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<mods:title>Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species</mods:title>
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<mods:title id="8DFFC1E45C81FF0A85C57A2A51068B4C">Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species</mods:title>
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<mods:namePart>Ranarilalatiana, Tolotra</mods:namePart>
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<mods:namePart>Raveloson Ravaomanarivo, Lala Harivelo</mods:namePart>
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<mods:namePart>Bergsten, Johannes</mods:namePart>
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<treatment ID-GBIF-Taxon="158995547" LSID="urn:lsid:plazi:treatment:C9FE8EA486D557DFA5B1A38A9F232743" httpUri="http://treatment.plazi.org/id/C9FE8EA486D557DFA5B1A38A9F232743" lastPageNumber="19" pageId="0" pageNumber="19">
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<taxonomicName LSID="c9fe8ea4-86d5-57df-a5b1-a38a9f232743" authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Copelatus marginipennis (Laporte, 1835)</taxonomicName>
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||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5A</figureCitation>
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<treatment id="C9FE8EA486D557DFA5B1A38A9F232743" ID-GBIF-Taxon="158995547" LSID="urn:lsid:plazi:treatment:C9FE8EA486D557DFA5B1A38A9F232743" httpUri="http://treatment.plazi.org/id/C9FE8EA486D557DFA5B1A38A9F232743" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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<subSubSection id="0B4BBB17CD30388B3F34EA5D434A1705" pageId="0" pageNumber="19" type="nomenclature">
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<paragraph id="7030B38D42273AF815807B8F75D15969" pageId="0" pageNumber="19">
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<taxonomicName id="9D978EC63A9E92287C896577FBA33B6E" ID-CoL="8LVBF" LSID="c9fe8ea4-86d5-57df-a5b1-a38a9f232743" authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Copelatus marginipennis (Laporte, 1835)</taxonomicName>
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||||
<figureCitation id="76A4E815FC9008E2840BD4E17D3FF046" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5A</figureCitation>
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||||
,
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||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8A</figureCitation>
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||||
<figureCitation id="354463A8C057D80FC5046FD4C16F8D34" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8A</figureCitation>
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
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||||
<paragraph pageId="0" pageNumber="19">
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||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Colymbetes marginipennis</taxonomicName>
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||||
<subSubSection id="304D81FC9973949772755C03CA4645AD" pageId="0" pageNumber="19" type="reference_group">
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||||
<paragraph id="C66E26B201407C151AB0ECF16F73D81D" pageId="0" pageNumber="19">
|
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<taxonomicName id="3AF0F871F7BC9E74BFFE5E1E9FA9796F" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Colymbetes marginipennis</taxonomicName>
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Laporte, 1835: 102.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">Copelatus aldabricus</taxonomicName>
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||||
<paragraph id="CCE098B9FFA1AC3FF913D202CCCF49BA" pageId="0" pageNumber="19">
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||||
<taxonomicName id="5339E026CA7E45C5A15B8A94F8EEABB5" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">Copelatus aldabricus</taxonomicName>
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J. Balfour-Browne, 1950: 368 syn. nov.; TL: Seychelles, Aldabra Islands.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Copelatus aldabricus var. simplex" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="variety" species="aldabricus" variety="simplex">Copelatus aldabricus var. simplex</taxonomicName>
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<bibRefCitation author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot 1952</bibRefCitation>
|
||||
<paragraph id="C6697E34FD3F167755222488044FDBA9" pageId="0" pageNumber="19">
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||||
<taxonomicName id="7C3056FA9B643B4D37AA6D542C5510F4" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Copelatus aldabricus var. simplex" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="variety" species="aldabricus" variety="simplex">Copelatus aldabricus var. simplex</taxonomicName>
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||||
<bibRefCitation id="F3A332DDA4BB226EB798D4588209A9D8" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot 1952</bibRefCitation>
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: 28 syn. nov.; TL: Madagascar.
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</paragraph>
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</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
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||||
<paragraph pageId="0" pageNumber="19">Senegal [possibly mislabelled].</paragraph>
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||||
<subSubSection id="185EC79E76B103A43B358EEDD4C14F9C" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="5C28815C979214C67523141CF21D1AD7" pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph id="3DA560428E5C8A08B8E12C58FE88A88D" pageId="0" pageNumber="19">Senegal [possibly mislabelled].</paragraph>
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</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
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||||
<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="B04CC2F74D79732D6909EA2C4F92BCBB" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="9AA5616F51EE767D2E0B666109C041B7" pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph id="93C0E2A5321570DF90A70F851A6A2D41" pageId="0" pageNumber="19">
|
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housed in Buquet collection and originating from Senegal; of
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<taxonomicName lsidName="aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
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<emphasis italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
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<taxonomicName id="B502DAD74A569C0A14A17CAEC7BAB5AF" lsidName="aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="38FA8E3484D28C1171332DDC968FFC2B" italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
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</taxonomicName>
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: based on male (holotype), J.C. Fryer collection, collected 1908-9 from Aldabra; of
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<taxonomicName lsidName="simplex" pageId="0" pageNumber="19" rank="species" species="simplex">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">simplex</emphasis>
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||||
<taxonomicName id="5F2B2ABD3EB2DE7DDDCF68AB27A805C6" lsidName="simplex" pageId="0" pageNumber="19" rank="species" species="simplex">
|
||||
<emphasis id="29B4C719C3ED278D1D7E4D990D584AD5" italics="true" pageId="0" pageNumber="19">simplex</emphasis>
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</taxonomicName>
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: based on male and female syntypes from Madagascar without further locality data.
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="08C6C7FE15C944ED1ED4B7F9B3F63A26" pageId="0" pageNumber="19" type="type material">
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||||
<paragraph id="0887C859A13CDC1FB1B741EA09D76748" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="53DB1315BF50FF2CEA1998728D471E46" pageId="0" pageNumber="19">
|
||||
-LT♂ (lectotype here designated) (
|
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<emphasis italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
<emphasis id="CE77F6AB050CB836B497AF45A7746FFB" italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
) (MNHN): // Data in NHRS | JLKB | 000065416 //
|
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<emphasis id="AE79EC3C109FFA50281BD889CDA4025B" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
Buquet | pulchellus var.
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
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||||
<normalizedToken id="78CED36A7AF089815781D471D80DF862" originalValue="Aubé">Aube</normalizedToken>
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| h. in Senegal D. Buquet // D. Sharp | Monogr. // Ex-Musaeo
|
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<normalizedToken originalValue="Déjean">Dejean</normalizedToken>
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<normalizedToken id="32163613568080D3FD6611B40A097F05" originalValue="Déjean">Dejean</normalizedToken>
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||||
//
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
<emphasis id="CF1A3DFDD046AD3E2B878B0F093F2736" italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
// Lectotype |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
<emphasis id="479F55808DB37B69242968F2D0DB558C" italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
| Laporte, 1835 | Des. Ranarilalatiana | & Bergsten, 2019// -HT♂ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
<emphasis id="DF8B72E3E6FF32EC65F267967B4AF69A" italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
) (NHMUK): // Aldabra, 08-9. J.C.F. Fryer // Perey Sladen Trust | expedition. | 1913-170. // Type [red round label] //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="3917CDE51E17CAC65E713A0CD5B0B776" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
|
||||
<emphasis id="CAF161D967B65D35B38E5EDFA598D060" italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
|
||||
Type | J. Balfour-Browne det. //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Andilamena, Moramanga</emphasis>
|
||||
<subSubSection id="023CE58822DAA831CF14723D2010F2AD" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="25D3AFE721C6E50D8FEEA58A9ECACE59" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="D3D58AC16D8008391A3B7D561C3E603F" pageId="0" pageNumber="19">
|
||||
<emphasis id="326DB43D2CD9BA2BFF1336B33FD479F0" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Andilamena, Moramanga</emphasis>
|
||||
: -1♀ (NMW): // Data in NHRS | JLKB | 000010726 // Madagascar 17.01.1995 | Ambatombe | nr. Andilamena 900 m | leg. Dunay &
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="54492356B448AC2D6963C9157424B50C" originalValue="Janák">Janak</normalizedToken>
|
||||
(10) // -1♀ (NHMUK): // BMNH-797894 // MAD: AMPA: Moramanga: Andasibe | Andasibe NP: Big Pond | P61BI15: N: -18.937: E:48.416: 940 m | 06/I/2007: Leg. Isambert et al. // DNA Voucher | BMNH <797894> | MSL294:B10 // -2♂ (GP), 15 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010587-8, 10799(Alc.) // Madagascar: Tamatave: Alaotra Mangoro: | Analamazaotra RS; Bas fond, non-permanent | pond near trail to "lac rouge"MAD15-1| 943 m,
|
||||
<geoCoordinate degrees="18" direction="south" minutes="56" orientation="latitude" precision="1" seconds="26.7" value="-18.940748">18°56'26.7"S</geoCoordinate>
|
||||
<geoCoordinate id="98D17BF88F1664DB06C22418D2C4ED22" degrees="18" direction="south" minutes="56" orientation="latitude" precision="1" seconds="26.7" value="-18.940748">18°56'26.7"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="048" direction="east" minutes="25" orientation="longitude" precision="1" seconds="03.9" value="48.41775">048°25'03.9"E</geoCoordinate>
|
||||
<geoCoordinate id="FEA2659B0C7B916B0677F6F07719F101" degrees="048" direction="east" minutes="25" orientation="longitude" precision="1" seconds="03.9" value="48.41775">048°25'03.9"E</geoCoordinate>
|
||||
, 16.III.2015 | Among vegetation and dead leaves in the pond, | Leg. T. Ranarilalatiana & H.J. Randriamihaja // -2♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010604-5, 10800(Alc.) // Madagascar: Tamatave: Alaotra Mangoro: Mantadia NP. | Non-permanent pond at PK18,50 m; E of Park road | 973 m,
|
||||
<geoCoordinate degrees="18" direction="south" minutes="46" orientation="latitude" precision="1" seconds="09.9" value="-18.769417">18°46'09.9"S</geoCoordinate>
|
||||
<geoCoordinate id="D9CFDB4BA14EFE14ABE29374CB6C0EF6" degrees="18" direction="south" minutes="46" orientation="latitude" precision="1" seconds="09.9" value="-18.769417">18°46'09.9"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="048" direction="east" minutes="26" orientation="longitude" precision="1" seconds="10.4" value="48.436222">048°26'10.4"E</geoCoordinate>
|
||||
<geoCoordinate id="CF30BCB56D487B7B30ED9179A5B0923E" degrees="048" direction="east" minutes="26" orientation="longitude" precision="1" seconds="10.4" value="48.436222">048°26'10.4"E</geoCoordinate>
|
||||
, 17.III.2015 | Under dead leaves & vegetation at the edge of the pond, | Leg.T. Ranarilalatiana, H.J. Randriamihaja; MAD15-5 // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010815 // Madagascar: Toamasina: Alaotra | Mangoro: RN2, Mangoro river | 10Km W of Moramanga: S18.92438 | E048.18273, 940 m. 06.XI.2011 | GB Nets and sieves: river and | pools: Field# MAD11-21 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | J.H. Randriamihaja // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010822 // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra SR: | close to park entrance: S18.9355 E | 048.41656, 970 m: 08.XI.2011: GB | Nets and sieves: dried up forest | pond: Field# MAD11-25 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -6 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010823 // Madagascar: Toamasina: Alaotra | Mangoro: by RN2 S border of | Analamazaotra reserve 1Km E | Antsampanana: S
|
||||
<geoCoordinate degrees="18.94987" direction="east" orientation="longitude" precision="1" value="18.94987">18.94987 E</geoCoordinate>
|
||||
<geoCoordinate id="22649880DE5457D2C571896563F76525" degrees="18.94987" direction="east" orientation="longitude" precision="1" value="18.94987">18.94987 E</geoCoordinate>
|
||||
048.42331 | 980m: 09.XI.2011: GB Nets & sieves | ditch next to road: Field# MAD11-29 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | J.H. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Analanjirofo: Maroantsetra</emphasis>
|
||||
<emphasis id="9F11A02F71E04667D246BBA866137CB2" bold="true" pageId="0" pageNumber="19">Toamasina. Analanjirofo: Maroantsetra</emphasis>
|
||||
: -3♂ (GP), 2♀ (NHMUK): // BMNH-797906-10 // MAD: TOAM: Maroantsetra: Masoala | Masoala NP: Pool | P58BI14: N: -15.758: E: 49.993: 10 m| 17/XI/2006: Leg. Isambert et al. // BMNH (DNA Voucher) // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010794 // MAD: TOAM: Maroantsetra: Masoala | Masoala NP: Pool | P58BI14: N: -15.758: E: 49.993: 10 m| 17/XI/2006: Leg. Isambert et al. // -1♂ (NMW): // Data in NHRS | JLKB | 000010728 // E-Madagascar: Fenerive | Foret de Tampolo | 28.12.1998 | leg. J. Moravec // -1♀(Alc.) (NHRS): // NHRS-JLKB | 000011122 // Madagascar: Toamasina: Analajinrofo: | Masoala NP: degraded lowalt. forest: | MAD18-47: small waterpools on path | ~0.5km NW of Marofototra village, | 15.7606S, 49.9926E, 15 m, 17.II.2018 | Leg. T. Ranarilalatiana //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toliara. Menabe: Morondava</emphasis>
|
||||
<emphasis id="DC7F309894A75F343A1DD695D2B75063" bold="true" pageId="0" pageNumber="19">Toliara. Menabe: Morondava</emphasis>
|
||||
: -3♂ (GP) 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010580, 10730 (JB197), 10751, 10802(Alc.) //Madagascar: Toliara: Menabe: | Kirindy RS: S20.07430 | E044.66307, 52 m.a.o. 12.XII.2009 | Water Net, Field# MAD09-46 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="F5EA70CC85CB0F5A6D8DD833BA43165A" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny, Mitsinjo</emphasis>
|
||||
<emphasis id="DCECF5BC65B28C8DD7A0EF0BE811DACB" bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny, Mitsinjo</emphasis>
|
||||
: -1♂ (NHRS): // NHRS-JLKB | 000010734 (JB201) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30350 | E046.81068, 87 m.a.o. 29.XI.2009 | Water Net, Field# MAD09-03 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="74E6DF92972C5642E79159A5FEEB8350" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -6♂ (GP), 3♀, 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010567-71, 10729 (JB196), 10732 (JB199), 10742-3, 10798(Alc.) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="5659AC8B31BCECD06D0739CA25CEC0D7" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♂ (NHRS): // NHRS-JLKB | 000010731 (JB198) // Madagascar: Mahajanga: Boeny | Ankarafantsika NP, S16.30270 | E046.80996; 75 m.a.o. 30.XI.2009 | 22W Black light, Field# MAD09-13 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="AABC492038F645AC8C584BA25E6ADDCE" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010737 (JB191) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.31418 | E046.81731, 30.XI.2009 | Hand picking, Field# MAD09-14 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="B864B763CE886DD6C67B4E9CE35C17FE" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -3♀, 1♂(GP), 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000065749 (JB809), 10738 (JB192), 10744-5, 10806(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.14653 | E045.94926, 9 m.a.o. 04.XII.2009 | Water net, Field# MAD09-24 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="3BE335E5BF7EE56F7D13961DA15EF9F2" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -2♂ (GP), 2♀, 18 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010578-9, 10739 (JB193), 10746, 10807(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.13337 | E045.95778, 19 m.a.o. 04.XII.2009 | Water net, Field# MAD09-25 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="6F951668A3E1DDAC74BF283A224AC2A1" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
| T. Ranarilalatiana, H.J. Randriamihaja // -5♂ (GP), 4♀, 79 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010572-7, 10740 (JB194), 10747-8, 10808 (Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.05776 | E045.80585, 22 m.a.o. 05.XII.2009 | Water net, Field# MAD09-28 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="045ECB2BD82B1BF6AC9ECA64789F0828" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♂, 1♀, 4 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010735 (JB202), 10749, 10809(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.06651 | E045.77672, 24 m.a.o. 05.XII.2009 | Water net, Field# MAD09-29 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="421461A10D4D4CCDF1C12DC9CFA6921D" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010741(JB189) // Madagascar: Mahajanga: Boeny | Mahavavy Kinkony RS, 16.05648S | 045.76371E; 55 m.a.o. 05.XII.2009 | Water net, Field# MAD09-30 | Leg: J. Bergsten, N. Jonsson, T. | Ranarilalatiana, H.J. Randriamihaja // -1♂, 6ex. (Alc.) (NHRS): // NHRS-JLKB | 000010750, 10810(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.01334 | E046.00376, 24 m.a.o. 06.XII.2009 | Water net, Field# MAD09-33 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="85A5D68193B95B3CB0E653CDA078FD74" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
<emphasis id="A997BBD1EA434E489FC36245FFB1B62A" bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
: -2♂ (GP) (CAS): // CASENT-8135015-6 // Madagascar: Mahajanga | Prov. Parc National Tsingy | de Bemaraha, 2.5 Km 62° ENE | Bekopaka, Ankidrodroa river | elev 100 m: 11-15 Nov 2001 //
|
||||
<geoCoordinate degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
<geoCoordinate id="CB2A082E5072E319EB57245B56FB84CB" degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
<geoCoordinate id="4750DD082B4288BB5BBC61C20C83E822" degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
| Coll: Fisher, Griswold et al. | California Acad. of Sciences | sifted litter - tropical dry forest | on Tsingy, code: BLF4340 // -1♂ (GP) (CAS): // CASENT-8131891 // Madagascar: Mahajanga | Prov. Parc National Tsingy | de Bemaraha, 2.5 Km 62° ENE | Bekopaka, Ankidrodroa river | elev 100 m: 11-15 Nov 2001 //
|
||||
<geoCoordinate degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
<geoCoordinate id="2F4B88CC9D5E9EB9084D1929EB6A65A5" degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
<geoCoordinate id="85B3B6EC9357492B5EC74269E7FA11F9" degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
| Coll: Fisher, Griswold et al. | California Acad. of Sciences | at light- tropical dry forest | on Tsingy, code: BLF4343 // -1♂ (GP) (CAS): // CASENT-8135006 // Madagascar Mahajanga | Prov. Foret de Tsimembo | 11.0Km 346° NNW Soatana | elev 50 m: 21-25 Nov 2001 |
|
||||
<geoCoordinate degrees="18" direction="south" minutes="59" orientation="latitude" precision="15" seconds="43" value="-18.995277">18°59'43"S</geoCoordinate>
|
||||
<geoCoordinate id="65222D2A9F0A10F584C501CF6A76FADF" degrees="18" direction="south" minutes="59" orientation="latitude" precision="15" seconds="43" value="-18.995277">18°59'43"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44" direction="east" minutes="26" orientation="longitude" precision="15" seconds="37" value="44.44361">44°26'37"E</geoCoordinate>
|
||||
<geoCoordinate id="66581E341180274907E76987111EB4A1" degrees="44" direction="east" minutes="26" orientation="longitude" precision="15" seconds="37" value="44.44361">44°26'37"E</geoCoordinate>
|
||||
// Coll: Fisher, Griswold et al. | California Acad. of Sciences | sifted litter (leaf mold, rotten wood) | in tropical dry forest | coll. Code: BLF4508 // -1♂ (GP), 2♀, 3 ex. (Alc.), 2♂(Alc.) (NHRS): // NHRS-JLKB | 000010581-2, 10752, 10803(Alc.), 10821(Alc.)// Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="85CF637937359426902609D6C3DF963F" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -3♂ GP, 1♀, 5 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010583-4, 10736 (JB203), 10754, 10805(Alc.) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S18.75643 | E044.71398, 119 m.a.o. 17.XII.2009 | Water Net, Field# MAD09-65 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="8F340692EA1922AD4E86C838C6E29B08" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -1♀, 1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010733 (JB200), 10753, 10804(Alc.) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: 19.03419S | 044.77499E, 41 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-59 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="E2FE18E1DDA3396DEA38DC0E9D9A89AD" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Ambanja, Ambilobe, Antsiranana II</emphasis>
|
||||
<emphasis id="3CD456EB3BDF7FFF8D08900D99E71FCE" bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Ambanja, Ambilobe, Antsiranana II</emphasis>
|
||||
: -1♂ (GP) (NHRS): // NHRS-JLKB | 000010585 // Madagascar: Antsiranana: Diana: | Ambilomagodra: Stream under the | bridge of the road RN6 in | Ambilomagodra village, S13.00780 | E49.13313, 139 m, 30.XI.2012, GB nets: | dried up stream with some pools in | the village: Field# MAD12-33 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010586, 10801(Alc.) // Madagascar: Antsiranana: Diana: | Andrafiabe: Antsoha stream | 200m from Andrafiabe, 12.93022S | 49.03466E, 32 m, 01.XII.2012, GB | nets: stream with some pools: | Field# MAD12-39 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♂ (Alc.) (NHRS): // NHRS-JLKB | 000011123 // Madagascar: Antsiranana: Diana: | Antsaba: 1km W of Antsaba, | 13.63474S; 48.72918E, 67 m, | 28.XI.2012, GB nets: forest | stream: Field# MAD12-32 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♀ (NMW): // Data in NHRS | JLKB | 000010725 // MADAGASCAR: Sakaramy | (M.
|
||||
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
<normalizedToken id="866689F90EA40C812AAB492B6C157F3D" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
, Antsiranana) | lake Farihy Makery | 29.03.2011 | leg. R. Gerecke (MD189) // riparian area near outflow |
|
||||
<geoCoordinate degrees="12" direction="south" minutes="26" orientation="latitude" precision="1" seconds="20.5" value="-12.439028">12°26'20.5"S</geoCoordinate>
|
||||
<geoCoordinate id="195ED4797187BA16C2E1E4EBEE969369" degrees="12" direction="south" minutes="26" orientation="latitude" precision="1" seconds="20.5" value="-12.439028">12°26'20.5"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="49" direction="east" minutes="14" orientation="longitude" precision="1" seconds="20.0" value="49.238888">49°14'20.0"E</geoCoordinate>
|
||||
<geoCoordinate id="30C2216EB6FFE2B317C41C2EC8D5EC2C" degrees="49" direction="east" minutes="14" orientation="longitude" precision="1" seconds="20.0" value="49.238888">49°14'20.0"E</geoCoordinate>
|
||||
| 377 m. 29.5 °C // -1♂ (NMW): // Data in NHRS | JLKB | 000010727 // N. Madagascar | Antseranana distr. | Sambirano riv. | Marovato vill. | 5-12.12.01, leg. J.
|
||||
<normalizedToken originalValue="Horák">Horak</normalizedToken>
|
||||
<normalizedToken id="371C4C88256B4F3B83D62E9FF902D7DE" originalValue="Horák">Horak</normalizedToken>
|
||||
// Antsira
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">nana. Sava: Sambava, Vohemar</emphasis>
|
||||
<emphasis id="360FFBF7E1307F2BDCB12DFB34CE89D2" bold="true" pageId="0" pageNumber="19">nana. Sava: Sambava, Vohemar</emphasis>
|
||||
: -1♀ (NHMUK): // BMNH-797876 // MAD: DIEG: Sambava: Marojejy | Marojejy NP: Pool | P57BI31: N: -14.457: E:49.79: 162m | 10/XI/2006; Leg. Isambert et al // DNA Voucher | BMNH <797876> | MSL294:A4 // -1♀ (CAS): // CASENT-8135007 // Madagascar Antsiranana | Foret
|
||||
<normalizedToken originalValue="d’Ampondrabe">d'Ampondrabe</normalizedToken>
|
||||
<normalizedToken id="F3CD69004472B38303F8DAA0CD583460" originalValue="d’Ampondrabe">d'Ampondrabe</normalizedToken>
|
||||
. | 26.3Km 10° NNE Daraina | elev 175m: 10 December 2003 //
|
||||
<geoCoordinate degrees="12" direction="south" minutes="58" orientation="latitude" precision="15" seconds="12" value="-12.969999">12°58'12"S</geoCoordinate>
|
||||
<geoCoordinate id="74F7732FC7FBE96D730899270EFF7281" degrees="12" direction="south" minutes="58" orientation="latitude" precision="15" seconds="12" value="-12.969999">12°58'12"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="049" direction="east" minutes="42" orientation="longitude" precision="15" seconds="00" value="49.7">049°42'00"E</geoCoordinate>
|
||||
<geoCoordinate id="67A03CC32C3650F2122F750BF7746B94" degrees="049" direction="east" minutes="42" orientation="longitude" precision="15" seconds="00" value="49.7">049°42'00"E</geoCoordinate>
|
||||
| California Acad. of Sciences | Coll: B.L.Fisher, sifted | litter (leaf mold, rotten wood) | tropical dry forest, BLF9974 //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<subSubSection id="A5278385F472E16C6B51B06059BFF6A1" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="D93265DFF2E87881B2A22209B6EC223C" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="807E77F728D547B1E56C050B34F06C65" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="4D14843D9E6E9B3F733E980BB1E8F1DA" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="2EAE1BDED6C48836C371EC8CBD305993" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from all other Malagasy
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="10F26C9A9ADDA3792107C7D111EE1987" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="CDB9F027812A0EFC3A8A98B4B9A2BBE2" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
except
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="46AA5BDB5F433D53BCD818B3CE0346FB" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="EDAB9F8C18EF63A738B9E661AD4D7389" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="FF2B27544466F15D5A488EAE6E6B527E" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="0BC5B19470AD244333FE099F30657272" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
by the presence of six discal elytral striae and a broadly oval body shape.
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<taxonomicName id="4CEA8D880BACE33A7A90F06832FC7990" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="E55ADC77273E2D204EAA9E002FFE74C1" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is most easily separated by the distinct shape of the male penis in lateral view (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
<figureCitation id="49802240D97C85BDD827A86E79D4869F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
).
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="3C049F12CDF06C1143041719079E3132" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="0FD4004696346D4B7F6D138A5E262B1B" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
(from Madagascar) lacks a transverse testaceous band basally on the elytra and females of
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="119A4C18D1D7197BF0BCF64B3FB70EC6" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="8C552C78095F946DA6E4F9DCA2E47A13" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
can usually be distinguished from
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="41396725A68A7EB63722EAFC390178B6" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="859C3E614D14503B2D845D8A2AFD2DAF" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
by the normally narrower testaceous band, a longer first elytral stria, and a broader body shape.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="97D665E1C9FF1350B0A37D63DD4611AE" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="62EBF1261A1ED5E6EA96408BBC0D33EC" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="FE18C026E615A57FE322D5F86ECA606A" pageId="0" pageNumber="19">
|
||||
Body length 5.2-6.6 mm. Body shape oval, rather convex and attenuate posteriorly, dark brown to brown ferrugineous. Head, pronotum and elytra in the same dark brown ferrugineous, covered with fine dense punctation. Lateral sides of pronotum more brownish, with short strioles and the widest striolate area in the posterior corners. Elytra dark brown to brown ferrugineous, with a testaceous transverse band at base (
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8A</figureCitation>
|
||||
<figureCitation id="4D3958076ABB90BF13A7F159FE6A964F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8A</figureCitation>
|
||||
). Six clearly impressed elytral striae present and one submarginal: first stria abbreviated and slightly shorter, starting at approx. 1/7th posterior of elytral base, second to sixth full length, and submarginal stria starting at approx. the middle; all striae approaching the apex of elytron except the sixth, which is abbreviated posteriorly and a little shorter. Sometimes, the first and second elytral striae, like fifth and sixth, unite posteriorly. Elytral surface covered with dense punctures.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Ventral side brownish to ferrugineous, metacoxa with microsculpture, densely and finely punctate. Metacoxa and abdominal ventrites striolate. Prosternal process rather short and spear-shaped, medially only weakly raised and rounded. Lateral parts of metaventrites rather broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae and palps both in the same brown colour. Pro- and mesothoracic legs brown ferrugineous. Metathoracic legs dark brown ferrugineous.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="D2719AAF67CE9384D0A02917E3DF8693" pageId="0" pageNumber="19">Ventral side brownish to ferrugineous, metacoxa with microsculpture, densely and finely punctate. Metacoxa and abdominal ventrites striolate. Prosternal process rather short and spear-shaped, medially only weakly raised and rounded. Lateral parts of metaventrites rather broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae and palps both in the same brown colour. Pro- and mesothoracic legs brown ferrugineous. Metathoracic legs dark brown ferrugineous.</paragraph>
|
||||
<paragraph id="D2875DF7433FE87BB515D04C40475829" pageId="0" pageNumber="19">
|
||||
Male: Protibia strongly angled basally and expanded in apical two thirds. Penis in ventral view with a small preapical tooth on right side; in lateral view very characteristic with a subbasal dorsal knob, and post-middle with a deep ventral invagination (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
<figureCitation id="9452F1FCE33EAC66D2BC57CD18068D47" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Females from Madagascar usually with elytral striolation rather weak and restricted to outer intervals, but rarely the elytra are entirely and distincly striolate. Females on average smaller than males.</paragraph>
|
||||
<paragraph id="8821ABB848C48DD7FDCBFF1DCA63069C" pageId="0" pageNumber="19">Females from Madagascar usually with elytral striolation rather weak and restricted to outer intervals, but rarely the elytra are entirely and distincly striolate. Females on average smaller than males.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="925BA09FCB74D22D13210D77727451EA" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="99F52898CF6BA7770870DDC4C23BB043" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="99AF1F00C286F5F511E6F116F3B8B08B" pageId="0" pageNumber="19">
|
||||
Endemic to the western Indian Ocean islands as far as is modernly known, but the 1835 type locality of the original description and labels read
|
||||
<normalizedToken originalValue="“Senegal”">"Senegal"</normalizedToken>
|
||||
<normalizedToken id="4A0DB330189D26E358E178B6FB9C5A5D" originalValue="“Senegal”">"Senegal"</normalizedToken>
|
||||
, which indicates either mislabeling or that the species in fact also occurs on continental Africa. Known from Madagascar, Reunion, Comores, Seychelles, and Aldabra Island (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
<bibRefCitation id="EB616DEA0BCC0D2480B5DDEBF7D48ABE" pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
). Widespread in Madagascar and recorded from Montagne
|
||||
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
<normalizedToken id="35F5BC8A22ACC0D57A111B8462FFC8C6" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
NP and Masoala NP in the north and northeast to Kirindy in the southwest (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
|
||||
<figureCitation id="528A6C3DBA15E7D760F3951BAD8CFA2C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
|
||||
), but apparently lacking from most of the central plateau. It can probably show up anywhere around the island below 1000 m in altitude.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="9B6DA779913403A8509332E9D9CC5A67" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="DBB001FCE0AFE5824C5EA9261F2EACAC" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="C19B39AEC996E30E65595A22398DDED3" pageId="0" pageNumber="19">
|
||||
This is a lowland species that seems to be most common in the deciduous western parts of Madagascar. It was common in the newly designated protected areas of Mahavavy Kinkony when we visited it in 2009. In the deciduous forest biome it has been recorded from Kirindy in the south to Ankarana NP in the north. The species seems to be a generalist and can as well show up on lowland humid east coast and at midaltitudes up to at least 1000 m. It is an apt flier collected with light traps and often in very temporary and small shallow pools including water-filled wheel tracks. Found in all kinds of temporary pools, as well as in streams and in residual pools in dried-up riverbeds. It occurred sympatrically with
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="FBBA20B14E8105E965C17C67107EA856" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="76D605315FEAFB319E330EB68B7F9CB8" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
in Betsabora river near Moramanga and at a 22W black light trap by a forest pool in Ankarafantsika NP.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
<subSubSection id="05068E3C3D362BEFE0BB0525254D72A3" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="CC731E42CECD27673ED5E5D877AE3FAC" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="B602CEB0976FFA5241375EC86CC7674B" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="604005BA75D1786CBE353FE1B242AC6A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis id="A88E9A0B59CA4F1698A62246BF738169" italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
was described by J.
|
||||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
<bibRefCitation id="D52A280BBE625EB7034642E02824AB57" author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
from Aldabra Island in the Seychelles.
|
||||
<bibRefCitation author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot (1952)</bibRefCitation>
|
||||
<bibRefCitation id="F2B42BC0C6C497353871F529F389FC76" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot (1952)</bibRefCitation>
|
||||
named a non-striolated female variety from Madagascar as var.
|
||||
<taxonomicName lsidName="var. simplex" pageId="0" pageNumber="19" rank="variety" variety="simplex">simplex</taxonomicName>
|
||||
<taxonomicName id="6320DB8B53B6AFC89E99AB3F09CF059B" lsidName="var. simplex" pageId="0" pageNumber="19" rank="variety" variety="simplex">simplex</taxonomicName>
|
||||
, and again in 1961 as var.
|
||||
<taxonomicName lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
<taxonomicName id="01055AEF5E9A68786C10E2BB1AA79C98" lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
(non-available infrasubspecific name) from Andranofotsy (E of Maroantsetra NE Madagascar), and stated (1961) that females of
|
||||
<taxonomicName lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
<taxonomicName id="E0AFB741120AB49700C34B48671AE951" lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="6897791BEF6ED4C97039D768C70AC4D6" italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
are normally striolated, but that on Madagascar the non-striolated var.
|
||||
<taxonomicName lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
<taxonomicName id="7202BF3258DC05BFC021BC6AC5D6EF39" lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
is most common or even ubiquitous. We have found at least one female on Madagascar (from Masoala NP) with the elytra entirely striolated (the
|
||||
<normalizedToken originalValue="“normal”">"normal"</normalizedToken>
|
||||
<normalizedToken id="C67781B249DEB4A091F5753FE653D09E" originalValue="“normal”">"normal"</normalizedToken>
|
||||
variety sensu
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
<bibRefCitation id="760BF322108614A1F77BFFD58181A600" pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
), but indeed the form where striolation is rather weak and only at outer elytral intervals is most common.
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">
|
||||
<normalizedToken originalValue="Guignot’s">Guignot's</normalizedToken>
|
||||
<bibRefCitation id="FDCA43709D0507432D091FB2E421EDBF" pageId="0" pageNumber="19" refId="B36">
|
||||
<normalizedToken id="F2FFC31FB7CA2FBBB3A943B7C98DA5F7" originalValue="Guignot’s">Guignot's</normalizedToken>
|
||||
(1961)
|
||||
</bibRefCitation>
|
||||
distinction between female forms either having or lacking strioles is, however, a simplification. All females we have seen have at least some weak tendencies of striolation on outer intervals and the extent can vary significantly between individuals.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<paragraph id="EACA3B26F522D91E4FE4E85D4C9F714D" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="209DE82B900B99FCE9D6A0520840FF48" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="FD35C807FCAB41D06ABCEA91D4987F40" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(Laporte, 1835) has been treated as a junior synonym of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="E1B64A0D43518FE1DE4D3F0C6A1EE2A3" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="ACC937B3BD55CA36C5B0868E5F8CCF53" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
Klug, 1834 since
|
||||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.59232" author="Aube, C" journalOrPublisher="Mequignon Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B2" refString="Aube, C, 1838. Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean. . Mequignon Pere et Fils, Paris" title="Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean." url="https://doi.org/10.5962/bhl.title.59232" year="1838">
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<bibRefCitation id="099505FB880E15E8DEEC8BF3B34644C6" DOI="https://doi.org/10.5962/bhl.title.59232" author="Aube, C" journalOrPublisher="Mequignon Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B2" refString="Aube, C, 1838. Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean. . Mequignon Pere et Fils, Paris" title="Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean." url="https://doi.org/10.5962/bhl.title.59232" year="1838">
|
||||
<normalizedToken id="049E4FC64F5F7C7FF44E6854AB8FAA69" originalValue="Aubé">Aube</normalizedToken>
|
||||
(1838)
|
||||
</bibRefCitation>
|
||||
. We discovered the type material of
|
||||
<normalizedToken originalValue="Laporte´s">Laporte's</normalizedToken>
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<normalizedToken id="67FE8665A3513315B47D81B2AA34A96C" originalValue="Laporte´s">Laporte's</normalizedToken>
|
||||
<taxonomicName id="39307F5FA5AC13C3B148AB2C428AED3A" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="E9C1799F5FE432BD7EEA503124A50A6A" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
in MNHN, Paris. This could be identified based on a folded blue label bearing the information "
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<taxonomicName id="7300DBA999DEFD0BACC4DDC5961F0053" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="95193FC12E03756C976F4EFF525C6EE9" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Buquet,
|
||||
<taxonomicName lsidName="pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
<taxonomicName id="5E8E15DD9F4635CF34AB30A8D20C7CE3" lsidName="pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="764A819E76E1B9792797BA06A105C6A6" italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
var
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<normalizedToken id="9A11627127FA3FDBED40A396C715FB1A" originalValue="Aubé">Aube</normalizedToken>
|
||||
, h. in Senegal, D. Buquet", which fits perfectly the description by
|
||||
<bibRefCitation author="Laporte, FLN" journalOrPublisher="Mequignon-Marvis Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B44" refString="Laporte, FLN, 1835. Caumont de 1834-1835: Etudes entomologiques. Premiere partie. Carnassiers. . Mequignon-Marvis Pere et Fils, Paris" title="Caumont de 1834 - 1835: Etudes entomologiques. Premiere partie. Carnassiers." year="1835">Laporte (1835)</bibRefCitation>
|
||||
<bibRefCitation id="7AC3E78DF52522E65ACA6B655A9DDF4B" author="Laporte, FLN" journalOrPublisher="Mequignon-Marvis Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B44" refString="Laporte, FLN, 1835. Caumont de 1834-1835: Etudes entomologiques. Premiere partie. Carnassiers. . Mequignon-Marvis Pere et Fils, Paris" title="Caumont de 1834 - 1835: Etudes entomologiques. Premiere partie. Carnassiers." year="1835">Laporte (1835)</bibRefCitation>
|
||||
. The genitalia of this male show without any doubt that it is a synonym of the younger name
|
||||
<taxonomicName lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
<taxonomicName id="1A0DD97D3CD3428B24D6AEC8FFE9190C" lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="A05034EE32E6C75CB5524F6C87DE469C" italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
and not of
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="DAEFDDE55C776EFC8029B7FAD09A23B6" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="30D6942F77EE06BCDF0B8925BDE4A27B" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
. The name
|
||||
<taxonomicName lsidName="marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">marginipennis</emphasis>
|
||||
<taxonomicName id="C825EA8D896BD65D52CD4D33628F54D2" lsidName="marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="1A7992EBA78AF6AD2DE6593AF00E4CEB" italics="true" pageId="0" pageNumber="19">marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
has to our knowledge not been used as a valid name after 1899, fulfilling the first of the two criteria for protecting a younger currently used synonym from an older available name (ICZN 23.9.1.1). However,
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
<taxonomicName id="2F0491F8DD3EE5F327FD2604B78EF7CA" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis id="C6BA2CB1E03EA15E89543863BFB4D72C" italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
is a relatively recent name from a region with a low-intensity taxonomic research and we do not believe it fulfils the second criteria of at least 25 papers published by at least ten authors in the immediately preceding 50 years and encompassing a span of not less than ten years (ICZN 23.9.1.2). Therefore it is not justifiable to protect the younger synonym
|
||||
<taxonomicName lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
<taxonomicName id="69B553B49CC3EB40A2A30670D015546A" lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="3A189CA69D689DA6361B60D72F66A84B" italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
and the older
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="79793047C0461D7254A33E3D8E0F13D6" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="ED7C7F59CF79685C47803294323E2B4E" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is therefore brought back as the valid name of this species. We designate the discovered syntype of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
<taxonomicName id="8B2C89E6A6A70BD9A882FA3C94E4270E" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="67D3B2173C51AFED1BE177027B516BB4" italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Laporte, 1835 in the MNHN collection as lectotype to preserve stability of the name. At the same time as this type specimen was discovered, also a likely
|
||||
<normalizedToken originalValue="“type”">"type"</normalizedToken>
|
||||
<normalizedToken id="CCA92B32A6C25541775A2C48643998AF" originalValue="“type”">"type"</normalizedToken>
|
||||
specimen of the nomen nudum
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus fimbriolatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriolatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus fimbriolatus</emphasis>
|
||||
<taxonomicName id="53DC810DA1BE766FCEBB4C90CD393F49" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus fimbriolatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriolatus">
|
||||
<emphasis id="8FA78D637B32EF632D0747C93F5D5F1D" italics="true" pageId="0" pageNumber="19">Copelatus fimbriolatus</emphasis>
|
||||
</taxonomicName>
|
||||
Dejean, 1837 from "Ile de France" [= Mauritius] was found with the same type of blue folded label (see supplementary file 1: NHRS-JLKB000065420). This nomen nudum is currently considered a synonym of
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="5B07E8C4A4934621100368EF9B2F8E33" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="E565493D9302D6A75261BB145B0E1594" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
Klug (Nilsson and Hajek 2018). Unfortunately, it is a female specimen, which is why its identity cannot be established with certainty, but it is more likely identical with
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="84DB7ADD31A4C5055F8620DF54A2C62F" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="FD98A15ED88DF2C14349CFD0B93D29C9" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
since
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="DC754C35892516A42D289FFCF76C5A98" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="3B2BB97A0CC29615593E99EDF92D7150" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
is not yet known from any of the Mascarene Islands (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
<bibRefCitation id="C9AD417EBAF400A89229AE9B54CFECE5" pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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