diff --git a/data/0E/57/D1/0E57D167F741FFE2FF66D87CFB739C98.xml b/data/0E/57/D1/0E57D167F741FFE2FF66D87CFB739C98.xml new file mode 100644 index 00000000000..5f2e2d70626 --- /dev/null +++ b/data/0E/57/D1/0E57D167F741FFE2FF66D87CFB739C98.xml @@ -0,0 +1,636 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion declaratum +Lieftinck, 1936 + +stat. nov. + + + + +Figures: 26–28 ( +syntype +♂ +, labels); 83 (head +♂ +); 100 (pterothorax +♂ +); 117a, b (abdomen +♂ +); 134a–c (caudal appendages +♂ +); + + + + +155 (field photo +♂ +). + +Pseudagrion pilidorsum +: + +Ris (1915b) +: 99, +Fig. 20 +( +♂ +appendages; +Flores +). + +Pseudagrion pilidorsum declaratum + +: +Lieftinck (1936) +: 130, 132–134, +Figs 13a, b +(description +♂ +♀ +; +♂ +appendages; key; + + +Lombok, Sumbawa, +Sumba +, +Flores +);— +Lieftinck (1953) +: 156–157 (Alor, Sawu);— +Kosterin (2014) +: 16, 21, +Figs 9 +, +13 +, +20 + + +( +Bali +, scarce; Lombok, abbundant; photos alive & habitat). + +Pseudagrion pilidorsum deflexum + +: +Irawan & Rahadi (2018) +: 87 ( +Sumba +). + + + + + +Name-bearing +type +specimens examined + +( +1♂ +, +1♀ +) + + + +1♂ +, +1♀ +syntype +, + +iv.1896 + +, +Sapit +, +Lombok +, +Indonesia +, + +2000 m +a.s.l. + +, +H. Fruhstorfer +leg., +ZMH +( +Figs 26–28 +) + +. + + +Lieftinck (1971: 83) +designated a +lectotype + +male with the data » +Labuan +Badjo +, + +100 m + +, + +xi.1937 + +, +J.K. de Jong +«. However, the +syntype + + +series contains only specimens from this locality and collector which were collected during » + +x.–xi.1929 + +«. +Furthermore +the respective envelope held at +RMNH +( +No. +JvT no. 2813) contains +two males +and a female. +Thus +the designation by +Lieftinck (1971) +is invalid because he did not choose a specimen from the +syntype + +series, and he even did not unambiguously select a particular specimen ( +ICZN 1999 +: 74.2 and 74.5). We do not consider it necessary to designate a +lectotype +, thus we propose to keep them as +syntypes +. + + + +FIGURES 26–28 +. + +Pseudagrion pilidorsum declaratum + +, syntype male: ( +26 +) habitus dorsal, ( +27 +) labels, ( +28 +) habitus lateral. Photos: M. Seehausen. + + + +Other specimens examined +( +29♂♂ +, +5♀♀ +) + + + + +Indonesia +, +Flores Island + +: +2♂♂ +, + +xi.1896 + +, +South +Flores, +R. Martin +leg., +SMF + +, no. 13973–13974;— + +1♂ +, + +28.ix1908 + +, +North +Flores +, +Rioeng Papang +, +river Alo Wai +, v.d. S. leg., +RMNH + +;— + +2♂♂ +, +1♀ +, + +vi.1937 + +, +Labuan Badjo +, +West +Flores +, +J.K. de Jong +leg., +RMNH + +, JvT no. 2813. + + + + +Indonesia +, +Lombok Island + +: +7♂♂ +, +1♀ +, + +iv.1896 + +, +Sapit +, + +2000 m +a.s.l. + +, +H. Fruhstorfer +leg., +SMF + +, no. 2314–2317, 13763–13766;— + +4♂♂ +, + +v.1896 + +, +Ekas +, +H. Fruhstorfer +leg., +SMF + +, no. 13761–13762, 13768–13769;— + +1♂ +, without particular locality, +SMF + +, no. 13767;— + +1♂ +, + +21.iv.1927 + +, +Selong +, +Sunda-Expedition Rensch +, +SMF + +, no. 1Ris01;— + +7♂♂ +, +2♀♀ +, + +18.ii.1993 + +, +Lendang Bajur +, +J. Belle +leg., +RMNH + +. + + + + +Indonesia +, +Sumba +Island + +: +2♂♂ +, + +xii.2015 + +, without particular locality, +MWNH + +. + + + + +Indonesia +, +Sumbawa Island + +: +1♂ +, + +29.v.1927 + +, Dompu, Sunda-Expedition Rensch, +SMF +, no. 1Ris03 + +. + + + + +Indonesia +, +Java +Island + +: +1♂ +, +August +, without particular locality, +NHMUK +, no. 012495866 + +;— + +1♀ +, + +vii.1918 + +, +Bagor +, +Dr W. Röpke +leg., +SMF +, no. 13947 + +. + + +Characterization of male + + +Head +( +Figs 26 +, +83 +, +155 +): Labrum, clypeus, antefrons and postfrons yellow-orange; colour of postfrons distinctly separated from red vertex around level of lateral ocelli; postocular spots red, separated by a very thin black line from postfrons; rear of head black, but broadly yellow-orange along the eye margins; base and scape of antennae yellow, following segments brownish; eyes when alive reddish, anteriorly washed-out yellow-orange. + + +Thorax +( +Figs 26, 28 +, +100 +, +155 +): Anterior, middle, and posterior lobe of pronotum dorsally red; middle pronotal lobe with a black patch on each side dorsolaterally, along the margin between anterior and middle lobe; sutures of middle lobe black, posterior margin of middle lobe broadly black; apical edge of posterior lobe convex. Pterothorax red with a thin black middorsal carina; a black stripe at posterior quarter of humeral suture and metapleural suture, both expanded in the middle; ventrally pale, may be covered with whitish pruinosity. Legs pale reddish; femora may have a blackish stripe outside, narrowing proximally; tibiae blackish along spines; spines black. + + +Abdomen +( +Figs 26, 28 +, +117a, b +, +155 +): S1 red; S2 red, often with a blackish marking at the posterior quarter, which may be connected by a blackish middorsal stripe with black posterior margin; S3–8 dorsally black, laterally pale whitish-yellow; S9 red with posterior edge black; S10 red with black dorsal “x”-like marking, variable in shape. + + +Caudal appendages +( +Figs 134a–c +): Cercus laterally washed-out darkish red, its edges black; slightly shorter than S10; in lateral view cercus subrectangular with open inter-lobe notch low on apex; in dorsal view cercus appearing narrow, apex of the superior lobe narrowly rounded and hooked inwards; a blunt interior projection (appearing subtriangular in dorsolateral view) is present at ca. the middle of the superior lobe; a tiny subbasal interior tooth-like projection on the inferior lobe at ca. the basal third; paraproct pale yellow-ochre, about half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, with black tooth at medial edge. + + +Characterization of female + + +Head +: As in male but red replaced by orange-ochre. + + +Thorax +: Pronotum and pterothorax as in male but red replaced by orange-ochre; posterior lobe of pronotum with two horns directed towards the head, almost reaching the centre of the middle lobe; pterothorax may be tinted greenish antehumerally. + + +Abdomen +: S1 orange-ochre; S2 orange-ochre, with black marking middorsally; S3–8 as in male; S9–10 laterally orange-ochre, dorsally with black markings variable in shape. Cercus washed-out dark brownish, as long as half of S10; paraproct brownish, in lateral view rounded; ovipositor pale yellowish, its apex not reaching to posterior end of S10; two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 11.5–13.5 px in forewing, and 10–12.5 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 38.5–47.0; abdomen length (without appendages) 30.4–37.9; hindwing 20.6–25.0; forewing 22.0–26.5; Pt in forewing 0.7–0.9; male cercus 0.6–0.8. + + + +Diagnosis + + +This species differs from almost all other species of the + +Pseudagrion + +red-group by the largely yellow-red head, bearing just a very thin black line around large red postocular spots ( +Figs 26 +, +83 +, +155 +). This colour pattern is shared only by + +P. deflexum + +( +Figs 29 +, +84 +), but living males of this species have the anterior half of the eyes distinctly yellow ( +Figs 157, 158 +). The laterally subrectangular male appendages of + +P. declaratum + +( +Fig. 134a +) separate this species from several congeners as well, especially from the similarly coloured + +P. deflexum + +, in which the cercus is distinctly arched in lateral view ( +Figs 135a +, +159 +). The male caudal appendages of + +P. obscurum + +and + +P. simalurum + +( +Figs 140a–c, 141a–c +) as well as of + +P. igneum + +and + +P. nigrihumerale + +( +Figs 138a–c, 139a–c +) have superficial similarities with those of + +P. declaratum + +, but these species (except for + +P. nigrihumerale + +) are separated by having S1–2 dorsally black ( +Figs 121a, b +, +123a, b, 124a, b +), black colouration around the postocular spots ( +Figs 87, 88, 89, 90 +), and regarding + +P. nigrihumerale + +by having a black humeral stripe ( +Fig. 105 +). + + + + +Distribution + + +This species is distributed from +Java +towards the +Lesser Sunda Islands +Bali +, Lombok, Sumbawa, +Flores +, +Sumba, Sawu +, and reaching Alor ( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F743FFE5FF66DBC4FC299A44.xml b/data/0E/57/D1/0E57D167F743FFE5FF66DBC4FC299A44.xml new file mode 100644 index 00000000000..1796898c75e --- /dev/null +++ b/data/0E/57/D1/0E57D167F743FFE5FF66DBC4FC299A44.xml @@ -0,0 +1,528 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion deflexum +Lieftinck, 1936 + +stat. nov. + + + + +Figures: 29–30 ( +lectotype +♂ +, envelope); 84 (head +♂ +); 101 (pterothorax +♂ +); 118a, b (abdomen +♂ +); 135a–c (caudal appendages +♂ +); 156–160 (field photos +♂ +). + + + + + + + +Pseudagrion pilidorsum deflexum +Lieftinck (1936) +: 130 + + +, 134, +Fig. 13c +(description +♂ +; +♂ +appendages; key; Timor);— + +Lieftinck (1953) +: 157 + +(Damar, Wetar);— + +Seehausen (2017) +: 5 + +, 8, +Fig. 6 +(Timor; photo +♂ +preserved dry specimen);— + + +Seehausen +et al +. (2018) + +: 9 + +, 11, 13–14, 17–18, 42, 56; +Figs 10 +, +13, 14 +, +18 +, +23, 24 +(Timor; descriptions & photos of habitats; photos +♂ +alive; map);— + + +da Silva Pinto +et al +. (2020) + +: 8 + +, 40 (Rote, Semau; map). + + + +Name-bearing type specimen examined +( +1♂ +) + + + + + +♂ +lectotype +, + +xii.1931 + +– + +i.1932 + +, Amarasi, Timor, +Indonesia +, +E. Handschin +leg., +RMNH +, JvT no. 2814 ( +Figs 29–30 +). +Lectotype +designation by +Lieftinck (1971: 83) +. + + + + + +FIGURES 29–30 +. + +Pseudagrion deflexum + +, lectotype male: ( +29 +) habitus lateral, ( +30 +) envelope. Photos: RMNH. + + + + + +Other +specimens examined + +( +38♂♂ +, +12♀♀ +) + + +Indonesia +, +West Timor + + +: + +2♂♂ +, +Kolbano +, +Timor Central +, ex-coll. +le Moult +, +IRSNB + +;— +1♂ +, +vii.1897 +, Atapupu, + + + + +West Timor +, A + +. Everett leg., +SMF +, no. 13770;— +7♂♂ +, +1♀ +, 1929, +Soe +, +Central Timor +, + +880 m +a.s.l. + +, +SMF +, no. 30113 + +– + + +30119;— + +18♂♂ +, +9♀♀ +, + +vi.1929 + +, +Soe +, +Timor Central +, ex-coll. +le Moult +, +IRSNB + +;— +1♀ +, +vi.1935 +, Nikiniki, C. Bühler + + +& Meyer, Basel Museum, + +MCZ +;— +1♂ +, + +xi.1992 + +, without particular locality, +Coll. von Rosen + +, + +ZSM +, no. +GvRosen +13022 + +. + + + + +Timor-Leste + +: +1♀ +, + +viii.1935 + +, +Baguia +, +C. Bühler +& +Meyer +, +Basel Museum +, +MCZ + +;— + +4♂♂ +, + +19.x.1973 + +, +Fatu Maca +, +10 km +south of +Baucau +, +Tom Weir +leg., +MAGNT + +;— + +2♂♂ +, + +25.v.2012 + +, +Kablaki Hotel +, +Same +, +8.999969°S +, +125.648072°E +, +AMS + +, no. K462953;— + +1♂ +, + +3.vi.2012 + +, +Gully + +0.5km +NE Laclubar + +, +8.74647°S +, +125.91497°E +, +AMS + +, no. K376343;— + +3♂♂ +, + +5.vi.2012 + +, spring, +3.8km +WNW +Manatuto +, +8.50847°N +, +125.98292°E +, +AMS + +, no. K376350. + + +Characterization of male + + +Head +( +Figs 29 +, +84 +, +157, 158 +): Labrum, clypeus, antefrons and postfrons bright yellow; colour of postfrons sharply separated from red vertex at level of lateral ocelli; postocular spots red, separated by a very thin black line from postfrons; rear of head black, but broadly yellow-orange along the eye margins; base and scape of antennae yellow, following segments brownish; eyes when alive red with anterior half yellow, continuing in line with the yellow frons. + + +Thorax +( +Figs 29 +, +101 +, +156, 157, 160 +): Anterior, middle, and posterior lobe of pronotum dorsally red; middle pronotal lobe with a black patch on each side dorsolaterally, along the margin between anterior and middle lobe; sutures of middle lobe black, posterior margin of middle lobe broadly black; apical edge of posterior lobe convex. Pterothorax red, with thin black middorsal carina; black stripes at posterior quarter of humeral suture and metapleural suture, both expanded in the middle; ventrally pale, may be covered with pruinosity; when immature basic colouration yellowish instead of red, with broadly brownish dorsum and humeral stripe ( +Fig. 160 +). Legs pale; femora with blackish stripe outside, narrowing proximally; tibiae blackish along spines; spines black. + + +Abdomen +( +Figs 29 +, +118a, b +, +156, 160 +): S1 red; S2 red, with black marking dorsally along posterior carina; S3–8 dorsally black, laterally whitish-yellow; S9 red with posterior edge black; S10 red, with black dorsal “x”-like marking, variable in shape; when immature basic colouration yellowish instead of red ( +Fig. 160 +). + + +Caudal appendages +( +Figs 135a–c +, +159 +): Cercus laterally blackish, slightly shorter than S10; in lateral view whole cercus distinctly arched, apex subrectangular with inter-lobe notch almost central; in dorsal view cercus appearing narrow, apex of the superior lobe narrowly rounded and turned inwards; a subtriangular interior projection present at ca. the centre of the superior lobe of the cercus; a tiny interior tooth-like projection before half-length on the inferior lobe of the cercus; paraproct pale yellow-ochre, slightly longer than half of the cercus, in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, medial edge without tooth. + + +Characterization of female + + +Head +: As in male but red replaced by orange-ochre. + + +Thorax +: Pronotum and pterothorax as in male but red replaced by orange-ochre; posterior lobe of pronotum with two horns directed towards the head, almost reaching the centre of the middle lobe; antehumeral pale, tinted ochre-greenish when alive. + + +Abdomen +: S1 orange-ochre; S2 orange-ochre, with black marking middorsally; S3–8 as in male; S9–10 laterally orange-ochre, dorsally with black markings variable in shape. Cercus washed-out dark brownish, as long as half of S10; paraproct brownish, in lateral view rounded; ovipositor pale yellowish, its apex not reaching to posterior end of S10; two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 10.5–12.5 px in forewing, and 9–11 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 38.0–45.5; abdomen length (without appendages) 32.5–37.5; hindwing 20.8–23.5; forewing 22.0–25.7; Pt in forewing 0.6–1.0; male cercus 0.5–0.6. + + + +Diagnosis + + +The morphology of the male appendages of + +P. deflexum + +is unmistakable: the cercus is arched in lateral view ( +Figs 135a +, +159 +), a unique character compared to all congeners. Additionally + +P. deflexum + +is distinctly different from all congeners, except for the adjacently distributed + +P. declaratum + +, in having a largely yellow-red head with just a very thin black line around the large red postocular spots ( +Figs 29 +, +84 +, +157 +). But the yellow colour of the frons is continuous in line to the anterior half of the eyes in live males of + +P. deflexum + +( +Figs 157, 158 +), whereas the eyes are reddish in + +P. declaratum + +( +Fig. 155 +). + + + + +Distribution + + +This species occurs on Timor, as well as on adjacent islands of Rote, Semau, Wetar, and Damar ( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F745FFE7FF66DAADFB6999D0.xml b/data/0E/57/D1/0E57D167F745FFE7FF66DAADFB6999D0.xml new file mode 100644 index 00000000000..07812167ad0 --- /dev/null +++ b/data/0E/57/D1/0E57D167F745FFE7FF66DAADFB6999D0.xml @@ -0,0 +1,327 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion enganoense +Lieftinck, 1948 + +stat. nov. + + + + +Figures: 31–32 ( +lectotype +♂ +, envelope); 85 (head +♂ +); 102 (pterothorax +♂ +); 119a, b (abdomen +♂ +); 136a–c (caudal appendages +♂ +). + + + + + + +Pseudagrion pilidorsum enganoense + +: + +Lieftinck (1948) +: 290 + +–291, +Fig. 6 +(description +♂ +♀ +; Engano). + + + +Name-bearing type specimen examined +( +1♂ +) + + + + + +♂ +lectotype +, + +30.v.1936 + +, +Buahbuah +, +Enggano Island +, +RMNH +, JvT no. 2816 ( +Figs 31–32 +) + +. + + +Lectotype +designation by +Lieftinck (1971: 86) +. +Lieftinck (1948) +spelled the island “Engano”, that’s why the scientific name is written with a single “g”. However, today the island is spelled “Enggano”. + + + + +FIGURES 31–32 +. + +Pseudagrion enganoense + +, lectotype male: ( +31 +) habitus lateral, ( +32 +) envelope. Photos: RMNH. + + + + +Other specimens examined +( +2♂♂ +) + + + + +Indonesia +, +Enggano Island +: + +2♂♂ +, + +30.v.1936 + +, Buahbuah, +RMNH +, no. 1658925, 1658926 + +. + + +Characterization of male + + +Head +( +Figs 31 +, +85 +): Labrum, clypeus, antefrons and postfrons yellow; colour of postfrons distinctly separated from vertex around the level of lateral ocelli; vertex red, with a small black spot posterior of median ocellus, and large black patches each along interior and posterior margin of lateral ocelli; postocular spots red, separated by black line from postfrons and vertex; rear of head black, but yellow-orange along the eye margins; base and scape of antennae yellow, following segments brownish. + + +Thorax +( +Figs 31 +, +102 +): Anterior lobe of pronotum yellowish, expanding in centre spot-like towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided yellowish patch in the middle, and on each side a yellowish spot dorsolaterally; propleuron black above, yellowish below and to rear; posterior lobe of pronotum yellowish; apical edge of posterior lobe convex. Pterothorax red, with thin black middorsal carina; humeral suture and metapleural suture with black streaks at posterior quarter; interpleural suture with a thin black line; ventrally pale, may be covered with whitish pruinosity. Legs pale orange-reddish; femora with blackish stripe outside, narrowing proximally; tibiae blackish along spines; spines black. + + +Abdomen +( +Figs 31 +, +119a, b +): S1 red, S2 red with its posterior margin black; S3–8 dorsally black, laterally pale orange-yellow; S9 black with anterior third to half red; S10 black, ventrally pale. + + +Caudal appendages +( +Figs 136a–c +): Cercus laterally obscure reddish; slightly shorter than S10, inferior lobe slightly longer than superior; in lateral view subrectangular, apex with inter-lobe notch deep and almost central; in dorsal view cercus appearing narrow, with terminus of superior lobe slightly turned in, bifurcated interiorly (visible in dorsolateral view); a small almost centrally-placed interior tooth-like projection on the inferior lobe of the cercus; paraproct pale ochre; around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, with blackish tubercular medial edge. + +Measurements (mm): Total length (with appendages) 42.3–44.5; abdomen length (without appendages) 35.7– 36.7; hindwing 23.7–24.3; forewing 25.0–26.0; Pt in forewing 0.7–1.0; male cercus 0.5–0.6. + +Characterization of female +[not examined, characterization adopted from +Lieftinck (1948) +] + + +Head +: As in male but ochraceous instead of yellow. + + +Thorax +: Pterothorax as in male but dorsally honey-yellowish, fading to cinnamon-buff laterally; markings at humeral and metapleural suture rusty. + + +Abdomen +: As in male but with S1–2 orange; S9 with a large yellow spot laterally; ovipositor yellowish. + + +Wings of both sexes with 14.5–17 px in forewing, and 13–16 px in hindwing [adopted from +Lieftinck (1948) +, and own data]. + + + + +Diagnosis + + + +Pseudagrion enganoense + +belongs to the group of endemic taxa of the island chain west of +Sumatra +. The morphology of the male cercus of + +P. enganoense + +is distinct: its inferior lobe is slightly longer than superior one ( +Figs 136a, b +), and in dorsolateral view the inner margin of superior lobe of cercus lacks an inward projection proximal of its bifurcated terminus ( +Fig. 136c +). Additionally both sexes of + +P. enganoense + +can be separated from congeners on adjacent islands by having S1–2 red in the male ( +Figs 31 +, +119a, b +), orange in the female. This character is shared with + +P. declaratum + +, + +P. deflexum + +, and + +P. nigrihumerale + +( +Figs 117a, b, 118a, b, 122a, b +). However + +P. nigrihumerale + +can also be separated by its broadly black antehumeral stripe ( +Fig. 105 +), while + +P. declaratum + +as well as + +P. deflexum + +have much larger red postocular spots than + +P. enganoense + +, not bordered by black except for a very small blackish line on vertex ( +Figs 26 +, +29 +, +31 +, +83, 84, 85 +, +155, 157 +). + + + + +Distribution + + +Endemic to Enggano ( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F746FFE8FF66DF14FB6498B3.xml b/data/0E/57/D1/0E57D167F746FFE8FF66DF14FB6498B3.xml new file mode 100644 index 00000000000..28efab72828 --- /dev/null +++ b/data/0E/57/D1/0E57D167F746FFE8FF66DF14FB6498B3.xml @@ -0,0 +1,398 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion halmaherae + +sp. nov. + + + + +Figures: 33 (habitus +holotype +♂ +); 34–36 (S9–10, caudal appendages +holotype +♂ +); 37 (head +holotype +♂ +); 38 (pronotum +holotype +♂ +); 39 (habitus +paratype +♀ +); 40 (head +paratype +♀ +); 41 (pronotum +paratype +♀ +); 42 (S9–10, ovipositor +paratype +♀ +); 86 (head +♂ +); 103 (pterothorax +♂ +); 120a, b (abdomen +♂ +); 137a–c (caudal appendages +♂ +). + + + + + +? + +Pseudagrion coriaceum + +: + +Selys (1878) +: 296 + +(Halmahera);— + +Selys (1879) +: 290 + +(Halmahera). + + + + + + + +Holotype + +: +♂ +, + +7.–12.x.1951 + +, +Biaur +, +Halmahera Island +, +North Maluku province +, +Indonesia +, + +600 m + +a.s.l, approximately +1.3°N +, +127.7°E +, no. 1519084. +Deposited +at +Naturalis Biodiversity Center Leiden +, the Netherlands. + + + + +Paratype + +( +1♀ +): +1♀ +, +7.–12.x.1951 +, + +Biaur +, +Halmahera Island +, +North Maluku province +, +Indonesia +, + +600 m +a.s.l. + +, approximately +1.3°N +, +127.7°E +, +RMNH +, no. 1519084 + +. + + + + +Etymology: +Named with regards to the +type +locality Halmahera Island. + + + + +Remarks +: Further specimens from Halmahera were discovered among unidentified material at the RMNH by V.J. Kalkman just prior to submission. For this reason we have not included them in the +type +series. + + + + + +Description of +holotype +male + + + +Head +( +Fig. 37 +, also compare +Fig. 86 +): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow; colour of postfrons sharply separated from black vertex at level of anterior margin of lateral ocelli; median ocellus with a small black spot posteriorly; vertex with a small yellowish spot situated at centre of occipital bar area; postocular spots orange-reddish, rectangle-like with posterior edge somewhat serrated, bordered by black; rear of head black, but yellowish along the eye margins; base and scape of antennae yellow, following segments ochre-brownish. + + +Thorax +( +Figs 33, 38 +, also compare +Fig. 103 +):Anterior lobe of pronotum with its centre orange, broadly surrounded by black, and orange patches laterally; middle pronotal lobe black with two parallel elongate orange patches in centre, and on each side a subtriangular orange spot dorsolaterally; propleuron black above, broadly orange below and to rear; posterior lobe of pronotum black, with an elongate orange spot in centre and broadly orange at each lateral margin; apical edge of posterior lobe convex. Pterothorax orange-reddish with its posterior margins black; a broad black middorsal stripe; a broad black humeral stripe, broadened posteriorly; interpleural suture with a black stripe reaching to metathoracic spiracle, broadened around its middle; metapleural suture with a black stripe, broadened posteriorly; mesinfraepisternum with a broad black diagonal stripe, lower margin pale orange; venter of pterothorax pale yellow-orange, covered with whitish pruinosity. Coxae and legs pale yellow-orange; outer side of femora with a washed-out blackish stripe along posterior two-third at foreleg, along posterior half at midleg and lacking at hindleg; tibiae with blackish along the black spines; tarsi washed-out blackish; tarsal claws ochre. + + +Wings +( +Fig. 33 +): Membrane hyaline; veins brown to black; 16.0–16.5 px in forewing; 14.5–15.0 px in hindwings; Pt dark reddish-brown, overlying one cell. + + + +FIGURES 33–38 +. + +Pseudagrion halmaherae + +, holotype male: ( +33) +habitus lateral, ( +34 +) appendages lateral, ( +35 +) appendages dorsal, ( +36 +) cercus dorsolateral, ( +37 +) head dorsal, ( +38 +) pronotum. Photos: M. Seehausen. + + + +Abdomen +( +Figs 33, 34, 35 +, also compare +Figs 120a, b +): S1 orange, dorsally with a goblet-like black patch; S2 orange, dorsally black, black colour expanded laterally at posterior end; S3–8 black dorsally; yellow-orange laterally; S9 orange, with a black middorsal stripe, and a quadrangle-like black patch covering posterior two-thirds of segment; laterally with the posterior third black; S10 orange, dorsally black. + + +Caudal appendages +( +Figs 34, 35, 36 +, also compare +Figs 137a–c +): Cercus washed-out blackish, slightly shorter than S10, inferior lobe shorter than superior lobe; cercus in lateral view subtriangular due to an almost straight upper margin of the cercus, and a convex lower margin; lower margin with a narrow but deep inter-lobe notch subapically; in dorsal view cercus almost straight, then turned slightly inward with its apex narrowly rounded and in dorsolateral view curved inward; in dorsal and dorsolateral view a cleft is visible between superior and inferior lobe; inferior lobe projected shelf-like to inward, a forward and inward directed subbasal interior projection is present, appearing larger in dorsolateral view; paraproct pale yellow-ochre, slightly longer than half of cercus; in lateral view subtriangular and tapered with a rounded apex; in dorsal view broadly rounded, concave bowl-like; with a small black tooth at medial edge. + +Measurements (mm): Total length (with appendages) 51.3; abdomen length (without appendages) 42.3; hindwing 28.8; forewing 30.5; Pt in forewing 0.8; cercus 0.6. + + +Description of female +paratype + + + +Head +( +Fig. 40 +): Labium, mandibles, genae, labrum, and anteclypeus yellowish-ochre; postclypeus black; antefrons, and postfrons yellowish with a large black patch at centre; yellow colour of postfrons sharply separated from vertex at level of anterior margin of lateral ocelli; vertex black; a black stripe expanded towards each antenna, and broadly towards the median ocellus, reaching level of lateral ocelli; occipital bar thin yellowish, its centre slightly expanded anteriorly; postocular spots yellow-orange, rectangle-like with their posterior edge somewhat serrated, bordered by black; rear of head black, but yellowish along the eye margins; base and scape of antennae yellowish-ochre, following segments brownish. + + +Thorax +( +Figs 39, 41 +): Anterior lobe of pronotum yellowish, yellowish colour expanding in centre towards middle lobe, its anterior margin black; middle pronotal lobe black, with a centrally divided large yellowish patch in the middle, and on either side a yellowish spot dorsolaterally; propleuron black above, broadly yellowish below and to rear; posterior lobe of pronotum black; a yellowish spot in centre, and washed-out dark yellowish at each lateral margin; apical edge of posterior lobe convex, bearing two darkish horns directed towards the head, almost reaching the centre of the middle lobe. Pterothorax yellowish with its posterior margins black; a broad black middorsal stripe; a broad black humeral stripe, broadened posteriorly; interpleural suture with a black stripe reaching to half of the pterothorax, its anterior terminus broadened; metapleural suture with a black stripe, broadened posteriorly; mesinfraepisternum with a broad black diagonal stripe, lower margin pale yellowish; venter of pterothorax pale yellowish, covered with whitish pruinosity. Coxae and legs pale yellowish; outer side of femora partly with a washed-out blackish stripe along posterior half; tibiae blackish along the black spines; tarsi obscure blackish; tarsal claws ochre. + + +Wings +( +Fig. 39 +): Membrane hyaline; veins brown to black; 17.5–18.5 px in forewing; 15.5 px in hindwings; Pt pale reddish-brown, overlying one cell. + + +Abdomen +( +Figs 39, 42 +): S1–8 dorsally black, laterally pale orange; each basal annulus pale yellowish-orange; S9 dorsally black, laterally pale yellowish-orange, this pale colour expanded dorsally in centre of segment; S10 dorsally black, laterally pale yellowish. Cercus in lateral view straight and tapered, blackish, about half as long as S10; paraproct pale yellow-brownish, short, in lateral view rounded; ovipositor yellowish-orange, not reaching posterior end of S10, two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Measurements (mm): Total length (with appendages) 50.7; abdomen length (without appendages) 40.5; hindwing 31.6; forewing 33.0; Pt in forewing 0.9. + + + +Diagnosis + + +The morphology of the male cercus ( +Figs 137a–c +) is distinct compared to all congeners. There are superficial similarities to the male cercus of + +P. coriaceum + +( +Figs 132a–c +), but + +P. halmaherae + +differs from the latter by the clearly visible inter-lobe notch in lateral view ( +Figs 132a, 137a +). Additionally the rectangular postocular spots of + +P. halmaherae + +, with their somewhat serrated posterior margin, separate this species in both sexes from congeners ( +Figs 33 +, +40 +, +86 +). Both sexes also have an obvious colour pattern of the pterothorax, in some aspects similar to that of + +P. crocops + +, but the antehumeral stripe of males is red in + +P. halmaherae + +( +Fig. 103 +) versus greenish in + +P. crocops + +( +Figs 99 +, +154 +). + + + + +FIGURES 39–42 +. + +Pseudagrion halmaherae + +, female paratype: ( +39 +) habitus lateral, ( +40 +) head dorsal, ( +41 +) pronotum, ( +42 +) ovipositor lateral. Photos: M. Seehausen. + + + + +Distribution + + +Probably endemic to Halmahera ( +Fig. 3 +). But as this region is little explored, it is possible that this species occurs on adjacent islands as well. + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F749FFEAFF66DFEDFA1E9AB1.xml b/data/0E/57/D1/0E57D167F749FFEAFF66DFEDFA1E9AB1.xml new file mode 100644 index 00000000000..ca302dc18a5 --- /dev/null +++ b/data/0E/57/D1/0E57D167F749FFEAFF66DFEDFA1E9AB1.xml @@ -0,0 +1,422 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion igneum + +sp. nov. + + + + +Figures: 43 (habitus +holotype +♂ +); 44–46 (S9–10, caudal appendages +holotype +♂ +); 47 (head +holotype +♂ +); 48 (pronotum +holotype +♂ +); 87 (head +♂ +); 104 (pterothorax +♂ +); 121a, b (abdomen +♂ +); 138a–c (caudal appendages +♂ +); 161–162 (field photos +♂ +). + + + + + +Pseudagrion pilidorsum + +: Hämäläinen +et al +. (1999): 28 (Middle Andaman; subspecific status remains to be studied);— + +Koparde +et al +. (2018) + +: 41, + +Figs +2g +, 3d + +(North Andaman; map; photo +♂ +alive, habitat photo). + + + + + + +Holotype + +: +♂ +, + +8.xii.1998 + +, +Chiriyatapu +, +South Andaman +, +India +, approximately +11.503°N +, +92.702°E +, ex. +Coll. M. Hämäläinen +, +Wen-Chi Yeh +, no. 1152912. +Deposited +at +Naturalis Biodiversity Center Leiden +, the Netherlands. + + + + +Paratypes + +( +2♂♂ +): +2♂♂ +, +9.xii.1976 +, + +rivulet near +Asmadabad +, +South Andaman +, +India +, +RMNH +, no. 1552773 + +. + + + + +Etymology: +The name refers to the fiery impression of the bright red thorax colouration of mature males ( +Figs 161, 162 +). + + + + +Remarks + + +Matjaž Bedjanič observed a male of this species at a stream along the Mayabunder-Diglipur section of the Great Andaman Trunk Road in the North Andaman ( +13.1872°N +, +92.8893°E +; +Figs 161, 162 +). This stream is about one metre wide, with sparse bank vegetation, and the surroundings are covered by forest. The locality is situated about +5 km +SW of the locality where the species has been observed a month later by + +Koparde +et al +. (2018) + +, and approximately +80 km +N of the species’ locality in the Middle Andaman, where a male of the species was collected in +June 1997 +, and reported by Hämäläinen +et al +. (1999). It is expected that with further targeted fieldwork in the right season and in suitable habitats, the species will be found at additional localities on the Andaman Islands. However, as the newly described + +Pseudagrion + +is a colourful and quite conspicuous damselfly, the fact that only records of +six males +are known to date, and that rather recent fieldtrips of M. Bedjanič and P. Koparde, P. Dawn and A. Sumanapala in monsoon season of 2017 yielded only one observation each, suggest that the species is rather local. Additional information on distribution, biology, phenology, and habitat requirements is needed for relevant assessment of threat status of this interesting new endemic. + + +Other species recorded at the locality were: + +Vestalis gracilis +( +Rambur, 1842 +) + +, + +Copera marginipes +( +Rambur, 1842 +) + +, + +Prodasineura verticalis andamanensis +( +Fraser, 1924 +) + +, + +Tetrathemis platyptera +Selys, 1878 + +, and + +Trithemis aurora +( +Burmeister, 1839 +) + +. + + + + + +Description of +holotype +male + + + +Head +( +Fig. 47 +, also compare +Fig. 87 +): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow; yellow of postfrons sharply separated at level of median ocellus; vertex black, bordered by an orange-reddish rhombus along the ocelli; a black spot posterior of the median ocellus; followed posteriorly by a large orange-reddish spot, elongated to level of posterior end of lateral ocelli; postocular spots orange-reddish, bordered by black; rear of head black, but narrow yellowish along the eye margins; base and scape of antennae yellow, pedicel brownish, rest of antennae missing. + + +Thorax +( +Figs 43, 48 +, also compare +Fig. 104 +): Anterior lobe of pronotum orange, with its anterior margin black; middle pronotal lobe black, an orange-reddish patch in the middle centrally divided by a black line, and on either side an orange-reddish patch dorsolaterally; propleuron black above, orange below and to rear; posterior lobe of pronotum orange; apical edge of posterior lobe convex. Pterothorax orange; middorsal carina thin black; mesopleural suture with an elongated black patch at posterior third; interpleural suture thin black, with a small dark spot anteriorly; metapleural suture with an elongated black patch at posterior third; mesinfraepisternum pale orange, its superior and posterior edge black; venter of pterothorax yellow-orange, covered with whitish pruinosity. Legs yellowish-ochre, outer side washed with a blackish stripe; tip of tarsi and tarsal claws brown; spines blackish. + + +Wings +( +Fig. 43 +): Membrane hyaline; veins brown to black; 12.5 px in right forewing, left forewing damaged; 11.5 px in both hindwings; Pt dark reddish-brown, overlying one cell. + + +Abdomen +( +Fig. 43 +, also compare +Figs 121a, b +): S1 dorsally washed out blackish, laterally orange; S2 dorsally black, apically the black expanded laterally, otherwise laterally orange; S3–8 dorsally black, apically on each segment the black expanded laterally; each segment laterally pale white-yellowish, except for S8 which is laterally orange; each segment with a narrow pale white-yellowish basal annulus; S9 orange, with its posterior third dorsally and laterally black; S10 ( +Figs 44, 45 +) with a black posterior margin, and a black middorsal stripe making a “Y”-like marking dorsally; laterally a faint and very narrow black stripe, fading anteriorly, separating two yellow-orange dorsolateral patches. + + +Caudal appendages +( +Figs 44–46 +, also compare +Figs 138a–c +): Cercus laterally washed-out blackish, slightly shorter than S10, inferior lobe appearing slightly shorter than superior lobe; in lateral view cercus subrectangular, apex with a broad inter-lobe notch; in dorsal view cercus appears narrow, apex of superior lobe slightly pointed and curved inwards, a subtriangular interior projection is situated just proximal of the middle of the cercus; a small, tooth-like subbasal interior projection before half-length of the inferior lobe of the cercus, visible in dorsal and dorsolateral views; paraproct pale yellow-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, with prominent black tooth at blackish medial edge. + +Measurements (mm): Total length (with appendages) 39.3; abdomen length (without appendages) 32.2; hindwing 21.5; forewing 22.8; Pt in forewing 0.8; cercus 0.6. + + +Variation in +paratype +males + + + +Wings +: 13.5–14 px in forewing, 10.5–12.5 px in hindwing. + + +Abdomen +: Dorsal black on S9 may be expanded to the posterior half. + +Measurements (mm): Total length (with appendages) 40.3–42.5; abdomen length (without appendages) 34.3– 35.0; hindwing 22.2; forewing 23.8; Pt in forewing 0.8; cercus 0.5–0.6. +Female unknown. + + + +FIGURES 43–48 +. + +Pseudagrion igneum + +, holotype male: ( +43) +habitus lateral, ( +44 +) appendages lateral, ( +45 +) appendages dorsal, ( +46 +) cercus dorsolateral, ( +47 +) head dorsal, ( +48 +) pronotum. Photos: M. Seehausen. + + + + +Diagnosis + + + +Pseudagrion igneum + +is separated from its Nicobar congeneric + +P. nigrihumerale + +by lacking a broad black humeral stripe ( +Figs 104, 105 +), S1–2 of the abdomen dorsally black, S10 red with black dorsal marking ( +Figs 121a, b, 122a, b +), as well as by red on vertex and postfrons ( +Figs 87, 88 +), whereas the male caudal appendages are identical in both species ( +Figs 138a–c, 139a–c +). Also the male cerci of + +P. simalurum + +and + +P. obscurum + +are almost identical, seemingly with only a tiny different: the cercus of + +P. igneum + +bears a small subbasal tooth before half-length of the inferior lobe, whereas in + +P. obscurum + +and + +P. simalurum + +there is a sharp edged bulge that only looks like a tooth in dorsal view ( +Figs 140a–c, 141a–c +). Additionally + +P. igneum + +lacks a darkened dorsum of the pterothorax as it is in + +P. obscurum + +( +Fig. 106 +). See also table 2 under + +P. obscurum + +for further information. All other species are separated by the morphology of the male appendages, and partly by the colour markings of head, thorax, and abdomen. + + + + +Distribution + + +Apparently endemic to the Andaman Islands, +India +( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F74CFFEFFF66DAADFA2F9A44.xml b/data/0E/57/D1/0E57D167F74CFFEFFF66DAADFA2F9A44.xml new file mode 100644 index 00000000000..fcb5e1b9fb6 --- /dev/null +++ b/data/0E/57/D1/0E57D167F74CFFEFFF66DAADFA2F9A44.xml @@ -0,0 +1,593 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion nigrihumerale + +sp. nov. + + + + +Figures: 49 (habitus +holotype +♂ +); 50–52 (S10, caudal appendages +holotype +♂ +); 53 (head +holotype +♂ +); 54 (pronotum +holotype +♂ +); 55 (habitus +paratype +♀ +); 56 (head +paratype +♀ +); 57 (pronotum +paratype +♀ +); 58 (S9–10, ovipositor +paratype +♀ +); 88 (head +♂ +); 105 (pterothorax +♂ +); 122a, b (abdomen +♂ +); 139a–c (caudal appendages +♂ +). + + + + + + +Pseudagrion pilidorsum + +: + + +Rajeshkumar +et al +. (2016) + +: 45 + +, +Fig. 2H +( +Nicobar Islands +; photo +♂ +alive). + + + + + + + +Holotype + +: +♂ +, + +8.i.2001 + +, +Munak +, +Kamorta Island +, +Nicobar Islands +, +India +, approximately +8.01°N +, +93.505°E +, +Dr Prashant Mohanraj +leg., ex. +Coll. M. Hämäläinen +, no. 1552778. +Deposited +at +Naturalis Biodiversity Center Leiden +, the Netherlands. + + + + +Paratypes + +( +8♂♂ +, +3♀♀ +): +1♂ +, +3♀♀ +, +23.xii.2000 +, + +27 km +E–W road, +Great Nicobar +, +India +, +Dr Prashant Mohanraj +leg., ex. +Coll. M. Hämäläinen +, +RMNH +, no. 1552782–1552784, 1552786 + +;— +3♂♂ +, +8.i.2001 +, + +Munak +, +Kamorta Island +, +Nicobar Islands +, +India +, +Dr Prashant Mohanraj +leg., ex. +Coll. M. Hämäläinen +, +RMNH +, no. 1552777, 1552779, 1552780 + +;— +4♂♂ +, +9.i.2001 +, + +Agriculture Department Farm +, +Kamorta Island +, +Nicobar Islands +, +India +, +Dr Prashant Mohanraj +leg., ex. +Coll. M. Hämäläinen +, +RMNH +, no. 1552774–1552776, 1552881 + +. + + + + +Other specimens + +: +1♂ +, + +19.xii.1996 + +, +Campbell Bay +, Great Nicobar, +India +, ex. +Coll. M. Hämäläinen +, Wen-Chi Yeh, +RMNH +, no. 1152932. This freshly emerged male belongs to this new species as well, but was excluded from the +type +series because of its damaged condition + +. + + + + +Etymology: +The name refers to the black humeral stripe on the thorax, a unique character of the revised species of the + +Pseudagrion + +red-group occurring in the western range. + + + + +Remarks + + +The colours when alive are adopted from the field photograph of a living male provided by + +Rajeshkumar +et al +. (2016: 43 + +, +Fig. 2H +). Additionally they provide the following data of this species (identified as + +P. pilidorsum + +): + +1♂ +, + +18.xii.2015 + +, +Near +dam, +Kamorta +, +Central Nicobar +( +8°8.242’N +, +93°29.256’E += +8.137367°N +, +93.4876°E +) + +; + +1♂ +, + +17.ii.2016 + +, near streams in interior forest, +Galathea +, +Great Nicobar Island +( +7°0.245’N +, +93°54.181’E += +7.004083°N +, +93.903017°E +) + +; +1♂ +, +1♀ +, +21.ii.2016 +; + +♀ +laying eggs on stream near +Laxmi Nagar +, +Great Nicobar Island +( +6°56.343’N +, +93°53.468’E += +6.93905°N +, +93.891133°E +) + +. + + + + + +Description of +holotype +male + + + +Head +( +Fig. 53 +, also compare +Fig. 88 +): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow; colour of postfrons sharply separated from vertex at anterior margin of lateral ocelli; vertex black; median ocellus with a black spot anteriorly, merged with black colour of vertex; occipital bar reddish; postocular spots reddish, bordered by black; rear of head black, but yellowish along the eye margins; base and scape of antennae yellow, following segments ochre-brownish. + + +Thorax +( +Figs 49, 54 +, also compare +Fig. 105 +): Anterior lobe of pronotum orange-reddish, expanding in centre towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large orange-reddish patch in the middle, and on either side a large orange-reddish patch dorsolaterally, fusing laterally with orange-reddish colour of anterior lobe; propleuron black above, orange-reddish below and to rear; posterior lobe orange-reddish; apical edge of posterior lobe convex. Pterothorax orange-reddish with a broad black middorsal stripe, and a broad black humeral stripe, narrowed posteriorly; metapleural suture with a small elongate black patch at posterior third; mesinfraepisternum with two-thirds black, in line with black of humeral stripe, lower third of mesinfraepisternum ochre and washed-out yellow-orange; venter of pterothorax pale yellowish. Coxae pale yellowish-ochre; outer side of femora with a washed-out blackish stripe along posterior half; tibiae with blackish along the black spines; tip of tarsi and tarsal claws ochre, washed-out blackish along joints. + + +Wings +( +Fig. 49 +): Membrane hyaline, slightly tinged yellowish; veins brown to black; 15.5–16.5 px in forewing; 14.0–14.5 px in hindwings; Pt dark reddish-brown, overlying one cell. + + +Abdomen +( +Fig. 49 +, also compare +Figs 122a, b +): S1 orange-reddish, with black along its posterior margin; S2 orange-reddish, dorsally with an anvil-like black marking in posterior third, and a washed-out blackish middorsal stripe; S3–8 dorsally black, the black colour expanded to lateral at each posterior end; laterally and each basal annulus pale yellowish; S9 with its anterior half laterally and dorsally orange-reddish, separated by a black middorsal stripe which merges with the black posterior half; the anterior margin of the black colour somewhat serrated; S10 black ( +Figs 50, 51 +). + + +Caudal appendages +( +Figs 50–52 +, also compare +Figs 139a–c +): Cercus laterally washed-out brownish, slightly shorter than S10, inferior lobe appears slightly shorter than superior lobe; in lateral view cercus subrectangular, apex with a broad inter-lobe notch; in dorsal view cercus appears narrow; apex of superior lobe of the cercus slightly pointed and curved inwards, a subtriangular interior projection is situated just proximal of the middle of the cercus; a small, tooth-like subbasal interior projection before half-length of the inferior lobe of the cercus, visible in dorsal and dorsolateral views; paraproct pale yellow-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; with prominent black tooth at blackish medial edge. + +Measurements (mm): Total length (with appendages) 41.2; abdomen length (without appendages) 34.3; hindwing 23.7; forewing 24.6; Pt in forewing 0.9; cercus 0.6. + + +FIGURES 49–54 +. + +Pseudagrion nigrihumerale + +, holotype male: ( +49) +habitus lateral, ( +50 +) appendages lateral, ( +51 +) appendages dorsal, ( +52 +) cercus dorsolateral, ( +53 +) head dorsal, ( +54 +) pronotum. Photos: M. Seehausen. + + + + +Variation in +paratype +males + + + +Head +: The reddish occipital bar is reduced in some specimens. + + +Wings +: 14.0–16.5 px in forewing, 12.5–14.5 px in hindwing. + + +Abdomen +: The middorsal marking on S9 does not reach S8, but middle of S +9 in +some specimens. + +Measurements (mm): Total length (with appendages) 37.8–43.1; abdomen length (without appendages) 31.4– 35.5; hindwing 21.0–24.0; forewing 22.2–25.3; Pt in forewing 0.7–0.9; cercus 0.6–0.7. + + +Description of female +paratypes + + + +Head +( +Fig. 56 +): Labium, mandibles, genae, labrum, anteclypeus, postclypeus, antefrons, and postfrons yellow-orange; postfrons dorsally with a small brownish dot in centre; yellow-orange colour of postfrons sharply separated from vertex at level of anterior margin of lateral ocelli; vertex black; median ocellus with a black spot posteriorly; postocular spots yellow-orange, bordered by black; small orange occipital bar; rear of head black, but yellowish along the eye margins; base and scape of antennae yellow, following segments ochre-brownish. + + +Thorax +( +Figs 55, 57 +): Anterior lobe of pronotum yellowish, expanding in centre subtriangular towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large yellowish patch in the middle, and on either side a yellowish spot dorsolaterally, fusing laterally with yellowish of the anterior lobe; propleuron black above, yellowish below and to rear; posterior lobe yellowish; apical edge of posterior lobe convex; bearing two yellowish horns directed towards head, approximately reaching the centre of the middle lobe. Pterothorax orange with posterior margins black; a broad black middorsal stripe; a broad black humeral stripe, narrowed posteriorly; metapleural suture with a small elongated black patch at posterior third; mesinfraepisternum with two-thirds washed-out brownish-black, in line with the black of humeral stripe, lower third of mesinfraepisternum yellowish-ochre; venter yellowish, may be slightly covered with whitish pruinosity. Coxae and legs pale yellowish-ochre; outer side of femora with a washed-out blackish stripe along posterior half; tibiae with blackish along the black spines; tip of tarsi and tarsal claws ochre, washed-out blackish along waists. + + +Wings +( +Fig. 55 +): Membrane hyaline; veins brown to black; 15.5–16.5 px in forewing; 14.0–15.5 px in hindwings; Pt brownish, overlying one cell. + + +Abdomen +( +Figs 55, 58 +): S1–2 orange with black markings dorsally; S3–8 dorsally black, laterally and each basal annulus pale yellowish; S9 dorsally black, laterally yellow-orange, this colour expanded dorsally in centre of segment, leaving anterior and posterior margin black; S10 black. Cercus in lateral view straight and tapered, washed-out brownish-black, half as long as S10; paraproct yellow-brownish, short, in lateral view rounded; ovipositor yellowish, almost reaching to the posterior end of S10, two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Measurements (mm): Total length (with appendages) 41.3–41.9; abdomen length (without appendages) 34.2– 34.5; hindwing 24.0–24.8; forewing 25.4–26.4; Pt in forewing 0.9. + + + +Diagnosis + + +Both sexes of + +P. nigrihumerale + +are separated from all adjacent species and also most congeners in this group by the obvious thorax marking, bearing a broad black humeral stripe ( +Figs 49 +, +55 +, +105 +). Only three species, namely the Sulawesian + +P. crocops + +, + +P. halmaherae + +from Halmahera, and + +P. buenafei + +from Mindanao, +Philippines +have a similar broad black humeral marking. Apart from their distant range and the morphological differences in the caudal appendages (compare +Figs 131a–c, 133a–c, 137a–c, 139a–c +), the males of the latter three species differ from + +P. nigrihumerale + +in their colour pattern as well. Beside the name-giving black humeral marking, + +P. nigrihumerale + +can be separated from its Andaman congener + +P. igneum + +by having S1–2 dorsally red and S10 entirely black ( +Figs 43 +, +49 +, +121a, b, 122a, b +), by having a black vertex and by lacking red colour on the postfrons ( +Figs 47 +, +53 +, +87, 88 +), whereas the male appendages of both species are identical ( +Figs 138a–c, 139a–c +). The southerly distributed species + +P. simalurum + +and + +P. obscurum + +differ by the same colour markings as listed for + +P. igneum + +, but also very slightly in the male appendages: the cercus of + +P. nigrihumerale + +bears a small subbasal tooth situated at the proximal third of the inferior lobe, whereas in + +P. simalurum + +and + +P. obscurum + +there is a sharp edged bulge that only looks like a tooth in dorsal view ( +Figs 140a–c, 141a–c +). + + + + +Distribution + + +Apparently endemic to the +Nicobar Islands +, +India +( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F74FFFD3FF66DFEAFA2F9DE0.xml b/data/0E/57/D1/0E57D167F74FFFD3FF66DFEAFA2F9DE0.xml new file mode 100644 index 00000000000..da8de9a7189 --- /dev/null +++ b/data/0E/57/D1/0E57D167F74FFFD3FF66DFEAFA2F9DE0.xml @@ -0,0 +1,442 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion obscurum +Lieftinck, 1936 + +stat. nov. + + + + +Figures: 59–60 ( +syntype +♂ +, envelope); 89a, b (head +♂ +); 106 (pterothorax +♂ +); 123a, b (abdomen +♂ +); 140a–c (caudal appendages +♂ +). + + + + + + +Pseudagrion pilidorsum obscurum + +: + +Lieftinck (1936) +: 128 + +, 131–132, +Fig. 12b +(key; description +♂ +♀ +; +♂ +appendages; Nias & probably Simalur). + + + + + + +Name-bearing +type +specimens examined + +( +2♂♂ +) + + + +2♂♂ +syntypes +, + +10.ix.1931 + +, Amadraja, +Nias Island +, +Indonesia +, +J.C. +v. d. +M. Mohr +leg., +RMNH +, JvT no. 2839. +Lieftinck (1971: 103) +designated a +lectotype +male, but there are + +two males +in the respective envelope (Figs + + +59–60). This +lectotype +designation is not valid because he has not explicitly designated a single specimen (ICZN + + +74.5). We do not consider it necessary to designate a +lectotype +, thus we propose to keep them as +syntypes +. + + + +FIGURES 59–60 +. + +Pseudagrion obscurum + +, syntype males: ( +59 +) habitus lateral, ( +60 +) envelope. Photos: RMNH. + + + +Other specimens examined +( +2♂♂ +, +1♀ +) + + + + +Indonesia +, +Nias Island + +: +2♂♂ +, +1♀ +, + +21.iv.–2.v.1972 + +, surroundings of +Hilisimaetanoe +, + +200 m +a.s.l. + +, +Roesler +& +Küppers +leg., +ZFMK +, no. 2009/1142.1–1142.3 + +. + + +Characterization of male + + +Head +( +Fig. 89a, b +): Labrum, clypeus, antefrons and postfrons yellow-orange; postclypeus with two elongated black spots, and a black centre along posterior edge; centre of frons may have a small darkish spot; in +one specimen +the postclypeus is completely black ( +Fig. 89b +); yellow-orange colour of the postfrons is separated from black and red colour of the vertex at level of anterior margin of lateral ocelli; a black spot at posterior margin of median ocellus; a small reddish spot is situated in centre of vertex around level of posterior margin of lateral ocelli; occipital bar reddish; postocular spots washed-out dark reddish, comparatively small due to obscurely blackish margins; broadly surrounded by black; rear of head black, but may be yellow along the eye margins; base of antennae yellow, scape of antennae anteriorly yellow, but posteriorly black, following segments brownish. + + +Thorax +( +Figs 59 +, +106 +): Anterior lobe of pronotum yellow, this colour expanding in centre towards middle lobe; margins of anterior lobe black; middle pronotal lobe black, with a centrally divided red patch in the middle, and on either side a red patch dorsolaterally; propleuron black above, yellowish below and to rear; posterior lobe yellowreddish, with its margins black; apical edge of posterior lobe convex. Pterothorax red; middorsal carina black; dorsum obscurely blackish, almost reaching to humeral suture; humeral suture with an elongated black patch at its posterior quarter, +one male +has also the anterior humeral suture thin black; interpleural suture with a black stripe at posterior quarter, and a black spot anteriorly; a black patch at posterior quarter of metapleural suture; a large black marking covering at least superior two-thirds of the mesinfraepisternum; venter pale, may be covered with whitish pruinosity. Legs pale, with outside of the femora obscurely blackish; tibiae blackish along spines; spines black. + + +Abdomen +( +Figs 59 +, +123a, b +): S1–2 dorsally black, laterally red; S3–8 dorsally black, laterally and each basal annulus yellow-orange; S9 red, with its posterior third to half black; S10 dorsally with black “x”-like marking, variable in shape; laterally red. + + +Caudal appendages +( +Figs 140a–c +): Cercus in lateral view darkish, shorter than S10, inferior lobe appears very slightly shorter than superior lobe; in lateral view cercus subrectangular with a shallow inter-lobe notch situated low on the apex; in dorsal view the terminus of the superior lobe of the cercus curved inwards; a small interior projection is situated around the centre of cercus; a small sharp-edged bulge-like interior projection before half-length on the inferior lobe of the cercus, visible in dorsal and dorsolateral views; paraproct pale brownish-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; bearing a small black tooth at medial edge. + + +Characterization of female + + +Head +: With dark markings as in male, but primary colour orange-ochre. + + +Thorax +: Pronotum markings as in male, but primary colour yellow-orange; posterior lobe of pronotum with two black horns directed towards head, almost reaching the centre of the middle lobe. Pterothorax entirely reddish; humeral suture with an elongated black patch at posterior quarter; black markings at interpleural and metapleural suture narrow. + + +Abdomen +: S1–10 dorsally black, laterally pale yellow-ochre; S9 pale yellow-ochre laterally, this expanded dorsally in centre; S10 with pale yellowish-ochre middorsal marking. Cercus blackish; paraproct brownish, in lateral view rounded; ovipositor pale yellowish-ochre, not reaching posterior margin of S10; two brownish styles reaching to beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 12.5–14.5 px in forewing, and 11.5–13 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 37.8–44.3; abdomen length (without appendages) 32.0–36.0; hindwing 21.8–24.5; forewing 22.9–27.5; Pt in forewing 0.8–1.0; male cercus 0.5–0.6. + + + +Remarks + + +The single female has an entirely reddish pterothorax and lacks the name-giving obscurely blackish dorsum, which is present in all +four males +examined. It is not clear if this obscuring is just due to older age in the males or gender-related. + + + + +Diagnosis + + + +Pseudagrion obscurum + +belongs to the group of endemic taxa of the island chain west of +Sumatra +. Due to scarcity of material we cannot give information on how to separate the females, but the males are separated from most congeners occurring on adjacent islands by having S1–2 dorsally black ( +Figs 59 +, +123a, b +). However this character is shared by + +P. igneum + +( +Figs 121a, b +) and + +P. simalurum + +( +Figs 124a, b +)—both are very similar to + +P. obscurum + +also in the colour of the head ( +Figs 87, 89a, b, 90 +), and the morphology of the male cercus ( +Figs 138a–c +, +140a–c, 141a–c +). We summarise a combination of characters for separating these three species in table 2. + + + + +TABLE 2. +Compilation of characters separating males of + +P. igneum + +, + +P. obscurum + +, and + +P. simalurum + +. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
+ +P. igneum + + + +P. obscurum + + + +P. simalurum + +
+Vertex +red, largely black alongindistinct red, may be hardlyred, less black along ocelli
ocelli (Fig. 87)visible, largely black along(Fig. 40)
ocelli (Fig. 89a, b)
+Postocular spot +red (Fig. 87)indistinct red, may be hardlyred (Fig. 40)
visible (Fig. 89a)
+Pterothorax +red, not obscurely blackishred, but dorsum obscurelyred, not obscurely blackish
(Fig. 104)blackish to different extent(Fig. 107)
(Fig. 106)
+Cercus +inferior lobe with a smallinferior lobe with a smallinferior lobe with a small
tooth-like interior projectionsharp edged bulge-likesharp edged bulge-like
before half-length, bestinterior projection beforeinterior projection before
visible in dorsolateral viewhalf-length, best visible inhalf-length, best visible in
(Fig. 138c)dorsolateral view (Fig. 140c)dorsolateral view (Fig. 141c)
+ + + +Distribution + + +This species is apparently endemic to Nias Island ( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F757FFFBFF66D9DCF8C998D4.xml b/data/0E/57/D1/0E57D167F757FFFBFF66D9DCF8C998D4.xml new file mode 100644 index 00000000000..5896257a962 --- /dev/null +++ b/data/0E/57/D1/0E57D167F757FFFBFF66D9DCF8C998D4.xml @@ -0,0 +1,498 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion acutidens + +sp. nov. + + + + +Figures: 5 (habitus +holotype +♂ +); 6–8 (S9–10, caudal appendages +holotype +♂ +); 9 (head +holotype +♂ +); 10 (pronotum +holotype +♂ +); 11 (habitus +paratype +♀ +); 12 (head +paratype +♀ +); 13 (pronotum +paratype +♀ +); 14 (S9–10, ovipositor +paratype +♀ +); 79 (head +♂ +); 96 (pterothorax +♂ +); 113a, b (abdomen +♂ +); 130a–c (caudal appendages +♂ +); 147–151 (field photos +♂ +). + + + + + + +Holotype +: + +♂ +, acq. + +v.1933 + +, +Tobelo +, +Halmahera Island +, +North Maluku +, +Indonesia +, approximately +1.724°N +, +127.997°E +, +van Diejen +leg., no. 1519102. +Deposited +at +Naturalis Biodiversity Center Leiden +, the Netherlands. + + + + +Paratypes + +( +4♂♂ +, +2♀♀ +): + +2♂♂ +, +1♀ +, acq. + +v.1933 + +, +Tobelo +, +Halmahera Island +, +North Maluku +, +Indonesia +, approximately +1.724°N +, +127.997°E +, +van Diejen +leg., +RMNH +, no. 1519101, 1519102 + +;— + +1♂ +, +1♀ +, + +8.vii.1985 + +, +Wayaua +, rivulet, edge of logged forest, +Bacan Island +, +North Maluku +, +Indonesia +, approximately - +0.741°N +, +127.641°E +, +F.G. Rozendaal +leg., +RMNH +, no. 1519079 + +;— + +1♂ +, + +1.viii.1997 + +, primary rainforest +9 km +of logging company camp +Chucumara +, +Morotai Island +, +North Maluku +, +Indonesia +, alt. + +400–480 m + +, +R. Verovnik +leg, +MB + +. + + + + +Etymology: +The name refers to the inferior lobe of the male cercus, which has a large tooth-like medially pointed projection, clearly visible in dorsal view. + + + + +Remarks +: Further specimens from the islands of Halmahera, Bacan, Buru, and Obi were discovered among unidentified material at the RMNH by V.J. Kalkman just prior to submission of the manuscript. For this reason we have not included them in the +type +series. + + + + + +Description of +holotype +male + + + +Head +( +Fig. 9 +, also compare +Fig. 79 +): Labium, mandibles, genae, labrum, anteclypeus and postclypeus yellow; antefrons and postfrons yellow, with posterior margin of postfrons washed out reddish, sharply separated from black vertex around level of anterior margin of lateral ocelli; a yellow-orange centre marking at vertex expands towards occipital bar; median ocellus with black spot posteriorly; occipital bar reduced to a small orange-reddish stripe in centre; triangular postocular spots orange-reddish, bordered by black; rear of head black, but pale along the eye margins; base and scape of antennae yellow, pedicel ochre-brownish, rest of antennae lacking. + + +Thorax +( +Figs 5, 10 +, also compare +Fig. 96 +): Anterior lobe of pronotum yellowish, with anterior margin black; the yellow colour projecting in centre as semi-circle towards middle lobe; middle pronotal lobe otherwise black, with a centrally divided yellow-orange patch in the middle, and on each side a large yellow-orange patch dorsolaterally; propleuron black above, yellow-orange below and to rear; posterior lobe yellow-orange; apical edge of posterior lobe convex. Pterothorax orange-reddish; middorsal carina black; moderately broad black humeral stripe, slightly widened at posterior third before peaked towards posterior margin of pterothorax; interpleural suture black with a black spot anteriorly; posterior half of metapleural suture black; mesinfraepisternum with black diagonal marking, continuing the black humeral stripe; venter pale, covered with whitish pruinosity. Coxae and legs pale orange-ochre; outer side of femur with a washed-out blackish stripe, covering almost complete femur in foreleg, almost complete posterior half of femur in midleg, and posterior quarter of femur in hind leg; tibiae blackish along the black spines; tarsi and tarsal claws orange-ochre, washed-out blackish along waists. + + +Wings +( +Fig. 5 +): Membrane hyaline, slightly tinged yellowish-brown; veins brown to black; 16.0–16.5 px in forewing; 14.5–15.0 px in hindwing; Pt reddish-brown, overlying one cell. + + +Abdomen +( +Figs 5–7 +, also compare +Figs 113a, b +): S1 orange-reddish, dorsally with a black quadrangular patch covering anterior two-thirds of segment, posterior margin black; S2 orange-reddish, dorsally with acute mushroom-like black marking at posterior third, and triangular black patch at anterior margin; both black markings almost connected by thin black middorsal stripe; S3–8 dorsally black, expanded to lateral half of segment; pale yellowish basal annulus; laterally and ventrally pale yellowish-orange; S9 with anterior half laterally and dorsally orange-reddish; posterior half of S9 black, with its anterior margin somewhat serrated with incomplete middorsal stripe; S10 orange-reddish, dorsally with broad black “x”-like marking; black colour of anterior and posterior margin expanded laterally. + + + +FIGURES 5–10 +. + +Pseudagrion acutidens + +, holotype male: ( +5) +habitus lateral, ( +6 +) appendages lateral, ( +7 +) appendages dorsal, ( +8 +) cercus dorsolateral, ( +9 +) head dorsal, ( +10 +) pronotum. Photos: M. Seehausen. + + + +Caudal appendages +( +Figs 6–8 +, also compare +Figs 130a–c +): Cercus laterally washed-out brownish, slightly shorter than S10; in lateral view cercus subrectangular, with inter-lobe notch centrally placed at the apex; in dorsal view apex of superior lobe of cercus rounded and curved in; inferior lobe expanded inwards, subtriangular, apex of expanded part blackish and tooth-like, curved towards body; paraproct blackish at base, apex orange-brownish; slightly longer than half of cercus; viewed laterally subtriangular and tapered, with apex rounded; viewed dorsally paraproct broadly rounded, concave bowl-like, with blackish tubercular medial edge. + +Measurements (mm): Total length (with appendages) 43.2; abdomen length (without appendages) 36.1; hindwing 23.6; forewing 25.2; Pt in forewing 0.8; cercus 0.5. + + +Variation in +paratype +males + + + +Head +: All males have a black spot at centre of posterior margin of postclypeus (as in +Figs 79 +, +148 +; not visible in +Fig. 9 +of the +holotype +because of dirt); the occipital bar is completely black in +two males +. + + +Thorax +: The males from Morotai and Bacan have the metapleural suture almost completely covered by a black stripe. + + +Wings: +14.5–17.5 px in forewing; 13.5–15.0 px in hindwing. + + +Abdomen +: +Two males +have the triangular black patch at anterior margin of S2 reduced; the male from Morotai has a complete thin black middorsal stripe on S2 (as in +Fig. 147 +); the male from Bacan has S8 dorsally black, but with two elongate triangular reddish patches at anterior third, each situated dorsolaterally, and about as long as one-third of S8; +one male +has S9 with a black dot each lateral of the incomplete middorsal stripe. + +Measurements (mm): Total length (with appendages) 42.3–45.3; abdomen length (without appendages) 35.7– 38.0; hindwing 23.3–24.2; forewing 25.0–25.6; Pt in forewing 0.8–0.9; cercus 0.5. + + +Description of female +paratypes + + + +Head +( +Fig. 12 +): Labium, mandibles, genae, labrum, anteclypeus and postclypeus yellow-orange; postclypeus with at least a blackish dot at centre of posterior margin, and an elongated blackish spot at each side, but may likewise be almost completely dark; antefrons and postfrons yellow-orange, with a black elongated patch in centre; yellow-orange colour of postfrons sharply separated from black vertex around level of lateral ocelli; median ocellus with a black spot posteriorly; lateral ocelli largely surrounded by black, less distinct or lacking at their anterior margins; triangular postocular spots orange-reddish, broadly bordered by black; rear of head black, but pale along the eye margins; base and scape of antennae orange-ochre, following segments brownish. + + +Thorax +( +Figs 11, 13 +): Pronotum with anterior lobe pale yellowish, its anterior margin black; middle pronotal lobe black with a centrally divided yellowish patch in the middle, and on either side a large yellowish patch dorsolaterally; propleuron black above, pale yellowish below and to rear; posterior lobe of pronotum yellow-brownish; apical edge of posterior lobe convex, bearing two dark horns directed towards head, approximately reaching the centre of the middle lobe. Pterothorax orange; middorsal carina black; moderately broad black humeral stripe, slightly widened in posterior third before peaking towards posterior margin of pterothorax; interpleural suture darkened with blackish spot anteriorly; metapleural suture at least with black elongated patch posteriorly, but may be almost completely blackish; mesinfraepisternum with black diagonal marking, continuing the black humeral stripe; venter of pterothorax pale yellowish, may be covered with whitish pruinosity. Coxae and legs pale yellow-ochre; outer side of femur with a washed-out blackish stripe along posterior two-thirds in foreleg, reduced to posterior half in midleg, and posterior quarter in hindleg; tibiae blackish along the black spines; tarsi and tarsal claws yellow-ochre, washed-out blackish along waists. + + +Wings +( +Fig. 11 +): Membrane hyaline, may be slightly tinged yellowish-brown; veins brown to black; 17.5–18.0 px in forewing; 14.5–16.5 px in hindwings; Pt ochre, overlying one cell. + + +Abdomen +( +Figs 11, 14 +): S1 yellow-orange, dorsum black; S2 yellow-orange, with black dorsal marking which is expanded laterally at posterior third, and may also be widened at anterior margin; S3–8 dorsally black, expanded to lateral half of segment, pale yellowish basal annulus; laterally and ventrally pale yellowish; S9–10 dorsally black, expanded to lateral along posterior and anterior margin; laterally and ventrally pale yellowish. Cercus dark, as long as half of S10; paraproct dark, rounded laterally; ovipositor pale yellowish, its apex not reaching to posterior end of S10; bearing two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Measurements (mm): Total length (with appendages) 41.6–42.0; abdomen length (without appendages) 33.8– 34.5; hindwing 25.1–25.3; forewing 26.6–26.8; Pt in forewing 0.8–1.0. + + + +FIGURES 11–14 +. + +Pseudagrion acutidens + +, female paratype: ( +11 +) habitus lateral, ( +12 +) head dorsal, ( +13 +) pronotum, ( +14 +) ovipositor lateral. Photos: M. Seehausen. + + + + +Diagnosis + + + +Pseudagrion acutidens + +overlaps in range on Halmahera with + +P. halmaherae + +described below, and on Buru with + +P. crocops + +and + +P. ustum + +. However, males of + +P. acutidens + +can easily be separated from all congeners west of Lydekker’s Line by the large medially projecting tooth on the cercus ( +Figs 7–8 +, +130b, c +, +149 +). + +Pseudagrion ustum + +has a tooth of similar size on the cercus, but in that species the tooth is projecting upward ( +Figs 146a–c +). Furthermore + +P. halmaherae + +has a broad humeral stripe in both sexes ( +Figs 33 +, +39 +, +103 +), whereas the humeral stripe is narrower in + +P. acutidens + +( +Figs 5 +, +11 +, +96 +, +147–148, 150–151 +). Furthermore both species differ in both sexes in the shape of the postocular spots, which are more rectangular and narrower, with a somewhat serrated posterior margin in + +P. halmaherae + +(compare +Figs 9 +, +12 +, +37 +, +40 +, +79, 86 +). + + + + +Distribution + + +This species is restricted to the northern +Maluku Islands +. Records are available from Morotai, Halmahera, Obi, Bacan, and Buru ( +Fig. 3 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F75AFFFDFF66DC09FBD39E68.xml b/data/0E/57/D1/0E57D167F75AFFFDFF66DC09FBD39E68.xml new file mode 100644 index 00000000000..11605fc22ee --- /dev/null +++ b/data/0E/57/D1/0E57D167F75AFFFDFF66DC09FBD39E68.xml @@ -0,0 +1,436 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion buenafei +Müller, 1996 + + + + + +Figures: 15–16 ( +holotype +♂ +, envelope); 80 (head +♂ +); 97 (pterothorax +♂ +); 114a, b (abdomen +♂ +); 131a–c (caudal appendages +♂ +). + + + + + + +Pseudagrion buenafei +: + + +Müller (1996) +: 377 + +–379, +Figs 1–3 +(description +♂ +♀ +; +♂ +appendages; Mindanao, +Philippines +);— + +Hämäläinen & Müller (1997) +: 261 + +(Mindanao). + + + + + + +Examined specimens from the +type +series + +( +4♂♂ +) + + + +♂ +holotype +, + +x.1991 + +, +Atongan River +, +Katanglad Mountains +, +Bukidnon +, +Mindanao +, +Philippines +, +Alex Buenafe +leg., ex. +Coll. Roland A. Müller +, +RMNH +, JvT no. 17838 ( +Figs 15–16 +) + +;— + +1♂ +paratype +, 24./ + +25.xi.1991 + +, +Muntian creek +, +Mt. Kalatungan +, +Olayan +, +Mabadiang +, +Bukidnon +, +Mindanao +, +Philippines +, + +1200–1400 m +a.s.l. + +, +Alex Buenafe +leg., ex. +Coll. Roland A. Müller +, +RMNH +, no. 1506118 + +;— + +1♂ +paratype +, + +ix.1995 + +, +Babang creek +, +Lake Agko +, +Ilomavis +, +Mt. Apo +, +North Cotabato +, +Mindanao +, +Philippines +, + +1600–1700 m +a.s.l. + +, +Alex Buenafe +leg., ex. +Coll. Roland A. Müller +, +RMNH +, no. 1506120 + +;— + +1♂ +paratype +, + +24.xi.1995 + +, +Mansabilan creek +, +Mt. Kalatungan +, +Dumatap +, +Mebadiang +, +Bukidnon +, +Mindanao +, +Philippines +, + +1000–1300 m +a.s.l. + +, +Alex Buenafe +leg., ex. +Coll. Roland A. Müller +, +RMNH +, no. 1506121 + +. + + +Müller (1996) +stated that the +holotype +should be deposited at +SMF +, but +Hämäläinen (2017) +corrected that statement: the +holotype +was deposited at +RMNH +. + + + +FIGURES 15–16 +. + +Pseudagrion buenafei + +, holotype male: ( +15 +) habitus lateral, ( +16 +) envelope. Photos: RMNH. + + + +Other specimens examined +( +1♂ +) + + + + +Philippines +, Mindanao Island + +: +1♂ +, + +x.1991 + +, +Atongan River +, +Katanglad Mountains +, +Bukidnon +, +Alex Buenafe +leg., ex. +Coll. Roland A. Müller +, +RMNH +, no. 1506113 + +. + + +Characterization of male + + +Head +( +Figs 15 +, +80 +): Labrum reddish brown to obscure blackish; anteclypeus red; postclypeus black; antefrons black, with anterior margin and laterally red; postfrons and vertex black; postocular spots and occipital bar red; rear of head black, but a pale patch along the centre of the eye margins; base and scape of antennae black, following segments brownish. + + +Thorax +( +Figs 15 +, +97 +): Pronotum with anterior lobe black, its posterior centre reddish, extending towards middle lobe; middle pronotal lobe otherwise black, with a centrally divided reddish rectangular patch in the middle, and on each side a large reddish patch dorsolaterally; propleuron black above, broadly reddish below and to rear; posterior lobe of pronotum blackish, with reddish centre; apical edge of posterior lobe convex. Pterothorax dorsally black, with antehumeral stripe red (may be interrupted); laterally red, with a black stripe each at posterior quarter of interpleural and metapleural suture, interpleural suture with a black spot anterior of the black stripe; ventrally pale, may be covered with whitish pruinosity. Legs blackish, with coxae and trochanters reddish. + + +Abdomen +( +Figs 15 +, +114a, b +): S1 red; S2 red with a small black triangle marking dorsally at posterior end; S3–7 dorsally black, laterally reddish; S8–10 red with black apical ring. + + +Caudal appendages +( +Figs 131a–c +): Cercus blackish; slightly shorter than S10; in lateral view subrectangular, inter-lobe notch placed low on apex; in dorsal view cercus narrow, with apex of superior lobe of cercus curved inward (more clearly so in dorsolateral view); an inward directed projection is situated at just after half-length on the superior lobe of the cercus; a small subbasal interior projection on inferior lobe of cercus, visible in dorsal and dorsolateral views, this bulge-like projection bearing a sharp proximal edge; paraproct reddish, around half as long as cercus, in lateral view somewhat triangular and tapered, with rounded apex; in dorsal view broadly rounded, concave bowl-like, with small black tooth at medial edge. + +Wings with 14.5–17.5 px in forewing, and 14.5–15.5 px in hindwing. +Measurements (mm): Total length (with appendages) 46.3–49.0; abdomen length (without appendages) 37.5– 39.4; hindwing 26.4–30.0; forewing 27.7–30.0; Pt in forewing 0.7–0.8; male cercus 0.7. + +Characterization of female +[not examined, characterization adopted from +Müller (1996) +] + + +Head: +As in male, but with a black triangular mark at frons. + + +Thorax +: Pronotum and pterothorax as in male, but red replaced by greenish-yellow; legs as in the male. + + +Abdomen: +S1 reddish-brown; S2 reddish-brown, with narrow black middorsal marking, widened apically; S3– 10 dorsally black, laterally reddish-brown. + + + + +Diagnosis + + +Both sexes of this species are separated from congeneric species, and especially the co-occurring + +P. pilidorsum + +, by the morphology of the male cercus (compare +Figs 131a–c +, +143a–c +), as well as by the combination of a largely black frons and vertex (compare +Figs 80, 92 +), a black humeral stripe (compare +Figs 97, 109 +), and blackish legs (compare +Figs 15 +, +63, 64 +, +165 +). + +Pseudagrion buenafei + +is also larger (total length +46–49 mm +) than + +P. pilidorsum + +( +38–46 mm +). Additionally, the male is separated from all congeners by the almost entirely red abdominal segment S8 ( +Figs 15 +, +114a, b +). + + + + +Distribution + + +Endemic to Mindanao, +Philippines +( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F75CFFFFFF66D994FA529E0C.xml b/data/0E/57/D1/0E57D167F75CFFFFFF66D994FA529E0C.xml new file mode 100644 index 00000000000..3ab5c2ea9c3 --- /dev/null +++ b/data/0E/57/D1/0E57D167F75CFFFFFF66D994FA529E0C.xml @@ -0,0 +1,420 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion coriaceum +Selys, 1876 + + + + + +Figures: 17–19 ( +lectotype +♂ +, labels); 81 (head +♂ +); 98 (pterothorax +♂ +); 115a, b (abdomen +♂ +); 132a–c (caudal appendages +♂ +); 152–153 (field photos +♂ +). + + + + + + +Pseudagrion coriaceum + +: + +Selys (1876) +: 513 + +(description +♂ +, +Ambon +);— + +Ris (1915b) +: 97 + +–98, +Fig. 19 +( +♂ +appendages; key; Seram);— + +Campion (1921) +: 263 + +( +Ambon +);— + +Ris (1929) +: 146 + +(description +♀ +; +Ambon +). + + + + + + +Examined specimens from the +type +series + +( +2♂♂ +) + + + +♂ +lectotype +, 1859, Amboina [ +Ambon +, +Indonesia +], +Dr Doleschal +leg., +NHMW +( +Figs 17–19 +) + +;— + +1♂ +paralectotype +, 1859, Amboina [ +Ambon +, +Indonesia +], +Dr Doleschal +leg., +NHMW + +. + + +Lectotype +designation by St. +Quentin (1970) +. Both males lack the last segments of the abdomen, including the caudal appendages. + + +Other specimens examined +( +6♂♂ +, +3♀♀ +) + + + + +Indonesia +, +Ambon Island +: + +1♀ +, without particular locality, +IRSNB + +;— + +3♂♂ +, + +23.iv.1941 + +, +Soengai +, +Waitoemoe +, +Mains +z lasoen, +RMNH + +, no. 1519109;— + +1♀ +, + +27.vii.1948 + +, without particular locality, +M.A. Lieftinck +leg., +RMNH + +, no. 1519104;— + +1♀ +, + +12.x.1949 + +, without particular locality, +M.A. Lieftinck +leg., +RMNH + +, no. 1658923. + + + + +Indonesia +, +Seram Island + +: +1♂ +, + +18.ix.1912 + +, without particular locality, +Elgner +leg., +SMF + +, no. 13924;— + +1♂ +, + +21.xi.1941 + +, +Wahai +, +van der Harre +leg., +RMNH + +, no. 1519260;— + +1♂ +, + +30.xi.1996 + +, stream + +500 m + +NE of +Logging Base Camp +, +1.5 km +SW +Roho village +, + +31 km +SW of Wahai + +, +M. Bedjanič +leg., +MB + +. + + +Characterization of male + + +Head +( +Figs 17 +, +81 +): Labrum, clypeus, antefrons and postfrons yellow; postfrons distinctly separated from black vertex around level of lateral ocelli; postocular spots large, reddish-brown, but they may be darkened and hardly visible; reddish occipital bar, but may be darkened and hardly visible; rear of head black, but broadly pale along the eye margins; eyes brownish, in life anteriorly yellow, continuing the yellow of the frons; base and scape antennae yellow, following segments brownish. + + +Thorax +( +Figs 17, 18 +, +98 +): Anterior lobe of pronotum red, this colour extending towards middle lobe in posterior centre; middle pronotal lobe otherwise black, with a centrally divided red patch in the middle, and on each side a large red patch dorsolaterally, these may fuse laterally with red of anterior lobe; propleuron black above, broadly yellowish-red below and to rear; posterior lobe of pronotum red; apical edge of posterior lobe almost convex. Pterothorax red with black middorsal stripe; a black humeral stripe of same width; interpleural suture with a black stripe, and a dark dot anteriorly; metapleural suture with a black stripe, widened posteriorly; ventrally pale. Legs orange-reddish; outer side of femora with a blackish stripe narrowing to proximal; along spines of tibiae, and the waists blackish; spines black. + + + +FIGURES 17–19 +. + +Pseudagrion coriaceum + +, lectotype male: ( +17 +) habitus dorsal, ( +18 +) pterothorax lateral, ( +19 +) labels. Photos: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner. + + + +Abdomen +( +Figs 115a, b +): S1 red, dorsally with black patch covering the anterior two-thirds of the segment, the patch may be divided in half by a red middorsal line; S2 red, but along posterior carina black; with blackish mushroom-like marking dorsally at posterior quarter, this marking may extend as a black middorsal stripe towards the segment base; S3–8 dorsally black, laterally pale yellow-red; S9 red, dorsally with a black middorsal stripe and with at least the posterior third to half black, the black margin may be serrated, but dorsum of S9 may likewise be completely black; S10 red, dorsally with a black “x”-like marking, variable in exact shape. + + +Caudal appendages +( +Figs 132a–c +): Cercus laterally dark, slightly shorter than S10; in lateral view cercus simple, subtriangular, inter-lobe notch not visible; in dorsal view apex of superior lobe rounded and deep inter-lobe notch visible; inferior lobe bulging smoothly inwards; paraproct pale yellow-ochre, as long as two-third of the cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like, its medial edge blackish tubercular. + + +Characterization of female + + +Head: +Labrum, and anteclypeus yellow-ochre; postclypeus blackish; antefrons and postfrons yellow-ochre, antefrons with blackish patch in centre and a black mid stripe towards postclypeus; vertex black from about level of lateral ocelli; postocular spots orange-ochre; occipital bar orange-ochre; rear of head black, but broadly pale along the eye margins; base and scape of antennae yellow-ochre, following segments brownish. + + +Thorax +: Anterior lobe of pronotum orange-ochre, its anterior margin black; middle pronotal lobe and posterior lobe as in male but orange-ochre; posterior lobe bearing two horns directed towards head, almost reaching the centre of the middle lobe; apical edge of posterior lobe almost convex. Pterothorax and legs as in male but orange-ochre. + + +Abdomen +: S1 orange, with a blackish dorsal marking at anterior two-third; S2 red, with a black middorsal stripe, widened to lateral at posterior quarter; S3–10 dorsally black, laterally orange-ochre; on S9–10 the orange-ochre expanded dorsally in centre, with the anterior and posterior margin, as well as the dorsum of each segment black. Cercus blackish, as long as half of S10; paraproct darkish ochre; in lateral view rounded; ovipositor pale yellowish-ochre, its apex not reaching to posterior end of S10; two yellow-brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 13.5–15.5 px in forewing, and 12.0–13.5 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 43.2–46.6; abdomen length (without appendages) 35.2–38.5; hindwing 24.0–26.7; forewing 25.4–28.0; Pt in forewing 0.8–1.0; male cercus 0.5. + + + +Diagnosis + + + +Pseudagrion coriaceum + +is currently the only known species of the + +Pseudagrion + +red-group which occurs on +Ambon +and Seram. Males can be separated from all congeners by the distinct morphology of the cercus ( +Figs 132a– c +), being subtriangularly tapered with inferior lobe and inter-lobe notch not visible in lateral view. The only species with a superficially similar morphology of the male cercus is + +P. halmaherae + +( +Figs 137a–c +), but in this species the cercus is less tapered in lateral view and an inter-lobe notch is situated subapically at the lower margin. The obvious pterothorax pattern of + +P. coriaceum + +( +Figs 17, 18 +, +98 +, +152, 153 +) separates this species from several congeners as well. + + + + +Distribution + + +This species is restricted to +Ambon +and Seram ( +Fig. 3 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F772FFD5FF66DB1CFB449DCC.xml b/data/0E/57/D1/0E57D167F772FFD5FF66DB1CFB449DCC.xml new file mode 100644 index 00000000000..bac9a393cad --- /dev/null +++ b/data/0E/57/D1/0E57D167F772FFD5FF66DB1CFB449DCC.xml @@ -0,0 +1,427 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion perfuscatum +Lieftinck, 1937 + + + + + +Figures: 61–62 ( +holotype +♂ +, envelope); 91 (head +♂ +); 108 (pterothorax +♂ +); 125a, b (abdomen +♂ +); 142a–c (caudal appendages +♂ +); 163–164 (field photos +♂ +). + + + + + + +Pseudagrion perfuscatum + +: + +Lieftinck 1937: 94–97 + +, +Fig. 21 +(description +♂ +♀ +; +♂ +appendages; West Borneo [ +West Kalimantan +, +Indonesia +]);— + +Orr (2001) +: 186 + +( +Brunei +);— + +Malkmus (2005) +: 11 + +(Mount Kinabalu, +Sabah +);— + +Dow (2005) +: 10 + +( +Sarawak +);— + + +Dow +et al +. (2024) + +: 37 + +(Kalimantan, +Brunei +, +Sarawak +, +Sabah +). + + + + + +Pseudagrion pruinosum + +: + +Laidlaw (1915) +: 275 + +(Mt. Merinjak [ +Sarawak +, +Malaysia +]). + + + + + + +Examined specimens from the +type +series + +( +1♂ +, +1♀ +) + + + +♂ +holotype +, + +5.–16.xi.1931 + +, Singkawang, weg naar Bengkajang, + +West Borneo +, L + +. +Coomans de Ruiter +leg., +RMNH +, JvT no. 2842 ( +Figs 61–62 +) + +;— + +1♀ +paratype +, + +5.–16.xi.1931 + +, Singkawang, weg naar Bengkajang, + +West Borneo +, L + +. +Coomans de Ruiter +leg., +RMNH +, JvT no. 2842 ( +Figs 61–62 +) + +. + + + +FIGURES 61–62 +. + +Pseudagrion perfuscatum + +, lectotype male, paralectotype female: ( +61 +) habitus lateral, ( +62 +) envelope. Photos: RMNH. + + + +Other specimens examined +( +3♂♂ +) + + + + +Indonesia +, +Borneo Island + +: +1♂ +, + +1.–7.i.1925 + +, +Lebang Hara +, +Sammelreise Prof Dr H. Winkler +, +ZMH + +;— + +2♂♂ +, + +16.xi.1931 + +, +Singkawang +, + +West Borneo +, L.C. + +d.R. leg., +RMNH + +, no. 1658921, 1658922. + + +Characterization of male + + +Head +( +Fig. 91 +): Labrum, clypeus, antefrons, postfrons, and vertex orange in preserved specimens but red in living individuals; a black spot posterior of median ocellus, and black patches along lateral ocelli towards orange colour of occipital bar; postocular spots orange-red, separated by black from postfrons and vertex; rear of head black, but yellow along the eye margins, may be covered with bluish pruinosity; base and scape of antennae orange-ochre, following segments brownish. + + +Thorax +( +Figs 61 +, +108 +):Anterior lobe of pronotum reddish, its margins washed-out blackish; middle pronotal lobe black, with a centrally divided red patch in the middle, and on each side a large red patch dorsolaterally; propleuron black above, pale orange below and to rear; posterior lobe of pronotum red, with its margins washed-out blackish; apical edge of posterior lobe convex. Pterothorax reddish-brown, with a black middorsal stripe; humeral suture at least with an elongate black patch at its posterior quarter, but may likewise be completely black; interpleural suture with a black line, a black spot may be present anteriorly; metapleural suture with a black streak situated at posterior quarter; venter pale; may be covered with bluish-white pruinosity up to interpleural suture. Legs pale orange; femora with black stripe outside; tibiae with black along spines; spines black. + + +Abdomen +( +Figs 61 +, +125a, b +): S1 red, with a blackish basal spot covering two-thirds of dorsum, segment may be completely covered with bluish-white pruinosity; S2 red, with black dorsally, may be completely covered with bluish-white pruinosity; S3–8 dorsally black, laterally pale yellowish; S9 red with its posterior third to half black; S10 black. + + +Caudal appendages +( +Figs 142a–c +): Cercus laterally blackish, but may be reddish at base; in lateral view cercus shorter than S10, the upper margin slightly convex, its apex almost square, inter-lobe notch barely or not apparent in this view; viewed dorsally the apical part of the superior lobe of the cercus curved inwards and runs as a ridge to the inter-lobe notch, with a separate inward directed projection proximal of this visible in dorsolateral view; another inwardly directed projection is situated more proximally at around the middle of the cercus; the inferior lobe of the cercus with a distinct ridge, best visible in dorsolateral view; a small bulge-like interior projection at the base of this ridge is visible in dorsal and dorsolateral view; paraproct pale orange, ventrally blackish; as long as two-thirds of the cercus; in lateral view subtriangular and tapered, with rounded apex; in dorsal view broadly rounded, concave bowl-like; blackish tubercular at medial edge. + + +Characterization of female +( +Fig. 61 +) + + +Head +: As in male but orange-brown instead of reddish; black colour on vertex reduced to patches proximal of each lateral ocellus; postocular spots surrounded by black. + + +Thorax +: Pronotum as in the male, but brownish-ochre instead of reddish; horns of posterior pronotal lobe directed towards head, reaching the centre of the middle lobe. Pterothorax as in male, but yellowish-ochre instead of reddish; humeral suture with a thin black line, and an elongated black patch at posterior quarter; upper third of mesinfraepisternum black. + + +Abdomen +: S1–2 dorsally black, laterally yellowish-ochre; S3–8 dorsally black, laterally yellowish; S9–10 dorsally black, lateral half yellowish-ochre. Cercus brownish; paraproct pale, in lateral view rounded; ovipositor pale yellowish-ochre, not reaching posterior margin of S10; two brownish styles reaching to beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 11.5–14.0 px in forewing, and 10.5–13.0 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 38.5–40.7; abdomen length (without appendages) 30.0–33.4; hindwing 19.0–23.0; forewing 22.0–23.8; Pt in forewing 0.8–0.9; male cercus 0.5. + + + +Diagnosis + + +The males of + +P. perfuscatum + +have the clypeus and frons red ( +Fig. 91 +), a character which separates this species from almost all congeners except for + +P. pilidorsum + +( +Figs 64 +, +92 +). The morphology of the male caudal appendages of + +P. perfuscatum + +and + +P. pilidorsum + +is almost identical ( +Figs 142a–c, 143a–c +). But at least in fully mature males the colour pattern separates both species: + +P. pilidorsum + +has less black on vertex and along the largely red postocular spots compared to + +P. perfuscatum + +( +Figs 91, 92 +). Additionally mature males of + +P. perfuscatum + +have the dorsum of S1–2 blackish and covered with pruinosity ( +Figs 61 +, +125a, b +, +163, 164 +), the lower half of the pterothorax is covered with pruinosity as well, and usually there is a black humeral stripe ( +Figs 61 +, +108 +, +163, 164 +)—making the colour pattern almost unmistakable; whereas + +P. pilidorsum + +has the dorsum of S1–2 reddish ( +Figs 126a, b +, +165, 166 +), the pruinosity on them is restricted to venter, and at least slightly expanded laterally, the pruinosity on the pterothorax is restricted to the venter and metepimeron, and a humeral stripe is lacking ( +Figs 63, 64 +, +109 +, +165, 166 +). [Note: See discussion section for further information]. + + + + +Distribution + + +This species is endemic to Borneo, and recorded from +Brunei +, Indonesian Kalimantan, as well as from Malaysian +Sarawak +and possibly +Sabah +( +Fig. 1 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F774FFD7FF66DB70FA06989B.xml b/data/0E/57/D1/0E57D167F774FFD7FF66DB70FA06989B.xml new file mode 100644 index 00000000000..845560c518b --- /dev/null +++ b/data/0E/57/D1/0E57D167F774FFD7FF66DB70FA06989B.xml @@ -0,0 +1,1373 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion pilidorsum +Brauer, 1868 + + + + + +Figures: 63–65 ( +syntype +♂ +, labels); 92 (head +♂ +); 109 (pterothorax +♂ +); 126a, b (abdomen +♂ +); 143a–c (caudal appendages +♂ +); 165–166, [167?] (field photos +♂ +). + + + + + + +Agrion pilidorsum + +: + + +Brauer (1868) +: 553 + +–554 (description +♂ +♀ +; +Manilla +, +Luzon +) + +. + + + + + +Pseudagrion pilidorsum + +: + +Selys (1876) +: 514 + +–515 (description +♂ +; +Manilla +, Luzon);— + +Selys (1882) +: 30 + +( +Manilla +, Luzon, Mindanao);— + +Selys (1891) +: 217 + +(Sibul, Irocin, +Bulacan +);— + +Ris (1915b) +: 97 + +, 99 (key; Negros Island, Taiwan);— + +Ris (1916) +: 43 + +–44 (Taiwan);— + +Lieftinck (1936) +: 129 + +, 132, +Fig. 12a +( +♂ +appendages; key; Mindanao & +Basilan +, +Philippines +);— + +Needham & Gyger (1939) +: 281 + +–283, plate 16, Figs 221 & 222 ( +♂ +appendages; several +Philippine Islands +);— + +Asahina (1964) +: 520 + +(Ryukyu Islands, +Japan +);— + +Hämäläinen & Müller (1997) +: 261 + +(25 +Philippine islands +listed);— + +Dow (2006) +: 10 + +( +Sarawak +, +Malaysia +);— + +Dow & Singa (2022) +: 20 + +( +Sarawak +& +Sabah +, +Malaysia +; +East Kalimantan +, +Indonesia +). + + + +Name-bearing type specimen examined +( +1♂ +) +♂ +syntype +, + +Manila +, +Luzon Island +, +Philippines +, +Brauer +det., +NHMW +( +Figs 63–65 +) + +. + + + + +FIGURES 63–65 +. + +Pseudagrion pilidorsum + +, syntype male: ( +63 +) habitus lateral, ( +64 +) pterothorax lateral, head dorsal, ( +65 +) labels. Photos 63, 65: © 2024 Natural History Museum Vienna, NOaS Image Collection/H. Bruckner; photo 64: M. Seehausen. + + + + +Other specimens examined +( +99♂♂ +, +39♀♀ +) + + + + +Japan +, Ishigaki Island + +: +1♂ +, + +15.vii.1969 + +, +Ishigaki Island +, +Tora river +, +Okinawa +, +H. Kuwahara +leg., +Coll. von Rosen +, +ZSM +, no. +GvRosen +3334 + +. + + + + +Philippines +, +Basilan +Island + +: +1♀ +, + +2.xii.1914 + +, +Basilan +, +G. Boettcher +leg., +SMF +, no. 13999 + +;— + +2♂♂ +, + +2.–8.xii.1914 + +, +Basilan +, +G. Boettcher +leg., +SMF +, no. 13992, 13993 + +;— + +1♂ +, +1♀ +, + +10.xii.1914 + +, +Basilan +, +G. Boettcher +leg., +SMF +, no. 13997, 13998 + +;— + +6♂♂ +, +1♀ +, + +12.xii.1914 + +, +Basilan +, +G. Boettcher +leg., +SMF +, no. 13988–13991, 13994–13996 + +. + + + + +Philippines +, +Camiguin +Island +: + +2♂♂ +, +1♀ +, + +22.vii.1916 + +, +Camiguin, G +. Boettcher leg., +SMF +, no. 13968–13970 + +. + + + + +Philippines +, +Leyte +Island + +: +1♀ +, + +iv.1915 + +, +Burauen, G +. Boettcher leg., +SMF +, no. 13987 + +;— + +2♂♂ +, +1♀ +, + +1.–7.v.1915 + +, +Burauen, G +. Boettcher leg., +SMF +, no. 13983–13985 + +;— + +1♀ +, + +7.v.1915 + +, +Burauen, G +. Boettcher leg., +SMF +, no. 13986 + +. + + + + +Philippines +, +Luzon Island + +: +1♂ +, +Hotspring +, +ZMUK + +;— + +1♂ +, +Lamao +, +ZMUK + +;— + +1♂ +, +1♀ +, +Mount Makiling +, +Baker +leg., +SMF + +, no. 19841, 29843;— + +2♂♂ +, + +21.x.1913 + +, +Limay +, +G. Boettcher +leg., +SMF + +, no. 13948, 13949;— + +1♂ +, + +24.ii.1914 + +, +Los Banos +, +G. Boettcher +leg., +SMF + +, no. 13982;— + +2♂♂ +, + +16.v.1914 + +, +Trinidad +, +G. Boettcher +leg., +SMF + +, no. 13953, 13954;— + +1♂ +, + +18.v.1914 + +, +Trinidad +, +G. Boettcher +leg., +SMF + +, no. 13960;— + +1♂ +, + +21.vii.1914 + +, +Los Banos +, +G. Boettcher +leg., +SMF + +, no. 13981;— + +1♂ +, + +23.vii.1914 + +, +Los Banos +, +G. Boettcher +leg., +SMF + +, no. 13980;— + +1♂ +, + +11.viii.1914 + +, +Mount Banahao +, +G. Boettcher +leg., +SMF + +, no. 13977;— + +1♂ +, +1♀ +, + +12.viii.1914 + +, +Mount Banahao +, +G. Boettcher +leg., +SMF + +, no. 13978, 13979;— + +1♀ +, + +30.viii.1915 + +, +Kasiguran +, +G. Boettcher +leg., +SMF + +, no. 13976;— + +1♂ +, + +24.v.1916 + +, +Imugan +, +G. Boettcher +leg., +SMF + +, no. 13975;— + +1♂ +, + +ix.1917 + +, +Mount Bulusan +, +G. Boettcher +leg., +SMF + +, no. 13967;— + +5♂♂ +, + +21.iii.1978 + +, +River +, hidden valley +N Alaminos +, + +120 m +a.s.l. + +, +G. von Rosen +leg., +ZSM + +, no. 5523–5527. + + + + +Philippines +, +Marinduque +Island + +: +14♂♂ +, +2♀♀ +, + +i.–ii.1997 + +, +Batingon river +, near +Buenavista +, + +150–200 m +a.s.l. + +, +Coll. von Rosen +, +ZSM + +, no. GvRosen 12562, 12565–12575, 12577, 12578, 15263–15264;— + +1♂ +, + +iv.1997 + +, +Mandiling +, +Buenavista +, +SMF + +. + + + + +Philippines +, +Masbate +Island + +: +1♂ +, + +30.viii.1916 + +, +Aroroy, G +. Boettcher leg., +SMF +, no. 13950 + +. + + + + +Philippines +, +Mindanao Island + +: +1♀ +, +Surigao +, +Baker +leg., +SMF +, no. 29842 + +;— + +1♂ +, +1♀ +, + +6.ii.1915 + +, +Dansalan +[Marawi], +G. Boettcher +leg., +SMF +, no. 14005, 14006 + +;— + +2♂♂ +, +2♀♀ +, + +7.ii.1915 + +, +Dansalan +[Marawi], +G. Boettcher +leg., +SMF +, no. 14003, 14004, 14012, 14013 + +;— + +1♂ +, +1♀ +, + +8.ii.1915 + +, +Dansalan +[Marawi], +G. Boettcher +leg., +SMF +, no. 14007, 14008 + +;— + +3♂♂ +, + +8.–10.ii.1915 + +, +Dansalan +[Marawi], +G. Boettcher +leg., +SMF +, no. 14000–14002 + +;— + +3♀♀ +, + +10.ii.1915 + +, +Dansalan +[Marawi], +G. Boettcher +leg., +SMF +, no. 14009–14011 + +;— + +2♀♀ +, + +20.ii.1915 + +, +Momungan +, +G. Boettcher +leg., +SMF +, no. 13964, 13965 + +;— + +1♂ +, + +18.vi.1915 + +, +Tankulan +, +G. Boettcher +leg., +SMF +, no. 13962 + +;— + +1♂ +, + +19.vi.1915 + +, +Tankulan +, +G. Boettcher +leg., +SMF +, no. 13961 + +;— + +14♂♂ +, +3♀♀ +, + +21.v.1915 + +, +Surigao +, +G. Boettcher +leg., +SMF +, no. 14027–14043 + +;— + +1♀ +, + +26.v.1915 + +, +Surigao +, +G. Boettcher +leg., +SMF +, no. 14044 + +;— + +1♀ +, + +14.vii.1915 + +, +Momungan +, +G. Boettcher +leg., +SMF +, no. 13945 + +;— + +2♀♀ +, + +11.xi.1915 + +, +Surigao +, +G. Boettcher +leg., +SMF +, no. 14045, 14046 + +;— + +9♂♂ +, +4♀♀ +, + +26.iii.1978 + +, +Eden, W. +Davao +, + +50 m +a.s.l. + +, +G. von Rosen +leg., +ZSM +, no. GvRosen 5543, 5544, 5566–5572, 5575–5578 + +. + + + + +Philippines +, +Mindoro Island + +: +1♂ +, + +25.i.1916 + +, +Subaan, G +. Boettcher leg., +SMF +, no. 13966 + +;— + +1♂ +, +1♀ +, + +xi.1917 + +, +Mangarin, G +. Boettcher leg., +SMF +, no. 13952, 13963 + +. + + + + +Philippines +, +Panaon Island +: + +1♀ +, + +8.xii.1915 + +, +Panaon, G +. Boettcher leg., +SMF +, no. 13946 + +. + + + + +Philippines +, +Polillo Island +: + +5♂♂ +, + +12.–20.viii.1915 + +, +Polillo, G +. Boettcher leg., +SMF +, no. 13955–13959 + +. + + + + +Philippines +, +Samar +Island + +: +2♂♂ +, + +18.iv.1915 + +, Catbalogan, +G. Boettcher +leg., +SMF +, no. 14014, 14015 + +;— + +2♀♀ +, + +18.–24.iv.1915 + +, Catbalogan, +G. Boettcher +leg., +SMF +, no. 14024, 14025 + +;— + +2♂♂ +, + +19.–22.iv.1915 + +, Catbalogan, +G. Boettcher +leg., +SMF +, no. 14020, 14021 + +;— + +2♂♂ +, + +21.iv.1915 + +, Catbalogan, +G. Boettcher +leg., +SMF +, no. 14022, 14023 + +;— + +4♂♂ +, +1♀ +, + +22.iv.1915 + +, Catbalogan, +G. Boettcher +leg., +SMF +, no. 14016–14019, 14026 + +. + + + + +Taiwan + +: +1♀ +, + +15.ix.1907 + +, +Koroton +, +Formosa +, +H. Sauter +leg., +UMB + +;— + +1♂ +, + +i.1908 + +, +Lambek Island +, +South-Formosa +, +H. Sauter +leg., +ZMH + +;— + +1♂ +, +1♀ +, + +viii.1908 + +, +Taihanroku +, +Formosa +, +Sauter +leg., +SMF + +, no. 13971, 13972. + + + +In addition to the specimens listed above there are more than 1830 +♂♂ +and +270 ♀♀ +from the +Philippines + + +at the +RMNH +originating from the following islands: +Balabac +, +Basilan +, +Batan +, +Biliran +, +Camiguin +, +Catanduan +, +Cebu +, +Dinagat +, +Homonhon +, +Leyte +, +Luzon +, +Marinduque +, +Masbate +, +Mindanao +, +Mindoro +, +Negros +, +Palawan +, +Panaon +, +Panay +, +Samar +, +Sibuyan +, +Siquijor +, +Sulu +Archipel +and Ticao. Data on these will be published as part of a forthcoming checklist and atlas of the dragonflies and damselflies of the +Philippines + +. + + +Characterization of male + + +Head +( +Figs 64 +, +92 +): Labrum, clypeus, antefrons, postfrons and vertex red; postocular spots red, separated by a thin black line from postfrons; rear of head black, with yellow along the eye margins; base and scape of antennae reddish, following segments brownish. + + +Thorax +( +Figs 63, 64 +, +109 +): Anterior lobe of pronotum red, this colour expanding in centre towards middle lobe; middle pronotal lobe black, with a centrally divided red patch in the middle, and on either side a large red patch dorsolaterally; posterior lobe of pronotum red; with its apical edge convex. Pterothorax red, with a thin black middorsal carina; humeral suture and metapleural suture each with a black stripe at posterior quarter, both slightly expanded in the middle dorsally and ventrally; metapleural suture may be covered with whitish pruinosity; interpleural suture with a thin black stripe at posterior quarter; venter pale, often covered with whitish pruinosity. Legs with femur and tibia pale reddish-white; femora bearing a black stripe outside reaching from distal margin to proximal half; tibiae blackish along spines; spines black. + + +Abdomen +( +Figs 63 +, +126a, b +, +165, 166 +): Variable in the whole range but generally with S1 red; S2 red, along its posterior carina black; S3 variable: in specimens from the +Philippines +, +Taiwan +, and +Japan +generally reddish, with variable washed-out blackish dorsal marking at posterior margin; in specimens from Borneo often dorsally black ( +Fig. 166 +); S4 variable: predominantly reddish, with variable washed-out blackish dorsal marking at posterior margin; in specimens from +Palawan +Islands/ +Philippines +it may be almost completely black dorsally, but usually at least the anterior margin remains washed-out reddish; in specimens from +Japan +S4 is dorsally black; S5 variable: Philippine specimens are predominantly black dorsally, but anterior edge may be washed-out with reddish in different extent; specimens from +Basilan +Island/ +Philippines +and from +Taiwan +have S5 reddish with a dark dorsal marking at posterior margin; S6–8 dorsally black; S9 red with posterior quarter to half black, sometimes with edges of black serrated; S10 red, predominantly with a black dorsal “x”-like marking, variable in exact shape; but segment may also be completely black dorsally, with only two small reddish spots (e.g. in a male from +Okinawa +, +Japan +). + + +Caudal appendages +( +Figs 143a–c +): Cercus laterally washed-out blackish, but may be reddish at base; in lateral view cercus shorter than S10, the upper margin slightly convex, its apex almost square, inter-lobe notch not apparent in this view; in dorsal view the apical part of the superior lobe of the cercus curved inwards and runs as a ridge to the inter-lobe notch, with a separate inward directed projection proximal of this visible in dorsolateral view; another inwardly directed projection is situated more proximally at before the middle of the cercus; the inferior lobe with a ridge, best visible in dorsolateral view; a small bulge-like interior projection at the base of the ridge is visble in dorsal and dorsolateral view; paraproct pale yellow-orange, slightly shorter than cercus; in lateral view subtriangular and tapered, with rounded apex; in dorsal view broadly rounded, concave bowl-like; bearing a black tooth at medial edge. + + +Characterization of female + + +Head: +Dark markings similar to male, but primary colours pale orange-ochre; postocular spots may be tinged greenish. + + +Thorax +: Pronotum markings as in male, but primary colour orange-ochre instead of reddish; posterior lobe of pronotum with two orange-ochre horns directed towards head, reaching the centre of the middle lobe. Pterothorax with primary colour orange-ochre; dark markings as in male; antehumeral area of pterothorax may be tinged greenish. + + +Abdomen +: S1–2 orange-ochre, with black dorsal markings, variable in size; S3 either orange-ochre, with a black dorsal marking posteriorly, variable in size, or completely black dorsally; S7–8 dorsally black, laterally pale yellowish; S9–10 either black dorsally, or black reduced to “x”-like marking, variable in exact shape; laterally yellow-ochre, this colour expanded dorsally in centre of segment, leaving anterior and posterior margin black. Cercus blackish; paraproct washed-out dark brownish; in lateral view rounded; ovipositor pale yellowish-ochre, not reaching posterior margin of S10, two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 11.5–14.5 px in forewing and 9.5–12.5 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 38.6–45.8; abdomen length (without appendages) 31.4–37.0; hindwing 21.3–25.7; forewing 22.7–27.8; Pt in forewing 0.7–1.0; male cercus 0.5–0.6. + + + +Diagnosis + + +The combination of a red clypeus and red frons ( +Figs 64 +, +92 +), a red pterothorax ( +Figs 63, 64 +, +109 +, +165, 166 +), and at least red S1–2, but up to S1–5 red ( +Figs 126a, b +, +165, 166 +), makes this species unmistakable. The red clypeus and frons are shared only by + +P. perfuscatum + +, but this species has smaller postocular spots and more extensive black on vertex ( +Figs 61 +, +91 +, +163, 164 +). Additionally mature males of + +P. perfuscatum + +have the pterothorax reddish-brown, covered with pruinosity up to interpleural suture, often a broad blackish humeral stripe ( +Figs 61 +, +108 +, +163, 164 +), and S1–2 are dorsally blackish and covered with pruinosity ( +Figs 61 +, +15a, b +, +163, 164 +). [Note: See discussion section for further details regarding populations from Borneo]. + + + + +Distribution + + +This species occurs throughout the +Philippines +and +Taiwan +, and reaches the southern Japanese Islands, as well as the Malaysian +Sabah +and +Sarawak +in northeast Borneo ( +Fig. 1 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F77CFFDFFF66DD6CFA0A9838.xml b/data/0E/57/D1/0E57D167F77CFFDFFF66DD6CFA0A9838.xml new file mode 100644 index 00000000000..dbe7e7a995c --- /dev/null +++ b/data/0E/57/D1/0E57D167F77CFFDFFF66DD6CFA0A9838.xml @@ -0,0 +1,301 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion simalurum +Lieftinck, 1948 + +stat. nov. + + + + +Figures: 76–77 ( +holotype +♂ +, envelope); 90 (head +♂ +); 107 (pterothorax +♂ +); 124a, b (abdomen +♂ +); 141a–c (caudal appendages +♂ +). + + + + + + +Pseudagrion pilidorsum +: + + +Ris (1915a) +: 12 + +–13 (description +♂ +; Simalur);— + +Ris (1915b) +: 99 + +(Simalur). + + + + + +Pseudagrion pilidorsum simalurum + +: + +Lieftinck (1948) +: 289 + +–290, +Fig. 5 +(description +♂ +; Simalur). + + + +Name-bearing type specimen examined +( +1♂ +) + + + + + +♂ +holotype +, + +i.1913 + +, Sinabang, +Simalur Island +, +Indonesia +, +E. Jacobson +leg., +RMNH +, JvT no. 2854 ( +Figs 76– 77 +) + +. + + +Lieftinck (1948: 290) +stated “Described from the +paratype +, in the Leiden Museum”, and +Lieftinck (1971: 112) +designated this specimen to be the +lectotype +. But the description is based on a single male, thus it is a +holotype +. + + + + +FIGURES 76–77 +. + +Pseudagrion simalurum + +, holotype male: ( +76 +) habitus lateral, ( +77 +) envelope. Photos: RMNH. + + + + +Other specimens examined +( +1♂ +) + + + + +Indonesia +, +Simalur Island +: + +1♂ +, + +i.1913 + +, +Sinabang, E +. Jacobson leg., +SMF +, no. 13951 + +. + + +Characterization of male + + +Head +( +Figs 76 +, +90 +): Labrum, clypeus, antefrons and postfrons yellowish; colour of postfrons separated from vertex around the level of median ocellus; vertex reddish, with a small black patch posterior of median ocellus; large black patches posterolateral of each lateral ocellus; postocular spots reddish, bordered by black; rear of head black, but narrow yellowish along the eye margins; base and scape of antennae yellow, following segments brownish ochre. + + +Thorax +( +Figs 76 +, +107 +): Anterior lobe of pronotum yellow-orange, this colour expanding in centre towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large yellow-orange patch in in the middle, and on either side a yellow-orange patch dorsolaterally; propleuron black above, yellow-orange below and to rear; posterior lobe of pronotum yellow-orange, with its margins black; apical edge of posterior lobe convex. Pterothorax orange-reddish; middorsal carina thin black; humeral suture with a thin black stripe which is expanded to an elongate black patch at posterior quarter; interpleural suture with a black stripe at posterior quarter, and a black spot anteriorly; metapleural suture with an elongate black patch at posterior quarter; a large black marking covering at least superior one-third to half of the mesinfraepisternum; venter pale, may be covered with whitish pruinosity. Legs pale orange; femora with black stripe outside, narrowing to proximal; tibiae with black along spines; spines black. + + +Abdomen +( +Figs 76 +, +124a, b +): S1–2 dorsally black, laterally orange; S3–8 dorsally black, laterally and each basal annulus pale yellowish; S9 black, with anterior third red; S10 dorsally with a broad black “x”-like marking, laterally red. + + +Caudal appendages +( +Figs 141a–c +): In lateral view cercus blackish, shorter than S10; subrectangular with an open inter-lobe notch around lower third; in dorsal view apex of the superior lobe of the cercus somewhat pointed, curved inwards; an inwards directed projection around the centre of the superior lobe of the cercus; a small bulge-like interior projection is situated at the base of a ridge on the inferior lobe of the cercus, visible in dorsal and dorsolateral views, proximal edge of this projection sharply sloping; paraproct brownish-ochre, around half as long as cercus; in lateral view subtriangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; bearing a small black tooth at medial edge. + +Wings with 14.5–15.0 px in forewing, and 12.5–12.5 px in hindwing. +Measurements (mm): Total length (with appendages) 45.0–45.4; abdomen length (without appendages) 37.3– 37.5; hindwing 24.3–25.0; forewing 25.5–26.5; Pt in forewing 0.8–1.0; male cercus 0.5–0.6. +Female unknown. + + + +Diagnosis + + + +Pseudagrion simalurum + +belongs to the group of endemic taxa of the island chain west of +Sumatra +. The males are separated from most congeners occurring at adjacent islands by having S1–2 dorsally black ( +Figs 76 +, +124a, b +). This character is elsewhere shared by + +P. igneum + +( +Figs 121a, b +) and + +P. obscurum + +( +Figs 123a, b +)—both are very similar to + +P. simalurum + +also in the colour of the head ( +Figs 87, 89a, b, 90 +), and the morphology of the male cercus ( +Figs 138a–c +, +140a–c, 141a–c +). We summarised the characters for separation in table 2. + + + + +Distribution + + +This species is apparently endemic to Simalur Island ( +Fig. 2 +). + + + + \ No newline at end of file diff --git a/data/0E/57/D1/0E57D167F77EFFC1FF66DF64FDAA9BD4.xml b/data/0E/57/D1/0E57D167F77EFFC1FF66DF64FDAA9BD4.xml new file mode 100644 index 00000000000..f18d40fddee --- /dev/null +++ b/data/0E/57/D1/0E57D167F77EFFC1FF66DF64FDAA9BD4.xml @@ -0,0 +1,409 @@ + + + +Revision of the Pseudagrion red-group occurring in Asia west of Lydekker’s line, with description of four new species (Odonata: Coenagrionidae) + + + +Author + +Seehausen, Malte +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + + + +Author + +Kalkman, Vincent J. +0000-0002-1484-7865 +Naturalis Biodiversity Center, 2300 RA Leiden, the Netherlands, vincent. kalkman @ naturalis. nl; https: // orcid. org / 0000 - 0002 - 1484 - 7865 +vincent.kalkman@naturalis.nl + + + +Author + +Bedjanič, Matjaž +0009-0000-6718-1078 +Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 & National Institute of Biology, Večna pot 121, SI- 1000, Ljubljana, Slovenia, matjaz. bedjanic @ nib. si; https: // orcid. org / 0000 - 0002 - 1926 - 0086 * Corresponding author & Zoologisches Museum der CAU zu Kiel, Hegewischstr. 3, 24105 Kiel, Germany, mseehausen @ zoolmuseum. uni-kiel. de; https: // orcid. org / 0009 - 0000 - 6718 - 1078 +mseehausen@zoolmuseum.uni-kiel.de + +text + + +Zootaxa + + +2025 + +2025-02-12 + + +5587 + + +1 + + +1 +66 + + + + +https://doi.org/10.11646/zootaxa.5587.1.1 + +journal article +10.11646/zootaxa.5587.1.1 +1175-5334 +B6F6D9C8-4423-4DC6-BC25-940725A83DB4 + + + + + + + +Pseudagrion ustum +Selys, 1876 + + + + + +Figures: 78 ( +holotype +♀ +); 95 (head +♂ +); 112 (pterothorax +♂ +); 129a, b (abdomen +♂ +); 146a–c (caudal appendages +♂ +); 168 (field photo +♂ +). + + + + + + +Pseudagrion ustum + +: + +Selys (1876) +: 515 + +(description +♀ +; Sulu Island, Malay Archipelago [actual Sula Archipelago, +Indonesia +]);— + +Ris (1915b) +: 97 + +, 99–100, +Fig. 21 +(key; description +♂ +; +Celebes +);— + +Lieftinck (1930a) +: 307 + +( +♂ +♀ +Buru). + + + +Name-bearing type specimen +(not examined) + + + + + +♀ +holotype +, +Sulu Island +, +Malay Archipelago +[actual Sula Archipelago, +Indonesia +], +IRSNB +( +Fig. 78 +) + +. + + + +Currently the +holotype +could not be found at the +IRSNB +( +J. Constant +, pers. comm.) + +. + + + + +Other specimens examined +( +15♂♂ +, +8♀♀ +) + + + + +Indonesia +, +Sulawesi + +: +3♂♂ +, +1♀ +, +Toli-Toli +, +SMF + +, no. 13754–13757;— + +1♂ +, + +March 1896 + +, +Lompa-Battau +, 3000’, +H. Fruhstorfer +leg., +ZMH + +;— + +5♂♂ +, +4♀♀ +, 1897, +Bonthain +, +SMF + +, no. 13743–13747, 13749, 13751–13753;— + +1♂ +, + +28.iv.1914 + +, +Maros +, +Dr L. Martin +leg., +SMF + +, no. 13933;— + +2♂♂ +, + +18.ix.-14.x.1983 + +, +Maros +, +E of Bantimurung area +, surroundings of +Bantimurung +waterfall, + +190 m +a.s.l. + +, +S.S. Pariwono +leg., ex. +Coll. M.A. Lieftinck +, +RMNH + +, no. 1658917, 1658918;— + +3♂♂ +, +3♀♀ +, + +21.ix.-10.x.1983 + +, +Maros +, +E of Bantimurung area +, QK94, +Patunuang Asue +, + +250 m +a.s.l. + +, +S.S. Pariwono +leg., +RMNH + +, no. 1519261, 1519262, 1519264. + + + +FIGURE 78 +. + +Pseudagrion ustum + +, holotype female, water colour by Selys, 20.viii.1875 (IRSNB Ag125a). Photo: IRSNB. + + + +Characterization of male + + +Head +( +Fig. 95 +): Labrum, clypeus, antefrons and postfrons yellow; colour of postfrons separated from vertex around level of lateral ocelli; vertex red; black posterolateral patches along each lateral ocellus; postocular spots red, separated by a black line from the postfrons; rear of head black, but pale along the eye margins; base and scape of the antennae yellow, following segments brownish ochre; eyes when alive reddish brown-orange. + + +Thorax +( +Figs 112 +, +168 +): Anterior lobe of pronotum orange, this colour expanding in centre towards middle lobe, anterior margin black; middle pronotal lobe black, with a centrally divided large orange patch in the middle, and on either side a large orange patch dorsolaterally; propleuron black above, orange below and to rear; posterior lobe of pronotum orange; apical edge of posterior lobe convex. Pterothorax orange-reddish; middorsal carina black; humeral suture and metapleural suture each with an elongate black patch at posterior quarter, both patches expanded in middle; interpleural suture may have a narrow blackish stripe, and a blackish spot anteriorly; venter pale, may be covered with pruinosity. Legs pale yellowish-reddish with a black stripe at posterior quarter to half of femora; tibiae with black stripe along black spines. + + +Abdomen +( +Figs 129a, b +, +168 +): S1 red, may have a washed-out blackish curly bracket-like patch dorsally; S2 red, but along posterior carina black, often with washed-out blackish marking dorsally at posterior quarter; S3–8 dorsally black, laterally pale yellow-orange; S9 red, with at least its posterior margin, but up to posterior one-third black; anterior margin of black colour may be serrated; some males additionally have two black spots situated dorsolaterally; S10 red, with a black dorsal “x”-like marking, variable in exact shape. + + +Caudal appendages +( +Figs 146a–c +): Cercus laterally blackish, slightly shorter than S10, inferior lobe subequal to superior lobe, its apex only just visible in this view; in lateral view superior lobe abruptly narrowing near apex, with inter-lobe notch very small; in dorsal view the cercus broadly rounded; the terminus of the superior lobe curved strongly inwards and somewhat bifurcated, this bifurcation more apparent in dorsolateral view; interior projection of inferior lobe of cercus very large, clearly visible in lateral view where it appears subtriangular and basally positioned; paraproct pale yellow-ochre, slightly longer than half of cercus; in lateral view somewhat triangular and tapered with rounded apex; in dorsal view broadly rounded, concave bowl-like; medial edge with blackish tooth. + + +Characterization of female +( +Fig. 78 +) + + +Head +: As in male, but orange-ochre instead of yellow and red. + + +Thorax +: Pronotum as in male, but pale yellowish instead of orange-reddish; posterior lobe with apical edge almost triangular-convex; bearing two yellowish horns directed towards the head, almost reaching the centre of the middle lobe. Pterothorax as in male, but when alive dorsum orange-ochre instead of red, and laterally pale greenish-yellow. + + +Abdomen +: S1 orange-ochre, with a black dorsal patch covering anterior two-thirds; S2 orange-ochre, with posterior margin black, and a black middorsal stripe which is expanded patch-like in posterior third; S3–7 dorsally black, laterally pale yellowish; S8 dorsally black, the black may be constricted in posterior third; S9 dorsally black, narrowing posteriorly, laterally pale yellowish; S10 pale yellowish, anterior margin may be black. Cercus pale; paraproct pale, in lateral view rounded; ovipositor pale yellowish, not reaching to posterior margin of S10, two brownish styles reaching beyond S10; inferior edge of ovipositor bearing several very small teeth. + +Wings of both sexes with 13.5–16.0 px in forewing, and 11.5–15.5 px in hindwing. +Measurements of both sexes (mm): Total length (with appendages) 44.5–50.8; abdomen length (without appendages) 36.8–42.4; hindwing 24.0–29.7; forewing 25.6–31.2; Pt in forewing 0.8–0.9; male cercus 0.6. + + + +Diagnosis + + +The morphology of the male cercus ( +Figs 146a–c +) separates this species from all congeners. + +Pseudagrion acutidens + +has also a cercus with a distinct tooth ( +Figs 130a–c +), but it is directed inward instead of upward as it is in + +P. ustum + +. Additionally the colour marking of the pterothorax and of the abdomen separates + +P. ustum + +( +Figs 78 +, +95 +, +112 +, +129a, b +, +168 +) in both sexes from + +P. crocops + +( +Figs 23, 25 +, +82 +, +116a, b +, +154 +), currently the only known other red + +Pseudagrion +species + +which occurs at +Sulawesi +. + + + + +Distribution + + +Records of this species are available from +Sulawesi +, the Sula Archipelago, and Buru ( +Fig. 3 +). + + + + \ No newline at end of file