diff --git a/data/01/49/66/0149666EE3E15D2FA4E180FD888742E5.xml b/data/01/49/66/0149666EE3E15D2FA4E180FD888742E5.xml index 726fe452d9f..f2a84ff8078 100644 --- a/data/01/49/66/0149666EE3E15D2FA4E180FD888742E5.xml +++ b/data/01/49/66/0149666EE3E15D2FA4E180FD888742E5.xml @@ -1,510 +1,510 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Copelatus pulchellus (Klug, 1834) -Figs 5C + + + +Copelatus pulchellus (Klug, 1834) +Figs 5C , -8D +8D - - -Agabus pulchellus -Klug 1834 + + +Agabus pulchellus +Klug 1834 : t. XXXIII: 7 - -Copelatus africanus + +Copelatus africanus Sharp, 1882: 583; TL: Botswana, Lake Ngami; - + ? -Copelatus basalis +Copelatus basalis Boheman, 1848: 244; TL: South Africa (Caffraria interiore); - -Copelatus discoideus + +Copelatus discoideus Sharp, 1882: 582; TL: Mesopotamia; - -Copelatus obtusus + +Copelatus obtusus Boheman, 1848: 242; TL: South Africa (Caffraria orientali); - -Copelatus strigulosus + +Copelatus strigulosus Sharp, 1882: 582; TL: Mesopotamia; - -Copelatus mimetes + +Copelatus mimetes Guignot, 1957: 73; TL: Madagascar, Sakaraha, Lambomakandro; syn. nov. - -Type locality. -Egypt, Sinai. + +Type locality. +Egypt, Sinai. - -Type material studied. - + +Type material studied. + -HT♂ (GP) ( -Copelatus mimetes +Copelatus mimetes ) (MNHN, "coll. Guignot"): // Data in NHRS | JLKB | 000030032 // Sakaraha | Lambomakandro | III-56 A.R. // Type [red label] // F. Guignot Det., 1956 | -Copelatus +Copelatus | -mimetes +mimetes n. sp.| Type ♂ // INSTITUT SCIENTIFIQUE MADAGASCAR // -LT♀ (lectotype here designated) ( -Copelatus obtusus +Copelatus obtusus ) (NHRS): // Caffra | ria. // J. Wahlb // Type. // HoloTypus [red label, -"Typus" +"Typus" printed, -"Holo" +"Holo" handwritten with small letters in front, possibly by J. Balfour-Browne who studied the type] // obtusus Boh. // -Cop. pulchellus +Cop. pulchellus var: | -obtusus +obtusus Boh. | J. Balfour-Browne det. // 5657 | E91 // NHRS-JLKB | 000065335 // Lectotype | -Copelatus obtusus +Copelatus obtusus | Boheman, 1842 | Des. Ranarilalatiana | & Bergsten, 2019// -? ST♂ ( -Copelatus basalis +Copelatus basalis ) (NHRS): // Caffra | ria. // J Wahlb // Type. // Typus [red label, printed] // 158 | 61 // Copelatus | basalis Bhn. // -Copelatus +Copelatus | -pulchellus +pulchellus | Klug. | Det. 19.iv.1961 | J. Omer-Cooper. // 5597 | E91 // NHRS-JLKB | 000065337 // -? ST♀ ( -Copelatus basalis +Copelatus basalis ) (NHRS): // Caffra | ria. // J. Wahlb // Paratypus [red label, printed] // 160 | 61 // 5596 | E91 // NHRS-JLKB | 000065338 // -? ST♀ ( -Copelatus basalis +Copelatus basalis ) (NHRS): // Caffra | ria. // J. Wahlb // Paratypus [red label, printed] // 159 | 61 // 5595 | E91 // NHRS-JLKB | 000065339 // - -Additional material studied. - -Antsiranana. Diana: Antsiranana + +Additional material studied. + +Antsiranana. Diana: Antsiranana : -1♂(GP) (MNHN): // Data in NHRS | JLKB | 000030259 // Madagascar | Diego-Suarez | Ch. Alluaud 1893 // -Mahajanga. Boeny: Mitsinjo +Mahajanga. Boeny: Mitsinjo : -1♀ (NHRS): // NHRS-JLKB | 000065733 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS: S 16.13337 | E 045.95778, 19 m.a.o. 04.XII.2009 | Water Net, Field# MAD09-25 | leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -Mahajanga. Melaky: Antsalova +Mahajanga. Melaky: Antsalova : -1♂(GP) (NHRS): // NHRS-JLKB | 000010695 // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg. J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // - -Diagnosis. - + +Diagnosis. + Similar to - -C. marginipennis + +C. marginipennis (Laporte, 1835) and - -C. mahajanga + +C. mahajanga in overall habitus, but body shape more like the elongated shape of - -C. mahajanga + +C. mahajanga . - -Copelatus pulchellus + +Copelatus pulchellus on Madagascar is ferrugineous black in overall colouration with or without a rather narrow or vague testaceous band basally on elytra ( -Fig. 8D +Fig. 8D ). Instead testaceous regions on elytra are limited to posterolateral and apical parts. The first elytral stria almost full length. Penis in lateral view rather abruptly curved past middle so that basal and apical parts are approximately at right angles ( -Fig. 5C +Fig. 5C ). Penis overall simpler compared to that of - -C. marginipennis + +C. marginipennis and - -C. mahajanga + +C. mahajanga and apex slightly twisted. - -Description. -(based on Malagasy specimens): -Body length 5.5-6.1 mm. Body shape oblong oval, rather convex and attenuate posteriorly, dark brown to blackish ferrugineous. Head infuscated brown ferrugineous, somewhat lighter posteriorly, covered with dense microreticulation and sparser punctation. -Pronotum dark brown to ferrugineous black with testaceous anterolateral corners. Disc covered with fine microsculpture forming regular cells and regularily spread small punctures of about same size as cells. Punctuation becomes coarser in posterolateral corners with a weak tendency to corrugate. - + +Description. +(based on Malagasy specimens): +Body length 5.5-6.1 mm. Body shape oblong oval, rather convex and attenuate posteriorly, dark brown to blackish ferrugineous. Head infuscated brown ferrugineous, somewhat lighter posteriorly, covered with dense microreticulation and sparser punctation. +Pronotum dark brown to ferrugineous black with testaceous anterolateral corners. Disc covered with fine microsculpture forming regular cells and regularily spread small punctures of about same size as cells. Punctuation becomes coarser in posterolateral corners with a weak tendency to corrugate. + Elytra predominantely dark brown to ferrugineous black on disc and along striae with or without a rather narrow and vague basal testaceous band ( -Fig. 8D +Fig. 8D ). Outer intervals and apical region testaceous to varying degrees, often a testaceous interval 4 stands out. Elytral surface covered with same type of microsculpture and punctures as on pronotum. Six clearly impressed elytral striae present on disc and one submarginal stria: second to sixth stria starting at base, first stria somewhat abbreviated anteriorly (by approximately 1/8th the length of elytra); submarginal stria starting at approx. middle. - + Ventral side ferrugineous dark brown, with testaceous spots laterally on abdominal ventrites. Metacoxa and ventrites with strioles. Prosternal process more elongate lanceolate and with blunter apex compared with - -C. marginipennis + +C. marginipennis and - -C. mahajanga + +C. mahajanga . Lateral parts of metaventrite medium broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae, palps, pro- and mesolegs testaceous, metalegs somewhat darker testaceous. - + Female: elytral striolation limited to the medial parts of the outer three elytral intervals in the single female studied from Madagascar. From other parts of the distribution a female form is known that has the entire elytra striolated ( -Sharp 1882 +Sharp 1882 ; - -Hajek + +Hajek and Reiter 2014 ). - + Male: protibia bisinuate and angled at base, distally expanded. Penis thin, strongly angled at middle in lateral view, and apex somewhat twisted to the left in ventral view ( -Fig. 5C +Fig. 5C ). Parameres more narrowly triangular apically compared to those in - -C. marginipennis + +C. marginipennis and - -C. mahajanga + +C. mahajanga . - - + + Figure 8. Habitus, dorsal view. -A +A Male: - -Copelatus marginipennis + +Copelatus marginipennis -B +B Female: - -Copelatus mahajanga + +Copelatus mahajanga -C +C Female: - -Copelatus + +Copelatus sp. 1 (Bemaraha) -D +D Male: - -Copelatus pulchellus + +Copelatus pulchellus . - -Distribution. - + +Distribution. + As the species - -C. pulchellus + +C. pulchellus is currently interpreted, this is a very widely distributed Afrotropical and Middle Eastern species. -Balfour-Browne (1950) +Balfour-Browne (1950) recorded the species from Senegal in the west, Tanzania in the east, South Africa in the south, and Mesopotamia (Iraq and adjacent regions) in the Middle East. - -Hajek + +Hajek and Reiter (2014) recorded the species from Yemen and stated that the likely distribution in the Middle East included the entire Arabian Peninsula. -Guignot (1961) +Guignot (1961) also gives India and Ceylon (= Sri Lanka) but this was likely based on the misidentification of closely related oriental species (see -Ghosh and Nilsson 2012 +Ghosh and Nilsson 2012 ; -Sheth et al. 2018 +Sheth et al. 2018 ). - -Copelatus pulchellus + +Copelatus pulchellus was not previously recorded from Madagascar following the revision by -Balfour-Browne (1950) +Balfour-Browne (1950) but as we synonymise - -C. mimetes + +C. mimetes with - -C. pulchellus + +C. pulchellus , Madagascar now forms part of the distribution. From Madagascar we have seen specimens from Antsiranana, Mahavavy Kinkony Reserve, Tsingy de Bemaraha National Park (Bekopaka), and from Lambomakandro, Sakaraha ( -Fig. 11C +Fig. 11C ). It can likely show up anywhere in lowland Madagascar, but especially in the western lowlands. - -Habitat and ecology. - + +Habitat and ecology. + On Madagascar we have collected the species associated with a small forest stream with sidepools in a karstic limestone area ( -"tsingy" +"tsingy" ) and in a muddy stagnant pool in a dried-out river bed. Both localities are in dry deciduous forests of lowland western Madagascar. - -Comment. - - -Copelatus pulchellus + +Comment. + + +Copelatus pulchellus forms part of a diverse species group with many externally very similar species. -Balfour-Browne (1950) +Balfour-Browne (1950) admitted that his previous treatment of the species ( -Balfour-Browne 1939 +Balfour-Browne 1939 ) was entirely wrong as he had then not studied the male genitalia. When he did so in 1950 it resulted in the description of several new species previously lumped under - -C. pulchellus + +C. pulchellus . - - -Copelatus pulchellus + + +Copelatus pulchellus is now interpreted as a widespread Afrotropical and Middle Eastern species with the male penis similar to that illustrated in -Figure 5C +Figure 5C . Dorsal colouration is interpreted as very variable, even consisting of several distinct colour forms such as the darker forms described as - -C. obtusus + +C. obtusus Boheman and later as - -C. africanus + +C. africanus Sharp (compare figure 24 with photographs in -Perissinotto et al. (2016 +Perissinotto et al. (2016 : fig. 24) and - -Hajek + +Hajek and Reiter (2014 : fig. 30)). Females are also interpreted as variable in the striolation pattern on the elytra. Sharp described a female form from Mesopotamia that was entirely striolated over the elytra under the name - -C. strigulosus + +C. strigulosus , which is interpreted today as intraspecific variation of - -C. pulchellus + +C. pulchellus (see photograph in - -Hajek + +Hajek and Reiter (2014 : fig. 30)). We have seen three males and one female from Madagascar. All males are of the darker colour form, lacking a basal transverse testaceous band on elytra, similar to -Boheman's - -C. obtusus +Boheman's + +C. obtusus . The female has a weak basal testaceous band but is otherwise also most similar to the dark colour form. Striolation is limited to the middle of the outer three intervals. We have no doubt that the three non-type specimens examined are conspecific with - -C. mimetes + +C. mimetes Guignot. But after our comparison of the male genitalia of the type with - -C. pulchellus + +C. pulchellus specimens from mainland Africa, we concluded that - -C. mimetes + +C. mimetes cannot be upheld as a separate species given how - -C. pulchellus + +C. pulchellus is interpreted. We noted very minor differences in the penis apex which are not greater than differences between east, south, and west African specimens of - -C. pulchellus + +C. pulchellus . The penis figured by -Guignot (1957 +Guignot (1957 : fig. 2, reproduced by -Nilsson et al. 1997 +Nilsson et al. 1997 : fig. 8) is very inaccurate, depicting a shape quite different from - -C. pulchellus + +C. pulchellus but that does not correspond with the holotype. - -Copelatus pulchellus + +Copelatus pulchellus may still be a species complex for future studies to solve but currently it is a widespread and variable species distributed over Africa, Madagascar, and the Middle East. - - -Copelatus basalis + + +Copelatus basalis Boheman, 1848 was synonymised with - -C. pulchellus + +C. pulchellus by Omer-Cooper (1965). We have studied the same type material as Omer-Cooper and agree with this conclusion. However, we are not convinced that the material housed at NHRS as these types are the correctly identified types of the name - -C. basalis + +C. basalis . -Boheman (1848) +Boheman (1848) describes in Latin four species from Johan August Wahlbergs collectings in Caffraria (South Africa) of which three are new, numbered 259 - - -Copelatus pulchellus + +Copelatus pulchellus Klug; 260 - - -Copelatus obtusus + +Copelatus obtusus ; 261 - - -Copelatus striatellus + +Copelatus striatellus ; and 262 - - -Copelatus basalis + +Copelatus basalis . For - -C. pulchellus + +C. pulchellus the disc of elytra is described as having six striae. For the longer description of - -C. obtusus + +C. obtusus this is further elaborated to detail that there are six discal stria but entire elytra has seven striae -"septem-striata" +"septem-striata" (six discal and one submarginal). This pattern and number of striae is consistent with the - -Copelatus pulchellus + +Copelatus pulchellus species group. The following species is correctly identified as - -C. striatellus + +C. striatellus with nine discal striae, and is clearly stated as such: "disco striis 9 tenuibus", of which the innermost is much abbreviated. This description matches very well with the types preserved at NHRS. Finally, - -C. basalis + +C. basalis is described as the last - -Copelatus + +Copelatus species in -Boheman's +Boheman's work and elytra is described as -"12-striata" +"12-striata" . The supposed types at NHRS for both - -C. obtusus + +C. obtusus and - -C. basalis + +C. basalis have six discal and one submarginal striae. The type for - -C. obtusus + +C. obtusus matches the original description of -"septem-striata" +"septem-striata" but the three types of - -C. basalis + +C. basalis with the same number of striae does not match the original description of -"12-striata" +"12-striata" . It would be very inconsistent of Boheman to describe the total number of striae on one elytron in one case -"elytra...septem-striata" +"elytra...septem-striata" , and in the other only the discal striae but summing up the number from both elytral halves -"elytra...12-striata" +"elytra...12-striata" . It would also be very illogical to place - -C. basalis + +C. basalis after the 9-striated - -C. striatellus + +C. striatellus if it has the same number of striae as - -C. pulchellus + +C. pulchellus and - -C. obtusus + +C. obtusus , which come first. Similarily, -Guignot (1961) +Guignot (1961) was confused about -Boheman's - -C. basalis +Boheman's + +C. basalis and listed the name both under the 12-striated - -C. mocquerysi + +C. mocquerysi -Regimbart +Regimbart with a question mark, and under - -C. pulchellus + +C. pulchellus . We consider the status of - -C. basalis + +C. basalis Boheman as uncertain but we have not found any alternative potential type material at NHRS. We designate the single undoubted syntype of - -C. obtusus + +C. obtusus Boheman, 1848 in the NHRS collection as lectotype to preserve the stability of the name. diff --git a/data/08/B5/ED/08B5ED96E2A559C1B8C12353E550A1A7.xml b/data/08/B5/ED/08B5ED96E2A559C1B8C12353E550A1A7.xml index 4a36993b62a..2a83d080db8 100644 --- a/data/08/B5/ED/08B5ED96E2A559C1B8C12353E550A1A7.xml +++ b/data/08/B5/ED/08B5ED96E2A559C1B8C12353E550A1A7.xml @@ -1,221 +1,221 @@ - - - -A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China + + + +A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China - - -Author + + +Author -Ge, Si Xun -https://orcid.org/0000-0003-3769-1530 -Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. +Ge, Si Xun +https://orcid.org/0000-0003-3769-1530 +Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. - - -Author + + +Author -Jiang, Zhuo Heng -School of Science, Westlake University, Hangzhou, China +Jiang, Zhuo Heng +School of Science, Westlake University, Hangzhou, China - - -Author + + +Author -Wang, Jia Qi -2288 Long, Hongxin Rd, Minhang District, Shanghai, China +Wang, Jia Qi +2288 Long, Hongxin Rd, Minhang District, Shanghai, China - - -Author + + +Author -Song, Kui -School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China +Song, Kui +School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China - - -Author + + +Author -Zhang, Chao -Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China +Zhang, Chao +Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China - - -Author + + +Author -Hu, Shao Ji -Institute of International Rivers and Eco-security, Yunnan University, Kunming, China -shaojihu@hotmail.com +Hu, Shao Ji +Institute of International Rivers and Eco-security, Yunnan University, Kunming, China +shaojihu@hotmail.com -text - - -Arthropod Systematics & amp; Phylogeny +text + + +Arthropod Systematics & amp; Phylogeny - -2023 - -2023-03-15 + +2023 + +2023-03-15 - -81 + +81 - -257 -287 + +257 +287 - -http://dx.doi.org/10.3897/asp.81.e85191 + +http://dx.doi.org/10.3897/asp.81.e85191 -journal article -http://dx.doi.org/10.3897/asp.81.e85191 -1864-8312-81-257 -CC0AB5E565324E9EB5D02FD08B5FFE2C -74CBFA410981516FBB87561614EFF561 +journal article +http://dx.doi.org/10.3897/asp.81.e85191 +1864-8312-81-257 +CC0AB5E565324E9EB5D02FD08B5FFE2C +74CBFA410981516FBB87561614EFF561 - - - - -Pieris euorientis euorientis Verity, 1908 -stat. rev. + + + + +Pieris euorientis euorientis Verity, 1908 +stat. rev. - - -Pieris napi var. euorientis + + +Pieris napi var. euorientis Verity, 1908; -Rhopalocera +Rhopalocera Palaearctica (13-14): 147, (17-20): pl. 32, f. 44; TL: -"Sajan" +"Sajan" (original description) - -Pieris euorientis + +Pieris euorientis Verity, 1908; -Eitschberger (1983) +Eitschberger (1983) : Herbipoliana 1(1-2): 131 (upgraded as distinct species) - -Pieris dulcinea euorientis + +Pieris dulcinea euorientis ; -Tuzov et al. (1997) +Tuzov et al. (1997) (as subspecies of -Pieris dulcinea +Pieris dulcinea ) - -Pieris napi euorientis + +Pieris napi euorientis ; -Tadokoro et al. (2014) +Tadokoro et al. (2014) : Butterflies 65: 26 (as subspecies of -Pieris napi +Pieris napi ) - -Description. - + +Description. + Both wings white on the upperside and a pale yellowish hue on the underside. Male (Fig. -11S, U +11S, U ): apical spot grey, merging as a small dark subtriangular spot on the apex or nearly absent. The 1st discal spot and outer spot present but rather faint on both upperside and underside, while the 2nd spot vaguely developed on the underside. Female (Fig. -11T, V +11T, V ): all spots distinctly developed on the upperside, while relatively faint on the underside. Brownish or blackish powder distinctly narrowed along all veins on both sides, relatively broader at the apex. - -Distribution. -Russia, Mongolia (Altai and Sayan Mts, Siberia and Transbaikalia) + +Distribution. +Russia, Mongolia (Altai and Sayan Mts, Siberia and Transbaikalia) - -Phenology. - + +Phenology. + Univoltine from June to July ( -Tuzov et al., 1997 +Tuzov et al., 1997 ). - -Male genitalia. - + +Male genitalia. + (Fig. -5J +5J ) basal part of uncus moderately narrower than the distal margin of tegumen, as a concave between uncus and tegumen; uncus slightly convex at its median part, apical half of uncus digitation. - -Female genitalia. -Not examined. + +Female genitalia. +Not examined. - -Note. - + +Note. + This species rather resembles the spring form of - -P. dulcinea + +P. dulcinea which can only be distinguished by a comparison of the apical spot (more or less merging in this species while always separated into several triangles in - -P. dulcinea + +P. dulcinea ) and the 1st discal spot (both species faint but in - -P. euorientis + +P. euorientis relatively more developed). Males can be easily distinguished from the summer form of - -P. dulcinea + +P. dulcinea in having: apical spot merged (separate in - -P. dulcinea + +P. dulcinea ), and 1st discal spot faint (strongly developed in - -P. dulcinea + +P. dulcinea ) and 2nd discal spot absent (present in - -P. dulcinea + +P. dulcinea ) on the upperside. This taxon shows monophyly in our phylogenetic analysis and its position is distant from - -P. napi + +P. napi ; therefore, we consider this taxon to be a distinct species. It is noteworthy that in the re-description of the species by -Eitschberger (1983) +Eitschberger (1983) a series of specimens from Tanu-Ola (Tanguura Mountains) were considered as belonging to this species, while in this study, we found morphological differences between topotypes of - -P. euorientis + +P. euorientis and specimens from regions south or west of the Tanguura Mountains (See notes in - -Pieris mihon + +Pieris mihon Yakovlev, 2006 stat. nov.). Combined with our phylogenetic results, this indicates that populations from the Tanguura Mountains and the regions to the south or west represent a distinct species (Fig. -12 +12 ): - -Pieris mihon + +Pieris mihon Yakovlev, 2006 stat. nov. As the redefined - -Pieris euorientis euorientis + +Pieris euorientis euorientis , we did not find female specimens of the summer form in the type locality or adjacent regions, thus this knowledge gap needs to be filled in future research. - - -Figure 12. + + +Figure 12. Distribution map of - -Pieris euorientis + +Pieris euorientis Verity, 1908 stat. rev. and - -Pieris mihon + +Pieris mihon Yakovlev, 2006 stat. nov. diff --git a/data/0A/69/4D/0A694D306A4456EB8ADB140766C419B7.xml b/data/0A/69/4D/0A694D306A4456EB8ADB140766C419B7.xml index 64e884139f9..439a6289aa6 100644 --- a/data/0A/69/4D/0A694D306A4456EB8ADB140766C419B7.xml +++ b/data/0A/69/4D/0A694D306A4456EB8ADB140766C419B7.xml @@ -1,124 +1,124 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Exocelina subjecta (Sharp, 1882) + + + +Exocelina subjecta (Sharp, 1882) - - -Copelatus subjectus + + +Copelatus subjectus Sharp, 1882: 568. - -Copelatus bilunatus + +Copelatus bilunatus Guignot, 1955: 73; TL: Madagascar [mislabelled, likely New Caledonia; see -Balke et al. (2014) +Balke et al. (2014) ] - -Type locality. -New Caledonia. + +Type locality. +New Caledonia. - -Remarks. - - -Copelatus bilunatus + +Remarks. + + +Copelatus bilunatus Guignot, 1955 is a synonym of - -Exocelina subjecta + +Exocelina subjecta (Sharp, 1882) following -Balke et al. (2014) +Balke et al. (2014) . - -Copelatus bilunatus + +Copelatus bilunatus was described by -Guignot (1955a) +Guignot (1955a) from -Zimmermann's +Zimmermann's collection based on a "♀ unique" from -"Madagascar" +"Madagascar" without any further locality data. The species was assigned to - -Copelatus + +Copelatus based on the presence of complete metacoxal lines and placed in the - -Copelatus hydroporoides + +Copelatus hydroporoides species group, as it lacked impressed elytral striae. -Balke et al. (2014) +Balke et al. (2014) concluded that the specimen is identical to - -Exocelina subjecta + +Exocelina subjecta (Sharp, 1882), a common species on New Caledonia, and synoymised the two assuming the specimen was mislabelled. Apparently the holotype is a male in contrast to what is stated in the original description. -Balke et al. (2014) +Balke et al. (2014) document the presence of a stout spine-like setae on the anterodistal angle of protarsomere IV, which is characteristic of the -Copelatinae +Copelatinae genus - -Exocelina + +Exocelina (but also of Malagasy - -Madaglymbus + +Madaglymbus ). We have not examined the type ourselves but it is clear that in any case it is not a - -Copelatus + +Copelatus species. diff --git a/data/0C/74/16/0C74161662CC517EA04E51E9940C49EC.xml b/data/0C/74/16/0C74161662CC517EA04E51E9940C49EC.xml index a02ffe95591..a6e4c9c7cfb 100644 --- a/data/0C/74/16/0C74161662CC517EA04E51E9940C49EC.xml +++ b/data/0C/74/16/0C74161662CC517EA04E51E9940C49EC.xml @@ -1,165 +1,165 @@ - - - -A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China + + + +A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China - - -Author + + +Author -Ge, Si Xun -https://orcid.org/0000-0003-3769-1530 -Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. +Ge, Si Xun +https://orcid.org/0000-0003-3769-1530 +Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. - - -Author + + +Author -Jiang, Zhuo Heng -School of Science, Westlake University, Hangzhou, China +Jiang, Zhuo Heng +School of Science, Westlake University, Hangzhou, China - - -Author + + +Author -Wang, Jia Qi -2288 Long, Hongxin Rd, Minhang District, Shanghai, China +Wang, Jia Qi +2288 Long, Hongxin Rd, Minhang District, Shanghai, China - - -Author + + +Author -Song, Kui -School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China +Song, Kui +School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China - - -Author + + +Author -Zhang, Chao -Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China +Zhang, Chao +Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China - - -Author + + +Author -Hu, Shao Ji -Institute of International Rivers and Eco-security, Yunnan University, Kunming, China -shaojihu@hotmail.com +Hu, Shao Ji +Institute of International Rivers and Eco-security, Yunnan University, Kunming, China +shaojihu@hotmail.com -text - - -Arthropod Systematics & amp; Phylogeny +text + + +Arthropod Systematics & amp; Phylogeny - -2023 - -2023-03-15 + +2023 + +2023-03-15 - -81 + +81 - -257 -287 + +257 +287 - -http://dx.doi.org/10.3897/asp.81.e85191 + +http://dx.doi.org/10.3897/asp.81.e85191 -journal article -http://dx.doi.org/10.3897/asp.81.e85191 -1864-8312-81-257 -CC0AB5E565324E9EB5D02FD08B5FFE2C -74CBFA410981516FBB87561614EFF561 +journal article +http://dx.doi.org/10.3897/asp.81.e85191 +1864-8312-81-257 +CC0AB5E565324E9EB5D02FD08B5FFE2C +74CBFA410981516FBB87561614EFF561 - - - -Pieris latouchei Mell, 1939 + + + +Pieris latouchei Mell, 1939 - - -Pieris extensa latouchei + + +Pieris extensa latouchei Mell, 1939; Dt. Ent. Z. Iris 52 (1938): 138-139. TL: "Kuatun, NW-Fukien" [N.W. of Shaowu City, Fujian] (original description) - -Pieris erutae latouchei + +Pieris erutae latouchei ; -Eitschberger (1983) +Eitschberger (1983) : Herbipoliana 1(1-2): 376 (as subspecies of -Pieris erutae +Pieris erutae Poujade, 1888) - -Pieris melete latouchei + +Pieris melete latouchei ; Wu (2010): Fauna Sinica -Insecta +Insecta 52: 300 (as subspecies of -Pieris melete -Menetries +Pieris melete +Menetries , 1857) - -Pieris latouchei + +Pieris latouchei ; -Tadokoro et al. (2014) +Tadokoro et al. (2014) : Butterflies 65: 25 (raised to distinct species from subspecies of -Pieris erutae +Pieris erutae ) - -Description. - + +Description. + Both wings white on the upperside while bright yellow on the underside (in autumn form males whitish). Spring form: medium size. Male (Fig. -10A +10A ): apical spot blackish with slightly whitish powder, merging as a small dark subtriangular spot on the apex. The 1st discal spot and outer spot rather faint or absent on the upperside. All spots absent on the underside, except the 2nd discal spot vaguely developed. Female: apical spot brownish, partly merging as a subtriangular spot on the apex. The 1st, 2nd and outer spot strongly developed on the upperside while faintly present on the underside (except outer spot absent). All veins narrowly brown powdered on both upperside and underside. Summer form: medium to large size. Resembles spring form but larger in size and rounder wing shape. The 1st discal spot of males (Fig. -10B +10B ) present on both sides and the brown powder along veins is stronger in summer form of females (Fig. -10D +10D ) than spring forms. Autumn form: large size. Male (Fig. -10C +10C ): resembles summer form but less yellowish on the underside (near whitish), the apical spot merging as a triangular spot and the 1st and 2nd discal spots rather distinct on both sides (the outer spot distinct on the upperside while absent on the underside). Female (Fig. -10E +10E ): resembles summer form but dark yellowish on the underside, all veins strongly brown powdered on the upperside while quite narrow on the underside. - -Distribution. -East China (Henan, Shandong, Jiangsu, Zhejiang, Fujian) + +Distribution. +East China (Henan, Shandong, Jiangsu, Zhejiang, Fujian) - -Phenology. -Multivoltine, occurs from April to September. + +Phenology. +Multivoltine, occurs from April to September. - -Male genitalia. - + +Male genitalia. + (Fig. -5K +5K ) tegumen relatively slender, with its median part slightly concave, the distal margin of tegumen without distinct convex; basal margin of the uncus slightly narrower than the tegumen; uncus with its apical obtusely digitation. - -Female genitalia. - + +Female genitalia. + (Fig. -7A +7A ) posterior apophysis slender, extended reach to the 8th tergum; inner distal of sterigma lobe-shaped, extend to center, with a distinct convex at the trailing edge; inner basal of sterigma near trapezoidal extend to the center, smoothly connected at the basal margin; signum cordiform-shaped, relatively slender, smoothly tapering to the basal. - -Note. - + +Note. + This taxon was elevated to species level by -Tadokoro et al. (2014) +Tadokoro et al. (2014) , based on results from his phylogenetic analysis, which is also supported by our molecular results. This species is closely related to - -P. erutae + +P. erutae and shows three different seasonal forms in generations, thus can be only distinguished from the bright yellow color of wings on the underside (while in autumn forms are -variable) and the basal 1/2 dark powdered discocell on the underside (only developed in autumn forms). diff --git a/data/0D/A7/78/0DA77899943B5226BBA57EB36328B122.xml b/data/0D/A7/78/0DA77899943B5226BBA57EB36328B122.xml index 6b00433fd3d..fe44b0a39f1 100644 --- a/data/0D/A7/78/0DA77899943B5226BBA57EB36328B122.xml +++ b/data/0D/A7/78/0DA77899943B5226BBA57EB36328B122.xml @@ -1,201 +1,201 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Copelatus baculiformis Guignot, 1955 -Fig. 7B + + + +Copelatus baculiformis Guignot, 1955 +Fig. 7B - - -Copelatus baculiformis + + +Copelatus baculiformis Guignot, 1955b: 193. - -Type locality. -Madagascar, Massif Ankaratra, Manjakatompo, alt. 1700-1800 m. + +Type locality. +Madagascar, Massif Ankaratra, Manjakatompo, alt. 1700-1800 m. - -Type information from original description. -based on a single female specimen (holotype), collected December 1951 by R. Benoist. + +Type information from original description. +based on a single female specimen (holotype), collected December 1951 by R. Benoist. - -Type material studied. - -Antananarivo. Vakinankaratra: Ambatolampy + +Type material studied. + +Antananarivo. Vakinankaratra: Ambatolampy : -HT ♀ (MNHN "coll. Guignot"): // Data in NHRS | JLKB | 000030226 // Madagascar: Massif An- | karatra 1700/1800 Man- | jakatompo XII-51 Benoist // Type [red label] // Guignot det., 1954 | -Copelatus +Copelatus | -baculiformis +baculiformis n. sp. | Type ♀ // - -Diagnosis. - + +Diagnosis. + Small size (4 mm). Elytra uniformly dark brown ferrugineous, without a basal testaceous area ( -Fig. 7B +Fig. 7B ) which separates the species from small species of - -Madaglymbus + +Madaglymbus . The absence of deep impressed elytral striae (remnants of four elytral striae present) separates this species from all other - -Copelatus + +Copelatus of Madagascar except - -Copelatus peridinus + +Copelatus peridinus , a larger species (5.7-6.6 mm). - -Description. -(based on holotype ♀): - + +Description. +(based on holotype ♀): + Body length 4 mm. Body shape elongate oval and dark brown to blackish ferrugineous. Head uniformly dark brown ferrugineous to slightly darker posteriorly inside eyes, with thin sparse punctation. Pronotum dark brown ferrugineous, same colour medially and laterally but darker along anterior and posterior third. Disc of pronotum less densely punctuated, posterolateral corners with dense superficial strioles. The entire dorsal surface covered with a microsculpture. Elytra uniformly coloured in same dark brown to blackish ferrugineous colour as anterior and posterior parts of pronotum ( -Fig. 7B +Fig. 7B ). Impressed striae absent but four elytral rows of impressed points present. Innermost, a presumed first row completely lacking, row 2, 3, and 4 visible from base but row 3 less distinct than 2 and 4. Row 5 very vague and only visible posteriorly. Elytra densely covered with punctures which laterally of approximately the third row of points is replaced by strioles (probably only in the female). Appendages testaceous. - - + + Figure 7. Habitus, dorsal view. -A +A Male: - -Copelatus peridinus + +Copelatus peridinus -B +B Female: - -Copelatus baculiformis + +Copelatus baculiformis -C +C Female: - -Copelatus + +Copelatus sp. 2 (Andasibe) -D +D Female: - -Copelatus distinguendus + +Copelatus distinguendus . - + Ventral side testaceous to weakly infuscated. Prosternal process carinate also onto apical process. Lateral parts of metaventrite ("metasternal wings") rather broad. Metacoxal lines anteriorly diverging and abbreviated before metaventral margin. Metacoxa with fine and long longitudinal strioles, continuing onto abdominal ventrites, and with 6-7 transverse -"wrinkles" +"wrinkles" anterolaterally. -Male: unknown. +Male: unknown. - -Distribution. - + +Distribution. + Madagascar, central highlands, only known from type locality Manjakatompo, Ankaratra Massif ( -Fig. 11A +Fig. 11A ). - -Habitat and ecology. - + +Habitat and ecology. + Unknown, but according to original description collected at at an altitude of 1700-1800 m. See -Hjalmarsson et al. (2013) +Hjalmarsson et al. (2013) for a description of the locality Manjakatompo and its conservation priority. - -Comments. - + +Comments. + No other specimen than the female holotype is known of this species and it is a bit of a "mystery species". We have conducted fieldwork at the type locality Manjakatompo multiple times (2011, 2012, 2014, and 2016) but never found any specimens resembling this species. The species belongs in the - -hydroporoides + +hydroporoides species group of - -Copelatus + +Copelatus , but two other Malagasy species placed in this group have turned out to be misidentified - -Madaglymbus + +Madaglymbus or - -Exocelina + +Exocelina (see above). - -Copelatus baculiformis + +Copelatus baculiformis was described by Guignot the same year (1955) that he described - -C. bilunatus + +C. bilunatus , considered mislabeled ( -Balke et al. 2014 +Balke et al. 2014 ) from the Zimmermann collection. It is certainly possible that also - -C. baculiformis + +C. baculiformis is based on a mislabeled specimen, but in contrast to - -C. bilunatus + +C. bilunatus the type locality data is more exact; Massif Ankaratra (mountain), Manjakatompo (locality), 1700-1800 m (altitude in meters), XII-51 (collecting month and year), and R. Benoist (collector), speaking against such a mistake. Based on general external morphology, body shape and lack of striae the species resembles - -C. peridinus + +C. peridinus . It is much smaller and therefore likely not conspecific, but they could be closely related, as well as with the female sequenced from Andasibe (NHRS-JLKB000065698). diff --git a/data/24/62/90/246290F17F4059918CAB1EF8A036B9D5.xml b/data/24/62/90/246290F17F4059918CAB1EF8A036B9D5.xml index 2d8a4e738d2..72d627d86a3 100644 --- a/data/24/62/90/246290F17F4059918CAB1EF8A036B9D5.xml +++ b/data/24/62/90/246290F17F4059918CAB1EF8A036B9D5.xml @@ -1,169 +1,169 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Copelatus befasicus Guignot, 1956 -Fig. 10D + + + +Copelatus befasicus Guignot, 1956 +Fig. 10D - - -Copelatus befasicus + + +Copelatus befasicus Guignot, 1956: 79. - -Type locality. -Madagascar, Morondava, forest south of Befasy. + +Type locality. +Madagascar, Morondava, forest south of Befasy. - -Type information from original description. -based on an unknown number of female type specimens but holotype and paratypes are distinguished in introduction. Collected in January 1956 by R. Paulian. + +Type information from original description. +based on an unknown number of female type specimens but holotype and paratypes are distinguished in introduction. Collected in January 1956 by R. Paulian. - -Type material studied. - -Toliara. Menabe: Morondava + +Type material studied. + +Toliara. Menabe: Morondava : -HT♀ (MNHN, "coll. IRSM"): // Data in NHRS | JLKB | 000030021 // Morondava | -foret +foret sud | de Befasy | I-56 R.P // Type [red label] // INSTITUT | SCIENTIFIQUE | MADAGASCAR // Guignot det., 1956 | -Copelatus +Copelatus | -befasicus +befasicus n. sp. | Type // -2PT ♀ (MNHN, "coll. Guignot"): // Data in NHRS | JLKB | 000030300-1// Morondava | -foret +foret sud | de Befasy // I-56 R.P // Paratype [red label] // [female symbol] // - -Additional material studied. - -Mahajanga. Melaky: Morafenobe + +Additional material studied. + +Mahajanga. Melaky: Morafenobe : -1♀ (NHRS): // NHRS-JLKB | 000010860 (JB204) // Madagascar: Mahajanga: Melaky | Btw. -Morafenobe-Ambohijanahary +Morafenobe-Ambohijanahary | S18.19091; E045.19986, 290 m.a.o | 19.XII.2009 Water Net, Field# MAD09-74 | Leg: J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // - -Diagnosis. - + +Diagnosis. + Similar to - -C. insuetus + +C. insuetus and related species in habitus by being small, elongate, and subparalell, but - -C. befasicus + +C. befasicus is distinguished from - -C. insuetus + +C. insuetus and from all other Malagasy - -Copelatus + +Copelatus species by the presence of only five elytral striae, and without submarginal striae. In addition, the first stria is shortened, present only in the posterior third ( -Fig. 10D +Fig. 10D ). - -Description. - + +Description. + Body length 4.1-4.2 mm. Body shape elongate and subparallel, dorsal surface reddish brown with a lighter elytral base. Head and pronotum uniformly rufus brown. Head, pronotum and elytra with dense punctation. Elytra and pronotum covered with dense punctures and the whole dorsal surface with a microsculpture. Lateral sides of pronotum striolate with the widest striolate area in the posterior corners. Elytra light brown with a distinct testaceous band basally ( -Fig. 10D +Fig. 10D ). First elytral stria shortened and present only in posterior third. Second to fifth elytral stria starting more or less at base and all striae approaching the apex of elytron, but the second and fourth a little shorter. Submarginal striae absent. Appendages testaceous. -Ventral side light brown. Metacoxa and abdominal ventrites punctate and striolate. Prosternal process rather short and medially raised, triangular in cross-section. Lateral parts of metaventrite medium broad. Metacoxal lines anteriorly diverging but rather weakly so, abbreviated well before metaventral margin. Antennae, palps and legs testaceous. -Male: unknown. +Ventral side light brown. Metacoxa and abdominal ventrites punctate and striolate. Prosternal process rather short and medially raised, triangular in cross-section. Lateral parts of metaventrite medium broad. Metacoxal lines anteriorly diverging but rather weakly so, abbreviated well before metaventral margin. Antennae, palps and legs testaceous. +Male: unknown. - -Distribution. - + +Distribution. + Known only from two localities in the western part of Madagascar, the deciduous forest south of Befasy, Morondava, and at one locality between Morafenobe and Beravina village ( -Fig. 12D +Fig. 12D ). - -Habitat and ecology. - + +Habitat and ecology. + Paulian collected the species in 1956 in the western dry deciduous forest south of Befasy, Morondava. We rediscovered the species in 2009, when we found one female specimen of - -C. befasicus + +C. befasicus also in the western part but a bit further north than the type locality, along the road between Morafenobe and Beravina village at an altitude of 290 m. This locality consisted of dry savannah with mixed wood and grassland ecosystem after deforestation. The habitat consisted of muddy/sandy residual pools with some dead leaves in a temporary stream after the rainy season. The dry deciduous forest ecosystem in western Madagascar has suffered immensely from deforestation and very little of this habitat remains ( -Ganzhorn et al. 2001 +Ganzhorn et al. 2001 ; -Whitehurst et al. 2009 +Whitehurst et al. 2009 ). The species is rare and likely threatened due to the disappearance of western deciduous forests in Madagascar. That all four known specimens are females may suggest an uneven sex ratio as a test of equal sex ratio is marginally non-significant (p = 0.0625) if considered randomly picked from the population. - -Comments. - + +Comments. + This is the only species in the - -Copelatus longicornis + +Copelatus longicornis group from Madagascar. The - -longicornis + +longicornis species group currently contains 38 species distributed mainly in the Neotropical and Afrotropical regions but also with few species present in Japan, New Guinea, and Fiji Islands ( - + Nilsson and -Hajek +Hajek 2018 ). The group as currently defined is certainly artificial from a phylogenetic perspective and the only character they have in common is the low number of elytral striae. diff --git a/data/25/AF/5D/25AF5D96C7285A06B37C8F9CC93796E0.xml b/data/25/AF/5D/25AF5D96C7285A06B37C8F9CC93796E0.xml index ea5354ffa07..7f2c5624a04 100644 --- a/data/25/AF/5D/25AF5D96C7285A06B37C8F9CC93796E0.xml +++ b/data/25/AF/5D/25AF5D96C7285A06B37C8F9CC93796E0.xml @@ -1,169 +1,169 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Copelatus peridinus Guignot, 1955 -Figs 4A + + + +Copelatus peridinus Guignot, 1955 +Figs 4A , -7A +7A - - -Copelatus peridinus + + +Copelatus peridinus Guignot, 1955c: 188. - -Copelatus seydeli + +Copelatus seydeli Guignot, 1958: 107; TL: Elisabethville, Zaire [DR Congo, Haut-Katanga, Lubumbashi]. - -Type locality. -Elisabethville, Zaire [DR Congo, Haut-Katanga, Lubumbashi]. + +Type locality. +Elisabethville, Zaire [DR Congo, Haut-Katanga, Lubumbashi]. - -Type material studied. - + +Type material studied. + -PT ♀ (MNHN, "Coll. Guignot"): // Data in NHRS | JLKB | 000030317 // Allotype [red label] // [female symbol] // Elisabethville | XI. 1951 // -Copelatus +Copelatus | -peridinus +peridinus | Allotype ♀ // - -Additional material studied. - + +Additional material studied. + -1♂(GP) (MNHN, "Coll. Guignot"): // Data in NHRS | JLKB | 000030318 // Congo Belge Lac | Edouard: Vitshumbi | U.V. 27.XI.1953 | 3091 // -Antananarivo. Analamanga: Antananarivo +Antananarivo. Analamanga: Antananarivo : -3♂(GP), 3♀ (MNHN, "Coll. Legros Magasin"): // Data in NHRS | JLKB | 000030319-24 // MADAGASCAR | TANANARIVE | BETONGOLO | 2 XII 1946 // -Piege +Piege | lumineux // Museum Paris | 1983 | Coll. Cl. Legros // -Toamasina. Alaotra Mangoro: Moramanga +Toamasina. Alaotra Mangoro: Moramanga : -1♂(GP), 2♀ (MNHN, "Coll. Paulian"): // Data in NHRS | JLKB | 000030316, 30325-6 // Madagascar Est | P.K.57-Rte -d'Anosibe +d'Anosibe | Moramanga | II.58 R. Vieu // -1♂(GP), 1♀ (NMPC, "Coll. J. -Hajek" +Hajek" ): // Data in NHRS | JLKB | 000030327, 30328 // Madagascar | Lokato, near | Andasibe Mantadia NP | M. Tryzna leg., 9-10.i.2007 // coll. Jiri -HAJEK +HAJEK | National Museum | Prague, Czech Republic // ["Lokato near Andasibe Mantadia NP" interpreted as the bifurcation of the road to Lakato which is near Andasibe Mantadia NP. Lakato itself is not near Andasibe Mantadia NP.] -2♂(GP), 1♀(teneral), 1♀, 1♂ (Alc. teneral) (NHRS): // NHRS-JLKB | 000010887, 65737, 65703, 10888(Alc.) // MAD: TOAM: Alaotra Mangoro: | Andasibe Mantadia NP, Analamazaotra: | 150m E of park entrance: Mad14-14 | shallow partly dried out forest pond: | 18.9355S | 48.4166E: 930m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja & T. Ranarilalatiana // - -Diagnosis. - + +Diagnosis. + Similar to - -C. baculiformis + +C. baculiformis based on shape and colouration, but - -C. peridinus + +C. peridinus is bigger with body length between 5.7 and 6.6 mm ( -Fig. 7A +Fig. 7A ). Elytra lacking deeply impressed striae, but elytral rows of impressed points present out of which two rows are most distinct. Penis in lateral view curved with two points at which it curves slightly more abruptly creating two gentle angles ( -Fig. 4A +Fig. 4A ); parameres rather broad at base and evenly curved ( -Fig. 4A +Fig. 4A ). - -Description. - + +Description. + Body length 5.7-6.6 mm. Body shape elongate oval, from midpoint uniformly tapering towards a rather pointed apex. Head and pronotum both in the same dark brown to blackish ferrugineous colouration; elytra anteriorly in the same colour as pronotum, but posterior part brown ferrugineous with largely testaceous lateral margins ( -Fig. 7A +Fig. 7A ). Head, pronotum and elytra with thin dense punctation, in addition pronotum and elytra with small and shallow punctures, and microsculptured. At posterolateral corners of pronotum, punctures almost a little joined and corrugated. Elytra lacking deeply impressed striae. Two continuous rows of punctures are most obvious and divide the elytra in three more or less equal intervals. A third and a fourth row of punctures also present, albeit less distinct and more fragmented. These occur between the two first distinct rows and between the second distinct row and the elytral margin. A fifth row suggested between the two outermost rows by a few spaced out punctures, almost unidentifiable in most Malagasy specimens seen but more distinct in specimens seen from mainland Africa. Appendages testaceous to rufus. -Ventral side dark ferrugineous. Prosternal process strongly carinate anteriorly and with a rather short process. Lateral parts of the metaventrite medium-broad. Metacoxal lines short, anteriorly diverging and abbreviated well before metaventral margin. Metacoxa with short fine strioles continuing onto abdominal ventrites. -Male: first three pro- and mesotarsomeres widened. Protibia modified, narrow at base and with an early abrupt bend, extended and broadened towards middle with a straight ventral side but angled dorsal side. Pro- and mesotarsal claws unmodified. - +Ventral side dark ferrugineous. Prosternal process strongly carinate anteriorly and with a rather short process. Lateral parts of the metaventrite medium-broad. Metacoxal lines short, anteriorly diverging and abbreviated well before metaventral margin. Metacoxa with short fine strioles continuing onto abdominal ventrites. +Male: first three pro- and mesotarsomeres widened. Protibia modified, narrow at base and with an early abrupt bend, extended and broadened towards middle with a straight ventral side but angled dorsal side. Pro- and mesotarsal claws unmodified. + Penis in ventral view with apical part more or less straight and even, gently pointed at apex but with the very apex minutely twisted to the right ( -Fig. 4A +Fig. 4A ). Penis in lateral view curved with two points (1/3rd and 2/3rd from base) at which it curves slightly more abruptly creating two gentle angles ( -Fig. 4A +Fig. 4A ). Parameres rather broad at base and evenly curved and tapering towards apex ( -Fig. 4A +Fig. 4A ). -Female: dorsal sculpture similar to male. +Female: dorsal sculpture similar to male. - -Distribution. - + +Distribution. + Likely a more widespread distribution in at least central and eastern continental Africa than the current records (Lubumbashi and Kivu in DR Congo) indicate. In Madagascar, only know from the eastern central parts: Betongolo (Antananarivo), the Analamazaotra NP, and P.K.57 Route -d'Anosibe +d'Anosibe [RN23] ( -Fig. 11C +Fig. 11C ). - -Habitat and ecology. - + +Habitat and ecology. + A series of six specimens collected with light trap ( -"piege +"piege lumineux") in the capital Antananarivo 1946, indicates flight capacity and anthropogenically disturbed habitats. All records from DR Congo are also from light trap catches ( -Bilardo 1982 +Bilardo 1982 ). We collected a second series of teneral specimens in a shallow forest pool with vegetation, near the entrance to the Analamazaotra NP. - -Comments. -This species was described from Lubumbashi, DR Congo, and has not been recorded from Madagascar before. Earliest record found is from November 1946. It seems to be a dispersive good flier and often collected at light so its presence in Madagascar is therefore not surprising. However, it is not widespread in Madagascar as far as we know. In fact, the known distribution is restricted to the surroundings of the capital and east of the capital along the main national route towards Toamasina, which could suggest a recent incidental human-mediated introduction from mainland Africa. - + +Comments. +This species was described from Lubumbashi, DR Congo, and has not been recorded from Madagascar before. Earliest record found is from November 1946. It seems to be a dispersive good flier and often collected at light so its presence in Madagascar is therefore not surprising. However, it is not widespread in Madagascar as far as we know. In fact, the known distribution is restricted to the surroundings of the capital and east of the capital along the main national route towards Toamasina, which could suggest a recent incidental human-mediated introduction from mainland Africa. + Note that it may be that this is a species with intraspecific variation with regards to elytral striation, ranging from five puncture lines out of which two are more distinct, to five weakly impressed striae (see further discussion under - -Copelatus + +Copelatus sp. 2 below). diff --git a/data/26/DA/2B/26DA2B91F7A55A8AB113C5C4AB99079A.xml b/data/26/DA/2B/26DA2B91F7A55A8AB113C5C4AB99079A.xml index cd8f71196cd..450be93c62e 100644 --- a/data/26/DA/2B/26DA2B91F7A55A8AB113C5C4AB99079A.xml +++ b/data/26/DA/2B/26DA2B91F7A55A8AB113C5C4AB99079A.xml @@ -1,198 +1,198 @@ - - - -A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China + + + +A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China - - -Author + + +Author -Ge, Si Xun -https://orcid.org/0000-0003-3769-1530 -Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. +Ge, Si Xun +https://orcid.org/0000-0003-3769-1530 +Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. - - -Author + + +Author -Jiang, Zhuo Heng -School of Science, Westlake University, Hangzhou, China +Jiang, Zhuo Heng +School of Science, Westlake University, Hangzhou, China - - -Author + + +Author -Wang, Jia Qi -2288 Long, Hongxin Rd, Minhang District, Shanghai, China +Wang, Jia Qi +2288 Long, Hongxin Rd, Minhang District, Shanghai, China - - -Author + + +Author -Song, Kui -School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China +Song, Kui +School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China - - -Author + + +Author -Zhang, Chao -Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China +Zhang, Chao +Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China - - -Author + + +Author -Hu, Shao Ji -Institute of International Rivers and Eco-security, Yunnan University, Kunming, China -shaojihu@hotmail.com +Hu, Shao Ji +Institute of International Rivers and Eco-security, Yunnan University, Kunming, China +shaojihu@hotmail.com -text - - -Arthropod Systematics & amp; Phylogeny +text + + +Arthropod Systematics & amp; Phylogeny - -2023 - -2023-03-15 + +2023 + +2023-03-15 - -81 + +81 - -257 -287 + +257 +287 - -http://dx.doi.org/10.3897/asp.81.e85191 + +http://dx.doi.org/10.3897/asp.81.e85191 -journal article -http://dx.doi.org/10.3897/asp.81.e85191 -1864-8312-81-257 -CC0AB5E565324E9EB5D02FD08B5FFE2C -74CBFA410981516FBB87561614EFF561 +journal article +http://dx.doi.org/10.3897/asp.81.e85191 +1864-8312-81-257 +CC0AB5E565324E9EB5D02FD08B5FFE2C +74CBFA410981516FBB87561614EFF561 - - - -Pieris chumbiensis gyantsensis Verity, 1911 + + + +Pieris chumbiensis gyantsensis Verity, 1911 - - -Pieris dubernardi var. chumbiensis r. gyantsensis + + +Pieris dubernardi var. chumbiensis r. gyantsensis Verity, 1911; Rho-pa-locera Palaearctica (30-36): 329, pl. 66, f. 21-23; TL: "Gyangtse, S. Tibet" (original description) - -Synchloe chumbiensis gyantsensis + +Synchloe chumbiensis gyantsensis ; -Bollow (1930) +Bollow (1930) : The Macrolepido-ptera of the World, supplement to Vol. 1: 339 pp, pl.16. (infrasubspecific name validated as subspecies of -Synchloe chumbiensis +Synchloe chumbiensis ) - -Pieris gyantsensis + +Pieris gyantsensis ; -Tadokoro et al. (2014) +Tadokoro et al. (2014) : Butterflies 65: 33 (as distinct species) - -Pieris chumbiensis gyantsensis + +Pieris chumbiensis gyantsensis ; -Tadokoro et al. (2016) +Tadokoro et al. (2016) : -Lepidoptera +Lepidoptera Science 67: 99-114 - -Pieris chumbiensis gyantsensis + +Pieris chumbiensis gyantsensis ; Huang (2019): Neue Entomologische Nachrichten 77: 203-255 (pictured, keyed) - -Description. - + +Description. + Medium size. Both wings whitish on the upperside while yellowish on the underside. Male (Fig. -13G +13G ): apical spot blackish, triangular extending along veins to outer margin with a trend of merging. A short black band at the end of the discocell. The 1st spot distinct and oval-shaped, with a trend of a spot faintly occurs on the cell between vein CuA1 and CuA2; outer spot less faintly present on the upperside. All veins of hindwings heavily brownish powdered on the underside, with a brownish sub-outer marginal band. Female (Fig. -13H +13H ): resembles male but with black markings more developed. The 1st spot distinctly developed and extends as a black band reaching the posterior margin. Hindwings with blackish sub-marginal band distinctly developed on both sides. - -Distribution. -Tibet (from Lhasa to Nyalam) + +Distribution. +Tibet (from Lhasa to Nyalam) - -Phenology. -Univoltine, occurs in June and July. + +Phenology. +Univoltine, occurs in June and July. - -Male genitalia. - + +Male genitalia. + (Fig. -5Q +5Q ) tegumen broad with its distal margin slightly narrower; basal margin of uncus slightly convex with a moderate concave between tegumen and uncus; uncus with its median part slightly convex, apical half of uncus digitation. - -Female genitalia. - + +Female genitalia. + (Fig. -8C +8C ) posterior apophysis long and slender, extended beyond the 8th tergum; inner distal of sterigma lobe shaped extend to center; inner basal of sterigma without distinct convex, smoothly connected at the basal margin; signum short and broad, almost disk shaped with smooth and shallowly concave on the terminal margin; basal part of signum tubular with distinct bending on the most basal part. - -Note. - + +Note. + The taxon - -Pieris gyantsensis + +Pieris gyantsensis was originally published by Verity (1911) as a quadrinomial race belonging to - -P. chum-biensis + +P. chum-biensis , thus unavailable due to being infrasub-specific. When -Bollow (1930) +Bollow (1930) re-defined this taxon as a subspecies of - -Synchloe chumbiensis + +Synchloe chumbiensis , the subspecies name became an available name. This taxon was once mistreated as belonging to the - -P. napi + +P. napi -complex by -Tadokoro et al. (2014) +Tadokoro et al. (2014) according to their phylogenetic results and observations on androconial characteristics. Our molecular results oppositely showed that this taxon constitues a distinct molecular lineage separated from the - -P. napi + +P. napi -complex. Moreover, this taxon has long been considered as a member of the - -P. dubernardi + +P. dubernardi -group, a unique species group with distinctive morphological characteristics and a high altitude habitat which includes - -P. dubernardi + +P. dubernardi , - -P. rothschildi + +P. rothschildi , - -P. wangi + +P. wangi as well as - -P. chumbiensis + +P. chumbiensis . diff --git a/data/45/AB/55/45AB55AB28805B229F01D47FDED687E7.xml b/data/45/AB/55/45AB55AB28805B229F01D47FDED687E7.xml index 22df0720186..031548cdab2 100644 --- a/data/45/AB/55/45AB55AB28805B229F01D47FDED687E7.xml +++ b/data/45/AB/55/45AB55AB28805B229F01D47FDED687E7.xml @@ -1,147 +1,147 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - - + + + + Madaglymbus unguicularis ( -Regimbart +Regimbart , 1903) -comb. nov. +comb. nov. - - -Copelatus unguicularis -Regimbart + + +Copelatus unguicularis +Regimbart , 1903:19. - -Type locality. -Madagascar, Suberbieville [= Maevatanana]. + +Type locality. +Madagascar, Suberbieville [= Maevatanana]. - -Type material examined. - -Mahajanga. Betsiboka: Maevatanana + +Type material examined. + +Mahajanga. Betsiboka: Maevatanana : -HT ♂ (GP) (MNHN, "coll. -Regimbart" +Regimbart" ): // Data in NHRS | JLKB | 000030225 // Suberbieville | Madag. Perier // Type [red label] // MUSEUM PARIS | Coll. Maurice -Regimbart +Regimbart | 1908 // -C. unguicularis +C. unguicularis | Type // -unguicularis -Reg +unguicularis +Reg . // - -Remarks. - - -Copelatus unguicularis + +Remarks. + + +Copelatus unguicularis was described by - -Regimbart + +Regimbart (1903) , based on a single male from Suberbieville [= Maevatanana]. It was classified by -Regimbart +Regimbart in the fifth species group of -Sharp (1882) +Sharp (1882) (the -consors +consors species group sensu -Guignot 1961 +Guignot 1961 ) based on ten discal but no submarginal elytral stria where it is still classified today ( -Guignot 1961 +Guignot 1961 ; - -Gueorguiev + +Gueorguiev 1968 ; -Nilsson et al. 1997 +Nilsson et al. 1997 ; -Nilsson 2001 +Nilsson 2001 ; - + Nilsson and -Hajek +Hajek 2018 ). We have examined the male holotype and like the type of - -Copelatus apicalis + +Copelatus apicalis , it has a stout spine-like setae on a protruding anterodistal corner of protarsomere IV and is not a - -Copelatus + +Copelatus . In addition, the elytral striae are very irregular, more like very elongate and deep strioles and possibly not homologous to the regular impressed striae in - -Copelatus + +Copelatus . We here transfer it to - -Madaglymbus + +Madaglymbus : - -Madaglymbus unguicularis + +Madaglymbus unguicularis ( -Regimbart +Regimbart , 1903) comb. nov. diff --git a/data/48/69/3E/48693E4733015386A1417218CDEAFB7A.xml b/data/48/69/3E/48693E4733015386A1417218CDEAFB7A.xml index 471460015cb..d1763cb10f1 100644 --- a/data/48/69/3E/48693E4733015386A1417218CDEAFB7A.xml +++ b/data/48/69/3E/48693E4733015386A1417218CDEAFB7A.xml @@ -1,260 +1,260 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - - + + + + Copelatus distinguendus -Regimbart +Regimbart , 1903 -Figs 4B +Figs 4B , -7D +7D - - -Copelatus distinguendus -Regimbart + + +Copelatus distinguendus +Regimbart , 1903: 19 [nom. nov., referring to his description of Malagasy material under the name -Copelatus duodecimstriatus -Aube +Copelatus duodecimstriatus +Aube in - -Regimbart + +Regimbart 1895 : 163] - -Type locality. -Environs de Tananarive [surroundings of Antananarivo] and Fianarantsoa, Madagascar. + +Type locality. +Environs de Tananarive [surroundings of Antananarivo] and Fianarantsoa, Madagascar. - -Type information from original description. -Based on an unknown number of specimens (syntypes) collected by Sikora (Antananarivo) and Perrot (Fianarantsoa). + +Type information from original description. +Based on an unknown number of specimens (syntypes) collected by Sikora (Antananarivo) and Perrot (Fianarantsoa). - -Type material studied. -Type material in MNHN not studied, as it was out on loan. + +Type material studied. +Type material in MNHN not studied, as it was out on loan. - -Additional material studied. - -Fianarantsoa. Matsiatra Ambony: Ambalavao, Ambohimahasoa, Lalangina + +Additional material studied. + +Fianarantsoa. Matsiatra Ambony: Ambalavao, Ambohimahasoa, Lalangina : -2♀, 1♂(GP) (NHMUK): // BMNH-792954-6 // MAD: FIAN: Andringitra | Zomandao R.: River edge: Bottle trap | P39EM08: N: -22.1043: E:46.92: 1420 m | 09/V/2006: Leg. Bergsten et al // BMNH (DNA Voucher) // -1♀ (NMW): // Data in NHRS | JLKB | 000010718 // Madagascar: Ambohimahasoa (Fianarantsoa) | RN7 (Km 378) | 16.04.2011: Leg. R. Gerecke (MD211) // spring area with meadow swamps Exp. E: -21°15'41.5"S +21°15'41.5"S , -47°14'10.9"E +47°14'10.9"E , 1500 m // -2♀ (NMW): // Data in NHRS | JLKB | 000065754-5 // Madagascar Est, 1100- | 1200m, P.N. Ranomafana // Vohiparara, 21.-24.1.1993 | J. -Janak +Janak lgt // -1♂(GP), 1♀, 5 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010618-9, 10787(Alc.) // Madagascar: Fianarantsoa: Matsiatra | Ambony: Ranomafana NP: | Sahamalaotra 2Km from Vohiparara: | S21.23807, E047.39489, 1140 m.| 01:XI:2011: stamping with sieves: forest | bog in rainforest: Field# MAD11-12 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♀(Alc.) (NHRS): // NHRS-JLKB | 000010830 // MAD: FIAN: Matsiatra Ambony | Ranomafana NP: 450m along | Sahamalaotra trail, left at the first | junction: Mad14-07: forestmarsh: | -21.2382S -49.3947E +21.2382S +49.3947E : 1130 m: 02.XI.2014 // Leg. J. Bergsten, | T. Ranarilalatiana | & S. Holmgren // -1♀ (NHMUK): // BMNH-670601_MSL007 | 06.xii.2004, Ranomafana, | Madagascar: lat - 21.2359 | Lon 47.3963 Coll Balke_M; | Monaghan_M // DNA Voucher | BMNH <670601> | MSL007:E07 // - + Fianarantsoa. -Amoron'i +Amoron'i Mania: Ambositra : -3♀ (NHMUK): // BMNH-792962-4 // Col de Tapias: Rte -Tana-Fianarantsoa +Tana-Fianarantsoa : Pond | P36C: N: -20.772: E:47.179: 1717 m | 06/V/2006: Leg. Bergsten et al // BMNH (DNA Voucher) // -1♀ (NHMUK): // BMNH-792912 // Ambositra: Ankazomivady forest | 01.xii.2005 // BMNH (DNA Voucher) // -9♀ (NHMUK), 8 ex. (Alc.) (NHRS): // BMNH-729890, 729893, 729896-7, 792976-80, 10793(Alc.) // 08.xii.2004, Col de Tapias, | P30MD33: lat -20.238 | Lon 47.1 Coll Balke_M | Monaghan_M // BMNH (DNA Voucher) // -Antananarivo. Vakinankaratra: Ambatolampy +Antananarivo. Vakinankaratra: Ambatolampy : -3♂(GP), 3♀, 10 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010620-5, 10786(Alc.) // Madagascar: Antananarivo: | Vakinankaratra: Manjakatompo Stn. | -forestiere +forestiere : 500m E Lac Froid by the | road: S -19.34485 E +19.34485 E 047.33381, 1620 m. | 04.XI.2011: GB Nets and sieves: pond | and inlet stream: MAD11-16 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♂(GP) (NHRS): // NHRS-JLKB | 000010626 // Madagascar: Antananarivo: | Vakinankaratra: Manjakatompo Stn. | -forestiere +forestiere : Analafandriana 500 m S | fish farm by the road: S19.36191 E | 47.31495, 1730 m, 03.XI.2011: GB | Nets: grassy pond: Field# MAD11-14 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♂(GP) (NHRS): // NHRS-JLKB | 000010660 // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-03: | "Lac froid": S-19.34292; E047.33893; | 1651 m: lake with grass at margins: | 03/02/2016 Leg. T. Ranarilalatiana // -4♂(GP), 4♀, 5 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010662-9, 10791(Alc.) // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-46: | "Lac froid": S-19.34292; E047.33893; | 1651 m: lake with grass at margins: | 17/09/2016 Leg. T. Ranarilalatiana // -1♂(GP) (NHRS): // NHRS-JLKB | 000010661 // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-10: | Ankafotra mountain: S-19.33753; | E047.24530; 2466 m: streampools: | 07/02/2016; Leg. T. Ranarilalatiana // -4♂(GP), 2♀, 18 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010670-5, 10792(Alc.) // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-47: | Ankafotra mountain: S-19.33753; | E047.24530; 2466 m: streampools: | 18/09/2016; Leg. T. Ranarilalatiana // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010832 // Madagascar: Antananarivo: | Vakinankaratra: Manjakatompo Stn | -forestiere +forestiere : Analamitana: S19.363972 E | 047.299083, 1757 m. 22:I:2012: swamp | near stream: Field# MJK12-02: Leg. T. | Ranarilalatiana & J.H. Randriamihaja // -Antananarivo. Analamanga: Anjozorobe, Ankazobe +Antananarivo. Analamanga: Anjozorobe, Ankazobe : -1♂(GP), 1♀ (NHRS): // NHRS-JLKB | 000010684-5 // Madagascar: Anjozorobe: MAD16-36: | Amboasarianala: S-18.45792; E047. | 93438; 1367 m: Ambatovikinina stream: | 04/04/2016; Leg. T. Ranarilalatiana // -1♂(GP), 1♀ (NHRS): // NHRS-JLKB | 000010686-7 // Madagascar: Anjozorobe: MAD16-43:| Amboasarianala, Antanambe stream: | S-18.4671; E047.93807; 1271 m: Stream | with sidepools: 07/04/2016 | Leg. T. Ranarilalatiana // -2♂(GP), 2♀, 1♀(Alc.) (NHRS): // NHRS-JLKB | 000010688-91, 10790(Alc.) // Madagascar: Anjozorobe: MAD16-44 | Amboasarianala, Mangarivotra stream: | S-18.4676; E047.92535; 1271 m: stream | with bedrock and grass at edge | 07/04/2016; Leg. T. Ranarilalatiana // -2 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010831 // MAD: ANTA: Analamanga: Anjoz | orobe forest reserve: Marsh next | to the stream by Saha forest, 10Km E of Anjozorobe: MAD14- | 78: forestmarsh: 18.4128S | 47.9439E; 1320 m; 23.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -2♂(GP), 2♀, 6 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010676-9, 10797(Alc.) // Madagascar: Ankazobe: MAD16-24: | Firarazana: S-18.13132; E047.23976; | 1551 m; Lake with grass at margins: | 12/03/2016; Leg. T. Ranarilalatiana // -2♀, 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010680-1, 10789(Alc.) // Madagascar: Ankazobe: MAD16-26: | Maharidaza, Large stream by the road to | military camp: S-18.22102; E047.27087; | 1547 m: stream and bog with grass: | 14/03/2016; Leg. T. Ranarilalatiana // -1♂(GP), 1♀, 7 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010682-3, 10788(Alc.) // Madagascar: Ankazobe: MAD16-29: | Firarazana, SW of Ambohitantely | reserve: S-18.16717; E047.26090; | 1532 m: Bog with grass: 17/03/2016; | Leg. T. Ranarilalatiana // -3 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010829 // MAD: ANTA: Analamanga: Mana- | nkazo river by the bridge of | RN4: Mad14-75: medium size river over bedrock: 18.158S | 47.2104E: 1450 m: 21.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -14 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010834 // MAD: ANTA: Analamanga: Andra- | nofeno river by the bridge of | RN4, next to Andranofeno Sud | village: Mad14-74: medium size, | slow flowing river: 18.0844S | 47.1776E: 1430 m: 21.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // - 1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010835 // Madagascar: Ankazobe: TR18L10: | Stream by the bridge S of Ambohitantely | reserve: S-18.2023; E047.2780; 1556 m: | Hygropetric rock and stagnant pools: | 11/04/2018 | Leg. T. Ranarilalatiana -1♀ (NMW): // Data in NHRS | JLKB | 000065756 // RM: Betsiboka Bas (PO533) | Andranofeno Sud Riv. | -47°10'46"E +47°10'46"E , -18°05'00"S +18°05'00"S | 06.11.1995 | Leg. Elouard, J.-M., Oliarinony. R. // -Toamasina. Alaotra Mangoro: Ambatondrazaka, Andilamena, Moramanga +Toamasina. Alaotra Mangoro: Ambatondrazaka, Andilamena, Moramanga : -1♀ (NMW): // Data in NHRS | JLKB | 000065759 // E-Madagascar (09) Ambaton- | drazaka Region, 5Km N Didy | 1100-1200 m. asl. 14-16.01.1995 | G. Dunay & J. -Janak +Janak coll. // -1♀ (NMW): // Data in NHRS | JLKB | 000065760 // E-Madagascar (10) | Ambatombe, near Andilamena | 900 m asl. 17.01.1995 | G. Dunay & J. -Janak +Janak coll. // -1♂ (NMW): // Data in NHRS | JLKB | 000065761 // E-Madagascar (11) Ampamoho | near Andilamena, 1200-1300 m. | asl. 18-20.01.1995 | G. Dunay & J. -Janak +Janak coll. // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010627 // MAD: TOAM: Alaotra Mangoro | Betsabora river by RN2 near | Antsapanana village, 6 Km W | of Moramanga: MAD14-81: river | with side pools: 18.9247S | 48.1828E; 900 m; 24.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♂(Alc.) (NHRS): // NHRS-JLKB | 000010833 // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra SR: bog | at S border of reserve: S18.95456 E | 048.44048, 910 m: 09.XI.2011: GB | Nets and sieves: bog with red mud: | Field# MAD11-27 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♀ (NHMUK): // BMNH-677103 // 11.xii.2004, Andasibe, | Madagascar: P27MD36 | Coll Balke_M; | Monaghan_M // (DNA Voucher) // -1♀ (NMW): // Data in NHRS | JLKB | 000065757 // MADAGASCAR (Md-4) | Andasibe, NP Perinet | 1150 m, -Pfuetze +Pfuetze auf waldwiese | und in Kleinen Bach | 7.12.2000, Leg. W. Dolin // -1♀ (NMW): // Data in NHRS | JLKB | 000065758 // E-Madagascar (07) | Andranokobaka, N Moramanga | 800 m. Asl, 13.01.1995 | G. Dunay & J. -Janak +Janak coll. // -Antsiranana. Sava: Andapa +Antsiranana. Sava: Andapa : -1♂ (GP), 1♂ (Alc.) (NHRS): // NHRS-JLKB | 000010827, 10828 (Alc.) // MAD: ANTS: Sava: Anjanaharibe | Sud NP: river Marolakana at the | crossing place: Mad14-64: larger | river with rocks; 14.7623S | 49.4834E: 920 m: 15.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // - -Diagnosis. - + +Diagnosis. + Body shape elongate oval, convex, and attenuate posteriorly, with uniform black colouration ( -Fig. 7D +Fig. 7D ). The pattern of abbreviation of the elytral striae separates this species from all other Malagasy - -Copelatus + +Copelatus species. Striae 1, 3, and 5 present only in posterior one third (1) or two thirds (3, 5); striae 2, 4, and 6 fragmented anteriorly and, except for stria 4, never reach the base as clearly impressed striae. Penis profile in lateral view is characteristic, resembling a -"pumpjack" +"pumpjack" ( -Fig. 4B +Fig. 4B ). - -Description. -Body length: 5.3-6.3 mm. Body shape elongate oval, convex, and attenuate posteriorly. Head, pronotum, and elytra all the same colour, ferrugineous black and finely punctate. - + +Description. +Body length: 5.3-6.3 mm. Body shape elongate oval, convex, and attenuate posteriorly. Head, pronotum, and elytra all the same colour, ferrugineous black and finely punctate. + Lateral margin of pronotum rusty ferrugineous, with short sparse strioles. Pronotum with puncture rows and microsculpture. Elytra narrowly testaceous to ferrugineous posterolaterally. Six elytral striae present and one submarginal stria ( -Fig. 7D +Fig. 7D ). Striae 2, 4, and 6 longer than striae 1, 3, and 5, at least as continued fragments; first stria shortest, present only in posterior fourth to posterior third. Second and third striae present as clearly impressed striae in posterior half or a little longer, but second stria continues, albeit fragmentary and less impressed, more distinctly all the way to base. Fourth stria the most complete, more or less present to base but less deeply impressed or fragmentary in anterior third. Fifth stria abbreviated a little less than third stria, present in approximately posterior two thirds, and like stria 3 has small isolated point remnants of stria more anteriorly. Sixth stria longer than fifth and with more distinct remnants of striae present anteriorly. Elytra covered with punctures in apical part. -Ventral side ferrugineous dark brown. Metacoxa and abdominal sternite II, III, IV, striolate. Prosternal process raised medially but rounded, not carinate. Lateral parts of metaventrite broad. Metacoxal lines short and strongly diverging anteriorly. Antenna, palps, pro- and mesothoracic legs brown to yellowish, but metathoracic legs dark brown. - +Ventral side ferrugineous dark brown. Metacoxa and abdominal sternite II, III, IV, striolate. Prosternal process raised medially but rounded, not carinate. Lateral parts of metaventrite broad. Metacoxal lines short and strongly diverging anteriorly. Antenna, palps, pro- and mesothoracic legs brown to yellowish, but metathoracic legs dark brown. + Male: protibia slightly widened at apex, somewhat curved and angulate basally. Penis in lateral view curved with two points where curvature is more abrupt, constricted before apex at a narrow -"neck" +"neck" and expanding to apical part ( -Fig. 4B +Fig. 4B ). As a whole in lateral view, the penis bears resemblance to the silhouette of a -"pumpjack" +"pumpjack" . Penis apex in ventral view left-turned. Parameres broad at base, curved and tapering towards apex, apical part with numerous setae on ventral margin ( -Fig. 4B +Fig. 4B ). -Female: dorsal sculpture similar to male. +Female: dorsal sculpture similar to male. - -Distribution. - + +Distribution. + Occurs on Madagascar and Mauritius ( -Gomy 2016 +Gomy 2016 ). -Guignot (1948) +Guignot (1948) tentatively refer a single female specimen from Lulua in DR Congo to - -C. distinguendus + +C. distinguendus but this was very likely a different species. In Madagascar, it has been recorded from the provinces of Antananarivo, Fianarantsoa Toamasina, and Antsiranana; more specifically from Anjozorobe Angavo reserve, around Ankazobe, Manankazo, Andranofeno Sud, around Ambohitantely reserve, Manjakatompo Ankaratra reserve; also from the eastern central area around Moramanga, Analamazaotra NP; in the southeast from Ambositra in col de Tapia, Ankazomivady forest, Ranomafana NP, Andringitra NP; and from the northeast in Anjanaharibe Sud reserve ( -Fig. 11D +Fig. 11D ). - -Habitat and ecology. -This species has been collected in various localities, mostly from open, partly deforested areas at mid- to high altitudes or open forest marshes. It occurs at altitudes above 900 m and is often associated with grass vegetation along lake shores and in marshes, found by stamping, and at vegetation-rich margins of rivers. + +Habitat and ecology. +This species has been collected in various localities, mostly from open, partly deforested areas at mid- to high altitudes or open forest marshes. It occurs at altitudes above 900 m and is often associated with grass vegetation along lake shores and in marshes, found by stamping, and at vegetation-rich margins of rivers. - -Comments. - - -Regimbart + +Comments. + + +Regimbart (1895) described this species under the name - -Copelatus duodecimstriatus + +Copelatus duodecimstriatus -Aube +Aube but realised later (1903) that what he had described from Madagascar was not conspecific with -Aube's - -C. duodecimstriatus +Aube's + +C. duodecimstriatus from the Mascarene islands and he gave it the new name - -C. distinguendus + +C. distinguendus , referring to his description from 1895. - -Copelatus duodecimstriatus + +Copelatus duodecimstriatus lacks submarginal striae, has a distinctly different penis shape, and is endemic to Mauritius ( -Vinson 1967 +Vinson 1967 ; -Guignot 1961 +Guignot 1961 ; -Gomy 2016 +Gomy 2016 ). The closest relatives of - -Copelatus distinguendus + +Copelatus distinguendus are likely a group of species on continental Africa with similar genitalia (e.g., - -C. ateles + +C. ateles Guignot, 1955, - -C. sylvaticus + +C. sylvaticus Guignot, 1952, - -C. andreinii + +C. andreinii -Regimbart +Regimbart , 1905, and additional species described by -Bilardo and Rocchi 2013 +Bilardo and Rocchi 2013 ). These have a variable number of elytral striae from five discal to ten discal plus one submarginal, but similar -"pumpjack" +"pumpjack" -shaped genitalia ( -Bilardo and Rocchi 2013 +Bilardo and Rocchi 2013 ). - -Copelatus distinguendus + +Copelatus distinguendus differs from these especially by the lack of a lateral tooth on the male penis in ventral view. diff --git a/data/57/3F/0C/573F0C84F70555E7A4F6FC4974616368.xml b/data/57/3F/0C/573F0C84F70555E7A4F6FC4974616368.xml index 35dbe0107c7..4e892c75880 100644 --- a/data/57/3F/0C/573F0C84F70555E7A4F6FC4974616368.xml +++ b/data/57/3F/0C/573F0C84F70555E7A4F6FC4974616368.xml @@ -1,236 +1,236 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Copelatus insuetus Guignot, 1941 -Figs 6A + + + +Copelatus insuetus Guignot, 1941 +Figs 6A , -9A +9A - - -Copelatus insuetus + + +Copelatus insuetus Guignot, 1941: 39. - -Type locality. -Madagascar, Perinet [= Analamazaotra NP]. + +Type locality. +Madagascar, Perinet [= Analamazaotra NP]. - -Type information from original description. -Based on single male specimen (holotype). Madagascar, Perinet. + +Type information from original description. +Based on single male specimen (holotype). Madagascar, Perinet. - -Type material studied. - -Toamasina. Alaotra Mangoro: Moramanga + +Type material studied. + +Toamasina. Alaotra Mangoro: Moramanga : -HT♂ (GP) (MNHN, "coll. Guignot"): Data in NHRS | JLKB | 000030227 // Madagascar | Perinet // Type [red label] // [male symbol] // -Copelatus +Copelatus | -insuetus +insuetus | guign. Type // - -Additional material studied. - -Toamasina. Alaotra Mangoro: Moramanga, Ambatondrazaka, Andilamena + +Additional material studied. + +Toamasina. Alaotra Mangoro: Moramanga, Ambatondrazaka, Andilamena : -2♀ (MNHN, "coll. Guignot"): Data in NHRS | JLKB | 000030228-9 // Perinet // INSTITUT | SCIENTIFIQUE | MADAGASCAR // [female symbol] // -5♀ (NMW): Data in NHRS | JLKB | 000030302-6 // Madagascar (Md 4/5) | Andasibe NP, Perinet | 1250 m -Pfuetze +Pfuetze im Urwald | 5.12.2000, leg. W. Dolin // -1♂ (NMCP, "coll. -Hajek" +Hajek" ): Data in NHRS | JLKB | 000030312 // Madagascar 10.1.2007 | Andasibe NP. | Lokato-near Andasibe | Mracek Z. leg. // coll. Jiri -HAJEK +HAJEK | National Museum | Prague, Czech Republic // -1♀, 1♂ (NMW): Data in NHRS | JLKB | 000030307-8 // E-Madagascar (10) | Ambatombe near Andilamena | 900 m asl. 17.01.1995 | G. Dunay & J. -Janak +Janak coll. -1♀, 1♂ (NMW): Data in NHRS | JLKB | 000030309-10 // E-Madagascar (11) Ampamoho | near Andilamena, 1200-1300 m. | asl. 18-20.01.1995 | G. Dunay & J. -Janak +Janak coll. // -1♀ (NMW): Data in NHRS | JLKB | 000030311 // Madagascar 18-20.1.1995 | 5km S Ampamoho | nr. Andilamena, 950-1000 m | Dunay & -Janak +Janak (11) // -1♀ (NHMUK): // BMNH-797884 // MAD: AMPA: Moramanga: Andasibe | Andasibe NP: Chanel| P61BI02: N: -18.935: E:48.418: 944 m | 04/I/2007 Leg. Isambert et al // DNA Voucher | BMNH <797884> | MSL294:A12 // -2♀ (NHRS): // NHRS-JLKB | 000010692-3 // Madagascar: Tamatave: Alaotra Mangoro: | Analamazaotra RS; Bas fond, non-permanent | pond near trail to "lac rouge"MAD15-1 | 943 m, -18°56'26.7"S +18°56'26.7"S , -048°25'03.9"E +048°25'03.9"E , 16.III.2015 | Among vegetation and dead leaves in the pond | Leg.T. Ranarilalatiana & H.J. Randriamihaja // -1♀, 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000065745, 10783 (Alc.) // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra RS: close to | park entrance: -18.93573S +18.93573S ; -048.41741E +048.41741E | 930 m. 08.XI.2011 GB Nets and sieves: | stagnant pools in tributary to | Analamazaotra river; Field# MAD11-26 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♂ (GP), 1♀, 3 ex. (1♂, 2♀) (Alc.) (NHRS): // NHRS-JLKB | 000010839, 65702, 10840 (Alc.) // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, Analamazaotra | 250m E of park entrance: Mad14-18: | dried up river bed with stagnant pools: | 18.9357S 48.4174E: 930 m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♀ (NHRS): // NHRS-JLKB | 000010841 // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, Analamazaotra | pond: 100m E from Lac vert: Mad14-15: | shallow forest pool: -18.9385S -48.4219E +18.9385S +48.4219E : | 940 m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♂ (GP), 1♀, 1♀ (Alc.) (NHRS): // NHRS-JLKB | 000065705, 65777, 10844(Alc.) // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, Analamazaotra | 150 m E of park entrance: Mad14-14: | shallow partly dried out forest pond: | -18.9355S -48.4166E +18.9355S +48.4166E : 930 m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♂ (GP), 34 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010842, 10843(Alc.) // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, | Mantadia: 12Km N of park | entrance: Mad14-83: | forest pond with some | vegetation: 18.7911S | 48.4259E: 960 m: 28.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -6♂ (GP), 4♀, 33 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010534-7, 10606-11, 10780(Alc.) // MAD: TOAM: Alaotra Mangoro: | Betsabora river by RN2 near | Antasmpanana village, 6Km W | of Moramanga: MAD14-81: river | with side pools: 18.9247S | 48.1828E: 900 m: 24.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♀ (NHMUK): // BMNH-797895 // MAD: TOAM: Ambatondrazaka | Zahamena: Zahamena NP: Stream | P60BI15:N: -17.52: E:48.721: 1075 m| 31:XI:2006: Leg. Isambert et al // DNA Voucher | BMNH <797895> | MSL294:B11 // -4♂ (GP), 1♀ 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010817, 10872-4, 65700, 10880(Alc.) // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | path towards Camp Cascade: S17.5166; | E048.7227; 1040 m. 07.III.2018; GB Nets, | white pan & sieves: Waterfilled goldigging | hole in forestswamp: Field# MAD18-80 | Leg. J. Bergsten, & T. Ranarilalatiana // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010885 // Madagascar: Toamasina: Ambatondrazaka | Zahamena NP: the way to Camp | Cascade: 19.III.2013, GB Nets & sieves: slow | stream, Leg. J.H. Randriamihaja & | T. Ranarialalatiana: Field# ZAH13-02 // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010881 // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Analamaitso forest: S17.5179; E048.7232; | 1050 m. 07.III.2018; GB Nets, white pan & | sieves: small foreststream: Field# MAD18-81 | Leg. J. Bergsten, & T. Ranarilalatiana // -3♀ (Alc.) (NHRS): // NHRS-JLKB | 000010882 // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Ambavahala forest: S17.5300; E048.7161; | 1070 m. 08.III.2018; GB Nets, white pan & | sieves: larger shaded forestswamp: Field# MAD18-90 | Leg. J. Bergsten, & T. Ranarilalatiana // -1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010875, 10883(Alc.) // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Pandanus Swamp: S17.5166; E048.7227; | 1040 m. 08.III.2018; GB Nets, white pan & | sieves: forestswamp: Field# MAD18-91 | Leg. J. Bergsten, & T. Ranarilalatiana -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010884 // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Analamaitso forest: S17.5225; E048.7250; | 1090 m. 08.III.2018; GB Nets, white pan & | sieves: waterfilled golddigging hole in flat: Field# MAD18-92 | Leg. J. Bergsten, & T. Ranarilalatiana // -2♂ (GP), 3♀, 6♀ (Alc.), 6ex. (3♂, 3♀) (Alc.), (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010816, 10876-9, 10886(Alc.), 65753(Alc.) // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: 150 m | upstream to Camp Cascade: S17.5458; E048.7244; | 1270 m. 10-11.III.2018; GB Nets, white pan & | sieves: waterpool with dead leaves: Field# MAD18-109, | Leg. J. Bergsten, & T. Ranarilalatiana // -Toamasina. Analanjirofo: Maroantsetra +Toamasina. Analanjirofo: Maroantsetra : -1♂, 1♀ (MNHN, "coll. Legros"): Data in NHRS | JLKB | 000030314-5 // Maroantsetra | Vadon IV.50 // 1/6 //? insuetus | guign. // [These two specimens were seen in MNHN, but their identity, after the discovery of additional very similar species in the complex, was not confirmed. We therefore only tentatively refer them to this species.] -Fianarantsoa. Vatovavy Fitovinany: Ifanadiana +Fianarantsoa. Vatovavy Fitovinany: Ifanadiana : -1♀ (CAS): // CASENT-8131927 // Madagascar: Province | Fianarantsoa, Parc National | Ranomafana, radio tower | at forest edge, elev 1130 m | 19-26 February 2002 // 21°15.05'S, 47°24.43' | coll: M. Irwin, R. -Harin'Hala +Harin'Hala | California Acad. of Sciences | malaise. Mixed tropical | forest MA-02-09B-17 // -1♂ (teneral) (NMPC, "coll. -Hajek" +Hajek" ): Data in NHRS | JLKB | 000030313 // Madagascar 26-31.I.2007 | Ranomafana NP | Ranomafana vill. env. | Z. Mracek leg. // coll. Jiri -HAJEK +HAJEK | National Museum | Prague, Czech Republic // -Fianarantsoa. Matsiatra Ambony: Lalangina +Fianarantsoa. Matsiatra Ambony: Lalangina : -1♀ (CAS): // CASENT-5004000 // Madagascar, Fianarantsoa | Province, Ranomafana National | Park, Vohiparara village, 1160m | mixed tropical forest | 2-22 January 2001 | 21.23906S / 47.38487E | COL-DHK-2001-003 // D.H. & K.M. Kavanagh | R.L. Brett, E. Elsom, F. Vargas, | R. Ranaivosolo, E.F. Randrianirina, | N. Rasoamanana, T.J. | Ravelomanana and H.C. | Raveloson collectors // -4♂ (CAS): // CASENT-5004056, 5004070-1, 5004075 // Madagascar, Fianarantsoa | Province, Ranomafana | National Park, Vohiparara | area, 1170m mixed tropical | forest 2-22 January 2001 | -21.22644S +21.22644S , -47.36979E +47.36979E , | Stop# DHK-01-004 // D.H. & K.M. Kavanagh | R.L. Brett, E. Elsom, F. | Vargas, R. Ranaivosolo, | E.F. Randrianirina, N. | Rasoamanana, T.J. | Ravelomanana and H.C. | Raveloson collectors // -Antsiranana. Diana: Nosy-Be +Antsiranana. Diana: Nosy-Be : -1♂(GP) (MNHN): // Data in NHRS | JLKB | 000030230 // Nosy-Be | Sokobe [=Lokobe NP] | V. 57 Hoyt Coll // ♂ [male symbol] // -Mahajanga. Boeny: Ambato-Boeny +Mahajanga. Boeny: Ambato-Boeny : -1♀ (NHRS): // NHRS-JLKB | 000010694 (JB206) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010781 (JB205) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30350 | E046.81068, 87 m.a.o. 29.XI.2009 | Water Net, Field# MAD09-03 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // - -Diagnosis. - + +Diagnosis. + Body shape elongate oval, elytra brown with basal testaceous band, with six discal and one submarginal stria ( -Fig. 9A +Fig. 9A ). - -Copelatus insuetus + +Copelatus insuetus can be distinguished from the following species based on the penis ventral outline in lateral view with a distinct -"shoulder" +"shoulder" ( -Fig. 6A +Fig. 6A ), an apex in left lateral view with a dorsal ridge crossing posterior dorsal margin and medial part longitudinally coarsely sulcate ventrolaterally in right lateral view ( -Fig. 6A +Fig. 6A ). - -Description. - + +Description. + Body length 4.3-5.0 mm. Body shape elongate, weakly oval, some specimens appearing subparalell. Head rufotestaceous anteriorly and posteriorly but infuscated by two blotches inbetween the eyes. Blotches usually meet in middle and form an M-mark, but sometimes infuscation reduced to nearly absent. Pronotum usually with extensive medial infuscation leaving only lateral, anterolateral and posterolateral areas testaceous. Some specimens with less extensive medial infuscation ( -Fig. 9A +Fig. 9A ). Elytra brown with an irregular testaceous band at base. Testaceous band with longest projection posteriorly in second interval. All appendages testaceous. -Elytra with six clearly impressed discal and one submarginal stria. Fifth and sixth striae abbreviated anteriorly, and also first stria may be somewhat abbreviated. Submarginal stria starting between one third from the base and midway between base and apex. Head, pronotum, and elytra microreticulate and finely micropunctate. Posterolateral corners of pronotum with strioles that extend along posterior surface, reduced or absent medially. - +Elytra with six clearly impressed discal and one submarginal stria. Fifth and sixth striae abbreviated anteriorly, and also first stria may be somewhat abbreviated. Submarginal stria starting between one third from the base and midway between base and apex. Head, pronotum, and elytra microreticulate and finely micropunctate. Posterolateral corners of pronotum with strioles that extend along posterior surface, reduced or absent medially. + Ventral side ferrugineous brown, except prothorax, epipleura, appendages, and gula of head which are testaceous. Area around metacoxal lines lighter (not all individuals) and also spots laterally on abdominal sternites -II-VII +II-VII and along some sternite margins are lighter. Metacoxa and abdominal sternites -II-IV +II-IV marked with strioles. Prosternal process short, broad, with blunt apex. Lateral parts of metaventrite broad. Metacoxal lines long, abbreviated only slightly before metaventral margin, diverging anteriorly. - + Male: first three pro- and mesotarsomeres widened and ventrally equipped with suction cups; number of suction cups per segment for -I-III +I-III : 7:4:4 for both pro- and mesotarsus. Protibia modified, narrow with a bisinuate angulate ventral margin at base, broadened distally. Pro- and mesotarsal claws unmodified. Penis in ventral view thin and simple, apical part slightly left-turned. Penis in lateral view angled after basal third forming a -"shoulder" +"shoulder" . Apex in left lateral view with a characteristic dorsal ridge crossing posterior dorsal margin ( -Fig. 6A +Fig. 6A ). Penis serrated with fine transverse ridges preapically on the left ventral ridge and halfway down the left side. Penis medially strongly longitudinally sulcate ventrolaterally in right lateral view ( -Fig. 6A +Fig. 6A ). Parameres as in -Figure 6A +Figure 6A . -Female: anterior half of elytra finely striolated from second or third elytral interval to external margin. Degree of elytral striolation in females quite variable between specimens. These strioles are finer than the strioles on pronotum found in both sexes. - - +Female: anterior half of elytra finely striolated from second or third elytral interval to external margin. Degree of elytral striolation in females quite variable between specimens. These strioles are finer than the strioles on pronotum found in both sexes. + + Figure 9. Habitus, dorsal view. -A +A Female: - -Copelatus insuetus + +Copelatus insuetus -B +B Male: - -Copelatus vokoka + +Copelatus vokoka sp. nov. -C +C Female: - -Copelatus kely + +Copelatus kely sp. nov. -D +D Female: - -Copelatus + +Copelatus sp. 3 (Ivohibe). - -Distribution. - + +Distribution. + Endemic to Madagascar. Known with certainty from the eastern escarpment, Ranomafana NP in the south, Analamazaotra NP, and Mantadia NP in the central region, and Zahamena NP and Andilamena further north ( -Fig. 12A +Fig. 12A ). We have also seen older material including males from Paris museum from lowland Lokobe NP (island of Nosy Be) in NW and Maroantsetra in NE (not reconfirmed) that correspond to the same species. Finally, two females (NHRS) from Ankarafantsika NP is likely this species. - -Habitat and ecology. -We have collected this species in Analamazaotra NP in stagnant forest pools with vegetation or with plentiful of dead leaves. Also found in a sidepool next to a river in semi-open degraded area near Moramanga. The eastern escarpment localities range in altitude between 900-1300 m, but if the localities of Nosy Be and Maroantsetra are correct this species can also occur at lowland sea level altitudes. It seems to be most abundant in tropical eastern forests, but if the Ankarafantsika females belong to this species, then it can also occur in western deciduous forests. + +Habitat and ecology. +We have collected this species in Analamazaotra NP in stagnant forest pools with vegetation or with plentiful of dead leaves. Also found in a sidepool next to a river in semi-open degraded area near Moramanga. The eastern escarpment localities range in altitude between 900-1300 m, but if the localities of Nosy Be and Maroantsetra are correct this species can also occur at lowland sea level altitudes. It seems to be most abundant in tropical eastern forests, but if the Ankarafantsika females belong to this species, then it can also occur in western deciduous forests. - -Comments. - + +Comments. + Eventual older records of - -C. insuetus + +C. insuetus should be regarded with caution in light of the habitually very similar new species described below. diff --git a/data/59/BB/65/59BB653E8C1950B0B863E8393226197D.xml b/data/59/BB/65/59BB653E8C1950B0B863E8393226197D.xml index bd1763d4262..83f68be86a1 100644 --- a/data/59/BB/65/59BB653E8C1950B0B863E8393226197D.xml +++ b/data/59/BB/65/59BB653E8C1950B0B863E8393226197D.xml @@ -1,234 +1,234 @@ - - - -A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China + + + +A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China - - -Author + + +Author -Ge, Si Xun -https://orcid.org/0000-0003-3769-1530 -Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. +Ge, Si Xun +https://orcid.org/0000-0003-3769-1530 +Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. - - -Author + + +Author -Jiang, Zhuo Heng -School of Science, Westlake University, Hangzhou, China +Jiang, Zhuo Heng +School of Science, Westlake University, Hangzhou, China - - -Author + + +Author -Wang, Jia Qi -2288 Long, Hongxin Rd, Minhang District, Shanghai, China +Wang, Jia Qi +2288 Long, Hongxin Rd, Minhang District, Shanghai, China - - -Author + + +Author -Song, Kui -School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China +Song, Kui +School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China - - -Author + + +Author -Zhang, Chao -Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China +Zhang, Chao +Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China - - -Author + + +Author -Hu, Shao Ji -Institute of International Rivers and Eco-security, Yunnan University, Kunming, China -shaojihu@hotmail.com +Hu, Shao Ji +Institute of International Rivers and Eco-security, Yunnan University, Kunming, China +shaojihu@hotmail.com -text - - -Arthropod Systematics & amp; Phylogeny +text + + +Arthropod Systematics & amp; Phylogeny - -2023 - -2023-03-15 + +2023 + +2023-03-15 - -81 + +81 - -257 -287 + +257 +287 - -http://dx.doi.org/10.3897/asp.81.e85191 + +http://dx.doi.org/10.3897/asp.81.e85191 -journal article -http://dx.doi.org/10.3897/asp.81.e85191 -1864-8312-81-257 -CC0AB5E565324E9EB5D02FD08B5FFE2C -74CBFA410981516FBB87561614EFF561 +journal article +http://dx.doi.org/10.3897/asp.81.e85191 +1864-8312-81-257 +CC0AB5E565324E9EB5D02FD08B5FFE2C +74CBFA410981516FBB87561614EFF561 - - - -Pieris narina Verity, 1908 -stat. rev. + + + +Pieris narina Verity, 1908 +stat. rev. - - -Pieris napi var. bryoniae r. narina + + +Pieris napi var. bryoniae r. narina Verity, 1908; -Rhopalocera +Rhopalocera Palaearctica (13-14): 145, (17-20): pl. 32, f. 28-29, pl. 33, f. 2; TL: -"Naryn" +"Naryn" . (original description) - -Pieris narina narina + +Pieris narina narina Verity, 1908; -Eitschberger (1983) +Eitschberger (1983) : Herbipoliana 1-(1--2): 221-222 (infrasubspecific name validated as distinct species). - -Pieris napi var. banghaasi + +Pieris napi var. banghaasi Sheljuzhko, 1910; -Eitschberger (1983) +Eitschberger (1983) : Herbipoliana 1 (1-2): 221-223 (as a synonym of -P. narina +P. narina ) - -Pieris ochsenheimeri narina + +Pieris ochsenheimeri narina Verity, 1908; -Tadokoro et al. (2014) +Tadokoro et al. (2014) : Butterflies 65: 20-25 (as subspecies of -P. ochsenheimeri +P. ochsenheimeri Staudinger, 1886) - -Pieris napi + +Pieris napi (Linnaeus, 1758); Wu & Hsu (2017): Butterflies of China vol. 1: 401-404, f. 15-16 [ -MisID +MisID ] - -Description. - + +Description. + small in size. Male (Fig. -9A +9A ): both wings white on the upperside and pale yellowish on the underside. Costal margin black; apical spot blackish, triangular, extending along veins to outer margin and sometimes reached vein CuA2. Outer margin border of forewing blackish. The 1st discal spot and outer spot variable, distinct to nearly absent on both upperside and underside. The 2nd discal spot absent or only faintly indicated on the underside and the 3rd discal spot absent or sometimes very vague on the upperside. Outer margin border of hindwings grey to blackish, sometimes triangularly extending along veins. Female (Fig. -9B, C +9B, C ): both wings white to pale yellowish with intensive brownish or grey suffusion on the upperside while yellowish on the underside, but clearly in the apical 1/3 of discocell on the upperside of forewing. Hindwing with dark powder strongly developed along veins. - - -Figure 9. + + +Figure 9. Habitus of - -P. napi + +P. napi -complex species occurring in China. Composite pictures on the left show the upperside, on the right show the underside. -A +A - -C - -Pieris narina +C + +Pieris narina stat. rev. Verity, 1908. -D +D - -I - -Pieris mihon +I + +Pieris mihon Yakovlev, 2006 stat. nov. -J +J - -M - -Pieris dulcinea dulcinea +M + +Pieris dulcinea dulcinea (Butler, 1882). Scale bar represents 10 mm. (L, M reference Yakovlev, 2006). - -Distribution. -Kazakhstan, Kyrgyzstan, Tajikistan (Northern and Inner Tian Shan, Dzhngarsky Alatau), China (Xinjiang) + +Distribution. +Kazakhstan, Kyrgyzstan, Tajikistan (Northern and Inner Tian Shan, Dzhngarsky Alatau), China (Xinjiang) - -Phenology. -Probably univoltine, based on a single form of adults observed in July. + +Phenology. +Probably univoltine, based on a single form of adults observed in July. - -Male genitalia. - + +Male genitalia. + (Fig. -5H +5H ) terminal part of tegumen with pairs of sickle shaped convex; uncus broad with its basal part as wide as terminal part of tegumen, apical half of uncus digitation. - -Female genitalia. - + +Female genitalia. + (Fig. -6D +6D ) posterior apophysis short and slender; inner distal of sterigma bilobate lobe shaped, extend to the center; inner basal of sterigma with a distinct tooth near the basal part, acute-angled connected at the basal margin; signum relatively broad cordiform shaped, moderately tapering to the base. - -Note. - + +Note. + The species rather resembles - -P. ochsenheimeri + +P. ochsenheimeri , but can be distinguished by comparing multiple morphological characteristics (see notes in - -P. ochsenheimeri + +P. ochsenheimeri ); the blackish outer marginal border on both wings and the relatively smaller size make it easily separated from other species of the - -Pieris napi + +Pieris napi -complex distributed in the same region. This taxon was formerly elevated by -Eitschberger (1983) +Eitschberger (1983) to species level while -Tadokoro et al. (2014) +Tadokoro et al. (2014) regarded it as a subspecies of - -P. ochsenheimeri + +P. ochsenheimeri . However, this taxon has unique morphological characteristics and a previous study has shown that there is deep differentiation between - -P. narina + +P. narina and - -P. ochsenheimeri + +P. ochsenheimeri for the COI gene ( -Lukhtanov et al. 2009 +Lukhtanov et al. 2009 ). Our phylogenetic results also indicated that this species is more closely related to - -P. bryoniae + +P. bryoniae than to - -P. napi + +P. napi or - -P. ochsenhelmeri + +P. ochsenhelmeri , thus we confirmed it as a distinct species. diff --git a/data/77/89/CF/7789CF7BC6545583A4D46C09B92D9673.xml b/data/77/89/CF/7789CF7BC6545583A4D46C09B92D9673.xml index 107d3035b96..648744ea6ef 100644 --- a/data/77/89/CF/7789CF7BC6545583A4D46C09B92D9673.xml +++ b/data/77/89/CF/7789CF7BC6545583A4D46C09B92D9673.xml @@ -1,196 +1,196 @@ - - - -A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China + + + +A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China - - -Author + + +Author -Ge, Si Xun -https://orcid.org/0000-0003-3769-1530 -Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. +Ge, Si Xun +https://orcid.org/0000-0003-3769-1530 +Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China. - - -Author + + +Author -Jiang, Zhuo Heng -School of Science, Westlake University, Hangzhou, China +Jiang, Zhuo Heng +School of Science, Westlake University, Hangzhou, China - - -Author + + +Author -Wang, Jia Qi -2288 Long, Hongxin Rd, Minhang District, Shanghai, China +Wang, Jia Qi +2288 Long, Hongxin Rd, Minhang District, Shanghai, China - - -Author + + +Author -Song, Kui -School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China +Song, Kui +School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China - - -Author + + +Author -Zhang, Chao -Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China +Zhang, Chao +Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China - - -Author + + +Author -Hu, Shao Ji -Institute of International Rivers and Eco-security, Yunnan University, Kunming, China -shaojihu@hotmail.com +Hu, Shao Ji +Institute of International Rivers and Eco-security, Yunnan University, Kunming, China +shaojihu@hotmail.com -text - - -Arthropod Systematics & amp; Phylogeny +text + + +Arthropod Systematics & amp; Phylogeny - -2023 - -2023-03-15 + +2023 + +2023-03-15 - -81 + +81 - -257 -287 + +257 +287 - -http://dx.doi.org/10.3897/asp.81.e85191 + +http://dx.doi.org/10.3897/asp.81.e85191 -journal article -http://dx.doi.org/10.3897/asp.81.e85191 -1864-8312-81-257 -CC0AB5E565324E9EB5D02FD08B5FFE2C -74CBFA410981516FBB87561614EFF561 +journal article +http://dx.doi.org/10.3897/asp.81.e85191 +1864-8312-81-257 +CC0AB5E565324E9EB5D02FD08B5FFE2C +74CBFA410981516FBB87561614EFF561 - - - - + + + + Pieris melaina -Roeber +Roeber , 1907 - - -Pieris melaina -Roeber + + +Pieris melaina +Roeber , 1907; Gross-Schmett. Erde 1: 48, pl. 20 g; TL: -"Tibet" +"Tibet" (original description) - -Pieris melaina melaina -Roeber + +Pieris melaina melaina +Roeber , 1907; -Eitschberger (1983) +Eitschberger (1983) : Herbipoliana 1(1-2): 406 (pictured) - -Pieris melete melete + +Pieris melete melete ra. -montana +montana Verity, 1908; -Eitschberger (1983) +Eitschberger (1983) : Herbipoliana 1(2): 406-408 (as a synonym of -P. melaina +P. melaina ) - -Pieris melaina -Roeber + +Pieris melaina +Roeber , 1907; -Tadokoro et al. (2017) +Tadokoro et al. (2017) : -Lepidoptera +Lepidoptera Science: 68(3-4): 81-91 (pictured) - -Description. - + +Description. + Both wings whitish on the upperside while pale yellowish on the underside. Male (Fig. -13A, C +13A, C ): apical spot blackish and sickle shaped. The 1st discal spot distinct and almost combined with the apical spot, the 2nd discal spot absent or faintly developed on the upperside while distinct on the underside. The outer spot distinct on the upperside while absent on the underside. Female (Fig. -13B, D +13B, D ): extremely dark powdered along veins on the upperside, with the 1st and 2nd discal spot merging with the apical spot, thus brownish suffusion on almost all outer marginal area except slightly whitish area in the space between vein CuA2 and 2A. - - -Figure 13. + + +Figure 13. Habitus of species similar to the - -P. napi + +P. napi -complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. -A +A - -D - -Pieris melaina +D + +Pieris melaina -Roeber +Roeber , 1907. -E +E - -F - -Pieris extensa +F + +Pieris extensa Poujade, 1888. -G +G - -H - -Pieris chumbiensis gyantsensis +H + +Pieris chumbiensis gyantsensis Verity, 1911. -I +I - -L - -Pieris melete +L + +Pieris melete -Menetries +Menetries , 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983) - -Distribution. -East Tibet (Yadong, Bomi), India (border to Tibet) + +Distribution. +East Tibet (Yadong, Bomi), India (border to Tibet) - -Phenology. -Univoltine in high altitude habitat (occurs in July and August above 4000 m) while probably bivoltine in low altitude habitat (occurs in April and July below 3000 m). + +Phenology. +Univoltine in high altitude habitat (occurs in July and August above 4000 m) while probably bivoltine in low altitude habitat (occurs in April and July below 3000 m). - -Male genitalia. - + +Male genitalia. + (Fig. -5A +5A ) tegumen broad, distinctly convex on its 4/5 apically; basal part of uncus moderately convex and with its apically distinctly narrowed towards tip. - -Female genitalia. - + +Female genitalia. + (Fig. -8A +8A ) sterigma with its inner distal bilobate lobe-shaped, shallowly expended to the centre; inner basal of sterigma arcuately protruding, smoothly connected at the basal margin; signum variable, cordiform to near cordiform-shaped. diff --git a/data/7B/3F/DE/7B3FDEB746D75239A167247A824B06E0.xml b/data/7B/3F/DE/7B3FDEB746D75239A167247A824B06E0.xml index da29d51960e..8388bbf01f0 100644 --- a/data/7B/3F/DE/7B3FDEB746D75239A167247A824B06E0.xml +++ b/data/7B/3F/DE/7B3FDEB746D75239A167247A824B06E0.xml @@ -1,156 +1,156 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Madaglymbus apicalis (Fairmaire, 1898) -comb. nov. + + + +Madaglymbus apicalis (Fairmaire, 1898) +comb. nov. - - -Copelatus apicalis + + +Copelatus apicalis Fairmaire, 1898: 465. - -Type locality. -Madagascar, Suberbieville [= Maevatanana]. + +Type locality. +Madagascar, Suberbieville [= Maevatanana]. - -Type material examined. - -Mahajanga. Betsiboka: Maevatanana + +Type material examined. + +Mahajanga. Betsiboka: Maevatanana : -ST ♂ (GP) (MNHN, "coll. -Regimbart" +Regimbart" ): // Data in NHRS | JLKB | 000030208 // Madag. Perrier | // Type [red writing] // Museum Paris | Coll. -Regimbart +Regimbart | apicalis // -Copelatus +Copelatus | -apicalis +apicalis n. sp. // - -Remarks. - - -Copelatus apicalis + +Remarks. + + +Copelatus apicalis was described by -Fairmaire (1898) +Fairmaire (1898) from Suberbieville [=Maevatanana]. It is classified in the genus - -Copelatus + +Copelatus and currently placed in the - -Copelatus hydroporoides + +Copelatus hydroporoides species group ( -Guignot 1961 +Guignot 1961 ; -Nilsson et al. 1997 +Nilsson et al. 1997 ; -Nilsson 2001 +Nilsson 2001 ; - + Nilsson and -Hajek +Hajek 2018 ). We have studied a male syntype of this species which was not found by -Guignot (1961 +Guignot (1961 : note 638) and conclude that is not a - -Copelatus + +Copelatus . The syntype male has a stout spine-like setae on a protruding anterodistal corner of protarsomere IV which places it in - -Madaglymbus + +Madaglymbus ( -Shaverdo et al. 2008 +Shaverdo et al. 2008 ) and we hereby transfer it to that genus: - -Madaglymbus apicalis + +Madaglymbus apicalis (Fairmaire, 1898) comb. nov.; in fact - -Gueorguiev + +Gueorguiev (1968 : 32) suggested it might belong to - -Aglymbus + +Aglymbus -"Peut-etre -c'est +"Peut-etre +c'est une - -Aglymbus + +Aglymbus ?" [Eng. translation "Maybe it is an - -Aglymbus + +Aglymbus "], which was correct since this was before - -Madaglymbus + +Madaglymbus was erected for the Madagascar species of - -Aglymbus + +Aglymbus ( -Shaverdo et al. 2008 +Shaverdo et al. 2008 ). diff --git a/data/BD/7D/9C/BD7D9CA1871E598AAC575F54EB7455FD.xml b/data/BD/7D/9C/BD7D9CA1871E598AAC575F54EB7455FD.xml index c3044946f65..2b72fb22dd9 100644 --- a/data/BD/7D/9C/BD7D9CA1871E598AAC575F54EB7455FD.xml +++ b/data/BD/7D/9C/BD7D9CA1871E598AAC575F54EB7455FD.xml @@ -1,214 +1,214 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - - + + + + Copelatus mahajanga Pederzani & -Hajek +Hajek , 2005 -Figs 5B +Figs 5B , -8B +8B - - -Copelatus mahajanga + + +Copelatus mahajanga Pederzani & -Hajek +Hajek , 2005: 104. - -Type locality. -Madagascar, Mahajanga Distr., Mahajanga env. [ca. 15°43'S, 46°18'E] + +Type locality. +Madagascar, Mahajanga Distr., Mahajanga env. [ca. 15°43'S, 46°18'E] - -Type information from OD. -Based on male (holotype), 3 male and 4 female (paratypes). I. Janis 1-10 December 1996. HT & 5PT in NMPC. Two paratypes in collection of Pederzani, Ravenna, Italy. + +Type information from OD. +Based on male (holotype), 3 male and 4 female (paratypes). I. Janis 1-10 December 1996. HT & 5PT in NMPC. Two paratypes in collection of Pederzani, Ravenna, Italy. - -Type material studied. - -Mahajanga. Boeny: Mahajanga I/Mahajanga II + +Type material studied. + +Mahajanga. Boeny: Mahajanga I/Mahajanga II [district cannot be verified based on original description or labels]: -HT♂(GP) (NMPC): // MADAGASCAR 1996, Mahajanga Distr., Mahajanga env., Dec. 1-10., I. -Janis +Janis leg. // HOLOTYPE, COPELATUS, -mahajanga +mahajanga sp. nov., F. Pederzani & J. -Hajek +Hajek det.2005 // - -Additional material studied. - -Mahajanga. Boeny: Ambato-Boeny, Mitsinjo, Soalala + +Additional material studied. + +Mahajanga. Boeny: Ambato-Boeny, Mitsinjo, Soalala : -3♂ (GP), 1♀ (NMW): // Data in NHRS | JLKB | 000010719-22 // RM: Betsiboka Bas (PO124) | Loc. Ambohimanatrika | Kamoro Riv. // 47°10'06", 16°28'55" | alt. 40 m 01.04.1993 // Leg. ORSTOM ANTANANARIVO // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010717 // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -2♀ (NHRS): // NHRS-JLKB | 000065747 (JB802), 65732 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.13337 | E045.95778, 19 m.a.o. 04.XII.2009 | Water net, Field# MAD09-25 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -2♀ (NHRS): // NHRS-JLKB | 000010724 (JB195), 65748 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.01334 | E046.00376, 24 m.a.o. 06.XII.2009 | Water net, Field# MAD09-33 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -2♂ (GP), 3♀ (CAS): // CASENT-8131811-5 // Madagascar: Mahajanga | Province, Parc National de | Namoroka 16.9 km 317° NW | Vilanandro 12-16 Nov 2002 // -16°24'24"S +16°24'24"S , -045°18'36"E +045°18'36"E // Colls. Fisher, Griswold et al.| California Acad. of Sciences | sifted litter, (leaf mold, rotten | wood) in tropical dry forest | elev 100 m code: BLF6582 // -1♀ (CAS): // CASENT-8135013 // Madagascar: Mahajanga | Prov. Parc National de | Namoroka 16.9 Km 317° NW | Vilanandro elev 100 m | 12-16 November 2002 // -16°24'24"S +16°24'24"S , -045°18'36"E +045°18'36"E | Coll. Fisher, Griswold et al. | California Acad. of Sciences | malaise trap- | tropical dry forest | collection code BLF6581 // -1♂ (GP) (CAS): // CASENT-8135001 // Madagascar: Mahajanga | Province Parc National de | Namoroka | 17.8 Km 329° WNW Vilanandro | elev 100 m: 08-12 Nov 2002 // -16°22'36"S +16°22'36"S , -045°19'36"E +045°19'36"E | Colls. Fisher, Griswold et al. | California Acad. of Sciences | pitfall trap- in tropical dry forest, | collection code BLF6506 // -Mahajanga. Melaky: Antsalova +Mahajanga. Melaky: Antsalova : -1♂ (GP) (NHRS): // NHRS-JLKB | 000010723 (JB190) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg. J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -Toamasina. Alaotra Mangoro: Moramanga +Toamasina. Alaotra Mangoro: Moramanga : -5♂ (GP), 3♀, 11 ex. (Alc) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010554-5, 10596-601, 10795 (Alc.) // MAD: TOAM: Alaotra Mangoro: | Betsabora river by RN2 near | Antasmpanana village, 6Km W | of Moramanga: MAD14-81: river | with side pools: 18.9247S | 48.1828E: 900 m: 24.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -3♂ (GP) 26 ex. (Alc.), 7 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010602-3, 10825, 10796(Alc.), 10826(Alc.) // Madagascar: Toamasina: Alaotra | Mangoro: RN2, Betsabora river | 10Km W of Moramanga: S18.92438 | E048.18273, 940 m. 06.XI.2011: | GB Nets and sieves: river and | pools: Field# MAD11-21 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // - -Diagnosis. - + +Diagnosis. + Close to - -C. marginipennis + +C. marginipennis in habitus appearance but body shape more elongate. On average - -C. mahajanga + +C. mahajanga have a broader transverse testaceous band at base of elytra ( -Fig. 8B +Fig. 8B ), and the first elytral stria is more abbreviated anteriorly, starting at approx. one quarter to one half posterior of base. Penis shape is unique in - -C. mahajanga + +C. mahajanga , bisinuate in ventral view with an expanded apex, not resembling that of any other - -Copelatus + +Copelatus species in the - -irinus + +irinus group from Madagascar. - -Description. - + +Description. + Body length 5.2-6.2 mm. Body shape oblong oval, rather convex, brown to dark brown. Head ferrugineous brown, paler in front, sometimes darker around eyes, finely microreticulate and punctate, two shallow depressions between eyes. Pronotum dark brown, paler at sides. Dorsal surface with fine microsculpture and scattered punctures, lateral sides of pronotum striolate with the widest striolate area in the posterior corners. Elytra brown, paler at sides and at apex, with a broad testaceous transverse band at base ( -Fig. 8B +Fig. 8B ). Elytral surface finely reticulate and punctate. Six clearly impressed elytral striae present and one fairly long submarginal stria; first stria abbreviated anteriorly, starting at approx. one quarter to one half posterior of elytral base; other striae (second to sixth) starting more or less at base; submarginal stria starting around middle; striae 1-5 approaching apex of the elytron, but the second and fourth are a little shorter; the sixth stria much abbreviated posteriorly and sometimes the sixth unites with fifth apically approx. 1/5 from apex. - + Ventral side ferrugineous brown, metacoxa and abdominal ventrites striolate and punctate. Prosternal process similar to - -C. marginipennis + +C. marginipennis but lateral parts of metaventrite slightly narrower and metacoxal lines less divergent anteriorly. Antennae, palps, pro- and mesothoracic legs brown, but metathoracic legs ferrugineous dark brown. - + Male: protibia modified, widened in front, strongly angled after base, with several long spines on the outer side of distal half. Tarsomeres -I-III +I-III of pro- and mesolegs enlarged, with pads of numerous setae. Penis unique: in ventral view with a bisinuate shape and a widening asymmetrical apex; in lateral view abruptly curved near middle, with a right angle of more or less 90 degrees, the basal part robust ( -Fig. 5B +Fig. 5B ). Parameres broad ( -Fig. 5B +Fig. 5B ). -Female: similar but smaller than the male, legs not modified. +Female: similar but smaller than the male, legs not modified. - -Distribution. - + +Distribution. + Endemic to Madagascar. Known from several places in Mahajanga province and near Moramanga ( -Fig. 11B +Fig. 11B ). We have seen material from around Mahajanga (Type series), Mahavavy Kinkony Reserve, Namoroka NP, Tsingy de Bemaraha NP, Ankarafantsika NP, Kamoro River south of Ankarafantsika NP and collected the species both in 2011 and 2014 at Betsabora River 6 km W of Moramanga which expanded the known distribution significantly. Note that the name Betsabora is uncertain and recovered from locals on site for this river which is a tributary to Mangoro River. Ampanihifana may be the correct name. - -Habitat and ecology. - + +Habitat and ecology. + Type series probably collected by light trap ( - + Pederzani and -Hajek +Hajek 2005 ), and we collected it with black light by a leaf-choked forest pool in Ankarafantsika NP. Also collected with pitfall traps and from leaf-litter in Namoroka NP before the rainy season in November, indicating the species can tolerate periods when the habitats dry up without immediately seeking new water. We collected the largest series at Betsabora River at an elevation of 940 m. This small river runs through a savannah type area with mixed woodland-grassland ecosystem subsequent to degradation and deforestation. The river site consisted of grassy, vegetation-rich margins, bays and side pools, further described in -Michat et al. (2017) +Michat et al. (2017) . Around the Mahajanga area all sites are lowaltitude, 100 m or lower. - -Comments. - - -Copelatus mahajanga + +Comments. + + +Copelatus mahajanga was the most recently described - -Copelatus + +Copelatus species from Madagascar (Peredzani and -Hajek +Hajek 2005) and was previously only known from the type series and lowland west type locality "Mahajanga env." without further details. The discovery of the species in Betsabora River at mid-altitude in the east indicates that the species may have a much wider distribution and ecological niche. As the species is very similar to - -C. marginipennis + +C. marginipennis habitually, it might be misidentified as such in collections if male genitalia were not examined. diff --git a/data/C9/FE/8E/C9FE8EA486D557DFA5B1A38A9F232743.xml b/data/C9/FE/8E/C9FE8EA486D557DFA5B1A38A9F232743.xml index 0a0050a2890..48bcd3a5e38 100644 --- a/data/C9/FE/8E/C9FE8EA486D557DFA5B1A38A9F232743.xml +++ b/data/C9/FE/8E/C9FE8EA486D557DFA5B1A38A9F232743.xml @@ -1,387 +1,387 @@ - - - -Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species + + + +Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species - - -Author + + +Author -Ranarilalatiana, Tolotra +Ranarilalatiana, Tolotra - - -Author + + +Author -Raveloson Ravaomanarivo, Lala Harivelo +Raveloson Ravaomanarivo, Lala Harivelo - - -Author + + +Author -Bergsten, Johannes +Bergsten, Johannes -text - - -ZooKeys +text + + +ZooKeys - -2019 - -869 + +2019 + +869 - -19 -90 + +19 +90 - -http://dx.doi.org/10.3897/zookeys.869.33997 + +http://dx.doi.org/10.3897/zookeys.869.33997 -journal article -http://dx.doi.org/10.3897/zookeys.869.33997 -1313-2970-869-19 -B7C88A64C06E4B67A352F2F9C8FB0D1C -C44C602AB0235FB2922DC2A4915C3288 +journal article +http://dx.doi.org/10.3897/zookeys.869.33997 +1313-2970-869-19 +B7C88A64C06E4B67A352F2F9C8FB0D1C +C44C602AB0235FB2922DC2A4915C3288 - - - -Copelatus marginipennis (Laporte, 1835) -Figs 5A + + + +Copelatus marginipennis (Laporte, 1835) +Figs 5A , -8A +8A - - -Colymbetes marginipennis + + +Colymbetes marginipennis Laporte, 1835: 102. - -Copelatus aldabricus + +Copelatus aldabricus J. Balfour-Browne, 1950: 368 syn. nov.; TL: Seychelles, Aldabra Islands. - -Copelatus aldabricus var. simplex -Guignot 1952 + +Copelatus aldabricus var. simplex +Guignot 1952 : 28 syn. nov.; TL: Madagascar. - -Type locality. -Senegal [possibly mislabelled]. + +Type locality. +Senegal [possibly mislabelled]. - -Type information from original description. - + +Type information from original description. + housed in Buquet collection and originating from Senegal; of - -aldabricus + +aldabricus : based on male (holotype), J.C. Fryer collection, collected 1908-9 from Aldabra; of - -simplex + +simplex : based on male and female syntypes from Madagascar without further locality data. - -Type material studied. - + +Type material studied. + -LT♂ (lectotype here designated) ( -Colymbetes marginipennis +Colymbetes marginipennis ) (MNHN): // Data in NHRS | JLKB | 000065416 // -Copelatus marginipennis +Copelatus marginipennis Buquet | pulchellus var. -Aube +Aube | h. in Senegal D. Buquet // D. Sharp | Monogr. // Ex-Musaeo -Dejean +Dejean // -pulchellus +pulchellus // Lectotype | -Colymbetes marginipennis +Colymbetes marginipennis | Laporte, 1835 | Des. Ranarilalatiana | & Bergsten, 2019// -HT♂ ( -Copelatus aldabricus +Copelatus aldabricus ) (NHMUK): // Aldabra, 08-9. J.C.F. Fryer // Perey Sladen Trust | expedition. | 1913-170. // Type [red round label] // -Copelatus +Copelatus | -aldabricus +aldabricus Type | J. Balfour-Browne det. // - -Additional material studied. - -Toamasina. Alaotra Mangoro: Andilamena, Moramanga + +Additional material studied. + +Toamasina. Alaotra Mangoro: Andilamena, Moramanga : -1♀ (NMW): // Data in NHRS | JLKB | 000010726 // Madagascar 17.01.1995 | Ambatombe | nr. Andilamena 900 m | leg. Dunay & -Janak +Janak (10) // -1♀ (NHMUK): // BMNH-797894 // MAD: AMPA: Moramanga: Andasibe | Andasibe NP: Big Pond | P61BI15: N: -18.937: E:48.416: 940 m | 06/I/2007: Leg. Isambert et al. // DNA Voucher | BMNH <797894> | MSL294:B10 // -2♂ (GP), 15 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010587-8, 10799(Alc.) // Madagascar: Tamatave: Alaotra Mangoro: | Analamazaotra RS; Bas fond, non-permanent | pond near trail to "lac rouge"MAD15-1| 943 m, -18°56'26.7"S +18°56'26.7"S , -048°25'03.9"E +048°25'03.9"E , 16.III.2015 | Among vegetation and dead leaves in the pond, | Leg. T. Ranarilalatiana & H.J. Randriamihaja // -2♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010604-5, 10800(Alc.) // Madagascar: Tamatave: Alaotra Mangoro: Mantadia NP. | Non-permanent pond at PK18,50 m; E of Park road | 973 m, -18°46'09.9"S +18°46'09.9"S , -048°26'10.4"E +048°26'10.4"E , 17.III.2015 | Under dead leaves & vegetation at the edge of the pond, | Leg.T. Ranarilalatiana, H.J. Randriamihaja; MAD15-5 // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010815 // Madagascar: Toamasina: Alaotra | Mangoro: RN2, Mangoro river | 10Km W of Moramanga: S18.92438 | E048.18273, 940 m. 06.XI.2011 | GB Nets and sieves: river and | pools: Field# MAD11-21 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | J.H. Randriamihaja // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010822 // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra SR: | close to park entrance: S18.9355 E | 048.41656, 970 m: 08.XI.2011: GB | Nets and sieves: dried up forest | pond: Field# MAD11-25 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -6 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010823 // Madagascar: Toamasina: Alaotra | Mangoro: by RN2 S border of | Analamazaotra reserve 1Km E | Antsampanana: S -18.94987 E +18.94987 E 048.42331 | 980m: 09.XI.2011: GB Nets & sieves | ditch next to road: Field# MAD11-29 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | J.H. Randriamihaja // -Toamasina. Analanjirofo: Maroantsetra +Toamasina. Analanjirofo: Maroantsetra : -3♂ (GP), 2♀ (NHMUK): // BMNH-797906-10 // MAD: TOAM: Maroantsetra: Masoala | Masoala NP: Pool | P58BI14: N: -15.758: E: 49.993: 10 m| 17/XI/2006: Leg. Isambert et al. // BMNH (DNA Voucher) // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010794 // MAD: TOAM: Maroantsetra: Masoala | Masoala NP: Pool | P58BI14: N: -15.758: E: 49.993: 10 m| 17/XI/2006: Leg. Isambert et al. // -1♂ (NMW): // Data in NHRS | JLKB | 000010728 // E-Madagascar: Fenerive | Foret de Tampolo | 28.12.1998 | leg. J. Moravec // -1♀(Alc.) (NHRS): // NHRS-JLKB | 000011122 // Madagascar: Toamasina: Analajinrofo: | Masoala NP: degraded lowalt. forest: | MAD18-47: small waterpools on path | ~0.5km NW of Marofototra village, | 15.7606S, 49.9926E, 15 m, 17.II.2018 | Leg. T. Ranarilalatiana // -Toliara. Menabe: Morondava +Toliara. Menabe: Morondava : -3♂ (GP) 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010580, 10730 (JB197), 10751, 10802(Alc.) //Madagascar: Toliara: Menabe: | Kirindy RS: S20.07430 | E044.66307, 52 m.a.o. 12.XII.2009 | Water Net, Field# MAD09-46 | Leg: J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -Mahajanga. Boeny: Ambato-Boeny, Mitsinjo +Mahajanga. Boeny: Ambato-Boeny, Mitsinjo : -1♂ (NHRS): // NHRS-JLKB | 000010734 (JB201) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30350 | E046.81068, 87 m.a.o. 29.XI.2009 | Water Net, Field# MAD09-03 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -6♂ (GP), 3♀, 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010567-71, 10729 (JB196), 10732 (JB199), 10742-3, 10798(Alc.) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -1♂ (NHRS): // NHRS-JLKB | 000010731 (JB198) // Madagascar: Mahajanga: Boeny | Ankarafantsika NP, S16.30270 | E046.80996; 75 m.a.o. 30.XI.2009 | 22W Black light, Field# MAD09-13 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010737 (JB191) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.31418 | E046.81731, 30.XI.2009 | Hand picking, Field# MAD09-14 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -3♀, 1♂(GP), 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000065749 (JB809), 10738 (JB192), 10744-5, 10806(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.14653 | E045.94926, 9 m.a.o. 04.XII.2009 | Water net, Field# MAD09-24 | Leg. J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -2♂ (GP), 2♀, 18 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010578-9, 10739 (JB193), 10746, 10807(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.13337 | E045.95778, 19 m.a.o. 04.XII.2009 | Water net, Field# MAD09-25 | Leg. J. Bergsten, N. -Joensson +Joensson | T. Ranarilalatiana, H.J. Randriamihaja // -5♂ (GP), 4♀, 79 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010572-7, 10740 (JB194), 10747-8, 10808 (Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.05776 | E045.80585, 22 m.a.o. 05.XII.2009 | Water net, Field# MAD09-28 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -1♂, 1♀, 4 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010735 (JB202), 10749, 10809(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.06651 | E045.77672, 24 m.a.o. 05.XII.2009 | Water net, Field# MAD09-29 | Leg. J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010741(JB189) // Madagascar: Mahajanga: Boeny | Mahavavy Kinkony RS, 16.05648S | 045.76371E; 55 m.a.o. 05.XII.2009 | Water net, Field# MAD09-30 | Leg: J. Bergsten, N. Jonsson, T. | Ranarilalatiana, H.J. Randriamihaja // -1♂, 6ex. (Alc.) (NHRS): // NHRS-JLKB | 000010750, 10810(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.01334 | E046.00376, 24 m.a.o. 06.XII.2009 | Water net, Field# MAD09-33 | Leg. J. Bergsten, N. -Joensson +Joensson , | T. Ranarilalatiana, H.J. Randriamihaja // -Mahajanga. Melaky: Antsalova +Mahajanga. Melaky: Antsalova : -2♂ (GP) (CAS): // CASENT-8135015-6 // Madagascar: Mahajanga | Prov. Parc National Tsingy | de Bemaraha, 2.5 Km 62° ENE | Bekopaka, Ankidrodroa river | elev 100 m: 11-15 Nov 2001 // -19°7'56"S +19°7'56"S , -44°48'53"E +44°48'53"E | Coll: Fisher, Griswold et al. | California Acad. of Sciences | sifted litter - tropical dry forest | on Tsingy, code: BLF4340 // -1♂ (GP) (CAS): // CASENT-8131891 // Madagascar: Mahajanga | Prov. Parc National Tsingy | de Bemaraha, 2.5 Km 62° ENE | Bekopaka, Ankidrodroa river | elev 100 m: 11-15 Nov 2001 // -19°7'56"S +19°7'56"S , -44°48'53"E +44°48'53"E | Coll: Fisher, Griswold et al. | California Acad. of Sciences | at light- tropical dry forest | on Tsingy, code: BLF4343 // -1♂ (GP) (CAS): // CASENT-8135006 // Madagascar Mahajanga | Prov. Foret de Tsimembo | 11.0Km 346° NNW Soatana | elev 50 m: 21-25 Nov 2001 | -18°59'43"S +18°59'43"S , -44°26'37"E +44°26'37"E // Coll: Fisher, Griswold et al. | California Acad. of Sciences | sifted litter (leaf mold, rotten wood) | in tropical dry forest | coll. Code: BLF4508 // -1♂ (GP), 2♀, 3 ex. (Alc.), 2♂(Alc.) (NHRS): // NHRS-JLKB | 000010581-2, 10752, 10803(Alc.), 10821(Alc.)// Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg: J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -3♂ GP, 1♀, 5 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010583-4, 10736 (JB203), 10754, 10805(Alc.) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S18.75643 | E044.71398, 119 m.a.o. 17.XII.2009 | Water Net, Field# MAD09-65 | Leg: J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -1♀, 1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010733 (JB200), 10753, 10804(Alc.) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: 19.03419S | 044.77499E, 41 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-59 | Leg: J. Bergsten, N. -Joensson +Joensson , T. | Ranarilalatiana, H.J. Randriamihaja // -Antsiranana. Diana: Ambanja, Ambilobe, Antsiranana II +Antsiranana. Diana: Ambanja, Ambilobe, Antsiranana II : -1♂ (GP) (NHRS): // NHRS-JLKB | 000010585 // Madagascar: Antsiranana: Diana: | Ambilomagodra: Stream under the | bridge of the road RN6 in | Ambilomagodra village, S13.00780 | E49.13313, 139 m, 30.XI.2012, GB nets: | dried up stream with some pools in | the village: Field# MAD12-33 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010586, 10801(Alc.) // Madagascar: Antsiranana: Diana: | Andrafiabe: Antsoha stream | 200m from Andrafiabe, 12.93022S | 49.03466E, 32 m, 01.XII.2012, GB | nets: stream with some pools: | Field# MAD12-39 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♂ (Alc.) (NHRS): // NHRS-JLKB | 000011123 // Madagascar: Antsiranana: Diana: | Antsaba: 1km W of Antsaba, | 13.63474S; 48.72918E, 67 m, | 28.XI.2012, GB nets: forest | stream: Field# MAD12-32 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♀ (NMW): // Data in NHRS | JLKB | 000010725 // MADAGASCAR: Sakaramy | (M. -d'Ambre +d'Ambre , Antsiranana) | lake Farihy Makery | 29.03.2011 | leg. R. Gerecke (MD189) // riparian area near outflow | -12°26'20.5"S +12°26'20.5"S , -49°14'20.0"E +49°14'20.0"E | 377 m. 29.5 °C // -1♂ (NMW): // Data in NHRS | JLKB | 000010727 // N. Madagascar | Antseranana distr. | Sambirano riv. | Marovato vill. | 5-12.12.01, leg. J. -Horak +Horak // Antsira -nana. Sava: Sambava, Vohemar +nana. Sava: Sambava, Vohemar : -1♀ (NHMUK): // BMNH-797876 // MAD: DIEG: Sambava: Marojejy | Marojejy NP: Pool | P57BI31: N: -14.457: E:49.79: 162m | 10/XI/2006; Leg. Isambert et al // DNA Voucher | BMNH <797876> | MSL294:A4 // -1♀ (CAS): // CASENT-8135007 // Madagascar Antsiranana | Foret -d'Ampondrabe +d'Ampondrabe . | 26.3Km 10° NNE Daraina | elev 175m: 10 December 2003 // -12°58'12"S +12°58'12"S , -049°42'00"E +049°42'00"E | California Acad. of Sciences | Coll: B.L.Fisher, sifted | litter (leaf mold, rotten wood) | tropical dry forest, BLF9974 // - -Diagnosis. - - -Copelatus marginipennis + +Diagnosis. + + +Copelatus marginipennis is distinguished from all other Malagasy - -Copelatus + +Copelatus except - -C. pulchellus + +C. pulchellus and - -C. mahajanga + +C. mahajanga by the presence of six discal elytral striae and a broadly oval body shape. - -Copelatus marginipennis + +Copelatus marginipennis is most easily separated by the distinct shape of the male penis in lateral view ( -Fig. 5A +Fig. 5A ). - -Copelatus pulchellus + +Copelatus pulchellus (from Madagascar) lacks a transverse testaceous band basally on the elytra and females of - -C. marginipennis + +C. marginipennis can usually be distinguished from - -C. mahajanga + +C. mahajanga by the normally narrower testaceous band, a longer first elytral stria, and a broader body shape. - -Description. - + +Description. + Body length 5.2-6.6 mm. Body shape oval, rather convex and attenuate posteriorly, dark brown to brown ferrugineous. Head, pronotum and elytra in the same dark brown ferrugineous, covered with fine dense punctation. Lateral sides of pronotum more brownish, with short strioles and the widest striolate area in the posterior corners. Elytra dark brown to brown ferrugineous, with a testaceous transverse band at base ( -Fig. 8A +Fig. 8A ). Six clearly impressed elytral striae present and one submarginal: first stria abbreviated and slightly shorter, starting at approx. 1/7th posterior of elytral base, second to sixth full length, and submarginal stria starting at approx. the middle; all striae approaching the apex of elytron except the sixth, which is abbreviated posteriorly and a little shorter. Sometimes, the first and second elytral striae, like fifth and sixth, unite posteriorly. Elytral surface covered with dense punctures. -Ventral side brownish to ferrugineous, metacoxa with microsculpture, densely and finely punctate. Metacoxa and abdominal ventrites striolate. Prosternal process rather short and spear-shaped, medially only weakly raised and rounded. Lateral parts of metaventrites rather broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae and palps both in the same brown colour. Pro- and mesothoracic legs brown ferrugineous. Metathoracic legs dark brown ferrugineous. - +Ventral side brownish to ferrugineous, metacoxa with microsculpture, densely and finely punctate. Metacoxa and abdominal ventrites striolate. Prosternal process rather short and spear-shaped, medially only weakly raised and rounded. Lateral parts of metaventrites rather broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae and palps both in the same brown colour. Pro- and mesothoracic legs brown ferrugineous. Metathoracic legs dark brown ferrugineous. + Male: Protibia strongly angled basally and expanded in apical two thirds. Penis in ventral view with a small preapical tooth on right side; in lateral view very characteristic with a subbasal dorsal knob, and post-middle with a deep ventral invagination ( -Fig. 5A +Fig. 5A ). -Females from Madagascar usually with elytral striolation rather weak and restricted to outer intervals, but rarely the elytra are entirely and distincly striolate. Females on average smaller than males. +Females from Madagascar usually with elytral striolation rather weak and restricted to outer intervals, but rarely the elytra are entirely and distincly striolate. Females on average smaller than males. - -Distribution. - + +Distribution. + Endemic to the western Indian Ocean islands as far as is modernly known, but the 1835 type locality of the original description and labels read -"Senegal" +"Senegal" , which indicates either mislabeling or that the species in fact also occurs on continental Africa. Known from Madagascar, Reunion, Comores, Seychelles, and Aldabra Island ( -Gomy 2016 +Gomy 2016 ). Widespread in Madagascar and recorded from Montagne -d'Ambre +d'Ambre NP and Masoala NP in the north and northeast to Kirindy in the southwest ( -Fig. 11A +Fig. 11A ), but apparently lacking from most of the central plateau. It can probably show up anywhere around the island below 1000 m in altitude. - -Habitat and ecology. - + +Habitat and ecology. + This is a lowland species that seems to be most common in the deciduous western parts of Madagascar. It was common in the newly designated protected areas of Mahavavy Kinkony when we visited it in 2009. In the deciduous forest biome it has been recorded from Kirindy in the south to Ankarana NP in the north. The species seems to be a generalist and can as well show up on lowland humid east coast and at midaltitudes up to at least 1000 m. It is an apt flier collected with light traps and often in very temporary and small shallow pools including water-filled wheel tracks. Found in all kinds of temporary pools, as well as in streams and in residual pools in dried-up riverbeds. It occurred sympatrically with - -C. mahajanga + +C. mahajanga in Betsabora river near Moramanga and at a 22W black light trap by a forest pool in Ankarafantsika NP. - -Comments. - - -Copelatus aldabricus + +Comments. + + +Copelatus aldabricus was described by J. -Balfour-Browne (1950) +Balfour-Browne (1950) from Aldabra Island in the Seychelles. -Guignot (1952) +Guignot (1952) named a non-striolated female variety from Madagascar as var. -simplex +simplex , and again in 1961 as var. -aequabilis +aequabilis (non-available infrasubspecific name) from Andranofotsy (E of Maroantsetra NE Madagascar), and stated (1961) that females of - -C. aldabricus + +C. aldabricus are normally striolated, but that on Madagascar the non-striolated var. -aequabilis +aequabilis is most common or even ubiquitous. We have found at least one female on Madagascar (from Masoala NP) with the elytra entirely striolated (the -"normal" +"normal" variety sensu -Guignot 1961 +Guignot 1961 ), but indeed the form where striolation is rather weak and only at outer elytral intervals is most common. - -Guignot's + +Guignot's (1961) distinction between female forms either having or lacking strioles is, however, a simplification. All females we have seen have at least some weak tendencies of striolation on outer intervals and the extent can vary significantly between individuals. - - -Copelatus marginipennis + + +Copelatus marginipennis (Laporte, 1835) has been treated as a junior synonym of - -Copelatus pulchellus + +Copelatus pulchellus Klug, 1834 since - -Aube + +Aube (1838) . We discovered the type material of -Laporte's - -C. marginipennis +Laporte's + +C. marginipennis in MNHN, Paris. This could be identified based on a folded blue label bearing the information " - -Copelatus marginipennis + +Copelatus marginipennis Buquet, - -pulchellus + +pulchellus var -Aube +Aube , h. in Senegal, D. Buquet", which fits perfectly the description by -Laporte (1835) +Laporte (1835) . The genitalia of this male show without any doubt that it is a synonym of the younger name - -C. aldabricus + +C. aldabricus and not of - -C. pulchellus + +C. pulchellus . The name - -marginipennis + +marginipennis has to our knowledge not been used as a valid name after 1899, fulfilling the first of the two criteria for protecting a younger currently used synonym from an older available name (ICZN 23.9.1.1). However, - -Copelatus aldabricus + +Copelatus aldabricus is a relatively recent name from a region with a low-intensity taxonomic research and we do not believe it fulfils the second criteria of at least 25 papers published by at least ten authors in the immediately preceding 50 years and encompassing a span of not less than ten years (ICZN 23.9.1.2). Therefore it is not justifiable to protect the younger synonym - -C. aldabricus + +C. aldabricus and the older - -C. marginipennis + +C. marginipennis is therefore brought back as the valid name of this species. We designate the discovered syntype of - -Colymbetes marginipennis + +Colymbetes marginipennis Laporte, 1835 in the MNHN collection as lectotype to preserve stability of the name. At the same time as this type specimen was discovered, also a likely -"type" +"type" specimen of the nomen nudum - -Copelatus fimbriolatus + +Copelatus fimbriolatus Dejean, 1837 from "Ile de France" [= Mauritius] was found with the same type of blue folded label (see supplementary file 1: NHRS-JLKB000065420). This nomen nudum is currently considered a synonym of - -C. pulchellus + +C. pulchellus Klug (Nilsson and Hajek 2018). Unfortunately, it is a female specimen, which is why its identity cannot be established with certainty, but it is more likely identical with - -C. marginipennis + +C. marginipennis since - -C. pulchellus + +C. pulchellus is not yet known from any of the Mascarene Islands ( -Gomy 2016 +Gomy 2016 ).