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<mods:title id="9AF5B1A451A4DB43708CCC4C195CED40">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:affiliation id="037737698457999C70362EFAB1A958F1">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
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<emphasis id="B48ABC63F741FFE0FF66D87CF8BC9F54" bold="true" box="[151,449,582,609]" italics="true" pageId="17" pageNumber="18">Pseudagrion declaratum</emphasis>
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Figures: 2628 (
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155 (field photo
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: 99,
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(
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appendages;
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<emphasis id="B48ABC63F741FFE0FF66D8DDFB729FCA" box="[151,527,743,767]" italics="true" pageId="17" pageNumber="18">Pseudagrion pilidorsum declaratum</emphasis>
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:
<bibRefCitation id="E26F1D80F741FFE0FDD3D8DDFBAC9FCA" author="Lieftinck, M. A." box="[546,721,743,767]" pageId="17" pageNumber="18" pagination="99 - 160" refId="ref40810" refString="Lieftinck, M. A. (1936) Die Odonaten der Kleinen Sunda-Inseln. Revue Suisse de Zoologie, 43, 99-160. https://doi.org/10.5962/bhl.part.145462" type="journal article" year="1936">Lieftinck (1936)</bibRefCitation>
: 130, 132134,
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(description
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appendages; key;
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Lombok, Sumbawa,
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,
<collectingCountry id="FEE920E1F741FFE0FE01D93CFB529E2B" box="[496,559,774,798]" name="Indonesia" pageId="17" pageNumber="18">Flores</collectingCountry>
);—
<bibRefCitation id="E26F1D80F741FFE0FDA5D93CFA7E9E2B" author="Lieftinck, M. A." box="[596,771,774,798]" pageId="17" pageNumber="18" pagination="118 - 228" refId="ref40941" refString="Lieftinck, M. A. (1953) The Odonata of the island Sumba with a survey of the dragonfly fauna of the Lesser Sunda Islands. Verhandlungen der Naturforschenden Gesellschaft in Basel, 64, 118-228." type="journal article" year="1953">Lieftinck (1953)</bibRefCitation>
: 156157 (Alor, Sawu);—
<bibRefCitation id="E26F1D80F741FFE0FBFFD93CFDCA9E2B" author="Kosterin, O. E." box="[1038,1207,774,798]" pageId="17" pageNumber="18" pagination="1 - 48" refId="ref40479" refString="Kosterin, O. E. (2014) Odonata briefly observed on the islands of Bali and Lombok, Lesser Sundas, Indonesia, in the late February 2014. International Dragonfly Fund-Report, 74, 1-48." type="journal article" year="2014">Kosterin (2014)</bibRefCitation>
: 16, 21,
<figureCitation id="1EC57CF4F741FFE0FAFED93CFC2D9E2B" box="[1295,1360,774,798]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="17" pageNumber="18">Figs 9</figureCitation>
,
<figureCitation id="1EC57CF4F741FFE0FAADD93CFC0A9E2B" box="[1372,1399,774,798]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="17" pageNumber="18">13</figureCitation>
,
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(
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, scarce; Lombok, abbundant; photos alive &amp; habitat).
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<emphasis id="B48ABC63F741FFE0FF66D979F8939E6E" box="[151,494,835,859]" italics="true" pageId="17" pageNumber="18">Pseudagrion pilidorsum deflexum</emphasis>
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:
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: 87 (
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Name-bearing
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)
</paragraph>
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<materialsCitation id="36966A2CF741FFE0FF36D997FC119EF2" box="[199,1388,941,968]" collectingDate="1896-04" collectionCode="ZMH" collectorName="H. Fruhstorfer" country="Indonesia" elevation="2000" location="Sapit" municipality="Lombok" pageId="17" pageNumber="18" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="90F8ABF8F741FFE0FF36D997F9919EF2" box="[199,236,941,967]" count="1" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
,
<specimenCount id="90F8ABF8F741FFE0FF09D997F8669EF2" box="[248,283,941,967]" count="1" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
<typeStatus id="5945DED3F741FFE0FED3D994F8019EFD" box="[290,380,942,968]" pageId="17" pageNumber="18" type="syntype">syntype</typeStatus>
,
<date id="F24046B1F741FFE0FE76D997F8A09EF2" box="[391,477,941,967]" pageId="17" pageNumber="18" value="1896-04">
<collectingDate id="E204BF59F741FFE0FE76D997F8A09EF2" box="[391,477,941,967]" pageId="17" pageNumber="18" value="1896-04">iv.1896</collectingDate>
</date>
,
<location id="832136AAF741FFE0FE19D997FB599EF2" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF741FFE0FE19D997FB599EF2" box="[488,548,941,967]" country="Indonesia" municipality="Lombok" name="Sapit" pageId="17" pageNumber="18">Sapit</location>
,
<collectingMunicipality id="6625FA0BF741FFE0FDC1D997FBEF9EF2" box="[560,658,941,967]" pageId="17" pageNumber="18">Lombok</collectingMunicipality>
,
<collectingCountry id="FEE920E1F741FFE0FD6CD997FA709EF2" box="[669,781,941,967]" name="Indonesia" pageId="17" pageNumber="18">Indonesia</collectingCountry>
,
<elevation id="0DD38742F741FFE0FCE9D997FAD49EF2" box="[792,937,941,968]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="2000.0">
<quantity id="4106CD94F741FFE0FCE9D997FA109EFD" box="[792,877,941,968]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F741FFE0FC44D994FD2B9EF2" box="[949,1110,941,968]" pageId="17" pageNumber="18">H. Fruhstorfer</collectorName>
leg.,
<collectionCode id="E0EFF8B4F741FFE0FB65D994FDAF9EFD" box="[1172,1234,942,968]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="17" pageNumber="18" type="Museum">ZMH</collectionCode>
(
<figureCitation id="1EC57CF4F741FFE0FB10D997FC199EF2" box="[1249,1380,941,967]" captionStart="FIGURES 2628" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[152,1432,1242,1879]" captionTargetId="figure-369@17.[151,1436,1241,1882]" captionTargetPageId="17" captionText="FIGURES 2628. Pseudagrion pilidorsum declaratum, syntype male: (26) habitus dorsal, (27) labels, (28) habitus lateral. Photos: M. Seehausen." pageId="17" pageNumber="18">Figs 2628</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F741FFE0FF36D9EBFD079995" blockId="17.[151,1437,905,1184]" pageId="17" pageNumber="18">
<bibRefCitation id="E26F1D80F741FFE0FF36D9EBF8D79EDE" author="Lieftinck, M. A." box="[199,426,977,1003]" pageId="17" pageNumber="18" pagination="65 - 139" refId="ref40982" refString="Lieftinck, M. A. (1971) A catalogue of the type-specimens of Odonata preserved in the Netherlands. With a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65-139, pls. 1 - 7." type="journal article" year="1971">Lieftinck (1971: 83)</bibRefCitation>
designated a
<typeStatus id="5945DED3F741FFE0FDB4D9EBFBD19EDE" box="[581,684,977,1003]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
<materialsCitation id="36966A2CF741FFE0FD42D9EBF8F09925" collectingDate="1937-11" collectorName="J. K. de Jong" elevation="100" location="Badjo" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Labuan" typeStatus="syntype">
male with the data »
<collectingRegion id="443AAE93F741FFE0FC64D9EBFA969EDE" box="[917,1003,977,1003]" country="Malaysia" name="Labuan" pageId="17" pageNumber="18">Labuan</collectingRegion>
<location id="832136AAF741FFE0FC03D9EBFD459EDE" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF741FFE0FC03D9EBFD459EDE" box="[1010,1080,977,1003]" name="Badjo" pageId="17" pageNumber="18" stateProvince="Labuan">Badjo</location>
,
<quantity id="4106CD94F741FFE0FBB5D9EBFDF09ED9" box="[1092,1165,977,1004]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">
<elevation id="0DD38742F741FFE0FBB5D9EBFDF09ED9" box="[1092,1165,977,1004]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="F24046B1F741FFE0FB68D9EBFD8D9EDE" box="[1177,1264,977,1003]" pageId="17" pageNumber="18" value="1937-11">
<collectingDate id="E204BF59F741FFE0FB68D9EBFD8D9EDE" box="[1177,1264,977,1003]" pageId="17" pageNumber="18" value="1937-11">xi.1937</collectingDate>
</date>
,
<collectorName id="2B0B05A7F741FFE0FB0DD9E8FCF69ED9" box="[1276,1419,977,1004]" pageId="17" pageNumber="18">J.K. de Jong</collectorName>
«. However, the
<typeStatus id="5945DED3F741FFE0FEC7D9CCF8F09925" box="[310,397,1014,1040]" pageId="17" pageNumber="18" type="syntype">syntype</typeStatus>
</materialsCitation>
<materialsCitation id="36966A2CF741FFE0FE65D9CFFCE1996D" collectingDate="1929-10" collectingDateMax="1929-11-31" collectingDateMin="1929-10" collectionCode="RMNH" collectorName="Furthermore &amp; No. &amp; Thus" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Labuan" typeStatus="syntype">
series contains only specimens from this locality and collector which were collected during »
<date id="F24046B1F741FFE0FF54DE23F85C9906" box="[165,289,1049,1075]" pageId="17" pageNumber="18" value="1929-10" valueMax="1929-11-31" valueMin="1929-10-00">
<collectingDate id="E204BF59F741FFE0FF54DE23F85C9906" box="[165,289,1049,1075]" pageId="17" pageNumber="18" value="1929-10" valueMax="1929-11-31" valueMin="1929-10-00">x.xi.1929</collectingDate>
</date>
«.
<collectorName id="2B0B05A7F741FFE0FECADE23F8BA9906" box="[315,455,1049,1075]" pageId="17" pageNumber="18">Furthermore</collectorName>
the respective envelope held at
<collectionCode id="E0EFF8B4F741FFE0FCCBDE20FAF39901" box="[826,910,1050,1076]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18" type="Museum">RMNH</collectionCode>
(
<collectorName id="2B0B05A7F741FFE0FC6EDE23FAB79906" box="[927,970,1049,1075]" pageId="17" pageNumber="18">No.</collectorName>
JvT no. 2813) contains
<specimenCount id="90F8ABF8F741FFE0FB10DE20FC289906" box="[1249,1365,1049,1076]" count="2" pageId="17" pageNumber="18" type="male">two males</specimenCount>
and a female.
<collectorName id="2B0B05A7F741FFE0FF00DE07F8549962" box="[241,297,1085,1111]" pageId="17" pageNumber="18">Thus</collectorName>
the designation by
<bibRefCitation id="E26F1D80F741FFE0FDF2DE07FBC69962" author="Lieftinck, M. A." box="[515,699,1085,1111]" pageId="17" pageNumber="18" pagination="65 - 139" refId="ref40982" refString="Lieftinck, M. A. (1971) A catalogue of the type-specimens of Odonata preserved in the Netherlands. With a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65-139, pls. 1 - 7." type="journal article" year="1971">Lieftinck (1971)</bibRefCitation>
is invalid because he did not choose a specimen from the
<typeStatus id="5945DED3F741FFE0FAB4DE04FCE1996D" box="[1349,1436,1086,1112]" pageId="17" pageNumber="18" type="syntype">syntype</typeStatus>
</materialsCitation>
series, and he even did not unambiguously select a particular specimen (
<bibRefCitation id="E26F1D80F741FFE0FC28DE5BFD1F994E" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[985,1122,1121,1147]" pageId="17" pageNumber="18" refId="ref40194" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: 74.2 and 74.5). We do not consider it necessary to designate a
<typeStatus id="5945DED3F741FFE0FDD9DEBFFBEF99AA" box="[552,658,1157,1183]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
, thus we propose to keep them as
<typeStatus id="5945DED3F741FFE0FBE1DEBCFD0B9995" box="[1040,1142,1158,1184]" pageId="17" pageNumber="18" type="syntype">syntypes</typeStatus>
.
</paragraph>
<caption id="D28130F9F741FFE0FF66DD4EF8029A85" pageId="17" pageNumber="18" startId="17.[151,264,1908,1932]" targetBox="[152,1432,1242,1879]" targetPageId="17" targetType="figure">
<paragraph id="86416071F741FFE0FF66DD4EF8029A85" blockId="17.[151,1437,1908,1968]" pageId="17" pageNumber="18">
<emphasis id="B48ABC63F741FFE0FF66DD4EF82D9AB9" bold="true" box="[151,336,1908,1932]" pageId="17" pageNumber="18">FIGURES 2628</emphasis>
.
<taxonomicName id="41FE1BF2F741FFE0FE91DD4EFBA89AB9" authorityName="Lieftinck" authorityYear="1936" box="[352,725,1908,1932]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="pilidorsum" subSpecies="declaratum">
<emphasis id="B48ABC63F741FFE0FE91DD4EFBA89AB9" box="[352,725,1908,1932]" italics="true" pageId="17" pageNumber="18">Pseudagrion pilidorsum declaratum</emphasis>
</taxonomicName>
, syntype male: (
<emphasis id="B48ABC63F741FFE0FC76DD4EFADD9AB9" bold="true" box="[903,928,1908,1932]" pageId="17" pageNumber="18">26</emphasis>
) habitus dorsal, (
<emphasis id="B48ABC63F741FFE0FBAADD4EFD099AB9" bold="true" box="[1115,1140,1908,1932]" pageId="17" pageNumber="18">27</emphasis>
) labels, (
<emphasis id="B48ABC63F741FFE0FB28DD4EFD8F9AB9" bold="true" box="[1241,1266,1908,1932]" pageId="17" pageNumber="18">28</emphasis>
) habitus lateral. Photos: M. Seehausen.
</paragraph>
</caption>
<paragraph id="86416071F742FFE3FF36DAACFBD29D84" blockId="18.[151,1437,150,2013]" box="[199,687,150,177]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DAACFB749D85" bold="true" box="[199,521,150,176]" pageId="18" pageNumber="19">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F742FFE3FDE9DAACFB1F9D85" box="[536,610,150,176]" count="29" pageId="18" pageNumber="19" type="male">29♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F742FFE3FD9CDAACFBD99D84" box="[621,676,150,177]" count="5" pageId="18" pageNumber="19" type="female">5♀♀</specimenCount>
)
</paragraph>
<paragraph id="86416071F742FFE3FF36DA80FB219C29" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<materialsCitation id="36966A2CF742FFE3FF36DA80FD759DE1" box="[199,1032,186,213]" collectingDate="1896-11" collectionCode="SMF" collectorName="R. Martin" country="Indonesia" location="South" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2">
<emphasis id="B48ABC63F742FFE3FF36DA80F89F9DE1" bold="true" box="[199,482,186,212]" pageId="18" pageNumber="19">
<collectingCountry id="FEE920E1F742FFE3FF36DA80F8409DE1" box="[199,317,186,212]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
,
<collectingCountry id="FEE920E1F742FFE3FEB9DA80F89F9DE1" box="[328,482,186,212]" name="Flores Island" pageId="18" pageNumber="19">Flores Island</collectingCountry>
</emphasis>
:
<specimenCount id="90F8ABF8F742FFE3FE1FDA80FB549DE1" box="[494,553,186,212]" count="2" pageId="18" pageNumber="19" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F742FFE3FDC3DA80FBF49DE1" box="[562,649,186,212]" pageId="18" pageNumber="19" value="1896-11">
<collectingDate id="E204BF59F742FFE3FDC3DA80FBF49DE1" box="[562,649,186,212]" pageId="18" pageNumber="19" value="1896-11">xi.1896</collectingDate>
</date>
,
<location id="832136AAF742FFE3FD63DA80FBAE9DE1" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF742FFE3FD63DA80FBAE9DE1" box="[658,723,186,212]" country="Indonesia" name="South" pageId="18" pageNumber="19">South</location>
Flores,
<collectorName id="2B0B05A7F742FFE3FCDBDA81FAE89DE1" box="[810,917,186,213]" pageId="18" pageNumber="19">R. Martin</collectorName>
leg.,
<collectionCode id="E0EFF8B4F742FFE3FC3EDA80FD759DE1" box="[975,1032,186,212]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
</materialsCitation>
, no. 1397313974;—
<materialsCitation id="36966A2CF742FFE3FB0EDA80FA139DCD" collectingDate="1908-09-28" collectionCode="RMNH" country="Indonesia" county="river Alo Wai" location="North" municipality="Rioeng Papang" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1">
<specimenCount id="90F8ABF8F742FFE3FB0EDA80FC619DE1" box="[1279,1308,186,212]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<date id="F24046B1F742FFE3FAD5DA80FCE59DE1" box="[1316,1432,186,212]" pageId="18" pageNumber="19" value="1908-09-28">
<collectingDate id="E204BF59F742FFE3FAD5DA80FCE59DE1" box="[1316,1432,186,212]" pageId="18" pageNumber="19" value="1908-09-28">28.ix1908</collectingDate>
</date>
,
<location id="832136AAF742FFE3FF66DAE4F9A59DCD" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF742FFE3FF66DAE4F9A59DCD" box="[151,216,222,248]" country="Indonesia" county="river Alo Wai" municipality="Rioeng Papang" name="North" pageId="18" pageNumber="19">North</location>
<collectingCountry id="FEE920E1F742FFE3FF2EDAE4F8559DCD" box="[223,296,222,248]" name="Indonesia" pageId="18" pageNumber="19">Flores</collectingCountry>
,
<collectingMunicipality id="6625FA0BF742FFE3FEC2DAE4F89D9DCC" box="[307,480,222,249]" pageId="18" pageNumber="19">Rioeng Papang</collectingMunicipality>
,
<collectingCounty id="6F2018FDF742FFE3FE1DDAE4FBF89DCD" box="[492,645,222,248]" pageId="18" pageNumber="19">river Alo Wai</collectingCounty>
, v.d. S. leg.,
<collectionCode id="E0EFF8B4F742FFE3FCE5DAE4FA139DCD" box="[788,878,222,248]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="19" type="Museum">RMNH</collectionCode>
</materialsCitation>
;—
<materialsCitation id="36966A2CF742FFE3FC60DAE4F8CA9C28" collectingDate="1937-06" collectionCode="RMNH" collectorName="J. K. de Jong" country="Indonesia" location="Labuan Badjo" municipality="West" pageId="18" pageNumber="19" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="90F8ABF8F742FFE3FC60DAE4FABA9DCD" box="[913,967,222,248]" count="2" pageId="18" pageNumber="19" type="male">2♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F742FFE3FC23DAE4FA889DCD" box="[978,1013,222,248]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
,
<date id="F24046B1F742FFE3FBF2DAE4FD279DCD" box="[1027,1114,222,248]" pageId="18" pageNumber="19" value="1937-06">
<collectingDate id="E204BF59F742FFE3FBF2DAE4FD279DCD" box="[1027,1114,222,248]" pageId="18" pageNumber="19" value="1937-06">vi.1937</collectingDate>
</date>
,
<location id="832136AAF742FFE3FB97DAE4FC7A9DCD" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF742FFE3FB97DAE4FC7A9DCD" box="[1126,1287,222,248]" country="Indonesia" municipality="West" name="Labuan Badjo" pageId="18" pageNumber="19">Labuan Badjo</location>
,
<collectingMunicipality id="6625FA0BF742FFE3FAE0DAE5FC359DCC" box="[1297,1352,223,249]" pageId="18" pageNumber="19">West</collectingMunicipality>
<collectingCountry id="FEE920E1F742FFE3FABEDAE4FCE59DCD" box="[1359,1432,222,248]" name="Indonesia" pageId="18" pageNumber="19">Flores</collectingCountry>
,
<collectorName id="2B0B05A7F742FFE3FF66DB39F85F9C28" box="[151,290,258,285]" pageId="18" pageNumber="19">J.K. de Jong</collectorName>
leg.,
<collectionCode id="E0EFF8B4F742FFE3FE91DB39F8CA9C28" box="[352,439,259,285]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="19" type="Museum">RMNH</collectionCode>
</materialsCitation>
, JvT no. 2813.
</paragraph>
<paragraph id="86416071F742FFE3FF36DB1CFB829C98" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<materialsCitation id="36966A2CF742FFE3FF36DB1CFD9E9C75" box="[199,1251,294,321]" collectingDate="1896-04" collectionCode="SMF" collectorName="Sapit &amp; H. Fruhstorfer" country="Indonesia" elevation="2000" location="Lombok Island" pageId="18" pageNumber="19" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Lombok Island">
<emphasis id="B48ABC63F742FFE3FF36DB1CFB7C9C75" bold="true" box="[199,513,294,320]" pageId="18" pageNumber="19">
<collectingCountry id="FEE920E1F742FFE3FF36DB1CF8409C75" box="[199,317,294,320]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F742FFE3FEBBDB1CFB7C9C75" box="[330,513,294,320]" pageId="18" pageNumber="19">Lombok Island</collectingRegion>
</emphasis>
:
<specimenCount id="90F8ABF8F742FFE3FDFEDB1DFB379C74" box="[527,586,295,321]" count="7" pageId="18" pageNumber="19" type="male">7♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F742FFE3FDA4DB1CFB059C75" box="[597,632,294,320]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
,
<date id="F24046B1F742FFE3FD77DB1CFBA69C75" box="[646,731,294,320]" pageId="18" pageNumber="19" value="1896-04">
<collectingDate id="E204BF59F742FFE3FD77DB1CFBA69C75" box="[646,731,294,320]" pageId="18" pageNumber="19" value="1896-04">iv.1896</collectingDate>
</date>
,
<collectorName id="2B0B05A7F742FFE3FD16DB1CFA5E9C75" box="[743,803,294,320]" pageId="18" pageNumber="19">Sapit</collectorName>
,
<elevation id="0DD38742F742FFE3FCDFDB1CFAC29C75" box="[814,959,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="m" value="2000.0">
<quantity id="4106CD94F742FFE3FCDFDB1CFAFE9C74" box="[814,899,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F742FFE3FC3DDB1DFD119C75" box="[972,1132,294,321]" pageId="18" pageNumber="19">H. Fruhstorfer</collectorName>
leg.,
<collectionCode id="E0EFF8B4F742FFE3FB5BDB1CFD9E9C75" box="[1194,1251,294,320]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
</materialsCitation>
, no. 23142317, 1376313766;—
<materialsCitation id="36966A2CF742FFE3FEA8DB70FA179C51" box="[345,874,330,357]" collectingDate="1896-05" collectionCode="SMF" collectorName="H. Fruhstorfer" country="Indonesia" location="Ekas" pageId="18" pageNumber="19" specimenCount="4" specimenCount-male="4" stateProvince="Lombok Island">
<specimenCount id="90F8ABF8F742FFE3FEA8DB70F8ED9C51" box="[345,400,330,356]" count="4" pageId="18" pageNumber="19" type="male">4♂♂</specimenCount>
,
<date id="F24046B1F742FFE3FE6EDB70F8909C51" box="[415,493,330,356]" pageId="18" pageNumber="19" value="1896-05">
<collectingDate id="E204BF59F742FFE3FE6EDB70F8909C51" box="[415,493,330,356]" pageId="18" pageNumber="19" value="1896-05">v.1896</collectingDate>
</date>
,
<location id="832136AAF742FFE3FE0CDB70FB4B9C51" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF742FFE3FE0CDB70FB4B9C51" box="[509,566,330,356]" country="Indonesia" name="Ekas" pageId="18" pageNumber="19" stateProvince="Lombok Island">Ekas</location>
,
<collectorName id="2B0B05A7F742FFE3FDB7DB71FB979C51" box="[582,746,330,357]" pageId="18" pageNumber="19">H. Fruhstorfer</collectorName>
leg.,
<collectionCode id="E0EFF8B4F742FFE3FCC0DB70FA179C51" box="[817,874,330,356]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
</materialsCitation>
, no. 1376113762, 1376813769;—
<materialsCitation id="36966A2CF742FFE3FAEADB70F8DC9CBD" collectionCode="SMF" country="Indonesia" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Lombok Island">
<specimenCount id="90F8ABF8F742FFE3FAEADB70FC4A9C51" box="[1307,1335,330,356]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
, without particular locality,
<collectionCode id="E0EFF8B4F742FFE3FE99DB54F8DC9CBD" box="[360,417,366,392]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
</materialsCitation>
, no. 13767;—
<materialsCitation id="36966A2CF742FFE3FDB7DB54FDCA9CBD" box="[582,1207,366,392]" collectingDate="1927-04-21" collectionCode="SMF" collectorName="Selong &amp; Sunda-Expedition Rensch" country="Indonesia" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Lombok Island">
<specimenCount id="90F8ABF8F742FFE3FDB7DB54FB199CBD" box="[582,612,366,392]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<date id="F24046B1F742FFE3FD9EDB54FB959CBD" box="[623,744,366,392]" pageId="18" pageNumber="19" value="1927-04-21">
<collectingDate id="E204BF59F742FFE3FD9EDB54FB959CBD" box="[623,744,366,392]" pageId="18" pageNumber="19" value="1927-04-21">21.iv.1927</collectingDate>
</date>
,
<collectorName id="2B0B05A7F742FFE3FD03DB54FA3E9CBD" box="[754,835,366,392]" pageId="18" pageNumber="19">Selong</collectorName>
,
<collectorName id="2B0B05A7F742FFE3FCBCDB54FD0E9CBD" box="[845,1139,366,392]" pageId="18" pageNumber="19">Sunda-Expedition Rensch</collectorName>
,
<collectionCode id="E0EFF8B4F742FFE3FB8FDB54FDCA9CBD" box="[1150,1207,366,392]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
</materialsCitation>
, no. 1Ris01;—
<materialsCitation id="36966A2CF742FFE3FAAEDB54FB869C98" collectingDate="1993-02-18" collectionCode="RMNH" collectorName="Lendang Bajur &amp; J. Belle" country="Indonesia" pageId="18" pageNumber="19" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Lombok Island">
<specimenCount id="90F8ABF8F742FFE3FAAEDB54FCE59CBD" box="[1375,1432,366,392]" count="7" pageId="18" pageNumber="19" type="male">7♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F742FFE3FF66DBA8F9AC9C99" box="[151,209,402,428]" count="2" pageId="18" pageNumber="19" type="female">2♀♀</specimenCount>
,
<date id="F24046B1F742FFE3FF2CDBA8F82C9C99" box="[221,337,402,428]" pageId="18" pageNumber="19" value="1993-02-18">
<collectingDate id="E204BF59F742FFE3FF2CDBA8F82C9C99" box="[221,337,402,428]" pageId="18" pageNumber="19" value="1993-02-18">18.ii.1993</collectingDate>
</date>
,
<collectorName id="2B0B05A7F742FFE3FEACDBA8FB7B9C99" box="[349,518,402,428]" pageId="18" pageNumber="19">Lendang Bajur</collectorName>
,
<collectorName id="2B0B05A7F742FFE3FDE0DBA9FB189C99" box="[529,613,402,429]" pageId="18" pageNumber="19">J. Belle</collectorName>
leg.,
<collectionCode id="E0EFF8B4F742FFE3FD52DBA9FB869C98" box="[675,763,403,429]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="19" type="Museum">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F742FFE3FF36DB8CFD3C9CE4" blockId="18.[151,1437,150,2013]" box="[199,1089,438,465]" pageId="18" pageNumber="19">
<materialsCitation id="36966A2CF742FFE3FF36DB8CFD409CE4" box="[199,1085,438,465]" collectingDate="2015-12" collectionCode="MWNH" country="Indonesia" location="Island" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2">
<emphasis id="B48ABC63F742FFE3FF36DB8CF88D9CE5" bold="true" box="[199,496,438,464]" pageId="18" pageNumber="19">
<collectingCountry id="FEE920E1F742FFE3FF36DB8CF8409CE5" box="[199,317,438,464]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
,
<taxonomicName id="41FE1BF2F742FFE3FEBBDB8CF8E39CE5" box="[330,414,438,464]" class="Insecta" family="Acrididae" genus="Sumba" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="54" phylum="Arthropoda" rank="genus">Sumba</taxonomicName>
<location id="832136AAF742FFE3FE54DB8CF88D9CE5" LSID="urn:lsid:plazi:treatment:0E57D167F741FFE2FF66D87CFB739C98:832136AAF742FFE3FE54DB8CF88D9CE5" box="[421,496,438,464]" country="Indonesia" name="Island" pageId="18" pageNumber="19">Island</location>
</emphasis>
:
<specimenCount id="90F8ABF8F742FFE3FE0FDB8CFB449CE5" box="[510,569,438,464]" count="2" pageId="18" pageNumber="19" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F742FFE3FDB5DB8CFBD99CE4" box="[580,676,438,465]" pageId="18" pageNumber="19" value="2015-12">
<collectingDate id="E204BF59F742FFE3FDB5DB8CFBD99CE4" box="[580,676,438,465]" pageId="18" pageNumber="19" value="2015-12">xii.2015</collectingDate>
</date>
, without particular locality,
<collectionCode id="E0EFF8B4F742FFE3FC2FDB8DFD409CE4" box="[990,1085,439,465]" country="Germany" httpUri="http://grbio.org/cool/zw2m-hsji" name="Museum Wiesbaden, Department of Natural Science" pageId="18" pageNumber="19">MWNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F742FFE3FF36DBE0FC5B9CC1" blockId="18.[151,1437,150,2013]" box="[199,1318,474,501]" pageId="18" pageNumber="19">
<materialsCitation id="36966A2CF742FFE3FF36DBE0FC5F9CC1" box="[199,1314,474,501]" collectingDate="1927-05-29" collectionCode="SMF" country="Indonesia" location="Sumbawa Island" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Sumbawa Island">
<emphasis id="B48ABC63F742FFE3FF36DBE0FB6F9CC1" bold="true" box="[199,530,474,500]" pageId="18" pageNumber="19">
<collectingCountry id="FEE920E1F742FFE3FF36DBE0F8409CC1" box="[199,317,474,500]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F742FFE3FEBBDBE0FB6F9CC1" box="[330,530,474,500]" pageId="18" pageNumber="19">Sumbawa Island</collectingRegion>
</emphasis>
:
<specimenCount id="90F8ABF8F742FFE3FDD0DBE0FB389CC1" box="[545,581,474,500]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<date id="F24046B1F742FFE3FDA3DBE0FBBF9CC1" box="[594,706,474,500]" pageId="18" pageNumber="19" value="1927-05-29">
<collectingDate id="E204BF59F742FFE3FDA3DBE0FBBF9CC1" box="[594,706,474,500]" pageId="18" pageNumber="19" value="1927-05-29">29.v.1927</collectingDate>
</date>
, Dompu, Sunda-Expedition Rensch,
<collectionCode id="E0EFF8B4F742FFE3FB90DBE0FDE79CC1" box="[1121,1178,474,500]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
, no. 1Ris03
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F742FFE3FF36DBC4FB089F09" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<materialsCitation id="36966A2CF742FFE3FF36DBC4FD8D9F2C" box="[199,1264,510,537]" collectionCode="NHMUK" collectorName="August" country="Indonesia" county="Island" location="Island" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Timur">
<emphasis id="B48ABC63F742FFE3FF36DBC4F8A69F2D" bold="true" box="[199,475,510,536]" pageId="18" pageNumber="19">
<collectingCountry id="FEE920E1F742FFE3FF36DBC4F8409F2D" box="[199,317,510,536]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F742FFE3FEBCDBC4F8F89F2D" box="[333,389,510,536]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="19">Java</collectingRegion>
<collectingCounty id="6F2018FDF742FFE3FE61DBC4F8A69F2D" box="[400,475,510,536]" pageId="18" pageNumber="19">Island</collectingCounty>
</emphasis>
:
<specimenCount id="90F8ABF8F742FFE3FE1DDBC4FB6C9F2D" box="[492,529,510,536]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectorName id="2B0B05A7F742FFE3FDEEDBC4FB0E9F2D" box="[543,627,510,536]" pageId="18" pageNumber="19">August</collectorName>
, without particular locality,
<collectionCode id="E0EFF8B4F742FFE3FC4BDBC5FD5A9F2C" box="[954,1063,511,537]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="19" type="Museum">NHMUK</collectionCode>
, no. 012495866
</materialsCitation>
;—
<materialsCitation id="36966A2CF742FFE3FAE2DBC4FB0C9F09" collectingDate="1918-07" collectionCode="SMF" collectorName="Bagor &amp; Dr W. Ropke" country="Indonesia" county="Island" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur">
<specimenCount id="90F8ABF8F742FFE3FAE2DBC4FC579F2C" box="[1299,1322,510,537]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
,
<date id="F24046B1F742FFE3FAC8DBC4FCEA9F2D" box="[1337,1431,510,536]" pageId="18" pageNumber="19" value="1918-07">
<collectingDate id="E204BF59F742FFE3FAC8DBC4FCEA9F2D" box="[1337,1431,510,536]" pageId="18" pageNumber="19" value="1918-07">vii.1918</collectingDate>
</date>
,
<collectorName id="2B0B05A7F742FFE3FF66D819F9A09F08" box="[151,221,547,573]" pageId="18" pageNumber="19">Bagor</collectorName>
,
<collectorName id="2B0B05A7F742FFE3FF19D819F8069F09" box="[232,379,546,573]" pageId="18" pageNumber="19">Dr W. Röpke</collectorName>
leg.,
<collectionCode id="E0EFF8B4F742FFE3FE4BD818F88E9F09" box="[442,499,546,572]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="19">SMF</collectionCode>
, no. 13947
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F742FFE3FF36D87CF88F9F55" blockId="18.[151,1437,150,2013]" box="[199,498,582,608]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36D87CF88F9F55" bold="true" box="[199,498,582,608]" pageId="18" pageNumber="19">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F742FFE3FF36D850FDCC9FC5" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36D850F8799FB1" box="[199,260,618,644]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
(
<figureCitation id="1EC57CF4F742FFE3FEE0D850F81A9FB1" box="[273,359,618,644]" captionStart="FIGURES 2628" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[152,1432,1242,1879]" captionTargetId="figure-369@17.[151,1436,1241,1882]" captionTargetPageId="17" captionText="FIGURES 2628. Pseudagrion pilidorsum declaratum, syntype male: (26) habitus dorsal, (27) labels, (28) habitus lateral. Photos: M. Seehausen." pageId="18" pageNumber="19">Figs 26</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FE83D850F8ED9FB1" box="[370,400,618,644]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="18" pageNumber="19">83</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FE6BD850F8B59FB0" box="[410,456,618,645]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="18" pageNumber="19">155</figureCitation>
): Labrum, clypeus, antefrons and postfrons yellow-orange; colour of postfrons distinctly separated from red vertex around level of lateral ocelli; postocular spots red, separated by a very thin black line from postfrons; rear of head black, but broadly yellow-orange along the eye margins; base and scape of antennae yellow, following segments brownish; eyes when alive reddish, anteriorly washed-out yellow-orange.
</paragraph>
<paragraph id="86416071F742FFE3FF36D8C0FDB59EFD" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36D8C0F8659E21" box="[199,280,762,788]" italics="true" pageId="18" pageNumber="19">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F742FFE3FED6D8C0F8D59E21" box="[295,424,762,788]" captionStart="FIGURES 2628" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[152,1432,1242,1879]" captionTargetId="figure-369@17.[151,1436,1241,1882]" captionTargetPageId="17" captionText="FIGURES 2628. Pseudagrion pilidorsum declaratum, syntype male: (26) habitus dorsal, (27) labels, (28) habitus lateral. Photos: M. Seehausen." pageId="18" pageNumber="19">Figs 26, 28</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FE42D8C0F8A29E21" box="[435,479,762,788]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="18" pageNumber="19">100</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FE1AD8C0FB649E20" box="[491,537,762,789]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="18" pageNumber="19">155</figureCitation>
): Anterior, middle, and posterior lobe of pronotum dorsally red; middle pronotal lobe with a black patch on each side dorsolaterally, along the margin between anterior and middle lobe; sutures of middle lobe black, posterior margin of middle lobe broadly black; apical edge of posterior lobe convex. Pterothorax red with a thin black middorsal carina; a black stripe at posterior quarter of humeral suture and metapleural suture, both expanded in the middle; ventrally pale, may be covered with whitish pruinosity. Legs pale reddish; femora may have a blackish stripe outside, narrowing proximally; tibiae blackish along spines; spines black.
</paragraph>
<paragraph id="86416071F742FFE3FF36D9E8F9A2996D" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36D9E8F84C9ED9" box="[199,305,978,1004]" italics="true" pageId="18" pageNumber="19">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F742FFE3FEB3D9E8F8BA9ED9" box="[322,455,978,1004]" captionStart="FIGURES 2628" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[152,1432,1242,1879]" captionTargetId="figure-369@17.[151,1436,1241,1882]" captionTargetPageId="17" captionText="FIGURES 2628. Pseudagrion pilidorsum declaratum, syntype male: (26) habitus dorsal, (27) labels, (28) habitus lateral. Photos: M. Seehausen." pageId="18" pageNumber="19">Figs 26, 28</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FE24D9E8FB579ED9" box="[469,554,978,1004]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="18" pageNumber="19">117a, b</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FDC9D9E8FB1B9ED8" box="[568,614,978,1005]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="18" pageNumber="19">155</figureCitation>
): S1 red; S2 red, often with a blackish marking at the posterior quarter, which may be connected by a blackish middorsal stripe with black posterior margin; S38 dorsally black, laterally pale whitish-yellow; S9 red with posterior edge black; S10 red with black dorsal “x”-like marking, variable in shape.
</paragraph>
<paragraph id="86416071F742FFE3FF36DE58F8BB9861" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DE58F8D89949" box="[199,421,1122,1148]" italics="true" pageId="18" pageNumber="19">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F742FFE3FE43DE58FB429949" box="[434,575,1122,1148]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="18" pageNumber="19">Figs 134ac</figureCitation>
): Cercus laterally washed-out darkish red, its edges black; slightly shorter than S10; in lateral view cercus subrectangular with open inter-lobe notch low on apex; in dorsal view cercus appearing narrow, apex of the superior lobe narrowly rounded and hooked inwards; a blunt interior projection (appearing subtriangular in dorsolateral view) is present at ca. the middle of the superior lobe; a tiny subbasal interior tooth-like projection on the inferior lobe at ca. the basal third; paraproct pale yellow-ochre, about half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, with black tooth at medial edge.
</paragraph>
<paragraph id="86416071F742FFE3FF36DF64FB7A984D" blockId="18.[151,1437,150,2013]" box="[199,519,1374,1400]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DF64FB7A984D" bold="true" box="[199,519,1374,1400]" pageId="18" pageNumber="19">Characterization of female</emphasis>
</paragraph>
<paragraph id="86416071F742FFE3FF36DFB8FA7698A9" blockId="18.[151,1437,150,2013]" box="[199,779,1410,1437]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DFB8F87998A9" box="[199,260,1410,1436]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
: As in male but red replaced by orange-ochre.
</paragraph>
<paragraph id="86416071F742FFE3FF36DF9CF8DE9B3D" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DF9CF86598F5" box="[199,280,1446,1472]" italics="true" pageId="18" pageNumber="19">Thorax</emphasis>
: Pronotum and pterothorax as in male but red replaced by orange-ochre; posterior lobe of pronotum with two horns directed towards the head, almost reaching the centre of the middle lobe; pterothorax may be tinted greenish antehumerally.
</paragraph>
<paragraph id="86416071F742FFE3FF36DC28FC119BAD" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DC28F84C9B19" box="[199,305,1554,1580]" italics="true" pageId="18" pageNumber="19">Abdomen</emphasis>
: S1 orange-ochre; S2 orange-ochre, with black marking middorsally; S38 as in male; S910 laterally orange-ochre, dorsally with black markings variable in shape. Cercus washed-out dark brownish, as long as half of S10; paraproct brownish, in lateral view rounded; ovipositor pale yellowish, its apex not reaching to posterior end of S10; two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F742FFE3FF36DC98FD289B89" blockId="18.[151,1437,150,2013]" box="[199,1109,1698,1725]" pageId="18" pageNumber="19">Wings of both sexes with 11.513.5 px in forewing, and 1012.5 px in hindwing.</paragraph>
<paragraph id="86416071F742FFE3FF36DCFDFC089A31" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="19">Measurements of both sexes (mm): Total length (with appendages) 38.547.0; abdomen length (without appendages) 30.437.9; hindwing 20.625.0; forewing 22.026.5; Pt in forewing 0.70.9; male cercus 0.60.8.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF742FFE2FF36DD34FA569C74" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="86416071F742FFE3FF36DD34F8479A1D" blockId="18.[151,1437,150,2013]" box="[199,314,1806,1832]" pageId="18" pageNumber="19">
<emphasis id="B48ABC63F742FFE3FF36DD34F8479A1D" bold="true" box="[199,314,1806,1832]" pageId="18" pageNumber="19">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F742FFE2FF36DD08FA569C74" blockId="18.[151,1437,150,2013]" lastBlockId="19.[151,1437,150,429]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
This species differs from almost all other species of the
<taxonomicName id="41FE1BF2F742FFE3FCC7DD08FAB89A79" authorityName="Selys" authorityYear="1876" box="[822,965,1842,1868]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B48ABC63F742FFE3FCC7DD08FAB89A79" box="[822,965,1842,1868]" italics="true" pageId="18" pageNumber="19">Pseudagrion</emphasis>
</taxonomicName>
red-group by the largely yellow-red head, bearing just a very thin black line around large red postocular spots (
<figureCitation id="1EC57CF4F742FFE3FC62DD6CFA949A45" box="[915,1001,1878,1904]" captionStart="FIGURES 2628" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[152,1432,1242,1879]" captionTargetId="figure-369@17.[151,1436,1241,1882]" captionTargetPageId="17" captionText="FIGURES 2628. Pseudagrion pilidorsum declaratum, syntype male: (26) habitus dorsal, (27) labels, (28) habitus lateral. Photos: M. Seehausen." pageId="18" pageNumber="19">Figs 26</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FC04DD6CFD6E9A45" box="[1013,1043,1878,1904]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="18" pageNumber="19">83</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FBEFDD6CFD319A44" box="[1054,1100,1878,1905]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="18" pageNumber="19">155</figureCitation>
). This colour pattern is shared only by
<taxonomicName id="41FE1BF2F742FFE3FF03DD41F80C9AA1" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[242,369,1914,1940]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F742FFE3FF03DD41F80C9AA1" box="[242,369,1914,1940]" italics="true" pageId="18" pageNumber="19">P. deflexum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F742FFE3FE71DD40F8AB9AA1" box="[384,470,1914,1940]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="18" pageNumber="19">Figs 29</figureCitation>
,
<figureCitation id="1EC57CF4F742FFE3FE10DD40FB7D9AA1" box="[481,512,1914,1940]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="18" pageNumber="19">84</figureCitation>
), but living males of this species have the anterior half of the eyes distinctly yellow (
<figureCitation id="1EC57CF4F742FFE3FF6EDDA4F83F9A8C" box="[159,322,1950,1977]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="18" pageNumber="19">Figs 157, 158</figureCitation>
). The laterally subrectangular male appendages of
<taxonomicName id="41FE1BF2F742FFE3FC79DDA5FD5B9A8D" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[904,1062,1950,1976]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="declaratum">
<emphasis id="B48ABC63F742FFE3FC79DDA5FD5B9A8D" box="[904,1062,1950,1976]" italics="true" pageId="18" pageNumber="19">P. declaratum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F742FFE3FBC7DDA4FDD89A8D" box="[1078,1189,1950,1976]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="18" pageNumber="19">Fig. 134a</figureCitation>
) separate this species from several congeners as well, especially from the similarly coloured
<taxonomicName id="41FE1BF2F742FFE3FC5FDDF9FD539AE9" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[942,1070,1986,2012]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F742FFE3FC5FDDF9FD539AE9" box="[942,1070,1986,2012]" italics="true" pageId="18" pageNumber="19">P. deflexum</emphasis>
</taxonomicName>
, in which the cercus is distinctly arched in lateral view (
<figureCitation id="1EC57CF4F743FFE2FE62DAADFB7E9D84" box="[403,515,150,177]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="19" pageNumber="20">Figs 135a</figureCitation>
,
<figureCitation id="1EC57CF4F743FFE2FDFFDAACFB419D84" box="[526,572,150,177]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="19" pageNumber="20">159</figureCitation>
). The male caudal appendages of
<taxonomicName id="41FE1BF2F743FFE2FC41DAADFD459D84" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[944,1080,151,177]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F743FFE2FC41DAADFD459D84" box="[944,1080,151,177]" italics="true" pageId="19" pageNumber="20">P. obscurum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F743FFE2FB9CDAADFD829D84" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[1133,1279,151,177]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F743FFE2FB9CDAADFD829D84" box="[1133,1279,151,177]" italics="true" pageId="19" pageNumber="20">P. simalurum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F743FFE2FAFCDAADF9949DE0" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="19" pageNumber="20">Figs 140ac, 141ac</figureCitation>
) as well as of
<taxonomicName id="41FE1BF2F743FFE2FE78DA81F88B9DE1" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[393,502,187,212]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F743FFE2FE78DA81F88B9DE1" box="[393,502,187,212]" italics="true" pageId="19" pageNumber="20">P. igneum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F743FFE2FDDDDA81FB9A9DE0" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[556,743,187,213]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F743FFE2FDDDDA81FB9A9DE0" box="[556,743,187,213]" italics="true" pageId="19" pageNumber="20">P. nigrihumerale</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F743FFE2FD07DA81FAA29DE0" box="[758,991,187,213]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="19" pageNumber="20">Figs 138ac, 139ac</figureCitation>
) have superficial similarities with those of
<taxonomicName id="41FE1BF2F743FFE2FF44DAE5F82C9DCC" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[181,337,223,249]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="declaratum">
<emphasis id="B48ABC63F743FFE2FF44DAE5F82C9DCC" box="[181,337,223,249]" italics="true" pageId="19" pageNumber="20">P. declaratum</emphasis>
</taxonomicName>
, but these species (except for
<taxonomicName id="41FE1BF2F743FFE2FD53DAE5FA209DCC" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[674,861,223,249]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F743FFE2FD53DAE5FA209DCC" box="[674,861,223,249]" italics="true" pageId="19" pageNumber="20">P. nigrihumerale</emphasis>
</taxonomicName>
) are separated by having S12 dorsally black (
<figureCitation id="1EC57CF4F743FFE2FA9BDAE5F9939C28" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="19" pageNumber="20">Figs 121a, b</figureCitation>
,
<figureCitation id="1EC57CF4F743FFE2FF0DDB39F8C49C28" box="[252,441,259,285]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="19" pageNumber="20">123a, b, 124a, b</figureCitation>
), black colouration around the postocular spots (
<figureCitation id="1EC57CF4F743FFE2FC1ADB39FDB79C28" box="[1003,1226,259,285]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="19" pageNumber="20">Figs 87, 88, 89, 90</figureCitation>
), and regarding
<taxonomicName id="41FE1BF2F743FFE2FA79DB39F84A9C74" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F743FFE2FA79DB39F84A9C74" italics="true" pageId="19" pageNumber="20">P. nigrihumerale</emphasis>
</taxonomicName>
by having a black humeral stripe (
<figureCitation id="1EC57CF4F743FFE2FD4DDB1DFA639C74" box="[700,798,294,321]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="19" pageNumber="20">Fig. 105</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF743FFE2FF36DB70FB739C98" pageId="19" pageNumber="20" type="distribution">
<paragraph id="86416071F743FFE2FF36DB70F8249C51" blockId="19.[151,1437,150,429]" box="[199,345,330,356]" pageId="19" pageNumber="20">
<emphasis id="B48ABC63F743FFE2FF36DB70F8249C51" bold="true" box="[199,345,330,356]" pageId="19" pageNumber="20">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F743FFE2FF36DB55FB739C98" blockId="19.[151,1437,150,429]" pageId="19" pageNumber="20">
This species is distributed from
<collectingRegion id="443AAE93F743FFE2FDD6DB55FB249CBC" box="[551,601,367,393]" country="Indonesia" name="Jawa Timur" pageId="19" pageNumber="20">Java</collectingRegion>
towards the
<collectingRegion id="443AAE93F743FFE2FD15DB55FAB09CBC" box="[740,973,367,393]" country="Indonesia" name="Nusa Tenggara Timur" pageId="19" pageNumber="20">Lesser Sunda Islands</collectingRegion>
<collectingRegion id="443AAE93F743FFE2FC22DB55FD799CBC" box="[979,1028,367,393]" country="Indonesia" name="Bali" pageId="19" pageNumber="20">Bali</collectingRegion>
, Lombok, Sumbawa,
<collectingCountry id="FEE920E1F743FFE2FB04DB55FC439CBC" box="[1269,1342,367,393]" name="Indonesia" pageId="19" pageNumber="20">Flores</collectingCountry>
,
<taxonomicName id="41FE1BF2F743FFE2FAB6DB55F9A59C98" authority=", Sawu" authorityName="Sawu" class="Insecta" family="Acrididae" genus="Sumba" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Sumba, Sawu</taxonomicName>
, and reaching Alor (
<figureCitation id="1EC57CF4F743FFE2FE4ADBA9FB7C9C98" box="[443,513,403,429]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="19" pageNumber="20">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="93F71AB4A3B7F1FFC90F514D3915A963">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:affiliation id="03B6C79023D0062D7BCDA1F1B43C2CA6">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
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Figures: 2930 (
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); 156160 (field photos
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<emphasis id="B48ABC63F743FFE2FF66D8BAF8939FAD" box="[151,494,640,664]" italics="true" pageId="19" pageNumber="20">Pseudagrion pilidorsum deflexum</emphasis>
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appendages; key; Timor);—
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(Damar, Wetar);—
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preserved dry specimen);—
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Seehausen
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,
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,
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,
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(Timor; descriptions &amp; photos of habitats; photos
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alive; map);—
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da Silva Pinto
<emphasis id="B48ABC63F743FFE2FE41D8E7F8A09FC1" box="[432,477,732,756]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
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<emphasis id="B48ABC63F743FFE2FF66D918FB119E09" bold="true" box="[151,620,802,829]" pageId="19" pageNumber="20">Name-bearing type specimen examined</emphasis>
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<typeStatus id="5945DED3F743FFE2FF42D97DF8609E54" box="[179,285,839,865]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
,
<date id="F24046B1F743FFE2FED9D97DF8F89E54" box="[296,389,839,865]" pageId="19" pageNumber="20" value="1931-12">
<collectingDate id="E204BF59F743FFE2FED9D97DF8F89E54" box="[296,389,839,865]" pageId="19" pageNumber="20" value="1931-12">xii.1931</collectingDate>
</date>
<date id="F24046B1F743FFE2FE64D97DF8A19E54" box="[405,476,839,865]" pageId="19" pageNumber="20" value="1932-01">
<collectingDate id="E204BF59F743FFE2FE64D97DF8A19E54" box="[405,476,839,865]" pageId="19" pageNumber="20" value="1932-01">i.1932</collectingDate>
</date>
, Amarasi, Timor,
<collectingCountry id="FEE920E1F743FFE2FD55D97DFA699E54" box="[676,788,839,865]" name="Indonesia" pageId="19" pageNumber="20">Indonesia</collectingCountry>
,
<collectorName id="2B0B05A7F743FFE2FCEED97DFACB9E54" box="[799,950,839,865]" pageId="19" pageNumber="20">E. Handschin</collectorName>
leg.,
<collectionCode id="E0EFF8B4F743FFE2FC05D97DFD369E54" box="[1012,1099,839,865]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20" type="Museum">RMNH</collectionCode>
, JvT no. 2814 (
<figureCitation id="1EC57CF4F743FFE2FB06D97DFC069E54" box="[1271,1403,839,865]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="19" pageNumber="20">Figs 2930</figureCitation>
).
<typeStatus id="5945DED3F743FFE2FF36D951F84A9EB0" box="[199,311,875,901]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
designation by
<bibRefCitation id="E26F1D80F743FFE2FE18D951FBB29EB0" author="Lieftinck, M. A." box="[489,719,875,901]" pageId="19" pageNumber="20" pagination="65 - 139" refId="ref40982" refString="Lieftinck, M. A. (1971) A catalogue of the type-specimens of Odonata preserved in the Netherlands. With a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65-139, pls. 1 - 7." type="journal article" year="1971">Lieftinck (1971: 83)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="D28130F9F743FFE2FF66DCCAFD8B9A3D" box="[151,1270,1776,1801]" pageId="19" pageNumber="20" startId="19.[151,264,1776,1800]" targetBox="[153,1433,925,1748]" targetPageId="19" targetType="figure">
<paragraph id="86416071F743FFE2FF66DCCAFD8B9A3D" blockId="19.[151,1270,1776,1801]" box="[151,1270,1776,1801]" pageId="19" pageNumber="20">
<emphasis id="B48ABC63F743FFE2FF66DCCAF8309A3D" bold="true" box="[151,333,1776,1800]" pageId="19" pageNumber="20">FIGURES 2930</emphasis>
.
<taxonomicName id="41FE1BF2F743FFE2FEABDCCAFB409A3D" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[346,573,1776,1800]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F743FFE2FEABDCCAFB409A3D" box="[346,573,1776,1800]" italics="true" pageId="19" pageNumber="20">Pseudagrion deflexum</emphasis>
</taxonomicName>
, lectotype male: (
<emphasis id="B48ABC63F743FFE2FD05DCCAFA709A3D" bold="true" box="[756,781,1776,1800]" pageId="19" pageNumber="20">29</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F743FFE2FC31DCCAFAA49A3D" bold="true" box="[960,985,1776,1800]" pageId="19" pageNumber="20">30</emphasis>
) envelope. Photos: RMNH.
</paragraph>
</caption>
<subSubSection id="CEE433FAF743FFE5FF36DD60FC089B18" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="86416071F743FFE2FF36DD60FCE19AAC" blockId="19.[151,1436,1882,2017]" pageId="19" pageNumber="20">
<emphasis id="B48ABC63F743FFE2FF36DD60FB749A41" bold="true" box="[199,521,1882,1908]" pageId="19" pageNumber="20">
<collectingRegion id="443AAE93F743FFE2FF36DD60F8739A41" box="[199,270,1882,1908]" pageId="19" pageNumber="20">Other</collectingRegion>
specimens examined
</emphasis>
(
<specimenCount id="90F8ABF8F743FFE2FDE9DD61FB1C9A40" box="[536,609,1883,1909]" count="38" pageId="19" pageNumber="20" type="male">38♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F743FFE2FD9CDD61FBCF9A40" box="[621,690,1883,1909]" count="12" pageId="19" pageNumber="20" type="female">12♀♀</specimenCount>
)
<materialsCitation id="36966A2CF743FFE2FF36DD44F8A69AAD" box="[199,475,1918,1945]" country="Indonesia" location="West Timor" pageId="19" pageNumber="20" specimenCount="1" stateProvince="West Timor">
<emphasis id="B48ABC63F743FFE2FF36DD44F8A69AAD" bold="true" box="[199,475,1918,1945]" pageId="19" pageNumber="20">
<collectingCountry id="FEE920E1F743FFE2FF36DD44F8409AAD" box="[199,317,1918,1944]" name="Indonesia" pageId="19" pageNumber="20">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F743FFE2FEBADD45F8A69AAD" box="[331,475,1918,1945]" pageId="19" pageNumber="20">West Timor</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="36966A2CF743FFE2FE1ADD45FD069AAC" box="[491,1147,1918,1945]" collectionCode="IRSNB" collectorName="Kolbano &amp; Timor Central &amp; le Moult" country="Indonesia" pageId="19" pageNumber="20" specimenCount="2" specimenCount-male="2" stateProvince="West Timor">
<specimenCount id="90F8ABF8F743FFE2FE1ADD45FB5B9AAC" box="[491,550,1919,1945]" count="2" pageId="19" pageNumber="20" type="male">2♂♂</specimenCount>
,
<collectorName id="2B0B05A7F743FFE2FDC2DD44FBEB9AAD" box="[563,662,1918,1944]" pageId="19" pageNumber="20">Kolbano</collectorName>
,
<collectorName id="2B0B05A7F743FFE2FD53DD44FA389AAD" box="[674,837,1918,1944]" pageId="19" pageNumber="20">Timor Central</collectorName>
, ex-coll.
<collectorName id="2B0B05A7F743FFE2FC41DD44FD699AAD" box="[944,1044,1918,1944]" pageId="19" pageNumber="20">le Moult</collectorName>
,
<collectionCode id="E0EFF8B4F743FFE2FBD0DD45FD069AAC" box="[1057,1147,1919,1945]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="19" pageNumber="20">IRSNB</collectionCode>
</materialsCitation>
;—
<specimenCount id="90F8ABF8F743FFE2FB51DD45FDC09AAC" box="[1184,1213,1919,1945]" count="1" pageId="19" pageNumber="20" type="male">1♂</specimenCount>
,
<date id="F24046B1F743FFE2FB3BDD44FC549AAD" box="[1226,1321,1918,1944]" pageId="19" pageNumber="20" value="1897-07">vii.1897</date>
, Atapupu,
</paragraph>
<paragraph id="86416071F743FFE2FF66DD99FCE19A88" blockId="19.[151,1436,1882,2017]" box="[151,1436,1954,1981]" pageId="19" pageNumber="20">
<materialsCitation id="36966A2CF743FFE2FF66DD99FCF39A88" box="[151,1422,1954,1981]" collectionCode="SMF" collectorName="West Timor, A &amp; A. Everett" county="Central Timor" elevation="880" location="West Timor" municipality="Soe" pageId="19" pageNumber="20" specimenCount="8" specimenCount-female="1" specimenCount-male="7">
<collectorName id="2B0B05A7F743FFE2FF66DD99F8419A88" box="[151,316,1954,1981]" pageId="19" pageNumber="20">
<location id="832136AAF743FFE2FF66DD99F8619A89" LSID="urn:lsid:plazi:treatment:0E57D167F743FFE5FF66DBC4FC299A44:832136AAF743FFE2FF66DD99F8619A89" box="[151,284,1954,1981]" county="Central Timor" municipality="Soe" name="West Timor" pageId="19" pageNumber="20">West Timor</location>
, A
</collectorName>
. Everett leg.,
<collectionCode id="E0EFF8B4F743FFE2FE27DD99FB729A88" box="[470,527,1955,1981]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="20">SMF</collectionCode>
, no. 13770;—
<specimenCount id="90F8ABF8F743FFE2FD40DD99FB959A88" box="[689,744,1955,1981]" count="7" pageId="19" pageNumber="20" type="male">7♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F743FFE2FD02DD99FA6B9A88" box="[755,790,1955,1981]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
, 1929,
<collectingMunicipality id="6625FA0BF743FFE2FC9BDD99FAEA9A88" box="[874,919,1955,1981]" pageId="19" pageNumber="20">Soe</collectingMunicipality>
,
<collectingCounty id="6F2018FDF743FFE2FC53DD98FD3C9A89" box="[930,1089,1954,1980]" pageId="19" pageNumber="20">Central Timor</collectingCounty>
,
<elevation id="0DD38742F743FFE2FBBDDD99FDB29A89" box="[1100,1231,1954,1981]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="19" pageNumber="20" unit="m" value="880.0">
<quantity id="4106CD94F743FFE2FBBDDD99FDEE9A88" box="[1100,1171,1955,1981]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="19" pageNumber="20" unit="m" value="880.0">880 m</quantity>
a.s.l.
</elevation>
,
<collectionCode id="E0EFF8B4F743FFE2FB2DDD99FC689A88" box="[1244,1301,1955,1981]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="20">SMF</collectionCode>
, no. 30113
</materialsCitation>
</paragraph>
<paragraph id="86416071F743FFE2FF66DDFDFCE19AD5" blockId="19.[151,1436,1882,2017]" box="[151,1436,1990,2017]" pageId="19" pageNumber="20">
30119;—
<materialsCitation id="36966A2CF743FFE2FEF7DDFDFD6E9AD4" box="[262,1043,1990,2017]" collectingDate="1929-06" collectionCode="IRSNB" collectorName="Soe &amp; Timor Central &amp; le Moult" pageId="19" pageNumber="20" specimenCount="27" specimenCount-female="9" specimenCount-male="18">
<specimenCount id="90F8ABF8F743FFE2FEF7DDFDF8349AD4" box="[262,329,1991,2017]" count="18" pageId="19" pageNumber="20" type="male">18♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F743FFE2FEA4DDFDF8F39AD4" box="[341,398,1991,2017]" count="9" pageId="19" pageNumber="20" type="female">9♀♀</specimenCount>
,
<date id="F24046B1F743FFE2FE6DDDFCF88E9AD5" box="[412,499,1990,2016]" pageId="19" pageNumber="20" value="1929-06">
<collectingDate id="E204BF59F743FFE2FE6DDDFCF88E9AD5" box="[412,499,1990,2016]" pageId="19" pageNumber="20" value="1929-06">vi.1929</collectingDate>
</date>
,
<collectorName id="2B0B05A7F743FFE2FDF1DDFDFB509AD4" box="[512,557,1991,2017]" pageId="19" pageNumber="20">Soe</collectorName>
,
<collectorName id="2B0B05A7F743FFE2FDC8DDFCFBA69AD5" box="[569,731,1990,2016]" pageId="19" pageNumber="20">Timor Central</collectorName>
, ex-coll.
<collectorName id="2B0B05A7F743FFE2FCB4DDFCFAD59AD5" box="[837,936,1990,2016]" pageId="19" pageNumber="20">le Moult</collectorName>
,
<collectionCode id="E0EFF8B4F743FFE2FC44DDFDFD6E9AD4" box="[949,1043,1991,2017]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="19" pageNumber="20">IRSNB</collectionCode>
</materialsCitation>
;—
<specimenCount id="90F8ABF8F743FFE2FBC8DDFDFD2D9AD4" box="[1081,1104,1991,2017]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
,
<date id="F24046B1F743FFE2FBAFDDFCFDC89AD4" box="[1118,1205,1990,2017]" pageId="19" pageNumber="20" value="1935-06">vi.1935</date>
, Nikiniki, C. Bühler
</paragraph>
<paragraph id="86416071F744FFE5FF66DAADF9989DE0" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
&amp; Meyer, Basel Museum,
<materialsCitation id="36966A2CF744FFE5FE36DAADFDCC9D84" box="[455,1201,151,177]" collectingDate="1992-11" collectionCode="MCZ" collectorName="Coll. von Rosen" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1">
<collectionCode id="E0EFF8B4F744FFE5FE36DAADFB769D84" box="[455,523,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="21" type="Museum">MCZ</collectionCode>
;—
<specimenCount id="90F8ABF8F744FFE5FDC1DAADFB309D84" box="[560,589,151,177]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<date id="F24046B1F744FFE5FDAADAADFBCF9D84" box="[603,690,151,177]" pageId="20" pageNumber="21" value="1992-11">
<collectingDate id="E204BF59F744FFE5FDAADAADFBCF9D84" box="[603,690,151,177]" pageId="20" pageNumber="21" value="1992-11">xi.1992</collectingDate>
</date>
, without particular locality,
<collectorName id="2B0B05A7F744FFE5FC04DAADFDCC9D84" box="[1013,1201,151,177]" pageId="20" pageNumber="21">Coll. von Rosen</collectorName>
</materialsCitation>
,
<materialsCitation id="36966A2CF744FFE5FB4FDAADF99C9DE0" collectionCode="ZSM" collectorName="GvRosen" pageId="20" pageNumber="21" specimenCount="1">
<collectionCode id="E0EFF8B4F744FFE5FB4FDAADFD879D84" box="[1214,1274,151,177]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="20" pageNumber="21" type="Museum">ZSM</collectionCode>
, no.
<collectorName id="2B0B05A7F744FFE5FAC5DAADFCE19D84" box="[1332,1436,151,177]" pageId="20" pageNumber="21">GvRosen</collectorName>
13022
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F744FFE5FF36DAE4FCFF9C50" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<materialsCitation id="36966A2CF744FFE5FF36DAE4FD009DCC" box="[199,1149,222,249]" collectingDate="1935-08" collectionCode="MCZ" collectorName="C. Buhler &amp; Meyer &amp; Basel Museum" country="Timor-Leste" location="Baguia" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1">
<emphasis id="B48ABC63F744FFE5FF36DAE4F8269DCD" bold="true" box="[199,347,222,248]" pageId="20" pageNumber="21">
<collectingCountry id="FEE920E1F744FFE5FF36DAE4F8269DCD" box="[199,347,222,248]" name="Timor-Leste" pageId="20" pageNumber="21">Timor-Leste</collectingCountry>
</emphasis>
:
<specimenCount id="90F8ABF8F744FFE5FE9EDAE5F8EF9DCC" box="[367,402,223,249]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
,
<date id="F24046B1F744FFE5FE54DAE4FB709DCC" box="[421,525,222,249]" pageId="20" pageNumber="21" value="1935-08">
<collectingDate id="E204BF59F744FFE5FE54DAE4FB709DCC" box="[421,525,222,249]" pageId="20" pageNumber="21" value="1935-08">viii.1935</collectingDate>
</date>
,
<location id="832136AAF744FFE5FDECDAE5FB129DCC" LSID="urn:lsid:plazi:treatment:0E57D167F743FFE5FF66DBC4FC299A44:832136AAF744FFE5FDECDAE5FB129DCC" box="[541,623,223,249]" country="Timor-Leste" name="Baguia" pageId="20" pageNumber="21">Baguia</location>
,
<collectorName id="2B0B05A7F744FFE5FD71DAE5FB8F9DCC" box="[640,754,223,249]" pageId="20" pageNumber="21">C. Bühler</collectorName>
&amp;
<collectorName id="2B0B05A7F744FFE5FCD1DAE5FA169DCC" box="[800,875,223,249]" pageId="20" pageNumber="21">Meyer</collectorName>
,
<collectorName id="2B0B05A7F744FFE5FC8DDAE5FD569DCC" box="[892,1067,223,249]" pageId="20" pageNumber="21">Basel Museum</collectorName>
,
<collectionCode id="E0EFF8B4F744FFE5FBCADAE5FD009DCC" box="[1083,1149,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="21" type="Museum">MCZ</collectionCode>
</materialsCitation>
;—
<materialsCitation id="36966A2CF744FFE5FB51DAE5FA569C28" collectingDate="1973-10-19" collectionCode="MAGNT" collectorName="Tom Weir" country="Timor-Leste" location="Fatu Maca" pageId="20" pageNumber="21" specimenCount="4" specimenCount-male="4" stateProvince="Baucau">
<specimenCount id="90F8ABF8F744FFE5FB51DAE5FDAB9DCC" box="[1184,1238,223,249]" count="4" pageId="20" pageNumber="21" type="male">4♂♂</specimenCount>
,
<date id="F24046B1F744FFE5FB16DAE5FC249DCC" box="[1255,1369,223,249]" pageId="20" pageNumber="21" value="1973-10-19">
<collectingDate id="E204BF59F744FFE5FB16DAE5FC249DCC" box="[1255,1369,223,249]" pageId="20" pageNumber="21" value="1973-10-19">19.x.1973</collectingDate>
</date>
,
<location id="832136AAF744FFE5FA9BDAE5F9A59C28" LSID="urn:lsid:plazi:treatment:0E57D167F743FFE5FF66DBC4FC299A44:832136AAF744FFE5FA9BDAE5F9A59C28" country="Timor-Leste" name="Fatu Maca" pageId="20" pageNumber="21" stateProvince="Baucau">Fatu Maca</location>
,
<quantity id="4106CD94F744FFE5FF16DB39F84C9C28" box="[231,305,259,285]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="km" value="10.0">10 km</quantity>
south of
<collectingRegion id="443AAE93F744FFE5FE52DB39F8879C28" box="[419,506,259,285]" country="Timor-Leste" name="Baucau" pageId="20" pageNumber="21">Baucau</collectingRegion>
,
<collectorName id="2B0B05A7F744FFE5FDF9DB39FB079C28" box="[520,634,259,285]" pageId="20" pageNumber="21">Tom Weir</collectorName>
leg.,
<collectionCode id="E0EFF8B4F744FFE5FD4FDB39FA569C28" box="[702,811,259,285]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34986" lsid="urn:lsid:biocol.org:col:34986" name="Museum and Art Gallery of the Northern Territory" pageId="20" pageNumber="21" type="Museum">MAGNT</collectionCode>
</materialsCitation>
;—
<materialsCitation id="36966A2CF744FFE5FCBFDB39F8F89C74" collectingDate="2012-05-25" collectionCode="AMS" country="Timor-Leste" latitude="-8.999969" location="Same" longLatPrecision="1" longitude="125.64807" municipality="Kablaki Hotel" pageId="20" pageNumber="21" specimenCount="2" specimenCount-male="2">
<specimenCount id="90F8ABF8F744FFE5FCBFDB39FAFF9C28" box="[846,898,259,285]" count="2" pageId="20" pageNumber="21" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F744FFE5FC60DB38FD7C9C28" box="[913,1025,258,285]" pageId="20" pageNumber="21" value="2012-05-25">
<collectingDate id="E204BF59F744FFE5FC60DB38FD7C9C28" box="[913,1025,258,285]" pageId="20" pageNumber="21" value="2012-05-25">25.v.2012</collectingDate>
</date>
,
<collectingMunicipality id="6625FA0BF744FFE5FBE1DB39FDC99C28" box="[1040,1204,259,285]" pageId="20" pageNumber="21">Kablaki Hotel</collectingMunicipality>
,
<location id="832136AAF744FFE5FB33DB39FC7E9C28" LSID="urn:lsid:plazi:treatment:0E57D167F743FFE5FF66DBC4FC299A44:832136AAF744FFE5FB33DB39FC7E9C28" box="[1218,1283,259,285]" country="Timor-Leste" latitude="-8.999969" longLatPrecision="1" longitude="125.64807" municipality="Kablaki Hotel" name="Same" pageId="20" pageNumber="21">Same</location>
,
<geoCoordinate id="E3CA06B6F744FFE5FAE0DB39FCE59C28" box="[1297,1432,259,285]" degrees="8.999969" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-8.999969">8.999969°S</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F744FFE5FF66DB1CF8469C74" box="[151,315,294,321]" degrees="125.648072" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="125.64807">125.648072°E</geoCoordinate>
,
<collectionCode id="E0EFF8B4F744FFE5FEB7DB1DF8F89C74" box="[326,389,295,321]" pageId="20" pageNumber="21">AMS</collectionCode>
</materialsCitation>
, no. K462953;—
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,
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<collectingDate id="E204BF59F744FFE5FD8ADB1DFB9A9C74" box="[635,743,295,321]" pageId="20" pageNumber="21" value="2012-06-03">3.vi.2012</collectingDate>
</date>
,
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NE Laclubar
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,
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, no. K376343;—
<materialsCitation id="36966A2CF744FFE5FEA2DB71FDA19C50" box="[339,1244,330,357]" collectingDate="2012-06-05" collectionCode="AMS" country="Timor-Leste" latitude="8.50847" location="Manatuto" longLatPrecision="1" longitude="125.98292" pageId="20" pageNumber="21" specimenCount="3" specimenCount-male="3" stateProvince="Manatuto">
<specimenCount id="90F8ABF8F744FFE5FEA2DB71F8FA9C50" box="[339,391,331,357]" count="3" pageId="20" pageNumber="21" type="male">3♂♂</specimenCount>
,
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, spring,
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WNW
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,
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,
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, no. K376350.
</paragraph>
<paragraph id="86416071F744FFE5FF36DB54F88F9CBC" blockId="20.[151,1437,151,1905]" box="[199,498,366,393]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DB54F88F9CBC" bold="true" box="[199,498,366,393]" pageId="20" pageNumber="21">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F744FFE5FF36DBA9F8519F08" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DBA9F8799C98" box="[199,260,403,429]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
(
<figureCitation id="1EC57CF4F744FFE5FEE7DBA9F80D9C98" box="[278,368,403,429]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="20" pageNumber="21">Figs 29</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FE8EDBA9F8E09C98" box="[383,413,403,429]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="20" pageNumber="21">84</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FE5DDBA8FB6B9C98" box="[428,534,402,429]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">157, 158</figureCitation>
): Labrum, clypeus, antefrons and postfrons bright yellow; colour of postfrons sharply separated from red vertex at level of lateral ocelli; postocular spots red, separated by a very thin black line from postfrons; rear of head black, but broadly yellow-orange along the eye margins; base and scape of antennae yellow, following segments brownish; eyes when alive red with anterior half yellow, continuing in line with the yellow frons.
</paragraph>
<paragraph id="86416071F744FFE5FF36D87DFC7A9E0C" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36D87DF8659F54" box="[199,280,583,609]" italics="true" pageId="20" pageNumber="21">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F744FFE5FED8D87DF8FC9F54" box="[297,385,583,609]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="20" pageNumber="21">Figs 29</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FE7ED87DF8C69F54" box="[399,443,583,609]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="20" pageNumber="21">101</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FE38D87CFB119F54" box="[457,620,582,609]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">156, 157, 160</figureCitation>
): Anterior, middle, and posterior lobe of pronotum dorsally red; middle pronotal lobe with a black patch on each side dorsolaterally, along the margin between anterior and middle lobe; sutures of middle lobe black, posterior margin of middle lobe broadly black; apical edge of posterior lobe convex. Pterothorax red, with thin black middorsal carina; black stripes at posterior quarter of humeral suture and metapleural suture, both expanded in the middle; ventrally pale, may be covered with pruinosity; when immature basic colouration yellowish instead of red, with broadly brownish dorsum and humeral stripe (
<figureCitation id="1EC57CF4F744FFE5FB45D8C1FC6B9E20" box="[1204,1302,763,789]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">Fig. 160</figureCitation>
). Legs pale; femora with blackish stripe outside, narrowing proximally; tibiae blackish along spines; spines black.
</paragraph>
<paragraph id="86416071F744FFE5FF36D979FDAA9E90" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36D979F84C9E68" box="[199,305,835,861]" italics="true" pageId="20" pageNumber="21">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F744FFE5FEB2D979F8E09E68" box="[323,413,835,861]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="20" pageNumber="21">Figs 29</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FE5DD979FB7F9E68" box="[428,514,835,861]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="20" pageNumber="21">118a, b</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FDE0D978FB019E68" box="[529,636,834,861]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">156, 160</figureCitation>
): S1 red; S2 red, with black marking dorsally along posterior carina; S38 dorsally black, laterally whitish-yellow; S9 red with posterior edge black; S10 red, with black dorsal “x”-like marking, variable in shape; when immature basic colouration yellowish instead of red (
<figureCitation id="1EC57CF4F744FFE5FB99D9B1FDB79E90" box="[1128,1226,907,933]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">Fig. 160</figureCitation>
).
</paragraph>
<paragraph id="86416071F744FFE5FF36D995FC099948" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36D995F8D99EFC" box="[199,420,943,969]" italics="true" pageId="20" pageNumber="21">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F744FFE5FE40D995FB479EFC" box="[433,570,942,969]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="20" pageNumber="21">Figs 135ac</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FDB5D994FB0F9EFC" box="[580,626,942,969]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">159</figureCitation>
): Cercus laterally blackish, slightly shorter than S10; in lateral view whole cercus distinctly arched, apex subrectangular with inter-lobe notch almost central; in dorsal view cercus appearing narrow, apex of the superior lobe narrowly rounded and turned inwards; a subtriangular interior projection present at ca. the centre of the superior lobe of the cercus; a tiny interior tooth-like projection before half-length on the inferior lobe of the cercus; paraproct pale yellow-ochre, slightly longer than half of the cercus, in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, medial edge without tooth.
</paragraph>
<paragraph id="86416071F744FFE5FF36DEBCFB7A9995" blockId="20.[151,1437,151,1905]" box="[199,519,1158,1184]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DEBCFB7A9995" bold="true" box="[199,519,1158,1184]" pageId="20" pageNumber="21">Characterization of female</emphasis>
</paragraph>
<paragraph id="86416071F744FFE5FF36DE91FA7699F0" blockId="20.[151,1437,151,1905]" box="[199,779,1195,1221]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DE91F87999F0" box="[199,260,1195,1221]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
: As in male but red replaced by orange-ochre.
</paragraph>
<paragraph id="86416071F744FFE5FF36DEF5F8BB9804" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DEF5F86599DC" box="[199,280,1231,1257]" italics="true" pageId="20" pageNumber="21">Thorax</emphasis>
: Pronotum and pterothorax as in male but red replaced by orange-ochre; posterior lobe of pronotum with two horns directed towards the head, almost reaching the centre of the middle lobe; antehumeral pale, tinted ochre-greenish when alive.
</paragraph>
<paragraph id="86416071F744FFE5FF36DF01FC1198F4" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DF01F84C9860" box="[199,305,1339,1365]" italics="true" pageId="20" pageNumber="21">Abdomen</emphasis>
: S1 orange-ochre; S2 orange-ochre, with black marking middorsally; S38 as in male; S910 laterally orange-ochre, dorsally with black markings variable in shape. Cercus washed-out dark brownish, as long as half of S10; paraproct brownish, in lateral view rounded; ovipositor pale yellowish, its apex not reaching to posterior end of S10; two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F744FFE5FF36DFF1FD4F98D0" blockId="20.[151,1437,151,1905]" box="[199,1074,1482,1509]" pageId="20" pageNumber="21">Wings of both sexes with 10.512.5 px in forewing, and 911 px in hindwing.</paragraph>
<paragraph id="86416071F744FFE5FF36DFD5FC089B18" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">Measurements of both sexes (mm): Total length (with appendages) 38.045.5; abdomen length (without appendages) 32.537.5; hindwing 20.823.5; forewing 22.025.7; Pt in forewing 0.61.0; male cercus 0.50.6.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF744FFE5FF36DC0CFB5A9A1C" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="86416071F744FFE5FF36DC0CF8479B65" blockId="20.[151,1437,151,1905]" box="[199,314,1590,1616]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DC0CF8479B65" bold="true" box="[199,314,1590,1616]" pageId="20" pageNumber="21">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F744FFE5FF36DC61FB5A9A1C" blockId="20.[151,1437,151,1905]" pageId="20" pageNumber="21">
The morphology of the male appendages of
<taxonomicName id="41FE1BF2F744FFE5FD32DC61FA389B40" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[707,837,1627,1653]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F744FFE5FD32DC61FA389B40" box="[707,837,1627,1653]" italics="true" pageId="20" pageNumber="21">P. deflexum</emphasis>
</taxonomicName>
is unmistakable: the cercus is arched in lateral view (
<figureCitation id="1EC57CF4F744FFE5FF6EDC45F86D9BAC" box="[159,272,1662,1689]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="20" pageNumber="21">Figs 135a</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FEEADC44F8359BAC" box="[283,328,1662,1689]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">159</figureCitation>
), a unique character compared to all congeners. Additionally
<taxonomicName id="41FE1BF2F744FFE5FC03DC45FD0F9BAC" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[1010,1138,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F744FFE5FC03DC45FD0F9BAC" box="[1010,1138,1663,1689]" italics="true" pageId="20" pageNumber="21">P. deflexum</emphasis>
</taxonomicName>
is distinctly different from all congeners, except for the adjacently distributed
<taxonomicName id="41FE1BF2F744FFE5FD2CDC99FA079B88" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[733,890,1699,1725]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="declaratum">
<emphasis id="B48ABC63F744FFE5FD2CDC99FA079B88" box="[733,890,1699,1725]" italics="true" pageId="20" pageNumber="21">P. declaratum</emphasis>
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, in having a largely yellow-red head with just a very thin black line around the large red postocular spots (
<figureCitation id="1EC57CF4F744FFE5FCDCDCFDFAF89BD4" box="[813,901,1735,1761]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="20" pageNumber="21">Figs 29</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FC63DCFDFACD9BD4" box="[914,944,1735,1761]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="20" pageNumber="21">84</figureCitation>
,
<figureCitation id="1EC57CF4F744FFE5FC4CDCFCFA969BD4" box="[957,1003,1734,1761]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">157</figureCitation>
). But the yellow colour of the frons is continuous in line to the anterior half of the eyes in live males of
<taxonomicName id="41FE1BF2F744FFE5FC82DCD1FA899A30" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[883,1012,1771,1797]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F744FFE5FC82DCD1FA899A30" box="[883,1012,1771,1797]" italics="true" pageId="20" pageNumber="21">P. deflexum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F744FFE5FBF2DCD1FDDC9A30" box="[1027,1185,1770,1797]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">Figs 157, 158</figureCitation>
), whereas the eyes are reddish in
<taxonomicName id="41FE1BF2F744FFE5FEFCDD35F8D49A1C" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[269,425,1807,1833]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="declaratum">
<emphasis id="B48ABC63F744FFE5FEFCDD35F8D49A1C" box="[269,425,1807,1833]" italics="true" pageId="20" pageNumber="21">P. declaratum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F744FFE5FE49DD35FB679A1C" box="[440,538,1806,1833]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="20" pageNumber="21">Fig. 155</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF744FFE5FF36DD08FC299A44" box="[199,1364,1842,1905]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="86416071F744FFE5FF36DD08F8249A79" blockId="20.[151,1437,151,1905]" box="[199,345,1842,1868]" pageId="20" pageNumber="21">
<emphasis id="B48ABC63F744FFE5FF36DD08F8249A79" bold="true" box="[199,345,1842,1868]" pageId="20" pageNumber="21">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F744FFE5FF36DD6DFC299A44" blockId="20.[151,1437,151,1905]" box="[199,1364,1879,1905]" pageId="20" pageNumber="21">
This species occurs on Timor, as well as on adjacent islands of Rote, Semau, Wetar, and Damar (
<figureCitation id="1EC57CF4F744FFE5FAF0DD6DFC3B9A44" box="[1281,1350,1879,1905]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="20" pageNumber="21">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,327 @@
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<mods:title id="F9A5E603136520117A891E761D2CD696">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<emphasis id="B48ABC63F745FFE4FF66DAADF8BA9D84" bold="true" box="[151,455,151,178]" italics="true" pageId="21" pageNumber="22">Pseudagrion enganoense</emphasis>
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Figures: 3132 (
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); 102 (pterothorax
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<emphasis id="B48ABC63F745FFE4FF66DB23FB769C04" box="[151,523,281,305]" italics="true" pageId="21" pageNumber="22">Pseudagrion pilidorsum enganoense</emphasis>
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:
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: 290
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291,
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(description
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; Engano).
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<emphasis id="B48ABC63F745FFE4FF66DB65FB119C4C" bold="true" box="[151,620,351,377]" pageId="21" pageNumber="22">Name-bearing type specimen examined</emphasis>
(
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)
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<materialsCitation id="36966A2CF745FFE4FF36DBBEFDE29CA8" box="[199,1183,387,414]" collectingDate="1936-05-30" collectionCode="RMNH" location="Buahbuah" municipality="Enggano Island" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<specimenCount id="90F8ABF8F745FFE4FF36DBBEF9A19CAB" box="[199,220,388,414]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
<typeStatus id="5945DED3F745FFE4FF12DBB9F8309CA8" box="[227,333,387,413]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
,
<date id="F24046B1F745FFE4FEA9DBB9F8B59CA8" box="[344,456,387,413]" pageId="21" pageNumber="22" value="1936-05-30">
<collectingDate id="E204BF59F745FFE4FEA9DBB9F8B59CA8" box="[344,456,387,413]" pageId="21" pageNumber="22" value="1936-05-30">30.v.1936</collectingDate>
</date>
,
<location id="832136AAF745FFE4FE25DBB9FB359CA8" LSID="urn:lsid:plazi:treatment:0E57D167F745FFE7FF66DAADFB6999D0:832136AAF745FFE4FE25DBB9FB359CA8" box="[468,584,387,413]" municipality="Enggano Island" name="Buahbuah" pageId="21" pageNumber="22">Buahbuah</location>
,
<collectingMunicipality id="6625FA0BF745FFE4FDA5DBBEFA789CA8" box="[596,773,387,414]" pageId="21" pageNumber="22">Enggano Island</collectingMunicipality>
,
<collectionCode id="E0EFF8B4F745FFE4FCE1DBBEFA1A9CAB" box="[784,871,388,414]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
, JvT no. 2816 (
<figureCitation id="1EC57CF4F745FFE4FBE5DBB9FDEA9CA8" box="[1044,1175,387,413]" captionStart="FIGURES 3132" captionStartId="21.[151,264,1458,1482]" captionTargetBox="[152,1433,510,1430]" captionTargetId="figure-103@21.[151,1436,507,1433]" captionTargetPageId="21" captionText="FIGURES 3132. Pseudagrion enganoense, lectotype male: (31) habitus lateral, (32) envelope. Photos: RMNH." pageId="21" pageNumber="22">Figs 3132</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F745FFE4FF36DB92FDDE9CD0" blockId="21.[151,1436,351,486]" pageId="21" pageNumber="22">
<typeStatus id="5945DED3F745FFE4FF36DB92F84A9CF7" box="[199,311,424,450]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
designation by
<bibRefCitation id="E26F1D80F745FFE4FE01DB9DFBA69CF4" author="Lieftinck, M. A." box="[496,731,423,449]" pageId="21" pageNumber="22" pagination="65 - 139" refId="ref40982" refString="Lieftinck, M. A. (1971) A catalogue of the type-specimens of Odonata preserved in the Netherlands. With a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65-139, pls. 1 - 7." type="journal article" year="1971">Lieftinck (1971: 86)</bibRefCitation>
.
<bibRefCitation id="E26F1D80F745FFE4FD19DB9DFADF9CF4" author="Lieftinck, M. A." box="[744,930,423,449]" pageId="21" pageNumber="22" pagination="279 - 304" refId="ref40875" refString="Lieftinck, M. A. (1948) The Odonta of Engano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279-304." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
spelled the island “Engano”, thats why the scientific name is written with a single “g”. However, today the island is spelled “Enggano”.
</paragraph>
</subSubSection>
<caption id="D28130F9F745FFE4FF66DF88FC6F98FF" box="[151,1298,1458,1483]" pageId="21" pageNumber="22" startId="21.[151,264,1458,1482]" targetBox="[152,1433,510,1430]" targetPageId="21" targetType="figure">
<paragraph id="86416071F745FFE4FF66DF88FC6F98FF" blockId="21.[151,1298,1458,1483]" box="[151,1298,1458,1483]" pageId="21" pageNumber="22">
<emphasis id="B48ABC63F745FFE4FF66DF88F83398FF" bold="true" box="[151,334,1458,1482]" pageId="21" pageNumber="22">FIGURES 3132</emphasis>
.
<taxonomicName id="41FE1BF2F745FFE4FEABDF89FB2798FF" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[346,602,1459,1483]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="enganoense">
<emphasis id="B48ABC63F745FFE4FEABDF89FB2798FF" box="[346,602,1459,1483]" italics="true" pageId="21" pageNumber="22">Pseudagrion enganoense</emphasis>
</taxonomicName>
, lectotype male: (
<emphasis id="B48ABC63F745FFE4FCE1DF88FA5498FF" bold="true" box="[784,809,1458,1482]" pageId="21" pageNumber="22">31</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F745FFE4FC2DDF88FA8898FF" bold="true" box="[988,1013,1458,1482]" pageId="21" pageNumber="22">32</emphasis>
) envelope. Photos: RMNH.
</paragraph>
</caption>
<subSubSection id="CEE433FAF745FFE7FF36DC26F8409E03" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="86416071F745FFE4FF36DC26FB279B02" blockId="21.[151,1437,1564,2023]" box="[199,602,1564,1591]" pageId="21" pageNumber="22">
<emphasis id="B48ABC63F745FFE4FF36DC26FB749B03" bold="true" box="[199,521,1564,1590]" pageId="21" pageNumber="22">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F745FFE4FDE9DC27FB2F9B02" box="[536,594,1565,1591]" count="2" pageId="21" pageNumber="22" type="male">2♂♂</specimenCount>
)
</paragraph>
<paragraph id="86416071F745FFE4FF36DC7AFDC09B69" blockId="21.[151,1437,1564,2023]" box="[199,1213,1600,1628]" pageId="21" pageNumber="22">
<materialsCitation id="36966A2CF745FFE4FF36DC7AFDC49B69" box="[199,1209,1600,1628]" collectingDate="1936-05-30" collectionCode="RMNH" country="Indonesia" location="Enggano Island" pageId="21" pageNumber="22" specimenCount="2" specimenCount-male="2" stateProvince="Enggano Island">
<emphasis id="B48ABC63F745FFE4FF36DC7AFB729B6E" bold="true" box="[199,527,1600,1627]" pageId="21" pageNumber="22">
<collectingCountry id="FEE920E1F745FFE4FF36DC7AF8409B6F" box="[199,317,1600,1626]" name="Indonesia" pageId="21" pageNumber="22">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F745FFE4FEBBDC7BFB779B6E" box="[330,522,1601,1627]" pageId="21" pageNumber="22">Enggano Island</collectingRegion>
:
</emphasis>
<specimenCount id="90F8ABF8F745FFE4FDE7DC7BFB2C9B6E" box="[534,593,1601,1627]" count="2" pageId="21" pageNumber="22" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F745FFE4FDADDC7BFBB19B6E" box="[604,716,1601,1627]" pageId="21" pageNumber="22" value="1936-05-30">
<collectingDate id="E204BF59F745FFE4FDADDC7BFBB19B6E" box="[604,716,1601,1627]" pageId="21" pageNumber="22" value="1936-05-30">30.v.1936</collectingDate>
</date>
, Buahbuah,
<collectionCode id="E0EFF8B4F745FFE4FCA6DC7BFAD39B6E" box="[855,942,1601,1627]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
, no. 1658925, 1658926
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F745FFE4FF36DC5EF88F9B4A" blockId="21.[151,1437,1564,2023]" box="[199,498,1636,1663]" pageId="21" pageNumber="22">
<emphasis id="B48ABC63F745FFE4FF36DC5EF88F9B4A" bold="true" box="[199,498,1636,1663]" pageId="21" pageNumber="22">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F745FFE4FF36DCB3FBB69A06" blockId="21.[151,1437,1564,2023]" pageId="21" pageNumber="22">
<emphasis id="B48ABC63F745FFE4FF36DCB3F8799B96" box="[199,260,1673,1699]" italics="true" pageId="21" pageNumber="22">Head</emphasis>
(
<figureCitation id="1EC57CF4F745FFE4FEE2DCB3F8169B96" box="[275,363,1673,1699]" captionStart="FIGURES 3132" captionStartId="21.[151,264,1458,1482]" captionTargetBox="[152,1433,510,1430]" captionTargetId="figure-103@21.[151,1436,507,1433]" captionTargetPageId="21" captionText="FIGURES 3132. Pseudagrion enganoense, lectotype male: (31) habitus lateral, (32) envelope. Photos: RMNH." pageId="21" pageNumber="22">Figs 31</figureCitation>
,
<figureCitation id="1EC57CF4F745FFE4FE89DCB3F8E59B91" box="[376,408,1673,1700]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="21" pageNumber="22">85</figureCitation>
): Labrum, clypeus, antefrons and postfrons yellow; colour of postfrons distinctly separated from vertex around the level of lateral ocelli; vertex red, with a small black spot posterior of median ocellus, and large black patches each along interior and posterior margin of lateral ocelli; postocular spots red, separated by black line from postfrons and vertex; rear of head black, but yellow-orange along the eye margins; base and scape of antennae yellow, following segments brownish.
</paragraph>
<paragraph id="86416071F745FFE7FF36DD07FAB29DE1" blockId="21.[151,1437,1564,2023]" lastBlockId="22.[151,1437,151,1253]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B48ABC63F745FFE4FF36DD07F8659A62" box="[199,280,1853,1879]" italics="true" pageId="21" pageNumber="22">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F745FFE4FEDBDD07F8F99A62" box="[298,388,1853,1879]" captionStart="FIGURES 3132" captionStartId="21.[151,264,1458,1482]" captionTargetBox="[152,1433,510,1430]" captionTargetId="figure-103@21.[151,1436,507,1433]" captionTargetPageId="21" captionText="FIGURES 3132. Pseudagrion enganoense, lectotype male: (31) habitus lateral, (32) envelope. Photos: RMNH." pageId="21" pageNumber="22">Figs 31</figureCitation>
,
<figureCitation id="1EC57CF4F745FFE4FE62DD07F8BF9A62" box="[403,450,1853,1879]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="21" pageNumber="22">102</figureCitation>
): Anterior lobe of pronotum yellowish, expanding in centre spot-like towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided yellowish patch in the middle, and on each side a yellowish spot dorsolaterally; propleuron black above, yellowish below and to rear; posterior lobe of pronotum yellowish; apical edge of posterior lobe convex. Pterothorax red, with thin black middorsal carina; humeral suture and metapleural suture with black streaks at posterior quarter; interpleural suture with a thin black line; ventrally pale, may be covered with whitish pruinosity. Legs pale orange-reddish; femora with blackish stripe outside, narrowing proximally; tibiae blackish along spines; spines black.
</paragraph>
<paragraph id="86416071F746FFE7FF36DAE4FD5F9C29" blockId="22.[151,1437,151,1253]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DAE4F84C9DCD" box="[199,305,222,248]" italics="true" pageId="22" pageNumber="23">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F746FFE7FEB1DAE4F8EA9DCD" box="[320,407,222,248]" captionStart="FIGURES 3132" captionStartId="21.[151,264,1458,1482]" captionTargetBox="[152,1433,510,1430]" captionTargetId="figure-103@21.[151,1436,507,1433]" captionTargetPageId="21" captionText="FIGURES 3132. Pseudagrion enganoense, lectotype male: (31) habitus lateral, (32) envelope. Photos: RMNH." pageId="22" pageNumber="23">Figs 31</figureCitation>
,
<figureCitation id="1EC57CF4F746FFE7FE55DAE4F8859DCD" box="[420,504,222,248]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="22" pageNumber="23">119a, b</figureCitation>
): S1 red, S2 red with its posterior margin black; S38 dorsally black, laterally pale orange-yellow; S9 black with anterior third to half red; S10 black, ventrally pale.
</paragraph>
<paragraph id="86416071F746FFE7FF36DB1CFD159CC6" blockId="22.[151,1437,151,1253]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DB1CF8D49C75" box="[199,425,294,320]" italics="true" pageId="22" pageNumber="23">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F746FFE7FE4ADB1CFB309C75" box="[443,589,294,320]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="22" pageNumber="23">Figs 136ac</figureCitation>
): Cercus laterally obscure reddish; slightly shorter than S10, inferior lobe slightly longer than superior; in lateral view subrectangular, apex with inter-lobe notch deep and almost central; in dorsal view cercus appearing narrow, with terminus of superior lobe slightly turned in, bifurcated interiorly (visible in dorsolateral view); a small almost centrally-placed interior tooth-like projection on the inferior lobe of the cercus; paraproct pale ochre; around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, with blackish tubercular medial edge.
</paragraph>
<paragraph id="86416071F746FFE7FF36DBC4FDDE9F09" blockId="22.[151,1437,151,1253]" pageId="22" pageNumber="23">Measurements (mm): Total length (with appendages) 42.344.5; abdomen length (without appendages) 35.7 36.7; hindwing 23.724.3; forewing 25.026.0; Pt in forewing 0.71.0; male cercus 0.50.6.</paragraph>
<paragraph id="86416071F746FFE7FF36D87FFDAB9F6A" blockId="22.[151,1437,151,1253]" box="[199,1238,581,608]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36D87FFB7A9F6A" bold="true" box="[199,519,581,607]" pageId="22" pageNumber="23">Characterization of female</emphasis>
[not examined, characterization adopted from
<bibRefCitation id="E26F1D80F746FFE7FBE5D87FFDB29F6A" author="Lieftinck, M. A." box="[1044,1231,581,607]" pageId="22" pageNumber="23" pagination="279 - 304" refId="ref40875" refString="Lieftinck, M. A. (1948) The Odonta of Engano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279-304." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
]
</paragraph>
<paragraph id="86416071F746FFE7FF36D853FA759FB6" blockId="22.[151,1437,151,1253]" box="[199,776,617,643]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36D853F8799FB6" box="[199,260,617,643]" italics="true" pageId="22" pageNumber="23">Head</emphasis>
: As in male but ochraceous instead of yellow.
</paragraph>
<paragraph id="86416071F746FFE7FF36D8B7FB439FFE" blockId="22.[151,1437,151,1253]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36D8B7F8659F92" box="[199,280,653,679]" italics="true" pageId="22" pageNumber="23">Thorax</emphasis>
: Pterothorax as in male but dorsally honey-yellowish, fading to cinnamon-buff laterally; markings at humeral and metapleural suture rusty.
</paragraph>
<paragraph id="86416071F746FFE7FF36D8EEFC309FDB" blockId="22.[151,1437,151,1253]" box="[199,1357,724,751]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36D8EEF84C9FDB" box="[199,305,724,750]" italics="true" pageId="22" pageNumber="23">Abdomen</emphasis>
: As in male but with S12 orange; S9 with a large yellow spot laterally; ovipositor yellowish.
</paragraph>
<paragraph id="86416071F746FFE7FF36D8C2F8409E03" blockId="22.[151,1437,151,1253]" pageId="22" pageNumber="23">
Wings of both sexes with 14.517 px in forewing, and 1316 px in hindwing [adopted from
<bibRefCitation id="E26F1D80F746FFE7FB2DD8C2FCEA9E27" author="Lieftinck, M. A." box="[1244,1431,760,786]" pageId="22" pageNumber="23" pagination="279 - 304" refId="ref40875" refString="Lieftinck, M. A. (1948) The Odonta of Engano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279-304." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
, and own data].
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE7FF36D97AFB9A99AB" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="86416071F746FFE7FF36D97AF8479E6F" blockId="22.[151,1437,151,1253]" box="[199,314,832,858]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36D97AF8479E6F" bold="true" box="[199,314,832,858]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F746FFE7FF36D95EFB9A99AB" blockId="22.[151,1437,151,1253]" pageId="22" pageNumber="23">
<taxonomicName id="41FE1BF2F746FFE7FF36D95EF89B9E4B" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[199,486,868,894]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="enganoense">
<emphasis id="B48ABC63F746FFE7FF36D95EF89B9E4B" box="[199,486,868,894]" italics="true" pageId="22" pageNumber="23">Pseudagrion enganoense</emphasis>
</taxonomicName>
belongs to the group of endemic taxa of the island chain west of
<collectingRegion id="443AAE93F746FFE7FAF0D95EFC1F9E4B" box="[1281,1378,868,894]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="23">Sumatra</collectingRegion>
. The morphology of the male cercus of
<taxonomicName id="41FE1BF2F746FFE7FDE0D9B3FBD29E97" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[529,687,905,930]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="enganoense">
<emphasis id="B48ABC63F746FFE7FDE0D9B3FBD29E97" box="[529,687,905,930]" italics="true" pageId="22" pageNumber="23">P. enganoense</emphasis>
</taxonomicName>
is distinct: its inferior lobe is slightly longer than superior one (
<figureCitation id="1EC57CF4F746FFE7FA9BD9B2F9919EF3" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="22" pageNumber="23">Figs 136a, b</figureCitation>
), and in dorsolateral view the inner margin of superior lobe of cercus lacks an inward projection proximal of its bifurcated terminus (
<figureCitation id="1EC57CF4F746FFE7FE37D9F5FB4B9EDC" box="[454,566,975,1001]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="22" pageNumber="23">Fig. 136c</figureCitation>
). Additionally both sexes of
<taxonomicName id="41FE1BF2F746FFE7FC8FD9EAFD629EDC" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[894,1055,976,1001]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="enganoense">
<emphasis id="B48ABC63F746FFE7FC8FD9EAFD629EDC" box="[894,1055,976,1001]" italics="true" pageId="22" pageNumber="23">P. enganoense</emphasis>
</taxonomicName>
can be separated from congeners on adjacent islands by having S12 red in the male (
<figureCitation id="1EC57CF4F746FFE7FD2AD9C9FA4C993B" box="[731,817,1011,1038]" captionStart="FIGURES 3132" captionStartId="21.[151,264,1458,1482]" captionTargetBox="[152,1433,510,1430]" captionTargetId="figure-103@21.[151,1436,507,1433]" captionTargetPageId="21" captionText="FIGURES 3132. Pseudagrion enganoense, lectotype male: (31) habitus lateral, (32) envelope. Photos: RMNH." pageId="22" pageNumber="23">Figs 31</figureCitation>
,
<figureCitation id="1EC57CF4F746FFE7FCCDD9CEFAF29938" box="[828,911,1011,1038]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="22" pageNumber="23">119a, b</figureCitation>
), orange in the female. This character is shared with
<taxonomicName id="41FE1BF2F746FFE7FF20DE22F8109904" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[209,365,1047,1073]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="declaratum">
<emphasis id="B48ABC63F746FFE7FF20DE22F8109904" box="[209,365,1047,1073]" italics="true" pageId="22" pageNumber="23">P. declaratum</emphasis>
</taxonomicName>
,
<taxonomicName id="41FE1BF2F746FFE7FE8DDE22F8809904" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[380,509,1047,1073]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F746FFE7FE8DDE22F8809904" box="[380,509,1047,1073]" italics="true" pageId="22" pageNumber="23">P. deflexum</emphasis>
</taxonomicName>
, and
<taxonomicName id="41FE1BF2F746FFE7FDCCDE22FB849904" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[573,761,1047,1073]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F746FFE7FDCCDE22FB849904" box="[573,761,1047,1073]" italics="true" pageId="22" pageNumber="23">P. nigrihumerale</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F746FFE7FCF8DE2DFD219904" box="[777,1116,1047,1073]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="22" pageNumber="23">Figs 117a, b, 118a, b, 122a, b</figureCitation>
). However
<taxonomicName id="41FE1BF2F746FFE7FB11DE22FCE19904" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[1248,1436,1047,1073]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F746FFE7FB11DE22FCE19904" box="[1248,1436,1047,1073]" italics="true" pageId="22" pageNumber="23">P. nigrihumerale</emphasis>
</taxonomicName>
can also be separated by its broadly black antehumeral stripe (
<figureCitation id="1EC57CF4F746FFE7FCBCDE01FAD39963" box="[845,942,1083,1110]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="22" pageNumber="23">Fig. 105</figureCitation>
), while
<taxonomicName id="41FE1BF2F746FFE7FBF6DE06FDDF9960" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[1031,1186,1083,1109]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="declaratum">
<emphasis id="B48ABC63F746FFE7FBF6DE06FDDF9960" box="[1031,1186,1083,1109]" italics="true" pageId="22" pageNumber="23">P. declaratum</emphasis>
</taxonomicName>
as well as
<taxonomicName id="41FE1BF2F746FFE7FAEDDE06FCE19960" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[1308,1436,1083,1109]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="deflexum">
<emphasis id="B48ABC63F746FFE7FAEDDE06FCE19960" box="[1308,1436,1083,1109]" italics="true" pageId="22" pageNumber="23">P. deflexum</emphasis>
</taxonomicName>
have much larger red postocular spots than
<taxonomicName id="41FE1BF2F746FFE7FD75DE5AFA58994C" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[644,805,1120,1145]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="enganoense">
<emphasis id="B48ABC63F746FFE7FD75DE5AFA58994C" box="[644,805,1120,1145]" italics="true" pageId="22" pageNumber="23">P. enganoense</emphasis>
</taxonomicName>
, not bordered by black except for a very small blackish line on vertex (
<figureCitation id="1EC57CF4F746FFE7FEB1DEB9F8EA99A8" box="[320,407,1155,1181]" captionStart="FIGURES 2628" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[152,1432,1242,1879]" captionTargetId="figure-369@17.[151,1436,1241,1882]" captionTargetPageId="17" captionText="FIGURES 2628. Pseudagrion pilidorsum declaratum, syntype male: (26) habitus dorsal, (27) labels, (28) habitus lateral. Photos: M. Seehausen." pageId="22" pageNumber="23">Figs 26</figureCitation>
,
<figureCitation id="1EC57CF4F746FFE7FE52DEB9F8BC99A8" box="[419,449,1155,1181]" captionStart="FIGURES 2930" captionStartId="19.[151,264,1776,1800]" captionTargetBox="[153,1433,925,1748]" captionTargetId="figure-250@19.[151,1436,922,1750]" captionTargetPageId="19" captionText="FIGURES 2930. Pseudagrion deflexum, lectotype male: (29) habitus lateral, (30) envelope. Photos: RMNH." pageId="22" pageNumber="23">29</figureCitation>
,
<figureCitation id="1EC57CF4F746FFE7FE3CDEB9F89699A8" box="[461,491,1155,1181]" captionStart="FIGURES 3132" captionStartId="21.[151,264,1458,1482]" captionTargetBox="[152,1433,510,1430]" captionTargetId="figure-103@21.[151,1436,507,1433]" captionTargetPageId="21" captionText="FIGURES 3132. Pseudagrion enganoense, lectotype male: (31) habitus lateral, (32) envelope. Photos: RMNH." pageId="22" pageNumber="23">31</figureCitation>
,
<figureCitation id="1EC57CF4F746FFE7FE06DEB9FB1499AB" box="[503,617,1155,1182]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="22" pageNumber="23">83, 84, 85</figureCitation>
,
<figureCitation id="1EC57CF4F746FFE7FD84DEB9FBA699AB" box="[629,731,1155,1182]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="22" pageNumber="23">155, 157</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE7FF36DE9CFB6999D0" box="[199,532,1190,1253]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="86416071F746FFE7FF36DE9CF82499F5" blockId="22.[151,1437,151,1253]" box="[199,345,1190,1216]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DE9CF82499F5" bold="true" box="[199,345,1190,1216]" pageId="22" pageNumber="23">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F746FFE7FF36DEF1FB6999D0" blockId="22.[151,1437,151,1253]" box="[199,532,1227,1253]" pageId="22" pageNumber="23">
Endemic to Enggano (
<figureCitation id="1EC57CF4F746FFE7FE30DEF1FB7A99D0" box="[449,519,1227,1253]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="22" pageNumber="23">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,398 @@
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<mods:title id="6ABC109E7145FDF16A4532B299ACE97B">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<emphasis id="B48ABC63F746FFE7FF66DF14F8B4987C" bold="true" box="[151,457,1326,1353]" italics="true" pageId="22" pageNumber="23">Pseudagrion halmaherae</emphasis>
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Figures: 33 (habitus
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); 3436 (S910, caudal appendages
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); 37 (head
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); 38 (pronotum
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); 39 (habitus
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); 40 (head
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); 41 (pronotum
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); 42 (S910, ovipositor
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); 86 (head
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); 103 (pterothorax
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); 120a, b (abdomen
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); 137ac (caudal appendages
<specimenCount id="90F8ABF8F746FFE7FC3ADF97FA9D98F0" box="[971,992,1453,1477]" count="1" pageId="22" pageNumber="23" type="male"></specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE7FF66DFF1FD3B98D6" box="[151,1094,1483,1507]" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="86416071F746FFE7FF66DFF1FD3B98D6" blockId="22.[151,1437,1391,1507]" box="[151,1094,1483,1507]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="A6EE475FF746FFE7FF66DFF1FD3B98D6" box="[151,1094,1483,1507]" pageId="22" pageNumber="23">
?
<taxonomicName id="41FE1BF2F746FFE7FF58DFF1F8E198D6" ID-CoL="6WD8B" authorityName="Selys" authorityYear="1876" box="[169,412,1483,1507]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F746FFE7FF58DFF1F8E198D6" box="[169,412,1483,1507]" italics="true" pageId="22" pageNumber="23">Pseudagrion coriaceum</emphasis>
</taxonomicName>
:
<treatmentCitation id="075F4660F746FFE7FE58DFF1FB2098D6" author="Selys Longchamps, E. de" box="[425,605,1483,1507]" page="296" pageId="22" pageNumber="23" year="1878">
<bibRefCitation id="E26F1D80F746FFE7FE58DFF1FB5398D6" author="Selys Longchamps, E. de" box="[425,558,1483,1507]" pageId="22" pageNumber="23" pagination="289 - 322" refId="ref42087" refString="Selys Longchamps, E. de (1878) Odonates de la region de la nouvelle Guinee. Mittheilungen aus dem Koniglichen Zoologischen Museum zu Dresden, 3, 289-322." type="journal article" year="1878">Selys (1878)</bibRefCitation>
: 296
</treatmentCitation>
(Halmahera);—
<treatmentCitation id="075F4660F746FFE7FCF4DFF1FAC598D6" author="Selys Longchamps, E. de" box="[773,952,1483,1507]" page="290" pageId="22" pageNumber="23" year="1879">
<bibRefCitation id="E26F1D80F746FFE7FCF4DFF1FAF598D6" author="Selys Longchamps, E. de" box="[773,904,1483,1507]" pageId="22" pageNumber="23" pagination="287 - 324" refId="ref42120" refString="Selys Longchamps, E. de (1879) Nouvelles observations sur les odonates de la region de la Nouvelle Guinee. Annali del museo civico di storia naturale, 14, 287-324." type="journal article" year="1879">Selys (1879)</bibRefCitation>
: 290
</treatmentCitation>
(Halmahera).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE7FF66DC37FB929BA1" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="86416071F746FFE7FF66DC37FD9E9B7E" blockId="22.[151,1437,1549,2042]" pageId="22" pageNumber="23">
<materialsCitation id="36966A2CF746FFE7FF66DC37FD9E9B7E" collectingDate="1951-10-07" collectingDateMax="1951-10-12" collectingDateMin="1951-10-07" country="Indonesia" county="Naturalis Biodiversity Center Leiden" elevation="600" latitude="1.3" location="Biaur" longLatPrecision="7855" longitude="127.7" municipality="Deposited" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Maluku Utara" typeStatus="holotype">
<emphasis id="B48ABC63F746FFE7FF66DC37F8799B12" bold="true" box="[151,260,1549,1575]" pageId="22" pageNumber="23">
<typeStatus id="5945DED3F746FFE7FF66DC37F8799B12" box="[151,260,1549,1575]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="90F8ABF8F746FFE7FEE0DC34F8559B1D" box="[273,296,1550,1576]" count="1" pageId="22" pageNumber="23" type="male"></specimenCount>
,
<date id="F24046B1F746FFE7FEC3DC37F8B59B1D" box="[306,456,1549,1576]" pageId="22" pageNumber="23" value="1951-10-07" valueMax="1951-10-12" valueMin="1951-10-07">
<collectingDate id="E204BF59F746FFE7FEC3DC37F8B59B1D" box="[306,456,1549,1576]" pageId="22" pageNumber="23" value="1951-10-07" valueMax="1951-10-12" valueMin="1951-10-07">7.12.x.1951</collectingDate>
</date>
,
<location id="832136AAF746FFE7FE23DC37FB6F9B12" LSID="urn:lsid:plazi:treatment:0E57D167F746FFE8FF66DF14FB6498B3:832136AAF746FFE7FE23DC37FB6F9B12" box="[466,530,1549,1575]" country="Indonesia" county="Naturalis Biodiversity Center Leiden" latitude="1.3" longLatPrecision="7855" longitude="127.7" municipality="Deposited" name="Biaur" pageId="22" pageNumber="23" stateProvince="Maluku Utara">Biaur</location>
,
<location id="832136AAF746FFE7FDEADC37FB9F9B12" LSID="urn:lsid:plazi:treatment:0E57D167F746FFE8FF66DF14FB6498B3:832136AAF746FFE7FDEADC37FB9F9B12" box="[539,738,1549,1575]" country="Indonesia" county="Naturalis Biodiversity Center Leiden" latitude="1.3" longLatPrecision="7855" longitude="127.7" municipality="Deposited" name="Halmahera Island" pageId="22" pageNumber="23" stateProvince="Maluku Utara">Halmahera Island</location>
,
<collectingRegion id="443AAE93F746FFE7FD1DDC37FA899B12" box="[748,1012,1549,1575]" country="Indonesia" name="Maluku Utara" pageId="22" pageNumber="23">North Maluku province</collectingRegion>
,
<collectingCountry id="FEE920E1F746FFE7FC0CDC37FD109B12" box="[1021,1133,1549,1575]" name="Indonesia" pageId="22" pageNumber="23">Indonesia</collectingCountry>
,
<quantity id="4106CD94F746FFE7FB86DC37FDC19B1D" box="[1143,1212,1549,1576]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="m" value="600.0">
<elevation id="0DD38742F746FFE7FB86DC37FDC19B1D" box="[1143,1212,1549,1576]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="m" value="600.0">600 m</elevation>
</quantity>
a.s.l, approximately
<geoCoordinate id="E3CA06B6F746FFE7FF66DC0BF9A19B7E" box="[151,220,1585,1611]" degrees="1.3" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="5555" value="1.3">1.3°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F746FFE7FF19DC0BF83B9B7E" box="[232,326,1585,1611]" degrees="127.7" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="127.7">127.7°E</geoCoordinate>
, no. 1519084.
<collectingMunicipality id="6625FA0BF746FFE7FE1ADC0BFB209B7E" box="[491,605,1585,1611]" pageId="22" pageNumber="23">Deposited</collectingMunicipality>
at
<collectingCounty id="6F2018FDF746FFE7FD8EDC0BFD5D9B7E" box="[639,1056,1585,1611]" pageId="22" pageNumber="23">Naturalis Biodiversity Center Leiden</collectingCounty>
, the Netherlands.
</materialsCitation>
</paragraph>
<paragraph id="86416071F746FFE7FF36DC6FFB929BA1" blockId="22.[151,1437,1549,2042]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DC6FF8499B5A" bold="true" box="[199,308,1621,1647]" pageId="22" pageNumber="23">
<typeStatus id="5945DED3F746FFE7FF36DC6FF8499B5A" box="[199,308,1621,1647]" pageId="22" pageNumber="23" type="paratype">Paratype</typeStatus>
</emphasis>
(
<specimenCount id="90F8ABF8F746FFE7FEB9DC6FF8149B5A" box="[328,361,1621,1647]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
):
<specimenCount id="90F8ABF8F746FFE7FE73DC6FF8D89B5A" box="[386,421,1621,1647]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
,
<date id="F24046B1F746FFE7FE45DC6FFB379B45" box="[436,586,1621,1648]" pageId="22" pageNumber="23" value="1951-10-07" valueMax="1951-10-12" valueMin="1951-10-07">7.12.x.1951</date>
,
<materialsCitation id="36966A2CF746FFE7FDA6DC6FFB919BA1" collectionCode="RMNH" country="Indonesia" elevation="600" latitude="1.3" location="Biaur" longLatPrecision="7855" longitude="127.7" municipality="Halmahera Island" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Maluku Utara" typeStatus="paratype">
<location id="832136AAF746FFE7FDA6DC6FFBEA9B5A" LSID="urn:lsid:plazi:treatment:0E57D167F746FFE8FF66DF14FB6498B3:832136AAF746FFE7FDA6DC6FFBEA9B5A" box="[599,663,1621,1647]" country="Indonesia" latitude="1.3" longLatPrecision="7855" longitude="127.7" municipality="Halmahera Island" name="Biaur" pageId="22" pageNumber="23" stateProvince="Maluku Utara">Biaur</location>
,
<collectingMunicipality id="6625FA0BF746FFE7FD52DC6FFA109B5A" box="[675,877,1621,1647]" pageId="22" pageNumber="23">Halmahera Island</collectingMunicipality>
,
<collectingRegion id="443AAE93F746FFE7FC8BDC6FFDF59B5A" box="[890,1160,1621,1647]" country="Indonesia" name="Maluku Utara" pageId="22" pageNumber="23">North Maluku province</collectingRegion>
,
<collectingCountry id="FEE920E1F746FFE7FB65DC6FFC799B5A" box="[1172,1284,1621,1647]" name="Indonesia" pageId="22" pageNumber="23">Indonesia</collectingCountry>
,
<elevation id="0DD38742F746FFE7FAE0DC6FFCEA9B5A" box="[1297,1431,1621,1647]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="m" value="600.0">
<quantity id="4106CD94F746FFE7FAE0DC6FFC249B5A" box="[1297,1369,1621,1647]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
, approximately
<geoCoordinate id="E3CA06B6F746FFE7FEB1DC43F8F99BA6" box="[320,388,1657,1683]" degrees="1.3" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="5555" value="1.3">1.3°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F746FFE7FE61DC43F8929BA6" box="[400,495,1657,1683]" degrees="127.7" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="127.7">127.7°E</geoCoordinate>
,
<collectionCode id="E0EFF8B4F746FFE7FE0BDC43FB2C9BA6" box="[506,593,1657,1683]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23" type="Museum">RMNH</collectionCode>
, no. 1519084
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE7FF36DCA7FA949B82" box="[199,1001,1693,1719]" pageId="22" pageNumber="23" type="etymology">
<paragraph id="86416071F746FFE7FF36DCA7FA949B82" blockId="22.[151,1437,1549,2042]" box="[199,1001,1693,1719]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DCA7F82F9B82" bold="true" box="[199,338,1693,1719]" pageId="22" pageNumber="23">Etymology:</emphasis>
Named with regards to the
<typeStatus id="5945DED3F746FFE7FD78DCA7FBC49B82" box="[649,697,1693,1719]" pageId="22" pageNumber="23">type</typeStatus>
locality Halmahera Island.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE7FF36DCFBFD8F9BCB" pageId="22" pageNumber="23" type="discussion">
<paragraph id="86416071F746FFE7FF36DCFBFD8F9BCB" blockId="22.[151,1437,1549,2042]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DCFBF8499BEE" bold="true" box="[199,308,1729,1755]" pageId="22" pageNumber="23">Remarks</emphasis>
: Further specimens from Halmahera were discovered among unidentified material at the RMNH by V.J. Kalkman just prior to submission. For this reason we have not included them in the
<typeStatus id="5945DED3F746FFE7FB82DCDFFDDE9BCA" box="[1139,1187,1765,1791]" pageId="22" pageNumber="23">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF746FFE9FF36DD32FBB89BEE" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="86416071F746FFE7FF36DD32FB629A17" blockId="22.[151,1437,1549,2042]" box="[199,543,1800,1826]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DD32FB629A17" bold="true" box="[199,543,1800,1826]" pageId="22" pageNumber="23">
Description of
<typeStatus id="5945DED3F746FFE7FE86DD32F8A39A17" box="[375,478,1800,1826]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="86416071F746FFE7FF36DD16F8C39ACC" blockId="22.[151,1437,1549,2042]" pageId="22" pageNumber="23">
<emphasis id="B48ABC63F746FFE7FF36DD16F8799A73" box="[199,260,1836,1862]" italics="true" pageId="22" pageNumber="23">Head</emphasis>
(
<figureCitation id="1EC57CF4F746FFE7FEE2DD16F81B9A73" box="[275,358,1836,1862]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="22" pageNumber="23">Fig. 37</figureCitation>
, also compare
<figureCitation id="1EC57CF4F746FFE7FDFFDD16FB1C9A73" box="[526,609,1836,1862]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="22" pageNumber="23">Fig. 86</figureCitation>
): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow; colour of postfrons sharply separated from black vertex at level of anterior margin of lateral ocelli; median ocellus with a small black spot posteriorly; vertex with a small yellowish spot situated at centre of occipital bar area; postocular spots orange-reddish, rectangle-like with posterior edge somewhat serrated, bordered by black; rear of head black, but yellowish along the eye margins; base and scape of antennae yellow, following segments ochre-brownish.
</paragraph>
<paragraph id="86416071F747FFE6FF36DAADFC2E9F2C" blockId="23.[151,1436,151,609]" pageId="23" pageNumber="24">
<emphasis id="B48ABC63F747FFE6FF36DAADF8659D84" box="[199,280,151,177]" italics="true" pageId="23" pageNumber="24">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F747FFE6FED5DAADF8E29D84" box="[292,415,151,177]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="23" pageNumber="24">Figs 33, 38</figureCitation>
, also compare
<figureCitation id="1EC57CF4F747FFE6FDCFDAADFBE09D84" box="[574,669,151,177]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="23" pageNumber="24">Fig. 103</figureCitation>
):Anterior lobe of pronotum with its centre orange, broadly surrounded by black, and orange patches laterally; middle pronotal lobe black with two parallel elongate orange patches in centre, and on each side a subtriangular orange spot dorsolaterally; propleuron black above, broadly orange below and to rear; posterior lobe of pronotum black, with an elongate orange spot in centre and broadly orange at each lateral margin; apical edge of posterior lobe convex. Pterothorax orange-reddish with its posterior margins black; a broad black middorsal stripe; a broad black humeral stripe, broadened posteriorly; interpleural suture with a black stripe reaching to metathoracic spiracle, broadened around its middle; metapleural suture with a black stripe, broadened posteriorly; mesinfraepisternum with a broad black diagonal stripe, lower margin pale orange; venter of pterothorax pale yellow-orange, covered with whitish pruinosity. Coxae and legs pale yellow-orange; outer side of femora with a washed-out blackish stripe along posterior two-third at foreleg, along posterior half at midleg and lacking at hindleg; tibiae with blackish along the black spines; tarsi washed-out blackish; tarsal claws ochre.
</paragraph>
<paragraph id="86416071F747FFE6FF36D819FB0C9F54" blockId="23.[151,1436,151,609]" pageId="23" pageNumber="24">
<emphasis id="B48ABC63F747FFE6FF36D819F8769F09" box="[199,267,547,572]" italics="true" pageId="23" pageNumber="24">Wings</emphasis>
(
<figureCitation id="1EC57CF4F747FFE6FEE8D819F8169F08" box="[281,363,547,573]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="23" pageNumber="24">Fig. 33</figureCitation>
): Membrane hyaline; veins brown to black; 16.016.5 px in forewing; 14.515.0 px in hindwings; Pt dark reddish-brown, overlying one cell.
</paragraph>
<caption id="D28130F9F747FFE6FF66DD48FD6B9A9B" pageId="23" pageNumber="24" startId="23.[151,264,1906,1930]" targetBox="[152,1430,672,1878]" targetPageId="23" targetType="figure">
<paragraph id="86416071F747FFE6FF66DD48FD6B9A9B" blockId="23.[151,1436,1906,1966]" pageId="23" pageNumber="24">
<emphasis id="B48ABC63F747FFE6FF66DD48F82C9ABF" bold="true" box="[151,337,1906,1930]" pageId="23" pageNumber="24">FIGURES 3338</emphasis>
.
<taxonomicName id="41FE1BF2F747FFE6FE90DD48FB189ABF" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[353,613,1906,1930]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F747FFE6FE90DD48FB189ABF" box="[353,613,1906,1930]" italics="true" pageId="23" pageNumber="24">Pseudagrion halmaherae</emphasis>
</taxonomicName>
, holotype male: (
<emphasis id="B48ABC63F747FFE6FCD3DD48FA3E9ABF" bold="true" box="[802,835,1906,1930]" pageId="23" pageNumber="24">33)</emphasis>
habitus lateral, (
<emphasis id="B48ABC63F747FFE6FC09DD48FD6C9ABF" bold="true" box="[1016,1041,1906,1930]" pageId="23" pageNumber="24">34</emphasis>
) appendages lateral, (
<emphasis id="B48ABC63F747FFE6FB0ADD48FC699ABF" bold="true" box="[1275,1300,1906,1930]" pageId="23" pageNumber="24">35</emphasis>
) appendages dorsal, (
<emphasis id="B48ABC63F747FFE6FF1BDDACF87E9A9B" bold="true" box="[234,259,1942,1966]" pageId="23" pageNumber="24">36</emphasis>
) cercus dorsolateral, (
<emphasis id="B48ABC63F747FFE6FE17DDACF8829A9B" bold="true" box="[486,511,1942,1966]" pageId="23" pageNumber="24">37</emphasis>
) head dorsal, (
<emphasis id="B48ABC63F747FFE6FD66DDACFBCD9A9B" bold="true" box="[663,688,1942,1966]" pageId="23" pageNumber="24">38</emphasis>
) pronotum. Photos: M. Seehausen.
</paragraph>
</caption>
<paragraph id="86416071F748FFE9FF36DAADFD769C29" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36DAADF84C9D84" box="[199,305,151,177]" italics="true" pageId="24" pageNumber="25">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F748FFE9FEB2DAADF88A9D84" box="[323,503,150,177]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="24" pageNumber="25">Figs 33, 34, 35</figureCitation>
, also compare
<figureCitation id="1EC57CF4F748FFE9FD56DAADFA479D84" box="[679,826,151,177]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="24" pageNumber="25">Figs 120a, b</figureCitation>
): S1 orange, dorsally with a goblet-like black patch; S2 orange, dorsally black, black colour expanded laterally at posterior end; S38 black dorsally; yellow-orange laterally; S9 orange, with a black middorsal stripe, and a quadrangle-like black patch covering posterior two-thirds of segment; laterally with the posterior third black; S10 orange, dorsally black.
</paragraph>
<paragraph id="86416071F748FFE9FF36DB1CF8BB9F6A" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36DB1CF8DA9C75" box="[199,423,294,320]" italics="true" pageId="24" pageNumber="25">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F748FFE9FE47DB1CFB1C9C75" box="[438,609,294,321]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="24" pageNumber="25">Figs 34, 35, 36</figureCitation>
, also compare
<figureCitation id="1EC57CF4F748FFE9FCF8DB1CFAEA9C75" box="[777,919,294,320]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="24" pageNumber="25">Figs 137ac</figureCitation>
): Cercus washed-out blackish, slightly shorter than S10, inferior lobe shorter than superior lobe; cercus in lateral view subtriangular due to an almost straight upper margin of the cercus, and a convex lower margin; lower margin with a narrow but deep inter-lobe notch subapically; in dorsal view cercus almost straight, then turned slightly inward with its apex narrowly rounded and in dorsolateral view curved inward; in dorsal and dorsolateral view a cleft is visible between superior and inferior lobe; inferior lobe projected shelf-like to inward, a forward and inward directed subbasal interior projection is present, appearing larger in dorsolateral view; paraproct pale yellow-ochre, slightly longer than half of cercus; in lateral view subtriangular and tapered with a rounded apex; in dorsal view broadly rounded, concave bowl-like; with a small black tooth at medial edge.
</paragraph>
<paragraph id="86416071F748FFE9FF36D853FA3B9F92" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">Measurements (mm): Total length (with appendages) 51.3; abdomen length (without appendages) 42.3; hindwing 28.8; forewing 30.5; Pt in forewing 0.8; cercus 0.6.</paragraph>
<paragraph id="86416071F748FFE9FF36D88AFB459FFE" blockId="24.[151,1437,150,2043]" box="[199,568,688,715]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36D88AFB459FFE" bold="true" box="[199,568,688,715]" pageId="24" pageNumber="25">
Description of female
<typeStatus id="5945DED3F748FFE9FE3CD88BFB459FFE" box="[461,568,689,715]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="86416071F748FFE9FF36D8EEFBD09EF3" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36D8EEF8799FDB" box="[199,260,724,750]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
(
<figureCitation id="1EC57CF4F748FFE9FEE8D8EEF8099FDA" box="[281,372,724,751]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="24" pageNumber="25">Fig. 40</figureCitation>
): Labium, mandibles, genae, labrum, and anteclypeus yellowish-ochre; postclypeus black; antefrons, and postfrons yellowish with a large black patch at centre; yellow colour of postfrons sharply separated from vertex at level of anterior margin of lateral ocelli; vertex black; a black stripe expanded towards each antenna, and broadly towards the median ocellus, reaching level of lateral ocelli; occipital bar thin yellowish, its centre slightly expanded anteriorly; postocular spots yellow-orange, rectangle-like with their posterior edge somewhat serrated, bordered by black; rear of head black, but yellowish along the eye margins; base and scape of antennae yellowish-ochre, following segments brownish.
</paragraph>
<paragraph id="86416071F748FFE9FF36D9EAF8599841" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36D9EAF8659EDF" box="[199,280,976,1002]" italics="true" pageId="24" pageNumber="25">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F748FFE9FEDAD9EAF8C89EDF" box="[299,437,976,1002]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="24" pageNumber="25">Figs 39, 41</figureCitation>
): Anterior lobe of pronotum yellowish, yellowish colour expanding in centre towards middle lobe, its anterior margin black; middle pronotal lobe black, with a centrally divided large yellowish patch in the middle, and on either side a yellowish spot dorsolaterally; propleuron black above, broadly yellowish below and to rear; posterior lobe of pronotum black; a yellowish spot in centre, and washed-out dark yellowish at each lateral margin; apical edge of posterior lobe convex, bearing two darkish horns directed towards the head, almost reaching the centre of the middle lobe. Pterothorax yellowish with its posterior margins black; a broad black middorsal stripe; a broad black humeral stripe, broadened posteriorly; interpleural suture with a black stripe reaching to half of the pterothorax, its anterior terminus broadened; metapleural suture with a black stripe, broadened posteriorly; mesinfraepisternum with a broad black diagonal stripe, lower margin pale yellowish; venter of pterothorax pale yellowish, covered with whitish pruinosity. Coxae and legs pale yellowish; outer side of femora partly with a washed-out blackish stripe along posterior half; tibiae blackish along the black spines; tarsi obscure blackish; tarsal claws ochre.
</paragraph>
<paragraph id="86416071F748FFE9FF36DF45FB2D9889" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36DF45F87698AD" box="[199,267,1407,1432]" italics="true" pageId="24" pageNumber="25">Wings</emphasis>
(
<figureCitation id="1EC57CF4F748FFE9FEEADF44F81298AD" box="[283,367,1406,1432]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="24" pageNumber="25">Fig. 39</figureCitation>
): Membrane hyaline; veins brown to black; 17.518.5 px in forewing; 15.5 px in hindwings; Pt pale reddish-brown, overlying one cell.
</paragraph>
<paragraph id="86416071F748FFE9FF36DFFCF82F9BA6" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36DFFCF84C98D5" box="[199,305,1478,1504]" italics="true" pageId="24" pageNumber="25">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F748FFE9FEB0DFFCF8BB98D5" box="[321,454,1478,1504]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="24" pageNumber="25">Figs 39, 42</figureCitation>
): S18 dorsally black, laterally pale orange; each basal annulus pale yellowish-orange; S9 dorsally black, laterally pale yellowish-orange, this pale colour expanded dorsally in centre of segment; S10 dorsally black, laterally pale yellowish. Cercus in lateral view straight and tapered, blackish, about half as long as S10; paraproct pale yellow-brownish, short, in lateral view rounded; ovipositor yellowish-orange, not reaching posterior end of S10, two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F748FFE9FF36DCA7FBB89BEE" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">Measurements (mm): Total length (with appendages) 50.7; abdomen length (without appendages) 40.5; hindwing 31.6; forewing 33.0; Pt in forewing 0.9.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF748FFE9FF36DCDEF8419ACE" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="86416071F748FFE9FF36DCDEF8479BCB" blockId="24.[151,1437,150,2043]" box="[199,314,1764,1790]" pageId="24" pageNumber="25">
<emphasis id="B48ABC63F748FFE9FF36DCDEF8479BCB" bold="true" box="[199,314,1764,1790]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F748FFE9FF36DD32F8419ACE" blockId="24.[151,1437,150,2043]" pageId="24" pageNumber="25">
The morphology of the male cercus (
<figureCitation id="1EC57CF4F748FFE9FD9BDD32FB8B9A16" box="[618,758,1800,1827]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="24" pageNumber="25">Figs 137ac</figureCitation>
) is distinct compared to all congeners. There are superficial similarities to the male cercus of
<taxonomicName id="41FE1BF2F748FFE9FDD0DD17FBCB9A73" authorityName="Selys" authorityYear="1876" box="[545,694,1837,1862]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F748FFE9FDD0DD17FBCB9A73" box="[545,694,1837,1862]" italics="true" pageId="24" pageNumber="25">P. coriaceum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F748FFE9FD39DD16FA249A72" box="[712,857,1836,1863]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="24" pageNumber="25">Figs 132ac</figureCitation>
), but
<taxonomicName id="41FE1BF2F748FFE9FC6EDD17FD399A73" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[927,1092,1836,1862]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F748FFE9FC6EDD17FD399A73" box="[927,1092,1836,1862]" italics="true" pageId="24" pageNumber="25">P. halmaherae</emphasis>
</taxonomicName>
differs from the latter by the clearly visible inter-lobe notch in lateral view (
<figureCitation id="1EC57CF4F748FFE9FD5EDD6AFA169A5F" box="[687,875,1872,1898]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="24" pageNumber="25">Figs 132a, 137a</figureCitation>
). Additionally the rectangular postocular spots of
<taxonomicName id="41FE1BF2F748FFE9FF66DD4FF8479ABB" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[151,314,1908,1934]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F748FFE9FF66DD4FF8479ABB" box="[151,314,1908,1934]" italics="true" pageId="24" pageNumber="25">P. halmaherae</emphasis>
</taxonomicName>
, with their somewhat serrated posterior margin, separate this species in both sexes from congeners (
<figureCitation id="1EC57CF4F748FFE9FF6EDDA2F98B9A87" box="[159,246,1944,1970]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="24" pageNumber="25">Figs 33</figureCitation>
,
<figureCitation id="1EC57CF4F748FFE9FEF3DDA2F85C9A87" box="[258,289,1944,1970]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="24" pageNumber="25">40</figureCitation>
,
<figureCitation id="1EC57CF4F748FFE9FEDDDDA2F8319A87" box="[300,332,1944,1970]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="24" pageNumber="25">86</figureCitation>
). Both sexes also have an obvious colour pattern of the pterothorax, in some aspects similar to that of
<taxonomicName id="41FE1BF2F748FFE9FF47DD87F8569AE3" authorityName="Selys" authorityYear="1876" box="[182,299,1981,2006]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crocops">
<emphasis id="B48ABC63F748FFE9FF47DD87F8569AE3" box="[182,299,1981,2006]" italics="true" pageId="24" pageNumber="25">P. crocops</emphasis>
</taxonomicName>
, but the antehumeral stripe of males is red in
<taxonomicName id="41FE1BF2F748FFE9FCC3DD87FAA89AE3" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[818,981,1980,2006]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F748FFE9FCC3DD87FAA89AE3" box="[818,981,1980,2006]" italics="true" pageId="24" pageNumber="25">P. halmaherae</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F748FFE9FC14DD86FD389AE3" box="[997,1093,1980,2006]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="24" pageNumber="25">Fig. 103</figureCitation>
) versus greenish in
<taxonomicName id="41FE1BF2F748FFE9FAD9DD87FCE09AE3" authorityName="Selys" authorityYear="1876" box="[1320,1437,1981,2006]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crocops">
<emphasis id="B48ABC63F748FFE9FAD9DD87FCE09AE3" box="[1320,1437,1981,2006]" italics="true" pageId="24" pageNumber="25">P. crocops</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F748FFE9FF6EDDDAF98B9ACF" box="[159,246,2016,2042]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="24" pageNumber="25">Figs 99</figureCitation>
,
<figureCitation id="1EC57CF4F748FFE9FEF3DDDAF84D9ACE" box="[258,304,2016,2043]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="24" pageNumber="25">154</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D28130F9F749FFE8FF66DEAFFB4599E7" pageId="25" pageNumber="26" startId="25.[151,264,1173,1197]" targetBox="[153,1432,182,1146]" targetPageId="25" targetType="figure">
<paragraph id="86416071F749FFE8FF66DEAFFB4599E7" blockId="25.[151,1436,1173,1234]" pageId="25" pageNumber="26">
<emphasis id="B48ABC63F749FFE8FF66DEAFF82F9998" bold="true" box="[151,338,1173,1197]" pageId="25" pageNumber="26">FIGURES 3942</emphasis>
.
<taxonomicName id="41FE1BF2F749FFE8FE92DEACFB15999B" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[355,616,1174,1198]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F749FFE8FE92DEACFB15999B" box="[355,616,1174,1198]" italics="true" pageId="25" pageNumber="26">Pseudagrion halmaherae</emphasis>
</taxonomicName>
, female paratype: (
<emphasis id="B48ABC63F749FFE8FCCBDEAFFA2E9998" bold="true" box="[826,851,1173,1197]" pageId="25" pageNumber="26">39</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F749FFE8FBE2DEAFFD519998" bold="true" box="[1043,1068,1173,1197]" pageId="25" pageNumber="26">40</emphasis>
) head dorsal, (
<emphasis id="B48ABC63F749FFE8FB23DEAFFD969998" bold="true" box="[1234,1259,1173,1197]" pageId="25" pageNumber="26">41</emphasis>
) pronotum, (
<emphasis id="B48ABC63F749FFE8FA8BDEAFFCEE9998" bold="true" box="[1402,1427,1173,1197]" pageId="25" pageNumber="26">42</emphasis>
) ovipositor lateral. Photos: M. Seehausen.
</paragraph>
</caption>
<subSubSection id="CEE433FAF749FFE8FF36DF1EFB6498B3" pageId="25" pageNumber="26" type="distribution">
<paragraph id="86416071F749FFE8FF36DF1EF824980B" blockId="25.[151,1436,1316,1414]" box="[199,345,1316,1342]" pageId="25" pageNumber="26">
<emphasis id="B48ABC63F749FFE8FF36DF1EF824980B" bold="true" box="[199,345,1316,1342]" pageId="25" pageNumber="26">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F749FFE8FF36DF72FB6498B3" blockId="25.[151,1436,1316,1414]" pageId="25" pageNumber="26">
Probably endemic to Halmahera (
<figureCitation id="1EC57CF4F749FFE8FDBBDF72FBEC9857" box="[586,657,1352,1378]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="25" pageNumber="26">Fig. 3</figureCitation>
). But as this region is little explored, it is possible that this species occurs on adjacent islands as well.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E09C5E6748900CDEF1D9053736C91FD8">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="F2C56259A748CD8B6C2B5E0163DA6AE2">Seehausen, Malte</mods:namePart>
<mods:nameIdentifier id="838B60675C85C6961670E5B9C206985D" type="ORCID">0009-0000-6718-1078</mods:nameIdentifier>
<mods:affiliation id="D6E526858E911C08BA6C3D559F225636">Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078</mods:affiliation>
<mods:nameIdentifier id="40487C31ABBA16735A573B2A8AED10DA" type="email">mseehausen@zoolmuseum.uni-kiel.de</mods:nameIdentifier>
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<mods:role id="40F8C47251D6980E7A26863A9DE6D2CE">
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<mods:namePart id="0FEF9519AE7E4FF407C2D0FE7BDB2B55">Kalkman, Vincent J.</mods:namePart>
<mods:nameIdentifier id="694BF3DDF4C0AEFBD020036FF0908E5F" type="ORCID">0000-0002-1484-7865</mods:nameIdentifier>
<mods:affiliation id="D9D40022230D368D6C8D52D284570F73">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
<mods:nameIdentifier id="113F945D4BFB4EC84E69965E58FB4975" type="email">vincent.kalkman@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name id="DF90AC0D50853261853CE35254640947" type="personal">
<mods:role id="0CB5DAFB567FBE911D14C18038F561BD">
<mods:roleTerm id="8BBD82BBE808736A24615FF9CD301282">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5355DFC196CCE1CA53ADD7539EE3E055">Bedjanič, Matjaž</mods:namePart>
<mods:nameIdentifier id="F07911B1DAF1785E90ADF5605F95351C" type="ORCID">0009-0000-6718-1078</mods:nameIdentifier>
<mods:affiliation id="DB7754D22B0B9D5E6D76044230F3DEAB">Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 &amp; National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author &amp; Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078</mods:affiliation>
<mods:nameIdentifier id="B2BCBCBAF777470553E77D2A796A0B51" type="email">mseehausen@zoolmuseum.uni-kiel.de</mods:nameIdentifier>
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<mods:date id="712FE8E313F6484EF8B280540FF1ED2E">2025</mods:date>
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<mods:number id="55486E3295E4D53C8D9F44CA32B76803">2025-02-12</mods:number>
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<mods:number id="79EDC2A93B03C74FCB1478764D9067CF">5587</mods:number>
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<mods:url id="A5F7769AD3D0A336C1C9EBDE77B887ED">https://doi.org/10.11646/zootaxa.5587.1.1</mods:url>
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<mods:classification id="5479EFC1C907BB1233ED156A5921E0D5">journal article</mods:classification>
<mods:identifier id="25E3DA30B29AFF5A8F759A65120FF99A" type="DOI">10.11646/zootaxa.5587.1.1</mods:identifier>
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<subSubSection id="CEE433FAF749FFE8FF66DFEDFCFE9B66" pageId="25" pageNumber="26" type="nomenclature">
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<taxonomicName id="41FE1BF2F749FFE8FF66DFEDF8EE98C7" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[151,403,1495,1522]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="igneum" status="sp. nov.">
<emphasis id="B48ABC63F749FFE8FF66DFEDF8EE98C7" bold="true" box="[151,403,1495,1522]" italics="true" pageId="25" pageNumber="26">Pseudagrion igneum</emphasis>
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Figures: 43 (habitus
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); 4446 (S910, caudal appendages
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); 47 (head
<typeStatus id="5945DED3F749FFE8FBDFDC26FDFA9B01" box="[1070,1159,1564,1588]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F749FFE8FB7DDC26FDDC9B01" box="[1164,1185,1564,1588]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
); 48 (pronotum
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<specimenCount id="90F8ABF8F749FFE8FF36DC01F9A19B66" box="[199,220,1595,1619]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
); 87 (head
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); 104 (pterothorax
<specimenCount id="90F8ABF8F749FFE8FDD6DC01FB419B66" box="[551,572,1595,1619]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
); 121a, b (abdomen
<specimenCount id="90F8ABF8F749FFE8FCFDDC01FA5C9B66" box="[780,801,1595,1619]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
); 138ac (caudal appendages
<specimenCount id="90F8ABF8F749FFE8FBA3DC01FD1A9B66" box="[1106,1127,1595,1619]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
); 161162 (field photos
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).
</paragraph>
</subSubSection>
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<taxonomicName id="41FE1BF2F749FFE8FF66DC60F8F09B47" authorityName="Brauer" authorityYear="1868" box="[151,397,1626,1650]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F749FFE8FF66DC60F8F09B47" box="[151,397,1626,1650]" italics="true" pageId="25" pageNumber="26">Pseudagrion pilidorsum</emphasis>
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: Hämäläinen
<emphasis id="B48ABC63F749FFE8FDEADC60FB359B47" box="[539,584,1626,1650]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. (1999): 28 (Middle Andaman; subspecific status remains to be studied);—
<bibRefCitation id="E26F1D80F749FFE8FAB2DC60F8359BA5" author="Koparde, P. &amp; Dawn, P. &amp; Sumanapala, A." pageId="25" pageNumber="26" pagination="37 - 41" refId="ref40432" refString="Koparde, P., Dawn, P. &amp; Sumanapala, A. (2018) Islands are calling - Short expedition to the Andaman Islands reveals five new spatial records and research gaps. Agrion, 22 (1), 37-41." type="journal article" year="2018">
Koparde
<emphasis id="B48ABC63F749FFE8FF36DC43F9899BA5" box="[199,244,1656,1680]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. (2018)
</bibRefCitation>
: 41,
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Figs
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, 3d
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(North Andaman; map; photo
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alive, habitat photo).
</paragraph>
</subSubSection>
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<materialsCitation id="36966A2CF749FFE8FF66DC85FC1F9BC9" collectingDate="1998-12-08" collectorName="Coll. M. Hamalainen &amp; Wen-Chi Yeh &amp; Deposited &amp; Naturalis Biodiversity Center Leiden" country="India" latitude="11.503" location="Chiriyatapu" longLatPrecision="77" longitude="92.702" municipality="South Andaman" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B48ABC63F749FFE8FF66DC85F8799BEC" bold="true" box="[151,260,1727,1753]" pageId="25" pageNumber="26">
<typeStatus id="5945DED3F749FFE8FF66DC85F8799BEC" box="[151,260,1727,1753]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
:
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,
<date id="F24046B1F749FFE8FECCDC84F8CC9BED" box="[317,433,1726,1752]" pageId="25" pageNumber="26" value="1998-12-08">
<collectingDate id="E204BF59F749FFE8FECCDC84F8CC9BED" box="[317,433,1726,1752]" pageId="25" pageNumber="26" value="1998-12-08">8.xii.1998</collectingDate>
</date>
,
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,
<collectingMunicipality id="6625FA0BF749FFE8FDA7DC84FA6C9BED" box="[598,785,1726,1752]" pageId="25" pageNumber="26">South Andaman</collectingMunicipality>
,
<collectingCountry id="FEE920E1F749FFE8FCEEDC84FA269BED" box="[799,859,1726,1752]" name="India" pageId="25" pageNumber="26">India</collectingCountry>
, approximately
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,
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, ex.
<collectorName id="2B0B05A7F749FFE8FACBDC84F85C9BC9" pageId="25" pageNumber="26">Coll. M. Hämäläinen</collectorName>
,
<collectorName id="2B0B05A7F749FFE8FEDDDCD8F8B89BC9" box="[300,453,1762,1788]" pageId="25" pageNumber="26">Wen-Chi Yeh</collectorName>
, no. 1152912.
<collectorName id="2B0B05A7F749FFE8FD98DCD8FBA69BC9" box="[617,731,1762,1788]" pageId="25" pageNumber="26">Deposited</collectorName>
at
<collectorName id="2B0B05A7F749FFE8FD0CDCD8FDE29BC9" box="[765,1183,1762,1789]" pageId="25" pageNumber="26">Naturalis Biodiversity Center Leiden</collectorName>
, the Netherlands.
</materialsCitation>
</paragraph>
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<emphasis id="B48ABC63F749FFE8FF36DD3DF8429A14" bold="true" box="[199,319,1799,1825]" pageId="25" pageNumber="26">
<typeStatus id="5945DED3F749FFE8FF36DD3DF8429A14" box="[199,319,1799,1825]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="90F8ABF8F749FFE8FEBEDD3DF8F69A14" box="[335,395,1799,1825]" count="2" pageId="25" pageNumber="26" type="male">2♂♂</specimenCount>
):
<specimenCount id="90F8ABF8F749FFE8FE6EDD3DF8A79A14" box="[415,474,1799,1825]" count="2" pageId="25" pageNumber="26" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F749FFE8FE14DD3CFB249A15" box="[485,601,1798,1824]" pageId="25" pageNumber="26" value="1976-12-09">9.xii.1976</date>
,
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rivulet near
<location id="832136AAF749FFE8FD1ADD3CFA0F9A15" LSID="urn:lsid:plazi:treatment:0E57D167F749FFEAFF66DFEDFA1E9AB1:832136AAF749FFE8FD1ADD3CFA0F9A15" box="[747,882,1798,1824]" country="India" municipality="South Andaman" name="Asmadabad" pageId="25" pageNumber="26">Asmadabad</location>
,
<collectingMunicipality id="6625FA0BF749FFE8FC8CDD3CFD499A15" box="[893,1076,1798,1824]" pageId="25" pageNumber="26">South Andaman</collectingMunicipality>
,
<collectingCountry id="FEE920E1F749FFE8FBCFDD3CFD079A15" box="[1086,1146,1798,1824]" name="India" pageId="25" pageNumber="26">India</collectingCountry>
,
<collectionCode id="E0EFF8B4F749FFE8FB77DD3DFDA09A14" box="[1158,1245,1799,1825]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26" type="Museum">RMNH</collectionCode>
, no. 1552773
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF749FFE8FF36DD11F8779A5D" pageId="25" pageNumber="26" type="etymology">
<paragraph id="86416071F749FFE8FF36DD11F8779A5D" blockId="25.[151,1437,1726,2041]" pageId="25" pageNumber="26">
<emphasis id="B48ABC63F749FFE8FF36DD11F82F9A70" bold="true" box="[199,338,1835,1861]" pageId="25" pageNumber="26">Etymology:</emphasis>
The name refers to the fiery impression of the bright red thorax colouration of mature males (
<figureCitation id="1EC57CF4F749FFE8FA9BDD10F9809A5D" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="25" pageNumber="26">Figs 161, 162</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF749FFEBFF36DD49F8BB9F08" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
<paragraph id="86416071F749FFE8FF36DD49F8499AB8" blockId="25.[151,1437,1726,2041]" box="[199,308,1907,1933]" pageId="25" pageNumber="26">
<emphasis id="B48ABC63F749FFE8FF36DD49F8499AB8" bold="true" box="[199,308,1907,1933]" pageId="25" pageNumber="26">Remarks</emphasis>
</paragraph>
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Matjaž Bedjanič observed a male of this species at a stream along the Mayabunder-Diglipur section of the Great Andaman Trunk Road in the North Andaman (
<geoCoordinate id="E3CA06B6F749FFE8FD1EDD81FA109AE0" box="[751,877,1979,2005]" degrees="13.1872" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="13.1872">13.1872°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F749FFE8FC8BDD81FA889AE0" box="[890,1013,1979,2005]" degrees="92.8893" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="92.8893">92.8893°E</geoCoordinate>
;
<figureCitation id="1EC57CF4F749FFE8FBF0DD80FDDC9AE1" box="[1025,1185,1978,2004]" captionStart="FIGURES 155162" captionStartId="63.[151,264,1636,1660]" captionTargetBox="[153,1431,183,1608]" captionTargetId="figure-18@63.[151,1436,179,1610]" captionTargetPageId="63" captionText="FIGURES 155162. Live males of the Pseudagrion red-group. (155) P. declaratum, 18.ii.2014, Banyar Hot Springs environs, Bali, Indonesia. Photo: O.E. Kosterin. (156) P. deflexum, 3.ii.2022, Aileu, Timor-Leste. Photo: M. Rose. (157159) P. deflexum, 17.viii.2018, Lokpin spring, Oecusse SAR, Timor-Leste. Photo: R.M. da Silva Pinto. (160) P. deflexum, immature, 28.vi.2022, Kupang Regency, Timor, Indonesia. Photo: M. Rose. (161162) P. igneum, 7.vii.2017, stream at the road Mayabunder-Diglipur, North Andaman Islands, India. Photos: M. Bedjanič." pageId="25" pageNumber="26">Figs 161, 162</figureCitation>
). This stream is about one metre wide, with sparse bank vegetation, and the surroundings are covered by forest. The locality is situated about
<quantity id="4106CD94F74AFFEBFF2FDAACF8649D84" box="[222,281,150,177]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="km" value="5.0">5 km</quantity>
SW of the locality where the species has been observed a month later by
<bibRefCitation id="E26F1D80F74AFFEBFB9CDAADFC1B9D84" author="Koparde, P. &amp; Dawn, P. &amp; Sumanapala, A." box="[1133,1382,151,177]" pageId="26" pageNumber="27" pagination="37 - 41" refId="ref40432" refString="Koparde, P., Dawn, P. &amp; Sumanapala, A. (2018) Islands are calling - Short expedition to the Andaman Islands reveals five new spatial records and research gaps. Agrion, 22 (1), 37-41." type="journal article" year="2018">
Koparde
<emphasis id="B48ABC63F74AFFEBFB26DAADFC779D84" box="[1239,1290,151,177]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. (2018)
</bibRefCitation>
, and approximately
<quantity id="4106CD94F74AFFEBFEB1DA81F8F59DE0" box="[320,392,187,213]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="27" unit="km" value="80.0">80 km</quantity>
N of the species locality in the Middle Andaman, where a male of the species was collected in
<date id="F24046B1F74AFFEBFF47DAE5F8519DCC" box="[182,300,223,249]" pageId="26" pageNumber="27" value="1997-06">June 1997</date>
, and reported by Hämäläinen
<emphasis id="B48ABC63F74AFFEBFD74DAE5FBCA9DCC" box="[645,695,223,249]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. (1999). It is expected that with further targeted fieldwork in the right season and in suitable habitats, the species will be found at additional localities on the Andaman Islands. However, as the newly described
<taxonomicName id="41FE1BF2F74AFFEBFDEADB1DFBD79C74" authorityName="Selys" authorityYear="1876" box="[539,682,295,321]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B48ABC63F74AFFEBFDEADB1DFBD79C74" box="[539,682,295,321]" italics="true" pageId="26" pageNumber="27">Pseudagrion</emphasis>
</taxonomicName>
is a colourful and quite conspicuous damselfly, the fact that only records of
<specimenCount id="90F8ABF8F74AFFEBFEE3DB71F8009C50" box="[274,381,331,357]" count="6" pageId="26" pageNumber="27" type="male">six males</specimenCount>
are known to date, and that rather recent fieldtrips of M. Bedjanič and P. Koparde, P. Dawn and A. Sumanapala in monsoon season of 2017 yielded only one observation each, suggest that the species is rather local. Additional information on distribution, biology, phenology, and habitat requirements is needed for relevant assessment of threat status of this interesting new endemic.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DBE1F8BB9F08" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Other species recorded at the locality were:
<taxonomicName id="41FE1BF2F74AFFEBFD38DBE1FD429CC0" authority="(Rambur, 1842)" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[713,1087,475,501]" class="Insecta" family="Calopterygidae" genus="Vestalis" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B48ABC63F74AFFEBFD38DBE1FA039CC0" box="[713,894,475,501]" italics="true" pageId="26" pageNumber="27">Vestalis gracilis</emphasis>
(
<bibRefCitation id="E26F1D80F74AFFEBFC7EDBE1FD4A9CC0" author="Rambur, M. P." box="[911,1079,475,501]" pageId="26" pageNumber="27" refId="ref41582" refString="Rambur, M. P. (1842) Histoire naturelle des Insectes. Nevropteres. Roret, Paris, 535 pp., 12 pls." type="book" year="1842">Rambur, 1842</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="41FE1BF2F74AFFEBFBBFDBE1F9A69F2C" authority="(Rambur, 1842)" baseAuthorityName="Rambur" baseAuthorityYear="1842" class="Insecta" family="Platycnemididae" genus="Copera" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="marginipes">
<emphasis id="B48ABC63F74AFFEBFBBFDBE1FC549CC1" box="[1102,1321,475,501]" italics="true" pageId="26" pageNumber="27">Copera marginipes</emphasis>
(
<bibRefCitation id="E26F1D80F74AFFEBFACBDBE1F9AE9F2C" author="Rambur, M. P." pageId="26" pageNumber="27" refId="ref41582" refString="Rambur, M. P. (1842) Histoire naturelle des Insectes. Nevropteres. Roret, Paris, 535 pp., 12 pls." type="book" year="1842">Rambur, 1842</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="41FE1BF2F74AFFEBFF1BDBC5FA339F2C" authority="(Fraser, 1924)" baseAuthorityName="Fraser" baseAuthorityYear="1924" box="[234,846,511,537]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="andamanensis">
<emphasis id="B48ABC63F74AFFEBFF1BDBC5FBDD9F2C" box="[234,672,511,537]" italics="true" pageId="26" pageNumber="27">Prodasineura verticalis andamanensis</emphasis>
(
<bibRefCitation id="E26F1D80F74AFFEBFD43DBC5FA3B9F2C" author="Fraser, F. C." box="[690,838,511,537]" pageId="26" pageNumber="27" pagination="409 - 413" refId="ref39735" refString="Fraser, F. C. (1924) Report on a collection of dragonflies (Odonata) from the Andaman Islands. Records of the Indian Museum, 26 (5), 409-413, pl. 24. https://doi.org/10.26515/rzsi/v26/i5/1924/162658" type="journal article" year="1924">Fraser, 1924</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="41FE1BF2F74AFFEBFCACDBC5FD8C9F2C" authority="Selys, 1878" authorityName="Selys" authorityYear="1878" box="[861,1265,511,537]" class="Insecta" family="Libellulidae" genus="Tetrathemis" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platyptera">
<emphasis id="B48ABC63F74AFFEBFCACDBC5FD239F2C" box="[861,1118,511,537]" italics="true" pageId="26" pageNumber="27">Tetrathemis platyptera</emphasis>
<bibRefCitation id="E26F1D80F74AFFEBFB99DBC5FD8C9F2C" author="Selys Longchamps, E. de" box="[1128,1265,511,537]" pageId="26" pageNumber="27" pagination="289 - 322" refId="ref42087" refString="Selys Longchamps, E. de (1878) Odonates de la region de la nouvelle Guinee. Mittheilungen aus dem Koniglichen Zoologischen Museum zu Dresden, 3, 289-322." type="journal article" year="1878">Selys, 1878</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="41FE1BF2F74AFFEBFAC3DBC5F8BF9F08" authority="(Burmeister, 1839)" baseAuthorityName="Burmeister" baseAuthorityYear="1839" class="Insecta" family="Libellulidae" genus="Trithemis" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="aurora">
<emphasis id="B48ABC63F74AFFEBFAC3DBC5F9999F09" italics="true" pageId="26" pageNumber="27">Trithemis aurora</emphasis>
(
<bibRefCitation id="E26F1D80F74AFFEBFF03D819F8C79F08" author="Burmeister, H." box="[242,442,547,573]" pageId="26" pageNumber="27" pagination="757 - 1050" refId="ref38953" refString="Burmeister, H. (1839) Handbuch der Entomologie 2. Theod. Chr. Fried. Enslin, Berlin, pp. 757-1050." type="book chapter" year="1839">Burmeister, 1839</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74AFFEBFF36D87CF8EC9AE8" pageId="26" pageNumber="27" type="description">
<paragraph id="86416071F74AFFEBFF36D87CFB629F54" blockId="26.[151,1437,150,2013]" box="[199,543,582,609]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36D87CFB629F54" bold="true" box="[199,543,582,609]" pageId="26" pageNumber="27">
Description of
<typeStatus id="5945DED3F74AFFEBFE86D87DF8A39F54" box="[375,478,583,609]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="86416071F74AFFEBFF36D851FBA89E0C" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36D851F8799FB0" box="[199,260,619,645]" italics="true" pageId="26" pageNumber="27">Head</emphasis>
(
<figureCitation id="1EC57CF4F74AFFEBFEE2D851F81B9FB0" box="[275,358,619,645]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="26" pageNumber="27">Fig. 47</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74AFFEBFDFFD851FB1C9FB0" box="[526,609,619,645]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="26" pageNumber="27">Fig. 87</figureCitation>
): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow; yellow of postfrons sharply separated at level of median ocellus; vertex black, bordered by an orange-reddish rhombus along the ocelli; a black spot posterior of the median ocellus; followed posteriorly by a large orange-reddish spot, elongated to level of posterior end of lateral ocelli; postocular spots orange-reddish, bordered by black; rear of head black, but narrow yellowish along the eye margins; base and scape of antennae yellow, pedicel brownish, rest of antennae missing.
</paragraph>
<paragraph id="86416071F74AFFEBFF36D979FC1E996C" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36D979F8659E68" box="[199,280,835,861]" italics="true" pageId="26" pageNumber="27">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F74AFFEBFEDAD979F8C89E68" box="[299,437,835,861]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="26" pageNumber="27">Figs 43, 48</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74AFFEBFD9BD979FBAC9E68" box="[618,721,835,861]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="26" pageNumber="27">Fig. 104</figureCitation>
): Anterior lobe of pronotum orange, with its anterior margin black; middle pronotal lobe black, an orange-reddish patch in the middle centrally divided by a black line, and on either side an orange-reddish patch dorsolaterally; propleuron black above, orange below and to rear; posterior lobe of pronotum orange; apical edge of posterior lobe convex. Pterothorax orange; middorsal carina thin black; mesopleural suture with an elongated black patch at posterior third; interpleural suture thin black, with a small dark spot anteriorly; metapleural suture with an elongated black patch at posterior third; mesinfraepisternum pale orange, its superior and posterior edge black; venter of pterothorax yellow-orange, covered with whitish pruinosity. Legs yellowish-ochre, outer side washed with a blackish stripe; tip of tarsi and tarsal claws brown; spines blackish.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DE59FAD99994" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36DE59F8769949" box="[199,267,1123,1148]" italics="true" pageId="26" pageNumber="27">Wings</emphasis>
(
<figureCitation id="1EC57CF4F74AFFEBFEEDDE59F80F9948" box="[284,370,1123,1149]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="26" pageNumber="27">Fig. 43</figureCitation>
): Membrane hyaline; veins brown to black; 12.5 px in right forewing, left forewing damaged; 11.5 px in both hindwings; Pt dark reddish-brown, overlying one cell.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DE91F80398A8" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36DE91F84C99F0" box="[199,305,1195,1221]" italics="true" pageId="26" pageNumber="27">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F74AFFEBFEB0DE91F8E999F0" box="[321,404,1195,1221]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="26" pageNumber="27">Fig. 43</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74AFFEBFDCFDE91FBB199F0" box="[574,716,1195,1221]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="26" pageNumber="27">Figs 121a, b</figureCitation>
): S1 dorsally washed out blackish, laterally orange; S2 dorsally black, apically the black expanded laterally, otherwise laterally orange; S38 dorsally black, apically on each segment the black expanded laterally; each segment laterally pale white-yellowish, except for S8 which is laterally orange; each segment with a narrow pale white-yellowish basal annulus; S9 orange, with its posterior third dorsally and laterally black; S10 (
<figureCitation id="1EC57CF4F74AFFEBFE5CDF01FB519860" box="[429,556,1338,1365]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="26" pageNumber="27">Figs 44, 45</figureCitation>
) with a black posterior margin, and a black middorsal stripe making a “Y”-like marking dorsally; laterally a faint and very narrow black stripe, fading anteriorly, separating two yellow-orange dorsolateral patches.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DF9DF8F69B88" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36DF9DF8D498F4" box="[199,425,1447,1473]" italics="true" pageId="26" pageNumber="27">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F74AFFEBFE4BDF9DFB4298F4" box="[442,575,1447,1473]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="26" pageNumber="27">Figs 4446</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74AFFEBFD1CDF9DFA0098F4" box="[749,893,1447,1473]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="26" pageNumber="27">Figs 138ac</figureCitation>
): Cercus laterally washed-out blackish, slightly shorter than S10, inferior lobe appearing slightly shorter than superior lobe; in lateral view cercus subrectangular, apex with a broad inter-lobe notch; in dorsal view cercus appears narrow, apex of superior lobe slightly pointed and curved inwards, a subtriangular interior projection is situated just proximal of the middle of the cercus; a small, tooth-like subbasal interior projection before half-length of the inferior lobe of the cercus, visible in dorsal and dorsolateral views; paraproct pale yellow-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, with prominent black tooth at blackish medial edge.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DCFDFA3B9A30" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">Measurements (mm): Total length (with appendages) 39.3; abdomen length (without appendages) 32.2; hindwing 21.5; forewing 22.8; Pt in forewing 0.8; cercus 0.6.</paragraph>
<paragraph id="86416071F74AFFEBFF36DD34FB6E9A1C" blockId="26.[151,1437,150,2013]" box="[199,531,1806,1833]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36DD34FB6E9A1C" bold="true" box="[199,531,1806,1833]" pageId="26" pageNumber="27">
Variation in
<typeStatus id="5945DED3F74AFFEBFEACDD35F8B59A1C" box="[349,456,1807,1833]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
males
</emphasis>
</paragraph>
<paragraph id="86416071F74AFFEBFF36DD09FA259A78" blockId="26.[151,1437,150,2013]" box="[199,856,1842,1869]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36DD09F8769A79" box="[199,267,1843,1868]" italics="true" pageId="26" pageNumber="27">Wings</emphasis>
: 13.514 px in forewing, 10.512.5 px in hindwing.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DD6DFAB69A44" blockId="26.[151,1437,150,2013]" box="[199,971,1879,1905]" pageId="26" pageNumber="27">
<emphasis id="B48ABC63F74AFFEBFF36DD6DF84C9A44" box="[199,305,1879,1905]" italics="true" pageId="26" pageNumber="27">Abdomen</emphasis>
: Dorsal black on S9 may be expanded to the posterior half.
</paragraph>
<paragraph id="86416071F74AFFEBFF36DD41FACA9A8C" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">Measurements (mm): Total length (with appendages) 40.342.5; abdomen length (without appendages) 34.3 35.0; hindwing 22.2; forewing 23.8; Pt in forewing 0.8; cercus 0.50.6.</paragraph>
<paragraph id="86416071F74AFFEBFF36DDF9F8EC9AE8" blockId="26.[151,1437,150,2013]" box="[199,401,1987,2013]" pageId="26" pageNumber="27">Female unknown.</paragraph>
</subSubSection>
<caption id="D28130F9F74BFFEAFF66DF75FAB198BE" pageId="27" pageNumber="28" startId="27.[151,264,1359,1383]" targetBox="[153,1433,185,1329]" targetPageId="27" targetType="figure">
<paragraph id="86416071F74BFFEAFF66DF75FAB198BE" blockId="27.[151,1436,1359,1419]" pageId="27" pageNumber="28">
<emphasis id="B48ABC63F74BFFEAFF66DF75F8339852" bold="true" box="[151,334,1359,1383]" pageId="27" pageNumber="28">FIGURES 4348</emphasis>
.
<taxonomicName id="41FE1BF2F74BFFEAFEADDF75FB4D9852" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[348,560,1359,1383]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F74BFFEAFEADDF75FB4D9852" box="[348,560,1359,1383]" italics="true" pageId="27" pageNumber="28">Pseudagrion igneum</emphasis>
</taxonomicName>
, holotype male: (
<emphasis id="B48ABC63F74BFFEAFD17DF75FA7A9852" bold="true" box="[742,775,1359,1383]" pageId="27" pageNumber="28">43)</emphasis>
habitus lateral, (
<emphasis id="B48ABC63F74BFFEAFC44DF75FAB39852" bold="true" box="[949,974,1359,1383]" pageId="27" pageNumber="28">44</emphasis>
) appendages lateral, (
<emphasis id="B48ABC63F74BFFEAFB40DF75FDB79852" bold="true" box="[1201,1226,1359,1383]" pageId="27" pageNumber="28">45</emphasis>
) appendages dorsal, (
<emphasis id="B48ABC63F74BFFEAFF51DF49F9C498BE" bold="true" box="[160,185,1395,1419]" pageId="27" pageNumber="28">46</emphasis>
) cercus dorsolateral, (
<emphasis id="B48ABC63F74BFFEAFE6ADF49F8C998BE" bold="true" box="[411,436,1395,1419]" pageId="27" pageNumber="28">47</emphasis>
) head dorsal, (
<emphasis id="B48ABC63F74BFFEAFDBCDF49FB1B98BE" bold="true" box="[589,614,1395,1419]" pageId="27" pageNumber="28">48</emphasis>
) pronotum. Photos: M. Seehausen.
</paragraph>
</caption>
<subSubSection id="CEE433FAF74BFFEAFF36DFE7FC419A0E" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="86416071F74BFFEAFF36DFE7F84798C2" blockId="27.[151,1437,1501,1924]" box="[199,314,1501,1527]" pageId="27" pageNumber="28">
<emphasis id="B48ABC63F74BFFEAFF36DFE7F84798C2" bold="true" box="[199,314,1501,1527]" pageId="27" pageNumber="28">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F74BFFEAFF36DC3BFC419A0E" blockId="27.[151,1437,1501,1924]" pageId="27" pageNumber="28">
<taxonomicName id="41FE1BF2F74BFFEAFF36DC3BF8C99B2E" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[199,436,1537,1563]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F74BFFEAFF36DC3BF8C99B2E" box="[199,436,1537,1563]" italics="true" pageId="27" pageNumber="28">Pseudagrion igneum</emphasis>
</taxonomicName>
is separated from its Nicobar congeneric
<taxonomicName id="41FE1BF2F74BFFEAFC58DC38FD179B2E" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[937,1130,1537,1563]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F74BFFEAFC58DC38FD179B2E" box="[937,1130,1537,1563]" italics="true" pageId="27" pageNumber="28">P. nigrihumerale</emphasis>
</taxonomicName>
by lacking a broad black humeral stripe (
<figureCitation id="1EC57CF4F74BFFEAFEB7DC1FF89E9B75" box="[326,483,1573,1600]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="27" pageNumber="28">Figs 104, 105</figureCitation>
), S12 of the abdomen dorsally black, S10 red with black dorsal marking (
<figureCitation id="1EC57CF4F74BFFEAFAD7DC1FF8789B56" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="27" pageNumber="28">Figs 121a, b, 122a, b</figureCitation>
), as well as by red on vertex and postfrons (
<figureCitation id="1EC57CF4F74BFFEAFD14DC73FA199B51" box="[741,868,1609,1636]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="27" pageNumber="28">Figs 87, 88</figureCitation>
), whereas the male caudal appendages are identical in both species (
<figureCitation id="1EC57CF4F74BFFEAFEA1DC57FB429BBD" box="[336,575,1645,1672]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="27" pageNumber="28">Figs 138ac, 139ac</figureCitation>
). Also the male cerci of
<taxonomicName id="41FE1BF2F74BFFEAFCA6DC54FA969BB2" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[855,1003,1645,1671]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F74BFFEAFCA6DC54FA969BB2" box="[855,1003,1645,1671]" italics="true" pageId="27" pageNumber="28">P. simalurum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F74BFFEAFBD5DC54FDD29BB2" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[1060,1199,1645,1671]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F74BFFEAFBD5DC54FDD29BB2" box="[1060,1199,1645,1671]" italics="true" pageId="27" pageNumber="28">P. obscurum</emphasis>
</taxonomicName>
are almost identical, seemingly with only a tiny different: the cercus of
<taxonomicName id="41FE1BF2F74BFFEAFD20DCA8FA429B9E" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[721,831,1682,1707]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F74BFFEAFD20DCA8FA429B9E" box="[721,831,1682,1707]" italics="true" pageId="27" pageNumber="28">P. igneum</emphasis>
</taxonomicName>
bears a small subbasal tooth before half-length of the inferior lobe, whereas in
<taxonomicName id="41FE1BF2F74BFFEAFE43DC8CFB419BFA" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[434,572,1717,1743]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F74BFFEAFE43DC8CFB419BFA" box="[434,572,1717,1743]" italics="true" pageId="27" pageNumber="28">P. obscurum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F74BFFEAFD85DC8CFA759BFA" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[628,776,1717,1743]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F74BFFEAFD85DC8CFA759BFA" box="[628,776,1717,1743]" italics="true" pageId="27" pageNumber="28">P. simalurum</emphasis>
</taxonomicName>
there is a sharp edged bulge that only looks like a tooth in dorsal view (
<figureCitation id="1EC57CF4F74BFFEAFED9DCE3FB689BC1" box="[296,533,1753,1780]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="27" pageNumber="28">Figs 140ac, 141ac</figureCitation>
). Additionally
<taxonomicName id="41FE1BF2F74BFFEAFD4ADCE0FA559BC6" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[699,808,1754,1779]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F74BFFEAFD4ADCE0FA559BC6" box="[699,808,1754,1779]" italics="true" pageId="27" pageNumber="28">P. igneum</emphasis>
</taxonomicName>
lacks a darkened dorsum of the pterothorax as it is in
<taxonomicName id="41FE1BF2F74BFFEAFA79DCE0F8789A22" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F74BFFEAFA79DCE0F8789A22" italics="true" pageId="27" pageNumber="28">P. obscurum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F74BFFEAFEE4DCC7F80A9A22" box="[277,375,1789,1815]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="27" pageNumber="28">Fig. 106</figureCitation>
). See also table 2 under
<taxonomicName id="41FE1BF2F74BFFEAFD76DCC4FA6C9A22" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[647,785,1789,1815]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F74BFFEAFD76DCC4FA6C9A22" box="[647,785,1789,1815]" italics="true" pageId="27" pageNumber="28">P. obscurum</emphasis>
</taxonomicName>
for further information. All other species are separated by the morphology of the male appendages, and partly by the colour markings of head, thorax, and abdomen.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74BFFEAFF36DD7FFA1E9AB1" box="[199,867,1861,1924]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="86416071F74BFFEAFF36DD7FF8249A6A" blockId="27.[151,1437,1501,1924]" box="[199,345,1861,1887]" pageId="27" pageNumber="28">
<emphasis id="B48ABC63F74BFFEAFF36DD7FF8249A6A" bold="true" box="[199,345,1861,1887]" pageId="27" pageNumber="28">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F74BFFEAFF36DD53FA1E9AB1" blockId="27.[151,1437,1501,1924]" box="[199,867,1897,1924]" pageId="27" pageNumber="28">
Apparently endemic to the Andaman Islands,
<collectingCountry id="FEE920E1F74BFFEAFD39DD53FA7F9AB6" box="[712,770,1897,1923]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
(
<figureCitation id="1EC57CF4F74BFFEAFCE1DD53FA2B9AB1" box="[784,854,1897,1924]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="27" pageNumber="28">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,593 @@
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<mods:title id="315523CEC3B4B0CAC1EAA350FBB6A6C6">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="CD2D2A38BCA27A7511F9CB10B140BA13">Seehausen, Malte</mods:namePart>
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<mods:namePart id="A94B2D68172304D85C8D23E16F1048F6">Kalkman, Vincent J.</mods:namePart>
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<mods:affiliation id="43F07FD133A7E0545D67D69B71BD3B6E">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
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<taxonomicName id="41FE1BF2F74CFFEDFF66DAADF89B9D87" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[151,486,151,178]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nigrihumerale" status="sp. nov.">
<emphasis id="B48ABC63F74CFFEDFF66DAADF89B9D87" bold="true" box="[151,486,151,178]" italics="true" pageId="28" pageNumber="29">Pseudagrion nigrihumerale</emphasis>
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Figures: 49 (habitus
<typeStatus id="5945DED3F74CFFEDFE9ADAE6F8B99DC1" box="[363,452,220,244]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
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); 5052 (S10, caudal appendages
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<specimenCount id="90F8ABF8F74CFFEDFC6FDAE6FACE9DC1" box="[926,947,220,244]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
); 53 (head
<typeStatus id="5945DED3F74CFFEDFBD9DAE6FDFC9DC1" box="[1064,1153,220,244]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F74CFFEDFB76DAE6FDE19DC1" box="[1159,1180,220,244]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
); 54 (pronotum
<typeStatus id="5945DED3F74CFFEDFAB5DAE6FCE09DC1" box="[1348,1437,220,244]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F74CFFEDFF36DAC0F9A19C27" box="[199,220,250,274]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
); 55 (habitus
<typeStatus id="5945DED3F74CFFEDFE94DAC1F8C19C26" box="[357,444,251,275]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F74CFFEDFE33DAC0F8A99C27" box="[450,468,250,274]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
); 56 (head
<typeStatus id="5945DED3F74CFFEDFDB6DAC1FBE39C26" box="[583,670,251,275]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F74CFFEDFD55DAC0FBCB9C27" box="[676,694,250,274]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
); 57 (pronotum
<typeStatus id="5945DED3F74CFFEDFCAADAC1FACF9C26" box="[859,946,251,275]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F74CFFEDFC49DAC0FAB79C27" box="[952,970,250,274]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
); 58 (S910, ovipositor
<typeStatus id="5945DED3F74CFFEDFB31DAC1FC6A9C26" box="[1216,1303,251,275]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F74CFFEDFAECDAC0FC529C27" box="[1309,1327,250,274]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
); 88 (head
<specimenCount id="90F8ABF8F74CFFEDFF36DB23F9A19C04" box="[199,220,281,305]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
); 105 (pterothorax
<specimenCount id="90F8ABF8F74CFFEDFE51DB23F8C89C04" box="[416,437,281,305]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
); 122a, b (abdomen
<specimenCount id="90F8ABF8F74CFFEDFD74DB23FBE79C04" box="[645,666,281,305]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
); 139ac (caudal appendages
<specimenCount id="90F8ABF8F74CFFEDFC3ADB23FA9D9C04" box="[971,992,281,305]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74CFFEDFF66DB02FD049C65" box="[151,1145,311,336]" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="86416071F74CFFEDFF66DB02FD049C65" blockId="28.[151,1437,219,336]" box="[151,1145,311,336]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="A6EE475FF74CFFEDFF66DB02FD049C65" box="[151,1145,311,336]" pageId="28" pageNumber="29">
<taxonomicName id="41FE1BF2F74CFFEDFF66DB02F8F09C65" ID-CoL="4NJDP" authorityName="Brauer" authorityYear="1868" box="[151,397,312,336]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F74CFFEDFF66DB02F8F09C65" box="[151,397,312,336]" italics="true" pageId="28" pageNumber="29">Pseudagrion pilidorsum</emphasis>
</taxonomicName>
:
<treatmentCitation id="075F4660F74CFFEDFE6BDB02FBB19C7A" author="Rajeshkumar, S. &amp; Raghunathan, C. &amp; Chandra, K." box="[410,716,311,336]" page="45" pageId="28" pageNumber="29" year="2016">
<bibRefCitation id="E26F1D80F74CFFEDFE6BDB02FBDB9C7A" author="Rajeshkumar, S. &amp; Raghunathan, C. &amp; Chandra, K." box="[410,678,311,336]" pageId="28" pageNumber="29" pagination="39 - 46" refId="ref41545" refString="Rajeshkumar, S., Raghunathan, C. &amp; Chandra, K. (2016) Additional records of Odonata from Andaman &amp; Nicobar Islands, India. Biosystematica, 10, 39-46." type="journal article" year="2016">
Rajeshkumar
<emphasis id="B48ABC63F74CFFEDFDD7DB02FB2E9C65" box="[550,595,312,336]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. (2016)
</bibRefCitation>
: 45
</treatmentCitation>
,
<figureCitation id="1EC57CF4F74CFFEDFD27DB02FA599C65" box="[726,804,312,336]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="28" pageNumber="29">Fig. 2H</figureCitation>
(
<collectingCountry id="FEE920E1F74CFFEDFCC1DB02FAA89C65" box="[816,981,312,336]" name="India" pageId="28" pageNumber="29">Nicobar Islands</collectingCountry>
; photo
<specimenCount id="90F8ABF8F74CFFEDFBEEDB02FD4F9C7A" box="[1055,1074,312,335]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
alive).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74CFFEDFF66DB44FBC69E35" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="86416071F74CFFEDFF66DB44F8329CD5" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<materialsCitation id="36966A2CF74CFFEDFF66DB44F8329CD5" collectingDate="2001-01-08" collectorName="Dr Prashant Mohanraj &amp; Coll. M. Hamalainen &amp; Deposited &amp; Naturalis Biodiversity Center Leiden" country="India" latitude="8.01" location="Munak" longLatPrecision="558" longitude="93.505" municipality="Kamorta Island" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B48ABC63F74CFFEDFF66DB44F8799CAD" bold="true" box="[151,260,382,408]" pageId="28" pageNumber="29">
<typeStatus id="5945DED3F74CFFEDFF66DB44F8799CAD" box="[151,260,382,408]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="90F8ABF8F74CFFEDFEE9DB44F8529CAD" box="[280,303,382,408]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
,
<date id="F24046B1F74CFFEDFEB0DB44F8E29CAD" box="[321,415,382,408]" pageId="28" pageNumber="29" value="2001-01-08">
<collectingDate id="E204BF59F74CFFEDFEB0DB44F8E29CAD" box="[321,415,382,408]" pageId="28" pageNumber="29" value="2001-01-08">8.i.2001</collectingDate>
</date>
,
<location id="832136AAF74CFFEDFE41DB44FB7F9CAD" LSID="urn:lsid:plazi:treatment:0E57D167F74CFFEFFF66DAADFA2F9A44:832136AAF74CFFEDFE41DB44FB7F9CAD" box="[432,514,382,408]" country="India" latitude="8.01" longLatPrecision="558" longitude="93.505" municipality="Kamorta Island" name="Munak" pageId="28" pageNumber="29">Munak</location>
,
<collectingMunicipality id="6625FA0BF74CFFEDFDE2DB44FBB59CAD" box="[531,712,382,408]" pageId="28" pageNumber="29">Kamorta Island</collectingMunicipality>
,
<collectingCountry id="FEE920E1F74CFFEDFD28DB44FAEF9CAD" box="[729,914,382,408]" name="India" pageId="28" pageNumber="29">Nicobar Islands</collectingCountry>
,
<collectingCountry id="FEE920E1F74CFFEDFC52DB44FAA29CAD" box="[931,991,382,408]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
, approximately
<geoCoordinate id="E3CA06B6F74CFFEDFB6FDB44FD8D9CAD" box="[1182,1264,382,408]" degrees="8.01" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="8.01">8.01°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F74CFFEDFAF3DB44FC139CAC" box="[1282,1390,382,409]" degrees="93.505" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="55" value="93.505">93.505°E</geoCoordinate>
,
<collectorName id="2B0B05A7F74CFFEDFA8EDB44F8169C89" pageId="28" pageNumber="29">Dr Prashant Mohanraj</collectorName>
leg., ex.
<collectorName id="2B0B05A7F74CFFEDFE20DB98FBBD9C89" box="[465,704,418,444]" pageId="28" pageNumber="29">Coll. M. Hämäläinen</collectorName>
, no. 1552778.
<collectorName id="2B0B05A7F74CFFEDFC95DB98FAAB9C89" box="[868,982,418,444]" pageId="28" pageNumber="29">Deposited</collectorName>
at
<collectorName id="2B0B05A7F74CFFEDFC09DB98FCE59C89" box="[1016,1432,418,444]" pageId="28" pageNumber="29">Naturalis Biodiversity Center Leiden</collectorName>
, the Netherlands.
</materialsCitation>
</paragraph>
<paragraph id="86416071F74CFFEDFF36DBD0FD469FA0" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DBD0F8429F31" bold="true" box="[199,319,490,516]" pageId="28" pageNumber="29">
<typeStatus id="5945DED3F74CFFEDFF36DBD0F8429F31" box="[199,319,490,516]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="90F8ABF8F74CFFEDFEBFDBD0F8F49F31" box="[334,393,490,516]" count="8" pageId="28" pageNumber="29" type="male">8♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F74CFFEDFE65DBD0F8B09F31" box="[404,461,490,516]" count="3" pageId="28" pageNumber="29" type="female">3♀♀</specimenCount>
):
<specimenCount id="90F8ABF8F74CFFEDFE15DBD0FB759F31" box="[484,520,490,516]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<specimenCount id="90F8ABF8F74CFFEDFDE5DBD0FB309F31" box="[532,589,490,516]" count="3" pageId="28" pageNumber="29" type="female">3♀♀</specimenCount>
,
<date id="F24046B1F74CFFEDFDABDBD0FBA19F31" box="[602,732,490,516]" pageId="28" pageNumber="29" value="2000-12-23">23.xii.2000</date>
,
<materialsCitation id="36966A2CF74CFFEDFD16DBD0FABD9F1C" collectionCode="RMNH" collectorName="Dr Prashant Mohanraj &amp; Coll. M. Hamalainen" country="India" location="Great Nicobar" pageId="28" pageNumber="29" specimenCount="1" typeStatus="paratype">
<quantity id="4106CD94F74CFFEDFD16DBD0FA509F31" box="[743,813,490,516]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="28" pageNumber="29" unit="km" value="27.0">27 km</quantity>
EW road,
<location id="832136AAF74CFFEDFC43DBD0FD2E9F31" LSID="urn:lsid:plazi:treatment:0E57D167F74CFFEFFF66DAADFA2F9A44:832136AAF74CFFEDFC43DBD0FD2E9F31" box="[946,1107,490,516]" country="India" name="Great Nicobar" pageId="28" pageNumber="29">Great Nicobar</location>
,
<collectingCountry id="FEE920E1F74CFFEDFBACDBD0FDE49F31" box="[1117,1177,490,516]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
,
<collectorName id="2B0B05A7F74CFFEDFB55DBD0FCE19F31" box="[1188,1436,490,516]" pageId="28" pageNumber="29">Dr Prashant Mohanraj</collectorName>
leg., ex.
<collectorName id="2B0B05A7F74CFFEDFF04D834F89E9F1D" box="[245,483,526,552]" pageId="28" pageNumber="29">Coll. M. Hämäläinen</collectorName>
,
<collectionCode id="E0EFF8B4F74CFFEDFE1FD834FB389F1D" box="[494,581,526,552]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
, no. 15527821552784, 1552786
</materialsCitation>
;—
<specimenCount id="90F8ABF8F74CFFEDFC12D834FD649F1C" box="[995,1049,526,553]" count="3" pageId="28" pageNumber="29" type="male">3♂♂</specimenCount>
,
<date id="F24046B1F74CFFEDFBD2D834FDFC9F1D" box="[1059,1153,526,552]" pageId="28" pageNumber="29" value="2001-01-08">8.i.2001</date>
,
<materialsCitation id="36966A2CF74CFFEDFB7DD834F9839F44" collectionCode="RMNH" collectorName="Dr Prashant Mohanraj &amp; Coll. M. Hamalainen" country="India" location="Munak" municipality="Kamorta Island" pageId="28" pageNumber="29" specimenCount="1" typeStatus="paratype">
<location id="832136AAF74CFFEDFB7DD834FDA39F1D" LSID="urn:lsid:plazi:treatment:0E57D167F74CFFEFFF66DAADFA2F9A44:832136AAF74CFFEDFB7DD834FDA39F1D" box="[1164,1246,526,552]" country="India" municipality="Kamorta Island" name="Munak" pageId="28" pageNumber="29">Munak</location>
,
<collectingMunicipality id="6625FA0BF74CFFEDFB19D834FCE59F1D" box="[1256,1432,526,552]" pageId="28" pageNumber="29">Kamorta Island</collectingMunicipality>
,
<collectingCountry id="FEE920E1F74CFFEDFF66D808F8329F79" box="[151,335,562,588]" name="India" pageId="28" pageNumber="29">Nicobar Islands</collectingCountry>
,
<collectingCountry id="FEE920E1F74CFFEDFEAED808F8E69F79" box="[351,411,562,588]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
,
<collectorName id="2B0B05A7F74CFFEDFE5AD808FBD09F79" box="[427,685,562,588]" pageId="28" pageNumber="29">Dr Prashant Mohanraj</collectorName>
leg., ex.
<collectorName id="2B0B05A7F74CFFEDFCD0D808FD679F79" box="[801,1050,562,588]" pageId="28" pageNumber="29">Coll. M. Hämäläinen</collectorName>
,
<collectionCode id="E0EFF8B4F74CFFEDFBD8D808FDFD9F79" box="[1065,1152,562,588]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
, no. 1552777, 1552779, 1552780
</materialsCitation>
;—
<specimenCount id="90F8ABF8F74CFFEDFED0D86CF8259F44" box="[289,344,598,625]" count="4" pageId="28" pageNumber="29" type="male">4♂♂</specimenCount>
,
<date id="F24046B1F74CFFEDFE96D86CF8BB9F45" box="[359,454,598,624]" pageId="28" pageNumber="29" value="2001-01-09">9.i.2001</date>
,
<materialsCitation id="36966A2CF74CFFEDFE25D86CFD4A9FA0" collectionCode="RMNH" collectorName="Dr Prashant Mohanraj &amp; Coll. M. Hamalainen" country="India" location="Agriculture Department Farm" municipality="Kamorta Island" pageId="28" pageNumber="29" specimenCount="1" typeStatus="paratype">
<location id="832136AAF74CFFEDFE25D86CFA509F45" LSID="urn:lsid:plazi:treatment:0E57D167F74CFFEFFF66DAADFA2F9A44:832136AAF74CFFEDFE25D86CFA509F45" box="[468,813,598,624]" country="India" municipality="Kamorta Island" name="Agriculture Department Farm" pageId="28" pageNumber="29">Agriculture Department Farm</location>
,
<collectingMunicipality id="6625FA0BF74CFFEDFCCCD86CFA8C9F45" box="[829,1009,598,624]" pageId="28" pageNumber="29">Kamorta Island</collectingMunicipality>
,
<collectingCountry id="FEE920E1F74CFFEDFBF1D86CFDC59F45" box="[1024,1208,598,624]" name="India" pageId="28" pageNumber="29">Nicobar Islands</collectingCountry>
,
<collectingCountry id="FEE920E1F74CFFEDFB36D86CFC7E9F45" box="[1223,1283,598,624]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
,
<collectorName id="2B0B05A7F74CFFEDFAE2D86CF8799FA1" pageId="28" pageNumber="29">Dr Prashant Mohanraj</collectorName>
leg., ex.
<collectorName id="2B0B05A7F74CFFEDFE9AD840FB279FA1" box="[363,602,634,660]" pageId="28" pageNumber="29">Coll. M. Hämäläinen</collectorName>
,
<collectionCode id="E0EFF8B4F74CFFEDFD94D840FBC19FA1" box="[613,700,634,660]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
, no. 15527741552776, 1552881
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F74CFFEDFF36D8A7FBC69E35" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<materialsCitation id="36966A2CF74CFFEDFF36D8A7FBC59E35" collectingDate="1996-12-19" collectionCode="RMNH" collectorName="Coll. M. Hamalainen" country="India" location="Campbell Bay" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1">
<emphasis id="B48ABC63F74CFFEDFF36D8A7F8ED9F82" bold="true" box="[199,400,669,695]" pageId="28" pageNumber="29">
<typeStatus id="5945DED3F74CFFEDFF36D8A7F8ED9F82" box="[199,400,669,695]" isOtherTypeStatus="true" pageId="28" pageNumber="29">Other specimens</typeStatus>
</emphasis>
:
<specimenCount id="90F8ABF8F74CFFEDFE50D8A4F8BB9F8D" box="[417,454,670,696]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<date id="F24046B1F74CFFEDFE25D8A4FB2B9F8D" box="[468,598,670,696]" pageId="28" pageNumber="29" value="1996-12-19">
<collectingDate id="E204BF59F74CFFEDFE25D8A4FB2B9F8D" box="[468,598,670,696]" pageId="28" pageNumber="29" value="1996-12-19">19.xii.1996</collectingDate>
</date>
,
<location id="832136AAF74CFFEDFD92D8A4FA7A9F8D" LSID="urn:lsid:plazi:treatment:0E57D167F74CFFEFFF66DAADFA2F9A44:832136AAF74CFFEDFD92D8A4FA7A9F8D" box="[611,775,670,696]" country="India" name="Campbell Bay" pageId="28" pageNumber="29">Campbell Bay</location>
, Great Nicobar,
<collectingCountry id="FEE920E1F74CFFEDFC34D8A4FD7C9F8D" box="[965,1025,670,696]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
, ex.
<collectorName id="2B0B05A7F74CFFEDFBC9D8A4FC569F8D" box="[1080,1323,670,696]" pageId="28" pageNumber="29">Coll. M. Hämäläinen</collectorName>
, Wen-Chi Yeh,
<collectionCode id="E0EFF8B4F74CFFEDFF23D8F8F8549FE9" box="[210,297,706,732]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
, no. 1152932. This freshly emerged male belongs to this new species as well, but was excluded from the
<typeStatus id="5945DED3F74CFFEDFF31D8DCF98D9E35" box="[192,240,742,768]" pageId="28" pageNumber="29">type</typeStatus>
series because of its damaged condition
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74CFFEDFF36D930FA3B9E7D" pageId="28" pageNumber="29" type="etymology">
<paragraph id="86416071F74CFFEDFF36D930FA3B9E7D" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36D930F82F9E11" bold="true" box="[199,338,778,804]" pageId="28" pageNumber="29">Etymology:</emphasis>
The name refers to the black humeral stripe on the thorax, a unique character of the revised species of the
<taxonomicName id="41FE1BF2F74CFFEDFF2ED914F8139E7D" authorityName="Selys" authorityYear="1876" box="[223,366,814,840]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B48ABC63F74CFFEDFF2ED914F8139E7D" box="[223,366,814,840]" italics="true" pageId="28" pageNumber="29">Pseudagrion</emphasis>
</taxonomicName>
red-group occurring in the western range.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74CFFEDFF36D968FB129971" pageId="28" pageNumber="29" type="discussion">
<paragraph id="86416071F74CFFEDFF36D968F8499E59" blockId="28.[151,1437,382,2028]" box="[199,308,850,876]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36D968F8499E59" bold="true" box="[199,308,850,876]" pageId="28" pageNumber="29">Remarks</emphasis>
</paragraph>
<paragraph id="86416071F74CFFEDFF36D94CFB129971" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
The colours when alive are adopted from the field photograph of a living male provided by
<bibRefCitation id="E26F1D80F74CFFEDFB1AD94CF8519E81" author="Rajeshkumar, S. &amp; Raghunathan, C. &amp; Chandra, K." pageId="28" pageNumber="29" pagination="39 - 46" refId="ref41545" refString="Rajeshkumar, S., Raghunathan, C. &amp; Chandra, K. (2016) Additional records of Odonata from Andaman &amp; Nicobar Islands, India. Biosystematica, 10, 39-46." type="journal article" year="2016">
Rajeshkumar
<emphasis id="B48ABC63F74CFFEDFA79D94DF9D09E81" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. (2016: 43
</bibRefCitation>
,
<figureCitation id="1EC57CF4F74CFFEDFECBD9A0F8E89E81" box="[314,405,922,948]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="28" pageNumber="29">Fig. 2H</figureCitation>
). Additionally they provide the following data of this species (identified as
<taxonomicName id="41FE1BF2F74CFFEDFB04D9A1FCF69E81" authorityName="Brauer" authorityYear="1868" box="[1269,1419,922,948]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F74CFFEDFB04D9A1FCF69E81" box="[1269,1419,922,948]" italics="true" pageId="28" pageNumber="29">P. pilidorsum</emphasis>
</taxonomicName>
):
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<specimenCount id="90F8ABF8F74CFFEDFF66D984F9C19EED" box="[151,188,958,984]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<date id="F24046B1F74CFFEDFF3BD984F8309EEC" box="[202,333,958,985]" pageId="28" pageNumber="29" value="2015-12-18">
<collectingDate id="E204BF59F74CFFEDFF3BD984F8309EEC" box="[202,333,958,985]" pageId="28" pageNumber="29" value="2015-12-18">18.xii.2015</collectingDate>
</date>
,
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dam,
<collectingMunicipality id="6625FA0BF74CFFEDFE28D984FB439EED" box="[473,574,958,984]" pageId="28" pageNumber="29">Kamorta</collectingMunicipality>
,
<collectingCounty id="6F2018FDF74CFFEDFDBAD984FA7D9EED" box="[587,768,958,984]" pageId="28" pageNumber="29">Central Nicobar</collectingCounty>
(
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,
<geoCoordinate id="E3CA06B6F74CFFEDFC69D984FD5A9EEC" box="[920,1063,958,985]" degrees="93" direction="east" minutes="29.256" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="93.4876">93°29.256E</geoCoordinate>
=
<geoCoordinate id="E3CA06B6F74CFFEDFBB8D984FDA99EED" box="[1097,1236,958,984]" degrees="8.137367" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="8.137367">8.137367°N</geoCoordinate>
,
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)
</materialsCitation>
;
<materialsCitation id="36966A2CF74CFFEDFA83D984F8459915" collectingDate="2016-02-17" latitude="7.004083" location="Galathea" longLatPrecision="1" longitude="93.903015" municipality="Great Nicobar Island" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1">
<specimenCount id="90F8ABF8F74CFFEDFA83D984FCEA9EED" box="[1394,1431,958,984]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<date id="F24046B1F74CFFEDFF66D9D8F8769EC9" box="[151,267,994,1020]" pageId="28" pageNumber="29" value="2016-02-17">
<collectingDate id="E204BF59F74CFFEDFF66D9D8F8769EC9" box="[151,267,994,1020]" pageId="28" pageNumber="29" value="2016-02-17">17.ii.2016</collectingDate>
</date>
, near streams in interior forest,
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,
<collectingMunicipality id="6625FA0BF74CFFEDFD10D9D8FAB69EC9" box="[737,971,994,1020]" pageId="28" pageNumber="29">Great Nicobar Island</collectingMunicipality>
(
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,
<geoCoordinate id="E3CA06B6F74CFFEDFB91D9D8FD939EC8" box="[1120,1262,994,1021]" degrees="93" direction="east" minutes="54.181" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="93.903015">93°54.181E</geoCoordinate>
=
<geoCoordinate id="E3CA06B6F74CFFEDFAFCD9D8FCE59EC9" box="[1293,1432,994,1020]" degrees="7.004083" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="7.004083">7.004083°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F74CFFEDFF66DE3CF84D9915" box="[151,304,1030,1056]" degrees="93.903017" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="93.903015">93.903017°E</geoCoordinate>
)
</materialsCitation>
;
<specimenCount id="90F8ABF8F74CFFEDFEB2DE3CF81A9915" box="[323,359,1030,1056]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<specimenCount id="90F8ABF8F74CFFEDFE85DE3CF8EA9915" box="[372,407,1030,1056]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
,
<date id="F24046B1F74CFFEDFE54DE3CFB679915" box="[421,538,1030,1056]" pageId="28" pageNumber="29" value="2016-02-21">21.ii.2016</date>
;
<materialsCitation id="36966A2CF74CFFEDFDD4DE3CFB119971" latitude="6.93905" location="Laxmi Nagar" longLatPrecision="1" longitude="93.891136" municipality="Great Nicobar Island" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1">
<specimenCount id="90F8ABF8F74CFFEDFDD4DE3CFB479915" box="[549,570,1030,1056]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
laying eggs on stream near
<location id="832136AAF74CFFEDFC84DE3CFD769915" LSID="urn:lsid:plazi:treatment:0E57D167F74CFFEFFF66DAADFA2F9A44:832136AAF74CFFEDFC84DE3CFD769915" box="[885,1035,1030,1056]" latitude="6.93905" longLatPrecision="1" longitude="93.891136" municipality="Great Nicobar Island" name="Laxmi Nagar" pageId="28" pageNumber="29">Laxmi Nagar</location>
,
<collectingMunicipality id="6625FA0BF74CFFEDFBE7DE3CFC7C9915" box="[1046,1281,1030,1056]" pageId="28" pageNumber="29">Great Nicobar Island</collectingMunicipality>
(
<geoCoordinate id="E3CA06B6F74CFFEDFAE1DE3CFCEA9914" box="[1296,1431,1030,1057]" degrees="6" direction="north" minutes="56.343" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="6.9390497">6°56.343N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F74CFFEDFF66DE10F85B9970" box="[151,294,1066,1093]" degrees="93" direction="east" minutes="53.468" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="93.891136">93°53.468E</geoCoordinate>
=
<geoCoordinate id="E3CA06B6F74CFFEDFEB5DE10F8BC9970" box="[324,449,1066,1093]" degrees="6.93905" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="6.93905">6.93905°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F74CFFEDFE3DDE10FB199971" box="[460,612,1066,1092]" degrees="93.891133" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="93.891136">93.891133°E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74CFFEFFF36DE77FAFE9804" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="86416071F74CFFEDFF36DE77FB62995D" blockId="28.[151,1437,382,2028]" box="[199,543,1101,1128]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DE77FB62995D" bold="true" box="[199,543,1101,1128]" pageId="28" pageNumber="29">
Description of
<typeStatus id="5945DED3F74CFFEDFE86DE74F8A3995D" box="[375,478,1102,1128]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="86416071F74CFFEDFF36DE48FA6E9829" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DE48F87999B9" box="[199,260,1138,1164]" italics="true" pageId="28" pageNumber="29">Head</emphasis>
(
<figureCitation id="1EC57CF4F74CFFEDFEE2DE48F81B99B8" box="[275,358,1138,1165]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="28" pageNumber="29">Fig. 53</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74CFFEDFDFFDE48FB1C99B9" box="[526,609,1138,1164]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="28" pageNumber="29">Fig. 88</figureCitation>
): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow; colour of postfrons sharply separated from vertex at anterior margin of lateral ocelli; vertex black; median ocellus with a black spot anteriorly, merged with black colour of vertex; occipital bar reddish; postocular spots reddish, bordered by black; rear of head black, but yellowish along the eye margins; base and scape of antennae yellow, following segments ochre-brownish.
</paragraph>
<paragraph id="86416071F74CFFEDFF36DF1CFDEB9BB1" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DF1CF8659875" box="[199,280,1318,1344]" italics="true" pageId="28" pageNumber="29">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F74CFFEDFED9DF1CF8D19874" box="[296,428,1318,1345]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="28" pageNumber="29">Figs 49, 54</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74CFFEDFDA9DF1CFBC69874" box="[600,699,1318,1345]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="28" pageNumber="29">Fig. 105</figureCitation>
): Anterior lobe of pronotum orange-reddish, expanding in centre towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large orange-reddish patch in the middle, and on either side a large orange-reddish patch dorsolaterally, fusing laterally with orange-reddish colour of anterior lobe; propleuron black above, orange-reddish below and to rear; posterior lobe orange-reddish; apical edge of posterior lobe convex. Pterothorax orange-reddish with a broad black middorsal stripe, and a broad black humeral stripe, narrowed posteriorly; metapleural suture with a small elongate black patch at posterior third; mesinfraepisternum with two-thirds black, in line with black of humeral stripe, lower third of mesinfraepisternum ochre and washed-out yellow-orange; venter of pterothorax pale yellowish. Coxae pale yellowish-ochre; outer side of femora with a washed-out blackish stripe along posterior half; tibiae with blackish along the black spines; tip of tarsi and tarsal claws ochre, washed-out blackish along joints.
</paragraph>
<paragraph id="86416071F74CFFEDFF36DCB5FAD69BF9" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DCB5F8769B9D" box="[199,267,1679,1704]" italics="true" pageId="28" pageNumber="29">Wings</emphasis>
(
<figureCitation id="1EC57CF4F74CFFEDFEEBDCB4F8129B9D" box="[282,367,1678,1704]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="28" pageNumber="29">Fig. 49</figureCitation>
): Membrane hyaline, slightly tinged yellowish; veins brown to black; 15.516.5 px in forewing; 14.014.5 px in hindwings; Pt dark reddish-brown, overlying one cell.
</paragraph>
<paragraph id="86416071F74CFFEDFF36DCECF8539A90" blockId="28.[151,1437,382,2028]" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DCECF84C9BC5" box="[199,305,1750,1776]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F74CFFEDFEB3DCECF8EA9BC5" box="[322,407,1750,1776]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="28" pageNumber="29">Fig. 49</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74CFFEDFDB2DCECFBA99BC5" box="[579,724,1750,1776]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="28" pageNumber="29">Figs 122a, b</figureCitation>
): S1 orange-reddish, with black along its posterior margin; S2 orange-reddish, dorsally with an anvil-like black marking in posterior third, and a washed-out blackish middorsal stripe; S38 dorsally black, the black colour expanded to lateral at each posterior end; laterally and each basal annulus pale yellowish; S9 with its anterior half laterally and dorsally orange-reddish, separated by a black middorsal stripe which merges with the black posterior half; the anterior margin of the black colour somewhat serrated; S10 black (
<figureCitation id="1EC57CF4F74CFFEDFF6EDDB0F85C9A90" box="[159,289,1930,1957]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="28" pageNumber="29">Figs 50, 51</figureCitation>
).
</paragraph>
<paragraph id="86416071F74CFFECFF36DD94F8F69C50" blockId="28.[151,1437,382,2028]" lastBlockId="29.[151,1436,151,429]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B48ABC63F74CFFEDFF36DD94F8D59AFD" box="[199,424,1966,1992]" italics="true" pageId="28" pageNumber="29">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F74CFFEDFE48DD94FB419AFC" box="[441,572,1966,1993]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="28" pageNumber="29">Figs 5052</figureCitation>
, also compare
<figureCitation id="1EC57CF4F74CFFEDFD19DD94FA0A9AFD" box="[744,887,1966,1992]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="28" pageNumber="29">Figs 139ac</figureCitation>
): Cercus laterally washed-out brownish, slightly shorter than S10, inferior lobe appears slightly shorter than superior lobe; in lateral view cercus subrectangular, apex with a broad inter-lobe notch; in dorsal view cercus appears narrow; apex of superior lobe of the cercus slightly pointed and curved inwards, a subtriangular interior projection is situated just proximal of the middle of the cercus; a small, tooth-like subbasal interior projection before half-length of the inferior lobe of the cercus, visible in dorsal and dorsolateral views; paraproct pale yellow-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; with prominent black tooth at blackish medial edge.
</paragraph>
<paragraph id="86416071F74DFFECFF36DB55FA3B9C98" blockId="29.[151,1436,151,429]" pageId="29" pageNumber="30">Measurements (mm): Total length (with appendages) 41.2; abdomen length (without appendages) 34.3; hindwing 23.7; forewing 24.6; Pt in forewing 0.9; cercus 0.6.</paragraph>
<caption id="D28130F9F74DFFECFF66DD9FFD6B9AD7" pageId="29" pageNumber="30" startId="29.[151,264,1957,1981]" targetBox="[153,1391,491,1928]" targetPageId="29" targetType="figure">
<paragraph id="86416071F74DFFECFF66DD9FFD6B9AD7" blockId="29.[151,1436,1957,2018]" pageId="29" pageNumber="30">
<emphasis id="B48ABC63F74DFFECFF66DD9FF8329A88" bold="true" box="[151,335,1957,1981]" pageId="29" pageNumber="30">FIGURES 4954</emphasis>
.
<taxonomicName id="41FE1BF2F74DFFECFEACDD9CFB0A9A8B" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[349,631,1958,1982]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F74DFFECFEACDD9CFB0A9A8B" box="[349,631,1958,1982]" italics="true" pageId="29" pageNumber="30">Pseudagrion nigrihumerale</emphasis>
</taxonomicName>
, holotype male: (
<emphasis id="B48ABC63F74DFFECFCDEDD9FFA2D9A88" bold="true" box="[815,848,1957,1981]" pageId="29" pageNumber="30">49)</emphasis>
habitus lateral, (
<emphasis id="B48ABC63F74DFFECFC0EDD9FFD659A88" bold="true" box="[1023,1048,1957,1981]" pageId="29" pageNumber="30">50</emphasis>
) appendages lateral, (
<emphasis id="B48ABC63F74DFFECFB0DDD9FFC689A88" bold="true" box="[1276,1301,1957,1981]" pageId="29" pageNumber="30">51</emphasis>
) appendages dorsal, (
<emphasis id="B48ABC63F74DFFECFF1BDDF3F87E9AD4" bold="true" box="[234,259,1993,2017]" pageId="29" pageNumber="30">52</emphasis>
) cercus dorsolateral, (
<emphasis id="B48ABC63F74DFFECFE17DDF3F8829AD4" bold="true" box="[486,511,1993,2017]" pageId="29" pageNumber="30">53</emphasis>
) head dorsal, (
<emphasis id="B48ABC63F74DFFECFD66DDF3FBCD9AD4" bold="true" box="[663,688,1993,2017]" pageId="29" pageNumber="30">54</emphasis>
) pronotum. Photos: M. Seehausen.
</paragraph>
</caption>
<paragraph id="86416071F74EFFEFFF36DAACFB6E9D84" blockId="30.[151,1437,150,1905]" box="[199,531,150,177]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DAACFB6E9D84" bold="true" box="[199,531,150,177]" pageId="30" pageNumber="31">
Variation in
<typeStatus id="5945DED3F74EFFEFFEACDAADF8B59D84" box="[349,456,151,177]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
males
</emphasis>
</paragraph>
<paragraph id="86416071F74EFFEFFF36DA81FAF89DE0" blockId="30.[151,1437,150,1905]" box="[199,901,187,213]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DA81F8799DE0" box="[199,260,187,213]" italics="true" pageId="30" pageNumber="31">Head</emphasis>
: The reddish occipital bar is reduced in some specimens.
</paragraph>
<paragraph id="86416071F74EFFEFFF36DAE5FA109DCC" blockId="30.[151,1437,150,1905]" box="[199,877,222,249]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DAE5F8769DCD" box="[199,267,223,248]" italics="true" pageId="30" pageNumber="31">Wings</emphasis>
: 14.016.5 px in forewing, 12.514.5 px in hindwing.
</paragraph>
<paragraph id="86416071F74EFFEFFF36DB39FC7A9C28" blockId="30.[151,1437,150,1905]" box="[199,1287,259,285]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DB39F84C9C28" box="[199,305,259,285]" italics="true" pageId="30" pageNumber="31">Abdomen</emphasis>
: The middorsal marking on S9 does not reach S8, but middle of S
<quantity id="4106CD94F74EFFEFFBE7DB39FD3C9C28" box="[1046,1089,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="30" pageNumber="31" unit="in" value="9.0">9 in</quantity>
some specimens.
</paragraph>
<paragraph id="86416071F74EFFEFFF36DB1DFD1B9C50" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">Measurements (mm): Total length (with appendages) 37.843.1; abdomen length (without appendages) 31.4 35.5; hindwing 21.024.0; forewing 22.225.3; Pt in forewing 0.70.9; cercus 0.60.7.</paragraph>
<paragraph id="86416071F74EFFEFFF36DB54FB3E9CBC" blockId="30.[151,1437,150,1905]" box="[199,579,366,393]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DB54FB3E9CBC" bold="true" box="[199,579,366,393]" pageId="30" pageNumber="31">
Description of female
<typeStatus id="5945DED3F74EFFEFFE3CDB55FB3E9CBC" box="[461,579,367,393]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="86416071F74EFFEFFF36DBA9FDB89F08" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DBA9F8799C98" box="[199,260,403,429]" italics="true" pageId="30" pageNumber="31">Head</emphasis>
(
<figureCitation id="1EC57CF4F74EFFEFFEE2DBA9F81A9C98" box="[275,359,402,429]" captionStart="FIGURES 5558" captionStartId="31.[151,264,1346,1370]" captionTargetBox="[153,1428,185,1318]" captionTargetId="figure-18@31.[151,1436,179,1321]" captionTargetPageId="31" captionText="FIGURES 5558. Pseudagrion nigrihumerale, female paratype: (55) habitus lateral, (56) head dorsal, (57) pronotum, (58) ovipositor lateral. Photos: M. Seehausen." pageId="30" pageNumber="31">Fig. 56</figureCitation>
): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow-orange; postfrons dorsally with a small brownish dot in centre; yellow-orange colour of postfrons sharply separated from vertex at level of anterior margin of lateral ocelli; vertex black; median ocellus with a black spot posteriorly; postocular spots yellow-orange, bordered by black; small orange occipital bar; rear of head black, but yellowish along the eye margins; base and scape of antennae yellow, following segments ochre-brownish.
</paragraph>
<paragraph id="86416071F74EFFEFFF36D87DFA299EFC" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36D87DF8659F54" box="[199,280,583,609]" italics="true" pageId="30" pageNumber="31">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F74EFFEFFED9D87DF8D19F54" box="[296,428,582,609]" captionStart="FIGURES 5558" captionStartId="31.[151,264,1346,1370]" captionTargetBox="[153,1428,185,1318]" captionTargetId="figure-18@31.[151,1436,179,1321]" captionTargetPageId="31" captionText="FIGURES 5558. Pseudagrion nigrihumerale, female paratype: (55) habitus lateral, (56) head dorsal, (57) pronotum, (58) ovipositor lateral. Photos: M. Seehausen." pageId="30" pageNumber="31">Figs 55, 57</figureCitation>
): Anterior lobe of pronotum yellowish, expanding in centre subtriangular towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large yellowish patch in the middle, and on either side a yellowish spot dorsolaterally, fusing laterally with yellowish of the anterior lobe; propleuron black above, yellowish below and to rear; posterior lobe yellowish; apical edge of posterior lobe convex; bearing two yellowish horns directed towards head, approximately reaching the centre of the middle lobe. Pterothorax orange with posterior margins black; a broad black middorsal stripe; a broad black humeral stripe, narrowed posteriorly; metapleural suture with a small elongated black patch at posterior third; mesinfraepisternum with two-thirds washed-out brownish-black, in line with the black of humeral stripe, lower third of mesinfraepisternum yellowish-ochre; venter yellowish, may be slightly covered with whitish pruinosity. Coxae and legs pale yellowish-ochre; outer side of femora with a washed-out blackish stripe along posterior half; tibiae with blackish along the black spines; tip of tarsi and tarsal claws ochre, washed-out blackish along waists.
</paragraph>
<paragraph id="86416071F74EFFEFFF36D9E9F8809924" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36D9E9F8769ED9" box="[199,267,979,1004]" italics="true" pageId="30" pageNumber="31">Wings</emphasis>
(
<figureCitation id="1EC57CF4F74EFFEFFEE8D9E9F8169ED8" box="[281,363,978,1005]" captionStart="FIGURES 5558" captionStartId="31.[151,264,1346,1370]" captionTargetBox="[153,1428,185,1318]" captionTargetId="figure-18@31.[151,1436,179,1321]" captionTargetPageId="31" captionText="FIGURES 5558. Pseudagrion nigrihumerale, female paratype: (55) habitus lateral, (56) head dorsal, (57) pronotum, (58) ovipositor lateral. Photos: M. Seehausen." pageId="30" pageNumber="31">Fig. 55</figureCitation>
): Membrane hyaline; veins brown to black; 15.516.5 px in forewing; 14.015.5 px in hindwings; Pt brownish, overlying one cell.
</paragraph>
<paragraph id="86416071F74EFFEFFF36DE21FBAB99DC" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DE21F84C9900" box="[199,305,1051,1077]" italics="true" pageId="30" pageNumber="31">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F74EFFEFFEB2DE21F8B69900" box="[323,459,1050,1077]" captionStart="FIGURES 5558" captionStartId="31.[151,264,1346,1370]" captionTargetBox="[153,1428,185,1318]" captionTargetId="figure-18@31.[151,1436,179,1321]" captionTargetPageId="31" captionText="FIGURES 5558. Pseudagrion nigrihumerale, female paratype: (55) habitus lateral, (56) head dorsal, (57) pronotum, (58) ovipositor lateral. Photos: M. Seehausen." pageId="30" pageNumber="31">Figs 55, 58</figureCitation>
): S12 orange with black markings dorsally; S38 dorsally black, laterally and each basal annulus pale yellowish; S9 dorsally black, laterally yellow-orange, this colour expanded dorsally in centre of segment, leaving anterior and posterior margin black; S10 black. Cercus in lateral view straight and tapered, washed-out brownish-black, half as long as S10; paraproct yellow-brownish, short, in lateral view rounded; ovipositor yellowish, almost reaching to the posterior end of S10, two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F74EFFEFFF36DEC9FAFE9804" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">Measurements (mm): Total length (with appendages) 41.341.9; abdomen length (without appendages) 34.2 34.5; hindwing 24.024.8; forewing 25.426.4; Pt in forewing 0.9.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74EFFEFFF36DF00FB439A1C" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="86416071F74EFFEFFF36DF00F8479861" blockId="30.[151,1437,150,1905]" box="[199,314,1338,1364]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DF00F8479861" bold="true" box="[199,314,1338,1364]" pageId="30" pageNumber="31">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F74EFFEFFF36DF65FB439A1C" blockId="30.[151,1437,150,1905]" pageId="30" pageNumber="31">
Both sexes of
<taxonomicName id="41FE1BF2F74EFFEFFE9BDF65FB5A984C" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[362,551,1375,1401]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F74EFFEFFE9BDF65FB5A984C" box="[362,551,1375,1401]" italics="true" pageId="30" pageNumber="31">P. nigrihumerale</emphasis>
</taxonomicName>
are separated from all adjacent species and also most congeners in this group by the obvious thorax marking, bearing a broad black humeral stripe (
<figureCitation id="1EC57CF4F74EFFEFFC53DFB9FA8498A8" box="[930,1017,1411,1437]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="30" pageNumber="31">Figs 49</figureCitation>
,
<figureCitation id="1EC57CF4F74EFFEFFBF5DFB8FD5F98A8" box="[1028,1058,1410,1437]" captionStart="FIGURES 5558" captionStartId="31.[151,264,1346,1370]" captionTargetBox="[153,1428,185,1318]" captionTargetId="figure-18@31.[151,1436,179,1321]" captionTargetPageId="31" captionText="FIGURES 5558. Pseudagrion nigrihumerale, female paratype: (55) habitus lateral, (56) head dorsal, (57) pronotum, (58) ovipositor lateral. Photos: M. Seehausen." pageId="30" pageNumber="31">55</figureCitation>
,
<figureCitation id="1EC57CF4F74EFFEFFBDCDFB8FD2698A8" box="[1069,1115,1410,1437]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="30" pageNumber="31">105</figureCitation>
). Only three species, namely the Sulawesian
<taxonomicName id="41FE1BF2F74EFFEFFEBDDF9DF8BE98F5" authorityName="Selys" authorityYear="1876" box="[332,451,1447,1472]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="crocops">
<emphasis id="B48ABC63F74EFFEFFEBDDF9DF8BE98F5" box="[332,451,1447,1472]" italics="true" pageId="30" pageNumber="31">P. crocops</emphasis>
</taxonomicName>
,
<taxonomicName id="41FE1BF2F74EFFEFFE22DF9DFB0A98F4" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[467,631,1447,1473]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F74EFFEFFE22DF9DFB0A98F4" box="[467,631,1447,1473]" italics="true" pageId="30" pageNumber="31">P. halmaherae</emphasis>
</taxonomicName>
from Halmahera, and
<taxonomicName id="41FE1BF2F74EFFEFFC8EDF9DFA8098F4" authorityName="Muller" authorityYear="1996" box="[895,1021,1447,1473]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="buenafei">
<emphasis id="B48ABC63F74EFFEFFC8EDF9DFA8098F4" box="[895,1021,1447,1473]" italics="true" pageId="30" pageNumber="31">P. buenafei</emphasis>
</taxonomicName>
from Mindanao,
<collectingCountry id="FEE920E1F74EFFEFFB38DF9DFC3A98F4" box="[1225,1351,1447,1473]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
have a similar broad black humeral marking. Apart from their distant range and the morphological differences in the caudal appendages (compare
<figureCitation id="1EC57CF4F74EFFEFFE67DFD5FA359B3C" box="[406,840,1519,1545]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="30" pageNumber="31">Figs 131ac, 133ac, 137ac, 139ac</figureCitation>
), the males of the latter three species differ from
<taxonomicName id="41FE1BF2F74EFFEFFA79DFD5F84A9B18" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F74EFFEFFA79DFD5F84A9B18" italics="true" pageId="30" pageNumber="31">P. nigrihumerale</emphasis>
</taxonomicName>
in their colour pattern as well. Beside the name-giving black humeral marking,
<taxonomicName id="41FE1BF2F74EFFEFFB2FDC29FCE19B18" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[1246,1436,1555,1581]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F74EFFEFFB2FDC29FCE19B18" box="[1246,1436,1555,1581]" italics="true" pageId="30" pageNumber="31">P. nigrihumerale</emphasis>
</taxonomicName>
can be separated from its Andaman congener
<taxonomicName id="41FE1BF2F74EFFEFFD6CDC0DFA719B65" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[669,780,1591,1616]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F74EFFEFFD6CDC0DFA719B65" box="[669,780,1591,1616]" italics="true" pageId="30" pageNumber="31">P. igneum</emphasis>
</taxonomicName>
by having S12 dorsally red and S10 entirely black (
<figureCitation id="1EC57CF4F74EFFEFFA9BDC0DF9C89B40" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="30" pageNumber="31">Figs 43</figureCitation>
,
<figureCitation id="1EC57CF4F74EFFEFFF30DC61F99D9B40" box="[193,224,1627,1653]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="30" pageNumber="31">49</figureCitation>
,
<figureCitation id="1EC57CF4F74EFFEFFF1ADC61F8DD9B40" box="[235,416,1627,1653]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="30" pageNumber="31">121a, b, 122a, b</figureCitation>
), by having a black vertex and by lacking red colour on the postfrons (
<figureCitation id="1EC57CF4F74EFFEFFB48DC61FC6D9B40" box="[1209,1296,1627,1653]" captionStart="FIGURES 4348" captionStartId="27.[151,264,1359,1383]" captionTargetBox="[153,1433,185,1329]" captionTargetId="figure-18@27.[151,1436,179,1333]" captionTargetPageId="27" captionText="FIGURES 4348. Pseudagrion igneum, holotype male: (43) habitus lateral, (44) appendages lateral, (45) appendages dorsal, (46) cercus dorsolateral, (47) head dorsal, (48) pronotum. Photos: M. Seehausen." pageId="30" pageNumber="31">Figs 47</figureCitation>
,
<figureCitation id="1EC57CF4F74EFFEFFAEDDC60FC479B40" box="[1308,1338,1626,1653]" captionStart="FIGURES 4954" captionStartId="29.[151,264,1957,1981]" captionTargetBox="[153,1391,491,1928]" captionTargetId="figure-131@29.[151,1436,486,1931]" captionTargetPageId="29" captionText="FIGURES 4954. Pseudagrion nigrihumerale, holotype male: (49) habitus lateral, (50) appendages lateral, (51) appendages dorsal, (52) cercus dorsolateral, (53) head dorsal, (54) pronotum. Photos: M. Seehausen." pageId="30" pageNumber="31">53</figureCitation>
,
<figureCitation id="1EC57CF4F74EFFEFFAB7DC61FCF29B40" box="[1350,1423,1627,1653]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="30" pageNumber="31">87, 88</figureCitation>
), whereas the male appendages of both species are identical (
<figureCitation id="1EC57CF4F74EFFEFFCDFDC45FD679BAC" box="[814,1050,1663,1689]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="30" pageNumber="31">Figs 138ac, 139ac</figureCitation>
). The southerly distributed species
<taxonomicName id="41FE1BF2F74EFFEFFF66DC99F8569B88" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[151,299,1699,1725]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F74EFFEFFF66DC99F8569B88" box="[151,299,1699,1725]" italics="true" pageId="30" pageNumber="31">P. simalurum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F74EFFEFFE92DC99F8909B88" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[355,493,1699,1725]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F74EFFEFFE92DC99F8909B88" box="[355,493,1699,1725]" italics="true" pageId="30" pageNumber="31">P. obscurum</emphasis>
</taxonomicName>
differ by the same colour markings as listed for
<taxonomicName id="41FE1BF2F74EFFEFFBE5DC99FDFF9B89" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[1044,1154,1699,1724]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F74EFFEFFBE5DC99FDFF9B89" box="[1044,1154,1699,1724]" italics="true" pageId="30" pageNumber="31">P. igneum</emphasis>
</taxonomicName>
, but also very slightly in the male appendages: the cercus of
<taxonomicName id="41FE1BF2F74EFFEFFDDADCFDFB9A9BD4" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[555,743,1735,1761]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nigrihumerale">
<emphasis id="B48ABC63F74EFFEFFDDADCFDFB9A9BD4" box="[555,743,1735,1761]" italics="true" pageId="30" pageNumber="31">P. nigrihumerale</emphasis>
</taxonomicName>
bears a small subbasal tooth situated at the proximal third of the inferior lobe, whereas in
<taxonomicName id="41FE1BF2F74EFFEFFE28DCD1FB119A30" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[473,620,1771,1797]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F74EFFEFFE28DCD1FB119A30" box="[473,620,1771,1797]" italics="true" pageId="30" pageNumber="31">P. simalurum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F74EFFEFFD53DCD1FA519A30" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[674,812,1771,1797]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F74EFFEFFD53DCD1FA519A30" box="[674,812,1771,1797]" italics="true" pageId="30" pageNumber="31">P. obscurum</emphasis>
</taxonomicName>
there is a sharp edged bulge that only looks like a tooth in dorsal view (
<figureCitation id="1EC57CF4F74EFFEFFEB4DD35FB4F9A1C" box="[325,562,1807,1833]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="30" pageNumber="31">Figs 140ac, 141ac</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF74EFFEFFF36DD08FA2F9A44" box="[199,850,1842,1905]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="86416071F74EFFEFFF36DD08F8249A79" blockId="30.[151,1437,150,1905]" box="[199,345,1842,1868]" pageId="30" pageNumber="31">
<emphasis id="B48ABC63F74EFFEFFF36DD08F8249A79" bold="true" box="[199,345,1842,1868]" pageId="30" pageNumber="31">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F74EFFEFFF36DD6DFA2F9A44" blockId="30.[151,1437,150,1905]" box="[199,850,1879,1905]" pageId="30" pageNumber="31">
Apparently endemic to the
<collectingCountry id="FEE920E1F74EFFEFFE09DD6DFBD69A44" box="[504,683,1879,1905]" name="India" pageId="30" pageNumber="31">Nicobar Islands</collectingCountry>
,
<collectingCountry id="FEE920E1F74EFFEFFD46DD6DFB8C9A44" box="[695,753,1879,1905]" name="India" pageId="30" pageNumber="31">India</collectingCountry>
(
<figureCitation id="1EC57CF4F74EFFEFFD0EDD6DFA389A44" box="[767,837,1879,1905]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="30" pageNumber="31">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,442 @@
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<mods:title id="C757DA6524CEB292D628B361780D16F6">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:affiliation id="2716A570397609AB6F8DC85ABB18AF00">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
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<heading id="DD09D71DF74FFFEEFF66DFEAFB9798DE" bold="true" box="[151,746,1488,1515]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
<emphasis id="B48ABC63F74FFFEEFF66DFEAFB9798DE" bold="true" box="[151,746,1488,1515]" pageId="31" pageNumber="32">
<taxonomicName id="41FE1BF2F74FFFEEFF66DFEAFB0E98DE" authority="Lieftinck, 1936" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[151,627,1488,1515]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obscurum" status="stat. nov.">
<emphasis id="B48ABC63F74FFFEEFF66DFEAF8CC98DE" bold="true" box="[151,433,1488,1515]" italics="true" pageId="31" pageNumber="32">Pseudagrion obscurum</emphasis>
<bibRefCitation id="E26F1D80F74FFFEEFE49DFEBFB0E98DE" author="Lieftinck, M. A." box="[440,627,1489,1515]" pageId="31" pageNumber="32" pagination="99 - 160" refId="ref40810" refString="Lieftinck, M. A. (1936) Die Odonaten der Kleinen Sunda-Inseln. Revue Suisse de Zoologie, 43, 99-160. https://doi.org/10.5962/bhl.part.145462" type="journal article" year="1936">Lieftinck, 1936</bibRefCitation>
</taxonomicName>
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Figures: 5960 (
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, envelope); 89a, b (head
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); 106 (pterothorax
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); 123a, b (abdomen
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); 140ac (caudal appendages
<specimenCount id="90F8ABF8F74FFFEEFF36DC0EF9A19B79" box="[199,220,1588,1612]" count="1" pageId="31" pageNumber="32" type="male"></specimenCount>
).
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<taxonomicName id="41FE1BF2F74FFFEEFF66DC69F8869B5E" ID-CoL="7HWZT" authorityName="Lieftinck" authorityYear="1936" box="[151,507,1619,1643]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="pilidorsum" subSpecies="obscurum">
<emphasis id="B48ABC63F74FFFEEFF66DC69F8869B5E" box="[151,507,1619,1643]" italics="true" pageId="31" pageNumber="32">Pseudagrion pilidorsum obscurum</emphasis>
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:
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<bibRefCitation id="E26F1D80F74FFFEEFDFADC69FBCA9B5E" author="Lieftinck, M. A." box="[523,695,1619,1643]" pageId="31" pageNumber="32" pagination="99 - 160" refId="ref40810" refString="Lieftinck, M. A. (1936) Die Odonaten der Kleinen Sunda-Inseln. Revue Suisse de Zoologie, 43, 99-160. https://doi.org/10.5962/bhl.part.145462" type="journal article" year="1936">Lieftinck (1936)</bibRefCitation>
: 128
</treatmentCitation>
, 131132,
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(key; description
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<specimenCount id="90F8ABF8F74FFFEEFB7FDC69FDDF9B5F" box="[1166,1186,1619,1642]" count="1" pageId="31" pageNumber="32" type="female"></specimenCount>
;
<specimenCount id="90F8ABF8F74FFFEEFB40DC69FDB99B5F" box="[1201,1220,1619,1642]" count="1" pageId="31" pageNumber="32" type="male"></specimenCount>
appendages; Nias &amp; probably Simalur).
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<emphasis id="B48ABC63F74FFFEEFF66DC8DFB0A9BE4" bold="true" box="[151,631,1719,1746]" pageId="31" pageNumber="32">
Name-bearing
<typeStatus id="5945DED3F74FFFEEFEB8DC82F8019BE7" box="[329,380,1720,1746]" pageId="31" pageNumber="32">type</typeStatus>
specimens examined
</emphasis>
(
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)
</paragraph>
<paragraph id="86416071F74FFFEEFF66DCE6FCE19A2C" blockId="31.[151,1436,1719,1890]" pageId="31" pageNumber="32">
<materialsCitation id="36966A2CF74FFFEEFF66DCE6FADB9A2F" collectingDate="1931-09-10" collectionCode="RMNH" collectorName="J.P. Caliman &amp; M. Mohr" country="Indonesia" location="Nias Island" pageId="31" pageNumber="32" specimenCount="3" specimenCount-male="2" typeStatus="syntype">
<specimenCount id="90F8ABF8F74FFFEEFF66DCE6F9B29BC3" box="[151,207,1756,1782]" count="2" pageId="31" pageNumber="32" type="male">2♂♂</specimenCount>
<typeStatus id="5945DED3F74FFFEEFF27DCE6F8469BC3" box="[214,315,1756,1782]" pageId="31" pageNumber="32" type="syntype">syntypes</typeStatus>
,
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<collectingDate id="E204BF59F74FFFEEFEB7DCE1F8BD9BC0" box="[326,448,1755,1781]" pageId="31" pageNumber="32" value="1931-09-10">10.ix.1931</collectingDate>
</date>
, Amadraja,
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,
<collectingCountry id="FEE920E1F74FFFEEFD25DCE1FA399BC0" box="[724,836,1755,1781]" name="Indonesia" pageId="31" pageNumber="32">Indonesia</collectingCountry>
,
<collectorName id="2B0B05A7F74FFFEEFCBFDCE6FA079BC3" box="[846,890,1756,1782]" name="J.P. Caliman" pageId="31" pageNumber="32">J.C.</collectorName>
v. d.
<collectorName id="2B0B05A7F74FFFEEFC46DCE6FD619BC0" box="[951,1052,1755,1782]" pageId="31" pageNumber="32">M. Mohr</collectorName>
leg.,
<collectionCode id="E0EFF8B4F74FFFEEFBABDCE6FDCC9BC3" box="[1114,1201,1756,1782]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="31" pageNumber="32" type="Museum">RMNH</collectionCode>
, JvT no. 2839.
<bibRefCitation id="E26F1D80F74FFFEEFF36DCC5F8C09A2F" author="Lieftinck, M. A." box="[199,445,1791,1818]" pageId="31" pageNumber="32" pagination="65 - 139" refId="ref40982" refString="Lieftinck, M. A. (1971) A catalogue of the type-specimens of Odonata preserved in the Netherlands. With a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65-139, pls. 1 - 7." type="journal article" year="1971">Lieftinck (1971: 103)</bibRefCitation>
designated a
<typeStatus id="5945DED3F74FFFEEFDAFDCC5FBB89A2C" box="[606,709,1791,1817]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
male, but there are
</materialsCitation>
<specimenCount id="90F8ABF8F74FFFEEFC41DD3AFD599A2C" box="[944,1060,1791,1818]" count="2" pageId="31" pageNumber="32" type="male">two males</specimenCount>
in the respective envelope (Figs
</paragraph>
<paragraph id="86416071F74FFFEEFF66DD19FCE19A0B" blockId="31.[151,1436,1719,1890]" box="[151,1436,1827,1854]" pageId="31" pageNumber="32">
5960). This
<typeStatus id="5945DED3F74FFFEEFEC1DD19F8EA9A08" box="[304,407,1827,1853]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
designation is not valid because he has not explicitly designated a single specimen (ICZN
</paragraph>
<paragraph id="86416071F74FFFEEFF66DD7DFC469A57" blockId="31.[151,1436,1719,1890]" box="[151,1339,1863,1890]" pageId="31" pageNumber="32">
74.5). We do not consider it necessary to designate a
<typeStatus id="5945DED3F74FFFEEFD1BDD7DFA299A54" box="[746,852,1863,1889]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
, thus we propose to keep them as
<typeStatus id="5945DED3F74FFFEEFB20DD72FC4A9A57" box="[1233,1335,1864,1890]" pageId="31" pageNumber="32" type="syntype">syntypes</typeStatus>
.
</paragraph>
<caption id="D28130F9F770FFD1FF66DEA7FD869980" box="[151,1275,1181,1206]" pageId="32" pageNumber="33" startId="32.[151,264,1181,1205]" targetBox="[155,1435,182,1151]" targetPageId="32" targetType="figure">
<paragraph id="86416071F770FFD1FF66DEA7FD869980" blockId="32.[151,1275,1181,1206]" box="[151,1275,1181,1206]" pageId="32" pageNumber="33">
<emphasis id="B48ABC63F770FFD1FF66DEA7F8309983" bold="true" box="[151,333,1181,1206]" pageId="32" pageNumber="33">FIGURES 5960</emphasis>
.
<taxonomicName id="41FE1BF2F770FFD1FEABDEA4FB3B9983" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[346,582,1182,1206]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F770FFD1FEABDEA4FB3B9983" box="[346,582,1182,1206]" italics="true" pageId="32" pageNumber="33">Pseudagrion obscurum</emphasis>
</taxonomicName>
, syntype males: (
<emphasis id="B48ABC63F770FFD1FD08DEA4FA6F9983" bold="true" box="[761,786,1182,1206]" pageId="32" pageNumber="33">59</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F770FFD1FC34DEA4FAA39983" bold="true" box="[965,990,1182,1206]" pageId="32" pageNumber="33">60</emphasis>
) envelope. Photos: RMNH.
</paragraph>
</caption>
<paragraph id="86416071F770FFD1FF36DF32FBF19817" blockId="32.[151,1437,1288,2034]" box="[199,652,1288,1314]" pageId="32" pageNumber="33">
<emphasis id="B48ABC63F770FFD1FF36DF32FB749817" bold="true" box="[199,521,1288,1314]" pageId="32" pageNumber="33">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F770FFD1FDE9DF32FB2E9817" box="[536,595,1288,1314]" count="2" pageId="32" pageNumber="33" type="male">2♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F770FFD1FDAEDF32FBFD9817" box="[607,640,1288,1314]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
)
</paragraph>
<paragraph id="86416071F770FFD1FF36DF16FBDA985F" blockId="32.[151,1437,1288,2034]" pageId="32" pageNumber="33">
<materialsCitation id="36966A2CF770FFD1FF36DF16FBD9985F" collectingDate="1972-04-21" collectingDateMax="1972-05-02" collectingDateMin="1972-04-21" collectionCode="ZFMK" collectorName="Hilisimaetanoe &amp; Roesler &amp; Kuppers" country="Indonesia" elevation="200" location="Nias Island" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Nias Island">
<emphasis id="B48ABC63F770FFD1FF36DF16F8A89873" bold="true" box="[199,469,1324,1350]" pageId="32" pageNumber="33">
<collectingCountry id="FEE920E1F770FFD1FF36DF16F8409873" box="[199,317,1324,1350]" name="Indonesia" pageId="32" pageNumber="33">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F770FFD1FEBDDF16F8A89873" box="[332,469,1324,1350]" pageId="32" pageNumber="33">Nias Island</collectingRegion>
</emphasis>
:
<specimenCount id="90F8ABF8F770FFD1FE14DF16FB5D9873" box="[485,544,1324,1350]" count="2" pageId="32" pageNumber="33" type="male">2♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F770FFD1FDDCDF16FB2D9873" box="[557,592,1324,1350]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
,
<date id="F24046B1F770FFD1FD90DF16FA729873" box="[609,783,1324,1350]" pageId="32" pageNumber="33" value="1972-04-21" valueMax="1972-05-02" valueMin="1972-04-21">
<collectingDate id="E204BF59F770FFD1FD90DF16FA729873" box="[609,783,1324,1350]" pageId="32" pageNumber="33" value="1972-04-21" valueMax="1972-05-02" valueMin="1972-04-21">21.iv.2.v.1972</collectingDate>
</date>
, surroundings of
<collectorName id="2B0B05A7F770FFD1FC29DF16FDF99873" box="[984,1156,1324,1350]" pageId="32" pageNumber="33">Hilisimaetanoe</collectorName>
,
<elevation id="0DD38742F770FFD1FB63DF16FC649873" box="[1170,1305,1324,1350]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="200.0">
<quantity id="4106CD94F770FFD1FB63DF16FDA69873" box="[1170,1243,1324,1350]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F770FFD1FAD9DF16FC039873" box="[1320,1406,1324,1350]" pageId="32" pageNumber="33">Roesler</collectorName>
&amp;
<collectorName id="2B0B05A7F770FFD1FF66DF6AF98B985F" box="[151,246,1360,1386]" pageId="32" pageNumber="33">Küppers</collectorName>
leg.,
<collectionCode id="E0EFF8B4F770FFD1FEC5DF6AF8F8985F" box="[308,389,1360,1386]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="32" pageNumber="33" type="Museum">ZFMK</collectionCode>
, no. 2009/1142.11142.3
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F770FFD1FF36DF4EF88F98BB" blockId="32.[151,1437,1288,2034]" box="[199,498,1396,1422]" pageId="32" pageNumber="33">
<emphasis id="B48ABC63F770FFD1FF36DF4EF88F98BB" bold="true" box="[199,498,1396,1422]" pageId="32" pageNumber="33">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F770FFD1FF36DFA2FD079B9B" blockId="32.[151,1437,1288,2034]" pageId="32" pageNumber="33">
<emphasis id="B48ABC63F770FFD1FF36DFA2F8799887" box="[199,260,1432,1458]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
(
<figureCitation id="1EC57CF4F770FFD1FEE5DFA2F8EF9887" box="[276,402,1432,1458]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="32" pageNumber="33">Fig. 89a, b</figureCitation>
): Labrum, clypeus, antefrons and postfrons yellow-orange; postclypeus with two elongated black spots, and a black centre along posterior edge; centre of frons may have a small darkish spot; in
<specimenCount id="90F8ABF8F770FFD1FAF5DF86FCE198E3" box="[1284,1436,1468,1494]" count="1" pageId="32" pageNumber="33" type="generic">one specimen</specimenCount>
the postclypeus is completely black (
<figureCitation id="1EC57CF4F770FFD1FDC8DFDAFBE098CF" box="[569,669,1504,1530]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="32" pageNumber="33">Fig. 89b</figureCitation>
); yellow-orange colour of the postfrons is separated from black and red colour of the vertex at level of anterior margin of lateral ocelli; a black spot at posterior margin of median ocellus; a small reddish spot is situated in centre of vertex around level of posterior margin of lateral ocelli; occipital bar reddish; postocular spots washed-out dark reddish, comparatively small due to obscurely blackish margins; broadly surrounded by black; rear of head black, but may be yellow along the eye margins; base of antennae yellow, scape of antennae anteriorly yellow, but posteriorly black, following segments brownish.
</paragraph>
<paragraph id="86416071F770FFD1FF36DC82FC0F9AC7" blockId="32.[151,1437,1288,2034]" pageId="32" pageNumber="33">
<emphasis id="B48ABC63F770FFD1FF36DC82F8659BE7" box="[199,280,1720,1746]" italics="true" pageId="32" pageNumber="33">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F770FFD1FED6DC82F8009BE6" box="[295,381,1720,1747]" captionStart="FIGURES 5960" captionStartId="32.[151,264,1181,1205]" captionTargetBox="[155,1435,182,1151]" captionTargetId="figure-18@32.[151,1436,179,1155]" captionTargetPageId="32" captionText="FIGURES 5960. Pseudagrion obscurum, syntype males: (59) habitus lateral, (60) envelope. Photos: RMNH." pageId="32" pageNumber="33">Figs 59</figureCitation>
,
<figureCitation id="1EC57CF4F770FFD1FE78DC82F8C59BE7" box="[393,440,1720,1746]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="32" pageNumber="33">106</figureCitation>
): Anterior lobe of pronotum yellow, this colour expanding in centre towards middle lobe; margins of anterior lobe black; middle pronotal lobe black, with a centrally divided red patch in the middle, and on either side a red patch dorsolaterally; propleuron black above, yellowish below and to rear; posterior lobe yellowreddish, with its margins black; apical edge of posterior lobe convex. Pterothorax red; middorsal carina black; dorsum obscurely blackish, almost reaching to humeral suture; humeral suture with an elongated black patch at its posterior quarter,
<specimenCount id="90F8ABF8F770FFD1FEAFDD56F8BE9AB3" box="[350,451,1900,1926]" count="1" pageId="32" pageNumber="33" type="male">one male</specimenCount>
has also the anterior humeral suture thin black; interpleural suture with a black stripe at posterior quarter, and a black spot anteriorly; a black patch at posterior quarter of metapleural suture; a large black marking covering at least superior two-thirds of the mesinfraepisternum; venter pale, may be covered with whitish pruinosity. Legs pale, with outside of the femora obscurely blackish; tibiae blackish along spines; spines black.
</paragraph>
<paragraph id="86416071F771FFD0FF36DAADF88F9DCC" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36DAADF84C9D84" box="[199,305,151,177]" italics="true" pageId="33" pageNumber="34">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F771FFD0FEB0DAADF8E79D84" box="[321,410,150,177]" captionStart="FIGURES 5960" captionStartId="32.[151,264,1181,1205]" captionTargetBox="[155,1435,182,1151]" captionTargetId="figure-18@32.[151,1436,179,1155]" captionTargetPageId="32" captionText="FIGURES 5960. Pseudagrion obscurum, syntype males: (59) habitus lateral, (60) envelope. Photos: RMNH." pageId="33" pageNumber="34">Figs 59</figureCitation>
,
<figureCitation id="1EC57CF4F771FFD0FE59DAADF8829D84" box="[424,511,151,177]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="33" pageNumber="34">123a, b</figureCitation>
): S12 dorsally black, laterally red; S38 dorsally black, laterally and each basal annulus yellow-orange; S9 red, with its posterior third to half black; S10 dorsally with black “x”-like marking, variable in shape; laterally red.
</paragraph>
<paragraph id="86416071F771FFD0FF36DB39FA289CC0" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36DB39F8DA9C28" box="[199,423,259,285]" italics="true" pageId="33" pageNumber="34">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F771FFD0FE47DB39FB389C28" box="[438,581,259,285]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="33" pageNumber="34">Figs 140ac</figureCitation>
): Cercus in lateral view darkish, shorter than S10, inferior lobe appears very slightly shorter than superior lobe; in lateral view cercus subrectangular with a shallow inter-lobe notch situated low on the apex; in dorsal view the terminus of the superior lobe of the cercus curved inwards; a small interior projection is situated around the centre of cercus; a small sharp-edged bulge-like interior projection before half-length on the inferior lobe of the cercus, visible in dorsal and dorsolateral views; paraproct pale brownish-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; bearing a small black tooth at medial edge.
</paragraph>
<paragraph id="86416071F771FFD0FF36DBC4FB7A9F2D" blockId="33.[151,1437,150,1365]" box="[199,519,510,536]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36DBC4FB7A9F2D" bold="true" box="[199,519,510,536]" pageId="33" pageNumber="34">Characterization of female</emphasis>
</paragraph>
<paragraph id="86416071F771FFD0FF36D819FA919F08" blockId="33.[151,1437,150,1365]" box="[199,1004,547,573]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36D819F8799F08" box="[199,260,547,573]" italics="true" pageId="33" pageNumber="34">Head</emphasis>
: With dark markings as in male, but primary colour orange-ochre.
</paragraph>
<paragraph id="86416071F771FFD0FF36D87DF8459FF8" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36D87DF8659F54" box="[199,280,583,609]" italics="true" pageId="33" pageNumber="34">Thorax</emphasis>
: Pronotum markings as in male, but primary colour yellow-orange; posterior lobe of pronotum with two black horns directed towards head, almost reaching the centre of the middle lobe. Pterothorax entirely reddish; humeral suture with an elongated black patch at posterior quarter; black markings at interpleural and metapleural suture narrow.
</paragraph>
<paragraph id="86416071F771FFD0FF36D8EDFA9C9E68" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36D8EDF84C9FC4" box="[199,305,727,753]" italics="true" pageId="33" pageNumber="34">Abdomen</emphasis>
: S110 dorsally black, laterally pale yellow-ochre; S9 pale yellow-ochre laterally, this expanded dorsally in centre; S10 with pale yellowish-ochre middorsal marking. Cercus blackish; paraproct brownish, in lateral view rounded; ovipositor pale yellowish-ochre, not reaching posterior margin of S10; two brownish styles reaching to beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F771FFD0FF36D95DFD289EB4" blockId="33.[151,1437,150,1365]" box="[199,1109,870,897]" pageId="33" pageNumber="34">Wings of both sexes with 12.514.5 px in forewing, and 11.513 px in hindwing.</paragraph>
<paragraph id="86416071F771FFD0FF36D9B1FC089EFC" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">Measurements of both sexes (mm): Total length (with appendages) 37.844.3; abdomen length (without appendages) 32.036.0; hindwing 21.824.5; forewing 22.927.5; Pt in forewing 0.81.0; male cercus 0.50.6.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF771FFD0FF36D9E9F83D996C" pageId="33" pageNumber="34" type="discussion">
<paragraph id="86416071F771FFD0FF36D9E9F8499ED8" blockId="33.[151,1437,150,1365]" box="[199,308,979,1005]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36D9E9F8499ED8" bold="true" box="[199,308,979,1005]" pageId="33" pageNumber="34">Remarks</emphasis>
</paragraph>
<paragraph id="86416071F771FFD0FF36D9CDF83D996C" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">
The single female has an entirely reddish pterothorax and lacks the name-giving obscurely blackish dorsum, which is present in all
<specimenCount id="90F8ABF8F771FFD0FE66DE21FB739900" box="[407,526,1051,1077]" count="4" pageId="33" pageNumber="34" type="male">four males</specimenCount>
examined. It is not clear if this obscuring is just due to older age in the males or gender-related.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF771FFD0FF36DE58FD159860" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="86416071F771FFD0FF36DE58F8479949" blockId="33.[151,1437,150,1365]" box="[199,314,1122,1148]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF36DE58F8479949" bold="true" box="[199,314,1122,1148]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F771FFD0FF36DEBDFD159860" blockId="33.[151,1437,150,1365]" pageId="33" pageNumber="34">
<taxonomicName id="41FE1BF2F771FFD0FF36DEBDF8B79994" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[199,458,1159,1185]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F771FFD0FF36DEBDF8B79994" box="[199,458,1159,1185]" italics="true" pageId="33" pageNumber="34">Pseudagrion obscurum</emphasis>
</taxonomicName>
belongs to the group of endemic taxa of the island chain west of
<collectingRegion id="443AAE93F771FFD0FB7CDEBDFD939994" box="[1165,1262,1159,1185]" country="Indonesia" name="Sumatera Utara" pageId="33" pageNumber="34">Sumatra</collectingRegion>
. Due to scarcity of material we cannot give information on how to separate the females, but the males are separated from most congeners occurring on adjacent islands by having S12 dorsally black (
<figureCitation id="1EC57CF4F771FFD0FC39DEF5FD6299DC" box="[968,1055,1230,1257]" captionStart="FIGURES 5960" captionStartId="32.[151,264,1181,1205]" captionTargetBox="[155,1435,182,1151]" captionTargetId="figure-18@32.[151,1436,179,1155]" captionTargetPageId="32" captionText="FIGURES 5960. Pseudagrion obscurum, syntype males: (59) habitus lateral, (60) envelope. Photos: RMNH." pageId="33" pageNumber="34">Figs 59</figureCitation>
,
<figureCitation id="1EC57CF4F771FFD0FBDDDEF5FDFF99DC" box="[1068,1154,1231,1257]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="33" pageNumber="34">123a, b</figureCitation>
). However this character is shared by
<taxonomicName id="41FE1BF2F771FFD0FED2DEC9F8ED9839" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[291,400,1267,1292]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F771FFD0FED2DEC9F8ED9839" box="[291,400,1267,1292]" italics="true" pageId="33" pageNumber="34">P. igneum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F771FFD0FE6EDEC9FB579838" box="[415,554,1267,1293]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="33" pageNumber="34">Figs 121a, b</figureCitation>
) and
<taxonomicName id="41FE1BF2F771FFD0FD99DEC9FB869838" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[616,763,1267,1293]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F771FFD0FD99DEC9FB869838" box="[616,763,1267,1293]" italics="true" pageId="33" pageNumber="34">P. simalurum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F771FFD0FCF8DEC9FAEE9838" box="[777,915,1267,1293]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="33" pageNumber="34">Figs 124a, b</figureCitation>
)—both are very similar to
<taxonomicName id="41FE1BF2F771FFD0FB32DEC9FC319838" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[1219,1356,1267,1293]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F771FFD0FB32DEC9FC319838" box="[1219,1356,1267,1293]" italics="true" pageId="33" pageNumber="34">P. obscurum</emphasis>
</taxonomicName>
also in the colour of the head (
<figureCitation id="1EC57CF4F771FFD0FE6BDF2DFB109804" box="[410,621,1303,1329]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="33" pageNumber="34">Figs 87, 89a, b, 90</figureCitation>
), and the morphology of the male cercus (
<figureCitation id="1EC57CF4F771FFD0FBB4DF2DFDAC9804" box="[1093,1233,1303,1329]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="33" pageNumber="34">Figs 138ac</figureCitation>
,
<figureCitation id="1EC57CF4F771FFD0FB2DDF2DFCED9804" box="[1244,1424,1303,1329]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="33" pageNumber="34">140ac, 141ac</figureCitation>
). We summarise a combination of characters for separating these three species in table 2.
</paragraph>
</subSubSection>
<caption id="D28130F9F771FFD0FF69DFBEFDB398A9" box="[152,1230,1412,1436]" pageId="33" pageNumber="34" startId="33.[152,244,1412,1436]" targetType="table">
<paragraph id="86416071F771FFD0FF69DFBEFDB398A9" blockId="33.[152,1230,1412,1436]" box="[152,1230,1412,1436]" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF69DFBEF87298A9" bold="true" box="[152,271,1412,1436]" pageId="33" pageNumber="34">TABLE 2.</emphasis>
Compilation of characters separating males of
<taxonomicName id="41FE1BF2F771FFD0FCF6DFBFFA0D98A9" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[775,880,1413,1436]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F771FFD0FCF6DFBFFA0D98A9" box="[775,880,1413,1436]" italics="true" pageId="33" pageNumber="34">P. igneum</emphasis>
</taxonomicName>
,
<taxonomicName id="41FE1BF2F771FFD0FC8CDFBFFD7D98A9" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[893,1024,1412,1436]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F771FFD0FC8CDFBFFD7D98A9" box="[893,1024,1412,1436]" italics="true" pageId="33" pageNumber="34">P. obscurum</emphasis>
</taxonomicName>
, and
<taxonomicName id="41FE1BF2F771FFD0FBCADFBFFDBA98A9" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[1083,1223,1412,1436]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F771FFD0FBCADFBFFDBA98A9" box="[1083,1223,1412,1436]" italics="true" pageId="33" pageNumber="34">P. simalurum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="86416071F771FFD0FE15DF97FCF09A9F" pageId="33" pageNumber="34">
<table id="F4FE92D1F771000EFF52DF96FCF09A9F" box="[163,1421,1452,1962]" gridcols="4" gridrows="14" pageId="33" pageNumber="34">
<tr id="38CE6233F771000EFF52DF96FCF098F1" box="[163,1421,1452,1476]" gridrow="0" pageId="33" pageNumber="34" rowspan-0="1">
<th id="7B1F0B4FF771000EFE15DF96FA7898F1" box="[484,773,1452,1476]" gridcol="1" gridrow="0" pageId="33" pageNumber="34">
<taxonomicName id="41FE1BF2F771FFD0FE15DF97FB3198F1" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[484,588,1452,1476]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F771FFD0FE15DF97FB3198F1" bold="true" box="[484,588,1452,1476]" italics="true" pageId="33" pageNumber="34">P. igneum</emphasis>
</taxonomicName>
</th>
<th id="7B1F0B4FF771000EFCD4DF96FD3698F1" box="[805,1099,1452,1476]" gridcol="2" gridrow="0" pageId="33" pageNumber="34">
<taxonomicName id="41FE1BF2F771FFD0FCD4DF97FADB98F1" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[805,934,1452,1476]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F771FFD0FCD4DF97FADB98F1" bold="true" box="[805,934,1452,1476]" italics="true" pageId="33" pageNumber="34">P. obscurum</emphasis>
</taxonomicName>
</th>
<th id="7B1F0B4FF771000EFB97DF96FCF098F1" box="[1126,1421,1452,1476]" gridcol="3" gridrow="0" pageId="33" pageNumber="34">
<taxonomicName id="41FE1BF2F771FFD0FB97DF97FD8C98F1" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[1126,1265,1452,1476]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F771FFD0FB97DF97FD8C98F1" bold="true" box="[1126,1265,1452,1476]" italics="true" pageId="33" pageNumber="34">P. simalurum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="38CE6233F771000EFF52DFEFFCF098D8" box="[163,1421,1493,1517]" gridrow="1" pageId="33" pageNumber="34">
<th id="7B1F0B4FF771000EFF52DFEFF83498D8" box="[163,329,1493,1517]" gridcol="0" gridrow="1" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF52DFEFF99798D8" bold="true" box="[163,234,1493,1517]" pageId="33" pageNumber="34">Vertex</emphasis>
</th>
<td id="7B1F0B4FF771000EFE15DFEFFA7898D8" box="[484,773,1493,1517]" gridcol="1" gridrow="1" pageId="33" pageNumber="34">red, largely black along</td>
<td id="7B1F0B4FF771000EFCD4DFEFFD3698D8" box="[805,1099,1493,1517]" gridcol="2" gridrow="1" pageId="33" pageNumber="34">indistinct red, may be hardly</td>
<td id="7B1F0B4FF771000EFB97DFEFFCF098D8" box="[1126,1421,1493,1517]" gridcol="3" gridrow="1" pageId="33" pageNumber="34">red, less black along ocelli</td>
</tr>
<tr id="38CE6233F771000EFF52DFC3FCF09B24" box="[163,1421,1529,1553]" gridrow="2" pageId="33" pageNumber="34" rowspan-0="1">
<td id="7B1F0B4FF771000EFE15DFC3FA789B24" box="[484,773,1529,1553]" gridcol="1" gridrow="2" pageId="33" pageNumber="34">ocelli (Fig. 87)</td>
<td id="7B1F0B4FF771000EFCD4DFC3FD369B24" box="[805,1099,1529,1553]" gridcol="2" gridrow="2" pageId="33" pageNumber="34">visible, largely black along</td>
<td id="7B1F0B4FF771000EFB97DFC3FCF09B24" box="[1126,1421,1529,1553]" gridcol="3" gridrow="2" pageId="33" pageNumber="34">(Fig. 40)</td>
</tr>
<tr id="38CE6233F771000EFF52DC27FCF09B00" box="[163,1421,1565,1589]" gridrow="3" pageId="33" pageNumber="34" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="7B1F0B4FF771000EFCD4DC27FD369B00" box="[805,1099,1565,1589]" gridcol="2" gridrow="3" pageId="33" pageNumber="34">ocelli (Fig. 89a, b)</td>
</tr>
<tr id="38CE6233F771000EFF52DC7FFCF09B6B" box="[163,1421,1605,1630]" gridrow="4" pageId="33" pageNumber="34">
<th id="7B1F0B4FF771000EFF52DC7FF8349B6B" box="[163,329,1605,1630]" gridcol="0" gridrow="4" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF52DC7FF8349B68" bold="true" box="[163,329,1605,1629]" pageId="33" pageNumber="34">Postocular spot</emphasis>
</th>
<td id="7B1F0B4FF771000EFE15DC7FFA789B6B" box="[484,773,1605,1630]" gridcol="1" gridrow="4" pageId="33" pageNumber="34">red (Fig. 87)</td>
<td id="7B1F0B4FF771000EFCD4DC7FFD369B6B" box="[805,1099,1605,1630]" gridcol="2" gridrow="4" pageId="33" pageNumber="34">indistinct red, may be hardly</td>
<td id="7B1F0B4FF771000EFB97DC7FFCF09B6B" box="[1126,1421,1605,1630]" gridcol="3" gridrow="4" pageId="33" pageNumber="34">red (Fig. 40)</td>
</tr>
<tr id="38CE6233F771000EFF52DC53FCF09BB4" box="[163,1421,1641,1665]" gridrow="5" pageId="33" pageNumber="34" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="7B1F0B4FF771000EFCD4DC53FD369BB4" box="[805,1099,1641,1665]" gridcol="2" gridrow="5" pageId="33" pageNumber="34">visible (Fig. 89a)</td>
</tr>
<tr id="38CE6233F771000EFF52DCABFCF09B9C" box="[163,1421,1681,1705]" gridrow="6" pageId="33" pageNumber="34">
<th id="7B1F0B4FF771000EFF52DCABF8349B9C" box="[163,329,1681,1705]" gridcol="0" gridrow="6" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF52DCABF8589B9C" bold="true" box="[163,293,1681,1705]" pageId="33" pageNumber="34">Pterothorax</emphasis>
</th>
<td id="7B1F0B4FF771000EFE15DCABFA789B9C" box="[484,773,1681,1705]" gridcol="1" gridrow="6" pageId="33" pageNumber="34">red, not obscurely blackish</td>
<td id="7B1F0B4FF771000EFCD4DCABFD369B9C" box="[805,1099,1681,1705]" gridcol="2" gridrow="6" pageId="33" pageNumber="34">red, but dorsum obscurely</td>
<td id="7B1F0B4FF771000EFB97DCABFCF09B9C" box="[1126,1421,1681,1705]" gridcol="3" gridrow="6" pageId="33" pageNumber="34">red, not obscurely blackish</td>
</tr>
<tr id="38CE6233F771000EFF52DC8FFCF09BFB" box="[163,1421,1717,1742]" gridrow="7" pageId="33" pageNumber="34" rowspan-0="1">
<td id="7B1F0B4FF771000EFE15DC8FFA789BFB" box="[484,773,1717,1742]" gridcol="1" gridrow="7" pageId="33" pageNumber="34">(Fig. 104)</td>
<td id="7B1F0B4FF771000EFCD4DC8FFD369BFB" box="[805,1099,1717,1742]" gridcol="2" gridrow="7" pageId="33" pageNumber="34">blackish to different extent</td>
<td id="7B1F0B4FF771000EFB97DC8FFCF09BFB" box="[1126,1421,1717,1742]" gridcol="3" gridrow="7" pageId="33" pageNumber="34">(Fig. 107)</td>
</tr>
<tr id="38CE6233F771000EFF52DCE3FCF09BC4" box="[163,1421,1753,1777]" gridrow="8" pageId="33" pageNumber="34" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="7B1F0B4FF771000EFCD4DCE3FD369BC4" box="[805,1099,1753,1777]" gridcol="2" gridrow="8" pageId="33" pageNumber="34">(Fig. 106)</td>
</tr>
<tr id="38CE6233F771000EFF52DD3BFCF09A2F" box="[163,1421,1793,1818]" gridrow="9" pageId="33" pageNumber="34">
<th id="7B1F0B4FF771000EFF52DD3BF8349A2F" box="[163,329,1793,1818]" gridcol="0" gridrow="9" pageId="33" pageNumber="34">
<emphasis id="B48ABC63F771FFD0FF52DD3BF9929A2C" bold="true" box="[163,239,1793,1817]" pageId="33" pageNumber="34">Cercus</emphasis>
</th>
<td id="7B1F0B4FF771000EFE15DD3BFA789A2F" box="[484,773,1793,1818]" gridcol="1" gridrow="9" pageId="33" pageNumber="34">inferior lobe with a small</td>
<td id="7B1F0B4FF771000EFCD4DD3BFD369A2F" box="[805,1099,1793,1818]" gridcol="2" gridrow="9" pageId="33" pageNumber="34">inferior lobe with a small</td>
<td id="7B1F0B4FF771000EFB97DD3BFCF09A2F" box="[1126,1421,1793,1818]" gridcol="3" gridrow="9" pageId="33" pageNumber="34">inferior lobe with a small</td>
</tr>
<tr id="38CE6233F771000EFF52DD1CFCF09A0B" box="[163,1421,1830,1854]" gridrow="10" pageId="33" pageNumber="34" rowspan-0="1">
<td id="7B1F0B4FF771000EFE15DD1CFA789A0B" box="[484,773,1830,1854]" gridcol="1" gridrow="10" pageId="33" pageNumber="34">tooth-like interior projection</td>
<td id="7B1F0B4FF771000EFCD4DD1CFD369A0B" box="[805,1099,1830,1854]" gridcol="2" gridrow="10" pageId="33" pageNumber="34">sharp edged bulge-like</td>
<td id="7B1F0B4FF771000EFB97DD1CFCF09A0B" box="[1126,1421,1830,1854]" gridcol="3" gridrow="10" pageId="33" pageNumber="34">sharp edged bulge-like</td>
</tr>
<tr id="38CE6233F771000EFF52DD70FCF09A57" box="[163,1421,1866,1890]" gridrow="11" pageId="33" pageNumber="34" rowspan-0="1">
<td id="7B1F0B4FF771000EFE15DD70FA789A57" box="[484,773,1866,1890]" gridcol="1" gridrow="11" pageId="33" pageNumber="34">before half-length, best</td>
<td id="7B1F0B4FF771000EFCD4DD70FD369A57" box="[805,1099,1866,1890]" gridcol="2" gridrow="11" pageId="33" pageNumber="34">interior projection before</td>
<td id="7B1F0B4FF771000EFB97DD70FCF09A57" box="[1126,1421,1866,1890]" gridcol="3" gridrow="11" pageId="33" pageNumber="34">interior projection before</td>
</tr>
<tr id="38CE6233F771000EFF52DD54FCF09AB3" box="[163,1421,1902,1926]" gridrow="12" pageId="33" pageNumber="34" rowspan-0="1">
<td id="7B1F0B4FF771000EFE15DD54FA789AB3" box="[484,773,1902,1926]" gridcol="1" gridrow="12" pageId="33" pageNumber="34">visible in dorsolateral view</td>
<td id="7B1F0B4FF771000EFCD4DD54FD369AB3" box="[805,1099,1902,1926]" gridcol="2" gridrow="12" pageId="33" pageNumber="34">half-length, best visible in</td>
<td id="7B1F0B4FF771000EFB97DD54FCF09AB3" box="[1126,1421,1902,1926]" gridcol="3" gridrow="12" pageId="33" pageNumber="34">half-length, best visible in</td>
</tr>
<tr id="38CE6233F771000EFF52DDA8FCF09A9F" box="[163,1421,1938,1962]" gridrow="13" pageId="33" pageNumber="34" rowspan-0="1">
<td id="7B1F0B4FF771000EFE15DDA8FA789A9F" box="[484,773,1938,1962]" gridcol="1" gridrow="13" pageId="33" pageNumber="34">(Fig. 138c)</td>
<td id="7B1F0B4FF771000EFCD4DDA8FD369A9F" box="[805,1099,1938,1962]" gridcol="2" gridrow="13" pageId="33" pageNumber="34">dorsolateral view (Fig. 140c)</td>
<td id="7B1F0B4FF771000EFB97DDA8FCF09A9F" box="[1126,1421,1938,1962]" gridcol="3" gridrow="13" pageId="33" pageNumber="34">dorsolateral view (Fig. 141c)</td>
</tr>
</table>
</paragraph>
<subSubSection id="CEE433FAF772FFD3FF36DAACFA2F9DE0" box="[199,850,150,213]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="86416071F772FFD3FF36DAACF8249D85" blockId="34.[199,850,150,213]" box="[199,345,150,176]" pageId="34" pageNumber="35">
<emphasis id="B48ABC63F772FFD3FF36DAACF8249D85" bold="true" box="[199,345,150,176]" pageId="34" pageNumber="35">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F772FFD3FF36DA81FA2F9DE0" blockId="34.[199,850,150,213]" box="[199,850,187,213]" pageId="34" pageNumber="35">
This species is apparently endemic to Nias Island (
<figureCitation id="1EC57CF4F772FFD3FD0EDA81FA389DE0" box="[767,837,187,213]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="34" pageNumber="35">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DEA93444039883933DBE1C78E8F5B739">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="DAE64FC1DA1182F7EA342A13F3D5D363">Seehausen, Malte</mods:namePart>
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<mods:affiliation id="464B444B04B4F00A1A527563FB45B512">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
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<mods:namePart id="25221E2C16D2872B34FFC4BB2DB8EAB6">Bedjanič, Matjaž</mods:namePart>
<mods:nameIdentifier id="4145A3DCB683CD93F4DF30F0B7C8C6C2" type="ORCID">0009-0000-6718-1078</mods:nameIdentifier>
<mods:affiliation id="76AE71F40D5CCD2209AAADEF8115FC73">Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 &amp; National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author &amp; Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078</mods:affiliation>
<mods:nameIdentifier id="AD81122FD0A503EC4D737E9C90C84B12" type="email">mseehausen@zoolmuseum.uni-kiel.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1E28E3E4BE6829C4168AE1712A044019">text</mods:typeOfResource>
<mods:relatedItem id="FDFCE95CDD8CFE42DB0ECC7EDA77BC26" type="host">
<mods:titleInfo id="D46281E69A009833F8D77A7A2E257541">
<mods:title id="3FD434CCAB7A5C5B359ECD102469C983">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B56D53874A7ECDB542D20F082BA0C573">
<mods:date id="B0F5E3DC34ADBDB1EF85AEE9FA0699A9">2025</mods:date>
<mods:detail id="DE8EF4B6668EFF11BA4A213B13B0DB8B" type="pubDate">
<mods:number id="026A3AF03E53A26C2413378C2EA6D534">2025-02-12</mods:number>
</mods:detail>
<mods:detail id="C6379563F04DEF6F7C4CDFD4EF90D8DB" type="volume">
<mods:number id="1F2DE1E1D30FC266DDBE6DDFAA800445">5587</mods:number>
</mods:detail>
<mods:detail id="B2E75F000E73B1B3BB1546DAECB5211B" type="issue">
<mods:number id="7C67B0D0A60A6326A8FD053895745730">1</mods:number>
</mods:detail>
<mods:extent id="F60780EAAC96C5690B2CF1632D573611" unit="page">
<mods:start id="38209C65ABA775F2E8E5FA20FC9D3368">1</mods:start>
<mods:end id="CFB464B3EF7F782D74B42D7D1A5E1C08">66</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="8B67626674BE86D2C27711AC4FC31DBA">
<mods:url id="2083CB54AA95B494BDF6233F338F0009">https://doi.org/10.11646/zootaxa.5587.1.1</mods:url>
</mods:location>
<mods:classification id="54EFD7BA1971C5A3B39C2C82539BDAC1">journal article</mods:classification>
<mods:identifier id="4E37D439435CA7A6A41420019B1D18CD" type="DOI">10.11646/zootaxa.5587.1.1</mods:identifier>
<mods:identifier id="C11FF8D1C093B22D2ADB9E7A65D7DEA5" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="00642F40B002E5CAF70256AFD984FB86" type="ZooBank">B6F6D9C8-4423-4DC6-BC25-940725A83DB4</mods:identifier>
</mods:mods>
<treatment id="0E57D167F757FFFBFF66D9DCF8C998D4" LSID="urn:lsid:plazi:treatment:0E57D167F757FFFBFF66D9DCF8C998D4" httpUri="http://treatment.plazi.org/id/0E57D167F757FFFBFF66D9DCF8C998D4" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<subSubSection id="CEE433FAF757FFF6FF66D9DCFDB899B4" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="86416071F757FFF6FF66D9DCFB6D9934" blockId="7.[151,528,998,1025]" box="[151,528,998,1025]" pageId="7" pageNumber="8">
<heading id="DD09D71DF757FFF6FF66D9DCFB6D9934" bold="true" box="[151,528,998,1025]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="4">
<emphasis id="B48ABC63F757FFF6FF66D9DCFB6D9934" bold="true" box="[151,528,998,1025]" pageId="7" pageNumber="8">
<taxonomicName id="41FE1BF2F757FFF6FF66D9DCF8D19934" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[151,428,998,1025]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acutidens" status="sp. nov.">
<emphasis id="B48ABC63F757FFF6FF66D9DCF8D19934" bold="true" box="[151,428,998,1025]" italics="true" pageId="7" pageNumber="8">Pseudagrion acutidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFB90118F757FFF6FE42D9DDFB6D9934" box="[435,528,999,1025]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="86416071F757FFF6FF66DE11FDB899B4" blockId="7.[151,1437,1067,1153]" pageId="7" pageNumber="8">
Figures: 5 (habitus
<typeStatus id="5945DED3F757FFF6FEAADE11F8C99976" box="[347,436,1067,1091]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FE4BDE16F8B29971" box="[442,463,1068,1092]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 68 (S910, caudal appendages
<typeStatus id="5945DED3F757FFF6FCDBDE11FAFE9976" box="[810,899,1067,1091]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FC79DE16FAE09971" box="[904,925,1068,1092]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 9 (head
<typeStatus id="5945DED3F757FFF6FBF3DE11FD269976" box="[1026,1115,1067,1091]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FB90DE16FD0B9971" box="[1121,1142,1068,1092]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 10 (pronotum
<typeStatus id="5945DED3F757FFF6FAEADE11FC099976" box="[1307,1396,1067,1091]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FA8BDE16FCF29971" box="[1402,1423,1068,1092]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 11 (habitus
<typeStatus id="5945DED3F757FFF6FECDDE71F8EE9956" box="[316,403,1099,1123]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FE68DE70F8D69957" box="[409,427,1098,1122]" count="1" pageId="7" pageNumber="8" type="female"></specimenCount>
); 12 (head
<typeStatus id="5945DED3F757FFF6FDEFDE71FB089956" box="[542,629,1099,1123]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FD8ADE70FBF09957" box="[635,653,1098,1122]" count="1" pageId="7" pageNumber="8" type="female"></specimenCount>
); 13 (pronotum
<typeStatus id="5945DED3F757FFF6FCC2DE71FAF79956" box="[819,906,1099,1123]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FC7EDE70FADC9957" box="[911,929,1098,1122]" count="1" pageId="7" pageNumber="8" type="female"></specimenCount>
); 14 (S910, ovipositor
<typeStatus id="5945DED3F757FFF6FB69DE71FD929956" box="[1176,1263,1099,1123]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
<specimenCount id="90F8ABF8F757FFF6FB04DE70FC7A9957" box="[1269,1287,1098,1122]" count="1" pageId="7" pageNumber="8" type="female"></specimenCount>
); 79 (head
<specimenCount id="90F8ABF8F757FFF6FA8BDE70FCF29957" box="[1402,1423,1098,1122]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 96 (pterothorax
<specimenCount id="90F8ABF8F757FFF6FE9BDE53F80299B4" box="[362,383,1129,1153]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 113a, b (abdomen
<specimenCount id="90F8ABF8F757FFF6FDBEDE53FB1999B4" box="[591,612,1129,1153]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 130ac (caudal appendages
<specimenCount id="90F8ABF8F757FFF6FC64DE53FAD799B4" box="[917,938,1129,1153]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
); 147151 (field photos
<specimenCount id="90F8ABF8F757FFF6FB52DE53FDC599B4" box="[1187,1208,1129,1153]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF757FFF6FF66DE95FDD79894" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="86416071F757FFF6FF66DE95FD9F99D8" blockId="7.[151,1437,1199,2017]" pageId="7" pageNumber="8">
<materialsCitation id="36966A2CF757FFF6FF66DE95FD9F99D8" collectingDate="1933-05" collectorName="van Diejen &amp; Deposited &amp; Naturalis Biodiversity Center Leiden" country="Indonesia" latitude="1.724" location="Tobelo" longLatPrecision="78" longitude="127.997" municipality="Halmahera Island" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Maluku Utara" typeStatus="holotype">
<emphasis id="B48ABC63F757FFF6FF66DE95F87099FC" bold="true" box="[151,269,1199,1225]" pageId="7" pageNumber="8">
<typeStatus id="5945DED3F757FFF6FF66DE95F87599FC" box="[151,264,1199,1225]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="90F8ABF8F757FFF6FEE2DE95F85799FC" box="[275,298,1199,1225]" count="1" pageId="7" pageNumber="8" type="male"></specimenCount>
, acq.
<date id="F24046B1F757FFF6FE95DE95F8CC99FC" box="[356,433,1199,1225]" pageId="7" pageNumber="8" value="1933-05">
<collectingDate id="E204BF59F757FFF6FE95DE95F8CC99FC" box="[356,433,1199,1225]" pageId="7" pageNumber="8" value="1933-05">v.1933</collectingDate>
</date>
,
<location id="832136AAF757FFF6FE4BDE95FB7799FC" LSID="urn:lsid:plazi:treatment:0E57D167F757FFFBFF66D9DCF8C998D4:832136AAF757FFF6FE4BDE95FB7799FC" box="[442,522,1199,1225]" country="Indonesia" latitude="1.724" longLatPrecision="78" longitude="127.997" municipality="Halmahera Island" name="Tobelo" pageId="7" pageNumber="8" stateProvince="Maluku Utara">Tobelo</location>
,
<collectingMunicipality id="6625FA0BF757FFF6FDE4DE95FBA199FC" box="[533,732,1199,1225]" pageId="7" pageNumber="8">Halmahera Island</collectingMunicipality>
,
<collectingRegion id="443AAE93F757FFF6FD17DE95FAF599FC" box="[742,904,1199,1225]" country="Indonesia" name="Maluku Utara" pageId="7" pageNumber="8">North Maluku</collectingRegion>
,
<collectingCountry id="FEE920E1F757FFF6FC63DE95FD7F99FC" box="[914,1026,1199,1225]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
, approximately
<geoCoordinate id="E3CA06B6F757FFF6FB42DE95FC6999FC" box="[1203,1300,1199,1225]" degrees="1.724" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="55" value="1.724">1.724°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F757FFF6FAEFDE95FCE599FC" box="[1310,1432,1199,1225]" degrees="127.997" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="55" value="127.997">127.997°E</geoCoordinate>
,
<collectorName id="2B0B05A7F757FFF6FF66DEE9F86F99D8" box="[151,274,1235,1261]" pageId="7" pageNumber="8">van Diejen</collectorName>
leg., no. 1519102.
<collectorName id="2B0B05A7F757FFF6FE18DEE9FB2699D8" box="[489,603,1235,1261]" pageId="7" pageNumber="8">Deposited</collectorName>
at
<collectorName id="2B0B05A7F757FFF6FD8CDEE9FD6299D8" box="[637,1055,1235,1261]" pageId="7" pageNumber="8">Naturalis Biodiversity Center Leiden</collectorName>
, the Netherlands.
</materialsCitation>
</paragraph>
<paragraph id="86416071F757FFF6FF36DECDFDD79894" blockId="7.[151,1437,1199,2017]" pageId="7" pageNumber="8">
<emphasis id="B48ABC63F757FFF6FF36DECDF8429824" bold="true" box="[199,319,1271,1297]" pageId="7" pageNumber="8">
<typeStatus id="5945DED3F757FFF6FF36DECDF8429824" box="[199,319,1271,1297]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="90F8ABF8F757FFF6FEA9DECDF8EE9824" box="[344,403,1271,1297]" count="4" pageId="7" pageNumber="8" type="male">4♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F757FFF6FE59DECDF89F9824" box="[424,482,1271,1297]" count="2" pageId="7" pageNumber="8" type="female">2♀♀</specimenCount>
):
<materialsCitation id="36966A2CF757FFF6FDF3DECDFD399803" collectingDate="1933-05" collectionCode="RMNH" collectorName="van Diejen" country="Indonesia" latitude="1.724" location="Tobelo" longLatPrecision="78" longitude="127.997" municipality="Halmahera Island" pageId="7" pageNumber="8" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Maluku Utara" typeStatus="paratype">
<specimenCount id="90F8ABF8F757FFF6FDF3DECDFB409824" box="[514,573,1271,1297]" count="2" pageId="7" pageNumber="8" type="male">2♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F757FFF6FDA0DECDFB099824" box="[593,628,1271,1297]" count="1" pageId="7" pageNumber="8" type="female">1♀</specimenCount>
, acq.
<date id="F24046B1F757FFF6FD4ADECDFA759824" box="[699,776,1271,1297]" pageId="7" pageNumber="8" value="1933-05">
<collectingDate id="E204BF59F757FFF6FD4ADECDFA759824" box="[699,776,1271,1297]" pageId="7" pageNumber="8" value="1933-05">v.1933</collectingDate>
</date>
,
<location id="832136AAF757FFF6FCEDDECDFA119824" LSID="urn:lsid:plazi:treatment:0E57D167F757FFFBFF66D9DCF8C998D4:832136AAF757FFF6FCEDDECDFA119824" box="[796,876,1271,1297]" country="Indonesia" latitude="1.724" longLatPrecision="78" longitude="127.997" municipality="Halmahera Island" name="Tobelo" pageId="7" pageNumber="8" stateProvince="Maluku Utara">Tobelo</location>
,
<collectingMunicipality id="6625FA0BF757FFF6FC70DECDFD2E9824" box="[897,1107,1271,1297]" pageId="7" pageNumber="8">Halmahera Island</collectingMunicipality>
,
<collectingRegion id="443AAE93F757FFF6FB99DECDFC699824" box="[1128,1300,1271,1297]" country="Indonesia" name="Maluku Utara" pageId="7" pageNumber="8">North Maluku</collectingRegion>
,
<collectingCountry id="FEE920E1F757FFF6FAD9DECDFCE59824" box="[1320,1432,1271,1297]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
, approximately
<geoCoordinate id="E3CA06B6F757FFF6FECFDF21F8DD9800" box="[318,416,1307,1333]" degrees="1.724" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="55" value="1.724">1.724°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F757FFF6FE58DF21FB5E9800" box="[425,547,1307,1333]" degrees="127.997" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="55" value="127.997">127.997°E</geoCoordinate>
,
<collectorName id="2B0B05A7F757FFF6FDDCDF21FBDB9800" box="[557,678,1307,1333]" pageId="7" pageNumber="8">van Diejen</collectorName>
leg.,
<collectionCode id="E0EFF8B4F757FFF6FD10DF21FA459800" box="[737,824,1307,1333]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="8" type="Museum">RMNH</collectionCode>
, no. 1519101, 1519102
</materialsCitation>
;—
<materialsCitation id="36966A2CF757FFF6FB99DF21FB27984B" collectingDate="1985-07-08" collectionCode="RMNH" collectorName="F. G. Rozendaal" country="Indonesia" latitude="0.741" location="Wayaua" longLatPrecision="78" longitude="127.641" municipality="Bacan Island" pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Maluku Utara" typeStatus="paratype">
<specimenCount id="90F8ABF8F757FFF6FB99DF21FDF89803" box="[1128,1157,1307,1334]" count="1" pageId="7" pageNumber="8" type="male">1♂</specimenCount>
,
<specimenCount id="90F8ABF8F757FFF6FB7FDF21FDCC9800" box="[1166,1201,1307,1333]" count="1" pageId="7" pageNumber="8" type="female">1♀</specimenCount>
,
<date id="F24046B1F757FFF6FB4FDF21FC4F9803" box="[1214,1330,1307,1334]" pageId="7" pageNumber="8" value="1985-07-08">
<collectingDate id="E204BF59F757FFF6FB4FDF21FC4F9803" box="[1214,1330,1307,1334]" pageId="7" pageNumber="8" value="1985-07-08">8.vii.1985</collectingDate>
</date>
,
<location id="832136AAF757FFF6FACDDF21FCE59800" LSID="urn:lsid:plazi:treatment:0E57D167F757FFFBFF66D9DCF8C998D4:832136AAF757FFF6FACDDF21FCE59800" box="[1340,1432,1307,1333]" country="Indonesia" latitude="0.741" longLatPrecision="78" longitude="127.641" municipality="Bacan Island" name="Wayaua" pageId="7" pageNumber="8" stateProvince="Maluku Utara">Wayaua</location>
, rivulet, edge of logged forest,
<collectingMunicipality id="6625FA0BF757FFF6FE02DF05FBF5986C" box="[499,648,1343,1369]" pageId="7" pageNumber="8">Bacan Island</collectingMunicipality>
,
<collectingRegion id="443AAE93F757FFF6FD67DF05FA47986C" box="[662,826,1343,1369]" country="Indonesia" name="Maluku Utara" pageId="7" pageNumber="8">North Maluku</collectingRegion>
,
<collectingCountry id="FEE920E1F757FFF6FCB6DF05FACA986C" box="[839,951,1343,1369]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
, approximately -
<geoCoordinate id="E3CA06B6F757FFF6FB87DF05FDA5986C" box="[1142,1240,1343,1369]" degrees="0.741" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="55" value="0.741">0.741°N</geoCoordinate>
,
<geoCoordinate id="E3CA06B6F757FFF6FB14DF05FC22986C" box="[1253,1375,1343,1369]" degrees="127.641" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="55" value="127.641">127.641°E</geoCoordinate>
,
<collectorName id="2B0B05A7F757FFF6FA9CDF05F8709848" pageId="7" pageNumber="8">F.G. Rozendaal</collectorName>
leg.,
<collectionCode id="E0EFF8B4F757FFF6FEA9DF59F8D29848" box="[344,431,1379,1405]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="8" type="Museum">RMNH</collectionCode>
, no. 1519079
</materialsCitation>
;—
<materialsCitation id="36966A2CF757FFF6FD8EDF59FDDB9894" collectingDate="1997-08-01" collectionCode="MB" collectorName="R. Verovnik" country="Indonesia" elevation="440" location="Chucumara" municipality="Morotai Island" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Maluku Utara" typeStatus="paratype">
<specimenCount id="90F8ABF8F757FFF6FD8EDF59FBE1984B" box="[639,668,1379,1406]" count="1" pageId="7" pageNumber="8" type="male">1♂</specimenCount>
,
<date id="F24046B1F757FFF6FD5CDF59FA549848" box="[685,809,1379,1405]" pageId="7" pageNumber="8" value="1997-08-01">
<collectingDate id="E204BF59F757FFF6FD5CDF59FA549848" box="[685,809,1379,1405]" pageId="7" pageNumber="8" value="1997-08-01">1.viii.1997</collectingDate>
</date>
, primary rainforest
<quantity id="4106CD94F757FFF6FBE8DF59FD259848" box="[1049,1112,1379,1405]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="km" value="9.0">9 km</quantity>
of logging company camp
<location id="832136AAF757FFF6FF66DFBDF8669894" LSID="urn:lsid:plazi:treatment:0E57D167F757FFFBFF66D9DCF8C998D4:832136AAF757FFF6FF66DFBDF8669894" box="[151,283,1415,1441]" country="Indonesia" municipality="Morotai Island" name="Chucumara" pageId="7" pageNumber="8" stateProvince="Maluku Utara">Chucumara</location>
,
<collectingMunicipality id="6625FA0BF757FFF6FED7DFBDF8B39894" box="[294,462,1415,1441]" pageId="7" pageNumber="8">Morotai Island</collectingMunicipality>
,
<collectingRegion id="443AAE93F757FFF6FE2BDFBDFB019894" box="[474,636,1415,1441]" country="Indonesia" name="Maluku Utara" pageId="7" pageNumber="8">North Maluku</collectingRegion>
,
<collectingCountry id="FEE920E1F757FFF6FD76DFBDFB8A9894" box="[647,759,1415,1441]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
, alt.
<quantity id="4106CD94F757FFF6FCDDDFBDFAD09894" box="[812,941,1415,1441]" metricMagnitude="2" metricUnit="m" metricValue="4.4" metricValueMax="4.8" metricValueMin="4.0" pageId="7" pageNumber="8" unit="m" value="440.0" valueMax="480.0" valueMin="400.0">
<elevation id="0DD38742F757FFF6FCDDDFBDFAD09894" box="[812,941,1415,1441]" metricMagnitude="2" metricUnit="m" metricValue="4.4" metricValueMax="4.8" metricValueMin="4.0" pageId="7" pageNumber="8" unit="m" value="440.0" valueMax="480.0" valueMin="400.0">400480 m</elevation>
</quantity>
,
<collectorName id="2B0B05A7F757FFF6FC48DFBDFD3D9894" box="[953,1088,1415,1441]" pageId="7" pageNumber="8">R. Verovnik</collectorName>
leg,
<collectionCode id="E0EFF8B4F757FFF6FB86DFBDFDDB9894" box="[1143,1190,1415,1441]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="8">MB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF757FFF6FF36DF91FBC698DC" pageId="7" pageNumber="8" type="etymology">
<paragraph id="86416071F757FFF6FF36DF91FBC698DC" blockId="7.[151,1437,1199,2017]" pageId="7" pageNumber="8">
<emphasis id="B48ABC63F757FFF6FF36DF91F82F98F0" bold="true" box="[199,338,1451,1477]" pageId="7" pageNumber="8">Etymology:</emphasis>
The name refers to the inferior lobe of the male cercus, which has a large tooth-like medially pointed projection, clearly visible in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF757FFF6FF36DFC9FB149B60" pageId="7" pageNumber="8" type="discussion">
<paragraph id="86416071F757FFF6FF36DFC9FB149B60" blockId="7.[151,1437,1199,2017]" pageId="7" pageNumber="8">
<emphasis id="B48ABC63F757FFF6FF36DFC9F8499B38" bold="true" box="[199,308,1523,1549]" pageId="7" pageNumber="8">Remarks</emphasis>
: Further specimens from the islands of Halmahera, Bacan, Buru, and Obi were discovered among unidentified material at the RMNH by V.J. Kalkman just prior to submission of the manuscript. For this reason we have not included them in the
<typeStatus id="5945DED3F757FFF6FE1BDC01FB679B60" box="[490,538,1595,1621]" pageId="7" pageNumber="8">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF757FFF8FF36DC65FAC99A8C" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph id="86416071F757FFF6FF36DC65FB629B4C" blockId="7.[151,1437,1199,2017]" box="[199,543,1631,1657]" pageId="7" pageNumber="8">
<emphasis id="B48ABC63F757FFF6FF36DC65FB629B4C" bold="true" box="[199,543,1631,1657]" pageId="7" pageNumber="8">
Description of
<typeStatus id="5945DED3F757FFF6FE86DC65F8A39B4C" box="[375,478,1631,1657]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="86416071F757FFF6FF36DCB9FDCB9A64" blockId="7.[151,1437,1199,2017]" pageId="7" pageNumber="8">
<emphasis id="B48ABC63F757FFF6FF36DCB9F8799BA8" box="[199,260,1667,1693]" italics="true" pageId="7" pageNumber="8">Head</emphasis>
(
<figureCitation id="1EC57CF4F757FFF6FEE5DCB9F8249BA8" box="[276,345,1667,1693]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="7" pageNumber="8">Fig. 9</figureCitation>
, also compare
<figureCitation id="1EC57CF4F757FFF6FDF3DCB9FB2B9BA8" box="[514,598,1667,1693]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="7" pageNumber="8">Fig. 79</figureCitation>
): Labium, mandibles, genae, labrum, anteclypeus and postclypeus yellow; antefrons and postfrons yellow, with posterior margin of postfrons washed out reddish, sharply separated from black vertex around level of anterior margin of lateral ocelli; a yellow-orange centre marking at vertex expands towards occipital bar; median ocellus with black spot posteriorly; occipital bar reduced to a small orange-reddish stripe in centre; triangular postocular spots orange-reddish, bordered by black; rear of head black, but pale along the eye margins; base and scape of antennae yellow, pedicel ochre-brownish, rest of antennae lacking.
</paragraph>
<paragraph id="86416071F757FFF9FF36DD61FAD49CBC" blockId="7.[151,1437,1199,2017]" lastBlockId="8.[151,1437,151,753]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B48ABC63F757FFF6FF36DD61F8659A40" box="[199,280,1883,1909]" italics="true" pageId="7" pageNumber="8">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F757FFF6FED7DD61F8EA9A40" box="[294,407,1883,1910]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="7" pageNumber="8">Figs 5, 10</figureCitation>
, also compare
<figureCitation id="1EC57CF4F757FFF6FDCDDD61FBF39A40" box="[572,654,1883,1909]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="7" pageNumber="8">Fig. 96</figureCitation>
): Anterior lobe of pronotum yellowish, with anterior margin black; the yellow colour projecting in centre as semi-circle towards middle lobe; middle pronotal lobe otherwise black, with a centrally divided yellow-orange patch in the middle, and on each side a large yellow-orange patch dorsolaterally; propleuron black above, yellow-orange below and to rear; posterior lobe yellow-orange; apical edge of posterior lobe convex. Pterothorax orange-reddish; middorsal carina black; moderately broad black humeral stripe, slightly widened at posterior third before peaked towards posterior margin of pterothorax; interpleural suture black with a black spot anteriorly; posterior half of metapleural suture black; mesinfraepisternum with black diagonal marking, continuing the black humeral stripe; venter pale, covered with whitish pruinosity. Coxae and legs pale orange-ochre; outer side of femur with a washed-out blackish stripe, covering almost complete femur in foreleg, almost complete posterior half of femur in midleg, and posterior quarter of femur in hind leg; tibiae blackish along the black spines; tarsi and tarsal claws orange-ochre, washed-out blackish along waists.
</paragraph>
<paragraph id="86416071F758FFF9FF36DBA9FAA19CE4" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="9">
<emphasis id="B48ABC63F758FFF9FF36DBA9F8769C99" box="[199,267,403,428]" italics="true" pageId="8" pageNumber="9">Wings</emphasis>
(
<figureCitation id="1EC57CF4F758FFF9FEEFDBA9F8189C98" box="[286,357,402,429]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="8" pageNumber="9">Fig. 5</figureCitation>
): Membrane hyaline, slightly tinged yellowish-brown; veins brown to black; 16.016.5 px in forewing; 14.515.0 px in hindwing; Pt reddish-brown, overlying one cell.
</paragraph>
<paragraph id="86416071F758FFF9FF36DBE1F8169FC4" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="9">
<emphasis id="B48ABC63F758FFF9FF36DBE1F84C9CC0" box="[199,305,475,501]" italics="true" pageId="8" pageNumber="9">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F758FFF9FEB0DBE1F8DB9CC0" box="[321,422,474,501]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="8" pageNumber="9">Figs 57</figureCitation>
, also compare
<figureCitation id="1EC57CF4F758FFF9FDA1DBE1FBA39CC0" box="[592,734,475,501]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="8" pageNumber="9">Figs 113a, b</figureCitation>
): S1 orange-reddish, dorsally with a black quadrangular patch covering anterior two-thirds of segment, posterior margin black; S2 orange-reddish, dorsally with acute mushroom-like black marking at posterior third, and triangular black patch at anterior margin; both black markings almost connected by thin black middorsal stripe; S38 dorsally black, expanded to lateral half of segment; pale yellowish basal annulus; laterally and ventrally pale yellowish-orange; S9 with anterior half laterally and dorsally orange-reddish; posterior half of S9 black, with its anterior margin somewhat serrated with incomplete middorsal stripe; S10 orange-reddish, dorsally with broad black “x”-like marking; black colour of anterior and posterior margin expanded laterally.
</paragraph>
<caption id="D28130F9F758FFF9FF66DD6EFAF29AA4" pageId="8" pageNumber="9" startId="8.[151,264,1876,1900]" targetBox="[153,1433,811,1847]" targetPageId="8" targetType="figure">
<paragraph id="86416071F758FFF9FF66DD6EFAF29AA4" blockId="8.[151,1436,1876,1937]" pageId="8" pageNumber="9">
<emphasis id="B48ABC63F758FFF9FF66DD6EF83C9A58" bold="true" box="[151,321,1876,1901]" pageId="8" pageNumber="9">FIGURES 510</emphasis>
.
<taxonomicName id="41FE1BF2F758FFF9FEBFDD6FFB459A58" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[334,568,1877,1901]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="acutidens">
<emphasis id="B48ABC63F758FFF9FEBFDD6FFB459A58" box="[334,568,1877,1901]" italics="true" pageId="8" pageNumber="9">Pseudagrion acutidens</emphasis>
</taxonomicName>
, holotype male: (
<emphasis id="B48ABC63F758FFF9FD1DDD6EFA7C9A59" bold="true" box="[748,769,1876,1900]" pageId="8" pageNumber="9">5)</emphasis>
habitus lateral, (
<emphasis id="B48ABC63F758FFF9FC5CDD6FFAC79A58" bold="true" box="[941,954,1877,1901]" pageId="8" pageNumber="9">6</emphasis>
) appendages lateral, (
<emphasis id="B48ABC63F758FFF9FB6ADD6FFDD59A58" bold="true" box="[1179,1192,1877,1901]" pageId="8" pageNumber="9">7</emphasis>
) appendages dorsal, (
<emphasis id="B48ABC63F758FFF9FA76DD6FFCE99A58" bold="true" box="[1415,1428,1877,1901]" pageId="8" pageNumber="9">8</emphasis>
) cercus dorsolateral, (
<emphasis id="B48ABC63F758FFF9FE9ADD43F8059AA4" bold="true" box="[363,376,1913,1937]" pageId="8" pageNumber="9">9</emphasis>
) head dorsal, (
<emphasis id="B48ABC63F758FFF9FDFEDD43FB559AA4" bold="true" box="[527,552,1913,1937]" pageId="8" pageNumber="9">10</emphasis>
) pronotum. Photos: M. Seehausen.
</paragraph>
</caption>
<paragraph id="86416071F759FFF8FF36DAADFD329C50" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36DAADF8D69D84" box="[199,427,151,177]" italics="true" pageId="9" pageNumber="10">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F759FFF8FE4CDAADFB5B9D84" box="[445,550,151,177]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="9" pageNumber="10">Figs 68</figureCitation>
, also compare
<figureCitation id="1EC57CF4F759FFF8FD2BDAADFA119D84" box="[730,876,151,177]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="9" pageNumber="10">Figs 130ac</figureCitation>
): Cercus laterally washed-out brownish, slightly shorter than S10; in lateral view cercus subrectangular, with inter-lobe notch centrally placed at the apex; in dorsal view apex of superior lobe of cercus rounded and curved in; inferior lobe expanded inwards, subtriangular, apex of expanded part blackish and tooth-like, curved towards body; paraproct blackish at base, apex orange-brownish; slightly longer than half of cercus; viewed laterally subtriangular and tapered, with apex rounded; viewed dorsally paraproct broadly rounded, concave bowl-like, with blackish tubercular medial edge.
</paragraph>
<paragraph id="86416071F759FFF8FF36DB55FA389C98" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">Measurements (mm): Total length (with appendages) 43.2; abdomen length (without appendages) 36.1; hindwing 23.6; forewing 25.2; Pt in forewing 0.8; cercus 0.5.</paragraph>
<paragraph id="86416071F759FFF8FF36DB8CFB6E9CE4" blockId="9.[151,1437,151,1977]" box="[199,531,438,465]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36DB8CFB6E9CE4" bold="true" box="[199,531,438,465]" pageId="9" pageNumber="10">
Variation in
<typeStatus id="5945DED3F759FFF8FEACDB8DF8B59CE4" box="[349,456,439,465]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
males
</emphasis>
</paragraph>
<paragraph id="86416071F759FFF8FF36DBE1FDE09F2C" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36DBE1F8799CC0" box="[199,260,475,501]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
: All males have a black spot at centre of posterior margin of postclypeus (as in
<figureCitation id="1EC57CF4F759FFF8FB7BDBE1FD9D9CC0" box="[1162,1248,475,501]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="9" pageNumber="10">Figs 79</figureCitation>
,
<figureCitation id="1EC57CF4F759FFF8FB1CDBE1FC679CC0" box="[1261,1306,475,501]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="9" pageNumber="10">148</figureCitation>
; not visible in
<figureCitation id="1EC57CF4F759FFF8FF45DBC5F9889F2C" box="[180,245,511,537]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="9" pageNumber="10">Fig. 9</figureCitation>
of the
<typeStatus id="5945DED3F759FFF8FEB5DBC5F8DB9F2C" box="[324,422,511,537]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
because of dirt); the occipital bar is completely black in
<specimenCount id="90F8ABF8F759FFF8FBD2DBC5FDE49F2C" box="[1059,1177,511,537]" count="2" pageId="9" pageNumber="10" type="male">two males</specimenCount>
.
</paragraph>
<paragraph id="86416071F759FFF8FF36D819F9A09F54" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36D819F8659F08" box="[199,280,547,573]" italics="true" pageId="9" pageNumber="10">Thorax</emphasis>
: The males from Morotai and Bacan have the metapleural suture almost completely covered by a black stripe.
</paragraph>
<paragraph id="86416071F759FFF8FF36D851FA129FB0" blockId="9.[151,1437,151,1977]" box="[199,879,618,645]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36D851F8689FB1" box="[199,277,619,644]" italics="true" pageId="9" pageNumber="10">Wings:</emphasis>
14.517.5 px in forewing; 13.515.0 px in hindwing.
</paragraph>
<paragraph id="86416071F759FFF8FF36D8B5FDD89E20" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36D8B5F84C9F9C" box="[199,305,655,681]" italics="true" pageId="9" pageNumber="10">Abdomen</emphasis>
:
<specimenCount id="90F8ABF8F759FFF8FECED8B5F8C49F9C" box="[319,441,655,681]" count="2" pageId="9" pageNumber="10" type="male">Two males</specimenCount>
have the triangular black patch at anterior margin of S2 reduced; the male from Morotai has a complete thin black middorsal stripe on S2 (as in
<figureCitation id="1EC57CF4F759FFF8FCFCD889FA129FF8" box="[781,879,691,717]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="9" pageNumber="10">Fig. 147</figureCitation>
); the male from Bacan has S8 dorsally black, but with two elongate triangular reddish patches at anterior third, each situated dorsolaterally, and about as long as one-third of S8;
<specimenCount id="90F8ABF8F759FFF8FEEFD8C1F8FE9E20" box="[286,387,763,789]" count="1" pageId="9" pageNumber="10" type="male">one male</specimenCount>
has S9 with a black dot each lateral of the incomplete middorsal stripe.
</paragraph>
<paragraph id="86416071F759FFF8FF36D925FD499E68" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">Measurements (mm): Total length (with appendages) 42.345.3; abdomen length (without appendages) 35.7 38.0; hindwing 23.324.2; forewing 25.025.6; Pt in forewing 0.80.9; cercus 0.5.</paragraph>
<paragraph id="86416071F759FFF8FF36D95CFB3E9EB4" blockId="9.[151,1437,151,1977]" box="[199,579,870,897]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36D95CFB3E9EB4" bold="true" box="[199,579,870,897]" pageId="9" pageNumber="10">
Description of female
<typeStatus id="5945DED3F759FFF8FE3CD95DFB3E9EB4" box="[461,579,871,897]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="86416071F759FFF8FF36D9B1FDFF9948" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36D9B1F8799E90" box="[199,260,907,933]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
(
<figureCitation id="1EC57CF4F759FFF8FEE4D9B1F8169E90" box="[277,363,907,933]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="9" pageNumber="10">Fig. 12</figureCitation>
): Labium, mandibles, genae, labrum, anteclypeus and postclypeus yellow-orange; postclypeus with at least a blackish dot at centre of posterior margin, and an elongated blackish spot at each side, but may likewise be almost completely dark; antefrons and postfrons yellow-orange, with a black elongated patch in centre; yellow-orange colour of postfrons sharply separated from black vertex around level of lateral ocelli; median ocellus with a black spot posteriorly; lateral ocelli largely surrounded by black, less distinct or lacking at their anterior margins; triangular postocular spots orange-reddish, broadly bordered by black; rear of head black, but pale along the eye margins; base and scape of antennae orange-ochre, following segments brownish.
</paragraph>
<paragraph id="86416071F759FFF8FF36DEBDFB6B9B18" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36DEBDF8659994" box="[199,280,1159,1185]" italics="true" pageId="9" pageNumber="10">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F759FFF8FED4DEBDF8DF9994" box="[293,418,1159,1185]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="9" pageNumber="10">Figs 11, 13</figureCitation>
): Pronotum with anterior lobe pale yellowish, its anterior margin black; middle pronotal lobe black with a centrally divided yellowish patch in the middle, and on either side a large yellowish patch dorsolaterally; propleuron black above, pale yellowish below and to rear; posterior lobe of pronotum yellow-brownish; apical edge of posterior lobe convex, bearing two dark horns directed towards head, approximately reaching the centre of the middle lobe. Pterothorax orange; middorsal carina black; moderately broad black humeral stripe, slightly widened in posterior third before peaking towards posterior margin of pterothorax; interpleural suture darkened with blackish spot anteriorly; metapleural suture at least with black elongated patch posteriorly, but may be almost completely blackish; mesinfraepisternum with black diagonal marking, continuing the black humeral stripe; venter of pterothorax pale yellowish, may be covered with whitish pruinosity. Coxae and legs pale yellow-ochre; outer side of femur with a washed-out blackish stripe along posterior two-thirds in foreleg, reduced to posterior half in midleg, and posterior quarter in hindleg; tibiae blackish along the black spines; tarsi and tarsal claws yellow-ochre, washed-out blackish along waists.
</paragraph>
<paragraph id="86416071F759FFF8FF36DC0DFABC9B40" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36DC0DF8769B65" box="[199,267,1591,1616]" italics="true" pageId="9" pageNumber="10">Wings</emphasis>
(
<figureCitation id="1EC57CF4F759FFF8FEEADC0DF8139B64" box="[283,366,1591,1617]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="9" pageNumber="10">Fig. 11</figureCitation>
): Membrane hyaline, may be slightly tinged yellowish-brown; veins brown to black; 17.518.0 px in forewing; 14.516.5 px in hindwings; Pt ochre, overlying one cell.
</paragraph>
<paragraph id="86416071F759FFF8FF36DC45F8649A44" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">
<emphasis id="B48ABC63F759FFF8FF36DC45F84C9BAC" box="[199,305,1663,1689]" italics="true" pageId="9" pageNumber="10">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F759FFF8FEB1DC45F8BE9BAC" box="[320,451,1663,1689]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="9" pageNumber="10">Figs 11, 14</figureCitation>
): S1 yellow-orange, dorsum black; S2 yellow-orange, with black dorsal marking which is expanded laterally at posterior third, and may also be widened at anterior margin; S38 dorsally black, expanded to lateral half of segment, pale yellowish basal annulus; laterally and ventrally pale yellowish; S910 dorsally black, expanded to lateral along posterior and anterior margin; laterally and ventrally pale yellowish. Cercus dark, as long as half of S10; paraproct dark, rounded laterally; ovipositor pale yellowish, its apex not reaching to posterior end of S10; bearing two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F759FFF8FF36DD41FAC99A8C" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="10">Measurements (mm): Total length (with appendages) 41.642.0; abdomen length (without appendages) 33.8 34.5; hindwing 25.125.3; forewing 26.626.8; Pt in forewing 0.81.0.</paragraph>
</subSubSection>
<caption id="D28130F9F75AFFFBFF66D997F8B69EDC" pageId="10" pageNumber="11" startId="10.[151,264,941,965]" targetBox="[154,1434,180,911]" targetPageId="10" targetType="figure">
<paragraph id="86416071F75AFFFBFF66D997F8B69EDC" blockId="10.[151,1436,941,1001]" pageId="10" pageNumber="11">
<emphasis id="B48ABC63F75AFFFBFF66D997F8369EF0" bold="true" box="[151,331,941,965]" pageId="10" pageNumber="11">FIGURES 1114</emphasis>
.
<taxonomicName id="41FE1BF2F75AFFFBFEA7D997FB409EF0" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[342,573,941,965]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">
<emphasis id="B48ABC63F75AFFFBFEA7D997FB409EF0" box="[342,573,941,965]" italics="true" pageId="10" pageNumber="11">Pseudagrion acutidens</emphasis>
</taxonomicName>
, female paratype: (
<emphasis id="B48ABC63F75AFFFBFD0FD997FA6B9EF0" bold="true" box="[766,790,941,965]" pageId="10" pageNumber="11">11</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F75AFFFBFC34D997FAA39EF0" bold="true" box="[965,990,941,965]" pageId="10" pageNumber="11">12</emphasis>
) head dorsal, (
<emphasis id="B48ABC63F75AFFFBFB83D997FDF69EF0" bold="true" box="[1138,1163,941,965]" pageId="10" pageNumber="11">13</emphasis>
) pronotum, (
<emphasis id="B48ABC63F75AFFFBFAFED997FC559EF0" bold="true" box="[1295,1320,941,965]" pageId="10" pageNumber="11">14</emphasis>
) ovipositor lateral. Photos: M. Seehausen.
</paragraph>
</caption>
<subSubSection id="CEE433FAF75AFFFBFF36DE01FB889840" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="86416071F75AFFFBFF36DE01F8479960" blockId="10.[151,1437,1083,1506]" box="[199,314,1083,1109]" pageId="10" pageNumber="11">
<emphasis id="B48ABC63F75AFFFBFF36DE01F8479960" bold="true" box="[199,314,1083,1109]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F75AFFFBFF36DE65FB889840" blockId="10.[151,1437,1083,1506]" pageId="10" pageNumber="11">
<taxonomicName id="41FE1BF2F75AFFFBFF36DE65F8B1994C" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[199,460,1119,1145]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">
<emphasis id="B48ABC63F75AFFFBFF36DE65F8B1994C" box="[199,460,1119,1145]" italics="true" pageId="10" pageNumber="11">Pseudagrion acutidens</emphasis>
</taxonomicName>
overlaps in range on Halmahera with
<taxonomicName id="41FE1BF2F75AFFFBFC60DE5AFD4B994C" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[913,1078,1119,1145]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F75AFFFBFC60DE5AFD4B994C" box="[913,1078,1119,1145]" italics="true" pageId="10" pageNumber="11">P. halmaherae</emphasis>
</taxonomicName>
described below, and on Buru with
<taxonomicName id="41FE1BF2F75AFFFBFF22DEBEF83799A8" authorityName="Selys" authorityYear="1876" box="[211,330,1156,1181]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="crocops">
<emphasis id="B48ABC63F75AFFFBFF22DEBEF83799A8" box="[211,330,1156,1181]" italics="true" pageId="10" pageNumber="11">P. crocops</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F75AFFFBFE77DEBEF89599A8" authorityName="Selys" authorityYear="1876" box="[390,488,1156,1181]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F75AFFFBFE77DEBEF89599A8" box="[390,488,1156,1181]" italics="true" pageId="10" pageNumber="11">P. ustum</emphasis>
</taxonomicName>
. However, males of
<taxonomicName id="41FE1BF2F75AFFFBFD2CDEBEFA1A99A8" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[733,871,1155,1181]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">
<emphasis id="B48ABC63F75AFFFBFD2CDEBEFA1A99A8" box="[733,871,1155,1181]" italics="true" pageId="10" pageNumber="11">P. acutidens</emphasis>
</taxonomicName>
can easily be separated from all congeners west of Lydekkers Line by the large medially projecting tooth on the cercus (
<figureCitation id="1EC57CF4F75AFFFBFC1BDE9DFD2E99F4" box="[1002,1107,1191,1217]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="10" pageNumber="11">Figs 78</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFB93DE9DFDC499F7" box="[1122,1209,1191,1218]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="10" pageNumber="11">130b, c</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFB38DE9DFD8A99F4" box="[1225,1271,1191,1217]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="10" pageNumber="11">149</figureCitation>
).
<taxonomicName id="41FE1BF2F75AFFFBFAFCDE9DF9A799D0" authorityName="Selys" authorityYear="1876" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F75AFFFBFAFCDE9DF9A799D0" italics="true" pageId="10" pageNumber="11">Pseudagrion ustum</emphasis>
</taxonomicName>
has a tooth of similar size on the cercus, but in that species the tooth is projecting upward (
<figureCitation id="1EC57CF4F75AFFFBFAF1DEF1FCED99D0" box="[1280,1424,1227,1253]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="10" pageNumber="11">Figs 146ac</figureCitation>
). Furthermore
<taxonomicName id="41FE1BF2F75AFFFBFEDBDECAF8B1983C" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[298,460,1263,1289]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F75AFFFBFEDBDECAF8B1983C" box="[298,460,1263,1289]" italics="true" pageId="10" pageNumber="11">P. halmaherae</emphasis>
</taxonomicName>
has a broad humeral stripe in both sexes (
<figureCitation id="1EC57CF4F75AFFFBFC53DED5FA85983C" box="[930,1016,1263,1289]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="10" pageNumber="11">Figs 33</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFBF5DED5FD5F983C" box="[1028,1058,1263,1289]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="10" pageNumber="11">39</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFBDFDED5FD26983C" box="[1070,1115,1263,1289]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="10" pageNumber="11">103</figureCitation>
), whereas the humeral stripe is narrower in
<taxonomicName id="41FE1BF2F75AFFFBFEC8DF2EF8C29818" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[313,447,1299,1325]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">
<emphasis id="B48ABC63F75AFFFBFEC8DF2EF8C29818" box="[313,447,1299,1325]" italics="true" pageId="10" pageNumber="11">P. acutidens</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F75AFFFBFE3FDF29FB6A981B" box="[462,535,1299,1326]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="10" pageNumber="11">Figs 5</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFDD3DF29FB429818" box="[546,575,1299,1325]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="10" pageNumber="11">11</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFDBADF29FB149818" box="[587,617,1299,1325]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="10" pageNumber="11">96</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFD84DF29FA36981B" box="[629,843,1299,1326]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="10" pageNumber="11">147148, 150151</figureCitation>
). Furthermore both species differ in both sexes in the shape of the postocular spots, which are more rectangular and narrower, with a somewhat serrated posterior margin in
<taxonomicName id="41FE1BF2F75AFFFBFF45DF66F8289840" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[180,341,1371,1397]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F75AFFFBFF45DF66F8289840" box="[180,341,1371,1397]" italics="true" pageId="10" pageNumber="11">P. halmaherae</emphasis>
</taxonomicName>
(compare
<figureCitation id="1EC57CF4F75AFFFBFE3CDF61FB699840" box="[461,532,1371,1397]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="10" pageNumber="11">Figs 9</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFDD1DF61FB439840" box="[544,574,1371,1397]" captionStart="FIGURES 1114" captionStartId="10.[151,264,941,965]" captionTargetBox="[154,1434,180,911]" captionTargetId="figure-18@10.[151,1436,179,915]" captionTargetPageId="10" captionText="FIGURES 1114. Pseudagrion acutidens, female paratype: (11) habitus lateral, (12) head dorsal, (13) pronotum, (14) ovipositor lateral. Photos: M. Seehausen." pageId="10" pageNumber="11">12</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFDBBDF61FB159840" box="[586,616,1371,1397]" captionStart="FIGURES 3338" captionStartId="23.[151,264,1906,1930]" captionTargetBox="[152,1430,672,1878]" captionTargetId="figure-223@23.[151,1436,666,1880]" captionTargetPageId="23" captionText="FIGURES 3338. Pseudagrion halmaherae, holotype male: (33) habitus lateral, (34) appendages lateral, (35) appendages dorsal, (36) cercus dorsolateral, (37) head dorsal, (38) pronotum. Photos: M. Seehausen." pageId="10" pageNumber="11">37</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFD85DF61FBEE9840" box="[628,659,1371,1397]" captionStart="FIGURES 3942" captionStartId="25.[151,264,1173,1197]" captionTargetBox="[153,1432,182,1146]" captionTargetId="figure-18@25.[151,1436,179,1148]" captionTargetPageId="25" captionText="FIGURES 3942. Pseudagrion halmaherae, female paratype: (39) habitus lateral, (40) head dorsal, (41) pronotum, (42) ovipositor lateral. Photos: M. Seehausen." pageId="10" pageNumber="11">40</figureCitation>
,
<figureCitation id="1EC57CF4F75AFFFBFD6FDF61FB959840" box="[670,744,1371,1397]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="10" pageNumber="11">79, 86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75AFFFBFF36DF45F8C998D4" pageId="10" pageNumber="11" type="distribution">
<paragraph id="86416071F75AFFFBFF36DF45F82498AC" blockId="10.[151,1437,1083,1506]" box="[199,345,1407,1433]" pageId="10" pageNumber="11">
<emphasis id="B48ABC63F75AFFFBFF36DF45F82498AC" bold="true" box="[199,345,1407,1433]" pageId="10" pageNumber="11">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F75AFFFBFF36DF99F8C998D4" blockId="10.[151,1437,1083,1506]" pageId="10" pageNumber="11">
This species is restricted to the northern
<collectingCountry id="FEE920E1F75AFFFBFD7ADF99FA419888" box="[651,828,1443,1469]" name="Indonesia" pageId="10" pageNumber="11">Maluku Islands</collectingCountry>
. Records are available from Morotai, Halmahera, Obi, Bacan, and Buru (
<figureCitation id="1EC57CF4F75AFFFBFE93DFFDF8DB98D4" box="[354,422,1479,1505]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="10" pageNumber="11">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7EDCEBC2BA24A32AF5C9044788D6C5AE">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="7240F7281305CB2105808AC847C2EA8D">Seehausen, Malte</mods:namePart>
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<mods:namePart id="D2C46FA8D3D358C5934ADE0EEA9CB3CE">Kalkman, Vincent J.</mods:namePart>
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<mods:affiliation id="61F30D377C7F31CEBEE053BAA4C71164">Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865</mods:affiliation>
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<mods:namePart id="52B424D96EFEAE5D8918AAD72ABF966C">Bedjanič, Matjaž</mods:namePart>
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<emphasis id="B48ABC63F75AFFFBFF66DC09F8DF9B7B" bold="true" box="[151,418,1587,1614]" italics="true" pageId="10" pageNumber="11">Pseudagrion buenafei</emphasis>
<bibRefCitation id="E26F1D80F75AFFFBFE58DC09FB3B9B78" author="Muller, R. A." box="[425,582,1587,1613]" pageId="10" pageNumber="11" pagination="377 - 379" refId="ref41132" refString="Muller, R. A. (1996) Pseudagrion buenafei spec. nov. from Mindanao, the Philippines (Zygoptera: Coenagrionidae). Odonatologica, 25 (4), 377-379." type="journal article" year="1996">Müller, 1996</bibRefCitation>
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Figures: 1516 (
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, envelope); 80 (head
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); 97 (pterothorax
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); 114a, b (abdomen
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); 131ac (caudal appendages
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).
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:
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379,
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(description
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;
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appendages; Mindanao,
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<treatmentCitation id="075F4660F75AFFFBFF36DCE9FB599BDE" author="Hamalainen, M. &amp; Muller, R. A." box="[199,548,1747,1772]" page="261" pageId="10" pageNumber="11" year="1997">
<bibRefCitation id="E26F1D80F75AFFFBFF36DCE9F8889BD9" author="Hamalainen, M. &amp; Muller, R. A." box="[199,501,1747,1772]" pageId="10" pageNumber="11" pagination="249 - 315" refId="ref40073" refString="Hamalainen, M. &amp; Muller, R. A. (1997) Synopsis of the Philippine Odonata, with lists of species recorded from forty islands. Odonatologica, 26 (3), 249-315." type="journal article" year="1997">Hämäläinen &amp; Müller (1997)</bibRefCitation>
: 261
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(Mindanao).
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Examined specimens from the
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series
</emphasis>
(
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)
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<collectingDate id="E204BF59F75AFFFBFEA9DD04F8D59A6D" box="[344,424,1854,1880]" pageId="10" pageNumber="11" value="1991-10">x.1991</collectingDate>
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,
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,
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,
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leg., ex.
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,
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, JvT no. 17838 (
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)
</materialsCitation>
;—
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, 24./
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<collectingDate id="E204BF59F75AFFFBFB47DD58FC539A48" box="[1206,1326,1890,1917]" pageId="10" pageNumber="11" value="1991-11-25">25.xi.1991</collectingDate>
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,
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,
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,
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<collectingCounty id="6F2018FDF75AFFFBFCFCDDBCFAFC9A95" box="[781,897,1926,1952]" pageId="10" pageNumber="11">Mindanao</collectingCounty>
,
<collectingCountry id="FEE920E1F75AFFFBFC7FDDBCFD729A95" box="[910,1039,1926,1952]" name="Philippines" pageId="10" pageNumber="11">Philippines</collectingCountry>
,
<elevation id="0DD38742F75AFFFBFBEADDBCFD899A95" box="[1051,1268,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="10" pageNumber="11" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">
<quantity id="4106CD94F75AFFFBFBEADDBCFDCA9A95" box="[1051,1207,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="10" pageNumber="11" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">12001400 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F75AFFFBFAF1DDBCFCE19A95" box="[1280,1436,1926,1952]" pageId="10" pageNumber="11">Alex Buenafe</collectorName>
leg., ex.
<collectorName id="2B0B05A7F75AFFFBFF04DD90F88B9AF1" box="[245,502,1962,1988]" pageId="10" pageNumber="11">Coll. Roland A. Müller</collectorName>
,
<collectionCode id="E0EFF8B4F75AFFFBFDF1DD90FB2A9AF1" box="[512,599,1962,1988]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="11" type="Museum">RMNH</collectionCode>
, no. 1506118
</materialsCitation>
;—
<materialsCitation id="36966A2CF75AFFFAFCE7DD90F89C9D84" collectingDate="1995-09" collectionCode="RMNH" collectorName="Alex Buenafe &amp; Coll. Roland A. Muller" country="Philippines" county="Mindanao" elevation="1650" lastPageId="11" lastPageNumber="12" location="Babang creek" municipality="Mt. Apo" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="North Cotabato" typeStatus="paratype">
<specimenCount id="90F8ABF8F75AFFFBFCE7DD90FA4C9AF0" box="[790,817,1962,1989]" count="1" pageId="10" pageNumber="11" type="male">1♂</specimenCount>
<typeStatus id="5945DED3F75AFFFBFCC9DD90FAE69AF1" box="[824,923,1962,1988]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
,
<date id="F24046B1F75AFFFBFC54DD90FA809AF0" box="[933,1021,1962,1989]" pageId="10" pageNumber="11" value="1995-09">
<collectingDate id="E204BF59F75AFFFBFC54DD90FA809AF0" box="[933,1021,1962,1989]" pageId="10" pageNumber="11" value="1995-09">ix.1995</collectingDate>
</date>
,
<location id="832136AAF75AFFFBFBF6DD90FDDF9AF1" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75AFFFBFBF6DD90FDDF9AF1" box="[1031,1186,1962,1988]" country="Philippines" county="Mindanao" municipality="Mt. Apo" name="Babang creek" pageId="10" pageNumber="11" stateProvince="North Cotabato">Babang creek</location>
,
<location id="832136AAF75AFFFBFB5DDD90FC549AF1" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75AFFFBFB5DDD90FC549AF1" box="[1196,1321,1962,1988]" country="Philippines" county="Mindanao" municipality="Mt. Apo" name="Lake Agko" pageId="10" pageNumber="11" stateProvince="North Cotabato">Lake Agko</location>
,
<location id="832136AAF75AFFFBFAC2DD90FCE59AF1" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75AFFFBFAC2DD90FCE59AF1" box="[1331,1432,1962,1988]" country="Philippines" county="Mindanao" municipality="Mt. Apo" name="Ilomavis" pageId="10" pageNumber="11" stateProvince="North Cotabato">Ilomavis</location>
,
<collectingMunicipality id="6625FA0BF75AFFFBFF66DDF4F9819ADD" box="[151,252,1998,2024]" pageId="10" pageNumber="11">Mt. Apo</collectingMunicipality>
,
<collectingRegion id="443AAE93F75AFFFBFEFADDF4F8BD9ADD" box="[267,448,1998,2024]" country="Philippines" name="North Cotabato" pageId="10" pageNumber="11">North Cotabato</collectingRegion>
,
<collectingCounty id="6F2018FDF75AFFFBFE3EDDF4FB3E9ADD" box="[463,579,1998,2024]" pageId="10" pageNumber="11">Mindanao</collectingCounty>
,
<collectingCountry id="FEE920E1F75AFFFBFDA2DDF4FBA99ADD" box="[595,724,1998,2024]" name="Philippines" pageId="10" pageNumber="11">Philippines</collectingCountry>
,
<elevation id="0DD38742F75AFFFBFD12DDF4FABE9ADD" box="[739,963,1998,2024]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="10" pageNumber="11" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<quantity id="4106CD94F75AFFFBFD12DDF4FAFF9ADD" box="[739,898,1998,2024]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="10" pageNumber="11" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">16001700 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F75AFFFBFC23DDF4FD0D9ADD" box="[978,1136,1998,2024]" pageId="10" pageNumber="11">Alex Buenafe</collectorName>
leg., ex.
<collectorName id="2B0B05A7F75AFFFAFB15DDF4F9989D84" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">Coll. Roland A. Müller</collectorName>
,
<collectionCode id="E0EFF8B4F75BFFFAFF1EDAADF83B9D84" box="[239,326,151,177]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12" type="Museum">RMNH</collectionCode>
, no. 1506120
</materialsCitation>
;—
<materialsCitation id="36966A2CF75BFFFAFDF4DAACF9809DCC" collectingDate="1995-11-24" collectionCode="RMNH" collectorName="Alex Buenafe &amp; Coll. Roland A. Muller" country="Philippines" county="Mindanao" elevation="1150" location="Mansabilan creek" municipality="Mebadiang" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Bukidnon" typeStatus="paratype">
<specimenCount id="90F8ABF8F75BFFFAFDF4DAACFB5D9D84" box="[517,544,150,177]" count="1" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
<typeStatus id="5945DED3F75BFFFAFDD7DAADFBF49D84" box="[550,649,151,177]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
,
<date id="F24046B1F75BFFFAFD62DAACFA709D84" box="[659,781,150,177]" pageId="11" pageNumber="12" value="1995-11-24">
<collectingDate id="E204BF59F75BFFFAFD62DAACFA709D84" box="[659,781,150,177]" pageId="11" pageNumber="12" value="1995-11-24">24.xi.1995</collectingDate>
</date>
,
<location id="832136AAF75BFFFAFCE9DAADFA9D9D84" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75BFFFAFCE9DAADFA9D9D84" box="[792,992,151,177]" country="Philippines" county="Mindanao" municipality="Mebadiang" name="Mansabilan creek" pageId="11" pageNumber="12" stateProvince="Bukidnon">Mansabilan creek</location>
,
<location id="832136AAF75BFFFAFC18DAADFDE79D84" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75BFFFAFC18DAADFDE79D84" box="[1001,1178,151,177]" country="Philippines" county="Mindanao" municipality="Mebadiang" name="Mt. Kalatungan" pageId="11" pageNumber="12" stateProvince="Bukidnon">Mt. Kalatungan</location>
,
<location id="832136AAF75BFFFAFB55DAADFC739D84" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75BFFFAFB55DAADFC739D84" box="[1188,1294,151,177]" country="Philippines" county="Mindanao" municipality="Mebadiang" name="Dumatap" pageId="11" pageNumber="12" stateProvince="Bukidnon">Dumatap</location>
,
<collectingMunicipality id="6625FA0BF75BFFFAFAE6DAADFCE59D84" box="[1303,1432,151,177]" pageId="11" pageNumber="12">Mebadiang</collectingMunicipality>
,
<collectingRegion id="443AAE93F75BFFFAFF66DA81F8759DE0" box="[151,264,187,213]" country="Philippines" name="Bukidnon" pageId="11" pageNumber="12">Bukidnon</collectingRegion>
,
<collectingCounty id="6F2018FDF75BFFFAFEE5DA81F8F59DE0" box="[276,392,187,213]" pageId="11" pageNumber="12">Mindanao</collectingCounty>
,
<collectingCountry id="FEE920E1F75BFFFAFE65DA81FB689DE0" box="[404,533,187,213]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
,
<elevation id="0DD38742F75BFFFAFDD1DA81FB859DE0" box="[544,760,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="11" pageNumber="12" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">
<quantity id="4106CD94F75BFFFAFDD1DA81FBC69DE0" box="[544,699,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="11" pageNumber="12" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">10001300 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F75BFFFAFCF2DA81FAE39DE0" box="[771,926,187,213]" pageId="11" pageNumber="12">Alex Buenafe</collectorName>
leg., ex.
<collectorName id="2B0B05A7F75BFFFAFBF7DA81FC769DE0" box="[1030,1291,187,213]" pageId="11" pageNumber="12">Coll. Roland A. Müller</collectorName>
,
<collectionCode id="E0EFF8B4F75BFFFAFAE6DA81FC139DE0" box="[1303,1390,187,213]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12" type="Museum">RMNH</collectionCode>
, no. 1506121
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F75BFFFAFF36DB39FBC29C74" blockId="11.[151,1436,150,321]" pageId="11" pageNumber="12">
<bibRefCitation id="E26F1D80F75BFFFAFF36DB39F8149C28" author="Muller, R. A." box="[199,361,259,285]" pageId="11" pageNumber="12" pagination="377 - 379" refId="ref41132" refString="Muller, R. A. (1996) Pseudagrion buenafei spec. nov. from Mindanao, the Philippines (Zygoptera: Coenagrionidae). Odonatologica, 25 (4), 377-379." type="journal article" year="1996">Müller (1996)</bibRefCitation>
stated that the
<typeStatus id="5945DED3F75BFFFAFDD2DB39FBF89C28" box="[547,645,259,285]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
should be deposited at
<collectionCode id="E0EFF8B4F75BFFFAFC51DB39FAA49C28" box="[928,985,259,285]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
, but
<bibRefCitation id="E26F1D80F75BFFFAFBE6DB39FD899C28" author="Hamalainen, M." box="[1047,1268,259,285]" pageId="11" pageNumber="12" pagination="369 - 392" refId="ref40033" refString="Hamalainen, M. (2017) Corrections to the depository records of 22 holotypes of Odonata, originally in the collection of Roland A. Muller. Notulae odonatologicae, 8 (10), 369-392." type="journal article" year="2017">Hämäläinen (2017)</bibRefCitation>
corrected that statement: the
<typeStatus id="5945DED3F75BFFFAFECADB1DF8E09C74" box="[315,413,295,321]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
was deposited at
<collectionCode id="E0EFF8B4F75BFFFAFD95DB1DFBC19C74" box="[612,700,295,321]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12" type="Museum">RMNH</collectionCode>
.
</paragraph>
<caption id="D28130F9F75BFFFAFF66DE93FD9399F4" box="[151,1262,1193,1217]" pageId="11" pageNumber="12" startId="11.[151,264,1193,1217]" targetBox="[154,1434,343,1167]" targetPageId="11" targetType="figure">
<paragraph id="86416071F75BFFFAFF66DE93FD9399F4" blockId="11.[151,1262,1193,1217]" box="[151,1262,1193,1217]" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF66DE93F83399F4" bold="true" box="[151,334,1193,1217]" pageId="11" pageNumber="12">FIGURES 1516</emphasis>
.
<taxonomicName id="41FE1BF2F75BFFFAFEABDE93FB4499F4" authorityName="Muller" authorityYear="1996" box="[346,569,1193,1217]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="buenafei">
<emphasis id="B48ABC63F75BFFFAFEABDE93FB4499F4" box="[346,569,1193,1217]" italics="true" pageId="11" pageNumber="12">Pseudagrion buenafei</emphasis>
</taxonomicName>
, holotype male: (
<emphasis id="B48ABC63F75BFFFAFD1DDE93FA7899F4" bold="true" box="[748,773,1193,1217]" pageId="11" pageNumber="12">15</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F75BFFFAFC49DE93FAAC99F4" bold="true" box="[952,977,1193,1217]" pageId="11" pageNumber="12">16</emphasis>
) envelope. Photos: RMNH.
</paragraph>
</caption>
<paragraph id="86416071F75BFFFAFF36DF31FB3B9810" blockId="11.[151,1437,1291,2038]" box="[199,582,1291,1317]" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF36DF31FB749810" bold="true" box="[199,521,1291,1317]" pageId="11" pageNumber="12">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F75BFFFAFDE8DF31FB419810" box="[537,572,1291,1317]" count="1" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
)
</paragraph>
<paragraph id="86416071F75BFFFAFF36DF15FB86985B" blockId="11.[151,1437,1291,2038]" pageId="11" pageNumber="12">
<materialsCitation id="36966A2CF75BFFFAFF36DF15FB85985B" collectingDate="1991-10" collectionCode="RMNH" collectorName="Alex Buenafe &amp; Coll. Roland A. Muller" country="Philippines" location="Katanglad Mountains" municipality="Atongan River" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Bukidnon">
<emphasis id="B48ABC63F75BFFFAFF36DF15FB56987C" bold="true" box="[199,555,1327,1353]" pageId="11" pageNumber="12">
<collectingCountry id="FEE920E1F75BFFFAFF36DF15F832987C" box="[199,335,1327,1353]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
, Mindanao Island
</emphasis>
:
<specimenCount id="90F8ABF8F75BFFFAFDCDDF15FB1D987C" box="[572,608,1327,1353]" count="1" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
,
<date id="F24046B1F75BFFFAFD9FDF15FBC3987C" box="[622,702,1327,1353]" pageId="11" pageNumber="12" value="1991-10">
<collectingDate id="E204BF59F75BFFFAFD9FDF15FBC3987C" box="[622,702,1327,1353]" pageId="11" pageNumber="12" value="1991-10">x.1991</collectingDate>
</date>
,
<collectingMunicipality id="6625FA0BF75BFFFAFD3BDF15FA0E987C" box="[714,883,1327,1353]" pageId="11" pageNumber="12">Atongan River</collectingMunicipality>
,
<location id="832136AAF75BFFFAFC71DF15FD0A987C" LSID="urn:lsid:plazi:treatment:0E57D167F75AFFFDFF66DC09FBD39E68:832136AAF75BFFFAFC71DF15FD0A987C" box="[896,1143,1327,1353]" country="Philippines" municipality="Atongan River" name="Katanglad Mountains" pageId="11" pageNumber="12" stateProvince="Bukidnon">Katanglad Mountains</location>
,
<collectingRegion id="443AAE93F75BFFFAFB72DF15FD89987C" box="[1155,1268,1327,1353]" country="Philippines" name="Bukidnon" pageId="11" pageNumber="12">Bukidnon</collectingRegion>
,
<collectorName id="2B0B05A7F75BFFFAFAF1DF15FCE1987C" box="[1280,1436,1327,1353]" pageId="11" pageNumber="12">Alex Buenafe</collectorName>
leg., ex.
<collectorName id="2B0B05A7F75BFFFAFF06DF69F8819858" box="[247,508,1363,1389]" pageId="11" pageNumber="12">Coll. Roland A. Müller</collectorName>
,
<collectionCode id="E0EFF8B4F75BFFFAFDF6DF6EFB23985B" box="[519,606,1364,1390]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12" type="Museum">RMNH</collectionCode>
, no. 1506113
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F75BFFFAFF36DF4DF88F98A4" blockId="11.[151,1437,1291,2038]" box="[199,498,1399,1425]" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF36DF4DF88F98A4" bold="true" box="[199,498,1399,1425]" pageId="11" pageNumber="12">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F75BFFFAFF36DFA1F80B9B14" blockId="11.[151,1437,1291,2038]" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF36DFA1F8799880" box="[199,260,1435,1461]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
(
<figureCitation id="1EC57CF4F75BFFFAFEE2DFA1F8169883" box="[275,363,1435,1462]" captionStart="FIGURES 1516" captionStartId="11.[151,264,1193,1217]" captionTargetBox="[154,1434,343,1167]" captionTargetId="figure-70@11.[151,1436,342,1167]" captionTargetPageId="11" captionText="FIGURES 1516. Pseudagrion buenafei, holotype male: (15) habitus lateral, (16) envelope. Photos: RMNH." pageId="11" pageNumber="12">Figs 15</figureCitation>
,
<figureCitation id="1EC57CF4F75BFFFAFE86DFA1F8EA9880" box="[375,407,1435,1461]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="11" pageNumber="12">80</figureCitation>
): Labrum reddish brown to obscure blackish; anteclypeus red; postclypeus black; antefrons black, with anterior margin and laterally red; postfrons and vertex black; postocular spots and occipital bar red; rear of head black, but a pale patch along the centre of the eye margins; base and scape of antennae black, following segments brownish.
</paragraph>
<paragraph id="86416071F75BFFFAFF36DC11FDEB9A28" blockId="11.[151,1437,1291,2038]" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF36DC11F8659B70" box="[199,280,1579,1605]" italics="true" pageId="11" pageNumber="12">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F75BFFFAFED7DC11F8019B73" box="[294,380,1579,1606]" captionStart="FIGURES 1516" captionStartId="11.[151,264,1193,1217]" captionTargetBox="[154,1434,343,1167]" captionTargetId="figure-70@11.[151,1436,342,1167]" captionTargetPageId="11" captionText="FIGURES 1516. Pseudagrion buenafei, holotype male: (15) habitus lateral, (16) envelope. Photos: RMNH." pageId="11" pageNumber="12">Figs 15</figureCitation>
,
<figureCitation id="1EC57CF4F75BFFFAFE79DC11F8D59B70" box="[392,424,1579,1605]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="11" pageNumber="12">97</figureCitation>
): Pronotum with anterior lobe black, its posterior centre reddish, extending towards middle lobe; middle pronotal lobe otherwise black, with a centrally divided reddish rectangular patch in the middle, and on each side a large reddish patch dorsolaterally; propleuron black above, broadly reddish below and to rear; posterior lobe of pronotum blackish, with reddish centre; apical edge of posterior lobe convex. Pterothorax dorsally black, with antehumeral stripe red (may be interrupted); laterally red, with a black stripe each at posterior quarter of interpleural and metapleural suture, interpleural suture with a black spot anterior of the black stripe; ventrally pale, may be covered with whitish pruinosity. Legs blackish, with coxae and trochanters reddish.
</paragraph>
<paragraph id="86416071F75BFFFAFF36DD1DFA069A50" blockId="11.[151,1437,1291,2038]" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF36DD1DF84C9A74" box="[199,305,1831,1857]" italics="true" pageId="11" pageNumber="12">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F75BFFFAFEB1DD1DF8EB9A77" box="[320,406,1831,1858]" captionStart="FIGURES 1516" captionStartId="11.[151,264,1193,1217]" captionTargetBox="[154,1434,343,1167]" captionTargetId="figure-70@11.[151,1436,342,1167]" captionTargetPageId="11" captionText="FIGURES 1516. Pseudagrion buenafei, holotype male: (15) habitus lateral, (16) envelope. Photos: RMNH." pageId="11" pageNumber="12">Figs 15</figureCitation>
,
<figureCitation id="1EC57CF4F75BFFFAFE53DD1DF88B9A74" box="[418,502,1831,1857]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="11" pageNumber="12">114a, b</figureCitation>
): S1 red; S2 red with a small black triangle marking dorsally at posterior end; S37 dorsally black, laterally reddish; S810 red with black apical ring.
</paragraph>
<paragraph id="86416071F75BFFFDFF36DD55FA629DCC" blockId="11.[151,1437,1291,2038]" lastBlockId="12.[151,1436,151,861]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B48ABC63F75BFFFAFF36DD55F8D59ABC" box="[199,424,1903,1929]" italics="true" pageId="11" pageNumber="12">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F75BFFFAFE49DD55FB359ABC" box="[440,584,1903,1929]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="11" pageNumber="12">Figs 131ac</figureCitation>
): Cercus blackish; slightly shorter than S10; in lateral view subrectangular, inter-lobe notch placed low on apex; in dorsal view cercus narrow, with apex of superior lobe of cercus curved inward (more clearly so in dorsolateral view); an inward directed projection is situated at just after half-length on the superior lobe of the cercus; a small subbasal interior projection on inferior lobe of cercus, visible in dorsal and dorsolateral views, this bulge-like projection bearing a sharp proximal edge; paraproct reddish, around half as long as cercus, in lateral view somewhat triangular and tapered, with rounded apex; in dorsal view broadly rounded, concave bowl-like, with small black tooth at medial edge.
</paragraph>
<paragraph id="86416071F75CFFFDFF36DB39FAAC9C28" blockId="12.[151,1436,151,861]" box="[199,977,258,285]" pageId="12" pageNumber="13">Wings with 14.517.5 px in forewing, and 14.515.5 px in hindwing.</paragraph>
<paragraph id="86416071F75CFFFDFF36DB1DFD0F9C50" blockId="12.[151,1436,151,861]" pageId="12" pageNumber="13">Measurements (mm): Total length (with appendages) 46.349.0; abdomen length (without appendages) 37.5 39.4; hindwing 26.430.0; forewing 27.730.0; Pt in forewing 0.70.8; male cercus 0.7.</paragraph>
<paragraph id="86416071F75CFFFDFF36DB54FDC19CBC" blockId="12.[151,1436,151,861]" box="[199,1212,366,393]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DB54FB7A9CBD" bold="true" box="[199,519,366,392]" pageId="12" pageNumber="13">Characterization of female</emphasis>
[not examined, characterization adopted from
<bibRefCitation id="E26F1D80F75CFFFDFBE5DB55FDC99CBC" author="Muller, R. A." box="[1044,1204,367,393]" pageId="12" pageNumber="13" pagination="377 - 379" refId="ref41132" refString="Muller, R. A. (1996) Pseudagrion buenafei spec. nov. from Mindanao, the Philippines (Zygoptera: Coenagrionidae). Odonatologica, 25 (4), 377-379." type="journal article" year="1996">Müller (1996)</bibRefCitation>
]
</paragraph>
<paragraph id="86416071F75CFFFDFF36DBA9FA189C98" blockId="12.[151,1436,151,861]" box="[199,869,403,429]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DBA9F8709C98" box="[199,269,403,429]" italics="true" pageId="12" pageNumber="13">Head:</emphasis>
As in male, but with a black triangular mark at frons.
</paragraph>
<paragraph id="86416071F75CFFFDFF36DB8DFC369CE4" blockId="12.[151,1436,151,861]" box="[199,1355,439,465]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DB8DF8659CE4" box="[199,280,439,465]" italics="true" pageId="12" pageNumber="13">Thorax</emphasis>
: Pronotum and pterothorax as in male, but red replaced by greenish-yellow; legs as in the male.
</paragraph>
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<emphasis id="B48ABC63F75CFFFDFF36DBE1F8479CC0" box="[199,314,475,501]" italics="true" pageId="12" pageNumber="13">Abdomen:</emphasis>
S1 reddish-brown; S2 reddish-brown, with narrow black middorsal marking, widened apically; S3 10 dorsally black, laterally reddish-brown.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75CFFFDFF36D818F8F59E20" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="86416071F75CFFFDFF36D818F8479F09" blockId="12.[151,1436,151,861]" box="[199,314,546,572]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36D818F8479F09" bold="true" box="[199,314,546,572]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F75CFFFDFF36D87DF8F59E20" blockId="12.[151,1436,151,861]" pageId="12" pageNumber="13">
Both sexes of this species are separated from congeneric species, and especially the co-occurring
<taxonomicName id="41FE1BF2F75CFFFDFAF0D87DFCE89F54" authorityName="Brauer" authorityYear="1868" box="[1281,1429,583,609]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F75CFFFDFAF0D87DFCE89F54" box="[1281,1429,583,609]" italics="true" pageId="12" pageNumber="13">P. pilidorsum</emphasis>
</taxonomicName>
, by the morphology of the male cercus (compare
<figureCitation id="1EC57CF4F75CFFFDFD31D851FA319FB0" box="[704,844,619,645]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="12" pageNumber="13">Figs 131ac</figureCitation>
,
<figureCitation id="1EC57CF4F75CFFFDFCA9D851FAD19FB0" box="[856,940,619,645]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="12" pageNumber="13">143ac</figureCitation>
), as well as by the combination of a largely black frons and vertex (compare
<figureCitation id="1EC57CF4F75CFFFDFDE4D8B5FBE79F9C" box="[533,666,655,681]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="12" pageNumber="13">Figs 80, 92</figureCitation>
), a black humeral stripe (compare
<figureCitation id="1EC57CF4F75CFFFDFBDCD8B5FDBD9F9C" box="[1069,1216,655,681]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="12" pageNumber="13">Figs 97, 109</figureCitation>
), and blackish legs (compare
<figureCitation id="1EC57CF4F75CFFFDFEFBD889F81F9FF8" box="[266,354,690,717]" captionStart="FIGURES 1516" captionStartId="11.[151,264,1193,1217]" captionTargetBox="[154,1434,343,1167]" captionTargetId="figure-70@11.[151,1436,342,1167]" captionTargetPageId="11" captionText="FIGURES 1516. Pseudagrion buenafei, holotype male: (15) habitus lateral, (16) envelope. Photos: RMNH." pageId="12" pageNumber="13">Figs 15</figureCitation>
,
<figureCitation id="1EC57CF4F75CFFFDFE81D889F8C79FF8" box="[368,442,691,717]" captionStart="FIGURES 6365" captionStartId="36.[151,264,1267,1291]" captionTargetBox="[158,1429,802,1237]" captionTargetId="figure-183@36.[151,1436,798,1241]" captionTargetPageId="36" captionText="FIGURES 6365. Pseudagrion pilidorsum, syntype male: (63) habitus lateral, (64) pterothorax lateral, head dorsal, (65) labels. Photos 63, 65: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner; photo 64: M. Seehausen." pageId="12" pageNumber="13">63, 64</figureCitation>
,
<figureCitation id="1EC57CF4F75CFFFDFE39D888F88B9FF8" box="[456,502,690,717]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="12" pageNumber="13">165</figureCitation>
).
<taxonomicName id="41FE1BF2F75CFFFDFDFDD889FA799FF8" authorityName="Muller" authorityYear="1996" box="[524,772,691,717]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="buenafei">
<emphasis id="B48ABC63F75CFFFDFDFDD889FA799FF8" box="[524,772,691,717]" italics="true" pageId="12" pageNumber="13">Pseudagrion buenafei</emphasis>
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is also larger (total length
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) than
<taxonomicName id="41FE1BF2F75CFFFDFAF4D889FCE19FF8" authorityName="Brauer" authorityYear="1868" box="[1285,1436,691,717]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F75CFFFDFAF4D889FCE19FF8" box="[1285,1436,691,717]" italics="true" pageId="12" pageNumber="13">P. pilidorsum</emphasis>
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(
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). Additionally, the male is separated from all congeners by the almost entirely red abdominal segment S8 (
<figureCitation id="1EC57CF4F75CFFFDFF35D8C1F8679E20" box="[196,282,762,789]" captionStart="FIGURES 1516" captionStartId="11.[151,264,1193,1217]" captionTargetBox="[154,1434,343,1167]" captionTargetId="figure-70@11.[151,1436,342,1167]" captionTargetPageId="11" captionText="FIGURES 1516. Pseudagrion buenafei, holotype male: (15) habitus lateral, (16) envelope. Photos: RMNH." pageId="12" pageNumber="13">Figs 15</figureCitation>
,
<figureCitation id="1EC57CF4F75CFFFDFED7D8C1F8069E20" box="[294,379,763,789]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="12" pageNumber="13">114a, b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75CFFFDFF36D924FBD39E68" box="[199,686,798,861]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="86416071F75CFFFDFF36D924F8249E0D" blockId="12.[151,1436,151,861]" box="[199,345,798,824]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36D924F8249E0D" bold="true" box="[199,345,798,824]" pageId="12" pageNumber="13">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F75CFFFDFF36D979FBD39E68" blockId="12.[151,1436,151,861]" box="[199,686,835,861]" pageId="12" pageNumber="13">
Endemic to Mindanao,
<collectingCountry id="FEE920E1F75CFFFDFE3FD979FB319E68" box="[462,588,835,861]" name="Philippines" pageId="12" pageNumber="13">Philippines</collectingCountry>
(
<figureCitation id="1EC57CF4F75CFFFDFDAAD979FBDC9E68" box="[603,673,835,861]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="12" pageNumber="13">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,420 @@
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<taxonomicName id="41FE1BF2F75CFFFDFF66D994FB3A9EFD" ID-CoL="6WD8B" authority="Selys, 1876" authorityName="Selys" authorityYear="1876" box="[151,583,942,969]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F75CFFFDFF66D994FB3A9EFD" bold="true" box="[151,583,942,969]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF66D994F8CA9EFC" bold="true" box="[151,439,942,969]" italics="true" pageId="12" pageNumber="13">Pseudagrion coriaceum</emphasis>
<bibRefCitation id="E26F1D80F75CFFFDFE4FD994FB3A9EFD" author="Selys Longchamps, E. de" box="[446,583,942,968]" pageId="12" pageNumber="13" pagination="247 - 322" refId="ref42034" refString="Selys Longchamps, E. de (1876) Synopsis des Agrionines (suite de la 5 me Legion: Agrion). Bulletins de l'academy royale des sciences, des lettres et des beaux-arts de Belgique, 45, Series 2, 42, 247-322 + 496 - 539." type="journal article" year="1876">Selys, 1876</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="86416071F75CFFFDFF66D9C9F8B2991F" blockId="12.[151,1436,1011,1127]" pageId="12" pageNumber="13">
Figures: 1719 (
<typeStatus id="5945DED3F75CFFFDFECED9C9F8E3993E" box="[319,414,1011,1035]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
<specimenCount id="90F8ABF8F75CFFFDFE54D9CEF8C7993E" box="[421,442,1012,1035]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
, labels); 81 (head
<specimenCount id="90F8ABF8F75CFFFDFD89D9C9FBF0993E" box="[632,653,1011,1035]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
); 98 (pterothorax
<specimenCount id="90F8ABF8F75CFFFDFCB6D9C9FA21993E" box="[839,860,1011,1035]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
); 115a, b (abdomen
<specimenCount id="90F8ABF8F75CFFFDFBC1D9C9FD38993E" box="[1072,1093,1011,1035]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
); 132ac (caudal appendages
<specimenCount id="90F8ABF8F75CFFFDFA8BD9C9FCF2993E" box="[1402,1423,1011,1035]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
); 152153 (field photos
<specimenCount id="90F8ABF8F75CFFFDFE5CDE28F8BF991F" box="[429,450,1042,1066]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75CFFFDFF66DE0AFD539952" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="86416071F75CFFFDFF66DE0AFD539952" blockId="12.[151,1436,1011,1127]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="A6EE475FF75CFFFDFF66DE0AFD539952" pageId="12" pageNumber="13">
<taxonomicName id="41FE1BF2F75CFFFDFF66DE0AF8F2997D" ID-CoL="6WD8B" authorityName="Selys" authorityYear="1876" box="[151,399,1072,1096]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F75CFFFDFF66DE0AF8F2997D" box="[151,399,1072,1096]" italics="true" pageId="12" pageNumber="13">Pseudagrion coriaceum</emphasis>
</taxonomicName>
:
<treatmentCitation id="075F4660F75CFFFDFE50DE0AFB1D997D" author="Selys Longchamps, E. de" box="[417,608,1072,1096]" page="513" pageId="12" pageNumber="13" year="1876">
<bibRefCitation id="E26F1D80F75CFFFDFE50DE0AFB56997D" author="Selys Longchamps, E. de" box="[417,555,1072,1096]" pageId="12" pageNumber="13" pagination="247 - 322" refId="ref42034" refString="Selys Longchamps, E. de (1876) Synopsis des Agrionines (suite de la 5 me Legion: Agrion). Bulletins de l'academy royale des sciences, des lettres et des beaux-arts de Belgique, 45, Series 2, 42, 247-322 + 496 - 539." type="journal article" year="1876">Selys (1876)</bibRefCitation>
: 513
</treatmentCitation>
(description
<specimenCount id="90F8ABF8F75CFFFDFD01DE0BFA78997D" box="[752,773,1073,1096]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
,
<collectingCountry id="FEE920E1F75CFFFDFCE2DE0AFA1F997D" box="[787,866,1072,1096]" name="Indonesia" pageId="12" pageNumber="13">Ambon</collectingCountry>
);—
<treatmentCitation id="075F4660F75CFFFDFC79DE0AFD4F997C" author="Ris, F." box="[904,1074,1072,1097]" page="97" pageId="12" pageNumber="13" year="1915">
<bibRefCitation id="E26F1D80F75CFFFDFC79DE0AFD74997D" author="Ris, F." box="[904,1033,1072,1096]" pageId="12" pageNumber="13" pagination="81 - 131" refId="ref41673" refString="Ris, F. (1915 b) Neuer Beitrag zur Kenntnis der Odonatenfauna der Neu-Guinea-Region. Nova Guinea, 13, 81-131." type="journal article" year="1915">Ris (1915b)</bibRefCitation>
: 97
</treatmentCitation>
98,
<figureCitation id="1EC57CF4F75CFFFDFB98DE0AFDC8997C" box="[1129,1205,1072,1097]" captionStart="FIGURES 1719" captionStartId="13.[151,264,1205,1229]" captionTargetBox="[158,1429,240,1177]" captionTargetId="figure-40@13.[151,1436,234,1179]" captionTargetPageId="13" captionText="FIGURES 1719. Pseudagrion coriaceum, lectotype male: (17) habitus dorsal, (18) pterothorax lateral, (19) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner." pageId="12" pageNumber="13">Fig. 19</figureCitation>
(
<specimenCount id="90F8ABF8F75CFFFDFB38DE0BFDA1997C" box="[1225,1244,1073,1097]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
appendages; key; Seram);—
<treatmentCitation id="075F4660F75CFFFDFEDFDE75FB769952" author="Campion, H." box="[302,523,1103,1127]" page="263" pageId="12" pageNumber="13" year="1921">
<bibRefCitation id="E26F1D80F75CFFFDFEDFDE75F8A69952" author="Campion, H." box="[302,475,1103,1127]" pageId="12" pageNumber="13" pagination="262 - 264" refId="ref39008" refString="Campion, H. (1921) A note on some dragonflies from Amboina. The entomologist, 54 (702), 262-264." type="journal article" year="1921">Campion (1921)</bibRefCitation>
: 263
</treatmentCitation>
(
<collectingCountry id="FEE920E1F75CFFFDFDE9DE75FB159952" box="[536,616,1103,1127]" name="Indonesia" pageId="12" pageNumber="13">Ambon</collectingCountry>
);—
<treatmentCitation id="075F4660F75CFFFDFD7FDE75FA509952" author="Ris, F." box="[654,813,1103,1127]" page="146" pageId="12" pageNumber="13" year="1929">
<bibRefCitation id="E26F1D80F75CFFFDFD7FDE75FB839952" author="Ris, F." box="[654,766,1103,1127]" pageId="12" pageNumber="13" pagination="139 - 147" refId="ref41731" refString="Ris, F. (1929) Fauna Buruana. Odonata gesammelt von L. J. Toxopeus auf Buru, 1921 - 1922, nebst einigen Odonaten von Amboina. (2. Teil, Zygoptera). Treubia Supplement, 7, 139-147." type="journal article" year="1929">Ris (1929)</bibRefCitation>
: 146
</treatmentCitation>
(description
<specimenCount id="90F8ABF8F75CFFFDFC42DE6AFABA9952" box="[947,967,1104,1127]" count="1" pageId="12" pageNumber="13" type="female"></specimenCount>
;
<collectingCountry id="FEE920E1F75CFFFDFC23DE75FD5E9952" box="[978,1059,1103,1127]" name="Indonesia" pageId="12" pageNumber="13">Ambon</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75CFFFFFF66DEAFFC399CE4" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="86416071F75CFFFDFF66DEAFFBA9999A" blockId="12.[151,1437,1173,2028]" box="[151,724,1173,1199]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF66DEAFFBFE999A" bold="true" box="[151,643,1173,1199]" pageId="12" pageNumber="13">
Examined specimens from the
<typeStatus id="5945DED3F75CFFFDFDF7DEAFFB44999A" box="[518,569,1173,1199]" pageId="12" pageNumber="13">type</typeStatus>
series
</emphasis>
(
<specimenCount id="90F8ABF8F75CFFFDFD63DEAFFBB1999A" box="[658,716,1173,1199]" count="2" pageId="12" pageNumber="13" type="male">2♂♂</specimenCount>
)
</paragraph>
<paragraph id="86416071F75CFFFDFF36DE80FA1399CD" blockId="12.[151,1437,1173,2028]" pageId="12" pageNumber="13">
<materialsCitation id="36966A2CF75CFFFDFF36DE80FDC899E6" box="[199,1205,1209,1236]" collectionCode="NHMW" collectorName="Dr Doleschal" country="Indonesia" location="Ambon" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<specimenCount id="90F8ABF8F75CFFFDFF36DE80F9A199E1" box="[199,220,1210,1236]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
<typeStatus id="5945DED3F75CFFFDFF13DE83F83199E6" box="[226,332,1209,1235]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
, 1859, Amboina [
<collectingCountry id="FEE920E1F75CFFFDFDFCDE83FB1A99E6" box="[525,615,1209,1235]" name="Indonesia" pageId="12" pageNumber="13">Ambon</collectingCountry>
,
<collectingCountry id="FEE920E1F75CFFFDFD81DE83FB9E99E6" box="[624,739,1209,1235]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
],
<collectorName id="2B0B05A7F75CFFFDFD02DE80FAFA99E6" box="[755,903,1209,1236]" pageId="12" pageNumber="13">Dr Doleschal</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFC33DE80FD6099E1" box="[962,1053,1210,1236]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13" type="Museum">NHMW</collectionCode>
(
<figureCitation id="1EC57CF4F75CFFFDFBDADE83FDD699E6" box="[1067,1195,1209,1235]" captionStart="FIGURES 1719" captionStartId="13.[151,264,1205,1229]" captionTargetBox="[158,1429,240,1177]" captionTargetId="figure-40@13.[151,1436,234,1179]" captionTargetPageId="13" captionText="FIGURES 1719. Pseudagrion coriaceum, lectotype male: (17) habitus dorsal, (18) pterothorax lateral, (19) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner." pageId="12" pageNumber="13">Figs 1719</figureCitation>
)
</materialsCitation>
;—
<materialsCitation id="36966A2CF75CFFFDFB2BDE83FA1799CD" collectionCode="NHMW" collectorName="Dr Doleschal" country="Indonesia" location="Ambon" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="90F8ABF8F75CFFFDFB2BDE83FD8499E6" box="[1242,1273,1209,1235]" count="1" pageId="12" pageNumber="13" type="male">1♂</specimenCount>
<typeStatus id="5945DED3F75CFFFDFB0FDE83FCE599E6" box="[1278,1432,1209,1235]" pageId="12" pageNumber="13" type="paralectotype">paralectotype</typeStatus>
, 1859, Amboina [
<collectingCountry id="FEE920E1F75CFFFDFEA0DEE7F8D699C2" box="[337,427,1245,1271]" name="Indonesia" pageId="12" pageNumber="13">Ambon</collectingCountry>
,
<collectingCountry id="FEE920E1F75CFFFDFE47DEE7FB5499C2" box="[438,553,1245,1271]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
],
<collectorName id="2B0B05A7F75CFFFDFDCBDEE4FBB299C2" box="[570,719,1245,1272]" pageId="12" pageNumber="13">Dr Doleschal</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFCFCDEE4FA1799CD" box="[781,874,1246,1272]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13" type="Museum">NHMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F75CFFFDFF36DF38F80F980A" blockId="12.[151,1437,1173,2028]" pageId="12" pageNumber="13">
<typeStatus id="5945DED3F75CFFFDFF36DF38F84A9829" box="[199,311,1282,1308]" pageId="12" pageNumber="13" type="lectotype">Lectotype</typeStatus>
designation by St.
<bibRefCitation id="E26F1D80F75CFFFDFDFEDF3BFBC3982E" author="Quentin, D." box="[527,702,1281,1307]" pageId="12" pageNumber="13" pagination="253 - 279" refId="ref42308" refString="St. Quentin, D. (1970) Katalog der Odonaten-Typen im Naturhistorischen Museum Wien. Annalen des Naturhistorischen Museum Wien, 74, 253-279." type="journal article" year="1970">Quentin (1970)</bibRefCitation>
. Both males lack the last segments of the abdomen, including the caudal appendages.
</paragraph>
<paragraph id="86416071F75CFFFDFF36DF73FBDC9856" blockId="12.[151,1437,1173,2028]" box="[199,673,1353,1379]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DF73FB749856" bold="true" box="[199,521,1353,1379]" pageId="12" pageNumber="13">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F75CFFFDFDE9DF73FB2E9856" box="[536,595,1353,1379]" count="6" pageId="12" pageNumber="13" type="male">6♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F75CFFFDFDAEDF73FBE89856" box="[607,661,1353,1379]" count="3" pageId="12" pageNumber="13" type="female">3♀♀</specimenCount>
)
</paragraph>
<paragraph id="86416071F75CFFFDFF36DF57FC7F98E5" blockId="12.[151,1437,1173,2028]" pageId="12" pageNumber="13">
<materialsCitation id="36966A2CF75CFFFDFF36DF57FABE98B2" box="[199,963,1389,1415]" collectionCode="IRSNB" country="Indonesia" location="Indonesia" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1">
<emphasis id="B48ABC63F75CFFFDFF36DF57F88398B2" bold="true" box="[199,510,1389,1415]" pageId="12" pageNumber="13">
<collectingCountry id="FEE920E1F75CFFFDFF36DF57F84098B2" box="[199,317,1389,1415]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
,
<collectingCountry id="FEE920E1F75CFFFDFEB8DF57F88498B2" box="[329,505,1389,1415]" name="Ambon Island" pageId="12" pageNumber="13">Ambon Island</collectingCountry>
:
</emphasis>
<specimenCount id="90F8ABF8F75CFFFDFDF6DF57FB5498B2" box="[519,553,1389,1415]" count="1" pageId="12" pageNumber="13" type="female">1♀</specimenCount>
, without particular locality,
<collectionCode id="E0EFF8B4F75CFFFDFC9ADF57FABE98B2" box="[875,963,1389,1415]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="12" pageNumber="13">IRSNB</collectionCode>
</materialsCitation>
;—
<materialsCitation id="36966A2CF75CFFFDFC16DF57F8DA9899" collectingDate="1941-04-23" collectionCode="RMNH" collectorName="Soengai &amp; Waitoemoe &amp; Mains" country="Ambon Island" pageId="12" pageNumber="13" specimenCount="3" specimenCount-male="3">
<specimenCount id="90F8ABF8F75CFFFDFC16DF57FD6198B2" box="[999,1052,1389,1415]" count="3" pageId="12" pageNumber="13" type="male">3♂♂</specimenCount>
,
<date id="F24046B1F75CFFFDFBD9DF57FDDC98B2" box="[1064,1185,1389,1415]" pageId="12" pageNumber="13" value="1941-04-23">
<collectingDate id="E204BF59F75CFFFDFBD9DF57FDDC98B2" box="[1064,1185,1389,1415]" pageId="12" pageNumber="13" value="1941-04-23">23.iv.1941</collectingDate>
</date>
,
<collectorName id="2B0B05A7F75CFFFDFB5FDF57FC7698B2" box="[1198,1291,1389,1415]" pageId="12" pageNumber="13">Soengai</collectorName>
,
<collectorName id="2B0B05A7F75CFFFDFAE7DF57FCE598B2" box="[1302,1432,1389,1415]" pageId="12" pageNumber="13">Waitoemoe</collectorName>
,
<collectorName id="2B0B05A7F75CFFFDFF66DFABF9A0989E" box="[151,221,1425,1451]" pageId="12" pageNumber="13">Mains</collectorName>
z lasoen,
<collectionCode id="E0EFF8B4F75CFFFDFEA1DFA8F8DA9899" box="[336,423,1426,1452]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1519109;—
<materialsCitation id="36966A2CF75CFFFDFD83DFABFCE59899" box="[626,1432,1425,1452]" collectingDate="1948-07-27" collectionCode="RMNH" collectorName="M. A. Lieftinck" country="Ambon Island" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1">
<specimenCount id="90F8ABF8F75CFFFDFD83DFABFBF79899" box="[626,650,1425,1452]" count="1" pageId="12" pageNumber="13" type="female">1♀</specimenCount>
,
<date id="F24046B1F75CFFFDFD67DFABFA65989E" box="[662,792,1425,1451]" pageId="12" pageNumber="13" value="1948-07-27">
<collectingDate id="E204BF59F75CFFFDFD67DFABFA65989E" box="[662,792,1425,1451]" pageId="12" pageNumber="13" value="1948-07-27">27.vii.1948</collectingDate>
</date>
, without particular locality,
<collectorName id="2B0B05A7F75CFFFDFBA6DFA8FC7C989E" box="[1111,1281,1425,1452]" pageId="12" pageNumber="13">M.A. Lieftinck</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFAB0DFA8FCE59899" box="[1345,1432,1426,1452]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1519104;—
<materialsCitation id="36966A2CF75CFFFDFEA4DF8FFD1E98E5" box="[341,1123,1461,1488]" collectingDate="1949-10-12" collectionCode="RMNH" collectorName="M. A. Lieftinck" country="Ambon Island" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1">
<specimenCount id="90F8ABF8F75CFFFDFEA4DF8FF81198E5" box="[341,364,1461,1488]" count="1" pageId="12" pageNumber="13" type="female">1♀</specimenCount>
,
<date id="F24046B1F75CFFFDFE89DF8FF89798FA" box="[376,490,1461,1487]" pageId="12" pageNumber="13" value="1949-10-12">
<collectingDate id="E204BF59F75CFFFDFE89DF8FF89798FA" box="[376,490,1461,1487]" pageId="12" pageNumber="13" value="1949-10-12">12.x.1949</collectingDate>
</date>
, without particular locality,
<collectorName id="2B0B05A7F75CFFFDFCD4DF8CFAB398FA" box="[805,974,1461,1488]" pageId="12" pageNumber="13">M.A. Lieftinck</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFBFDDF8CFD1E98E5" box="[1036,1123,1462,1488]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1658923.
</paragraph>
<paragraph id="86416071F75CFFFDFF36DFE3FA859B09" blockId="12.[151,1437,1173,2028]" pageId="12" pageNumber="13">
<materialsCitation id="36966A2CF75CFFFDFF36DFE3FDB698C6" box="[199,1227,1497,1524]" collectingDate="1912-09-18" collectionCode="SMF" collectorName="Elgner" country="Indonesia" location="Seram Island" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Seram Island">
<emphasis id="B48ABC63F75CFFFDFF36DFE3F88F98C6" bold="true" box="[199,498,1497,1523]" pageId="12" pageNumber="13">
<collectingCountry id="FEE920E1F75CFFFDFF36DFE3F84098C6" box="[199,317,1497,1523]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F75CFFFDFEBFDFE3F88F98C6" box="[334,498,1497,1523]" pageId="12" pageNumber="13">Seram Island</collectingRegion>
</emphasis>
:
<specimenCount id="90F8ABF8F75CFFFDFDF4DFE3FB5498C6" box="[517,553,1497,1523]" count="1" pageId="12" pageNumber="13" type="male">1♂</specimenCount>
,
<date id="F24046B1F75CFFFDFDCBDFE3FBC998C6" box="[570,692,1497,1523]" pageId="12" pageNumber="13" value="1912-09-18">
<collectingDate id="E204BF59F75CFFFDFDCBDFE3FBC998C6" box="[570,692,1497,1523]" pageId="12" pageNumber="13" value="1912-09-18">18.ix.1912</collectingDate>
</date>
, without particular locality,
<collectorName id="2B0B05A7F75CFFFDFBF1DFE3FD3698C6" box="[1024,1099,1497,1523]" pageId="12" pageNumber="13">Elgner</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFB63DFE3FDB698C6" box="[1170,1227,1497,1523]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="13">SMF</collectionCode>
</materialsCitation>
, no. 13924;—
<materialsCitation id="36966A2CF75CFFFDFA8BDFE3FBDF9B2D" collectingDate="1941-11-21" collectionCode="RMNH" collectorName="van der Harre" country="Indonesia" location="Wahai" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Seram Island">
<specimenCount id="90F8ABF8F75CFFFDFA8BDFE3FCEA98C6" box="[1402,1431,1497,1523]" count="1" pageId="12" pageNumber="13" type="male">1♂</specimenCount>
,
<date id="F24046B1F75CFFFDFF66DFC7F86F9B22" box="[151,274,1533,1559]" pageId="12" pageNumber="13" value="1941-11-21">
<collectingDate id="E204BF59F75CFFFDFF66DFC7F86F9B22" box="[151,274,1533,1559]" pageId="12" pageNumber="13" value="1941-11-21">21.xi.1941</collectingDate>
</date>
,
<location id="832136AAF75CFFFDFEECDFC7F81A9B22" LSID="urn:lsid:plazi:treatment:0E57D167F75CFFFFFF66D994FA529E0C:832136AAF75CFFFDFEECDFC7F81A9B22" box="[285,359,1533,1559]" country="Indonesia" name="Wahai" pageId="12" pageNumber="13" stateProvince="Seram Island">Wahai</location>
,
<collectorName id="2B0B05A7F75CFFFDFE83DFC4FB719B2D" box="[370,524,1533,1560]" pageId="12" pageNumber="13">van der Harre</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFDBADFC4FBDF9B2D" box="[587,674,1534,1560]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1519260;—
<materialsCitation id="36966A2CF75CFFFDFC97DFC7FA899B09" collectingDate="1996-11-30" collectionCode="MB" collectorName="M. Bedjanic" country="Indonesia" elevation="500" location="Roho village" municipality="Logging Base Camp" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Seram Island">
<specimenCount id="90F8ABF8F75CFFFDFC97DFC7FAFE9B2D" box="[870,899,1533,1560]" count="1" pageId="12" pageNumber="13" type="male">1♂</specimenCount>
,
<date id="F24046B1F75CFFFDFC7FDFC7FD749B22" box="[910,1033,1533,1559]" pageId="12" pageNumber="13" value="1996-11-30">
<collectingDate id="E204BF59F75CFFFDFC7FDFC7FD749B22" box="[910,1033,1533,1559]" pageId="12" pageNumber="13" value="1996-11-30">30.xi.1996</collectingDate>
</date>
, stream
<quantity id="4106CD94F75CFFFDFB97DFC7FDD09B2D" box="[1126,1197,1533,1560]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="m" value="500.0">
<elevation id="0DD38742F75CFFFDFB97DFC7FDD09B2D" box="[1126,1197,1533,1560]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="m" value="500.0">500 m</elevation>
</quantity>
NE of
<collectingMunicipality id="6625FA0BF75CFFFDFAF1DFC7F9A09B0E" pageId="12" pageNumber="13">Logging Base Camp</collectingMunicipality>
,
<quantity id="4106CD94F75CFFFDFF19DC1BF84B9B0E" box="[232,310,1569,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="km" value="1.5">1.5 km</quantity>
SW
<location id="832136AAF75CFFFDFE9CDC1BFB7D9B0E" LSID="urn:lsid:plazi:treatment:0E57D167F75CFFFFFF66D994FA529E0C:832136AAF75CFFFDFE9CDC1BFB7D9B0E" box="[365,512,1569,1595]" country="Indonesia" municipality="Logging Base Camp" name="Roho village" pageId="12" pageNumber="13" stateProvince="Seram Island">Roho village</location>
,
<locationDeviation id="97F37953F75CFFFDFDFADC1BFB8C9B0E" box="[523,753,1569,1595]" location="Wahai" pageId="12" pageNumber="13">
<quantity id="4106CD94F75CFFFDFDFADC1BFB2F9B0E" box="[523,594,1569,1595]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="13" unit="km" value="31.0">31 km</quantity>
SW of Wahai
</locationDeviation>
,
<collectorName id="2B0B05A7F75CFFFDFD0DDC18FAFA9B0E" box="[764,903,1569,1596]" pageId="12" pageNumber="13">M. Bedjanič</collectorName>
leg.,
<collectionCode id="E0EFF8B4F75CFFFDFC34DC18FA899B09" box="[965,1012,1570,1596]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="12" pageNumber="13">MB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F75CFFFDFF36DC7FF88F9B6A" blockId="12.[151,1437,1173,2028]" box="[199,498,1605,1631]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DC7FF88F9B6A" bold="true" box="[199,498,1605,1631]" pageId="12" pageNumber="13">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F75CFFFDFF36DC53FB3E9A26" blockId="12.[151,1437,1173,2028]" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DC53F8799BB6" box="[199,260,1641,1667]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
(
<figureCitation id="1EC57CF4F75CFFFDFEE3DC53F8159BB6" box="[274,360,1641,1667]" captionStart="FIGURES 1719" captionStartId="13.[151,264,1205,1229]" captionTargetBox="[158,1429,240,1177]" captionTargetId="figure-40@13.[151,1436,234,1179]" captionTargetPageId="13" captionText="FIGURES 1719. Pseudagrion coriaceum, lectotype male: (17) habitus dorsal, (18) pterothorax lateral, (19) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner." pageId="12" pageNumber="13">Figs 17</figureCitation>
,
<figureCitation id="1EC57CF4F75CFFFDFE82DC53F8EF9BB6" box="[371,402,1641,1667]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="12" pageNumber="13">81</figureCitation>
): Labrum, clypeus, antefrons and postfrons yellow; postfrons distinctly separated from black vertex around level of lateral ocelli; postocular spots large, reddish-brown, but they may be darkened and hardly visible; reddish occipital bar, but may be darkened and hardly visible; rear of head black, but broadly pale along the eye margins; eyes brownish, in life anteriorly yellow, continuing the yellow of the frons; base and scape antennae yellow, following segments brownish.
</paragraph>
<paragraph id="86416071F75CFFFCFF36DD27F89F9DE0" blockId="12.[151,1437,1173,2028]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B48ABC63F75CFFFDFF36DD27F8659A02" box="[199,280,1821,1847]" italics="true" pageId="12" pageNumber="13">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F75CFFFDFED7DD27F8DB9A02" box="[294,422,1821,1847]" captionStart="FIGURES 1719" captionStartId="13.[151,264,1205,1229]" captionTargetBox="[158,1429,240,1177]" captionTargetId="figure-40@13.[151,1436,234,1179]" captionTargetPageId="13" captionText="FIGURES 1719. Pseudagrion coriaceum, lectotype male: (17) habitus dorsal, (18) pterothorax lateral, (19) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner." pageId="12" pageNumber="13">Figs 17, 18</figureCitation>
,
<figureCitation id="1EC57CF4F75CFFFDFE40DD27F8AC9A02" box="[433,465,1821,1847]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="12" pageNumber="13">98</figureCitation>
): Anterior lobe of pronotum red, this colour extending towards middle lobe in posterior centre; middle pronotal lobe otherwise black, with a centrally divided red patch in the middle, and on each side a large red patch dorsolaterally, these may fuse laterally with red of anterior lobe; propleuron black above, broadly yellowish-red below and to rear; posterior lobe of pronotum red; apical edge of posterior lobe almost convex. Pterothorax red with black middorsal stripe; a black humeral stripe of same width; interpleural suture with a black stripe, and a dark dot anteriorly; metapleural suture with a black stripe, widened posteriorly; ventrally pale. Legs orange-reddish; outer side of femora with a blackish stripe narrowing to proximal; along spines of tibiae, and the waists blackish; spines black.
</paragraph>
<caption id="D28130F9F75DFFFCFF66DE8FFAD899C4" pageId="13" pageNumber="14" startId="13.[151,264,1205,1229]" targetBox="[158,1429,240,1177]" targetPageId="13" targetType="figure">
<paragraph id="86416071F75DFFFCFF66DE8FFAD899C4" blockId="13.[151,1436,1205,1266]" pageId="13" pageNumber="14">
<emphasis id="B48ABC63F75DFFFCFF66DE8FF83399F8" bold="true" box="[151,334,1205,1229]" pageId="13" pageNumber="14">FIGURES 1719</emphasis>
.
<taxonomicName id="41FE1BF2F75DFFFCFEABDE8FFB3099F8" authorityName="Selys" authorityYear="1876" box="[346,589,1205,1229]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F75DFFFCFEABDE8FFB3099F8" box="[346,589,1205,1229]" italics="true" pageId="13" pageNumber="14">Pseudagrion coriaceum</emphasis>
</taxonomicName>
, lectotype male: (
<emphasis id="B48ABC63F75DFFFCFCF5DE8FFA6099F8" bold="true" box="[772,797,1205,1229]" pageId="13" pageNumber="14">17</emphasis>
) habitus dorsal, (
<emphasis id="B48ABC63F75DFFFCFC3EDE8FFA9599F8" bold="true" box="[975,1000,1205,1229]" pageId="13" pageNumber="14">18</emphasis>
) pterothorax lateral, (
<emphasis id="B48ABC63F75DFFFCFB36DE8FFD9D99F8" bold="true" box="[1223,1248,1205,1229]" pageId="13" pageNumber="14">19</emphasis>
) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner.
</paragraph>
</caption>
<paragraph id="86416071F75DFFFCFF36DF79FDF29B24" blockId="13.[151,1437,1347,2022]" pageId="13" pageNumber="14">
<emphasis id="B48ABC63F75DFFFCFF36DF79F84C9868" box="[199,305,1347,1373]" italics="true" pageId="13" pageNumber="14">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F75DFFFCFEB2DF79F8A89868" box="[323,469,1347,1374]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="13" pageNumber="14">Figs 115a, b</figureCitation>
): S1 red, dorsally with black patch covering the anterior two-thirds of the segment, the patch may be divided in half by a red middorsal line; S2 red, but along posterior carina black; with blackish mushroom-like marking dorsally at posterior quarter, this marking may extend as a black middorsal stripe towards the segment base; S38 dorsally black, laterally pale yellow-red; S9 red, dorsally with a black middorsal stripe and with at least the posterior third to half black, the black margin may be serrated, but dorsum of S9 may likewise be completely black; S10 red, dorsally with a black “x”-like marking, variable in exact shape.
</paragraph>
<paragraph id="86416071F75DFFFCFF36DC21FB7F9BF0" blockId="13.[151,1437,1347,2022]" pageId="13" pageNumber="14">
<emphasis id="B48ABC63F75DFFFCFF36DC21F8D89B00" box="[199,421,1563,1589]" italics="true" pageId="13" pageNumber="14">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F75DFFFCFE43DC21FB429B03" box="[434,575,1563,1590]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="13" pageNumber="14">Figs 132ac</figureCitation>
): Cercus laterally dark, slightly shorter than S10; in lateral view cercus simple, subtriangular, inter-lobe notch not visible; in dorsal view apex of superior lobe rounded and deep inter-lobe notch visible; inferior lobe bulging smoothly inwards; paraproct pale yellow-ochre, as long as two-third of the cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, its medial edge blackish tubercular.
</paragraph>
<paragraph id="86416071F75DFFFCFF36DCF5FB7A9BDC" blockId="13.[151,1437,1347,2022]" box="[199,519,1743,1769]" pageId="13" pageNumber="14">
<emphasis id="B48ABC63F75DFFFCFF36DCF5FB7A9BDC" bold="true" box="[199,519,1743,1769]" pageId="13" pageNumber="14">Characterization of female</emphasis>
</paragraph>
<paragraph id="86416071F75DFFFCFF36DCC9FDFF9A4C" blockId="13.[151,1437,1347,2022]" pageId="13" pageNumber="14">
<emphasis id="B48ABC63F75DFFFCFF36DCC9F8709A38" box="[199,269,1779,1805]" italics="true" pageId="13" pageNumber="14">Head:</emphasis>
Labrum, and anteclypeus yellow-ochre; postclypeus blackish; antefrons and postfrons yellow-ochre, antefrons with blackish patch in centre and a black mid stripe towards postclypeus; vertex black from about level of lateral ocelli; postocular spots orange-ochre; occipital bar orange-ochre; rear of head black, but broadly pale along the eye margins; base and scape of antennae yellow-ochre, following segments brownish.
</paragraph>
<paragraph id="86416071F75DFFFCFF36DDB9FCE89AD0" blockId="13.[151,1437,1347,2022]" pageId="13" pageNumber="14">
<emphasis id="B48ABC63F75DFFFCFF36DDB9F8659AA8" box="[199,280,1923,1949]" italics="true" pageId="13" pageNumber="14">Thorax</emphasis>
: Anterior lobe of pronotum orange-ochre, its anterior margin black; middle pronotal lobe and posterior lobe as in male but orange-ochre; posterior lobe bearing two horns directed towards head, almost reaching the centre of the middle lobe; apical edge of posterior lobe almost convex. Pterothorax and legs as in male but orange-ochre.
</paragraph>
<paragraph id="86416071F75EFFFFFF36DAADFBE79C50" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="15">
<emphasis id="B48ABC63F75EFFFFFF36DAADF84C9D84" box="[199,305,151,177]" italics="true" pageId="14" pageNumber="15">Abdomen</emphasis>
: S1 orange, with a blackish dorsal marking at anterior two-third; S2 red, with a black middorsal stripe, widened to lateral at posterior quarter; S310 dorsally black, laterally orange-ochre; on S910 the orange-ochre expanded dorsally in centre, with the anterior and posterior margin, as well as the dorsum of each segment black. Cercus blackish, as long as half of S10; paraproct darkish ochre; in lateral view rounded; ovipositor pale yellowish-ochre, its apex not reaching to posterior end of S10; two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F75EFFFFFF36DB55FD169CBC" blockId="14.[151,1437,151,825]" box="[199,1131,366,393]" pageId="14" pageNumber="15">Wings of both sexes with 13.515.5 px in forewing, and 12.013.5 px in hindwing.</paragraph>
<paragraph id="86416071F75EFFFFFF36DBA9FC399CE4" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="15">Measurements of both sexes (mm): Total length (with appendages) 43.246.6; abdomen length (without appendages) 35.238.5; hindwing 24.026.7; forewing 25.428.0; Pt in forewing 0.81.0; male cercus 0.5.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75EFFFFFF36DBE0F9B29FC4" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="86416071F75EFFFFFF36DBE0F8479CC1" blockId="14.[151,1437,151,825]" box="[199,314,474,500]" pageId="14" pageNumber="15">
<emphasis id="B48ABC63F75EFFFFFF36DBE0F8479CC1" bold="true" box="[199,314,474,500]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F75EFFFFFF36DBC5F9B29FC4" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="15">
<taxonomicName id="41FE1BF2F75EFFFFFF36DBC5F8AB9F2D" authorityName="Selys" authorityYear="1876" box="[199,470,511,537]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F75EFFFFFF36DBC5F8AB9F2D" box="[199,470,511,537]" italics="true" pageId="14" pageNumber="15">Pseudagrion coriaceum</emphasis>
</taxonomicName>
is currently the only known species of the
<taxonomicName id="41FE1BF2F75EFFFFFC3EDBC5FD239F2C" authorityName="Selys" authorityYear="1876" box="[975,1118,511,537]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B48ABC63F75EFFFFFC3EDBC5FD239F2C" box="[975,1118,511,537]" italics="true" pageId="14" pageNumber="15">Pseudagrion</emphasis>
</taxonomicName>
red-group which occurs on
<collectingCountry id="FEE920E1F75EFFFFFF66D819F9969F08" box="[151,235,547,573]" name="Indonesia" pageId="14" pageNumber="15">Ambon</collectingCountry>
and Seram. Males can be separated from all congeners by the distinct morphology of the cercus (
<figureCitation id="1EC57CF4F75EFFFFFAD1D819F9D89F54" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="14" pageNumber="15">Figs 132a c</figureCitation>
), being subtriangularly tapered with inferior lobe and inter-lobe notch not visible in lateral view. The only species with a superficially similar morphology of the male cercus is
<taxonomicName id="41FE1BF2F75EFFFFFCA3D851FA889FB0" authorityName="Seehausen, Kalkman &amp; Bedjanic" authorityYear="2025" box="[850,1013,619,645]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="halmaherae">
<emphasis id="B48ABC63F75EFFFFFCA3D851FA889FB0" box="[850,1013,619,645]" italics="true" pageId="14" pageNumber="15">P. halmaherae</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F75EFFFFFBF5D851FDEF9FB0" box="[1028,1170,619,645]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="14" pageNumber="15">Figs 137ac</figureCitation>
), but in this species the cercus is less tapered in lateral view and an inter-lobe notch is situated subapically at the lower margin. The obvious pterothorax pattern of
<taxonomicName id="41FE1BF2F75EFFFFFE68D889FB509FF9" authorityName="Selys" authorityYear="1876" box="[409,557,691,716]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coriaceum">
<emphasis id="B48ABC63F75EFFFFFE68D889FB509FF9" box="[409,557,691,716]" italics="true" pageId="14" pageNumber="15">P. coriaceum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F75EFFFFFDCED889FBB99FF8" box="[575,708,691,717]" captionStart="FIGURES 1719" captionStartId="13.[151,264,1205,1229]" captionTargetBox="[158,1429,240,1177]" captionTargetId="figure-40@13.[151,1436,234,1179]" captionTargetPageId="13" captionText="FIGURES 1719. Pseudagrion coriaceum, lectotype male: (17) habitus dorsal, (18) pterothorax lateral, (19) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner." pageId="14" pageNumber="15">Figs 17, 18</figureCitation>
,
<figureCitation id="1EC57CF4F75EFFFFFD23D889FB8D9FF8" box="[722,752,691,717]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="14" pageNumber="15">98</figureCitation>
,
<figureCitation id="1EC57CF4F75EFFFFFD0ED888FA199FF8" box="[767,868,690,717]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="14" pageNumber="15">152, 153</figureCitation>
) separates this species from several congeners as well.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF75EFFFFFF36D8C0FA529E0C" box="[199,815,762,825]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="86416071F75EFFFFFF36D8C0F8249E21" blockId="14.[151,1437,151,825]" box="[199,345,762,788]" pageId="14" pageNumber="15">
<emphasis id="B48ABC63F75EFFFFFF36D8C0F8249E21" bold="true" box="[199,345,762,788]" pageId="14" pageNumber="15">Distribution</emphasis>
</paragraph>
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This species is restricted to
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and Seram (
<figureCitation id="1EC57CF4F75EFFFFFD2CD925FA5F9E0C" box="[733,802,799,825]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="14" pageNumber="15">Fig. 3</figureCitation>
).
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View file

@ -0,0 +1,427 @@
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<mods:title id="806C5B70B3A3D46D642048D3DB9379DA">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<emphasis id="B48ABC63F772FFD3FF66DB1CF8AD9C75" bold="true" box="[151,464,294,321]" italics="true" pageId="34" pageNumber="35">Pseudagrion perfuscatum</emphasis>
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Figures: 6162 (
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appendages; West Borneo [
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(Mount Kinabalu,
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: 10
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(
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Dow
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: 37
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(Kalimantan,
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,
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,
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<emphasis id="B48ABC63F772FFD3FF66D83EF8F69F29" box="[151,395,516,540]" italics="true" pageId="34" pageNumber="35">Pseudagrion pruinosum</emphasis>
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:
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: 275
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(Mt. Merinjak [
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,
<collectingCountry id="FEE920E1F772FFD3FC83D83EFAAB9F29" box="[882,982,516,540]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
]).
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</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF772FFD2FF66D870FC399B79" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="86416071F772FFD3FF66D870FB8C9F50" blockId="34.[151,1436,586,721]" box="[151,753,586,613]" pageId="34" pageNumber="35">
<emphasis id="B48ABC63F772FFD3FF66D870FBFE9F51" bold="true" box="[151,643,586,613]" pageId="34" pageNumber="35">
Examined specimens from the
<typeStatus id="5945DED3F772FFD3FDF7D871FB449F50" box="[518,569,587,613]" pageId="34" pageNumber="35">type</typeStatus>
series
</emphasis>
(
<specimenCount id="90F8ABF8F772FFD3FD65D871FBC59F50" box="[660,696,587,613]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
,
<specimenCount id="90F8ABF8F772FFD3FD35D871FB989F50" box="[708,741,587,613]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
)
</paragraph>
<paragraph id="86416071F772FFD3FF36D855FABD9FE4" blockId="34.[151,1436,586,721]" pageId="34" pageNumber="35">
<materialsCitation id="36966A2CF772FFD3FF36D855FB3A9F98" collectingDate="1931-11-05" collectingDateMax="1931-11-16" collectingDateMin="1931-11-05" collectionCode="RMNH" collectorName="West Borneo, L &amp; L. Coomans de Ruiter" location="West Borneo" municipality="Coomans de Ruiter" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="90F8ABF8F772FFD3FF36D855F9A19FBC" box="[199,220,623,649]" count="1" pageId="34" pageNumber="35" type="male"></specimenCount>
<typeStatus id="5945DED3F772FFD3FF19D855F8309FBC" box="[232,333,623,649]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
,
<date id="F24046B1F772FFD3FEADD854F8849FBC" box="[348,505,622,649]" pageId="34" pageNumber="35" value="1931-11-05" valueMax="1931-11-16" valueMin="1931-11-05">
<collectingDate id="E204BF59F772FFD3FEADD854F8849FBC" box="[348,505,622,649]" pageId="34" pageNumber="35" value="1931-11-05" valueMax="1931-11-16" valueMin="1931-11-05">5.16.xi.1931</collectingDate>
</date>
, Singkawang, weg naar Bengkajang,
<collectorName id="2B0B05A7F772FFD3FC45D855FD0D9FBC" box="[948,1136,623,649]" pageId="34" pageNumber="35">
<location id="832136AAF772FFD3FC45D855FD319FBC" LSID="urn:lsid:plazi:treatment:0E57D167F772FFD5FF66DB1CFB449DCC:832136AAF772FFD3FC45D855FD319FBC" box="[948,1100,623,649]" municipality="Coomans de Ruiter" name="West Borneo" pageId="34" pageNumber="35">West Borneo</location>
, L
</collectorName>
.
<collectingMunicipality id="6625FA0BF772FFD3FB8ED855FC1C9FBC" box="[1151,1377,623,649]" pageId="34" pageNumber="35">Coomans de Ruiter</collectingMunicipality>
leg.,
<collectionCode id="E0EFF8B4F772FFD3FF66D8A9F9939F98" box="[151,238,659,685]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="34" pageNumber="35" type="Museum">RMNH</collectionCode>
, JvT no. 2842 (
<figureCitation id="1EC57CF4F772FFD3FE40D8A9FB439F98" box="[433,574,659,685]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="34" pageNumber="35">Figs 6162</figureCitation>
)
</materialsCitation>
;—
<materialsCitation id="36966A2CF772FFD3FD9FD8A9FAC19FE4" collectingDate="1931-11-05" collectingDateMax="1931-11-16" collectingDateMin="1931-11-05" collectionCode="RMNH" collectorName="West Borneo, L &amp; L. Coomans de Ruiter" location="West Borneo" municipality="Coomans de Ruiter" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="90F8ABF8F772FFD3FD9FD8A9FBFB9F98" box="[622,646,659,685]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
<typeStatus id="5945DED3F772FFD3FD62D8A9FB8B9F98" box="[659,758,659,685]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
,
<date id="F24046B1F772FFD3FCF6D8A8FAD99F98" box="[775,932,658,685]" pageId="34" pageNumber="35" value="1931-11-05" valueMax="1931-11-16" valueMin="1931-11-05">
<collectingDate id="E204BF59F772FFD3FCF6D8A8FAD99F98" box="[775,932,658,685]" pageId="34" pageNumber="35" value="1931-11-05" valueMax="1931-11-16" valueMin="1931-11-05">5.16.xi.1931</collectingDate>
</date>
, Singkawang, weg naar Bengkajang,
<collectorName id="2B0B05A7F772FFD3FA94D8A9F8719FE4" pageId="34" pageNumber="35">
<location id="832136AAF772FFD3FA94D8A9F9919FE4" LSID="urn:lsid:plazi:treatment:0E57D167F772FFD5FF66DB1CFB449DCC:832136AAF772FFD3FA94D8A9F9919FE4" municipality="Coomans de Ruiter" name="West Borneo" pageId="34" pageNumber="35">West Borneo</location>
, L
</collectorName>
.
<collectingMunicipality id="6625FA0BF772FFD3FEE6D88DF8929FE4" box="[279,495,695,721]" pageId="34" pageNumber="35">Coomans de Ruiter</collectingMunicipality>
leg.,
<collectionCode id="E0EFF8B4F772FFD3FDDCD88DFBF99FE4" box="[557,644,695,721]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="34" pageNumber="35" type="Museum">RMNH</collectionCode>
, JvT no. 2842 (
<figureCitation id="1EC57CF4F772FFD3FCC1D88DFACE9FE4" box="[816,947,695,721]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="34" pageNumber="35">Figs 6162</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="D28130F9F772FFD3FF66DCF1F9969A32" pageId="34" pageNumber="35" startId="34.[151,264,1739,1763]" targetBox="[154,1434,744,1711]" targetPageId="34" targetType="figure">
<paragraph id="86416071F772FFD3FF66DCF1F9969A32" blockId="34.[151,1437,1739,1799]" pageId="34" pageNumber="35">
<emphasis id="B48ABC63F772FFD3FF66DCF1F8309BD6" bold="true" box="[151,333,1739,1763]" pageId="34" pageNumber="35">FIGURES 6162</emphasis>
.
<taxonomicName id="41FE1BF2F772FFD3FEABDCF6FB239BD1" authorityName="Lieftinck" authorityYear="1937" box="[346,606,1740,1764]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="perfuscatum">
<emphasis id="B48ABC63F772FFD3FEABDCF6FB239BD1" box="[346,606,1740,1764]" italics="true" pageId="34" pageNumber="35">Pseudagrion perfuscatum</emphasis>
</taxonomicName>
, lectotype male, paralectotype female: (
<emphasis id="B48ABC63F772FFD3FC04DCF1FD739BD6" bold="true" box="[1013,1038,1739,1763]" pageId="34" pageNumber="35">61</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F772FFD3FB31DCF1FDA49BD6" bold="true" box="[1216,1241,1739,1763]" pageId="34" pageNumber="35">62</emphasis>
) envelope. Photos: RMNH.
</paragraph>
</caption>
<paragraph id="86416071F772FFD3FF36DD60FB269A41" blockId="34.[151,1436,1882,1981]" box="[199,603,1882,1908]" pageId="34" pageNumber="35">
<emphasis id="B48ABC63F772FFD3FF36DD60FB749A41" bold="true" box="[199,521,1882,1908]" pageId="34" pageNumber="35">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F772FFD3FDE9DD60FB2C9A41" box="[536,593,1882,1908]" count="3" pageId="34" pageNumber="35" type="male">3♂♂</specimenCount>
)
</paragraph>
<paragraph id="86416071F772FFD3FF36DD44FD259A88" blockId="34.[151,1436,1882,1981]" pageId="34" pageNumber="35">
<materialsCitation id="36966A2CF772FFD3FF36DD44FC439AAD" box="[199,1342,1918,1945]" collectingDate="1925-01-01" collectingDateMax="1925-01-07" collectingDateMin="1925-01-01" collectionCode="ZMH" collectorName="Lebang Hara &amp; Sammelreise Prof Dr H. Winkler" country="Indonesia" location="Borneo Island" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Borneo Island">
<emphasis id="B48ABC63F772FFD3FF36DD44F8859AAD" bold="true" box="[199,504,1918,1944]" pageId="34" pageNumber="35">
<collectingCountry id="FEE920E1F772FFD3FF36DD44F8409AAD" box="[199,317,1918,1944]" name="Indonesia" pageId="34" pageNumber="35">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F772FFD3FEBCDD44F8859AAD" box="[333,504,1918,1944]" pageId="34" pageNumber="35">Borneo Island</collectingRegion>
</emphasis>
:
<specimenCount id="90F8ABF8F772FFD3FDF8DD44FB539AAD" box="[521,558,1918,1944]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
,
<date id="F24046B1F772FFD3FDCCDD44FBC39AAC" box="[573,702,1918,1945]" pageId="34" pageNumber="35" value="1925-01-01" valueMax="1925-01-07" valueMin="1925-01-01">
<collectingDate id="E204BF59F772FFD3FDCCDD44FBC39AAC" box="[573,702,1918,1945]" pageId="34" pageNumber="35" value="1925-01-01" valueMax="1925-01-07" valueMin="1925-01-01">1.7.i.1925</collectingDate>
</date>
,
<collectorName id="2B0B05A7F772FFD3FD3CDD44FA1E9AAD" box="[717,867,1918,1944]" pageId="34" pageNumber="35">Lebang Hara</collectorName>
,
<collectorName id="2B0B05A7F772FFD3FC80DD44FD939AAD" box="[881,1262,1918,1944]" pageId="34" pageNumber="35">Sammelreise Prof Dr H. Winkler</collectorName>
,
<collectionCode id="E0EFF8B4F772FFD3FB0ADD44FC439AAD" box="[1275,1342,1918,1944]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="34" pageNumber="35" type="Museum">ZMH</collectionCode>
</materialsCitation>
;—
<materialsCitation id="36966A2CF772FFD3FA93DD44FA349A89" collectingDate="1931-11-16" collectionCode="RMNH" collectorName="Singkawang &amp; West Borneo, L. C." country="Indonesia" location="West Borneo" pageId="34" pageNumber="35" specimenCount="2" specimenCount-male="2" stateProvince="Borneo Island">
<specimenCount id="90F8ABF8F772FFD3FA93DD44FCEA9AAD" box="[1378,1431,1918,1944]" count="2" pageId="34" pageNumber="35" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F772FFD3FF66DD98F86F9A89" box="[151,274,1954,1980]" pageId="34" pageNumber="35" value="1931-11-16">
<collectingDate id="E204BF59F772FFD3FF66DD98F86F9A89" box="[151,274,1954,1980]" pageId="34" pageNumber="35" value="1931-11-16">16.xi.1931</collectingDate>
</date>
,
<collectorName id="2B0B05A7F772FFD3FEECDD98F8D79A89" box="[285,426,1954,1980]" pageId="34" pageNumber="35">Singkawang</collectorName>
,
<collectorName id="2B0B05A7F772FFD3FE44DD98FBF89A89" box="[437,645,1954,1980]" pageId="34" pageNumber="35">
<location id="832136AAF772FFD3FE44DD98FB359A89" LSID="urn:lsid:plazi:treatment:0E57D167F772FFD5FF66DB1CFB449DCC:832136AAF772FFD3FE44DD98FB359A89" box="[437,584,1954,1980]" country="Indonesia" name="West Borneo" pageId="34" pageNumber="35" stateProvince="Borneo Island">West Borneo</location>
, L.C.
</collectorName>
d.R. leg.,
<collectionCode id="E0EFF8B4F772FFD3FD03DD98FA349A89" box="[754,841,1954,1980]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="34" pageNumber="35" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1658921, 1658922.
</paragraph>
<paragraph id="86416071F773FFD2FF36DAACF88F9D84" blockId="35.[151,1437,150,2042]" box="[199,498,150,177]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DAACF88F9D84" bold="true" box="[199,498,150,177]" pageId="35" pageNumber="36">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F773FFD2FF36DA80FB4B9C51" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DA80F8799DE1" box="[199,260,186,212]" italics="true" pageId="35" pageNumber="36">Head</emphasis>
(
<figureCitation id="1EC57CF4F773FFD2FEE4DA80F8119DE0" box="[277,364,186,213]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">Fig. 91</figureCitation>
): Labrum, clypeus, antefrons, postfrons, and vertex orange in preserved specimens but red in living individuals; a black spot posterior of median ocellus, and black patches along lateral ocelli towards orange colour of occipital bar; postocular spots orange-red, separated by black from postfrons and vertex; rear of head black, but yellow along the eye margins, may be covered with bluish pruinosity; base and scape of antennae orange-ochre, following segments brownish.
</paragraph>
<paragraph id="86416071F773FFD2FF36DB54FA819FB6" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DB54F8659CBD" box="[199,280,366,392]" italics="true" pageId="35" pageNumber="36">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F773FFD2FED4DB54F8079CBD" box="[293,378,366,392]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="35" pageNumber="36">Figs 61</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FE75DB54F8CF9CBD" box="[388,434,366,392]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">108</figureCitation>
):Anterior lobe of pronotum reddish, its margins washed-out blackish; middle pronotal lobe black, with a centrally divided red patch in the middle, and on each side a large red patch dorsolaterally; propleuron black above, pale orange below and to rear; posterior lobe of pronotum red, with its margins washed-out blackish; apical edge of posterior lobe convex. Pterothorax reddish-brown, with a black middorsal stripe; humeral suture at least with an elongate black patch at its posterior quarter, but may likewise be completely black; interpleural suture with a black line, a black spot may be present anteriorly; metapleural suture with a black streak situated at posterior quarter; venter pale; may be covered with bluish-white pruinosity up to interpleural suture. Legs pale orange; femora with black stripe outside; tibiae with black along spines; spines black.
</paragraph>
<paragraph id="86416071F773FFD2FF36D8B7F8739E27" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36D8B7F84C9F92" box="[199,305,653,679]" italics="true" pageId="35" pageNumber="36">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F773FFD2FEB0D8B7F8E79F92" box="[321,410,653,679]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="35" pageNumber="36">Figs 61</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FE56D8B7F8839F92" box="[423,510,653,680]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="35" pageNumber="36">125a, b</figureCitation>
): S1 red, with a blackish basal spot covering two-thirds of dorsum, segment may be completely covered with bluish-white pruinosity; S2 red, with black dorsally, may be completely covered with bluish-white pruinosity; S38 dorsally black, laterally pale yellowish; S9 red with its posterior third to half black; S10 black.
</paragraph>
<paragraph id="86416071F773FFD2FF36D926FBE49960" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36D926F8DB9E03" box="[199,422,796,822]" italics="true" pageId="35" pageNumber="36">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F773FFD2FE44D926FB399E03" box="[437,580,796,822]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="35" pageNumber="36">Figs 142ac</figureCitation>
): Cercus laterally blackish, but may be reddish at base; in lateral view cercus shorter than S10, the upper margin slightly convex, its apex almost square, inter-lobe notch barely or not apparent in this view; viewed dorsally the apical part of the superior lobe of the cercus curved inwards and runs as a ridge to the inter-lobe notch, with a separate inward directed projection proximal of this visible in dorsolateral view; another inwardly directed projection is situated more proximally at around the middle of the cercus; the inferior lobe of the cercus with a distinct ridge, best visible in dorsolateral view; a small bulge-like interior projection at the base of this ridge is visible in dorsal and dorsolateral view; paraproct pale orange, ventrally blackish; as long as two-thirds of the cercus; in lateral view subtriangular and tapered, with rounded apex; in dorsal view broadly rounded, concave bowl-like; blackish tubercular at medial edge.
</paragraph>
<paragraph id="86416071F773FFD2FF36DE65FB0C994C" blockId="35.[151,1437,150,2042]" box="[199,625,1119,1145]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DE65FB7A994C" bold="true" box="[199,519,1119,1145]" pageId="35" pageNumber="36">Characterization of female</emphasis>
(
<figureCitation id="1EC57CF4F773FFD2FDE6DE65FB15994C" box="[535,616,1119,1145]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="35" pageNumber="36">Fig. 61</figureCitation>
)
</paragraph>
<paragraph id="86416071F773FFD2FF36DEB9FA5899F4" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DEB9F87999A8" box="[199,260,1155,1181]" italics="true" pageId="35" pageNumber="36">Head</emphasis>
: As in male but orange-brown instead of reddish; black colour on vertex reduced to patches proximal of each lateral ocellus; postocular spots surrounded by black.
</paragraph>
<paragraph id="86416071F773FFD2FF36DEF1F8BC9865" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DEF1F86599D0" box="[199,280,1227,1253]" italics="true" pageId="35" pageNumber="36">Thorax</emphasis>
: Pronotum as in the male, but brownish-ochre instead of reddish; horns of posterior pronotal lobe directed towards head, reaching the centre of the middle lobe. Pterothorax as in male, but yellowish-ochre instead of reddish; humeral suture with a thin black line, and an elongated black patch at posterior quarter; upper third of mesinfraepisternum black.
</paragraph>
<paragraph id="86416071F773FFD2FF36DF60FBAB98D5" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DF60F84C9841" box="[199,305,1370,1396]" italics="true" pageId="35" pageNumber="36">Abdomen</emphasis>
: S12 dorsally black, laterally yellowish-ochre; S38 dorsally black, laterally yellowish; S910 dorsally black, lateral half yellowish-ochre. Cercus brownish; paraproct pale, in lateral view rounded; ovipositor pale yellowish-ochre, not reaching posterior margin of S10; two brownish styles reaching to beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F773FFD2FF36DFD3FD179B36" blockId="35.[151,1437,150,2042]" box="[199,1130,1513,1540]" pageId="35" pageNumber="36">Wings of both sexes with 11.514.0 px in forewing, and 10.513.0 px in hindwing.</paragraph>
<paragraph id="86416071F773FFD2FF36DC34FC399B79" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">Measurements of both sexes (mm): Total length (with appendages) 38.540.7; abdomen length (without appendages) 30.033.4; hindwing 19.023.0; forewing 22.023.8; Pt in forewing 0.80.9; male cercus 0.5.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF773FFD2FF36DC6FFB049ACF" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="86416071F773FFD2FF36DC6FF8479B5A" blockId="35.[151,1437,150,2042]" box="[199,314,1621,1647]" pageId="35" pageNumber="36">
<emphasis id="B48ABC63F773FFD2FF36DC6FF8479B5A" bold="true" box="[199,314,1621,1647]" pageId="35" pageNumber="36">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F773FFD2FF36DC43FB049ACF" blockId="35.[151,1437,150,2042]" pageId="35" pageNumber="36">
The males of
<taxonomicName id="41FE1BF2F773FFD2FE93DC40FB759BA6" authorityName="Lieftinck" authorityYear="1937" box="[354,520,1657,1683]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="perfuscatum">
<emphasis id="B48ABC63F773FFD2FE93DC40FB759BA6" box="[354,520,1657,1683]" italics="true" pageId="35" pageNumber="36">P. perfuscatum</emphasis>
</taxonomicName>
have the clypeus and frons red (
<figureCitation id="1EC57CF4F773FFD2FC86DC43FAB79BA6" box="[887,970,1657,1683]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">Fig. 91</figureCitation>
), a character which separates this species from almost all congeners except for
<taxonomicName id="41FE1BF2F773FFD2FDCEDCA4FBA99B82" authorityName="Brauer" authorityYear="1868" box="[575,724,1693,1719]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F773FFD2FDCEDCA4FBA99B82" box="[575,724,1693,1719]" italics="true" pageId="35" pageNumber="36">P. pilidorsum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F773FFD2FD15DCA7FA419B82" box="[740,828,1693,1719]" captionStart="FIGURES 6365" captionStartId="36.[151,264,1267,1291]" captionTargetBox="[158,1429,802,1237]" captionTargetId="figure-183@36.[151,1436,798,1241]" captionTargetPageId="36" captionText="FIGURES 6365. Pseudagrion pilidorsum, syntype male: (63) habitus lateral, (64) pterothorax lateral, head dorsal, (65) labels. Photos 63, 65: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner; photo 64: M. Seehausen." pageId="35" pageNumber="36">Figs 64</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FCB8DCA7FA149B82" box="[841,873,1693,1719]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">92</figureCitation>
). The morphology of the male caudal appendages of
<taxonomicName id="41FE1BF2F773FFD2FF44DCF8F8279BEE" authorityName="Lieftinck" authorityYear="1937" box="[181,346,1729,1755]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="perfuscatum">
<emphasis id="B48ABC63F773FFD2FF44DCF8F8279BEE" box="[181,346,1729,1755]" italics="true" pageId="35" pageNumber="36">P. perfuscatum</emphasis>
</taxonomicName>
and
<taxonomicName id="41FE1BF2F773FFD2FE61DCF8FB599BEE" authorityName="Brauer" authorityYear="1868" box="[400,548,1729,1755]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F773FFD2FE61DCF8FB599BEE" box="[400,548,1729,1755]" italics="true" pageId="35" pageNumber="36">P. pilidorsum</emphasis>
</taxonomicName>
is almost identical (
<figureCitation id="1EC57CF4F773FFD2FCF5DCFBFA8D9BEE" box="[772,1008,1729,1755]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="35" pageNumber="36">Figs 142ac, 143ac</figureCitation>
). But at least in fully mature males the colour pattern separates both species:
<taxonomicName id="41FE1BF2F773FFD2FDB8DCDFFBA29BCA" authorityName="Brauer" authorityYear="1868" box="[585,735,1765,1791]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F773FFD2FDB8DCDFFBA29BCA" box="[585,735,1765,1791]" italics="true" pageId="35" pageNumber="36">P. pilidorsum</emphasis>
</taxonomicName>
has less black on vertex and along the largely red postocular spots compared to
<taxonomicName id="41FE1BF2F773FFD2FE9FDD33FB699A16" authorityName="Lieftinck" authorityYear="1937" box="[366,532,1801,1827]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="perfuscatum">
<emphasis id="B48ABC63F773FFD2FE9FDD33FB699A16" box="[366,532,1801,1827]" italics="true" pageId="35" pageNumber="36">P. perfuscatum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F773FFD2FDD4DD32FBD79A16" box="[549,682,1800,1827]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">Figs 91, 92</figureCitation>
). Additionally mature males of
<taxonomicName id="41FE1BF2F773FFD2FBE3DD33FDC49A16" authorityName="Lieftinck" authorityYear="1937" box="[1042,1209,1801,1827]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="perfuscatum">
<emphasis id="B48ABC63F773FFD2FBE3DD33FDC49A16" box="[1042,1209,1801,1827]" italics="true" pageId="35" pageNumber="36">P. perfuscatum</emphasis>
</taxonomicName>
have the dorsum of S12 blackish and covered with pruinosity (
<figureCitation id="1EC57CF4F773FFD2FD70DD16FBA59A73" box="[641,728,1836,1862]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="35" pageNumber="36">Figs 61</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FD12DD16FA4A9A73" box="[739,823,1836,1863]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="35" pageNumber="36">125a, b</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FCB2DD16FADA9A73" box="[835,935,1836,1862]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="35" pageNumber="36">163, 164</figureCitation>
), the lower half of the pterothorax is covered with pruinosity as well, and usually there is a black humeral stripe (
<figureCitation id="1EC57CF4F773FFD2FC52DD6AFA819A5F" box="[931,1020,1872,1898]" captionStart="FIGURES 6162" captionStartId="34.[151,264,1739,1763]" captionTargetBox="[154,1434,744,1711]" captionTargetId="figure-153@34.[151,1436,742,1713]" captionTargetPageId="34" captionText="FIGURES 6162. Pseudagrion perfuscatum, lectotype male, paralectotype female: (61) habitus lateral, (62) envelope. Photos: RMNH." pageId="35" pageNumber="36">Figs 61</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FBF8DD6AFD489A5F" box="[1033,1077,1872,1898]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">108</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FBB2DD6AFDD89A5F" box="[1091,1189,1872,1898]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="35" pageNumber="36">163, 164</figureCitation>
)—making the colour pattern almost unmistakable; whereas
<taxonomicName id="41FE1BF2F773FFD2FDBBDD4FFB9D9ABB" authorityName="Brauer" authorityYear="1868" box="[586,736,1908,1934]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pilidorsum">
<emphasis id="B48ABC63F773FFD2FDBBDD4FFB9D9ABB" box="[586,736,1908,1934]" italics="true" pageId="35" pageNumber="36">P. pilidorsum</emphasis>
</taxonomicName>
has the dorsum of S12 reddish (
<figureCitation id="1EC57CF4F773FFD2FB90DD4EFD8C9ABB" box="[1121,1265,1908,1934]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="35" pageNumber="36">Figs 126a, b</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FB0FDD4EFC199ABB" box="[1278,1380,1908,1935]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="35" pageNumber="36">165, 166</figureCitation>
), the pruinosity on them is restricted to venter, and at least slightly expanded laterally, the pruinosity on the pterothorax is restricted to the venter and metepimeron, and a humeral stripe is lacking (
<figureCitation id="1EC57CF4F773FFD2FC19DD86FD149AE3" box="[1000,1129,1980,2006]" captionStart="FIGURES 6365" captionStartId="36.[151,264,1267,1291]" captionTargetBox="[158,1429,802,1237]" captionTargetId="figure-183@36.[151,1436,798,1241]" captionTargetPageId="36" captionText="FIGURES 6365. Pseudagrion pilidorsum, syntype male: (63) habitus lateral, (64) pterothorax lateral, head dorsal, (65) labels. Photos 63, 65: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner; photo 64: M. Seehausen." pageId="35" pageNumber="36">Figs 63, 64</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FB85DD86FDDD9AE3" box="[1140,1184,1980,2006]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="35" pageNumber="36">109</figureCitation>
,
<figureCitation id="1EC57CF4F773FFD2FB5DDD86FC6F9AE3" box="[1196,1298,1980,2007]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="35" pageNumber="36">165, 166</figureCitation>
). [Note: See discussion section for further information].
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF774FFD5FF36DAACFB449DCC" pageId="36" pageNumber="37" type="distribution">
<paragraph id="86416071F774FFD5FF36DAACF8249D85" blockId="36.[151,1437,150,249]" box="[199,345,150,176]" pageId="36" pageNumber="37">
<emphasis id="B48ABC63F774FFD5FF36DAACF8249D85" bold="true" box="[199,345,150,176]" pageId="36" pageNumber="37">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F774FFD5FF36DA81FB449DCC" blockId="36.[151,1437,150,249]" pageId="36" pageNumber="37">
This species is endemic to Borneo, and recorded from
<collectingCountry id="FEE920E1F774FFD5FCE9DA81FA1A9DE0" box="[792,871,187,213]" name="Brunei" pageId="36" pageNumber="37">Brunei</collectingCountry>
, Indonesian Kalimantan, as well as from Malaysian
<collectingCountry id="FEE920E1F774FFD5FF66DAE5F98A9DCC" box="[151,247,223,249]" name="Malaysia" pageId="36" pageNumber="37">Sarawak</collectingCountry>
and possibly
<collectingRegion id="443AAE93F774FFD5FE63DAE5F8AB9DCC" box="[402,470,223,249]" country="Malaysia" name="Sabah" pageId="36" pageNumber="37">Sabah</collectingRegion>
(
<figureCitation id="1EC57CF4F774FFD5FE17DAE5FB5A9DCC" box="[486,551,223,249]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="36" pageNumber="37">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1B9C9D9AB1EB1F53BFFECB7525C40787">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="5665F15CA2E305B081175DC3162E8DF6">Bedjanič, Matjaž</mods:namePart>
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<mods:affiliation id="A49E3C6E8CF7C81A56DDCC70C2796DBC">Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 &amp; National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author &amp; Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078</mods:affiliation>
<mods:nameIdentifier id="841C4DC23ED00E74B1DBA0FCE45A1713" type="email">mseehausen@zoolmuseum.uni-kiel.de</mods:nameIdentifier>
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<mods:typeOfResource id="7F36EB06D25421B341ED2638D5C7EE31">text</mods:typeOfResource>
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<mods:title id="7FAE872322E7FF7453950B644387D7E4">Zootaxa</mods:title>
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<mods:date id="BB8072FEDFEC39A025109EA3B0532638">2025</mods:date>
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<mods:number id="936020C37A43F914C17DE23059E69E69">5587</mods:number>
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<subSubSection id="CEE433FAF77CFFDDFF66DD6CF9949AE7" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="86416071F77CFFDDFF66DD6CFB889A44" blockId="44.[151,757,1878,1905]" box="[151,757,1878,1905]" pageId="44" pageNumber="45">
<heading id="DD09D71DF77CFFDDFF66DD6CFB889A44" bold="true" box="[151,757,1878,1905]" fontSize="11" level="2" pageId="44" pageNumber="45" reason="4">
<emphasis id="B48ABC63F77CFFDDFF66DD6CFB889A44" bold="true" box="[151,757,1878,1905]" pageId="44" pageNumber="45">
<taxonomicName id="41FE1BF2F77CFFDDFF66DD6CFB029A45" authority="Lieftinck, 1948" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[151,639,1878,1905]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="simalurum" status="stat. nov.">
<emphasis id="B48ABC63F77CFFDDFF66DD6CF8C19A44" bold="true" box="[151,444,1878,1905]" italics="true" pageId="44" pageNumber="45">Pseudagrion simalurum</emphasis>
<bibRefCitation id="E26F1D80F77CFFDDFE32DD6CFB029A45" author="Lieftinck, M. A." box="[451,639,1878,1904]" pageId="44" pageNumber="45" pagination="279 - 304" refId="ref40875" refString="Lieftinck, M. A. (1948) The Odonta of Engano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279-304." type="journal article" year="1948">Lieftinck, 1948</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="AFB90118F77CFFDDFD77DD6DFB889A44" box="[646,757,1879,1905]" pageId="44" pageNumber="45">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="86416071F77CFFDDFF66DDA1F9949AE7" blockId="44.[151,1436,1947,2033]" pageId="44" pageNumber="45">
Figures: 7677 (
<typeStatus id="5945DED3F77CFFDDFECEDDA1F8E49A86" box="[319,409,1947,1971]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F77CFFDDFE50DDA6F8CB9A86" box="[417,438,1948,1971]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
, envelope); 90 (head
<specimenCount id="90F8ABF8F77CFFDDFD65DDA1FBD49A86" box="[660,681,1947,1971]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
); 107 (pterothorax
<specimenCount id="90F8ABF8F77CFFDDFC81DDA1FAF89A86" box="[880,901,1947,1971]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
); 124a, b (abdomen
<specimenCount id="90F8ABF8F77CFFDDFBABDDA1FD129A86" box="[1114,1135,1947,1971]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
); 141ac (caudal appendages
<specimenCount id="90F8ABF8F77CFFDDFF36DD80F9A19AE7" box="[199,220,1978,2002]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77CFFDCFF66DDE2FBD49DC2" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="86416071F77CFFDDFF66DDE2FDF19AC5" blockId="44.[151,1436,1947,2033]" box="[151,1164,2008,2033]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="A6EE475FF77CFFDDFF66DDE2FDF19AC5" box="[151,1164,2008,2033]" pageId="44" pageNumber="45">
<emphasis id="B48ABC63F77CFFDDFF66DDE2F8EB9AC5" box="[151,406,2008,2032]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="41FE1BF2F77CFFDDFF66DDE2F8ED9AC5" ID-CoL="4NJDP" authorityName="Brauer" authorityYear="1868" box="[151,400,2008,2032]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pilidorsum">Pseudagrion pilidorsum</taxonomicName>
:
</emphasis>
<treatmentCitation id="075F4660F77CFFDDFE6DDDE2FB449AC4" author="Ris, F." box="[412,569,2008,2033]" page="12" pageId="44" pageNumber="45" year="1915">
<bibRefCitation id="E26F1D80F77CFFDDFE6DDDE2FB6B9AC5" author="Ris, F." box="[412,534,2008,2032]" pageId="44" pageNumber="45" pagination="5 - 21" refId="ref41650" refString="Ris, F. (1915 a) Fauna simalurensis. Odonata. Tijdschrift voor entomologie, 58, 5-21." type="journal article" year="1915">Ris (1915a)</bibRefCitation>
: 12
</treatmentCitation>
13 (description
<specimenCount id="90F8ABF8F77CFFDDFD14DDE3FB879AC5" box="[741,762,2009,2032]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
; Simalur);—
<treatmentCitation id="075F4660F77CFFDDFC8FDDE2FD609AC4" author="Ris, F." box="[894,1053,2008,2033]" page="99" pageId="44" pageNumber="45" year="1915">
<bibRefCitation id="E26F1D80F77CFFDDFC8FDDE2FA869AC5" author="Ris, F." box="[894,1019,2008,2032]" pageId="44" pageNumber="45" pagination="81 - 131" refId="ref41673" refString="Ris, F. (1915 b) Neuer Beitrag zur Kenntnis der Odonatenfauna der Neu-Guinea-Region. Nova Guinea, 13, 81-131." type="journal article" year="1915">Ris (1915b)</bibRefCitation>
: 99
</treatmentCitation>
(Simalur).
</treatmentCitationGroup>
</paragraph>
<paragraph id="86416071F77DFFDCFF66DAADFD1A9D9A" blockId="45.[151,1127,150,175]" box="[151,1127,150,175]" pageId="45" pageNumber="46">
<treatmentCitationGroup id="A6EE475FF77DFFDCFF66DAADFD1A9D9A" box="[151,1127,150,175]" pageId="45" pageNumber="46">
<taxonomicName id="41FE1BF2F77DFFDCFF66DAADF8829D9A" ID-CoL="7HWZW" authorityName="Lieftinck" authorityYear="1948" box="[151,511,151,175]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="pilidorsum" subSpecies="simalurum">
<emphasis id="B48ABC63F77DFFDCFF66DAADF8829D9A" box="[151,511,151,175]" italics="true" pageId="45" pageNumber="46">Pseudagrion pilidorsum simalurum</emphasis>
</taxonomicName>
:
<treatmentCitation id="075F4660F77DFFDCFDFCDAADFB959D9A" author="Lieftinck, M. A." box="[525,744,151,175]" page="289" pageId="45" pageNumber="46" year="1948">
<bibRefCitation id="E26F1D80F77DFFDCFDFCDAADFBC59D9A" author="Lieftinck, M. A." box="[525,696,151,175]" pageId="45" pageNumber="46" pagination="279 - 304" refId="ref40875" refString="Lieftinck, M. A. (1948) The Odonta of Engano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279-304." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
: 289
</treatmentCitation>
290,
<figureCitation id="1EC57CF4F77DFFDCFCD7DAADFA1F9D9B" box="[806,866,150,175]" captionStart="FIGURES 510" captionStartId="8.[151,264,1876,1900]" captionTargetBox="[153,1433,811,1847]" captionTargetId="figure-260@8.[151,1436,810,1850]" captionTargetPageId="8" captionText="FIGURES 510. Pseudagrion acutidens, holotype male: (5) habitus lateral, (6) appendages lateral, (7) appendages dorsal, (8) cercus dorsolateral, (9) head dorsal, (10) pronotum. Photos: M. Seehausen." pageId="45" pageNumber="46">Fig. 5</figureCitation>
(description
<specimenCount id="90F8ABF8F77DFFDCFC16DAADFA819D9B" box="[999,1020,151,174]" count="1" pageId="45" pageNumber="46" type="male"></specimenCount>
; Simalur).
</treatmentCitationGroup>
</paragraph>
<paragraph id="86416071F77DFFDCFF66DAE7FBD49DC2" blockId="45.[151,1437,221,391]" box="[151,681,221,247]" pageId="45" pageNumber="46">
<emphasis id="B48ABC63F77DFFDCFF66DAE7FB119DC2" bold="true" box="[151,620,221,247]" pageId="45" pageNumber="46">Name-bearing type specimen examined</emphasis>
(
<specimenCount id="90F8ABF8F77DFFDCFD8CDAE7FBDD9DC2" box="[637,672,221,247]" count="1" pageId="45" pageNumber="46" type="male">1♂</specimenCount>
)
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77DFFDCFF36DB3BFC0A9CB2" pageId="45" pageNumber="46" type="description">
<paragraph id="86416071F77DFFDCFF36DB3BF9B99C0A" blockId="45.[151,1437,221,391]" pageId="45" pageNumber="46">
<materialsCitation id="36966A2CF77DFFDCFF36DB3BF9BD9C0A" collectingDate="1913-01" collectionCode="RMNH" collectorName="E. Jacobson" country="Indonesia" location="Simalur Island" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="90F8ABF8F77DFFDCFF36DB3BF9A19C2E" box="[199,220,257,283]" count="1" pageId="45" pageNumber="46" type="male"></specimenCount>
<typeStatus id="5945DED3F77DFFDCFF17DB3BF8369C2E" box="[230,331,257,283]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
,
<date id="F24046B1F77DFFDCFEA9DB3BF8DC9C2E" box="[344,417,257,283]" pageId="45" pageNumber="46" value="1913-01">
<collectingDate id="E204BF59F77DFFDCFEA9DB3BF8DC9C2E" box="[344,417,257,283]" pageId="45" pageNumber="46" value="1913-01">i.1913</collectingDate>
</date>
, Sinabang,
<location id="832136AAF77DFFDCFDD8DB3BFBAF9C2E" LSID="urn:lsid:plazi:treatment:0E57D167F77CFFDFFF66DD6CFA0A9838:832136AAF77DFFDCFDD8DB3BFBAF9C2E" box="[553,722,257,283]" country="Indonesia" name="Simalur Island" pageId="45" pageNumber="46">Simalur Island</location>
,
<collectingCountry id="FEE920E1F77DFFDCFD11DB3BFA2D9C2E" box="[736,848,257,283]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
,
<collectorName id="2B0B05A7F77DFFDCFCACDB3BFA9B9C2E" box="[861,998,257,283]" pageId="45" pageNumber="46">E. Jacobson</collectorName>
leg.,
<collectionCode id="E0EFF8B4F77DFFDCFBD8DB3BFDFD9C2E" box="[1065,1152,257,283]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="45" pageNumber="46" type="Museum">RMNH</collectionCode>
, JvT no. 2854 (
<figureCitation id="1EC57CF4F77DFFDCFAC6DB3BF9CA9C0A" captionStart="FIGURES 7677" captionStartId="45.[151,264,1624,1648]" captionTargetBox="[154,1435,413,1595]" captionTargetId="figure-89@45.[151,1436,413,1599]" captionTargetPageId="45" captionText="FIGURES 7677. Pseudagrion simalurum, holotype male: (76) habitus lateral, (77) envelope. Photos: RMNH." pageId="45" pageNumber="46">Figs 76 77</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F77DFFDCFF36DB73FC0A9CB2" blockId="45.[151,1437,221,391]" pageId="45" pageNumber="46">
<bibRefCitation id="E26F1D80F77DFFDCFF36DB73F8C49C56" author="Lieftinck, M. A." box="[199,441,329,355]" pageId="45" pageNumber="46" pagination="279 - 304" refId="ref40875" refString="Lieftinck, M. A. (1948) The Odonta of Engano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279-304." type="journal article" year="1948">Lieftinck (1948: 290)</bibRefCitation>
stated “Described from the
<typeStatus id="5945DED3F77DFFDCFD04DB73FA259C56" box="[757,856,329,355]" pageId="45" pageNumber="46" type="paratype">paratype</typeStatus>
, in the Leiden Museum”, and
<bibRefCitation id="E26F1D80F77DFFDCFB5DDB73FCE19C56" author="Lieftinck, M. A." box="[1196,1436,329,355]" pageId="45" pageNumber="46" pagination="65 - 139" refId="ref40982" refString="Lieftinck, M. A. (1971) A catalogue of the type-specimens of Odonata preserved in the Netherlands. With a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65-139, pls. 1 - 7." type="journal article" year="1971">Lieftinck (1971: 112)</bibRefCitation>
designated this specimen to be the
<typeStatus id="5945DED3F77DFFDCFDEFDB57FBF59CB2" box="[542,648,365,391]" pageId="45" pageNumber="46" type="lectotype">lectotype</typeStatus>
. But the description is based on a single male, thus it is a
<typeStatus id="5945DED3F77DFFDCFAFCDB57FC0E9CB2" box="[1293,1395,365,391]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="D28130F9F77DFFDCFF66DC62FC7E9B45" box="[151,1283,1624,1649]" pageId="45" pageNumber="46" startId="45.[151,264,1624,1648]" targetBox="[154,1435,413,1595]" targetPageId="45" targetType="figure">
<paragraph id="86416071F77DFFDCFF66DC62FC7E9B45" blockId="45.[151,1283,1624,1649]" box="[151,1283,1624,1649]" pageId="45" pageNumber="46">
<emphasis id="B48ABC63F77DFFDCFF66DC62F8309B45" bold="true" box="[151,333,1624,1648]" pageId="45" pageNumber="46">FIGURES 7677</emphasis>
.
<taxonomicName id="41FE1BF2F77DFFDCFEABDC62FB339B45" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[346,590,1624,1648]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F77DFFDCFEABDC62FB339B45" box="[346,590,1624,1648]" italics="true" pageId="45" pageNumber="46">Pseudagrion simalurum</emphasis>
</taxonomicName>
, holotype male: (
<emphasis id="B48ABC63F77DFFDCFCF0DC62FA679B45" bold="true" box="[769,794,1624,1648]" pageId="45" pageNumber="46">76</emphasis>
) habitus lateral, (
<emphasis id="B48ABC63F77DFFDCFC3CDC63FA9B9B44" bold="true" box="[973,998,1625,1649]" pageId="45" pageNumber="46">77</emphasis>
) envelope. Photos: RMNH.
</paragraph>
</caption>
<subSubSection id="CEE433FAF77DFFDFFF36DCF8F8EC9ED8" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="86416071F77DFFDCFF36DCF8FB3B9BE8" blockId="45.[151,1436,1730,2009]" box="[199,582,1730,1757]" pageId="45" pageNumber="46">
<emphasis id="B48ABC63F77DFFDCFF36DCF8FB749BE9" bold="true" box="[199,521,1730,1756]" pageId="45" pageNumber="46">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F77DFFDCFDE8DCF9FB419BE8" box="[537,572,1731,1757]" count="1" pageId="45" pageNumber="46" type="male">1♂</specimenCount>
)
</paragraph>
<paragraph id="86416071F77DFFDCFF36DCDCFDFA9A37" blockId="45.[151,1436,1730,2009]" box="[199,1159,1766,1794]" pageId="45" pageNumber="46">
<materialsCitation id="36966A2CF77DFFDCFF36DCDCFDFE9A37" box="[199,1155,1766,1794]" collectingDate="1913-01" collectionCode="SMF" collectorName="Sinabang, E &amp; E. Jacobson" country="Indonesia" location="Simalur Island" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Simalur Island">
<emphasis id="B48ABC63F77DFFDCFF36DCDCFB789A34" bold="true" box="[199,517,1766,1793]" pageId="45" pageNumber="46">
<collectingCountry id="FEE920E1F77DFFDCFF36DCDCF8409A35" box="[199,317,1766,1792]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F77DFFDCFEBBDCDCFB7D9A34" box="[330,512,1766,1793]" pageId="45" pageNumber="46">Simalur Island</collectingRegion>
:
</emphasis>
<specimenCount id="90F8ABF8F77DFFDCFDFDDCDDFB4D9A34" box="[524,560,1767,1793]" count="1" pageId="45" pageNumber="46" type="male">1♂</specimenCount>
,
<date id="F24046B1F77DFFDCFDCCDCDDFBFB9A34" box="[573,646,1767,1793]" pageId="45" pageNumber="46" value="1913-01">
<collectingDate id="E204BF59F77DFFDCFDCCDCDDFBFB9A34" box="[573,646,1767,1793]" pageId="45" pageNumber="46" value="1913-01">i.1913</collectingDate>
</date>
,
<collectorName id="2B0B05A7F77DFFDCFD63DCDDFA619A34" box="[658,796,1767,1793]" pageId="45" pageNumber="46">Sinabang, E</collectorName>
. Jacobson leg.,
<collectionCode id="E0EFF8B4F77DFFDCFC3DDCDDFD789A34" box="[972,1029,1767,1793]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="45" pageNumber="46">SMF</collectionCode>
, no. 13951
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F77DFFDCFF36DD30F88F9A10" blockId="45.[151,1436,1730,2009]" box="[199,498,1802,1829]" pageId="45" pageNumber="46">
<emphasis id="B48ABC63F77DFFDCFF36DD30F88F9A10" bold="true" box="[199,498,1802,1829]" pageId="45" pageNumber="46">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F77DFFDCFF36DD15F9A09AEC" blockId="45.[151,1436,1730,2009]" pageId="45" pageNumber="46">
<emphasis id="B48ABC63F77DFFDCFF36DD15F8799A7C" box="[199,260,1839,1865]" italics="true" pageId="45" pageNumber="46">Head</emphasis>
(
<figureCitation id="1EC57CF4F77DFFDCFEE4DD15F8139A7C" box="[277,366,1839,1865]" captionStart="FIGURES 7677" captionStartId="45.[151,264,1624,1648]" captionTargetBox="[154,1435,413,1595]" captionTargetId="figure-89@45.[151,1436,413,1599]" captionTargetPageId="45" captionText="FIGURES 7677. Pseudagrion simalurum, holotype male: (76) habitus lateral, (77) envelope. Photos: RMNH." pageId="45" pageNumber="46">Figs 76</figureCitation>
,
<figureCitation id="1EC57CF4F77DFFDCFE8ADD15F8E19A7C" box="[379,412,1839,1865]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="45" pageNumber="46">90</figureCitation>
): Labrum, clypeus, antefrons and postfrons yellowish; colour of postfrons separated from vertex around the level of median ocellus; vertex reddish, with a small black patch posterior of median ocellus; large black patches posterolateral of each lateral ocellus; postocular spots reddish, bordered by black; rear of head black, but narrow yellowish along the eye margins; base and scape of antennae yellow, following segments brownish ochre.
</paragraph>
<paragraph id="86416071F77EFFDFFF36DAADF8BF9CC0" blockId="46.[151,1437,151,1293]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF36DAADF8659D84" box="[199,280,151,177]" italics="true" pageId="46" pageNumber="47">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F77EFFDFFEDADAADF8FA9D84" box="[299,391,151,177]" captionStart="FIGURES 7677" captionStartId="45.[151,264,1624,1648]" captionTargetBox="[154,1435,413,1595]" captionTargetId="figure-89@45.[151,1436,413,1599]" captionTargetPageId="45" captionText="FIGURES 7677. Pseudagrion simalurum, holotype male: (76) habitus lateral, (77) envelope. Photos: RMNH." pageId="46" pageNumber="47">Figs 76</figureCitation>
,
<figureCitation id="1EC57CF4F77EFFDFFE67DAADF8B99D84" box="[406,452,151,177]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="46" pageNumber="47">107</figureCitation>
): Anterior lobe of pronotum yellow-orange, this colour expanding in centre towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large yellow-orange patch in in the middle, and on either side a yellow-orange patch dorsolaterally; propleuron black above, yellow-orange below and to rear; posterior lobe of pronotum yellow-orange, with its margins black; apical edge of posterior lobe convex. Pterothorax orange-reddish; middorsal carina thin black; humeral suture with a thin black stripe which is expanded to an elongate black patch at posterior quarter; interpleural suture with a black stripe at posterior quarter, and a black spot anteriorly; metapleural suture with an elongate black patch at posterior quarter; a large black marking covering at least superior one-third to half of the mesinfraepisternum; venter pale, may be covered with whitish pruinosity. Legs pale orange; femora with black stripe outside, narrowing to proximal; tibiae with black along spines; spines black.
</paragraph>
<paragraph id="86416071F77EFFDFFF36DBC5F9BE9F54" blockId="46.[151,1437,151,1293]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF36DBC5F84C9F2C" box="[199,305,511,537]" italics="true" pageId="46" pageNumber="47">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F77EFFDFFECEDBC5F8EB9F2C" box="[319,406,511,537]" captionStart="FIGURES 7677" captionStartId="45.[151,264,1624,1648]" captionTargetBox="[154,1435,413,1595]" captionTargetId="figure-89@45.[151,1436,413,1599]" captionTargetPageId="45" captionText="FIGURES 7677. Pseudagrion simalurum, holotype male: (76) habitus lateral, (77) envelope. Photos: RMNH." pageId="46" pageNumber="47">Figs 76</figureCitation>
,
<figureCitation id="1EC57CF4F77EFFDFFE51DBC5F8889F2C" box="[416,501,511,537]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="46" pageNumber="47">124a, b</figureCitation>
): S12 dorsally black, laterally orange; S38 dorsally black, laterally and each basal annulus pale yellowish; S9 black, with anterior third red; S10 dorsally with a broad black “x”-like marking, laterally red.
</paragraph>
<paragraph id="86416071F77EFFDFFF36D851FB8F9E68" blockId="46.[151,1437,151,1293]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF36D851F8D49FB0" box="[199,425,619,645]" italics="true" pageId="46" pageNumber="47">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F77EFFDFFE48D851FB379FB0" box="[441,586,619,645]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="46" pageNumber="47">Figs 141ac</figureCitation>
): In lateral view cercus blackish, shorter than S10; subrectangular with an open inter-lobe notch around lower third; in dorsal view apex of the superior lobe of the cercus somewhat pointed, curved inwards; an inwards directed projection around the centre of the superior lobe of the cercus; a small bulge-like interior projection is situated at the base of a ridge on the inferior lobe of the cercus, visible in dorsal and dorsolateral views, proximal edge of this projection sharply sloping; paraproct brownish-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; bearing a small black tooth at medial edge.
</paragraph>
<paragraph id="86416071F77EFFDFFF36D95DFAAC9EB4" blockId="46.[151,1437,151,1293]" box="[199,977,870,897]" pageId="46" pageNumber="47">Wings with 14.515.0 px in forewing, and 12.512.5 px in hindwing.</paragraph>
<paragraph id="86416071F77EFFDFFF36D9B1FDDE9EFC" blockId="46.[151,1437,151,1293]" pageId="46" pageNumber="47">Measurements (mm): Total length (with appendages) 45.045.4; abdomen length (without appendages) 37.3 37.5; hindwing 24.325.0; forewing 25.526.5; Pt in forewing 0.81.0; male cercus 0.50.6.</paragraph>
<paragraph id="86416071F77EFFDFFF36D9E9F8EC9ED8" blockId="46.[151,1437,151,1293]" box="[199,401,979,1005]" pageId="46" pageNumber="47">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77EFFDFFF36D9CCFD1799F0" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="86416071F77EFFDFFF36D9CCF8479925" blockId="46.[151,1437,151,1293]" box="[199,314,1014,1040]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF36D9CCF8479925" bold="true" box="[199,314,1014,1040]" pageId="46" pageNumber="47">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F77EFFDFFF36DE21FD1799F0" blockId="46.[151,1437,151,1293]" pageId="46" pageNumber="47">
<taxonomicName id="41FE1BF2F77EFFDFFF36DE21F8AB9900" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[199,470,1051,1077]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F77EFFDFFF36DE21F8AB9900" box="[199,470,1051,1077]" italics="true" pageId="46" pageNumber="47">Pseudagrion simalurum</emphasis>
</taxonomicName>
belongs to the group of endemic taxa of the island chain west of
<collectingRegion id="443AAE93F77EFFDFFB4DDE21FC609900" box="[1212,1309,1051,1077]" country="Indonesia" name="Sumatera Utara" pageId="46" pageNumber="47">Sumatra</collectingRegion>
. The males are separated from most congeners occurring at adjacent islands by having S12 dorsally black (
<figureCitation id="1EC57CF4F77EFFDFFB07DE05FC2C996C" box="[1270,1361,1087,1113]" captionStart="FIGURES 7677" captionStartId="45.[151,264,1624,1648]" captionTargetBox="[154,1435,413,1595]" captionTargetId="figure-89@45.[151,1436,413,1599]" captionTargetPageId="45" captionText="FIGURES 7677. Pseudagrion simalurum, holotype male: (76) habitus lateral, (77) envelope. Photos: RMNH." pageId="46" pageNumber="47">Figs 76</figureCitation>
,
<figureCitation id="1EC57CF4F77EFFDFFAAEDE05F9D59948" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="46" pageNumber="47">124a, b</figureCitation>
). This character is elsewhere shared by
<taxonomicName id="41FE1BF2F77EFFDFFD9DDE59FBA69949" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" box="[620,731,1123,1148]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="igneum">
<emphasis id="B48ABC63F77EFFDFFD9DDE59FBA69949" box="[620,731,1123,1148]" italics="true" pageId="46" pageNumber="47">P. igneum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F77EFFDFFD1ADE59FA049948" box="[747,889,1123,1149]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="46" pageNumber="47">Figs 121a, b</figureCitation>
) and
<taxonomicName id="41FE1BF2F77EFFDFFC4BDE59FD399948" authorityName="Lieftinck" authorityYear="1936" baseAuthorityName="Lieftinck" baseAuthorityYear="1936" box="[954,1092,1123,1149]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="obscurum">
<emphasis id="B48ABC63F77EFFDFFC4BDE59FD399948" box="[954,1092,1123,1149]" italics="true" pageId="46" pageNumber="47">P. obscurum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F77EFFDFFBA5DE59FD9C9948" box="[1108,1249,1123,1149]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="46" pageNumber="47">Figs 123a, b</figureCitation>
)—both are very similar to
<taxonomicName id="41FE1BF2F77EFFDFFEF8DEBDF8E09994" authorityName="Lieftinck" authorityYear="1948" baseAuthorityName="Lieftinck" baseAuthorityYear="1948" box="[265,413,1159,1185]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="simalurum">
<emphasis id="B48ABC63F77EFFDFFEF8DEBDF8E09994" box="[265,413,1159,1185]" italics="true" pageId="46" pageNumber="47">P. simalurum</emphasis>
</taxonomicName>
also in the colour of the head (
<figureCitation id="1EC57CF4F77EFFDFFD0ADEBDFAAD9994" box="[763,976,1159,1185]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="46" pageNumber="47">Figs 87, 89a, b, 90</figureCitation>
), and the morphology of the male cercus (
<figureCitation id="1EC57CF4F77EFFDFFF6EDE91F85699F0" box="[159,299,1195,1221]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="46" pageNumber="47">Figs 138ac</figureCitation>
,
<figureCitation id="1EC57CF4F77EFFDFFEC6DE91F89699F0" box="[311,491,1195,1221]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="46" pageNumber="47">140ac, 141ac</figureCitation>
). We summarised the characters for separation in table 2.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77EFFDFFF36DEF4FA0A9838" box="[199,887,1230,1293]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="86416071F77EFFDFFF36DEF4F82499DD" blockId="46.[151,1437,151,1293]" box="[199,345,1230,1256]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF36DEF4F82499DD" bold="true" box="[199,345,1230,1256]" pageId="46" pageNumber="47">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F77EFFDFFF36DEC9FA0A9838" blockId="46.[151,1437,151,1293]" box="[199,887,1267,1293]" pageId="46" pageNumber="47">
This species is apparently endemic to Simalur Island (
<figureCitation id="1EC57CF4F77EFFDFFCD4DEC9FA179838" box="[805,874,1267,1293]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="46" pageNumber="47">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,409 @@
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<mods:title id="13345B4102ED01BE8B7763C3B25DAB95">Revision of the Pseudagrion red-group occurring in Asia west of Lydekkers line, with description of four new species (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="635CB1067BF7EB2E5B2ABC3F6AFCCF7E">Seehausen, Malte</mods:namePart>
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</mods:mods>
<treatment id="0E57D167F77EFFC1FF66DF64FDAA9BD4" LSID="urn:lsid:plazi:treatment:0E57D167F77EFFC1FF66DF64FDAA9BD4" httpUri="http://treatment.plazi.org/id/0E57D167F77EFFC1FF66DF64FDAA9BD4" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="CEE433FAF77EFFDFFF66DF64F85498EA" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="86416071F77EFFDFFF66DF64FB6E984D" blockId="46.[151,531,1374,1401]" box="[151,531,1374,1401]" pageId="46" pageNumber="47">
<heading id="DD09D71DF77EFFDFFF66DF64FB6E984D" bold="true" box="[151,531,1374,1401]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
<taxonomicName id="41FE1BF2F77EFFDFFF66DF64FB6E984D" ID-CoL="4NJF6" authority="Selys, 1876" authorityName="Selys" authorityYear="1876" box="[151,531,1374,1401]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F77EFFDFFF66DF64FB6E984D" bold="true" box="[151,531,1374,1401]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF66DF64F8FE984C" bold="true" box="[151,387,1374,1401]" italics="true" pageId="46" pageNumber="47">Pseudagrion ustum</emphasis>
<bibRefCitation id="E26F1D80F77EFFDFFE7BDF64FB6E984D" author="Selys Longchamps, E. de" box="[394,531,1374,1400]" pageId="46" pageNumber="47" pagination="247 - 322" refId="ref42034" refString="Selys Longchamps, E. de (1876) Synopsis des Agrionines (suite de la 5 me Legion: Agrion). Bulletins de l'academy royale des sciences, des lettres et des beaux-arts de Belgique, 45, Series 2, 42, 247-322 + 496 - 539." type="journal article" year="1876">Selys, 1876</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="86416071F77EFFDFFF66DF99F85498EA" blockId="46.[151,1436,1443,1565]" pageId="46" pageNumber="47">
Figures: 78 (
<typeStatus id="5945DED3F77EFFDFFEE8DF99F80E988E" box="[281,371,1443,1467]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
<specimenCount id="90F8ABF8F77EFFDFFE8BDF99F8F1988E" box="[378,396,1443,1467]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
); 95 (head
<specimenCount id="90F8ABF8F77EFFDFFDF2DF99FB65988E" box="[515,536,1443,1467]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
); 112 (pterothorax
<specimenCount id="90F8ABF8F77EFFDFFD2CDF99FB8F988E" box="[733,754,1443,1467]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
); 129a, b (abdomen
<specimenCount id="90F8ABF8F77EFFDFFC37DF99FAA6988E" box="[966,987,1443,1467]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
); 146ac (caudal appendages
<specimenCount id="90F8ABF8F77EFFDFFAFEDF99FC59988E" box="[1295,1316,1443,1467]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
); 168 (field photo
<specimenCount id="90F8ABF8F77EFFDFFEF6DFFDF86198EA" box="[263,284,1479,1503]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77EFFDFFF66DFDCFBDE9B50" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="86416071F77EFFDFFF66DFDCFDB09B28" blockId="46.[151,1436,1443,1565]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="A6EE475FF77EFFDFFF66DFDCFDB09B28" pageId="46" pageNumber="47">
<taxonomicName id="41FE1BF2F77EFFDFFF66DFDCF82698CB" ID-CoL="4NJF6" authorityName="Selys" authorityYear="1876" box="[151,347,1510,1534]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F77EFFDFFF66DFDCF82698CB" box="[151,347,1510,1534]" italics="true" pageId="46" pageNumber="47">Pseudagrion ustum</emphasis>
</taxonomicName>
:
<treatmentCitation id="075F4660F77EFFDFFE96DFDCFB6798CB" author="Selys Longchamps, E. de" box="[359,538,1510,1534]" page="515" pageId="46" pageNumber="47" year="1876">
<bibRefCitation id="E26F1D80F77EFFDFFE96DFDCF89698CB" author="Selys Longchamps, E. de" box="[359,491,1510,1534]" pageId="46" pageNumber="47" pagination="247 - 322" refId="ref42034" refString="Selys Longchamps, E. de (1876) Synopsis des Agrionines (suite de la 5 me Legion: Agrion). Bulletins de l'academy royale des sciences, des lettres et des beaux-arts de Belgique, 45, Series 2, 42, 247-322 + 496 - 539." type="journal article" year="1876">Selys (1876)</bibRefCitation>
: 515
</treatmentCitation>
(description
<specimenCount id="90F8ABF8F77EFFDFFD6FDFDCFBCF98C8" box="[670,690,1510,1533]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
; Sulu Island, Malay Archipelago [actual Sula Archipelago,
<collectingCountry id="FEE920E1F77EFFDFFAF9DFDCFC0D98CB" box="[1288,1392,1510,1534]" name="Indonesia" pageId="46" pageNumber="47">Indonesia</collectingCountry>
]);—
<treatmentCitation id="075F4660F77EFFDFFF36DC3EF8149B28" author="Ris, F." box="[199,361,1540,1565]" page="97" pageId="46" pageNumber="47" year="1915">
<bibRefCitation id="E26F1D80F77EFFDFFF36DC3EF8399B29" author="Ris, F." box="[199,324,1540,1564]" pageId="46" pageNumber="47" pagination="81 - 131" refId="ref41673" refString="Ris, F. (1915 b) Neuer Beitrag zur Kenntnis der Odonatenfauna der Neu-Guinea-Region. Nova Guinea, 13, 81-131." type="journal article" year="1915">Ris (1915b)</bibRefCitation>
: 97
</treatmentCitation>
, 99100,
<figureCitation id="1EC57CF4F77EFFDFFE3DDC3EFB6E9B28" box="[460,531,1540,1565]" captionStart="FIGURES 2022" captionStartId="15.[151,264,964,988]" captionTargetBox="[158,1434,181,935]" captionTargetId="figure-18@15.[151,1436,179,938]" captionTargetPageId="15" captionText="FIGURES 2022. Pseudagrion crocops, holotype male: (20) habitus dorsal, (21) labels, (22) habitus lateral. Photos: © The Trustees of the Natural History Museum, London. http://creativecommons.org/licenses/by/4.0." pageId="46" pageNumber="47">Fig. 21</figureCitation>
(key; description
<specimenCount id="90F8ABF8F77EFFDFFD3ADC3FFB9D9B29" box="[715,736,1541,1564]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
;
<collectingRegion id="443AAE93F77EFFDFFD1ADC3EFA469B29" box="[747,827,1540,1564]" country="Indonesia" name="Sulawesi Utara" pageId="46" pageNumber="47">Celebes</collectingRegion>
);—
<treatmentCitation id="075F4660F77EFFDFFC91DC3EFD379B28" author="Lieftinck, M. A." box="[864,1098,1540,1565]" page="307" pageId="46" pageNumber="47" year="1930">
<bibRefCitation id="E26F1D80F77EFFDFFC91DC3EFD669B28" author="Lieftinck, M. A." box="[864,1051,1540,1565]" pageId="46" pageNumber="47" pagination="305 - 330" refId="ref40581" refString="Lieftinck, M. A. (1930 a) Fauna Buruana. Odonata 3. A review of the dragonfly-fauna of Boeroe, in the Moluccas, with descriptions of new or interesting species, and an account of their larvae. Treubia Supplement, 7, 305-330." type="journal article" year="1930">Lieftinck (1930a)</bibRefCitation>
: 307
</treatmentCitation>
(
<specimenCount id="90F8ABF8F77EFFDFFBA9DC3FFD169B28" box="[1112,1131,1541,1565]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
<specimenCount id="90F8ABF8F77EFFDFFB83DC3FFDF89B29" box="[1138,1157,1541,1564]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
Buru).
</treatmentCitationGroup>
</paragraph>
<paragraph id="86416071F77EFFDFFF66DC70FBDE9B50" blockId="46.[151,1437,1610,1961]" box="[151,675,1610,1637]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF66DC70F88F9B51" bold="true" box="[151,498,1610,1637]" pageId="46" pageNumber="47">Name-bearing type specimen</emphasis>
(not examined)
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77EFFDFFF36DC55FD0F9B98" pageId="46" pageNumber="47" type="description">
<paragraph id="86416071F77EFFDFFF36DC55FC559BBC" blockId="46.[151,1437,1610,1961]" box="[199,1320,1647,1673]" pageId="46" pageNumber="47">
<materialsCitation id="36966A2CF77EFFDFFF36DC55FC599BBC" box="[199,1316,1647,1673]" collectionCode="IRSNB" country="Indonesia" location="Sulu Island" municipality="Malay Archipelago" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="90F8ABF8F77EFFDFFF36DC55F9A19BBC" box="[199,220,1647,1673]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
<typeStatus id="5945DED3F77EFFDFFF12DC55F8359BBC" box="[227,328,1647,1673]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
,
<location id="832136AAF77EFFDFFEA2DC55F8A89BBC" LSID="urn:lsid:plazi:treatment:0E57D167F77EFFC1FF66DF64FDAA9BD4:832136AAF77EFFDFFEA2DC55F8A89BBC" box="[339,469,1647,1673]" country="Indonesia" municipality="Malay Archipelago" name="Sulu Island" pageId="46" pageNumber="47">Sulu Island</location>
,
<collectingMunicipality id="6625FA0BF77EFFDFFE11DC55FBCA9BBC" box="[480,695,1647,1673]" pageId="46" pageNumber="47">Malay Archipelago</collectingMunicipality>
[actual Sula Archipelago,
<collectingCountry id="FEE920E1F77EFFDFFC10DC55FD299BBC" box="[993,1108,1647,1673]" name="Indonesia" pageId="46" pageNumber="47">Indonesia</collectingCountry>
],
<collectionCode id="E0EFF8B4F77EFFDFFB94DC55FDCA9BBC" box="[1125,1207,1647,1673]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="46" pageNumber="47">IRSNB</collectionCode>
(
<figureCitation id="1EC57CF4F77EFFDFFB36DC55FC669BBC" box="[1223,1307,1647,1673]" captionStart="FIGURE 78" captionStartId="47.[152,255,1845,1869]" captionTargetBox="[155,1434,182,1816]" captionTargetId="figure-18@47.[151,1436,179,1820]" captionTargetPageId="47" captionText="FIGURE 78. Pseudagrion ustum, holotype female, water colour by Selys, 20.viii.1875 (IRSNB Ag125a). Photo: IRSNB." pageId="46" pageNumber="47">Fig. 78</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="86416071F77EFFDFFF36DCA9FD0F9B98" blockId="46.[151,1437,1610,1961]" box="[199,1138,1683,1709]" pageId="46" pageNumber="47">
<materialsCitation id="36966A2CF77EFFDFFF36DCA9FD109B98" box="[199,1133,1683,1709]" collectionCode="IRSNB" collectorName="J. Constant" pageId="46" pageNumber="47" specimenCount="1" typeStatus="holotype">
Currently the
<typeStatus id="5945DED3F77EFFDFFE92DCA9F8B89B98" box="[355,453,1683,1709]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
could not be found at the
<collectionCode id="E0EFF8B4F77EFFDFFD1BDCA9FA419B98" box="[746,828,1683,1709]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="46" pageNumber="47">IRSNB</collectionCode>
(
<collectorName id="2B0B05A7F77EFFDFFCBCDCA9FAB19B98" box="[845,972,1683,1709]" pageId="46" pageNumber="47">J. Constant</collectorName>
, pers. comm.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF77EFFC1FF3FDC8CFC3998F4" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="86416071F77EFFDFFF3FDC8CFBCB9BE4" blockId="46.[151,1437,1610,1961]" box="[206,694,1718,1745]" pageId="46" pageNumber="47">
<emphasis id="B48ABC63F77EFFDFFF3FDC8CFB6D9BE5" bold="true" box="[206,528,1718,1744]" pageId="46" pageNumber="47">Other specimens examined</emphasis>
(
<specimenCount id="90F8ABF8F77EFFDFFDEEDC8CFB149BE4" box="[543,617,1718,1745]" count="15" pageId="46" pageNumber="47" type="male">15♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F77EFFDFFD85DC8DFBD79BE4" box="[628,682,1719,1745]" count="8" pageId="46" pageNumber="47" type="female">8♀♀</specimenCount>
)
</paragraph>
<paragraph id="86416071F77EFFDFFF36DCE0FAF09A9C" blockId="46.[151,1437,1610,1961]" pageId="46" pageNumber="47">
<materialsCitation id="36966A2CF77EFFDFFF36DCE0FB989BC0" box="[199,741,1754,1781]" collectionCode="SMF" collectorName="Toli-Toli" country="Indonesia" location="Sulawesi" pageId="46" pageNumber="47" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Sulawesi Utara">
<emphasis id="B48ABC63F77EFFDFFF36DCE0F8CE9BC1" bold="true" box="[199,435,1754,1780]" pageId="46" pageNumber="47">
<collectingCountry id="FEE920E1F77EFFDFFF36DCE0F8409BC1" box="[199,317,1754,1780]" name="Indonesia" pageId="46" pageNumber="47">Indonesia</collectingCountry>
,
<collectingRegion id="443AAE93F77EFFDFFEBADCE0F8CE9BC1" box="[331,435,1754,1780]" country="Indonesia" name="Sulawesi Utara" pageId="46" pageNumber="47">Sulawesi</collectingRegion>
</emphasis>
:
<specimenCount id="90F8ABF8F77EFFDFFE32DCE1F8809BC0" box="[451,509,1755,1781]" count="3" pageId="46" pageNumber="47" type="male">3♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F77EFFDFFDF8DCE1FB519BC0" box="[521,556,1755,1781]" count="1" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
,
<collectorName id="2B0B05A7F77EFFDFFDCADCE1FBDC9BC0" box="[571,673,1755,1781]" pageId="46" pageNumber="47">Toli-Toli</collectorName>
,
<collectionCode id="E0EFF8B4F77EFFDFFD5DDCE1FB989BC0" box="[684,741,1755,1781]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="46" pageNumber="47">SMF</collectionCode>
</materialsCitation>
, no. 1375413757;—
<materialsCitation id="36966A2CF77EFFDFFC12DCE0F8BB9A2C" collectingDate="1896-03" collectionCode="ZMH" collectorName="H. Fruhstorfer" country="Indonesia" location="Lompa-Battau" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
<specimenCount id="90F8ABF8F77EFFDFFC12DCE0FA829BC0" box="[995,1023,1754,1781]" count="1" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
<date id="F24046B1F77EFFDFFBFADCE1FDEA9BC0" box="[1035,1175,1755,1781]" pageId="46" pageNumber="47" value="1896-03">
<collectingDate id="E204BF59F77EFFDFFBFADCE1FDEA9BC0" box="[1035,1175,1755,1781]" pageId="46" pageNumber="47" value="1896-03">March 1896</collectingDate>
</date>
,
<location id="832136AAF77EFFDFFB52DCE1FC359BC0" LSID="urn:lsid:plazi:treatment:0E57D167F77EFFC1FF66DF64FDAA9BD4:832136AAF77EFFDFFB52DCE1FC359BC0" box="[1187,1352,1755,1781]" country="Indonesia" name="Lompa-Battau" pageId="46" pageNumber="47" stateProvince="Sulawesi Utara">Lompa-Battau</location>
, 3000,
<collectorName id="2B0B05A7F77EFFDFFF66DCC5F8469A2C" box="[151,315,1791,1817]" pageId="46" pageNumber="47">H. Fruhstorfer</collectorName>
leg.,
<collectionCode id="E0EFF8B4F77EFFDFFE73DCC4F8BB9A2C" box="[386,454,1790,1817]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="46" pageNumber="47" type="Museum">ZMH</collectionCode>
</materialsCitation>
;—
<materialsCitation id="36966A2CF77EFFDFFE18DCC4FA0E9A2C" box="[489,883,1790,1817]" collectionCode="SMF" collectorName="Bonthain" country="Indonesia" pageId="46" pageNumber="47" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Sulawesi Utara">
<specimenCount id="90F8ABF8F77EFFDFFE18DCC4FB629A2C" box="[489,543,1790,1817]" count="5" pageId="46" pageNumber="47" type="male">5♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F77EFFDFFDDFDCC5FB1A9A2C" box="[558,615,1791,1817]" count="4" pageId="46" pageNumber="47" type="female">4♀♀</specimenCount>
, 1897,
<collectorName id="2B0B05A7F77EFFDFFD33DCC5FA519A2C" box="[706,812,1791,1817]" pageId="46" pageNumber="47">Bonthain</collectorName>
,
<collectionCode id="E0EFF8B4F77EFFDFFCCBDCC5FA0E9A2C" box="[826,883,1791,1817]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="46" pageNumber="47">SMF</collectionCode>
</materialsCitation>
, no. 1374313747, 13749, 1375113753;—
<materialsCitation id="36966A2CF77EFFDFFA8ADCC4FBF99A08" collectingDate="1914-04-28" collectionCode="SMF" collectorName="Dr L. Martin" country="Indonesia" location="Maros" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
<specimenCount id="90F8ABF8F77EFFDFFA8ADCC4FCE59A2C" box="[1403,1432,1790,1817]" count="1" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
<date id="F24046B1F77EFFDFFF66DD19F8729A08" box="[151,271,1827,1853]" pageId="46" pageNumber="47" value="1914-04-28">
<collectingDate id="E204BF59F77EFFDFFF66DD19F8729A08" box="[151,271,1827,1853]" pageId="46" pageNumber="47" value="1914-04-28">28.iv.1914</collectingDate>
</date>
,
<location id="832136AAF77EFFDFFEECDD19F81A9A08" LSID="urn:lsid:plazi:treatment:0E57D167F77EFFC1FF66DF64FDAA9BD4:832136AAF77EFFDFFEECDD19F81A9A08" box="[285,359,1827,1853]" country="Indonesia" name="Maros" pageId="46" pageNumber="47" stateProvince="Sulawesi Utara">Maros</location>
,
<collectorName id="2B0B05A7F77EFFDFFE84DD19FB759A08" box="[373,520,1827,1853]" pageId="46" pageNumber="47">Dr L. Martin</collectorName>
leg.,
<collectionCode id="E0EFF8B4F77EFFDFFDBADD19FBF99A08" box="[587,644,1827,1853]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="46" pageNumber="47">SMF</collectionCode>
</materialsCitation>
, no. 13933;—
<materialsCitation id="36966A2CF77EFFDFFCDBDD19FC169A54" collectingDate="1983-09-18" collectingDateMax="1983-10-14" collectingDateMin="1983-09-18" collectionCode="RMNH" collectorName="S. S. Pariwono &amp; Coll. M. A. Lieftinck" country="Indonesia" elevation="190" location="Bantimurung" municipality="Maros" pageId="46" pageNumber="47" specimenCount="2" specimenCount-male="2" stateProvince="Sulawesi Utara">
<specimenCount id="90F8ABF8F77EFFDFFCDBDD19FA1C9A08" box="[810,865,1827,1853]" count="2" pageId="46" pageNumber="47" type="male">2♂♂</specimenCount>
,
<date id="F24046B1F77EFFDFFC9EDD19FD579A08" box="[879,1066,1827,1853]" pageId="46" pageNumber="47" value="1983-09-18" valueMax="1983-10-14" valueMin="1983-09-18">
<collectingDate id="E204BF59F77EFFDFFC9EDD19FD579A08" box="[879,1066,1827,1853]" pageId="46" pageNumber="47" value="1983-09-18" valueMax="1983-10-14" valueMin="1983-09-18">18.ix.-14.x.1983</collectingDate>
</date>
,
<collectingMunicipality id="6625FA0BF77EFFDFFBC9DD19FDFF9A08" box="[1080,1154,1827,1853]" pageId="46" pageNumber="47">Maros</collectingMunicipality>
,
<locationDeviation id="97F37953F77EFFDFFB7EDD19FCE59A08" box="[1167,1432,1827,1853]" location="Bantimurung area" pageId="46" pageNumber="47">E of Bantimurung area</locationDeviation>
, surroundings of
<location id="832136AAF77EFFDFFEA4DD7DF8949A54" LSID="urn:lsid:plazi:treatment:0E57D167F77EFFC1FF66DF64FDAA9BD4:832136AAF77EFFDFFEA4DD7DF8949A54" box="[341,489,1863,1889]" country="Indonesia" municipality="Maros" name="Bantimurung" pageId="46" pageNumber="47" stateProvince="Sulawesi Utara">Bantimurung</location>
waterfall,
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<quantity id="4106CD94F77EFFDFFD99DD7DFBCF9A54" box="[616,690,1863,1889]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="46" pageNumber="47" unit="m" value="190.0">190 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F77EFFDFFCF3DD7DFAD99A54" box="[770,932,1863,1889]" pageId="46" pageNumber="47">S.S. Pariwono</collectorName>
leg., ex.
<collectorName id="2B0B05A7F77EFFDFFBE4DD7DFC7B9A54" box="[1045,1286,1863,1889]" pageId="46" pageNumber="47">Coll. M.A. Lieftinck</collectorName>
,
<collectionCode id="E0EFF8B4F77EFFDFFAE5DD7DFC169A54" box="[1300,1387,1863,1889]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="46" pageNumber="47" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1658917, 1658918;—
<materialsCitation id="36966A2CF77EFFDFFE63DD50FB739A9C" collectingDate="1983-09-21" collectingDateMax="1983-10-10" collectingDateMin="1983-09-21" collectionCode="RMNH" collectorName="S. S. Pariwono" country="Indonesia" elevation="250" location="Patunuang Asue" municipality="Maros" pageId="46" pageNumber="47" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Sulawesi Utara">
<specimenCount id="90F8ABF8F77EFFDFFE63DD50F8B59AB0" box="[402,456,1898,1925]" count="3" pageId="46" pageNumber="47" type="male">3♂♂</specimenCount>
,
<specimenCount id="90F8ABF8F77EFFDFFE23DD51FB769AB0" box="[466,523,1899,1925]" count="3" pageId="46" pageNumber="47" type="female">3♀♀</specimenCount>
,
<date id="F24046B1F77EFFDFFDE9DD51FBA99AB0" box="[536,724,1899,1925]" pageId="46" pageNumber="47" value="1983-09-21" valueMax="1983-10-10" valueMin="1983-09-21">
<collectingDate id="E204BF59F77EFFDFFDE9DD51FBA99AB0" box="[536,724,1899,1925]" pageId="46" pageNumber="47" value="1983-09-21" valueMax="1983-10-10" valueMin="1983-09-21">21.ix.-10.x.1983</collectingDate>
</date>
,
<collectingMunicipality id="6625FA0BF77EFFDFFD2EDD51FA549AB0" box="[735,809,1899,1925]" pageId="46" pageNumber="47">Maros</collectingMunicipality>
,
<locationDeviation id="97F37953F77EFFDFFCC5DD51FD4B9AB0" box="[820,1078,1899,1925]" location="Bantimurung area" pageId="46" pageNumber="47">E of Bantimurung area</locationDeviation>
, QK94,
<location id="832136AAF77EFFDFFB62DD51FC379AB0" LSID="urn:lsid:plazi:treatment:0E57D167F77EFFC1FF66DF64FDAA9BD4:832136AAF77EFFDFFB62DD51FC379AB0" box="[1171,1354,1899,1925]" country="Indonesia" municipality="Maros" name="Patunuang Asue" pageId="46" pageNumber="47" stateProvince="Sulawesi Utara">Patunuang Asue</location>
,
<elevation id="0DD38742F77EFFDFFAA7DD50F9B09A9C" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="47" unit="m" value="250.0">
<quantity id="4106CD94F77EFFDFFAA7DD50FCE19AB0" box="[1366,1436,1898,1925]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="47" unit="m" value="250.0">250 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2B0B05A7F77EFFDFFF28DDB5F8059A9C" box="[217,376,1935,1961]" pageId="46" pageNumber="47">S.S. Pariwono</collectorName>
leg.,
<collectionCode id="E0EFF8B4F77EFFDFFE46DDB5FB739A9C" box="[439,526,1935,1961]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="46" pageNumber="47" type="Museum">RMNH</collectionCode>
</materialsCitation>
, no. 1519261, 1519262, 1519264.
</paragraph>
<caption id="D28130F9F77FFFDEFF69DD0FFC139A7B" box="[152,1390,1845,1870]" pageId="47" pageNumber="48" startId="47.[152,255,1845,1869]" targetBox="[155,1434,182,1816]" targetPageId="47" targetType="figure">
<paragraph id="86416071F77FFFDEFF69DD0FFC139A7B" blockId="47.[152,1390,1845,1870]" box="[152,1390,1845,1870]" pageId="47" pageNumber="48">
<emphasis id="B48ABC63F77FFFDEFF69DD0FF8639A78" bold="true" box="[152,286,1845,1869]" pageId="47" pageNumber="48">FIGURE 78</emphasis>
.
<taxonomicName id="41FE1BF2F77FFFDEFEDADD0CF88D9A78" authorityName="Selys" authorityYear="1876" box="[299,496,1846,1870]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F77FFFDEFEDADD0CF88D9A78" box="[299,496,1846,1870]" italics="true" pageId="47" pageNumber="48">Pseudagrion ustum</emphasis>
</taxonomicName>
, holotype female, water colour by Selys, 20.viii.1875 (IRSNB Ag125a). Photo: IRSNB.
</paragraph>
</caption>
<paragraph id="86416071F77FFFDEFF36DDADF88F9A87" blockId="47.[151,1436,1943,2042]" box="[199,498,1943,1970]" pageId="47" pageNumber="48">
<emphasis id="B48ABC63F77FFFDEFF36DDADF88F9A87" bold="true" box="[199,498,1943,1970]" pageId="47" pageNumber="48">Characterization of male</emphasis>
</paragraph>
<paragraph id="86416071F77FFFC1FF36DD86FDA69DE0" blockId="47.[151,1436,1943,2042]" lastBlockId="48.[151,1437,151,1761]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="B48ABC63F77FFFDEFF36DD86F8799AE3" box="[199,260,1980,2006]" italics="true" pageId="47" pageNumber="48">Head</emphasis>
(
<figureCitation id="1EC57CF4F77FFFDEFEE4DD86F8119AE2" box="[277,364,1980,2007]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="47" pageNumber="48">Fig. 95</figureCitation>
): Labrum, clypeus, antefrons and postfrons yellow; colour of postfrons separated from vertex around level of lateral ocelli; vertex red; black posterolateral patches along each lateral ocellus; postocular spots red, separated by a black line from the postfrons; rear of head black, but pale along the eye margins; base and scape of the antennae yellow, following segments brownish ochre; eyes when alive reddish brown-orange.
</paragraph>
<paragraph id="86416071F760FFC1FF36DAE5FB529CC0" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36DAE5F8659DCC" box="[199,280,223,249]" italics="true" pageId="48" pageNumber="49">Thorax</emphasis>
(
<figureCitation id="1EC57CF4F760FFC1FEDBDAE5F8EF9DCC" box="[298,402,223,249]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="48" pageNumber="49">Figs 112</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FE51DAE5F8B39DCC" box="[416,462,223,249]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="48" pageNumber="49">168</figureCitation>
): Anterior lobe of pronotum orange, this colour expanding in centre towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large orange patch in the middle, and on either side a large orange patch dorsolaterally; propleuron black above, orange below and to rear; posterior lobe of pronotum orange; apical edge of posterior lobe convex. Pterothorax orange-reddish; middorsal carina black; humeral suture and metapleural suture each with an elongate black patch at posterior quarter, both patches expanded in middle; interpleural suture may have a narrow blackish stripe, and a blackish spot anteriorly; venter pale, may be covered with pruinosity. Legs pale yellowish-reddish with a black stripe at posterior quarter to half of femora; tibiae with black stripe along black spines.
</paragraph>
<paragraph id="86416071F760FFC1FF36DBC5FD2E9F9C" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36DBC5F84C9F2C" box="[199,305,511,537]" italics="true" pageId="48" pageNumber="49">Abdomen</emphasis>
(
<figureCitation id="1EC57CF4F760FFC1FEB3DBC5F8AE9F2C" box="[322,467,511,537]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="48" pageNumber="49">Figs 129a, b</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FE10DBC5FB729F2C" box="[481,527,511,537]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="48" pageNumber="49">168</figureCitation>
): S1 red, may have a washed-out blackish curly bracket-like patch dorsally; S2 red, but along posterior carina black, often with washed-out blackish marking dorsally at posterior quarter; S38 dorsally black, laterally pale yellow-orange; S9 red, with at least its posterior margin, but up to posterior one-third black; anterior margin of black colour may be serrated; some males additionally have two black spots situated dorsolaterally; S10 red, with a black dorsal “x”-like marking, variable in exact shape.
</paragraph>
<paragraph id="86416071F760FFC1FF36D889FCFB9E90" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36D889F8D59FF8" box="[199,424,691,717]" italics="true" pageId="48" pageNumber="49">Caudal appendages</emphasis>
(
<figureCitation id="1EC57CF4F760FFC1FE46D889FB3A9FF8" box="[439,583,691,717]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="48" pageNumber="49">Figs 146ac</figureCitation>
): Cercus laterally blackish, slightly shorter than S10, inferior lobe subequal to superior lobe, its apex only just visible in this view; in lateral view superior lobe abruptly narrowing near apex, with inter-lobe notch very small; in dorsal view the cercus broadly rounded; the terminus of the superior lobe curved strongly inwards and somewhat bifurcated, this bifurcation more apparent in dorsolateral view; interior projection of inferior lobe of cercus very large, clearly visible in lateral view where it appears subtriangular and basally positioned; paraproct pale yellow-ochre, slightly longer than half of cercus; in lateral view somewhat triangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; medial edge with blackish tooth.
</paragraph>
<paragraph id="86416071F760FFC1FF36D994FB0D9EFC" blockId="48.[151,1437,151,1761]" box="[199,624,942,969]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36D994FB7A9EFD" bold="true" box="[199,519,942,968]" pageId="48" pageNumber="49">Characterization of female</emphasis>
(
<figureCitation id="1EC57CF4F760FFC1FDE6D995FB159EFC" box="[535,616,943,969]" captionStart="FIGURE 78" captionStartId="47.[152,255,1845,1869]" captionTargetBox="[155,1434,182,1816]" captionTargetId="figure-18@47.[151,1436,179,1820]" captionTargetPageId="47" captionText="FIGURE 78. Pseudagrion ustum, holotype female, water colour by Selys, 20.viii.1875 (IRSNB Ag125a). Photo: IRSNB." pageId="48" pageNumber="49">Fig. 78</figureCitation>
)
</paragraph>
<paragraph id="86416071F760FFC1FF36D9E9FAFD9ED8" blockId="48.[151,1437,151,1761]" box="[199,896,979,1005]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36D9E9F8799ED8" box="[199,260,979,1005]" italics="true" pageId="48" pageNumber="49">Head</emphasis>
: As in male, but orange-ochre instead of yellow and red.
</paragraph>
<paragraph id="86416071F760FFC1FF36D9CDF9949948" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36D9CDF8659924" box="[199,280,1015,1041]" italics="true" pageId="48" pageNumber="49">Thorax</emphasis>
: Pronotum as in male, but pale yellowish instead of orange-reddish; posterior lobe with apical edge almost triangular-convex; bearing two yellowish horns directed towards the head, almost reaching the centre of the middle lobe. Pterothorax as in male, but when alive dorsum orange-ochre instead of red, and laterally pale greenish-yellow.
</paragraph>
<paragraph id="86416071F760FFC1FF36DEBDFD9F9860" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36DEBDF84C9994" box="[199,305,1159,1185]" italics="true" pageId="48" pageNumber="49">Abdomen</emphasis>
: S1 orange-ochre, with a black dorsal patch covering anterior two-thirds; S2 orange-ochre, with posterior margin black, and a black middorsal stripe which is expanded patch-like in posterior third; S37 dorsally black, laterally pale yellowish; S8 dorsally black, the black may be constricted in posterior third; S9 dorsally black, narrowing posteriorly, laterally pale yellowish; S10 pale yellowish, anterior margin may be black. Cercus pale; paraproct pale, in lateral view rounded; ovipositor pale yellowish, not reaching to posterior margin of S10, two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth.
</paragraph>
<paragraph id="86416071F760FFC1FF36DF65FD17984C" blockId="48.[151,1437,151,1761]" box="[199,1130,1374,1401]" pageId="48" pageNumber="49">Wings of both sexes with 13.516.0 px in forewing, and 11.515.5 px in hindwing.</paragraph>
<paragraph id="86416071F760FFC1FF36DFB9FC3998F4" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">Measurements of both sexes (mm): Total length (with appendages) 44.550.8; abdomen length (without appendages) 36.842.4; hindwing 24.029.7; forewing 25.631.2; Pt in forewing 0.80.9; male cercus 0.6.</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF760FFC1FF36DFF0FBD49BAC" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="86416071F760FFC1FF36DFF0F84798D1" blockId="48.[151,1437,151,1761]" box="[199,314,1482,1508]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36DFF0F84798D1" bold="true" box="[199,314,1482,1508]" pageId="48" pageNumber="49">Diagnosis</emphasis>
</paragraph>
<paragraph id="86416071F760FFC1FF36DFD5FBD49BAC" blockId="48.[151,1437,151,1761]" pageId="48" pageNumber="49">
The morphology of the male cercus (
<figureCitation id="1EC57CF4F760FFC1FD8CDFD5FA709B3C" box="[637,781,1519,1545]" captionStart="FIGURES 140146" captionStartId="61.[151,264,1645,1669]" captionTargetBox="[153,1432,203,1613]" captionTargetId="figure-18@61.[151,1436,198,1619]" captionTargetPageId="61" captionText="FIGURES 140146. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="48" pageNumber="49">Figs 146ac</figureCitation>
) separates this species from all congeners.
<taxonomicName id="41FE1BF2F760FFC1FAFCDFD5F87F9B18" authorityName="Seehausen &amp; Kalkman &amp; Bedjanič" authorityYear="2025" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="acutidens">
<emphasis id="B48ABC63F760FFC1FAFCDFD5F87F9B18" italics="true" pageId="48" pageNumber="49">Pseudagrion acutidens</emphasis>
</taxonomicName>
has also a cercus with a distinct tooth (
<figureCitation id="1EC57CF4F760FFC1FD49DC29FA389B18" box="[696,837,1555,1581]" captionStart="FIGURES 130139" captionStartId="60.[151,264,1943,1967]" captionTargetBox="[156,1428,185,1914]" captionTargetId="figure-18@60.[154,1433,179,1917]" captionTargetPageId="60" captionText="FIGURES 130139. Male appendages of the Pseudagrion red-group in lateral view (a) and dorsal view (b), cercus in dorsolateral view (c). Illustrations: M. Seehausen." pageId="48" pageNumber="49">Figs 130ac</figureCitation>
), but it is directed inward instead of upward as it is in
<taxonomicName id="41FE1BF2F760FFC1FF66DC0DF98A9B65" authorityName="Selys" authorityYear="1876" box="[151,247,1591,1616]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F760FFC1FF66DC0DF98A9B65" box="[151,247,1591,1616]" italics="true" pageId="48" pageNumber="49">P. ustum</emphasis>
</taxonomicName>
. Additionally the colour marking of the pterothorax and of the abdomen separates
<taxonomicName id="41FE1BF2F760FFC1FB54DC0DFC799B65" authorityName="Selys" authorityYear="1876" box="[1189,1284,1591,1616]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ustum">
<emphasis id="B48ABC63F760FFC1FB54DC0DFC799B65" box="[1189,1284,1591,1616]" italics="true" pageId="48" pageNumber="49">P. ustum</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F760FFC1FAE5DC0DFC119B64" box="[1300,1388,1591,1617]" captionStart="FIGURE 78" captionStartId="47.[152,255,1845,1869]" captionTargetBox="[155,1434,182,1816]" captionTargetId="figure-18@47.[151,1436,179,1820]" captionTargetPageId="47" captionText="FIGURE 78. Pseudagrion ustum, holotype female, water colour by Selys, 20.viii.1875 (IRSNB Ag125a). Photo: IRSNB." pageId="48" pageNumber="49">Figs 78</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FA88DC0CFCEA9B64" box="[1401,1431,1590,1617]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="48" pageNumber="49">95</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FF66DC61F9BE9B40" box="[151,195,1627,1653]" captionStart="FIGURES 96112" captionStartId="57.[151,264,1487,1511]" captionTargetBox="[153,1435,183,1455]" captionTargetId="figure-18@57.[151,1436,179,1461]" captionTargetPageId="57" captionText="FIGURES 96112. Male pterothorax of the Pseudagrion red-group in lateral view. Illustrations: M. Seehausen." pageId="48" pageNumber="49">112</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FF3FDC61F85F9B40" box="[206,290,1627,1653]" captionStart="FIGURES 123129" captionStartId="59.[151,264,1344,1368]" captionTargetBox="[153,1436,184,1313]" captionTargetId="figure-18@59.[151,1436,179,1319]" captionTargetPageId="59" captionText="FIGURES 123129. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="48" pageNumber="49">129a, b</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FEDFDC61F8249B40" box="[302,345,1627,1653]" captionStart="FIGURES 163170" captionStartId="64.[151,264,1734,1758]" captionTargetBox="[155,1431,184,1704]" captionTargetId="figure-18@64.[151,1436,179,1708]" captionTargetPageId="64" captionText="FIGURES 163170. Live males of the Pseudagrion red-group. (163) P. perfuscatum, 13.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia.Photo: J. Hoves. (164) P.perfuscatum, 15.iii.2007, Sanggau Regency, West Kalimantan, Borneo, Indonesia. Photo: J. Hoves. (165) P. pilidorsum, 9.ix.2023, Laiban falls, Tanay, Luzon, Philippines. Photo: M.J. Silvius. (166) P. pilidorsum, 20.vii.2013, Lehmba Danun, Sabah, Malaysia. Photo: M.F.C. Ng. (167) P. cf. pilidorsum [?], 16.i.2023, Papar, Sabah, Malaysia. Photo: D. Matthey. (168) P. ustum, 31.vii.2011, Pattunuang Asue, SW Sulawesi, Indonesia. Photo:A. Günther. (169170) P. spec., ii.2023, southern Taliabu Island, Indonesia. Photos: G. Downing." pageId="48" pageNumber="49">168</figureCitation>
) in both sexes from
<taxonomicName id="41FE1BF2F760FFC1FDCEDC61FBCF9B41" authorityName="Selys" authorityYear="1876" box="[575,690,1627,1652]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="crocops">
<emphasis id="B48ABC63F760FFC1FDCEDC61FBCF9B41" box="[575,690,1627,1652]" italics="true" pageId="48" pageNumber="49">P. crocops</emphasis>
</taxonomicName>
(
<figureCitation id="1EC57CF4F760FFC1FD31DC61FA3C9B40" box="[704,833,1626,1653]" captionStart="FIGURES 2325" captionStartId="15.[151,264,1850,1874]" captionTargetBox="[153,1433,1085,1820]" captionTargetId="figure-58@15.[151,1436,1082,1824]" captionTargetPageId="15" captionText="FIGURES 2325. Pseudagrion semicolon, syntype male: (23) habitus dorsal, (24) head, (25) pronotum. Photos: T. Eldredge, UMMZ." pageId="48" pageNumber="49">Figs 23, 25</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FCBCDC61FA169B40" box="[845,875,1627,1653]" captionStart="FIGURES 7995" captionStartId="56.[151,264,1538,1562]" captionTargetBox="[153,1435,183,1506]" captionTargetId="figure-18@56.[151,1436,179,1512]" captionTargetPageId="56" captionText="FIGURES 7995. Male head of the Pseudagrion red-group in dorsal view. Illustrations: M. Seehausen." pageId="48" pageNumber="49">82</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FC86DC61FAB79B40" box="[887,970,1627,1653]" captionStart="FIGURES 113122" captionStartId="58.[151,264,1841,1865]" captionTargetBox="[153,1436,185,1809]" captionTargetId="figure-18@58.[151,1436,179,1815]" captionTargetPageId="58" captionText="FIGURES 113122. Male abdomen of the Pseudagrion red-group in lateral view (a) and dorsal view (b). Illustrations: M. Seehausen." pageId="48" pageNumber="49">116a, b</figureCitation>
,
<figureCitation id="1EC57CF4F760FFC1FC27DC60FD7E9B40" box="[982,1027,1626,1653]" captionStart="FIGURES 147154" captionStartId="62.[151,264,1572,1596]" captionTargetBox="[154,1435,181,1543]" captionTargetId="figure-18@62.[151,1436,179,1546]" captionTargetPageId="62" captionText="FIGURES 147154. Live males of the Pseudagrion red-group. (147149) P. acutidens, 24.ii.2013, NE of Tatam, Halmahera Timur, Indonesia. Photos: B. Segerer. (150) P. acutidens, 26.xi.2018, SW of Laiwui, Obi, Indonesia. Photo: M. Matthiessen. (151) P. acutidens, 22.viii.2012, Binagara, Halmahera, Indonesia. Photo: B. Schweinhart. (152) P. coriaceum, 22.iii.2017, SW of Opin, Seram, Indonesia. Photo: B. Segerer. (153) P. coriaceum, 30.xi.1996, Stream 5.5 km NW of Roho village, Seram, Indonesia. Photo: M. Bedjanič. (154) P. crocops, Sunai Wera, NW Lake Poso, Central Sulawesi. Photo: A. Günther." pageId="48" pageNumber="49">154</figureCitation>
), currently the only known other red
<taxonomicName id="41FE1BF2F760FFC1FF66DC45F8039BAC" box="[151,382,1663,1689]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF66DC45F85B9BAC" box="[151,294,1663,1689]" italics="true" pageId="48" pageNumber="49">Pseudagrion</emphasis>
species
</taxonomicName>
which occurs at
<collectingRegion id="443AAE93F760FFC1FDCDDC45FBD89BAC" box="[572,677,1663,1689]" country="Indonesia" name="Sulawesi Utara" pageId="48" pageNumber="49">Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="CEE433FAF760FFC1FF36DC98FDAA9BD4" box="[199,1239,1698,1761]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="86416071F760FFC1FF36DC98F8249B89" blockId="48.[151,1437,151,1761]" box="[199,345,1698,1724]" pageId="48" pageNumber="49">
<emphasis id="B48ABC63F760FFC1FF36DC98F8249B89" bold="true" box="[199,345,1698,1724]" pageId="48" pageNumber="49">Distribution</emphasis>
</paragraph>
<paragraph id="86416071F760FFC1FF36DCFDFDAA9BD4" blockId="48.[151,1437,151,1761]" box="[199,1239,1735,1761]" pageId="48" pageNumber="49">
Records of this species are available from
<collectingRegion id="443AAE93F760FFC1FD53DCFDFA779BD4" box="[674,778,1735,1761]" country="Indonesia" name="Sulawesi Utara" pageId="48" pageNumber="49">Sulawesi</collectingRegion>
, the Sula Archipelago, and Buru (
<figureCitation id="1EC57CF4F760FFC1FB74DCFDFDB49BD4" box="[1157,1225,1735,1761]" captionStart="FIGURES 13" captionStartId="5.[152,270,1877,1901]" captionTargetBox="[174,1412,802,1844]" captionTargetId="figure-308@5.[170,1417,798,1850]" captionTargetPageId="5" captionText="FIGURES 13. Distribution maps of the species of the Pseudagrion red-group occurring west of Lydekkers line (black line): (1) P. perfuscatum (green), P. pilidorsum (yellow line), P. p. pruinosum (dark blue), P. p. elongatum (light blue). (2) P. buenafei (dark green), P. declaratum (orange), P. deflexum (light blue), P. enganoense (purple), P. igneum (yellow), P. nigrihumerale (red), P. obscurum (dark blue), P. simalurum (light green). (3) P. acutidens (green line), P. halmaherae (dark blue), P. coriaceum (red), P. crocops (yellow), P. ustum (purple line)." pageId="48" pageNumber="49">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>