Add updates up until 2025-02-25 08:52:13

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KUFOS-AN-2024.12, 205.0 mm SL, collected from a local fisher at the
, KUFOS-AN-2024.12, 205.0 mm SL, collected from a local fisher at the
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, near Gorubathan,
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,
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&amp;
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SL, 2 ex, collection location same as above,
@ -155,7 +152,7 @@ See
,
<figureCitation id="3C922A33FFE74C6CFDFDFAE874689229" box="[540,636,1374,1401]" captionStart="FIGURE 7" captionStartId="8.[152,255,846,870]" captionTargetBox="[340,1247,181,822]" captionTargetId="figure-21@8.[340,1247,181,822]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of head, (A &amp; B) Channa amphibeus, ca. 270 and 500 mm SL, showing neon-blue portion (square region) and maroon blotches; C &amp; D) Channa barca, ca. 200 &amp; 500 mm SL, showing black blotches and absence of neon-blue portion (Photo credits, C &amp; D: Heok Hui Tan and Dominik Niemeier)." figureDoi="http://doi.org/10.5281/zenodo.14798030" httpUri="https://zenodo.org/record/14798030/files/figure.png" pageId="2" pageNumber="89">7A &amp; B</figureCitation>
for general appearance and
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for morphometric data. A large snakehead species reaching at least
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SL (present study), but reported by local fishers to reach
@ -286,7 +283,7 @@ See
. In 10% formalin: Dorsum, sides of head and body dark-grey with black spots. Ventral surface of body pale-grey to creamy-white. Bars on sides beige. Spots on head pale-grey. Pectoral fin greyish, with 3 narrow, semi-circular grey bands. Dorsal fin dark-grey with narrow white distal margin. Anal fin pale grey with 23 horizontal rows of black spots. Caudal fin pale-grey with black fin rays and whitish narrow margin. Pelvic fin pale-grey with dark-grey tips. Lower jaw ventral surface and belly creamy-white, branchiostegal membrane dark grey to black.
</paragraph>
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<emphasis id="96DDEAA4FFE34C68FF26FF6874AA97A9" bold="true" box="[199,702,222,249]" pageId="6" pageNumber="93">Distribution, habitat, and ecological notes.</emphasis>
Currently,

View file

@ -0,0 +1,654 @@
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<taxonomicName id="D4D8E9D729206A73FF7AE595FD87FC10" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[151,562,978,1005]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="minax">
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<emphasis id="21AC4E4629206A73FF7AE595FECCFC11" bold="true" box="[151,377,978,1005]" italics="true" pageId="2" pageNumber="73">Metallactus minax</emphasis>
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<paragraph id="1367925429206A73FF7AE5B0FE17FBED" blockId="2.[151,562,978,1041]" box="[151,418,1014,1041]" pageId="2" pageNumber="73">
(
<figureCitation id="8BE38ED129206A73FF72E5B0FF52FBED" box="[159,231,1015,1041]" captionStart="FIGURE 1" captionStartId="4.[151,250,1902,1926]" captionTargetBox="[167,1417,181,1861]" captionTargetId="figure-33@4.[151,1435,181,1879]" captionTargetPageId="4" captionText="FIGURE 1. Metallactus minax. ab: habitus, dorsal and lateral view (LT); c: frontal view, male (LT); d: frontal view, female (PLT, MNHUB); en: colour variations (e: female PLT, MNHUB; f: male, PT of M. mosei, Piracicaba, São Paulo, Brazil; g: male, “São Paulo”, Brazil; h: male, La Colmena, Paraguarí, Paraguay; k: male, “São Paulo”, Brazil; m: female, “Corocò”, Concepción, Paraguay; n: male, Jaguariaiva, Paraná, Brazil). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.14798004" httpUri="https://zenodo.org/record/14798004/files/figure.png" pageId="2" pageNumber="73">Figs 1</figureCitation>
;
<figureCitation id="8BE38ED129206A73FF18E5B0FE9AFBED" box="[245,303,1015,1041]" captionStart="FIGURE 4" captionStartId="11.[151,250,1594,1618]" captionTargetBox="[171,1416,309,1554]" captionTargetId="figure-80@11.[171,1416,309,1570]" captionTargetPageId="11" captionText="FIGURE 4. ad: Metallactus minax, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg: M. tommasoi, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="2" pageNumber="73">4ad</figureCitation>
;
<figureCitation id="8BE38ED129206A73FED7E5B1FEC6FBED" box="[314,371,1014,1041]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="2" pageNumber="73">5ab</figureCitation>
;
<figureCitation id="8BE38ED129206A73FE92E5B0FE2FFBED" box="[383,410,1015,1041]" captionStart="FIGURE 6" captionStartId="13.[151,250,948,972]" captionTargetBox="[151,1436,273,910]" captionTargetId="figure-48@13.[151,1436,273,924]" captionTargetPageId="13" captionText="FIGURE 6. Distribution maps for the Metallactus minax species-group. a: M. minax; b: M. abstersus; c: M. tommasoi. Circles: data from specimens analysed during the study. Star in fig. b: the available information allows to point out the presence in the state but without more precise data on the location. Star in fig. c: Identification uncertain based on females only, therefore excluded from the type series. In fig. a: record of the species for the state of Pará, indicated on the label of one of the paratypes (see text), is not shown on the map because this data is considered to require further confirmation." figureDoi="http://doi.org/10.5281/zenodo.14798014" httpUri="https://zenodo.org/record/14798014/files/figure.png" pageId="2" pageNumber="73">6a</figureCitation>
)
</paragraph>
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<treatmentCitation id="9279B44529206A73FF7AE27CFD97FBAF" author="Suffrian, E." box="[151,546,1083,1107]" page="249" pageId="2" pageNumber="73" year="1866">
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<emphasis id="21AC4E4629206A73FF7AE27CFEE6FBAF" box="[151,339,1083,1107]" italics="true" pageId="2" pageNumber="73">Metallactus minax</emphasis>
<bibRefCitation id="7749EFA529206A73FEB4E27CFD97FBAF" author="Suffrian, E." box="[345,546,1083,1107]" pageId="2" pageNumber="73" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866: 249</bibRefCitation>
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;
<treatmentCitation id="9279B44529206A73FDC1E27CFD4DFBAF" author="Clavareau, C. H." box="[556,760,1083,1107]" page="93" pageId="2" pageNumber="73" year="1913">
<bibRefCitation id="7749EFA529206A73FDC1E27CFD4DFBAF" author="Clavareau, C. H." box="[556,760,1083,1107]" pageId="2" pageNumber="73" pagination="1 - 278" refId="ref10059" refString="Clavareau, C. H. (1913) Pars 53: Chrysomelidae: Megascelinae, Megalopodinae, Clytrinae, Cryptocephalinae, Chlamydinae, Lamprosominae. In: Schenkling S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1-278." type="book chapter" year="1913">Clavareau, 1913: 93</bibRefCitation>
</treatmentCitation>
(catalogue);
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<bibRefCitation id="7749EFA529206A73FC93E27CFBC5FBAF" author="Blackwelder, R. E." box="[894,1136,1083,1107]" pageId="2" pageNumber="73" pagination="551 - 763" refId="ref10002" refString="Blackwelder, R. E. (1946) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 4. Bulletin of the United States National Museum, 185, 551-763. https://doi.org/10.5479/si.03629236.185.4" type="journal article" year="1946">Blackwelder, 1946: 641</bibRefCitation>
</treatmentCitation>
(catalogue).
<emphasis id="21AC4E4629206A73FF2AE21CFF6DFB8E" box="[199,216,1115,1138]" italics="true" pageId="2" pageNumber="73">=</emphasis>
<treatmentCitation id="9279B44529206A73FF32E21DFDE2FB8E" author="Scholler, M." box="[223,599,1114,1138]" page="71" pageId="2" pageNumber="73" year="2003">
<taxonomicName id="D4D8E9D729206A73FF32E21DFDE2FB8E" authority="Scholler, 2003: 71" authorityName=", Scholler" authorityPageNumber="71" authorityYear="2003" box="[223,599,1114,1138]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629206A73FF32E21DFE2DFB8E" box="[223,408,1114,1138]" italics="true" pageId="2" pageNumber="73">Metallactus mosei</emphasis>
<bibRefCitation id="7749EFA529206A73FE73E21DFDE2FB8E" author="Scholler, M." box="[414,599,1114,1138]" pageId="2" pageNumber="73" pagination="71 - 74" refId="ref10639" refString="Scholler, M. (2003) A new species of Metallactus Suffrian, 1866 (Coleoptera: Chrysomelidae: Cryptocephalinae) from Brazil. Russian Entomological Journal, 12 (1), 71-74." type="journal article" year="2003">Schöller, 2003: 71</bibRefCitation>
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s
<emphasis id="21AC4E4629206A73FD85E21DFD0EFB8E" bold="true" box="[616,699,1114,1138]" pageId="2" pageNumber="73">yn. nov.</emphasis>
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<emphasis id="21AC4E4629206A73FF7AE2E7FF55FB46" bold="true" box="[151,224,1184,1210]" italics="true" pageId="2" pageNumber="73">Types.</emphasis>
<bibRefCitation id="7749EFA529206A73FF05E2E7FE20FB46" author="Suffrian, E." box="[232,405,1184,1210]" pageId="2" pageNumber="73" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian (1866)</bibRefCitation>
did not mention the number of the studied specimens, but he stated that they all came both from Berlin Museum and from Deyrolles and Clarks collections, both of them at present hosted in BMNH. Three putative
<typeStatus id="CC632CF629206A73FF14E2AEFEEBFAFF" box="[249,350,1257,1283]" pageId="2" pageNumber="73" type="syntype">syntypes</typeStatus>
,
<specimenCount id="05DE59DD29206A73FE87E2AEFE65FAFE" box="[362,464,1256,1283]" count="1" pageId="2" pageNumber="73" type="male">one male</specimenCount>
and
<specimenCount id="05DE59DD29206A73FDE5E2AEFD26FAFE" box="[520,659,1256,1283]" count="2" pageId="2" pageNumber="73" type="female">two females</specimenCount>
, were tracked down in BMNH and
<specimenCount id="05DE59DD29206A73FBCBE2AFFB64FAFE" box="[1062,1233,1256,1282]" count="5" pageId="2" pageNumber="73" type="generic">five specimens</specimenCount>
, all females, were available in MNHUB. All these specimens match the original description of the colour pattern and its variation. Suffrian clearly stated that he had only females available for the study. However, as mentioned above, one of the specimens from London is a male. This would suggest excluding at least this specimen from the type series. Nevertheless, certain circumstances led me to a different decision, and this specimen has instead been designated the
<typeStatus id="CC632CF629206A73FF2EE3DBFE9FFA4A" box="[195,298,1436,1462]" pageId="2" pageNumber="73" type="lectotype">lectotype</typeStatus>
of the species. The reasons are as follows. In the species group under study, sexual dimorphism is very reduced. Specifically, the interocular distance is only slightly different when comparing males and females. Additionally, males do not exhibit any easily evident development of the tarsi or anterior tibiae. Furthermore, of notable importance, the fifth anal sternite in males has an unusually deep depression at the apex, curiously similar to that typically observed in females of the majority of Cryptocephalinae. Again, in the original description, Suffrian focuses on the characteristics of the reddish spot in the basal region of the elytra, stating: “
<emphasis id="21AC4E4629206A73FB79E016FEB3F92A" italics="true" pageId="2" pageNumber="73">Die Farbe schwarz, ein die schwarz bleibende Schulterbeule umziehender, den Seitenlappen mit einschliessender, nach Innen bis an den dritten Zwischenraum reichender Schulterfleck und die ziemlich breite Spitze, letztere in noch bleicherer Färbung, ziegelroth</emphasis>
”. This description, particularly the detail of the brick-red shoulder spot separated from the apical spot and encircling the black humeral callus, corresponds only to the
<specimenCount id="05DE59DD29206A73FC84E0A6FBBAF906" box="[873,1039,1760,1787]" count="2" pageId="2" pageNumber="73" type="generic">two specimens</specimenCount>
from the Clark collection. Besides, only the male exhibits the pronotal coloration that Suffrian describes as &quot;typical&quot;, specifically black with lateral margins and part of the anterior margin yellow. No other specimens from London or Berlin display this combination of colours on the pronotum. Furthermore, the male from the Clark collection has the typical handwritten note “a D. Suffrian. determinat.”, which is commonly found on the material studied by Suffrian in the preparation of his monograph (1866). As a matter of fact, there is also a further handwritten note “S Paul” that Suffrian does not mention. However, it should be noted that Suffrian was often not very precise in recording all the collection data, as can be seen in many other instances (see, for example,
<bibRefCitation id="7749EFA529206A73FCCEE19BFC2AF80A" author="Sassi, D." box="[803,927,2012,2038]" pageId="2" pageNumber="73" pagination="57 - 110" refId="ref10255" refString="Sassi, D. (2018) Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4413 (1), 57-110. https://doi.org/10.11646/zootaxa.4413.1.2" type="journal article" year="2018">Sassi 2018</bibRefCitation>
,
<bibRefCitation id="7749EFA529206A73FC47E19BFC50F80A" author="Sassi, D." box="[938,997,2012,2038]" pageId="2" pageNumber="73" pagination="251 - 282" refId="ref10460" refString="Sassi, D. (2022) Revision of the Metallactus taeniatellus species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 5125 (3), 251-282. https://doi.org/10.11646/zootaxa.5125.3.1" type="journal article" year="2022">2022</bibRefCitation>
,
<bibRefCitation id="7749EFA529206A73FC1CE19BFB9BF80A" author="Sassi, D." box="[1009,1070,2012,2038]" pageId="2" pageNumber="73" pagination="201 - 237" refId="ref10505" refString="Sassi, D. (2024) Revision of the Griburius posticatus (Suffrian, 1866) species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 5406 (2), 201-237. https://doi.org/10.11646/zootaxa.5406.2.1" type="journal article" year="2024">2024</bibRefCitation>
). Therefore, the most reasonable interpretation of the observed situation is that Suffrian overlooked the presence of a male among the materials he studied. This interpretation is supported by the fact that the German entomologist made the same mistake on other occasions; see, for example, the treatment of the following species and what was stated in the designation of the
<typeStatus id="CC632CF629216A72FF7AE744FEBCFEE1" box="[151,265,259,285]" pageId="3" pageNumber="74" type="lectotype">lectotypes</typeStatus>
of
<taxonomicName id="D4D8E9D729216A72FEC5E744FD05FEE1" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[296,688,259,285]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="insitivus">
<emphasis id="21AC4E4629216A72FEC5E744FDB8FEE0" box="[296,525,259,285]" italics="true" pageId="3" pageNumber="74">Metallactus insitivus</emphasis>
<bibRefCitation id="7749EFA529216A72FDFFE744FD05FEE1" author="Suffrian, E." box="[530,688,259,285]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D4D8E9D729216A72FD0CE744FB94FEE1" authority="Suffrian 1866" authorityName="Suffrian" authorityYear="1866" box="[737,1057,259,285]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="taeniatellus">
<emphasis id="21AC4E4629216A72FD0CE744FC33FEE1" box="[737,902,259,285]" italics="true" pageId="3" pageNumber="74">M.taeniatellus</emphasis>
<bibRefCitation id="7749EFA529216A72FC67E744FB94FEE1" author="Suffrian, E." box="[906,1057,259,285]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian 1866</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="7749EFA529216A72FBC0E744FB1CFEE1" author="Sassi, D." box="[1069,1193,259,285]" pageId="3" pageNumber="74" pagination="251 - 282" refId="ref10460" refString="Sassi, D. (2022) Revision of the Metallactus taeniatellus species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 5125 (3), 251-282. https://doi.org/10.11646/zootaxa.5125.3.1" type="journal article" year="2022">Sassi 2022</bibRefCitation>
). For all these reasons, the typification has been made as follows.
<typeStatus id="CC632CF629216A72FD6DE760FCB5FEBD" box="[640,768,295,321]" pageId="3" pageNumber="74" type="lectotype">LECTOTYPE</typeStatus>
:
<specimenCount id="05DE59DD29216A72FCE6E760FC97FEBD" box="[779,802,295,321]" count="1" pageId="3" pageNumber="74" type="male"></specimenCount>
, glued, aedeagus extracted and glued on the same label bearing the specimen, abdomen glued on a separate label, // “250” [blue label, handwritten] // “S Paul” [one second unreadable word, white label, handwritten] // “E. Coll
<superScript id="E4AD3F1C29216A72FD19E728FC81FE81" attach="left" box="[756,820,367,393]" fontSize="6" pageId="3" pageNumber="74">Chevt</superScript>
.” [white label, printed] // “65.56” [white label, printed] // “a D. Suffrian. determinat.” [white label, printed] // “
<taxonomicName id="D4D8E9D729216A72FD19E7D5FCC7FE50" box="[756,882,402,428]" genus="Met" pageId="3" pageNumber="74" rank="species" species="minax">Met minax</taxonomicName>
” [white label, handwritten] // “
<typeStatus id="CC632CF629216A72FB2AE7D5FA92FE50" box="[1223,1319,402,428]" pageId="3" pageNumber="74" type="syntype">Syntype</typeStatus>
” [rounded white label with blue margin, printed] // “
<taxonomicName id="D4D8E9D729216A72FD9CE7F1FC59FE2C" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[625,1004,438,464]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FD9CE7F1FCF4FE2C" box="[625,833,438,464]" italics="true" pageId="3" pageNumber="74">Metallactus minax</emphasis>
<bibRefCitation id="7749EFA529216A72FCA4E7F1FC59FE2C" author="Suffrian, E." box="[841,1004,438,464]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
<typeStatus id="CC632CF629216A72FC19E7F1FBC9FE2C" box="[1012,1148,438,464]" pageId="3" pageNumber="74" type="lectotype">LECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] // (BMNH).
<typeStatus id="CC632CF629216A72FE6EE79DFDF6FE08" box="[387,579,474,500]" pageId="3" pageNumber="74" type="paralectotype">PARALECTOTYPES</typeStatus>
:
<specimenCount id="05DE59DD29216A72FDA1E79DFDDAFE08" box="[588,623,474,500]" count="1" pageId="3" pageNumber="74" type="female">1♀</specimenCount>
, pinned // ”
<taxonomicName id="D4D8E9D729216A72FD1DE79DFBB1FE08" authority="Suffr" authorityName="Suffrian" authorityYear="1866" box="[752,1028,474,500]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">Metallactus minax Suffr</taxonomicName>
<collectingCountry id="6BCFD2C429216A72FBE6E79DFBE6FE08" box="[1035,1107,474,500]" name="Brazil" pageId="3" pageNumber="74">Brazil</collectingCountry>
” [white label, handwritten) // “Type Suffr Coll. Deyrolle” [white label, handwritten] // “Baly Coll.” [white label, printed] // “
<typeStatus id="CC632CF629216A72FB44E7B9FABCFDE4" box="[1193,1289,510,536]" pageId="3" pageNumber="74" type="syntype">Syntype</typeStatus>
” [white label with blue border, printed] // (BMNH);
<specimenCount id="05DE59DD29216A72FDA8E466FDDDFDC7" box="[581,616,545,571]" count="1" pageId="3" pageNumber="74" type="female">1♀</specimenCount>
, pinned // “
<collectingRegion id="D11C5CB629216A72FD1DE466FC91FDC7" box="[752,804,545,571]" country="Brazil" name="Para" pageId="3" pageNumber="74">Para</collectingRegion>
” [white label, handwritten) // “
<taxonomicName id="D4D8E9D729216A72FB91E466FAB4FDC7" authorityName="Suffrian" authorityYear="1866" box="[1148,1281,545,571]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">Met. minax</taxonomicName>
” [white label, handwritten] // &quot;E Coll Laferté&quot; [white label, printed] // &quot;67.56&quot; [white label, printed] // &quot;116&quot; [blue label, printed] // &quot;
<typeStatus id="CC632CF629216A72FF5AE42EFEAFFD7F" box="[183,282,617,643]" pageId="3" pageNumber="74" type="syntype">Syntype</typeStatus>
&quot; [white with blue border, printed] // (BMNH);
<specimenCount id="05DE59DD29216A72FCD9E42EFCECFD7F" box="[820,857,617,643]" count="4" pageId="3" pageNumber="74" type="female">4♀</specimenCount>
, pinned // “23948” [white label, printed] //
<taxonomicName id="D4D8E9D729216A72FAA7E42EFF6CFD5A" authority="Suffr." authorityName="Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">“minax Suffr.</taxonomicName>
* c. variet.
<collectingCountry id="6BCFD2C429216A72FEBDE4CBFE26FD5A" box="[336,403,652,678]" name="Brazil" pageId="3" pageNumber="74">Brasil</collectingCountry>
C. Sell.” [blue label, handwritten] // (MNHUB);
<specimenCount id="05DE59DD29216A72FC58E4CAFC6DFD5B" box="[949,984,653,679]" count="1" pageId="3" pageNumber="74" type="female">1♀</specimenCount>
, pinned // “23949” [white label, printed] //
<taxonomicName id="D4D8E9D729216A72FF42E4F7FEB7FD36" authorityName="Suffrian" authorityYear="1866" box="[175,258,688,714]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">“minax</taxonomicName>
var. Suffr.*
<collectingCountry id="6BCFD2C429216A72FE7FE4F7FE60FD36" box="[402,469,688,714]" name="Brazil" pageId="3" pageNumber="74">Brasil</collectingCountry>
C. Sell.” [blue label, handwritten] // (MNHUB). The label information for these last
<specimenCount id="05DE59DD29216A72FF7AE493FEF5FD12" box="[151,320,724,750]" count="5" pageId="3" pageNumber="74" type="generic">five specimens</specimenCount>
matches the registration data from the old catalogue of the MNHUB (“23948 [
<taxonomicName id="D4D8E9D729216A72FB52E493FAFDFD12" authorityName="Suffrian" authorityYear="1866" box="[1215,1352,724,750]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
]
<taxonomicName id="D4D8E9D729216A72FABBE493FF6CFCEE" authority="Suffr." authorityName="Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">minax Suffr.</taxonomicName>
* 4.
<collectingCountry id="6BCFD2C429216A72FEE8E4BFFEFDFCEE" box="[261,328,760,786]" name="Brazil" pageId="3" pageNumber="74">Brasil</collectingCountry>
Sello.” and “23949 [
<taxonomicName id="D4D8E9D729216A72FDC3E4BFFCB6FCEE" authorityName="Suffrian" authorityYear="1866" box="[558,771,760,786]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">Metallactus minax</taxonomicName>
] var. Suffr.* 1. [
<collectingCountry id="6BCFD2C429216A72FC5EE4BFFC4CFCEE" box="[947,1017,760,786]" name="Brazil" pageId="3" pageNumber="74">Brasil</collectingCountry>
Sello].”). All
<typeStatus id="CC632CF629216A72FB79E4BFFA83FCEE" box="[1172,1334,760,786]" pageId="3" pageNumber="74" type="paralectotype">paralectotypes</typeStatus>
labelled: “
<taxonomicName id="D4D8E9D729216A72FF49E55CFDAEFCC9" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[164,539,795,821]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FF49E55CFEC6FCC9" box="[164,371,795,821]" italics="true" pageId="3" pageNumber="74">Metallactus minax</emphasis>
<bibRefCitation id="7749EFA529216A72FE97E55CFDAEFCC9" author="Suffrian, E." box="[378,539,795,821]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
<typeStatus id="CC632CF629216A72FDCFE55BFD55FCCA" box="[546,736,796,822]" pageId="3" pageNumber="74" type="paralectotype">PARALECTOTYPUS</typeStatus>
D. Sassi det.” [red label, printed].
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29216A72FF1AE578FE1DFBAF" pageId="3" pageNumber="74" type="discussion">
<paragraph id="1367925429216A72FF1AE578FE1DFBAF" blockId="3.[151,1437,151,2037]" pageId="3" pageNumber="74">
As regard
<taxonomicName id="D4D8E9D729216A72FE82E578FD4BFCA5" authority=", Scholler (2003)" authorityName=", Scholler" authorityYear="2003" box="[367,766,831,858]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629216A72FE82E578FD88FCA5" box="[367,573,831,857]" italics="true" pageId="3" pageNumber="74">Metallactus mosei</emphasis>
,
<bibRefCitation id="7749EFA529216A72FDA0E578FD4BFCA5" author="Scholler, M." box="[589,766,831,857]" pageId="3" pageNumber="74" pagination="71 - 74" refId="ref10639" refString="Scholler, M. (2003) A new species of Metallactus Suffrian, 1866 (Coleoptera: Chrysomelidae: Cryptocephalinae) from Brazil. Russian Entomological Journal, 12 (1), 71-74." type="journal article" year="2003">Schöller (2003)</bibRefCitation>
</taxonomicName>
described it based on 44 (47?) specimens. The description was made comparing the external habitus of the specimens with the
<typeStatus id="CC632CF629216A72FC57E524FBA9FC81" box="[954,1052,867,893]" pageId="3" pageNumber="74" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="D4D8E9D729216A72FBA5E523FB00FC81" authorityName="Suffrian" authorityYear="1866" box="[1096,1205,868,893]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FBA5E523FB00FC81" box="[1096,1205,868,893]" italics="true" pageId="3" pageNumber="74">M. minax</emphasis>
</taxonomicName>
hosted in MNHUB (which included only females) and highlighting a series of differences primarily in the colour pattern. Unfortunately, the material of the historical collection in MNHUB did not provide a complete picture of the variation range of
<taxonomicName id="D4D8E9D729216A72FF7AE588FEB7FC14" authorityName="Suffrian" authorityYear="1866" box="[151,258,975,1000]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FF7AE588FEB7FC14" box="[151,258,975,1000]" italics="true" pageId="3" pageNumber="74">M. minax</emphasis>
</taxonomicName>
, and the lack of male specimens prevented a comparison of the male genitalia. I had the opportunity to study
<typeStatus id="CC632CF629216A72FF37E5B5FEF0FBF0" box="[218,325,1010,1036]" pageId="3" pageNumber="74" type="paratype">paratypes</typeStatus>
(of both sexes) of
<taxonomicName id="D4D8E9D729216A72FDF8E5B4FDCFFBF0" authorityName=", Scholler" authorityYear="2003" box="[533,634,1011,1036]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629216A72FDF8E5B4FDCFFBF0" box="[533,634,1011,1036]" italics="true" pageId="3" pageNumber="74">M. mosei</emphasis>
</taxonomicName>
and I did not find any significative differences between the two species. For this reason, the following synonymy is here proposed:
<taxonomicName id="D4D8E9D729216A72FCD9E252FB1AFBD3" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[820,1199,1045,1071]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FCD9E252FBB1FBD3" box="[820,1028,1045,1071]" italics="true" pageId="3" pageNumber="74">Metallactus minax</emphasis>
<bibRefCitation id="7749EFA529216A72FBE0E252FB1AFBD3" author="Suffrian, E." box="[1037,1199,1045,1071]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="D4D8E9D729216A72FB22E252FE8CFBAF" authority="Scholler, 2003" authorityName=", Scholler" authorityYear="2003" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="mosei" status="syn. nov.">
<emphasis id="21AC4E4629216A72FB22E252FA29FBD3" box="[1231,1436,1045,1071]" italics="true" pageId="3" pageNumber="74">Metallactus mosei</emphasis>
<bibRefCitation id="7749EFA529216A72FF7AE27EFE8CFBAF" author="Scholler, M." box="[151,313,1081,1107]" pageId="3" pageNumber="74" pagination="71 - 74" refId="ref10639" refString="Scholler, M. (2003) A new species of Metallactus Suffrian, 1866 (Coleoptera: Chrysomelidae: Cryptocephalinae) from Brazil. Russian Entomological Journal, 12 (1), 71-74." type="journal article" year="2003">Schöller, 2003</bibRefCitation>
</taxonomicName>
<emphasis id="21AC4E4629216A72FEADE27EFE1DFBAF" bold="true" box="[320,424,1081,1107]" pageId="3" pageNumber="74">
<taxonomicNameLabel id="3A9FF33D29216A72FEADE27EFE1DFBAF" box="[320,424,1081,1107]" pageId="3" pageNumber="74">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29216A72FF2AE21AFEBBF9B4" pageId="3" pageNumber="74" type="materials_examined">
<paragraph id="1367925429216A72FF2AE21AFD88FB8B" blockId="3.[151,1437,151,2037]" box="[199,573,1116,1143]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE21AFED5FB8B" bold="true" box="[199,352,1116,1143]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE21AFEECFB8B" bold="true" box="[199,345,1116,1143]" italics="true" pageId="3" pageNumber="74">
<typeStatus id="CC632CF629216A72FF2AE21AFF4BFB8B" box="[199,254,1117,1143]" pageId="3" pageNumber="74">Type</typeStatus>
locality
</emphasis>
.
</emphasis>
<collectingRegion id="D11C5CB629216A72FE8AE21AFE6DFB8B" box="[359,472,1117,1143]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="74">São Paulo</collectingRegion>
(
<collectingCountry id="6BCFD2C429216A72FE0AE21AFD84FB8B" box="[487,561,1117,1143]" name="Brazil" pageId="3" pageNumber="74">Brazil</collectingCountry>
).
</paragraph>
<paragraph id="1367925429216A72FF2AE2C7FE0EF9FC" blockId="3.[151,1437,151,2037]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE2C7FD6DFB66" bold="true" box="[199,728,1152,1179]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE2C7FD93FB67" bold="true" box="[199,550,1152,1179]" italics="true" pageId="3" pageNumber="74">Additional material examined.</emphasis>
<collectingCountry id="6BCFD2C429216A72FDC0E2C7FD6DFB66" box="[557,728,1152,1178]" name="Argentina" pageId="3" pageNumber="74">ARGENTINA</collectingCountry>
</emphasis>
:
<emphasis id="21AC4E4629216A72FD0AE2C7FCE4FB66" bold="true" box="[743,849,1152,1178]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FD0AE2C7FCE4FB66" box="[743,849,1152,1178]" country="Argentina" name="Misiones" pageId="3" pageNumber="74">Misiones</collectingRegion>
</emphasis>
: Santa Ana
<date id="6766B49429216A72FC34E2C6FB94FB60" box="[985,1057,1153,1180]" pageId="3" pageNumber="74" value="1952-01">I.1952</date>
(1,
<collectionCode id="75C90A9129216A72FBA3E2C6FB78FB67" box="[1102,1229,1153,1179]" pageId="3" pageNumber="74">USNMNH</collectionCode>
);
<quantity id="D4203FB129216A72FB32E2C6FA92FB67" box="[1247,1319,1153,1179]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="74" unit="km" value="12.0">12 km</quantity>
SW Santa Ana
<date id="6766B49429216A72FF26E2E2FE83FB43" box="[203,310,1189,1215]" pageId="3" pageNumber="74" value="1989-01-21">21.I.1989</date>
(1,
<collectionCode id="75C90A9129216A72FEB2E2E2FE2AFB43" box="[351,415,1189,1215]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="3" pageNumber="74" type="University or college">BYU</collectionCode>
);
<emphasis id="21AC4E4629216A72FE5CE2E3FDA9FB42" bold="true" box="[433,540,1188,1214]" pageId="3" pageNumber="74">
<collectingCountry id="6BCFD2C429216A72FE5CE2E3FDA9FB42" box="[433,540,1188,1214]" name="Brazil" pageId="3" pageNumber="74">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="21AC4E4629216A72FDC4E2E2FD77FB42" bold="true" box="[553,706,1188,1215]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FDC4E2E2FD77FB42" box="[553,706,1188,1215]" country="Brazil" name="Mato Grosso" pageId="3" pageNumber="74">Mato Grosso</collectingRegion>
</emphasis>
: Maracaju
<date id="6766B49429216A72FCD3E2E2FC25FB43" box="[830,912,1189,1215]" pageId="3" pageNumber="74" value="1937-02">II.1937</date>
(1,
<collectionCode id="75C90A9129216A72FC54E2E2FB8DFB43" box="[953,1080,1189,1215]" pageId="3" pageNumber="74">USNMNH</collectionCode>
).
<emphasis id="21AC4E4629216A72FBA5E2E3FB2AFB43" bold="true" box="[1096,1183,1188,1215]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FBA5E2E3FB2AFB43" box="[1096,1183,1188,1215]" country="Brazil" name="Parana" pageId="3" pageNumber="74">Paraná</collectingRegion>
</emphasis>
: Jaguariaiva
<date id="6766B49429216A72FADCE2E2FA29FB43" box="[1329,1436,1189,1215]" pageId="3" pageNumber="74" value="1974-01-29">29.I.1974</date>
(1,
<collectionCode id="75C90A9129216A72FF52E28FFEAFFB1E" box="[191,282,1224,1250]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="74">AMNH</collectionCode>
).
<emphasis id="21AC4E4629216A72FEDDE28FFE1EFB1E" bold="true" box="[304,427,1224,1250]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FEDDE28FFE1EFB1E" box="[304,427,1224,1250]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="74">São Paulo</collectingRegion>
</emphasis>
: “
<collectingRegion id="D11C5CB629216A72FE25E28FFDF4FB1E" box="[456,577,1224,1250]" inferredBySuffix="true" pageId="3" pageNumber="74">São Paulo</collectingRegion>
” (15,
<collectionCode id="75C90A9129216A72FD63E28FFD68FB1E" box="[654,733,1224,1250]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="74">NMPC</collectionCode>
&amp;
<collectionCode id="75C90A9129216A72FCE4E28FFCD1FB1E" box="[777,868,1224,1250]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="74">MNHN</collectionCode>
); Piracicaba
<date id="6766B49429216A72FC16E28FFBF9FB1E" box="[1019,1100,1224,1250]" pageId="3" pageNumber="74" value="2002-02">II.2002</date>
Lab. Rearing on
<taxonomicName id="D4D8E9D729216A72FACDE28FFEB6FAFA" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="urophylla">
<emphasis id="21AC4E4629216A72FACDE28FFEB6FAFA" italics="true" pageId="3" pageNumber="74">Eucalyptus urophylla</emphasis>
</taxonomicName>
(3,
<collectionCode id="75C90A9129216A72FEDDE2ABFECCFAFA" box="[304,377,1260,1286]" pageId="3" pageNumber="74">DSPC</collectionCode>
,
<collectionCode id="75C90A9129216A72FE69E2AAFE16FAFB" box="[388,419,1261,1287]" country="Canada" name="Université de Montréal" pageId="3" pageNumber="74" type="Herbarium">LT</collectionCode>
of
<taxonomicName id="D4D8E9D729216A72FE24E2ABFD20FAFA" authorityName=", Scholler" authorityYear="2003" box="[457,661,1260,1286]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629216A72FE24E2ABFD20FAFA" box="[457,661,1260,1286]" italics="true" pageId="3" pageNumber="74">Metallactus mosei</emphasis>
</taxonomicName>
).
<emphasis id="21AC4E4629216A72FD40E2ABFC64FAFA" bold="true" box="[685,977,1260,1286]" pageId="3" pageNumber="74">
<collectingCountry id="6BCFD2C429216A72FD40E2ABFCFEFAFA" box="[685,843,1260,1286]" name="Paraguay" pageId="3" pageNumber="74">PARAGUAY</collectingCountry>
:
<collectingRegion id="D11C5CB629216A72FCB8E2ABFC64FAFA" box="[853,977,1260,1286]" country="Paraguay" name="Amambay" pageId="3" pageNumber="74">Amambay</collectingRegion>
</emphasis>
: Cerro Cora National Park
<date id="6766B49429216A72FAEAE2ABFA28FAFA" box="[1287,1437,1260,1286]" pageId="3" pageNumber="74" value="2001-01-19" valueMax="2001-01-21" valueMin="2001-01-19">1921.I.2001</date>
(1,
<collectionCode id="75C90A9129216A72FF56E357FEA5FAD6" box="[187,272,1296,1322]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="74">NMPC</collectionCode>
).
<emphasis id="21AC4E4629216A72FECCE348FE24FAD6" bold="true" box="[289,401,1295,1322]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FECCE348FE24FAD6" box="[289,401,1295,1322]" country="Paraguay" name="Asuncion" pageId="3" pageNumber="74">Asunción</collectingRegion>
</emphasis>
: 1891 (7,
<collectionCode id="75C90A9129216A72FDEDE357FDEEFAD6" box="[512,603,1296,1322]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="74">MNHN</collectionCode>
).
<emphasis id="21AC4E4629216A72FD81E348FD58FAD6" bold="true" box="[620,749,1295,1322]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FD81E348FD5DFAD6" box="[620,744,1295,1322]" country="Paraguay" name="Caaguazu" pageId="3" pageNumber="74">Caaguazú</collectingRegion>
:
</emphasis>
Caaguazú
<date id="6766B49429216A72FC8AE357FC78FAD6" box="[871,973,1296,1322]" pageId="3" pageNumber="74" value="1943-12">XII.1943</date>
(1,
<collectionCode id="75C90A9129216A72FC1AE357FBC3FAD6" box="[1015,1142,1296,1322]" pageId="3" pageNumber="74">USNMNH</collectionCode>
).
<emphasis id="21AC4E4629216A72FB6BE348FB57FAD5" bold="true" box="[1158,1250,1295,1321]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FB6BE348FB57FAD5" box="[1158,1250,1295,1321]" country="Paraguay" name="Central" pageId="3" pageNumber="74">Central</collectingRegion>
</emphasis>
: Capiatá
<date id="6766B49429216A72FAA6E348FA29FAD6" box="[1355,1436,1295,1322]" pageId="3" pageNumber="74" value="1995-02">II.1995</date>
(1,
<collectionCode id="75C90A9129216A72FF52E373FEAFFAB2" box="[191,282,1332,1358]" pageId="3" pageNumber="74">MSNM</collectionCode>
); “Central” 1885 (1,
<collectionCode id="75C90A9129216A72FDE6E373FDD3FAB2" box="[523,614,1332,1358]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="74">MNHN</collectionCode>
).
<emphasis id="21AC4E4629216A72FD97E375FCB3FAB1" bold="true" box="[634,774,1330,1357]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FD97E375FCB3FAB1" box="[634,774,1330,1357]" country="Paraguay" name="Concepcion" pageId="3" pageNumber="74">Concepción</collectingRegion>
</emphasis>
:
<quantity id="D4203FB129216A72FCFAE373FCDBFAB1" box="[791,878,1331,1358]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="74" unit="km" value="120.0">120 km</quantity>
NEE Concepción
<date id="6766B49429216A72FBA8E373FB7FFAB2" box="[1093,1226,1332,1358]" pageId="3" pageNumber="74" value="2008-02-02" valueMax="2008-02-03" valueMin="2008-02-02">23-II.2008</date>
<quantity id="D4203FB129216A72FB3EE374FAA8FAB2" box="[1235,1309,1331,1358]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="74" unit="m" value="175.0">175 m</quantity>
(5,
<collectionCode id="75C90A9129216A72FAA0E373FA29FAB2" box="[1357,1436,1332,1358]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="74">NMPC</collectionCode>
&amp;
<collectionCode id="75C90A9129216A72FF5AE310FEB7FA8D" box="[183,258,1367,1393]" pageId="3" pageNumber="74">DSPC</collectionCode>
); Serranía San Luis
<date id="6766B49429216A72FE1CE310FDE8FA8D" box="[497,605,1367,1393]" pageId="3" pageNumber="74" value="2001-01-23">23.I.2001</date>
(4,
<collectionCode id="75C90A9129216A72FD7DE310FD78FA8D" box="[656,717,1367,1393]" pageId="3" pageNumber="74">JPPC</collectionCode>
&amp;
<collectionCode id="75C90A9129216A72FD1AE310FCF8FA8D" box="[759,845,1367,1393]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="3" pageNumber="74">MSNG</collectionCode>
); Concepción (1,
<collectionCode id="75C90A9129216A72FBF0E310FB3FFA8D" box="[1053,1162,1367,1393]" pageId="3" pageNumber="74">HNHMB</collectionCode>
); “Concepción Corocò”
<date id="6766B49429216A72FF7AE33CFF5DFA69" box="[151,232,1403,1429]" pageId="3" pageNumber="74" value="1997-02">II.1997</date>
(3,
<collectionCode id="75C90A9129216A72FEFFE33CFEE8FA69" box="[274,349,1403,1429]" pageId="3" pageNumber="74">DSPC</collectionCode>
); Orquela Pochon (Horqueta?)
<date id="6766B49429216A72FD58E33CFD48FA69" box="[693,765,1403,1429]" pageId="3" pageNumber="74" value="1936-01">I.1936</date>
(1,
<collectionCode id="75C90A9129216A72FCCAE33CFC35FA69" box="[807,896,1403,1429]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="74">NHMB</collectionCode>
).
<emphasis id="21AC4E4629216A72FC7DE33CFBB9FA69" bold="true" box="[912,1036,1403,1429]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FC7DE33CFBB9FA69" box="[912,1036,1403,1429]" country="Paraguay" name="Cordillera" pageId="3" pageNumber="74">Cordillera</collectingRegion>
</emphasis>
: Naranjo
<date id="6766B49429216A72FB97E33CFB7EFA6A" box="[1146,1227,1403,1430]" pageId="3" pageNumber="74" value="1995-02">II.1995</date>
(3,
<collectionCode id="75C90A9129216A72FB18E33CFAE5FA69" box="[1269,1360,1403,1429]" pageId="3" pageNumber="74">MSNM</collectionCode>
); Altos
<date id="6766B49429216A72FF7AE3D8FF47FA45" box="[151,242,1439,1465]" pageId="3" pageNumber="74" value="1991-03">III.1991</date>
(2,
<collectionCode id="75C90A9129216A72FEF3E3D8FEDCFA45" box="[286,361,1439,1465]" pageId="3" pageNumber="74">DSPC</collectionCode>
); San Bernardino (3,
<collectionCode id="75C90A9129216A72FDBBE3D8FD22FA45" box="[598,663,1439,1465]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="74" type="Museum">NMV</collectionCode>
&amp;
<collectionCode id="75C90A9129216A72FD57E3D8FC92FA45" box="[698,807,1439,1465]" pageId="3" pageNumber="74">MNHUB</collectionCode>
).
<emphasis id="21AC4E4629216A72FCD7E3D9FC06FA45" bold="true" box="[826,947,1438,1465]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FCD7E3D9FC06FA45" box="[826,947,1438,1465]" country="Paraguay" name="Paraguari" pageId="3" pageNumber="74">Paraguarí</collectingRegion>
</emphasis>
: 5 Km W La Colmena
<quantity id="D4203FB129216A72FB5AE3D8FB42FA45" box="[1207,1271,1439,1465]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="74" unit="m" value="200.0">200m</quantity>
2122,
<date id="6766B49429216A72FAA7E3D8FA28FA45" box="[1354,1437,1439,1465]" pageId="3" pageNumber="74" value="2008-02">II.2008</date>
(2,
<collectionCode id="75C90A9129216A72FF50E385FEA3FA20" box="[189,278,1474,1500]" pageId="3" pageNumber="74">MMPC</collectionCode>
); Sapucai 1903 (3,
<collectionCode id="75C90A9129216A72FE00E384FDF4FA21" box="[493,577,1475,1501]" pageId="3" pageNumber="74">BMNH</collectionCode>
&amp;
<collectionCode id="75C90A9129216A72FD88E385FD51FA20" box="[613,740,1474,1500]" pageId="3" pageNumber="74">USNMNH</collectionCode>
).
<emphasis id="21AC4E4629216A72FD1BE385FCC7FA20" bold="true" box="[758,882,1474,1500]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FD1BE385FCC7FA20" box="[758,882,1474,1500]" country="Paraguay" name="San Pedro" pageId="3" pageNumber="74">San Pedro</collectingRegion>
</emphasis>
:
<quantity id="D4203FB129216A72FC6DE385FC0FFA20" box="[896,954,1474,1501]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="74" unit="km" value="5.0">5 km</quantity>
N Taquati
<date id="6766B49429216A72FBDBE385FB24FA20" box="[1078,1169,1474,1500]" pageId="3" pageNumber="74" value="1989-03">III.1989</date>
(1,
<collectionCode id="75C90A9129216A72FB53E385FABCFA20" box="[1214,1289,1474,1500]" pageId="3" pageNumber="74">DSPC</collectionCode>
). “Paraguay” (15,
<collectionCode id="75C90A9129216A72FF26E3A0FE86F9FD" box="[203,307,1511,1537]" pageId="3" pageNumber="74">MNHUB</collectionCode>
&amp;
<collectionCode id="75C90A9129216A72FEBAE3A1FE05F9FC" box="[343,432,1510,1536]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="74">NHMB</collectionCode>
).
</paragraph>
<paragraph id="1367925429216A72FF2AE04EFEBBF9B4" blockId="3.[151,1437,151,2037]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE04EFD9EF9D8" bold="true" box="[199,555,1545,1572]" italics="true" pageId="3" pageNumber="74">Additional data from literature</emphasis>
.
<emphasis id="21AC4E4629216A72FDD4E04DFD11F9D8" bold="true" box="[569,676,1546,1572]" pageId="3" pageNumber="74">
<collectingCountry id="6BCFD2C429216A72FDD4E04DFD11F9D8" box="[569,676,1546,1572]" name="Brazil" pageId="3" pageNumber="74">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="21AC4E4629216A72FD5EE04DFC9FF9D8" bold="true" box="[691,810,1546,1572]" pageId="3" pageNumber="74">
<collectingRegion id="D11C5CB629216A72FD5EE04DFC9FF9D8" box="[691,810,1546,1572]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="74">São Paulo</collectingRegion>
</emphasis>
: Boa Esperança do Sul (
<bibRefCitation id="7749EFA529216A72FBD2E04DFB55F9D8" author="Scholler, M." box="[1087,1248,1546,1572]" pageId="3" pageNumber="74" pagination="71 - 74" refId="ref10639" refString="Scholler, M. (2003) A new species of Metallactus Suffrian, 1866 (Coleoptera: Chrysomelidae: Cryptocephalinae) from Brazil. Russian Entomological Journal, 12 (1), 71-74." type="journal article" year="2003">Schöller 2003</bibRefCitation>
, locus typicus of
<taxonomicName id="D4D8E9D729216A72FF7AE069FF4BF9BB" authorityName=", Scholler" authorityYear="2003" box="[151,254,1582,1607]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629216A72FF7AE069FF4BF9BB" box="[151,254,1582,1607]" italics="true" pageId="3" pageNumber="74">M. mosei</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29216A72FF2AE016FB76F973" pageId="3" pageNumber="74" type="distribution">
<paragraph id="1367925429216A72FF2AE016FB76F973" blockId="3.[151,1437,151,2037]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE016FEE6F990" bold="true" box="[199,339,1617,1644]" italics="true" pageId="3" pageNumber="74">Distribution</emphasis>
.
<collectingCountry id="6BCFD2C429216A72FE89E016FE63F997" box="[356,470,1617,1643]" name="Argentina" pageId="3" pageNumber="74">Argentina</collectingCountry>
,
<collectingCountry id="6BCFD2C429216A72FE08E016FD99F997" box="[485,556,1617,1643]" name="Brazil" pageId="3" pageNumber="74">Brazil</collectingCountry>
,
<collectingCountry id="6BCFD2C429216A72FDD1E015FD13F990" box="[572,678,1618,1644]" name="Paraguay" pageId="3" pageNumber="74">Paraguay</collectingCountry>
. New for
<collectingCountry id="6BCFD2C429216A72FCF2E016FC3BF997" box="[799,910,1617,1643]" name="Argentina" pageId="3" pageNumber="74">Argentina</collectingCountry>
and
<collectingCountry id="6BCFD2C429216A72FC20E015FB82F990" box="[973,1079,1618,1644]" name="Paraguay" pageId="3" pageNumber="74">Paraguay</collectingCountry>
. In my opinion, the indication “Para” on the label of one of the females housed in BMNH would require further confirmation.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29216A72FF2AE0DEFE47F809" pageId="3" pageNumber="74" type="discussion">
<paragraph id="1367925429216A72FF2AE0DEFE47F809" blockId="3.[151,1437,151,2037]" pageId="3" pageNumber="74">
<emphasis id="21AC4E4629216A72FF2AE0DEFE2AF94F" bold="true" box="[199,415,1689,1715]" italics="true" pageId="3" pageNumber="74">Comparative notes</emphasis>
. Based on the available material, compared to the other two species in the group, the dorsal black coloration of this species is more extensive, predominantly featuring a pattern with a black pronotum and yellow elytra with a common, large, elliptical black spot in the center. However, this black pattern is often reduced and fragmented, and the pronotum can also be extensively yellow. Due to the elliptical elytral spot and the black coloration of the legs,
<taxonomicName id="D4D8E9D729216A72FE78E16FFDD0F8BD" authorityName="Suffrian" authorityYear="1866" box="[405,613,1832,1858]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FE78E16FFDD0F8BD" box="[405,613,1832,1858]" italics="true" pageId="3" pageNumber="74">Metallactus minax</emphasis>
</taxonomicName>
could resemble some species of
<taxonomicName id="D4D8E9D729216A72FC37E16FFBFDF8BE" authorityName="Haldeman" authorityYear="1849" box="[986,1096,1832,1858]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="21AC4E4629216A72FC37E16FFBFDF8BE" box="[986,1096,1832,1858]" italics="true" pageId="3" pageNumber="74">Griburius</emphasis>
</taxonomicName>
in the
<taxonomicName id="D4D8E9D729216A72FB7AE16FFABCF8BD" authorityName="Suffrian" authorityYear="1866" box="[1175,1289,1832,1857]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="posticatus">
<emphasis id="21AC4E4629216A72FB7AE16FFABCF8BD" box="[1175,1289,1832,1857]" italics="true" pageId="3" pageNumber="74">posticatus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="7749EFA529216A72FA8CE16FFF66F89A" author="Sassi, D." pageId="3" pageNumber="74" pagination="201 - 237" refId="ref10505" refString="Sassi, D. (2024) Revision of the Griburius posticatus (Suffrian, 1866) species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 5406 (2), 201-237. https://doi.org/10.11646/zootaxa.5406.2.1" type="journal article" year="2024">Sassi 2024</bibRefCitation>
), such as
<taxonomicName id="D4D8E9D729216A72FEA5E10BFD17F899" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[328,674,1867,1894]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="21AC4E4629216A72FEA5E10BFE44F899" box="[328,497,1867,1894]" italics="true" pageId="3" pageNumber="74">G. ephippiatus</emphasis>
<bibRefCitation id="7749EFA529216A72FE17E10CFD17F899" author="Suffrian, E." box="[506,674,1867,1893]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="D4D8E9D729216A72FD0FE10BFB9FF899" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[738,1066,1867,1894]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="posticatus">
<emphasis id="21AC4E4629216A72FD0FE10BFCCCF899" box="[738,889,1868,1894]" italics="true" pageId="3" pageNumber="74">G. posticatus</emphasis>
<bibRefCitation id="7749EFA529216A72FC6EE10CFB9FF899" author="Suffrian, E." box="[899,1066,1867,1893]" pageId="3" pageNumber="74" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
. Nevertheless,
<taxonomicName id="D4D8E9D729216A72FB37E10BFAF3F899" authorityName="Suffrian" authorityYear="1866" box="[1242,1350,1868,1893]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FB37E10BFAF3F899" box="[1242,1350,1868,1893]" italics="true" pageId="3" pageNumber="74">M. minax</emphasis>
</taxonomicName>
can be easily distinguished from them by its more cylindrical profile, which is not dorsally flattened as seen in the typical
<taxonomicName id="D4D8E9D729216A72FF7AE1D4FEB0F851" authorityName="Haldeman" authorityYear="1849" box="[151,261,1939,1965]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="21AC4E4629216A72FF7AE1D4FEB0F851" box="[151,261,1939,1965]" italics="true" pageId="3" pageNumber="74">Griburius</emphasis>
</taxonomicName>
habitus of the aforementioned species. Additionally,
<taxonomicName id="D4D8E9D729216A72FC80E1D3FC6CF851" authorityName="Suffrian" authorityYear="1866" box="[877,985,1940,1965]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629216A72FC80E1D3FC6CF851" box="[877,985,1940,1965]" italics="true" pageId="3" pageNumber="74">M. minax</emphasis>
</taxonomicName>
is clearly distinct in terms of aedeagal morphology and the shape of the spermatheca, where the ductus, contrary to what is generally observed in
<taxonomicName id="D4D8E9D729216A72FACAE1F0FA20F82D" authorityName="Haldeman" authorityYear="1849" box="[1319,1429,1975,2001]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="21AC4E4629216A72FACAE1F0FA20F82D" box="[1319,1429,1975,2001]" italics="true" pageId="3" pageNumber="74">Griburius</emphasis>
</taxonomicName>
, is long, frail and tightly coiled.
</paragraph>
</subSubSection>
<caption id="47A7C2DC29266A75FF7AE129FC7AF80F" ID-DOI="http://doi.org/10.5281/zenodo.14798004" ID-Zenodo-Dep="14798004" httpUri="https://zenodo.org/record/14798004/files/figure.png" pageId="4" pageNumber="75" startId="4.[151,250,1902,1926]" targetBox="[167,1417,181,1861]" targetPageId="4" targetType="figure">
<paragraph id="1367925429266A75FF7AE129FC7AF80F" blockId="4.[151,1436,1902,2035]" pageId="4" pageNumber="75">
<emphasis id="21AC4E4629266A75FF7AE129FEA1F87A" bold="true" box="[151,276,1902,1927]" pageId="4" pageNumber="75">FIGURE 1.</emphasis>
<emphasis id="21AC4E4629266A75FEF1E128FE55F87A" box="[284,480,1903,1927]" italics="true" pageId="4" pageNumber="75">
<taxonomicName id="D4D8E9D729266A75FEF1E128FE69F87A" authorityName="Suffrian" authorityYear="1866" box="[284,476,1903,1927]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="minax">Metallactus minax</taxonomicName>
.
</emphasis>
ab: habitus, dorsal and lateral view (LT); c: frontal view, male (LT); d: frontal view, female (PLT, MNHUB); en: colour variations (e: female PLT, MNHUB; f: male, PT of
<taxonomicName id="D4D8E9D729266A75FC0CE1D4FB0FF857" authority=", Piracicaba" authorityName="Piracicaba" box="[993,1210,1939,1963]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629266A75FC0CE1D4FBF5F856" box="[993,1088,1939,1962]" italics="true" pageId="4" pageNumber="75">M. mosei</emphasis>
, Piracicaba
</taxonomicName>
, São Paulo, Brazil; g: male, “São Paulo”, Brazil; h: male, La Colmena, Paraguarí, Paraguay; k: male, “São Paulo”, Brazil; m: female, “Corocò”, Concepción, Paraguay; n: male, Jaguariaiva, Paraná, Brazil). Images not to scale.
</paragraph>
</caption>
<subSubSection id="5BC2C1DF29276A77FF2AE6D0FAC2FEE1" lastPageId="6" lastPageNumber="77" pageId="5" pageNumber="76" type="description">
<paragraph id="1367925429276A74FF2AE6D0FCB3FF29" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE6D0FE04FF4D" bold="true" box="[199,433,150,177]" italics="true" pageId="5" pageNumber="76">Description of male.</emphasis>
Habitus in figs 1ab (HT). BL =
<quantity id="D4203FB129276A74FCC6E6D1FC01FF4D" box="[811,948,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.7" metricValueMin="4.8" pageId="5" pageNumber="76" unit="mm" value="5.25" valueMax="5.7" valueMin="4.8">4.85.7 mm</quantity>
, BW =
<quantity id="D4203FB129276A74FBE6E6D0FB21FF4D" box="[1035,1172,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="5" pageNumber="76" unit="mm" value="3.15" valueMax="3.4" valueMin="2.9">2.93.4 mm</quantity>
, PL = 1.52.0 mm, PW =
<quantity id="D4203FB129276A74FF43E6FDFE8DFF29" box="[174,312,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="5" pageNumber="76" unit="mm" value="2.8" valueMax="3.1" valueMin="2.5">2.53.1 mm</quantity>
. Interocular distance 12.314.6 % of BL.
</paragraph>
<paragraph id="1367925429276A74FF2AE698FE7EFE09" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE698FEB1FF05" box="[199,260,223,249]" italics="true" pageId="5" pageNumber="76">Head</emphasis>
: usually completely black, occasionally with small, burred round yellow spot on frontoclypeal suture. Labrum light yellow. Surface of upper frons and vertex quite convex, head punctation sparse but well impressed on vertex, denser on clypeus and along inner rim of eyes, setosity almost missing on vertex and frons, but, again, thick and adpressed along lower part of inner eye margins. Mid-cranial suture apparent between upper half of eyes. Upper lobes of eyes well separated along median line. Ocular lines narrow, strictly adhering to ocular rim, marked by rows of small punctures. Ocular canthus rounded, scarcely setose. Antennae rather long, slender, blackish with first five antennomeres partly yellowish, 35 bright, subcylindrical, the subsequent ones dull, moderately flattened and more diffusedly setose.
</paragraph>
<paragraph id="1367925429276A74FF2AE7B8FBACFCE9" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE7B8FE83FDE4" box="[199,310,511,536]" italics="true" pageId="5" pageNumber="76">Pronotum</emphasis>
: black, often with longitudinal median yellow stripe extended from anterior to posterior margin, slightly larger posteriorly, spotted with small black marking just in front of scutellum. Lateral margins and external part of anterior and posterior ones yellow as well. Often yellow median stripe can be reduced so not attaining anterior and posterior margins. At times yellow pattern coalesces so black colour reduced to couple of large, oblique, wavy markings at sides of midline. In single specimen from Jaguariaíva (
<collectingRegion id="D11C5CB629276A74FC6DE4C8FC64FD55" box="[896,977,655,681]" country="Brazil" name="Parana" pageId="5" pageNumber="76">Paraná</collectingRegion>
,
<collectingCountry id="6BCFD2C429276A74FC31E4C8FB97FD55" box="[988,1058,655,681]" name="Brazil" pageId="5" pageNumber="76">Brazil</collectingCountry>
) pronotum totally black (
<figureCitation id="8BE38ED129276A74FAD1E4C8FA3BFD55" box="[1340,1422,655,681]" captionStart="FIGURE 1" captionStartId="4.[151,250,1902,1926]" captionTargetBox="[167,1417,181,1861]" captionTargetId="figure-33@4.[151,1435,181,1879]" captionTargetPageId="4" captionText="FIGURE 1. Metallactus minax. ab: habitus, dorsal and lateral view (LT); c: frontal view, male (LT); d: frontal view, female (PLT, MNHUB); en: colour variations (e: female PLT, MNHUB; f: male, PT of M. mosei, Piracicaba, São Paulo, Brazil; g: male, “São Paulo”, Brazil; h: male, La Colmena, Paraguarí, Paraguay; k: male, “São Paulo”, Brazil; m: female, “Corocò”, Concepción, Paraguay; n: male, Jaguariaiva, Paraná, Brazil). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.14798004" httpUri="https://zenodo.org/record/14798004/files/figure.png" pageId="5" pageNumber="76">Fig. 1n</figureCitation>
). Pronotal outline transverse, regularly convex. Lateral margins narrow, regularly curved with maximum width just behind half of length. Posterolateral impression short, shallow. Surface moderately lustrous with scattered, well-impressed punctation, denser on sides, almost obliterated on central part of disc.
</paragraph>
<paragraph id="1367925429276A74FF2AE558FB76FCA1" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<taxonomicName id="D4D8E9D729276A74FF2AE558FE80FCC5" box="[199,309,799,825]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="21AC4E4629276A74FF2AE558FE80FCC5" box="[199,309,799,825]" italics="true" pageId="5" pageNumber="76">Scutellum</emphasis>
</taxonomicName>
: completely black, moderately raised. Apex truncated in a straight line or mildly rounded. Surface smooth, minutely and sparsely punctured, with few, short, whitish setae above all on basal half.
</paragraph>
<paragraph id="1367925429276A74FF2AE520FBD1FB81" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE520FEB8FC7D" box="[199,269,871,897]" italics="true" pageId="5" pageNumber="76">Elytra</emphasis>
: yellow with common, large, transverse, elliptical black spot at middle of disc. Such black spot can be reduced or broken into separate smaller markings or, as in
<typeStatus id="CC632CF629276A74FCF0E5CCFC32FC59" box="[797,903,907,933]" pageId="5" pageNumber="76" type="lectotype">lectotype</typeStatus>
, prolonged along suture in broad stripe reaching anterior margin. In single studied specimen (
<figureCitation id="8BE38ED129276A74FD7BE5E8FD52FC35" box="[662,743,943,969]" captionStart="FIGURE 1" captionStartId="4.[151,250,1902,1926]" captionTargetBox="[167,1417,181,1861]" captionTargetId="figure-33@4.[151,1435,181,1879]" captionTargetPageId="4" captionText="FIGURE 1. Metallactus minax. ab: habitus, dorsal and lateral view (LT); c: frontal view, male (LT); d: frontal view, female (PLT, MNHUB); en: colour variations (e: female PLT, MNHUB; f: male, PT of M. mosei, Piracicaba, São Paulo, Brazil; g: male, “São Paulo”, Brazil; h: male, La Colmena, Paraguarí, Paraguay; k: male, “São Paulo”, Brazil; m: female, “Corocò”, Concepción, Paraguay; n: male, Jaguariaiva, Paraná, Brazil). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.14798004" httpUri="https://zenodo.org/record/14798004/files/figure.png" pageId="5" pageNumber="76">Fig. 1n</figureCitation>
) from Jaguariaíva (
<collectingRegion id="D11C5CB629276A74FC24E5E8FBAFFC35" box="[969,1050,943,969]" country="Brazil" name="Parana" pageId="5" pageNumber="76">Paraná</collectingRegion>
,
<collectingCountry id="6BCFD2C429276A74FBCBE5E8FBD9FC35" box="[1062,1132,943,969]" name="Brazil" pageId="5" pageNumber="76">Brazil</collectingCountry>
) black pattern extended to cover most of elytral surface leaving yellow colour only on humeral area and apex. Humeral callus always spotted with black even in lighter specimens. Elytral outline cylindrical with sides almost parallel. Lateral margins narrow, scarcely visible from above. Scutellar area not raised. Humeral callus scarcely prominent, not punctured. Elytral punctation fine and weak, rather close, irregularly arranged on anterior half, but hints of more regular rows toward apex. Intervals flat. Epipleura flat or slightly convex, with smooth, impunctate surface.
</paragraph>
<paragraph id="1367925429276A74FF2AE2C0FAACFB5D" blockId="5.[151,1437,150,2013]" box="[199,1305,1159,1185]" pageId="5" pageNumber="76">
<taxonomicName id="D4D8E9D729276A74FF2AE2C0FE87FB5D" authorityName="Meyen" authorityYear="1834" box="[199,306,1159,1185]" family="Trichomycteridae" genus="Pygidium" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="76" phylum="Chordata" rank="genus">
<emphasis id="21AC4E4629276A74FF2AE2C0FE87FB5D" box="[199,306,1159,1185]" italics="true" pageId="5" pageNumber="76">Pygidium</emphasis>
</taxonomicName>
: black, smooth, covered by thick, adpressed, whitish setae and sparse shallow punctures.
</paragraph>
<paragraph id="1367925429276A74FF2AE2ECFDBCFAA9" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE2ECFE89FB39" box="[199,316,1195,1221]" italics="true" pageId="5" pageNumber="76">Underside</emphasis>
: ventral surface black, matt, covered with long, thick, adpressed, yellowish setae, except shiny, scarcely setose, sparsely punctured posterior part of hypomera. Prosternal process narrow, fairly impressed at middle, with apex roughly triangular, slightly raised at tip. Surface of prosternal process covered with long, thick, semi-erect whitish setae. Fifth abdominal ventrite with deep, heart-shaped, bald, impunctate median depression and deeply notched posterior margin.
</paragraph>
<paragraph id="1367925429276A74FF2AE318FC59FA85" blockId="5.[151,1437,150,2013]" box="[199,1004,1375,1401]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE318FF49FA84" box="[199,252,1375,1400]" italics="true" pageId="5" pageNumber="76">Legs</emphasis>
: black with yellow stripe along anterior margin of anterior femora.
</paragraph>
<paragraph id="1367925429276A74FF2AE3C4FCDCF91D" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE3C4FEDEFA61" box="[199,363,1411,1437]" italics="true" pageId="5" pageNumber="76">Male genitalia</emphasis>
. Median lobe of aedeagus (
<figureCitation id="8BE38ED129276A74FD73E3C4FCBCFA61" box="[670,777,1411,1437]" captionStart="FIGURE 4" captionStartId="11.[151,250,1594,1618]" captionTargetBox="[171,1416,309,1554]" captionTargetId="figure-80@11.[171,1416,309,1570]" captionTargetPageId="11" captionText="FIGURE 4. ad: Metallactus minax, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg: M. tommasoi, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="5" pageNumber="76">Fig. 4ad</figureCitation>
) subcylindrical with apex blunt, flattened, spoon-shaped in ventral and dorsal view, almost straight in lateral view. Setose depressions shifted to very lateral border of ventral surface, in form of narrow, lengthened callosities, with short, scattered, barely visible setae. Ventral surface just below apex raised in large, squat, pointed median bulge whose distal margin, in lateral view, is fairly concave. At times, such bulge showing two symmetrical indentations (sd in fig. 4d) on apical side so that its base connected to apex by short carina. This pattern is not always evident, and in many cases, surface regularly slanted (
<figureCitation id="8BE38ED129276A74FAD1E070FA3AF9AD" box="[1340,1423,1591,1617]" captionStart="FIGURE 4" captionStartId="11.[151,250,1594,1618]" captionTargetBox="[171,1416,309,1554]" captionTargetId="figure-80@11.[171,1416,309,1570]" captionTargetPageId="11" captionText="FIGURE 4. ad: Metallactus minax, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg: M. tommasoi, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="5" pageNumber="76">Fig. 4a</figureCitation>
). Endophallus (
<figureCitation id="8BE38ED129276A74FEDBE01CFE3CF989" box="[310,393,1626,1653]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="5" pageNumber="76">Fig. 5a</figureCitation>
) with sclerite I well developed, with large straight-directed denticule. Dorsal spicule well developed, finger-like, with well pigmented apex. Sclerite II slender, slightly convex. Sclerite III robust, sickle shaped. Branches of sclerite IV with surface minutely wrinkled, shorter than sclerite III in folded-up structure, parallel-sided with rounded apex and long sharp process at base.
</paragraph>
<paragraph id="1367925429276A74FF2AE0ACFEC3F8D5" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">
<emphasis id="21AC4E4629276A74FF2AE0ACFEAEF8F9" box="[199,283,1771,1797]" italics="true" pageId="5" pageNumber="76">Female</emphasis>
. BL =
<quantity id="D4203FB129276A74FE9FE0ACFE4BF8F9" box="[370,510,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="7.1" metricValueMin="6.1" pageId="5" pageNumber="76" unit="mm" value="6.6" valueMax="7.1" valueMin="6.1">6.17.1 mm</quantity>
, BW =
<quantity id="D4203FB129276A74FDB0E0ACFD5DF8F9" box="[605,744,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="5" pageNumber="76" unit="mm" value="4.1" valueMax="4.3" valueMin="3.9">3.94.3 mm</quantity>
, PL = 2.0
<quantity id="D4203FB129276A74FC81E0ACFC73F8F9" box="[876,966,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="76" unit="mm" value="2.3">2.3 mm</quantity>
, PW =
<quantity id="D4203FB129276A74FBCCE0ACFB1BF8F9" box="[1057,1198,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="5" pageNumber="76" unit="mm" value="3.55" valueMax="3.8" valueMin="3.3">3.33.8 mm</quantity>
. Interocular distance 16.918.0 % of BL.
</paragraph>
<paragraph id="1367925429276A74FF2AE174FACCF869" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="76">Female differs in the larger interocular distance, in the pronotal colouration, which is usually totally black, reduced and finer punctation on the frontoclypeal surface. Additionally, often the ventral yellowish setosity is not as gaudy as in male and does not hide the surface of inferior parts. Besides, the anterior tarsi are slightly narrower.</paragraph>
<paragraph id="1367925429276A77FF2AE1D8FAC2FEE1" blockId="5.[151,1437,150,2013]" lastBlockId="6.[151,1437,151,609]" lastPageId="6" lastPageNumber="77" pageId="5" pageNumber="76">
The fifth abdominal ventrite in females has a large and deep pit. The bottom of the pit is glabrous, matt, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (
<figureCitation id="8BE38ED129276A74FBFEE184FBD2F821" box="[1043,1127,1986,2013]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="5" pageNumber="76">Fig. 5b</figureCitation>
) is moderately pigmented, sickle-shaped with slightly twisted, not swollen proximal lobe. The distal lobe is rather short, slender, tapered with an acute apex slightly bent downwards. The ampulla is not pigmented, short, sitting just at the basal apex of the vasculum. The duct and the sperm gland insertions are perceptibly distinct. The duct is very long, uniform in size, slender, floppy, strictly coiled, with its insertion on the bursa copulatrix forming a rigid, well pigmented sleeve.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29246A77FF2AE761FDFAFE75" pageId="6" pageNumber="77" type="biology_ecology">
<paragraph id="1367925429246A77FF2AE761FDFAFE75" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="77">
<emphasis id="21AC4E4629246A77FF2AE761FE30FEBD" bold="true" box="[199,389,294,321]" italics="true" pageId="6" pageNumber="77">Biological notes</emphasis>
.
<bibRefCitation id="7749EFA529246A77FE7BE760FDFCFEBD" author="Scholler, M." box="[406,585,295,321]" pageId="6" pageNumber="77" pagination="71 - 74" refId="ref10639" refString="Scholler, M. (2003) A new species of Metallactus Suffrian, 1866 (Coleoptera: Chrysomelidae: Cryptocephalinae) from Brazil. Russian Entomological Journal, 12 (1), 71-74." type="journal article" year="2003">Schöller (2003)</bibRefCitation>
reported some specimens of the
<typeStatus id="CC632CF629246A77FC22E760FC4AFEBD" box="[975,1023,295,321]" pageId="6" pageNumber="77">type</typeStatus>
series of
<taxonomicName id="D4D8E9D729246A77FB9BE760FB55FEBC" box="[1142,1248,295,320]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="mosei">
<emphasis id="21AC4E4629246A77FB9BE760FB55FEBC" box="[1142,1248,295,320]" italics="true" pageId="6" pageNumber="77">M. mosei</emphasis>
</taxonomicName>
to be reared on
<taxonomicName id="D4D8E9D729246A77FF7AE70CFEE1FE98" box="[151,340,331,357]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="77" phylum="Tracheophyta" rank="species" species="guajava">
<emphasis id="21AC4E4629246A77FF7AE70CFEE1FE98" box="[151,340,331,357]" italics="true" pageId="6" pageNumber="77">Psidium guajava</emphasis>
</taxonomicName>
(
<taxonomicName id="D4D8E9D729246A77FE8FE70CFE69FE99" box="[354,476,331,357]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="77" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) and
<taxonomicName id="D4D8E9D729246A77FDF7E70CFCBCFE99" box="[538,777,331,357]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="77" phylum="Tracheophyta" rank="species" species="urophylla">
<emphasis id="21AC4E4629246A77FDF7E70CFCBCFE99" box="[538,777,331,357]" italics="true" pageId="6" pageNumber="77">Eucalyptus urophylla</emphasis>
</taxonomicName>
(
<taxonomicName id="D4D8E9D729246A77FCFAE70CFC24FE99" box="[791,913,331,357]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="77" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) (see also
<bibRefCitation id="7749EFA529246A77FBE8E70CFB65FE99" author="Agrain, F. A. &amp; Vento, B. &amp; Flinte, V. &amp; Reid, C. A. M. &amp; Chaboo, C. S." box="[1029,1232,331,357]" pageId="6" pageNumber="77" pagination="1 - 20" refId="ref9923" refString="Agrain, F. A., Vento, B., Flinte, V., Reid, C. A. M. &amp; Chaboo, C. S. (2024) Global macroecological patterns in host plant associations of Cryptocephalinae case-bearer leaf beetles (Coleoptera: Chrysomelidae), Biological Journal of the Linnean Society, 20, 1-20. https://doi.org/10.1093/biolinnean/blae041" type="journal article" year="2024">
Agrain
<emphasis id="21AC4E4629246A77FBB7E70CFB3EFE99" box="[1114,1163,331,357]" italics="true" pageId="6" pageNumber="77">et al</emphasis>
. 2024
</bibRefCitation>
for the evaluation of the degree in host plant association).
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29246A77FF2AE7D5FE71FD9D" pageId="6" pageNumber="77" type="discussion">
<paragraph id="1367925429246A77FF2AE7D5FE71FD9D" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="77">
<emphasis id="21AC4E4629246A77FF2AE7D5FE80FE51" bold="true" box="[199,309,402,429]" italics="true" pageId="6" pageNumber="77">Remarks.</emphasis>
Regarding the variability mentioned above, concerning the appearance of the ventral surface of the aedeagal median lobe (
<figureCitation id="8BE38ED129246A77FE77E7F0FDF1FE2D" box="[410,580,439,465]" captionStart="FIGURE 4" captionStartId="11.[151,250,1594,1618]" captionTargetBox="[171,1416,309,1554]" captionTargetId="figure-80@11.[171,1416,309,1570]" captionTargetPageId="11" captionText="FIGURE 4. ad: Metallactus minax, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg: M. tommasoi, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="6" pageNumber="77">Figs 4a and 4d</figureCitation>
), careful comparisons have revealed that this variability is distributed rather randomly and shows no correlation with either the external coloration or possible partitions of the distribution range. Additionally, different intermediate forms are present, which cannot be attributed to distinct alternative patterns. Based on current knowledge, it does not seem possible to assign taxonomic significance to the variations in this structure within this taxon.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,164 @@
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<subSubSection id="5BC2C1DF29236A70FF7AE2C0FC61FB5D" box="[151,980,1159,1185]" pageId="1" pageNumber="72" type="nomenclature">
<paragraph id="1367925429236A70FF7AE2C0FC61FB5D" blockId="1.[151,980,1159,1185]" box="[151,980,1159,1185]" pageId="1" pageNumber="72">
Species belonging to the
<taxonomicName id="D4D8E9D729236A70FE5CE2C0FC9CFB5D" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[433,809,1159,1185]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629236A70FE5CE2C0FD35FB5C" box="[433,640,1159,1185]" italics="true" pageId="1" pageNumber="72">Metallactus minax</emphasis>
<bibRefCitation id="7749EFA529236A70FD6AE2C0FC9CFB5D" author="Suffrian, E." box="[647,809,1159,1185]" pageId="1" pageNumber="72" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
species-group:
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29236A70FF7AE288FE45FA85" pageId="1" pageNumber="72" type="taxon_list">
<paragraph id="1367925429236A70FF7AE288FDBAFB15" blockId="1.[151,732,1231,1401]" box="[151,527,1231,1257]" pageId="1" pageNumber="72">
<taxonomicName id="D4D8E9D729236A70FF7AE288FDBAFB15" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[151,527,1231,1257]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629236A70FF7AE288FED3FB14" box="[151,358,1231,1257]" italics="true" pageId="1" pageNumber="72">Metallactus minax</emphasis>
<bibRefCitation id="7749EFA529236A70FE83E288FDBAFB15" author="Suffrian, E." box="[366,527,1231,1257]" pageId="1" pageNumber="72" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1367925429236A70FF2AE2B4FD70FAF1" blockId="1.[151,732,1231,1401]" box="[199,709,1267,1293]" pageId="1" pageNumber="72">
<emphasis id="21AC4E4629236A70FF2AE2B4FF6FFAF0" box="[199,218,1267,1292]" italics="true" pageId="1" pageNumber="72">=</emphasis>
<taxonomicName id="D4D8E9D729236A70FF0CE2B4FDE2FAF1" authority="Scholler, 2003" authorityName=", Scholler" authorityYear="2003" box="[225,599,1267,1293]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="mosei" status="syn. nov.">
<emphasis id="21AC4E4629236A70FF0CE2B4FE18FAF0" box="[225,429,1267,1293]" italics="true" pageId="1" pageNumber="72">Metallactus mosei</emphasis>
<bibRefCitation id="7749EFA529236A70FE58E2B4FDE2FAF1" author="Scholler, M." box="[437,599,1267,1293]" pageId="1" pageNumber="72" pagination="71 - 74" refId="ref10639" refString="Scholler, M. (2003) A new species of Metallactus Suffrian, 1866 (Coleoptera: Chrysomelidae: Cryptocephalinae) from Brazil. Russian Entomological Journal, 12 (1), 71-74." type="journal article" year="2003">Schöller, 2003</bibRefCitation>
</taxonomicName>
<emphasis id="21AC4E4629236A70FDB3E2B4FD70FAF1" bold="true" box="[606,709,1267,1293]" pageId="1" pageNumber="72">
<taxonomicNameLabel id="3A9FF33D29236A70FDB3E2B4FD70FAF1" box="[606,709,1267,1293]" pageId="1" pageNumber="72">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="1367925429236A70FF7AE350FD7BFACD" blockId="1.[151,732,1231,1401]" box="[151,718,1303,1329]" pageId="1" pageNumber="72">
<taxonomicName id="D4D8E9D729236A70FF7AE350FDF2FACD" authority="(Suffrian, 1866)" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[151,583,1303,1329]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="abstersus" status="comb. nov.">
<emphasis id="21AC4E4629236A70FF7AE350FE39FACD" box="[151,396,1303,1329]" italics="true" pageId="1" pageNumber="72">Metallactus abstersus</emphasis>
(
<bibRefCitation id="7749EFA529236A70FE76E350FD8AFACD" author="Suffrian, E." box="[411,575,1303,1329]" pageId="1" pageNumber="72" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
<emphasis id="21AC4E4629236A70FDA3E350FD7BFACD" bold="true" box="[590,718,1303,1329]" pageId="1" pageNumber="72">
<taxonomicNameLabel id="3A9FF33D29236A70FDA3E350FD7BFACD" box="[590,718,1303,1329]" pageId="1" pageNumber="72" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="1367925429236A70FF2AE37CFD69FAA9" blockId="1.[151,732,1231,1401]" box="[199,732,1339,1365]" pageId="1" pageNumber="72">
<emphasis id="21AC4E4629236A70FF2AE37CFF6FFAA8" box="[199,218,1339,1364]" italics="true" pageId="1" pageNumber="72">=</emphasis>
<taxonomicName id="D4D8E9D729236A70FF0CE37CFDD8FAA9" authority="Suffrian, 1866" authorityName=", Suffrian" authorityYear="1866" box="[225,621,1339,1365]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="incultus" status="syn. nov.">
<emphasis id="21AC4E4629236A70FF0CE37CFE70FAA9" box="[225,453,1339,1365]" italics="true" pageId="1" pageNumber="72">Metallactus incultus</emphasis>
<bibRefCitation id="7749EFA529236A70FE21E37CFDD8FAA9" author="Suffrian, E." box="[460,621,1339,1365]" pageId="1" pageNumber="72" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
<emphasis id="21AC4E4629236A70FD99E37CFD69FAA9" bold="true" box="[628,732,1339,1365]" pageId="1" pageNumber="72">
<taxonomicNameLabel id="3A9FF33D29236A70FD99E37CFD69FAA9" box="[628,732,1339,1365]" pageId="1" pageNumber="72">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="1367925429236A70FF7AE318FE45FA85" blockId="1.[151,732,1231,1401]" box="[151,496,1375,1401]" pageId="1" pageNumber="72">
<taxonomicName id="D4D8E9D729236A70FF7AE318FE3AFA84" authorityName="Sassi" authorityYear="2025" box="[151,399,1375,1401]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="tommasoi" status="sp. nov.">
<emphasis id="21AC4E4629236A70FF7AE318FE3AFA84" box="[151,399,1375,1401]" italics="true" pageId="1" pageNumber="72">Metallactus tommasoi</emphasis>
</taxonomicName>
<emphasis id="21AC4E4629236A70FE7BE318FE45FA85" bold="true" box="[406,496,1375,1401]" pageId="1" pageNumber="72">
<taxonomicNameLabel id="3A9FF33D29236A70FE7BE318FE45FA85" box="[406,496,1375,1401]" pageId="1" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29236A73FF7AE38DFEC2FC59" lastPageId="2" lastPageNumber="73" pageId="1" pageNumber="72" type="description">
<paragraph id="1367925429236A70FF7AE38DFDC2FA19" blockId="1.[151,631,1482,1509]" box="[151,631,1482,1509]" pageId="1" pageNumber="72">
<heading id="482F253829236A70FF7AE38DFDC2FA19" bold="true" box="[151,631,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="72" reason="1">
<emphasis id="21AC4E4629236A70FF7AE38DFDC2FA19" bold="true" box="[151,631,1482,1509]" italics="true" pageId="1" pageNumber="72">General characters of the species group</emphasis>
</heading>
</paragraph>
<paragraph id="1367925429236A70FF7AE04AFD27F992" blockId="1.[151,1437,1548,2043]" pageId="1" pageNumber="72">
In terms of overall external appearance, size, and dorsal coloration pattern, the species belonging to this group are comparable to those recently assigned to the
<taxonomicName id="D4D8E9D729236A70FD7DE077FBB5F9B6" authority="(Perty, 1832)" baseAuthorityName="Perty" baseAuthorityYear="1832" box="[656,1024,1584,1610]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="21AC4E4629236A70FD7DE077FCD2F9B6" box="[656,871,1584,1610]" italics="true" pageId="1" pageNumber="72">Metallactus kollari</emphasis>
(
<bibRefCitation id="7749EFA529236A70FC98E077FC42F9B6" author="Perty, M." box="[885,1015,1584,1610]" pageId="1" pageNumber="72" refId="ref10186" refString="Perty, M. (1832) Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII - MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis augustissimi peracto collegerunt Dr. JB de Spix et Dr. CF Ph. de Martius: accedit dissertatio de insectorum in America Meridionali habitantium vitae genere moribus et distributione geographica. Impensis Editoris, Monachii, 224 pp." type="book" year="1832">Perty, 1832</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D4D8E9D729236A70FBDAE077FF7AF992" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="generosus">
<emphasis id="21AC4E4629236A70FBDAE077FA86F9B6" box="[1079,1331,1584,1610]" italics="true" pageId="1" pageNumber="72">Metallactus generosus</emphasis>
<bibRefCitation id="7749EFA529236A70FAD7E077FF7AF992" author="Suffrian, E." pageId="1" pageNumber="72" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
species complexes (
<bibRefCitation id="7749EFA529236A70FE58E013FD87F992" author="Sassi, D." box="[437,562,1620,1646]" pageId="1" pageNumber="72" pagination="57 - 110" refId="ref10255" refString="Sassi, D. (2018) Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4413 (1), 57-110. https://doi.org/10.11646/zootaxa.4413.1.2" type="journal article" year="2018">Sassi 2018</bibRefCitation>
,
<bibRefCitation id="7749EFA529236A70FDD0E013FD33F992" author="Sassi, D." box="[573,646,1620,1646]" pageId="1" pageNumber="72" pagination="148 - 162" refId="ref10350" refString="Sassi, D. (2020 a) Revision of the Metallactus generosus species-group with a preliminary evaluation of the effectiveness of the endophallus morphology in distinguishing critical sibling species (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4877 (1), 148-162. https://doi.org/10.11646/zootaxa.4877.1.6" type="journal article" year="2020">2020a</bibRefCitation>
).
</paragraph>
<paragraph id="1367925429236A70FF2AE03FFC1AF896" blockId="1.[151,1437,1548,2043]" pageId="1" pageNumber="72">Distinctive features of the species group proposed here are primarily based on the shape of the median lobe of the aedeagus, which has a rounded spoon-like apex and a characteristic more or less pointed tubercle below the apex on the ventral surface. In two of the three species, this tubercle is particularly developed, forming a conspicuous sharp, conical process. Additionally, there is an unusually deep, heart-shaped depression on the fifth abdominal ventrite of the male. On the contrary, in the above-mentioned species complexes, the ventral aedeagal surface is basically smooth and devoid of particular structures, and the depression on the male fifth anal sternite is definitely shallower, with its rear margin only weakly notched or almost straight.</paragraph>
<paragraph id="1367925429236A73FF2AE133FA87FEE1" blockId="1.[151,1437,1548,2043]" lastBlockId="2.[151,1437,151,825]" lastPageId="2" lastPageNumber="73" pageId="1" pageNumber="72">
The overall dorsal coloration shows significant individual variability, which can create considerable difficulties in correctly identifying the species when examination of the median lobe of the aedeagus is not possible. However, a trait that seems to characterize this group of species quite well, compared to those mentioned above, is the coloration of the legs and ventral parts, which are generally totally or almost totally black, contrary to what is observed, for example, in the
<taxonomicName id="D4D8E9D729206A73FEA4E6D0FDAAFF4D" baseAuthorityName="Perty" baseAuthorityYear="1832" box="[329,543,151,177]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="21AC4E4629206A73FEA4E6D0FDAAFF4D" box="[329,543,151,177]" italics="true" pageId="2" pageNumber="73">Metallactus kollari</emphasis>
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group, where specimens with predominantly light coloration on the ventral side and legs prevail. Additionally, the almost entirely black coloration of the forehead (rarely there is a small, blurred light marking between the antennal insertions), in both males and females, distinguishes the species
<taxonomicName id="D4D8E9D729206A73FAEFE698FAD9FF04" authorityName="Suffrian" authorityYear="1866" box="[1282,1388,223,248]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629206A73FAEFE698FAD9FF04" box="[1282,1388,223,248]" italics="true" pageId="2" pageNumber="73">M. minax</emphasis>
</taxonomicName>
and
<taxonomicName id="D4D8E9D729206A73FF7AE744FE92FEE1" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[151,295,259,285]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629206A73FF7AE744FE92FEE1" box="[151,295,259,285]" italics="true" pageId="2" pageNumber="73">M. abstersus</emphasis>
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, while in the third species (
<taxonomicName id="D4D8E9D729206A73FDB0E744FD5AFEE0" authorityName="Sassi" authorityYear="2025" box="[605,751,259,284]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="tommasoi">
<emphasis id="21AC4E4629206A73FDB0E744FD5AFEE0" box="[605,751,259,284]" italics="true" pageId="2" pageNumber="73">M. tommasoi</emphasis>
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) the clypeofrontal surface is predominantly yellow.
</paragraph>
<paragraph id="1367925429206A73FF2AE760FADFFDC1" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="73">The pronotal colouration can be totally black to black with margins and a longitudinal median stripe yellow. The elytral coloration is variable but falls into patterns that are quite common within the entire genus, namely a bicolored motif, with light parts in a yellowish or brick-red color, and dark parts represented by black spots variable in number, shape, and size, but with a predominant pattern consisting of a common large elliptical spot located in the center of the two elytra. This spot can appear significantly reduced, turning into a more or less oblique, separated pattern on each elytron, or it can be represented by a broad sutural band accompanied by other spots arranged longitudinally along the elytral disc. Specimens with very reduced and almost absent black patterns, as well as nearly completely melanistic specimens, have been observed, and might be not uncommon within all the species of the complex.</paragraph>
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The endophallus (
<figureCitation id="8BE38ED129206A73FE7FE400FE56FD9D" box="[402,483,582,609]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="2" pageNumber="73">Fig. 5a</figureCitation>
) shows the typical organization already described for all the other groups (see
<bibRefCitation id="7749EFA529206A73FA8EE400FF7AFD79" author="Sassi, D." pageId="2" pageNumber="73" pagination="57 - 110" refId="ref10255" refString="Sassi, D. (2018) Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4413 (1), 57-110. https://doi.org/10.11646/zootaxa.4413.1.2" type="journal article" year="2018">Sassi 2018</bibRefCitation>
for the terminology and figures) and regarded as an important trait in the delimitation of the genus. In this structure, the dorsal spicules are well-developed, with the apex aligned with the upper edge of the first sclerite in the folded-up structure. The fourth sclerite is broad, with wrinkled surface.
</paragraph>
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The vasculum of the spermatheca (
<figureCitation id="8BE38ED129206A73FDB3E490FD66FD0D" box="[606,723,726,753]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="2" pageNumber="73">Figs 5bd</figureCitation>
) is C-shaped, with non-swollen lobes, the distal lobe slightly shorter than the basal one. The ductus is thin, rather long, and tightly coiled. The insertion of the ductus onto the bursa of the spermatheca is visibly sclerotized and elongated to form a pigmented sleeve.
</paragraph>
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<emphasis id="21AC4E4629206A73FF7AE5CCFEC2FC59" bold="true" box="[151,375,907,933]" italics="true" pageId="2" pageNumber="73">Species treatments</emphasis>
</paragraph>
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@ -0,0 +1,367 @@
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<taxonomicName id="D4D8E9D729246A77FF7AE4F5FDDCFD30" authority="(Suffrian, 1866)" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[151,617,690,717]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="abstersus" status="comb. nov.">
<emphasis id="21AC4E4629246A77FF7AE4F5FE29FD31" bold="true" box="[151,412,690,717]" italics="true" pageId="6" pageNumber="77">Metallactus abstersus</emphasis>
(
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)
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<taxonomicNameLabel id="3A9FF33D29246A77FD9CE4F4FD42FD31" box="[625,759,691,717]" pageId="6" pageNumber="77" rank="species">comb. nov.</taxonomicNameLabel>
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(
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;
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;
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<emphasis id="21AC4E4629246A77FF7AE55CFEC5FCCF" box="[151,368,795,819]" italics="true" pageId="6" pageNumber="77">Scolochrus abstersus</emphasis>
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[
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<emphasis id="21AC4E4629246A77FCDEE55CFC22FCCF" box="[819,919,795,819]" italics="true" pageId="6" pageNumber="77">Griburius</emphasis>
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[
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<emphasis id="21AC4E4629246A77FADDE55CFA21FCCF" box="[1328,1428,795,819]" italics="true" pageId="6" pageNumber="77">Griburius</emphasis>
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<treatmentCitation id="9279B44529246A77FF0CE51FFD33FC8C" author="Suffrian, E." box="[225,646,856,880]" page="358" pageId="6" pageNumber="77" year="1866">
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<emphasis id="21AC4E4629246A77FF0CE51FFE04FC8C" box="[225,433,856,880]" italics="true" pageId="6" pageNumber="77">Metallactus incultus</emphasis>
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;
<treatmentCitation id="9279B44529246A77FD7EE51FFCD6FC8D" author="Clavareau, C. H." box="[659,867,856,881]" page="93" pageId="6" pageNumber="77" year="1913">
<bibRefCitation id="7749EFA529246A77FD7EE51FFCD6FC8D" author="Clavareau, C. H." box="[659,867,856,881]" pageId="6" pageNumber="77" pagination="1 - 278" refId="ref10059" refString="Clavareau, C. H. (1913) Pars 53: Chrysomelidae: Megascelinae, Megalopodinae, Clytrinae, Cryptocephalinae, Chlamydinae, Lamprosominae. In: Schenkling S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1-278." type="book chapter" year="1913">Clavareau, 1913: 93</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="9279B44529246A77FC03E51FFB51FC8C" author="Blackwelder, R. E." box="[1006,1252,856,880]" page="641" pageId="6" pageNumber="77" year="1946">
<bibRefCitation id="7749EFA529246A77FC03E51FFB51FC8C" author="Blackwelder, R. E." box="[1006,1252,856,880]" pageId="6" pageNumber="77" pagination="551 - 763" refId="ref10002" refString="Blackwelder, R. E. (1946) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 4. Bulletin of the United States National Museum, 185, 551-763. https://doi.org/10.5479/si.03629236.185.4" type="journal article" year="1946">Blackwelder, 1946: 641</bibRefCitation>
</treatmentCitation>
(catalogue).
<emphasis id="21AC4E4629246A77FA80E51FFF46FC73" bold="true" pageId="6" pageNumber="77">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29246A76FF7AE5F9FB0CFEBD" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77" type="discussion">
<paragraph id="1367925429246A77FF7AE5F9FC00FA9F" blockId="6.[151,1437,957,2028]" pageId="6" pageNumber="77">
<emphasis id="21AC4E4629246A77FF7AE5F9FF55FC24" bold="true" box="[151,224,958,984]" italics="true" pageId="6" pageNumber="77">Types.</emphasis>
As regard
<taxonomicName id="D4D8E9D729246A77FEB0E5FAFCB3FC2B" authority=", Suffrian (1866)" authorityName=", Suffrian" authorityYear="1866" box="[349,774,957,984]" genus="Scolochrus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="77" rank="species" species="abstersus">
<emphasis id="21AC4E4629246A77FEB0E5FAFDFEFC2B" box="[349,587,957,983]" italics="true" pageId="6" pageNumber="77">Scolochrus abstersus</emphasis>
,
<bibRefCitation id="7749EFA529246A77FDB4E5FAFCB3FC2B" author="Suffrian, E." box="[601,774,957,983]" pageId="6" pageNumber="77" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian (1866)</bibRefCitation>
</taxonomicName>
did not mention the number of specimens under study, but he reported that he could examine only females, and the material (hosted in MNHUB) was collected by Virmond in
<collectingCountry id="6BCFD2C429246A77FF7AE242FF6AFBE3" box="[151,223,1029,1055]" name="Brazil" pageId="6" pageNumber="77">Brazil</collectingCountry>
. One single specimen matching the original description was tracked down in MNHUB. Curiously, and once again, the specimen turned out to be male. For the same reasons pointed out concerning the typification of
<taxonomicName id="D4D8E9D729246A77FADEE26DFA29FBBF" authorityName="Suffrian" authorityYear="1866" box="[1331,1436,1066,1091]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629246A77FADEE26DFA29FBBF" box="[1331,1436,1066,1091]" italics="true" pageId="6" pageNumber="77">M. minax</emphasis>
</taxonomicName>
(see above), it seemed reasonable to assume that Suffrian erroneously believed the likely single available specimen to be a female. Therefore, the specimen in MNHUB, whose external habitus perfectly matches the original description, has been designated as the
<typeStatus id="CC632CF629246A77FE27E2D2FD81FB53" box="[458,564,1173,1199]" pageId="6" pageNumber="77" type="lectotype">lectotype</typeStatus>
. The typification was made as follows.
<typeStatus id="CC632CF629246A77FC18E2D2FBC0FB53" box="[1013,1141,1173,1199]" pageId="6" pageNumber="77" type="lectotype">LECTOTYPE</typeStatus>
:
<specimenCount id="05DE59DD29246A77FB92E2D1FB23FB4C" box="[1151,1174,1174,1200]" count="1" pageId="6" pageNumber="77" type="male"></specimenCount>
, glued, aedeagus glued on the same card, abdomen glued on a separate card, //
<taxonomicName id="D4D8E9D729246A77FCF9E2FEFC67FB2F" authority="Suffr." authorityName=", Suffrian" authorityYear="1866" box="[788,978,1209,1235]" genus="Scolochrus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="77" rank="species" species="abstersus">“abstersus Suffr.</taxonomicName>
*
<collectingCountry id="6BCFD2C429246A77FC08E2FEFB9DFB2F" box="[997,1064,1209,1235]" name="Brazil" pageId="6" pageNumber="77">Brasil</collectingCountry>
Virm.” [blue label, handwritten] // “24027” [white label, printed] // “
<taxonomicName id="D4D8E9D729246A77FDD5E29AFC6FFB0B" authority="(Suffrian, 1866)" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[568,986,1245,1271]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629246A77FDD5E29AFCAFFB0B" box="[568,794,1245,1271]" italics="true" pageId="6" pageNumber="77">Griburius abstersus</emphasis>
(
<bibRefCitation id="7749EFA529246A77FCC6E29AFC67FB0B" author="Suffrian, E." box="[811,978,1245,1271]" pageId="6" pageNumber="77" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
(
<taxonomicName id="D4D8E9D729246A77FC00E29AFB68FB0B" authorityName=", Suffrian" authorityYear="1866" box="[1005,1245,1245,1271]" genus="Scolochrus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="77" rank="species" species="abstersus">
<emphasis id="21AC4E4629246A77FC00E29AFB68FB0B" box="[1005,1245,1245,1271]" italics="true" pageId="6" pageNumber="77">Scolochrus abstersus</emphasis>
</taxonomicName>
)
<typeStatus id="CC632CF629246A77FB02E29AFAC2FB0B" box="[1263,1399,1245,1271]" pageId="6" pageNumber="77" type="lectotype">LECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] // “
<taxonomicName id="D4D8E9D729246A77FDFBE346FB99FAE7" authority="(Suffrian, 1866) D. Sassi" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[534,1068,1281,1308]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629246A77FDFBE346FCB9FAE7" box="[534,780,1281,1307]" italics="true" pageId="6" pageNumber="77">Metallactus abstersus</emphasis>
(
<bibRefCitation id="7749EFA529246A77FCF7E346FC0AFAE7" author="Suffrian, E." box="[794,959,1281,1307]" pageId="6" pageNumber="77" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
) D. Sassi
</taxonomicName>
det. 2024” [white label, printed] // (MNHUB). The information on the original label, matches the registration data from the old catalogue of the MNHUB (“24027 [
<taxonomicName id="D4D8E9D729246A77FE9DE30EFE47FA9F" authorityName="Suffrian" authorityYear="1852" box="[368,498,1353,1379]" genus="Scolochrus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="77" rank="genus">Scolochrus</taxonomicName>
]
<taxonomicName id="D4D8E9D729246A77FE12E30EFD1AFA9F" authority="Suffr." authorityName="Suffr." box="[511,687,1353,1379]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="abstersus">abstersus Suffr.</taxonomicName>
* 1.
<collectingCountry id="6BCFD2C429246A77FD31E30EFC97FA9F" box="[732,802,1353,1379]" name="Brazil" pageId="6" pageNumber="77">Brasil</collectingCountry>
, Virmund.”).
</paragraph>
<paragraph id="1367925429246A76FF1AE32AFB0CFEBD" blockId="6.[151,1437,957,2028]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77">
Regarding
<taxonomicName id="D4D8E9D729246A77FE99E32AFD71FA7B" authority=", Suffrian" authorityName=", Suffrian" authorityYear="1866" box="[372,708,1389,1416]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="incultus">
<emphasis id="21AC4E4629246A77FE99E32AFDEFFA7B" box="[372,602,1389,1415]" italics="true" pageId="6" pageNumber="77">Metallactus incultus</emphasis>
, Suffrian
</taxonomicName>
once again does not indicate the exact number of specimens he studied, but states that he saw only females. From the description provided, it can be inferred that there were at least three. Additionally, he lists MNHUB and NMV, as well as the Clark collection, currently housed in BMNH, as depositories of the material. My research has located
<specimenCount id="05DE59DD29246A77FD0FE39EFCC9FA08" box="[738,892,1497,1524]" count="4" pageId="6" pageNumber="77" type="generic" typeStatus="syntype">four syntypes</specimenCount>
, all females: one in MNHUB, one in BMNH, and two in NMV. The Berlin specimen matches perfectly with the description of the coloration that Suffrian seems to consider as 'typical,' and for this reason, it has been designated as the
<typeStatus id="CC632CF629246A77FC7DE066FC42F9C7" box="[912,1015,1569,1595]" pageId="6" pageNumber="77" type="lectotype">lectotype</typeStatus>
of the species. The information on the original label, matches the registration data from the old catalogue of the MNHUB (“23982 [
<taxonomicName id="D4D8E9D729246A77FB40E002FA83F9A3" authorityName="Suffrian" authorityYear="1866" box="[1197,1334,1605,1631]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
]
<taxonomicName id="D4D8E9D729246A77FAA9E002FF66F97F" authority="Suffr" authorityName=", Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="incultus">incultus Suffr</taxonomicName>
* 1. [
<collectingCountry id="6BCFD2C429246A77FEEAE02EFEE4F97F" box="[263,337,1641,1667]" name="Brazil" pageId="6" pageNumber="77">Brasil</collectingCountry>
], v. Olf.&quot;). The specimens from London and Vienna correspond, with some minor approximations, to what the German entomologist reports as variations in colour (essentially a reduction of the black elytral pattern, up to completely yellow elytra). These
<specimenCount id="05DE59DD29246A77FD8EE0F6FCAEF937" box="[611,795,1713,1739]" count="3" pageId="6" pageNumber="77" type="generic">three specimens</specimenCount>
have been designated as
<typeStatus id="CC632CF629246A77FBA5E0F6FB58F937" box="[1096,1261,1713,1739]" pageId="6" pageNumber="77" type="paralectotype">paralectotypes</typeStatus>
. Regarding the labels on the specimens from Vienna, which do not match what
<bibRefCitation id="7749EFA529246A77FCB2E092FBBEF913" author="Suffrian, E." box="[863,1035,1749,1775]" pageId="6" pageNumber="77" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian (1866)</bibRefCitation>
reported in the original description, it should be emphasized (as already pointed out above) that this issue pertains to all the material from Vienna used by Suffrian in his monograph. This matter has been discussed multiple times in previous works, to which the reader is referred for details (
<bibRefCitation id="7749EFA529246A77FE7CE106FDBBF8A7" author="Sassi, D." box="[401,526,1857,1883]" pageId="6" pageNumber="77" pagination="57 - 110" refId="ref10255" refString="Sassi, D. (2018) Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4413 (1), 57-110. https://doi.org/10.11646/zootaxa.4413.1.2" type="journal article" year="2018">Sassi 2018</bibRefCitation>
,
<bibRefCitation id="7749EFA529246A77FDF4E106FDE3F8A7" author="Sassi, D." box="[537,598,1857,1883]" pageId="6" pageNumber="77" pagination="251 - 282" refId="ref10460" refString="Sassi, D. (2022) Revision of the Metallactus taeniatellus species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 5125 (3), 251-282. https://doi.org/10.11646/zootaxa.5125.3.1" type="journal article" year="2022">2022</bibRefCitation>
). The typification was made as follows.
<typeStatus id="CC632CF629246A77FBF7E106FB2FF8A7" box="[1050,1178,1857,1883]" pageId="6" pageNumber="77" type="lectotype">LECTOTYPE</typeStatus>
:
<specimenCount id="05DE59DD29246A77FB49E105FB0CF8A0" box="[1188,1209,1858,1884]" count="1" pageId="6" pageNumber="77" type="female"></specimenCount>
, pinned, // &quot;
<taxonomicName id="D4D8E9D729246A77FAD2E106FF60F883" authority="Suffr" authorityName=", Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="incultus">incultus Suffr</taxonomicName>
.*
<collectingCountry id="6BCFD2C429246A77FF01E122FE9AF883" box="[236,303,1893,1919]" name="Brazil" pageId="6" pageNumber="77">Brasil</collectingCountry>
v. Olf.&quot; [blue label, handwritten] // &quot;23982&quot; [white label, printed] // “
<taxonomicName id="D4D8E9D729246A77FBA1E122FF7AF85F" authority="Suffrian, 1866" authorityName=", Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="incultus">
<emphasis id="21AC4E4629246A77FBA1E122FA87F883" box="[1100,1330,1893,1919]" italics="true" pageId="6" pageNumber="77">Metallactus incultus</emphasis>
<bibRefCitation id="7749EFA529246A77FAD7E122FF7AF85F" author="Suffrian, E." pageId="6" pageNumber="77" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
<typeStatus id="CC632CF629246A77FF3AE1CEFEEAF85F" box="[215,351,1929,1955]" pageId="6" pageNumber="77" type="lectotype">LECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] // “
<taxonomicName id="D4D8E9D729246A77FCE4E1CEFA95F85F" authority="(Suffrian, 1866) D. Sassi" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[777,1312,1929,1956]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629246A77FCE4E1CEFC4AF85F" box="[777,1023,1929,1955]" italics="true" pageId="6" pageNumber="77">Metallactus abstersus</emphasis>
(
<bibRefCitation id="7749EFA529246A77FBE3E1CEFB06F85F" author="Suffrian, E." box="[1038,1203,1929,1955]" pageId="6" pageNumber="77" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
) D. Sassi
</taxonomicName>
det. 2024” [white label, printed] //. (MNHUB). The information on the original label, matches the registration data from the old catalogue of the MNHUB (“23982 [
<taxonomicName id="D4D8E9D729246A77FDB9E196FD68F817" authorityName="Suffrian" authorityYear="1866" box="[596,733,2001,2027]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
]
<taxonomicName id="D4D8E9D729246A77FD07E196FC33F817" authority="Suffr" authorityName=", Suffrian" authorityYear="1866" box="[746,902,2001,2027]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="incultus">incultus Suffr</taxonomicName>
* 1. [
<collectingCountry id="6BCFD2C429246A77FC57E196FBB1F817" box="[954,1028,2001,2027]" name="Brazil" pageId="6" pageNumber="77">Brasil</collectingCountry>
], v. Olf.”).
<typeStatus id="CC632CF629246A77FB6CE196FA8EF817" box="[1153,1339,2001,2027]" pageId="6" pageNumber="77" type="paralectotype">PARALECTOTYPES</typeStatus>
(3):
<specimenCount id="05DE59DD29246A77FA9FE196FA20F810" box="[1394,1429,2001,2028]" count="1" pageId="6" pageNumber="77" type="female">1♀</specimenCount>
, pinned, // “brasilia” [white label, handwritten] // “162” [blue label, handwritten] // “E. Coll. Chev.
<superScript id="E4AD3F1C29256A76FB18E6D0FB4FFF59" attach="none" box="[1269,1274,151,165]" fontSize="6" pageId="7" pageNumber="78">t</superScript>
” [white label, printed] // “
<taxonomicName id="D4D8E9D729256A76FEF5E6FCFE1CFF29" box="[280,425,187,213]" genus="Met" pageId="7" pageNumber="78" rank="species" species="incultus">Met incultus</taxonomicName>
” [white label, handwritten] // “a D. Saffrian. [sic!] determinat.” [white label, handwritten] // “67.56” [white label, printed] // (BMNH);
<specimenCount id="05DE59DD29256A76FD7BE698FD0CFF05" box="[662,697,223,249]" count="2" pageId="7" pageNumber="78" type="female">2♀</specimenCount>
, pinned, // “Ytaranga
<collectingCountry id="6BCFD2C429256A76FC5AE698FBA9FF05" box="[951,1052,223,249]" name="Brazil" pageId="7" pageNumber="78">Brasilien</collectingCountry>
Natterer” [white label, printed] // (NMV). All
<typeStatus id="CC632CF629256A76FECCE744FE76FEE1" box="[289,451,259,285]" pageId="7" pageNumber="78" type="paralectotype">paralectotypes</typeStatus>
labelled: “
<taxonomicName id="D4D8E9D729256A76FDD6E744FC70FEE1" authority="Suffrian, 1866" authorityName=", Suffrian" authorityYear="1866" box="[571,965,259,285]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="incultus">
<emphasis id="21AC4E4629256A76FDD6E744FCAAFEE1" box="[571,799,259,285]" italics="true" pageId="7" pageNumber="78">Metallactus incultus</emphasis>
<bibRefCitation id="7749EFA529256A76FCC9E744FC70FEE1" author="Suffrian, E." box="[804,965,259,285]" pageId="7" pageNumber="78" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
<typeStatus id="CC632CF629256A76FC26E744FB3CFEE1" box="[971,1161,259,285]" pageId="7" pageNumber="78" type="paralectotype">PARALECTOTYPUS</typeStatus>
D. Sassi det.” [red label, printed] // “
<taxonomicName id="D4D8E9D729256A76FEF7E760FC98FEBD" authority="(Suffrian, 1866) D. Sassi" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[282,813,295,321]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629256A76FEF7E760FDBAFEBD" box="[282,527,295,321]" italics="true" pageId="7" pageNumber="78">Metallactus abstersus</emphasis>
(
<bibRefCitation id="7749EFA529256A76FDF0E760FD74FEBD" author="Suffrian, E." box="[541,705,295,321]" pageId="7" pageNumber="78" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
) D. Sassi
</taxonomicName>
det. 2024” [white label, printed] //.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29256A76FF2AE70CFDE1FE75" pageId="7" pageNumber="78" type="materials_examined">
<paragraph id="1367925429256A76FF2AE70CFE7EFE99" blockId="7.[151,1437,151,2013]" box="[199,459,330,357]" pageId="7" pageNumber="78">
<emphasis id="21AC4E4629256A76FF2AE70CFEEAFE99" bold="true" box="[199,351,330,357]" italics="true" pageId="7" pageNumber="78">
<typeStatus id="CC632CF629256A76FF2AE70CFF4BFE99" box="[199,254,331,357]" pageId="7" pageNumber="78">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="6BCFD2C429256A76FE9CE70CFE09FE99" box="[369,444,331,357]" name="Brazil" pageId="7" pageNumber="78">Brazil</collectingCountry>
”.
</paragraph>
<paragraph id="1367925429256A76FF2AE729FDE1FE75" blockId="7.[151,1437,151,2013]" box="[199,596,366,393]" pageId="7" pageNumber="78">
<emphasis id="21AC4E4629256A76FF2AE729FDB4FE75" bold="true" box="[199,513,366,393]" italics="true" pageId="7" pageNumber="78">Further material examined</emphasis>
. None.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29256A76FF2AE7D5FBD2FDE5" pageId="7" pageNumber="78" type="distribution">
<paragraph id="1367925429256A76FF2AE7D5FBD2FDE5" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="78">
<emphasis id="21AC4E4629256A76FF2AE7D5FEE6FE51" bold="true" box="[199,339,402,429]" italics="true" pageId="7" pageNumber="78">Distribution</emphasis>
.
<collectingCountry id="6BCFD2C429256A76FE8DE7D4FE1DFE51" box="[352,424,403,429]" name="Brazil" pageId="7" pageNumber="78">Brazil</collectingCountry>
.At present, it seems impossible to identify the exact geographic location of the only locality mentioned on the labels of the material preserved at NMV (Ytaranga). However, it is reasonable to assume that this is an old toponym from southern
<collectingCountry id="6BCFD2C429256A76FDACE79CFD3DFE09" box="[577,648,475,501]" name="Brazil" pageId="7" pageNumber="78">Brazil</collectingCountry>
, possibly from the state of
<collectingRegion id="D11C5CB629256A76FC51E79CFB84FE09" box="[956,1073,475,501]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="78">São Paulo</collectingRegion>
, where it is known that Natterer conducted extensive research and collected a large amount of material (
<bibRefCitation id="7749EFA529256A76FC5AE7B8FBEFFDE5" author="Schifter, H." box="[951,1114,511,537]" pageId="7" pageNumber="78" pagination="155 - 180" refId="ref10605" refString="Schifter, H. (1993) Johann Natterer und seine ornithologischen Entdeckungen in Brasilien, 1817 - 1835. Kataloge des Oberosterreichischen Landesmuseums, New Series, 61, 155-180." type="journal article" year="1993">Schifter, 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29256A76FF2AE464FCD2FAF1" pageId="7" pageNumber="78" type="discussion">
<paragraph id="1367925429256A76FF2AE464FCD2FAF1" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="78">
<emphasis id="21AC4E4629256A76FF2AE464FE12FDC1" bold="true" box="[199,423,547,573]" italics="true" pageId="7" pageNumber="78">Comparative notes.</emphasis>
For its dorsal coloration,
<taxonomicName id="D4D8E9D729256A76FD20E464FC76FDC1" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[717,963,547,573]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629256A76FD20E464FC76FDC1" box="[717,963,547,573]" italics="true" pageId="7" pageNumber="78">Metallactus abstersus</emphasis>
</taxonomicName>
closely resembles some species in the
<taxonomicName id="D4D8E9D729256A76FA93E464FECBFD9D" authority="(Perty, 1832)" baseAuthorityName="Perty" baseAuthorityYear="1832" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="21AC4E4629256A76FA93E464FF57FD9D" italics="true" pageId="7" pageNumber="78">M. kollari</emphasis>
(
<bibRefCitation id="7749EFA529256A76FF1FE400FEC0FD9D" author="Perty, M." box="[242,373,583,609]" pageId="7" pageNumber="78" refId="ref10186" refString="Perty, M. (1832) Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII - MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis augustissimi peracto collegerunt Dr. JB de Spix et Dr. CF Ph. de Martius: accedit dissertatio de insectorum in America Meridionali habitantium vitae genere moribus et distributione geographica. Impensis Editoris, Monachii, 224 pp." type="book" year="1832">Perty, 1832</bibRefCitation>
)
</taxonomicName>
species-group (
<bibRefCitation id="7749EFA529256A76FDDFE400FD04FD9D" author="Sassi, D." box="[562,689,583,609]" pageId="7" pageNumber="78" pagination="57 - 110" refId="ref10255" refString="Sassi, D. (2018) Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4413 (1), 57-110. https://doi.org/10.11646/zootaxa.4413.1.2" type="journal article" year="2018">Sassi 2018</bibRefCitation>
), particularly in specimens where there is a reduced black pattern on the elytra, as sometimes seen in
<taxonomicName id="D4D8E9D729256A76FDC6E42CFC22FD79" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[555,919,619,645]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="dodecastictus">
<emphasis id="21AC4E4629256A76FDC6E42CFD5CFD79" box="[555,745,619,645]" italics="true" pageId="7" pageNumber="78">M. dodecastictus</emphasis>
<bibRefCitation id="7749EFA529256A76FD1CE42CFC22FD79" author="Suffrian, E." box="[753,919,619,645]" pageId="7" pageNumber="78" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
. However, in the species of the
<taxonomicName id="D4D8E9D729256A76FAECE42CFAF9FD79" baseAuthorityName="Perty" baseAuthorityYear="1832" box="[1281,1356,619,645]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="21AC4E4629256A76FAECE42CFAF9FD79" box="[1281,1356,619,645]" italics="true" pageId="7" pageNumber="78">kollari</emphasis>
</taxonomicName>
group, the head, legs, and ventral parts typically have extensive or predominantly yellow colouration while in
<taxonomicName id="D4D8E9D729256A76FAE3E4C8FA29FD55" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1294,1436,655,681]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629256A76FAE3E4C8FA29FD55" box="[1294,1436,655,681]" italics="true" pageId="7" pageNumber="78">M. abstersus</emphasis>
</taxonomicName>
the ventral parts and the legs are completely black, and the head almost entirely black, sometimes with a hint of a dirty yellow spot on the lower part of the clypeus. Conversely, the upper lip is consistently a rather bright yellow, contrasting sharply with the dark colour of the rest of the head. This combination of colours is not found in other
<taxonomicName id="D4D8E9D729256A76FF7AE558FEAEFCC5" authorityName="Suffrian" authorityYear="1866" box="[151,283,799,825]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="21AC4E4629256A76FF7AE558FEAEFCC5" box="[151,283,799,825]" italics="true" pageId="7" pageNumber="78">Metallactus</emphasis>
</taxonomicName>
species-groups. The only species in the
<taxonomicName id="D4D8E9D729256A76FD33E558FC9CFCC5" baseAuthorityName="Perty" baseAuthorityYear="1832" box="[734,809,799,825]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="21AC4E4629256A76FD33E558FC9CFCC5" box="[734,809,799,825]" italics="true" pageId="7" pageNumber="78">kollari</emphasis>
</taxonomicName>
group with entirely black head and legs is
<taxonomicName id="D4D8E9D729256A76FAE4E558FE8BFCA1" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="21AC4E4629256A76FAE4E558FA29FCC5" box="[1289,1436,799,825]" italics="true" pageId="7" pageNumber="78">M. languidus</emphasis>
<bibRefCitation id="7749EFA529256A76FF7AE504FE8BFCA1" author="Suffrian, E." box="[151,318,835,861]" pageId="7" pageNumber="78" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
, but here the sides of the abdomen are bordered with yellow, and the elytral pattern observed so far is clearly different, being characterized by a series of small, rounded black spots (up to six), arranged in two rows on each elytron. Again, in
<taxonomicName id="D4D8E9D729256A76FE10E5CCFC9BFC59" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[509,814,907,933]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="albifrons">
<emphasis id="21AC4E4629256A76FE10E5CCFD33FC59" box="[509,646,907,933]" italics="true" pageId="7" pageNumber="78">M. albifrons</emphasis>
<bibRefCitation id="7749EFA529256A76FD60E5CCFC9BFC59" author="Suffrian, E." box="[653,814,907,933]" pageId="7" pageNumber="78" pagination="1 - 483" refId="ref10677" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1-483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
the abdomen and the legs are completely black, but the frons is extensive yellow. Anyway, the morphology of the aedeagus is always decisive, as the large tubercle on the ventral face of the median lobe of the aedeagus, characteristic of species in the
<taxonomicName id="D4D8E9D729256A76FBF1E594FBD5FC10" authorityName="Suffrian" authorityYear="1866" box="[1052,1120,979,1004]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629256A76FBF1E594FBD5FC10" box="[1052,1120,979,1004]" italics="true" pageId="7" pageNumber="78">minax</emphasis>
</taxonomicName>
group, is never observed in the
<taxonomicName id="D4D8E9D729256A76FF2FE5B0FEB8FBED" baseAuthorityName="Perty" baseAuthorityYear="1832" box="[194,269,1015,1041]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="21AC4E4629256A76FF2FE5B0FEB8FBED" box="[194,269,1015,1041]" italics="true" pageId="7" pageNumber="78">kollari</emphasis>
</taxonomicName>
group of species. The combination of the chromatic characters mentioned above (legs, head, upper lip and ventral parts) allows this species to be closely aligned with
<taxonomicName id="D4D8E9D729256A76FC8AE25CFC65FBC8" authorityName="Suffrian" authorityYear="1866" box="[871,976,1051,1076]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629256A76FC8AE25CFC65FBC8" box="[871,976,1051,1076]" italics="true" pageId="7" pageNumber="78">M. minax</emphasis>
</taxonomicName>
, whose affinity is especially evident due to the morphology of the aedeagus and the shape of the anal depression in the male. Males of
<taxonomicName id="D4D8E9D729256A76FB51E278FAF9FBA5" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1212,1356,1087,1113]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629256A76FB51E278FAF9FBA5" box="[1212,1356,1087,1113]" italics="true" pageId="7" pageNumber="78">M. abstersus</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="D4D8E9D729256A76FE35E224FDF6FB80" authorityName="Suffrian" authorityYear="1866" box="[472,579,1123,1148]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629256A76FE35E224FDF6FB80" box="[472,579,1123,1148]" italics="true" pageId="7" pageNumber="78">M. minax</emphasis>
</taxonomicName>
not only by the different color pattern but also by the shape of the median lobe of the aedeagus. In particular, the curvature of the median ventral bulge, whose apical profile in lateral view is distinctly convex in
<taxonomicName id="D4D8E9D729256A76FE4EE2ECFD83FB39" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[419,566,1195,1221]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629256A76FE4EE2ECFD83FB39" box="[419,566,1195,1221]" italics="true" pageId="7" pageNumber="78">M. abstersus</emphasis>
</taxonomicName>
, whereas it is concave in
<taxonomicName id="D4D8E9D729256A76FC86E2ECFC6DFB38" authorityName="Suffrian" authorityYear="1866" box="[875,984,1195,1220]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629256A76FC86E2ECFC6DFB38" box="[875,984,1195,1220]" italics="true" pageId="7" pageNumber="78">M. minax</emphasis>
</taxonomicName>
. Lastly, the shared colour pattern with females previously assigned by Suffrian to a different taxon
<emphasis id="21AC4E4629256A76FCAEE288FC79FB15" box="[835,972,1231,1257]" italics="true" pageId="7" pageNumber="78">
(
<taxonomicName id="D4D8E9D729256A76FCA1E288FC79FB15" authorityName=", Suffrian" authorityYear="1866" box="[844,972,1231,1257]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="incultus">M. incultus</taxonomicName>
</emphasis>
), and the substantial similarity in overall appearance, further justifies proposing the following synonymy:
</paragraph>
</subSubSection>
</treatment>
</document>