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<document id="5746118F73EE5D47B3FDEE3F4ADC79F8" ID-DOI="10.3897/herpetozoa.37.e122825" ID-ZooBank="11B82D36-D464-429B-9C3B-D22423B8FBC6" ID-publisher-id="122825" URI-arpha="DE04D54E-C773-5EF0-95A6-23528E54505A" URI-zoobank="http://zoobank.org/11B82D36-D464-429B-9C3B-D22423B8FBC6" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1731951019993" checkinUser="pensoft" docAuthor="Naveen, R. S., Chandramouli, S. R., Babu, S., Ryndongsngi, A. M., Karunakaran, P. V. &amp; Kumara, Honnavalli N." docDate="2024" docId="20BC62AC2BF75AFEA9C4959A922DBEE0" docLanguage="en" docName="Herpetozoa 37: 359-372" docOrigin="Herpetozoa 37" docSource="https://herpetozoa.pensoft.net/article/122825/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Raorchestes asakgrensis Naveen, Chandramouli &amp; Babu 2024, sp. nov." docType="treatment" docUuid="3AC56A6A-DBAE-4F49-9B8C-D425E38E0B79" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="372" masterDocId="DE04D54EC7735EF095A623528E54505A" masterDocTitle="Rediscovery and redescription of Ixalus garo Boulenger, 1919, and Ixalus kempiae Boulenger, 1919, with a reassessment of the taxonomic status of Raorchestes cangyuanensis Wu, Suwannapoom, Xu, Murphy &amp; Che, 2019 and the description of a new species from the Garo Hills of Meghalaya" masterLastPageNumber="372" masterPageNumber="359" pageNumber="359" updateTime="1731956179489" updateUser="admin">
<mods:mods id="FD4A3D9A7AB7BE64C266F27A3075B737" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC26E62DD64FD445F7C1C9939D9B6937">
<mods:title id="3F238EF8451BC55630A61E7C0E5997EC">Rediscovery and redescription of Ixalus garo Boulenger, 1919, and Ixalus kempiae Boulenger, 1919, with a reassessment of the taxonomic status of Raorchestes cangyuanensis Wu, Suwannapoom, Xu, Murphy &amp; Che, 2019 and the description of a new species from the Garo Hills of Meghalaya</mods:title>
</mods:titleInfo>
<mods:name id="CAAA542E6CB74A7A7EE1C040B9897652" type="personal">
<mods:role id="A8E3C1320E10A444D2B91394D0B1A30D">
<mods:roleTerm id="DBD83D7FDAC1C0B2B2D7932AC98863B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B206083CCBCF7DD0D50D79C67A9F68B">Naveen, R. S.</mods:namePart>
<mods:affiliation id="F61A2F2B46696255339835D4B6603604">Sálim Ali Centre for Ornithology and Natural History (South India Centre of Wildlife Institute of India), Anaikatty, Coimbatore - 641108, Tamil Nadu, India &amp; EDGE of Existence Programme, Conservation and Policy, Zoological Society of London, London, NW 1 4 RY, UK</mods:affiliation>
</mods:name>
<mods:name id="AF4100690EC3B017C279215983A7B2CF" type="personal">
<mods:role id="D82F87F8722593478E4425DD36A21822">
<mods:roleTerm id="414451126DC64635AAD179EB0D412726">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8313C4EDD11570464E17E2A7E5E0D643">Chandramouli, S. R.</mods:namePart>
<mods:nameIdentifier id="4304B331AD7EB1C89B64DEE4BAFA2967" type="ORCID">0000-0003-0626-0527</mods:nameIdentifier>
<mods:affiliation id="57D634B542416AD35C81D61DE0462882">Department of Ecology and Environmental Sciences, School of Life Sciences, Pondicherry University, Puducherry, India</mods:affiliation>
</mods:name>
<mods:name id="9DE5AB3ACF64E5B74D475600E1E5F1E9" type="personal">
<mods:role id="9E6792F05FF1E99AE57B3FA09F71465A">
<mods:roleTerm id="A800D4FDA8F7DE497BE72383DE0814B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A5C4C5A9017165E3510F21715AD7BDD">Babu, S.</mods:namePart>
<mods:affiliation id="71BE38A8C98889386F059441AFA27107">Sálim Ali Centre for Ornithology and Natural History (South India Centre of Wildlife Institute of India), Anaikatty, Coimbatore - 641108, Tamil Nadu, India</mods:affiliation>
</mods:name>
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<mods:role id="F0E5222ABCA710C0456D8C3AAA402FF4">
<mods:roleTerm id="712378B10BE58A99E20E6175CB8158C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E1EDC00F62052BC955E4AED52A32B86">Ryndongsngi, A. M.</mods:namePart>
<mods:affiliation id="DF70B0DD8490A3A6C668F4512C988AE2">House No. 80, Iewmawlong, Nongpoh, Ri Bhoi District, Meghalaya- 793102, India</mods:affiliation>
</mods:name>
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<mods:roleTerm id="6E0FC3BF675486275273BD1D59110117">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E1234596EB081D6964BCA50A6800FD3">Karunakaran, P. V.</mods:namePart>
<mods:nameIdentifier id="4078D3E565B7C162F8D5229217720A6B" type="ORCID">0000-0003-4376-6068</mods:nameIdentifier>
<mods:affiliation id="DA7CD935514F03E64AF1561002EC9D9C">Sálim Ali Centre for Ornithology and Natural History (South India Centre of Wildlife Institute of India), Anaikatty, Coimbatore - 641108, Tamil Nadu, India</mods:affiliation>
</mods:name>
<mods:name id="6C84B62202C27CF2F3FA42C197E566E8" type="personal">
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</mods:role>
<mods:namePart id="8F176970054A9DB0A7D9C5EA2D7A0D84">Kumara, Honnavalli N.</mods:namePart>
<mods:affiliation id="577CF5FF01B35AD85F0659A834CA7570">Sálim Ali Centre for Ornithology and Natural History (South India Centre of Wildlife Institute of India), Anaikatty, Coimbatore - 641108, Tamil Nadu, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8B11432B128BB2B45C9896DEF8900440">text</mods:typeOfResource>
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<mods:titleInfo id="D8D8A4BE76EEFB7F514A5CE72099A30C">
<mods:title id="A03340B11657038C98D85F7DA4F24749">Herpetozoa</mods:title>
</mods:titleInfo>
<mods:part id="E96E4325FD4C059BF45A1D0A724AAA7D">
<mods:date id="FA4066B2A27DB80A3139BC34E6EBD883">2024</mods:date>
<mods:detail id="3EE2D82D11383306C06819887331C407" type="pubDate">
<mods:number id="684A64F1B602D16B7C9F74CE20F428FB">2024-11-18</mods:number>
</mods:detail>
<mods:detail id="9D7355305C682AD3187053C6562DB122" type="volume">
<mods:number id="491EB077DE6B76C9AA33D868A0F877D2">37</mods:number>
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<mods:classification id="A4AF8B9027249DD323FF6449B3F18EDE">journal article</mods:classification>
<mods:identifier id="19CD5E12671CEB2A44E4459EDF59C506" type="DOI">10.3897/herpetozoa.37.e122825</mods:identifier>
<mods:identifier id="C9028440FBB2268E5A578F85D9DD1DB0" type="ZooBank">11B82D36-D464-429B-9C3B-D22423B8FBC6</mods:identifier>
</mods:mods>
<treatment id="20BC62AC2BF75AFEA9C4959A922DBEE0" ID-arpha="20BC62AC-2BF7-5AFE-A9C4-959A922DBEE0" ID-zoobank="https://zoobank.org/3AC56A6A-DBAE-4F49-9B8C-D425E38E0B79" LSID="urn:lsid:zoobank.org:act:3AC56A6A-DBAE-4F49-9B8C-D425E38E0B79" LSID-ZBK="urn:lsid:zoobank.org:act:3AC56A6A-DBAE-4F49-9B8C-D425E38E0B79" httpUri="http://treatment.plazi.org/id/20BC62AC2BF75AFEA9C4959A922DBEE0">
<subSubSection id="078CEC15B4178112A86183BB7FC8313D" type="nomenclature">
<paragraph id="4716D186C7A5E47B3395ADD49A2BB23A">
<taxonomicName id="21FB291EF4E626E37CB691019BED7C79" authority="Naveen, Chandramouli &amp; Babu" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis" status="sp. nov.">
<emphasis id="B4BD3EE9531706E840A874C4C7B63083" italics="true">Raorchestes asakgrensis</emphasis>
Naveen, Chandramouli &amp; Babu
</taxonomicName>
<taxonomicNameLabel id="158E363A1CE9FDA2BA7C969ECDC376B7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFZBI" type="material">
<paragraph id="236A8392F646AA1ED548E13E6BCB0B0B">
<heading id="F172CE862A2AB7A9B2FFAD26DCB2E9E5" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="0BCFFB64CF47264ACC17544C1B5A4140">
<materialsCitation id="FC8338BB44FF78CCEE6CE362CE5817F5" collectingDate="2023-08-15" collectionCode="SACON" country="India" elevation="174" latitude="25.36788" location="Eman Asakgre Community Reserve" longLatPrecision="1" longitude="90.54344" specimenCode="VA 805" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<emphasis id="5A2D8D72E2EFFCFDFB228A5956451897" bold="true" italics="true">
<typeStatus id="3F387201057766ACE6371BD67FECD5AE" type="holotype">Holotype</typeStatus>
</emphasis>
: An
<specimenCount id="F90D8D4511EFB570F3A33993F3BF213A" count="1" type="female">adult female</specimenCount>
,
<collectionCode id="31873396327C60AC08534D1A165F2C91" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
<specimenCode id="5342FF7080847C8DB830DD0653B0A608" collectionCode="SACON" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">VA 805</specimenCode>
(Fig.
<figureCitation id="5E8CD58807E0A9F19213F636C862A523" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Adult female holotype (SACON VA 805) and adult male paratype (SACON VA 343) of Raorchestes asakgrensis sp. nov. in preservative, A. dorsal; C. ventral of holotype, and B. dorsal; D. ventral of paratype. Scale bars: 10 mm." figureDoi="10.3897/herpetozoa.37.e122825.figure6" httpUri="https://binary.pensoft.net/fig/1177124">6</figureCitation>
) from
<location id="E1E4F34EE5A81735DA8138AF25DD2F72" LSID="urn:lsid:plazi:treatment:20BC62AC2BF75AFEA9C4959A922DBEE0:E1E4F34EE5A81735DA8138AF25DD2F72" country="India" latitude="25.36788" longLatPrecision="1" longitude="90.54344" name="Eman Asakgre Community Reserve" stateProvince="Meghalaya">Eman Asakgre Community Reserve</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXZBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.543440,25.367880]}">
<geoCoordinate id="5B09FED8767FAB948B61D2A8E2A2565E" degrees="25.36788" direction="north" orientation="latitude" precision="1" value="25.36788">25.36788 ° N</geoCoordinate>
,
<geoCoordinate id="A23D1785C220F7112246380F542F3895" degrees="90.54344" direction="east" orientation="longitude" precision="1" value="90.54344">90.54344 ° E</geoCoordinate>
</named-content>
,
<quantity id="35B67242B42FB151B32DFA4510FD0CA2" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">
<elevation id="C13F9AEFAC5AFA4EF6F2C80C6C50C77A" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">174 m</elevation>
</quantity>
),
<collectingRegion id="919EB40D085F4417636C8FBF78DD3652" country="India" name="Meghalaya">Meghalaya</collectingRegion>
,
<collectingCountry id="CBCC34641D60A3AED7A2CA9469A61C44" name="India">India</collectingCountry>
, collected by RSN on
<date id="EC3DBFF775E31CEE952B1A306CF623DF" value="2023-08-15">
<collectingDate id="3252BE4A1648B4477432E84D2B52F98D" value="2023-08-15">
August 15
<superScript id="1B1B019275680B0E650E04C913451530">th</superScript>
, 2023
</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="A0344705E70E3ABF8FB90E364397EA2B">
<materialsCitation id="0F5CCD2D66B7EA9F72A8AEDC0DCD25F9" collectingDate="2023-08-15" collectionCode="SACON" collectorName="RSN" country="India" elevation="174" latitude="25.36788" location="Eman Asakgre Community Reserve" longLatPrecision="1" longitude="90.54344" specimenCode="VA 339, VA 343" specimenCount="3" specimenCount-male="3" stateProvince="Meghalaya" typeStatus="paratype">
<emphasis id="2C538A3872C3B4A1371C07B033753838" bold="true" italics="true">
<typeStatus id="751C92F7FDC4A3D5E7724526B9A17039" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="8277B62074F0B0AAFA6566CEC122B100" count="3" type="male">Three adult males</specimenCount>
,
<collectionCode id="20AF91427CE6F3CCDB578149C29868CF" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
<specimenCode id="54E6003B88073AEEF958D9E8413A81ED">VA 339</specimenCode>
, 340, and
<specimenCode id="652C208078D0A9EBF777BD64C3FBAA17">VA 343</specimenCode>
, from
<location id="5A6984402BBC1CA9015B120680E71804" LSID="urn:lsid:plazi:treatment:20BC62AC2BF75AFEA9C4959A922DBEE0:5A6984402BBC1CA9015B120680E71804" country="India" latitude="25.36788" longLatPrecision="1" longitude="90.54344" name="Eman Asakgre Community Reserve" stateProvince="Meghalaya">Eman Asakgre Community Reserve</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EI1BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.543440,25.367880]}">
<geoCoordinate id="630A0D12544485670B475A3B71ACA05E" degrees="25.36788" direction="north" orientation="latitude" precision="1" value="25.36788">25.36788 ° N</geoCoordinate>
,
<geoCoordinate id="1298F7E0A1DCF8D26D5BB5C0FCDE2C07" degrees="90.54344" direction="east" orientation="longitude" precision="1" value="90.54344">90.54344 ° E</geoCoordinate>
</named-content>
,
<quantity id="AF61E9092A99481E0EBB162E931520D3" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">
<elevation id="DAE3DAFC20662BE6C18378721E503ACD" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">174 m</elevation>
</quantity>
),
<collectingRegion id="13C9A80C08EC8ACCDAB2620C6362A1BF" country="India" name="Meghalaya">Meghalaya</collectingRegion>
,
<collectingCountry id="B366275985E90C9FE0FD2E01718DA2A9" name="India">India</collectingCountry>
, collected by
<collectorName id="BDA590B82A78C8F148C9E53C8DAAFD31">RSN</collectorName>
on
<date id="FFFD91985C18BB45DBC945580B7F720D" value="2023-08-15">
<collectingDate id="F4A50D90B387A16809289B8A87E40734" value="2023-08-15">
August 15
<superScript id="A9310A66D851F2B6DBB8C7A7270A8854">th</superScript>
, 2023
</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EP1BI" type="diagnosis">
<paragraph id="B7971F66960AC7F2F42C226574CFA320">
<heading id="8B98595F417F9C5BB610570A29E3BAED" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="571AA35E2A474777A8DB3FC552E84241">
<taxonomicName id="27CC03740EE011AAC85C3D158C7F64D9" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis" status="sp. nov.">
<emphasis id="38C957FD438731579823B7EE4F3F3F52" italics="true">Raorchestes asakgrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0111CBB925B4476709532109408FB8C6" rank="species">sp. nov.</taxonomicNameLabel>
is placed in the genus
<taxonomicName id="EE091FA1ED2B51D28E180F5F7DB7BC6B" authorityName="Biju, Shouche, Dubois, Dutta &amp; Bossuyt" authorityYear="2010" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="95E04897A4BC775A18D18E24E76E952C" italics="true">Raorchestes</emphasis>
</taxonomicName>
due to the combination of the following characters: small body size, vomerine teeth absent, single translucent external subgular vocal sac present, and tips of all fingers and toes expanded into discs with circum-marginal grooves (see
<bibRefCitation id="E76D13B1601D151D4DA9CDFDC2F78E95" author="Biju" etAl="et al." firstAuthor="Biju" journalOrPublisher="Current Science, Bangalore" pagination="1119-1125" refId="B3" refString="Biju SD, Shouche YS, Dubois A, Dutta SK, Bossuy F (2010) A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India. Current Science, Bangalore 98: 11191125." title="A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India." volume="98" year="2010">Biju et al. 2010</bibRefCitation>
). The species can be distinguished from congeners in the Indo-China region by the following combination of characters: A small-sized (mean
<abbrev id="ABBRID0EL2BI" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="174B3309C1919BEEC22183BCD49787EE" metricMagnitude="-2" metricUnit="m" metricValue="2.049" unit="mm" value="20.49">20.49 mm</quantity>
) arboreal species with an obtusely pointed snout; tympanum fairly visible with a visible supra-tympanic, relatively weakly developed; vomerine teeth absent; single semi-transparent gular vocal sac; tongue without papilla; ventral aspect of the hind limb uniform white as the venter; finger and toe discs well developed; toe webbing I 1-2 II 2 - 2 III 2 - 1 IV 1 - 2 V. Phylogenetically,
<taxonomicName id="EF4682E9A0038ED2495B9981ED0E76DB" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis">
<emphasis id="CBC55A4FB763B86CBEEC1430E97C9684" italics="true">R. asakgrensis</emphasis>
</taxonomicName>
forms a well-supported clade (
<collectionCode id="B35FEAA502A1245D590DA7ACCC25DF8F">BS</collectionCode>
: 100) consisting of two more Genbank sequences from India (
<ext-link id="A1772105362EB5E5A47588EB4C7D4B5D" ext-link-type="gen" xlink_href="JX092712.1" xlink_type="simple">
<specimenCode id="873998CD7F21B26E01FEC08102E3DFF1">
<collectionCode id="DB70815F461DBEDA2AAD0DFD516692AA">JX</collectionCode>
092712.1
</specimenCode>
</ext-link>
) and Nongkhyllem,
<collectingRegion id="87ECBC034244A4CCDFA2985FB2579AE2" country="India" name="Meghalaya">Meghalaya</collectingRegion>
(
<ext-link id="5C367FA109095470EF47F36777F673FA" ext-link-type="gen" xlink_href="MN524578.1" xlink_type="simple">
<specimenCode id="4C46DB743A1A86FF93A324BCE9DD5F53">
<collectionCode id="C0696596BD39F6A00A98D245F7C57846" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro">MN</collectionCode>
524578.1
</specimenCode>
</ext-link>
).
</paragraph>
<caption id="160A7557D3836556A258419192525026" ID-DOI="10.3897/herpetozoa.37.e122825.figure6" ID-arpha="64AA2432-BA07-5971-AE90-14B5B998423F" httpUri="https://binary.pensoft.net/fig/1177124" startId="F6">
<paragraph id="7ACE306E647BD4F394FD667E54C7774C">
<label id="6EFE707FC2DAA985AC7FCA1ADF34286B">Figure 6.</label>
</paragraph>
<paragraph id="63B87B6156BD7AA097782957EC68B833">
Adult female holotype (SACON VA 805) and adult male paratype (SACON VA 343) of
<taxonomicName id="D4C2583C328E127A5F77897CD56E4CAA" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis" status="sp. nov.">
<emphasis id="DB2DE0B9D1B38D1D800CF0846A867336" italics="true">Raorchestes asakgrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAE43AD68E59ADA0F5A30881E6F1E016" rank="species">sp. nov.</taxonomicNameLabel>
in preservative,
<emphasis id="98D78A300331118DC506A32FB0757B0A" bold="true">A.</emphasis>
dorsal;
<emphasis id="2A01077B1C641D1657B499349E81DFB9" bold="true">C.</emphasis>
ventral of holotype, and
<emphasis id="D54FB4C9E8F137A341A144B3A74D4ADF" bold="true">B.</emphasis>
dorsal;
<emphasis id="E88F95610182A54DBBE0FF5736362240" bold="true">D.</emphasis>
ventral of paratype. Scale bars: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EE4BI" type="description">
<paragraph id="D943E4A79A67D8FBA405E9F1EAEE5565">
<heading id="37EA1F3CD090A95945F4DFD3526397A3" reason="title">
Description of the
<typeStatus id="75E087FBF1C9106C482E807952906896" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="FA9D6A87D9565F3EFEED459D5A1ABDB2">
A small-sized
<taxonomicName id="21E30FD4756FA0A69CA4067617A1310D" authorityName="Biju, Shouche, Dubois, Dutta &amp; Bossuyt" authorityYear="2010" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="F499DE33C203880D09684BFC1529F4EA" italics="true">Raorchestes</emphasis>
</taxonomicName>
(
<abbrev id="ABBRID0ER4BI" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="82ED6F0A6EA3631197185A2E94E5A08F" metricMagnitude="-2" metricUnit="m" metricValue="2.28" unit="mm" value="22.8">22.8 mm</quantity>
), dorsal and lateral skin smooth, venter granulated. Head wider than long (
<abbrev id="ABBRID0EV4BI" xlink_title="head length">HL</abbrev>
:
<abbrev id="ABBRID0EZ4BI" xlink_title="head width">HW</abbrev>
0.82); large (
<abbrev id="ABBRID0E44BI" xlink_title="head length">HL</abbrev>
:
<abbrev id="ABBRID0EB5BI" xlink_title="snoutvent length">SVL</abbrev>
0.28). A short snout (
<abbrev id="ABBRID0EF5BI" xlink_title="eye diameter">ED</abbrev>
:
<abbrev id="ABBRID0EJ5BI" xlink_title="eyesnout distance">ES</abbrev>
0.9), obtusely pointed in dorsal view. Canthus rostralis evident and rounded; loreal region concave. Trunk short (AG:
<abbrev id="ABBRID0EN5BI" xlink_title="snoutvent length">SVL</abbrev>
0.43) and less gracile (AG:
<abbrev id="ABBRID0ER5BI" xlink_title="body width">BW</abbrev>
0.95); eyes large (
<abbrev id="ABBRID0EV5BI" xlink_title="eye diameter">ED</abbrev>
:
<abbrev id="ABBRID0EZ5BI" xlink_title="head length">HL</abbrev>
0.42); nostrils rounded, situated closer to the snout tip than to the eyes (
<abbrev id="ABBRID0E45BI" xlink_title="eyenostril distance">EN</abbrev>
:
<abbrev id="ABBRID0EB6BI" xlink_title="eyesnout distance">ES</abbrev>
0.53), directed laterally with no visible rim. Tympanum fairly visible, rounded in shape (
<abbrev id="ABBRID0EF6BI" xlink_title="tympanum diameter">TYD</abbrev>
<quantity id="EBFC9A741ADBEBCD9B57FEEFCC0AC54A" metricMagnitude="-3" metricUnit="m" metricValue="1.92" unit="mm" value="1.92">1.92 mm</quantity>
); Supra-tympanic fold relatively weakly developed; Inter-orbital space broader than inter-narial space (
<abbrev id="ABBRID0EJ6BI" xlink_title="interorbital distance">IO</abbrev>
:
<abbrev id="ABBRID0EN6BI" xlink_title="internarial distance">IN</abbrev>
1.14). Upper arms short (
<abbrev id="ABBRID0ER6BI" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0EV6BI" xlink_title="snoutvent length">SVL</abbrev>
0.18), shorter than lower arms (
<abbrev id="ABBRID0EZ6BI" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0E46BI" xlink_title="lower arm length">LAL</abbrev>
0.79); palm shorter than the upper arms (
<abbrev id="ABBRID0EBACI" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0EFACI" xlink_title="palm length">PAL</abbrev>
0.75); relative finger lengths III&gt; IV&gt; II&gt; I. Subarticular tubercles distinct in both palmar and plantar surfaces, rounded; Supernumerary tubercles present in the palm; no webbing between fingers; fingers with well-developed rounded discs. Thighs short, nearly half as long as the body (
<abbrev id="ABBRID0EJACI" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0ENACI" xlink_title="snoutvent length">SVL</abbrev>
0.48), slightly shorter than the tibia (
<abbrev id="ABBRID0ERACI" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0EVACI" xlink_title="tibia length">TBL</abbrev>
0.89). Foot slightly shorter than the thigh (
<abbrev id="ABBRID0EZACI" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0E4ACI" xlink_title="foot length">FOL</abbrev>
1.12), toes, short with well-developed rounded discs and partial webbing, webbing formula I 1-2 II 2 - 2 III 2 - 1 IV 1 - 2 V. In life dorsum uniformly brown with a faint “) - (“ mark; loreal yellowish brown; iris golden brown. Venter uniformly white, pale white eggs visible through ventral surface. Forearm and hindlimbs with dark brown bands (Fig.
<figureCitation id="690BAA317CAE1E4CFEE827AA6C7942A7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Raorchestes asakgrensis sp. nov. holotype, SACON VA 805, an adult female dorso-lateral view in life (above); paratype, SACON VA 340, an adult male dorso-lateral view in life (below)." figureDoi="10.3897/herpetozoa.37.e122825.figure7" httpUri="https://binary.pensoft.net/fig/1177125">7</figureCitation>
).
</paragraph>
<caption id="0934452035BDBFF10572F42F31711B84" ID-DOI="10.3897/herpetozoa.37.e122825.figure7" ID-arpha="A7B40314-F932-5431-95D4-093044DF9A48" httpUri="https://binary.pensoft.net/fig/1177125" startId="F7">
<paragraph id="8678489930C488BD3DE0C0E5A7420527">
<label id="42BAFBB1A5B446BD639A9B44A91A9B1D">Figure 7.</label>
</paragraph>
<paragraph id="401510116A1E13798D9C8A1031F85AA1">
<taxonomicName id="51280FD5295925B5B36CC4E0E3A4555F" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis" status="sp. nov.">
<emphasis id="D73066695089F9FF11C57E69D8CF038A" italics="true">Raorchestes asakgrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E47390ADA71CFF047984F43C31BC78D3" rank="species">sp. nov.</taxonomicNameLabel>
holotype, SACON VA 805, an adult female dorso-lateral view in life (above); paratype, SACON VA 340, an adult male dorso-lateral view in life (below).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E4BCI" type="variation">
<paragraph id="A7C452B09A8CB6C66B487D2F45107628">
<heading id="7CAE25EE1CAEBA7B504D2D1D17CCD41E" reason="title">Variation.</heading>
</paragraph>
<paragraph id="B7C33F5F11A4D039034EFED1EB3A4EE8">
Measurement of the
<typeStatus id="8FAD55EF0F538727FC04020633BD2093" type="paratype">paratypes</typeStatus>
presented in Table
<tableCitation id="F74E3301D6B9EB612A1FB49251AFB300" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric measurements (in mm) of R. asakgrensis sp. nov." httpUri="http://table.plazi.org/id/AE9FEFE530F8A974EF9C9F163A4772F6" tableUuid="AE9FEFE530F8A974EF9C9F163A4772F6">4</tableCitation>
. Variations noted during examination of preserved specimens and live individuals observed in the field. The single known female (
<typeStatus id="E1D87671D72EEBC7625681380F486600" type="holotype">Holotype</typeStatus>
VA 805) is larger (
<abbrev id="ABBRID0EHCCI" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="EF80F6B88C7CF4A8038B8CB60A7B6177" metricMagnitude="-2" metricUnit="m" metricValue="2.28" unit="mm" value="22.8">22.8 mm</quantity>
) than all the male specimens examined during this study (range
<quantity id="F36E2A707D8F1134876BA225A03950E2" metricMagnitude="-2" metricUnit="m" metricValue="2.009" metricValueMax="2.15" metricValueMin="1.8679999999999999" unit="mm" value="20.09" valueMax="21.5" valueMin="18.68">18.6821.5 mm</quantity>
; mean
<abbrev id="ABBRID0ELCCI" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="5E65AE019EE4FC9451ACAAEB285D90A4" metricMagnitude="-2" metricUnit="m" metricValue="1.972" unit="mm" value="19.72">19.72 mm</quantity>
). One of the male
<typeStatus id="9204EB74CB93F6609B9A0992F821E907" type="paratype">paratypes</typeStatus>
(VA 340) with a thin yellow line passing dorsally from the tip of the snout to the cloaca and extending through the thighs and shank (Fig.
<figureCitation id="1A83D16708674DFD2E951D78AF6008EA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Raorchestes asakgrensis sp. nov. holotype, SACON VA 805, an adult female dorso-lateral view in life (above); paratype, SACON VA 340, an adult male dorso-lateral view in life (below)." figureDoi="10.3897/herpetozoa.37.e122825.figure7" httpUri="https://binary.pensoft.net/fig/1177125">7</figureCitation>
). Dorsal coloration pale to dark brown with or without a faint or dark hourglass pattern, “) (“.
</paragraph>
<caption id="AE9FEFE530F8A974EF9C9F163A4772F6" ID-Table-UUID="AE9FEFE530F8A974EF9C9F163A4772F6" httpUri="http://table.plazi.org/id/AE9FEFE530F8A974EF9C9F163A4772F6" startId="T4" targetIsTable="true">
<paragraph id="8C8F991C120CEA2CB01E3162895733AD">
<label id="FE61CCB15ACFBC415E853C91839F5C58">Table 4.</label>
</paragraph>
<paragraph id="09714C0CC879A07C3420935C74447996">
Morphometric measurements (in mm) of
<taxonomicName id="260757E78A66C19CDE9309FE12829D1E" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis" status="sp. nov.">
<emphasis id="68EB5B7D26B1D67494320595E3FEF703" italics="true">R. asakgrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F70ACA0C117B9782BEA6E3A2CDC532BF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3D20562F39622E14CB1F627BE0545F72">
<table id="TID0EN6CI" rules="all">
<tbody id="B4995D49107E6F9BA452BADA7D2E4FE0">
<tr id="FC859AD84BDB61CCD8B2AC101DDBA770">
<th id="0136486415D4C38B12E56C0ACDE488E4" colspan="1" rowspan="1">Voucher no:</th>
<th id="FDDB562362397517C1C0BE5FADFBF9A6" colspan="1" rowspan="1">Sex</th>
<th id="AF54CBC7536EFAB1BAF3DC08CB356576" colspan="1" rowspan="1">
<abbrev id="ABBRID0EWDCI" xlink_title="snoutvent length">SVL</abbrev>
</th>
<th id="4847BBCC404064B5B04C5D8FCF3D6369" colspan="1" rowspan="1">AG</th>
<th id="C1BC38BF3E2D213298330203668A9765" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBECI" xlink_title="body width">BW</abbrev>
</th>
<th id="B609C003DC31169D050D81296B33E787" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJECI" xlink_title="head length">HL</abbrev>
</th>
<th id="2ED513424089DA4550433F078DFAAA07" colspan="1" rowspan="1">
<abbrev id="ABBRID0ERECI" xlink_title="head width">HW</abbrev>
</th>
<th id="21A87AB4B3F68E4D4FEB794865021D02" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZECI" xlink_title="head depth">HD</abbrev>
</th>
<th id="F5E76F668CAFF035A3C862BFA57E8E40" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBFCI" xlink_title="eye diameter">ED</abbrev>
</th>
<th id="79BDB4E983C7ED35DC99A604392A2F89" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJFCI" xlink_title="eyenostril distance">EN</abbrev>
</th>
<th id="6CD91EC615194B3541F8B29B1F1ADD6F" colspan="1" rowspan="1">
<abbrev id="ABBRID0ERFCI" xlink_title="eyesnout distance">ES</abbrev>
</th>
<th id="90DB368A9F850B918270D3C86E630D0B" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZFCI" xlink_title="tympanum diameter">TYD</abbrev>
</th>
<th id="A780720BB2F37E4FD390138C4B6CECC0" colspan="1" rowspan="1">ET</th>
<th id="96DE565EBFF5ED9D660B12AB517089CC" colspan="1" rowspan="1">
<abbrev id="ABBRID0EEGCI" xlink_title="upper eyelid width">UEW</abbrev>
</th>
<th id="D6FDAA85B7FC854B57FBFEDCE2FD53B7" colspan="1" rowspan="1">
<abbrev id="ABBRID0EMGCI" xlink_title="interorbital distance">IO</abbrev>
</th>
<th id="27B654976CA91F19E9E5BC29804DB126" colspan="1" rowspan="1">
<abbrev id="ABBRID0EUGCI" xlink_title="internarial distance">IN</abbrev>
</th>
<th id="D38CA28BBE68BDB8F9B0D89D6A6118F4" colspan="1" rowspan="1">
<abbrev id="ABBRID0E3GCI" xlink_title="upper arm length">UAL</abbrev>
</th>
<th id="45BE7D201DD30D377BED89CA3F478FCE" colspan="1" rowspan="1">
<abbrev id="ABBRID0EEHCI" xlink_title="lower arm length">LAL</abbrev>
</th>
<th id="7A75AA8B7DC7261592D2FCB2100D9865" colspan="1" rowspan="1">
<abbrev id="ABBRID0EMHCI" xlink_title="palm length">PAL</abbrev>
</th>
<th id="FA0D8AF096E2F88930A874D9D03B0652" colspan="1" rowspan="1">
<abbrev id="ABBRID0EUHCI" xlink_title="femur length">FEL</abbrev>
</th>
<th id="651A703CB467A98397446ECF63D3ECBF" colspan="1" rowspan="1">
<abbrev id="ABBRID0E3HCI" xlink_title="tibia length">TBL</abbrev>
</th>
<th id="F21230116A185D7D758C8B81211A0F41" colspan="1" rowspan="1">TRL</th>
<th id="0014C01C3009DC138D537AD196F4D2CA" colspan="1" rowspan="1">
<abbrev id="ABBRID0EHICI" xlink_title="foot length">FOL</abbrev>
</th>
</tr>
<tr id="7012ACCB1E3BE135823DC2DD68B92CC4">
<td id="603DF68CE89EBB8E9DC002940C444DE0" colspan="1" rowspan="1">SACON VA 805 Holotype</td>
<td id="AE64FFAC606C83692D96E6E4E4FC66E5" colspan="1" rowspan="1">F</td>
<td id="C50E013AB841DC65CE8239852299FCA7" colspan="1" rowspan="1">22.8</td>
<td id="9F8E488B9956967D41696C04B966269B" colspan="1" rowspan="1">10.02</td>
<td id="6090CF3E714AA33E17F5D9C19758836A" colspan="1" rowspan="1">10.5</td>
<td id="35603E54116C186894C6BB9EB62CD67A" colspan="1" rowspan="1">6.4</td>
<td id="2600CDD5B06D3DBE70892E9E7F640569" colspan="1" rowspan="1">7.72</td>
<td id="5D4A91BBB31A74C72CDAD7038BECE1DD" colspan="1" rowspan="1">4.5</td>
<td id="29C2C737AA8AADBC0638C2DBC7F95856" colspan="1" rowspan="1">2.7</td>
<td id="1522766360346F47FCF67F84AB1D111A" colspan="1" rowspan="1">1.6</td>
<td id="17E22F001F87D92BCD93723C84C4BB1B" colspan="1" rowspan="1">3</td>
<td id="CAB68B1DDF6E886E8A2866D312497139" colspan="1" rowspan="1">1.92</td>
<td id="97466E23E788EB70BC667BF1EB03D777" colspan="1" rowspan="1">0.44</td>
<td id="83F18D0E6348F949AE6ECC6EABBB4054" colspan="1" rowspan="1">1.6</td>
<td id="B41B227BAB8500745683623B7C217A5F" colspan="1" rowspan="1">3.2</td>
<td id="99279CC704D3891F0FDAE5A7C5403388" colspan="1" rowspan="1">2.8</td>
<td id="0A04462443AED709450AB6C88FFE33DB" colspan="1" rowspan="1">4.12</td>
<td id="63C26CCC3BCC59EDD27A9D034AB1D930" colspan="1" rowspan="1">5.18</td>
<td id="DDE6A05BD9084D4F6C249C4DB6DB8FAF" colspan="1" rowspan="1">5.5</td>
<td id="04B82803C44DDC94607E538879AE726D" colspan="1" rowspan="1">11.02</td>
<td id="0183517926094941DB3D7DF03C42C684" colspan="1" rowspan="1">12.42</td>
<td id="4136FA70DE4CAAF5959D79584B3E271A" colspan="1" rowspan="1">7.3</td>
<td id="D9A7422AD1296C042B1DA1F3FDEF9DCB" colspan="1" rowspan="1">9.82</td>
</tr>
<tr id="2EFE92FA1D6FCADBED9DBE562A001BB5">
<td id="E4F2552799B0401E3389E707349AA582" colspan="1" rowspan="1">SACON VA 343</td>
<td id="A97139E05E03F2CCB8A206CFED9C5E8C" colspan="1" rowspan="1">M</td>
<td id="B35F87CDD36F8C264DFFDB8D6C6DAF9D" colspan="1" rowspan="1">21.5</td>
<td id="D4099BEBE466ECB1CD06D0ACAC0BA472" colspan="1" rowspan="1">10.74</td>
<td id="CE4EC1B761D48BF923E6C1E4EFF2CF54" colspan="1" rowspan="1">8.14</td>
<td id="C41BD8E4AACDE87CBE1DF04F5E024018" colspan="1" rowspan="1">5.82</td>
<td id="1872996EF34B419FDBB11CDD205E45AF" colspan="1" rowspan="1">7.6</td>
<td id="58C4F1105CEDCB1A093232DD0B9C0363" colspan="1" rowspan="1">4.68</td>
<td id="AB840773F93C6DDBD5FB85F718EC3827" colspan="1" rowspan="1">2.6</td>
<td id="A985E59A8A3D70D30AEF77C5674AF118" colspan="1" rowspan="1">1.42</td>
<td id="4A124F64F25D21F9A2E9880C05EDD0BF" colspan="1" rowspan="1">3.18</td>
<td id="9C5CAC1037DAE27273E0EF479DC391C9" colspan="1" rowspan="1">1.18</td>
<td id="10BDE2AD2AB4FAB0B78E1C878EB64733" colspan="1" rowspan="1">0.52</td>
<td id="B73B0B60F51AE71EBCB00B4EF1C4A1B5" colspan="1" rowspan="1">1.84</td>
<td id="5F486A958E170B973F3EE9779036A66A" colspan="1" rowspan="1">2.98</td>
<td id="AFD0468B761CF3995288911FB1001BB4" colspan="1" rowspan="1">1.98</td>
<td id="D03092B041A987DF0F94F433AE182BB0" colspan="1" rowspan="1">3.82</td>
<td id="660BB40BEE663BCEA7783F3024D2B665" colspan="1" rowspan="1">5.08</td>
<td id="6D80D682BF93E7181534492D784EE982" colspan="1" rowspan="1">5.2</td>
<td id="D5194587F53297EE789CA065D16820BF" colspan="1" rowspan="1">10.64</td>
<td id="4B5191861C2A78E3BC20CAE838EACA54" colspan="1" rowspan="1">11.64</td>
<td id="81F0DFCC7867051DCF7798A1F7E2B884" colspan="1" rowspan="1">6.98</td>
<td id="1DD70362CA779863DA5B0FD34DD6C810" colspan="1" rowspan="1">8.3</td>
</tr>
<tr id="766633C28CEF975497E0926F1993B9FA">
<td id="8D3C54DB922B3C6EB6C73118186ADCC7" colspan="1" rowspan="1">SACON VA 340</td>
<td id="A00DDEA96B6AD2F18B35BC74BF8DA2B3" colspan="1" rowspan="1">M</td>
<td id="1C8B8149A1583054FED114BE77E1EC53" colspan="1" rowspan="1">18.68</td>
<td id="8E1AB02C719804F57FCF866502577959" colspan="1" rowspan="1">8.24</td>
<td id="72EBBFAF845F37392483C115C824F44A" colspan="1" rowspan="1">7.18</td>
<td id="68E4495688DB24E1F7DA4FD70C5B4F8A" colspan="1" rowspan="1">5.04</td>
<td id="714826F44265F983FBECDE2E4B069C3E" colspan="1" rowspan="1">7.14</td>
<td id="184224F613357D1DE19B6A19B26B14FF" colspan="1" rowspan="1">4.54</td>
<td id="41A1D4EF40A03763A983C8967CD47FED" colspan="1" rowspan="1">2.72</td>
<td id="C96A7D183C735B7D0769B282DA87E3FD" colspan="1" rowspan="1">1.42</td>
<td id="734C4FFB8D3C24BFB365A5647B781420" colspan="1" rowspan="1">2.54</td>
<td id="59C55B20CB0C58F9F4E3CC01DB506711" colspan="1" rowspan="1">0.8</td>
<td id="0A1328C11417F40325E8D19E7B37B1BA" colspan="1" rowspan="1">0.42</td>
<td id="46A56B01C391ADBA78BC6411673B10DE" colspan="1" rowspan="1">1.42</td>
<td id="31B55C2E3A43E416AEA89955439BBC6A" colspan="1" rowspan="1">2.42</td>
<td id="CA89D786EFDA10C61831DE4BEEC2D0F3" colspan="1" rowspan="1">1.8</td>
<td id="A94EADCFF96745393138E4C85061F68B" colspan="1" rowspan="1">3.68</td>
<td id="8B6100DA6B8F36911B789F36D83BFC34" colspan="1" rowspan="1">4.12</td>
<td id="B2BCE8C942506B1B0D59219ABC774CC0" colspan="1" rowspan="1">5</td>
<td id="F00181D3B68CC8A56C5D49D5C282AE68" colspan="1" rowspan="1">10</td>
<td id="25488A5296902F8A85DB1D596C778EA6" colspan="1" rowspan="1">10.68</td>
<td id="93291831C69D89782EAB267BB4DCE639" colspan="1" rowspan="1">6.44</td>
<td id="8D71EF4FC1CD2C0605222B072950033B" colspan="1" rowspan="1">7.24</td>
</tr>
<tr id="712E9D8B6E52E78E4EC13E89D400BB09">
<td id="4768F7972D7DFF10C9BBAC23230AE318" colspan="1" rowspan="1">SACON VA 339</td>
<td id="FF3A748F974B958983E5422C057A8696" colspan="1" rowspan="1">M</td>
<td id="C0751E06054F5E3BE9B35EF810A56B58" colspan="1" rowspan="1">18.98</td>
<td id="418F01C8C683279020F5C89FC2EF93DD" colspan="1" rowspan="1">7.92</td>
<td id="A4FFE368A0156DFD63C9DE62A83FF032" colspan="1" rowspan="1">8.68</td>
<td id="538A0708B32D2F0D07008627B704437D" colspan="1" rowspan="1">6.34</td>
<td id="D4434DF1DAB66CF9650CD3F57A5A26D9" colspan="1" rowspan="1">7.02</td>
<td id="B752C0E568968F84346EF30DB9617D70" colspan="1" rowspan="1">5.14</td>
<td id="89B168D074306888D90579674DFBE044" colspan="1" rowspan="1">3.12</td>
<td id="E738E91795E58D344DF5C27ED27C1744" colspan="1" rowspan="1">1.44</td>
<td id="362575E27497CC0EFB40F42B88DC444E" colspan="1" rowspan="1">3.02</td>
<td id="1FAD352BA2DDF84D57757DFEA30CAEE3" colspan="1" rowspan="1">0.92</td>
<td id="38084E6A2ACD34278FFF0ECB4B24A077" colspan="1" rowspan="1">0.52</td>
<td id="BC502EDB1F08236B905F0C0BCA02BDE3" colspan="1" rowspan="1">1.44</td>
<td id="9E812E6448F347142C78E6DAC6C44E1E" colspan="1" rowspan="1">2.34</td>
<td id="4C48FDC80C995D74E61F88C2152BC16C" colspan="1" rowspan="1">2.4</td>
<td id="9CE857864CB76B571CB15C328A044DE8" colspan="1" rowspan="1">4.94</td>
<td id="44C5F6AEB7CD781E86FB2ECE71EF5DF6" colspan="1" rowspan="1">4.12</td>
<td id="87D2759D2D8348D9C2886DB733475211" colspan="1" rowspan="1">4.22</td>
<td id="EC6879A399CF00D7F2B491F7EF9E3F7B" colspan="1" rowspan="1">9.3</td>
<td id="321496576FFDEE203DB042E14B1C8461" colspan="1" rowspan="1">10.54</td>
<td id="6D1C2D91D4DBEBACDBE7BD49C570525D" colspan="1" rowspan="1">6.6</td>
<td id="44E0FA8176DD67D8570D78AC8738C018" colspan="1" rowspan="1">7.24</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDRCI" type="comparisons">
<paragraph id="875805AE9CA88281D36A5A74CA5CC363">
<heading id="924F87044B9B4E71B07DB0C92B994A71" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="1EF4E8E89FEF66FA15AF404C0477899F">
<taxonomicName id="D8D0F8D6AC5A1071D01517F4A11A3563" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrensis" status="sp. nov.">
<emphasis id="DE9CAEC5A83F5FB364DD49263D7E12FE" italics="true">Raorchestes asakgrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="699E5D7E69586837BBC907ACB4F2AD41" rank="species">sp. nov.</taxonomicNameLabel>
is genetically divergent (5.810.8 %) from congeners, with the closest species being
<taxonomicName id="53B23A26B7D521DE0ABE6BBE4371BAD0" baseAuthorityName="Pillai &amp; Chanda" baseAuthorityYear="1973" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shillongensis">
<emphasis id="590937D28BDF06417FFC35AB742999CA" italics="true">R. shillongensis</emphasis>
</taxonomicName>
with a divergence of 5.8 %. On the phylogenetic tree, it clusters with a clade comprising
<taxonomicName id="338632E5C4D8B5B41A6D4635E0F6D4AA" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="77EEC6F9DEDA68BB31E30F607F3F1776" italics="true">R. garo</emphasis>
</taxonomicName>
and
<taxonomicName id="0224C848635D5107D866A6DBF4BA43B3" baseAuthorityName="Pillai &amp; Chanda" baseAuthorityYear="1973" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shillongensis">
<emphasis id="8F7A3E9CCCB9167F36C0D7844C7D8F06" italics="true">R. shillongensis</emphasis>
</taxonomicName>
, with a moderate node support (79). From species occurring in the Indo-China region, it can be distinguished by: having an obtusely pointed snout (vs. truncate and scarcely projecting in
<taxonomicName id="EAF1FC9FA4E52F797F0915CC1A94CE74" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="474EDCB423780ACE1E0F09D2FC416DDB" italics="true">R. garo</emphasis>
</taxonomicName>
, rounded in
<taxonomicName id="F69644AB6D4CD597EA87B74E0E275503" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="503B067DAF070E1350BEA73DC012907F" italics="true">R. kempiae</emphasis>
</taxonomicName>
,
<taxonomicName id="857BAFF4249B01FE10310E943AA4EB31" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="DF77DA6FD025A5B7336956543F27AA11" italics="true">R. menglaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EA7E917BAF8E184D9FAEC17295F443FB" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hillisi">
<emphasis id="9A1D5947687E81DEE52F696AB9A1ED61" italics="true">R. hillisi</emphasis>
</taxonomicName>
,
<taxonomicName id="CBDB74945FA6A0F6DB5DF57F41B257AA" authorityName="Du, Xu, Liu &amp; Yu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hekouensis">
<emphasis id="9E65881A487A435233C25A85203C0659" italics="true">R. hekouensis</emphasis>
</taxonomicName>
,
<taxonomicName id="31B0742E2B2FDB7659F3CCAFC2BD5067" authorityName="Huang, Liu, Du, Bernstein, Liu, Yang, Yu &amp; Wu" authorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="malipoensis">
<emphasis id="8EEE8D80175933B79315912E9C1E66DF" italics="true">R. malipoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="38E3B935E57C1FC7ACF9D7C99B87B91A" baseAuthorityName="Yang &amp; Li" baseAuthorityYear="1978" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longchuanensis">
<emphasis id="C5FB03AE7D9C9956C0FB19E57D65D296" italics="true">R. longchuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="958A8594FD88537939FA5322693A24D4" authorityName="Wu, Liu, Gao, Wang, Li, Zhou, Yuan &amp; Che" authorityYear="2021" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="dulongensis">
<emphasis id="D0D04214AF952DE7572B04DFC6B44B15" italics="true">R. dulongensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="B36FA0AF010AA9522C0D90879D03F666" authorityName="Jiang, Wang, Ren &amp; Li" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="huanglianshan">
<emphasis id="296071CDC6AE2EC17ED09D05D1F309C5" italics="true">R. huanglianshan</emphasis>
</taxonomicName>
and sharply projecting in
<taxonomicName id="64D2B442A0F523E743C32A1A793FE1B1" baseAuthorityName="Boulenger" baseAuthorityYear="1906" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="annandalii">
<emphasis id="D42711F690DCD4A35E9ACD7F89D260B7" italics="true">R. annandalii</emphasis>
</taxonomicName>
and
<taxonomicName id="54FDDBFF2FB3D3104DBE7686D50C0459" authorityName="Zhang, Shu, Liu, Dong &amp; Guo" authorityYear="2022" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yadongensis">
<emphasis id="A9F9900E1ED4765928D3F8CC65B139D5" italics="true">R. yadongensis</emphasis>
</taxonomicName>
), a fairly visible tympanum (vs. not externally visible in
<taxonomicName id="7B4EEB04E44E35FF9EF3F3B6A7AB6EF9" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="C23CEE7B77FEE3EE55B17D63CC22F318" italics="true">R. kempiae</emphasis>
</taxonomicName>
,
<taxonomicName id="8687CD23289A67B8BC3E11C4D4E2C242" baseAuthorityName="Pillai &amp; Chanda" baseAuthorityYear="1973" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shillongensis">
<emphasis id="8D04767C5111307B386A7C1B58FFBDAE" italics="true">R. shillongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EC058478EF80DFFE9B96FD265F8C9332" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="52A4F13D6728C63CE0D9DFB925CE6163" italics="true">R. rezakhani</emphasis>
</taxonomicName>
,
<taxonomicName id="E3E5073DE83B4D1B6A2B52C791AFDE57" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="E4B31F87CD9DE228E290686444F9CA82" italics="true">R. menglaensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="C821CD80174AB5B2593D5F0A539BD751" baseAuthorityName="Boulenger" baseAuthorityYear="1893" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="parvulus">
<emphasis id="390F18359FA66262ACD22C91AEFB72A3" italics="true">R. parvulus</emphasis>
</taxonomicName>
); and the presence of supernumerary tubercles on the palmar surface (vs. absent in
<taxonomicName id="5BA25D85379682FC563C5D087A7EE55D" authorityName="Wu, Liu, Gao, Wang, Li, Zhou, Yuan &amp; Che" authorityYear="2021" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="dulongensis">
<emphasis id="98E20676F1B6FDDE62522AB95C260DB7" italics="true">R. dulongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="ED90CB7CAC7698C8F5238647E761DE80" authorityName="Zhang, Shu, Liu, Dong &amp; Guo" authorityYear="2022" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yadongensis">
<emphasis id="26E0B7A9BE1725575AB3C1D8FB26CD37" italics="true">R. yadongensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="92C35815E07F9BFE61F60C78F894903B" authorityName="Du, Xu, Liu &amp; Yu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hekouensis">
<emphasis id="8EC22AB419D576257382F8972A22A8D3" italics="true">R. hekouensis</emphasis>
</taxonomicName>
). Ventral aspect of the hind limb uniformly creamy white in color (vs. dark orange in
<taxonomicName id="DEB168BADF50597F22700947269E37CB" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="ECE6A0BA49D7EB7DD797F44144C3270D" italics="true">R. garo</emphasis>
</taxonomicName>
and
<taxonomicName id="7A4CC0CC6FBFA80EC3E3B64BBA3BC09E" baseAuthorityName="Boulenger" baseAuthorityYear="1906" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="annandalii">
<emphasis id="CEA9CEFE3C67FC70414BD7662CA5F2CB" italics="true">R. annandalii</emphasis>
</taxonomicName>
, yellow with brown to black blotches in
<taxonomicName id="8A693DDE78EEDC54C4042D080E9D8FFB" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="838B0D4C3FECF34D98D66F3E2482A584" italics="true">R. kempiae</emphasis>
</taxonomicName>
, and pale brown with white flecks in
<taxonomicName id="78F9198C9A8BE39E3BE7A8729B7BF1BF" baseAuthorityName="Boulenger" baseAuthorityYear="1893" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="parvulus">
<emphasis id="921C91BF5E330F185053363CE698413F" italics="true">R. parvulus</emphasis>
</taxonomicName>
); toe webbing formula I 1-2 II 2-2 III 2-1 IV 1-2 V (vs. I 1-1 II 1-1 III 3-3 IV 3-2.5 V in
<taxonomicName id="24F988C860C744333489B73955E7245B" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="242A2BE18B582DD897AA25FA1E48012C" italics="true">R. garo</emphasis>
</taxonomicName>
, I 1-2 II 2 - 1 III 1 - 1 IV 1 - 2 V in
<taxonomicName id="BEE5C396E399C8EA55B5EA4E6AE00465" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="30DC0511E0DF2634593E380BA01D3CB6" italics="true">R. kempiae</emphasis>
</taxonomicName>
, I 22 II 22 III 23 IV 32 V in
<taxonomicName id="C729727A4D7BB54DFAB4350DA7BB65BB" authorityName="Huang, Liu, Du, Bernstein, Liu, Yang, Yu &amp; Wu" authorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="malipoensis">
<emphasis id="EC48A9069881816DB6EC5A5503467CE8" italics="true">R. malipoensis</emphasis>
</taxonomicName>
, II 12 III 12.5 IV 2.51 V in
<taxonomicName id="B690B6599BB330561A9F0BB27FB8FB19" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="79B3FA0B9733996CFB93E97E2970EF96" italics="true">R. menglaensis</emphasis>
</taxonomicName>
, II 12 III 12.5 IV 2.51 V in
<taxonomicName id="3BC4602E01C0F3C7BB8A61EDC0DED3FC" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hillisi">
<emphasis id="2334F901E4FFD69261976DCA98F7B203" italics="true">R. hillisi</emphasis>
</taxonomicName>
, II 12 III 12 IV 21 V in
<taxonomicName id="D83B95F3178DCD2A76DC506ABC114F77" authorityName="Jiang, Wang, Ren &amp; Li" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="huanglianshan">
<emphasis id="E6C259D51FE7DCB6A358A4B2CCA32A2E" italics="true">R. huanglianshan</emphasis>
</taxonomicName>
, I 22 II 1.5 3.25 III 2 3.5 IV 3.25 2 V in
<taxonomicName id="020638C3322916677F7D173B7A0F2AD7" baseAuthorityName="Boulenger" baseAuthorityYear="1893" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="parvulus">
<emphasis id="6711EF6C92DCF8756B64579B9A7EB44C" italics="true">R. parvulus</emphasis>
</taxonomicName>
and I 22 II 1.752 III 1.53 IV 2.752 V in
<taxonomicName id="584CD50A41C5927530C6382DC4B09C62" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="1C7A557FCE110C76AAF5CE6B4F3E05BC" italics="true">R. rezakhani</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK4CI" type="remarks">
<paragraph id="79E3A54BC0AC1E8573B863A50D8D543B">
<heading id="1FF10EE80947D6C4422F4996F3260317" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="CBE2BBA113135ADA449D52B844C42AAB">
The new species is clearly differentiated by a distinct set of morphological characters, particularly the presence of an externally visible tympanum and snout shape (see Fig.
<figureCitation id="06937EB10B97645C1688D2D43B94D949" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Raorchestes asakgrensis sp. nov. holotype, SACON VA 805, an adult female dorso-lateral view in life (above); paratype, SACON VA 340, an adult male dorso-lateral view in life (below)." figureDoi="10.3897/herpetozoa.37.e122825.figure7" httpUri="https://binary.pensoft.net/fig/1177125">7</figureCitation>
).
<bibRefCitation id="104E0C637195408B1E232DAAA348D2D7" DOI="10.11606/issn.2316-9079.v17i1p3-20" author="Boruah B &amp; Raj P &amp; Dutta SK &amp; Das A" etAl="et al." firstAuthor="Boruah" issue="1" journalOrPublisher="Phyllomedusa, Journal of Herpetology" pagination="3-20" refId="B5" refString="Boruah B, Raj P, Dutta SK, Das A (2018) Redescription and geographic distribution of Raorchestes shillongensis (Anura, Rhacophoridae) from Meghalaya, northeast India. Phyllomedusa, Journal of Herpetology 17 (1): 320. https://doi.org/10.11606/issn.2316-9079.v17i1p3-20" title="Redescription and geographic distribution of Raorchestes shillongensis (Anura, Rhacophoridae) from Meghalaya, northeast India." volume="17" year="2018">Boruah et al. (2018)</bibRefCitation>
reported a
<taxonomicName id="92FCBDDF5D90AF95E97E970B68B97BA8" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="85127AEE9C4FBC00EE5C3931267AA0AA" italics="true">Raorchestes</emphasis>
sp.
</taxonomicName>
from regions north of the Khasi Hills, and it is likely that this could be
<taxonomicName id="CCEAD20481798EB27E216BBE3BFD8850" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="asakgrenesis">
<emphasis id="DBE3F35A0AC565D63DB88D8D49B91AC7" italics="true">R. asakgrenesis</emphasis>
</taxonomicName>
, as we also recorded this species from near Nongkhyllem Wildlife Sanctuary.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK5CI" type="distribution">
<paragraph id="B5AF986FEF36419DFD5C812E8E2A013A">
<heading id="00BC2260CC051CB5193CB5CD7DCB37B5" reason="title">Natural history and distribution.</heading>
</paragraph>
<paragraph id="7527CE75E0394FFF44A6012D7DFE6649">
The frogs were seen calling at dusk from bushes in and around the Eman Asakgre Community Reserve. A single amplecting pair was recorded at around 19: 30 h perched on a bush
<quantity id="62BCC871620FCD17F540948932CB8DDD" metricMagnitude="0" metricUnit="m" metricValue="1.5" unit="m" value="1.5">1.5 m</quantity>
above the ground during the pre-monsoon season following the first shower in May. The sequence generated during the current study also matches closely with a sequence from Nongkhyllem (
<ext-link id="F5DE4FEFB961DAAA740B06A664F41DD7" ext-link-type="gen" xlink_href="MN524577.1" xlink_type="simple">
<specimenCode id="D4DE280E13C7E38F219CDDC21AC9A24F">
<collectionCode id="830B2C51409AB0FE0E0A74E389FA3795" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro">MN</collectionCode>
524577.1
</specimenCode>
</ext-link>
), foothills of Khasi hills (about
<quantity id="58677A0093F8F74C30FA6369743005CF" metricMagnitude="5" metricUnit="m" metricValue="1.5" unit="km" value="150.0">150 km</quantity>
away from the type locality) in
<collectingRegion id="FED443E2B4F89B9A1B69CF7496AC558F" country="India" name="Meghalaya">Meghalaya</collectingRegion>
. In addition to this, during this study this species was also recorded from various locations across Garo hills (Sasatgre, Oragitok, and Dangkipara) and low-elevation regions of Khasi hills (Umling and Nongpoh). We calculated the extent of occurrence of this species with the current data from this study, and it was estimated to be
<quantity id="C893D8ECC843E6AB358F1F58591C8A1B" metricMagnitude="6" metricUnit="m" metricValue="1.625" unit="km" value="1625.0">1,625 km</quantity>
<superScript id="84DB13482CFDDD29E0246DB4FA43A3A8">2</superScript>
across an elevational range of
<quantity id="F9F983AE6D2890B2C5D493287EE6F4B5" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
asl to
<quantity id="3DDA851C7D2F7462796AE00299B4F489" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</quantity>
asl; however, further surveys are required to robustly estimate the distribution range of this species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EX5CI" type="etymology">
<paragraph id="FF990F4030040960A719B6410C02C292">
<heading id="3544E4B9F4E92ED700C500FACAD4FAC8" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5E465C67536E05B97A0261C2BCDC42EB">The species is named after the type locality, Eman Asakgre Community Reserve, to honor the residents who greatly supported the surveys during which the type specimens were collected.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,821 @@
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<mods:title id="E0A54B53EDF21BAB724BFE65CE5ED15A">Rediscovery and redescription of Ixalus garo Boulenger, 1919, and Ixalus kempiae Boulenger, 1919, with a reassessment of the taxonomic status of Raorchestes cangyuanensis Wu, Suwannapoom, Xu, Murphy &amp; Che, 2019 and the description of a new species from the Garo Hills of Meghalaya</mods:title>
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<mods:affiliation id="D16C16146418E32D02D955D85162818F">Department of Ecology and Environmental Sciences, School of Life Sciences, Pondicherry University, Puducherry, India</mods:affiliation>
</mods:name>
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<mods:namePart id="F34A0B9D027675AECD765C7E849A3978">Kumara, Honnavalli N.</mods:namePart>
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<mods:title id="26F8CFDE919624BD33DD90C39A587253">Herpetozoa</mods:title>
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<mods:date id="EC397FD3C8A330BF8819E3D1904707D1">2024</mods:date>
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<treatment id="408622B7D1C75C9781B19D71ED1CE674" ID-arpha="408622B7-D1C7-5C97-81B1-9D71ED1CE674" LSID="urn:lsid:plazi:treatment:408622B7D1C75C9781B19D71ED1CE674" httpUri="http://treatment.plazi.org/id/408622B7D1C75C9781B19D71ED1CE674">
<subSubSection id="46B546A3F92662492FADA123431DFA75" type="nomenclature">
<paragraph id="95B25F64C2542930EFBBAB8AF045CBC2">
<taxonomicName id="21F4C1716F5934629EC3549F23332BBD" authority="(Boulenger, 1919)" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo" status="comb. nov.">
<emphasis id="B2A7AC5339CD9207FEFF093720209D47" italics="true">Raorchestes garo</emphasis>
(
<bibRefCitation id="8737CE263E71E7530305694E884ED0A9" DOI="10.26515/rzsi/v16/i2/1919/162810" author="Boulenger GA" issue="2" journalOrPublisher="Records of the Zoological Survey of India" pagination="207-208" refId="B7" refString="Boulenger GA (1919) XII Descriptions of Three new Batrachians from the Garo Hills, Assam. Records of the Zoological Survey of India 16 (2): 207 - 208. https: // doi. org / 10.26515 / rzsi / v 16 / i 2 / 1919 / 162810" title="XII Descriptions of Three new Batrachians from the Garo Hills, Assam." volume="16" year="1919">Boulenger, 1919</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="78789FFB8CC2EEAF049E27F64EC00D1A" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
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<paragraph id="C17FAC75F85ECDC8CFC7AF7743E6D830">
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<treatmentCitation id="CD68B2ADD18566051D25884BB5F77651">
<taxonomicName id="AA7ED3DD8422589D681358AF3C8E7A95" ID-CoL="3QFBY" authority="Boulenger 1919" authorityName="Boulenger" authorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Ixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="DA19173F9C3F0F4C4DEABA791E1F03DE" italics="true">Ixalus garo</emphasis>
<bibRefCitation id="6AEB3D9E2D99F20A4B1CECD784CB7E94" DOI="10.26515/rzsi/v16/i2/1919/162810" author="Boulenger" firstAuthor="Boulenger" issue="2" journalOrPublisher="Records of the Zoological Survey of India" pagination="207-208" refId="B7" refString="Boulenger GA (1919) XII Descriptions of Three new Batrachians from the Garo Hills, Assam. Records of the Zoological Survey of India 16 (2): 207208. https://doi.org/10.26515/rzsi/v16/i2/1919/162810" title="XII Descriptions of Three new Batrachians from the Garo Hills, Assam." volume="16" year="1919">Boulenger 1919</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="A30B0F9F94DBC075DE12DED0B9BB9BA8">
<treatmentCitationGroup id="42E391D58D0773F15C3920C2EE795638">
<treatmentCitation id="71FA1ECE0CEB3BEFA7BB9B68DCD7D39C">
<taxonomicName id="1E6C0D5E26350CF41C4FF12F2F4506BC" ID-CoL="6X659" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Rhacophorus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo" subGenus="Philautus">
<emphasis id="441A85C7247745BE094F54EBF982507E" italics="true">Rhacophorus</emphasis>
(
<emphasis id="1E6B99BFF6A95EBD2B20C8BB6758F691" italics="true">Philautus</emphasis>
)
<emphasis id="DF7737853AC1971253F2BADEEE9F7B5C" italics="true">garo</emphasis>
</taxonomicName>
<bibRefCitation id="50D26D85FAA552604331A517E22C393D" author="Ahl" firstAuthor="Ahl" refId="B1" refString="Ahl E (1931) Amphibia, Anura III, Polypedatidae. Das Tierreich 55: xvi + 477." year="1931">Ahl 1931</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F12130102D2107B0D5407EBEC938CAC0">
<treatmentCitationGroup id="4CEA1C53C8A38E1CBA4DFB12DDC8184C">
<treatmentCitation id="63949CAC05D4F849A98F46E6EB7F3852">
<taxonomicName id="294A34C75984333A599D5D0AD4F727AF" ID-CoL="4G5G9" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="F42BA5CD85CEC9FDFAE13426D8CD2D83" italics="true">Philautus garo</emphasis>
</taxonomicName>
<bibRefCitation id="F01333880ACAEE5094ED9B10989E0A5B" author="Bourret R" firstAuthor="Bourret" journalOrPublisher="Hanoi: Institut Océanographique de lIndochine" pagination="1-547" refId="B8" refString="Bourret R (1942) Les Batraciens de lIndochine. Hanoi: Institut Océanographique de lIndochine 6: 1547." title="Les Batraciens de lIndochine." volume="6" year="1942">Bourret (1942)</bibRefCitation>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0ECCBG" type="material">
<paragraph id="E913BA0E89F264B3B5D3F2AA0E76343F">
<heading id="7EFC2808F54AB1E0C6C7F6A2B0E7FB24" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="687C704F9370226A3E3610C8470A3381">
<materialsCitation id="24A8F390C8FA8195541C01B1CEB2D4D7" collectionCode="ZSI" collectorName="S. W. Kemp" location="Above Tura" specimenCode="ZSI 19187" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
<emphasis id="83987F1EE1655E879D45E8AEBB482BE9" bold="true" italics="true">
<typeStatus id="E51F221DA8E3643BF1D9EC8EB4046F86" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="1025E04E745CC48232A537976E44C2E6">ZSI 19187</specimenCode>
(Fig.
<figureCitation id="F59898C57A686A0E75500C7B19E44C9D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Ixalus garo (ZSI 19187) (above) (not to scale); SACON VA 809 in preservation (below); A. Dorsal view; and B. Ventral view. Scale bars: 10 mm." figureDoi="10.3897/herpetozoa.37.e122825.figure2" httpUri="https://binary.pensoft.net/fig/1177120">2</figureCitation>
above)
<specimenCount id="E10256E007F3E8D580192AEBD1B218F0" count="1" type="adult">sub adult</specimenCount>
, collected by
<collectorName id="7352F6A57F5025B13A7C97C31B4C3C72">S. W. Kemp</collectorName>
, from “
<location id="A85BF26AB834FDBAB79C46BBF206D496" LSID="urn:lsid:plazi:treatment:408622B7D1C75C9781B19D71ED1CE674:A85BF26AB834FDBAB79C46BBF206D496" name="Above Tura">Above Tura</location>
</materialsCitation>
.
</paragraph>
<paragraph id="F58D0FA0526C3D2CD7B5CFB208082DF8">
<materialsCitation id="DD65B7B9786E185C794AD5C27CD8C031" collectingDate="2022-04-26" collectionCode="SACON" collectorName="RSN" elevation="895" latitude="25.51" longLatPrecision="748" longitude="90.38" specimenCode="VA 809" specimenCount="308" specimenCount-male="308" typeStatus="topotype">
<emphasis id="406FC925AB1845BB5E615687E3532C84" bold="true" italics="true">
<typeStatus id="0927DDC86025BFAE185143A557ECF69E" type="topotype">Topotypes</typeStatus>
</emphasis>
:
<collectionCode id="B472176545E08FA270B1793FFD4559C1" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
<specimenCode id="593B78D40878C9149F7A4B1A27C2C312">VA 809</specimenCode>
(Fig.
<figureCitation id="3660F6F673682F1BDBCBDC7C01A4C334" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Ixalus garo (ZSI 19187) (above) (not to scale); SACON VA 809 in preservation (below); A. Dorsal view; and B. Ventral view. Scale bars: 10 mm." figureDoi="10.3897/herpetozoa.37.e122825.figure2" httpUri="https://binary.pensoft.net/fig/1177120">2</figureCitation>
below), 163, 305, 306,
<specimenCount id="CDDE377763F029804E46E103204FB10F" count="308" type="male">308 adult males</specimenCount>
, collected by
<collectorName id="4F956B367EE8E838B24C68DA1D02D0E6">RSN</collectorName>
on
<date id="1C220B94FFC822D0B705C873DD40A098" value="2022-04-26">
<collectingDate id="08EFC5B9CE422B6496A1552DF28B8800" value="2022-04-26">
April 26
<superScript id="6C5959A93D37EDB7DE92810E012C7879">th</superScript>
, 2022
</collectingDate>
</date>
from (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGDBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.380000,25.510000]}">
<geoCoordinate id="37FA804DA63E4799A2FD437635755E9D" degrees="25.51" direction="north" orientation="latitude" precision="555" value="25.51">25.51 ° N</geoCoordinate>
,
<geoCoordinate id="5B9E4889711758AC4AD49821B5D46694" degrees="90.38" direction="east" orientation="longitude" precision="555" value="90.38">90.38 ° E</geoCoordinate>
</named-content>
,
<quantity id="47A28A25BAACC52D3D82BF73D2675419" metricMagnitude="2" metricUnit="m" metricValue="8.95" unit="m" value="895.0">
<elevation id="F60CFF7C277136DBFDF2001033A6E9FE" metricMagnitude="2" metricUnit="m" metricValue="8.95" unit="m" value="895.0">895 m</elevation>
</quantity>
)
</materialsCitation>
, and
<materialsCitation id="F5FFAB04EA7B12B305AAA8975CE8243B" collectingDate="2020-06-23" collectionCode="SACON" country="India" county="East Garo Hills" elevation="1200" latitude="25.47297" location="Daribokgre Community Reserve" longLatPrecision="5" longitude="90.3148" specimenCode="VA 129" specimenCount="1" specimenCount-female="1" stateProvince="Meghalaya">
<collectionCode id="A7CAC15B974E1E2A3C401353F4E3FB80" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
<specimenCode id="6058B4EDB4543902066C025CFE4FA088">VA 129</specimenCode>
, an
<specimenCount id="DDEE468719B53D65EDAE6D7E49948A8F" count="1" type="female">adult female</specimenCount>
, on
<date id="10EEF1E2F7239DDAD19CCFBEB9AB813E" value="2020-06-23">
<collectingDate id="0CD5D8689CA4FDEA72BFA3604BE3CDE8" value="2020-06-23">
June 23
<superScript id="4110F067E8D3306B583460F4903D2075">rd</superScript>
, 2020
</collectingDate>
</date>
from
<location id="CC5CCEDD5F26222E83B4680A81CC70FE" LSID="urn:lsid:plazi:treatment:408622B7D1C75C9781B19D71ED1CE674:CC5CCEDD5F26222E83B4680A81CC70FE" country="India" county="East Garo Hills" latitude="25.47297" longLatPrecision="5" longitude="90.3148" name="Daribokgre Community Reserve" stateProvince="Meghalaya">Daribokgre Community Reserve</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQDBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.314800,25.472970]}">
<geoCoordinate id="A60F1FBDB5F0813A4361F714466DF7DF" degrees="25.47297" direction="north" orientation="latitude" precision="1" value="25.47297">25.47297 ° N</geoCoordinate>
,
<geoCoordinate id="F86E4FD8CCBD078F2273A02082626E78" degrees="90.3148" direction="east" orientation="longitude" precision="5" value="90.3148">90.3148 ° E</geoCoordinate>
</named-content>
,
<quantity id="3EE5ABE44977ABB0B2566D006BFC11E5" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="8B6AE39C075E8FCD19BFB1216BD015C1" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
),
<collectingCounty id="8AF32D67F29AE5CDAA3ACE833F78A31A">East Garo Hills</collectingCounty>
,
<collectingRegion id="7750BB03D8BD16605C5A7A829F16B250" country="India" name="Meghalaya">Meghalaya</collectingRegion>
,
<collectingCountry id="EF89EDB282A89D0B35856AF785F76F70" name="India">India</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVDBG" type="diagnosis">
<paragraph id="63AB02751D4DD67F33DE4DD9DE5CBADF">
<heading id="CBD9CDC3D5E1FA824291F8CCA4350B12" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="76006C8368C70DB930C5095648F1F78F">
<taxonomicName id="8E8C4B13257FB78FEFC7E36FAB0B66F4" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo" status="comb. nov.">
<emphasis id="4BA5D7FC3D153A7E8C745BD925E987C8" italics="true">Raorchestes garo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02D2C3CD1841D201CA346671807E8F06" rank="species">comb nov.</taxonomicNameLabel>
is placed in the genus
<taxonomicName id="7C25217935C2C8D7E001CDC1BA77665A" authorityName="Biju, Shouche, Dubois, Dutta &amp; Bossuyt" authorityYear="2010" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="916FE51A571C4113EFEA840D4B228EE3" italics="true">Raorchestes</emphasis>
</taxonomicName>
due to the combination of following characters: small body size, vomerine teeth absent, single translucent external subgular vocal sac present, and tips of all fingers and toes expanded into discs with circum-marginal grooves (see
<bibRefCitation id="12E6EB7D240859F0AF7E54A3A264AB03" author="Biju" etAl="et al." firstAuthor="Biju" journalOrPublisher="Current Science, Bangalore" pagination="1119-1125" refId="B3" refString="Biju SD, Shouche YS, Dubois A, Dutta SK, Bossuy F (2010) A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India. Current Science, Bangalore 98: 11191125." title="A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India." volume="98" year="2010">Biju et al. 2010</bibRefCitation>
). The species can be distinguished from congeners in the Indo-China region by the following combination of characters: A medium-sized (mean
<abbrev id="ABBRID0EREBG" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="329388885E3AFDC587A8A8B58C754814" metricMagnitude="-2" metricUnit="m" metricValue="2.092" unit="mm" value="20.92">20.92 mm</quantity>
) arboreal species with a truncated snout scarcely projecting beyond the mandible; tympanum small, fairly distinct, ovoid, and supra-tympanic fold evident; tongue without papilla; ventral aspect of the hind limb dark orange in color; finger and toe discs well developed; tips of all fingers and toes brown to yellowish orange; toe webbing I 1-1 II 1 - 1 III 3 - 3 IV 3 - 2.5 V. Phylogenetically,
<taxonomicName id="88F54744B54279C4619FB46007C8EC48" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="F14376CF8A479948B4581C59967CC926" italics="true">R. garo</emphasis>
</taxonomicName>
clusters with
<taxonomicName id="05C438FA28249583A1689B21D2716DD1" baseAuthorityName="Pillai &amp; Chanda" baseAuthorityYear="1973" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shillongensis">
<emphasis id="6B625BDC826E2E4D36B369E0C42F1D5F" italics="true">R. shillongensis</emphasis>
</taxonomicName>
with a low bootstrap support (ML BS: 51) and shows the lowest genetic divergence from
<taxonomicName id="55ACE3069B1542555F7D6EC42A232C30" baseAuthorityName="Yang &amp; Li" baseAuthorityYear="1978" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longchuanensis">
<emphasis id="3FCF861BB2F5E57C38317119EDB3A8BD" italics="true">R. longchuanensis</emphasis>
</taxonomicName>
(4.28 %).
</paragraph>
<caption id="18B307E36D56D78355F77ED5A8817593" ID-DOI="10.3897/herpetozoa.37.e122825.figure2" ID-arpha="17A84580-7DE6-5BBD-8A2C-0A15A436A2DC" httpUri="https://binary.pensoft.net/fig/1177120" startId="F2">
<paragraph id="7A7D8E7E94F4ECB5BEE57C9425EE6783">
<label id="72101FA3AAE25E2A9E94BFF272A0266D">Figure 2.</label>
</paragraph>
<paragraph id="C6D38A34AE2ED77CFCCF019D6F804584">
The holotype of
<taxonomicName id="72D621E06835D4A29626D4939CE5F5F6" authorityName="Boulenger" authorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Ixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="4FC9F832F2FA4F8BFFCCDD6D227ACBE1" italics="true">Ixalus garo</emphasis>
</taxonomicName>
(ZSI 19187) (above) (not to scale); SACON VA 809 in preservation (below);
<emphasis id="DB3E29E1C34B9FCBDEBDAB6FAEB242AF" bold="true">A.</emphasis>
Dorsal view; and
<emphasis id="0A721F8A43080775880181194997515C" bold="true">B.</emphasis>
Ventral view. Scale bars: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESGBG" type="description">
<paragraph id="C5B7EFF645747F4DEBA864D39BC6FA28">
<heading id="9C1102139F748263EEAD8F0AF734F0F3" reason="title">
Description of a
<typeStatus id="F72CA9F1D4EB494BCBBA8AE2D1EE5027" type="topotype">topotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="988406B357C2B5BAE310127AA1710B0D">
<emphasis id="6A67C24F43DEA730C4E657F36F653ECF" bold="true">
<collectionCode id="DD3BB8DE79602B108DA88B663875F67D" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
VA 809
</emphasis>
Medium-sized adult male (
<abbrev id="ABBRID0E1GBG" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="2A211C9BEF50666D10792FEF28FC3A9C" metricMagnitude="-2" metricUnit="m" metricValue="2.38" unit="mm" value="23.8">23.8 mm</quantity>
), dorsal skin mostly smooth with a few irregularly scattered spinules and lateral skin completely smooth, ventral skin slightly granulose, increasingly granulose towards the lower body. Head large (
<abbrev id="ABBRID0E5GBG" xlink_title="head length">HL</abbrev>
:
<abbrev id="ABBRID0ECHBG" xlink_title="snoutvent length">SVL</abbrev>
0.30), wider than long (
<abbrev id="ABBRID0EGHBG" xlink_title="head length">HL</abbrev>
:
<abbrev id="ABBRID0EKHBG" xlink_title="head width">HW</abbrev>
0.86). A short snout (
<abbrev id="ABBRID0EOHBG" xlink_title="eye diameter">ED</abbrev>
:
<abbrev id="ABBRID0ESHBG" xlink_title="eyesnout distance">ES</abbrev>
0.95), sharply pointed in dorsal view. Canthus rostralis evident and rounded; loreal region slightly concave. Trunk short (AG:
<abbrev id="ABBRID0EWHBG" xlink_title="snoutvent length">SVL</abbrev>
0.54) and stout (AG:
<abbrev id="ABBRID0E1HBG" xlink_title="body width">BW</abbrev>
1.20); eyes large (
<abbrev id="ABBRID0E5HBG" xlink_title="eye diameter">ED</abbrev>
:
<abbrev id="ABBRID0ECIBG" xlink_title="head length">HL</abbrev>
0.43); nostrils, rounded, with no visible rim, situated almost midway between the snout and the eye (
<abbrev id="ABBRID0EGIBG" xlink_title="eyenostril distance">EN</abbrev>
:
<abbrev id="ABBRID0EKIBG" xlink_title="eyesnout distance">ES</abbrev>
0.58), directed laterally; with a weakly developed rim around them. Tympanum distinct, ovoid (
<abbrev id="ABBRID0EOIBG" xlink_title="tympanum diameter">TYD</abbrev>
1.36). Inter-orbital space broader than inter-narial space (
<abbrev id="ABBRID0ESIBG" xlink_title="interorbital distance">IO</abbrev>
:
<abbrev id="ABBRID0EWIBG" xlink_title="internarial distance">IN</abbrev>
1.59). Upper arms short (
<abbrev id="ABBRID0E1IBG" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0E5IBG" xlink_title="snoutvent length">SVL</abbrev>
0.20), shorter than lower arms (
<abbrev id="ABBRID0ECJBG" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0EGJBG" xlink_title="lower arm length">LAL</abbrev>
0.75); palm shorter than the upper arms (
<abbrev id="ABBRID0EKJBG" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0EOJBG" xlink_title="palm length">PAL</abbrev>
0.74); relative finger lengths III&gt; IV&gt; II&gt; I. Subarticular tubercles distinct in both palmar and plantar surfaces, rounded; no webbing between fingers; fingers with well-developed ovoid-shaped discs. Thighs short, nearly half as long as the body (
<abbrev id="ABBRID0ESJBG" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0EWJBG" xlink_title="snoutvent length">SVL</abbrev>
0.46), nearly as long as the tibia (
<abbrev id="ABBRID0E1JBG" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0E5JBG" xlink_title="tibia length">TBL</abbrev>
0.97). Foot slightly shorter than the thigh (
<abbrev id="ABBRID0ECKBG" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0EGKBG" xlink_title="foot length">FOL</abbrev>
1.29), toes short, toe discs well developed and ovoid, smaller than finger discs, toes with partial webbing, webbing formula I 1-1 II 1 - 1 III 3 - 3 IV 3 - 2.5 V. Dorsum uniformly brown with yellowish white spinules; loreal region dark brown; iris golden brown. Venter white, speckled with black and gray spots, and lower trunk grayish brown. Groin orange (bordered by golden yellow) turning red as it reaches the thighs (Fig.
<figureCitation id="137837B26BB6A3C08D108AB079454D30" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Raorchestes garo (dorsolateral view of SACON VA 809, on top and dorsal view of SACON VA 809, bottom left and dorsal view of VA 305, adult male bottom right, showing hourglass pattern) from Garo Hills in life." figureDoi="10.3897/herpetozoa.37.e122825.figure3" httpUri="https://binary.pensoft.net/fig/1177121">3</figureCitation>
).
</paragraph>
<caption id="DF8E22F842F58726C423FE8FF0FE9EA2" ID-DOI="10.3897/herpetozoa.37.e122825.figure3" ID-arpha="5CF4C877-5A5E-5D4B-8996-72A2009E2812" httpUri="https://binary.pensoft.net/fig/1177121" startId="F3">
<paragraph id="65B96EDE6362EE9E0A35F68E66BB0823">
<label id="F395BF3D5C442D85FAC073B53F117DD2">Figure 3.</label>
</paragraph>
<paragraph id="9EDF59772DF8BF95E1EF06A4E89A8FD0">
<taxonomicName id="37FBE4544102627EE71F02E975AD5726" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="0F370DF5E91B30C34CF4A6B096D680E9" italics="true">Raorchestes garo</emphasis>
</taxonomicName>
(dorsolateral view of SACON VA 809, on top and dorsal view of SACON VA 809, bottom left and dorsal view of VA 305, adult male bottom right, showing hourglass pattern) from Garo Hills in life.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGLBG" type="variation">
<paragraph id="73A7B72BDDFF1A392B2FC71CBDF5DC75">
<heading id="6DBB25D34512EE4480788B1F2C94C70C" reason="title">Variation.</heading>
</paragraph>
<paragraph id="F3E1E183200CE401174EE0F563A80D3F">
Morphometric measurements of the
<typeStatus id="194762BBD5FA8E5BA90EB39BB62F5D18" type="holotype">holotypes</typeStatus>
and other preserved
<typeStatus id="B697E72F38ABF42883A7F0662FECA598" type="topotype">topotypes</typeStatus>
examined are presented in Table
<tableCitation id="15F9BBAD9926B3D115F36A8D2F6E452C" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric measurements (in mm) of Raorchestes garo (abbreviations explained in Material and methods)." httpUri="http://table.plazi.org/id/4FD57B1E0973AEFE169D0F91743A865A" tableUuid="4FD57B1E0973AEFE169D0F91743A865A">2</tableCitation>
. Variations noted during examination of preserved specimens and live individuals observed in the field. The single female specimen collected during this study is larger (
<abbrev id="ABBRID0EQLBG" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="BD196B5870471A0001E386CCE24FA090" metricMagnitude="-2" metricUnit="m" metricValue="2.61" unit="mm" value="26.1">26.1 mm</quantity>
) than all the males (range
<quantity id="46BEFE0E28587CC3BC892297BF270898" metricMagnitude="-2" metricUnit="m" metricValue="1.969" metricValueMax="2.38" metricValueMin="1.558" unit="mm" value="19.69" valueMax="23.8" valueMin="15.58">15.5823.8 mm</quantity>
; mean
<abbrev id="ABBRID0EULBG" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="E9062FF74DD4AB1C8A826CA4E1A410E6" metricMagnitude="-2" metricUnit="m" metricValue="2.092" unit="mm" value="20.92">20.92 mm</quantity>
). Dorsal coloration variable from light to reddish or dark brown. Some individuals with an hourglass pattern, intensity of the dorsal hourglass pattern variable from feeble to bold and black (usually absent in males during the breeding season). Some individuals with a darker hourglass pattern and light background (in VA 305), and some with lighter hourglass pattern and darker background (in VA 308). Bands on the limbs feeble to bold. Ventral coloration uniform in all individuals.
</paragraph>
<caption id="4FD57B1E0973AEFE169D0F91743A865A" ID-Table-UUID="4FD57B1E0973AEFE169D0F91743A865A" httpUri="http://table.plazi.org/id/4FD57B1E0973AEFE169D0F91743A865A" startId="T2" targetIsTable="true">
<paragraph id="326D741E8B77A621C6EEF8B0C1078935">
<label id="40CE2989528CCF4836785DB7A506B066">Table 2.</label>
</paragraph>
<paragraph id="5365BBE1BED69A42AE0380ABAAC400A6">
Morphometric measurements (in mm) of
<taxonomicName id="2B8E1F583204A9035080B5FF35E31948" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="FA57E7E349BD1AEC301E4B5AAB9B4C49" italics="true">Raorchestes garo</emphasis>
</taxonomicName>
(abbreviations explained in Material and methods).
</paragraph>
</caption>
<paragraph id="DA0769888B75E3535380655353EEAC19">
<table id="TID0EKLBI" rules="all">
<tbody id="E641CCFB3D32AA9F03D7005F61890D9C">
<tr id="A50E1A8E10D3937F31DEA7E8EF4ADCE3">
<th id="B74C8DB0C881DBDBE6ADF03CEFEF29DF" colspan="1" rowspan="1">Voucher no:</th>
<th id="D23A0A5FA4A0E331FBAB341E093540F7" colspan="1" rowspan="1">Sex</th>
<th id="B285AB7F1BD58993B7C834BF5F70C03B" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2MBG" xlink_title="snoutvent length">SVL</abbrev>
</th>
<th id="51AFCDCD0473E2385AD779173DE9BA77" colspan="1" rowspan="1">AG</th>
<th id="4A88FD26C1B6D44CFA2AB4554523302D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EGNBG" xlink_title="body width">BW</abbrev>
</th>
<th id="8BEB0C9F32630E75664600EB93233AA3" colspan="1" rowspan="1">
<abbrev id="ABBRID0EONBG" xlink_title="head length">HL</abbrev>
</th>
<th id="0289682D61446321B72CB17A17D00D81" colspan="1" rowspan="1">
<abbrev id="ABBRID0EWNBG" xlink_title="head width">HW</abbrev>
</th>
<th id="3AC3F5B79D078AC43BC97FF1C6722015" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5NBG" xlink_title="head depth">HD</abbrev>
</th>
<th id="688E32E59B89927686D9BB91E49DF36C" colspan="1" rowspan="1">
<abbrev id="ABBRID0EGOBG" xlink_title="eye diameter">ED</abbrev>
</th>
<th id="FEA29807A41DB8024C52E85D0B586CAE" colspan="1" rowspan="1">
<abbrev id="ABBRID0EOOBG" xlink_title="eyenostril distance">EN</abbrev>
</th>
<th id="FFD062AD684DDB32BF9FA2C0C60B71EE" colspan="1" rowspan="1">
<abbrev id="ABBRID0EWOBG" xlink_title="eyesnout distance">ES</abbrev>
</th>
<th id="941A79ED514786E9175CD970AD6EF1EF" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5OBG" xlink_title="tympanum diameter">TYD</abbrev>
</th>
<th id="E6BDC52D09CBD6815718F6DCF3044DB2" colspan="1" rowspan="1">ET</th>
<th id="797E00BD92D8E694200C2ED1BA3D1B68" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJPBG" xlink_title="upper eyelid width">UEW</abbrev>
</th>
<th id="32B136D2F85FE7337FF659D44E9E303B" colspan="1" rowspan="1">
<abbrev id="ABBRID0ERPBG" xlink_title="interorbital distance">IO</abbrev>
</th>
<th id="CA61ABDC426318E9069B154DCE8E05DF" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZPBG" xlink_title="internarial distance">IN</abbrev>
</th>
<th id="5CB566E5655D021C3B21432960B2F1DA" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBQBG" xlink_title="upper arm length">UAL</abbrev>
</th>
<th id="9795D86CF581169AA77812CABB6B824E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJQBG" xlink_title="lower arm length">LAL</abbrev>
</th>
<th id="EED6F82AC2D90726A36839C5A7B91429" colspan="1" rowspan="1">
<abbrev id="ABBRID0ERQBG" xlink_title="palm length">PAL</abbrev>
</th>
<th id="E1A0C52CF1B5D2587E8461FEC45A7114" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZQBG" xlink_title="femur length">FEL</abbrev>
</th>
<th id="AE3F833C101E1ABCE63E17C908DD4542" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBRBG" xlink_title="tibia length">TBL</abbrev>
</th>
<th id="35F8EC971AAF70A5DB801DE36AF5E083" colspan="1" rowspan="1">TRL</th>
<th id="E814D77D2A7C43EF7847AB47C36BFB91" colspan="1" rowspan="1">
<abbrev id="ABBRID0EMRBG" xlink_title="foot length">FOL</abbrev>
</th>
</tr>
<tr id="EE3F20799A27FFADCF437005C91CF134">
<td id="D98D1824091DC0B699BF9EE2E5F28347" colspan="1" rowspan="1">Holotype (ZSI 19187)</td>
<td id="E020ED675F2E0151311DE2D5D452119B" colspan="1" rowspan="1">NA</td>
<td id="A69B944DD81DA33E57009B5C8CB4AAA7" colspan="1" rowspan="1">12.6</td>
<td id="3FCE15BB7FCDF4A24BD0A020873F4762" colspan="1" rowspan="1">5.82</td>
<td id="7D7ACB91C799BB07402ED89AECEFA07D" colspan="1" rowspan="1">5.3</td>
<td id="36FACC210EF2EC54054A2EFF8EC4C290" colspan="1" rowspan="1">2.62</td>
<td id="BF531B320B7A75D2F3137FF2C6936919" colspan="1" rowspan="1">5.92</td>
<td id="84993773556D654791834FDF66E8D620" colspan="1" rowspan="1">3.5</td>
<td id="6DE2B375ECFA2720630528D120069B5F" colspan="1" rowspan="1">2</td>
<td id="0FFE54DE4F2FB9F47FA99BDADB60294D" colspan="1" rowspan="1">1.2</td>
<td id="38A80C39B6CC8F8E1112B2DE8C1E2B9D" colspan="1" rowspan="1">2.4</td>
<td id="D36DCB0D05F5C7D001A8DDE6B924F648" colspan="1" rowspan="1">0.3</td>
<td id="90191DFB0590AE3499015F7F151AA3AE" colspan="1" rowspan="1">0.4</td>
<td id="28E622F6C173865E3C0AF7246C159271" colspan="1" rowspan="1">0.72</td>
<td id="F72B36586C9482865178182FCFF90179" colspan="1" rowspan="1">1.7</td>
<td id="211FCB981364241B0D1A19B3186A28AE" colspan="1" rowspan="1">1.32</td>
<td id="1FCF1E5608A30E8ADA5CE31EE35C2DD2" colspan="1" rowspan="1">2.62</td>
<td id="39B6EF5BAAF8506683CBD8966C1838AF" colspan="1" rowspan="1">2.74</td>
<td id="76196AD4F14686A787E2076003451A76" colspan="1" rowspan="1">3.45</td>
<td id="72669C7200C68010404FDC1A8E5C7E2F" colspan="1" rowspan="1">7.26</td>
<td id="77CADC7BB1395FA73DD3383F6F24DEC4" colspan="1" rowspan="1">7.24</td>
<td id="47805879D5E109EE5C65197093E56F3E" colspan="1" rowspan="1">3.74</td>
<td id="FE6F15B6ACAEA5DF0E6806EA9D193664" colspan="1" rowspan="1">4.52</td>
</tr>
<tr id="874F62C78CB8FA081C74905BD3F6B2AE">
<td id="79CE0C5C2B77E833172A0832DF372323" colspan="1" rowspan="1">SACON VA- 809</td>
<td id="5588FEB3222E1CE806D868EF575C4835" colspan="1" rowspan="1">M</td>
<td id="982E7171BAA4FB58123E2F8A667F35CC" colspan="1" rowspan="1">23.8</td>
<td id="9FCBC04DD66724B73059881BD9DE0C85" colspan="1" rowspan="1">12.92</td>
<td id="9E5421D196ABC5D158391D946446A4B3" colspan="1" rowspan="1">10.8</td>
<td id="781FD1C35CB24F21007546BEAAAA4DE6" colspan="1" rowspan="1">7.18</td>
<td id="FE57F817728BF225CF27368FB5032BFD" colspan="1" rowspan="1">8.34</td>
<td id="34799F9FBA1FEC846ABF629C238D352D" colspan="1" rowspan="1">4.36</td>
<td id="A534A3D3DC9381225A10E6914E43C245" colspan="1" rowspan="1">3.12</td>
<td id="8B3AB7BEE6FB14DABA0DAECB738F5024" colspan="1" rowspan="1">1.92</td>
<td id="CB6B073367E340B7139241103D83695D" colspan="1" rowspan="1">3.3</td>
<td id="FC4B92C9C6D59F8C5ADEC31B76482313" colspan="1" rowspan="1">1.36</td>
<td id="AC60FE9279C525300C657370D41BF722" colspan="1" rowspan="1">0.7</td>
<td id="0C3136935DDD6BED6A7F89B4D5016FCA" colspan="1" rowspan="1">2.28</td>
<td id="745B4FCAC5B0E5465B929890B414CD12" colspan="1" rowspan="1">3.56</td>
<td id="CA8AC3B6C49549BD63B69525FABBF8AA" colspan="1" rowspan="1">2.24</td>
<td id="B9426CACE0FCE46891CFF944CC65844C" colspan="1" rowspan="1">4.66</td>
<td id="5871F4BC9F9891D574C1FE7FA4191567" colspan="1" rowspan="1">6.2</td>
<td id="AE8E9A3B05A5C702F5D112775B39B652" colspan="1" rowspan="1">6.32</td>
<td id="0EB800F6CE8A19E6E10883AD4F88994E" colspan="1" rowspan="1">10.98</td>
<td id="FC1FE39FEFBDB8713AA46D3B2D51FE3A" colspan="1" rowspan="1">11.34</td>
<td id="5AA26C195DB554C904D084E0BBCC598E" colspan="1" rowspan="1">7.3</td>
<td id="226CE6458979BEEBF50DB2146855AE6A" colspan="1" rowspan="1">8.54</td>
</tr>
<tr id="5916A336729C7CC23E31EA71CFBF2FA3">
<td id="32CE07D647891B2FF72FF48BD79117AC" colspan="1" rowspan="1">SACON VA 163</td>
<td id="1412734EA27888AD664562711CB911F7" colspan="1" rowspan="1">M</td>
<td id="7A769467E516105F0DD23AC68EE0471E" colspan="1" rowspan="1">21.38</td>
<td id="7AD8C085B2D56B1CF399759F5637D833" colspan="1" rowspan="1">9.72</td>
<td id="EED085403EC4422FA62BF4193D0AF1F4" colspan="1" rowspan="1">8.02</td>
<td id="AF4F747B00E4D62AC2F7894D8358EC46" colspan="1" rowspan="1">7.22</td>
<td id="DC8798CB42F6F98D1E85C8BB5154086B" colspan="1" rowspan="1">8.24</td>
<td id="182220F72199F082DD2DEBDFEE1AC0F9" colspan="1" rowspan="1">4.26</td>
<td id="924A7B463DF5D9C28148D8D9E0BA64A1" colspan="1" rowspan="1">3.04</td>
<td id="2EBC95504E69DFA858E73EEE4E3E69D4" colspan="1" rowspan="1">2.12</td>
<td id="9CB4E40B39C1AA82548345AB439DFBF3" colspan="1" rowspan="1">3.46</td>
<td id="B0B0CDCC18B93DE7697AC456D10F0180" colspan="1" rowspan="1">1.44</td>
<td id="2306F8CB715591026A27C6E9619C5864" colspan="1" rowspan="1">0.3</td>
<td id="8A179E86D745F3CD795E817B937F22C8" colspan="1" rowspan="1">1.68</td>
<td id="CF7269A1544FC7132C06FEA14DFC05DD" colspan="1" rowspan="1">3.08</td>
<td id="1EA1C6B04BD7A1CDC899F3D746FCD1E7" colspan="1" rowspan="1">2.3</td>
<td id="9A96A81B3B7FC3C6FF73E7A312C625B9" colspan="1" rowspan="1">4.28</td>
<td id="64F0C31ED951CE521397B374B3524AD3" colspan="1" rowspan="1">6.1</td>
<td id="EEBBF49E4DF5A427D587695D10867347" colspan="1" rowspan="1">5.38</td>
<td id="A3FF079A556643BEED2B61FD4EA5D4C5" colspan="1" rowspan="1">9.82</td>
<td id="B1CD0ACFB7804F04BC89126F0F2D7FAA" colspan="1" rowspan="1">11.1</td>
<td id="24035B890DF8DEABA02744E3A17D96BC" colspan="1" rowspan="1">6.36</td>
<td id="21940518B1F88619B7361F6DA8F41A13" colspan="1" rowspan="1">7.24</td>
</tr>
<tr id="279405E2E5A81BEC6821C84F08A05E02">
<td id="117204BCAE4BF386BF5DA53A18971EB7" colspan="1" rowspan="1">SACON VA- 305</td>
<td id="153D56D723C4B25EE58379DF1388CF7E" colspan="1" rowspan="1">M</td>
<td id="40EE9D92E6DDEB475DD83F8D998A0857" colspan="1" rowspan="1">22.16</td>
<td id="C234C77D8CE2F0B5F5D1B63534CD4BAA" colspan="1" rowspan="1">11.4</td>
<td id="BD32DED157174089FB876AA0E95FA449" colspan="1" rowspan="1">10.54</td>
<td id="8332CA23C40F562EFCC3989FA7D42774" colspan="1" rowspan="1">7.76</td>
<td id="8FFFAE2994088EDC105FA347785650E7" colspan="1" rowspan="1">8.5</td>
<td id="772B2D7DC9BB7C0BAF03B7EF06FEE69A" colspan="1" rowspan="1">4.98</td>
<td id="0F0AE5B32B74F9B10CF87B2A2639743F" colspan="1" rowspan="1">3.1</td>
<td id="C58B326EDDD3A5962249A27919A4F510" colspan="1" rowspan="1">1.4</td>
<td id="88B4FC1843DE17613EEF7D5158C4440F" colspan="1" rowspan="1">3.58</td>
<td id="8C882FBC014B96A2F1F3C48294163F82" colspan="1" rowspan="1">1</td>
<td id="585A759A3BA7634FC8045A8CF1A9058A" colspan="1" rowspan="1">0.32</td>
<td id="88F54AB7C19136A22C739B1D7DD5FC16" colspan="1" rowspan="1">1.76</td>
<td id="B651984A61D819B5F86A6F955771F0DD" colspan="1" rowspan="1">2.76</td>
<td id="23EB15296531218C8E904D8A0CC89D97" colspan="1" rowspan="1">2.06</td>
<td id="C01B4A20F40497E856E623CC1DAAAAB3" colspan="1" rowspan="1">4.62</td>
<td id="485AC1FC068CBD556F135FD4A5A8CA5C" colspan="1" rowspan="1">5.44</td>
<td id="26F04FADDF896DB22FDC40C6532D96C1" colspan="1" rowspan="1">6.54</td>
<td id="3D9A496ECAF09F7BA6813E61B47F7577" colspan="1" rowspan="1">10.34</td>
<td id="41499C7B1F8982B53059B6AE650818AF" colspan="1" rowspan="1">11.36</td>
<td id="E23873C622F45F5F6EE2A26E7A04120C" colspan="1" rowspan="1">6.8</td>
<td id="49E3D5B67FF964956C375EB546642A25" colspan="1" rowspan="1">9.06</td>
</tr>
<tr id="20C74A5CA5C97D714035874F6F62DAAE">
<td id="FB93E328E5A2307D0E34F6F3608C9813" colspan="1" rowspan="1">SACON VA- 306</td>
<td id="A5225E1FE3049B4FC6FDC3ADB598FE89" colspan="1" rowspan="1">M</td>
<td id="E43506D5C717AA99F14BAF2F2123D3C1" colspan="1" rowspan="1">15.58</td>
<td id="45E3EE575619BAB3E10DDD483DD8318B" colspan="1" rowspan="1">7.8</td>
<td id="B11195EF5DB97BD904D52BBE64A1C175" colspan="1" rowspan="1">7.02</td>
<td id="533568479F921229152A88BC019A2A62" colspan="1" rowspan="1">4.7</td>
<td id="0F7BFD0B8D209F15C42AA78CCF1EDF5C" colspan="1" rowspan="1">5.2</td>
<td id="C0FB839F5F514D0B0590EC745D7CAE3A" colspan="1" rowspan="1">3.92</td>
<td id="1B3D5D45722A72E2FE458BA50E891807" colspan="1" rowspan="1">2.3</td>
<td id="F1084551161E39FB1854CB1718DE4B0C" colspan="1" rowspan="1">1.3</td>
<td id="88044F6BEBF43B2175B286A60D7BAB15" colspan="1" rowspan="1">2.34</td>
<td id="3624D3008443E74E5160274F7446129D" colspan="1" rowspan="1">0.82</td>
<td id="134DA36D6C09ABA0CC1A685007BE2C09" colspan="1" rowspan="1">0</td>
<td id="6FD968299BEC8860287939D0BFEEA345" colspan="1" rowspan="1">1.42</td>
<td id="2E0C314FA4D32F13CB18A5AACEFD1F31" colspan="1" rowspan="1">2.06</td>
<td id="F0F0464DD731660DDD1495FE2CB4DACC" colspan="1" rowspan="1">1.5</td>
<td id="A12BC5A33E0362EF071CF7B63AB88B7F" colspan="1" rowspan="1">2.8</td>
<td id="EE2758150BE79CD6DAAFF7842D761627" colspan="1" rowspan="1">3.68</td>
<td id="BF24FF793AA35B1016C41A97C3493AB2" colspan="1" rowspan="1">3.5</td>
<td id="BE752079F842168D38F543F7AF346CD6" colspan="1" rowspan="1">7.2</td>
<td id="1CB2483653E9C8F5D8F93BAAEB707211" colspan="1" rowspan="1">7.96</td>
<td id="CBB5FF0DB5BF6954DE2B0B69FCD20A0B" colspan="1" rowspan="1">4.42</td>
<td id="2931AD1D3DE45A3A05FC5268DF8C0438" colspan="1" rowspan="1">5.46</td>
</tr>
<tr id="2ACAF3B15AA2D3E6D1610695A2C37C02">
<td id="C115EE8446DC9F0A2594D0C2BE0AA0EC" colspan="1" rowspan="1">SACON VA- 308</td>
<td id="89BD536437B34F75FA967F2F0078FB65" colspan="1" rowspan="1">M</td>
<td id="8FD8025C67B64D430A07EFEBB967BF22" colspan="1" rowspan="1">21.66</td>
<td id="F03027C8D9CA91D6309DA91A8ECF0FEB" colspan="1" rowspan="1">10.98</td>
<td id="A0EAF60699113111B85DDCA38F0439FB" colspan="1" rowspan="1">8.4</td>
<td id="6FD5B555F12B0903A9B7E8E26A7D1F10" colspan="1" rowspan="1">6.6</td>
<td id="825D10F5AF78A994A16CBFDC5C680BE7" colspan="1" rowspan="1">7.8</td>
<td id="1C8BAC40F751B1F8553FB8B8CCFCC735" colspan="1" rowspan="1">3.98</td>
<td id="355E8E3D8F772E16A6FEC67428792712" colspan="1" rowspan="1">2.94</td>
<td id="A78B7F234928D893C66BE701B796F57A" colspan="1" rowspan="1">1.4</td>
<td id="939268D49F5D2745039299E668F14955" colspan="1" rowspan="1">3.06</td>
<td id="D42F273D34F274094D5B677EA086C7B8" colspan="1" rowspan="1">1.4</td>
<td id="2CF34DE78363DE80CF3897630CD3FD04" colspan="1" rowspan="1">0</td>
<td id="7EF26D7AECE77F14D49F6D11E082314D" colspan="1" rowspan="1">1.68</td>
<td id="8034B3C7C350D5B3441D426B5740E3B1" colspan="1" rowspan="1">2.76</td>
<td id="36BD0B5921013B29EE0CA875362E9DAC" colspan="1" rowspan="1">2.24</td>
<td id="0DA528B2DE1A3B48B7EA235BD105AF5C" colspan="1" rowspan="1">3.44</td>
<td id="8962DA23663B3381F47CD967114CEDC3" colspan="1" rowspan="1">5.3</td>
<td id="BB4AF27C1FC745E15C962462D6D8624E" colspan="1" rowspan="1">6.1</td>
<td id="C74FCB375EF5D517EB31B2A0142E1955" colspan="1" rowspan="1">9.8</td>
<td id="3C6F8F9C4D0B5AED972BD9236733EA83" colspan="1" rowspan="1">10</td>
<td id="B425E6C3D778BF2A1EFC887D705C875B" colspan="1" rowspan="1">5.5</td>
<td id="50B9D0755D0ED51BC21C8B398CB45C73" colspan="1" rowspan="1">7.26</td>
</tr>
<tr id="7A74D0CD93885493152B3CEF208C9ACA">
<td id="0FADD98253E9FB14CCE05F126BF415ED" colspan="1" rowspan="1">SACON VA 129</td>
<td id="1F746D5FECD25BDA4FEB83868E81201C" colspan="1" rowspan="1">F</td>
<td id="CFD3BA2B83F029BBA1DC80DA84A04591" colspan="1" rowspan="1">26.1</td>
<td id="3AE9D64DF95F42176E352B3F056A8A0C" colspan="1" rowspan="1">13.2</td>
<td id="DF40A985B9EA4FDEF415D696AB1DCEA1" colspan="1" rowspan="1">11.9</td>
<td id="98EA468AA672A43447575082A06A9310" colspan="1" rowspan="1">8.62</td>
<td id="48199DB4A6FCA3814AA4B566BE68AB19" colspan="1" rowspan="1">8.46</td>
<td id="A971031E52CEC3004986E5C6C030AA50" colspan="1" rowspan="1">4.9</td>
<td id="B3E387C21018E1E802FA5673A800DF40" colspan="1" rowspan="1">3.5</td>
<td id="1DB077B48FDB3683E7ED6E956A43800C" colspan="1" rowspan="1">1.6</td>
<td id="A040F337FD0D995052343B80ECB74167" colspan="1" rowspan="1">4.02</td>
<td id="696836819B259FA972DD2DECE41A4572" colspan="1" rowspan="1">1.7</td>
<td id="4B5C0F943329764391FCEB1C3145FFA8" colspan="1" rowspan="1">0.86</td>
<td id="FF7173088869990009A80401B22864E4" colspan="1" rowspan="1">2.46</td>
<td id="11A6D5F31AA9F084EFF831D434834B4D" colspan="1" rowspan="1">4.8</td>
<td id="B8EDEB029B958955FD44E9CDFA805B76" colspan="1" rowspan="1">2.48</td>
<td id="92AD7FFD5664C8CC9DFE44B10390393A" colspan="1" rowspan="1">4.32</td>
<td id="F55796B0984E6AE840445F9554423DFD" colspan="1" rowspan="1">5.02</td>
<td id="39045C6B77428EA59EB0B8971CFCD6BA" colspan="1" rowspan="1">5.8</td>
<td id="6149653AA85FA001820D501E23B65407" colspan="1" rowspan="1">11.84</td>
<td id="27DCD43BA21998CBD522B8FE01C491F1" colspan="1" rowspan="1">12.42</td>
<td id="9A9166F3E54FB4EFA30B74EEC9F9EE70" colspan="1" rowspan="1">7.12</td>
<td id="9E32C8C1DF409351E8A2A29AF5A99C41" colspan="1" rowspan="1">8.58</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2AAI" type="comparisons">
<paragraph id="14E11F86B695288B3C435C22090CCD49">
<heading id="FD3A0101FB836A07E39DAA4EB2E62480" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="06BAB7388517016449A24D27F09B0585">
<taxonomicName id="82DB7833EA7E162DD885F60D0C0EC055" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="87C08E6BECB7F1D4A22FEF8075ABB1B0" italics="true">Raorchestes garo</emphasis>
</taxonomicName>
is genetically divergent (4.288.34 % on the 16 S rRNA gene) from congeners; the least divergent species is
<taxonomicName id="6FA1F817E62E28568ACCB518B3138DAF" baseAuthorityName="Yang &amp; Li" baseAuthorityYear="1978" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longchuanensis">
<emphasis id="46709FD7CB94BCEEBD747B5AD92703AD" italics="true">R. longchuanensis</emphasis>
</taxonomicName>
, with a divergence of 4.28 %. However, on the phylogenetic tree, it clusters with
<taxonomicName id="18B0C7DE49ADF6E9D2615662A2B0B19C" baseAuthorityName="Pillai &amp; Chanda" baseAuthorityYear="1973" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shillongensis">
<emphasis id="55C06E531D345FCD43771AFA133F452B" italics="true">R. shillongensis</emphasis>
</taxonomicName>
, with a low node support (51). From congeners in the Indo-China region, it can be easily distinguished by: having an externally visible tympanum (absent in
<taxonomicName id="A75DD22BADAC7F13EE831EABEF7C1B1C" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="DEE41BE56763F538D5BA3193025F730A" italics="true">R. kempiae</emphasis>
</taxonomicName>
,
<taxonomicName id="E9EB3AB12113F66E813094D8071ACC69" baseAuthorityName="Pillai &amp; Chanda" baseAuthorityYear="1973" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="shillongensis">
<emphasis id="EAB19ECA490DFEA00089A6EA4E214EB6" italics="true">R. shillongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="3C2D1FC4D77E6D43FA2146C5D5D59063" baseAuthorityName="Kou" baseAuthorityYear="1990" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="E8E880F17912FD83B33950E4050FCE16" italics="true">R. menglaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A4C54E1122C6A78C3B335C88122352EE" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="E3BB5F7194D7FD91505F511CC44772DF" italics="true">R. rezakhani</emphasis>
</taxonomicName>
); ventral aspect of the hind limb dark orange in color (vs. uniform creamy white in
<taxonomicName id="10B8537378297637FD0E420F59A68D1C" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="847FBA840BD112AC8609E6FFC67AC61F" italics="true">R. kempiae</emphasis>
</taxonomicName>
and dark gray in
<taxonomicName id="DF89EE257C75DDCFA9F59FD74C8B0A32" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="301AA4632D588EDE390FF67988F773E9" italics="true">R. rezakhani</emphasis>
</taxonomicName>
); toe webbing I 1-1 II 1 - 1 III 3 - 3 IV 3 - 2.5 V in
<taxonomicName id="267968AC35322C54D942D516FB98E6F6" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="96C438EEBF951E3321F01CEA8E4D60F0" italics="true">R. garo</emphasis>
</taxonomicName>
(vs. I 1-2 II 2 - 1 III 1 - 1 IV 1 - 2 V in
<taxonomicName id="165DDFE25644FC70A5A37C155232EA50" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="6E0F6C6B463FE08406E804064D359694" italics="true">R. kempiae</emphasis>
</taxonomicName>
, I 22 II 22 III 23 IV 32 V in
<taxonomicName id="3ECD7FC0FC0A5271B409A29C1F2EEDC7" authority=", II" authorityName="Huang, Liu, Du, Bernstein, Liu, Yang, Yu &amp; Wu" authorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="malipoensis">
<emphasis id="DDD3B21C8748E1AF1649A51FD8C50D02" italics="true">R. malipoensis</emphasis>
, II
</taxonomicName>
12 III 12.5 IV 2.51 V in
<taxonomicName id="1AD51D85129BCFDEBB40C869CE4BED05" authority=", II" authorityName="II" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="9FAF0FEB3CA324433498CB505B2ED55E" italics="true">R. menglaensis</emphasis>
, II
</taxonomicName>
12 III 12.5 IV 2.51 V in
<taxonomicName id="6F83B62D2A5454783C7797E9CB067ED8" authority=", II" authorityName="II" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hillisi">
<emphasis id="1167F40EA11923719176D57F807EA66A" italics="true">R. hillisi</emphasis>
, II
</taxonomicName>
12 III 12 IV 21 V in
<taxonomicName id="20F57AAE4B136268644E04A28C578C4D" authorityName="Jiang, Wang, Ren &amp; Li" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="huanglianshan">
<emphasis id="6370AC5BF42A4D77B19FBC53D87A2B9B" italics="true">R. huanglianshan</emphasis>
</taxonomicName>
and I 22 II 1.752 III 1.53 IV 2.752 V in
<taxonomicName id="E52175CAFAE03C541EBF8C983138DEF7" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="C1F10A32935ABBDCF989BF2DBD3F46BA" italics="true">R. rezakhani</emphasis>
</taxonomicName>
); snout short and truncate in
<taxonomicName id="FC8448462B082374C7A743D2997361E8" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="263D0CD541061D1AF62128604D5DCE23" italics="true">R. garo</emphasis>
</taxonomicName>
(vs. rounded in
<taxonomicName id="EE942D6BFE49C7D1C9DFE50E6BC15C5A" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="C765C29117D3309FD0EC3AEF7A8318FF" italics="true">R. kempiae</emphasis>
</taxonomicName>
,
<taxonomicName id="852482587DF60320C04B1A2FAD335FEC" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="EBE8A70CE531740514DF55870D633E2A" italics="true">R. menglaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="62860058D2F962C5AF8D10CC983EB14E" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hillisi">
<emphasis id="6C3E7E6F42C6ACBB8CE50907FE54284B" italics="true">R. hillisi</emphasis>
</taxonomicName>
,
<taxonomicName id="9A14D2D429D69079AAB133FDAF7C7663" authorityName="Du, Xu, Liu &amp; Yu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hekouensis">
<emphasis id="C6A17AF17957F54B95B657BA7D059364" italics="true">R. hekouensis</emphasis>
</taxonomicName>
,
<taxonomicName id="0A4202067B858CDAE86EBD6B4E8253EE" authorityName="Huang, Liu, Du, Bernstein, Liu, Yang, Yu &amp; Wu" authorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="malipoensis">
<emphasis id="2953F0C238106900247B4625E75B5FF2" italics="true">R. malipoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6B30675ABD704A3BA4F93194B2D778A6" baseAuthorityName="Yang &amp; Li" baseAuthorityYear="1978" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longchuanensis">
<emphasis id="9F4BD5EA70E198D734348E8F247605A6" italics="true">R. longchuanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B5BC0C28EC4E9CF55892CAAE91A4FB0C" authorityName="Jiang, Wang, Ren &amp; Li" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="huanglianshan">
<emphasis id="F43B96961316F39EAF9E555665912957" italics="true">R. huanglianshan</emphasis>
</taxonomicName>
and sharply projecting in
<taxonomicName id="2989AA7CF47E3091BB4EB0C62AC145DD" baseAuthorityName="Boulenger" baseAuthorityYear="1906" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="annandalii">
<emphasis id="84022CE7986C831A00236B832DA2FBCA" italics="true">R. annandalii</emphasis>
</taxonomicName>
and
<taxonomicName id="9EC97D39200EE0818B259C802ECE8639" authorityName="Zhang, Shu, Liu, Dong &amp; Guo" authorityYear="2022" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yadongensis">
<emphasis id="C4A7D9949BD3D4615AEB5107DE4C511F" italics="true">R. yadongensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6JAI" type="distribution">
<paragraph id="309AD96BDF998B70974A855F971CB204">
<heading id="08543B9BE68E6FB29A15FC56FBD6496C" reason="title">Natural history and distribution.</heading>
</paragraph>
<paragraph id="F6615D504036714B42E471B4B1D37C1E">
These frogs can be heard calling at dusk. Males were seen to be actively vocalizing from April to September. Most individuals recorded during this study were observed to be calling from shrubs (usually higher&gt;
<quantity id="6A1478CA0E1B3AD8BDC32F1CC150F39D" metricMagnitude="0" metricUnit="m" metricValue="2.5" unit="m" value="2.5">2.5 m</quantity>
above the ground). The species was infrequently encountered and was only recorded from areas above
<quantity id="3D7418F72216ED716797E5840BDFA915" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
asl during this study. This species was recorded from three different localities in and around the West Garo Hills region: Sakalgre (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIKAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.380000,25.510000]}">
<geoCoordinate id="C0CE51800CD8D83D2049A8A15E33179C" degrees="25.51" direction="north" orientation="latitude" precision="555" value="25.51">25.51 ° N</geoCoordinate>
,
<geoCoordinate id="9E576BB8F259B7FFDF8A383824ED304C" degrees="90.38" direction="east" orientation="longitude" precision="555" value="90.38">90.38 ° E</geoCoordinate>
</named-content>
;
<quantity id="A1A8AA9F1C0155BD1F3858C2A1FA65A7" metricMagnitude="2" metricUnit="m" metricValue="8.95" unit="m" value="895.0">895 m</quantity>
); Daribokgre (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQKAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.310000,25.470000]}">
<geoCoordinate id="C2080BA2351D217A93AF8E8FE0338443" degrees="25.47" direction="north" orientation="latitude" precision="555" value="25.47">25.47 ° N</geoCoordinate>
,
<geoCoordinate id="94FB47749641E9156BE56B25D68869F9" degrees="90.31" direction="east" orientation="longitude" precision="555" value="90.31">90.31 ° E</geoCoordinate>
</named-content>
;
<quantity id="9DEC3A7BBEBCD6084B1EC2E1EAD5798A" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
); and Mandalgre (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYKAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.410000,25.480000]}">
<geoCoordinate id="48A4E604B7E662BCB9C4BDB41C6156B5" degrees="25.48" direction="north" orientation="latitude" precision="555" value="25.48">25.48 ° N</geoCoordinate>
,
<geoCoordinate id="D20BB474150BC5ECB8F3BDE7746AC450" degrees="90.41" direction="east" orientation="longitude" precision="555" value="90.41">90.41 ° E</geoCoordinate>
</named-content>
;
<quantity id="25341DCA431DF6CD09F3514BA7FDF41B" metricMagnitude="3" metricUnit="m" metricValue="1.081" unit="m" value="1081.0">1081 m</quantity>
). Based on the locations from the study, we calculated the extent of occurrence of this species with the current data from this study, and it was estimated to be
<quantity id="0722363F766D691ED8D7C1E0DE17CAAF" metricMagnitude="4" metricUnit="m" metricValue="1.146" unit="km" value="11.46">11.46 km</quantity>
<superScript id="1FFE595F435ADAB93F45392777B15FD5">2</superScript>
. However, further surveys are required to robustly estimate the distribution range of this species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6KAI" type="remarks">
<paragraph id="6AA9C0EE3D8C8F8966CCBC81D9CC390B">
<heading id="0D76637B63DB13B1FB20120164517106" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="BC188A7AFDD4AF8984A19C953EE07357">
<bibRefCitation id="83C0385F5C69D1D9BF2E225AAF8F909E" DOI="10.26515/rzsi/v16/i2/1919/162810" author="Boulenger GA" firstAuthor="Boulenger" issue="2" journalOrPublisher="Records of the Zoological Survey of India" pagination="207-208" refId="B7" refString="Boulenger GA (1919) XII Descriptions of Three new Batrachians from the Garo Hills, Assam. Records of the Zoological Survey of India 16 (2): 207208. https://doi.org/10.26515/rzsi/v16/i2/1919/162810" title="XII Descriptions of Three new Batrachians from the Garo Hills, Assam." volume="16" year="1919">Boulenger (1919)</bibRefCitation>
described this species from the Garo Hills along with
<taxonomicName id="C080ACD74D8B891463DD6CE827F9F063" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Bufonidae" genus="Bufoides" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempi">
<emphasis id="77C6D519FA830ABC6282CFF84192E34D" italics="true">Bufoides kempi</emphasis>
</taxonomicName>
and
<taxonomicName id="6744D0958B856277DDF1709DA5EB0609" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="F9AB0AB9449E2E972FC2E2E07E0C22F4" italics="true">R. kempiae</emphasis>
</taxonomicName>
. Our topotypical specimens are in agreement with the
<typeStatus id="D155C9614A3A051BDD6C5BC2E574D1E3" type="holotype">holotype</typeStatus>
and with Boulengers description in most characters.
<bibRefCitation id="08E2434C7ACA69807FD26C8D700ABC9D" author="Kharkongor IJ &amp; Saikia B &amp; Deb R" etAl="et al." firstAuthor="Kharkongor" issue="4" journalOrPublisher="Records of the Zoological Survey of India" pagination="439-444" refId="B20" refString="Kharkongor IJ, Saikia B, Deb R (2016) Detailed study on Philautus garo (Boulenger, 1919) (Amphibia, Anura, Rhacophoridae) - An extremely rare and endemic species of North-East India. Records of the Zoological Survey of India 116 (4): 439444." title="Detailed study on Philautus garo (Boulenger, 1919) (Amphibia, Anura, Rhacophoridae) - An extremely rare and endemic species of North-East India." volume="116" year="2016">Kharkongor et al. (2016)</bibRefCitation>
reported a population of
<taxonomicName id="65809F62892F9EFC8B150406BDE1FA29" authorityName="Biju, Shouche, Dubois, Dutta &amp; Bossuyt" authorityYear="2010" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="358FF639D8E27D55AA0B6127F7A8C9C7" italics="true">Raorchestes</emphasis>
</taxonomicName>
from Upper Shillong with detailed morphological description as
<emphasis id="9822AD4FA6591C376A9D37360042279D" italics="true">
<taxonomicName id="47FF4B1A89BA52C37E08BF90E1ECF025" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">Philautus garo</taxonomicName>
.
</emphasis>
Although their descriptions of these specimens closely match the description of
<taxonomicName id="F349B5953628D1A8F36A46E1999E1650" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="8175AE0A8F3DC7F8C61835B067EC7FC6" italics="true">R. garo</emphasis>
</taxonomicName>
, since these are not topotypical populations, i. e., the collection locations occur almost
<quantity id="92C8159D83CF8883734CB07421BAD8B9" metricMagnitude="5" metricUnit="m" metricValue="2.0" unit="km" value="200.0">200 km</quantity>
(aerial distance) from the type locality and from a different elevational range, further studies based on molecular analysis are required to see if this population is conspecific with
<emphasis id="704F6A566244AFAAB14DE6FEFC727BFC" italics="true">
<taxonomicName id="57C44D30C0B79202AFF10D5067EE0C2C" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">Raorchestes garo</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,763 @@
<document id="F08BB8E6C865E3CC96305D7C5264D6A4" ID-DOI="10.3897/herpetozoa.37.e122825" ID-ZooBank="11B82D36-D464-429B-9C3B-D22423B8FBC6" ID-publisher-id="122825" URI-arpha="DE04D54E-C773-5EF0-95A6-23528E54505A" URI-zoobank="http://zoobank.org/11B82D36-D464-429B-9C3B-D22423B8FBC6" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1731951019993" checkinUser="pensoft" docAuthor="Naveen, R. S., Chandramouli, S. R., Babu, S., Ryndongsngi, A. M., Karunakaran, P. V. &amp; Kumara, Honnavalli N." docDate="2024" docId="EFEA61F555B55FAEAD0B49110787CB93" docLanguage="en" docName="Herpetozoa 37: 359-372" docOrigin="Herpetozoa 37" docSource="https://herpetozoa.pensoft.net/article/122825/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Raorchestes kempiae Naveen, Chandramouli &amp; Babu 2024, comb. nov." docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="372" masterDocId="DE04D54EC7735EF095A623528E54505A" masterDocTitle="Rediscovery and redescription of Ixalus garo Boulenger, 1919, and Ixalus kempiae Boulenger, 1919, with a reassessment of the taxonomic status of Raorchestes cangyuanensis Wu, Suwannapoom, Xu, Murphy &amp; Che, 2019 and the description of a new species from the Garo Hills of Meghalaya" masterLastPageNumber="372" masterPageNumber="359" pageNumber="359" updateTime="1731956028832" updateUser="admin">
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<mods:title id="060751290ECAD5EF3A22536F82631009">Rediscovery and redescription of Ixalus garo Boulenger, 1919, and Ixalus kempiae Boulenger, 1919, with a reassessment of the taxonomic status of Raorchestes cangyuanensis Wu, Suwannapoom, Xu, Murphy &amp; Che, 2019 and the description of a new species from the Garo Hills of Meghalaya</mods:title>
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<mods:affiliation id="F1E5622B8F23E25C050772CD4E5BE095">Department of Ecology and Environmental Sciences, School of Life Sciences, Pondicherry University, Puducherry, India</mods:affiliation>
</mods:name>
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<mods:identifier id="621F67DD294840AE6922F6B2CB407835" type="DOI">10.3897/herpetozoa.37.e122825</mods:identifier>
<mods:identifier id="00337F22DB93A0A98F7AB7172B0CF7DA" type="ZooBank">11B82D36-D464-429B-9C3B-D22423B8FBC6</mods:identifier>
</mods:mods>
<treatment id="EFEA61F555B55FAEAD0B49110787CB93" ID-arpha="EFEA61F5-55B5-5FAE-AD0B-49110787CB93" LSID="urn:lsid:plazi:treatment:EFEA61F555B55FAEAD0B49110787CB93" httpUri="http://treatment.plazi.org/id/EFEA61F555B55FAEAD0B49110787CB93">
<subSubSection id="37B85140643E9DF3FD15C0F2A4F653A8" type="nomenclature">
<paragraph id="9550CFAF2EA2A0E4DEB3A63386F6246F">
<taxonomicName id="05CC27183B5A6EEC08D440DDD67139B1" authority="(Boulenger, 1919)" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae" status="comb. nov.">
<emphasis id="4EEB68DB9499171B411A09275B7DF556" italics="true">Raorchestes kempiae</emphasis>
(
<bibRefCitation id="F607D71E4E096751AB2451AE2C72D2EC" DOI="10.26515/rzsi/v16/i2/1919/162810" author="Boulenger GA" issue="2" journalOrPublisher="Records of the Zoological Survey of India" pagination="207-208" refId="B7" refString="Boulenger GA (1919) XII Descriptions of Three new Batrachians from the Garo Hills, Assam. Records of the Zoological Survey of India 16 (2): 207 - 208. https: // doi. org / 10.26515 / rzsi / v 16 / i 2 / 1919 / 162810" title="XII Descriptions of Three new Batrachians from the Garo Hills, Assam." volume="16" year="1919">Boulenger, 1919</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="9D08268114486A6A1F4C02C621DDEE0F" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5B5A2CD3D225842DCBAC687C37949BF6" type="reference_group">
<paragraph id="FBEADA2A80B151990B41C71B02E8B494">
<treatmentCitationGroup id="8E2178374764FEC7863C26834D787B56">
<treatmentCitation id="89C2234F51A38043E517D870E0D91E1E">
<taxonomicName id="F032EB9DDAF6BDE7003ED6B7CB4C84CB" ID-CoL="3QFCB" authority="Boulenger, 1919" authorityName=", Boulenger" authorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Ixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="E9540D39173C5D643B70577F3A2A8B56" italics="true">Ixalus kempiae</emphasis>
<bibRefCitation id="E58DCCFF3323D8DC37F3F0B244927CBF" DOI="10.26515/rzsi/v16/i2/1919/162810" author="Boulenger GA" issue="2" journalOrPublisher="Records of the Zoological Survey of India" pagination="207-208" refId="B7" refString="Boulenger GA (1919) XII Descriptions of Three new Batrachians from the Garo Hills, Assam. Records of the Zoological Survey of India 16 (2): 207 - 208. https: // doi. org / 10.26515 / rzsi / v 16 / i 2 / 1919 / 162810" title="XII Descriptions of Three new Batrachians from the Garo Hills, Assam." volume="16" year="1919">Boulenger, 1919</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="BE3A60AFCB2271A4A64B69C6CF365E87">
<treatmentCitationGroup id="916AC8A439D6B4EF47E7C98BDFC9C6E1">
<treatmentCitation id="4B505C61650FE79FB6283A37100EBD51">
<taxonomicName id="BACCD111DA880D2D1E543B1E3BBBDE4E" ID-CoL="4RXG2" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Rhacophorus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae" subGenus="Philautus">
<emphasis id="A8F5399D1503840148BE168FC15AF074" italics="true">Rhacophorus</emphasis>
(
<emphasis id="E2792DAF0EFB6D3E9A618CC8079275FD" italics="true">Philautus</emphasis>
)
<emphasis id="2A52969AA569773369DFC0CEA549690C" italics="true">kempiae</emphasis>
</taxonomicName>
<bibRefCitation id="0ADB97DF15935DD0AF94F2D41193F5F2" author="Ahl E" refId="B1" refString="Ahl E (1931) Amphibia, Anura III, Polypedatidae. Das Tierreich 55: xvi + 477." year="1931">Ahl, 1931</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="C083E49A25C2DF6D67AE8285EAEA26AB">
<treatmentCitationGroup id="ADC3CFEC1E3F2A2D25848C6BCA664557">
<treatmentCitation id="55B3E2497AED32A9BCF7CA9A7BBD3D33">
<taxonomicName id="ED68007F23FA52EA6926DF3644FF9B85" ID-CoL="4G5H8" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="0AAC1071698495DD28E1766ED582AF50" italics="true">Philautus kempiae</emphasis>
</taxonomicName>
<bibRefCitation id="08B029022AF8CCB2A2119ED51F3BF3B2" author="Bourret R" firstAuthor="Bourret" journalOrPublisher="Hanoi: Institut Océanographique de lIndochine" pagination="1-547" refId="B8" refString="Bourret R (1942) Les Batraciens de lIndochine. Hanoi: Institut Océanographique de lIndochine 6: 1547." title="Les Batraciens de lIndochine." volume="6" year="1942">Bourret (1942)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC740A3D0205313B38EF67D37C009CB6">
<treatmentCitationGroup id="378A4522CBF1FB2208A0F60EFA0FAE3A">
<treatmentCitation id="B94ADBAABC9581A1202E322CEE0B5334">
<taxonomicName id="C51E7E5A7060BD0838AEFBD2EF157721" authorityName="Wu, Suwannapoom, Xu, Murphy &amp; Che" authorityYear="2019" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cangyuanensis">
<emphasis id="28CC04B5793B09889F0126756FF864DB" italics="true">Raorchestes cangyuanensis</emphasis>
</taxonomicName>
<bibRefCitation id="4CFF5792E5452A20E1BB235029302E42" DOI="10.24272/j.issn.2095-8137.2019.066" author="Wu YH &amp; Suwannapoom C &amp; Xu K &amp; Chen JM &amp; Jin JQ &amp; Chen HM &amp; Murphy RW &amp; Che J" issue="6" journalOrPublisher="Zoological Research" pagination="558-563" refId="B33" refString="Wu YH, Suwannapoom C, Xu K, Chen JM, Jin JQ, Chen HM, Murphy RW, Che J (2019) A new species of the genus Raorchestes (Anura, Rhacophoridae) from Yunnan Province, China. Zoological Research 40 (6): 558 - 563. https: // doi. org / 10.24272 / j. issn. 2095 - 8137.2019. 066" title="A new species of the genus Raorchestes (Anura, Rhacophoridae) from Yunnan Province, China." volume="40" year="2019">Wu, Suwannapoom, Xu, Murphy &amp; Che, 2019</bibRefCitation>
</treatmentCitation>
,
<taxonomicNameLabel id="84A34F639286DF262B48BF4A839C2F87">syn. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIQAI" type="material">
<paragraph id="8ADA75B619877AA2A9D720054AA18245">
<heading id="BEF5DD4FCFD3F4D350066CCA58BC767A" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="B81A259A194C6B39BC95DDA768782782">
<materialsCitation id="2C9EEA17A45DBAB2CC69A136D7736838" collectionCode="ZSI" collectorName="Agnes Kemp" location="Above Tura" specimenCode="ZSI 18859" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
<emphasis id="BE55B4641F54EB96C4DE1E2ACA0A07A9" bold="true" italics="true">
<typeStatus id="602386F2BAEC50C8D3A2A9AADD145C16" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="1FCA576556A653BC5573C5F57CB8D58B">ZSI 18859</specimenCode>
(Fig.
<figureCitation id="5CC5479B27036A4A02EC8663E200A282" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype of Ixalus kempiae (ZSI 18859) - not to scale (above); SACON VA 806 in preservation (below). A. Dorsal view; and B. Ventral view. Scale bars: 10 mm." figureDoi="10.3897/herpetozoa.37.e122825.figure4" httpUri="https://binary.pensoft.net/fig/1177122">4</figureCitation>
above);
<specimenCount id="3C5AEF4B3317211721C3559D234FB9B1" count="1" type="adult">sub adult</specimenCount>
, collected by
<collectorName id="963C9032996B4D2732A2C688B6495C78">Agnes Kemp</collectorName>
, from “
<location id="02A7C0596B98D49F33C47E67B88E761C" LSID="urn:lsid:plazi:treatment:EFEA61F555B55FAEAD0B49110787CB93:02A7C0596B98D49F33C47E67B88E761C" name="Above Tura">Above Tura</location>
</materialsCitation>
.
</paragraph>
<paragraph id="72632BD6CB04871E8FEBE97488B7000E">
<materialsCitation id="75EF58A83600B746B1F7021856471EA5" collectingDate="2021-01-23" collectionCode="SACON" collectorName="RSN" country="India" county="South Garo Hills" elevation="174" latitude="25.43366" location="Mikadogre Community Reserve" longLatPrecision="1" longitude="90.39898" specimenCode="VA 806" specimenCount="5" specimenCount-male="5" stateProvince="Meghalaya" typeStatus="topotype">
<emphasis id="A0B8DAE94A8234467DB353647938208E" bold="true" italics="true">
<typeStatus id="0B7453250CCD7FF20BE091D0F26658E6" type="topotype">Topotypes</typeStatus>
</emphasis>
:
<specimenCount id="37CB889D8777E1EF484A1D41CD5EA1D2" count="5" type="male">Five adult males</specimenCount>
,
<collectionCode id="0A31CC89C52A2A38C1C2036D88715302" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
<specimenCode id="5242C446689E81A5E657F76301F5EB52">VA 806</specimenCode>
(Fig.
<figureCitation id="363E241C37284B7314D0898BFEA93850" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype of Ixalus kempiae (ZSI 18859) - not to scale (above); SACON VA 806 in preservation (below). A. Dorsal view; and B. Ventral view. Scale bars: 10 mm." figureDoi="10.3897/herpetozoa.37.e122825.figure4" httpUri="https://binary.pensoft.net/fig/1177122">4</figureCitation>
below), 301, 302, 297, 299 from
<location id="567CDC21854E6E1F3A74FA4E88C07A56" LSID="urn:lsid:plazi:treatment:EFEA61F555B55FAEAD0B49110787CB93:567CDC21854E6E1F3A74FA4E88C07A56" country="India" county="South Garo Hills" latitude="25.43366" longLatPrecision="1" longitude="90.39898" name="Mikadogre Community Reserve" stateProvince="Meghalaya">Mikadogre Community Reserve</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKRAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.398981,25.433660]}">
<geoCoordinate id="10F5D006CD0FBA6792A6EC10921E214E" degrees="25.433660" direction="north" orientation="latitude" precision="1" value="25.43366">25.433660 ° N</geoCoordinate>
,
<geoCoordinate id="23A49C04608B9AF26E702D0840DA290C" degrees="90.398981" direction="east" orientation="longitude" precision="1" value="90.39898">90.398981 ° E</geoCoordinate>
</named-content>
,
<quantity id="2E1D6A313383DBF3D91D5DA9B7325EAB" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">
<elevation id="B9DA142897D0C8EE3475A254C86DCC18" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">174 m</elevation>
</quantity>
),
<collectingCounty id="2A1346B6981A622923A397D14C12EB3B">South Garo Hills</collectingCounty>
,
<collectingRegion id="6B9FCA36B362B48195151778236FD3A4" country="India" name="Meghalaya">Meghalaya</collectingRegion>
,
<collectingCountry id="B4FA1EE741FC795A36E1AFE0768ACEFF" name="India">India</collectingCountry>
, collected by
<collectorName id="5EC20BF93D5A092061FE73F93159B472">RSN</collectorName>
on
<date id="5920327FF81BDE6F78F449F7EE366C87" value="2021-01-23">
<collectingDate id="FBC6DAC3CACD8C95E97EA4552B9BFD31" value="2021-01-23">
January 23
<superScript id="94C1B090B409254E79702BA751E21DFA">rd</superScript>
2021
</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="E77B617E1E5E775F4AAAD6E6EEE7296D" ID-DOI="10.3897/herpetozoa.37.e122825.figure4" ID-arpha="3CA7FBCE-641D-5A6B-9FC2-B2DCD9F642F1" httpUri="https://binary.pensoft.net/fig/1177122" startId="F4">
<paragraph id="40D43B0218F52B03EE27F02EC26BCE9D">
<label id="AF833270E5F0AC5EE54EB3B59E4507CE">Figure 4.</label>
</paragraph>
<paragraph id="E960850F1F5EB36110015B3A00C22001">
The holotype of
<taxonomicName id="9B6CDFC5B7661F44FE675033F606D997" authorityName=", Boulenger" authorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Ixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="ECD0536E838FC7B2A5ED12A5CB15AE7B" italics="true">Ixalus kempiae</emphasis>
</taxonomicName>
(ZSI 18859) - not to scale (above); SACON VA 806 in preservation (below).
<emphasis id="D4E581373B41A38743B94A95811CEC16" bold="true">A.</emphasis>
Dorsal view; and
<emphasis id="126F4F27480E147DBE5C8D362CBF754D" bold="true">B.</emphasis>
Ventral view. Scale bars: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ENSAI" type="diagnosis">
<paragraph id="5EEEFEC24D9EFD395EB218FB787349F5">
<heading id="F21BDF05A7926D3DAB5A67F0C0735C0B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A31BAA161D6A12415AA3F3CB361DDB7A">
<taxonomicName id="9170B0090322DE740D16C84C01DBD826" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae" status="comb. nov.">
<emphasis id="DE35074A213F8F78E699029F969F9A88" italics="true">Raorchestes kempiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2D366A5289E9A10629353880E1032595" rank="species">comb nov.</taxonomicNameLabel>
is placed in the genus
<taxonomicName id="E646F5370B932D708B70F69F1749763D" authorityName="Biju, Shouche, Dubois, Dutta &amp; Bossuyt" authorityYear="2010" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="23A71AAE3467B3FD13B297C0FA91F70A" italics="true">Raorchestes</emphasis>
</taxonomicName>
due to the combination of following characters: small body size, vomerine teeth absent, single translucent external subgular vocal sac present, and tips of all fingers and toes expanded into discs with circum-marginal grooves (see
<bibRefCitation id="E98FB1A0A67959748FBAEA3ABF8D988B" author="Biju" etAl="et al." firstAuthor="Biju" journalOrPublisher="Current Science, Bangalore" pagination="1119-1125" refId="B3" refString="Biju SD, Shouche YS, Dubois A, Dutta SK, Bossuy F (2010) A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India. Current Science, Bangalore 98: 11191125." title="A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India." volume="98" year="2010">Biju et al. 2010</bibRefCitation>
). The species can be distinguished from congeners in the Indo-China region by the following combination of characters: A small-sized (mean
<abbrev id="ABBRID0EJTAI" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="D70364C66BD8AB95831AC4011B1C469A" metricMagnitude="-2" metricUnit="m" metricValue="2.41" unit="mm" value="24.1">24.1 mm</quantity>
in adult males) arboreal species with a rounded snout; tympanum indistinct and supratympanic fold prominent; vomerine teeth absent; single semi-transparent gular vocal sac; tongue without papilla; ventral aspect of the hind limb and groin yellow with black or brown spots; finger and toe discs well developed; toe webbing I 1-2 II 2 - 1 III 1 - 1 IV 1 - 2 V.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENTAI" type="description">
<paragraph id="300FB4D6EC658F154699C37141C599BC">
<heading id="861BE6DD343167A4E9D7513660A3AB28" reason="title">Description</heading>
</paragraph>
<paragraph id="E8F331B2A748C910546AABA2D00F3012">
(based on an adult male
<collectionCode id="07E8779CB3FF4F0EAF73EC8CE6725479" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" type="">SACON</collectionCode>
VA 806). Small-sized frog (
<abbrev id="ABBRID0ETTAI" xlink_title="snoutvent length">SVL</abbrev>
<quantity id="CDE3DDE42039EEE90FD4E51935CA65A6" metricMagnitude="-2" metricUnit="m" metricValue="2.38" unit="mm" value="23.8">23.8 mm</quantity>
), with robust body. Dorsal skin mostly smooth with a few scattered spinules, ventral skin slightly granulose. Head broader than long (
<abbrev id="ABBRID0EXTAI" xlink_title="head length">HL</abbrev>
:
<abbrev id="ABBRID0E2TAI" xlink_title="head width">HW</abbrev>
0.83); large (
<abbrev id="ABBRID0E6TAI" xlink_title="head length">HL</abbrev>
:
<abbrev id="ABBRID0EDUAI" xlink_title="snoutvent length">SVL</abbrev>
0.33); with a short-rounded snout (
<abbrev id="ABBRID0EHUAI" xlink_title="eye diameter">ED</abbrev>
:
<abbrev id="ABBRID0ELUAI" xlink_title="eyesnout distance">ES</abbrev>
0.72). Canthus rostralis evident and rounded; loreal region concave. Trunk short (AG:
<abbrev id="ABBRID0EPUAI" xlink_title="snoutvent length">SVL</abbrev>
0.41) and stout (AG:
<abbrev id="ABBRID0ETUAI" xlink_title="body width">BW</abbrev>
1.02); eyes large (
<abbrev id="ABBRID0EXUAI" xlink_title="eye diameter">ED</abbrev>
:
<abbrev id="ABBRID0E2UAI" xlink_title="head length">HL</abbrev>
0.38); nostrils, ovoid, with no visible rim, situated closer to the snout tip than to the eyes (
<abbrev id="ABBRID0E6UAI" xlink_title="eyenostril distance">EN</abbrev>
:
<abbrev id="ABBRID0EDVAI" xlink_title="eyesnout distance">ES</abbrev>
0.61), directed laterally. Inter-orbital space broader than inter-narial space (
<abbrev id="ABBRID0EHVAI" xlink_title="interorbital distance">IO</abbrev>
:
<abbrev id="ABBRID0ELVAI" xlink_title="internarial distance">IN</abbrev>
0.69). Tympanum indistinct. Upper arms short (
<abbrev id="ABBRID0EPVAI" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0ETVAI" xlink_title="snoutvent length">SVL</abbrev>
0.28), shorter than lower arms (
<abbrev id="ABBRID0EXVAI" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0E2VAI" xlink_title="lower arm length">LAL</abbrev>
0.57); palm slightly shorter than the upper arms (
<abbrev id="ABBRID0E6VAI" xlink_title="upper arm length">UAL</abbrev>
:
<abbrev id="ABBRID0EDWAI" xlink_title="palm length">PAL</abbrev>
0.54); relative finger lengths III&gt; IV&gt; II&gt; I; Subarticular tubercles distinct in both palmar and plantar surfaces, rounded; Supernumerary tubercles present in the palm; no webbing between fingers; fingers with well-developed rounded discs. Thighs short, nearly half as long as the body (
<abbrev id="ABBRID0EHWAI" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0ELWAI" xlink_title="snoutvent length">SVL</abbrev>
0.32), slightly shorter than the tibia (
<abbrev id="ABBRID0EPWAI" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0ETWAI" xlink_title="tibia length">TBL</abbrev>
0.83). Foot shorter than the thigh (
<abbrev id="ABBRID0EXWAI" xlink_title="femur length">FEL</abbrev>
:
<abbrev id="ABBRID0E2WAI" xlink_title="foot length">FOL</abbrev>
2.83), toes short with well-developed rounded discs, partial webbing, webbing formula I 1-2 II 2 - 1 III 1 - 1 IV 1 - 2 V. Dorsum uniformly brown with a faint hourglass pattern. Loreals pale brown, iris dark golden brown. Venter granular, pale gray, spotted with gray and brown in lower trunk and thighs. Lateral aspect of the hind limb and groin orangish yellow with black and brown blotches (Fig.
<figureCitation id="A7D90EF68A3DBDD5C92757957ADA82A5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Raorchestes kempiae (dorso-lateral view of SACON VA 806 on top and dorso-lateral view of an uncollected male, bottom) from Garo Hills in life." figureDoi="10.3897/herpetozoa.37.e122825.figure5" httpUri="https://binary.pensoft.net/fig/1177123">5</figureCitation>
).
</paragraph>
<caption id="1A3E8588D9C88DF87276EBFD5650ED36" ID-DOI="10.3897/herpetozoa.37.e122825.figure5" ID-arpha="504E2828-EA0E-5834-819A-5BF6CEB1D969" httpUri="https://binary.pensoft.net/fig/1177123" startId="F5">
<paragraph id="BF6A1195816EE97605AC8673165A4295">
<label id="941256C08B7042FF41A5BEDD397D3BE1">Figure 5.</label>
</paragraph>
<paragraph id="18E5626BC3C8DF1ABCCFAB1430E2878B">
<taxonomicName id="64FA49567EC5B6FDB99EAA01077F20D3" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="5A967559F6819EA44C5F4F155DDA53DD" italics="true">Raorchestes kempiae</emphasis>
</taxonomicName>
(dorso-lateral view of SACON VA 806 on top and dorso-lateral view of an uncollected male, bottom) from Garo Hills in life.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E2XAI" type="variation">
<paragraph id="3AF41985E76FF29B2D467C3B331A9C16">
<heading id="8BF1855D82DCB9F306658B33C4EC31F9" reason="title">Variation.</heading>
</paragraph>
<paragraph id="5231DA5704205B4F5125DC57E4B1A1F0">
Morphometric measurements of the
<typeStatus id="534954F75D5F1F7063ACA974CF1D397E" type="holotype">holotypes</typeStatus>
and other preserved
<typeStatus id="ACDF8BA0230FB95773F82AC8CAC5A047" type="topotype">topotypes</typeStatus>
examined are presented in Table
<tableCitation id="56939FDC7F09070295C0ED7B7622833D" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric measurements (in mm) of R. kempiae." httpUri="http://table.plazi.org/id/2CBC684D6BFAAC65AB8CB42DF48FE3FC" tableUuid="2CBC684D6BFAAC65AB8CB42DF48FE3FC">3</tableCitation>
. Variations noted during examination of preserved specimens and live individuals observed in the field. Dorsal coloration ranging from light brown to brownish black with or without an “) (“ pattern (present in VA 301 and VA 302). Brown or black stripes over the thighs and tibia. Some individuals have a bright yellow to white band across the interorbital region (Fig.
<figureCitation id="D6CBABC1DC92361137FE2D487A0B483E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Raorchestes kempiae (dorso-lateral view of SACON VA 806 on top and dorso-lateral view of an uncollected male, bottom) from Garo Hills in life." figureDoi="10.3897/herpetozoa.37.e122825.figure5" httpUri="https://binary.pensoft.net/fig/1177123">5</figureCitation>
).
</paragraph>
<caption id="2CBC684D6BFAAC65AB8CB42DF48FE3FC" ID-Table-UUID="2CBC684D6BFAAC65AB8CB42DF48FE3FC" httpUri="http://table.plazi.org/id/2CBC684D6BFAAC65AB8CB42DF48FE3FC" startId="T3" targetIsTable="true">
<paragraph id="8096CE863A1807EC162F63718E5CE07A">
<label id="D199490C80FF1A4A37FB911557AA59A9">Table 3.</label>
</paragraph>
<paragraph id="4E4FC7D189DAB90D13CE7458DE229A85">
Morphometric measurements (in mm) of
<taxonomicName id="E570C2D649D9CE76663D5DDCCDE5BD1E" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="FD0AA871D1C9C7BBCBAC3239D2513EAF" italics="true">R. kempiae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="17464A989609B9D26C3BEADDC9CEF057">
<table id="TID0EZGCI" rules="all">
<tbody id="942209B0B14A0F44956248C0E4F8CEEE">
<tr id="F7A0622D76CABA14792A12BE305E1742">
<th id="46C60157FD517922EE832D6D73A16EAB" colspan="1" rowspan="1">Voucher no:</th>
<th id="BA7FC09C7238281F757892C78D191A23" colspan="1" rowspan="1">Sex</th>
<th id="36DBFE450BDF1801831358A4A32707C6" colspan="1" rowspan="1">
<abbrev id="ABBRID0EMZAI" xlink_title="snoutvent length">SVL</abbrev>
</th>
<th id="EB878F701203FAD11DB2135603714F0C" colspan="1" rowspan="1">AG</th>
<th id="54B43460AF43B20F4775D923B402B91A" colspan="1" rowspan="1">
<abbrev id="ABBRID0EXZAI" xlink_title="body width">BW</abbrev>
</th>
<th id="4C844AB9F909F23CF72904809B4B00AC" colspan="1" rowspan="1">
<abbrev id="ABBRID0E6ZAI" xlink_title="head length">HL</abbrev>
</th>
<th id="007243F504AFAC9C688CAB27CA1D1027" colspan="1" rowspan="1">
<abbrev id="ABBRID0EH1AI" xlink_title="head width">HW</abbrev>
</th>
<th id="F6D175CC00A5F352D7BEA6F43558B1BB" colspan="1" rowspan="1">
<abbrev id="ABBRID0EP1AI" xlink_title="head depth">HD</abbrev>
</th>
<th id="CB301B402E49B4BC8A388D6C1B009FD4" colspan="1" rowspan="1">
<abbrev id="ABBRID0EX1AI" xlink_title="eye diameter">ED</abbrev>
</th>
<th id="77D98D64B3DA3B1CF9E609B191474C6F" colspan="1" rowspan="1">
<abbrev id="ABBRID0E61AI" xlink_title="eyenostril distance">EN</abbrev>
</th>
<th id="30BCC783A8683B00B33EDDED442F13E3" colspan="1" rowspan="1">
<abbrev id="ABBRID0EH2AI" xlink_title="eyesnout distance">ES</abbrev>
</th>
<th id="6F887069AB8F872741D2D0CC1E9992C6" colspan="1" rowspan="1">
<abbrev id="ABBRID0EP2AI" xlink_title="tympanum diameter">TYD</abbrev>
</th>
<th id="B84CEB8878ACE98BB6303D8C62513758" colspan="1" rowspan="1">ET</th>
<th id="D1C8430FB034789E7D56062CAE3B8F84" colspan="1" rowspan="1">
<abbrev id="ABBRID0E12AI" xlink_title="upper eyelid width">UEW</abbrev>
</th>
<th id="8FF3C0DEF4A7F3C29C0ACE4254DE5945" colspan="1" rowspan="1">
<abbrev id="ABBRID0EC3AI" xlink_title="interorbital distance">IO</abbrev>
</th>
<th id="4613108C7ABE7CD01B09735771E51CC2" colspan="1" rowspan="1">
<abbrev id="ABBRID0EK3AI" xlink_title="internarial distance">IN</abbrev>
</th>
<th id="588B8C4883F7BAD9F92E1CB270A63A44" colspan="1" rowspan="1">
<abbrev id="ABBRID0ES3AI" xlink_title="upper arm length">UAL</abbrev>
</th>
<th id="5F036F11CBED6D0EB474248C67286981" colspan="1" rowspan="1">
<abbrev id="ABBRID0E13AI" xlink_title="lower arm length">LAL</abbrev>
</th>
<th id="F191BB0AE6B2074C1EB90D415AD6BEBC" colspan="1" rowspan="1">
<abbrev id="ABBRID0EC4AI" xlink_title="palm length">PAL</abbrev>
</th>
<th id="F77F584688D921FD66FCA2E4EAB9195A" colspan="1" rowspan="1">
<abbrev id="ABBRID0EK4AI" xlink_title="femur length">FEL</abbrev>
</th>
<th id="63AB3A9164D66269DE5F23869DB27D60" colspan="1" rowspan="1">
<abbrev id="ABBRID0ES4AI" xlink_title="tibia length">TBL</abbrev>
</th>
<th id="15261C5B16A61977AE5C1A351011DDC2" colspan="1" rowspan="1">TRL</th>
<th id="E5E730E0B46B77506DEF8A7A5236162B" colspan="1" rowspan="1">
<abbrev id="ABBRID0E44AI" xlink_title="foot length">FOL</abbrev>
</th>
</tr>
<tr id="363F6C56BADA04ED5013380212C819D1">
<td id="C284131BB24EB4166FF1658786751DEA" colspan="1" rowspan="1">Holotype (ZSI 18859)</td>
<td id="027AF5ECDB85FD20C3806885D072605B" colspan="1" rowspan="1">NA</td>
<td id="185E49F85F534DFD416F07CF45780A39" colspan="1" rowspan="1">16.3</td>
<td id="1CBEA45272B6F80131D58F372299BFE9" colspan="1" rowspan="1">6.02</td>
<td id="697F418BC55C7580EED14B305B1AF68C" colspan="1" rowspan="1">6.14</td>
<td id="476DC5E374DD34A8457AD11F646EB426" colspan="1" rowspan="1">5.22</td>
<td id="B46EF5D1D93750B8067D9C9580B8B8A0" colspan="1" rowspan="1">5.76</td>
<td id="BAE63A7287C762768BFB59D5577F3CA7" colspan="1" rowspan="1">3.14</td>
<td id="EE6B181C14E60984F2D560ECB8773A53" colspan="1" rowspan="1">2.44</td>
<td id="4082E1E49CC62BD715DFE863D56C8626" colspan="1" rowspan="1">1.7</td>
<td id="2FF307AC001EA99EEE266170E19F8B1D" colspan="1" rowspan="1">2.04</td>
<td id="0093C4B7C08F4D1E77C039A7F3E9A698" colspan="1" rowspan="1">0</td>
<td id="356C844E1AF7607B1D7954DC1EF8AA78" colspan="1" rowspan="1">0</td>
<td id="56BD54C8A24169298B541F6CFFE42C3D" colspan="1" rowspan="1">1.4</td>
<td id="A37D96C886DB48AA493032B62209F370" colspan="1" rowspan="1">1.7</td>
<td id="81D34C45279D9801CB4176044C26DB1C" colspan="1" rowspan="1">1.4</td>
<td id="5A44C62D4891C973E97BCEBFF6551738" colspan="1" rowspan="1">2.42</td>
<td id="DB2700D7CD6FC3C16F9984316914BED0" colspan="1" rowspan="1">2.82</td>
<td id="2CD9B6DA6DA82384F6C05FCF557EE171" colspan="1" rowspan="1">2.46</td>
<td id="983B26830A53675D72B9404C0D7B3B49" colspan="1" rowspan="1">6.7</td>
<td id="60F048A553FADE28A310D51A9D1C2954" colspan="1" rowspan="1">7.92</td>
<td id="1EBBC26BD115E74C011D2C1A3975F82E" colspan="1" rowspan="1">5</td>
<td id="0AF42BCD065A08C47F6E49C03A0D6D83" colspan="1" rowspan="1">U / N</td>
</tr>
<tr id="FC03A0DF3AFEB33504ECC0FE8ACBE3F4">
<td id="F8B0D2E204371AC44DE2DC05920A936F" colspan="1" rowspan="1">SACON VA- 806</td>
<td id="32B2FBDE37BD40721B12CD6AC0BC50FE" colspan="1" rowspan="1">Male</td>
<td id="EBD4704D2E8053C78C981A53B0D87035" colspan="1" rowspan="1">23.8</td>
<td id="850C8DAA3EE8E015DDC03803236A31E3" colspan="1" rowspan="1">9.7</td>
<td id="48DACB4CB413C37A7C7123CEC0576D7D" colspan="1" rowspan="1">9.52</td>
<td id="91B7BB34A428EC8669B2257692EE655E" colspan="1" rowspan="1">7.76</td>
<td id="92265DD168B029B19534CA1C9A645A4E" colspan="1" rowspan="1">9.36</td>
<td id="9FC6489C768A07D3CA8D00209C71C33B" colspan="1" rowspan="1">5.34</td>
<td id="C3164BD56A231FBC393F8BEDF3D46181" colspan="1" rowspan="1">2.96</td>
<td id="06713AA57DA7145CA01BE705823DEC2F" colspan="1" rowspan="1">2.5</td>
<td id="D9AD89B3FF7389548FA662CE7D5FF27F" colspan="1" rowspan="1">4.1</td>
<td id="1F499B3F550FB8EA92262828649A9D4A" colspan="1" rowspan="1">0</td>
<td id="ED221B84ACE60A477F817C6331A45703" colspan="1" rowspan="1">0</td>
<td id="04D457E3617507A14130CAE1DDA4BDD9" colspan="1" rowspan="1">2.12</td>
<td id="25D2DBF4988F04F53D8BE92D25E64463" colspan="1" rowspan="1">4.5</td>
<td id="E6CD85134123F53799BBEBD2B305406A" colspan="1" rowspan="1">6.5</td>
<td id="3B8097D43025038F953FE378CAFCF13B" colspan="1" rowspan="1">6.58</td>
<td id="23A4923841F8024D4E8A39FE14E4A62F" colspan="1" rowspan="1">11.42</td>
<td id="FC647A3B634D8E74E10EB10FEF35E687" colspan="1" rowspan="1">12.16</td>
<td id="869269BB820C30C1657EB71BA51D3E30" colspan="1" rowspan="1">7.7</td>
<td id="F07093E6D76FDBB4D78DCBE577942296" colspan="1" rowspan="1">9.26</td>
<td id="C204CACA05FB5A0F65B8836F3EA82079" colspan="1" rowspan="1">2.68</td>
<td id="014448CA86DE29310F982A15FBF09441" colspan="1" rowspan="1">2.72</td>
</tr>
<tr id="D80FC5DD41FA3AB268299DE0D9A697E7">
<td id="6DEC0691D7FD6462A886A0DE3CE8B9B9" colspan="1" rowspan="1">SACON VA- 301</td>
<td id="B2E8F7A547C631C1BED027F76509524D" colspan="1" rowspan="1">Male</td>
<td id="649A369162B25E3535905248F07717DC" colspan="1" rowspan="1">21.9</td>
<td id="0E804CA2C7FFB503EFF926FBEED8AED7" colspan="1" rowspan="1">7.94</td>
<td id="04CDEC92F57C32A22B50ED594E206B7D" colspan="1" rowspan="1">9.82</td>
<td id="01383B8E69FDBFE3A5D85E0BFEF8EE18" colspan="1" rowspan="1">6.42</td>
<td id="D028E4A2299291D80DE7B2B68A8D69BB" colspan="1" rowspan="1">8.58</td>
<td id="557C238BA265DB07A776F2B7AA39A536" colspan="1" rowspan="1">4.3</td>
<td id="276BB3D27EE4056EAAE28E156EBA457E" colspan="1" rowspan="1">3.24</td>
<td id="629D323260A7EA9B7225A8F0797F1850" colspan="1" rowspan="1">1.88</td>
<td id="90D309E7479631D3BD636713D85169B9" colspan="1" rowspan="1">3.52</td>
<td id="1365E00E7C940306550C772ED866C430" colspan="1" rowspan="1">0</td>
<td id="08AF0C30E4AF50CC680F6B0BBCE3789E" colspan="1" rowspan="1">0</td>
<td id="4CF6535BB2953CEB22CAF3E23AB19960" colspan="1" rowspan="1">1.88</td>
<td id="2BB793559A8DCA8B39478162BDF593F7" colspan="1" rowspan="1">3.8</td>
<td id="1EAD5BC7A5BEE42FD4FFAC3932C11570" colspan="1" rowspan="1">4.9</td>
<td id="DA32749BE9845F9F041FC4A021856658" colspan="1" rowspan="1">5.78</td>
<td id="D7553DDD3B1C07F78877F0BE20C6BA43" colspan="1" rowspan="1">9.9</td>
<td id="CB24EB25E4B320293705278415E58502" colspan="1" rowspan="1">10.86</td>
<td id="438BA48B114B719C159F8277780A92F6" colspan="1" rowspan="1">6.5</td>
<td id="130C9206CB55B271F3B3196501E6A29A" colspan="1" rowspan="1">9.2</td>
<td id="69C28806D2F15707B73331EF4F4080E7" colspan="1" rowspan="1">2.3</td>
<td id="05549FB5FD78C02BFF0E2CB74F935AC6" colspan="1" rowspan="1">2.92</td>
</tr>
<tr id="30497527F8F8B1DBEA14FA2357F51D2F">
<td id="07BEFF2FDFA7AC91F2F705679949D69E" colspan="1" rowspan="1">SACON VA- 302</td>
<td id="20C4AA5998D09AF9C9FA8ADC19634B3E" colspan="1" rowspan="1">Male</td>
<td id="1518662DD33DA4C385FA0132875AE24E" colspan="1" rowspan="1">25</td>
<td id="09A957BC040A44144B7066B70C7D2F37" colspan="1" rowspan="1">11.92</td>
<td id="77112AF78F464A805D018E50A5AEA1E7" colspan="1" rowspan="1">11.58</td>
<td id="8B2CEFA6D66E86FF3E9522D645374258" colspan="1" rowspan="1">7.78</td>
<td id="91AD839155CC6BAD73CF7474F7F13E67" colspan="1" rowspan="1">9.6</td>
<td id="A278911B00C17FD7224366E4A3E90206" colspan="1" rowspan="1">4.96</td>
<td id="50D49CE9D5A2D6E2C0E7103A853AD6F1" colspan="1" rowspan="1">3.72</td>
<td id="1BB7044CA1C7A1EAF0089E9D9A461F87" colspan="1" rowspan="1">1.66</td>
<td id="8028AF01E9EF84B77CC91267AE7F5B21" colspan="1" rowspan="1">3.6</td>
<td id="CD91CCA6E2464A0C20D9293DEF28DA6B" colspan="1" rowspan="1">0</td>
<td id="C8C07DE2593B778550D8BE2BF8FB0113" colspan="1" rowspan="1">0</td>
<td id="68C0E708AE0342CE0AA105864FA5EFF1" colspan="1" rowspan="1">2.9</td>
<td id="1C156B0686A5F0140FCCEED6A01AC435" colspan="1" rowspan="1">4.8</td>
<td id="9F6D3771170A885E3848992BBE650C86" colspan="1" rowspan="1">5.44</td>
<td id="B20C4927B2B3ED55D9869AB13B0004B0" colspan="1" rowspan="1">6.2</td>
<td id="80E1D5CA91A9D1A97A46210DB772204E" colspan="1" rowspan="1">12.54</td>
<td id="C47CF0A506E0B255A15B44ACCADBCAEF" colspan="1" rowspan="1">12.7</td>
<td id="CD4E14B91C0B7F58C4663F2BC554CB59" colspan="1" rowspan="1">7.18</td>
<td id="9431166713930CCF8924B840DBF93316" colspan="1" rowspan="1">9.44</td>
<td id="9221E6DCE1DEC55D86D7E466E444962D" colspan="1" rowspan="1">2.14</td>
<td id="51E779E8081DFB7FD4F42A9BF136B275" colspan="1" rowspan="1">3.32</td>
</tr>
<tr id="5306DDE9A6774B26512F163BCB5CAB97">
<td id="7157DB5F4A10DC8E9F0F460B9C58A32C" colspan="1" rowspan="1">SACON VA- 297</td>
<td id="89C1646343AC700C3F0322346D29AE46" colspan="1" rowspan="1">Male</td>
<td id="36F2CEC81487E7EDBD88D9AF28F1F88A" colspan="1" rowspan="1">25.2</td>
<td id="6A5B12ED870F8B843AC770A795A195D6" colspan="1" rowspan="1">13</td>
<td id="009C58F0A5B27E01CBEB6CE1DDF6083C" colspan="1" rowspan="1">11.66</td>
<td id="BF301D6B5E167DB01DA117AB5112BEBC" colspan="1" rowspan="1">8.14</td>
<td id="19F4E339BE124697709A843AC1E498D1" colspan="1" rowspan="1">9.56</td>
<td id="280F89D6702E56E49BE3EC1ABA6F6E08" colspan="1" rowspan="1">5.98</td>
<td id="B7BEBD84DEC90EE29E41A599F66EAB5D" colspan="1" rowspan="1">4.16</td>
<td id="08D9B56E51CA393A2DA8EF10845AEC21" colspan="1" rowspan="1">1.92</td>
<td id="42EDAD3DEE3E24E8B6FE5C688CC08A88" colspan="1" rowspan="1">3.86</td>
<td id="C587A4413A43888DF5C26122EA14E253" colspan="1" rowspan="1">0</td>
<td id="A12777829FC1355EF87E91BA1763D5FC" colspan="1" rowspan="1">0</td>
<td id="4155B90FC1DB6A4BA4C710DACA9CB9AC" colspan="1" rowspan="1">2.6</td>
<td id="385324B59F404E93100E58BE94B967C1" colspan="1" rowspan="1">3.94</td>
<td id="7AC13E0BFDE2DC85B88B92CB75D6EECF" colspan="1" rowspan="1">5.26</td>
<td id="65321EFE8B26C24038C98D636552D132" colspan="1" rowspan="1">6.12</td>
<td id="2C30783F3362A19BE06BF581C21D82C4" colspan="1" rowspan="1">12.2</td>
<td id="AC6BDD6F4DA9E6D16CE41C2D4C8511A5" colspan="1" rowspan="1">13.12</td>
<td id="65DF98DBACF82DB441B0FD77D956F8CB" colspan="1" rowspan="1">7.4</td>
<td id="4C88D2B42082048C8DE5061EDE95224F" colspan="1" rowspan="1">10.26</td>
<td id="1A03F300FC6104D6518BDC173DE6A280" colspan="1" rowspan="1">2.24</td>
<td id="D615C267155E68B350BB66EF28178297" colspan="1" rowspan="1">3.32</td>
</tr>
<tr id="95E65478A4834C56E184C648DC3332BF">
<td id="ECB950AD8F1CEC4574DC5604527D8243" colspan="1" rowspan="1">SACON VA- 299</td>
<td id="2D1D68AC3700877555A011EA41035823" colspan="1" rowspan="1">Male</td>
<td id="BBF0D576FC9D1940484164028D5EA7BE" colspan="1" rowspan="1">24.6</td>
<td id="DE2EA454A01D233D949E1042B0B76F76" colspan="1" rowspan="1">11.52</td>
<td id="3937DE69DF3C636446676147DEB11E8B" colspan="1" rowspan="1">10.6</td>
<td id="89F487DFA54134A4586140DB65700FC7" colspan="1" rowspan="1">7.52</td>
<td id="08975FDA9B1BCCC9C1ED94B04B4B2198" colspan="1" rowspan="1">10.68</td>
<td id="8DD7E2A66D1CD07B3EB6F5919EE5BAA6" colspan="1" rowspan="1">4.76</td>
<td id="D822D77EA3F8FAD8505C2EE253A29DD2" colspan="1" rowspan="1">3.62</td>
<td id="000EC4B0F202432DD3825115FA561579" colspan="1" rowspan="1">1.2</td>
<td id="7182EC724FD8003348FCC1226DDD3A4E" colspan="1" rowspan="1">3.34</td>
<td id="2C3FED15D444B23CC79E1202DFA1C581" colspan="1" rowspan="1">0</td>
<td id="4F96BF49932E7EE276370173C5F02716" colspan="1" rowspan="1">0</td>
<td id="C02F85F5B5CC1009C0D2059FA1C2ECE4" colspan="1" rowspan="1">2.2</td>
<td id="5BD8EB834E0DEDFD21475BD6AA20C4C1" colspan="1" rowspan="1">4.6</td>
<td id="3B893C46893BA07367404E423515A65E" colspan="1" rowspan="1">6.02</td>
<td id="743C82B4B597526DBDD01C0F09D395C3" colspan="1" rowspan="1">7.6</td>
<td id="14193DE80B8713222506546B6E7C3EE1" colspan="1" rowspan="1">12.48</td>
<td id="147C2DEC685462DD0C91551456EFDAA1" colspan="1" rowspan="1">12.72</td>
<td id="F60F0A03FC60AFDD324D12F81FF45BDA" colspan="1" rowspan="1">7.1</td>
<td id="602F2D4740E55F89782AF6A8B6EF51AF" colspan="1" rowspan="1">9.82</td>
<td id="71DD822AC00CFE3DF363DB3A32F2EE83" colspan="1" rowspan="1">2.2</td>
<td id="F8862B28980196EA1139B7FA3A78AD08" colspan="1" rowspan="1">3.00</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFLBI" type="distribution">
<paragraph id="8819A773957B2B85757C3468997A0F9A">
<heading id="B1D9A8900684C227B04A22A7AD0B8A85" reason="title">Natural history and distribution.</heading>
</paragraph>
<paragraph id="244EF303D947339CA8A4C812B2AE88BD">
Males were usually observed to start calling at dusk from bamboo bushes. The species was recorded from Mikadogre Community Reserve (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOLBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.399000,25.434000]}">
<geoCoordinate id="8FC596ED7B52466AE520269832C18167" degrees="25.434" direction="north" orientation="latitude" precision="55" value="25.434">25.434 ° N</geoCoordinate>
,
<geoCoordinate id="08326D8E4543C8A04BA4060863ABE5C4" degrees="90.399" direction="east" orientation="longitude" precision="55" value="90.399">90.399 ° E</geoCoordinate>
</named-content>
,
<quantity id="29D13EADA089E77CD89DF6DC5FF82200" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" unit="m" value="174.0">174 m</quantity>
) and Bollonggre village (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWLBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[89.995000,25.489000]}">
<geoCoordinate id="333230E38665DB53B15C3D9D8C1CC65E" degrees="25.489" direction="north" orientation="latitude" precision="55" value="25.489">25.489 ° N</geoCoordinate>
,
<geoCoordinate id="14BD358E78B1D987FB1BE29C5250F6C5" degrees="89.995" direction="east" orientation="longitude" precision="55" value="89.995">89.995 ° E</geoCoordinate>
</named-content>
,
<quantity id="725249FFC79340EE2253F835AEFD2F61" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
) from South Garo Hills and Nengmandalgre Fish Sanctuary (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5LBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[90.548000,25.494000]}">
<geoCoordinate id="A40189A50374101B386F567D8CF0D9DB" degrees="25.494" direction="north" orientation="latitude" precision="55" value="25.494">25.494 ° N</geoCoordinate>
,
<geoCoordinate id="2E5C7387997D7D0D4E3D002B422B407D" degrees="90.548" direction="east" orientation="longitude" precision="55" value="90.548">90.548 ° E</geoCoordinate>
</named-content>
,
<quantity id="C965DD6FE237CA519A8704AA65E9C40F" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</quantity>
) from East Garo Hills. Based on these locations and the locality provided by
<bibRefCitation id="39D7CCDBF210D78E2140E508EF6968C3" DOI="10.24272/j.issn.2095-8137.2019.066" author="Wu YH &amp; Suwannapoom C &amp; Xu K &amp; Chen JM &amp; Jin JQ &amp; Chen HM &amp; Murphy RW &amp; Che J" etAl="et al." firstAuthor="Wu" issue="6" journalOrPublisher="Zoological Research" pagination="558-563" refId="B33" refString="Wu YH, Suwannapoom C, Xu K, Chen JM, Jin JQ, Chen HM, Murphy RW, Che J (2019) A new species of the genus Raorchestes (Anura, Rhacophoridae) from Yunnan Province, China. Zoological Research 40 (6): 558563. https://doi.org/10.24272/j.issn.2095-8137.2019.066" title="A new species of the genus Raorchestes (Anura, Rhacophoridae) from Yunnan Province, China." volume="40" year="2019">Wu et al. (2019)</bibRefCitation>
, we estimated the extent of occurrence of this species to be
<quantity id="EB45C3955B6AFC5079DF414097CDCE0E" metricMagnitude="8" metricUnit="m" metricValue="2.4249099999999997" unit="km" value="242491.0">242,491 km</quantity>
². We observed it to be relatively abundant across an elevational gradient of
<quantity id="5F7150FFC05659DD13EAFD1EDDFA0BFD" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
asl to
<quantity id="48FD95DFB99B414AA709AA6C7E59C830" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
asl. Therefore, we recommend that this species be downgraded from Critically Endangered to the least concern category.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHMBI" type="remarks">
<paragraph id="761BABE6F6DBDF150DF83C0243F7EB1D">
<heading id="CD5375A0778D4FA5987C3F2226A24D7C" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="4CA7CCD849681723FC5BD5EC1E007829">
<bibRefCitation id="F97B1B57AC185FD947836E6BD2AEA816" DOI="10.24272/j.issn.2095-8137.2019.066" author="Wu YH &amp; Suwannapoom C &amp; Xu K &amp; Chen JM &amp; Jin JQ &amp; Chen HM &amp; Murphy RW &amp; Che J" etAl="et al." firstAuthor="Wu" issue="6" journalOrPublisher="Zoological Research" pagination="558-563" refId="B33" refString="Wu YH, Suwannapoom C, Xu K, Chen JM, Jin JQ, Chen HM, Murphy RW, Che J (2019) A new species of the genus Raorchestes (Anura, Rhacophoridae) from Yunnan Province, China. Zoological Research 40 (6): 558563. https://doi.org/10.24272/j.issn.2095-8137.2019.066" title="A new species of the genus Raorchestes (Anura, Rhacophoridae) from Yunnan Province, China." volume="40" year="2019">Wu et al. (2019)</bibRefCitation>
described
<taxonomicName id="FBC432F99C0054B064C69D3FBB88F927" authorityName="Wu, Suwannapoom, Xu, Murphy &amp; Che" authorityYear="2019" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cangyuanensis">
<emphasis id="54EA4DC60B5EE264EEE913C14EF6183D" italics="true">Raorchestes cangyuanensis</emphasis>
</taxonomicName>
from
<collectingRegion id="A749CE682532209F48A2D532E23E2270" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCountry id="43B9880C6623D1C767E2AA7942959BCB" name="China">China</collectingCountry>
. However, they did not compare this species with the senior taxon
<taxonomicName id="A1425D36ECE2CDD386113449BEF9667D" authorityName=", Boulenger" authorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Ixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="1BE93A48E344A8298861B025E65B72E6" italics="true">Ixalus kempiae</emphasis>
</taxonomicName>
, likely because
<taxonomicName id="1170D9F11CC6719CF377538D848D6A12" authorityName=", Boulenger" authorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Ixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="A62AF268E8691F17FED4F9F1BB404596" italics="true">I. kempiae</emphasis>
</taxonomicName>
had remained poorly known since its description and there had been no literature available on this species based on new sightings since it was first described. It is also unusual for frogs of the genus
<taxonomicName id="D3A5A5FE8736BC415BB60DCDC2145CCA" authorityName="Biju, Shouche, Dubois, Dutta &amp; Bossuyt" authorityYear="2010" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="A94E1C674D059406CB5BE328E1EF01AE" italics="true">Raorchestes</emphasis>
</taxonomicName>
to have such wide distribution ranges (see
<bibRefCitation id="CF4AFFE68D825A4B34FE483F186E753C" DOI="10.11646/zootaxa.3893.4.1" author="Vijayakumar" etAl="et al." firstAuthor="Vijayakumar" issue="4" journalOrPublisher="Zootaxa" pagination="451-488" refId="B31" refString="Vijayakumar SP, Dinesh KP, Prabhu MV, Shanker K (2014) Lineage delimitation and description of nine new species of bush frogs (Anura, Raorchestes, Rhacophoridae) from the Western Ghats Escarpment. Zootaxa 3893 (4): 451488. https://doi.org/10.11646/zootaxa.3893.4.1" title="Lineage delimitation and description of nine new species of bush frogs (Anura, Raorchestes, Rhacophoridae) from the Western Ghats Escarpment." volume="3893" year="2014">Vijayakumar et al. 2014</bibRefCitation>
). We conclude that
<taxonomicName id="363B89ADB9C6A13FEEADA460A8AA5C3A" authorityName="Wu, Suwannapoom, Xu, Murphy &amp; Che" authorityYear="2019" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cangyuanensis">
<emphasis id="7B3715F7E242BC964262AD7141B33778" italics="true">Raorchestes cangyuanensis</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="B8A53983A695AB8DCBA4D3E48BD5DDF3" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="6788FA547513B88D7200FFCAC2F7360E" italics="true">Raorchestes kempiae</emphasis>
</taxonomicName>
based on molecular and morphological evidence (see Tables
<tableCitation id="4F2141027224BDC8F0286BEF33195BCC" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Uncorrected 16 s pairwise genetic divergences between the Rhacophoridae species studied." httpUri="http://table.plazi.org/id/A4D4B456602712C6C9AE660F1C1D9331" tableUuid="A4D4B456602712C6C9AE660F1C1D9331">1</tableCitation>
,
<tableCitation id="72DD3B70DB2B93C2E289054D887AA9F9" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric measurements (in mm) of R. kempiae." httpUri="http://table.plazi.org/id/2CBC684D6BFAAC65AB8CB42DF48FE3FC" tableUuid="2CBC684D6BFAAC65AB8CB42DF48FE3FC">3</tableCitation>
). In addition to this,
<bibRefCitation id="6CE344150CB2BA6048F18BB50966F2D3" DOI="10.11609/jott.8590.15.10.24016-24031" author="Gouda S &amp; Decemson Ht &amp; Zoramkhuma &amp; Malsawmdawngliana F &amp; Biakzuala L &amp; Lalremsanga HT" etAl="et al." firstAuthor="Gouda" issue="10" journalOrPublisher="Journal of Threatened Taxa" pagination="24016-24031" refId="B16" refString="Gouda S, Decemson Ht, Zoramkhuma, Malsawmdawngliana F, Biakzuala L, Lalremsanga HT (2023) Threat assessment and conservation challenges for the herpetofaunal diversity of Dampa Tiger Reserve, Mizoram, India. Journal of Threatened Taxa 15 (10): 2401624031. https://doi.org/10.11609/jott.8590.15.10.24016-24031" title="Threat assessment and conservation challenges for the herpetofaunal diversity of Dampa Tiger Reserve, Mizoram, India." volume="15" year="2023">Gouda et al. (2023)</bibRefCitation>
regarded
<taxonomicName id="5DE0CB1DB1FA80539E3B65DB8DFF7111" authorityName="Wu, Suwannapoom, Xu, Murphy &amp; Che" authorityYear="2019" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cangyuanensis">
<emphasis id="E5F2B7632D991EABACC35A99CC5C9E55" italics="true">Raorchestes cangyuanensis</emphasis>
</taxonomicName>
as a subjective junior synonym of
<taxonomicName id="91BC3C04BC921E3891112F632EBA695F" authority="(Mathew &amp; Sen, 2009)" baseAuthorityName="Mathew &amp; Sen" baseAuthorityYear="2009" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="manipurensis">
<emphasis id="BFC84E863E45869989C6C0197E109C4A" italics="true">R. manipurensis</emphasis>
(Mathew &amp; Sen, 2009)
</taxonomicName>
. Our analysis also revealed that specimens referable to both these names actually show characters that are diagnostic of
<taxonomicName id="6D9326741CBA1475C9CCA9012034C58C" authority="(Boulenger 1919)" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="F10C81AFD03D35A520D1BFBCF1D32652" italics="true">Raorchestes kempiae</emphasis>
(
<bibRefCitation id="27AF632FBA21C3CC57DF7ECDD2441236" DOI="10.26515/rzsi/v16/i2/1919/162810" author="Boulenger" firstAuthor="Boulenger" issue="2" journalOrPublisher="Records of the Zoological Survey of India" pagination="207-208" refId="B7" refString="Boulenger GA (1919) XII Descriptions of Three new Batrachians from the Garo Hills, Assam. Records of the Zoological Survey of India 16 (2): 207208. https://doi.org/10.26515/rzsi/v16/i2/1919/162810" title="XII Descriptions of Three new Batrachians from the Garo Hills, Assam." volume="16" year="1919">Boulenger 1919</bibRefCitation>
)
</taxonomicName>
, a species described much earlier but overlooked until now due to improper and deficient comparisons. However, since we did not have molecular data or access to type specimens of
<taxonomicName id="3F54368E3EE692937E1D326F511B93AE" baseAuthorityName="Mathew &amp; Sen" baseAuthorityYear="2009" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="manipurensis">
<emphasis id="A1FBDA4BF087A75EB210F530FEFCACC9" italics="true">R. manipurensis</emphasis>
</taxonomicName>
, we do not address this in the current work and recommend that resolving this taxonomic uncertainty should be a priority for further study.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPQBI" type="comparisons">
<paragraph id="BE848F7219D92D111DEEA7D420429FFC">
<heading id="4ADDEE78D4EA23BCD73DD5B7AA6F7055" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="123C1DFB6D92E38616F1E19577728577">
<taxonomicName id="ED2E540CE6D93507C79E756C7BFDC738" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="BCDC713DAF40F136ECF8A94EC42FE13A" italics="true">Raorchestes kempiae</emphasis>
</taxonomicName>
is genetically divergent (5.259.47 % on the 16 S rRNA gene) from congeners, with the least divergent species being Western Ghats endemic
<taxonomicName id="45ED1BC5EE54269B30B8CF1A0A7CCFFE" authorityName="Padhye, Sayyed, Jadhav &amp; Dahanukar" authorityYear="2013" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ghatei">
<emphasis id="8CB15BFE31C16DB6D8BABEEDA3E74012" italics="true">R. ghatei</emphasis>
</taxonomicName>
, with a divergence of 5.77 %. From congeners in this region, it can be easily distinguished by: absence of an externally visible tympanum (vs. present in
<taxonomicName id="78A06C28EF1F12F8A3140CDB14BCD239" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="B3EBF3593E762D5D04A7FEE9D6A0262F" italics="true">R. garo</emphasis>
</taxonomicName>
,
<taxonomicName id="B95F5B042DA3E8E4DA21E594BAACDE9C" authorityName="Du, Xu, Liu &amp; Yu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hekouensis">
<emphasis id="AECD228FCA796283A4E7718E9634DAAD" italics="true">R. hekouensis</emphasis>
</taxonomicName>
,
<taxonomicName id="70F6F380C30AAE858CB1316A3796FB00" authorityName="Huang, Liu, Du, Bernstein, Liu, Yang, Yu &amp; Wu" authorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="malipoensis">
<emphasis id="863FF7F27556D8582CB77124F05911DB" italics="true">R. malipoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="379985E949BF902D43DF3A62515EA63C" baseAuthorityName="Boulenger" baseAuthorityYear="1893" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="parvulus">
<emphasis id="267B15C76E061C9F2F67FED5AC4120FA" italics="true">R. parvulus</emphasis>
</taxonomicName>
,
<taxonomicName id="68DD8DA62237BBEB193E2C3FE739162C" authorityName="Wu, Liu, Gao, Wang, Li, Zhou, Yuan &amp; Che" authorityYear="2021" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="dulongensis">
<emphasis id="AA5D14BB1DCE77F8F3889EE27A493DF7" italics="true">R. dulongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="7E6C3B1F1D8050B0E3251D955746E8F7" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hillisi">
<emphasis id="56226712623511F21183ABCDC7F30FD7" italics="true">R. hillisi</emphasis>
</taxonomicName>
,
<taxonomicName id="692145C752B7451C6BE7CF2ED3F54C17" authorityName="Jiang, Wang, Ren &amp; Li" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="huanglianshan">
<emphasis id="A3AAA6B418A50461223509216FB5B103" italics="true">R. huanglianshan</emphasis>
</taxonomicName>
); ventral aspect of the hind limb uniform creamy white (vs. dark gray in
<taxonomicName id="B823F1040D2ADED3B9F1D81E7D132C60" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="7E20B78AC8A8FF544DB8B99F8E4D06B6" italics="true">R. rezakhani</emphasis>
</taxonomicName>
); ventral surface mostly smooth with slight granulation (vs. granulated and well developed whitish tubercles in
<taxonomicName id="8A1E2BC8C7B4FC776F167CE43F5D08FB" authorityName="Du, Xu, Liu &amp; Yu" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hekouensis">
<emphasis id="7167CBBD92127B239B6454BAB993E095" italics="true">R. hekouensis</emphasis>
</taxonomicName>
); toe webbing I 1-2 II 2 - 1 III 1 - 1 IV 1 - 2 V in
<taxonomicName id="680936A0782CF861F088FFBAB96694BE" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="2173F7A2195210827BFD38ED10FABCBA" italics="true">R. kempiae</emphasis>
</taxonomicName>
(vs. I 1-1 II 1 - 1 III 3 - 3 IV 3 - 2.5 V in
<taxonomicName id="1980CBECF8938BBA5F4AAEDA489422F4" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="A1BD4797D935C951DB962A3A5CAAC328" italics="true">R. garo</emphasis>
</taxonomicName>
, I 22 II 22 III 23 IV 32 V in
<taxonomicName id="4BD70459BC380DC7AC6958894F614613" authority=", II" authorityName="Huang, Liu, Du, Bernstein, Liu, Yang, Yu &amp; Wu" authorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="malipoensis">
<emphasis id="93D8808D9A0700AC36B6CB0000D84543" italics="true">R. malipoensis</emphasis>
, II
</taxonomicName>
12 III 12.5 IV 2.51 V in
<taxonomicName id="FC86D929BD1E73A543881338457086CE" authority=", II" authorityName="II" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="menglaensis">
<emphasis id="AC4E5D77E077F4ACD86CD1C6E783D3EC" italics="true">R. menglaensis</emphasis>
, II
</taxonomicName>
12 III 12.5 IV 2.51 V in
<taxonomicName id="10EE18720509F515C1BB139F29CA9BF5" authority=", II" authorityName="II" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="hillisi">
<emphasis id="88A4CFE888F0D3DF27BC4613394E669E" italics="true">R. hillisi</emphasis>
, II
</taxonomicName>
12 III 12 IV 21 V in
<taxonomicName id="457A319EFC16A337612678EA196DB685" authorityName="Jiang, Wang, Ren &amp; Li" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="huanglianshan">
<emphasis id="2F9C37A764B758E0F829C690569A592D" italics="true">R. huanglianshan</emphasis>
</taxonomicName>
and I 22 II 1.752 III 1.53 IV 2.752 V in
<taxonomicName id="345E50B0CF84DBACAC4E7DB8F55170C2" authorityName="Al-Razi, Maria &amp; Muzaffar" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rezakhani">
<emphasis id="009F5283965B52CFB68B3806C5F88A68" italics="true">R. rezakhani</emphasis>
</taxonomicName>
); snout rounded in
<taxonomicName id="1214636B8258A050A9D5D6C0EC0A0073" authorityName="Naveen, Chandramouli &amp; Babu" authorityYear="2024" baseAuthorityName="Boulenger" baseAuthorityYear="1919" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kempiae">
<emphasis id="A068D0128C688C35EA2C9B098DACDB93" italics="true">R. kempiae</emphasis>
</taxonomicName>
(vs. pointed in
<taxonomicName id="A0D0E393D7C51DEB89FEFD6DC6B27EF9" baseAuthorityName="Boulenger" baseAuthorityYear="1906" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="annandalii">
<emphasis id="E04459CBEB3C6456D66B855762BB473D" italics="true">R. annandalii</emphasis>
</taxonomicName>
and short and truncate in
<taxonomicName id="AA8EC3F22D7DBC8F61AE0D6C4E5D7849" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="garo">
<emphasis id="44A4AB7ABA1D516551A262457FD14055" italics="true">R. garo</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>