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<mods:title id="EA2F5CB3E6D0E3A15E4AEACA9B912D93">Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini)</mods:title>
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<mods:namePart>Derkarabetian, Shahan</mods:namePart>
<mods:affiliation>Department of Biology, San Diego State University, San Diego, CA 92182, USA &amp; Department of Organismic and Evolutionary Biology, Harvard University, Cambridge MA 02138, USA</mods:affiliation>
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<mods:namePart>Hedin, Marshal</mods:namePart>
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<treatment ID-GBIF-Taxon="163540048" LSID="urn:lsid:zoobank.org:act:4C656386-8B15-4B5C-BF3F-27805897C65F" httpUri="http://treatment.plazi.org/id/10AE52273E2F52CFA4AD23408A5379DA" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="10AE5227-3E2F-52CF-A4AD-23408A5379DA" authority="W. Maddison" authorityName="W. Maddison" authorityYear="2020" family="Salticidae" genus="Tomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomis manabita" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manabita" status="sp. nov.">Tomis manabita W. Maddison</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">Figures 120-128</figureCitation>
<treatment id="10AE52273E2F52CFA4AD23408A5379DA" ID-GBIF-Taxon="163540048" LSID="urn:lsid:zoobank.org:act:4C656386-8B15-4B5C-BF3F-27805897C65F" httpUri="http://treatment.plazi.org/id/10AE52273E2F52CFA4AD23408A5379DA" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Salticidae" scope_order="Araneae" scope_subTribe="Sitticina" scope_tribe="Sitticini">
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<taxonomicName id="60B61653CDCDF70049C8B6893673CDFF" ID-CoL="94BK6" LSID="10AE5227-3E2F-52CF-A4AD-23408A5379DA" authority="W. Maddison" authorityName="W. Maddison" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Tomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomis manabita" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manabita" status="sp. nov.">Tomis manabita W. Maddison</taxonomicName>
<taxonomicNameLabel id="BF5158D81F4CDAE0C80380AE66257AEA" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="9DAA6CE2C6100EF22D9BA592C25ADD0D" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">Figures 120-128</figureCitation>
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
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<paragraph id="2FB9D2DACD9E115091C4132F0001BA9F" pageId="0" pageNumber="1">Type material.</paragraph>
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Male holotype, 10 male and 8 female paratypes from Ecuador:
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="642BE0045B46B2BF024CBD656D59C213" originalValue="Manabí">Manabi</normalizedToken>
: Puerto Rico,
<normalizedToken originalValue="Cabañas">Cabanas</normalizedToken>
<normalizedToken id="B4BC7D9AC94B7F71DAB12B7F9838D6F4" originalValue="Cabañas">Cabanas</normalizedToken>
Alandaluz 5 May 1994 W. Maddison WPM#94-031. The holotype (specimen UBC-SEM AR00217) pertains to the Museum of Zoology, Pontificia Universidad Católica, Quito, Ecuador (QCAZ), but is currently held in the Spencer Entomological Collection at the Beaty Biodiversity Museum, University of British Columbia (UBC-SEM).
</paragraph>
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Based on the type locality; the form is the adjective in Spanish (masculine or feminine).</paragraph>
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<paragraph id="DC0A6C31B77A50B1C7A4873E35C5C959" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="56C47FD02A3316F58FF3AAC42765E106" pageId="0" pageNumber="1">Based on the type locality; the form is the adjective in Spanish (masculine or feminine).</paragraph>
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A species on the beaches of coastal Ecuador, resembling
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<emphasis italics="true" pageId="0" pageNumber="1">Attulus</emphasis>
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in habitus. It was used in the molecular phylogenetic study of
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<bibRefCitation id="F0F03C6E8F923B21C7DCB90397003E1A" DOI="https://doi.org/10.1071/IS02044" author="Maddison, WP" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="1" pagination="529 - 549" refId="B39" refString="Maddison, WP, Hedin, MC, 2003. Jumping spider phylogeny (Araneae: Salticidae). Invertebrate Systematics 17: 529 - 549, DOI: https://doi.org/10.1071/IS02044" title="Jumping spider phylogeny (Araneae: Salticidae)." url="https://doi.org/10.1071/IS02044" volume="17" year="2003">Maddison and Hedin (2003)</bibRefCitation>
under the name &quot;
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<emphasis italics="true" pageId="0" pageNumber="1">Sitticus</emphasis>
<taxonomicName id="949C2C96096AEBBBA81A5D9CAA038891" authorityName="Simon" authorityYear="1901" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="2B77F834138B40228A4CEDA0F33947BD" italics="true" pageId="0" pageNumber="1">Sitticus</emphasis>
</taxonomicName>
sp.&quot; (voucher S220) from
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="DC263BF471EB42192D28C9A634C16B3C" originalValue="Manabí">Manabi</normalizedToken>
, Ecuador.
</paragraph>
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="FF2BC83A904A02AF053171B33D4C77D1" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="2E155E69B32FC870399B26458D14D584" pageId="0" pageNumber="1">
Palp closely resembles that of
<taxonomicName family="Salticidae" genus="Tomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomis pavidus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pavidus">
<emphasis italics="true" pageId="0" pageNumber="1">Tomis pavidus</emphasis>
<taxonomicName id="26B85273DCBC9355549B06739C1F620C" class="Arachnida" family="Salticidae" genus="Tomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomis pavidus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pavidus">
<emphasis id="84C224CBBB1DFE80769127A34835F275" italics="true" pageId="0" pageNumber="1">Tomis pavidus</emphasis>
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, from which it differs in the smaller tibia and longer RTA.
</paragraph>
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
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<paragraph id="41BCF3675976BE1A38DDADDBDA01FF32" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="84EE3AC3B53D8C9BA90F33A1E11305C6" pageId="0" pageNumber="1">
<emphasis id="2D8FC017D0A9F2CBF3AC8F489F845056" bold="true" pageId="0" pageNumber="1">Male</emphasis>
(holotype). Carapace length 1.58; abdomen length 1.51.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
<emphasis id="20C648DFFE74A0BACD175F3DF348154D" bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
(Figs
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<figureCitation id="B70F0C1325895CCC5925594BC0755176" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">124</figureCitation>
,
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">127</figureCitation>
<figureCitation id="21B38568449FC0EF0D69AD3320653574" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">127</figureCitation>
) medium brown with recumbent brown setae except for thin medial longitudinal band of white setae on thorax, two spots of pale setae in ocular area, and a marginal band of white setae that is broad on the thorax but narrows to the anterior, disappearing before the clypeus is reached.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Clypeus</emphasis>
<emphasis id="7A4FF678C7EF10BE0AF59B1D80F5520F" bold="true" italics="true" pageId="0" pageNumber="1">Clypeus</emphasis>
brown, with a few brownish setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Chelicerae</emphasis>
<emphasis id="BDC13C54E1A87CE2F831A059587ABDAD" bold="true" italics="true" pageId="0" pageNumber="1">Chelicerae</emphasis>
vertical, with a few pale setae near the clypeus. Retromarginal cheliceral teeth lacking.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Palp</emphasis>
<emphasis id="7DF2685E5B3C95F030F172ADF806781A" bold="true" italics="true" pageId="0" pageNumber="1">Palp</emphasis>
clothed with white setae dorsally, but prolaterally with darker integument and setae from tip of femur to cymbium; cymbium mostly dark brown.
<taxonomicName class="Holothuroidea" family="Molpadiidae" genus="Embolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolus" order="Molpadida" pageId="0" pageNumber="1" phylum="Echinodermata" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Embolus</emphasis>
<taxonomicName id="21C46F0FB80188E65A8E1F86B6E5FF11" class="Holothuroidea" family="Molpadiidae" genus="Embolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolus" order="Molpadida" pageId="0" pageNumber="1" phylum="Echinodermata" rank="genus">
<emphasis id="A034D981360A91A5413BC233E07F8E9D" bold="true" italics="true" pageId="0" pageNumber="1">Embolus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">120</figureCitation>
<figureCitation id="AFC4E86637C23C872E6D2C945B37B6D3" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">120</figureCitation>
) arises broadly, more centrally beneath bulb (and less peripherally) than is typical, then narrows abruptly at ca. 9
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
<normalizedToken id="D8DE20E481842B9531A23498C1F79F78" originalValue="oclock">o'clock</normalizedToken>
. RTA extremely long and thin, parallel to axis of the palp.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
<emphasis id="3E627494C3F0407962FDB999A0EA13B2" bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
honey-coloured, with notably darker annulae at the tarsus-metatarsus joints, and black stripe on prolateral-ventral face of first patella and tibia.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
<emphasis id="D68381DBAEA476AC4E687522095369EF" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Figs
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">124</figureCitation>
<figureCitation id="5EB8427075F9E05566B43191864D6289" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">124</figureCitation>
,
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">127</figureCitation>
<figureCitation id="E64DB317337D20AA4679F9769E7330B2" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">127</figureCitation>
) brown with two lateral and one median longitudinal bands of paler setae, the medial band less distinct, wavy, and accompanied by a lateral extension that forms a cross.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
<paragraph id="71E5399115714BDF7D8AB5E87D4E35CD" pageId="0" pageNumber="1">
<emphasis id="8C11AFA22C13271581D3FA339E34961D" bold="true" pageId="0" pageNumber="1">Female</emphasis>
(paratype # UBC-SEM AR00218).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
<emphasis id="54DB5746AAE144F8B56AFD928DAE7EA1" bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
length 1.76;
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">abdomen</emphasis>
<emphasis id="ED6498681E9BEBDCD6B291707DE926E1" bold="true" italics="true" pageId="0" pageNumber="1">abdomen</emphasis>
length 2.22.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
<emphasis id="CC8DF2975A717BBE281E78174206600D" bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
(Figs
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">126</figureCitation>
<figureCitation id="A53A7129909F6D36B3D2F241AC26654C" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">126</figureCitation>
,
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">128</figureCitation>
<figureCitation id="00F8DD27C1B5B1DA58F3717E93D5E94A" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">128</figureCitation>
) brown, covered unevenly with recumbent cream-coloured setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Clypeus</emphasis>
<emphasis id="C7F5A80D539E61CF21ED7873983103C1" bold="true" italics="true" pageId="0" pageNumber="1">Clypeus</emphasis>
with white setae, longest at midline where they overhang the chelicerae. Chelicerae brown, with a few setae near clypeus.
<taxonomicName genus="Palps" lsidName="Palps" pageId="0" pageNumber="1" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Palps</emphasis>
<taxonomicName id="55F0AC6D382F541219884456C2B3E129" genus="Palps" lsidName="Palps" pageId="0" pageNumber="1" rank="genus">
<emphasis id="7D0EE19E7FD545A49193AEDAA3AD2639" bold="true" italics="true" pageId="0" pageNumber="1">Palps</emphasis>
</taxonomicName>
and
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">legs</emphasis>
<emphasis id="CEC7ADFF7698EC7BD949F1BF430DBBFE" bold="true" italics="true" pageId="0" pageNumber="1">legs</emphasis>
honey coloured, with weak annulae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
<emphasis id="D961DF7DF15B6427C3A60E5A0E7534C7" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
brown, marked as in male except bands are less distinct (Figs
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">126</figureCitation>
<figureCitation id="2A5B995B566542298BE76E823CFCBB2F" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">126</figureCitation>
,
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">128</figureCitation>
<figureCitation id="4446CBC8808E0921D6D090BBA54BDFE2" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">128</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Epigynum</emphasis>
<emphasis id="618812EF1D6D92665E708189488E9FCA" bold="true" italics="true" pageId="0" pageNumber="1">Epigynum</emphasis>
with an anterior atrium from which short copulatory ducts lead diagonally to the spermathecae (Figs
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">122</figureCitation>
<figureCitation id="66A63B0071FE3CAA4DEDE3CB729EB4BC" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">122</figureCitation>
,
<figureCitation captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">123</figureCitation>
<figureCitation id="8D74BB47CFB1A99F2481B5BC55559204" captionStart="Figures 120128" captionStartId="F12" captionText="Figures 120 - 128. Tomis manabita, sp. nov. 120, 121 Left palp of holotype 120 ventral view 121 retrolateral view 122 ventral view of epigyne of paratype 123 dorsal view of same, cleared 124 - 128 specimens from type locality 124 male 125 male 126 female 127 male holotype 128 female paratype." figureDoi="10.3897/zookeys.925.39691.figures120-128" httpUri="https://binary.pensoft.net/fig/397994" pageId="0" pageNumber="1">123</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="93500E49F6668CD39DFCD9EE2CFC9CAA" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="111C8F7316FE9C0F2A77E10F59BF8991" pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph id="8337D0E9608A590B545F6A6696C2050B" pageId="0" pageNumber="1">
15 males and 7 females from Ecuador:
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="FAA6AEF91C87BC89E467194728CED404" originalValue="Manabí">Manabi</normalizedToken>
: Machalilla National Park, Salaite, between HWY and coast 6 May 1994 W. Maddison WPM#94-032 (4 males, 2 females); Ecuador:
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="DEBE1F44A85B2E37C05374AF6BF1F8AB" originalValue="Manabí">Manabi</normalizedToken>
: Machalilla National Park, Salaite, 1 km inland along trail from HWY. 6 May 1994 W. Maddison WPM#94-033 (3 males); Ecuador:
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="C06B6ED8AE560729338F198D2D6AB75E" originalValue="Manabí">Manabi</normalizedToken>
: Machalilla National Park, trail between Agua Blanca &amp; San Sebastien 50-400 m dry forest 7 May 1994 W. Maddison WPM#94-034 (1 male); Ecuador:
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="0E6010B22A2B141B4FCF4DA01F4BA979" originalValue="Manabí">Manabi</normalizedToken>
: Crucita. 30 August 1988 W. Maddison WPM#88-040 (2 males 4 females); Ecuador: Del Oro:
<normalizedToken originalValue="Jambelí">Jambeli</normalizedToken>
<normalizedToken id="8D8D2C9C762633EE26AF7223394FE292" originalValue="Jambelí">Jambeli</normalizedToken>
13 August 1989 W. Maddison WPM#89-040 (3 males); Ecuador:
<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
<normalizedToken id="D1DCFD44CD158DE551E20232077461A5" originalValue="Manabí">Manabi</normalizedToken>
: Puerto Lopez.
<geoCoordinate degrees="1.5497" direction="north" orientation="latitude" precision="5" value="1.5497">1.5497</geoCoordinate>
<geoCoordinate id="22F246BDB567F88DC5FC27CBDCDE0391" degrees="1.5497" direction="north" orientation="latitude" precision="5" value="1.5497">1.5497</geoCoordinate>
,
<geoCoordinate degrees="80.8104" direction="west" orientation="longitude" precision="5" value="-80.8104">-80.8104</geoCoordinate>
<geoCoordinate id="D708A4C15627549762DDC9242ECAEFB0" degrees="80.8104" direction="west" orientation="longitude" precision="5" value="-80.8104">-80.8104</geoCoordinate>
; 5 m el. 1-5 August 2004 W. Maddison. WPM#04-067 (2 males 1 female).
</paragraph>
</subSubSection>