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<mods:title id="A3399BA1B2B270883F5726F5919FA2AA">Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species</mods:title>
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<mods:namePart>Raveloson Ravaomanarivo, Lala Harivelo</mods:namePart>
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<mods:namePart>Bergsten, Johannes</mods:namePart>
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<treatment ID-GBIF-Taxon="158995562" LSID="urn:lsid:plazi:treatment:0149666EE3E15D2FA4E180FD888742E5" httpUri="http://treatment.plazi.org/id/0149666EE3E15D2FA4E180FD888742E5" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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||||
<taxonomicName LSID="0149666e-e3e1-5d2f-a4e1-80fd888742e5" authority="Klug, 1834" authorityName="Klug" authorityYear="1834" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">Copelatus pulchellus (Klug, 1834)</taxonomicName>
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||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5C</figureCitation>
|
||||
<treatment id="0149666EE3E15D2FA4E180FD888742E5" ID-GBIF-Taxon="158995562" LSID="urn:lsid:plazi:treatment:0149666EE3E15D2FA4E180FD888742E5" httpUri="http://treatment.plazi.org/id/0149666EE3E15D2FA4E180FD888742E5" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
|
||||
<subSubSection id="EAFFA82271D53807FE8D3DCE30ED4C2A" pageId="0" pageNumber="19" type="nomenclature">
|
||||
<paragraph id="EC13FBB874482003FAEE4A56BE739F37" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="8555DD4DAC052C31A5D5048FCA2FB3AF" ID-CoL="5ZWX4" LSID="0149666e-e3e1-5d2f-a4e1-80fd888742e5" authority="Klug, 1834" authorityName="Klug" authorityYear="1834" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">Copelatus pulchellus (Klug, 1834)</taxonomicName>
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<figureCitation id="8233557871AA6C258F8D203034ED75D0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5C</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8D</figureCitation>
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||||
<figureCitation id="D73F64A49C0B76C572CB0F60C7C5C7DB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8D</figureCitation>
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">Agabus pulchellus</taxonomicName>
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<bibRefCitation author="Klug, JCF" journalOrPublisher="Officina Academica, Berolini" pageId="0" pageNumber="19" refId="B42" refString="Klug, JCF, 1834. Symbolae physicae, seu icones et descriptiones Insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. -Insecta. Decas quarta. . Officina Academica, Berolini" title="Symbolae physicae, seu icones et descriptiones Insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. - Insecta. Decas quarta." year="1834">Klug 1834</bibRefCitation>
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||||
<subSubSection id="DF5EBFADD812A01BF151F653F44F7750" pageId="0" pageNumber="19" type="reference_group">
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||||
<paragraph id="04D37AC5354C2E1CF834BEC30373C0C0" pageId="0" pageNumber="19">
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||||
<taxonomicName id="B100AD5F47EE41AFB25AAF8A8BF04C2E" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">Agabus pulchellus</taxonomicName>
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||||
<bibRefCitation id="DF774F9FF7A97B4A1F5530613CCA0D78" author="Klug, JCF" journalOrPublisher="Officina Academica, Berolini" pageId="0" pageNumber="19" refId="B42" refString="Klug, JCF, 1834. Symbolae physicae, seu icones et descriptiones Insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. -Insecta. Decas quarta. . Officina Academica, Berolini" title="Symbolae physicae, seu icones et descriptiones Insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. - Insecta. Decas quarta." year="1834">Klug 1834</bibRefCitation>
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: t. XXXIII: 7
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus africanus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="africanus">Copelatus africanus</taxonomicName>
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<paragraph id="38EF7015D47354174AEE39478E02D935" pageId="0" pageNumber="19">
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<taxonomicName id="31EB1BBD1A9C369B34BF76E86CD477E2" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus africanus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="africanus">Copelatus africanus</taxonomicName>
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Sharp, 1882: 583; TL: Botswana, Lake Ngami;
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<paragraph id="1B6AB6CA67D38E4C9A5FB1A92CA94B5A" pageId="0" pageNumber="19">
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?
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">Copelatus basalis</taxonomicName>
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<taxonomicName id="BCA113D401D54206CAF82D447943F002" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">Copelatus basalis</taxonomicName>
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Boheman, 1848: 244; TL: South Africa (Caffraria interiore);
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus discoideus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="discoideus">Copelatus discoideus</taxonomicName>
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<paragraph id="9F3CD1A8A5CC5F99DAEED0F7D4DAAAFB" pageId="0" pageNumber="19">
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<taxonomicName id="06A4D4CE3638AF1B7B28A2C58355A63F" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus discoideus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="discoideus">Copelatus discoideus</taxonomicName>
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Sharp, 1882: 582; TL: Mesopotamia;
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus obtusus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusus">Copelatus obtusus</taxonomicName>
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<paragraph id="4AD06C15E381E3E942427FBB5532CA9B" pageId="0" pageNumber="19">
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<taxonomicName id="1CE04A09FF7BD6B7E9AB306F458D20B1" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus obtusus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusus">Copelatus obtusus</taxonomicName>
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Boheman, 1848: 242; TL: South Africa (Caffraria orientali);
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus strigulosus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="strigulosus">Copelatus strigulosus</taxonomicName>
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<paragraph id="1E73F0EFC5B94C0543AC82FE043F7721" pageId="0" pageNumber="19">
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<taxonomicName id="BEB4F16C26871AA2B31530D8D5EC45E4" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus strigulosus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="strigulosus">Copelatus strigulosus</taxonomicName>
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Sharp, 1882: 582; TL: Mesopotamia;
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mimetes" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mimetes">Copelatus mimetes</taxonomicName>
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<paragraph id="0E8F2ED01F1E85D41264688E26B8E81F" pageId="0" pageNumber="19">
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<taxonomicName id="E82E0FA8E3BA1DD7FE737C671C305D1B" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mimetes" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mimetes">Copelatus mimetes</taxonomicName>
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Guignot, 1957: 73; TL: Madagascar, Sakaraha, Lambomakandro; syn. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type locality">
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<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="19">Egypt, Sinai.</paragraph>
|
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<subSubSection id="BDF23C4D0E80249C63CAB33B95C17009" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="9C4B247B0D273CBC4F89259688124E90" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="E731DABDC2C4A0336A01A92F64C62807" pageId="0" pageNumber="19">Egypt, Sinai.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="45628AF81A4ACCAD23EF1C34BE81C177" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="78079C315D12894071A74715A52B6C7C" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="EF3A9EEA84D5456799157531078823B2" pageId="0" pageNumber="19">
|
||||
-HT♂ (GP) (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus mimetes</emphasis>
|
||||
<emphasis id="BFC8C85F3F276F1DC584D2E5F2DADEC5" italics="true" pageId="0" pageNumber="19">Copelatus mimetes</emphasis>
|
||||
) (MNHN, "coll. Guignot"): // Data in NHRS | JLKB | 000030032 // Sakaraha | Lambomakandro | III-56 A.R. // Type [red label] // F. Guignot Det., 1956 |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="D338FBA80B7CD12A5A28B20539854F2B" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">mimetes</emphasis>
|
||||
<emphasis id="6B1BA1C61CAC4A833C6D15B249B27679" italics="true" pageId="0" pageNumber="19">mimetes</emphasis>
|
||||
n. sp.| Type ♂ // INSTITUT SCIENTIFIQUE MADAGASCAR // -LT♀ (lectotype here designated) (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||||
<emphasis id="509AD187E6FD6FDA8F0C26973D02C766" italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||||
) (NHRS): // Caffra | ria. // J. Wahlb // Type. // HoloTypus [red label,
|
||||
<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
|
||||
<normalizedToken id="132EFF666FD7317B5796FCBEDEDD1AB0" originalValue="“Typus”">"Typus"</normalizedToken>
|
||||
printed,
|
||||
<normalizedToken originalValue="“Holo”">"Holo"</normalizedToken>
|
||||
<normalizedToken id="5355DEF0153E807E8FDCD892FD129A6D" originalValue="“Holo”">"Holo"</normalizedToken>
|
||||
handwritten with small letters in front, possibly by J. Balfour-Browne who studied the type] // obtusus Boh. //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Cop. pulchellus</emphasis>
|
||||
<emphasis id="494B885783AEE84E96DB708C6A37C26A" italics="true" pageId="0" pageNumber="19">Cop. pulchellus</emphasis>
|
||||
var: |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">obtusus</emphasis>
|
||||
<emphasis id="AA4CF4F7530E963FC26F0A9B01E93F05" italics="true" pageId="0" pageNumber="19">obtusus</emphasis>
|
||||
Boh. | J. Balfour-Browne det. // 5657 | E91 // NHRS-JLKB | 000065335 // Lectotype |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||||
<emphasis id="229B150E860ADD430D750A667F41D2AA" italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||||
| Boheman, 1842 | Des. Ranarilalatiana | & Bergsten, 2019// -? ST♂ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
<emphasis id="15B926AB784F964838DAE3F8F5B00649" italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
) (NHRS): // Caffra | ria. // J Wahlb // Type. // Typus [red label, printed] // 158 | 61 // Copelatus | basalis Bhn. //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="1F91AE9D03E182CD90AB96F384B34B89" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
<emphasis id="9A9D14BA3740504C38C5CA277A92E295" italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
| Klug. | Det. 19.iv.1961 | J. Omer-Cooper. // 5597 | E91 // NHRS-JLKB | 000065337 // -? ST♀ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
<emphasis id="41531706170C7CC03941368BEF2B219C" italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
) (NHRS): // Caffra | ria. // J. Wahlb // Paratypus [red label, printed] // 160 | 61 // 5596 | E91 // NHRS-JLKB | 000065338 // -? ST♀ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
<emphasis id="43D794BD542FAD50C3A60298DFBBCEEE" italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
) (NHRS): // Caffra | ria. // J. Wahlb // Paratypus [red label, printed] // 159 | 61 // 5595 | E91 // NHRS-JLKB | 000065339 //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Antsiranana</emphasis>
|
||||
<subSubSection id="6AEF72431F8A481F69E3EF0C14B27EBB" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="213B96F6842A62424005760F5A80257F" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="7BB245E4F0F752D721B93DBBD6725005" pageId="0" pageNumber="19">
|
||||
<emphasis id="9A63526CDAF47C41948E7F0B808BB21B" bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Antsiranana</emphasis>
|
||||
: -1♂(GP) (MNHN): // Data in NHRS | JLKB | 000030259 // Madagascar | Diego-Suarez | Ch. Alluaud 1893 //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Mitsinjo</emphasis>
|
||||
<emphasis id="FA79F1B5A0F81C88FFF8422A0B64FCE1" bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Mitsinjo</emphasis>
|
||||
: -1♀ (NHRS): // NHRS-JLKB | 000065733 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS: S 16.13337 | E 045.95778, 19 m.a.o. 04.XII.2009 | Water Net, Field# MAD09-25 | leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="E36DCC1C94789BA1CB2218851BBDCDE9" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
<emphasis id="92842A1519F435DC34219127C3112DBE" bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
: -1♂(GP) (NHRS): // NHRS-JLKB | 000010695 // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="690246AD7F6EA2ED52F42DB1E65351A8" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="D205215A76B3E042787102132C05C700" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="DD3DCAF2602995579AE7397A6B02A311" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="9E47B6F97B3D4749B9294643BF055360" pageId="0" pageNumber="19">
|
||||
Similar to
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="0AB9AED32B0DA4DAC0E3907306FDDC4E" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="84690330BB1B53E8BC62F7F9963C3BE6" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(Laporte, 1835) and
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="6C9D5DB5CEF707D612B24CBB44E08297" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="78AA32481AB4BC7E7DAD7D000CD8CB5E" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
in overall habitus, but body shape more like the elongated shape of
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="B91F2B9FBFC6800E1BE75E0F6FE139A9" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="C71736872AA37F460CBAC4E88CD6F2A4" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="668E08AC9574FC3363375B16E51F3EB8" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="C7D0D283A9E393B7205E62D2BF83038C" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
on Madagascar is ferrugineous black in overall colouration with or without a rather narrow or vague testaceous band basally on elytra (
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8D</figureCitation>
|
||||
<figureCitation id="0E05B249AF4BD06E0E3E134484EF2C7A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8D</figureCitation>
|
||||
). Instead testaceous regions on elytra are limited to posterolateral and apical parts. The first elytral stria almost full length. Penis in lateral view rather abruptly curved past middle so that basal and apical parts are approximately at right angles (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5C</figureCitation>
|
||||
<figureCitation id="3C5F3F32524B1F620F698C2F1D9873A3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5C</figureCitation>
|
||||
). Penis overall simpler compared to that of
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="A5BA8B2377EF8B55A27E55FCD22EB252" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="0DD1D64F12B495DC5BFA863D48997400" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="786EE714E63A4BB5D8BEB826B47EF12E" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="6FE431555FF3FE64FCDE8749A043D529" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
and apex slightly twisted.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">(based on Malagasy specimens):</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Body length 5.5-6.1 mm. Body shape oblong oval, rather convex and attenuate posteriorly, dark brown to blackish ferrugineous. Head infuscated brown ferrugineous, somewhat lighter posteriorly, covered with dense microreticulation and sparser punctation.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Pronotum dark brown to ferrugineous black with testaceous anterolateral corners. Disc covered with fine microsculpture forming regular cells and regularily spread small punctures of about same size as cells. Punctuation becomes coarser in posterolateral corners with a weak tendency to corrugate.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="72A999E43B2CD7C8F9B610630C5F44C2" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="4902F8E2B5575E0BB0B42BC549B799DF" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="ED35090C136D6A888C9B7C93D2BEF6F4" pageId="0" pageNumber="19">(based on Malagasy specimens):</paragraph>
|
||||
<paragraph id="9C53EAFA5ED5EB87907F35EA1B2A99E4" pageId="0" pageNumber="19">Body length 5.5-6.1 mm. Body shape oblong oval, rather convex and attenuate posteriorly, dark brown to blackish ferrugineous. Head infuscated brown ferrugineous, somewhat lighter posteriorly, covered with dense microreticulation and sparser punctation.</paragraph>
|
||||
<paragraph id="6E827889128DF4B6249A97462CB1D2B9" pageId="0" pageNumber="19">Pronotum dark brown to ferrugineous black with testaceous anterolateral corners. Disc covered with fine microsculpture forming regular cells and regularily spread small punctures of about same size as cells. Punctuation becomes coarser in posterolateral corners with a weak tendency to corrugate.</paragraph>
|
||||
<paragraph id="A0914066DF7A4A8CA7E084E0F177E6FE" pageId="0" pageNumber="19">
|
||||
Elytra predominantely dark brown to ferrugineous black on disc and along striae with or without a rather narrow and vague basal testaceous band (
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8D</figureCitation>
|
||||
<figureCitation id="9BB6B8A481A6F31547A11BABAB080C44" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8D</figureCitation>
|
||||
). Outer intervals and apical region testaceous to varying degrees, often a testaceous interval 4 stands out. Elytral surface covered with same type of microsculpture and punctures as on pronotum. Six clearly impressed elytral striae present on disc and one submarginal stria: second to sixth stria starting at base, first stria somewhat abbreviated anteriorly (by approximately 1/8th the length of elytra); submarginal stria starting at approx. middle.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="452535735E73DCC12F5379734432516E" pageId="0" pageNumber="19">
|
||||
Ventral side ferrugineous dark brown, with testaceous spots laterally on abdominal ventrites. Metacoxa and ventrites with strioles. Prosternal process more elongate lanceolate and with blunter apex compared with
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="75708F5077DE06A878CBF26228A811E8" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="9C814098A7B8ED0AFE1BD37CD6BBF3B6" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="057A0D6FE900FD52FD464A584FBF2303" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="E0A8E4723E04960F87817AA4CBFAC156" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
. Lateral parts of metaventrite medium broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae, palps, pro- and mesolegs testaceous, metalegs somewhat darker testaceous.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="D42989C208BABABF550D3CADB0287A6D" pageId="0" pageNumber="19">
|
||||
Female: elytral striolation limited to the medial parts of the outer three elytral intervals in the single female studied from Madagascar. From other parts of the distribution a female form is known that has the entire elytra striolated (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B68">Sharp 1882</bibRefCitation>
|
||||
<bibRefCitation id="F0457CEA72E1573750AA962FB10101F9" pageId="0" pageNumber="19" refId="B68">Sharp 1882</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<bibRefCitation id="4D0DCC0E42F4885A7ED326387CE80494" pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken id="E1DC647082DDBDEC9C4468D8CAC7C3D9" originalValue="Hájek">Hajek</normalizedToken>
|
||||
and Reiter 2014
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="473355C6DCFD59B0BC48AA03E08E7B09" pageId="0" pageNumber="19">
|
||||
Male: protibia bisinuate and angled at base, distally expanded. Penis thin, strongly angled at middle in lateral view, and apex somewhat twisted to the left in ventral view (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5C</figureCitation>
|
||||
<figureCitation id="95A88775FD8449BFCBA7279EB0DF34C1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5C</figureCitation>
|
||||
). Parameres more narrowly triangular apically compared to those in
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="B5A78D3539EE4738423BB610EA9D88EE" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="916BBE80E94783053218DEFAC6BBC5FC" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="621122C6245E66AEF0AE9CAEFFDC5FC0" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="9AC495CFB1FCFDA7A6BA39ACC18A4603" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19" start="Figure 8" startId="F8">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<caption id="EEABAD8C030E5A86DC0013594E93E90D" doi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19" start="Figure 8" startId="F8">
|
||||
<paragraph id="125D6D65BD9D148F2C89749B87693815" pageId="0" pageNumber="19">
|
||||
Figure 8. Habitus, dorsal view.
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
|
||||
<emphasis id="90B8C71933539CFC4137E2B289538574" bold="true" pageId="0" pageNumber="19">A</emphasis>
|
||||
Male:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<taxonomicName id="BA38DBC761118BCA49E6BD7B9DCCB65D" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="773F90122DF9E274DFC077A92CEC9E5D" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||||
<emphasis id="E4913E517408CE66474AC1ED175F4715" bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus mahajanga</emphasis>
|
||||
<taxonomicName id="51BD48BD9095A12639A985FA768EC15A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||||
<emphasis id="CF85A6295D5A6A9A152E3A2AA833A5BE" italics="true" pageId="0" pageNumber="19">Copelatus mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||||
<emphasis id="3FEC063894467F59B1A14863B6152FBF" bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="92671443339910B5F475155C8F151510" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B288BBD5926532894A236CB9147CC90C" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 1 (Bemaraha)
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||||
<emphasis id="36476D93EB7209EBC454B045254A1DF9" bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||||
Male:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="1B34C25108AE2F7D67329E927CD63101" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="4F26913B2FB4A8320AEF17DCFDE50044" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="F6BA842A8AC2587B066503A9E3DA903A" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="2CF8DE15D57E3784C89DC70B8042D885" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="941FFC6A6C01AA3AC55319526A8C5410" pageId="0" pageNumber="19">
|
||||
As the species
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="86FAEB8545201DCD7F33BB13750AC1BF" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="BD454AA834E30704E4A964A9704E840E" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
is currently interpreted, this is a very widely distributed Afrotropical and Middle Eastern species.
|
||||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
<bibRefCitation id="4F5DA26310FB5909A71A9D53EBCAF677" author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
recorded the species from Senegal in the west, Tanzania in the east, South Africa in the south, and Mesopotamia (Iraq and adjacent regions) in the Middle East.
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<bibRefCitation id="966A0E1A8DF7AEDF22A17403F3FF1D75" pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken id="574CAA5AEF9B845C49CE01003C6E20EB" originalValue="Hájek">Hajek</normalizedToken>
|
||||
and Reiter (2014)
|
||||
</bibRefCitation>
|
||||
recorded the species from Yemen and stated that the likely distribution in the Middle East included the entire Arabian Peninsula.
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot (1961)</bibRefCitation>
|
||||
<bibRefCitation id="9E96EF9CB6015A83A12303F7F7C44921" pageId="0" pageNumber="19" refId="B36">Guignot (1961)</bibRefCitation>
|
||||
also gives India and Ceylon (= Sri Lanka) but this was likely based on the misidentification of closely related oriental species (see
|
||||
<bibRefCitation author="Ghosh, SK" journalOrPublisher="Skoervnoepparn Supplement" pageId="0" pageNumber="19" pagination="1 - 77" publicationUrl="http://www.norrent.se/images/stories/upload/SN_Supplement_03_hq.pdf" refId="B20" refString="Ghosh, SK, Nilsson, AN, 2012. Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae). . Skoervnoepparn Supplement 3: 1 - 77" title="Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae)." url="http://www.norrent.se/images/stories/upload/SN_Supplement_03_hq.pdf" volume="3" year="2012">Ghosh and Nilsson 2012</bibRefCitation>
|
||||
<bibRefCitation id="5CBD11001A34EFCE1DEEDE083F46432C" author="Ghosh, SK" journalOrPublisher="Skoervnoepparn Supplement" pageId="0" pageNumber="19" pagination="1 - 77" publicationUrl="http://www.norrent.se/images/stories/upload/SN_Supplement_03_hq.pdf" refId="B20" refString="Ghosh, SK, Nilsson, AN, 2012. Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae). . Skoervnoepparn Supplement 3: 1 - 77" title="Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae)." url="http://www.norrent.se/images/stories/upload/SN_Supplement_03_hq.pdf" volume="3" year="2012">Ghosh and Nilsson 2012</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4459.2.2" author="Sheth, SD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="19" pagination="235 - 260" refId="B70" refString="Sheth, SD, Ghate, HV, Hajek, J, 2018. Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). . Zootaxa 4459: 235 - 260" title="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)." url="https://doi.org/10.11646/zootaxa.4459.2.2" volume="4459" year="2018">Sheth et al. 2018</bibRefCitation>
|
||||
<bibRefCitation id="DED48DEB1D7CB059CDE76076547522B1" DOI="https://doi.org/10.11646/zootaxa.4459.2.2" author="Sheth, SD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="19" pagination="235 - 260" refId="B70" refString="Sheth, SD, Ghate, HV, Hajek, J, 2018. Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). . Zootaxa 4459: 235 - 260" title="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)." url="https://doi.org/10.11646/zootaxa.4459.2.2" volume="4459" year="2018">Sheth et al. 2018</bibRefCitation>
|
||||
).
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="B3B5AF7F1BA328436237C9CA127C76A9" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="5EA0B6E1704136F5FACFEB930D5ECC1A" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
was not previously recorded from Madagascar following the revision by
|
||||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
<bibRefCitation id="E682093D610F2E24A0F58250EE2B36BF" author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
but as we synonymise
|
||||
<taxonomicName lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||||
<taxonomicName id="B575046BFEEF9C127E155520EB6FD65E" lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||||
<emphasis id="F0D9F9D81963D550EA67AA1E9CA41923" italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||||
</taxonomicName>
|
||||
with
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="BA20267D925C2479FE666FF54691EE5D" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="F275D888A275F7590DF58F07847D40E4" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
, Madagascar now forms part of the distribution. From Madagascar we have seen specimens from Antsiranana, Mahavavy Kinkony Reserve, Tsingy de Bemaraha National Park (Bekopaka), and from Lambomakandro, Sakaraha (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11C</figureCitation>
|
||||
<figureCitation id="440B0B9BB34136B3C5F28A1ED0B7AE2E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11C</figureCitation>
|
||||
). It can likely show up anywhere in lowland Madagascar, but especially in the western lowlands.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="90B653CFB010A0D3D10A554E4EC0BBA5" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="EB17E93DD8B8F0EAD8A3058568E5DE61" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="C018F322FF979C56E513ABFD57E9E3A8" pageId="0" pageNumber="19">
|
||||
On Madagascar we have collected the species associated with a small forest stream with sidepools in a karstic limestone area (
|
||||
<normalizedToken originalValue="“tsingy”">"tsingy"</normalizedToken>
|
||||
<normalizedToken id="F93E4F5B8CFCF6B4CB2F192166B63E29" originalValue="“tsingy”">"tsingy"</normalizedToken>
|
||||
) and in a muddy stagnant pool in a dried-out river bed. Both localities are in dry deciduous forests of lowland western Madagascar.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comment.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<subSubSection id="7F09AE95E64A7C306D35EC3CDA9D005E" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="9770A8B01C9A18FD6BC55C3E9D78F26E" pageId="0" pageNumber="19">Comment.</paragraph>
|
||||
<paragraph id="50CDEBD105F2D2CE1527D486BC3666DA" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="3BCA9A99A52A4E358D319ABDCC816844" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="5CA43AB979629183CBEF26942A7D4878" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
forms part of a diverse species group with many externally very similar species.
|
||||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
<bibRefCitation id="29CB8FCB74F0DEFE40069386B5B5A46E" author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
admitted that his previous treatment of the species (
|
||||
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" author="Balfour-Browne, J" journalOrPublisher="The Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="19" pagination="57 - 88" refId="B3" refString="Balfour-Browne, J, 1939. On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with description of new species. . The Transactions of the Royal Entomological Society of London 88: 57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with description of new species." url="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
|
||||
<bibRefCitation id="D5140CC074118421D59E52C0AFD1964F" DOI="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" author="Balfour-Browne, J" journalOrPublisher="The Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="19" pagination="57 - 88" refId="B3" refString="Balfour-Browne, J, 1939. On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with description of new species. . The Transactions of the Royal Entomological Society of London 88: 57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with description of new species." url="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
|
||||
) was entirely wrong as he had then not studied the male genitalia. When he did so in 1950 it resulted in the description of several new species previously lumped under
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="182F289E37E100C6315B0041ADF31785" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="6997884B6DAE8E1D90EE8C515929FCEE" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<paragraph id="8BA176555F5AF574BDFE7977808A0963" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="8315050BB247C2110AE8C7981BDA253C" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="5E15F47220F76CF78CAF234CFBA9BE72" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
is now interpreted as a widespread Afrotropical and Middle Eastern species with the male penis similar to that illustrated in
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figure 5C</figureCitation>
|
||||
<figureCitation id="A13FA1D460E0601B9F41E958A9D2704F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figure 5C</figureCitation>
|
||||
. Dorsal colouration is interpreted as very variable, even consisting of several distinct colour forms such as the darker forms described as
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<taxonomicName id="C2E93117383924E4363ABB9C3F82E7AD" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="B4B8AA39A560D7E6868EF3826634F145" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
Boheman and later as
|
||||
<taxonomicName lsidName="C. africanus" pageId="0" pageNumber="19" rank="species" species="africanus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. africanus</emphasis>
|
||||
<taxonomicName id="AE334FFF107A4419A92015BDC91CF66C" lsidName="C. africanus" pageId="0" pageNumber="19" rank="species" species="africanus">
|
||||
<emphasis id="F6734F60D95937017DA753DDB48C8356" italics="true" pageId="0" pageNumber="19">C. africanus</emphasis>
|
||||
</taxonomicName>
|
||||
Sharp (compare figure 24 with photographs in
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.595.8614" author="Perissinotto, R" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="85 - 135" refId="B60" refString="Perissinotto, R, Bird, MS, Bilton, DT, 2016. Predaceous water beetles (Coleoptera, Hydradephaga) of the Lake St Lucia system, South Africa: biodiversity, community ecology and conservation implications. . ZooKeys 595: 85 - 135" title="Predaceous water beetles (Coleoptera, Hydradephaga) of the Lake St Lucia system, South Africa: biodiversity, community ecology and conservation implications." url="https://doi.org/10.3897/zookeys.595.8614" volume="595" year="2016">Perissinotto et al. (2016</bibRefCitation>
|
||||
<bibRefCitation id="680E25C977CBCD46EC1FBD6CFF3F359F" DOI="https://doi.org/10.3897/zookeys.595.8614" author="Perissinotto, R" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="85 - 135" refId="B60" refString="Perissinotto, R, Bird, MS, Bilton, DT, 2016. Predaceous water beetles (Coleoptera, Hydradephaga) of the Lake St Lucia system, South Africa: biodiversity, community ecology and conservation implications. . ZooKeys 595: 85 - 135" title="Predaceous water beetles (Coleoptera, Hydradephaga) of the Lake St Lucia system, South Africa: biodiversity, community ecology and conservation implications." url="https://doi.org/10.3897/zookeys.595.8614" volume="595" year="2016">Perissinotto et al. (2016</bibRefCitation>
|
||||
: fig. 24) and
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<bibRefCitation id="FAF469273935F30444BBE698BDAE5DFF" pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken id="2E585D2D847A1876027A87805F5E47A8" originalValue="Hájek">Hajek</normalizedToken>
|
||||
and Reiter (2014
|
||||
</bibRefCitation>
|
||||
: fig. 30)). Females are also interpreted as variable in the striolation pattern on the elytra. Sharp described a female form from Mesopotamia that was entirely striolated over the elytra under the name
|
||||
<taxonomicName lsidName="C. strigulosus" pageId="0" pageNumber="19" rank="species" species="strigulosus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. strigulosus</emphasis>
|
||||
<taxonomicName id="3B14BEB0BE46C4195A72DFE6B213B691" lsidName="C. strigulosus" pageId="0" pageNumber="19" rank="species" species="strigulosus">
|
||||
<emphasis id="9D3FBE95085CD5E6352D26F7DDB1602D" italics="true" pageId="0" pageNumber="19">C. strigulosus</emphasis>
|
||||
</taxonomicName>
|
||||
, which is interpreted today as intraspecific variation of
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="06C4D4882A9E401EAF3DC01F8742038A" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="69D39041089BD3D8EB31DA88511DDEB5" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
(see photograph in
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<bibRefCitation id="BF2F6D8A51E2F9433A1B891B4054C47A" pageId="0" pageNumber="19" refId="B37">
|
||||
<normalizedToken id="BE9B82285B3EE481758B2630A211033A" originalValue="Hájek">Hajek</normalizedToken>
|
||||
and Reiter (2014
|
||||
</bibRefCitation>
|
||||
: fig. 30)). We have seen three males and one female from Madagascar. All males are of the darker colour form, lacking a basal transverse testaceous band on elytra, similar to
|
||||
<normalizedToken originalValue="Boheman’s">Boheman's</normalizedToken>
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<normalizedToken id="40AA8FF9651A5EBAC94B24A02F86D001" originalValue="Boheman’s">Boheman's</normalizedToken>
|
||||
<taxonomicName id="1A6772676DF0D8B38E12B35B9F412374" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="896B555D42023E37C5DAF308D0A99D81" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
. The female has a weak basal testaceous band but is otherwise also most similar to the dark colour form. Striolation is limited to the middle of the outer three intervals. We have no doubt that the three non-type specimens examined are conspecific with
|
||||
<taxonomicName lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||||
<taxonomicName id="DC5A39C1B4A4994B7FD2C50182F67DF8" lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||||
<emphasis id="4DE844410DA33C25573A05C8290B6105" italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||||
</taxonomicName>
|
||||
Guignot. But after our comparison of the male genitalia of the type with
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="3D09F94479433CCBC730459B16808EF2" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="859AF1613CF594287A9403AD3DB440A9" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
specimens from mainland Africa, we concluded that
|
||||
<taxonomicName lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||||
<taxonomicName id="8A5D1CF86422EA2BA0EACED85EA4E6D8" lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||||
<emphasis id="22A03E41A3F0C53EC7A80263D4BE5548" italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||||
</taxonomicName>
|
||||
cannot be upheld as a separate species given how
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="EDD2C18C39EAB16F6F12BE1D9AC73AB5" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="321B2202F952FB63B8FFEE3A134F2E41" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
is interpreted. We noted very minor differences in the penis apex which are not greater than differences between east, south, and west African specimens of
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="16AF089CE9D5E3ED6B57D0BCDB26846B" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="76696D9ED4DBD7D04E28966603D961BC" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
. The penis figured by
|
||||
<bibRefCitation author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="0" pageNumber="19" pagination="71 - 74" refId="B33" refString="Guignot, F, 1957. Un haliplide et deux dytiscides nouveaux de Madagascar. . Le Naturaliste Malgache 9: 71 - 74" title="Un haliplide et deux dytiscides nouveaux de Madagascar." volume="9" year="1957">Guignot (1957</bibRefCitation>
|
||||
<bibRefCitation id="416CD02BED91793FEDFDF93A5C5A5CE4" author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="0" pageNumber="19" pagination="71 - 74" refId="B33" refString="Guignot, F, 1957. Un haliplide et deux dytiscides nouveaux de Madagascar. . Le Naturaliste Malgache 9: 71 - 74" title="Un haliplide et deux dytiscides nouveaux de Madagascar." volume="9" year="1957">Guignot (1957</bibRefCitation>
|
||||
: fig. 2, reproduced by
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
|
||||
<bibRefCitation id="2451ACECDBF4AC85993B003ED71E62F3" DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
|
||||
: fig. 8) is very inaccurate, depicting a shape quite different from
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="0572B8B86F1869665222AD682E9CF977" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="F16A5379FC5D9FB994674AB5479AD173" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
but that does not correspond with the holotype.
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="61E0C03FA9C73562D004A70E7BD3608A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="7842C845B6D77C55E70BBC645FB056F0" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
may still be a species complex for future studies to solve but currently it is a widespread and variable species distributed over Africa, Madagascar, and the Middle East.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
<paragraph id="7814C6CB6ADA136D2F98ACAABBC3B5EB" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="4D35B50DBD35366FAE7375384214EED0" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">
|
||||
<emphasis id="067F8871CE778F211AA78FA999A8C140" italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
</taxonomicName>
|
||||
Boheman, 1848 was synonymised with
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="729038F8E4E90DB67DD3CD68D81891AE" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="0B9A4F9FB059C5EF47C095E69B8F8E59" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
by Omer-Cooper (1965). We have studied the same type material as Omer-Cooper and agree with this conclusion. However, we are not convinced that the material housed at NHRS as these types are the correctly identified types of the name
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<taxonomicName id="77455785B4A43EBED61587A90C04B038" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="2FAC6065139087C410F6DBC7983CE8C8" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.9375" author="Boheman, CH" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="19" refId="B11" refString="Boheman, CH, 1848. . https://doi.org/10.5962/bhl.title.9375" url="https://doi.org/10.5962/bhl.title.9375" year="1848">Boheman (1848)</bibRefCitation>
|
||||
<bibRefCitation id="21A105E85B73D98B262DB2E1DFDD5FF9" DOI="https://doi.org/10.5962/bhl.title.9375" author="Boheman, CH" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="19" refId="B11" refString="Boheman, CH, 1848. . https://doi.org/10.5962/bhl.title.9375" url="https://doi.org/10.5962/bhl.title.9375" year="1848">Boheman (1848)</bibRefCitation>
|
||||
describes in Latin four species from Johan August Wahlbergs collectings in Caffraria (South Africa) of which three are new, numbered 259 -
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="54EEDA1FBA0B84BED3400126E55A159F" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="F1744DA5BA4686761905FE5DDC5FD700" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
Klug; 260 -
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus obtusus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||||
<taxonomicName id="7FA0D7293389DFF93285329D264308DA" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus obtusus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusus">
|
||||
<emphasis id="FE80542756B1590B3F75D0C85EFD1B5E" italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
; 261 -
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus striatellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="striatellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus striatellus</emphasis>
|
||||
<taxonomicName id="9A318F905B989E604FF982404B9A5DCB" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus striatellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="striatellus">
|
||||
<emphasis id="1D4CA3EFFC0CEA25BB7F2F51873EEBB2" italics="true" pageId="0" pageNumber="19">Copelatus striatellus</emphasis>
|
||||
</taxonomicName>
|
||||
; and 262 -
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
<taxonomicName id="21CE6FD6DAFF94161CFCFE9AD5F7AC72" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">
|
||||
<emphasis id="4878BA40AC0475309AC667C02C501FA0" italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||||
</taxonomicName>
|
||||
. For
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="EA340825A1D8214B8B464CEA0B90B466" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="50A9DCF5B608C590E8B7B1814F6B4854" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
the disc of elytra is described as having six striae. For the longer description of
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<taxonomicName id="C7EE6095F5D28A7E3F25455E8E797D14" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="76C1DEC2B58CFEEC50C339E1A21440BF" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
this is further elaborated to detail that there are six discal stria but entire elytra has seven striae
|
||||
<normalizedToken originalValue="“septem-striata”">"septem-striata"</normalizedToken>
|
||||
<normalizedToken id="59A10DDC1E6D478F80F52875D8FC57E5" originalValue="“septem-striata”">"septem-striata"</normalizedToken>
|
||||
(six discal and one submarginal). This pattern and number of striae is consistent with the
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="7A9C8D64C07190BC81C2CF5A32DFB600" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="46242379D049A56700A8704F4C5DFDDA" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
species group. The following species is correctly identified as
|
||||
<taxonomicName lsidName="C. striatellus" pageId="0" pageNumber="19" rank="species" species="striatellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. striatellus</emphasis>
|
||||
<taxonomicName id="D72A833F1D4C49E1735DFDB1154D5703" lsidName="C. striatellus" pageId="0" pageNumber="19" rank="species" species="striatellus">
|
||||
<emphasis id="AEF4E442347BE5E4CAA8929DF3239EE2" italics="true" pageId="0" pageNumber="19">C. striatellus</emphasis>
|
||||
</taxonomicName>
|
||||
with nine discal striae, and is clearly stated as such: "disco striis 9 tenuibus", of which the innermost is much abbreviated. This description matches very well with the types preserved at NHRS. Finally,
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<taxonomicName id="32152B7A285038D77D8E6AEAF0EA808D" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="6056B3E2D7231E4187D1793FFE50BD9B" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
is described as the last
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="09ADB8530BFDA5D40586D57EB5BA9BE5" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="62AE42F87246A59DBA37A38C1F1AFE90" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
species in
|
||||
<normalizedToken originalValue="Boheman’s">Boheman's</normalizedToken>
|
||||
<normalizedToken id="B96CC66227B16AF9C32BF99E58D7A59C" originalValue="Boheman’s">Boheman's</normalizedToken>
|
||||
work and elytra is described as
|
||||
<normalizedToken originalValue="“12-striata”">"12-striata"</normalizedToken>
|
||||
<normalizedToken id="4B72E450877A97B299E9FE27EE886684" originalValue="“12-striata”">"12-striata"</normalizedToken>
|
||||
. The supposed types at NHRS for both
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<taxonomicName id="F1DF643DC32DD4956F3683FC1644E152" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="9577E4CB42489FC29C65B187BD73ADED" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<taxonomicName id="978E50C208A74D8361B30C69BA5CD0B8" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="2A94B6DEEED8F53CBD0849AE02B73D4E" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
have six discal and one submarginal striae. The type for
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<taxonomicName id="85602C37C14584C15C5C7A513DFE731F" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="D5D35893AD1DCD25A069AF853BFDE578" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
matches the original description of
|
||||
<normalizedToken originalValue="“septem-striata”">"septem-striata"</normalizedToken>
|
||||
<normalizedToken id="081152F6E6E1420D5B386CEA01CE7AA4" originalValue="“septem-striata”">"septem-striata"</normalizedToken>
|
||||
but the three types of
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<taxonomicName id="2F3AC921A99C88E5DBE66313B60F3AAB" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="A094AED5A6D7953C7406C9CDAD83E898" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
with the same number of striae does not match the original description of
|
||||
<normalizedToken originalValue="“12-striata”">"12-striata"</normalizedToken>
|
||||
<normalizedToken id="7D7C25CCC2E8AF821BBA767DCA753D6D" originalValue="“12-striata”">"12-striata"</normalizedToken>
|
||||
. It would be very inconsistent of Boheman to describe the total number of striae on one elytron in one case
|
||||
<normalizedToken originalValue="“elytra…septem-striata”">"elytra...septem-striata"</normalizedToken>
|
||||
<normalizedToken id="D6785E10AD56FB03A41F04186C08BE6C" originalValue="“elytra…septem-striata”">"elytra...septem-striata"</normalizedToken>
|
||||
, and in the other only the discal striae but summing up the number from both elytral halves
|
||||
<normalizedToken originalValue="“elytra…12-striata”">"elytra...12-striata"</normalizedToken>
|
||||
<normalizedToken id="BB0F4C1A76691E70DA692A3A6E235F36" originalValue="“elytra…12-striata”">"elytra...12-striata"</normalizedToken>
|
||||
. It would also be very illogical to place
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<taxonomicName id="2581AD23FA880B06F1500031E0053585" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="D4243A976C78E5B41B143A5A0C82BAD0" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
after the 9-striated
|
||||
<taxonomicName lsidName="C. striatellus" pageId="0" pageNumber="19" rank="species" species="striatellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. striatellus</emphasis>
|
||||
<taxonomicName id="2F5B5539E768A6F7641EBB8F281D1E1E" lsidName="C. striatellus" pageId="0" pageNumber="19" rank="species" species="striatellus">
|
||||
<emphasis id="0405CD1B6369396940C25A5175738278" italics="true" pageId="0" pageNumber="19">C. striatellus</emphasis>
|
||||
</taxonomicName>
|
||||
if it has the same number of striae as
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="A4EDB9F95C5C6B8A149FFBB1EFA5AB6C" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="58247C297446E628AA747669A247D860" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<taxonomicName id="099D13D4FC8EA2CF9E2A30021334212B" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="1723D656F91D54EDCA0B4B2C60AEC80C" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
, which come first. Similarily,
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot (1961)</bibRefCitation>
|
||||
<bibRefCitation id="7092573FEBB57A1F901BC13648530FF7" pageId="0" pageNumber="19" refId="B36">Guignot (1961)</bibRefCitation>
|
||||
was confused about
|
||||
<normalizedToken originalValue="Boheman’s">Boheman's</normalizedToken>
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<normalizedToken id="10952B8F660C81312EE6CEB330CBF6F2" originalValue="Boheman’s">Boheman's</normalizedToken>
|
||||
<taxonomicName id="71F789391BB21A2A7C95A8B7A287A53A" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="E652DF546069DA55395D1D0AB0EF201E" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
and listed the name both under the 12-striated
|
||||
<taxonomicName lsidName="C. mocquerysi" pageId="0" pageNumber="19" rank="species" species="mocquerysi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mocquerysi</emphasis>
|
||||
<taxonomicName id="3AC147E26BA581743B54AFBDA603FB84" lsidName="C. mocquerysi" pageId="0" pageNumber="19" rank="species" species="mocquerysi">
|
||||
<emphasis id="E3B1215F93DB2B69DE6068B82EC7B64E" italics="true" pageId="0" pageNumber="19">C. mocquerysi</emphasis>
|
||||
</taxonomicName>
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<normalizedToken id="F1DF7B816D929CF285F2231265CA2C84" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
with a question mark, and under
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="28C33FC4557A36D911B1F7439F1DD85A" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="2462EB73D499C355A722669980B7AF4C" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
. We consider the status of
|
||||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
<taxonomicName id="8F8CB9990A25A5BC7F7F889EBFABE28F" lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||||
<emphasis id="DF4C1FEA905AABA371992420AF0A246B" italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||||
</taxonomicName>
|
||||
Boheman as uncertain but we have not found any alternative potential type material at NHRS. We designate the single undoubted syntype of
|
||||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
<taxonomicName id="51BE4E45426E3A0CCD694054CFA06D08" lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||||
<emphasis id="39D57638A31CD25AC207B4E09C954D08" italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||||
</taxonomicName>
|
||||
Boheman, 1848 in the NHRS collection as lectotype to preserve the stability of the name.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,221 +1,221 @@
|
|||
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China</mods:title>
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|
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||||
</mods:role>
|
||||
<mods:namePart>Ge, Si Xun</mods:namePart>
|
||||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3769-1530</mods:nameIdentifier>
|
||||
<mods:affiliation>Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China.</mods:affiliation>
|
||||
<mods:namePart id="8C553A352EDF622C08B664DE956B0B8B">Ge, Si Xun</mods:namePart>
|
||||
<mods:nameIdentifier id="3DF7B2DE8079E83EA5763335FC651615" type="ORCID">https://orcid.org/0000-0003-3769-1530</mods:nameIdentifier>
|
||||
<mods:affiliation id="D979814D6B0F9D91A266920850B9AEC6">Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China.</mods:affiliation>
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||||
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|
||||
<mods:name type="personal">
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||||
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|
||||
<mods:namePart>Jiang, Zhuo Heng</mods:namePart>
|
||||
<mods:affiliation>School of Science, Westlake University, Hangzhou, China</mods:affiliation>
|
||||
<mods:namePart id="0EAC2CB814CED009D7CABD8FB54C70B3">Jiang, Zhuo Heng</mods:namePart>
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||||
<mods:affiliation id="8684ACD2DC9AA947D42036C699BB229B">School of Science, Westlake University, Hangzhou, China</mods:affiliation>
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</mods:name>
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||||
<mods:name type="personal">
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||||
<mods:namePart>Wang, Jia Qi</mods:namePart>
|
||||
<mods:affiliation>2288 Long, Hongxin Rd, Minhang District, Shanghai, China</mods:affiliation>
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<mods:namePart id="92AEABCF872B8B439AF9E74C08E59AAD">Wang, Jia Qi</mods:namePart>
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<mods:affiliation id="E86CC461AC91245074FDA57763976C8E">2288 Long, Hongxin Rd, Minhang District, Shanghai, China</mods:affiliation>
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||||
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<mods:roleTerm id="A86EB3890E99E8910C0BAC2C34C3EE4F">Author</mods:roleTerm>
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</mods:role>
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||||
<mods:namePart>Song, Kui</mods:namePart>
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||||
<mods:affiliation>School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China</mods:affiliation>
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<mods:namePart id="95CF17F934D8C4A315B7B67B1FB4670F">Song, Kui</mods:namePart>
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<mods:affiliation id="963AF4CCA2FE0D11FABA16C1142201B7">School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China</mods:affiliation>
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<mods:namePart>Zhang, Chao</mods:namePart>
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<mods:affiliation>Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China</mods:affiliation>
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<mods:namePart id="B2FE03109A9549978C469BC6C2EA71FE">Zhang, Chao</mods:namePart>
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<mods:affiliation id="383067540F013257897C928AFF4F6516">Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China</mods:affiliation>
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<mods:namePart>Hu, Shao Ji</mods:namePart>
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<mods:affiliation>Institute of International Rivers and Eco-security, Yunnan University, Kunming, China</mods:affiliation>
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<mods:nameIdentifier type="email">shaojihu@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="CA9F50524EE82442A5C8D733293B6277">Hu, Shao Ji</mods:namePart>
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<mods:affiliation id="30EA46E9E7E99BDF3AAFED595EA58498">Institute of International Rivers and Eco-security, Yunnan University, Kunming, China</mods:affiliation>
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<mods:nameIdentifier id="CDB74EC6C09B9AFDD02E6CD6411DEC86" type="email">shaojihu@hotmail.com</mods:nameIdentifier>
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<mods:date>2023</mods:date>
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<mods:number>81</mods:number>
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<mods:number id="EE1AF95BBE715721A14B8194AA48B97F">81</mods:number>
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<subSection pageId="0" pageNumber="257" type="3. results">
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<treatment LSID="urn:lsid:plazi:treatment:08B5ED96E2A559C1B8C12353E550A1A7" httpUri="http://treatment.plazi.org/id/08B5ED96E2A559C1B8C12353E550A1A7" lastPageNumber="257" pageId="0" pageNumber="257">
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<taxonomicName LSID="08B5ED96-E2A5-59C1-B8C1-2353E550A1A7" authority="euorientis Verity, 1908" authorityName="euorientis Verity" authorityYear="1908" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="euorientis" status="stat. rev." subSpecies="euorientis">Pieris euorientis euorientis Verity, 1908</taxonomicName>
|
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<taxonomicNameLabel pageId="0" pageNumber="257">stat. rev.</taxonomicNameLabel>
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<subSection id="94A2FE4B8F7E1CEFC1A0C95BCF3D5275" pageId="0" pageNumber="257" type="3. results">
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<treatment id="08B5ED96E2A559C1B8C12353E550A1A7" ID-GBIF-Taxon="209450344" LSID="urn:lsid:plazi:treatment:08B5ED96E2A559C1B8C12353E550A1A7" httpUri="http://treatment.plazi.org/id/08B5ED96E2A559C1B8C12353E550A1A7" lastPageNumber="257" pageId="0" pageNumber="257" scope_family="Pieridae" scope_order="Lepidoptera">
|
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<subSubSection id="B6221DC7E850A6ECA1976862FBAA0F5E" pageId="0" pageNumber="257" type="nomenclature">
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<taxonomicName id="05B9FDDDC295C402888105FFDFD4A161" LSID="08B5ED96-E2A5-59C1-B8C1-2353E550A1A7" authority="euorientis Verity, 1908" authorityName="euorientis Verity" authorityYear="1908" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="euorientis" status="stat. rev." subSpecies="euorientis">Pieris euorientis euorientis Verity, 1908</taxonomicName>
|
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<taxonomicNameLabel id="248E4FE63ECBB6C50C7B21C59530B1D3" pageId="0" pageNumber="257">stat. rev.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="257" type="reference_group">
|
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<paragraph pageId="0" pageNumber="257">
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<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. var. var. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="variety" species="napi" status="stat. rev." subSpecies="var." variety="euorientis">Pieris napi var. euorientis</taxonomicName>
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<subSubSection id="4663FEACE586B5301582E6ED021DC1FD" pageId="0" pageNumber="257" type="reference_group">
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<paragraph id="A9A51B3A469D1414F3457C126D686B42" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="D2C0CBF49374D20EF83013D464B3B032" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. var. var. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="variety" species="napi" status="stat. rev." subSpecies="var." variety="euorientis">Pieris napi var. euorientis</taxonomicName>
|
||||
Verity, 1908;
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="257" rank="subOrder" status="stat. rev." subOrder="Rhopalocera">Rhopalocera</taxonomicName>
|
||||
<taxonomicName id="455A0342599540B1BD9DC22B6941CC58" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subOrder" status="stat. rev." subOrder="Rhopalocera">Rhopalocera</taxonomicName>
|
||||
Palaearctica (13-14): 147, (17-20): pl. 32, f. 44; TL:
|
||||
<normalizedToken originalValue="“Sajan”">"Sajan"</normalizedToken>
|
||||
<normalizedToken id="21251F4C35670FD607B17FFA1DCEC98B" originalValue="“Sajan”">"Sajan"</normalizedToken>
|
||||
(original description)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="1908" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="euorientis" status="stat. rev.">Pieris euorientis</taxonomicName>
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||||
<paragraph id="E0EDFE1E9049C82C0ADE3D6CE342C87D" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="F3714AA341562292364BCE8074C4B2EB" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="1908" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="euorientis" status="stat. rev.">Pieris euorientis</taxonomicName>
|
||||
Verity, 1908;
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="6F9A2F5C54A825E701D2A80B1ECA1A49" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
: Herbipoliana 1(1-2): 131 (upgraded as distinct species)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris dulcinea subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="dulcinea" status="stat. rev." subSpecies="euorientis">Pieris dulcinea euorientis</taxonomicName>
|
||||
<paragraph id="3C501FBD46747C131490B3C80D892EEE" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="2CA624623608A4971799B93287000600" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris dulcinea subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="dulcinea" status="stat. rev." subSpecies="euorientis">Pieris dulcinea euorientis</taxonomicName>
|
||||
;
|
||||
<bibRefCitation author="Tuzov, VK" journalOrPublisher="Molecular biology and evulotion" pageId="0" pageNumber="257" refId="B27" refString="Tuzov, VK, Bogdanov, PV, Devyatkin, AL, Kaabak, LV, Korolev, VA, Murzin, VS, Samodurov, GD, Tarasov, EA, 1997. Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." title="Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." year="1997">Tuzov et al. (1997)</bibRefCitation>
|
||||
<bibRefCitation id="E2D5688267EB655C3692B3EB833090AE" author="Tuzov, VK" journalOrPublisher="Molecular biology and evulotion" pageId="0" pageNumber="257" refId="B27" refString="Tuzov, VK, Bogdanov, PV, Devyatkin, AL, Kaabak, LV, Korolev, VA, Murzin, VS, Samodurov, GD, Tarasov, EA, 1997. Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." title="Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." year="1997">Tuzov et al. (1997)</bibRefCitation>
|
||||
(as subspecies of
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="dulcinea (Butler" baseAuthorityYear="1882" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea" status="stat. rev.">Pieris dulcinea</taxonomicName>
|
||||
<taxonomicName id="EB19411CB591058AAAF9D685DF67A417" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="dulcinea (Butler" baseAuthorityYear="1882" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea" status="stat. rev.">Pieris dulcinea</taxonomicName>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="napi" status="stat. rev." subSpecies="euorientis">Pieris napi euorientis</taxonomicName>
|
||||
<paragraph id="6D7B69DD061FDF18A9F3AE93EB730983" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="1E07358DCE1723388C034C2A4ABE24A5" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="napi" status="stat. rev." subSpecies="euorientis">Pieris napi euorientis</taxonomicName>
|
||||
;
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="47E5C7F3D309AED863B8AB5407405750" author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
: Butterflies 65: 26 (as subspecies of
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="napi (Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi" status="stat. rev.">Pieris napi</taxonomicName>
|
||||
<taxonomicName id="8E0B06DB4D7A4E0096F5DB78B26A4507" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="napi (Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi" status="stat. rev.">Pieris napi</taxonomicName>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="description">
|
||||
<paragraph pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="389836F410A2ED600F8932047A7EE1B5" pageId="0" pageNumber="257" type="description">
|
||||
<paragraph id="E5D0F59D12FF29394DF7778E242655B9" pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph id="DE0B1CC35FADCBC35CDEF1991CCC295E" pageId="0" pageNumber="257">
|
||||
Both wings white on the upperside and a pale yellowish hue on the underside. Male (Fig.
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of species related to Chinese species of Pieris napi - complex. Composite pictures on the left show the upperside, on the right show the underside. A - B Pieris ochsenheimeri ochsenheimeri Staudinger, 1886. C - E Pieris napi sauron Yakovlev, 2004 comb. nov. F - H Pieris napi bryonides Sheljuzhko, 1910 comb. rev. I - L Pieris napi muchei Eitschberger, 1983. M - N Pieris napi napi (Linnaeus, 1758). O - R Pieris bryoniae wolfsbergeri Eitschberger, 1983. S - V Pieris euorientis euorientis Verity, 1908 stat. rev. Scale bar represents 10 mm. (B, F, I, K, L, O-R, V reference Eitschberger, 1983; T and U reference Tshikolovets et al., 2009; C reference Yakovlev, 2004 as black and white photos consistent with original description)." figureDoi="10.3897/asp.81.e85191.figure11" httpUri="https://binary.pensoft.net/fig/825206" pageId="0" pageNumber="257">11S, U</figureCitation>
|
||||
<figureCitation id="DD1476F5F5A72C9EB8E5C698C5415529" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of species related to Chinese species of Pieris napi - complex. Composite pictures on the left show the upperside, on the right show the underside. A - B Pieris ochsenheimeri ochsenheimeri Staudinger, 1886. C - E Pieris napi sauron Yakovlev, 2004 comb. nov. F - H Pieris napi bryonides Sheljuzhko, 1910 comb. rev. I - L Pieris napi muchei Eitschberger, 1983. M - N Pieris napi napi (Linnaeus, 1758). O - R Pieris bryoniae wolfsbergeri Eitschberger, 1983. S - V Pieris euorientis euorientis Verity, 1908 stat. rev. Scale bar represents 10 mm. (B, F, I, K, L, O-R, V reference Eitschberger, 1983; T and U reference Tshikolovets et al., 2009; C reference Yakovlev, 2004 as black and white photos consistent with original description)." figureDoi="10.3897/asp.81.e85191.figure11" httpUri="https://binary.pensoft.net/fig/825206" pageId="0" pageNumber="257">11S, U</figureCitation>
|
||||
): apical spot grey, merging as a small dark subtriangular spot on the apex or nearly absent. The 1st discal spot and outer spot present but rather faint on both upperside and underside, while the 2nd spot vaguely developed on the underside. Female (Fig.
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of species related to Chinese species of Pieris napi - complex. Composite pictures on the left show the upperside, on the right show the underside. A - B Pieris ochsenheimeri ochsenheimeri Staudinger, 1886. C - E Pieris napi sauron Yakovlev, 2004 comb. nov. F - H Pieris napi bryonides Sheljuzhko, 1910 comb. rev. I - L Pieris napi muchei Eitschberger, 1983. M - N Pieris napi napi (Linnaeus, 1758). O - R Pieris bryoniae wolfsbergeri Eitschberger, 1983. S - V Pieris euorientis euorientis Verity, 1908 stat. rev. Scale bar represents 10 mm. (B, F, I, K, L, O-R, V reference Eitschberger, 1983; T and U reference Tshikolovets et al., 2009; C reference Yakovlev, 2004 as black and white photos consistent with original description)." figureDoi="10.3897/asp.81.e85191.figure11" httpUri="https://binary.pensoft.net/fig/825206" pageId="0" pageNumber="257">11T, V</figureCitation>
|
||||
<figureCitation id="00F13FCA4D4238BC7118923A119F771A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of species related to Chinese species of Pieris napi - complex. Composite pictures on the left show the upperside, on the right show the underside. A - B Pieris ochsenheimeri ochsenheimeri Staudinger, 1886. C - E Pieris napi sauron Yakovlev, 2004 comb. nov. F - H Pieris napi bryonides Sheljuzhko, 1910 comb. rev. I - L Pieris napi muchei Eitschberger, 1983. M - N Pieris napi napi (Linnaeus, 1758). O - R Pieris bryoniae wolfsbergeri Eitschberger, 1983. S - V Pieris euorientis euorientis Verity, 1908 stat. rev. Scale bar represents 10 mm. (B, F, I, K, L, O-R, V reference Eitschberger, 1983; T and U reference Tshikolovets et al., 2009; C reference Yakovlev, 2004 as black and white photos consistent with original description)." figureDoi="10.3897/asp.81.e85191.figure11" httpUri="https://binary.pensoft.net/fig/825206" pageId="0" pageNumber="257">11T, V</figureCitation>
|
||||
): all spots distinctly developed on the upperside, while relatively faint on the underside. Brownish or blackish powder distinctly narrowed along all veins on both sides, relatively broader at the apex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Russia, Mongolia (Altai and Sayan Mts, Siberia and Transbaikalia)</paragraph>
|
||||
<subSubSection id="83DC5AB6353B5142DE2D01A95E2D92C4" pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph id="5615B10440B7EEDB05D7F42155A565B2" pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph id="A5C77AE4395F354430EEFFD515AF44EF" pageId="0" pageNumber="257">Russia, Mongolia (Altai and Sayan Mts, Siberia and Transbaikalia)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="3E5B11A17626A6F58A7D7977BFB880C0" pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph id="ABE8CB73CC66751C94F0E2D58D761BE2" pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph id="3AB34B7C0E517C67E205F34F51B43BE3" pageId="0" pageNumber="257">
|
||||
Univoltine from June to July (
|
||||
<bibRefCitation author="Tuzov, VK" journalOrPublisher="Molecular biology and evulotion" pageId="0" pageNumber="257" refId="B27" refString="Tuzov, VK, Bogdanov, PV, Devyatkin, AL, Kaabak, LV, Korolev, VA, Murzin, VS, Samodurov, GD, Tarasov, EA, 1997. Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." title="Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." year="1997">Tuzov et al., 1997</bibRefCitation>
|
||||
<bibRefCitation id="3DD4A2DAAEEEDB3EC13143360743185C" author="Tuzov, VK" journalOrPublisher="Molecular biology and evulotion" pageId="0" pageNumber="257" refId="B27" refString="Tuzov, VK, Bogdanov, PV, Devyatkin, AL, Kaabak, LV, Korolev, VA, Murzin, VS, Samodurov, GD, Tarasov, EA, 1997. Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." title="Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera). Pensoft, Sofia." year="1997">Tuzov et al., 1997</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="838399FA3126326887920140B1275731" pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph id="80A47B7A7C870D50ABB4FFA8585DB416" pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph id="8883C44B0FD2A3532EAFD4AB64D089F2" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5J</figureCitation>
|
||||
<figureCitation id="58EC2EF54E711FD3B374C072E85B54FE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5J</figureCitation>
|
||||
) basal part of uncus moderately narrower than the distal margin of tegumen, as a concave between uncus and tegumen; uncus slightly convex at its median part, apical half of uncus digitation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Not examined.</paragraph>
|
||||
<subSubSection id="5E06AA0ED369E2FB3441889FB3F18C99" pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph id="549FEC65C96FB9617D9680093843443A" pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph id="5CC7835BC523DD0775307F63447FDB08" pageId="0" pageNumber="257">Not examined.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="21F56A7783ABAE5FBACBA9CFAF00A1E6" pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph id="7916752F3D2678495FC676A5D516C811" pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph id="B8321669EFB13E3F49E59FEEA8240E99" pageId="0" pageNumber="257">
|
||||
This species rather resembles the spring form of
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
<taxonomicName id="977540EF397D5B416CA0B962020703B7" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea">
|
||||
<emphasis id="168D5B3E24F3881F54BD434CAFD1BB44" italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
which can only be distinguished by a comparison of the apical spot (more or less merging in this species while always separated into several triangles in
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
<taxonomicName id="64FA0C592A6403D106BD7F55D72589C1" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea">
|
||||
<emphasis id="54D97D137FF9C417F930E8E60C3D5C92" italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
) and the 1st discal spot (both species faint but in
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="euorientis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. euorientis</emphasis>
|
||||
<taxonomicName id="B9861C40F5F04F510254F925260FEC0A" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="euorientis">
|
||||
<emphasis id="51CF056889250E01BED3EA5F77770C7C" italics="true" pageId="0" pageNumber="257">P. euorientis</emphasis>
|
||||
</taxonomicName>
|
||||
relatively more developed). Males can be easily distinguished from the summer form of
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
<taxonomicName id="9AF675DC7B3ED0DCF40DF1E930909A64" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea">
|
||||
<emphasis id="1ADEA21A02E01E36832A5590CE4BFAF9" italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
in having: apical spot merged (separate in
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
<taxonomicName id="366E4D3DF8750A5C390DAB8DDFA4BC7F" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea">
|
||||
<emphasis id="25E9D1389144870722BBD7062878D241" italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
), and 1st discal spot faint (strongly developed in
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
<taxonomicName id="ADC3B7646F3CD079F103511AB591A2FB" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea">
|
||||
<emphasis id="C37038E631579471A3DC6DFF498B9FA1" italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
) and 2nd discal spot absent (present in
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
<taxonomicName id="E1E51569CE368F8EC52CB1F6D5042977" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dulcinea">
|
||||
<emphasis id="EDBD983ED544ECCC8E77496C86B58D49" italics="true" pageId="0" pageNumber="257">P. dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
) on the upperside. This taxon shows monophyly in our phylogenetic analysis and its position is distant from
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
<taxonomicName id="BA49B89E5F7CF197FC65C6259D2889AA" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="F50BD132A5967281F0DB4DF6A1AAED21" italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
</taxonomicName>
|
||||
; therefore, we consider this taxon to be a distinct species. It is noteworthy that in the re-description of the species by
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="88B18B6E676FE6B79180C4FE1878149A" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
a series of specimens from Tanu-Ola (Tanguura Mountains) were considered as belonging to this species, while in this study, we found morphological differences between topotypes of
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="euorientis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. euorientis</emphasis>
|
||||
<taxonomicName id="47FE1E66CF2D04EC0283634BC950242C" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="euorientis">
|
||||
<emphasis id="69513FAFFC1865275C2E7B8D5AC11F6B" italics="true" pageId="0" pageNumber="257">P. euorientis</emphasis>
|
||||
</taxonomicName>
|
||||
and specimens from regions south or west of the Tanguura Mountains (See notes in
|
||||
<taxonomicName authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
<taxonomicName id="392A09DDB820418DC0C342057B5E8122" authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis id="2DB92CC5847C055956B94C378E479DCE" italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
</taxonomicName>
|
||||
Yakovlev, 2006 stat. nov.). Combined with our phylogenetic results, this indicates that populations from the Tanguura Mountains and the regions to the south or west represent a distinct species (Fig.
|
||||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution map of Pieris euorientis Verity, 1908 stat. rev. and Pieris mihon Yakovlev, 2006 stat. nov." figureDoi="10.3897/asp.81.e85191.figure12" httpUri="https://binary.pensoft.net/fig/825207" pageId="0" pageNumber="257">12</figureCitation>
|
||||
<figureCitation id="110134A51DFE62BE05F11D8E46FF352D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution map of Pieris euorientis Verity, 1908 stat. rev. and Pieris mihon Yakovlev, 2006 stat. nov." figureDoi="10.3897/asp.81.e85191.figure12" httpUri="https://binary.pensoft.net/fig/825207" pageId="0" pageNumber="257">12</figureCitation>
|
||||
):
|
||||
<taxonomicName authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
<taxonomicName id="419F53BD608E3BB64D9AE8C3D69633E5" authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis id="05F5E1D84ACD08D6E37BE54A9FB6F6C2" italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
</taxonomicName>
|
||||
Yakovlev, 2006 stat. nov. As the redefined
|
||||
<taxonomicName authority="euorientis" authorityName="euorientis Verity" authorityYear="1908" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="euorientis" subSpecies="euorientis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris euorientis euorientis</emphasis>
|
||||
<taxonomicName id="36E6A56A1DE7E6EEAFF0F8A394031AEF" authority="euorientis" authorityName="euorientis Verity" authorityYear="1908" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis subsp. euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="euorientis" subSpecies="euorientis">
|
||||
<emphasis id="087600B5FAA97E3D1E84241F04A9A663" italics="true" pageId="0" pageNumber="257">Pieris euorientis euorientis</emphasis>
|
||||
</taxonomicName>
|
||||
, we did not find female specimens of the summer form in the type locality or adjacent regions, thus this knowledge gap needs to be filled in future research.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/asp.81.e85191.figure12" httpUri="https://binary.pensoft.net/fig/825207" pageId="0" pageNumber="257" start="Figure 12" startId="F12">
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">Figure 12.</emphasis>
|
||||
<caption id="6B796C74C7FADC3944552532D015D5A6" doi="10.3897/asp.81.e85191.figure12" httpUri="https://binary.pensoft.net/fig/825207" pageId="0" pageNumber="257" start="Figure 12" startId="F12">
|
||||
<paragraph id="6DE2E770A5AB9137904BD9F1D26667F5" pageId="0" pageNumber="257">
|
||||
<emphasis id="FF1CADEA66FD89D72A8A233F4C1E4787" bold="true" pageId="0" pageNumber="257">Figure 12.</emphasis>
|
||||
Distribution map of
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="1908" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="euorientis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris euorientis</emphasis>
|
||||
<taxonomicName id="FCEDE8DC3D9E982CE3413116306E9812" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="1908" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris euorientis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="euorientis">
|
||||
<emphasis id="BBB3093F65F5CDB187A8FB3BAEC4D1D3" italics="true" pageId="0" pageNumber="257">Pieris euorientis</emphasis>
|
||||
</taxonomicName>
|
||||
Verity, 1908 stat. rev. and
|
||||
<taxonomicName authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
<taxonomicName id="A0E208D6F2EEDF87FFAD29E4E16D7F5B" authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis id="0C5FA28BF8AF64B8F115D6BD34C1161A" italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
</taxonomicName>
|
||||
Yakovlev, 2006 stat. nov.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,124 +1,124 @@
|
|||
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<treatment ID-GBIF-Taxon="158995560" LSID="urn:lsid:plazi:treatment:0A694D306A4456EB8ADB140766C419B7" httpUri="http://treatment.plazi.org/id/0A694D306A4456EB8ADB140766C419B7" lastPageNumber="19" pageId="0" pageNumber="19">
|
||||
<subSubSection pageId="0" pageNumber="19" type="nomenclature">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName LSID="0a694d30-6a44-56eb-8adb-140766c419b7" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina subjecta" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjecta">Exocelina subjecta (Sharp, 1882)</taxonomicName>
|
||||
<treatment id="0A694D306A4456EB8ADB140766C419B7" ID-GBIF-Taxon="158995560" LSID="urn:lsid:plazi:treatment:0A694D306A4456EB8ADB140766C419B7" httpUri="http://treatment.plazi.org/id/0A694D306A4456EB8ADB140766C419B7" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
|
||||
<subSubSection id="EF9D44072485F76BB37593BD77CA07AE" pageId="0" pageNumber="19" type="nomenclature">
|
||||
<paragraph id="1C56B4263A4D15F4623296F6FDEEB61C" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="D99E2BB561D0FCE1B3EB9B078A39D865" ID-CoL="3DLRJ" LSID="0a694d30-6a44-56eb-8adb-140766c419b7" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina subjecta" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjecta">Exocelina subjecta (Sharp, 1882)</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus subjectus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjectus">Copelatus subjectus</taxonomicName>
|
||||
<subSubSection id="2A03FFF01D313CD6556A969D37157C8F" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="08BBCF1F6DD37FA2FB7160994009B4F2" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="26322227833C6C93C5DD50C8BB65BEBD" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus subjectus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjectus">Copelatus subjectus</taxonomicName>
|
||||
Sharp, 1882: 568.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bilunatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="bilunatus">Copelatus bilunatus</taxonomicName>
|
||||
<paragraph id="93CFEE09E7FABAB5234CA73A92FF7155" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="D0406A95DEBEC38C90F466C6BFFBA85C" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bilunatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="bilunatus">Copelatus bilunatus</taxonomicName>
|
||||
Guignot, 1955: 73; TL: Madagascar [mislabelled, likely New Caledonia; see
|
||||
<bibRefCitation author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="53247DEC689792DE08CF203FCCE8B808" author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
]
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">New Caledonia.</paragraph>
|
||||
<subSubSection id="53C40ACD1D2B2BF19467D6A373B99C7B" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="978599A38139F44A1E6318759CA2BB7C" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="DF844EE58DD09E1920E6F8D02E990274" pageId="0" pageNumber="19">New Caledonia.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bilunatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="bilunatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus bilunatus</emphasis>
|
||||
<subSubSection id="94DB2803CA96E01172C05DB03A12D5C1" pageId="0" pageNumber="19" type="remarks">
|
||||
<paragraph id="DD31105310D2D3F9877F0132DDA78EB7" pageId="0" pageNumber="19">Remarks.</paragraph>
|
||||
<paragraph id="348C46DB63FA505719477C31ECCBB53E" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="F8341E4543C7AC64E4CC7F58E1FB96DC" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bilunatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="bilunatus">
|
||||
<emphasis id="6570160848C84E43356B7B4EE17D261D" italics="true" pageId="0" pageNumber="19">Copelatus bilunatus</emphasis>
|
||||
</taxonomicName>
|
||||
Guignot, 1955 is a synonym of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina subjecta" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjecta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Exocelina subjecta</emphasis>
|
||||
<taxonomicName id="27D1FFA3831E1C4961C95F022D8DF79D" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina subjecta" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjecta">
|
||||
<emphasis id="60CF6EB09DBD000E99B80B59ED2CE791" italics="true" pageId="0" pageNumber="19">Exocelina subjecta</emphasis>
|
||||
</taxonomicName>
|
||||
(Sharp, 1882) following
|
||||
<bibRefCitation author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="ED8401D290DADE9DF5FAC5C7AF3D4D2A" author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
.
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bilunatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="bilunatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus bilunatus</emphasis>
|
||||
<taxonomicName id="18369241A2869643050BFD173949453B" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bilunatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="bilunatus">
|
||||
<emphasis id="D3F3FC470EF396D06DF4FCBECD5C146D" italics="true" pageId="0" pageNumber="19">Copelatus bilunatus</emphasis>
|
||||
</taxonomicName>
|
||||
was described by
|
||||
<bibRefCitation author="Guignot, F" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="19" pagination="67 - 73" refId="B29" refString="Guignot, F, 1955a. Dytiscides inedits de la collection Zimmermann (Col. Dytiscidae). . Annales de la Societe Entomologique de France 123: 67 - 73" title="Dytiscides inedits de la collection Zimmermann (Col. Dytiscidae)." volume="123" year="1955 a">Guignot (1955a)</bibRefCitation>
|
||||
<bibRefCitation id="ECBB39D6E3442DE851342B1DDFEB5A1A" author="Guignot, F" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="19" pagination="67 - 73" refId="B29" refString="Guignot, F, 1955a. Dytiscides inedits de la collection Zimmermann (Col. Dytiscidae). . Annales de la Societe Entomologique de France 123: 67 - 73" title="Dytiscides inedits de la collection Zimmermann (Col. Dytiscidae)." volume="123" year="1955 a">Guignot (1955a)</bibRefCitation>
|
||||
from
|
||||
<normalizedToken originalValue="Zimmermann’s">Zimmermann's</normalizedToken>
|
||||
<normalizedToken id="1211D947DE0C4FD32EB924C53E13606B" originalValue="Zimmermann’s">Zimmermann's</normalizedToken>
|
||||
collection based on a "♀ unique" from
|
||||
<normalizedToken originalValue="“Madagascar”">"Madagascar"</normalizedToken>
|
||||
<normalizedToken id="2A8601708364DA18302D11D3A5C52D7B" originalValue="“Madagascar”">"Madagascar"</normalizedToken>
|
||||
without any further locality data. The species was assigned to
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="88252B7501E68C16A232653966ABE90F" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="D1436FBB10FDD25E46B79AD556A029E1" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
based on the presence of complete metacoxal lines and placed in the
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus hydroporoides" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="hydroporoides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus hydroporoides</emphasis>
|
||||
<taxonomicName id="BCCA5BF49809C6E7E981F6B2574B3952" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus hydroporoides" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="hydroporoides">
|
||||
<emphasis id="1C6C77B14FF1469F99577B7057E8F619" italics="true" pageId="0" pageNumber="19">Copelatus hydroporoides</emphasis>
|
||||
</taxonomicName>
|
||||
species group, as it lacked impressed elytral striae.
|
||||
<bibRefCitation author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="81A7F32F16FF8160DC546A0FA26D86C7" author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
concluded that the specimen is identical to
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina subjecta" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjecta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Exocelina subjecta</emphasis>
|
||||
<taxonomicName id="590324C6122464F95F3A80AC968D798B" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina subjecta" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="subjecta">
|
||||
<emphasis id="FB1E3AEFEA7072EBA3448A3028796D2B" italics="true" pageId="0" pageNumber="19">Exocelina subjecta</emphasis>
|
||||
</taxonomicName>
|
||||
(Sharp, 1882), a common species on New Caledonia, and synoymised the two assuming the specimen was mislabelled. Apparently the holotype is a male in contrast to what is stated in the original description.
|
||||
<bibRefCitation author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="03C495F86CF05D259ED37B209B95B642" author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. (2014)</bibRefCitation>
|
||||
document the presence of a stout spine-like setae on the anterodistal angle of protarsomere IV, which is characteristic of the
|
||||
<taxonomicName lsidName="" pageId="0" pageNumber="19" rank="subfamily" subfamily="Copelatinae">Copelatinae</taxonomicName>
|
||||
<taxonomicName id="F629184DE5F0F03D1ECD6615A99D2420" lsidName="" pageId="0" pageNumber="19" rank="subfamily" subfamily="Copelatinae">Copelatinae</taxonomicName>
|
||||
genus
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Exocelina</emphasis>
|
||||
<taxonomicName id="EFB08FE5E4798C3FCE39D7E79C87BCE4" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="CAA627E46C8B390320FE740E6D14159D" italics="true" pageId="0" pageNumber="19">Exocelina</emphasis>
|
||||
</taxonomicName>
|
||||
(but also of Malagasy
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
<taxonomicName id="3DCF57F04A08C9D4CBA22D6583C8373C" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9C88565C234CD2B76E28EB960EF1F74B" italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
). We have not examined the type ourselves but it is clear that in any case it is not a
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="B1489026882ADDA5FE9B9761F3A91F7A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1A3C9FC5C9B6815783518FC930C0ECB9" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
species.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,165 +1,165 @@
|
|||
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||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China</mods:title>
|
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<mods:title id="308CE6B148643D599BF988CF0684BCB1">A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China</mods:title>
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<mods:name type="personal">
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<mods:role>
|
||||
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<mods:name id="A998EF6577263CF8ED503B8824A65FF1" type="personal">
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<mods:role id="528060E5B4D847E6ACC47E6C4000836C">
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||||
</mods:role>
|
||||
<mods:namePart>Ge, Si Xun</mods:namePart>
|
||||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3769-1530</mods:nameIdentifier>
|
||||
<mods:affiliation>Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China.</mods:affiliation>
|
||||
<mods:namePart id="4BBE8325D2A5DB4DEC2E698A98213436">Ge, Si Xun</mods:namePart>
|
||||
<mods:nameIdentifier id="5B2CA27C6FFACB0899C1C6114770AF25" type="ORCID">https://orcid.org/0000-0003-3769-1530</mods:nameIdentifier>
|
||||
<mods:affiliation id="76175D0B48C192996D33B286E5EC631B">Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China.</mods:affiliation>
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||||
</mods:name>
|
||||
<mods:name type="personal">
|
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<mods:role>
|
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:role id="12D35B436F4C9C63348531189DC0ED5D">
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<mods:roleTerm id="79013E8F77113D5B2017EFAC88B3A0EB">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart>Jiang, Zhuo Heng</mods:namePart>
|
||||
<mods:affiliation>School of Science, Westlake University, Hangzhou, China</mods:affiliation>
|
||||
<mods:namePart id="FF1C82EF329DD0A7FD5A541BB31A205E">Jiang, Zhuo Heng</mods:namePart>
|
||||
<mods:affiliation id="E50093F4595051E5418ACCB8FDAB15F8">School of Science, Westlake University, Hangzhou, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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<mods:name id="88F084061B091F3414100C99B64E9F63" type="personal">
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<mods:role id="89B2B982E264E4547F5D31058873B9D0">
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<mods:roleTerm id="B1AE6BC10E58C7CD63A9D51C1338893F">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Wang, Jia Qi</mods:namePart>
|
||||
<mods:affiliation>2288 Long, Hongxin Rd, Minhang District, Shanghai, China</mods:affiliation>
|
||||
<mods:namePart id="172920B14AE4C713E17C5D873FD2F62B">Wang, Jia Qi</mods:namePart>
|
||||
<mods:affiliation id="6B3800029FF50CE313126E4395D79EE4">2288 Long, Hongxin Rd, Minhang District, Shanghai, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
<mods:name id="660EBD0864B1736BF4EEECD91655A44D" type="personal">
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<mods:role id="9CA60CC804CC51F64F4B299899E96BD5">
|
||||
<mods:roleTerm id="D10519F2EC942C154B33CA9949370ACA">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Song, Kui</mods:namePart>
|
||||
<mods:affiliation>School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China</mods:affiliation>
|
||||
<mods:namePart id="FB8C2C9EFFDFD69F1F58DBB08A6EB17B">Song, Kui</mods:namePart>
|
||||
<mods:affiliation id="E116C815B7062CF0EA0FF687A9C916AE">School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhang, Chao</mods:namePart>
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<mods:affiliation>Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China</mods:affiliation>
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<mods:namePart id="F4F5F154CF4816C7B689C9A3373351EF">Zhang, Chao</mods:namePart>
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<mods:affiliation id="81448C2837B6E0DDDD16A1B2A2C4908E">Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China</mods:affiliation>
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<mods:namePart>Hu, Shao Ji</mods:namePart>
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<mods:affiliation>Institute of International Rivers and Eco-security, Yunnan University, Kunming, China</mods:affiliation>
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<mods:nameIdentifier type="email">shaojihu@hotmail.com</mods:nameIdentifier>
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<mods:affiliation id="137459AC89C12A8BDE6A48F25A119549">Institute of International Rivers and Eco-security, Yunnan University, Kunming, China</mods:affiliation>
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<mods:number>81</mods:number>
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<treatment LSID="urn:lsid:plazi:treatment:0C74161662CC517EA04E51E9940C49EC" httpUri="http://treatment.plazi.org/id/0C74161662CC517EA04E51E9940C49EC" lastPageNumber="257" pageId="0" pageNumber="257">
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<taxonomicName LSID="0C741616-62CC-517E-A04E-51E9940C49EC" authority="Mell, 1939" authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="latouchei">Pieris latouchei Mell, 1939</taxonomicName>
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<treatment id="0C74161662CC517EA04E51E9940C49EC" ID-GBIF-Taxon="209450352" LSID="urn:lsid:plazi:treatment:0C74161662CC517EA04E51E9940C49EC" httpUri="http://treatment.plazi.org/id/0C74161662CC517EA04E51E9940C49EC" lastPageNumber="257" pageId="0" pageNumber="257" scope_family="Pieridae" scope_order="Lepidoptera">
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<taxonomicName id="4415352486663620BB12F2E437E73553" LSID="0C741616-62CC-517E-A04E-51E9940C49EC" authority="Mell, 1939" authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="latouchei">Pieris latouchei Mell, 1939</taxonomicName>
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</paragraph>
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<paragraph pageId="0" pageNumber="257">
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<taxonomicName authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris extensa subsp. latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="extensa" subSpecies="latouchei">Pieris extensa latouchei</taxonomicName>
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<subSubSection id="FD658CFCEA60522FD333AD0F52CE7125" pageId="0" pageNumber="257" type="reference_group">
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<paragraph id="649120C6E85166565617707444FE78D2" pageId="0" pageNumber="257">
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<taxonomicName id="3BF32C565D88C3AA7793F76FB4D461F4" authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris extensa subsp. latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="extensa" subSpecies="latouchei">Pieris extensa latouchei</taxonomicName>
|
||||
Mell, 1939; Dt. Ent. Z. Iris 52 (1938): 138-139. TL: "Kuatun, NW-Fukien" [N.W. of Shaowu City, Fujian] (original description)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
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<taxonomicName class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris erutae subsp. latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="erutae" subSpecies="latouchei">Pieris erutae latouchei</taxonomicName>
|
||||
<paragraph id="5AFC16708ED0D9ACB04A3AF365D76E14" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="175830C82B1BC5F326B2C9FBDA461D66" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris erutae subsp. latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="erutae" subSpecies="latouchei">Pieris erutae latouchei</taxonomicName>
|
||||
;
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="D06FD871CD39EB4CE67D98AEF054B2BF" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
: Herbipoliana 1(1-2): 376 (as subspecies of
|
||||
<taxonomicName authorityName="Poujade" authorityYear="1888" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris erutae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="erutae">Pieris erutae</taxonomicName>
|
||||
<taxonomicName id="71991B5F75E4D82DB9E12A394513C79A" authorityName="Poujade" authorityYear="1888" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris erutae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="erutae">Pieris erutae</taxonomicName>
|
||||
Poujade, 1888)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete subsp. latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="melete" subSpecies="latouchei">Pieris melete latouchei</taxonomicName>
|
||||
<paragraph id="BAA6E9F4D539529FDBDE9ED6FEBB1E06" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="07019218AAE37923F02BF615FE8F1692" authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete subsp. latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="melete" subSpecies="latouchei">Pieris melete latouchei</taxonomicName>
|
||||
; Wu (2010): Fauna Sinica
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" pageId="0" pageNumber="257" rank="class">Insecta</taxonomicName>
|
||||
<taxonomicName id="BD42AD2C3CF7403F1F19302DC8AE1A76" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" pageId="0" pageNumber="257" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
|
||||
52: 300 (as subspecies of
|
||||
<taxonomicName authorityName="Menetries" authorityYear="1857" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melete">Pieris melete</taxonomicName>
|
||||
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
|
||||
<taxonomicName id="6E18121DB74CA986C1E883DBAEDDADD4" authorityName="Menetries" authorityYear="1857" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melete">Pieris melete</taxonomicName>
|
||||
<normalizedToken id="53CEF64876595BC670700D8047485818" originalValue="Ménétriés">Menetries</normalizedToken>
|
||||
, 1857)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="latouchei">Pieris latouchei</taxonomicName>
|
||||
<paragraph id="DFC026E716D57B0D56F27556BE84FD01" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="ED7DDC209BC0BEB543B48CF05E13678C" authorityName="Mell" authorityYear="1939" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris latouchei" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="latouchei">Pieris latouchei</taxonomicName>
|
||||
;
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="D2BD530036229630890B16EC59F8E450" author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
: Butterflies 65: 25 (raised to distinct species from subspecies of
|
||||
<taxonomicName authorityName="Poujade" authorityYear="1888" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris erutae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="erutae">Pieris erutae</taxonomicName>
|
||||
<taxonomicName id="5F5D88A851C815F1D6F311477C395CD0" authorityName="Poujade" authorityYear="1888" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris erutae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="erutae">Pieris erutae</taxonomicName>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="description">
|
||||
<paragraph pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="C7A522C7AA5B84099BFCA92CD15A9C6F" pageId="0" pageNumber="257" type="description">
|
||||
<paragraph id="EA0934CFE8F8A7E4EF07DD302E1AA560" pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph id="72FBBDD249F575DC70098515BBF02D2E" pageId="0" pageNumber="257">
|
||||
Both wings white on the upperside while bright yellow on the underside (in autumn form males whitish). Spring form: medium size. Male (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10A</figureCitation>
|
||||
<figureCitation id="9FF21B428C6A6220CB0D7B6B150085EE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10A</figureCitation>
|
||||
): apical spot blackish with slightly whitish powder, merging as a small dark subtriangular spot on the apex. The 1st discal spot and outer spot rather faint or absent on the upperside. All spots absent on the underside, except the 2nd discal spot vaguely developed. Female: apical spot brownish, partly merging as a subtriangular spot on the apex. The 1st, 2nd and outer spot strongly developed on the upperside while faintly present on the underside (except outer spot absent). All veins narrowly brown powdered on both upperside and underside. Summer form: medium to large size. Resembles spring form but larger in size and rounder wing shape. The 1st discal spot of males (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10B</figureCitation>
|
||||
<figureCitation id="0F85EC800706659D3DCF3F8DBB682038" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10B</figureCitation>
|
||||
) present on both sides and the brown powder along veins is stronger in summer form of females (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10D</figureCitation>
|
||||
<figureCitation id="14144B050C7CF888440FA7FCE4CD5F45" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10D</figureCitation>
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||||
) than spring forms. Autumn form: large size. Male (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10C</figureCitation>
|
||||
<figureCitation id="29B8803081A8908143B6A85C6F7FD454" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10C</figureCitation>
|
||||
): resembles summer form but less yellowish on the underside (near whitish), the apical spot merging as a triangular spot and the 1st and 2nd discal spots rather distinct on both sides (the outer spot distinct on the upperside while absent on the underside). Female (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10E</figureCitation>
|
||||
<figureCitation id="645C1C3E1D7C53C2325766D4BECFF7F4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upper sid, on the right show the underside. A - E Pieris latouchei Mell, 1939. F - K Pieris erutae erutae Poujade, 1888. L - O Pieris erutae kneitzi Eitschberger, 1983 comb. rev. P - R Pieris erutae reissingeri Eitschberger, 1983. S - T P. steinigeri Eitschberger, 1983. Scale bar represents 10 mm. (All type specimens reference Eitschberger, 1983)." figureDoi="10.3897/asp.81.e85191.figure10" httpUri="https://binary.pensoft.net/fig/825205" pageId="0" pageNumber="257">10E</figureCitation>
|
||||
): resembles summer form but dark yellowish on the underside, all veins strongly brown powdered on the upperside while quite narrow on the underside.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">East China (Henan, Shandong, Jiangsu, Zhejiang, Fujian)</paragraph>
|
||||
<subSubSection id="9E3564720FD6AF1114D3D2D49572F7F4" pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph id="4A01287179D2C060C93A6D8446097926" pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph id="0FAD313F373E1458C214FA63BB213C2A" pageId="0" pageNumber="257">East China (Henan, Shandong, Jiangsu, Zhejiang, Fujian)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Multivoltine, occurs from April to September.</paragraph>
|
||||
<subSubSection id="C553A68BF637093239C35CC6E95C6CE5" pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph id="E9C2F9127F24459FEBA2FD54377B0472" pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph id="80E81B515C8716A6900BB0A849850D9C" pageId="0" pageNumber="257">Multivoltine, occurs from April to September.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="15D813E5C76033AD2F773EAC6549CC35" pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph id="B14D3F2CDA225C26E04E586457EDBFE1" pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph id="C6D8E81A91687E85AA2B00315D4A3B16" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5K</figureCitation>
|
||||
<figureCitation id="76223A610EEA9C7D18A826D00B4ECF80" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5K</figureCitation>
|
||||
) tegumen relatively slender, with its median part slightly concave, the distal margin of tegumen without distinct convex; basal margin of the uncus slightly narrower than the tegumen; uncus with its apical obtusely digitation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="00ECF23FE8746CAE7EED858EAC9F20FC" pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph id="01B298AC09C8CE3B0C978C7DC72EC8D0" pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph id="C22A8CB6D5B47EF99BBD91A3C48ED2EB" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female genitalia of P. napi - complex species. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure7" httpUri="https://binary.pensoft.net/fig/825202" pageId="0" pageNumber="257">7A</figureCitation>
|
||||
<figureCitation id="0BF59539BFCF29258F1F4C71D3260741" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female genitalia of P. napi - complex species. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure7" httpUri="https://binary.pensoft.net/fig/825202" pageId="0" pageNumber="257">7A</figureCitation>
|
||||
) posterior apophysis slender, extended reach to the 8th tergum; inner distal of sterigma lobe-shaped, extend to center, with a distinct convex at the trailing edge; inner basal of sterigma near trapezoidal extend to the center, smoothly connected at the basal margin; signum cordiform-shaped, relatively slender, smoothly tapering to the basal.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="F139384A8D98C9D52ECA0EF2209AA195" pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph id="560EA007CAF0A5A0B0B01C8B7F09523B" pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph id="76E0669DC32E7D1A8212F4355FEF62B6" pageId="0" pageNumber="257">
|
||||
This taxon was elevated to species level by
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="433A3DAF7D40772A75B4628BD0D6A133" author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
, based on results from his phylogenetic analysis, which is also supported by our molecular results. This species is closely related to
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. erutae" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="erutae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. erutae</emphasis>
|
||||
<taxonomicName id="CC34A28895D84ACC9AE07EB8D9816015" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. erutae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="erutae">
|
||||
<emphasis id="27970782CE984361322FB617CF1E4636" italics="true" pageId="0" pageNumber="257">P. erutae</emphasis>
|
||||
</taxonomicName>
|
||||
and shows three different seasonal forms in generations, thus can be only distinguished from the bright yellow color of wings on the underside (while in autumn forms are -variable) and the basal 1/2 dark powdered discocell on the underside (only developed in autumn forms).
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,201 +1,201 @@
|
|||
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<treatment ID-GBIF-Taxon="158995558" LSID="urn:lsid:plazi:treatment:0DA77899943B5226BBA57EB36328B122" httpUri="http://treatment.plazi.org/id/0DA77899943B5226BBA57EB36328B122" lastPageNumber="19" pageId="0" pageNumber="19">
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<taxonomicName LSID="0da77899-943b-5226-bba5-7eb36328b122" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">Copelatus baculiformis Guignot, 1955</taxonomicName>
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
|
||||
<treatment id="0DA77899943B5226BBA57EB36328B122" ID-GBIF-Taxon="158995558" LSID="urn:lsid:plazi:treatment:0DA77899943B5226BBA57EB36328B122" httpUri="http://treatment.plazi.org/id/0DA77899943B5226BBA57EB36328B122" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
|
||||
<subSubSection id="F55ED81927E7F28655280C44BF5CFF56" pageId="0" pageNumber="19" type="nomenclature">
|
||||
<paragraph id="853CA0CC962C350C8D6ECFEA6D8D862C" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="AC7128F43C1C27E00853CAE1F72EC4F0" ID-CoL="Y343" LSID="0da77899-943b-5226-bba5-7eb36328b122" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">Copelatus baculiformis Guignot, 1955</taxonomicName>
|
||||
<figureCitation id="15FA9B2F717E305710597AA15A4C920A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">Copelatus baculiformis</taxonomicName>
|
||||
<subSubSection id="F315B2D4CA39EDC581C0ADBB69CAF5A7" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="375892D98E884D9843EF4EA671D0CB13" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="BD2B815D47EB831D3191264C525B27AD" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">Copelatus baculiformis</taxonomicName>
|
||||
Guignot, 1955b: 193.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Madagascar, Massif Ankaratra, Manjakatompo, alt. 1700-1800 m.</paragraph>
|
||||
<subSubSection id="45359DBFF0251179F09753FB893FEE57" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="55F8E2D4C3C5A472D15E915B23AB01C2" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="E59DF82135E5B5EA3FF033267C026633" pageId="0" pageNumber="19">Madagascar, Massif Ankaratra, Manjakatompo, alt. 1700-1800 m.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">based on a single female specimen (holotype), collected December 1951 by R. Benoist.</paragraph>
|
||||
<subSubSection id="ECD1BD41C3D95B5260E3F12EA955252E" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="C102CE970C78173764C34BC39A238E08" pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph id="E7012ECC22358558383B237A5A152378" pageId="0" pageNumber="19">based on a single female specimen (holotype), collected December 1951 by R. Benoist.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antananarivo. Vakinankaratra: Ambatolampy</emphasis>
|
||||
<subSubSection id="0EE7852BD6BC85E9C1F91E627DC048C4" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="123EF1E76913F8F7A759FB963D62DF9D" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="0D6DB93E3F435AF0D0D3363707B24795" pageId="0" pageNumber="19">
|
||||
<emphasis id="ED654689947DD34E272F98CB95A47040" bold="true" pageId="0" pageNumber="19">Antananarivo. Vakinankaratra: Ambatolampy</emphasis>
|
||||
: -HT ♀ (MNHN "coll. Guignot"): // Data in NHRS | JLKB | 000030226 // Madagascar: Massif An- | karatra 1700/1800 Man- | jakatompo XII-51 Benoist // Type [red label] // Guignot det., 1954 |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="0A9D482544CB8669427AA1FCC83FEEF2" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">baculiformis</emphasis>
|
||||
<emphasis id="AA45E54D1F3E3B6903C1DBBDD96B4928" italics="true" pageId="0" pageNumber="19">baculiformis</emphasis>
|
||||
n. sp. | Type ♀ //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="7EC34ECC67A9A75F529185D3A8CDE493" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="80752B9A70D5EE82773734C42D9E4DC3" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="B7B5A25A78412C8ECAC946275690C67D" pageId="0" pageNumber="19">
|
||||
Small size (4 mm). Elytra uniformly dark brown ferrugineous, without a basal testaceous area (
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
|
||||
<figureCitation id="83257F73029348C3205AFF87AA2769E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
|
||||
) which separates the species from small species of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
<taxonomicName id="9BEFF41AE0A8E2126B8AFE6C66C66421" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="953141D28673FE213770CB92EE428AFC" italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
. The absence of deep impressed elytral striae (remnants of four elytral striae present) separates this species from all other
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="7706AA9A9C4F3FEE7EC9A335490B2D73" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C4AC143598E4783A290A3DD25701CF97" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
of Madagascar except
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus peridinus</emphasis>
|
||||
<taxonomicName id="8E7CAA8F3DAEAD3EA5D77C83771DD235" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">
|
||||
<emphasis id="A122DFEB3A33558CC6531C6C0A634F8A" italics="true" pageId="0" pageNumber="19">Copelatus peridinus</emphasis>
|
||||
</taxonomicName>
|
||||
, a larger species (5.7-6.6 mm).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">(based on holotype ♀):</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="A96350E21DE272A52123CF18D8415C2C" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="EE459DD8128AECDA44F082FB88D0CFE4" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="123DD4C4A4E214A5795EF66B28DF91CC" pageId="0" pageNumber="19">(based on holotype ♀):</paragraph>
|
||||
<paragraph id="702DBC8BEE8D9F35BAEC1A9AE67437B6" pageId="0" pageNumber="19">
|
||||
Body length 4 mm. Body shape elongate oval and dark brown to blackish ferrugineous. Head uniformly dark brown ferrugineous to slightly darker posteriorly inside eyes, with thin sparse punctation. Pronotum dark brown ferrugineous, same colour medially and laterally but darker along anterior and posterior third. Disc of pronotum less densely punctuated, posterolateral corners with dense superficial strioles. The entire dorsal surface covered with a microsculpture. Elytra uniformly coloured in same dark brown to blackish ferrugineous colour as anterior and posterior parts of pronotum (
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
|
||||
<figureCitation id="4CBCF989E55F897C142C5FED5E9E7CF1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
|
||||
). Impressed striae absent but four elytral rows of impressed points present. Innermost, a presumed first row completely lacking, row 2, 3, and 4 visible from base but row 3 less distinct than 2 and 4. Row 5 very vague and only visible posteriorly. Elytra densely covered with punctures which laterally of approximately the third row of points is replaced by strioles (probably only in the female). Appendages testaceous.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19" start="Figure 7" startId="F7">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<caption id="8408EB86BC53376521099B37F74A5F57" doi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19" start="Figure 7" startId="F7">
|
||||
<paragraph id="231B09A0E8AA2ED9B918AD88A8FAAA71" pageId="0" pageNumber="19">
|
||||
Figure 7. Habitus, dorsal view.
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
|
||||
<emphasis id="8253EF529D59AEAA6BAD65754C653CAC" bold="true" pageId="0" pageNumber="19">A</emphasis>
|
||||
Male:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus peridinus</emphasis>
|
||||
<taxonomicName id="3A55C8B85B366D44C92C39E9A75512DA" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">
|
||||
<emphasis id="3D8A2976FCA7435854EDA4F3A9EB5CA4" italics="true" pageId="0" pageNumber="19">Copelatus peridinus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||||
<emphasis id="C29E3D8B9CCAD067196D55C3379C8BFE" bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus baculiformis</emphasis>
|
||||
<taxonomicName id="202717AA772A59ACD3146970A0ADFF8F" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">
|
||||
<emphasis id="F08CF67A98F8A729F2FD9AB68D95988E" italics="true" pageId="0" pageNumber="19">Copelatus baculiformis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||||
<emphasis id="652155FFF730CE556CF89C7484CE1AAA" bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="4665350360AD1539B165F913DF3BDB1E" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="6D545971C30B2B0A620515111D1F7570" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 2 (Andasibe)
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||||
<emphasis id="5FD3BCE65D8A74D850B1DD6A641FDFBB" bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
|
||||
<taxonomicName id="D745AFE0DE847DCF0C89844E0DF5568C" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
|
||||
<emphasis id="5A8F406974AF28D20150C70FD79140F8" italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="70292BB0CCE5D5AF9092AA39A36182F9" pageId="0" pageNumber="19">
|
||||
Ventral side testaceous to weakly infuscated. Prosternal process carinate also onto apical process. Lateral parts of metaventrite ("metasternal wings") rather broad. Metacoxal lines anteriorly diverging and abbreviated before metaventral margin. Metacoxa with fine and long longitudinal strioles, continuing onto abdominal ventrites, and with 6-7 transverse
|
||||
<normalizedToken originalValue="“wrinkles”">"wrinkles"</normalizedToken>
|
||||
<normalizedToken id="2204FD980A782E83E8B7F7BA7FE5067C" originalValue="“wrinkles”">"wrinkles"</normalizedToken>
|
||||
anterolaterally.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Male: unknown.</paragraph>
|
||||
<paragraph id="8D489EA1940FEFE2D8037270BECE4BF2" pageId="0" pageNumber="19">Male: unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="15100923DBFEC1022820DA72FE2B8ABA" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="F2A876E6F534A5A720A6BCDD6AD05C25" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="68D0E65D1CA0BDAC758F82652C565B2D" pageId="0" pageNumber="19">
|
||||
Madagascar, central highlands, only known from type locality Manjakatompo, Ankaratra Massif (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
|
||||
<figureCitation id="6EA2657E23778C0BC6A9C59481978960" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="CBAD4D942EE89B4066EE963808F5ED87" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="452031907482BA58FBC4D365767A5088" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="2BD7CBC323399E6E9E7518A7DCE6D874" pageId="0" pageNumber="19">
|
||||
Unknown, but according to original description collected at at an altitude of 1700-1800 m. See
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.350.6127" author="Hjalmarsson, AE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="21 - 45" refId="B38" refString="Hjalmarsson, AE, Bukontaite, R, Ranarilalatiana, T, Randriamihaja, JH, Bergsten, J, 2013. Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority. . ZooKeys 350: 21 - 45" title="Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority." url="https://doi.org/10.3897/zookeys.350.6127" volume="350" year="2013">Hjalmarsson et al. (2013)</bibRefCitation>
|
||||
<bibRefCitation id="6D6F6016DB11FED58793C3B5368DE3D0" DOI="https://doi.org/10.3897/zookeys.350.6127" author="Hjalmarsson, AE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="21 - 45" refId="B38" refString="Hjalmarsson, AE, Bukontaite, R, Ranarilalatiana, T, Randriamihaja, JH, Bergsten, J, 2013. Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority. . ZooKeys 350: 21 - 45" title="Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority." url="https://doi.org/10.3897/zookeys.350.6127" volume="350" year="2013">Hjalmarsson et al. (2013)</bibRefCitation>
|
||||
for a description of the locality Manjakatompo and its conservation priority.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="48C0C15060B479C70E179DEBE188DF19" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="A6DE2B3D746BD3AC42E311BF1F510276" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="076C2BADACA9CF7DC417E797046F2915" pageId="0" pageNumber="19">
|
||||
No other specimen than the female holotype is known of this species and it is a bit of a "mystery species". We have conducted fieldwork at the type locality Manjakatompo multiple times (2011, 2012, 2014, and 2016) but never found any specimens resembling this species. The species belongs in the
|
||||
<taxonomicName lsidName="hydroporoides" pageId="0" pageNumber="19" rank="species" species="hydroporoides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">hydroporoides</emphasis>
|
||||
<taxonomicName id="57C3E7F9D308BC26689A197E6C13A382" lsidName="hydroporoides" pageId="0" pageNumber="19" rank="species" species="hydroporoides">
|
||||
<emphasis id="59281B693D12315741232B0AE1237326" italics="true" pageId="0" pageNumber="19">hydroporoides</emphasis>
|
||||
</taxonomicName>
|
||||
species group of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="C1AE445D145AC917FA7DB62D81B616AE" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="2AE2EEA36FD2EA7DEC3507903D810AFB" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
, but two other Malagasy species placed in this group have turned out to be misidentified
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
<taxonomicName id="60E87A232CB8B36F2BB25AEE14137BFA" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9F4A374FE0E435E3D8EFBBE455119CBB" italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Exocelina</emphasis>
|
||||
<taxonomicName id="54CDFE968613999FC981854EA3272ECF" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="42FB2F1878349A11041FF3B0DD88A43D" italics="true" pageId="0" pageNumber="19">Exocelina</emphasis>
|
||||
</taxonomicName>
|
||||
(see above).
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus baculiformis</emphasis>
|
||||
<taxonomicName id="6E647614E16EFCF0624DAE9CD54D0890" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">
|
||||
<emphasis id="FFDDD318E5D2388EE2AC4046C749B3E5" italics="true" pageId="0" pageNumber="19">Copelatus baculiformis</emphasis>
|
||||
</taxonomicName>
|
||||
was described by Guignot the same year (1955) that he described
|
||||
<taxonomicName lsidName="C. bilunatus" pageId="0" pageNumber="19" rank="species" species="bilunatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. bilunatus</emphasis>
|
||||
<taxonomicName id="3EA18588347122C732AEEE83ED40F03E" lsidName="C. bilunatus" pageId="0" pageNumber="19" rank="species" species="bilunatus">
|
||||
<emphasis id="B69D21D0062E1957B4BE3C02F7DB4A23" italics="true" pageId="0" pageNumber="19">C. bilunatus</emphasis>
|
||||
</taxonomicName>
|
||||
, considered mislabeled (
|
||||
<bibRefCitation author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. 2014</bibRefCitation>
|
||||
<bibRefCitation id="80D8D88B84A8A203B1920D94E20DEE31" author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. 2014</bibRefCitation>
|
||||
) from the Zimmermann collection. It is certainly possible that also
|
||||
<taxonomicName lsidName="C. baculiformis" pageId="0" pageNumber="19" rank="species" species="baculiformis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. baculiformis</emphasis>
|
||||
<taxonomicName id="512AAD6FFECBB34CDE33E2FCEEDE304E" lsidName="C. baculiformis" pageId="0" pageNumber="19" rank="species" species="baculiformis">
|
||||
<emphasis id="94EE59D9FB79E4DCD4EBF13F378D60B7" italics="true" pageId="0" pageNumber="19">C. baculiformis</emphasis>
|
||||
</taxonomicName>
|
||||
is based on a mislabeled specimen, but in contrast to
|
||||
<taxonomicName lsidName="C. bilunatus" pageId="0" pageNumber="19" rank="species" species="bilunatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. bilunatus</emphasis>
|
||||
<taxonomicName id="EAF2355E7BB663270699C1ECEC6B090C" lsidName="C. bilunatus" pageId="0" pageNumber="19" rank="species" species="bilunatus">
|
||||
<emphasis id="4C034784EA05B3F1B9F8DDD0BBEC9072" italics="true" pageId="0" pageNumber="19">C. bilunatus</emphasis>
|
||||
</taxonomicName>
|
||||
the type locality data is more exact; Massif Ankaratra (mountain), Manjakatompo (locality), 1700-1800 m (altitude in meters), XII-51 (collecting month and year), and R. Benoist (collector), speaking against such a mistake. Based on general external morphology, body shape and lack of striae the species resembles
|
||||
<taxonomicName lsidName="C. peridinus" pageId="0" pageNumber="19" rank="species" species="peridinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. peridinus</emphasis>
|
||||
<taxonomicName id="915D4B7D79F994E860110B49AE9B30AC" lsidName="C. peridinus" pageId="0" pageNumber="19" rank="species" species="peridinus">
|
||||
<emphasis id="525C3F645B19E087864F50977DE2F72B" italics="true" pageId="0" pageNumber="19">C. peridinus</emphasis>
|
||||
</taxonomicName>
|
||||
. It is much smaller and therefore likely not conspecific, but they could be closely related, as well as with the female sequenced from Andasibe (NHRS-JLKB000065698).
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,169 +1,169 @@
|
|||
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<treatment ID-GBIF-Taxon="158995557" LSID="urn:lsid:plazi:treatment:246290F17F4059918CAB1EF8A036B9D5" httpUri="http://treatment.plazi.org/id/246290F17F4059918CAB1EF8A036B9D5" lastPageNumber="19" pageId="0" pageNumber="19">
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<taxonomicName LSID="246290f1-7f40-5991-8cab-1ef8a036b9d5" authority="Guignot, 1956" authorityName="Guignot" authorityYear="1956" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus befasicus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="befasicus">Copelatus befasicus Guignot, 1956</taxonomicName>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus, dorsal view. A Female: Copelatus ankaratra sp. nov. B Male: Copelatus pseudostriatus sp. nov. C Male: Copelatus safiotra sp. nov. D Female: Copelatus befasicus." figureDoi="10.3897/zookeys.869.33997.figure10" httpUri="https://binary.pensoft.net/fig/322935" pageId="0" pageNumber="19">Fig. 10D</figureCitation>
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<treatment id="246290F17F4059918CAB1EF8A036B9D5" ID-GBIF-Taxon="158995557" LSID="urn:lsid:plazi:treatment:246290F17F4059918CAB1EF8A036B9D5" httpUri="http://treatment.plazi.org/id/246290F17F4059918CAB1EF8A036B9D5" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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|
||||
<taxonomicName id="8BAA35D97BDDB389B5EFBAEFA39B4F5E" ID-CoL="Y34D" LSID="246290f1-7f40-5991-8cab-1ef8a036b9d5" authority="Guignot, 1956" authorityName="Guignot" authorityYear="1956" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus befasicus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="befasicus">Copelatus befasicus Guignot, 1956</taxonomicName>
|
||||
<figureCitation id="B5CFE0962A829D991512878D76145A5A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus, dorsal view. A Female: Copelatus ankaratra sp. nov. B Male: Copelatus pseudostriatus sp. nov. C Male: Copelatus safiotra sp. nov. D Female: Copelatus befasicus." figureDoi="10.3897/zookeys.869.33997.figure10" httpUri="https://binary.pensoft.net/fig/322935" pageId="0" pageNumber="19">Fig. 10D</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus befasicus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="befasicus">Copelatus befasicus</taxonomicName>
|
||||
<subSubSection id="0C01BE6D0090DBDBC35E7BE840C6E349" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="694FDA7F7F281BA5CBA6A021EB017B2C" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="80CA223DE03330768030F05A3982C58A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus befasicus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="befasicus">Copelatus befasicus</taxonomicName>
|
||||
Guignot, 1956: 79.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Madagascar, Morondava, forest south of Befasy.</paragraph>
|
||||
<subSubSection id="D71572E285E364C5DEEEEB50C58A8232" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="519DBD3E6C33A10E182600510645118F" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="8D620EBB290B8A1E8D9FA5C2420E514D" pageId="0" pageNumber="19">Madagascar, Morondava, forest south of Befasy.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">based on an unknown number of female type specimens but holotype and paratypes are distinguished in introduction. Collected in January 1956 by R. Paulian.</paragraph>
|
||||
<subSubSection id="9F3890C86ED3EB25081CF4045BF84EF9" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="8C270E4056D7515440CF133F312E1BBC" pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph id="5A9192B5526D42421905B9F156B44536" pageId="0" pageNumber="19">based on an unknown number of female type specimens but holotype and paratypes are distinguished in introduction. Collected in January 1956 by R. Paulian.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toliara. Menabe: Morondava</emphasis>
|
||||
<subSubSection id="FEA3A08FE79613705192BAEA0F2E6FF4" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="7B047F1B2227F3FA3B0643AD81ACC48F" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="CEC56179E27BF96317A2AC2B7A7C206F" pageId="0" pageNumber="19">
|
||||
<emphasis id="89A1677272A3408D2813F8EA3EFA3B31" bold="true" pageId="0" pageNumber="19">Toliara. Menabe: Morondava</emphasis>
|
||||
: -HT♀ (MNHN, "coll. IRSM"): // Data in NHRS | JLKB | 000030021 // Morondava |
|
||||
<normalizedToken originalValue="fôret">foret</normalizedToken>
|
||||
<normalizedToken id="FE2B2F862B3F3B3F65B5817F5879243B" originalValue="fôret">foret</normalizedToken>
|
||||
sud | de Befasy | I-56 R.P // Type [red label] // INSTITUT | SCIENTIFIQUE | MADAGASCAR // Guignot det., 1956 |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="1F76B9290B1873CE9BF370903D6931AE" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">befasicus</emphasis>
|
||||
<emphasis id="54033EBD2FFA472DDF07E5CEA1CE27AF" italics="true" pageId="0" pageNumber="19">befasicus</emphasis>
|
||||
n. sp. | Type // -2PT ♀ (MNHN, "coll. Guignot"): // Data in NHRS | JLKB | 000030300-1// Morondava |
|
||||
<normalizedToken originalValue="fôret">foret</normalizedToken>
|
||||
<normalizedToken id="A16BBC825C09CABFF2AB04040CEE689F" originalValue="fôret">foret</normalizedToken>
|
||||
sud | de Befasy // I-56 R.P // Paratype [red label] // [female symbol] //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Morafenobe</emphasis>
|
||||
<subSubSection id="57F1B4C30FC9BBC2FC4896B318D1F139" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="701A1EAB4CAA3B3A0FAFC25E49A1F487" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="4013D807D928DC7FFFC0561BCFD8B5A2" pageId="0" pageNumber="19">
|
||||
<emphasis id="1102C5B5D1856287F7E152DBBE6ABCA1" bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Morafenobe</emphasis>
|
||||
: -1♀ (NHRS): // NHRS-JLKB | 000010860 (JB204) // Madagascar: Mahajanga: Melaky | Btw.
|
||||
<normalizedToken originalValue="Morafenobe–Ambohijanahary">Morafenobe-Ambohijanahary</normalizedToken>
|
||||
<normalizedToken id="2595351A7A63F70ECF078614504163E9" originalValue="Morafenobe–Ambohijanahary">Morafenobe-Ambohijanahary</normalizedToken>
|
||||
| S18.19091; E045.19986, 290 m.a.o | 19.XII.2009 Water Net, Field# MAD09-74 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="960483D82BFAC909D821785A83329278" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="ACD5CE45CEAB5C9E318EA123CDCD78D5" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="42D5AE5F433FE9E92DE051B6F5C2F39D" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="F37D1ED3AD7432FFF1D08F18694D1668" pageId="0" pageNumber="19">
|
||||
Similar to
|
||||
<taxonomicName lsidName="C. insuetus" pageId="0" pageNumber="19" rank="species" species="insuetus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. insuetus</emphasis>
|
||||
<taxonomicName id="9681130C6366B26138FF41FF857FF691" lsidName="C. insuetus" pageId="0" pageNumber="19" rank="species" species="insuetus">
|
||||
<emphasis id="BD3AB6B07164DDE41CC26A84ED78459B" italics="true" pageId="0" pageNumber="19">C. insuetus</emphasis>
|
||||
</taxonomicName>
|
||||
and related species in habitus by being small, elongate, and subparalell, but
|
||||
<taxonomicName lsidName="C. befasicus" pageId="0" pageNumber="19" rank="species" species="befasicus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. befasicus</emphasis>
|
||||
<taxonomicName id="25D9A4F7406CC5B60D294FEB63D62345" lsidName="C. befasicus" pageId="0" pageNumber="19" rank="species" species="befasicus">
|
||||
<emphasis id="B55CBCC6347B673E7B420138A0BF46AC" italics="true" pageId="0" pageNumber="19">C. befasicus</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from
|
||||
<taxonomicName lsidName="C. insuetus" pageId="0" pageNumber="19" rank="species" species="insuetus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. insuetus</emphasis>
|
||||
<taxonomicName id="1FE92B68F46573A0034B9052AB4ADBD0" lsidName="C. insuetus" pageId="0" pageNumber="19" rank="species" species="insuetus">
|
||||
<emphasis id="2094BA547351AE9631711CDE6F75EF42" italics="true" pageId="0" pageNumber="19">C. insuetus</emphasis>
|
||||
</taxonomicName>
|
||||
and from all other Malagasy
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="804E516C354B052F50FC3F5243F204AB" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3F6598150B367032D69A13C2C4B4FD66" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
species by the presence of only five elytral striae, and without submarginal striae. In addition, the first stria is shortened, present only in the posterior third (
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus, dorsal view. A Female: Copelatus ankaratra sp. nov. B Male: Copelatus pseudostriatus sp. nov. C Male: Copelatus safiotra sp. nov. D Female: Copelatus befasicus." figureDoi="10.3897/zookeys.869.33997.figure10" httpUri="https://binary.pensoft.net/fig/322935" pageId="0" pageNumber="19">Fig. 10D</figureCitation>
|
||||
<figureCitation id="31962BAE972F265B1CAB3C752E6EF873" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus, dorsal view. A Female: Copelatus ankaratra sp. nov. B Male: Copelatus pseudostriatus sp. nov. C Male: Copelatus safiotra sp. nov. D Female: Copelatus befasicus." figureDoi="10.3897/zookeys.869.33997.figure10" httpUri="https://binary.pensoft.net/fig/322935" pageId="0" pageNumber="19">Fig. 10D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="A291ABA0057EA097821012109BDF58ED" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="237619F7EEAD87C2BE6CABFB51C0AA21" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="235FC94E617DCF58E7D27F3DB5ACEE02" pageId="0" pageNumber="19">
|
||||
Body length 4.1-4.2 mm. Body shape elongate and subparallel, dorsal surface reddish brown with a lighter elytral base. Head and pronotum uniformly rufus brown. Head, pronotum and elytra with dense punctation. Elytra and pronotum covered with dense punctures and the whole dorsal surface with a microsculpture. Lateral sides of pronotum striolate with the widest striolate area in the posterior corners. Elytra light brown with a distinct testaceous band basally (
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus, dorsal view. A Female: Copelatus ankaratra sp. nov. B Male: Copelatus pseudostriatus sp. nov. C Male: Copelatus safiotra sp. nov. D Female: Copelatus befasicus." figureDoi="10.3897/zookeys.869.33997.figure10" httpUri="https://binary.pensoft.net/fig/322935" pageId="0" pageNumber="19">Fig. 10D</figureCitation>
|
||||
<figureCitation id="9B44A1FE11D34D63F2862E44C0348EF3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus, dorsal view. A Female: Copelatus ankaratra sp. nov. B Male: Copelatus pseudostriatus sp. nov. C Male: Copelatus safiotra sp. nov. D Female: Copelatus befasicus." figureDoi="10.3897/zookeys.869.33997.figure10" httpUri="https://binary.pensoft.net/fig/322935" pageId="0" pageNumber="19">Fig. 10D</figureCitation>
|
||||
). First elytral stria shortened and present only in posterior third. Second to fifth elytral stria starting more or less at base and all striae approaching the apex of elytron, but the second and fourth a little shorter. Submarginal striae absent. Appendages testaceous.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Ventral side light brown. Metacoxa and abdominal ventrites punctate and striolate. Prosternal process rather short and medially raised, triangular in cross-section. Lateral parts of metaventrite medium broad. Metacoxal lines anteriorly diverging but rather weakly so, abbreviated well before metaventral margin. Antennae, palps and legs testaceous.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Male: unknown.</paragraph>
|
||||
<paragraph id="055EBBA3D5C274CAAD6F661846E9E3AC" pageId="0" pageNumber="19">Ventral side light brown. Metacoxa and abdominal ventrites punctate and striolate. Prosternal process rather short and medially raised, triangular in cross-section. Lateral parts of metaventrite medium broad. Metacoxal lines anteriorly diverging but rather weakly so, abbreviated well before metaventral margin. Antennae, palps and legs testaceous.</paragraph>
|
||||
<paragraph id="B0DE742A8FEE654698A2148F0AC16097" pageId="0" pageNumber="19">Male: unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="26E92E524012CB7BFD1322C42E607FE2" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="E25CFA9CFB105EF3D2FB8751A078FB67" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="67CD4429D81EAA8B56DA0C8D748E89C2" pageId="0" pageNumber="19">
|
||||
Known only from two localities in the western part of Madagascar, the deciduous forest south of Befasy, Morondava, and at one locality between Morafenobe and Beravina village (
|
||||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution maps of Copelatus species. A C. insuetus (circle), C. pseudostriatus sp. nov. (square) B C. kely sp. nov. (circle), C. vokoka sp. nov. (square) C C. ankaratra sp. nov. (circle), C. sp. 3 (square) D C. safiotra sp. nov. (circle), C. befasicus (square)." figureDoi="10.3897/zookeys.869.33997.figure12" httpUri="https://binary.pensoft.net/fig/322937" pageId="0" pageNumber="19">Fig. 12D</figureCitation>
|
||||
<figureCitation id="335F9BCFC01D9750AE3AC0BA6C1A0028" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution maps of Copelatus species. A C. insuetus (circle), C. pseudostriatus sp. nov. (square) B C. kely sp. nov. (circle), C. vokoka sp. nov. (square) C C. ankaratra sp. nov. (circle), C. sp. 3 (square) D C. safiotra sp. nov. (circle), C. befasicus (square)." figureDoi="10.3897/zookeys.869.33997.figure12" httpUri="https://binary.pensoft.net/fig/322937" pageId="0" pageNumber="19">Fig. 12D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="D594F8B28EF31F69355BA9CA67519996" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="9F1B705B852D2DE28ACD3305CBC25364" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="68E2FAD60F72109B633F7CC1769A91D3" pageId="0" pageNumber="19">
|
||||
Paulian collected the species in 1956 in the western dry deciduous forest south of Befasy, Morondava. We rediscovered the species in 2009, when we found one female specimen of
|
||||
<taxonomicName lsidName="C. befasicus" pageId="0" pageNumber="19" rank="species" species="befasicus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. befasicus</emphasis>
|
||||
<taxonomicName id="D3BC807ADFBC697981D1E98A5225C11E" lsidName="C. befasicus" pageId="0" pageNumber="19" rank="species" species="befasicus">
|
||||
<emphasis id="5D31705E7044F91DCD3B67FC76A9FBFE" italics="true" pageId="0" pageNumber="19">C. befasicus</emphasis>
|
||||
</taxonomicName>
|
||||
also in the western part but a bit further north than the type locality, along the road between Morafenobe and Beravina village at an altitude of 290 m. This locality consisted of dry savannah with mixed wood and grassland ecosystem after deforestation. The habitat consisted of muddy/sandy residual pools with some dead leaves in a temporary stream after the rainy season. The dry deciduous forest ecosystem in western Madagascar has suffered immensely from deforestation and very little of this habitat remains (
|
||||
<bibRefCitation DOI="https://doi.org/10.1046/j.1365-3008.2001.00201.x" author="Ganzhorn, JU" journalOrPublisher="Oryx" pageId="0" pageNumber="19" pagination="346 - 348" refId="B19" refString="Ganzhorn, JU, Lowry, PP, Schatz, GE, Sommer, A, 2001. The biodiversity of Madagascar: one of the world's hottest hotspots on its way out. . Oryx 35: 346 - 348" title="The biodiversity of Madagascar: one of the world's hottest hotspots on its way out." url="https://doi.org/10.1046/j.1365-3008.2001.00201.x" volume="35" year="2001">Ganzhorn et al. 2001</bibRefCitation>
|
||||
<bibRefCitation id="7EAA43B361832C515E27B993F8799465" DOI="https://doi.org/10.1046/j.1365-3008.2001.00201.x" author="Ganzhorn, JU" journalOrPublisher="Oryx" pageId="0" pageNumber="19" pagination="346 - 348" refId="B19" refString="Ganzhorn, JU, Lowry, PP, Schatz, GE, Sommer, A, 2001. The biodiversity of Madagascar: one of the world's hottest hotspots on its way out. . Oryx 35: 346 - 348" title="The biodiversity of Madagascar: one of the world's hottest hotspots on its way out." url="https://doi.org/10.1046/j.1365-3008.2001.00201.x" volume="35" year="2001">Ganzhorn et al. 2001</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation DOI="https://doi.org/10.1017/S0030605309001756" author="Whitehurst, AS" journalOrPublisher="Oryx" pageId="0" pageNumber="19" pagination="275 - 283" refId="B75" refString="Whitehurst, AS, Sexton, JO, Dollar, L, 2009. Land cover change in western Madagascar's dry deciduous forests: a comparison of forest changes in and around Kirindy Mite National Park. . Oryx 43: 275 - 283" title="Land cover change in western Madagascar's dry deciduous forests: a comparison of forest changes in and around Kirindy Mite National Park." url="https://doi.org/10.1017/S0030605309001756" volume="43" year="2009">Whitehurst et al. 2009</bibRefCitation>
|
||||
<bibRefCitation id="E41C55AC802ADD61C5C59C44E4F08760" DOI="https://doi.org/10.1017/S0030605309001756" author="Whitehurst, AS" journalOrPublisher="Oryx" pageId="0" pageNumber="19" pagination="275 - 283" refId="B75" refString="Whitehurst, AS, Sexton, JO, Dollar, L, 2009. Land cover change in western Madagascar's dry deciduous forests: a comparison of forest changes in and around Kirindy Mite National Park. . Oryx 43: 275 - 283" title="Land cover change in western Madagascar's dry deciduous forests: a comparison of forest changes in and around Kirindy Mite National Park." url="https://doi.org/10.1017/S0030605309001756" volume="43" year="2009">Whitehurst et al. 2009</bibRefCitation>
|
||||
). The species is rare and likely threatened due to the disappearance of western deciduous forests in Madagascar. That all four known specimens are females may suggest an uneven sex ratio as a test of equal sex ratio is marginally non-significant (p = 0.0625) if considered randomly picked from the population.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="D7F2CA4540C19C930D959B278E49CB80" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="BB90C2B89FDFEF1EC80D8666D3C94899" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="23F9F3D8FB7BA4B96D6B4D152FD12537" pageId="0" pageNumber="19">
|
||||
This is the only species in the
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus longicornis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="longicornis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus longicornis</emphasis>
|
||||
<taxonomicName id="D317F81430308FB2D2EE89BB65134227" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus longicornis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="longicornis">
|
||||
<emphasis id="E2116487D8626C74D8F2CD7B00747867" italics="true" pageId="0" pageNumber="19">Copelatus longicornis</emphasis>
|
||||
</taxonomicName>
|
||||
group from Madagascar. The
|
||||
<taxonomicName lsidName="longicornis" pageId="0" pageNumber="19" rank="species" species="longicornis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">longicornis</emphasis>
|
||||
<taxonomicName id="D8FB6A9C26E027F4B25D923477E7A12A" lsidName="longicornis" pageId="0" pageNumber="19" rank="species" species="longicornis">
|
||||
<emphasis id="FBD759C2092C2DE4C9B78631965D4BA6" italics="true" pageId="0" pageNumber="19">longicornis</emphasis>
|
||||
</taxonomicName>
|
||||
species group currently contains 38 species distributed mainly in the Neotropical and Afrotropical regions but also with few species present in Japan, New Guinea, and Fiji Islands (
|
||||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" publicationUrl="http://www.waterbeetles.eu" refId="B57" refString="Nilsson, AN, Hajek, J, 2018. . http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2018">
|
||||
<bibRefCitation id="E4E87F0924092505D558DA7E9026BBFE" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" publicationUrl="http://www.waterbeetles.eu" refId="B57" refString="Nilsson, AN, Hajek, J, 2018. . http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2018">
|
||||
Nilsson and
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<normalizedToken id="C8D29D23014B68D5E96F0DE41DD49AC7" originalValue="Hájek">Hajek</normalizedToken>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
). The group as currently defined is certainly artificial from a phylogenetic perspective and the only character they have in common is the low number of elytral striae.
|
||||
|
|
|
|||
|
|
@ -1,169 +1,169 @@
|
|||
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<treatment ID-GBIF-Taxon="158995556" LSID="urn:lsid:plazi:treatment:25AF5D96C7285A06B37C8F9CC93796E0" httpUri="http://treatment.plazi.org/id/25AF5D96C7285A06B37C8F9CC93796E0" lastPageNumber="19" pageId="0" pageNumber="19">
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||||
<taxonomicName LSID="25af5d96-c728-5a06-b37c-8f9cc93796e0" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">Copelatus peridinus Guignot, 1955</taxonomicName>
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Figs 4A</figureCitation>
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<treatment id="25AF5D96C7285A06B37C8F9CC93796E0" ID-GBIF-Taxon="158995556" LSID="urn:lsid:plazi:treatment:25AF5D96C7285A06B37C8F9CC93796E0" httpUri="http://treatment.plazi.org/id/25AF5D96C7285A06B37C8F9CC93796E0" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
|
||||
<subSubSection id="4A01619061090BD8712AE0E885FD14DA" pageId="0" pageNumber="19" type="nomenclature">
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<paragraph id="F987F8E8C0416E35508319312C43011D" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="CA229992BF0D71BAACBFFF82029505AB" ID-CoL="5ZX8G" LSID="25af5d96-c728-5a06-b37c-8f9cc93796e0" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">Copelatus peridinus Guignot, 1955</taxonomicName>
|
||||
<figureCitation id="84BDF2C8C306F936D159F1A9918D96D4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Figs 4A</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">7A</figureCitation>
|
||||
<figureCitation id="EA20EE32BE55AD03E5EF925AF4A0DDFE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">7A</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">Copelatus peridinus</taxonomicName>
|
||||
<subSubSection id="EE97F873F97C1C2D3BB84B3DD01AC910" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="B5A7F3B9FA85F3289999079DE29894F7" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="4269D97C1218F5A815AA818FADAEC1D2" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">Copelatus peridinus</taxonomicName>
|
||||
Guignot, 1955c: 188.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus seydeli" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="seydeli">Copelatus seydeli</taxonomicName>
|
||||
<paragraph id="68DC891E919837D94F85AF754088D1CC" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="7003F20991FCEE529F45813AA4DBD51E" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus seydeli" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="seydeli">Copelatus seydeli</taxonomicName>
|
||||
Guignot, 1958: 107; TL: Elisabethville, Zaire [DR Congo, Haut-Katanga, Lubumbashi].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Elisabethville, Zaire [DR Congo, Haut-Katanga, Lubumbashi].</paragraph>
|
||||
<subSubSection id="468EBB9AC9703A632E8F08BF791726A3" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="DB7FA94B837708D5BC1ECE3A0C3B47A4" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="336737CED999F264D8718E24718EB35A" pageId="0" pageNumber="19">Elisabethville, Zaire [DR Congo, Haut-Katanga, Lubumbashi].</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="C0CDA50AC187EA9552591151886C57A8" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="217357E907427EB43100E97210D1439A" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="81C50F256D040C413EE2E830A8ED1945" pageId="0" pageNumber="19">
|
||||
-PT ♀ (MNHN, "Coll. Guignot"): // Data in NHRS | JLKB | 000030317 // Allotype [red label] // [female symbol] // Elisabethville | XI. 1951 //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="FD9498F4D8AEAA81767BD7B384AAF346" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">peridinus</emphasis>
|
||||
<emphasis id="D70714A2F7B9D82F49CA95806848EEE9" italics="true" pageId="0" pageNumber="19">peridinus</emphasis>
|
||||
| Allotype ♀ //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="9CDDFF0002AA3E58FD8AF0C8AEF0CBF5" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="98EA8AC5891119539136ED1266505318" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="1FE005C86F0118E6ECD14A4EC83B73FA" pageId="0" pageNumber="19">
|
||||
-1♂(GP) (MNHN, "Coll. Guignot"): // Data in NHRS | JLKB | 000030318 // Congo Belge Lac | Edouard: Vitshumbi | U.V. 27.XI.1953 | 3091 //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antananarivo. Analamanga: Antananarivo</emphasis>
|
||||
<emphasis id="9B514C288511C8834D6F73469B4F4BF1" bold="true" pageId="0" pageNumber="19">Antananarivo. Analamanga: Antananarivo</emphasis>
|
||||
: -3♂(GP), 3♀ (MNHN, "Coll. Legros Magasin"): // Data in NHRS | JLKB | 000030319-24 // MADAGASCAR | TANANARIVE | BETONGOLO | 2 XII 1946 //
|
||||
<normalizedToken originalValue="Piège">Piege</normalizedToken>
|
||||
<normalizedToken id="57B9131127DA4C7F7B59574F754E3699" originalValue="Piège">Piege</normalizedToken>
|
||||
| lumineux // Museum Paris | 1983 | Coll. Cl. Legros //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga</emphasis>
|
||||
<emphasis id="67028A5063E9D0B82985337D4397865E" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga</emphasis>
|
||||
: -1♂(GP), 2♀ (MNHN, "Coll. Paulian"): // Data in NHRS | JLKB | 000030316, 30325-6 // Madagascar Est | P.K.57-Rte
|
||||
<normalizedToken originalValue="d’Anosibe">d'Anosibe</normalizedToken>
|
||||
<normalizedToken id="D423C70DBFB5E08605DC817274B052C0" originalValue="d’Anosibe">d'Anosibe</normalizedToken>
|
||||
| Moramanga | II.58 R. Vieu // -1♂(GP), 1♀ (NMPC, "Coll. J.
|
||||
<normalizedToken originalValue="Hájek”">Hajek"</normalizedToken>
|
||||
<normalizedToken id="43D01124193BE6AA3B35A53DA93B7D08" originalValue="Hájek”">Hajek"</normalizedToken>
|
||||
): // Data in NHRS | JLKB | 000030327, 30328 // Madagascar | Lokato, near | Andasibe Mantadia NP | M. Tryzna leg., 9-10.i.2007 // coll. Jiri
|
||||
<normalizedToken originalValue="HÁJEK">HAJEK</normalizedToken>
|
||||
<normalizedToken id="6D682DEF9DF5757B93C8D6FF9FE2CA01" originalValue="HÁJEK">HAJEK</normalizedToken>
|
||||
| National Museum | Prague, Czech Republic // ["Lokato near Andasibe Mantadia NP" interpreted as the bifurcation of the road to Lakato which is near Andasibe Mantadia NP. Lakato itself is not near Andasibe Mantadia NP.] -2♂(GP), 1♀(teneral), 1♀, 1♂ (Alc. teneral) (NHRS): // NHRS-JLKB | 000010887, 65737, 65703, 10888(Alc.) // MAD: TOAM: Alaotra Mangoro: | Andasibe Mantadia NP, Analamazaotra: | 150m E of park entrance: Mad14-14 | shallow partly dried out forest pond: | 18.9355S | 48.4166E: 930m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja & T. Ranarilalatiana //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="DA76CEE47442A2B705F7D5668C3DC7AB" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="17C92D8D4C7B5F03F4F88A0B733413EB" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="78C373B0CCA7084BB7993B0C7FEF621F" pageId="0" pageNumber="19">
|
||||
Similar to
|
||||
<taxonomicName lsidName="C. baculiformis" pageId="0" pageNumber="19" rank="species" species="baculiformis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. baculiformis</emphasis>
|
||||
<taxonomicName id="0D9F0E051BADA5456B7CF084354B1343" lsidName="C. baculiformis" pageId="0" pageNumber="19" rank="species" species="baculiformis">
|
||||
<emphasis id="B336B003A222535DBB32280E6E18CD26" italics="true" pageId="0" pageNumber="19">C. baculiformis</emphasis>
|
||||
</taxonomicName>
|
||||
based on shape and colouration, but
|
||||
<taxonomicName lsidName="C. peridinus" pageId="0" pageNumber="19" rank="species" species="peridinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. peridinus</emphasis>
|
||||
<taxonomicName id="D18D6EB6EB1D2046DDD0A0D7050BDC18" lsidName="C. peridinus" pageId="0" pageNumber="19" rank="species" species="peridinus">
|
||||
<emphasis id="F42FE73D219D922EEBFBD2FE23303E83" italics="true" pageId="0" pageNumber="19">C. peridinus</emphasis>
|
||||
</taxonomicName>
|
||||
is bigger with body length between 5.7 and 6.6 mm (
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7A</figureCitation>
|
||||
<figureCitation id="6D5B186B04732DF19E3BB5D83B471D75" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7A</figureCitation>
|
||||
). Elytra lacking deeply impressed striae, but elytral rows of impressed points present out of which two rows are most distinct. Penis in lateral view curved with two points at which it curves slightly more abruptly creating two gentle angles (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
<figureCitation id="A9382FE24A37F09D1D9DCEA3A9F5DBF7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
); parameres rather broad at base and evenly curved (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
<figureCitation id="3AAC23934063C32F38642A4AADA9FC38" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="569F81730ED0DBBDBECE7E1B8533FA6E" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="79A3453D8CAF0D405DD2ED8150B84ED5" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="3D826CAD00411A678B3AD3A5595BF1B7" pageId="0" pageNumber="19">
|
||||
Body length 5.7-6.6 mm. Body shape elongate oval, from midpoint uniformly tapering towards a rather pointed apex. Head and pronotum both in the same dark brown to blackish ferrugineous colouration; elytra anteriorly in the same colour as pronotum, but posterior part brown ferrugineous with largely testaceous lateral margins (
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7A</figureCitation>
|
||||
<figureCitation id="5F9FCFE96CC9D53E58985C6F74E1F317" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7A</figureCitation>
|
||||
). Head, pronotum and elytra with thin dense punctation, in addition pronotum and elytra with small and shallow punctures, and microsculptured. At posterolateral corners of pronotum, punctures almost a little joined and corrugated. Elytra lacking deeply impressed striae. Two continuous rows of punctures are most obvious and divide the elytra in three more or less equal intervals. A third and a fourth row of punctures also present, albeit less distinct and more fragmented. These occur between the two first distinct rows and between the second distinct row and the elytral margin. A fifth row suggested between the two outermost rows by a few spaced out punctures, almost unidentifiable in most Malagasy specimens seen but more distinct in specimens seen from mainland Africa. Appendages testaceous to rufus.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Ventral side dark ferrugineous. Prosternal process strongly carinate anteriorly and with a rather short process. Lateral parts of the metaventrite medium-broad. Metacoxal lines short, anteriorly diverging and abbreviated well before metaventral margin. Metacoxa with short fine strioles continuing onto abdominal ventrites.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Male: first three pro- and mesotarsomeres widened. Protibia modified, narrow at base and with an early abrupt bend, extended and broadened towards middle with a straight ventral side but angled dorsal side. Pro- and mesotarsal claws unmodified.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="59980FF0AB7CA27D37DD7C99E3903F80" pageId="0" pageNumber="19">Ventral side dark ferrugineous. Prosternal process strongly carinate anteriorly and with a rather short process. Lateral parts of the metaventrite medium-broad. Metacoxal lines short, anteriorly diverging and abbreviated well before metaventral margin. Metacoxa with short fine strioles continuing onto abdominal ventrites.</paragraph>
|
||||
<paragraph id="ECABA57072CDF36665BC3A61B5FF2FF8" pageId="0" pageNumber="19">Male: first three pro- and mesotarsomeres widened. Protibia modified, narrow at base and with an early abrupt bend, extended and broadened towards middle with a straight ventral side but angled dorsal side. Pro- and mesotarsal claws unmodified.</paragraph>
|
||||
<paragraph id="F7CAB181B7505E7F67449891BA3C7F03" pageId="0" pageNumber="19">
|
||||
Penis in ventral view with apical part more or less straight and even, gently pointed at apex but with the very apex minutely twisted to the right (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
<figureCitation id="F9BF79B160DDF294954387E64A42C500" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
). Penis in lateral view curved with two points (1/3rd and 2/3rd from base) at which it curves slightly more abruptly creating two gentle angles (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
<figureCitation id="9DD24AB72C28E7F49A050D94234880F1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
). Parameres rather broad at base and evenly curved and tapering towards apex (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
<figureCitation id="B407C76EEFB970DABFA154526FFC6478" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Female: dorsal sculpture similar to male.</paragraph>
|
||||
<paragraph id="BBD50F80959D5578F687B01526F44B16" pageId="0" pageNumber="19">Female: dorsal sculpture similar to male.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="B78A4024C85750B87CB6D9AFBAB5285B" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="04ACDEBDFC20EE5A2DB5B9102BB65B96" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="BF22E840632F3089248192CEF07C65FF" pageId="0" pageNumber="19">
|
||||
Likely a more widespread distribution in at least central and eastern continental Africa than the current records (Lubumbashi and Kivu in DR Congo) indicate. In Madagascar, only know from the eastern central parts: Betongolo (Antananarivo), the Analamazaotra NP, and P.K.57 Route
|
||||
<normalizedToken originalValue="d’Anosibe">d'Anosibe</normalizedToken>
|
||||
<normalizedToken id="848CA2E3D9BAA71096660C8CF10A8770" originalValue="d’Anosibe">d'Anosibe</normalizedToken>
|
||||
[RN23] (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11C</figureCitation>
|
||||
<figureCitation id="294BBD8B12952E40D59629D0ABA228BB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="C8F7873B325F312DF25A65617099B45C" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="2087D11D91C99FE563DB59269F8F1A3D" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="AE1FDCE66792066091EBBA747BD36B35" pageId="0" pageNumber="19">
|
||||
A series of six specimens collected with light trap (
|
||||
<normalizedToken originalValue="“piège">"piege</normalizedToken>
|
||||
<normalizedToken id="3B6875E186315A92C4BBA810D33C43FE" originalValue="“piège">"piege</normalizedToken>
|
||||
lumineux") in the capital Antananarivo 1946, indicates flight capacity and anthropogenically disturbed habitats. All records from DR Congo are also from light trap catches (
|
||||
<bibRefCitation author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali del Museo Civico di Storia Naturale di Milano" pageId="0" pageNumber="19" pagination="441 - 461" refId="B7" refString="Bilardo, A, 1982. Contributo alla conoscenza dei Dytiscidae del Cameroun (Coleoptera). . Atti della Societa Italiana di Scienze Naturali del Museo Civico di Storia Naturale di Milano 123: 441 - 461" title="Contributo alla conoscenza dei Dytiscidae del Cameroun (Coleoptera)." volume="123" year="1982">Bilardo 1982</bibRefCitation>
|
||||
<bibRefCitation id="F908EE890863CCB9738FB98E2B62DA28" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali del Museo Civico di Storia Naturale di Milano" pageId="0" pageNumber="19" pagination="441 - 461" refId="B7" refString="Bilardo, A, 1982. Contributo alla conoscenza dei Dytiscidae del Cameroun (Coleoptera). . Atti della Societa Italiana di Scienze Naturali del Museo Civico di Storia Naturale di Milano 123: 441 - 461" title="Contributo alla conoscenza dei Dytiscidae del Cameroun (Coleoptera)." volume="123" year="1982">Bilardo 1982</bibRefCitation>
|
||||
). We collected a second series of teneral specimens in a shallow forest pool with vegetation, near the entrance to the Analamazaotra NP.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">This species was described from Lubumbashi, DR Congo, and has not been recorded from Madagascar before. Earliest record found is from November 1946. It seems to be a dispersive good flier and often collected at light so its presence in Madagascar is therefore not surprising. However, it is not widespread in Madagascar as far as we know. In fact, the known distribution is restricted to the surroundings of the capital and east of the capital along the main national route towards Toamasina, which could suggest a recent incidental human-mediated introduction from mainland Africa.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="69F0DF1F68763AA3EC221FAE05363026" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="10881E20982E3EBAD5797D8AD87B0434" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="405BA574C829F2FD05BC4539A7C5B0AE" pageId="0" pageNumber="19">This species was described from Lubumbashi, DR Congo, and has not been recorded from Madagascar before. Earliest record found is from November 1946. It seems to be a dispersive good flier and often collected at light so its presence in Madagascar is therefore not surprising. However, it is not widespread in Madagascar as far as we know. In fact, the known distribution is restricted to the surroundings of the capital and east of the capital along the main national route towards Toamasina, which could suggest a recent incidental human-mediated introduction from mainland Africa.</paragraph>
|
||||
<paragraph id="5DF16173BDDB6F6ACA097087E14CB75D" pageId="0" pageNumber="19">
|
||||
Note that it may be that this is a species with intraspecific variation with regards to elytral striation, ranging from five puncture lines out of which two are more distinct, to five weakly impressed striae (see further discussion under
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="721AC8174F9BF154AF3B8426AAA7C662" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7534CDC2E1ADBBBD8EDD949F5D2C8527" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 2 below).
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,198 +1,198 @@
|
|||
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<mods:titleInfo>
|
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<mods:title>A revision of the Pieris napi - complex (Lepidoptera: Pieridae) and similar species with distribution in China</mods:title>
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Verity, 1911; Rho-pa-locera Palaearctica (30-36): 329, pl. 66, f. 21-23; TL: "Gyangtse, S. Tibet" (original description)
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;
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: The Macrolepido-ptera of the World, supplement to Vol. 1: 339 pp, pl.16. (infrasubspecific name validated as subspecies of
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)
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<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
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: Butterflies 65: 33 (as distinct species)
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<taxonomicName authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris chumbiensis subsp. gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="chumbiensis" subSpecies="gyantsensis">Pieris chumbiensis gyantsensis</taxonomicName>
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;
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Lepidoptera Science" pageId="0" pageNumber="257" pagination="99 - 114" refId="B20" refString="Tadokoro, T, Inomata, T, Wang, M, 2016. Taxonomic review of the Pieris dubernardi -group, with description of a new subspecies from Northern Sichuan, China (Lepidoptera, Pieridae). Lepidoptera Science 67 (3-4): 99 - 114" title="Taxonomic review of the Pieris dubernardi - group, with description of a new subspecies from Northern Sichuan, China (Lepidoptera, Pieridae)." volume="67" year="2016">Tadokoro et al. (2016)</bibRefCitation>
|
||||
<bibRefCitation id="BA3FA548BA402D2EE7846EE0BD7E400D" author="Tadokoro, T" journalOrPublisher="Lepidoptera Science" pageId="0" pageNumber="257" pagination="99 - 114" refId="B20" refString="Tadokoro, T, Inomata, T, Wang, M, 2016. Taxonomic review of the Pieris dubernardi -group, with description of a new subspecies from Northern Sichuan, China (Lepidoptera, Pieridae). Lepidoptera Science 67 (3-4): 99 - 114" title="Taxonomic review of the Pieris dubernardi - group, with description of a new subspecies from Northern Sichuan, China (Lepidoptera, Pieridae)." volume="67" year="2016">Tadokoro et al. (2016)</bibRefCitation>
|
||||
:
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="257" rank="order">Lepidoptera</taxonomicName>
|
||||
<taxonomicName id="600BA127115FE621DC8A618A6E523BA4" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
|
||||
Science 67: 99-114
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris chumbiensis subsp. gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="chumbiensis" subSpecies="gyantsensis">Pieris chumbiensis gyantsensis</taxonomicName>
|
||||
<paragraph id="B49AC2CFB3BED4789CDA3C0546BB64CE" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="9FB9861A42686F672A2BF7D2EB469649" authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris chumbiensis subsp. gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="chumbiensis" subSpecies="gyantsensis">Pieris chumbiensis gyantsensis</taxonomicName>
|
||||
; Huang (2019): Neue Entomologische Nachrichten 77: 203-255 (pictured, keyed)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="description">
|
||||
<paragraph pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="F4E27D0890B78B82065BC434C735D57E" pageId="0" pageNumber="257" type="description">
|
||||
<paragraph id="DA88AAC49379AD14287D223EFFD279B8" pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph id="0C93331F64FFEA259E4100969647651F" pageId="0" pageNumber="257">
|
||||
Medium size. Both wings whitish on the upperside while yellowish on the underside. Male (Fig.
|
||||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13G</figureCitation>
|
||||
<figureCitation id="401850E2B47C3BA68BAAD2142E9C6446" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13G</figureCitation>
|
||||
): apical spot blackish, triangular extending along veins to outer margin with a trend of merging. A short black band at the end of the discocell. The 1st spot distinct and oval-shaped, with a trend of a spot faintly occurs on the cell between vein CuA1 and CuA2; outer spot less faintly present on the upperside. All veins of hindwings heavily brownish powdered on the underside, with a brownish sub-outer marginal band. Female (Fig.
|
||||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13H</figureCitation>
|
||||
<figureCitation id="8D3D766F674BDB0A3182FC7FDA904326" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13H</figureCitation>
|
||||
): resembles male but with black markings more developed. The 1st spot distinctly developed and extends as a black band reaching the posterior margin. Hindwings with blackish sub-marginal band distinctly developed on both sides.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Tibet (from Lhasa to Nyalam)</paragraph>
|
||||
<subSubSection id="25B977719C1F7348BFBACAC2D5980C21" pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph id="CAAC02B51A74BDB7BDF72021164B1C68" pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph id="115FD3893EEDC5FD2AAD280FD7918A23" pageId="0" pageNumber="257">Tibet (from Lhasa to Nyalam)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Univoltine, occurs in June and July.</paragraph>
|
||||
<subSubSection id="11EFCC4F12A08474703F6C4D6D3ED0EA" pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph id="CDFFFE1540893AC4F6F77169D0D9F954" pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph id="78D87B5C099C029969E437CD54C578CB" pageId="0" pageNumber="257">Univoltine, occurs in June and July.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="CDAFB67C2D334AB8BBDAC5EFD514D6D3" pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph id="CBEE162CF8CC92CE99312E598EA4BD76" pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph id="84AE5D2C575C92BF39F95A977E5F7A8F" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5Q</figureCitation>
|
||||
<figureCitation id="1FD37C20C5A41E5B77C04239CBB92016" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5Q</figureCitation>
|
||||
) tegumen broad with its distal margin slightly narrower; basal margin of uncus slightly convex with a moderate concave between tegumen and uncus; uncus with its median part slightly convex, apical half of uncus digitation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="E2666707864D82B2A1221B854AC561B8" pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph id="CB2491EE4DBFCE75C4BC8184937B064D" pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph id="4D65CABAE2EA73131443817B850B43FF" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of species similar to P. napi - complex distributed in China. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure8" httpUri="https://binary.pensoft.net/fig/825203" pageId="0" pageNumber="257">8C</figureCitation>
|
||||
<figureCitation id="CDC54041DF88F0928FD000084D50A634" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of species similar to P. napi - complex distributed in China. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure8" httpUri="https://binary.pensoft.net/fig/825203" pageId="0" pageNumber="257">8C</figureCitation>
|
||||
) posterior apophysis long and slender, extended beyond the 8th tergum; inner distal of sterigma lobe shaped extend to center; inner basal of sterigma without distinct convex, smoothly connected at the basal margin; signum short and broad, almost disk shaped with smooth and shallowly concave on the terminal margin; basal part of signum tubular with distinct bending on the most basal part.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="C740A8F0CF80CA40609EEC7035CB2C90" pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph id="EAA2A5C76C74D6C229D746D388B12A98" pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph id="1D77B06E97AB1A2C1D271452D9EC0FD6" pageId="0" pageNumber="257">
|
||||
The taxon
|
||||
<taxonomicName authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="gyantsensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris gyantsensis</emphasis>
|
||||
<taxonomicName id="4C43FF1525535FA1A71A60DFDB669703" authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="gyantsensis">
|
||||
<emphasis id="F8ED1D9A01D3BD0AD030FA64E12EA880" italics="true" pageId="0" pageNumber="257">Pieris gyantsensis</emphasis>
|
||||
</taxonomicName>
|
||||
was originally published by Verity (1911) as a quadrinomial race belonging to
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. chumbiensis" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="chumbiensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. chum-biensis</emphasis>
|
||||
<taxonomicName id="24A80D0F598CC274579A9F518C8D033A" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. chumbiensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="chumbiensis">
|
||||
<emphasis id="7F4C1C02715416FBDD1AEE8056D1FA16" italics="true" pageId="0" pageNumber="257">P. chum-biensis</emphasis>
|
||||
</taxonomicName>
|
||||
, thus unavailable due to being infrasub-specific. When
|
||||
<bibRefCitation author="Bollow, C" journalOrPublisher="In Seitz A (Ed) Alfred Kernen, Stuttgart" pageId="0" pageNumber="257" refId="B5" refString="Bollow, C, 1930. 1930-1932: The Macrolepidoptera of the World, supplement to Vol. 1 - the Palaearctic butterflies. In Seitz A (Ed) Alfred Kernen, Stuttgart" title="1930 - 1932: The Macrolepidoptera of the World, supplement to Vol. 1 - the Palaearctic butterflies." year="1930">Bollow (1930)</bibRefCitation>
|
||||
<bibRefCitation id="C34EE5AA72DE4D35DE35221D0CE76128" author="Bollow, C" journalOrPublisher="In Seitz A (Ed) Alfred Kernen, Stuttgart" pageId="0" pageNumber="257" refId="B5" refString="Bollow, C, 1930. 1930-1932: The Macrolepidoptera of the World, supplement to Vol. 1 - the Palaearctic butterflies. In Seitz A (Ed) Alfred Kernen, Stuttgart" title="1930 - 1932: The Macrolepidoptera of the World, supplement to Vol. 1 - the Palaearctic butterflies." year="1930">Bollow (1930)</bibRefCitation>
|
||||
re-defined this taxon as a subspecies of
|
||||
<taxonomicName class="Insecta" family="Pieridae" genus="Synchloe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synchloe chumbiensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="chumbiensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Synchloe chumbiensis</emphasis>
|
||||
<taxonomicName id="03CC2F97B8A8C5A6AB4460498E7023C4" class="Insecta" family="Pieridae" genus="Synchloe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synchloe chumbiensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="chumbiensis">
|
||||
<emphasis id="2EC671D69627B42827E86F5C3E872B6F" italics="true" pageId="0" pageNumber="257">Synchloe chumbiensis</emphasis>
|
||||
</taxonomicName>
|
||||
, the subspecies name became an available name. This taxon was once mistreated as belonging to the
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
<taxonomicName id="FB64014AE466C157873D95B275E72934" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="F8FD9B610550596F4A1E65697885A87C" italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
</taxonomicName>
|
||||
-complex by
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="039E1349FE5FFCAFA32E177A163C2971" author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
according to their phylogenetic results and observations on androconial characteristics. Our molecular results oppositely showed that this taxon constitues a distinct molecular lineage separated from the
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
<taxonomicName id="3E4364A5B623A756454698E9D8436D94" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="414A5E2D3BA988B7F7BC92E50AFCE762" italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
</taxonomicName>
|
||||
-complex. Moreover, this taxon has long been considered as a member of the
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dubernardi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dubernardi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dubernardi</emphasis>
|
||||
<taxonomicName id="BC10080C44353F3C5CE5AE5487C58D7D" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dubernardi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dubernardi">
|
||||
<emphasis id="22CFBDAE6D8D18B3DFB253FBAF9F9A9B" italics="true" pageId="0" pageNumber="257">P. dubernardi</emphasis>
|
||||
</taxonomicName>
|
||||
-group, a unique species group with distinctive morphological characteristics and a high altitude habitat which includes
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. dubernardi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="dubernardi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. dubernardi</emphasis>
|
||||
<taxonomicName id="3F10AAEE20E9705EADE9B0B01898C36F" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dubernardi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="dubernardi">
|
||||
<emphasis id="018708AECCD015173627653D6DE73090" italics="true" pageId="0" pageNumber="257">P. dubernardi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. rothschildi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="rothschildi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. rothschildi</emphasis>
|
||||
<taxonomicName id="4D230105D30E5BCB776A17FC1CEC8040" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. rothschildi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="rothschildi">
|
||||
<emphasis id="3F7BE36AD113EA394B06A92E12ED932A" italics="true" pageId="0" pageNumber="257">P. rothschildi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. wangi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="wangi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. wangi</emphasis>
|
||||
<taxonomicName id="719B73E7EEE1C3A759A4C294FB0B7B6B" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. wangi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="wangi">
|
||||
<emphasis id="884F9056BD0EAFB438E54787BEA2C45C" italics="true" pageId="0" pageNumber="257">P. wangi</emphasis>
|
||||
</taxonomicName>
|
||||
as well as
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. chumbiensis" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="chumbiensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. chumbiensis</emphasis>
|
||||
<taxonomicName id="3E9FF924CE8B331CC407CB7C73702822" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. chumbiensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="chumbiensis">
|
||||
<emphasis id="F76FEC03E9CC4C1403CE453D68B04AF2" italics="true" pageId="0" pageNumber="257">P. chumbiensis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,147 +1,147 @@
|
|||
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<treatment ID-GBIF-Taxon="158995545" LSID="urn:lsid:plazi:treatment:45AB55AB28805B229F01D47FDED687E7" httpUri="http://treatment.plazi.org/id/45AB55AB28805B229F01D47FDED687E7" lastPageNumber="19" pageId="0" pageNumber="19">
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<treatment id="45AB55AB28805B229F01D47FDED687E7" ID-GBIF-Taxon="158995545" LSID="urn:lsid:plazi:treatment:45AB55AB28805B229F01D47FDED687E7" httpUri="http://treatment.plazi.org/id/45AB55AB28805B229F01D47FDED687E7" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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Madaglymbus unguicularis (
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, 1903)
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<taxonomicNameLabel pageId="0" pageNumber="19">comb. nov.</taxonomicNameLabel>
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<taxonomicNameLabel id="0E39F3BAB96AD4B6A9864DDEFE7D4857" pageId="0" pageNumber="19">comb. nov.</taxonomicNameLabel>
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<taxonomicName id="BAB6DA12CF120500EBA36676F3A48D52" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus unguicularis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="unguicularis">Copelatus unguicularis</taxonomicName>
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, 1903:19.
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<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="19">Madagascar, Suberbieville [= Maevatanana].</paragraph>
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<paragraph id="FBBB719660FADCE75A4C7B539A94F0EC" pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph id="7BDC11120B1A1D5FB960776747C9B13E" pageId="0" pageNumber="19">Madagascar, Suberbieville [= Maevatanana].</paragraph>
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<paragraph pageId="0" pageNumber="19">Type material examined.</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Betsiboka: Maevatanana</emphasis>
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<paragraph id="30F7C8802CB88269B721E423BFE2FD62" pageId="0" pageNumber="19">Type material examined.</paragraph>
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<emphasis id="4FF5F401D170C60BC118B07EAEDED13B" bold="true" pageId="0" pageNumber="19">Mahajanga. Betsiboka: Maevatanana</emphasis>
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: -HT ♂ (GP) (MNHN, "coll.
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): // Data in NHRS | JLKB | 000030225 // Suberbieville | Madag. Perier // Type [red label] // MUSEUM PARIS | Coll. Maurice
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<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
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| 1908 //
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<emphasis italics="true" pageId="0" pageNumber="19">C. unguicularis</emphasis>
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<emphasis id="D4F12A567151C89534F18D8C8EC8D025" italics="true" pageId="0" pageNumber="19">C. unguicularis</emphasis>
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| Type //
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<emphasis italics="true" pageId="0" pageNumber="19">unguicularis</emphasis>
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. //
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<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus unguicularis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="unguicularis">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus unguicularis</emphasis>
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<paragraph id="05132FC506F0278EE3FFA53A0EBEE511" pageId="0" pageNumber="19">Remarks.</paragraph>
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<taxonomicName id="41ED1CB8BF9E2F820BFCE88D8C96F956" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus unguicularis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="unguicularis">
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<emphasis id="0257448ACF9AAC6CDB7A885663E80092" italics="true" pageId="0" pageNumber="19">Copelatus unguicularis</emphasis>
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</taxonomicName>
|
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was described by
|
||||
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="19" pagination="1 - 51" refId="B64" refString="Regimbart, M, 1903. Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le sud de Madagascar par M. Ch. Alluaud (juillet 1900-mai 1901). . Annales de la Societe Entomologique de France 72: 1 - 51" title="Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le sud de Madagascar par M. Ch. Alluaud (juillet 1900 - mai 1901)." volume="72" year="1903">
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<bibRefCitation id="39B0F5FFDB16A2AAA3E9D5146C2857A9" author="Regimbart, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="19" pagination="1 - 51" refId="B64" refString="Regimbart, M, 1903. Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le sud de Madagascar par M. Ch. Alluaud (juillet 1900-mai 1901). . Annales de la Societe Entomologique de France 72: 1 - 51" title="Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le sud de Madagascar par M. Ch. Alluaud (juillet 1900 - mai 1901)." volume="72" year="1903">
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(1903)
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, based on a single male from Suberbieville [= Maevatanana]. It was classified by
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<normalizedToken id="0F9F02549EEA0541E9B571224716BD75" originalValue="Régimbart">Regimbart</normalizedToken>
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in the fifth species group of
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<bibRefCitation pageId="0" pageNumber="19" refId="B68">Sharp (1882)</bibRefCitation>
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<bibRefCitation id="8CF9BC4F5DB6964A51B80C1422286274" pageId="0" pageNumber="19" refId="B68">Sharp (1882)</bibRefCitation>
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(the
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<emphasis italics="true" pageId="0" pageNumber="19">consors</emphasis>
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<emphasis id="6DE7CE5983900DD68F100CBCD9A69CE4" italics="true" pageId="0" pageNumber="19">consors</emphasis>
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species group sensu
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<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
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<bibRefCitation id="E85C1F3C068BD7C4B027A5D29E4D7977" pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
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) based on ten discal but no submarginal elytral stria where it is still classified today (
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<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
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<bibRefCitation id="38674127696835BA6334893586A31C67" pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
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;
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||||
<bibRefCitation author="Gueorguiev, VB" journalOrPublisher="Izvestija na Zoologitjeskija Instituts Musei Sofia" pageId="0" pageNumber="19" pagination="5 - 45" refId="B25" refString="Gueorguiev, VB, 1968. Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae). . Izvestija na Zoologitjeskija Instituts Musei Sofia 28: 5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
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<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
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<bibRefCitation id="4D133287CD0B3754E6BACD60B2F81DED" author="Gueorguiev, VB" journalOrPublisher="Izvestija na Zoologitjeskija Instituts Musei Sofia" pageId="0" pageNumber="19" pagination="5 - 45" refId="B25" refString="Gueorguiev, VB, 1968. Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae). . Izvestija na Zoologitjeskija Instituts Musei Sofia 28: 5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
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1968
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</bibRefCitation>
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;
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||||
<bibRefCitation DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
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<bibRefCitation id="E07F0C14EB48A01AB9A508431E539820" DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="19" refId="B55" refString="Nilsson, AN, 2001. World catalogue of insects. Volume 3. Dytiscidae (Coleoptera). . Apollo Books, Stenstrup" title="World catalogue of insects. Volume 3. Dytiscidae (Coleoptera)." year="2001">Nilsson 2001</bibRefCitation>
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<bibRefCitation id="FE3490C6A0336990BE1E248DCA69B17B" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="19" refId="B55" refString="Nilsson, AN, 2001. World catalogue of insects. Volume 3. Dytiscidae (Coleoptera). . Apollo Books, Stenstrup" title="World catalogue of insects. Volume 3. Dytiscidae (Coleoptera)." year="2001">Nilsson 2001</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" publicationUrl="http://www.waterbeetles.eu" refId="B57" refString="Nilsson, AN, Hajek, J, 2018. . http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2018">
|
||||
<bibRefCitation id="10C4E0ABAAFA3270083252550F91150A" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" publicationUrl="http://www.waterbeetles.eu" refId="B57" refString="Nilsson, AN, Hajek, J, 2018. . http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2018">
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Nilsson and
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
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<normalizedToken id="4EC9C9CFCB18DA9064ECF64E1793A56F" originalValue="Hájek">Hajek</normalizedToken>
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2018
|
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|
||||
). We have examined the male holotype and like the type of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus apicalis</emphasis>
|
||||
<taxonomicName id="A663A0F8BFDE56AF48C3A972DB911631" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">
|
||||
<emphasis id="CFA7CFA18F392C06D12DB42719C10934" italics="true" pageId="0" pageNumber="19">Copelatus apicalis</emphasis>
|
||||
</taxonomicName>
|
||||
, it has a stout spine-like setae on a protruding anterodistal corner of protarsomere IV and is not a
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="46461A3A2848AE8FE055AF028391A8C7" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F7125CE631916536C1F28032E73C80AF" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
. In addition, the elytral striae are very irregular, more like very elongate and deep strioles and possibly not homologous to the regular impressed striae in
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="44DFD0EBBB56636626E5259A585CDFFD" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3FBD49838A75153E51AA85490FF607B1" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
. We here transfer it to
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
<taxonomicName id="91CD770A64E6C6D7AD997A5E64F8EBB0" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4A234232A832AC190FDE3D6BD878267C" italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus unguicularis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="unguicularis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus unguicularis</emphasis>
|
||||
<taxonomicName id="2F1F40959214A3BBC2999909D75D1A6F" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus unguicularis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="unguicularis">
|
||||
<emphasis id="83D0D8BB6C4FF513B4685144A5D5C0D9" italics="true" pageId="0" pageNumber="19">Madaglymbus unguicularis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<normalizedToken id="840F93623F392E9B7D000BEAA0C5AC48" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
, 1903) comb. nov.
|
||||
</paragraph>
|
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</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,260 +1,260 @@
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<treatment ID-GBIF-Taxon="158995554" LSID="urn:lsid:plazi:treatment:48693E4733015386A1417218CDEAFB7A" httpUri="http://treatment.plazi.org/id/48693E4733015386A1417218CDEAFB7A" lastPageNumber="19" pageId="0" pageNumber="19">
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<treatment id="48693E4733015386A1417218CDEAFB7A" ID-GBIF-Taxon="158995554" LSID="urn:lsid:plazi:treatment:48693E4733015386A1417218CDEAFB7A" httpUri="http://treatment.plazi.org/id/48693E4733015386A1417218CDEAFB7A" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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Copelatus distinguendus
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<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
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, 1903
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Figs 4B</figureCitation>
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<figureCitation id="C114EA2594DB171574E7066CC6877BC3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Figs 4B</figureCitation>
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,
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">7D</figureCitation>
|
||||
<figureCitation id="9D68800342C695925A0DC1D8DD9C4118" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">7D</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">Copelatus distinguendus</taxonomicName>
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<subSubSection id="E85FBA5B70CB78C2C0553E00122A3938" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="68E75A149C952260B0AC28A4D7A42609" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="AA86554E88FE2C70E21C420D234707FE" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">Copelatus distinguendus</taxonomicName>
|
||||
<normalizedToken id="58E50914802E803CC7D0DB7614536C2A" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
, 1903: 19 [nom. nov., referring to his description of Malagasy material under the name
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus duodecimstriatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="duodecimstriatus">Copelatus duodecimstriatus</taxonomicName>
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<taxonomicName id="6527A848B3FA89DF392E30673A5B0801" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus duodecimstriatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="duodecimstriatus">Copelatus duodecimstriatus</taxonomicName>
|
||||
<normalizedToken id="CAD2E0C9704C7B550F9616436F24586B" originalValue="Aubé">Aube</normalizedToken>
|
||||
in
|
||||
<bibRefCitation author="Regimbart, M" journalOrPublisher="Memoires de la Societe Entomologique de Belgique" pageId="0" pageNumber="19" pagination="1 - 244" refId="B63" refString="Regimbart, M, 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. . Memoires de la Societe Entomologique de Belgique 4: 1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines." volume="4" year="1895">
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<bibRefCitation id="1065F8F2F6147BFF2EE05D8A29CF78AE" author="Regimbart, M" journalOrPublisher="Memoires de la Societe Entomologique de Belgique" pageId="0" pageNumber="19" pagination="1 - 244" refId="B63" refString="Regimbart, M, 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. . Memoires de la Societe Entomologique de Belgique 4: 1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines." volume="4" year="1895">
|
||||
<normalizedToken id="56F20425CF3CF61BAA9497C2ED770E50" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
1895
|
||||
</bibRefCitation>
|
||||
: 163]
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Environs de Tananarive [surroundings of Antananarivo] and Fianarantsoa, Madagascar.</paragraph>
|
||||
<subSubSection id="774EA2EB4F8447A1D96443711EC2F21C" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="5F34C4EE4E007C954788F0C404873951" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="1D5BC11F69C7E1DC1238DA5FE3255EA1" pageId="0" pageNumber="19">Environs de Tananarive [surroundings of Antananarivo] and Fianarantsoa, Madagascar.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Based on an unknown number of specimens (syntypes) collected by Sikora (Antananarivo) and Perrot (Fianarantsoa).</paragraph>
|
||||
<subSubSection id="09916A20FD580A2AB36B4F7BB2DBABBA" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="CCE18264BD205B5A1D5FCA4EE26FFF86" pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph id="0F97F665BEB11B06D596F41959892FB8" pageId="0" pageNumber="19">Based on an unknown number of specimens (syntypes) collected by Sikora (Antananarivo) and Perrot (Fianarantsoa).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Type material in MNHN not studied, as it was out on loan.</paragraph>
|
||||
<subSubSection id="DF7952C1F87B11430C9CFD91000D20FC" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="67533735630C92806B862CB4137C86C3" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="D0B7C78624CC018E9046C0F9D3567BC0" pageId="0" pageNumber="19">Type material in MNHN not studied, as it was out on loan.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Fianarantsoa. Matsiatra Ambony: Ambalavao, Ambohimahasoa, Lalangina</emphasis>
|
||||
<subSubSection id="69115B4271491AC371B38EED4D8363FD" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="62F6ED7104FC9948245DF42F53BDC99B" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="CCB60D1D6698078AB92604D2E3022535" pageId="0" pageNumber="19">
|
||||
<emphasis id="95344FCE56DDCBCF2E9A13208605C2E3" bold="true" pageId="0" pageNumber="19">Fianarantsoa. Matsiatra Ambony: Ambalavao, Ambohimahasoa, Lalangina</emphasis>
|
||||
: -2♀, 1♂(GP) (NHMUK): // BMNH-792954-6 // MAD: FIAN: Andringitra | Zomandao R.: River edge: Bottle trap | P39EM08: N: -22.1043: E:46.92: 1420 m | 09/V/2006: Leg. Bergsten et al // BMNH (DNA Voucher) // -1♀ (NMW): // Data in NHRS | JLKB | 000010718 // Madagascar: Ambohimahasoa (Fianarantsoa) | RN7 (Km 378) | 16.04.2011: Leg. R. Gerecke (MD211) // spring area with meadow swamps Exp. E:
|
||||
<geoCoordinate degrees="21" direction="south" minutes="15" orientation="latitude" precision="1" seconds="41.5" value="-21.261528">21°15'41.5"S</geoCoordinate>
|
||||
<geoCoordinate id="73A89ED2B307F9CA9E87365B6D6555B4" degrees="21" direction="south" minutes="15" orientation="latitude" precision="1" seconds="41.5" value="-21.261528">21°15'41.5"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="47" direction="east" minutes="14" orientation="longitude" precision="1" seconds="10.9" value="47.236362">47°14'10.9"E</geoCoordinate>
|
||||
<geoCoordinate id="B4B656B9288490B7692B91A1786CBA8F" degrees="47" direction="east" minutes="14" orientation="longitude" precision="1" seconds="10.9" value="47.236362">47°14'10.9"E</geoCoordinate>
|
||||
, 1500 m // -2♀ (NMW): // Data in NHRS | JLKB | 000065754-5 // Madagascar Est, 1100- | 1200m, P.N. Ranomafana // Vohiparara, 21.-24.1.1993 | J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="8D0863A791EF9010F5D36EB880C18C7F" originalValue="Janák">Janak</normalizedToken>
|
||||
lgt // -1♂(GP), 1♀, 5 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010618-9, 10787(Alc.) // Madagascar: Fianarantsoa: Matsiatra | Ambony: Ranomafana NP: | Sahamalaotra 2Km from Vohiparara: | S21.23807, E047.39489, 1140 m.| 01:XI:2011: stamping with sieves: forest | bog in rainforest: Field# MAD11-12 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♀(Alc.) (NHRS): // NHRS-JLKB | 000010830 // MAD: FIAN: Matsiatra Ambony | Ranomafana NP: 450m along | Sahamalaotra trail, left at the first | junction: Mad14-07: forestmarsh: |
|
||||
<geoCoordinate degrees="21.2382" direction="south" orientation="latitude" precision="5" value="-21.2382">21.2382S</geoCoordinate>
|
||||
<geoCoordinate degrees="49.3947" direction="east" orientation="longitude" precision="5" value="49.3947">49.3947E</geoCoordinate>
|
||||
<geoCoordinate id="E4656E9FC3852FCC23F4F4CE372C3912" degrees="21.2382" direction="south" orientation="latitude" precision="5" value="-21.2382">21.2382S</geoCoordinate>
|
||||
<geoCoordinate id="EFBC465BE78C6527A92A877612378FA0" degrees="49.3947" direction="east" orientation="longitude" precision="5" value="49.3947">49.3947E</geoCoordinate>
|
||||
: 1130 m: 02.XI.2014 // Leg. J. Bergsten, | T. Ranarilalatiana | & S. Holmgren // -1♀ (NHMUK): // BMNH-670601_MSL007 | 06.xii.2004, Ranomafana, | Madagascar: lat - 21.2359 | Lon 47.3963 Coll Balke_M; | Monaghan_M // DNA Voucher | BMNH <670601> | MSL007:E07 //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">
|
||||
<emphasis id="AF3CCC6DE8DFF3F956FEBE274B8B1425" bold="true" pageId="0" pageNumber="19">
|
||||
Fianarantsoa.
|
||||
<normalizedToken originalValue="Amoron’i">Amoron'i</normalizedToken>
|
||||
<normalizedToken id="D3533210FD7BC4E6C1C53C8271F63EEB" originalValue="Amoron’i">Amoron'i</normalizedToken>
|
||||
Mania: Ambositra
|
||||
</emphasis>
|
||||
: -3♀ (NHMUK): // BMNH-792962-4 // Col de Tapias: Rte
|
||||
<normalizedToken originalValue="Tana–Fianarantsoa">Tana-Fianarantsoa</normalizedToken>
|
||||
<normalizedToken id="135CECAB6E815060CD395EC87B985FE0" originalValue="Tana–Fianarantsoa">Tana-Fianarantsoa</normalizedToken>
|
||||
: Pond | P36C: N: -20.772: E:47.179: 1717 m | 06/V/2006: Leg. Bergsten et al // BMNH (DNA Voucher) // -1♀ (NHMUK): // BMNH-792912 // Ambositra: Ankazomivady forest | 01.xii.2005 // BMNH (DNA Voucher) // -9♀ (NHMUK), 8 ex. (Alc.) (NHRS): // BMNH-729890, 729893, 729896-7, 792976-80, 10793(Alc.) // 08.xii.2004, Col de Tapias, | P30MD33: lat -20.238 | Lon 47.1 Coll Balke_M | Monaghan_M // BMNH (DNA Voucher) //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antananarivo. Vakinankaratra: Ambatolampy</emphasis>
|
||||
<emphasis id="75148EAD046D495C90C2763E904B188E" bold="true" pageId="0" pageNumber="19">Antananarivo. Vakinankaratra: Ambatolampy</emphasis>
|
||||
: -3♂(GP), 3♀, 10 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010620-5, 10786(Alc.) // Madagascar: Antananarivo: | Vakinankaratra: Manjakatompo Stn. |
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="7D5E289B9E7CACC3898C18966B9FAC3A" originalValue="forestière">forestiere</normalizedToken>
|
||||
: 500m E Lac Froid by the | road: S
|
||||
<geoCoordinate degrees="19.34485" direction="east" orientation="longitude" precision="1" value="19.34485">19.34485 E</geoCoordinate>
|
||||
<geoCoordinate id="B0FF1C8E37353C15607525FFC0692C3E" degrees="19.34485" direction="east" orientation="longitude" precision="1" value="19.34485">19.34485 E</geoCoordinate>
|
||||
047.33381, 1620 m. | 04.XI.2011: GB Nets and sieves: pond | and inlet stream: MAD11-16 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♂(GP) (NHRS): // NHRS-JLKB | 000010626 // Madagascar: Antananarivo: | Vakinankaratra: Manjakatompo Stn. |
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="FA05AEFED77265FF886A378CC11ADA24" originalValue="forestière">forestiere</normalizedToken>
|
||||
: Analafandriana 500 m S | fish farm by the road: S19.36191 E | 47.31495, 1730 m, 03.XI.2011: GB | Nets: grassy pond: Field# MAD11-14 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♂(GP) (NHRS): // NHRS-JLKB | 000010660 // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-03: | "Lac froid": S-19.34292; E047.33893; | 1651 m: lake with grass at margins: | 03/02/2016 Leg. T. Ranarilalatiana // -4♂(GP), 4♀, 5 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010662-9, 10791(Alc.) // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-46: | "Lac froid": S-19.34292; E047.33893; | 1651 m: lake with grass at margins: | 17/09/2016 Leg. T. Ranarilalatiana // -1♂(GP) (NHRS): // NHRS-JLKB | 000010661 // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-10: | Ankafotra mountain: S-19.33753; | E047.24530; 2466 m: streampools: | 07/02/2016; Leg. T. Ranarilalatiana // -4♂(GP), 2♀, 18 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010670-5, 10792(Alc.) // Madagascar: Ambatolampy: Manjaka- | tompo Ankaratra Reserve: MAD16-47: | Ankafotra mountain: S-19.33753; | E047.24530; 2466 m: streampools: | 18/09/2016; Leg. T. Ranarilalatiana // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010832 // Madagascar: Antananarivo: | Vakinankaratra: Manjakatompo Stn |
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="DE83C38121CB928802D45F8B4A197F49" originalValue="forestière">forestiere</normalizedToken>
|
||||
: Analamitana: S19.363972 E | 047.299083, 1757 m. 22:I:2012: swamp | near stream: Field# MJK12-02: Leg. T. | Ranarilalatiana & J.H. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antananarivo. Analamanga: Anjozorobe, Ankazobe</emphasis>
|
||||
<emphasis id="6214638CFCCF5A0C754668DFD77D768A" bold="true" pageId="0" pageNumber="19">Antananarivo. Analamanga: Anjozorobe, Ankazobe</emphasis>
|
||||
: -1♂(GP), 1♀ (NHRS): // NHRS-JLKB | 000010684-5 // Madagascar: Anjozorobe: MAD16-36: | Amboasarianala: S-18.45792; E047. | 93438; 1367 m: Ambatovikinina stream: | 04/04/2016; Leg. T. Ranarilalatiana // -1♂(GP), 1♀ (NHRS): // NHRS-JLKB | 000010686-7 // Madagascar: Anjozorobe: MAD16-43:| Amboasarianala, Antanambe stream: | S-18.4671; E047.93807; 1271 m: Stream | with sidepools: 07/04/2016 | Leg. T. Ranarilalatiana // -2♂(GP), 2♀, 1♀(Alc.) (NHRS): // NHRS-JLKB | 000010688-91, 10790(Alc.) // Madagascar: Anjozorobe: MAD16-44 | Amboasarianala, Mangarivotra stream: | S-18.4676; E047.92535; 1271 m: stream | with bedrock and grass at edge | 07/04/2016; Leg. T. Ranarilalatiana // -2 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010831 // MAD: ANTA: Analamanga: Anjoz | orobe forest reserve: Marsh next | to the stream by Saha forest, 10Km E of Anjozorobe: MAD14- | 78: forestmarsh: 18.4128S | 47.9439E; 1320 m; 23.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -2♂(GP), 2♀, 6 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010676-9, 10797(Alc.) // Madagascar: Ankazobe: MAD16-24: | Firarazana: S-18.13132; E047.23976; | 1551 m; Lake with grass at margins: | 12/03/2016; Leg. T. Ranarilalatiana // -2♀, 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010680-1, 10789(Alc.) // Madagascar: Ankazobe: MAD16-26: | Maharidaza, Large stream by the road to | military camp: S-18.22102; E047.27087; | 1547 m: stream and bog with grass: | 14/03/2016; Leg. T. Ranarilalatiana // -1♂(GP), 1♀, 7 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010682-3, 10788(Alc.) // Madagascar: Ankazobe: MAD16-29: | Firarazana, SW of Ambohitantely | reserve: S-18.16717; E047.26090; | 1532 m: Bog with grass: 17/03/2016; | Leg. T. Ranarilalatiana // -3 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010829 // MAD: ANTA: Analamanga: Mana- | nkazo river by the bridge of | RN4: Mad14-75: medium size river over bedrock: 18.158S | 47.2104E: 1450 m: 21.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -14 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010834 // MAD: ANTA: Analamanga: Andra- | nofeno river by the bridge of | RN4, next to Andranofeno Sud | village: Mad14-74: medium size, | slow flowing river: 18.0844S | 47.1776E: 1430 m: 21.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // - 1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010835 // Madagascar: Ankazobe: TR18L10: | Stream by the bridge S of Ambohitantely | reserve: S-18.2023; E047.2780; 1556 m: | Hygropetric rock and stagnant pools: | 11/04/2018 | Leg. T. Ranarilalatiana -1♀ (NMW): // Data in NHRS | JLKB | 000065756 // RM: Betsiboka Bas (PO533) | Andranofeno Sud Riv. |
|
||||
<geoCoordinate degrees="47" direction="east" minutes="10" orientation="longitude" precision="15" seconds="46" value="47.179447">47°10'46"E</geoCoordinate>
|
||||
<geoCoordinate id="DB2D0D0769DBE10AA6D52795D446A448" degrees="47" direction="east" minutes="10" orientation="longitude" precision="15" seconds="46" value="47.179447">47°10'46"E</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="18" direction="south" minutes="05" orientation="latitude" precision="15" seconds="00" value="-18.083334">18°05'00"S</geoCoordinate>
|
||||
<geoCoordinate id="3A0CCBD3C7F88A96350EC34876BCB2DE" degrees="18" direction="south" minutes="05" orientation="latitude" precision="15" seconds="00" value="-18.083334">18°05'00"S</geoCoordinate>
|
||||
| 06.11.1995 | Leg. Elouard, J.-M., Oliarinony. R. //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Ambatondrazaka, Andilamena, Moramanga</emphasis>
|
||||
<emphasis id="C461776B6F21ABA1913F4BA3031950EB" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Ambatondrazaka, Andilamena, Moramanga</emphasis>
|
||||
: -1♀ (NMW): // Data in NHRS | JLKB | 000065759 // E-Madagascar (09) Ambaton- | drazaka Region, 5Km N Didy | 1100-1200 m. asl. 14-16.01.1995 | G. Dunay & J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="023FDC3A7085BB680308FD79FFCB464A" originalValue="Janák">Janak</normalizedToken>
|
||||
coll. // -1♀ (NMW): // Data in NHRS | JLKB | 000065760 // E-Madagascar (10) | Ambatombe, near Andilamena | 900 m asl. 17.01.1995 | G. Dunay & J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="93E2813196A985529E67AC650AB59CCE" originalValue="Janák">Janak</normalizedToken>
|
||||
coll. // -1♂ (NMW): // Data in NHRS | JLKB | 000065761 // E-Madagascar (11) Ampamoho | near Andilamena, 1200-1300 m. | asl. 18-20.01.1995 | G. Dunay & J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="9B3587E447A257CA0199B5E18334A3A5" originalValue="Janák">Janak</normalizedToken>
|
||||
coll. // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010627 // MAD: TOAM: Alaotra Mangoro | Betsabora river by RN2 near | Antsapanana village, 6 Km W | of Moramanga: MAD14-81: river | with side pools: 18.9247S | 48.1828E; 900 m; 24.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♂(Alc.) (NHRS): // NHRS-JLKB | 000010833 // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra SR: bog | at S border of reserve: S18.95456 E | 048.44048, 910 m: 09.XI.2011: GB | Nets and sieves: bog with red mud: | Field# MAD11-27 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♀ (NHMUK): // BMNH-677103 // 11.xii.2004, Andasibe, | Madagascar: P27MD36 | Coll Balke_M; | Monaghan_M // (DNA Voucher) // -1♀ (NMW): // Data in NHRS | JLKB | 000065757 // MADAGASCAR (Md-4) | Andasibe, NP Perinet | 1150 m,
|
||||
<normalizedToken originalValue="Pfütze">Pfuetze</normalizedToken>
|
||||
<normalizedToken id="1C5C166B9DC47A07A9D75297B547FC3F" originalValue="Pfütze">Pfuetze</normalizedToken>
|
||||
auf waldwiese | und in Kleinen Bach | 7.12.2000, Leg. W. Dolin // -1♀ (NMW): // Data in NHRS | JLKB | 000065758 // E-Madagascar (07) | Andranokobaka, N Moramanga | 800 m. Asl, 13.01.1995 | G. Dunay & J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="3C585165628D8A93E88DA0D173F0EA81" originalValue="Janák">Janak</normalizedToken>
|
||||
coll. //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antsiranana. Sava: Andapa</emphasis>
|
||||
<emphasis id="C8E2731EB1CAF443306BC3402F870956" bold="true" pageId="0" pageNumber="19">Antsiranana. Sava: Andapa</emphasis>
|
||||
: -1♂ (GP), 1♂ (Alc.) (NHRS): // NHRS-JLKB | 000010827, 10828 (Alc.) // MAD: ANTS: Sava: Anjanaharibe | Sud NP: river Marolakana at the | crossing place: Mad14-64: larger | river with rocks; 14.7623S | 49.4834E: 920 m: 15.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="01DC61D5119A239D84ABC4A349F8568C" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="9A42EAFED07EBCDA0AB6D309792D6098" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="D653FE86B318741B10C8B555EFEE0A09" pageId="0" pageNumber="19">
|
||||
Body shape elongate oval, convex, and attenuate posteriorly, with uniform black colouration (
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7D</figureCitation>
|
||||
<figureCitation id="BAF852434168129D2DD999B3154EA99C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7D</figureCitation>
|
||||
). The pattern of abbreviation of the elytral striae separates this species from all other Malagasy
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="80F0967799232A5E79CAEF5049FFE03F" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="39A64DB075D6632724EBA99ED0F73D4F" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
species. Striae 1, 3, and 5 present only in posterior one third (1) or two thirds (3, 5); striae 2, 4, and 6 fragmented anteriorly and, except for stria 4, never reach the base as clearly impressed striae. Penis profile in lateral view is characteristic, resembling a
|
||||
<normalizedToken originalValue="“pumpjack”">"pumpjack"</normalizedToken>
|
||||
<normalizedToken id="E68A456D93027D6A9B2E77C496171ECF" originalValue="“pumpjack”">"pumpjack"</normalizedToken>
|
||||
(
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4B</figureCitation>
|
||||
<figureCitation id="ED7487EA70448C0E950B214C4CB0CC76" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Body length: 5.3-6.3 mm. Body shape elongate oval, convex, and attenuate posteriorly. Head, pronotum, and elytra all the same colour, ferrugineous black and finely punctate.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="D16DEB16FB42CA6F408631214EA496A9" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="45991719C5F5171D06F94280EF4C7EF5" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="442F9DFE807791E846C7C4E2ED456CD8" pageId="0" pageNumber="19">Body length: 5.3-6.3 mm. Body shape elongate oval, convex, and attenuate posteriorly. Head, pronotum, and elytra all the same colour, ferrugineous black and finely punctate.</paragraph>
|
||||
<paragraph id="9CED74E277AFDC303F3364A1DE7AA6B7" pageId="0" pageNumber="19">
|
||||
Lateral margin of pronotum rusty ferrugineous, with short sparse strioles. Pronotum with puncture rows and microsculpture. Elytra narrowly testaceous to ferrugineous posterolaterally. Six elytral striae present and one submarginal stria (
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7D</figureCitation>
|
||||
<figureCitation id="17E2CD67DCE33B6BBC9AF8C3E78D507F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7D</figureCitation>
|
||||
). Striae 2, 4, and 6 longer than striae 1, 3, and 5, at least as continued fragments; first stria shortest, present only in posterior fourth to posterior third. Second and third striae present as clearly impressed striae in posterior half or a little longer, but second stria continues, albeit fragmentary and less impressed, more distinctly all the way to base. Fourth stria the most complete, more or less present to base but less deeply impressed or fragmentary in anterior third. Fifth stria abbreviated a little less than third stria, present in approximately posterior two thirds, and like stria 3 has small isolated point remnants of stria more anteriorly. Sixth stria longer than fifth and with more distinct remnants of striae present anteriorly. Elytra covered with punctures in apical part.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Ventral side ferrugineous dark brown. Metacoxa and abdominal sternite II, III, IV, striolate. Prosternal process raised medially but rounded, not carinate. Lateral parts of metaventrite broad. Metacoxal lines short and strongly diverging anteriorly. Antenna, palps, pro- and mesothoracic legs brown to yellowish, but metathoracic legs dark brown.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="7449BFA0CD7F668F5B76AA4639C6B557" pageId="0" pageNumber="19">Ventral side ferrugineous dark brown. Metacoxa and abdominal sternite II, III, IV, striolate. Prosternal process raised medially but rounded, not carinate. Lateral parts of metaventrite broad. Metacoxal lines short and strongly diverging anteriorly. Antenna, palps, pro- and mesothoracic legs brown to yellowish, but metathoracic legs dark brown.</paragraph>
|
||||
<paragraph id="2657EF4E177816342A6A5D8DDA0DD788" pageId="0" pageNumber="19">
|
||||
Male: protibia slightly widened at apex, somewhat curved and angulate basally. Penis in lateral view curved with two points where curvature is more abrupt, constricted before apex at a narrow
|
||||
<normalizedToken originalValue="“neck”">"neck"</normalizedToken>
|
||||
<normalizedToken id="37FFE0176E5CB4FD07BD3E19580D3497" originalValue="“neck”">"neck"</normalizedToken>
|
||||
and expanding to apical part (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4B</figureCitation>
|
||||
<figureCitation id="C149BF11688CF949D3088A5AD74B3021" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4B</figureCitation>
|
||||
). As a whole in lateral view, the penis bears resemblance to the silhouette of a
|
||||
<normalizedToken originalValue="“pumpjack”">"pumpjack"</normalizedToken>
|
||||
<normalizedToken id="BEE14E4FA1BB80C31848532110BAE22B" originalValue="“pumpjack”">"pumpjack"</normalizedToken>
|
||||
. Penis apex in ventral view left-turned. Parameres broad at base, curved and tapering towards apex, apical part with numerous setae on ventral margin (
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4B</figureCitation>
|
||||
<figureCitation id="E3FDBB9382127DC8AE1D323BC21C0D6C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus peridinus B Copelatus distinguendus C Copelatus pseudostriatus sp. nov." figureDoi="10.3897/zookeys.869.33997.figure4" httpUri="https://binary.pensoft.net/fig/322929" pageId="0" pageNumber="19">Fig. 4B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Female: dorsal sculpture similar to male.</paragraph>
|
||||
<paragraph id="EC4F8AA585511F6F038375331D7070C9" pageId="0" pageNumber="19">Female: dorsal sculpture similar to male.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="1E56645411DF316C69BFF26D825B7D9F" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="0BC44B89C20E115C9B246336F516FDF6" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="B6D1A02D5A4F33719B172CF4EA379CAB" pageId="0" pageNumber="19">
|
||||
Occurs on Madagascar and Mauritius (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
<bibRefCitation id="BA791EDC603D73F250A4AEF977D4E3B7" pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
).
|
||||
<bibRefCitation author="Guignot, F" journalOrPublisher="Bulletin de Musee Royal d'Histoire Naturelle de Belgique" pageId="0" pageNumber="19" pagination="1 - 14" refId="B27" refString="Guignot, F, 1948. Vingt-sixieme note sur les hydrocanthares. Dytiscides et Gyrinides nouveaux ou peu connus du Congo belge. . Bulletin de Musee Royal d'Histoire Naturelle de Belgique 24: 1 - 14" title="Vingt-sixieme note sur les hydrocanthares. Dytiscides et Gyrinides nouveaux ou peu connus du Congo belge." volume="24" year="1948">Guignot (1948)</bibRefCitation>
|
||||
<bibRefCitation id="9DF5D2F4230CFD6FFA7A9DE7C4998ED6" author="Guignot, F" journalOrPublisher="Bulletin de Musee Royal d'Histoire Naturelle de Belgique" pageId="0" pageNumber="19" pagination="1 - 14" refId="B27" refString="Guignot, F, 1948. Vingt-sixieme note sur les hydrocanthares. Dytiscides et Gyrinides nouveaux ou peu connus du Congo belge. . Bulletin de Musee Royal d'Histoire Naturelle de Belgique 24: 1 - 14" title="Vingt-sixieme note sur les hydrocanthares. Dytiscides et Gyrinides nouveaux ou peu connus du Congo belge." volume="24" year="1948">Guignot (1948)</bibRefCitation>
|
||||
tentatively refer a single female specimen from Lulua in DR Congo to
|
||||
<taxonomicName lsidName="C. distinguendus" pageId="0" pageNumber="19" rank="species" species="distinguendus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. distinguendus</emphasis>
|
||||
<taxonomicName id="771C2C56E469874D77183CCB60C2115E" lsidName="C. distinguendus" pageId="0" pageNumber="19" rank="species" species="distinguendus">
|
||||
<emphasis id="3773F38D0381641DD6A64280A2DE13CD" italics="true" pageId="0" pageNumber="19">C. distinguendus</emphasis>
|
||||
</taxonomicName>
|
||||
but this was very likely a different species. In Madagascar, it has been recorded from the provinces of Antananarivo, Fianarantsoa Toamasina, and Antsiranana; more specifically from Anjozorobe Angavo reserve, around Ankazobe, Manankazo, Andranofeno Sud, around Ambohitantely reserve, Manjakatompo Ankaratra reserve; also from the eastern central area around Moramanga, Analamazaotra NP; in the southeast from Ambositra in col de Tapia, Ankazomivady forest, Ranomafana NP, Andringitra NP; and from the northeast in Anjanaharibe Sud reserve (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11D</figureCitation>
|
||||
<figureCitation id="37F19392833F28554D7F8055DCB27015" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">This species has been collected in various localities, mostly from open, partly deforested areas at mid- to high altitudes or open forest marshes. It occurs at altitudes above 900 m and is often associated with grass vegetation along lake shores and in marshes, found by stamping, and at vegetation-rich margins of rivers.</paragraph>
|
||||
<subSubSection id="D9FE6A2849D1B77CA72DC93F7E9BE896" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="4D7473AC928683B2082AD905FC3BB451" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="FF01B1E13ED586A39BA5FEBC4F959268" pageId="0" pageNumber="19">This species has been collected in various localities, mostly from open, partly deforested areas at mid- to high altitudes or open forest marshes. It occurs at altitudes above 900 m and is often associated with grass vegetation along lake shores and in marshes, found by stamping, and at vegetation-rich margins of rivers.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<bibRefCitation author="Regimbart, M" journalOrPublisher="Memoires de la Societe Entomologique de Belgique" pageId="0" pageNumber="19" pagination="1 - 244" refId="B63" refString="Regimbart, M, 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. . Memoires de la Societe Entomologique de Belgique 4: 1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines." volume="4" year="1895">
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<subSubSection id="80FAB152FA67CCD4EF5699263391C249" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="B93FDE09E14F0ED90C7D0DCA8C4C0ADB" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="3BA2846E922047146445C09CE955A231" pageId="0" pageNumber="19">
|
||||
<bibRefCitation id="469108084293779AE2F49A1B56F4A5E0" author="Regimbart, M" journalOrPublisher="Memoires de la Societe Entomologique de Belgique" pageId="0" pageNumber="19" pagination="1 - 244" refId="B63" refString="Regimbart, M, 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. . Memoires de la Societe Entomologique de Belgique 4: 1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines." volume="4" year="1895">
|
||||
<normalizedToken id="367000149BA6E73B6FB743A70B5342B5" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
(1895)
|
||||
</bibRefCitation>
|
||||
described this species under the name
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus duodecimstriatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="duodecimstriatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus duodecimstriatus</emphasis>
|
||||
<taxonomicName id="A897B2688260ACAFF7E74AF89D577CA8" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus duodecimstriatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="duodecimstriatus">
|
||||
<emphasis id="7AFF91267FABF13C62D37CC6476F94F9" italics="true" pageId="0" pageNumber="19">Copelatus duodecimstriatus</emphasis>
|
||||
</taxonomicName>
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<normalizedToken id="7AA1E45AD75EDC6F2A78F67FE1E89215" originalValue="Aubé">Aube</normalizedToken>
|
||||
but realised later (1903) that what he had described from Madagascar was not conspecific with
|
||||
<normalizedToken originalValue="Aubé’s">Aube's</normalizedToken>
|
||||
<taxonomicName lsidName="C. duodecimstriatus" pageId="0" pageNumber="19" rank="species" species="duodecimstriatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. duodecimstriatus</emphasis>
|
||||
<normalizedToken id="E4E782BDF2165BCEB9460F5D515E472F" originalValue="Aubé’s">Aube's</normalizedToken>
|
||||
<taxonomicName id="FFF26BFEF3773CF63C43C2AB867DDEB0" lsidName="C. duodecimstriatus" pageId="0" pageNumber="19" rank="species" species="duodecimstriatus">
|
||||
<emphasis id="9D30B188AD81E82A8F753A5E663B3566" italics="true" pageId="0" pageNumber="19">C. duodecimstriatus</emphasis>
|
||||
</taxonomicName>
|
||||
from the Mascarene islands and he gave it the new name
|
||||
<taxonomicName lsidName="C. distinguendus" pageId="0" pageNumber="19" rank="species" species="distinguendus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. distinguendus</emphasis>
|
||||
<taxonomicName id="25A7DE109838F09480A2F45213FC2E1C" lsidName="C. distinguendus" pageId="0" pageNumber="19" rank="species" species="distinguendus">
|
||||
<emphasis id="97D483B35C6C72851C098CD655BE6816" italics="true" pageId="0" pageNumber="19">C. distinguendus</emphasis>
|
||||
</taxonomicName>
|
||||
, referring to his description from 1895.
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus duodecimstriatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="duodecimstriatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus duodecimstriatus</emphasis>
|
||||
<taxonomicName id="18C15EAA5FBDB59ED28C19317FCDC4C9" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus duodecimstriatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="duodecimstriatus">
|
||||
<emphasis id="6B51989EAED28A4E4B4333D565EA5834" italics="true" pageId="0" pageNumber="19">Copelatus duodecimstriatus</emphasis>
|
||||
</taxonomicName>
|
||||
lacks submarginal striae, has a distinctly different penis shape, and is endemic to Mauritius (
|
||||
<bibRefCitation author="Vinson, J" journalOrPublisher="The Mauritius Institute Bulletin" pageId="0" pageNumber="19" pagination="299 - 372" refId="B73" refString="Vinson, J, 1967. Liste chorologique des Coleopteres des Mascareignes. . The Mauritius Institute Bulletin 4: 299 - 372" title="Liste chorologique des Coleopteres des Mascareignes." volume="4" year="1967">Vinson 1967</bibRefCitation>
|
||||
<bibRefCitation id="227285372F404AE7BDFF2009172047D1" author="Vinson, J" journalOrPublisher="The Mauritius Institute Bulletin" pageId="0" pageNumber="19" pagination="299 - 372" refId="B73" refString="Vinson, J, 1967. Liste chorologique des Coleopteres des Mascareignes. . The Mauritius Institute Bulletin 4: 299 - 372" title="Liste chorologique des Coleopteres des Mascareignes." volume="4" year="1967">Vinson 1967</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
<bibRefCitation id="3B3B3AE5F2D6AF977841E0A027E4998F" pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
<bibRefCitation id="26DA2F224A611B486CF043863BD52AA8" pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
). The closest relatives of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
|
||||
<taxonomicName id="CC9E06A92EA419517583DF5E006D8027" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
|
||||
<emphasis id="DA1D468F38131EAAF12C91A4EA07E708" italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
|
||||
</taxonomicName>
|
||||
are likely a group of species on continental Africa with similar genitalia (e.g.,
|
||||
<taxonomicName lsidName="C. ateles" pageId="0" pageNumber="19" rank="species" species="ateles">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. ateles</emphasis>
|
||||
<taxonomicName id="B81D669762930C6815BC4945F5CA81D2" lsidName="C. ateles" pageId="0" pageNumber="19" rank="species" species="ateles">
|
||||
<emphasis id="60C27A854257B0AEE7081E76DF2DC879" italics="true" pageId="0" pageNumber="19">C. ateles</emphasis>
|
||||
</taxonomicName>
|
||||
Guignot, 1955,
|
||||
<taxonomicName lsidName="C. sylvaticus" pageId="0" pageNumber="19" rank="species" species="sylvaticus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. sylvaticus</emphasis>
|
||||
<taxonomicName id="EC581F60C383952A677E6AEF6E5B7518" lsidName="C. sylvaticus" pageId="0" pageNumber="19" rank="species" species="sylvaticus">
|
||||
<emphasis id="7556D40AEBAA2CCEC5BCC1A2B67302F1" italics="true" pageId="0" pageNumber="19">C. sylvaticus</emphasis>
|
||||
</taxonomicName>
|
||||
Guignot, 1952,
|
||||
<taxonomicName lsidName="C. andreinii" pageId="0" pageNumber="19" rank="species" species="andreinii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. andreinii</emphasis>
|
||||
<taxonomicName id="E419C05DAB1366EF1AC8B3912AB2DA23" lsidName="C. andreinii" pageId="0" pageNumber="19" rank="species" species="andreinii">
|
||||
<emphasis id="88D5D051AF0AC087ADC40E3662F96899" italics="true" pageId="0" pageNumber="19">C. andreinii</emphasis>
|
||||
</taxonomicName>
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<normalizedToken id="8AC520CDBBBB00D40EB92FF2E7577386" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
, 1905, and additional species described by
|
||||
<bibRefCitation DOI="https://doi.org/10.4081/nhs.2013.131" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="0" pageNumber="19" pagination="131 - 155" refId="B9" refString="Bilardo, A, Rocchi, S, 2013. Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012). . Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 154: 131 - 155" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9 eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012)." url="https://doi.org/10.4081/nhs.2013.131" volume="154" year="2013">Bilardo and Rocchi 2013</bibRefCitation>
|
||||
<bibRefCitation id="154A46B9A94843E64277B470C7FB61B4" DOI="https://doi.org/10.4081/nhs.2013.131" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="0" pageNumber="19" pagination="131 - 155" refId="B9" refString="Bilardo, A, Rocchi, S, 2013. Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012). . Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 154: 131 - 155" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9 eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012)." url="https://doi.org/10.4081/nhs.2013.131" volume="154" year="2013">Bilardo and Rocchi 2013</bibRefCitation>
|
||||
). These have a variable number of elytral striae from five discal to ten discal plus one submarginal, but similar
|
||||
<normalizedToken originalValue="“pumpjack”">"pumpjack"</normalizedToken>
|
||||
<normalizedToken id="789B0735443C59D58D99F165391D7D79" originalValue="“pumpjack”">"pumpjack"</normalizedToken>
|
||||
-shaped genitalia (
|
||||
<bibRefCitation DOI="https://doi.org/10.4081/nhs.2013.131" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="0" pageNumber="19" pagination="131 - 155" refId="B9" refString="Bilardo, A, Rocchi, S, 2013. Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012). . Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 154: 131 - 155" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9 eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012)." url="https://doi.org/10.4081/nhs.2013.131" volume="154" year="2013">Bilardo and Rocchi 2013</bibRefCitation>
|
||||
<bibRefCitation id="87C9122132419FB2C90060A11B0C7943" DOI="https://doi.org/10.4081/nhs.2013.131" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="0" pageNumber="19" pagination="131 - 155" refId="B9" refString="Bilardo, A, Rocchi, S, 2013. Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012). . Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 154: 131 - 155" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9 eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012)." url="https://doi.org/10.4081/nhs.2013.131" volume="154" year="2013">Bilardo and Rocchi 2013</bibRefCitation>
|
||||
).
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
|
||||
<taxonomicName id="92BBB77CD27BF10F40F58A027BF011FF" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
|
||||
<emphasis id="2C94B51ADEEDD200B671B55FDB7F2ECB" italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
|
||||
</taxonomicName>
|
||||
differs from these especially by the lack of a lateral tooth on the male penis in ventral view.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,236 +1,236 @@
|
|||
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<taxonomicName LSID="573f0c84-f705-55e7-a4f6-fc4974616368" authority="Guignot, 1941" authorityName="Guignot" authorityYear="1941" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">Copelatus insuetus Guignot, 1941</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Figs 6A</figureCitation>
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|
||||
<figureCitation id="93E3D8C7F80DC5667E80B205EAFD1D3E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Figs 6A</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus, dorsal view. A Female: Copelatus insuetus B Male: Copelatus vokoka sp. nov. C Female: Copelatus kely sp. nov. D Female: Copelatus sp. 3 (Ivohibe)." figureDoi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19">9A</figureCitation>
|
||||
<figureCitation id="49748540A33475FE14A9FD3D57920560" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus, dorsal view. A Female: Copelatus insuetus B Male: Copelatus vokoka sp. nov. C Female: Copelatus kely sp. nov. D Female: Copelatus sp. 3 (Ivohibe)." figureDoi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19">9A</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">Copelatus insuetus</taxonomicName>
|
||||
<subSubSection id="E2E5F9F12AF963C446BE076CAE3F110E" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="42A56B8CC60CCB5D694005F1A15D4D00" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="7A601CBDD011C0278C90AA2C57785334" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">Copelatus insuetus</taxonomicName>
|
||||
Guignot, 1941: 39.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Madagascar, Perinet [= Analamazaotra NP].</paragraph>
|
||||
<subSubSection id="49D56D118D67F6DB09CAEF398AD9AD3A" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="B3B716353167DFB3A495DE7413DAF905" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="33C839623C0EBF9879DFBA323B1D2678" pageId="0" pageNumber="19">Madagascar, Perinet [= Analamazaotra NP].</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Based on single male specimen (holotype). Madagascar, Perinet.</paragraph>
|
||||
<subSubSection id="F44DB443D11F65F0E4722EC85DFB178A" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="007D238048FB07788D0B98BDD108EBA1" pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph id="3E7A48FB43D936DD515DB4CF6E6ACD1C" pageId="0" pageNumber="19">Based on single male specimen (holotype). Madagascar, Perinet.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga</emphasis>
|
||||
<subSubSection id="2194ADE73580737827D5A463A92F757F" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="ADCD41CA92801F2AB91927546EB85949" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="A5A4F75D07F7EC67A85B2C295F0FAC66" pageId="0" pageNumber="19">
|
||||
<emphasis id="FB1DA05766D71556C36217AE255AB04A" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga</emphasis>
|
||||
: -HT♂ (GP) (MNHN, "coll. Guignot"): Data in NHRS | JLKB | 000030227 // Madagascar | Perinet // Type [red label] // [male symbol] //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="DD1389162BAEED25936264F1F941FCB2" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">insuetus</emphasis>
|
||||
<emphasis id="F9CF952BCBF59C74078B7EA3925CB403" italics="true" pageId="0" pageNumber="19">insuetus</emphasis>
|
||||
| guign. Type //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga, Ambatondrazaka, Andilamena</emphasis>
|
||||
<subSubSection id="CFE0DDD331DF19E9326AF2DC16ACD450" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="2633134B0FB2AFED53046247CD0D1606" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="047B2209F39DA080A2BCCBAD9CB6093A" pageId="0" pageNumber="19">
|
||||
<emphasis id="C4CFDCDE451AB842A0F514E3A2B76299" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga, Ambatondrazaka, Andilamena</emphasis>
|
||||
: -2♀ (MNHN, "coll. Guignot"): Data in NHRS | JLKB | 000030228-9 // Perinet // INSTITUT | SCIENTIFIQUE | MADAGASCAR // [female symbol] // -5♀ (NMW): Data in NHRS | JLKB | 000030302-6 // Madagascar (Md 4/5) | Andasibe NP, Perinet | 1250 m
|
||||
<normalizedToken originalValue="Pfütze">Pfuetze</normalizedToken>
|
||||
<normalizedToken id="C7849894833CB34F89918DFA06E5B010" originalValue="Pfütze">Pfuetze</normalizedToken>
|
||||
im Urwald | 5.12.2000, leg. W. Dolin // -1♂ (NMCP, "coll.
|
||||
<normalizedToken originalValue="Hájek”">Hajek"</normalizedToken>
|
||||
<normalizedToken id="326DC3FA310DA602E93C4913C7DB52EC" originalValue="Hájek”">Hajek"</normalizedToken>
|
||||
): Data in NHRS | JLKB | 000030312 // Madagascar 10.1.2007 | Andasibe NP. | Lokato-near Andasibe | Mracek Z. leg. // coll. Jiri
|
||||
<normalizedToken originalValue="HÁJEK">HAJEK</normalizedToken>
|
||||
<normalizedToken id="89D8A1C9D6B13B5A1EC5D4CCBD7B9F5C" originalValue="HÁJEK">HAJEK</normalizedToken>
|
||||
| National Museum | Prague, Czech Republic // -1♀, 1♂ (NMW): Data in NHRS | JLKB | 000030307-8 // E-Madagascar (10) | Ambatombe near Andilamena | 900 m asl. 17.01.1995 | G. Dunay & J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="7AF8D430A5005325674EEA578831BE7C" originalValue="Janák">Janak</normalizedToken>
|
||||
coll. -1♀, 1♂ (NMW): Data in NHRS | JLKB | 000030309-10 // E-Madagascar (11) Ampamoho | near Andilamena, 1200-1300 m. | asl. 18-20.01.1995 | G. Dunay & J.
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="D0CD0D40E1F830B64C59C1A61F91A13E" originalValue="Janák">Janak</normalizedToken>
|
||||
coll. // -1♀ (NMW): Data in NHRS | JLKB | 000030311 // Madagascar 18-20.1.1995 | 5km S Ampamoho | nr. Andilamena, 950-1000 m | Dunay &
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="AB907DCEBD6063FF92467B4140D47B73" originalValue="Janák">Janak</normalizedToken>
|
||||
(11) // -1♀ (NHMUK): // BMNH-797884 // MAD: AMPA: Moramanga: Andasibe | Andasibe NP: Chanel| P61BI02: N: -18.935: E:48.418: 944 m | 04/I/2007 Leg. Isambert et al // DNA Voucher | BMNH <797884> | MSL294:A12 // -2♀ (NHRS): // NHRS-JLKB | 000010692-3 // Madagascar: Tamatave: Alaotra Mangoro: | Analamazaotra RS; Bas fond, non-permanent | pond near trail to "lac rouge"MAD15-1 | 943 m,
|
||||
<geoCoordinate degrees="18" direction="south" minutes="56" orientation="latitude" precision="1" seconds="26.7" value="-18.940748">18°56'26.7"S</geoCoordinate>
|
||||
<geoCoordinate id="5891F2DD232427386E8C6922C3E9C3BD" degrees="18" direction="south" minutes="56" orientation="latitude" precision="1" seconds="26.7" value="-18.940748">18°56'26.7"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="048" direction="east" minutes="25" orientation="longitude" precision="1" seconds="03.9" value="48.41775">048°25'03.9"E</geoCoordinate>
|
||||
<geoCoordinate id="815734A96E32EC0466647C23D46764F7" degrees="048" direction="east" minutes="25" orientation="longitude" precision="1" seconds="03.9" value="48.41775">048°25'03.9"E</geoCoordinate>
|
||||
, 16.III.2015 | Among vegetation and dead leaves in the pond | Leg.T. Ranarilalatiana & H.J. Randriamihaja // -1♀, 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000065745, 10783 (Alc.) // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra RS: close to | park entrance:
|
||||
<geoCoordinate degrees="18.93573" direction="south" orientation="latitude" precision="1" value="-18.93573">18.93573S</geoCoordinate>
|
||||
<geoCoordinate id="7CE69A106E87BD583C5BE5A94668660B" degrees="18.93573" direction="south" orientation="latitude" precision="1" value="-18.93573">18.93573S</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate degrees="048.41741" direction="east" orientation="longitude" precision="1" value="48.41741">048.41741E</geoCoordinate>
|
||||
<geoCoordinate id="01734A837A72288ECA949D77AB38F279" degrees="048.41741" direction="east" orientation="longitude" precision="1" value="48.41741">048.41741E</geoCoordinate>
|
||||
| 930 m. 08.XI.2011 GB Nets and sieves: | stagnant pools in tributary to | Analamazaotra river; Field# MAD11-26 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -1♂ (GP), 1♀, 3 ex. (1♂, 2♀) (Alc.) (NHRS): // NHRS-JLKB | 000010839, 65702, 10840 (Alc.) // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, Analamazaotra | 250m E of park entrance: Mad14-18: | dried up river bed with stagnant pools: | 18.9357S 48.4174E: 930 m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♀ (NHRS): // NHRS-JLKB | 000010841 // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, Analamazaotra | pond: 100m E from Lac vert: Mad14-15: | shallow forest pool:
|
||||
<geoCoordinate degrees="18.9385" direction="south" orientation="latitude" precision="5" value="-18.9385">18.9385S</geoCoordinate>
|
||||
<geoCoordinate degrees="48.4219" direction="east" orientation="longitude" precision="5" value="48.4219">48.4219E</geoCoordinate>
|
||||
<geoCoordinate id="AD24973079B547453EBEBFB3AEA88FCD" degrees="18.9385" direction="south" orientation="latitude" precision="5" value="-18.9385">18.9385S</geoCoordinate>
|
||||
<geoCoordinate id="346A212186772E47EC887E731A0D70DA" degrees="48.4219" direction="east" orientation="longitude" precision="5" value="48.4219">48.4219E</geoCoordinate>
|
||||
: | 940 m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♂ (GP), 1♀, 1♀ (Alc.) (NHRS): // NHRS-JLKB | 000065705, 65777, 10844(Alc.) // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, Analamazaotra | 150 m E of park entrance: Mad14-14: | shallow partly dried out forest pond: |
|
||||
<geoCoordinate degrees="18.9355" direction="south" orientation="latitude" precision="5" value="-18.9355">18.9355S</geoCoordinate>
|
||||
<geoCoordinate degrees="48.4166" direction="east" orientation="longitude" precision="5" value="48.4166">48.4166E</geoCoordinate>
|
||||
<geoCoordinate id="159B403816BA724A6ED81A9F1C4451FD" degrees="18.9355" direction="south" orientation="latitude" precision="5" value="-18.9355">18.9355S</geoCoordinate>
|
||||
<geoCoordinate id="3776063AEF4D77D6B0E4E53D9779F5A4" degrees="48.4166" direction="east" orientation="longitude" precision="5" value="48.4166">48.4166E</geoCoordinate>
|
||||
: 930 m: 27.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♂ (GP), 34 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010842, 10843(Alc.) // MAD: TOAM: Alaotra Mangoro | Andasibe Mantadia NP, | Mantadia: 12Km N of park | entrance: Mad14-83: | forest pond with some | vegetation: 18.7911S | 48.4259E: 960 m: 28.XI.2014 // Leg. J. Bergsten, R. | Bukontaite, S. Holmgren, | J.H. Randriamihaja | & T. Ranarilalatiana // -6♂ (GP), 4♀, 33 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010534-7, 10606-11, 10780(Alc.) // MAD: TOAM: Alaotra Mangoro: | Betsabora river by RN2 near | Antasmpanana village, 6Km W | of Moramanga: MAD14-81: river | with side pools: 18.9247S | 48.1828E: 900 m: 24.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -1♀ (NHMUK): // BMNH-797895 // MAD: TOAM: Ambatondrazaka | Zahamena: Zahamena NP: Stream | P60BI15:N: -17.52: E:48.721: 1075 m| 31:XI:2006: Leg. Isambert et al // DNA Voucher | BMNH <797895> | MSL294:B11 // -4♂ (GP), 1♀ 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010817, 10872-4, 65700, 10880(Alc.) // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | path towards Camp Cascade: S17.5166; | E048.7227; 1040 m. 07.III.2018; GB Nets, | white pan & sieves: Waterfilled goldigging | hole in forestswamp: Field# MAD18-80 | Leg. J. Bergsten, & T. Ranarilalatiana // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010885 // Madagascar: Toamasina: Ambatondrazaka | Zahamena NP: the way to Camp | Cascade: 19.III.2013, GB Nets & sieves: slow | stream, Leg. J.H. Randriamihaja & | T. Ranarialalatiana: Field# ZAH13-02 // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010881 // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Analamaitso forest: S17.5179; E048.7232; | 1050 m. 07.III.2018; GB Nets, white pan & | sieves: small foreststream: Field# MAD18-81 | Leg. J. Bergsten, & T. Ranarilalatiana // -3♀ (Alc.) (NHRS): // NHRS-JLKB | 000010882 // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Ambavahala forest: S17.5300; E048.7161; | 1070 m. 08.III.2018; GB Nets, white pan & | sieves: larger shaded forestswamp: Field# MAD18-90 | Leg. J. Bergsten, & T. Ranarilalatiana // -1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010875, 10883(Alc.) // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Pandanus Swamp: S17.5166; E048.7227; | 1040 m. 08.III.2018; GB Nets, white pan & | sieves: forestswamp: Field# MAD18-91 | Leg. J. Bergsten, & T. Ranarilalatiana -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010884 // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: | Analamaitso forest: S17.5225; E048.7250; | 1090 m. 08.III.2018; GB Nets, white pan & | sieves: waterfilled golddigging hole in flat: Field# MAD18-92 | Leg. J. Bergsten, & T. Ranarilalatiana // -2♂ (GP), 3♀, 6♀ (Alc.), 6ex. (3♂, 3♀) (Alc.), (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010816, 10876-9, 10886(Alc.), 65753(Alc.) // Madagascar: Toamasina: Alaotra Mangoro: | Zahamena NP: Sect. Antanandava: 150 m | upstream to Camp Cascade: S17.5458; E048.7244; | 1270 m. 10-11.III.2018; GB Nets, white pan & | sieves: waterpool with dead leaves: Field# MAD18-109, | Leg. J. Bergsten, & T. Ranarilalatiana //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Analanjirofo: Maroantsetra</emphasis>
|
||||
<emphasis id="0919E0E8465432812D363D262445A63B" bold="true" pageId="0" pageNumber="19">Toamasina. Analanjirofo: Maroantsetra</emphasis>
|
||||
: -1♂, 1♀ (MNHN, "coll. Legros"): Data in NHRS | JLKB | 000030314-5 // Maroantsetra | Vadon IV.50 // 1/6 //? insuetus | guign. // [These two specimens were seen in MNHN, but their identity, after the discovery of additional very similar species in the complex, was not confirmed. We therefore only tentatively refer them to this species.]
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Fianarantsoa. Vatovavy Fitovinany: Ifanadiana</emphasis>
|
||||
<emphasis id="2DDB8D9733436261D9EF6B9E7EA85A72" bold="true" pageId="0" pageNumber="19">Fianarantsoa. Vatovavy Fitovinany: Ifanadiana</emphasis>
|
||||
: -1♀ (CAS): // CASENT-8131927 // Madagascar: Province | Fianarantsoa, Parc National | Ranomafana, radio tower | at forest edge, elev 1130 m | 19-26 February 2002 // 21°15.05'S, 47°24.43' | coll: M. Irwin, R.
|
||||
<normalizedToken originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||||
<normalizedToken id="84FF49B5D3F66AB625C0DFF22D822627" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||||
| California Acad. of Sciences | malaise. Mixed tropical | forest MA-02-09B-17 // -1♂ (teneral) (NMPC, "coll.
|
||||
<normalizedToken originalValue="Hájek”">Hajek"</normalizedToken>
|
||||
<normalizedToken id="206053FEE6621BCEA897262729ADBE65" originalValue="Hájek”">Hajek"</normalizedToken>
|
||||
): Data in NHRS | JLKB | 000030313 // Madagascar 26-31.I.2007 | Ranomafana NP | Ranomafana vill. env. | Z. Mracek leg. // coll. Jiri
|
||||
<normalizedToken originalValue="HÁJEK">HAJEK</normalizedToken>
|
||||
<normalizedToken id="F36E253B3172BD8383C1E39885E1C887" originalValue="HÁJEK">HAJEK</normalizedToken>
|
||||
| National Museum | Prague, Czech Republic //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Fianarantsoa. Matsiatra Ambony: Lalangina</emphasis>
|
||||
<emphasis id="E69CFF8D8955BB4D397EC39F58D0913E" bold="true" pageId="0" pageNumber="19">Fianarantsoa. Matsiatra Ambony: Lalangina</emphasis>
|
||||
: -1♀ (CAS): // CASENT-5004000 // Madagascar, Fianarantsoa | Province, Ranomafana National | Park, Vohiparara village, 1160m | mixed tropical forest | 2-22 January 2001 | 21.23906S / 47.38487E | COL-DHK-2001-003 // D.H. & K.M. Kavanagh | R.L. Brett, E. Elsom, F. Vargas, | R. Ranaivosolo, E.F. Randrianirina, | N. Rasoamanana, T.J. | Ravelomanana and H.C. | Raveloson collectors // -4♂ (CAS): // CASENT-5004056, 5004070-1, 5004075 // Madagascar, Fianarantsoa | Province, Ranomafana | National Park, Vohiparara | area, 1170m mixed tropical | forest 2-22 January 2001 |
|
||||
<geoCoordinate degrees="21.22644" direction="south" orientation="latitude" precision="1" value="-21.22644">21.22644S</geoCoordinate>
|
||||
<geoCoordinate id="5F45D3628D6554571DD87DBAAC45D08E" degrees="21.22644" direction="south" orientation="latitude" precision="1" value="-21.22644">21.22644S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="47.36979" direction="east" orientation="longitude" precision="1" value="47.36979">47.36979E</geoCoordinate>
|
||||
<geoCoordinate id="AABDD819702A226FFA065830723A0F78" degrees="47.36979" direction="east" orientation="longitude" precision="1" value="47.36979">47.36979E</geoCoordinate>
|
||||
, | Stop# DHK-01-004 // D.H. & K.M. Kavanagh | R.L. Brett, E. Elsom, F. | Vargas, R. Ranaivosolo, | E.F. Randrianirina, N. | Rasoamanana, T.J. | Ravelomanana and H.C. | Raveloson collectors //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Nosy-Be</emphasis>
|
||||
<emphasis id="350AC792B479B4C393F4D2A810F36699" bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Nosy-Be</emphasis>
|
||||
: -1♂(GP) (MNHN): // Data in NHRS | JLKB | 000030230 // Nosy-Be | Sokobe [=Lokobe NP] | V. 57 Hoyt Coll // ♂ [male symbol] //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny</emphasis>
|
||||
<emphasis id="F789512BFC4B6B9866AD7A598B1536FD" bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny</emphasis>
|
||||
: -1♀ (NHRS): // NHRS-JLKB | 000010694 (JB206) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="A2E2F91507F0296D21AB60B8F3E80258" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010781 (JB205) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30350 | E046.81068, 87 m.a.o. 29.XI.2009 | Water Net, Field# MAD09-03 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="98051A532CF0034A49156F9278F4E56F" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="DA30F2F5D477936A6DEBADE292C54C5C" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="C2A6E85598D2F47B95AA6FB45BD776FF" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="9AF641851E3BE8B5778C8010DB9A5B58" pageId="0" pageNumber="19">
|
||||
Body shape elongate oval, elytra brown with basal testaceous band, with six discal and one submarginal stria (
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus, dorsal view. A Female: Copelatus insuetus B Male: Copelatus vokoka sp. nov. C Female: Copelatus kely sp. nov. D Female: Copelatus sp. 3 (Ivohibe)." figureDoi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19">Fig. 9A</figureCitation>
|
||||
<figureCitation id="5E7B3B3ED3FADABB3A52C1A378302352" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus, dorsal view. A Female: Copelatus insuetus B Male: Copelatus vokoka sp. nov. C Female: Copelatus kely sp. nov. D Female: Copelatus sp. 3 (Ivohibe)." figureDoi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19">Fig. 9A</figureCitation>
|
||||
).
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus insuetus</emphasis>
|
||||
<taxonomicName id="D8B317D298046973DB7AC553F74E4E8E" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">
|
||||
<emphasis id="64B1BF893B5B31C1416913FF48AFC423" italics="true" pageId="0" pageNumber="19">Copelatus insuetus</emphasis>
|
||||
</taxonomicName>
|
||||
can be distinguished from the following species based on the penis ventral outline in lateral view with a distinct
|
||||
<normalizedToken originalValue="“shoulder”">"shoulder"</normalizedToken>
|
||||
<normalizedToken id="57273FAD0BB7E012B19942853572347F" originalValue="“shoulder”">"shoulder"</normalizedToken>
|
||||
(
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
<figureCitation id="0140CC8F728C5233AA38B849E59B5055" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
), an apex in left lateral view with a dorsal ridge crossing posterior dorsal margin and medial part longitudinally coarsely sulcate ventrolaterally in right lateral view (
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
<figureCitation id="D2FC7BAFDEE2DB5FD015E62F739D1D85" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="7AF55337DDEB6797493B8D1E9F66E8D5" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="67A0D40BF735A82D57EF39E22CF0E175" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="9472231FB10908C950DE535B08CB814A" pageId="0" pageNumber="19">
|
||||
Body length 4.3-5.0 mm. Body shape elongate, weakly oval, some specimens appearing subparalell. Head rufotestaceous anteriorly and posteriorly but infuscated by two blotches inbetween the eyes. Blotches usually meet in middle and form an M-mark, but sometimes infuscation reduced to nearly absent. Pronotum usually with extensive medial infuscation leaving only lateral, anterolateral and posterolateral areas testaceous. Some specimens with less extensive medial infuscation (
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus, dorsal view. A Female: Copelatus insuetus B Male: Copelatus vokoka sp. nov. C Female: Copelatus kely sp. nov. D Female: Copelatus sp. 3 (Ivohibe)." figureDoi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19">Fig. 9A</figureCitation>
|
||||
<figureCitation id="99380ADF2FF6C471C122FADDAE6AA56D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus, dorsal view. A Female: Copelatus insuetus B Male: Copelatus vokoka sp. nov. C Female: Copelatus kely sp. nov. D Female: Copelatus sp. 3 (Ivohibe)." figureDoi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19">Fig. 9A</figureCitation>
|
||||
). Elytra brown with an irregular testaceous band at base. Testaceous band with longest projection posteriorly in second interval. All appendages testaceous.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Elytra with six clearly impressed discal and one submarginal stria. Fifth and sixth striae abbreviated anteriorly, and also first stria may be somewhat abbreviated. Submarginal stria starting between one third from the base and midway between base and apex. Head, pronotum, and elytra microreticulate and finely micropunctate. Posterolateral corners of pronotum with strioles that extend along posterior surface, reduced or absent medially.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="641398E3F6DE98EBCEAE4E90DDE31040" pageId="0" pageNumber="19">Elytra with six clearly impressed discal and one submarginal stria. Fifth and sixth striae abbreviated anteriorly, and also first stria may be somewhat abbreviated. Submarginal stria starting between one third from the base and midway between base and apex. Head, pronotum, and elytra microreticulate and finely micropunctate. Posterolateral corners of pronotum with strioles that extend along posterior surface, reduced or absent medially.</paragraph>
|
||||
<paragraph id="9705E4125F0A44BF13ECF04CDC7A5AA6" pageId="0" pageNumber="19">
|
||||
Ventral side ferrugineous brown, except prothorax, epipleura, appendages, and gula of head which are testaceous. Area around metacoxal lines lighter (not all individuals) and also spots laterally on abdominal sternites
|
||||
<normalizedToken originalValue="II–VII">II-VII</normalizedToken>
|
||||
<normalizedToken id="E854B40B060F8DFBAC5F5F2CD27E4868" originalValue="II–VII">II-VII</normalizedToken>
|
||||
and along some sternite margins are lighter. Metacoxa and abdominal sternites
|
||||
<normalizedToken originalValue="II–IV">II-IV</normalizedToken>
|
||||
<normalizedToken id="0131EA59E780CCE1B5A2B9298C60224D" originalValue="II–IV">II-IV</normalizedToken>
|
||||
marked with strioles. Prosternal process short, broad, with blunt apex. Lateral parts of metaventrite broad. Metacoxal lines long, abbreviated only slightly before metaventral margin, diverging anteriorly.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="856DCED7924C090D4EA9AC38CA11560F" pageId="0" pageNumber="19">
|
||||
Male: first three pro- and mesotarsomeres widened and ventrally equipped with suction cups; number of suction cups per segment for
|
||||
<normalizedToken originalValue="I–III">I-III</normalizedToken>
|
||||
<normalizedToken id="A33A76343F2F0C979F0BEFCFD4D9D7FF" originalValue="I–III">I-III</normalizedToken>
|
||||
: 7:4:4 for both pro- and mesotarsus. Protibia modified, narrow with a bisinuate angulate ventral margin at base, broadened distally. Pro- and mesotarsal claws unmodified. Penis in ventral view thin and simple, apical part slightly left-turned. Penis in lateral view angled after basal third forming a
|
||||
<normalizedToken originalValue="“shoulder”">"shoulder"</normalizedToken>
|
||||
<normalizedToken id="56D7F7DB617520C60F53E4BB18A6B56F" originalValue="“shoulder”">"shoulder"</normalizedToken>
|
||||
. Apex in left lateral view with a characteristic dorsal ridge crossing posterior dorsal margin (
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
<figureCitation id="9780A8FEC6A7000F21D8814A04A26266" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
). Penis serrated with fine transverse ridges preapically on the left ventral ridge and halfway down the left side. Penis medially strongly longitudinally sulcate ventrolaterally in right lateral view (
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
<figureCitation id="BEBC8E30E023AA192D88AEC175F8C999" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Fig. 6A</figureCitation>
|
||||
). Parameres as in
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Figure 6A</figureCitation>
|
||||
<figureCitation id="ED7C917949621568EEE54ABE82EE8F03" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov." figureDoi="10.3897/zookeys.869.33997.figure6" httpUri="https://binary.pensoft.net/fig/322931" pageId="0" pageNumber="19">Figure 6A</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Female: anterior half of elytra finely striolated from second or third elytral interval to external margin. Degree of elytral striolation in females quite variable between specimens. These strioles are finer than the strioles on pronotum found in both sexes.</paragraph>
|
||||
<caption doi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19" start="Figure 9" startId="F9">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="194D45AA70C62F2FF3B0D8ADC1F466C0" pageId="0" pageNumber="19">Female: anterior half of elytra finely striolated from second or third elytral interval to external margin. Degree of elytral striolation in females quite variable between specimens. These strioles are finer than the strioles on pronotum found in both sexes.</paragraph>
|
||||
<caption id="2EAC0C05A6DEE469BBA1FAC9B8FA8070" doi="10.3897/zookeys.869.33997.figure9" httpUri="https://binary.pensoft.net/fig/322934" pageId="0" pageNumber="19" start="Figure 9" startId="F9">
|
||||
<paragraph id="35994126B33A76388D0231F9A431017A" pageId="0" pageNumber="19">
|
||||
Figure 9. Habitus, dorsal view.
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
|
||||
<emphasis id="14C6065296FC843170A4A7F51DE68678" bold="true" pageId="0" pageNumber="19">A</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus insuetus</emphasis>
|
||||
<taxonomicName id="EACC0919FF4F61581E35BF51F011D49B" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus insuetus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="insuetus">
|
||||
<emphasis id="4CD0FA52D5FD07D5F731CF7DEEFC47C0" italics="true" pageId="0" pageNumber="19">Copelatus insuetus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||||
<emphasis id="6EB50428D53E4BA10A34CD1135BE62F4" bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||||
Male:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus vokoka" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="vokoka">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus vokoka</emphasis>
|
||||
<taxonomicName id="10BA20CE7397C4E7EAC36B829AD5048D" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus vokoka" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="vokoka">
|
||||
<emphasis id="3BF97DC00987DF46DBB03AE0EC4D1210" italics="true" pageId="0" pageNumber="19">Copelatus vokoka</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||||
<emphasis id="AFDD768CBA10F2EDD8FEDD9F7E1BDBFB" bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus kely" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="kely">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus kely</emphasis>
|
||||
<taxonomicName id="2A8ECB0CF9DFA11DBD28AD39E22C33C5" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus kely" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="kely">
|
||||
<emphasis id="3C9256F394E412675266E6B4D221C376" italics="true" pageId="0" pageNumber="19">Copelatus kely</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||||
<emphasis id="5C2D4627CD3604C2F72E52349B9607A6" bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||||
Female:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="B3678870DDF1916D3C8E2026D02A4CF7" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A9619BC7E32F0B503F41458D3E59934D" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 3 (Ivohibe).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="6004935508411B8B9616629A062A7A51" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="45E3FD9AF2183109049675C8BD13BD1E" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="610CC132CC64C8CC6A46F23AFF5A1B46" pageId="0" pageNumber="19">
|
||||
Endemic to Madagascar. Known with certainty from the eastern escarpment, Ranomafana NP in the south, Analamazaotra NP, and Mantadia NP in the central region, and Zahamena NP and Andilamena further north (
|
||||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution maps of Copelatus species. A C. insuetus (circle), C. pseudostriatus sp. nov. (square) B C. kely sp. nov. (circle), C. vokoka sp. nov. (square) C C. ankaratra sp. nov. (circle), C. sp. 3 (square) D C. safiotra sp. nov. (circle), C. befasicus (square)." figureDoi="10.3897/zookeys.869.33997.figure12" httpUri="https://binary.pensoft.net/fig/322937" pageId="0" pageNumber="19">Fig. 12A</figureCitation>
|
||||
<figureCitation id="C14C3765B394E29129A02FD6DD811274" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution maps of Copelatus species. A C. insuetus (circle), C. pseudostriatus sp. nov. (square) B C. kely sp. nov. (circle), C. vokoka sp. nov. (square) C C. ankaratra sp. nov. (circle), C. sp. 3 (square) D C. safiotra sp. nov. (circle), C. befasicus (square)." figureDoi="10.3897/zookeys.869.33997.figure12" httpUri="https://binary.pensoft.net/fig/322937" pageId="0" pageNumber="19">Fig. 12A</figureCitation>
|
||||
). We have also seen older material including males from Paris museum from lowland Lokobe NP (island of Nosy Be) in NW and Maroantsetra in NE (not reconfirmed) that correspond to the same species. Finally, two females (NHRS) from Ankarafantsika NP is likely this species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">We have collected this species in Analamazaotra NP in stagnant forest pools with vegetation or with plentiful of dead leaves. Also found in a sidepool next to a river in semi-open degraded area near Moramanga. The eastern escarpment localities range in altitude between 900-1300 m, but if the localities of Nosy Be and Maroantsetra are correct this species can also occur at lowland sea level altitudes. It seems to be most abundant in tropical eastern forests, but if the Ankarafantsika females belong to this species, then it can also occur in western deciduous forests.</paragraph>
|
||||
<subSubSection id="38AE33222B9B3F763802013F5B025B54" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="5B50F64204E6FB0ECCE26CDD3D85ABC6" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="2B90FDA14A9CBB27AE217A5554A430EC" pageId="0" pageNumber="19">We have collected this species in Analamazaotra NP in stagnant forest pools with vegetation or with plentiful of dead leaves. Also found in a sidepool next to a river in semi-open degraded area near Moramanga. The eastern escarpment localities range in altitude between 900-1300 m, but if the localities of Nosy Be and Maroantsetra are correct this species can also occur at lowland sea level altitudes. It seems to be most abundant in tropical eastern forests, but if the Ankarafantsika females belong to this species, then it can also occur in western deciduous forests.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="3A3565A11B8D920C0BA06D1A06CBF3B2" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="91244E2E3500AA95F400787DC3AE3170" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="57D0B2886397B7914093FFD472D426CC" pageId="0" pageNumber="19">
|
||||
Eventual older records of
|
||||
<taxonomicName lsidName="C. insuetus" pageId="0" pageNumber="19" rank="species" species="insuetus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. insuetus</emphasis>
|
||||
<taxonomicName id="C688B23CAC9D7575CE262B02938A0316" lsidName="C. insuetus" pageId="0" pageNumber="19" rank="species" species="insuetus">
|
||||
<emphasis id="B4F409F041BD5655A31657EDDC278E37" italics="true" pageId="0" pageNumber="19">C. insuetus</emphasis>
|
||||
</taxonomicName>
|
||||
should be regarded with caution in light of the habitually very similar new species described below.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,234 +1,234 @@
|
|||
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<taxonomicName authority="r. narina" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. r. var. bryoniae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="napi" status="stat. rev." subSpecies="r.">Pieris napi var. bryoniae r. narina</taxonomicName>
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Verity, 1908;
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Palaearctica (13-14): 145, (17-20): pl. 32, f. 28-29, pl. 33, f. 2; TL:
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. (original description)
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|
||||
<taxonomicName authority="narina" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris narina subsp. narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="narina" status="stat. rev." subSpecies="narina">Pieris narina narina</taxonomicName>
|
||||
<paragraph id="049B5CDBF0F76242D26BB2C1100C6006" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="EF51868E67F3A9E3AD5EA8A3F0BE7B42" authority="narina" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris narina subsp. narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="narina" status="stat. rev." subSpecies="narina">Pieris narina narina</taxonomicName>
|
||||
Verity, 1908;
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="FE8814FB83605C137DDC8F05BEE8AE35" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
: Herbipoliana 1-(1--2): 221-222 (infrasubspecific name validated as distinct species).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. var. var. banghaasi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="variety" species="napi" status="stat. rev." subSpecies="var." variety="banghaasi">Pieris napi var. banghaasi</taxonomicName>
|
||||
<paragraph id="6A16B739220A478FB220EAAEC6D39587" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="5EA47BD6855D0E28CA7D77D6C3703BBD" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi subsp. var. var. banghaasi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="variety" species="napi" status="stat. rev." subSpecies="var." variety="banghaasi">Pieris napi var. banghaasi</taxonomicName>
|
||||
Sheljuzhko, 1910;
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="905AEA9C2D422F942D9B7D9FAD0B15CE" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
: Herbipoliana 1 (1-2): 221-223 (as a synonym of
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. narina" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="narina" status="stat. rev.">P. narina</taxonomicName>
|
||||
<taxonomicName id="A5F56C8AA897E6C7B9F36FCD1EECADE8" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="narina" status="stat. rev.">P. narina</taxonomicName>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris ochsenheimeri subsp. narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="ochsenheimeri" status="stat. rev." subSpecies="narina">Pieris ochsenheimeri narina</taxonomicName>
|
||||
<paragraph id="3618B8E8DB57B8D93966A528F5D93C17" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="B0EE7291118902CA8413C39B375AD873" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris ochsenheimeri subsp. narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="ochsenheimeri" status="stat. rev." subSpecies="narina">Pieris ochsenheimeri narina</taxonomicName>
|
||||
Verity, 1908;
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="E810991018D1A7E3E53F5A98E709CE23" author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
: Butterflies 65: 20-25 (as subspecies of
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="ochsenheimeri" status="stat. rev.">P. ochsenheimeri</taxonomicName>
|
||||
<taxonomicName id="6B7675E3DE51CB89FAFA3B4143FB6AFB" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="ochsenheimeri" status="stat. rev.">P. ochsenheimeri</taxonomicName>
|
||||
Staudinger, 1886)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="napi (Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi" status="stat. rev.">Pieris napi</taxonomicName>
|
||||
<paragraph id="9BD0F6F8657C1AF8127F048D450D4576" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="F6EDA1A076A125CE3486F6CF06DEC7BC" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="napi (Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi" status="stat. rev.">Pieris napi</taxonomicName>
|
||||
(Linnaeus, 1758); Wu & Hsu (2017): Butterflies of China vol. 1: 401-404, f. 15-16 [
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">MisID</emphasis>
|
||||
<emphasis id="2D1DEEC6A18B7CCA72BDD9462745FBC3" bold="true" pageId="0" pageNumber="257">MisID</emphasis>
|
||||
]
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="description">
|
||||
<paragraph pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="183B05E4F43EDC6CF00BF1F028B6B4D1" pageId="0" pageNumber="257" type="description">
|
||||
<paragraph id="0D2C31B573263A1D0CAE2C8A16B3AC0A" pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph id="CD73AB0DAB13B7B1344ED71F88F52526" pageId="0" pageNumber="257">
|
||||
small in size. Male (Fig.
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upperside, on the right show the underside. A - C Pieris narina stat. rev. Verity, 1908. D - I Pieris mihon Yakovlev, 2006 stat. nov. J - M Pieris dulcinea dulcinea (Butler, 1882). Scale bar represents 10 mm. (L, M reference Yakovlev, 2006)." figureDoi="10.3897/asp.81.e85191.figure9" httpUri="https://binary.pensoft.net/fig/825204" pageId="0" pageNumber="257">9A</figureCitation>
|
||||
<figureCitation id="704B88222FECE169064D4A4EFDF691D2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upperside, on the right show the underside. A - C Pieris narina stat. rev. Verity, 1908. D - I Pieris mihon Yakovlev, 2006 stat. nov. J - M Pieris dulcinea dulcinea (Butler, 1882). Scale bar represents 10 mm. (L, M reference Yakovlev, 2006)." figureDoi="10.3897/asp.81.e85191.figure9" httpUri="https://binary.pensoft.net/fig/825204" pageId="0" pageNumber="257">9A</figureCitation>
|
||||
): both wings white on the upperside and pale yellowish on the underside. Costal margin black; apical spot blackish, triangular, extending along veins to outer margin and sometimes reached vein CuA2. Outer margin border of forewing blackish. The 1st discal spot and outer spot variable, distinct to nearly absent on both upperside and underside. The 2nd discal spot absent or only faintly indicated on the underside and the 3rd discal spot absent or sometimes very vague on the upperside. Outer margin border of hindwings grey to blackish, sometimes triangularly extending along veins. Female (Fig.
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upperside, on the right show the underside. A - C Pieris narina stat. rev. Verity, 1908. D - I Pieris mihon Yakovlev, 2006 stat. nov. J - M Pieris dulcinea dulcinea (Butler, 1882). Scale bar represents 10 mm. (L, M reference Yakovlev, 2006)." figureDoi="10.3897/asp.81.e85191.figure9" httpUri="https://binary.pensoft.net/fig/825204" pageId="0" pageNumber="257">9B, C</figureCitation>
|
||||
<figureCitation id="2818A701C9C6AB805FDC5CA66A19E0C2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of P. napi - complex species occurring in China. Composite pictures on the left show the upperside, on the right show the underside. A - C Pieris narina stat. rev. Verity, 1908. D - I Pieris mihon Yakovlev, 2006 stat. nov. J - M Pieris dulcinea dulcinea (Butler, 1882). Scale bar represents 10 mm. (L, M reference Yakovlev, 2006)." figureDoi="10.3897/asp.81.e85191.figure9" httpUri="https://binary.pensoft.net/fig/825204" pageId="0" pageNumber="257">9B, C</figureCitation>
|
||||
): both wings white to pale yellowish with intensive brownish or grey suffusion on the upperside while yellowish on the underside, but clearly in the apical 1/3 of discocell on the upperside of forewing. Hindwing with dark powder strongly developed along veins.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/asp.81.e85191.figure9" httpUri="https://binary.pensoft.net/fig/825204" pageId="0" pageNumber="257" start="Figure 9" startId="F9">
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">Figure 9.</emphasis>
|
||||
<caption id="5CE23A3351695675C5BC739826721C8B" doi="10.3897/asp.81.e85191.figure9" httpUri="https://binary.pensoft.net/fig/825204" pageId="0" pageNumber="257" start="Figure 9" startId="F9">
|
||||
<paragraph id="FA67A0FED1F0B4185DDC4ECD2551C496" pageId="0" pageNumber="257">
|
||||
<emphasis id="CCF94280677308A45B7A3DB58466196A" bold="true" pageId="0" pageNumber="257">Figure 9.</emphasis>
|
||||
Habitus of
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
<taxonomicName id="D5570A59890C1E2A2D80787C6A60F413" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="C30B67A6DC44B1276A6A4A8F2709F124" italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
</taxonomicName>
|
||||
-complex species occurring in China. Composite pictures on the left show the upperside, on the right show the underside.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">A</emphasis>
|
||||
<emphasis id="1994E6C5086A433A93134CE6B5D4C16B" bold="true" pageId="0" pageNumber="257">A</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">C</emphasis>
|
||||
<taxonomicName authorityName="Verity" authorityYear="1908" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="narina">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris narina</emphasis>
|
||||
<emphasis id="72D89F924DAFE6C8B4C6EF48CDB4156A" bold="true" pageId="0" pageNumber="257">C</emphasis>
|
||||
<taxonomicName id="3C5BADECF5A1B29EDFD46A04FB34A1C3" authorityName="Verity" authorityYear="1908" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="narina">
|
||||
<emphasis id="F1753EF4BF5EAA8A6AC62B4EE93F0E1F" italics="true" pageId="0" pageNumber="257">Pieris narina</emphasis>
|
||||
</taxonomicName>
|
||||
stat. rev. Verity, 1908.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">D</emphasis>
|
||||
<emphasis id="F26D3AAEFDF30C48F13223430FA3B0CB" bold="true" pageId="0" pageNumber="257">D</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">I</emphasis>
|
||||
<taxonomicName authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
<emphasis id="740EBA9A163C59BF40372472B487A339" bold="true" pageId="0" pageNumber="257">I</emphasis>
|
||||
<taxonomicName id="2C3BFDAFDAE3AFA6ABD607310A8608EA" authorityName="Yakovlev" authorityYear="2006" baseAuthorityName="Ge & Jiang & Wang & Song & Zhang & Hu" baseAuthorityYear="2023" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris mihon" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="mihon">
|
||||
<emphasis id="8B4C2BBA991A43B78C6C82F471AECCBB" italics="true" pageId="0" pageNumber="257">Pieris mihon</emphasis>
|
||||
</taxonomicName>
|
||||
Yakovlev, 2006 stat. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">J</emphasis>
|
||||
<emphasis id="D2D907A4425685A3197EF54BEA6B6F6A" bold="true" pageId="0" pageNumber="257">J</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">M</emphasis>
|
||||
<taxonomicName authority="dulcinea" authorityName="dulcinea (Butler" authorityYear="1882" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris dulcinea subsp. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="dulcinea" subSpecies="dulcinea">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris dulcinea dulcinea</emphasis>
|
||||
<emphasis id="824E8A7C8EDAE3CCEE0627A422FEACB2" bold="true" pageId="0" pageNumber="257">M</emphasis>
|
||||
<taxonomicName id="4C4FF28817793E14A87A801C33101ABB" authority="dulcinea" authorityName="dulcinea (Butler" authorityYear="1882" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris dulcinea subsp. dulcinea" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="dulcinea" subSpecies="dulcinea">
|
||||
<emphasis id="4BC64F0873982C5FB2284E895BBA85E0" italics="true" pageId="0" pageNumber="257">Pieris dulcinea dulcinea</emphasis>
|
||||
</taxonomicName>
|
||||
(Butler, 1882). Scale bar represents 10 mm. (L, M reference Yakovlev, 2006).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Kazakhstan, Kyrgyzstan, Tajikistan (Northern and Inner Tian Shan, Dzhngarsky Alatau), China (Xinjiang)</paragraph>
|
||||
<subSubSection id="BC5CE450B2D5873468DCE56D347962E6" pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph id="9EC6771136B6CFC37CB32E8EB11A2A56" pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph id="C39E4CD689A685EB635CA2B372F68922" pageId="0" pageNumber="257">Kazakhstan, Kyrgyzstan, Tajikistan (Northern and Inner Tian Shan, Dzhngarsky Alatau), China (Xinjiang)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Probably univoltine, based on a single form of adults observed in July.</paragraph>
|
||||
<subSubSection id="91FC1E525C0EA1E1029861978BBF4A9D" pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph id="BA96ECF7786C320E3CF0FFA4E5111B06" pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph id="B4CF75FE4295108652FD3908DA6AEA99" pageId="0" pageNumber="257">Probably univoltine, based on a single form of adults observed in July.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="C7886EA5C231371429CEA1C600D906B4" pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph id="783BC4C7F1D046D24D76DE0AB3959B12" pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph id="5DF4A1270170C47E9E0F543805C6BD6F" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5H</figureCitation>
|
||||
<figureCitation id="A11A73229853BEF854B5388EAB215E36" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5H</figureCitation>
|
||||
) terminal part of tegumen with pairs of sickle shaped convex; uncus broad with its basal part as wide as terminal part of tegumen, apical half of uncus digitation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="E432DF8EE77311A7754B483C422B78F3" pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph id="ED0C070C8B0638AED11488542C4BA7B5" pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph id="28F43A0ACEC5B91F7562E2F7C8F0CC98" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of P. napi - complex species. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure6" httpUri="https://binary.pensoft.net/fig/825201" pageId="0" pageNumber="257">6D</figureCitation>
|
||||
<figureCitation id="11E2DB35E60373ACC0E98B977A77DD21" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of P. napi - complex species. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure6" httpUri="https://binary.pensoft.net/fig/825201" pageId="0" pageNumber="257">6D</figureCitation>
|
||||
) posterior apophysis short and slender; inner distal of sterigma bilobate lobe shaped, extend to the center; inner basal of sterigma with a distinct tooth near the basal part, acute-angled connected at the basal margin; signum relatively broad cordiform shaped, moderately tapering to the base.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="8DC7E3280DB7D47D6186A94754A5FB2E" pageId="0" pageNumber="257" type="notes">
|
||||
<paragraph id="9C6E32895C05B71F2B7AE456B5C9EF92" pageId="0" pageNumber="257">Note.</paragraph>
|
||||
<paragraph id="0B85A1EC35FA01C2126A1AC654EBCAAB" pageId="0" pageNumber="257">
|
||||
The species rather resembles
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="ochsenheimeri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
<taxonomicName id="75B849DBAE4EBB3948B18CEF632DCE90" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="ochsenheimeri">
|
||||
<emphasis id="73D58924016C6F308BAEAD4C8098E78F" italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
</taxonomicName>
|
||||
, but can be distinguished by comparing multiple morphological characteristics (see notes in
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="ochsenheimeri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
<taxonomicName id="582423060C0BE242BD86E25A97F7CB35" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="ochsenheimeri">
|
||||
<emphasis id="64411DED5F4079BC88E4E985477B0EC8" italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
</taxonomicName>
|
||||
); the blackish outer marginal border on both wings and the relatively smaller size make it easily separated from other species of the
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="napi (Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris napi</emphasis>
|
||||
<taxonomicName id="C648281E5CBE8BAD524E7B09239BC239" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" baseAuthorityName="napi (Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="803AF188AF5683E6C40139D190C0AEF8" italics="true" pageId="0" pageNumber="257">Pieris napi</emphasis>
|
||||
</taxonomicName>
|
||||
-complex distributed in the same region. This taxon was formerly elevated by
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="10C81CC11DE3885084E0CB753084FA04" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
to species level while
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="6DE560D037ACC534ACF38F1ED93050CD" author="Tadokoro, T" journalOrPublisher="Butterflies" pageId="0" pageNumber="257" pagination="20 - 35" refId="B22" refString="Tadokoro, T, Shinkawa, T, Wang, M, 2014. Primary study of Pieris napi --group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution. Butterflies 65: 20 - 35" title="Primary study of Pieris napi -- group in East Asia (Part II): Phylogenetic analysis, morphological characteristics and geographical distribution." volume="65" year="2014">Tadokoro et al. (2014)</bibRefCitation>
|
||||
regarded it as a subspecies of
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="ochsenheimeri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
<taxonomicName id="A5EE545C12B5F9C2A60AED3B2E727519" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="ochsenheimeri">
|
||||
<emphasis id="75661117CDC0D07556F3337B031F0FA3" italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
</taxonomicName>
|
||||
. However, this taxon has unique morphological characteristics and a previous study has shown that there is deep differentiation between
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" kingdom="Animalia" lsidName="P. narina" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="narina">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. narina</emphasis>
|
||||
<taxonomicName id="8FEABB8ED45846F996107D16CE9483DD" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. narina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="narina">
|
||||
<emphasis id="BEB357785E0521F5533C37C885843549" italics="true" pageId="0" pageNumber="257">P. narina</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" family="Pieridae" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="ochsenheimeri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
<taxonomicName id="9C9948B8EFE5214EE24E64C689B5CE4C" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. ochsenheimeri" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="ochsenheimeri">
|
||||
<emphasis id="AAECA71E1FD539764B0B92B8DE1D2EEC" italics="true" pageId="0" pageNumber="257">P. ochsenheimeri</emphasis>
|
||||
</taxonomicName>
|
||||
for the COI gene (
|
||||
<bibRefCitation DOI="https://doi.org/10.1111/j.1755-0998.2009.02577.x" author="Lukhtanov, VA" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="257" pagination="1302 - 1310" refId="B13" refString="Lukhtanov, VA, Sourakov, A, Zakharov, EV, Hebert, PD, 2009. DNA barcoding Central Asian butterflies: increasing geographical dimension does not significantly reduce the success of species identification. Molecular Ecology Resources 9 (5): 1302 - 1310, DOI: https://doi.org/10.1111/j.1755-0998.2009.02577.x" title="DNA barcoding Central Asian butterflies: increasing geographical dimension does not significantly reduce the success of species identification." url="https://doi.org/10.1111/j.1755-0998.2009.02577.x" volume="9" year="2009">Lukhtanov et al. 2009</bibRefCitation>
|
||||
<bibRefCitation id="4664ED8E818DC39967F598D00D71F418" DOI="https://doi.org/10.1111/j.1755-0998.2009.02577.x" author="Lukhtanov, VA" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="257" pagination="1302 - 1310" refId="B13" refString="Lukhtanov, VA, Sourakov, A, Zakharov, EV, Hebert, PD, 2009. DNA barcoding Central Asian butterflies: increasing geographical dimension does not significantly reduce the success of species identification. Molecular Ecology Resources 9 (5): 1302 - 1310, DOI: https://doi.org/10.1111/j.1755-0998.2009.02577.x" title="DNA barcoding Central Asian butterflies: increasing geographical dimension does not significantly reduce the success of species identification." url="https://doi.org/10.1111/j.1755-0998.2009.02577.x" volume="9" year="2009">Lukhtanov et al. 2009</bibRefCitation>
|
||||
). Our phylogenetic results also indicated that this species is more closely related to
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. bryoniae" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="bryoniae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. bryoniae</emphasis>
|
||||
<taxonomicName id="44AEE0ECE6BC08417069FF2B6E43E286" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bryoniae" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="bryoniae">
|
||||
<emphasis id="F3A0D3C91AEFD578E1CBD60E5F26FC78" italics="true" pageId="0" pageNumber="257">P. bryoniae</emphasis>
|
||||
</taxonomicName>
|
||||
than to
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
<taxonomicName id="BF2B95A991B3B438BA5A630421887BA5" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="CD634EE2FB81DEF2DA395BDB4B0D56B7" italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
</taxonomicName>
|
||||
or
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. ochsenhelmeri" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="ochsenhelmeri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. ochsenhelmeri</emphasis>
|
||||
<taxonomicName id="EE77800212E96A6D5F54D4371E43AEEA" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. ochsenhelmeri" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="ochsenhelmeri">
|
||||
<emphasis id="99F493F8421F936D6B7AA003C5FC7638" italics="true" pageId="0" pageNumber="257">P. ochsenhelmeri</emphasis>
|
||||
</taxonomicName>
|
||||
, thus we confirmed it as a distinct species.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,196 +1,196 @@
|
|||
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<mods:namePart>Ge, Si Xun</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3769-1530</mods:nameIdentifier>
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<mods:affiliation>Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China.</mods:affiliation>
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<mods:namePart id="D871A035B2C41C5201066A970A712267">Ge, Si Xun</mods:namePart>
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<mods:affiliation id="7E7CEE2CCC2B87D35D67011F5A4160F5">Beijing Key Laboratory for Forest Pest Control, Beijing Forestry University, Beijing 100083, P. R. China.</mods:affiliation>
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<mods:namePart>Jiang, Zhuo Heng</mods:namePart>
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<mods:affiliation>School of Science, Westlake University, Hangzhou, China</mods:affiliation>
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<mods:namePart id="4773DC9CBD01052EDED7D7977FD82AF4">Jiang, Zhuo Heng</mods:namePart>
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<mods:affiliation id="C7D6BDA7F55F3E7DB3B58DFE896483FE">School of Science, Westlake University, Hangzhou, China</mods:affiliation>
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</mods:role>
|
||||
<mods:namePart>Wang, Jia Qi</mods:namePart>
|
||||
<mods:affiliation>2288 Long, Hongxin Rd, Minhang District, Shanghai, China</mods:affiliation>
|
||||
<mods:namePart id="EF3E2779E47D6256E465035E5756B114">Wang, Jia Qi</mods:namePart>
|
||||
<mods:affiliation id="9AD9EEFEABEBB13CBE95A3F738E41AC7">2288 Long, Hongxin Rd, Minhang District, Shanghai, China</mods:affiliation>
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|
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</mods:role>
|
||||
<mods:namePart>Song, Kui</mods:namePart>
|
||||
<mods:affiliation>School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China</mods:affiliation>
|
||||
<mods:namePart id="02C83C2B50B85F558A8519A0DA79A7EA">Song, Kui</mods:namePart>
|
||||
<mods:affiliation id="8CA994BC0BE9FE814D00B6263D610286">School of Economic and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining 810007, Qinghai, P. R. China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
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<mods:role id="6B57F70FEFAFF97A435743B3183DB92F">
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<mods:roleTerm id="97041CE002DF142A47E086F0CF11ADB9">Author</mods:roleTerm>
|
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</mods:role>
|
||||
<mods:namePart>Zhang, Chao</mods:namePart>
|
||||
<mods:affiliation>Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China</mods:affiliation>
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<mods:namePart id="A1ABF52E255B27436E9AC2D5BC2C06AD">Zhang, Chao</mods:namePart>
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<mods:affiliation id="0A44DA5C30FAF6594C9FA8011BB39F82">Simianshan Forest Resource Service Center, Jiangjin District, Chongqing, 402296, P. R. China</mods:affiliation>
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<mods:namePart>Hu, Shao Ji</mods:namePart>
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<mods:affiliation>Institute of International Rivers and Eco-security, Yunnan University, Kunming, China</mods:affiliation>
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<mods:nameIdentifier type="email">shaojihu@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="23367AD037B0F8E8174A9CF0D99AE535">Hu, Shao Ji</mods:namePart>
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<mods:affiliation id="FE54E39FC03BA70EAED668143EB1DEA1">Institute of International Rivers and Eco-security, Yunnan University, Kunming, China</mods:affiliation>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:title id="16C8ABE180709C5236C187A5A72465B0">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date>2023</mods:date>
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<treatment LSID="urn:lsid:plazi:treatment:7789CF7BC6545583A4D46C09B92D9673" httpUri="http://treatment.plazi.org/id/7789CF7BC6545583A4D46C09B92D9673" lastPageNumber="257" pageId="0" pageNumber="257">
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<taxonomicName LSID="7789CF7B-C654-5583-A4D4-6C09B92D9673" authority="Roeber, 1907" authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">
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<treatment id="7789CF7BC6545583A4D46C09B92D9673" ID-GBIF-Taxon="209450347" LSID="urn:lsid:plazi:treatment:7789CF7BC6545583A4D46C09B92D9673" httpUri="http://treatment.plazi.org/id/7789CF7BC6545583A4D46C09B92D9673" lastPageNumber="257" pageId="0" pageNumber="257" scope_family="Pieridae" scope_order="Lepidoptera">
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<taxonomicName id="D8D3CE8D30B2489CDF70DADD4B1D2718" LSID="7789CF7B-C654-5583-A4D4-6C09B92D9673" authority="Roeber, 1907" authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">
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Pieris melaina
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<normalizedToken originalValue="Röber">Roeber</normalizedToken>
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<normalizedToken id="B007D15ABA2A4714CF923B5AA61E73DB" originalValue="Röber">Roeber</normalizedToken>
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, 1907
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</taxonomicName>
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<taxonomicName authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">Pieris melaina</taxonomicName>
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<normalizedToken originalValue="Röber">Roeber</normalizedToken>
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<paragraph id="806110A52474F37316B0AA9FCCADE583" pageId="0" pageNumber="257">
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<taxonomicName id="3EA20D6042B45543449B5779FB71470A" authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">Pieris melaina</taxonomicName>
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<normalizedToken id="2574C18512DC701234D9F90293E8F444" originalValue="Röber">Roeber</normalizedToken>
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, 1907; Gross-Schmett. Erde 1: 48, pl. 20 g; TL:
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<normalizedToken originalValue="“Tibet”">"Tibet"</normalizedToken>
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<normalizedToken id="1F4DFD707955E80A56832AAF7643326E" originalValue="“Tibet”">"Tibet"</normalizedToken>
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(original description)
|
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</paragraph>
|
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<paragraph pageId="0" pageNumber="257">
|
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<taxonomicName authority="melaina" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina subsp. melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="melaina" subSpecies="melaina">Pieris melaina melaina</taxonomicName>
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<normalizedToken originalValue="Röber">Roeber</normalizedToken>
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<paragraph id="FDE4246056C8B9434306C6F3DFEF9E74" pageId="0" pageNumber="257">
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<taxonomicName id="7C2FBE4A8E90686186CFB41B13A36D3D" authority="melaina" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina subsp. melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="melaina" subSpecies="melaina">Pieris melaina melaina</taxonomicName>
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<normalizedToken id="77EADA14B3E83B522BDAFF7A569603AF" originalValue="Röber">Roeber</normalizedToken>
|
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, 1907;
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
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<bibRefCitation id="06AAA471C44C7FA362E7F9DB8DE36E63" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
: Herbipoliana 1(1-2): 406 (pictured)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authority="melete" authorityName="melete Menetries" authorityYear="1857" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete subsp. melete" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="melete" subSpecies="melete">Pieris melete melete</taxonomicName>
|
||||
<paragraph id="BCE12033A7DD010A0DA9BAFF630897C4" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="84E73E32FE73377DD8D3FBD86EC42AD0" authority="melete" authorityName="melete Menetries" authorityYear="1857" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete subsp. melete" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="melete" subSpecies="melete">Pieris melete melete</taxonomicName>
|
||||
ra.
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">montana</emphasis>
|
||||
<emphasis id="90659DCBCA7E910EE4F79F563356EBF9" italics="true" pageId="0" pageNumber="257">montana</emphasis>
|
||||
Verity, 1908;
|
||||
<bibRefCitation author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
<bibRefCitation id="9E387DE9D495836A8CF81601E4ADBC3A" author="Eitschberger, U" journalOrPublisher="Advances in Ecological Research" pageId="0" pageNumber="257" refId="B10" refString="Eitschberger, U, 1983. Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana 1(1): I-XXII, 1-504; 1(2): 1-601." title="Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s. l.). Herbipoliana 1 (1): I-XXII, 1 - 504; 1 (2): 1 - 601." year="1983">Eitschberger (1983)</bibRefCitation>
|
||||
: Herbipoliana 1(2): 406-408 (as a synonym of
|
||||
<taxonomicName family="Pieridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. melaina" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="melaina">P. melaina</taxonomicName>
|
||||
<taxonomicName id="13387D19B53FBF526DB7768249DEEDE6" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">P. melaina</taxonomicName>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<taxonomicName authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">Pieris melaina</taxonomicName>
|
||||
<normalizedToken originalValue="Röber">Roeber</normalizedToken>
|
||||
<paragraph id="57B2D72428B762B4B2BC86744197CDF9" pageId="0" pageNumber="257">
|
||||
<taxonomicName id="EEF944EC23915CF67F8A296FBCACB081" authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">Pieris melaina</taxonomicName>
|
||||
<normalizedToken id="2964780F082983864DB180368B44FFCF" originalValue="Röber">Roeber</normalizedToken>
|
||||
, 1907;
|
||||
<bibRefCitation author="Tadokoro, T" journalOrPublisher="Lepidoptera Science" pageId="0" pageNumber="257" pagination="81 - 91" refId="B23" refString="Tadokoro, T, Wangchuk, S, Wangdi, S, Wangdi, K, Sherub, , Wangdi, R, Drukupa, S, Harada, M, Saito, M, Aoki, T, Yamaguchi, S, Igarashi, Y, Watanabe, Y, Yago, M, 2017. Description of a new subspecies of Pieris erutae Poujade, 1888 from Eastern Bhutan, with taxonomic notes on the Pieris napi group from the Himalayas (Lepidoptera, Pieridae). Lepidoptera Science 68 (3-4): 81 - 91" title="Description of a new subspecies of Pieris erutae Poujade, 1888 from Eastern Bhutan, with taxonomic notes on the Pieris napi group from the Himalayas (Lepidoptera, Pieridae)." volume="68" year="2017">Tadokoro et al. (2017)</bibRefCitation>
|
||||
<bibRefCitation id="632AAEC6CFEC2627F5585E9BFC8D7B9C" author="Tadokoro, T" journalOrPublisher="Lepidoptera Science" pageId="0" pageNumber="257" pagination="81 - 91" refId="B23" refString="Tadokoro, T, Wangchuk, S, Wangdi, S, Wangdi, K, Sherub, , Wangdi, R, Drukupa, S, Harada, M, Saito, M, Aoki, T, Yamaguchi, S, Igarashi, Y, Watanabe, Y, Yago, M, 2017. Description of a new subspecies of Pieris erutae Poujade, 1888 from Eastern Bhutan, with taxonomic notes on the Pieris napi group from the Himalayas (Lepidoptera, Pieridae). Lepidoptera Science 68 (3-4): 81 - 91" title="Description of a new subspecies of Pieris erutae Poujade, 1888 from Eastern Bhutan, with taxonomic notes on the Pieris napi group from the Himalayas (Lepidoptera, Pieridae)." volume="68" year="2017">Tadokoro et al. (2017)</bibRefCitation>
|
||||
:
|
||||
<taxonomicName authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="257" rank="order">Lepidoptera</taxonomicName>
|
||||
<taxonomicName id="70D7FFE32DF17813A08A410B8ADFEFB1" authorityName="Ge & Jiang & Wang & Song & Zhang & Hu" authorityYear="2023" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
|
||||
Science: 68(3-4): 81-91 (pictured)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="description">
|
||||
<paragraph pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="570D7AD5AEE06E4E1B4B0495CB96CFCE" pageId="0" pageNumber="257" type="description">
|
||||
<paragraph id="E30134861EDBD95F3441F725AC99637E" pageId="0" pageNumber="257">Description.</paragraph>
|
||||
<paragraph id="E6CD46C0E0C3E9603393A421065D19FA" pageId="0" pageNumber="257">
|
||||
Both wings whitish on the upperside while pale yellowish on the underside. Male (Fig.
|
||||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13A, C</figureCitation>
|
||||
<figureCitation id="8827843630A7F706CF39CE7E8A8B7565" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13A, C</figureCitation>
|
||||
): apical spot blackish and sickle shaped. The 1st discal spot distinct and almost combined with the apical spot, the 2nd discal spot absent or faintly developed on the upperside while distinct on the underside. The outer spot distinct on the upperside while absent on the underside. Female (Fig.
|
||||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13B, D</figureCitation>
|
||||
<figureCitation id="8DCFC353F5846A7A35DDF990FB662C0E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of species similar to the P. napi - complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside. A - D Pieris melaina Roeber, 1907. E - F Pieris extensa Poujade, 1888. G - H Pieris chumbiensis gyantsensis Verity, 1911. I - L Pieris melete Menetries, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)" figureDoi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257">13B, D</figureCitation>
|
||||
): extremely dark powdered along veins on the upperside, with the 1st and 2nd discal spot merging with the apical spot, thus brownish suffusion on almost all outer marginal area except slightly whitish area in the space between vein CuA2 and 2A.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257" start="Figure 13" startId="F13">
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">Figure 13.</emphasis>
|
||||
<caption id="C3A13725DF685021F3E1994329932B5B" doi="10.3897/asp.81.e85191.figure13" httpUri="https://binary.pensoft.net/fig/825208" pageId="0" pageNumber="257" start="Figure 13" startId="F13">
|
||||
<paragraph id="0CD1029E1317B7EF1F16789D684D9615" pageId="0" pageNumber="257">
|
||||
<emphasis id="2F7F4A0AF79073A438465DECD4BFBDEA" bold="true" pageId="0" pageNumber="257">Figure 13.</emphasis>
|
||||
Habitus of species similar to the
|
||||
<taxonomicName family="Pieridae" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" rank="species" species="napi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
<taxonomicName id="769CB4A89D52E413EB68BE71399EC5CB" class="Insecta" family="Pieridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. napi" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="napi">
|
||||
<emphasis id="CBDE8C9EE46F044FB68AE3D6AB9CCD0F" italics="true" pageId="0" pageNumber="257">P. napi</emphasis>
|
||||
</taxonomicName>
|
||||
-complex, distributed in China. Composite pictures on the left show the upperside, on the right show the underside.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">A</emphasis>
|
||||
<emphasis id="D90AE09AA21588B709AA05CFB092967B" bold="true" pageId="0" pageNumber="257">A</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">D</emphasis>
|
||||
<taxonomicName authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris melaina</emphasis>
|
||||
<emphasis id="40ED978490985E62B786E2EF86BFC7CC" bold="true" pageId="0" pageNumber="257">D</emphasis>
|
||||
<taxonomicName id="FC63A43B54E12083CE852D3D8F078812" authorityName="Roeber" authorityYear="1907" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melaina" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melaina">
|
||||
<emphasis id="E477433D12855E674E7803622F17B835" italics="true" pageId="0" pageNumber="257">Pieris melaina</emphasis>
|
||||
</taxonomicName>
|
||||
<normalizedToken originalValue="Röber">Roeber</normalizedToken>
|
||||
<normalizedToken id="066891503A2FECCD9019BA2FC40E5A82" originalValue="Röber">Roeber</normalizedToken>
|
||||
, 1907.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">E</emphasis>
|
||||
<emphasis id="02C029C9ACFF7CAD278BDCB9B7CDFCB6" bold="true" pageId="0" pageNumber="257">E</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">F</emphasis>
|
||||
<taxonomicName authorityName="Poujade" authorityYear="1888" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris extensa" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="extensa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris extensa</emphasis>
|
||||
<emphasis id="95859D1C12DC2C780E977F083A6253EB" bold="true" pageId="0" pageNumber="257">F</emphasis>
|
||||
<taxonomicName id="362167CD88A32CCB5FD452A0227F811D" authorityName="Poujade" authorityYear="1888" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris extensa" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="extensa">
|
||||
<emphasis id="4DF12CBDBEA20940B0F95538D2FD4F76" italics="true" pageId="0" pageNumber="257">Pieris extensa</emphasis>
|
||||
</taxonomicName>
|
||||
Poujade, 1888.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">G</emphasis>
|
||||
<emphasis id="A302FD1D66A23CE5C3CF227C23D3BD4B" bold="true" pageId="0" pageNumber="257">G</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">H</emphasis>
|
||||
<taxonomicName authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris chumbiensis subsp. gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="chumbiensis" subSpecies="gyantsensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris chumbiensis gyantsensis</emphasis>
|
||||
<emphasis id="E860F23F0E551D1AF37036EF205B85F5" bold="true" pageId="0" pageNumber="257">H</emphasis>
|
||||
<taxonomicName id="3C7020BD250F0F21564906FDCF7CBC9F" authorityName="Verity" authorityYear="1911" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris chumbiensis subsp. gyantsensis" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="subSpecies" species="chumbiensis" subSpecies="gyantsensis">
|
||||
<emphasis id="040ADD611BD5DAE320465A80C5FC1D19" italics="true" pageId="0" pageNumber="257">Pieris chumbiensis gyantsensis</emphasis>
|
||||
</taxonomicName>
|
||||
Verity, 1911.
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">I</emphasis>
|
||||
<emphasis id="27D91E3E7AD4F8ED872FF5411303C5C7" bold="true" pageId="0" pageNumber="257">I</emphasis>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="257">L</emphasis>
|
||||
<taxonomicName authorityName="Menetries" authorityYear="1857" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melete">
|
||||
<emphasis italics="true" pageId="0" pageNumber="257">Pieris melete</emphasis>
|
||||
<emphasis id="04F55B39C5C2CEF01782FF9B0440B34F" bold="true" pageId="0" pageNumber="257">L</emphasis>
|
||||
<taxonomicName id="AC04578D7D7FC6BC6A2502C65D8B08FC" authorityName="Menetries" authorityYear="1857" class="Insecta" family="Pieridae" genus="Pieris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pieris melete" order="Lepidoptera" pageId="0" pageNumber="257" phylum="Arthropoda" rank="species" species="melete">
|
||||
<emphasis id="3D65F0725108736C0328FB1B102DA3F4" italics="true" pageId="0" pageNumber="257">Pieris melete</emphasis>
|
||||
</taxonomicName>
|
||||
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
|
||||
<normalizedToken id="E042DF9B7100C57394DC9BACBA71E5D6" originalValue="Ménétriés">Menetries</normalizedToken>
|
||||
, 1857. Scale bar represents 10 mm. (C, D reference Eitschberger, 1983)
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">East Tibet (Yadong, Bomi), India (border to Tibet)</paragraph>
|
||||
<subSubSection id="924C3AA6B937588A1F6F86999AC9CE5D" pageId="0" pageNumber="257" type="distribution">
|
||||
<paragraph id="92487C1B64BC19A88D24B72A8EA00A06" pageId="0" pageNumber="257">Distribution.</paragraph>
|
||||
<paragraph id="F961AEA3F28F558DE1379ADBC1246193" pageId="0" pageNumber="257">East Tibet (Yadong, Bomi), India (border to Tibet)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">Univoltine in high altitude habitat (occurs in July and August above 4000 m) while probably bivoltine in low altitude habitat (occurs in April and July below 3000 m).</paragraph>
|
||||
<subSubSection id="E12E5A6DC21ABCA734B837A5348FBB7F" pageId="0" pageNumber="257" type="phenology">
|
||||
<paragraph id="72F16B61EAD67A46D8AE1C57E4B82382" pageId="0" pageNumber="257">Phenology.</paragraph>
|
||||
<paragraph id="EEB2F54A747C49D4DD9DB83FC0BF5A6C" pageId="0" pageNumber="257">Univoltine in high altitude habitat (occurs in July and August above 4000 m) while probably bivoltine in low altitude habitat (occurs in April and July below 3000 m).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="701C0A69641E7762AE3712A2C5582B60" pageId="0" pageNumber="257" type="male genitalia">
|
||||
<paragraph id="E5AF0CD91BD16FD5926F2E9996FFFBE0" pageId="0" pageNumber="257">Male genitalia.</paragraph>
|
||||
<paragraph id="DA3ACD6AC7F519158F88B5FFCCB9A431" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5A</figureCitation>
|
||||
<figureCitation id="03CDBCD13A4206EF60DC5C8AD533FDDE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tegumen and uncus of male genitalia in dorsal view. Scale bar represents 1 mm." figureDoi="10.3897/asp.81.e85191.figure5" httpUri="https://binary.pensoft.net/fig/825200" pageId="0" pageNumber="257">5A</figureCitation>
|
||||
) tegumen broad, distinctly convex on its 4/5 apically; basal part of uncus moderately convex and with its apically distinctly narrowed towards tip.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="257">
|
||||
<subSubSection id="067F6E4EA025C5A83476CA3A0163BE82" pageId="0" pageNumber="257" type="female genitalia">
|
||||
<paragraph id="7CC0AD2DF71BDF6D6DAEC2B2C9032513" pageId="0" pageNumber="257">Female genitalia.</paragraph>
|
||||
<paragraph id="221870C0FF08F607EF77AD2FCE71B85A" pageId="0" pageNumber="257">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of species similar to P. napi - complex distributed in China. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure8" httpUri="https://binary.pensoft.net/fig/825203" pageId="0" pageNumber="257">8A</figureCitation>
|
||||
<figureCitation id="F0800CBDAFCA469B5700972D80EAD54E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of species similar to P. napi - complex distributed in China. Scale bar represents 0.5 mm." figureDoi="10.3897/asp.81.e85191.figure8" httpUri="https://binary.pensoft.net/fig/825203" pageId="0" pageNumber="257">8A</figureCitation>
|
||||
) sterigma with its inner distal bilobate lobe-shaped, shallowly expended to the centre; inner basal of sterigma arcuately protruding, smoothly connected at the basal margin; signum variable, cordiform to near cordiform-shaped.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,156 +1,156 @@
|
|||
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<treatment ID-GBIF-Taxon="158995544" LSID="urn:lsid:plazi:treatment:7B3FDEB746D75239A167247A824B06E0" httpUri="http://treatment.plazi.org/id/7B3FDEB746D75239A167247A824B06E0" lastPageNumber="19" pageId="0" pageNumber="19">
|
||||
<subSubSection pageId="0" pageNumber="19" type="nomenclature">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName LSID="7b3fdeb7-46d7-5239-a167-247a824b06e0" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">Madaglymbus apicalis (Fairmaire, 1898)</taxonomicName>
|
||||
<taxonomicNameLabel pageId="0" pageNumber="19">comb. nov.</taxonomicNameLabel>
|
||||
<treatment id="7B3FDEB746D75239A167247A824B06E0" ID-GBIF-Taxon="158995544" LSID="urn:lsid:plazi:treatment:7B3FDEB746D75239A167247A824B06E0" httpUri="http://treatment.plazi.org/id/7B3FDEB746D75239A167247A824B06E0" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
|
||||
<subSubSection id="0CFE7C4CA36D52050302B4E008F94FB6" pageId="0" pageNumber="19" type="nomenclature">
|
||||
<paragraph id="2AC1580FFC55A86F9CDB8D628D5E37BA" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="36D88C3FA2CD98A3F7F967932EA6B13E" ID-CoL="8LZGS" LSID="7b3fdeb7-46d7-5239-a167-247a824b06e0" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">Madaglymbus apicalis (Fairmaire, 1898)</taxonomicName>
|
||||
<taxonomicNameLabel id="1738826B11521B928ADDEAB8ECB4318F" pageId="0" pageNumber="19">comb. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">Copelatus apicalis</taxonomicName>
|
||||
<subSubSection id="C399D134FCEF4A30B5CB9B3F15C987E6" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="09CDD22BDD5C9FF5DABCE70837B60030" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="9E46BDB0D25FC839103BF32945A86A89" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">Copelatus apicalis</taxonomicName>
|
||||
Fairmaire, 1898: 465.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Madagascar, Suberbieville [= Maevatanana].</paragraph>
|
||||
<subSubSection id="9AF150F52528A545BF763FF461EBB10C" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="3EADDD0BF73C9F6969B4D3FE8DF697DE" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="C903C6DFB057609B0051407FDB877E8B" pageId="0" pageNumber="19">Madagascar, Suberbieville [= Maevatanana].</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Betsiboka: Maevatanana</emphasis>
|
||||
<subSubSection id="9DC8943740D69883A2694A26CFA74CC9" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="B9A4B9498D2F8E8106887B10F38E0EC5" pageId="0" pageNumber="19">Type material examined.</paragraph>
|
||||
<paragraph id="2505764D0DDD2B5A7E2C59A84B365B50" pageId="0" pageNumber="19">
|
||||
<emphasis id="56637FF28EDB18130D0062F6D7F5CFA1" bold="true" pageId="0" pageNumber="19">Mahajanga. Betsiboka: Maevatanana</emphasis>
|
||||
: -ST ♂ (GP) (MNHN, "coll.
|
||||
<normalizedToken originalValue="Régimbart”">Regimbart"</normalizedToken>
|
||||
<normalizedToken id="CF57CA27937B2D46C4914E2FA8871A3D" originalValue="Régimbart”">Regimbart"</normalizedToken>
|
||||
): // Data in NHRS | JLKB | 000030208 // Madag. Perrier | // Type [red writing] // Museum Paris | Coll.
|
||||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
<normalizedToken id="77A84C3A2FE064DED8CAC6D38CADC6AE" originalValue="Régimbart">Regimbart</normalizedToken>
|
||||
| apicalis //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="DCBAF3BCF79C82EF0CDF6947301551C7" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">apicalis</emphasis>
|
||||
<emphasis id="EB51594244845B8C179D383E74740ABC" italics="true" pageId="0" pageNumber="19">apicalis</emphasis>
|
||||
n. sp. //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus apicalis</emphasis>
|
||||
<subSubSection id="9FC5B1501AF217272F8C9F4D2D2C4919" pageId="0" pageNumber="19" type="remarks">
|
||||
<paragraph id="1B50D3EEA0BF4E075FD0438BD634BD71" pageId="0" pageNumber="19">Remarks.</paragraph>
|
||||
<paragraph id="C8609E4D8049C3074D70D9F32511C14F" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="ADF2C39A648223FCFA1580C3E337E70B" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">
|
||||
<emphasis id="6566B1964203A646722BAB03508299D6" italics="true" pageId="0" pageNumber="19">Copelatus apicalis</emphasis>
|
||||
</taxonomicName>
|
||||
was described by
|
||||
<bibRefCitation author="Fairmaire, L" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="19" pagination="463 - 499" refId="B17" refString="Fairmaire, L, 1898. Materiaux pour la faune coleopterique de la region malgache. 7e note. . Annales de la Societe Entomologique de Belgique 42: 463 - 499" title="Materiaux pour la faune coleopterique de la region malgache. 7 e note." volume="42" year="1898">Fairmaire (1898)</bibRefCitation>
|
||||
<bibRefCitation id="A78F39D342886AD04FB0A6C85DE744BD" author="Fairmaire, L" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="19" pagination="463 - 499" refId="B17" refString="Fairmaire, L, 1898. Materiaux pour la faune coleopterique de la region malgache. 7e note. . Annales de la Societe Entomologique de Belgique 42: 463 - 499" title="Materiaux pour la faune coleopterique de la region malgache. 7 e note." volume="42" year="1898">Fairmaire (1898)</bibRefCitation>
|
||||
from Suberbieville [=Maevatanana]. It is classified in the genus
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="8845FC02941A0BA18BC2B4C6CE92462B" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A965449C80238150A6A136D4B293CCCC" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
and currently placed in the
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus hydroporoides" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="hydroporoides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus hydroporoides</emphasis>
|
||||
<taxonomicName id="3104C1EFA145652C3CE2337D4C9A041A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus hydroporoides" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="hydroporoides">
|
||||
<emphasis id="55FA64A507370FC685F2EB15A35CF824" italics="true" pageId="0" pageNumber="19">Copelatus hydroporoides</emphasis>
|
||||
</taxonomicName>
|
||||
species group (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
<bibRefCitation id="5394E40DD3B6AE84A93EAFD65E6C1503" pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
|
||||
<bibRefCitation id="530893982769E46435A23E2CE4FFA908" DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="19" refId="B55" refString="Nilsson, AN, 2001. World catalogue of insects. Volume 3. Dytiscidae (Coleoptera). . Apollo Books, Stenstrup" title="World catalogue of insects. Volume 3. Dytiscidae (Coleoptera)." year="2001">Nilsson 2001</bibRefCitation>
|
||||
<bibRefCitation id="800BA12A0E3AA7139917353A0D3DAF08" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="19" refId="B55" refString="Nilsson, AN, 2001. World catalogue of insects. Volume 3. Dytiscidae (Coleoptera). . Apollo Books, Stenstrup" title="World catalogue of insects. Volume 3. Dytiscidae (Coleoptera)." year="2001">Nilsson 2001</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" publicationUrl="http://www.waterbeetles.eu" refId="B57" refString="Nilsson, AN, Hajek, J, 2018. . http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2018">
|
||||
<bibRefCitation id="69A66C1448B41BDA9D6B9B59294C95C4" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" publicationUrl="http://www.waterbeetles.eu" refId="B57" refString="Nilsson, AN, Hajek, J, 2018. . http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2018">
|
||||
Nilsson and
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<normalizedToken id="893A4E43E2A4567E051ADDFE8FC572FC" originalValue="Hájek">Hajek</normalizedToken>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
). We have studied a male syntype of this species which was not found by
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot (1961</bibRefCitation>
|
||||
<bibRefCitation id="1C4B2DAC2C709B656EAB719949A8C0D8" pageId="0" pageNumber="19" refId="B36">Guignot (1961</bibRefCitation>
|
||||
: note 638) and conclude that is not a
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="C9076909D8BCE2186A59DD4036917183" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F736F0476C650ADCED02D0028814CA40" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
. The syntype male has a stout spine-like setae on a protruding anterodistal corner of protarsomere IV which places it in
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
<taxonomicName id="89294194D2A333C654BFB011B1058942" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="5F4C3EF9D4D42E004E56FDCF22F275E8" italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation DOI="https://doi.org/10.1017/S1477200007002599" author="Shaverdo, HV" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="19" pagination="43 - 51" refId="B69" refString="Shaverdo, HV, Monaghan, MT, Lees, DC, Ranaivosolo, R, Balke, M, 2008. Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae). . Systematics and Biodiversity 6: 43 - 51" title="Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae)." url="https://doi.org/10.1017/S1477200007002599" volume="6" year="2008">Shaverdo et al. 2008</bibRefCitation>
|
||||
<bibRefCitation id="934CCB8D1146EAFD0E7F4F58CAE9FA6E" DOI="https://doi.org/10.1017/S1477200007002599" author="Shaverdo, HV" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="19" pagination="43 - 51" refId="B69" refString="Shaverdo, HV, Monaghan, MT, Lees, DC, Ranaivosolo, R, Balke, M, 2008. Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae). . Systematics and Biodiversity 6: 43 - 51" title="Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae)." url="https://doi.org/10.1017/S1477200007002599" volume="6" year="2008">Shaverdo et al. 2008</bibRefCitation>
|
||||
) and we hereby transfer it to that genus:
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus apicalis</emphasis>
|
||||
<taxonomicName id="E288B8421360A1E2E669CF9E201D7AA5" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus apicalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="apicalis">
|
||||
<emphasis id="D947A1870D95C96D65A8BF0971B6D37D" italics="true" pageId="0" pageNumber="19">Madaglymbus apicalis</emphasis>
|
||||
</taxonomicName>
|
||||
(Fairmaire, 1898) comb. nov.; in fact
|
||||
<bibRefCitation author="Gueorguiev, VB" journalOrPublisher="Izvestija na Zoologitjeskija Instituts Musei Sofia" pageId="0" pageNumber="19" pagination="5 - 45" refId="B25" refString="Gueorguiev, VB, 1968. Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae). . Izvestija na Zoologitjeskija Instituts Musei Sofia 28: 5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
|
||||
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||||
<bibRefCitation id="6EA9AF4412EB247F998F493E8D2CAE29" author="Gueorguiev, VB" journalOrPublisher="Izvestija na Zoologitjeskija Instituts Musei Sofia" pageId="0" pageNumber="19" pagination="5 - 45" refId="B25" refString="Gueorguiev, VB, 1968. Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae). . Izvestija na Zoologitjeskija Instituts Musei Sofia 28: 5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
|
||||
<normalizedToken id="D9A1D70077DF88FBC78A1826ECC966A7" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||||
(1968
|
||||
</bibRefCitation>
|
||||
: 32) suggested it might belong to
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
<taxonomicName id="4EC7692271685BA6583278B7E1B7E255" class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="754B207FCCF157BFD5AD6662F1014656" italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
<normalizedToken originalValue="“Peut-être">"Peut-etre</normalizedToken>
|
||||
<normalizedToken originalValue="c’est">c'est</normalizedToken>
|
||||
<normalizedToken id="19CC77FEAA8EF328A8EC2D9552F4D07D" originalValue="“Peut-être">"Peut-etre</normalizedToken>
|
||||
<normalizedToken id="FB8A083E1013A391B4DFB06189EB7CC5" originalValue="c’est">c'est</normalizedToken>
|
||||
une
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
<taxonomicName id="B5FCF62250FC2FF8828E4EC4F0F287D6" class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="85E116B5DAD2D20AF8C22603E2996002" italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
?" [Eng. translation "Maybe it is an
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
<taxonomicName id="A19E9F3E134A091F5E01E0D37B8F58E7" class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="80411EA38CAE7544A8C9F1F320F08418" italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
"], which was correct since this was before
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
<taxonomicName id="5CAB11EF7AA3F07BBB8540766E595BDD" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="5C246641856F5594B868A68CC2915957" italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
was erected for the Madagascar species of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
<taxonomicName id="345776D5867C818EB3BBC372DC4C2A6C" class="Insecta" family="Dytiscidae" genus="Aglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="5D9A80D115413BBAE06D950AF0959DC2" italics="true" pageId="0" pageNumber="19">Aglymbus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation DOI="https://doi.org/10.1017/S1477200007002599" author="Shaverdo, HV" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="19" pagination="43 - 51" refId="B69" refString="Shaverdo, HV, Monaghan, MT, Lees, DC, Ranaivosolo, R, Balke, M, 2008. Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae). . Systematics and Biodiversity 6: 43 - 51" title="Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae)." url="https://doi.org/10.1017/S1477200007002599" volume="6" year="2008">Shaverdo et al. 2008</bibRefCitation>
|
||||
<bibRefCitation id="DC2406DDDB8296E52F48316091556DA9" DOI="https://doi.org/10.1017/S1477200007002599" author="Shaverdo, HV" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="19" pagination="43 - 51" refId="B69" refString="Shaverdo, HV, Monaghan, MT, Lees, DC, Ranaivosolo, R, Balke, M, 2008. Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae). . Systematics and Biodiversity 6: 43 - 51" title="Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae)." url="https://doi.org/10.1017/S1477200007002599" volume="6" year="2008">Shaverdo et al. 2008</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,214 +1,214 @@
|
|||
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<treatment ID-GBIF-Taxon="158995549" LSID="urn:lsid:plazi:treatment:BD7D9CA1871E598AAC575F54EB7455FD" httpUri="http://treatment.plazi.org/id/BD7D9CA1871E598AAC575F54EB7455FD" lastPageNumber="19" pageId="0" pageNumber="19">
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|
||||
<taxonomicName LSID="bd7d9ca1-871e-598a-ac57-5f54eb7455fd" authority="Pederzani & Hajek, 2005" authorityName="Pederzani & Hajek" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||||
<treatment id="BD7D9CA1871E598AAC575F54EB7455FD" ID-GBIF-Taxon="158995549" LSID="urn:lsid:plazi:treatment:BD7D9CA1871E598AAC575F54EB7455FD" httpUri="http://treatment.plazi.org/id/BD7D9CA1871E598AAC575F54EB7455FD" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
|
||||
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|
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<paragraph id="A905975BCE20F8B26D9FACEDB1DB02CC" pageId="0" pageNumber="19">
|
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<taxonomicName id="0B1416949BB89D4BC46A9AD4A0A9364C" ID-CoL="Y3BP" LSID="bd7d9ca1-871e-598a-ac57-5f54eb7455fd" authority="Pederzani & Hajek, 2005" authorityName="Pederzani & Hajek" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||||
Copelatus mahajanga Pederzani &
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<normalizedToken id="82D269E86CB83BC287C3CF254261D63E" originalValue="Hájek">Hajek</normalizedToken>
|
||||
, 2005
|
||||
</taxonomicName>
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5B</figureCitation>
|
||||
<figureCitation id="104BB79C9C14812E6811DC4171C8BC6C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5B</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8B</figureCitation>
|
||||
<figureCitation id="8273F7877E7389FE78EE3C8D2AC95F43" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8B</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">Copelatus mahajanga</taxonomicName>
|
||||
<subSubSection id="651B3E2C3AC700180B0E65C467585EF7" pageId="0" pageNumber="19" type="reference_group">
|
||||
<paragraph id="8DBFA02F3F524ECD04463DFD2418B179" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="56E188F4A9DDA580CB27F426449EDBD3" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">Copelatus mahajanga</taxonomicName>
|
||||
Pederzani &
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<normalizedToken id="D0D57193189CC2901A1FFAF74D2258E9" originalValue="Hájek">Hajek</normalizedToken>
|
||||
, 2005: 104.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Madagascar, Mahajanga Distr., Mahajanga env. [ca. 15°43'S, 46°18'E]</paragraph>
|
||||
<subSubSection id="E484D5508D8A855539EE4BB4747CB4D2" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="BFD9993263DDBE4DA1EAECFBE2A266B7" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="776D8DF804CE06C2F75185AEA8A652F7" pageId="0" pageNumber="19">Madagascar, Mahajanga Distr., Mahajanga env. [ca. 15°43'S, 46°18'E]</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type information from od">
|
||||
<paragraph pageId="0" pageNumber="19">Type information from OD.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Based on male (holotype), 3 male and 4 female (paratypes). I. Janis 1-10 December 1996. HT & 5PT in NMPC. Two paratypes in collection of Pederzani, Ravenna, Italy.</paragraph>
|
||||
<subSubSection id="3D3D374D308FFF08EFF8088B7D0BDA0D" pageId="0" pageNumber="19" type="type information from od">
|
||||
<paragraph id="91C8942D268737BD0388385A7AEA38DF" pageId="0" pageNumber="19">Type information from OD.</paragraph>
|
||||
<paragraph id="CB9F35298F1ACB71020CF9FCC8E20954" pageId="0" pageNumber="19">Based on male (holotype), 3 male and 4 female (paratypes). I. Janis 1-10 December 1996. HT & 5PT in NMPC. Two paratypes in collection of Pederzani, Ravenna, Italy.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Mahajanga I/Mahajanga II</emphasis>
|
||||
<subSubSection id="DB37B293A1F6F26FAB419EF308FC5D95" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="E3F7438DB1D861B0FFAE093CDA9E66ED" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="839A76FD86E785F04480C995F77ED952" pageId="0" pageNumber="19">
|
||||
<emphasis id="E0C34F623084EBE4F3279928B4997C8A" bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Mahajanga I/Mahajanga II</emphasis>
|
||||
[district cannot be verified based on original description or labels]: -HT♂(GP) (NMPC): // MADAGASCAR 1996, Mahajanga Distr., Mahajanga env., Dec. 1-10., I.
|
||||
<normalizedToken originalValue="Janiš">Janis</normalizedToken>
|
||||
<normalizedToken id="23B2BBE2F5A6586D59985CE6262B3F96" originalValue="Janiš">Janis</normalizedToken>
|
||||
leg. // HOLOTYPE, COPELATUS,
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">mahajanga</emphasis>
|
||||
<emphasis id="D9E1429037A366950DB8CF99F0319BFE" italics="true" pageId="0" pageNumber="19">mahajanga</emphasis>
|
||||
sp. nov., F. Pederzani & J.
|
||||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||||
<normalizedToken id="D91399F6BEF03A6599BB5E40BDBE5028" originalValue="Hájek">Hajek</normalizedToken>
|
||||
det.2005 //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny, Mitsinjo, Soalala</emphasis>
|
||||
<subSubSection id="ED56DE505AADC4E39115B673B5C9462A" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="B3DB12E23BDD2728FDD7DB00212A8CB3" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="06316465AB2A45FE30FDBF81496C3620" pageId="0" pageNumber="19">
|
||||
<emphasis id="270C841F74AA3B2F055D8CCD3D00923D" bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny, Mitsinjo, Soalala</emphasis>
|
||||
: -3♂ (GP), 1♀ (NMW): // Data in NHRS | JLKB | 000010719-22 // RM: Betsiboka Bas (PO124) | Loc. Ambohimanatrika | Kamoro Riv. // 47°10'06", 16°28'55" | alt. 40 m 01.04.1993 // Leg. ORSTOM ANTANANARIVO // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010717 // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="3B569F23F07B4DBD7D1138E5B315EC11" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -2♀ (NHRS): // NHRS-JLKB | 000065747 (JB802), 65732 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.13337 | E045.95778, 19 m.a.o. 04.XII.2009 | Water net, Field# MAD09-25 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="375FE8E319631D97B122E5C4085A25EA" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -2♀ (NHRS): // NHRS-JLKB | 000010724 (JB195), 65748 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.01334 | E046.00376, 24 m.a.o. 06.XII.2009 | Water net, Field# MAD09-33 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="0BE5D2D2688418916AD613BA61DAC86F" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -2♂ (GP), 3♀ (CAS): // CASENT-8131811-5 // Madagascar: Mahajanga | Province, Parc National de | Namoroka 16.9 km 317° NW | Vilanandro 12-16 Nov 2002 //
|
||||
<geoCoordinate degrees="16" direction="south" minutes="24" orientation="latitude" precision="15" seconds="24" value="-16.406666">16°24'24"S</geoCoordinate>
|
||||
<geoCoordinate id="ED56636F42084CCFA84532D2E3D6A330" degrees="16" direction="south" minutes="24" orientation="latitude" precision="15" seconds="24" value="-16.406666">16°24'24"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="045" direction="east" minutes="18" orientation="longitude" precision="15" seconds="36" value="45.309998">045°18'36"E</geoCoordinate>
|
||||
<geoCoordinate id="33C436EE24B45146B6BAC9713985551C" degrees="045" direction="east" minutes="18" orientation="longitude" precision="15" seconds="36" value="45.309998">045°18'36"E</geoCoordinate>
|
||||
// Colls. Fisher, Griswold et al.| California Acad. of Sciences | sifted litter, (leaf mold, rotten | wood) in tropical dry forest | elev 100 m code: BLF6582 // -1♀ (CAS): // CASENT-8135013 // Madagascar: Mahajanga | Prov. Parc National de | Namoroka 16.9 Km 317° NW | Vilanandro elev 100 m | 12-16 November 2002 //
|
||||
<geoCoordinate degrees="16" direction="south" minutes="24" orientation="latitude" precision="15" seconds="24" value="-16.406666">16°24'24"S</geoCoordinate>
|
||||
<geoCoordinate id="8888641553D6D816E1FD3881F771AE96" degrees="16" direction="south" minutes="24" orientation="latitude" precision="15" seconds="24" value="-16.406666">16°24'24"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="045" direction="east" minutes="18" orientation="longitude" precision="15" seconds="36" value="45.309998">045°18'36"E</geoCoordinate>
|
||||
<geoCoordinate id="197ADEBD41401BC4D79124DCCFFEAC31" degrees="045" direction="east" minutes="18" orientation="longitude" precision="15" seconds="36" value="45.309998">045°18'36"E</geoCoordinate>
|
||||
| Coll. Fisher, Griswold et al. | California Acad. of Sciences | malaise trap- | tropical dry forest | collection code BLF6581 // -1♂ (GP) (CAS): // CASENT-8135001 // Madagascar: Mahajanga | Province Parc National de | Namoroka | 17.8 Km 329° WNW Vilanandro | elev 100 m: 08-12 Nov 2002 //
|
||||
<geoCoordinate degrees="16" direction="south" minutes="22" orientation="latitude" precision="15" seconds="36" value="-16.376667">16°22'36"S</geoCoordinate>
|
||||
<geoCoordinate id="49CCFCB3DB58D057B94F7ED82A7C3E3B" degrees="16" direction="south" minutes="22" orientation="latitude" precision="15" seconds="36" value="-16.376667">16°22'36"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="045" direction="east" minutes="19" orientation="longitude" precision="15" seconds="36" value="45.326664">045°19'36"E</geoCoordinate>
|
||||
<geoCoordinate id="137CFC24074607B0E2E12189FF74C100" degrees="045" direction="east" minutes="19" orientation="longitude" precision="15" seconds="36" value="45.326664">045°19'36"E</geoCoordinate>
|
||||
| Colls. Fisher, Griswold et al. | California Acad. of Sciences | pitfall trap- in tropical dry forest, | collection code BLF6506 //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
<emphasis id="F9896EB6A5D49BE299CF840B1E620794" bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
: -1♂ (GP) (NHRS): // NHRS-JLKB | 000010723 (JB190) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="82DF297D676E1C7B855A29A5F49306CF" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga</emphasis>
|
||||
<emphasis id="9B978424EC6AFD24D33A5C47E1842249" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Moramanga</emphasis>
|
||||
: -5♂ (GP), 3♀, 11 ex. (Alc) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010554-5, 10596-601, 10795 (Alc.) // MAD: TOAM: Alaotra Mangoro: | Betsabora river by RN2 near | Antasmpanana village, 6Km W | of Moramanga: MAD14-81: river | with side pools: 18.9247S | 48.1828E: 900 m: 24.XI.2014 // Leg. J. Bergsten, | J.H. Randriamihaja | & T. Ranarilalatiana // -3♂ (GP) 26 ex. (Alc.), 7 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010602-3, 10825, 10796(Alc.), 10826(Alc.) // Madagascar: Toamasina: Alaotra | Mangoro: RN2, Betsabora river | 10Km W of Moramanga: S18.92438 | E048.18273, 940 m. 06.XI.2011: | GB Nets and sieves: river and | pools: Field# MAD11-21 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="AFBC6A8140B705945F28C410EF714E0E" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="DD073C3D04A596DC5430924B229FE58F" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="55979F59641E004645E5D76549C9C6BC" pageId="0" pageNumber="19">
|
||||
Close to
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="266CAD50FE2540036160BEE51C83B188" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="0BE7F2B8011717EA7A2E1F7E9222BBE5" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
in habitus appearance but body shape more elongate. On average
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="462E1FDEC7BC386CB4F578CDF41ED370" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="6A36FF5851550F2A93C0FCF54CF3C1CE" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
have a broader transverse testaceous band at base of elytra (
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8B</figureCitation>
|
||||
<figureCitation id="0CBF72D5D731E0DE71F7DD1051028D5C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8B</figureCitation>
|
||||
), and the first elytral stria is more abbreviated anteriorly, starting at approx. one quarter to one half posterior of base. Penis shape is unique in
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="AE19176677B97A7FEF47BC2CCF0DF685" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="FDB2EDA01352865120A122DE8D44143B" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
, bisinuate in ventral view with an expanded apex, not resembling that of any other
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="875ECA51A5A33FCC64C8F6AD025D11A2" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="84FB7673FCD4AA01F63AED9F3B5330B9" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
species in the
|
||||
<taxonomicName lsidName="irinus" pageId="0" pageNumber="19" rank="species" species="irinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">irinus</emphasis>
|
||||
<taxonomicName id="C79962E2148224DA84EB654CE448224D" lsidName="irinus" pageId="0" pageNumber="19" rank="species" species="irinus">
|
||||
<emphasis id="F5BE68A3E6D1AFB05C2F94643FCA6E2C" italics="true" pageId="0" pageNumber="19">irinus</emphasis>
|
||||
</taxonomicName>
|
||||
group from Madagascar.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="FE751FE864BEF9944D3F8D8441769E9F" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="8D709C5B0C3F39768FE8D9523597EE7E" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="0EF0F85E6D30D186CB7A0E1296F83F30" pageId="0" pageNumber="19">
|
||||
Body length 5.2-6.2 mm. Body shape oblong oval, rather convex, brown to dark brown. Head ferrugineous brown, paler in front, sometimes darker around eyes, finely microreticulate and punctate, two shallow depressions between eyes. Pronotum dark brown, paler at sides. Dorsal surface with fine microsculpture and scattered punctures, lateral sides of pronotum striolate with the widest striolate area in the posterior corners. Elytra brown, paler at sides and at apex, with a broad testaceous transverse band at base (
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8B</figureCitation>
|
||||
<figureCitation id="AF473D617CD2F7AAF0264BDC254842F6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8B</figureCitation>
|
||||
). Elytral surface finely reticulate and punctate. Six clearly impressed elytral striae present and one fairly long submarginal stria; first stria abbreviated anteriorly, starting at approx. one quarter to one half posterior of elytral base; other striae (second to sixth) starting more or less at base; submarginal stria starting around middle; striae 1-5 approaching apex of the elytron, but the second and fourth are a little shorter; the sixth stria much abbreviated posteriorly and sometimes the sixth unites with fifth apically approx. 1/5 from apex.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="694B43C96D6F0EF54AA2FC5C2A301B14" pageId="0" pageNumber="19">
|
||||
Ventral side ferrugineous brown, metacoxa and abdominal ventrites striolate and punctate. Prosternal process similar to
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="EC71AD86EA867C426A82404F61579AB4" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="147614380229FD9A99A07F87218C70F7" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
but lateral parts of metaventrite slightly narrower and metacoxal lines less divergent anteriorly. Antennae, palps, pro- and mesothoracic legs brown, but metathoracic legs ferrugineous dark brown.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="9A89CF805AAE13EDE8CF5193C40C9061" pageId="0" pageNumber="19">
|
||||
Male: protibia modified, widened in front, strongly angled after base, with several long spines on the outer side of distal half. Tarsomeres
|
||||
<normalizedToken originalValue="I–III">I-III</normalizedToken>
|
||||
<normalizedToken id="F2B1E5872B60DE2E29B8081BA8890F9A" originalValue="I–III">I-III</normalizedToken>
|
||||
of pro- and mesolegs enlarged, with pads of numerous setae. Penis unique: in ventral view with a bisinuate shape and a widening asymmetrical apex; in lateral view abruptly curved near middle, with a right angle of more or less 90 degrees, the basal part robust (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5B</figureCitation>
|
||||
<figureCitation id="354163ACD2018B957C24ECEF84C5B5D8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5B</figureCitation>
|
||||
). Parameres broad (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5B</figureCitation>
|
||||
<figureCitation id="60E6AB578DFAC0DF1D6BDADEE91B1BA4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Female: similar but smaller than the male, legs not modified.</paragraph>
|
||||
<paragraph id="1592A19A36B09D3D8B0B08584E4F63B8" pageId="0" pageNumber="19">Female: similar but smaller than the male, legs not modified.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="0B620C155BB6FA3BB0B839A62D86370E" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="558B809F8B710612CB2CB12736000DCD" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="78500ECD93AB0E0B7B1C60FC49A108C6" pageId="0" pageNumber="19">
|
||||
Endemic to Madagascar. Known from several places in Mahajanga province and near Moramanga (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11B</figureCitation>
|
||||
<figureCitation id="04D84E7844A34F78A3DBF84493DB69BE" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11B</figureCitation>
|
||||
). We have seen material from around Mahajanga (Type series), Mahavavy Kinkony Reserve, Namoroka NP, Tsingy de Bemaraha NP, Ankarafantsika NP, Kamoro River south of Ankarafantsika NP and collected the species both in 2011 and 2014 at Betsabora River 6 km W of Moramanga which expanded the known distribution significantly. Note that the name Betsabora is uncertain and recovered from locals on site for this river which is a tributary to Mangoro River. Ampanihifana may be the correct name.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="C8FA0AB2FD2A5C8E8AA34DD444B52F16" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="289B37DFCABC0FA4FB8203BC5FFDDFB3" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="9DE60F029334A1DA6F85AC4FBD9E4E63" pageId="0" pageNumber="19">
|
||||
Type series probably collected by light trap (
|
||||
<bibRefCitation author="Pederzani, F" journalOrPublisher="Quaderno di Studi e Notizie di Storia Naturale della Romagna" pageId="0" pageNumber="19" pagination="103 - 107" refId="B59" refString="Pederzani, F, Hajek, J, 2005. Copelatusmahajanga sp. nov. from Madagascar (InsectaColeopteraDytiscidae). . Quaderno di Studi e Notizie di Storia Naturale della Romagna 21: 103 - 107" title="Copelatusmahajanga sp. nov. from Madagascar (InsectaColeopteraDytiscidae)." volume="21" year="2005">
|
||||
<bibRefCitation id="D5163639646C323CEA8C29FAF029AF8C" author="Pederzani, F" journalOrPublisher="Quaderno di Studi e Notizie di Storia Naturale della Romagna" pageId="0" pageNumber="19" pagination="103 - 107" refId="B59" refString="Pederzani, F, Hajek, J, 2005. Copelatusmahajanga sp. nov. from Madagascar (InsectaColeopteraDytiscidae). . Quaderno di Studi e Notizie di Storia Naturale della Romagna 21: 103 - 107" title="Copelatusmahajanga sp. nov. from Madagascar (InsectaColeopteraDytiscidae)." volume="21" year="2005">
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Pederzani and
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<normalizedToken id="19302CBADA40B70F98B87E690931BC4F" originalValue="Hájek">Hajek</normalizedToken>
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2005
|
||||
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|
||||
), and we collected it with black light by a leaf-choked forest pool in Ankarafantsika NP. Also collected with pitfall traps and from leaf-litter in Namoroka NP before the rainy season in November, indicating the species can tolerate periods when the habitats dry up without immediately seeking new water. We collected the largest series at Betsabora River at an elevation of 940 m. This small river runs through a savannah type area with mixed woodland-grassland ecosystem subsequent to degradation and deforestation. The river site consisted of grassy, vegetation-rich margins, bays and side pools, further described in
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.718.20726" author="Michat, MC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="95 - 114" refId="B46" refString="Michat, MC, Gustafson, GT, Bergsten, J, 2017. Larval description and chaetotaxic analysis of Dineutus sinuosipennis Laporte, 1840, with a key for the identification of larvae of the tribe Dineutini (Coleoptera, Gyrinidae). . ZooKeys 718: 95 - 114" title="Larval description and chaetotaxic analysis of Dineutus sinuosipennis Laporte, 1840, with a key for the identification of larvae of the tribe Dineutini (Coleoptera, Gyrinidae)." url="https://doi.org/10.3897/zookeys.718.20726" volume="718" year="2017">Michat et al. (2017)</bibRefCitation>
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<bibRefCitation id="85CA132D27381D7158231C528CE1F296" DOI="https://doi.org/10.3897/zookeys.718.20726" author="Michat, MC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="95 - 114" refId="B46" refString="Michat, MC, Gustafson, GT, Bergsten, J, 2017. Larval description and chaetotaxic analysis of Dineutus sinuosipennis Laporte, 1840, with a key for the identification of larvae of the tribe Dineutini (Coleoptera, Gyrinidae). . ZooKeys 718: 95 - 114" title="Larval description and chaetotaxic analysis of Dineutus sinuosipennis Laporte, 1840, with a key for the identification of larvae of the tribe Dineutini (Coleoptera, Gyrinidae)." url="https://doi.org/10.3897/zookeys.718.20726" volume="718" year="2017">Michat et al. (2017)</bibRefCitation>
|
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. Around the Mahajanga area all sites are lowaltitude, 100 m or lower.
|
||||
</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus mahajanga</emphasis>
|
||||
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<paragraph id="37D5167EAE7CCA028399E36D8E643D1E" pageId="0" pageNumber="19">Comments.</paragraph>
|
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<paragraph id="CABCA2C4DEDBDA1E5B41CC6630BD365A" pageId="0" pageNumber="19">
|
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<taxonomicName id="6F0032E3C09D1D17A6E14BBF55FE473E" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||||
<emphasis id="5F0BB8B630949E2C655137B82C1CBBF3" italics="true" pageId="0" pageNumber="19">Copelatus mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
was the most recently described
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="CEEC19B4F8A3438A2870C91415206C79" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="36F85E179CB34703CB0F03403188D616" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
species from Madagascar (Peredzani and
|
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|
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<normalizedToken id="1FBEDC8E59A4B4BAFC1794F992F1D68F" originalValue="Hájek">Hajek</normalizedToken>
|
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2005) and was previously only known from the type series and lowland west type locality "Mahajanga env." without further details. The discovery of the species in Betsabora River at mid-altitude in the east indicates that the species may have a much wider distribution and ecological niche. As the species is very similar to
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="4F09F9F51A124F7D1149BF071B8F2CB1" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
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<emphasis id="C525EB86D7CB702A05B776D0BD57ACCB" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
habitually, it might be misidentified as such in collections if male genitalia were not examined.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,387 +1,387 @@
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<treatment ID-GBIF-Taxon="158995547" LSID="urn:lsid:plazi:treatment:C9FE8EA486D557DFA5B1A38A9F232743" httpUri="http://treatment.plazi.org/id/C9FE8EA486D557DFA5B1A38A9F232743" lastPageNumber="19" pageId="0" pageNumber="19">
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<taxonomicName LSID="c9fe8ea4-86d5-57df-a5b1-a38a9f232743" authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Copelatus marginipennis (Laporte, 1835)</taxonomicName>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5A</figureCitation>
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<treatment id="C9FE8EA486D557DFA5B1A38A9F232743" ID-GBIF-Taxon="158995547" LSID="urn:lsid:plazi:treatment:C9FE8EA486D557DFA5B1A38A9F232743" httpUri="http://treatment.plazi.org/id/C9FE8EA486D557DFA5B1A38A9F232743" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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<taxonomicName id="9D978EC63A9E92287C896577FBA33B6E" ID-CoL="8LVBF" LSID="c9fe8ea4-86d5-57df-a5b1-a38a9f232743" authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Copelatus marginipennis (Laporte, 1835)</taxonomicName>
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<figureCitation id="76A4E815FC9008E2840BD4E17D3FF046" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5A</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8A</figureCitation>
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<figureCitation id="354463A8C057D80FC5046FD4C16F8D34" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8A</figureCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Colymbetes marginipennis</taxonomicName>
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<paragraph id="C66E26B201407C151AB0ECF16F73D81D" pageId="0" pageNumber="19">
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<taxonomicName id="3AF0F871F7BC9E74BFFE5E1E9FA9796F" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">Colymbetes marginipennis</taxonomicName>
|
||||
Laporte, 1835: 102.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">Copelatus aldabricus</taxonomicName>
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|
||||
<taxonomicName id="5339E026CA7E45C5A15B8A94F8EEABB5" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">Copelatus aldabricus</taxonomicName>
|
||||
J. Balfour-Browne, 1950: 368 syn. nov.; TL: Seychelles, Aldabra Islands.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Copelatus aldabricus var. simplex" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="variety" species="aldabricus" variety="simplex">Copelatus aldabricus var. simplex</taxonomicName>
|
||||
<bibRefCitation author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot 1952</bibRefCitation>
|
||||
<paragraph id="C6697E34FD3F167755222488044FDBA9" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="7C3056FA9B643B4D37AA6D542C5510F4" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Copelatus aldabricus var. simplex" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="variety" species="aldabricus" variety="simplex">Copelatus aldabricus var. simplex</taxonomicName>
|
||||
<bibRefCitation id="F3A332DDA4BB226EB798D4588209A9D8" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot 1952</bibRefCitation>
|
||||
: 28 syn. nov.; TL: Madagascar.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Senegal [possibly mislabelled].</paragraph>
|
||||
<subSubSection id="185EC79E76B103A43B358EEDD4C14F9C" pageId="0" pageNumber="19" type="type locality">
|
||||
<paragraph id="5C28815C979214C67523141CF21D1AD7" pageId="0" pageNumber="19">Type locality.</paragraph>
|
||||
<paragraph id="3DA560428E5C8A08B8E12C58FE88A88D" pageId="0" pageNumber="19">Senegal [possibly mislabelled].</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="B04CC2F74D79732D6909EA2C4F92BCBB" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="9AA5616F51EE767D2E0B666109C041B7" pageId="0" pageNumber="19">Type information from original description.</paragraph>
|
||||
<paragraph id="93C0E2A5321570DF90A70F851A6A2D41" pageId="0" pageNumber="19">
|
||||
housed in Buquet collection and originating from Senegal; of
|
||||
<taxonomicName lsidName="aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
|
||||
<taxonomicName id="B502DAD74A569C0A14A17CAEC7BAB5AF" lsidName="aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="38FA8E3484D28C1171332DDC968FFC2B" italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
: based on male (holotype), J.C. Fryer collection, collected 1908-9 from Aldabra; of
|
||||
<taxonomicName lsidName="simplex" pageId="0" pageNumber="19" rank="species" species="simplex">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">simplex</emphasis>
|
||||
<taxonomicName id="5F2B2ABD3EB2DE7DDDCF68AB27A805C6" lsidName="simplex" pageId="0" pageNumber="19" rank="species" species="simplex">
|
||||
<emphasis id="29B4C719C3ED278D1D7E4D990D584AD5" italics="true" pageId="0" pageNumber="19">simplex</emphasis>
|
||||
</taxonomicName>
|
||||
: based on male and female syntypes from Madagascar without further locality data.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="08C6C7FE15C944ED1ED4B7F9B3F63A26" pageId="0" pageNumber="19" type="type material">
|
||||
<paragraph id="0887C859A13CDC1FB1B741EA09D76748" pageId="0" pageNumber="19">Type material studied.</paragraph>
|
||||
<paragraph id="53DB1315BF50FF2CEA1998728D471E46" pageId="0" pageNumber="19">
|
||||
-LT♂ (lectotype here designated) (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
<emphasis id="CE77F6AB050CB836B497AF45A7746FFB" italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
) (MNHN): // Data in NHRS | JLKB | 000065416 //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<emphasis id="AE79EC3C109FFA50281BD889CDA4025B" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
Buquet | pulchellus var.
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<normalizedToken id="78CED36A7AF089815781D471D80DF862" originalValue="Aubé">Aube</normalizedToken>
|
||||
| h. in Senegal D. Buquet // D. Sharp | Monogr. // Ex-Musaeo
|
||||
<normalizedToken originalValue="Déjean">Dejean</normalizedToken>
|
||||
<normalizedToken id="32163613568080D3FD6611B40A097F05" originalValue="Déjean">Dejean</normalizedToken>
|
||||
//
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
<emphasis id="CF1A3DFDD046AD3E2B878B0F093F2736" italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
// Lectotype |
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
<emphasis id="479F55808DB37B69242968F2D0DB558C" italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
| Laporte, 1835 | Des. Ranarilalatiana | & Bergsten, 2019// -HT♂ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
<emphasis id="DF8B72E3E6FF32EC65F267967B4AF69A" italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
) (NHMUK): // Aldabra, 08-9. J.C.F. Fryer // Perey Sladen Trust | expedition. | 1913-170. // Type [red round label] //
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<emphasis id="3917CDE51E17CAC65E713A0CD5B0B776" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
|
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
|
||||
<emphasis id="CAF161D967B65D35B38E5EDFA598D060" italics="true" pageId="0" pageNumber="19">aldabricus</emphasis>
|
||||
Type | J. Balfour-Browne det. //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Andilamena, Moramanga</emphasis>
|
||||
<subSubSection id="023CE58822DAA831CF14723D2010F2AD" pageId="0" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="25D3AFE721C6E50D8FEEA58A9ECACE59" pageId="0" pageNumber="19">Additional material studied.</paragraph>
|
||||
<paragraph id="D3D58AC16D8008391A3B7D561C3E603F" pageId="0" pageNumber="19">
|
||||
<emphasis id="326DB43D2CD9BA2BFF1336B33FD479F0" bold="true" pageId="0" pageNumber="19">Toamasina. Alaotra Mangoro: Andilamena, Moramanga</emphasis>
|
||||
: -1♀ (NMW): // Data in NHRS | JLKB | 000010726 // Madagascar 17.01.1995 | Ambatombe | nr. Andilamena 900 m | leg. Dunay &
|
||||
<normalizedToken originalValue="Janák">Janak</normalizedToken>
|
||||
<normalizedToken id="54492356B448AC2D6963C9157424B50C" originalValue="Janák">Janak</normalizedToken>
|
||||
(10) // -1♀ (NHMUK): // BMNH-797894 // MAD: AMPA: Moramanga: Andasibe | Andasibe NP: Big Pond | P61BI15: N: -18.937: E:48.416: 940 m | 06/I/2007: Leg. Isambert et al. // DNA Voucher | BMNH <797894> | MSL294:B10 // -2♂ (GP), 15 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010587-8, 10799(Alc.) // Madagascar: Tamatave: Alaotra Mangoro: | Analamazaotra RS; Bas fond, non-permanent | pond near trail to "lac rouge"MAD15-1| 943 m,
|
||||
<geoCoordinate degrees="18" direction="south" minutes="56" orientation="latitude" precision="1" seconds="26.7" value="-18.940748">18°56'26.7"S</geoCoordinate>
|
||||
<geoCoordinate id="98D17BF88F1664DB06C22418D2C4ED22" degrees="18" direction="south" minutes="56" orientation="latitude" precision="1" seconds="26.7" value="-18.940748">18°56'26.7"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="048" direction="east" minutes="25" orientation="longitude" precision="1" seconds="03.9" value="48.41775">048°25'03.9"E</geoCoordinate>
|
||||
<geoCoordinate id="FEA2659B0C7B916B0677F6F07719F101" degrees="048" direction="east" minutes="25" orientation="longitude" precision="1" seconds="03.9" value="48.41775">048°25'03.9"E</geoCoordinate>
|
||||
, 16.III.2015 | Among vegetation and dead leaves in the pond, | Leg. T. Ranarilalatiana & H.J. Randriamihaja // -2♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010604-5, 10800(Alc.) // Madagascar: Tamatave: Alaotra Mangoro: Mantadia NP. | Non-permanent pond at PK18,50 m; E of Park road | 973 m,
|
||||
<geoCoordinate degrees="18" direction="south" minutes="46" orientation="latitude" precision="1" seconds="09.9" value="-18.769417">18°46'09.9"S</geoCoordinate>
|
||||
<geoCoordinate id="D9CFDB4BA14EFE14ABE29374CB6C0EF6" degrees="18" direction="south" minutes="46" orientation="latitude" precision="1" seconds="09.9" value="-18.769417">18°46'09.9"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="048" direction="east" minutes="26" orientation="longitude" precision="1" seconds="10.4" value="48.436222">048°26'10.4"E</geoCoordinate>
|
||||
<geoCoordinate id="CF30BCB56D487B7B30ED9179A5B0923E" degrees="048" direction="east" minutes="26" orientation="longitude" precision="1" seconds="10.4" value="48.436222">048°26'10.4"E</geoCoordinate>
|
||||
, 17.III.2015 | Under dead leaves & vegetation at the edge of the pond, | Leg.T. Ranarilalatiana, H.J. Randriamihaja; MAD15-5 // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010815 // Madagascar: Toamasina: Alaotra | Mangoro: RN2, Mangoro river | 10Km W of Moramanga: S18.92438 | E048.18273, 940 m. 06.XI.2011 | GB Nets and sieves: river and | pools: Field# MAD11-21 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | J.H. Randriamihaja // -1♀ (Alc.) (NHRS): // NHRS-JLKB | 000010822 // Madagascar: Toamasina: Alaotra | Mangoro: Analamazaotra SR: | close to park entrance: S18.9355 E | 048.41656, 970 m: 08.XI.2011: GB | Nets and sieves: dried up forest | pond: Field# MAD11-25 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | H.J. Randriamihaja // -6 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010823 // Madagascar: Toamasina: Alaotra | Mangoro: by RN2 S border of | Analamazaotra reserve 1Km E | Antsampanana: S
|
||||
<geoCoordinate degrees="18.94987" direction="east" orientation="longitude" precision="1" value="18.94987">18.94987 E</geoCoordinate>
|
||||
<geoCoordinate id="22649880DE5457D2C571896563F76525" degrees="18.94987" direction="east" orientation="longitude" precision="1" value="18.94987">18.94987 E</geoCoordinate>
|
||||
048.42331 | 980m: 09.XI.2011: GB Nets & sieves | ditch next to road: Field# MAD11-29 // Leg. J. Bergsten, R. | Bukontaite, T. | Ranarilalatiana & | J.H. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toamasina. Analanjirofo: Maroantsetra</emphasis>
|
||||
<emphasis id="9F11A02F71E04667D246BBA866137CB2" bold="true" pageId="0" pageNumber="19">Toamasina. Analanjirofo: Maroantsetra</emphasis>
|
||||
: -3♂ (GP), 2♀ (NHMUK): // BMNH-797906-10 // MAD: TOAM: Maroantsetra: Masoala | Masoala NP: Pool | P58BI14: N: -15.758: E: 49.993: 10 m| 17/XI/2006: Leg. Isambert et al. // BMNH (DNA Voucher) // -1♂ (GP) (NHRS): // NHRS-JLKB | 000010794 // MAD: TOAM: Maroantsetra: Masoala | Masoala NP: Pool | P58BI14: N: -15.758: E: 49.993: 10 m| 17/XI/2006: Leg. Isambert et al. // -1♂ (NMW): // Data in NHRS | JLKB | 000010728 // E-Madagascar: Fenerive | Foret de Tampolo | 28.12.1998 | leg. J. Moravec // -1♀(Alc.) (NHRS): // NHRS-JLKB | 000011122 // Madagascar: Toamasina: Analajinrofo: | Masoala NP: degraded lowalt. forest: | MAD18-47: small waterpools on path | ~0.5km NW of Marofototra village, | 15.7606S, 49.9926E, 15 m, 17.II.2018 | Leg. T. Ranarilalatiana //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Toliara. Menabe: Morondava</emphasis>
|
||||
<emphasis id="DC7F309894A75F343A1DD695D2B75063" bold="true" pageId="0" pageNumber="19">Toliara. Menabe: Morondava</emphasis>
|
||||
: -3♂ (GP) 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010580, 10730 (JB197), 10751, 10802(Alc.) //Madagascar: Toliara: Menabe: | Kirindy RS: S20.07430 | E044.66307, 52 m.a.o. 12.XII.2009 | Water Net, Field# MAD09-46 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="F5EA70CC85CB0F5A6D8DD833BA43165A" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny, Mitsinjo</emphasis>
|
||||
<emphasis id="DCECF5BC65B28C8DD7A0EF0BE811DACB" bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Ambato-Boeny, Mitsinjo</emphasis>
|
||||
: -1♂ (NHRS): // NHRS-JLKB | 000010734 (JB201) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30350 | E046.81068, 87 m.a.o. 29.XI.2009 | Water Net, Field# MAD09-03 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="74E6DF92972C5642E79159A5FEEB8350" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -6♂ (GP), 3♀, 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010567-71, 10729 (JB196), 10732 (JB199), 10742-3, 10798(Alc.) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.30341 | E046.81073, 74 m.a.o. 29.XI.2009 | 22W Black Light, Field# MAD09-07 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="5659AC8B31BCECD06D0739CA25CEC0D7" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♂ (NHRS): // NHRS-JLKB | 000010731 (JB198) // Madagascar: Mahajanga: Boeny | Ankarafantsika NP, S16.30270 | E046.80996; 75 m.a.o. 30.XI.2009 | 22W Black light, Field# MAD09-13 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="AABC492038F645AC8C584BA25E6ADDCE" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010737 (JB191) // Madagascar: Mahajanga: Boeny: | Ankarafantsika NP. S16.31418 | E046.81731, 30.XI.2009 | Hand picking, Field# MAD09-14 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="B864B763CE886DD6C67B4E9CE35C17FE" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -3♀, 1♂(GP), 11 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000065749 (JB809), 10738 (JB192), 10744-5, 10806(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.14653 | E045.94926, 9 m.a.o. 04.XII.2009 | Water net, Field# MAD09-24 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="3BE335E5BF7EE56F7D13961DA15EF9F2" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -2♂ (GP), 2♀, 18 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010578-9, 10739 (JB193), 10746, 10807(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.13337 | E045.95778, 19 m.a.o. 04.XII.2009 | Water net, Field# MAD09-25 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="6F951668A3E1DDAC74BF283A224AC2A1" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
| T. Ranarilalatiana, H.J. Randriamihaja // -5♂ (GP), 4♀, 79 ex. (Alc.) (NHRS, DEUA & PBZT/MBC): // NHRS-JLKB | 000010572-7, 10740 (JB194), 10747-8, 10808 (Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.05776 | E045.80585, 22 m.a.o. 05.XII.2009 | Water net, Field# MAD09-28 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="045ECB2BD82B1BF6AC9ECA64789F0828" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja // -1♂, 1♀, 4 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010735 (JB202), 10749, 10809(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.06651 | E045.77672, 24 m.a.o. 05.XII.2009 | Water net, Field# MAD09-29 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="421461A10D4D4CCDF1C12DC9CFA6921D" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -1♀ (NHRS): // NHRS-JLKB | 000010741(JB189) // Madagascar: Mahajanga: Boeny | Mahavavy Kinkony RS, 16.05648S | 045.76371E; 55 m.a.o. 05.XII.2009 | Water net, Field# MAD09-30 | Leg: J. Bergsten, N. Jonsson, T. | Ranarilalatiana, H.J. Randriamihaja // -1♂, 6ex. (Alc.) (NHRS): // NHRS-JLKB | 000010750, 10810(Alc.) // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS. S16.01334 | E046.00376, 24 m.a.o. 06.XII.2009 | Water net, Field# MAD09-33 | Leg. J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="85A5D68193B95B3CB0E653CDA078FD74" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, | T. Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
<emphasis id="A997BBD1EA434E489FC36245FFB1B62A" bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
|
||||
: -2♂ (GP) (CAS): // CASENT-8135015-6 // Madagascar: Mahajanga | Prov. Parc National Tsingy | de Bemaraha, 2.5 Km 62° ENE | Bekopaka, Ankidrodroa river | elev 100 m: 11-15 Nov 2001 //
|
||||
<geoCoordinate degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
<geoCoordinate id="CB2A082E5072E319EB57245B56FB84CB" degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
<geoCoordinate id="4750DD082B4288BB5BBC61C20C83E822" degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
| Coll: Fisher, Griswold et al. | California Acad. of Sciences | sifted litter - tropical dry forest | on Tsingy, code: BLF4340 // -1♂ (GP) (CAS): // CASENT-8131891 // Madagascar: Mahajanga | Prov. Parc National Tsingy | de Bemaraha, 2.5 Km 62° ENE | Bekopaka, Ankidrodroa river | elev 100 m: 11-15 Nov 2001 //
|
||||
<geoCoordinate degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
<geoCoordinate id="2F4B88CC9D5E9EB9084D1929EB6A65A5" degrees="19" direction="south" minutes="7" orientation="latitude" precision="15" seconds="56" value="-19.132223">19°7'56"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
<geoCoordinate id="85B3B6EC9357492B5EC74269E7FA11F9" degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53"E</geoCoordinate>
|
||||
| Coll: Fisher, Griswold et al. | California Acad. of Sciences | at light- tropical dry forest | on Tsingy, code: BLF4343 // -1♂ (GP) (CAS): // CASENT-8135006 // Madagascar Mahajanga | Prov. Foret de Tsimembo | 11.0Km 346° NNW Soatana | elev 50 m: 21-25 Nov 2001 |
|
||||
<geoCoordinate degrees="18" direction="south" minutes="59" orientation="latitude" precision="15" seconds="43" value="-18.995277">18°59'43"S</geoCoordinate>
|
||||
<geoCoordinate id="65222D2A9F0A10F584C501CF6A76FADF" degrees="18" direction="south" minutes="59" orientation="latitude" precision="15" seconds="43" value="-18.995277">18°59'43"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44" direction="east" minutes="26" orientation="longitude" precision="15" seconds="37" value="44.44361">44°26'37"E</geoCoordinate>
|
||||
<geoCoordinate id="66581E341180274907E76987111EB4A1" degrees="44" direction="east" minutes="26" orientation="longitude" precision="15" seconds="37" value="44.44361">44°26'37"E</geoCoordinate>
|
||||
// Coll: Fisher, Griswold et al. | California Acad. of Sciences | sifted litter (leaf mold, rotten wood) | in tropical dry forest | coll. Code: BLF4508 // -1♂ (GP), 2♀, 3 ex. (Alc.), 2♂(Alc.) (NHRS): // NHRS-JLKB | 000010581-2, 10752, 10803(Alc.), 10821(Alc.)// Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="85CF637937359426902609D6C3DF963F" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -3♂ GP, 1♀, 5 ex. (Alc.) (NHRS): // NHRS-JLKB | 000010583-4, 10736 (JB203), 10754, 10805(Alc.) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S18.75643 | E044.71398, 119 m.a.o. 17.XII.2009 | Water Net, Field# MAD09-65 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="8F340692EA1922AD4E86C838C6E29B08" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja // -1♀, 1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010733 (JB200), 10753, 10804(Alc.) // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: 19.03419S | 044.77499E, 41 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-59 | Leg: J. Bergsten, N.
|
||||
<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
|
||||
<normalizedToken id="E2FE18E1DDA3396DEA38DC0E9D9A89AD" originalValue="Jönsson">Joensson</normalizedToken>
|
||||
, T. | Ranarilalatiana, H.J. Randriamihaja //
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Ambanja, Ambilobe, Antsiranana II</emphasis>
|
||||
<emphasis id="3CD456EB3BDF7FFF8D08900D99E71FCE" bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Ambanja, Ambilobe, Antsiranana II</emphasis>
|
||||
: -1♂ (GP) (NHRS): // NHRS-JLKB | 000010585 // Madagascar: Antsiranana: Diana: | Ambilomagodra: Stream under the | bridge of the road RN6 in | Ambilomagodra village, S13.00780 | E49.13313, 139 m, 30.XI.2012, GB nets: | dried up stream with some pools in | the village: Field# MAD12-33 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♂ (GP), 2♀ (Alc.) (NHRS): // NHRS-JLKB | 000010586, 10801(Alc.) // Madagascar: Antsiranana: Diana: | Andrafiabe: Antsoha stream | 200m from Andrafiabe, 12.93022S | 49.03466E, 32 m, 01.XII.2012, GB | nets: stream with some pools: | Field# MAD12-39 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♂ (Alc.) (NHRS): // NHRS-JLKB | 000011123 // Madagascar: Antsiranana: Diana: | Antsaba: 1km W of Antsaba, | 13.63474S; 48.72918E, 67 m, | 28.XI.2012, GB nets: forest | stream: Field# MAD12-32 // Leg. J. Bergsten, R. | Bukontaite, T | Ranarilalatiana & | J.H. Randriamihaja // -1♀ (NMW): // Data in NHRS | JLKB | 000010725 // MADAGASCAR: Sakaramy | (M.
|
||||
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
<normalizedToken id="866689F90EA40C812AAB492B6C157F3D" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
, Antsiranana) | lake Farihy Makery | 29.03.2011 | leg. R. Gerecke (MD189) // riparian area near outflow |
|
||||
<geoCoordinate degrees="12" direction="south" minutes="26" orientation="latitude" precision="1" seconds="20.5" value="-12.439028">12°26'20.5"S</geoCoordinate>
|
||||
<geoCoordinate id="195ED4797187BA16C2E1E4EBEE969369" degrees="12" direction="south" minutes="26" orientation="latitude" precision="1" seconds="20.5" value="-12.439028">12°26'20.5"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="49" direction="east" minutes="14" orientation="longitude" precision="1" seconds="20.0" value="49.238888">49°14'20.0"E</geoCoordinate>
|
||||
<geoCoordinate id="30C2216EB6FFE2B317C41C2EC8D5EC2C" degrees="49" direction="east" minutes="14" orientation="longitude" precision="1" seconds="20.0" value="49.238888">49°14'20.0"E</geoCoordinate>
|
||||
| 377 m. 29.5 °C // -1♂ (NMW): // Data in NHRS | JLKB | 000010727 // N. Madagascar | Antseranana distr. | Sambirano riv. | Marovato vill. | 5-12.12.01, leg. J.
|
||||
<normalizedToken originalValue="Horák">Horak</normalizedToken>
|
||||
<normalizedToken id="371C4C88256B4F3B83D62E9FF902D7DE" originalValue="Horák">Horak</normalizedToken>
|
||||
// Antsira
|
||||
<emphasis bold="true" pageId="0" pageNumber="19">nana. Sava: Sambava, Vohemar</emphasis>
|
||||
<emphasis id="360FFBF7E1307F2BDCB12DFB34CE89D2" bold="true" pageId="0" pageNumber="19">nana. Sava: Sambava, Vohemar</emphasis>
|
||||
: -1♀ (NHMUK): // BMNH-797876 // MAD: DIEG: Sambava: Marojejy | Marojejy NP: Pool | P57BI31: N: -14.457: E:49.79: 162m | 10/XI/2006; Leg. Isambert et al // DNA Voucher | BMNH <797876> | MSL294:A4 // -1♀ (CAS): // CASENT-8135007 // Madagascar Antsiranana | Foret
|
||||
<normalizedToken originalValue="d’Ampondrabe">d'Ampondrabe</normalizedToken>
|
||||
<normalizedToken id="F3CD69004472B38303F8DAA0CD583460" originalValue="d’Ampondrabe">d'Ampondrabe</normalizedToken>
|
||||
. | 26.3Km 10° NNE Daraina | elev 175m: 10 December 2003 //
|
||||
<geoCoordinate degrees="12" direction="south" minutes="58" orientation="latitude" precision="15" seconds="12" value="-12.969999">12°58'12"S</geoCoordinate>
|
||||
<geoCoordinate id="74F7732FC7FBE96D730899270EFF7281" degrees="12" direction="south" minutes="58" orientation="latitude" precision="15" seconds="12" value="-12.969999">12°58'12"S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="049" direction="east" minutes="42" orientation="longitude" precision="15" seconds="00" value="49.7">049°42'00"E</geoCoordinate>
|
||||
<geoCoordinate id="67A03CC32C3650F2122F750BF7746B94" degrees="049" direction="east" minutes="42" orientation="longitude" precision="15" seconds="00" value="49.7">049°42'00"E</geoCoordinate>
|
||||
| California Acad. of Sciences | Coll: B.L.Fisher, sifted | litter (leaf mold, rotten wood) | tropical dry forest, BLF9974 //
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<subSubSection id="A5278385F472E16C6B51B06059BFF6A1" pageId="0" pageNumber="19" type="diagnosis">
|
||||
<paragraph id="D93265DFF2E87881B2A22209B6EC223C" pageId="0" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph id="807E77F728D547B1E56C050B34F06C65" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="4D14843D9E6E9B3F733E980BB1E8F1DA" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="2EAE1BDED6C48836C371EC8CBD305993" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from all other Malagasy
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
<taxonomicName id="10F26C9A9ADDA3792107C7D111EE1987" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="CDB9F027812A0EFC3A8A98B4B9A2BBE2" italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||||
</taxonomicName>
|
||||
except
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="46AA5BDB5F433D53BCD818B3CE0346FB" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="EDAB9F8C18EF63A738B9E661AD4D7389" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="FF2B27544466F15D5A488EAE6E6B527E" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="0BC5B19470AD244333FE099F30657272" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
by the presence of six discal elytral striae and a broadly oval body shape.
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<taxonomicName id="4CEA8D880BACE33A7A90F06832FC7990" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="E55ADC77273E2D204EAA9E002FFE74C1" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is most easily separated by the distinct shape of the male penis in lateral view (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
<figureCitation id="49802240D97C85BDD827A86E79D4869F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
).
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="3C049F12CDF06C1143041719079E3132" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="0FD4004696346D4B7F6D138A5E262B1B" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
(from Madagascar) lacks a transverse testaceous band basally on the elytra and females of
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="119A4C18D1D7197BF0BCF64B3FB70EC6" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="8C552C78095F946DA6E4F9DCA2E47A13" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
can usually be distinguished from
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="41396725A68A7EB63722EAFC390178B6" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="859C3E614D14503B2D845D8A2AFD2DAF" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
by the normally narrower testaceous band, a longer first elytral stria, and a broader body shape.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="description">
|
||||
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="97D665E1C9FF1350B0A37D63DD4611AE" pageId="0" pageNumber="19" type="description">
|
||||
<paragraph id="62EBF1261A1ED5E6EA96408BBC0D33EC" pageId="0" pageNumber="19">Description.</paragraph>
|
||||
<paragraph id="FE18C026E615A57FE322D5F86ECA606A" pageId="0" pageNumber="19">
|
||||
Body length 5.2-6.6 mm. Body shape oval, rather convex and attenuate posteriorly, dark brown to brown ferrugineous. Head, pronotum and elytra in the same dark brown ferrugineous, covered with fine dense punctation. Lateral sides of pronotum more brownish, with short strioles and the widest striolate area in the posterior corners. Elytra dark brown to brown ferrugineous, with a testaceous transverse band at base (
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8A</figureCitation>
|
||||
<figureCitation id="4D3958076ABB90BF13A7F159FE6A964F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8A</figureCitation>
|
||||
). Six clearly impressed elytral striae present and one submarginal: first stria abbreviated and slightly shorter, starting at approx. 1/7th posterior of elytral base, second to sixth full length, and submarginal stria starting at approx. the middle; all striae approaching the apex of elytron except the sixth, which is abbreviated posteriorly and a little shorter. Sometimes, the first and second elytral striae, like fifth and sixth, unite posteriorly. Elytral surface covered with dense punctures.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Ventral side brownish to ferrugineous, metacoxa with microsculpture, densely and finely punctate. Metacoxa and abdominal ventrites striolate. Prosternal process rather short and spear-shaped, medially only weakly raised and rounded. Lateral parts of metaventrites rather broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae and palps both in the same brown colour. Pro- and mesothoracic legs brown ferrugineous. Metathoracic legs dark brown ferrugineous.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<paragraph id="D2719AAF67CE9384D0A02917E3DF8693" pageId="0" pageNumber="19">Ventral side brownish to ferrugineous, metacoxa with microsculpture, densely and finely punctate. Metacoxa and abdominal ventrites striolate. Prosternal process rather short and spear-shaped, medially only weakly raised and rounded. Lateral parts of metaventrites rather broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae and palps both in the same brown colour. Pro- and mesothoracic legs brown ferrugineous. Metathoracic legs dark brown ferrugineous.</paragraph>
|
||||
<paragraph id="D2875DF7433FE87BB515D04C40475829" pageId="0" pageNumber="19">
|
||||
Male: Protibia strongly angled basally and expanded in apical two thirds. Penis in ventral view with a small preapical tooth on right side; in lateral view very characteristic with a subbasal dorsal knob, and post-middle with a deep ventral invagination (
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
<figureCitation id="9452F1FCE33EAC66D2BC57CD18068D47" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">Females from Madagascar usually with elytral striolation rather weak and restricted to outer intervals, but rarely the elytra are entirely and distincly striolate. Females on average smaller than males.</paragraph>
|
||||
<paragraph id="8821ABB848C48DD7FDCBFF1DCA63069C" pageId="0" pageNumber="19">Females from Madagascar usually with elytral striolation rather weak and restricted to outer intervals, but rarely the elytra are entirely and distincly striolate. Females on average smaller than males.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="925BA09FCB74D22D13210D77727451EA" pageId="0" pageNumber="19" type="distribution">
|
||||
<paragraph id="99F52898CF6BA7770870DDC4C23BB043" pageId="0" pageNumber="19">Distribution.</paragraph>
|
||||
<paragraph id="99AF1F00C286F5F511E6F116F3B8B08B" pageId="0" pageNumber="19">
|
||||
Endemic to the western Indian Ocean islands as far as is modernly known, but the 1835 type locality of the original description and labels read
|
||||
<normalizedToken originalValue="“Senegal”">"Senegal"</normalizedToken>
|
||||
<normalizedToken id="4A0DB330189D26E358E178B6FB9C5A5D" originalValue="“Senegal”">"Senegal"</normalizedToken>
|
||||
, which indicates either mislabeling or that the species in fact also occurs on continental Africa. Known from Madagascar, Reunion, Comores, Seychelles, and Aldabra Island (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
<bibRefCitation id="EB616DEA0BCC0D2480B5DDEBF7D48ABE" pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
). Widespread in Madagascar and recorded from Montagne
|
||||
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
<normalizedToken id="35F5BC8A22ACC0D57A111B8462FFC8C6" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||||
NP and Masoala NP in the north and northeast to Kirindy in the southwest (
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
|
||||
<figureCitation id="528A6C3DBA15E7D760F3951BAD8CFA2C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
|
||||
), but apparently lacking from most of the central plateau. It can probably show up anywhere around the island below 1000 m in altitude.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<subSubSection id="9B6DA779913403A8509332E9D9CC5A67" pageId="0" pageNumber="19" type="habitat">
|
||||
<paragraph id="DBB001FCE0AFE5824C5EA9261F2EACAC" pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||||
<paragraph id="C19B39AEC996E30E65595A22398DDED3" pageId="0" pageNumber="19">
|
||||
This is a lowland species that seems to be most common in the deciduous western parts of Madagascar. It was common in the newly designated protected areas of Mahavavy Kinkony when we visited it in 2009. In the deciduous forest biome it has been recorded from Kirindy in the south to Ankarana NP in the north. The species seems to be a generalist and can as well show up on lowland humid east coast and at midaltitudes up to at least 1000 m. It is an apt flier collected with light traps and often in very temporary and small shallow pools including water-filled wheel tracks. Found in all kinds of temporary pools, as well as in streams and in residual pools in dried-up riverbeds. It occurred sympatrically with
|
||||
<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
<taxonomicName id="FBBA20B14E8105E965C17C67107EA856" lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
|
||||
<emphasis id="76D605315FEAFB319E330EB68B7F9CB8" italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
|
||||
</taxonomicName>
|
||||
in Betsabora river near Moramanga and at a 22W black light trap by a forest pool in Ankarafantsika NP.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
<subSubSection id="05068E3C3D362BEFE0BB0525254D72A3" pageId="0" pageNumber="19" type="comments">
|
||||
<paragraph id="CC731E42CECD27673ED5E5D877AE3FAC" pageId="0" pageNumber="19">Comments.</paragraph>
|
||||
<paragraph id="B602CEB0976FFA5241375EC86CC7674B" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="604005BA75D1786CBE353FE1B242AC6A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis id="A88E9A0B59CA4F1698A62246BF738169" italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
was described by J.
|
||||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
<bibRefCitation id="D52A280BBE625EB7034642E02824AB57" author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||||
from Aldabra Island in the Seychelles.
|
||||
<bibRefCitation author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot (1952)</bibRefCitation>
|
||||
<bibRefCitation id="F2B42BC0C6C497353871F529F389FC76" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="19" pagination="17 - 31" refId="B28" refString="Guignot, F, 1952. Description de dytiscides inedits de la collection Regimbart. . Revue Francaise d'Entomologie 19: 17 - 31" title="Description de dytiscides inedits de la collection Regimbart." volume="19" year="1952">Guignot (1952)</bibRefCitation>
|
||||
named a non-striolated female variety from Madagascar as var.
|
||||
<taxonomicName lsidName="var. simplex" pageId="0" pageNumber="19" rank="variety" variety="simplex">simplex</taxonomicName>
|
||||
<taxonomicName id="6320DB8B53B6AFC89E99AB3F09CF059B" lsidName="var. simplex" pageId="0" pageNumber="19" rank="variety" variety="simplex">simplex</taxonomicName>
|
||||
, and again in 1961 as var.
|
||||
<taxonomicName lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
<taxonomicName id="01055AEF5E9A68786C10E2BB1AA79C98" lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
(non-available infrasubspecific name) from Andranofotsy (E of Maroantsetra NE Madagascar), and stated (1961) that females of
|
||||
<taxonomicName lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
<taxonomicName id="E0AFB741120AB49700C34B48671AE951" lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="6897791BEF6ED4C97039D768C70AC4D6" italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
are normally striolated, but that on Madagascar the non-striolated var.
|
||||
<taxonomicName lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
<taxonomicName id="7202BF3258DC05BFC021BC6AC5D6EF39" lsidName="var. aequabilis" pageId="0" pageNumber="19" rank="variety" variety="aequabilis">aequabilis</taxonomicName>
|
||||
is most common or even ubiquitous. We have found at least one female on Madagascar (from Masoala NP) with the elytra entirely striolated (the
|
||||
<normalizedToken originalValue="“normal”">"normal"</normalizedToken>
|
||||
<normalizedToken id="C67781B249DEB4A091F5753FE653D09E" originalValue="“normal”">"normal"</normalizedToken>
|
||||
variety sensu
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
<bibRefCitation id="760BF322108614A1F77BFFD58181A600" pageId="0" pageNumber="19" refId="B36">Guignot 1961</bibRefCitation>
|
||||
), but indeed the form where striolation is rather weak and only at outer elytral intervals is most common.
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">
|
||||
<normalizedToken originalValue="Guignot’s">Guignot's</normalizedToken>
|
||||
<bibRefCitation id="FDCA43709D0507432D091FB2E421EDBF" pageId="0" pageNumber="19" refId="B36">
|
||||
<normalizedToken id="F2FFC31FB7CA2FBBB3A943B7C98DA5F7" originalValue="Guignot’s">Guignot's</normalizedToken>
|
||||
(1961)
|
||||
</bibRefCitation>
|
||||
distinction between female forms either having or lacking strioles is, however, a simplification. All females we have seen have at least some weak tendencies of striolation on outer intervals and the extent can vary significantly between individuals.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="19">
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<paragraph id="EACA3B26F522D91E4FE4E85D4C9F714D" pageId="0" pageNumber="19">
|
||||
<taxonomicName id="209DE82B900B99FCE9D6A0520840FF48" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="FD35C807FCAB41D06ABCEA91D4987F40" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(Laporte, 1835) has been treated as a junior synonym of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
<taxonomicName id="E1B64A0D43518FE1DE4D3F0C6A1EE2A3" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||||
<emphasis id="ACC937B3BD55CA36C5B0868E5F8CCF53" italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
Klug, 1834 since
|
||||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.59232" author="Aube, C" journalOrPublisher="Mequignon Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B2" refString="Aube, C, 1838. Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean. . Mequignon Pere et Fils, Paris" title="Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean." url="https://doi.org/10.5962/bhl.title.59232" year="1838">
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<bibRefCitation id="099505FB880E15E8DEEC8BF3B34644C6" DOI="https://doi.org/10.5962/bhl.title.59232" author="Aube, C" journalOrPublisher="Mequignon Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B2" refString="Aube, C, 1838. Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean. . Mequignon Pere et Fils, Paris" title="Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean." url="https://doi.org/10.5962/bhl.title.59232" year="1838">
|
||||
<normalizedToken id="049E4FC64F5F7C7FF44E6854AB8FAA69" originalValue="Aubé">Aube</normalizedToken>
|
||||
(1838)
|
||||
</bibRefCitation>
|
||||
. We discovered the type material of
|
||||
<normalizedToken originalValue="Laporte´s">Laporte's</normalizedToken>
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<normalizedToken id="67FE8665A3513315B47D81B2AA34A96C" originalValue="Laporte´s">Laporte's</normalizedToken>
|
||||
<taxonomicName id="39307F5FA5AC13C3B148AB2C428AED3A" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="E9C1799F5FE432BD7EEA503124A50A6A" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
in MNHN, Paris. This could be identified based on a folded blue label bearing the information "
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
<taxonomicName id="7300DBA999DEFD0BACC4DDC5961F0053" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="95193FC12E03756C976F4EFF525C6EE9" italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Buquet,
|
||||
<taxonomicName lsidName="pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
<taxonomicName id="5E8E15DD9F4635CF34AB30A8D20C7CE3" lsidName="pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="764A819E76E1B9792797BA06A105C6A6" italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
var
|
||||
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
|
||||
<normalizedToken id="9A11627127FA3FDBED40A396C715FB1A" originalValue="Aubé">Aube</normalizedToken>
|
||||
, h. in Senegal, D. Buquet", which fits perfectly the description by
|
||||
<bibRefCitation author="Laporte, FLN" journalOrPublisher="Mequignon-Marvis Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B44" refString="Laporte, FLN, 1835. Caumont de 1834-1835: Etudes entomologiques. Premiere partie. Carnassiers. . Mequignon-Marvis Pere et Fils, Paris" title="Caumont de 1834 - 1835: Etudes entomologiques. Premiere partie. Carnassiers." year="1835">Laporte (1835)</bibRefCitation>
|
||||
<bibRefCitation id="7AC3E78DF52522E65ACA6B655A9DDF4B" author="Laporte, FLN" journalOrPublisher="Mequignon-Marvis Pere et Fils, Paris" pageId="0" pageNumber="19" refId="B44" refString="Laporte, FLN, 1835. Caumont de 1834-1835: Etudes entomologiques. Premiere partie. Carnassiers. . Mequignon-Marvis Pere et Fils, Paris" title="Caumont de 1834 - 1835: Etudes entomologiques. Premiere partie. Carnassiers." year="1835">Laporte (1835)</bibRefCitation>
|
||||
. The genitalia of this male show without any doubt that it is a synonym of the younger name
|
||||
<taxonomicName lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
<taxonomicName id="1A0DD97D3CD3428B24D6AEC8FFE9190C" lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="A05034EE32E6C75CB5524F6C87DE469C" italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
and not of
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="DAEFDDE55C776EFC8029B7FAD09A23B6" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="30D6942F77EE06BCDF0B8925BDE4A27B" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
. The name
|
||||
<taxonomicName lsidName="marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">marginipennis</emphasis>
|
||||
<taxonomicName id="C825EA8D896BD65D52CD4D33628F54D2" lsidName="marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="1A7992EBA78AF6AD2DE6593AF00E4CEB" italics="true" pageId="0" pageNumber="19">marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
has to our knowledge not been used as a valid name after 1899, fulfilling the first of the two criteria for protecting a younger currently used synonym from an older available name (ICZN 23.9.1.1). However,
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
<taxonomicName id="2F0491F8DD3EE5F327FD2604B78EF7CA" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus aldabricus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="aldabricus">
|
||||
<emphasis id="C6BA2CB1E03EA15E89543863BFB4D72C" italics="true" pageId="0" pageNumber="19">Copelatus aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
is a relatively recent name from a region with a low-intensity taxonomic research and we do not believe it fulfils the second criteria of at least 25 papers published by at least ten authors in the immediately preceding 50 years and encompassing a span of not less than ten years (ICZN 23.9.1.2). Therefore it is not justifiable to protect the younger synonym
|
||||
<taxonomicName lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
<taxonomicName id="69B553B49CC3EB40A2A30670D015546A" lsidName="C. aldabricus" pageId="0" pageNumber="19" rank="species" species="aldabricus">
|
||||
<emphasis id="3A189CA69D689DA6361B60D72F66A84B" italics="true" pageId="0" pageNumber="19">C. aldabricus</emphasis>
|
||||
</taxonomicName>
|
||||
and the older
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="79793047C0461D7254A33E3D8E0F13D6" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="ED7C7F59CF79685C47803294323E2B4E" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is therefore brought back as the valid name of this species. We designate the discovered syntype of
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
<taxonomicName id="8B2C89E6A6A70BD9A882FA3C94E4270E" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||||
<emphasis id="67D3B2173C51AFED1BE177027B516BB4" italics="true" pageId="0" pageNumber="19">Colymbetes marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Laporte, 1835 in the MNHN collection as lectotype to preserve stability of the name. At the same time as this type specimen was discovered, also a likely
|
||||
<normalizedToken originalValue="“type”">"type"</normalizedToken>
|
||||
<normalizedToken id="CCA92B32A6C25541775A2C48643998AF" originalValue="“type”">"type"</normalizedToken>
|
||||
specimen of the nomen nudum
|
||||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus fimbriolatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriolatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus fimbriolatus</emphasis>
|
||||
<taxonomicName id="53DC810DA1BE766FCEBB4C90CD393F49" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus fimbriolatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriolatus">
|
||||
<emphasis id="8FA78D637B32EF632D0747C93F5D5F1D" italics="true" pageId="0" pageNumber="19">Copelatus fimbriolatus</emphasis>
|
||||
</taxonomicName>
|
||||
Dejean, 1837 from "Ile de France" [= Mauritius] was found with the same type of blue folded label (see supplementary file 1: NHRS-JLKB000065420). This nomen nudum is currently considered a synonym of
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="5B07E8C4A4934621100368EF9B2F8E33" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="E565493D9302D6A75261BB145B0E1594" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
Klug (Nilsson and Hajek 2018). Unfortunately, it is a female specimen, which is why its identity cannot be established with certainty, but it is more likely identical with
|
||||
<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
<taxonomicName id="84DB7ADD31A4C5055F8620DF54A2C62F" lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
|
||||
<emphasis id="FD98A15ED88DF2C14349CFD0B93D29C9" italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
|
||||
</taxonomicName>
|
||||
since
|
||||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
<taxonomicName id="DC754C35892516A42D289FFCF76C5A98" lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||||
<emphasis id="3B2BB97A0CC29615593E99EDF92D7150" italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||||
</taxonomicName>
|
||||
is not yet known from any of the Mascarene Islands (
|
||||
<bibRefCitation pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
<bibRefCitation id="C9AD417EBAF400A89229AE9B54CFECE5" pageId="0" pageNumber="19" refId="B21">Gomy 2016</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
Loading…
Reference in a new issue