Add updates up until 2025-02-25 09:02:04
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<document id="E8FE575DB8C11A131864A4E64BA85790" ID-DOI="10.11646/zootaxa.5583.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14798039" ID-ZooBank="5BD9EECB-50CD-4810-AE0A-9868F0645681" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1738624864826" checkinUser="plazi" docAuthor="Zhang, Hai-Xin, Zhang, Yan-Ping, Chen, Wen-Jing & Zhang, E." docDate="2025" docId="951A62731117FFB2FF79F9F82B5BF84A" docLanguage="en" docName="zootaxa.5583.1.5.pdf" docOrigin="Zootaxa 5583 (1)" docSource="https://doi.org/10.11646/zootaxa.5583.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Niwaella tigrina Zhang, Zhang, Chen & Zhang, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="107" masterDocId="69231A0B1114FFB4FFEEFFFA2938FF9B" masterDocTitle="Niwaella tigrina, a new species of spined loach from the Poyang Lake system of Jiangxi Province, East China" masterLastPageNumber="112" masterPageNumber="101" pageNumber="104" updateTime="1740474121457" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C36718AA8EC5EF92086F6FD000B6CDD3">
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<mods:title id="EE4F86982B1A7FAF9BD3720026D6741E">Niwaella tigrina, a new species of spined loach from the Poyang Lake system of Jiangxi Province, East China</mods:title>
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<mods:roleTerm id="BBAC8D559883B05D35E1721CEB5F3F97">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3527C50130EFE70B8E2CE59DF4DCFFF2">Zhang, Hai-Xin</mods:namePart>
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<mods:affiliation id="0B5F6CE1363FE802FD4F9FDD1F2ED1C4">Jiangxi Provincial Institute of Fisheries Science, Nanchang 330039, China & Jiangxi Provincial Fishery Resources and Ecological Environment Monitoring Center, Nanchang 330039, China.</mods:affiliation>
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<mods:namePart id="3F78C4593CEDC6FB40EA95364DEC6F3C">Zhang, Yan-Ping</mods:namePart>
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<mods:affiliation id="2176065A8EC33EEECFB19A355F17F69F">Jiangxi Provincial Institute of Fisheries Science, Nanchang 330039, China</mods:affiliation>
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</mods:role>
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<mods:namePart id="DC224A285A146ABCEEB95D6A78957F98">Chen, Wen-Jing</mods:namePart>
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<mods:affiliation id="44208F6CF11F77E6FB16BCC3B5F65E4B">Jiangxi Provincial Institute of Fisheries Science, Nanchang 330039, China</mods:affiliation>
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</mods:role>
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<mods:namePart id="755071D0F219D3B4D16321F58FB7FEDF">Zhang, E.</mods:namePart>
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<mods:affiliation id="5C343F01D81A6EE389DC60DA8F974164">Institute of hydrobiology, Chinese academy of sciences, Hubei 430072, China.</mods:affiliation>
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<mods:title id="C14BF802D962CC145F669E8861CD505B">Zootaxa</mods:title>
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<mods:date id="E2F568886390A90BF89E8454E8F8E452">2025</mods:date>
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<mods:number id="A8AD6FFF35F0B2CD9AD07FF5495AF7BD">2025-01-31</mods:number>
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<mods:number id="63CB8662818B5A2242C63CCA9E968E3E">5583</mods:number>
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<mods:number id="96A7CA66961CD28DBB333811577DA566">1</mods:number>
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<mods:start id="DAED725DA774CAC658EBF916B96C3010">101</mods:start>
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<mods:url id="884BA9F2FCFEFA2780AD6CFD3BB7DEAC">https://doi.org/10.11646/zootaxa.5583.1.5</mods:url>
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<mods:classification id="29D943089E1681DFE460F8811F54E5EB">journal article</mods:classification>
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<mods:identifier id="E089FCCC74CB3473A07EC13D02DECD6E" type="DOI">10.11646/zootaxa.5583.1.5</mods:identifier>
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<mods:identifier id="6AAE7DBC795F4FC12829589CE7049392" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7B1D02B2E0374D8B39771859DDB4FD9F" type="Zenodo-Dep">14798039</mods:identifier>
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<mods:identifier id="8FF0BDAF2AA926A476342EC37CCDDBC8" type="ZooBank">5BD9EECB-50CD-4810-AE0A-9868F0645681</mods:identifier>
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</mods:mods>
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<treatment id="951A62731117FFB2FF79F9F82B5BF84A" LSID="urn:lsid:plazi:treatment:951A62731117FFB2FF79F9F82B5BF84A" httpUri="http://treatment.plazi.org/id/951A62731117FFB2FF79F9F82B5BF84A" lastPageId="6" lastPageNumber="107" pageId="3" pageNumber="104">
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<subSubSection id="55A980EE1117FFB7FF79F9F828DDF987" box="[151,485,1537,1564]" pageId="3" pageNumber="104" type="nomenclature">
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<paragraph id="1D0CD3651117FFB7FF79F9F828DDF987" blockId="3.[151,485,1537,1600]" box="[151,485,1537,1564]" pageId="3" pageNumber="104">
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<heading id="464464091117FFB7FF79F9F828DDF987" bold="true" box="[151,485,1537,1564]" fontSize="11" level="1" pageId="3" pageNumber="104" reason="1">
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<emphasis id="2FC70F771117FFB7FF79F9F828DDF987" bold="true" box="[151,485,1537,1564]" pageId="3" pageNumber="104">
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||||
3.1
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<taxonomicName id="DAB3A8E61117FFB7FF2DF9FB28B8F987" box="[195,384,1537,1564]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="104" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771117FFB7FF2DF9FB28B8F987" bold="true" box="[195,384,1537,1564]" italics="true" pageId="3" pageNumber="104">Niwaellatigrina</emphasis>
|
||||
</taxonomicName>
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||||
<taxonomicNameLabel id="34F4B20C1117FFB7FE66F9F828DDF987" box="[392,485,1538,1564]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
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||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="55A980EE1117FFB7FF79F9DC29D4F9DB" box="[151,236,1574,1600]" pageId="3" pageNumber="104" type="description">
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<paragraph id="1D0CD3651117FFB7FF79F9DC29D4F9DB" blockId="3.[151,485,1537,1600]" box="[151,236,1574,1600]" pageId="3" pageNumber="104">
|
||||
(
|
||||
<figureCitation id="8588CFE01117FFB7FF71F9DC29DBF9DB" box="[159,227,1574,1600]" captionStart="FIGURE 2" captionStartId="4.[151,250,1786,1810]" captionTargetBox="[184,1403,921,1763]" captionTargetId="figure-378@4.[184,1403,921,1763]" captionTargetPageId="4" captionText="FIGURE 2. Niwaella tigrina, JPIFS20230401, holotype, 55.7 mm SL, Ding-Jiang, Jiangxi Province, China. (a) lateral view, (b) dorsal view, (c) ventral view. Scale bar=10 mm." figureDoi="http://doi.org/10.5281/zenodo.14798044" httpUri="https://zenodo.org/record/14798044/files/figure.png" pageId="3" pageNumber="104">Fig. 2</figureCitation>
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)
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="55A980EE1117FFB7FF79F9942A07F883" pageId="3" pageNumber="104" type="materials_examined">
|
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<paragraph id="1D0CD3651117FFB7FF79F9942AD4F937" blockId="3.[151,1437,1646,2032]" pageId="3" pageNumber="104">
|
||||
<materialsCitation id="ADDBD9381117FFB7FF79F9942AD4F937" collectingDate="2023-04" collectionCode="JPIFS" collectorName="Hai-Xin Zhang" county="Tonggu County" latitude="28.515278" location="Ding-Jiang flowing into Xiu-Shui" longLatPrecision="20" longitude="114.28306" pageId="3" pageNumber="104" specimenCode="JPIFS20230401" specimenCount="1" stateProvince="Jiangxi Province" typeStatus="holotype">
|
||||
<emphasis id="2FC70F771117FFB7FF79F9942833F913" bold="true" box="[151,267,1646,1672]" pageId="3" pageNumber="104">
|
||||
<typeStatus id="C2086DC71117FFB7FF79F994283EF913" box="[151,262,1646,1672]" pageId="3" pageNumber="104" type="holotype">Holotype</typeStatus>
|
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.
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</emphasis>
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||||
<specimenCode id="4D157B1E1117FFB7FEFDF99428F2F913" box="[275,458,1646,1672]" pageId="3" pageNumber="104">JPIFS20230401</specimenCode>
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||||
,
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||||
<quantity id="DA4B7E801117FFB7FE38F9942B0AF912" box="[470,562,1646,1673]" metricMagnitude="-2" metricUnit="m" metricValue="5.57" pageId="3" pageNumber="104" unit="mm" value="55.7">55.7mm</quantity>
|
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SL, from
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<location id="186C85BE1117FFB7FD47F9942D11F913" LSID="urn:lsid:plazi:treatment:951A62731117FFB2FF79F9F82B5BF84A:186C85BE1117FFB7FD47F9942D11F913" box="[681,1065,1646,1672]" county="Tonggu County" latitude="28.515278" longLatPrecision="20" longitude="114.28306" name="Ding-Jiang flowing into Xiu-Shui" pageId="3" pageNumber="104" stateProvince="Jiangxi Province">Ding-Jiang flowing into Xiu-Shui</location>
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in
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<collectingCounty id="F46DABE91117FFB7FBA0F9942DC6F913" box="[1102,1278,1646,1672]" pageId="3" pageNumber="104">Tonggu County</collectingCounty>
|
||||
(
|
||||
<geoCoordinate id="7887B5A21117FFB7FAE1F9952CA0F911" box="[1295,1432,1647,1674]" degrees="28" direction="north" minutes="30" orientation="latitude" pageId="3" pageNumber="104" precision="15" seconds="55" value="28.515278">28°30′55″N</geoCoordinate>
|
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,
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<geoCoordinate id="7887B5A21117FFB7FF79F9692812F935" box="[151,298,1683,1710]" degrees="114" direction="east" minutes="16" orientation="longitude" pageId="3" pageNumber="104" precision="15" seconds="59" value="114.28306">114°16′59″E</geoCoordinate>
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),
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<collectingRegion id="DF771D871117FFB7FED0F96828C4F937" box="[318,508,1682,1708]" country="China" name="Jiangxi" pageId="3" pageNumber="104">Jiangxi Province</collectingRegion>
|
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; collected by
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<collectorName id="B046B6B31117FFB7FD79F9682A7CF937" box="[663,836,1682,1708]" pageId="3" pageNumber="104">Hai-Xin Zhang</collectorName>
|
||||
, in
|
||||
<date id="690DF5A51117FFB7FC84F9682AD0F937" box="[874,1000,1682,1708]" pageId="3" pageNumber="104" value="2023-04">
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<collectingDate id="79490C4D1117FFB7FC84F9682AD0F937" box="[874,1000,1682,1708]" pageId="3" pageNumber="104" value="2023-04">April 2023</collectingDate>
|
||||
</date>
|
||||
.
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</materialsCitation>
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</paragraph>
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<paragraph id="1D0CD3651117FFB7FF29F94C2A07F883" blockId="3.[151,1437,1646,2032]" pageId="3" pageNumber="104">
|
||||
<materialsCitation id="ADDBD9381117FFB7FF29F94C2AC4F94B" box="[199,1020,1718,1745]" collectingDate="2023-04" collectionCode="IHB, JPIFS" collectorName="Hai-Xin Zhang" country="China" county="Tonggu County" latitude="28.733612" location="Tonggu County" longLatPrecision="20" longitude="114.21861" pageId="3" pageNumber="104" specimenCode="IHB 20230402, JPIFS 20230803" specimenCount="18" stateProvince="Jiangxi Province" typeStatus="paratype">
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<emphasis id="2FC70F771117FFB7FF29F94C287EF94B" bold="true" box="[199,326,1718,1744]" pageId="3" pageNumber="104">
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<typeStatus id="C2086DC71117FFB7FF29F94C2879F94B" box="[199,321,1718,1744]" pageId="3" pageNumber="104" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCode id="4D157B1E1117FFB7FEBEF94C28C6F94B" box="[336,510,1718,1744]" pageId="3" pageNumber="104">IHB 20230402</specimenCode>
|
||||
,
|
||||
<quantity id="DA4B7E801117FFB7FDE2F94C2B4CF94B" box="[524,628,1718,1745]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" pageId="3" pageNumber="104" unit="mm" value="59.5">59.5 mm</quantity>
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SL, other data same as holotype;
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</materialsCitation>
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<materialsCitation id="ADDBD9381117FFB7FBE8F94C2A07F883" collectingDate="2023-04" collectionCode="IHB, JPIFS" collectorName="Hai-Xin Zhang" country="China" county="Tonggu County" latitude="28.733612" location="Tonggu County" longLatPrecision="20" longitude="114.21861" pageId="3" pageNumber="104" specimenCode="IHB 20230402, JPIFS 20230803" specimenCount="18" stateProvince="Jiangxi Province" typeStatus="paratype">
|
||||
<specimenCode id="4D157B1E1117FFB7FBE8F94C2DF0F94B" box="[1030,1224,1718,1744]" pageId="3" pageNumber="104">JPIFS 20230803</specimenCode>
|
||||
-20,
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<specimenCount id="0BB518EC1117FFB7FB14F94C2CA1F94B" box="[1274,1433,1718,1744]" count="18" pageId="3" pageNumber="104" type="generic">18 specimens</specimenCount>
|
||||
,
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||||
<quantity id="DA4B7E801117FFB7FF79F920280AF96F" box="[151,306,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="5.534999999999999" metricValueMax="6.39" metricValueMin="4.68" pageId="3" pageNumber="104" unit="mm" value="55.349999999999994" valueMax="63.9" valueMin="46.8">46.8–63.9mm</quantity>
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SL, from Jinsha-He draining into Xiu-Shui in
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<collectingCounty id="F46DABE91117FFB7FCB0F9202D29F96F" box="[862,1041,1754,1780]" pageId="3" pageNumber="104">Tonggu County</collectingCounty>
|
||||
(
|
||||
<geoCoordinate id="7887B5A21117FFB7FBCBF9212DA7F96D" box="[1061,1183,1755,1782]" degrees="28" direction="north" minutes="44" orientation="latitude" pageId="3" pageNumber="104" precision="15" seconds="1" value="28.733612">28°44′1″N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="7887B5A21117FFB7FB41F9212C0BF96D" box="[1199,1331,1755,1782]" degrees="114" direction="east" minutes="13" orientation="longitude" pageId="3" pageNumber="104" precision="15" seconds="7" value="114.21861">114°13′7″E</geoCoordinate>
|
||||
),
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<collectingRegion id="DF771D871117FFB7FAA5F92029C5F883" country="China" name="Jiangxi" pageId="3" pageNumber="104">Jiangxi Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="65A493F51117FFB7FEE7F9042877F883" box="[265,335,1790,1816]" name="China" pageId="3" pageNumber="104">China</collectingCountry>
|
||||
; collected by
|
||||
<collectorName id="B046B6B31117FFB7FE07F9042BAEF883" box="[489,662,1790,1816]" pageId="3" pageNumber="104">Hai-Xin Zhang</collectorName>
|
||||
, in
|
||||
<date id="690DF5A51117FFB7FD53F9042A03F883" box="[701,827,1790,1816]" pageId="3" pageNumber="104" value="2023-04">
|
||||
<collectingDate id="79490C4D1117FFB7FD53F9042A03F883" box="[701,827,1790,1816]" pageId="3" pageNumber="104" value="2023-04">April 2023</collectingDate>
|
||||
</date>
|
||||
.
|
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</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="55A980EE1117FFB0FF29F8D82DC2FEDA" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104" type="diagnosis">
|
||||
<paragraph id="1D0CD3651117FFB0FF29F8D82DC2FEDA" blockId="3.[151,1437,1646,2032]" lastBlockId="4.[151,1437,151,897]" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
|
||||
<emphasis id="2FC70F771117FFB7FF29F8D8287AF8A7" bold="true" box="[199,322,1826,1852]" pageId="3" pageNumber="104">Diagnosis.</emphasis>
|
||||
<taxonomicName id="DAB3A8E61117FFB7FEA6F8D82B38F8A7" box="[328,512,1826,1852]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="104" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771117FFB7FEA6F8D82B38F8A7" box="[328,512,1826,1852]" italics="true" pageId="3" pageNumber="104">Niwaella tigrina</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from all Chinese congeneric species by its unique body coloration featuring 20–25 transverse black bars on the dorsum and upper portion of the flank and an indistinct blackish mid-dorsal stripe from the nape to the caudal-fin (vs. a row of transverse black blotches on the dorsum and vermicular black marks on the upper portion of the flank, or a conspicuous dark brown mid-dorsal stripe and numerous vermicular marks on the dorsum and upper portion of the flank). It is similar to
|
||||
<taxonomicName id="DAB3A8E61117FFB7FBE2F8492DF7F857" authorityName="Chen & Chen" authorityYear="2005" box="[1036,1231,1971,1996]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="104" phylum="Chordata" rank="species" species="xingjiangensis">
|
||||
<emphasis id="2FC70F771117FFB7FBE2F8492DF7F857" box="[1036,1231,1971,1996]" italics="true" pageId="3" pageNumber="104">N. xingjiangensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="DAB3A8E61117FFB7FAEAF8492CA4F857" authorityName="Chen & Chen" authorityYear="2017" box="[1284,1436,1970,1996]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="104" phylum="Chordata" rank="species" species="brevipinna">
|
||||
<emphasis id="2FC70F771117FFB7FAEAF8492CA4F857" box="[1284,1436,1970,1996]" italics="true" pageId="3" pageNumber="104">N. brevipinna</emphasis>
|
||||
</taxonomicName>
|
||||
in possessing a lower jaw fully covered by the lower lip, a trait not shared with other congeneric species.
|
||||
<taxonomicName id="DAB3A8E61117FFB0FAD9F82C29DBFF2B" family="Cobitidae" genus="Niwaella" kingdom="Animalia" lastPageId="4" lastPageNumber="105" order="Cypriniformes" pageId="3" pageNumber="104" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771117FFB0FAD9F82C29DBFF2B" italics="true" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">Niwaella tigrina</emphasis>
|
||||
</taxonomicName>
|
||||
is further distinguished from
|
||||
<taxonomicName id="DAB3A8E61110FFB0FDD8FF6D2BC1FF2B" authorityName="Chen & Chen" authorityYear="2005" box="[566,761,151,176]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="105" phylum="Chordata" rank="species" species="xingjiangensis">
|
||||
<emphasis id="2FC70F771110FFB0FDD8FF6D2BC1FF2B" box="[566,761,151,176]" italics="true" pageId="4" pageNumber="105">N. xingjiangensis</emphasis>
|
||||
</taxonomicName>
|
||||
by having 17–20 brownish or black vertical bars below the mid-lateral line of the body (vs. 16–18, with a few anterior bars fused to form a dark stripe along the mid-lateral line). Additionally it has a thicker (vs. slender) curved suborbital spine with a shorter (vs. longer) processus latero-caudalis, which is less than one-fifth (vs. one-third) of the processus medio-caudalis. Furthermore, it differs from
|
||||
<taxonomicName id="DAB3A8E61110FFB0FA6DFEF92837FEDA" authorityName="Chen & Chen" authorityYear="2017" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="105" phylum="Chordata" rank="species" species="brevipinna">
|
||||
<emphasis id="2FC70F771110FFB0FA6DFEF92837FEDA" italics="true" pageId="4" pageNumber="105">N. brevipinna</emphasis>
|
||||
</taxonomicName>
|
||||
in having five (vs. six) and six (vs. four) branched dorsal- and anal-fin rays, respectively.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="55A980EE1110FFB1FF29FEB02C1CF810" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105" type="description">
|
||||
<paragraph id="1D0CD3651110FFB0FF29FEB02891FE4A" blockId="4.[151,1437,151,897]" pageId="4" pageNumber="105">
|
||||
<emphasis id="2FC70F771110FFB0FF29FEB02861FEFF" bold="true" box="[199,345,330,356]" pageId="4" pageNumber="105">Description.</emphasis>
|
||||
Morphometric measurements for
|
||||
<typeStatus id="C2086DC71110FFB0FD27FEB12BC1FEFE" box="[713,761,331,357]" pageId="4" pageNumber="105">type</typeStatus>
|
||||
specimens were provided in
|
||||
<tableCitation id="5031E6DE1110FFB0FBC3FEB12DB8FEFE" box="[1069,1152,331,357]" captionStart="TABLE 2" captionStartId="6.[151,239,296,320]" captionTargetPageId="6" captionText="TABLE 2. Morphometric measurements and meristic counts of Niwaella tigrina." pageId="4" pageNumber="105">Table 2</tableCitation>
|
||||
. General body appearance and structures of different mouthpart depicted in
|
||||
<figureCitation id="8588CFE01110FFB0FD22FE952A37FE12" box="[716,783,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1786,1810]" captionTargetBox="[184,1403,921,1763]" captionTargetId="figure-378@4.[184,1403,921,1763]" captionTargetPageId="4" captionText="FIGURE 2. Niwaella tigrina, JPIFS20230401, holotype, 55.7 mm SL, Ding-Jiang, Jiangxi Province, China. (a) lateral view, (b) dorsal view, (c) ventral view. Scale bar=10 mm." figureDoi="http://doi.org/10.5281/zenodo.14798044" httpUri="https://zenodo.org/record/14798044/files/figure.png" pageId="4" pageNumber="105">Fig. 2</figureCitation>
|
||||
and
|
||||
<figureCitation id="8588CFE01110FFB0FCA4FE952AA5FE12" box="[842,925,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,943,967]" captionTargetBox="[303,1284,417,919]" captionTargetId="figure-142@5.[303,1285,417,919]" captionTargetPageId="5" captionText="FIGURE 3. Niwaella tigrina, JPIFS 20230401, holotype. (a) mouth, (b) pre-dorsal scale removed from upper trunk." figureDoi="http://doi.org/10.5281/zenodo.14798046" httpUri="https://zenodo.org/record/14798046/files/figure.png" pageId="4" pageNumber="105">Fig. 3a</figureCitation>
|
||||
, respectively. The body is moderately small, slender, and slightly compressed laterally; depth is nearly uniform anterior to anal-fin origin, slightly decreasing towards caudal-fin base.
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD3651110FFB0FF29FE212B37FC1A" blockId="4.[151,1437,151,897]" pageId="4" pageNumber="105">
|
||||
The head is relatively shorter but longer than the maximum vertical depth of the body. The snout looks slightly obtuse in lateral view and triangular or bluntly pointed in dorsal and ventral view, almost equal to postorbital head length. The eyes are small, positioned dorso-laterally in the middle of the head, with convex and narrow interorbital space; diameter nearly equal to interorbital width. The mouth is small and inferior, with three pairs of short barbels: one pair of rostral barbels not reaching the corners of the mouth, one pair of maxillary barbels not reaching the anterior margin of the eye, and one pair of maxillomandibular barbels. The lips are fleshy and smooth. The upper lip is broadened and continuous with a thick lower lip. The lower lip is divided into two well-developed mental lobes (
|
||||
<figureCitation id="8588CFE01110FFB0FF71FD2D29CBFD6A" box="[159,243,727,753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1786,1810]" captionTargetBox="[184,1403,921,1763]" captionTargetId="figure-378@4.[184,1403,921,1763]" captionTargetPageId="4" captionText="FIGURE 2. Niwaella tigrina, JPIFS20230401, holotype, 55.7 mm SL, Ding-Jiang, Jiangxi Province, China. (a) lateral view, (b) dorsal view, (c) ventral view. Scale bar=10 mm." figureDoi="http://doi.org/10.5281/zenodo.14798044" httpUri="https://zenodo.org/record/14798044/files/figure.png" pageId="4" pageNumber="105">Fig. 2a</figureCitation>
|
||||
;
|
||||
<figureCitation id="8588CFE01110FFB0FEEEFD2D286BFD6A" box="[256,339,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,943,967]" captionTargetBox="[303,1284,417,919]" captionTargetId="figure-142@5.[303,1285,417,919]" captionTargetPageId="5" captionText="FIGURE 3. Niwaella tigrina, JPIFS 20230401, holotype. (a) mouth, (b) pre-dorsal scale removed from upper trunk." figureDoi="http://doi.org/10.5281/zenodo.14798046" httpUri="https://zenodo.org/record/14798046/files/figure.png" pageId="4" pageNumber="105">Fig. 3a</figureCitation>
|
||||
). The lower jaw is fully covered by the lower lip. The subocular spine is thick and curved, with processus latero-caudalis being shorter than one-fifth of its processus medio-caudalis (
|
||||
<figureCitation id="8588CFE01110FFB0FB9CFD012D87FC8E" box="[1138,1215,763,789]" captionStart="FIGURE 4" captionStartId="5.[151,250,1367,1391]" captionTargetBox="[151,1435,1052,1343]" captionTargetId="figure-162@5.[151,1435,1052,1343]" captionTargetPageId="5" captionText="FIGURE 4. Niwaella tigrina, JPIFS20230401, holotype. (a) X-ray, (b) Micro-CT scan of suborbital spine." figureDoi="http://doi.org/10.5281/zenodo.14798048" httpUri="https://zenodo.org/record/14798048/files/figure.png" pageId="4" pageNumber="105">Fig.4b</figureCitation>
|
||||
). The lateral line is short, not reaching to the tip of the pectoral fin (
|
||||
<figureCitation id="8588CFE01110FFB0FD44FCE52BC3FCA2" box="[682,763,799,825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1786,1810]" captionTargetBox="[184,1403,921,1763]" captionTargetId="figure-378@4.[184,1403,921,1763]" captionTargetPageId="4" captionText="FIGURE 2. Niwaella tigrina, JPIFS20230401, holotype, 55.7 mm SL, Ding-Jiang, Jiangxi Province, China. (a) lateral view, (b) dorsal view, (c) ventral view. Scale bar=10 mm." figureDoi="http://doi.org/10.5281/zenodo.14798044" httpUri="https://zenodo.org/record/14798044/files/figure.png" pageId="4" pageNumber="105">Fig. 2a</figureCitation>
|
||||
). Body covered with fine and rounded scales. Each scale has a large focal area, with 23–26 primary radial grooves and a few secondary radial grooves (
|
||||
<figureCitation id="8588CFE01110FFB0FB71FCB92DD4FCC6" box="[1183,1260,835,861]" captionStart="FIGURE 3" captionStartId="5.[151,250,943,967]" captionTargetBox="[303,1284,417,919]" captionTargetId="figure-142@5.[303,1285,417,919]" captionTargetPageId="5" captionText="FIGURE 3. Niwaella tigrina, JPIFS 20230401, holotype. (a) mouth, (b) pre-dorsal scale removed from upper trunk." figureDoi="http://doi.org/10.5281/zenodo.14798046" httpUri="https://zenodo.org/record/14798046/files/figure.png" pageId="4" pageNumber="105">Fig.3b</figureCitation>
|
||||
). The vertebrae arrangement is 4+41+1 (
|
||||
<tableCitation id="5031E6DE1110FFB0FE44FC9D2B39FC1A" box="[426,513,871,897]" captionStart="TABLE 2" captionStartId="6.[151,239,296,320]" captionTargetPageId="6" captionText="TABLE 2. Morphometric measurements and meristic counts of Niwaella tigrina." pageId="4" pageNumber="105">Table 2</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="49CC83ED1110FFB0FF79F9002B45F8AD" ID-DOI="http://doi.org/10.5281/zenodo.14798044" ID-Zenodo-Dep="14798044" httpUri="https://zenodo.org/record/14798044/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,250,1786,1810]" targetBox="[184,1403,921,1763]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="1D0CD3651110FFB0FF79F9002B45F8AD" blockId="4.[151,1436,1786,1846]" pageId="4" pageNumber="105">
|
||||
<emphasis id="2FC70F771110FFB0FF79F900282AF889" bold="true" box="[151,274,1786,1810]" pageId="4" pageNumber="105">FIGURE 2.</emphasis>
|
||||
<taxonomicName id="DAB3A8E61110FFB0FEF6F9002B3FF889" authority=", JPIFS" authorityName="JPIFS" box="[280,519,1786,1810]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="105" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771110FFB0FEF6F9002886F889" box="[280,446,1786,1810]" italics="true" pageId="4" pageNumber="105">Niwaella tigrina</emphasis>
|
||||
, JPIFS
|
||||
</taxonomicName>
|
||||
20230401, holotype, 55.7 mm SL, Ding-Jiang, Jiangxi Province, China. (a) lateral view, (b) dorsal view, (c) ventral view. Scale bar=10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="1D0CD3651110FFB1FF29F8722878FE12" blockId="4.[151,1436,1928,2027]" lastBlockId="5.[151,1436,151,393]" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105">
|
||||
The dorsal fin contains three unbranched and five branched rays, making it moderately short. The distal margin is slightly concave, inserted just behind the midpoint between the tip of the snout and the caudal-fin base, or opposite to pelvic-fin insertion. The pectoral fins are with one unbranched and six or seven branched rays, extending from the posterior margins of the gill cover and stretched backward around one-third distance from the pectoral to the pelvic-fin base. The pelvic fins are with one unbranched and six branched rays. These are closer to the caudal fin than to the pelvic fin insertion and extended slightly beyond the vertical line at the posterior end of the dorsal fin but do not reach one-third of the distance to the origin of the anal fin. The anal fin contains three unbranched and branched rays, originating closer to the caudal-fin base than to the pelvic-fin insertion and extending to backward direction about to one-half distance to the caudal-fin base. Finally, the caudal fin is truncated or slightly notched at the tip (
|
||||
<figureCitation id="8588CFE01111FFB1FF03FE95280BFE12" box="[237,307,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1786,1810]" captionTargetBox="[184,1403,921,1763]" captionTargetId="figure-378@4.[184,1403,921,1763]" captionTargetPageId="4" captionText="FIGURE 2. Niwaella tigrina, JPIFS20230401, holotype, 55.7 mm SL, Ding-Jiang, Jiangxi Province, China. (a) lateral view, (b) dorsal view, (c) ventral view. Scale bar=10 mm." figureDoi="http://doi.org/10.5281/zenodo.14798044" httpUri="https://zenodo.org/record/14798044/files/figure.png" pageId="5" pageNumber="106">Fig. 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="49CC83ED1111FFB1FF79FC552C0CFC5C" ID-DOI="http://doi.org/10.5281/zenodo.14798046" ID-Zenodo-Dep="14798046" box="[151,1332,943,967]" httpUri="https://zenodo.org/record/14798046/files/figure.png" pageId="5" pageNumber="106" startId="5.[151,250,943,967]" targetBox="[303,1284,417,919]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="1D0CD3651111FFB1FF79FC552C0CFC5C" blockId="5.[151,1332,943,967]" box="[151,1332,943,967]" pageId="5" pageNumber="106">
|
||||
<emphasis id="2FC70F771111FFB1FF79FC55282BFC5C" bold="true" box="[151,275,943,967]" pageId="5" pageNumber="106">FIGURE 3.</emphasis>
|
||||
<taxonomicName id="DAB3A8E61111FFB1FEF7FC552B32FC5C" authority=", JPIFS" authorityName="JPIFS" box="[281,522,943,967]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771111FFB1FEF7FC5528F8FC5C" box="[281,448,943,967]" italics="true" pageId="5" pageNumber="106">Niwaella tigrina</emphasis>
|
||||
, JPIFS
|
||||
</taxonomicName>
|
||||
20230401, holotype. (a) mouth, (b) pre-dorsal scale removed from upper trunk.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="49CC83ED1111FFB1FF79FAAD2DE9FAF4" ID-DOI="http://doi.org/10.5281/zenodo.14798048" ID-Zenodo-Dep="14798048" box="[151,1233,1367,1392]" httpUri="https://zenodo.org/record/14798048/files/figure.png" pageId="5" pageNumber="106" startId="5.[151,250,1367,1391]" targetBox="[151,1435,1052,1343]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="1D0CD3651111FFB1FF79FAAD2DE9FAF4" blockId="5.[151,1233,1367,1392]" box="[151,1233,1367,1392]" pageId="5" pageNumber="106">
|
||||
<emphasis id="2FC70F771111FFB1FF79FAAD282BFAF4" bold="true" box="[151,275,1367,1391]" pageId="5" pageNumber="106">FIGURE 4.</emphasis>
|
||||
<taxonomicName id="DAB3A8E61111FFB1FEF7FAAD2B33FAF4" authority=", JPIFS" authorityName="JPIFS" box="[281,523,1367,1391]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771111FFB1FEF7FAAD28F8FAF4" box="[281,448,1367,1391]" italics="true" pageId="5" pageNumber="106">Niwaella tigrina</emphasis>
|
||||
, JPIFS
|
||||
</taxonomicName>
|
||||
20230401, holotype. (a) X-ray, (b) Micro-CT scan of suborbital spine.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="1D0CD3651111FFB1FF29FA3B2803F8FC" blockId="5.[151,1437,1473,2040]" pageId="5" pageNumber="106">
|
||||
<emphasis id="2FC70F771111FFB1FF29FA3B2877FA40" bold="true" box="[199,335,1473,1499]" pageId="5" pageNumber="106">Coloration.</emphasis>
|
||||
The formalin-preserved specimens are yellowish on the dorsal surface of the body, while lighter ventrally. The top of the head, opercle, and sub-opercle are grayish, with a touch of grayish-yellow, all sparsely scattered with black spots. The ventral side of the head is grayish-white. A black stripe extends from the nape, through the eye, to the insertion of the first rostral barbel. An indistinct blackish band advances from the posterior margin of the head to the caudal-fin base. Additionally, the dorsum and upper portion of the flank possess 20–25 transverse brown bars creating a tiger-striped pattern. Most of these brown bars extended downwards beyond a faint blackish band located just above the mid-lateral line of the body, reaching into the lower portion of the flank. However, the bars did not stretch to the ventral surface of the body. The dorsal fin exhibits two or three striations characterized by black spotting. The caudal fin has two or three striations running across the fin rays with distinct dark black spots on the upper portion of the caudal-fin base. A few blackish flecks are present on the distal portion of the pectoral fins, the proximal portion of the pelvic fins, and the basal portion, as well as on several leading rays of the anal fin.
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD3651111FFB1FF29F88B2C1CF810" blockId="5.[151,1437,1473,2040]" box="[199,1316,1905,1932]" pageId="5" pageNumber="106">
|
||||
<emphasis id="2FC70F771111FFB1FF29F88B288FF810" bold="true" box="[199,439,1905,1931]" pageId="5" pageNumber="106">Sexual dimorphism.</emphasis>
|
||||
No obvious signs of sexual dimorphism appeared in the external morphology.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="55A980EE1111FFB1FF29F86F2BC5F86C" pageId="5" pageNumber="106" type="distribution">
|
||||
<paragraph id="1D0CD3651111FFB1FF29F86F2BC5F86C" blockId="5.[151,1437,1473,2040]" pageId="5" pageNumber="106">
|
||||
<emphasis id="2FC70F771111FFB1FF29F86F28CCF834" bold="true" box="[199,500,1941,1968]" pageId="5" pageNumber="106">Distribution and habitat.</emphasis>
|
||||
<taxonomicName id="DAB3A8E61111FFB1FE12F86F2B8EF834" box="[508,694,1941,1967]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F771111FFB1FE12F86F2B8EF834" box="[508,694,1941,1967]" italics="true" pageId="5" pageNumber="106">Niwaella tigrina</emphasis>
|
||||
</taxonomicName>
|
||||
has been observed from Ding-Jiang and Jinsha-He, both flowing into Xiu-Shui, an affluent of Lake Poyang in
|
||||
<collectingRegion id="DF771D871111FFB1FD78F8432A6AF848" box="[662,850,1977,2003]" country="China" name="Jiangxi" pageId="5" pageNumber="106">Jiangxi Province</collectingRegion>
|
||||
of East
|
||||
<collectingCountry id="65A493F51111FFB1FC5EF8432ACBF848" box="[944,1011,1977,2003]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
|
||||
(
|
||||
<figureCitation id="8588CFE01111FFB1FBEDF8432D7CF84F" box="[1027,1092,1977,2004]" captionStart="FIGURE 1" captionStartId="1.[151,250,2013,2037]" captionTargetBox="[182,1418,283,1986]" captionTargetId="figure-69@1.[169,1418,270,1986]" captionTargetPageId="1" captionText="FIGURE 1. Type localities of all recognized species of Niwaella in East Asia." figureDoi="http://doi.org/10.5281/zenodo.14798041" httpUri="https://zenodo.org/record/14798041/files/figure.png" pageId="5" pageNumber="106">Fig. 1</figureCitation>
|
||||
). It prefers streams with clean water, strong currents, and gravelly bottoms as habitat.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="55A980EE1112FFB2FF29FF6D28C7FF62" pageId="6" pageNumber="107" type="etymology">
|
||||
<paragraph id="1D0CD3651112FFB2FF29FF6D28C7FF62" blockId="6.[151,1436,151,249]" pageId="6" pageNumber="107">
|
||||
<emphasis id="2FC70F771112FFB2FF29FF6D2876FF2A" bold="true" box="[199,334,151,177]" pageId="6" pageNumber="107">Etymology.</emphasis>
|
||||
The epithet, used as an adjective, is derived from the Latin word
|
||||
<emphasis id="2FC70F771112FFB2FBF1FF6D2D4EFF2B" box="[1055,1142,151,176]" italics="true" pageId="6" pageNumber="107">tigrinus</emphasis>
|
||||
(= tiger-striped), referring to the presence of tiger stripes on the dorsum and upper part of the flank. The Chinese name here suggested for this new species is“
|
||||
<emphasis id="2FC70F771112FFB2FEAAFF2528D4FF6D" bold="true" box="[324,492,223,246]" pageId="6" pageNumber="107">njử后ǾAEƟ</emphasis>
|
||||
”.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="49CC83ED1112FFB2FF79FED22AC3FEDB" box="[151,1019,296,320]" pageId="6" pageNumber="107" startId="6.[151,239,296,320]" targetBox="[163,1338,336,1568]" targetIsTable="true" targetPageId="6" targetType="table">
|
||||
<paragraph id="1D0CD3651112FFB2FF79FED22AC3FEDB" blockId="6.[151,1019,296,320]" box="[151,1019,296,320]" pageId="6" pageNumber="107">
|
||||
<emphasis id="2FC70F771112FFB2FF79FED22832FEDB" bold="true" box="[151,266,296,320]" pageId="6" pageNumber="107">TABLE 2.</emphasis>
|
||||
Morphometric measurements and meristic counts of
|
||||
<taxonomicName id="DAB3A8E61112FFB2FCAAFED22ACFFEDB" box="[836,1015,296,320]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="tigrina">Niwaella tigrina</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="1D0CD3651112FFB2FF4DFE9F2D12F9BB" pageId="6" pageNumber="107">
|
||||
<table id="6FB321C51112004BFF4DFEAA2C02F9BB" box="[163,1338,336,1568]" gridcols="4" gridrows="30" pageId="6" pageNumber="107">
|
||||
<tr id="A383D1271112004BFF4DFEAA2C02FE0A" box="[163,1338,336,401]" gridrow="0" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFEAA2BF9FE0A" box="[163,705,336,401]" gridcol="0" gridrow="0" pageId="6" pageNumber="107">Morphometric measurements</th>
|
||||
<th id="E052B85B1112004BFCE9FEAA2A52FE0A" box="[775,874,336,401]" gridcol="1" gridrow="0" pageId="6" pageNumber="107">
|
||||
<emphasis id="2FC70F771112FFB2FCE9FE9F2A52FEE6" bold="true" box="[775,874,357,381]" pageId="6" pageNumber="107">Holotype</emphasis>
|
||||
</th>
|
||||
<th id="E052B85B1112004BFC34FEAA2DABFE0A" box="[986,1171,336,401]" gridcol="2" gridrow="0" pageId="6" pageNumber="107">
|
||||
<emphasis id="2FC70F771112FFB2FC34FEAA2DABFEF3" bold="true" box="[986,1171,336,360]" pageId="6" pageNumber="107">Paratypes (n=19)</emphasis>
|
||||
Range
|
||||
</th>
|
||||
<th id="E052B85B1112004BFB2BFEAA2C02FE0A" box="[1221,1338,336,401]" gridcol="3" gridrow="0" pageId="6" pageNumber="107">Mean ± SD</th>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFE5B2C02FE22" box="[163,1338,417,441]" gridrow="1" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFE5B2BF9FE22" box="[163,705,417,441]" gridcol="0" gridrow="1" pageId="6" pageNumber="107">Standard length (SL, mm)</th>
|
||||
<td id="E052B85B1112004BFCE9FE5B2A52FE22" box="[775,874,417,441]" gridcol="1" gridrow="1" pageId="6" pageNumber="107">55.7</td>
|
||||
<td id="E052B85B1112004BFC34FE5B2DABFE22" box="[986,1171,417,441]" gridcol="2" gridrow="1" pageId="6" pageNumber="107">46.8-63.9</td>
|
||||
<td id="E052B85B1112004BFB2BFE5B2C02FE22" box="[1221,1338,417,441]" gridcol="3" gridrow="1" pageId="6" pageNumber="107">55.1±4.6</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFE332C02FE7A" box="[163,1338,457,481]" gridrow="2" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFE332C02FE7A" box="[163,1338,457,481]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="6" pageNumber="107">In percents of SL(%)</th>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFE0B2C02FD92" box="[163,1338,497,521]" gridrow="3" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFE0B2BF9FD92" box="[163,705,497,521]" gridcol="0" gridrow="3" pageId="6" pageNumber="107">Head length (HL)</th>
|
||||
<td id="E052B85B1112004BFCE9FE0B2A52FD92" box="[775,874,497,521]" gridcol="1" gridrow="3" pageId="6" pageNumber="107">17.3</td>
|
||||
<td id="E052B85B1112004BFC34FE0B2DABFD92" box="[986,1171,497,521]" gridcol="2" gridrow="3" pageId="6" pageNumber="107">16.1-18.3</td>
|
||||
<td id="E052B85B1112004BFB2BFE0B2C02FD92" box="[1221,1338,497,521]" gridcol="3" gridrow="3" pageId="6" pageNumber="107">16.8±0.7</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFDE32C02FDA9" box="[163,1338,537,562]" gridrow="4" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFDE32BF9FDA9" box="[163,705,537,562]" gridcol="0" gridrow="4" pageId="6" pageNumber="107">Body depth</th>
|
||||
<td id="E052B85B1112004BFCE9FDE32A52FDA9" box="[775,874,537,562]" gridcol="1" gridrow="4" pageId="6" pageNumber="107">11.8</td>
|
||||
<td id="E052B85B1112004BFC34FDE32DABFDA9" box="[986,1171,537,562]" gridcol="2" gridrow="4" pageId="6" pageNumber="107">11.1-12.8</td>
|
||||
<td id="E052B85B1112004BFB2BFDE32C02FDA9" box="[1221,1338,537,562]" gridcol="3" gridrow="4" pageId="6" pageNumber="107">12.1±0.6</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFDB82C02FDC1" box="[163,1338,578,602]" gridrow="5" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFDB82BF9FDC1" box="[163,705,578,602]" gridcol="0" gridrow="5" pageId="6" pageNumber="107">Distance between pectoral fin and pelvic fin insertion</th>
|
||||
<td id="E052B85B1112004BFCE9FDB82A52FDC1" box="[775,874,578,602]" gridcol="1" gridrow="5" pageId="6" pageNumber="107">32.7</td>
|
||||
<td id="E052B85B1112004BFC34FDB82DABFDC1" box="[986,1171,578,602]" gridcol="2" gridrow="5" pageId="6" pageNumber="107">32.7-38.5</td>
|
||||
<td id="E052B85B1112004BFB2BFDB82C02FDC1" box="[1221,1338,578,602]" gridcol="3" gridrow="5" pageId="6" pageNumber="107">34.7±1.5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFD902C02FD19" box="[163,1338,618,642]" gridrow="6" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFD902BF9FD19" box="[163,705,618,642]" gridcol="0" gridrow="6" pageId="6" pageNumber="107">Caudal-peduncle length</th>
|
||||
<td id="E052B85B1112004BFCE9FD902A52FD19" box="[775,874,618,642]" gridcol="1" gridrow="6" pageId="6" pageNumber="107">14.7</td>
|
||||
<td id="E052B85B1112004BFC34FD902DABFD19" box="[986,1171,618,642]" gridcol="2" gridrow="6" pageId="6" pageNumber="107">13.8-15.2</td>
|
||||
<td id="E052B85B1112004BFB2BFD902C02FD19" box="[1221,1338,618,642]" gridcol="3" gridrow="6" pageId="6" pageNumber="107">15.0±0.7</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFD682C02FD31" box="[163,1338,658,682]" gridrow="7" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFD682BF9FD31" box="[163,705,658,682]" gridcol="0" gridrow="7" pageId="6" pageNumber="107">Caudal-peduncle depth</th>
|
||||
<td id="E052B85B1112004BFCE9FD682A52FD31" box="[775,874,658,682]" gridcol="1" gridrow="7" pageId="6" pageNumber="107">7.8</td>
|
||||
<td id="E052B85B1112004BFC34FD682DABFD31" box="[986,1171,658,682]" gridcol="2" gridrow="7" pageId="6" pageNumber="107">7.3-8.8</td>
|
||||
<td id="E052B85B1112004BFB2BFD682C02FD31" box="[1221,1338,658,682]" gridcol="3" gridrow="7" pageId="6" pageNumber="107">8.2±0.4</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFD402C02FD49" box="[163,1338,698,722]" gridrow="8" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFD402BF9FD49" box="[163,705,698,722]" gridcol="0" gridrow="8" pageId="6" pageNumber="107">Dorsal-fin length</th>
|
||||
<td id="E052B85B1112004BFCE9FD402A52FD49" box="[775,874,698,722]" gridcol="1" gridrow="8" pageId="6" pageNumber="107">14.5</td>
|
||||
<td id="E052B85B1112004BFC34FD402DABFD49" box="[986,1171,698,722]" gridcol="2" gridrow="8" pageId="6" pageNumber="107">13.0-15.7</td>
|
||||
<td id="E052B85B1112004BFB2BFD402C02FD49" box="[1221,1338,698,722]" gridcol="3" gridrow="8" pageId="6" pageNumber="107">14.2±0.6</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFD192C02FD60" box="[163,1338,739,763]" gridrow="9" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFD192BF9FD60" box="[163,705,739,763]" gridcol="0" gridrow="9" pageId="6" pageNumber="107">Pectoral-fin length</th>
|
||||
<td id="E052B85B1112004BFCE9FD192A52FD60" box="[775,874,739,763]" gridcol="1" gridrow="9" pageId="6" pageNumber="107">9.8</td>
|
||||
<td id="E052B85B1112004BFC34FD192DABFD60" box="[986,1171,739,763]" gridcol="2" gridrow="9" pageId="6" pageNumber="107">9.8-11.5</td>
|
||||
<td id="E052B85B1112004BFB2BFD192C02FD60" box="[1221,1338,739,763]" gridcol="3" gridrow="9" pageId="6" pageNumber="107">10.1±0.5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFCF12C02FCB8" box="[163,1338,779,803]" gridrow="10" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFCF12BF9FCB8" box="[163,705,779,803]" gridcol="0" gridrow="10" pageId="6" pageNumber="107">Pelvic-fin length</th>
|
||||
<td id="E052B85B1112004BFCE9FCF12A52FCB8" box="[775,874,779,803]" gridcol="1" gridrow="10" pageId="6" pageNumber="107">9.6</td>
|
||||
<td id="E052B85B1112004BFC34FCF12DABFCB8" box="[986,1171,779,803]" gridcol="2" gridrow="10" pageId="6" pageNumber="107">8.7-10.0</td>
|
||||
<td id="E052B85B1112004BFB2BFCF12C02FCB8" box="[1221,1338,779,803]" gridcol="3" gridrow="10" pageId="6" pageNumber="107">9.4±0.4</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFCC92C02FCD0" box="[163,1338,819,843]" gridrow="11" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFCC92BF9FCD0" box="[163,705,819,843]" gridcol="0" gridrow="11" pageId="6" pageNumber="107">Anal-fin length</th>
|
||||
<td id="E052B85B1112004BFCE9FCC92A52FCD0" box="[775,874,819,843]" gridcol="1" gridrow="11" pageId="6" pageNumber="107">11.1</td>
|
||||
<td id="E052B85B1112004BFC34FCC92DABFCD0" box="[986,1171,819,843]" gridcol="2" gridrow="11" pageId="6" pageNumber="107">9.8-11.6</td>
|
||||
<td id="E052B85B1112004BFB2BFCC92C02FCD0" box="[1221,1338,819,843]" gridcol="3" gridrow="11" pageId="6" pageNumber="107">10.8±0.8</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFCA12C02FCEF" box="[163,1338,859,884]" gridrow="12" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFCA12BF9FCEF" box="[163,705,859,884]" gridcol="0" gridrow="12" pageId="6" pageNumber="107">Caudal-fin length</th>
|
||||
<td id="E052B85B1112004BFCE9FCA12A52FCEF" box="[775,874,859,884]" gridcol="1" gridrow="12" pageId="6" pageNumber="107">13.8</td>
|
||||
<td id="E052B85B1112004BFC34FCA12DABFCEF" box="[986,1171,859,884]" gridcol="2" gridrow="12" pageId="6" pageNumber="107">13.1-15.6</td>
|
||||
<td id="E052B85B1112004BFB2BFCA12C02FCEF" box="[1221,1338,859,884]" gridcol="3" gridrow="12" pageId="6" pageNumber="107">13.9±0.7</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFC7E2C02FC07" box="[163,1338,900,924]" gridrow="13" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFC7E2BF9FC07" box="[163,705,900,924]" gridcol="0" gridrow="13" pageId="6" pageNumber="107">Predorsal length</th>
|
||||
<td id="E052B85B1112004BFCE9FC7E2A52FC07" box="[775,874,900,924]" gridcol="1" gridrow="13" pageId="6" pageNumber="107">53.1</td>
|
||||
<td id="E052B85B1112004BFC34FC7E2DABFC07" box="[986,1171,900,924]" gridcol="2" gridrow="13" pageId="6" pageNumber="107">51.2-55.5</td>
|
||||
<td id="E052B85B1112004BFB2BFC7E2C02FC07" box="[1221,1338,900,924]" gridcol="3" gridrow="13" pageId="6" pageNumber="107">54.0±1.5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFC562C02FC5F" box="[163,1338,940,964]" gridrow="14" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFC562BF9FC5F" box="[163,705,940,964]" gridcol="0" gridrow="14" pageId="6" pageNumber="107">Prepelvic length</th>
|
||||
<td id="E052B85B1112004BFCE9FC562A52FC5F" box="[775,874,940,964]" gridcol="1" gridrow="14" pageId="6" pageNumber="107">53.1</td>
|
||||
<td id="E052B85B1112004BFC34FC562DABFC5F" box="[986,1171,940,964]" gridcol="2" gridrow="14" pageId="6" pageNumber="107">52.3-56.1</td>
|
||||
<td id="E052B85B1112004BFB2BFC562C02FC5F" box="[1221,1338,940,964]" gridcol="3" gridrow="14" pageId="6" pageNumber="107">54.7±1.5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFC2E2C02FC77" box="[163,1338,980,1004]" gridrow="15" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFC2E2BF9FC77" box="[163,705,980,1004]" gridcol="0" gridrow="15" pageId="6" pageNumber="107">Preanal length</th>
|
||||
<td id="E052B85B1112004BFCE9FC2E2A52FC77" box="[775,874,980,1004]" gridcol="1" gridrow="15" pageId="6" pageNumber="107">80.9</td>
|
||||
<td id="E052B85B1112004BFC34FC2E2DABFC77" box="[986,1171,980,1004]" gridcol="2" gridrow="15" pageId="6" pageNumber="107">80.8-84.4</td>
|
||||
<td id="E052B85B1112004BFB2BFC2E2C02FC77" box="[1221,1338,980,1004]" gridcol="3" gridrow="15" pageId="6" pageNumber="107">82.1±1.4</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFC062C02FB8E" box="[163,1338,1020,1045]" gridrow="16" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFC062BF9FB8E" box="[163,705,1020,1045]" gridcol="0" gridrow="16" pageId="6" pageNumber="107">Dorsal-fin base length</th>
|
||||
<td id="E052B85B1112004BFCE9FC062A52FB8E" box="[775,874,1020,1045]" gridcol="1" gridrow="16" pageId="6" pageNumber="107">8.9</td>
|
||||
<td id="E052B85B1112004BFC34FC062DABFB8E" box="[986,1171,1020,1045]" gridcol="2" gridrow="16" pageId="6" pageNumber="107">7.6-9.4</td>
|
||||
<td id="E052B85B1112004BFB2BFC062C02FB8E" box="[1221,1338,1020,1045]" gridcol="3" gridrow="16" pageId="6" pageNumber="107">8.6±0.5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFBDE2C02FBA6" box="[163,1338,1060,1085]" gridrow="17" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFBDE2BF9FBA6" box="[163,705,1060,1085]" gridcol="0" gridrow="17" pageId="6" pageNumber="107">Anal-fin base length</th>
|
||||
<td id="E052B85B1112004BFCE9FBDE2A52FBA6" box="[775,874,1060,1085]" gridcol="1" gridrow="17" pageId="6" pageNumber="107">6.6</td>
|
||||
<td id="E052B85B1112004BFC34FBDE2DABFBA6" box="[986,1171,1060,1085]" gridcol="2" gridrow="17" pageId="6" pageNumber="107">5.3-6.8</td>
|
||||
<td id="E052B85B1112004BFB2BFBDE2C02FBA6" box="[1221,1338,1060,1085]" gridcol="3" gridrow="17" pageId="6" pageNumber="107">5.9±0.4</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFBB72C02FBFE" box="[163,1338,1101,1125]" gridrow="18" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFBB72C02FBFE" box="[163,1338,1101,1125]" colspan="4" colspanRight="3" gridcol="0" gridrow="18" pageId="6" pageNumber="107">In percents of HL(%)</th>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFB8F2C02FB16" box="[163,1338,1141,1165]" gridrow="19" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFB8F2BF9FB16" box="[163,705,1141,1165]" gridcol="0" gridrow="19" pageId="6" pageNumber="107">Snout length</th>
|
||||
<td id="E052B85B1112004BFCE9FB8F2A52FB16" box="[775,874,1141,1165]" gridcol="1" gridrow="19" pageId="6" pageNumber="107">46.8</td>
|
||||
<td id="E052B85B1112004BFC34FB8F2DABFB16" box="[986,1171,1141,1165]" gridcol="2" gridrow="19" pageId="6" pageNumber="107">46.8-52.7</td>
|
||||
<td id="E052B85B1112004BFB2BFB8F2C02FB16" box="[1221,1338,1141,1165]" gridcol="3" gridrow="19" pageId="6" pageNumber="107">48.8±1.5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFB672C02FB2D" box="[163,1338,1181,1206]" gridrow="20" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFB672BF9FB2D" box="[163,705,1181,1206]" gridcol="0" gridrow="20" pageId="6" pageNumber="107">Eye diameter</th>
|
||||
<td id="E052B85B1112004BFCE9FB672A52FB2D" box="[775,874,1181,1206]" gridcol="1" gridrow="20" pageId="6" pageNumber="107">17.9</td>
|
||||
<td id="E052B85B1112004BFC34FB672DABFB2D" box="[986,1171,1181,1206]" gridcol="2" gridrow="20" pageId="6" pageNumber="107">16.4-20.5</td>
|
||||
<td id="E052B85B1112004BFB2BFB672C02FB2D" box="[1221,1338,1181,1206]" gridcol="3" gridrow="20" pageId="6" pageNumber="107">17.6±1.6</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFB3C2C02FB45" box="[163,1338,1222,1246]" gridrow="21" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFB3C2BF9FB45" box="[163,705,1222,1246]" gridcol="0" gridrow="21" pageId="6" pageNumber="107">Interorbital width</th>
|
||||
<td id="E052B85B1112004BFCE9FB3C2A52FB45" box="[775,874,1222,1246]" gridcol="1" gridrow="21" pageId="6" pageNumber="107">16.4</td>
|
||||
<td id="E052B85B1112004BFC34FB3C2DABFB45" box="[986,1171,1222,1246]" gridcol="2" gridrow="21" pageId="6" pageNumber="107">11.1-18.3</td>
|
||||
<td id="E052B85B1112004BFB2BFB3C2C02FB45" box="[1221,1338,1222,1246]" gridcol="3" gridrow="21" pageId="6" pageNumber="107">14.8±1.8</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFB142C02FA9D" box="[163,1338,1262,1286]" gridrow="22" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFB142BF9FA9D" box="[163,705,1262,1286]" gridcol="0" gridrow="22" pageId="6" pageNumber="107">CPL/CPD</th>
|
||||
<td id="E052B85B1112004BFCE9FB142A52FA9D" box="[775,874,1262,1286]" gridcol="1" gridrow="22" pageId="6" pageNumber="107">1.8</td>
|
||||
<td id="E052B85B1112004BFC34FB142DABFA9D" box="[986,1171,1262,1286]" gridcol="2" gridrow="22" pageId="6" pageNumber="107">1.7-2.1</td>
|
||||
<td id="E052B85B1112004BFB2BFB142C02FA9D" box="[1221,1338,1262,1286]" gridcol="3" gridrow="22" pageId="6" pageNumber="107">1.8±0.1</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFAEC2C02FAB5" box="[163,1338,1302,1326]" gridrow="23" pageId="6" pageNumber="107">
|
||||
<th id="E052B85B1112004BFF4DFAEC2C02FAB5" box="[163,1338,1302,1326]" colspan="4" colspanRight="3" gridcol="0" gridrow="23" pageId="6" pageNumber="107">Meristic counts</th>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFAC42C02FACC" box="[163,1338,1342,1367]" gridrow="24" pageId="6" pageNumber="107" rowspan-3="1">
|
||||
<th id="E052B85B1112004BFF4DFAC42BF9FACC" box="[163,705,1342,1367]" gridcol="0" gridrow="24" pageId="6" pageNumber="107">Dorsal-fin spines and rays</th>
|
||||
<td id="E052B85B1112004BFCE9FAC42A52FACC" box="[775,874,1342,1367]" gridcol="1" gridrow="24" pageId="6" pageNumber="107">ii-5</td>
|
||||
<td id="E052B85B1112004BFC34FAC42DABFACC" box="[986,1171,1342,1367]" gridcol="2" gridrow="24" pageId="6" pageNumber="107">ii-5</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFA9D2C02FAE4" box="[163,1338,1383,1407]" gridrow="25" pageId="6" pageNumber="107" rowspan-3="1">
|
||||
<th id="E052B85B1112004BFF4DFA9D2BF9FAE4" box="[163,705,1383,1407]" gridcol="0" gridrow="25" pageId="6" pageNumber="107">Anal-fin spines and rays</th>
|
||||
<td id="E052B85B1112004BFCE9FA9D2A52FAE4" box="[775,874,1383,1407]" gridcol="1" gridrow="25" pageId="6" pageNumber="107">i-6</td>
|
||||
<td id="E052B85B1112004BFC34FA9D2DABFAE4" box="[986,1171,1383,1407]" gridcol="2" gridrow="25" pageId="6" pageNumber="107">i-6</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFA752C02FA3C" box="[163,1338,1423,1447]" gridrow="26" pageId="6" pageNumber="107" rowspan-3="1">
|
||||
<th id="E052B85B1112004BFF4DFA752BF9FA3C" box="[163,705,1423,1447]" gridcol="0" gridrow="26" pageId="6" pageNumber="107">Pectoral-fin spines and rays</th>
|
||||
<td id="E052B85B1112004BFCE9FA752A52FA3C" box="[775,874,1423,1447]" gridcol="1" gridrow="26" pageId="6" pageNumber="107">i-7</td>
|
||||
<td id="E052B85B1112004BFC34FA752DABFA3C" box="[986,1171,1423,1447]" gridcol="2" gridrow="26" pageId="6" pageNumber="107">i-6~7</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFA4D2C02FA4B" box="[163,1338,1463,1488]" gridrow="27" pageId="6" pageNumber="107" rowspan-3="1">
|
||||
<th id="E052B85B1112004BFF4DFA4D2BF9FA4B" box="[163,705,1463,1488]" gridcol="0" gridrow="27" pageId="6" pageNumber="107">Caudal-fin spines and rays</th>
|
||||
<td id="E052B85B1112004BFCE9FA4D2A52FA4B" box="[775,874,1463,1488]" gridcol="1" gridrow="27" pageId="6" pageNumber="107">iv-14-iv</td>
|
||||
<td id="E052B85B1112004BFC34FA4D2DABFA4B" box="[986,1171,1463,1488]" gridcol="2" gridrow="27" pageId="6" pageNumber="107">iv-14-16-iv</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DFA252C02FA63" box="[163,1338,1503,1528]" gridrow="28" pageId="6" pageNumber="107" rowspan-3="1">
|
||||
<th id="E052B85B1112004BFF4DFA252BF9FA63" box="[163,705,1503,1528]" gridcol="0" gridrow="28" pageId="6" pageNumber="107">Pelvic-fin spines and rays</th>
|
||||
<td id="E052B85B1112004BFCE9FA252A52FA63" box="[775,874,1503,1528]" gridcol="1" gridrow="28" pageId="6" pageNumber="107">i-6</td>
|
||||
<td id="E052B85B1112004BFC34FA252DABFA63" box="[986,1171,1503,1528]" gridcol="2" gridrow="28" pageId="6" pageNumber="107">i-6</td>
|
||||
</tr>
|
||||
<tr id="A383D1271112004BFF4DF9F22C02F9BB" box="[163,1338,1544,1568]" gridrow="29" pageId="6" pageNumber="107" rowspan-3="1">
|
||||
<th id="E052B85B1112004BFF4DF9F22BF9F9BB" box="[163,705,1544,1568]" gridcol="0" gridrow="29" pageId="6" pageNumber="107">Vertebrae</th>
|
||||
<td id="E052B85B1112004BFCE9F9F22A52F9BB" box="[775,874,1544,1568]" gridcol="1" gridrow="29" pageId="6" pageNumber="107">4+41+1</td>
|
||||
<td id="E052B85B1112004BFC34F9F22DABF9BB" box="[986,1171,1544,1568]" gridcol="2" gridrow="29" pageId="6" pageNumber="107">4+41+1</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<subSubSection id="55A980EE1112FFB2FF79F9892B5BF84A" pageId="6" pageNumber="107" type="multiple">
|
||||
<paragraph id="1D0CD3651112FFB2FF79F9892BCDF916" blockId="6.[151,757,1651,1677]" box="[151,757,1651,1677]" pageId="6" pageNumber="107">
|
||||
<heading id="464464091112FFB2FF79F9892BCDF916" bold="true" box="[151,757,1651,1677]" fontSize="11" level="1" pageId="6" pageNumber="107" reason="1">
|
||||
<emphasis id="2FC70F771112FFB2FF79F9892BCDF916" bold="true" box="[151,757,1651,1677]" pageId="6" pageNumber="107">3.2 Sequence variation and molecular phylogeny</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD3651112FFB2FF79F94128AFF8D9" blockId="6.[151,1437,1723,2002]" pageId="6" pageNumber="107">A total length of 1, 023 bp was retained after trimming and used as subsequent alignment. This included 1020 conserved sites, three variable sites, zero parsimony in formative sites, and three singleton sites. The mean nucleotide frequencies of the described species were: A=26.3%, T=32.6%, C=26.4%, and G=14.8%, with an A-T rich base composition of 58.8%.</paragraph>
|
||||
<paragraph id="1D0CD3651112FFB2FF29F8B12B5BF84A" blockId="6.[151,1437,1723,2002]" pageId="6" pageNumber="107">
|
||||
The phylogenetic tree constructed using the Cyt
|
||||
<emphasis id="2FC70F771112FFB2FD1AF8B12A3AF8FE" box="[756,770,1867,1893]" italics="true" pageId="6" pageNumber="107">b</emphasis>
|
||||
gene for all the analyzed
|
||||
<taxonomicName id="DAB3A8E61112FFB2FBC1F8B12DD6F8FE" box="[1071,1262,1867,1893]" pageId="6" pageNumber="107">
|
||||
<emphasis id="2FC70F771112FFB2FBC1F8B12DACF8FE" box="[1071,1172,1867,1893]" italics="true" pageId="6" pageNumber="107">Niwaella</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
is presented in
|
||||
<figureCitation id="8588CFE01112FFB2FF79F89529E2F811" box="[151,218,1903,1930]" captionStart="FIGURE 5" captionStartId="7.[151,250,1148,1172]" captionTargetBox="[198,1385,189,1113]" captionTargetId="figure-20@7.[190,1398,181,1124]" captionTargetPageId="7" captionText="FIGURE 5. The Cyt b gene-based ML tree for Niwaella tigrina and 29 related species. The genetic distances among all sampled species of Niwaella, calculated by utilizing the Kimura 2-Parameter Distance Model, are in Table 3. The intraspecific genetic distance of N. tigrina was zero. In contrast, the interspecific genetic divergence of this new species compared to all Chinese congeneric species ranged from 4.2% to 6.4%." figureDoi="http://doi.org/10.5281/zenodo.14798050" httpUri="https://zenodo.org/record/14798050/files/figure.png" pageId="6" pageNumber="107">Fig. 5</figureCitation>
|
||||
, based on ML methods. In the topology of these trees, six samples of the newly described species exclusively clustered into a single lineage. These samples were closely related to those of other Chinese
|
||||
<taxonomicName id="DAB3A8E61112FFB2FB61F8692C76F836" box="[1167,1358,1939,1965]" pageId="6" pageNumber="107">
|
||||
<emphasis id="2FC70F771112FFB2FB61F8692DCCF836" box="[1167,1268,1939,1965]" italics="true" pageId="6" pageNumber="107">Niwaella</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
, which collectively formed a well-defined clade.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
177
data/95/1A/62/951A6273111EFFBEFF79FF6D2CA5FD7F.xml
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177
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@ -0,0 +1,177 @@
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<mods:title id="4BB9910E80C5E35A948287BEEE9ED37F">Niwaella tigrina, a new species of spined loach from the Poyang Lake system of Jiangxi Province, East China</mods:title>
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<paragraph id="1D0CD365111EFFBEFF79FF6D2B7DFF29" blockId="10.[151,581,151,178]" box="[151,581,151,178]" pageId="10" pageNumber="111">
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Key to Chinese species of
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<taxonomicName id="DAB3A8E6111EFFBEFE37FF6D2B7DFF29" ID-CoL="64JB" authorityName="Nalbant" authorityYear="1963" box="[473,581,151,178]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="genus">
|
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<emphasis id="2FC70F77111EFFBEFE37FF6D2B7DFF29" bold="true" box="[473,581,151,178]" italics="true" pageId="10" pageNumber="111">Niwaella</emphasis>
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|
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<paragraph id="1D0CD365111EFFBEFF79FF262CA4FF6A" blockId="10.[151,1437,220,741]" box="[151,1436,220,242]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FF262CA4FF6A" box="[151,1436,220,242]" pageId="10" pageNumber="111">1a. Lower jaw unexposed or fully covered by lower lip.......................................................... 2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FF032CA4FE94" blockId="10.[151,1437,220,741]" box="[151,1436,249,271]" pageId="10" pageNumber="111">
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||||
<keyLead id="A6427050111EFFBEFF79FF032CA4FE94" box="[151,1436,249,271]" pageId="10" pageNumber="111">1b. Lower jaw exposed or separated from lower lip............................................................. 4</keyLead>
|
||||
</paragraph>
|
||||
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|
||||
<keyStep id="A647CBC0111EFFBEFF79FEED2CA4FED2" pageId="10" pageNumber="111">
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||||
<keyLead id="A6427050111EFFBEFF79FEED2CA4FEB7" box="[151,1436,279,301]" pageId="10" pageNumber="111">
|
||||
2a. Transverse black bars on dorsum and upper part of flank (Ding-Jiang)....................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFAABFEED2CA4FEB7" authorityName="Zhang & Zhang & Chen & Zhang" authorityYear="2025" box="[1349,1436,279,300]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="tigrina">
|
||||
<emphasis id="2FC70F77111EFFBEFAABFEED2CA4FEB7" box="[1349,1436,279,300]" italics="true" pageId="10" pageNumber="111">N.tigrina</emphasis>
|
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</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FECE2CA4FED2" blockId="10.[151,1437,220,741]" box="[151,1436,308,330]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FECE2CA4FED2" box="[151,1436,308,330]" pageId="10" pageNumber="111">2b. A row of transverse black blotches on dorsum and vermicular black marks on upper part of flank...................... 3</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="A647CBC0111EFFBEFF79FEA82CA5FE1F" pageId="10" pageNumber="111">
|
||||
<paragraph id="1D0CD365111EFFBEFF79FEA82CA5FEFD" blockId="10.[151,1437,220,741]" box="[151,1437,337,359]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FEA82CA5FEFD" box="[151,1437,337,359]" pageId="10" pageNumber="111">
|
||||
3a. Four branched anal-fin rays (Dongtiao-Xi)........................................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFAF1FEAB2CA5FEFD" authorityName="Chen & Chen" authorityYear="2017" box="[1311,1437,337,358]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="brevipinna">
|
||||
<emphasis id="2FC70F77111EFFBEFAF1FEAB2CA5FEFD" box="[1311,1437,337,358]" italics="true" pageId="10" pageNumber="111">N.brevipinna</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FE952CA5FE1F" blockId="10.[151,1437,220,741]" box="[151,1437,367,389]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FE952CA5FE1F" box="[151,1437,367,389]" pageId="10" pageNumber="111">
|
||||
3b. Six branched anal-fin rays (Xin-Jiang).........................................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFAECFE952CA5FE1F" ID-CoL="47K98" box="[1282,1437,367,388]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="xinjiangensis">
|
||||
<emphasis id="2FC70F77111EFFBEFAECFE952CA5FE1F" box="[1282,1437,367,388]" italics="true" pageId="10" pageNumber="111">N. xinjiangensis</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="A647CBC0111EFFBEFF79FE762CA4FE61" pageId="10" pageNumber="111">
|
||||
<paragraph id="1D0CD365111EFFBEFF79FE762CA4FE24" blockId="10.[151,1437,220,741]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FE762CA4FE24" pageId="10" pageNumber="111">4a. A distinct blackish mid-dorsal stripe from behind head to caudal-fin basin, and 8–10 brownish or black vertical bars on lower part of flank......................................................................................... 5</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FE3D2CA4FE61" blockId="10.[151,1437,220,741]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FE3D2CA4FE61" pageId="10" pageNumber="111">4b. An indistinct blackish mid-dorsal stripe from behind head to caudal-fin basin, and more no fewer than 15 brownish or black vertical bars on lower part of flank........................................................................ 6</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="A647CBC0111EFFBEFF79FDFB2CA4FDF5" pageId="10" pageNumber="111">
|
||||
<paragraph id="1D0CD365111EFFBEFF79FDFB2CA4FDAF" blockId="10.[151,1437,220,741]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FDFB2CA4FDAF" pageId="10" pageNumber="111">
|
||||
5a. Suborbital spine straight, with shorter processus latero-caudalis less than one-fifth of processus medio-caudalis (Dongtiao-Xi)..........................................................................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFAC9FDE52CA4FDAF" authorityName="Chen & Chen" authorityYear="2017" box="[1319,1436,543,564]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="fimbriata">
|
||||
<emphasis id="2FC70F77111EFFBEFAC9FDE52CA4FDAF" box="[1319,1436,543,564]" italics="true" pageId="10" pageNumber="111">N. fimbriata</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FDC62CA4FDF5" blockId="10.[151,1437,220,741]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FDC62CA4FDF5" pageId="10" pageNumber="111">
|
||||
5b. Suborbital spine curved, with longer processus latero-caudalis more than one-third of processus medio-caudalis (Shuai-Shui)...........................................................................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFACDFDA32CA4FDF5" authorityName="Chen & Chen" authorityYear="2017" box="[1315,1436,601,622]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="nigrolinea">
|
||||
<emphasis id="2FC70F77111EFFBEFACDFDA32CA4FDF5" box="[1315,1436,601,622]" italics="true" pageId="10" pageNumber="111">N.nigrolinea</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="A647CBC0111EFFBEFF79FD8D2CA4FD31" pageId="10" pageNumber="111">
|
||||
<paragraph id="1D0CD365111EFFBEFF79FD8D2CA4FD17" blockId="10.[151,1437,220,741]" box="[151,1436,631,653]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FD8D2CA4FD17" box="[151,1436,631,653]" pageId="10" pageNumber="111">
|
||||
6a. Seven branched dorsal-fin rays (Qu-Jiang)......................................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFAE9FD8D2CA4FD17" box="[1287,1436,631,652]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="qujiangensis">
|
||||
<emphasis id="2FC70F77111EFFBEFAE9FD8D2CA4FD17" box="[1287,1436,631,652]" italics="true" pageId="10" pageNumber="111">N. qujiangensis</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FD6E2CA4FD31" blockId="10.[151,1437,220,741]" box="[151,1436,660,682]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FD6E2CA4FD31" box="[151,1436,660,682]" pageId="10" pageNumber="111">6b. Six branched dorsal-fin rays............................................................................ 7</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="A647CBC0111EFFBEFF79FD482CA5FD7F" pageId="10" pageNumber="111">
|
||||
<paragraph id="1D0CD365111EFFBEFF79FD482CA4FD5D" blockId="10.[151,1437,220,741]" box="[151,1436,689,711]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FD482CA4FD5D" box="[151,1436,689,711]" pageId="10" pageNumber="111">
|
||||
7a. Barbels shorter than eye diameter(Yong-Jiang).................................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFB1DFD482CA4FD5D" ID-CoL="47K95" baseAuthorityName="Yan & Zheng" baseAuthorityYear="1984" box="[1267,1436,689,711]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="laterimaculata">
|
||||
<emphasis id="2FC70F77111EFFBEFB1DFD482CA4FD5D" box="[1267,1436,689,711]" italics="true" pageId="10" pageNumber="111">N. laterimaculata</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="1D0CD365111EFFBEFF79FD352CA5FD7F" blockId="10.[151,1437,220,741]" box="[151,1437,719,741]" pageId="10" pageNumber="111">
|
||||
<keyLead id="A6427050111EFFBEFF79FD352CA5FD7F" box="[151,1437,719,741]" pageId="10" pageNumber="111">
|
||||
7b. Barbels longer than or equal to eye diameter (Cao’e-Jiang)..........................................
|
||||
<taxonomicName id="DAB3A8E6111EFFBEFAF8FD352CA5FD7F" ID-CoL="47K96" authorityName="Chen & Chen" authorityYear="2005" box="[1302,1437,719,740]" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="longibarba">
|
||||
<emphasis id="2FC70F77111EFFBEFAF8FD352CA5FD7F" box="[1302,1437,719,740]" italics="true" pageId="10" pageNumber="111">N. longibarba</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue