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<document id="86BC8CED9051ACD23004DC940B921376" ID-DOI="10.11646/zootaxa.5507.1.9" ID-ISSN="1175-5326" ID-ZooBank="801FE442-0B77-4A70-A321-80971F811C60" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726056266076" checkinUser="plazi" docAuthor="Jeong, In-Won &amp; Kim, Sora" docDate="2024" docId="14115A56FFF8FFA8FF0CFEC022D9FB81" docLanguage="en" docName="zootaxa.5507.1.9.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyrhabda benearena Jeong &amp; Kim 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="184" masterDocId="E828222EFFFBFFADFF9BFFDF222CFFF2" masterDocTitle="A new species of Pachyrhabda Meyrick (Lepidoptera: Stathmopodidae) from the Korean Peninsula" masterLastPageNumber="186" masterPageNumber="179" pageNumber="182" updateTime="1726056325044" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="9BC2667D08396DE7C3011EC00B8C685B">A new species of Pachyrhabda Meyrick (Lepidoptera: Stathmopodidae) from the Korean Peninsula</mods:title>
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<mods:namePart id="0C57B3465E35B3AB68070EDCDC4C49DA">Jeong, In-Won</mods:namePart>
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<mods:affiliation id="F39C9699BB9ACCB491BE6EC39F1595B4">Department of Agricultural Convergence Technology, Jeonbuk National University</mods:affiliation>
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<mods:namePart id="3162C79EF01F942C459F85E03FEE6352">Kim, Sora</mods:namePart>
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<mods:affiliation id="768AA0D169D000B4EE3668A91AC6B776">Department of Agricultural Convergence Technology, Jeonbuk National University</mods:affiliation>
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<paragraph id="9C07EB40FFF8FFAEFF0CFEC0204FFEC8" blockId="3.[151,611,287,314]" box="[151,611,287,314]" pageId="3" pageNumber="182">
<heading id="C74F5C2CFFF8FFAEFF0CFEC0204FFEC8" bold="true" box="[151,611,287,314]" fontSize="11" level="1" pageId="3" pageNumber="182" reason="1">
<emphasis id="AECC3752FFF8FFAEFF0CFEC0204FFEC8" bold="true" box="[151,611,287,314]" pageId="3" pageNumber="182">
<taxonomicName id="5BB890C3FFF8FFAEFF0CFEC023D1FECB" authority="Kim" authorityName="Jeong &amp; Kim" authorityYear="2024" box="[151,509,287,314]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="benearena" status="sp. nov.">
<emphasis id="AECC3752FFF8FFAEFF0CFEC02393FEC8" bold="true" box="[151,447,287,314]" italics="true" pageId="3" pageNumber="182">Pachyrhabda benearena</emphasis>
Kim
</taxonomicName>
<taxonomicNameLabel id="B5FF8A29FFF8FFAEFD9EFEFF204FFEC8" box="[517,611,288,314]" pageId="3" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4A2B8CBFFF8FFAEFF0CFEB720C8FE54" pageId="3" pageNumber="182" type="materials_examined">
<paragraph id="9C07EB40FFF8FFAEFF0CFEB720C8FE54" blockId="3.[151,1437,359,422]" pageId="3" pageNumber="182">
<emphasis id="AECC3752FFF8FFAEFF0CFEB7236BFE73" bold="true" box="[151,327,359,386]" pageId="3" pageNumber="182">Type material.</emphasis>
<materialsCitation id="2CD0E11DFFF8FFAEFECAFEB820C8FE54" collectingDate="2011-07-10" collectionCode="JBNU" collectorName="Sora Kim" country="South Korea" county="Ganghyeon" location="Gangwan Province" municipality="Yangyang" pageId="3" pageNumber="182" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="430355E2FFF8FFAEFECAFEB82392FE73" box="[337,446,359,385]" pageId="3" pageNumber="182" type="holotype">Holotype</typeStatus>
:
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,
<collectingCountry id="E4AFABD0FFF8FFAEFE6BFEB820EAFE70" box="[496,710,359,386]" name="South Korea" pageId="3" pageNumber="182">Republic of Korea</collectingCountry>
,
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,
<collectingMunicipality id="7C63713AFFF8FFAEFC27FEB72603FE70" box="[956,1071,360,386]" pageId="3" pageNumber="182">Yangyang</collectingMunicipality>
,
<collectingCounty id="756693CCFFF8FFAEFBA7FEB826ECFE73" box="[1084,1216,359,385]" pageId="3" pageNumber="182">Ganghyeon</collectingCounty>
,
<date id="E806CD80FFF8FFAEFB55FEB7274AFE70" box="[1230,1382,359,386]" pageId="3" pageNumber="182" value="2011-07-10">
<collectingDate id="F8423468FFF8FFAEFB55FEB7274AFE70" box="[1230,1382,359,386]" pageId="3" pageNumber="182" value="2011-07-10">10 July 2011</collectingDate>
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, leg.
<collectorName id="314D8E96FFF8FFAEFF0CFE53232BFE57" box="[151,263,395,422]" pageId="3" pageNumber="182">Sora Kim</collectorName>
, gen. slide no. 9157; [deposited in
<collectionCode id="FAA97385FFF8FFAEFD16FE5320F5FE54" box="[653,729,396,422]" pageId="3" pageNumber="182">JBNU</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4A2B8CBFFF8FFAEFF0CFE2822C4FC6F" pageId="3" pageNumber="182" type="diagnosis">
<paragraph id="9C07EB40FFF8FFAEFF0CFE28233CFDE3" blockId="3.[151,272,503,529]" box="[151,272,503,529]" pageId="3" pageNumber="182">
<emphasis id="AECC3752FFF8FFAEFF0CFE28233CFDE3" bold="true" box="[151,272,503,529]" pageId="3" pageNumber="182">Diagnosis</emphasis>
</paragraph>
<paragraph id="9C07EB40FFF8FFAEFF0CFDE02022FD1B" blockId="3.[151,1437,575,925]" pageId="3" pageNumber="182">
The new species is superficially similar to
<taxonomicName id="5BB890C3FFF8FFAEFD20FDE0269AFDAB" authority="Terada, 2016" authorityName="Terada" authorityYear="2016" box="[699,1206,575,601]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="argyrocosmos">
<emphasis id="AECC3752FFF8FFAEFD20FDE02625FDAB" box="[699,1033,575,601]" italics="true" pageId="3" pageNumber="182">Pachyrhabda argyrocosmos</emphasis>
<bibRefCitation id="F82996B1FFF8FFAEFB8CFDE0269AFDAB" author="Terada, T." box="[1047,1206,575,601]" pageId="3" pageNumber="182" refId="ref4185" refString="Terada, T. (2016) Family Stathmopodidae. In: The Insects of Japan. Vol. 7. Entomological Society of Japan, Touka Shobo Co. Ltd., Fukuoka, pp. 104 - 120." type="book" year="2016">Terada, 2016</bibRefCitation>
</taxonomicName>
but can be easily distinguished by the ground colour and markings of the forewing. The latter species has paler forewings with large two darker blotches and hind tibia with dense tufts, while in the new species
<taxonomicName id="5BB890C3FFF8FFAEFBE6FD5822E6FD37" authority="Kim" authorityName="Jeong &amp; Kim" authorityYear="2024" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="benearena" status="sp. nov.">
<emphasis id="AECC3752FFF8FFAEFBE6FD5827B1FD53" box="[1149,1437,647,673]" italics="true" pageId="3" pageNumber="182">Pachyrhabda benearena</emphasis>
Kim
</taxonomicName>
<taxonomicNameLabel id="B5FF8A29FFF8FFAEFF49FD732302FD34" box="[210,302,684,710]" pageId="3" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
many irregular dark brown spots are dispersed in a disordered manner on the forewing; hairs on the hind tibia are almost absent.
</paragraph>
<paragraph id="9C07EB40FFF8FFAEFF5CFD2C22C4FC6F" blockId="3.[151,1437,575,925]" pageId="3" pageNumber="182">
Male genitalia of the new species are close to
<taxonomicName id="5BB890C3FFF8FFAEFD61FD2B2649FCFF" authority="Terada, 2016" authorityName="Terada" authorityYear="2016" box="[762,1125,755,782]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="fuscimaculata">
<emphasis id="AECC3752FFF8FFAEFD61FD2B2127FCFF" box="[762,779,756,781]" italics="true" pageId="3" pageNumber="182">P</emphasis>
.
<emphasis id="AECC3752FFF8FFAEFC86FD2C21EDFCFF" box="[797,961,755,781]" italics="true" pageId="3" pageNumber="182">fuscimaculata</emphasis>
<bibRefCitation id="F82996B1FFF8FFAEFC50FD2C2649FCFF" author="Terada, T." box="[971,1125,755,781]" pageId="3" pageNumber="182" refId="ref4185" refString="Terada, T. (2016) Family Stathmopodidae. In: The Insects of Japan. Vol. 7. Entomological Society of Japan, Touka Shobo Co. Ltd., Fukuoka, pp. 104 - 120." type="book" year="2016">Terada, 2016</bibRefCitation>
</taxonomicName>
but differ in the shape of uncus and aedeagus. Uncus of the new species is tapering towards the apex with a swollen part at the middle and mainly has few setae at the middle, aedeagus has a sclerotized plate and a tooth-shaped cornutus at vesical. While uncus of
<taxonomicName id="5BB890C3FFF8FFAEFECFFCBF2036FC8B" authorityName="Terada" authorityYear="2016" box="[340,538,863,890]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="fuscimaculata">
<emphasis id="AECC3752FFF8FFAEFECFFCBF2349FC8B" box="[340,357,864,889]" italics="true" pageId="3" pageNumber="182">P</emphasis>
.
<emphasis id="AECC3752FFF8FFAEFEEDFC802036FC8B" box="[374,538,863,889]" italics="true" pageId="3" pageNumber="182">fuscimaculata</emphasis>
</taxonomicName>
is bell-shaped and densely setose; aedeagus has only a sclerotized plated at vesica.
</paragraph>
</subSubSection>
<subSubSection id="D4A2B8CBFFF8FFAEFF0CFC302026F89B" pageId="3" pageNumber="182" type="description">
<paragraph id="9C07EB40FFF8FFAEFF0CFC302304FBFB" blockId="3.[151,296,1007,1033]" box="[151,296,1007,1033]" pageId="3" pageNumber="182">
<emphasis id="AECC3752FFF8FFAEFF0CFC302304FBFB" bold="true" box="[151,296,1007,1033]" pageId="3" pageNumber="182">Description</emphasis>
</paragraph>
<paragraph id="9C07EB40FFF8FFAEFF0CFBE821CBFADB" blockId="3.[151,1437,1079,1898]" pageId="3" pageNumber="182">
Male (
<figureCitation id="0483F7C5FFF8FFAEFF79FBE82309FBA0" box="[226,293,1079,1106]" captionStart="FIGURE 1" captionStartId="4.[152,255,1313,1337]" captionTargetBox="[241,1346,269,1288]" captionTargetId="figure-33@4.[241,1346,179,1288]" captionTargetPageId="4" captionText="FIGURE 1. Pachyrhabda benearena sp. nov. A, adult; B, wing; C, head; D, antenna." pageId="3" pageNumber="182">Fig. 1</figureCitation>
).
<emphasis id="AECC3752FFF8FFAEFEA4FBE82350FBA3" box="[319,380,1079,1105]" italics="true" pageId="3" pageNumber="182">Head</emphasis>
. Vertex and frons yellowish ochreous, covered with white scales shaped rounded at the base and tapering toward the apex. Occiput of the same colour as vertex and covered with scales of the same shape as on vertex. Antennal flagellomeres are dark ochreous without cilia. Flagellum up to 54
<superScript id="6BCD4608FFF8FFAEFB1AFBA026A2FB7F" attach="left" box="[1153,1166,1151,1165]" fontSize="6" pageId="3" pageNumber="182">th</superScript>
flagellomere, ochreous interchanging with dark ochreous flagellomere one by one, but from 55
<superScript id="6BCD4608FFF8FFAEFC34FB7C2190FB43" attach="left" box="[943,956,1187,1201]" fontSize="6" pageId="3" pageNumber="182">th</superScript>
flagellomere, one ochreous flagellomere is followed by three dark ochreous flagellomeres and such interchangeable pattern of flagellomere colour is repeated up to the apex. Antenna ca. 4/5 the length of forewing; scape pale ochreous. Labial palpus pale ochreous, slightly upcurved with acute apex; terminal palpomere partially brownish ochreous.
</paragraph>
<paragraph id="9C07EB40FFF8FFAEFF5CFAEC21D8F9F3" blockId="3.[151,1437,1079,1898]" pageId="3" pageNumber="182">
<emphasis id="AECC3752FFF8FFAEFF5CFAEC2334FABF" box="[199,280,1331,1357]" italics="true" pageId="3" pageNumber="182">Thorax</emphasis>
. Mostly yellowish ochreous Wing span ca.
<quantity id="5B4046A5FFF8FFAEFC9AFAEC2144FABC" box="[769,872,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="182" unit="mm" value="10.5">10.5 mm</quantity>
. Forewing with irregular dark brownish spots on a pale ochreous background; pale yellowish cilia along termen. The ground colour of the hindwing is pale ochreous with darker shading towards the apex; cilia pale yellowish. Fore tibia dark ochreous. Mid tibia is mostly ochreous; a pair of brownish long and thick spurs present at the apical part of the mid tibia. Hind tibia is dorsally dark brown and pale ochreous ventrally with two pairs of ochreous long and thick spurs at the basis and at the apex of the tibia. Each tarsus is dark brown dorsally with a pair of short brownish erect scales.
</paragraph>
<paragraph id="9C07EB40FFF8FFAEFF5CF9D4203BF8B7" blockId="3.[151,1437,1079,1898]" pageId="3" pageNumber="182">
<emphasis id="AECC3752FFF8FFAEFF5CF9D42340F9D7" box="[199,364,1547,1573]" italics="true" pageId="3" pageNumber="182">Male genitalia</emphasis>
(
<figureCitation id="0483F7C5FFF8FFAEFEE7F9D423EEF9D4" box="[380,450,1547,1574]" captionStart="FIGURE 2" captionStartId="4.[152,255,2015,2039]" captionTargetBox="[263,1325,1374,1981]" captionTargetId="figure-52@4.[250,1338,1362,1991]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of Pachyrhabda benearena sp. nov. A, male genitalia; B, aedeagus. Scale bar, 0.5 mm." pageId="3" pageNumber="182">Fig. 2</figureCitation>
). Uncus straight, tapering apically, with a swollen part at the middle and some sparsely distributed setae laterally. Gnathos consists of two parts: posterior round-shaped part, and the anterior part that is broadly tongue-shaped, as long as uncus; with short dense setae at the caudal anterior part. Tegumen is as long as half of valval length. Valva broad, slightly arched with gently rounded cucullus; dorsal margin of cucullus smoothly arched; sacculus convex caudally, sclerotized at the base which is covered by short setae, gently concave near the dorsal margin, the inner surface of sacculus with numerous setae. Vinculum short, broad, and irregularly shaped. Aedeagus as long as a genital capsule, narrow tubular shaped and long sclerotized cornutus on vesica, a tooth-shaped sclerotized structure present near the base of vesica, long, narrow, lineal cornutus stretches along almost the entire length of aedeagus dorsally.
</paragraph>
<paragraph id="9C07EB40FFF8FFAEFF5CF8902026F89B" blockId="3.[151,1437,1079,1898]" box="[199,522,1871,1898]" pageId="3" pageNumber="182">
<emphasis id="AECC3752FFF8FFAEFF5CF89023AAF89B" box="[199,390,1871,1897]" italics="true" pageId="3" pageNumber="182">Female genitalia</emphasis>
. Unknown.
</paragraph>
</subSubSection>
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<paragraph id="9C07EB40FFFFFFA9FF03FAFE21D1FACB" blockId="4.[152,1021,1313,1338]" box="[152,1021,1313,1338]" pageId="4" pageNumber="183">
<emphasis id="AECC3752FFFFFFA9FF03FAFE2335FACB" bold="true" box="[152,281,1313,1337]" pageId="4" pageNumber="183">FIGURE 1.</emphasis>
<taxonomicName id="5BB890C3FFFFFFA9FE84FAFD203BFAC8" authorityName="Jeong &amp; Kim" authorityYear="2024" box="[287,535,1314,1338]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="benearena" status="sp. nov.">
<emphasis id="AECC3752FFFFFFA9FE84FAFD203BFAC8" box="[287,535,1314,1338]" italics="true" pageId="4" pageNumber="183">Pachyrhabda benearena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5FF8A29FFFFFFA9FD86FAFD2040FACB" box="[541,620,1314,1337]" pageId="4" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
A, adult; B, wing; C, head; D, antenna.
</paragraph>
</caption>
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<paragraph id="9C07EB40FFFFFFA9FF03F80026C3F805" blockId="4.[152,1263,2015,2039]" box="[152,1263,2015,2039]" pageId="4" pageNumber="183">
<emphasis id="AECC3752FFFFFFA9FF03F8002334F805" bold="true" box="[152,280,2015,2039]" pageId="4" pageNumber="183">FIGURE 2.</emphasis>
Genitalia of
<taxonomicName id="5BB890C3FFFFFFA9FE06F80020BAF805" authorityName="Jeong &amp; Kim" authorityYear="2024" box="[413,662,2015,2039]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="benearena" status="sp. nov.">
<emphasis id="AECC3752FFFFFFA9FE06F80020BAF805" box="[413,662,2015,2039]" italics="true" pageId="4" pageNumber="183">Pachyrhabda benearena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5FF8A29FFFFFFA9FD07F83F20C7F805" box="[668,747,2016,2039]" pageId="4" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
A, male genitalia; B, aedeagus. Scale bar, 0.5 mm.
</paragraph>
</caption>
<subSubSection id="D4A2B8CBFFFEFFA8FF0CFF48270FFF0B" box="[151,1315,151,250]" pageId="5" pageNumber="184" type="distribution">
<paragraph id="9C07EB40FFFEFFA8FF0CFF48231CFF43" blockId="5.[151,304,151,177]" box="[151,304,151,177]" pageId="5" pageNumber="184">
<emphasis id="AECC3752FFFEFFA8FF0CFF48231CFF43" bold="true" box="[151,304,151,177]" pageId="5" pageNumber="184">Distribution</emphasis>
</paragraph>
<paragraph id="9C07EB40FFFEFFA8FF0CFF00270FFF0B" blockId="5.[151,1315,223,250]" box="[151,1315,223,250]" pageId="5" pageNumber="184">
<materialsCitation id="2CD0E11DFFFEFFA8FF0CFF002733FF0B" box="[151,1311,223,250]" country="South Korea" location="Gangwon Province" pageId="5" pageNumber="184" specimenCount="1" stateProvince="Gangwon">
Palaearctic:
<collectingCountry id="E4AFABD0FFFEFFA8FEBAFF3F2346FF08" box="[289,362,224,250]" name="South Korea" pageId="5" pageNumber="184">Korea</collectingCountry>
: known only from the
<typeStatus id="430355E2FFFEFFA8FDFCFF3F20BBFF08" box="[615,663,224,250]" pageId="5" pageNumber="184">type</typeStatus>
locality in
<collectingRegion id="5E7C25A2FFFEFFA8FC8DFF0021DCFF0B" box="[790,1008,223,249]" country="South Korea" name="Gangwon" pageId="5" pageNumber="184">Gangwon Province</collectingRegion>
, Korean Peninsula (
<figureCitation id="0483F7C5FFFEFFA8FB4AFF002739FF0B" box="[1233,1301,223,249]" captionStart="FIGURE 3" captionStartId="5.[152,255,790,814]" captionTargetBox="[151,1432,273,767]" captionTargetId="figure-39@5.[151,1437,273,767]" captionTargetPageId="5" captionText="FIGURE 3. Collection site (in yellow circle) of Pachyrhabda benearena sp. nov. A, type locality of new species in Korean Peninsula by QGIS 3.30.0 using an EPSG:4326-WGS 84 coordinate system; B, the topography of the collecting site using QGIS 3.30.0 and data from the National Geographic Information and National Spatial Information Portal." pageId="5" pageNumber="184">Fig. 3</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="C8C7BBC8FFFEFFA8FF03FCC926ABFC85" pageId="5" pageNumber="184" startId="5.[152,255,790,814]" targetBox="[151,1432,273,767]" targetPageId="5" targetType="figure">
<paragraph id="9C07EB40FFFEFFA8FF03FCC926ABFC85" blockId="5.[151,1437,790,887]" pageId="5" pageNumber="184">
<emphasis id="AECC3752FFFEFFA8FF03FCC92336FCDD" bold="true" box="[152,282,790,815]" pageId="5" pageNumber="184">FIGURE 3.</emphasis>
Collection site (in yellow circle) of
<taxonomicName id="5BB890C3FFFEFFA8FD0EFCC821A3FCDD" authorityName="Jeong &amp; Kim" authorityYear="2024" box="[661,911,791,815]" class="Insecta" family="Stathmopodidae" genus="Pachyrhabda" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="benearena" status="sp. nov.">
<emphasis id="AECC3752FFFEFFA8FD0EFCC821A3FCDD" box="[661,911,791,815]" italics="true" pageId="5" pageNumber="184">Pachyrhabda benearena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5FF8A29FFFEFFA8FC0CFCC821C4FCDC" box="[919,1000,791,814]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
A, type locality of new species in Korean Peninsula by QGIS 3.30.0 using an EPSG:4326-WGS 84 coordinate system; B, the topography of the collecting site using QGIS 3.30.0 and data from the National Geographic Information and National Spatial Information Portal.
</paragraph>
</caption>
<subSubSection id="D4A2B8CBFFFEFFA8FF0CFC1622D9FB81" pageId="5" pageNumber="184" type="etymology">
<paragraph id="9C07EB40FFFEFFA8FF0CFC162332FC11" blockId="5.[151,286,969,995]" box="[151,286,969,995]" pageId="5" pageNumber="184">
<emphasis id="AECC3752FFFEFFA8FF0CFC162332FC11" bold="true" box="[151,286,969,995]" pageId="5" pageNumber="184">Etymology</emphasis>
</paragraph>
<paragraph id="9C07EB40FFFEFFA8FF0CFBCE22D9FB81" blockId="5.[151,1437,1041,1139]" pageId="5" pageNumber="184">The species name is a composite word derived from the Latin, “bene” and “arena”, meaning “well” and “sand”. The species-group name indicates the ground colour of the forewing with darker spots, a pattern reminiscent of sand in a desert.</paragraph>
</subSubSection>
</treatment>
</document>