphus [Clairville], 1798 (Coleoptera: Curculionidae). Zootaxa 5169 (4): 371 - 380, DOI: https://doi.org/10.11646/zootaxa.5169.4.6" title="In search of some type-specimens of Rhamphus [Clairville], 1798 (Coleoptera: Curculionidae)." url="https://doi.org/10.11646/zootaxa.5169.4.6" volume="5169" year="2022">Caldara et al. 2022</bibRefCitation>
|
||||
). We upgrade
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||||
<taxonomicName id="4D40237552CCBA2ABE4E6E2BBF97CA45" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis">
|
||||
<emphasis id="701EFBDF00A1AA09247116A5475855FA" italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||||
</taxonomicName>
|
||||
as a valid species and consequently treat
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="tamdaoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. tamdaoensis</emphasis>
|
||||
<taxonomicName id="7011828641634FD62109A6A9A8BBC5C3" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. tamdaoensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="tamdaoensis">
|
||||
<emphasis id="DC225456C58B6075F87ABEF165411B25" italics="true" pageId="0" pageNumber="61">C. tamdaoensis</emphasis>
|
||||
</taxonomicName>
|
||||
as synonym of
|
||||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||||
<taxonomicName id="BE2F78A2720B65D7EF2C1D135FCDD7AD" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis">
|
||||
<emphasis id="39CF953BB405A8A643F609FF6828F98B" italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
|
|||
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