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<mods:title id="DD0C7F2985E54EC6A0A71FCF8AC3C2E1">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<taxonomicName id="3EBDC4F9F248141773B5EC3155082E96" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,301,1653,1680]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F248141773B5EC3155082E96" bold="true" box="[151,301,1653,1680]" italics="true" pageId="33" pageNumber="34">Calvariopsis</emphasis>
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sp., female,
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(
<figureCitation id="6186A3FFF248141773BDECDE54DD2EB2" box="[159,248,1690,1716]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="33" pageNumber="34">Figs 1Y</figureCitation>
,
<figureCitation id="6186A3FFF24814177226ECDE55052EB2" box="[260,288,1690,1716]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="33" pageNumber="34">27</figureCitation>
A–B)
</paragraph>
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<paragraph id="F902BF7AF248141773B5ECA5570A2F26" blockId="33.[151,1437,1761,2041]" pageId="33" pageNumber="34">
<emphasis id="CBC96368F248141773B5ECA555A72EFA" bold="true" box="[151,386,1761,1788]" pageId="33" pageNumber="34">Material examined.</emphasis>
Female (SEMC): “
<collectingCountry id="81AAFFEAF2481417717FECA656F22EFA" box="[605,727,1762,1788]" name="Bolivia" pageId="33" pageNumber="34">BOLIVIA</collectingCountry>
: El
<collectingRegion id="3B797198F24814177021ECA6571D2EFA" box="[771,824,1762,1788]" country="Bolivia" name="Beni" pageId="33" pageNumber="34">Beni</collectingRegion>
\ El Povenir Stn \ NE of San Borja \
<date id="8D0399BAF248141777C1ECA651A82EFA" box="[1251,1421,1762,1788]" pageId="33" pageNumber="34" value="1988-08-08">8 August 1988</date>
”; “BIOLAT-SI/MAB \ ex., malaise trap \ Robert W. Brooks”.
</paragraph>
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<emphasis id="CBC96368F248141773E5ED6E55E92F42" bold="true" box="[199,460,1834,1860]" pageId="33" pageNumber="34">Notes on morphology.</emphasis>
Small (TL
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), body yellow, with single adsutural, black spot in posterior portion of each elytron. Gonocoxites narrow, membranous, gonostyli not preserved. Bursal sclerite (L
<quantity id="3E45129FF248141777F1ED0A511E2F6E" box="[1235,1339,1870,1896]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="33" pageNumber="34" unit="mm" value="0.32">0.32 mm</quantity>
,
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W
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mm,
<figureCitation id="6186A3FFF248141773F3ED3655112F8A" box="[209,308,1906,1932]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="33" pageNumber="34">Fig. 27B</figureCitation>
) elongate, with small denticle in posterior portion.
</paragraph>
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<subSubSection id="B1A7ECF1F248141773E5EDD255302FFF" pageId="33" pageNumber="34" type="discussion">
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<emphasis id="CBC96368F248141773E5EDD2551E2FB6" bold="true" box="[199,315,1942,1968]" pageId="33" pageNumber="34">Remarks.</emphasis>
Included in group II due to the following characters: small body, membranous gonocoxites and lack of foveate excitators (black elytral spots which are covered with black setae directed toward suture are probably excitators).
</paragraph>
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</treatment>
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<taxonomicName id="3EBDC4F9F249141673B5EE7755B22C48" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,407,1075,1102]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="wittmeri" status="sp. nov.">
<emphasis id="CBC96368F249141673B5EE7755B22C48" bold="true" box="[151,407,1075,1102]" italics="true" pageId="32" pageNumber="33">Calvariopsis wittmeri</emphasis>
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(
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Figs
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,
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)
</paragraph>
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<subSubSection id="B1A7ECF1F249141673B5EEE455C82CD8" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="F902BF7AF249141673B5EEE455C82CD8" blockId="32.[151,1437,1183,2038]" pageId="32" pageNumber="33">
<emphasis id="CBC96368F249141673B5EEE455612CBC" bold="true" box="[151,324,1183,1210]" pageId="32" pageNumber="33">Type material.</emphasis>
<materialsCitation id="49D5B527F24914167268EEE4501E2CBC" ID-GBIF-Occurrence="2239706616" box="[330,1083,1184,1210]" collectionCode="ANIC" country="Brazil" county="Wittmer" location="Rio" municipality="Janeiro" pageId="32" pageNumber="33" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F24914167268EEE455902CBC" box="[330,437,1184,1210]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
, male (
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): “
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de \ 8/49 \
<collectingMunicipality id="19662500F24914167030EEE457462CBC" box="[786,867,1184,1210]" pageId="32" pageNumber="33">Janeiro</collectingMunicipality>
\
<collectingCountry id="81AAFFEAF24914167055EEE4579F2CBC" box="[887,954,1184,1210]" name="Brazil" pageId="32" pageNumber="33">Brasil</collectingCountry>
\
<collectingCounty id="1063C7F6F249141670ECEEE450142CBC" box="[974,1073,1184,1210]" pageId="32" pageNumber="33">Wittmer</collectingCounty>
”
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.
<materialsCitation id="49D5B527F24914167766EEE455CF2CD8" ID-GBIF-Occurrence="2239706615" collectionCode="ANIC" collectorName="R. de Janeiro &amp; W. Wittmer" country="Brazil" location="Brazil" pageId="32" pageNumber="33" specimenCount="1" typeStatus="paratype">
<typeStatus id="260601D8F24914167766EEE4508C2CBC" box="[1092,1193,1184,1210]" pageId="32" pageNumber="33" type="paratype">Paratype</typeStatus>
, male (
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<collectorName id="5448DAACF24914167643EEE454CD2CD8" pageId="32" pageNumber="33">R. de Janeiro</collectorName>
\
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\
<collectorName id="5448DAACF2491416727AEE8055C52CD8" box="[344,480,1220,1246]" pageId="32" pageNumber="33">W. Wittmer</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F249141673E5EEA3514C2D68" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="F902BF7AF249141673E5EEA3514C2D68" blockId="32.[151,1437,1183,2038]" pageId="32" pageNumber="33">
<emphasis id="CBC96368F249141673E5EEA355672D04" bold="true" box="[199,322,1255,1282]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
Small (TL
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), body widest in anterior half of elytra, externally resembling
<emphasis id="CBC96368F2491416763DEEAC54F42D20" italics="true" pageId="32" pageNumber="33">C. saopau- lense</emphasis>
<emphasis id="CBC96368F249141673FAEF4855172D20" bold="true" box="[216,306,1292,1318]" pageId="32" pageNumber="33">sp. nov.</emphasis>
Identification is possible only on the basis of morphology of male genitalia: basal portion of tegmen rectangular, penis curved in lateral view (unique among known members of the genus), pala of penis rectangular, parameroids relatively wide in apical portions, tergite VIII with widely U-shaped emargination in apical plate.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F249141773E5EF3356EB2942" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="F902BF7AF249141673E5EF3356232ED0" blockId="32.[151,1437,1183,2038]" pageId="32" pageNumber="33">
<emphasis id="CBC96368F249141673E5EF33557C2D94" bold="true" box="[199,345,1399,1426]" pageId="32" pageNumber="33">Description.</emphasis>
<typeStatus id="260601D8F24914167240EF3C55E82D94" box="[354,461,1400,1426]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
, male. Body oblong oval, convex, covered with whitish, suberect setae. Head brown, pronotum, scutellar shield and elytra light brown, legs, antennae and mouthparts yellowish. Head very broad, 1.7× wider than width of interocular space, with fine punctation, punctures small, not granulate, separated by 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, not granulate punctation, punctures separated by ca. 3.0–4.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest slightly before the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation much stronger than on pronotum, punctures elongate, distance between punctures ca. 0.7–1.0× diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF249141673E5ECA457E92FF0" blockId="32.[151,1437,1183,2038]" pageId="32" pageNumber="33">
Male genitalia. Penis (L
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,
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W
<quantity id="3E45129FF2491416714CECA456BA2EFC" box="[622,671,1760,1786]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="33" unit="mm" value="0.17">0.17</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF249141671C2ECA457162EFC" box="[736,819,1759,1786]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="32" pageNumber="33">Figs 26</figureCitation>
A–B) distinctly curved in lateral view, with subparallel sides, parameroids subrectangular in basal portion, with subtriangular apices, trigonium subtriangular, with rhomboidal basal part. Tegmen (L
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,
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W
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mm,
<figureCitation id="6186A3FFF249141671FCED6C57642F44" box="[734,833,1831,1858]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="32" pageNumber="33">Fig. 26C</figureCitation>
) U-shaped, without apodemes in basal portion, apices of parameres pointed, not setose. Sternite IX (L
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,
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W
<quantity id="3E45129FF24914167067ED0857532F60" box="[837,886,1868,1894]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="32" pageNumber="33" unit="mm" value="0.21">0.21</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF24914167094ED0850392F60" box="[950,1052,1867,1894]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="32" pageNumber="33">Fig. 26D</figureCitation>
) oval, with single apodeme which is only slightly shorter than apical plate, numerous long setae present on apical and lateral margins. Tergite VIII (L
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,
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W
<quantity id="3E45129FF2491416726EEDD055582FA8" box="[332,381,1940,1966]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="32" pageNumber="33" unit="mm" value="0.28">0.28</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2491416729DEDD056072FA8" box="[447,546,1939,1966]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="32" pageNumber="33">Fig. 26E</figureCitation>
) with wide, U-shaped emargination of apical margin, apical margin not setose, apodemes as long as apical plate. Tergite IX (L
<quantity id="3E45129FF24914167197EDFC57392FD4" box="[693,796,1976,2002]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="32" pageNumber="33" unit="mm" value="0.32">0.32 mm</quantity>
,
<geoCoordinate id="9C89D9BDF2491416700AEDFC575E2FD4" box="[808,891,1976,2002]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="555" value="-0.32">
W
<quantity id="3E45129FF24914167068EDFC575E2FD4" box="[842,891,1976,2002]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="32" pageNumber="33" unit="mm" value="0.32">0.32</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2491416709CEDFC503A2FD4" box="[958,1055,1975,2002]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="32" pageNumber="33">Fig. 26F</figureCitation>
) with membranous and relatively narrow apical plate, apodemes as long as apical plate, strongly diverging.
</paragraph>
<paragraph id="F902BF7AF248141773E5EADD55BB28B5" blockId="33.[151,1436,153,467]" box="[199,414,153,179]" pageId="33" pageNumber="34">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF248141773E5EAF950B228D1" blockId="33.[151,1436,153,467]" box="[199,1175,189,215]" pageId="33" pageNumber="34">
Variability. Colouration variable,
<typeStatus id="260601D8F2481417711DEAF956BA28D1" box="[575,671,189,215]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
much darker than
<typeStatus id="260601D8F24814177053EAF957F328D1" box="[881,982,189,215]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
, brownish-black.
</paragraph>
<paragraph id="F902BF7AF248141773E5EAA556EB2942" blockId="33.[151,1436,153,467]" pageId="33" pageNumber="34">
<emphasis id="CBC96368F248141773E5EAA555DA28FD" bold="true" box="[199,511,224,251]" pageId="33" pageNumber="34">Measurements and ratios.</emphasis>
Males (n = 2) TL 2.07–2.40 (2.24) mm, PL 0.47–0.50 (0.49) mm, PW 1.00–1.10 (1.05) mm, EL 1.77–2.07 (1.92) mm, EW 1.33–1.47 (1.40) mm, TL/EW 1.55–1.64 (1.59), PW/PL 2.14–2.20 (2.17), EL/EW 1.33–1.41 (1.37), EL/PL 3.79–4.13 (3.96).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F248141773E5EB0857162961" box="[199,819,332,359]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="F902BF7AF248141773E5EB0857162961" blockId="33.[151,1436,153,467]" box="[199,819,332,359]" pageId="33" pageNumber="34">
<emphasis id="CBC96368F248141773E5EB0855442961" bold="true" box="[199,353,332,359]" pageId="33" pageNumber="34">Distribution.</emphasis>
Known only from
<collectingRegion id="3B797198F24814177115EB0956F92961" box="[567,732,333,359]" country="Brazil" name="Rio de Janeiro" pageId="33" pageNumber="34">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="81AAFFEAF248141771C5EB09570A2961" box="[743,815,333,359]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F248141773E5EB3554FF29D5" pageId="33" pageNumber="34" type="etymology">
<paragraph id="F902BF7AF248141773E5EB3554FF29D5" blockId="33.[151,1436,153,467]" pageId="33" pageNumber="34">
<emphasis id="CBC96368F248141773E5EB35556B298D" bold="true" box="[199,334,369,395]" pageId="33" pageNumber="34">Etymology.</emphasis>
Named after Walter Wittmer (
<date id="8D0399BAF2481417718AEB345709298D" box="[680,812,368,395]" pageId="33" pageNumber="34" value="1915" valueMax="1998">1915–1998</date>
), curator in the Naturhistorisches Museum Basel, a devoted coleopterist and specialist in the taxonomy of several families of beetles and collector of the
<typeStatus id="260601D8F2481417763DEBD1516A29A9" box="[1311,1359,405,431]" pageId="33" pageNumber="34">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

94
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<heading id="A24A0816F24B141473B5E82956752A81" bold="true" box="[151,592,621,648]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="CBC96368F24B141473B5E82956752A81" bold="true" box="[151,592,621,648]" pageId="34" pageNumber="35">
<taxonomicName id="3EBDC4F9F24B141473B5E82955082A8E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,301,621,648]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F24B141473B5E82955082A8E" bold="true" box="[151,301,621,648]" italics="true" pageId="34" pageNumber="35">Calvariopsis</emphasis>
</taxonomicName>
sp., female,
<collectingCountry id="81AAFFEAF24B141472EAE82956752A81" box="[456,592,621,647]" name="Costa Rica" pageId="34" pageNumber="35">Costa Rica</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473B5E8D555AB2AAD" box="[151,398,657,684]" pageId="34" pageNumber="35" type="description">
<paragraph id="F902BF7AF24B141473B5E8D555AB2AAD" blockId="34.[151,592,621,684]" box="[151,398,657,684]" pageId="34" pageNumber="35">
(
<figureCitation id="6186A3FFF24B141473BDE8D554DE2AAD" box="[159,251,657,683]" captionStart="FIGURE 2" captionStartId="4.[151,250,1057,1082]" captionTargetBox="[164,1423,199,1014]" captionTargetId="figure@4.[164,1423,199,1014]" captionTargetPageId="4" captionText="FIGURE 2. Calvariopsis gen. nov., females, dorsal view.A) C. sp., Colombia; B) C. sp., Costa Rica; C) C. sp., Ecuador; D) C. sp., Ecuador; E) C. sp., French Guiana; F) C. sp., Panama; G) C. sp., Paraguay; H) C. sp., Peru. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783686/files/figure.png" pageId="34" pageNumber="35">Figs 2B</figureCitation>
,
<figureCitation id="6186A3FFF24B14147224E8D6550C2AAA" box="[262,297,658,684]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="34" pageNumber="35">7B</figureCitation>
,
<figureCitation id="6186A3FFF24B14147217E8D555752AAD" box="[309,336,657,683]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">27</figureCitation>
E–G)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473B5E89D55F22B85" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="F902BF7AF24B141473B5E89D50D02AF5" blockId="34.[151,1437,729,971]" box="[151,1269,729,756]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473B5E89D55A42AF2" bold="true" box="[151,385,729,756]" pageId="34" pageNumber="35">Material examined.</emphasis>
Female (MNHN): “MUSEUM PARIS \
<collectingCountry id="81AAFFEAF24B1414706BE89D57CC2AF5" box="[841,1001,729,755]" name="Costa Rica" pageId="34" pageNumber="35">COSTA RICA</collectingCountry>
\ PAUL SERRE 1920”.
</paragraph>
<paragraph id="F902BF7AF24B141473E5E8BA55F22B85" blockId="34.[151,1437,729,971]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473E5E8BA55E92B1E" bold="true" box="[199,460,766,792]" pageId="34" pageNumber="35">Notes on morphology.</emphasis>
Body small (TL
<quantity id="3E45129FF24B141471AEE8B956C32B1E" box="[652,742,765,792]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="34" pageNumber="35" unit="mm" value="2.3">2.3 mm</quantity>
), dark brown, each elytron with one round concavity in apical portion (
<figureCitation id="6186A3FFF24B141473D5E965556A2B3A" box="[247,335,801,828]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="34" pageNumber="35">Fig. 7B</figureCitation>
), area situated near concavity covered with short setae. Gonocoxites moderately wide, subtriangular, with subapical gonostyli (
<figureCitation id="6186A3FFF24B141472FDE90156662B59" box="[479,579,837,863]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">Fig. 27E</figureCitation>
). Bursal sclerite (
<figureCitation id="6186A3FFF24B14147026E901574C2B59" box="[772,873,837,863]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">Fig. 27G</figureCitation>
) suboval, with truncate posterior margin and small denticle in posterior portion.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473E5E9CA55022BCD" pageId="34" pageNumber="35" type="discussion">
<paragraph id="F902BF7AF24B141473E5E9CA55022BCD" blockId="34.[151,1437,729,971]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473E5E9CA551E2BAE" bold="true" box="[199,315,910,936]" pageId="34" pageNumber="35">Remarks.</emphasis>
Included in group I due to the following characters: elytra with distinct foveate excitators, bursal sclerite oval.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="794BF991D6A9FF8567DE7201CD020963">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F24B141473B5EADD573028B2" box="[151,789,153,180]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="F902BF7AF24B141473B5EADD573028B2" blockId="34.[151,789,153,215]" box="[151,789,153,180]" pageId="34" pageNumber="35">
<heading id="A24A0816F24B141473B5EADD573028B2" bold="true" box="[151,789,153,180]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="CBC96368F24B141473B5EADD573028B2" bold="true" box="[151,789,153,180]" pageId="34" pageNumber="35">
<taxonomicName id="3EBDC4F9F24B141473B5EADD550828B2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,301,153,180]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F24B141473B5EADD550828B2" bold="true" box="[151,301,153,180]" italics="true" pageId="34" pageNumber="35">Calvariopsis</emphasis>
</taxonomicName>
sp., female,
<collectingCountry id="81AAFFEAF24B141472EAEADD566128B5" box="[456,580,153,179]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
, Gorgona Island
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473B5EAF955B428D1" box="[151,401,189,215]" pageId="34" pageNumber="35" type="description">
<paragraph id="F902BF7AF24B141473B5EAF955B428D1" blockId="34.[151,789,153,215]" box="[151,401,189,215]" pageId="34" pageNumber="35">
(
<figureCitation id="6186A3FFF24B141473BDEAF954D928D1" box="[159,252,189,215]" captionStart="FIGURE 2" captionStartId="4.[151,250,1057,1082]" captionTargetBox="[164,1423,199,1014]" captionTargetId="figure@4.[164,1423,199,1014]" captionTargetPageId="4" captionText="FIGURE 2. Calvariopsis gen. nov., females, dorsal view.A) C. sp., Colombia; B) C. sp., Costa Rica; C) C. sp., Ecuador; D) C. sp., Ecuador; E) C. sp., French Guiana; F) C. sp., Panama; G) C. sp., Paraguay; H) C. sp., Peru. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783686/files/figure.png" pageId="34" pageNumber="35">Figs 2A</figureCitation>
,
<figureCitation id="6186A3FFF24B1414722AEAF9550E28D1" box="[264,299,189,215]" captionStart="FIGURE 8" captionStartId="10.[151,250,1352,1377]" captionTargetBox="[153,1435,181,1302]" captionTargetId="figure@10.[153,1438,181,1302]" captionTargetPageId="10" captionText="FIGURE 8. Calvariopsis gen. nov., group I (A, C, E–F) and group II (B, D), females, apical portion of elytra, excitators. A) C. pittieri; B) C. brasiliense, an arrow indicates patch of modified setae; C) C. sp., Colombia, Gorgona Island; D) C. sp., Ecuador, an arrow indicates patch of modified setae; E) C. sp., Paraguay; F) C. sp., Peru." httpUri="https://zenodo.org/record/2783720/files/figure.png" pageId="34" pageNumber="35">8C</figureCitation>
,
<figureCitation id="6186A3FFF24B14147215EAF9557628D1" box="[311,339,189,215]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">27</figureCitation>
C–D)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473B5EB41559429D5" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="F902BF7AF24B141473B5EB4156CF2945" blockId="34.[151,1437,261,539]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473B5EB4155A42926" bold="true" box="[151,385,261,288]" pageId="34" pageNumber="35">Material examined.</emphasis>
Female (NHM): “
<collectingCountry id="81AAFFEAF24B14147173EB4156E22919" box="[593,711,261,287]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
: \ Gorgona Island. \
<date id="8D0399BAF24B1414708CEB4150382919" box="[942,1053,261,287]" pageId="34" pageNumber="35" value="1924-10">Oct. 1924</date>
\ St. George Expedn. \ C. L. Col- lenette”; “On \ herbage”; “Brit. Mus. \ 1924.—488.”.
</paragraph>
<paragraph id="F902BF7AF24B141473E5EB0A559429D5" blockId="34.[151,1437,261,539]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473E5EB0A55EE296E" bold="true" box="[199,459,334,360]" pageId="34" pageNumber="35">Notes on morphology.</emphasis>
Body of moderate size (TL
<quantity id="3E45129FF24B14147021EB095779296E" box="[771,860,333,360]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="34" pageNumber="35" unit="mm" value="2.8">2.8 mm</quantity>
), brown, each elytron with two round concavities in apical portion (
<figureCitation id="6186A3FFF24B14147262EB3555B3298D" box="[320,406,369,395]" captionStart="FIGURE 8" captionStartId="10.[151,250,1352,1377]" captionTargetBox="[153,1435,181,1302]" captionTargetId="figure@10.[153,1438,181,1302]" captionTargetPageId="10" captionText="FIGURE 8. Calvariopsis gen. nov., group I (A, C, E–F) and group II (B, D), females, apical portion of elytra, excitators. A) C. pittieri; B) C. brasiliense, an arrow indicates patch of modified setae; C) C. sp., Colombia, Gorgona Island; D) C. sp., Ecuador, an arrow indicates patch of modified setae; E) C. sp., Paraguay; F) C. sp., Peru." httpUri="https://zenodo.org/record/2783720/files/figure.png" pageId="34" pageNumber="35">Fig. 8C</figureCitation>
), small hooked tuft of setae present in adsutural portion of each elytron. Bursal sclerite missing in the studied specimen, only ovipositor and sternite VIII present. Gonocoxites widely triangular (
<figureCitation id="6186A3FFF24B14147605EBD151AA29A9" box="[1319,1423,405,431]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">Fig. 27C</figureCitation>
), with subapical gonostyli.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473E5EB9A561A2A1D" pageId="34" pageNumber="35" type="discussion">
<paragraph id="F902BF7AF24B141473E5EB9A561A2A1D" blockId="34.[151,1437,261,539]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473E5EB9A551E29FE" bold="true" box="[199,315,478,504]" pageId="34" pageNumber="35">Remarks.</emphasis>
Included in group I due to the following characters: elytra with distinct foveate excitators, wide gonocoxites with subapical gonostyli.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="CBC96368F24B141473B5EE5955082C3E" bold="true" box="[151,301,1053,1080]" italics="true" pageId="34" pageNumber="35">Calvariopsis</emphasis>
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sp., female,
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(
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,
<figureCitation id="6186A3FFF24B14147224EE05551D2C5D" box="[262,312,1089,1115]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">27H</figureCitation>
)
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<emphasis id="CBC96368F24B141473B5EECD55A42CA2" bold="true" box="[151,385,1161,1188]" pageId="34" pageNumber="35">Material examined.</emphasis>
Female (SEMC): “
<collectingCountry id="81AAFFEAF24B1414717BEECD56CD2CA5" box="[601,744,1161,1187]" name="Ecuador" pageId="34" pageNumber="35">ECUADOR</collectingCountry>
:
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\ Sacha Lodge, 0.5°S, \ 76.5°W,
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,
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\ 1994, Hibbs, ex: malaise”.
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<emphasis id="CBC96368F24B141473E5EE9655EA2CEA" bold="true" box="[199,463,1234,1260]" pageId="34" pageNumber="35">Notes on morphology.</emphasis>
Body of moderate size (TL
<quantity id="3E45129FF24B14147036EE95574A2CEA" box="[788,879,1233,1260]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="34" pageNumber="35" unit="mm" value="2.7">2.7 mm</quantity>
), dark brown with red humeral portions of elytra and yellowish suture in apical portion, excitators absent. Gonocoxites wide, subtriangular, with subapical gonostyli (
<figureCitation id="6186A3FFF24B14147382EF5D552D2D35" box="[160,264,1305,1331]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">Fig. 27H</figureCitation>
). Bursal sclerite elongate, with denticle in central portion.
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<emphasis id="CBC96368F24B141473E5EF7A551E2D5E" bold="true" box="[199,315,1342,1368]" pageId="34" pageNumber="35">Remarks.</emphasis>
Size and morphology of gonocoxites suggest this is a member of group I although it lacks a foveate elytral excitator. On a map (
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) marked as a female of unknown group.
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103
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sp., female,
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,
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,
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<date id="8D0399BAF24B1414721AEC51557B2E29" box="[312,350,1557,1583]" pageId="34" pageNumber="35" value="1927-01">27I</date>
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)
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<paragraph id="F902BF7AF24B141473B5EC1955F52E9D" blockId="34.[151,1437,1629,1871]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473B5EC1955A42E7E" bold="true" box="[151,385,1629,1656]" pageId="34" pageNumber="35">Material examined.</emphasis>
Female (SEMC): “
<collectingCountry id="81AAFFEAF24B1414717BEC1956CD2E71" box="[601,744,1629,1655]" name="Ecuador" pageId="34" pageNumber="35">ECUADOR</collectingCountry>
:
<collectingRegion id="3B797198F24B141471D1EC19574A2E71" box="[755,879,1629,1655]" country="Ecuador" name="Sucumbios" pageId="34" pageNumber="35">Sucumbios</collectingRegion>
\ Sacha Lodge, 0.5°S, \ 76.5°W,
<quantity id="3E45129FF24B141477C1EC1951002E71" box="[1251,1317,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="34" pageNumber="35" unit="m" value="270.0">270m</quantity>
,
<date id="8D0399BAF24B14147613EC1951B92E71" box="[1329,1436,1629,1655]" bridgedPair="-" pageId="34" pageNumber="35" value="2014-03-24">14–24-III</date>
\ 1994, Hibbs, ex: malaise”.
</paragraph>
<paragraph id="F902BF7AF24B141473E5ECE256F02F01" blockId="34.[151,1437,1629,1871]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473E5ECE255E82EC6" bold="true" box="[199,461,1702,1728]" pageId="34" pageNumber="35">Notes on morphology.</emphasis>
Body very small (TL
<quantity id="3E45129FF24B141471EAECE157142EC6" box="[712,817,1701,1728]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="34" pageNumber="35" unit="mm" value="1.85">1.85 mm</quantity>
), brownish black, covered with erect setae. Apical por- tion of each elytron with small opaque area covered by dense, stout, black setae. Gonocoxites membranous, gonostyli apical (
<figureCitation id="6186A3FFF24B14147203ECA9555B2F01" box="[289,382,1773,1799]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="34" pageNumber="35">
Fig.
<date id="8D0399BAF24B14147277ECA9555B2F01" box="[341,382,1773,1799]" pageId="34" pageNumber="35" value="1927-01">27I</date>
</figureCitation>
), bursal sclerite rod-like, long.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24B141473E5ED5655A62F49" pageId="34" pageNumber="35" type="discussion">
<paragraph id="F902BF7AF24B141473E5ED5655A62F49" blockId="34.[151,1437,1629,1871]" pageId="34" pageNumber="35">
<emphasis id="CBC96368F24B141473E5ED56551E2F2A" bold="true" box="[199,315,1810,1836]" pageId="34" pageNumber="35">Remarks.</emphasis>
Included in group II due to the following characters: small body, membranous gonocoxites and lack of foveate excitators.
</paragraph>
</subSubSection>
</treatment>
</document>

213
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<mods:title id="D22706C840B6886548072400FFA48227">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F24C141373B5EADD566C28B2" box="[151,585,153,180]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="F902BF7AF24C141373B5EADD566C28B2" blockId="37.[151,585,153,180]" box="[151,585,153,180]" pageId="37" pageNumber="38">
<heading id="A24A0816F24C141373B5EADD566C28B2" bold="true" box="[151,585,153,180]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="CBC96368F24C141373B5EADD566C28B2" bold="true" box="[151,585,153,180]" pageId="37" pageNumber="38">
Check-list of
<taxonomicName id="3EBDC4F9F24C1413721FEADD55F628B2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[317,467,153,180]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F24C1413721FEADD55F628B2" bold="true" box="[317,467,153,180]" italics="true" pageId="37" pageNumber="38">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F24C141372F9EADE566C28B2" box="[475,585,154,180]" pageId="37" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24C141373B5EAA5576F29D5" pageId="37" pageNumber="38" type="description">
<paragraph id="F902BF7AF24C141373B5EAA554CA28FA" blockId="37.[151,893,225,576]" box="[151,239,225,252]" pageId="37" pageNumber="38">Group I</paragraph>
<paragraph id="F902BF7AF24C141373B5EB4157112919" blockId="37.[151,893,225,576]" box="[151,820,261,288]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EB4155B92919" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,412,261,287]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5EB4155B92919" box="[151,412,261,287]" italics="true" pageId="37" pageNumber="38">Calvariopsis borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C14137281EB4255D82926" bold="true" box="[419,509,262,288]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C14137281EB4255D82926" box="[419,509,262,288]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5EB4157112919" box="[727,820,261,287]" name="Ecuador" pageId="37" pageNumber="38">Ecuador</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5EB6D57582945" blockId="37.[151,893,225,576]" box="[151,893,297,324]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EB6D55E32942" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,454,297,324]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5EB6D55E32942" box="[151,454,297,324]" italics="true" pageId="37" pageNumber="38">Calvariopsis fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372EFEB6E56022942" bold="true" box="[461,551,298,324]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372EFEB6E56022942" box="[461,551,298,324]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5EB6D57582945" box="[727,893,297,323]" name="French Guiana" pageId="37" pageNumber="38">French Guiana</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5EB095715296E" blockId="37.[151,893,225,576]" box="[151,816,333,360]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EB09558E2961" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,427,333,359]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="panamense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5EB09558E2961" box="[151,427,333,359]" italics="true" pageId="37" pageNumber="38">Calvariopsis panamense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C14137290EB0A5629296E" bold="true" box="[434,524,334,360]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C14137290EB0A5629296E" box="[434,524,334,360]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5EB0A5715296E" box="[727,816,334,360]" name="Panama" pageId="37" pageNumber="38">Panama</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5EB35572F298A" blockId="37.[151,893,225,576]" box="[151,778,369,396]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EB3555BA298D" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,415,369,395]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5EB3555BA298D" box="[151,415,369,395]" italics="true" pageId="37" pageNumber="38">Calvariopsis peruviana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C14137284EB3655DA298A" bold="true" box="[422,511,370,396]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C14137284EB3655DA298A" box="[422,511,370,396]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5EB36572F298A" box="[727,778,370,396]" name="Peru" pageId="37" pageNumber="38">Peru</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5EBD1570C29A9" blockId="37.[151,893,225,576]" box="[151,809,405,432]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EBD1554029A9" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,357,405,431]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="picta" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5EBD1554029A9" box="[151,357,405,431]" italics="true" pageId="37" pageNumber="38">Calvariopsis picta</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C1413724EEBD255E329B6" bold="true" box="[364,454,406,432]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C1413724EEBD255E329B6" box="[364,454,406,432]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5EBD1570C29A9" box="[727,809,405,431]" name="Bolivia" pageId="37" pageNumber="38">Bolivia</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5EBFD576F29D5" blockId="37.[151,893,225,576]" box="[151,842,441,468]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EBFD555C29D5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,377,441,467]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5EBFD555C29D5" box="[151,377,441,467]" italics="true" pageId="37" pageNumber="38">Calvariopsis pittieri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372A3EBFE55FF29D2" bold="true" box="[385,474,442,468]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372A3EBFE55FF29D2" box="[385,474,442,468]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5EBFD576F29D5" box="[727,842,441,467]" name="Venezuela" pageId="37" pageNumber="38">Venezuela</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24C141373B5EB99576F2A1D" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="F902BF7AF24C141373B5EB99573E29F1" blockId="37.[151,893,225,576]" box="[151,795,477,503]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5EB99561B29F1" authority="(Pic, 1931)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[151,574,477,503]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sculptipenne">
<emphasis id="CBC96368F24C141373B5EB99559E29F1" box="[151,443,477,503]" italics="true" pageId="37" pageNumber="38">Calvariopsis sculptipenne</emphasis>
(
<bibRefCitation id="9D2CC28BF24C141372E8EB99561329F1" author="Pic, M." box="[458,566,477,503]" pageId="37" pageNumber="38" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
)
</taxonomicName>
<collectingCountry id="81AAFFEAF24C141371F5EB99573E29F1" box="[727,795,477,503]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5E845576F2A1D" blockId="37.[151,893,225,576]" box="[151,842,513,540]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E845559A2A1D" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,447,513,539]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E845559A2A1D" box="[151,447,513,539]" italics="true" pageId="37" pageNumber="38">Calvariopsis venezuelense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372E4E84656052A1A" bold="true" box="[454,544,514,540]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372E4E84656052A1A" box="[454,544,514,540]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E845576F2A1D" box="[727,842,513,539]" name="Venezuela" pageId="37" pageNumber="38">Venezuela</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24C141373B5E86157582AAD" pageId="37" pageNumber="38" type="description">
<paragraph id="F902BF7AF24C141373B5E86157112A39" blockId="37.[151,893,225,576]" box="[151,820,549,576]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E86155EF2A39" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,458,549,575]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E86155EF2A39" box="[151,458,549,575]" italics="true" pageId="37" pageNumber="38">Calvariopsis yanayacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372F3E862560E2A46" bold="true" box="[465,555,550,576]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372F3E862560E2A46" box="[465,555,550,576]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E86157112A39" box="[727,820,549,575]" name="Ecuador" pageId="37" pageNumber="38">Ecuador</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5E82954DC2A8E" blockId="37.[151,893,621,900]" box="[151,249,621,648]" pageId="37" pageNumber="38">Group II</paragraph>
<paragraph id="F902BF7AF24C141373B5E8D557582AAD" blockId="37.[151,893,621,900]" box="[151,893,657,684]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E8D555E62AAD" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,451,657,683]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E8D555E62AAD" box="[151,451,657,683]" italics="true" pageId="37" pageNumber="38">Calvariopsis bituberculata</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372E8E8D656012AAA" bold="true" box="[458,548,658,684]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372E8E8D656012AAA" box="[458,548,658,684]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E8D557582AAD" box="[727,893,657,683]" name="French Guiana" pageId="37" pageNumber="38">French Guiana</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24C141373B5E8F1570B2AF5" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="F902BF7AF24C141373B5E8F1573E2AC9" blockId="37.[151,893,621,900]" box="[151,795,693,720]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E8F156092AD6" authority="(Pic, 1916)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[151,556,693,720]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F24C141373B5E8F1558D2AC9" box="[151,424,693,719]" italics="true" pageId="37" pageNumber="38">Calvariopsis brasiliense</emphasis>
(
<bibRefCitation id="9D2CC28BF24C14137295E8F156062AD6" author="Pic, M." box="[439,547,693,720]" pageId="37" pageNumber="38" pagination="1 - 20" refId="ref23088" refString="Pic, M. (1916) Descriptions abregees diverses. Melanges exotico-entomologiques, 20 (1), 1 - 20." type="journal article" year="1916">Pic, 1916</bibRefCitation>
)
</taxonomicName>
<collectingCountry id="81AAFFEAF24C141371F5E8F1573E2AC9" box="[727,795,693,719]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5E89D570B2AF5" blockId="37.[151,893,621,900]" box="[151,814,729,756]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E89D55862AF5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,419,729,755]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E89D55862AF5" box="[151,419,729,755]" italics="true" pageId="37" pageNumber="38">Calvariopsis guyanense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C14137288E89E56212AF2" bold="true" box="[426,516,730,756]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C14137288E89E56212AF2" box="[426,516,730,756]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E89D570B2AF5" box="[727,814,729,755]" name="Guyana" pageId="37" pageNumber="38">Guyana</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24C141373B5E8B9573E2B85" pageId="37" pageNumber="38" type="description">
<paragraph id="F902BF7AF24C141373B5E8B957582B11" blockId="37.[151,893,621,900]" box="[151,893,765,792]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E8B955A92B11" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,396,765,791]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="kawense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E8B955A92B11" box="[151,396,765,791]" italics="true" pageId="37" pageNumber="38">Calvariopsis kawense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372B1E8BA55C82B1E" bold="true" box="[403,493,766,792]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372B1E8BA55C82B1E" box="[403,493,766,792]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E8B957582B11" box="[727,893,765,791]" name="French Guiana" pageId="37" pageNumber="38">French Guiana</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5E965573E2B3D" blockId="37.[151,893,621,900]" box="[151,795,801,828]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E96555402B3D" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,357,801,827]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E96555402B3D" box="[151,357,801,827]" italics="true" pageId="37" pageNumber="38">Calvariopsis nana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C1413724EE96655E32B3A" bold="true" box="[364,454,802,828]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C1413724EE96655E32B3A" box="[364,454,802,828]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E965573E2B3D" box="[727,795,801,827]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5E901573E2B59" blockId="37.[151,893,621,900]" box="[151,795,837,864]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E90155922B59" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,439,837,863]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="saopaulense" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E90155922B59" box="[151,439,837,863]" italics="true" pageId="37" pageNumber="38">Calvariopsis saopaulense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C1413729DE902563D2B66" bold="true" box="[447,536,838,864]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C1413729DE902563D2B66" box="[447,536,838,864]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E901573E2B59" box="[727,795,837,863]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
</paragraph>
<paragraph id="F902BF7AF24C141373B5E92D573E2B85" blockId="37.[151,893,621,900]" box="[151,795,873,900]" pageId="37" pageNumber="38">
<taxonomicName id="3EBDC4F9F24C141373B5E92D55AF2B85" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,394,873,899]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="wittmeri" status="sp. nov.">
<emphasis id="CBC96368F24C141373B5E92D55AF2B85" box="[151,394,873,899]" italics="true" pageId="37" pageNumber="38">Calvariopsis wittmeri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24C141372B0E92E55CE2B82" bold="true" box="[402,491,874,900]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D0FADE13F24C141372B0E92E55CE2B82" box="[402,491,874,900]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="81AAFFEAF24C141371F5E92D573E2B85" box="[727,795,873,899]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5DBCB702C417D7FEFBF3EAAB13C84821">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<treatment id="71140E6CF24D141273B5E829556A2BF6" ID-DOI="http://doi.org/10.5281/zenodo.5921195" ID-GBIF-Taxon="157135225" ID-Zenodo-Dep="5921195" LSID="urn:lsid:plazi:treatment:71140E6CF24D141273B5E829556A2BF6" httpUri="http://treatment.plazi.org/id/71140E6CF24D141273B5E829556A2BF6" lastPageNumber="37" pageId="36" pageNumber="37">
<subSubSection id="B1A7ECF1F24D141273B5E829560A2A8E" box="[151,559,621,648]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="F902BF7AF24D141273B5E829560A2A8E" blockId="36.[151,559,621,683]" box="[151,559,621,648]" pageId="36" pageNumber="37">
<heading id="A24A0816F24D141273B5E829560A2A8E" bold="true" box="[151,559,621,648]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="CBC96368F24D141273B5E829560A2A8E" bold="true" box="[151,559,621,648]" pageId="36" pageNumber="37">
<taxonomicName id="3EBDC4F9F24D141273B5E82955082A8E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,301,621,648]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F24D141273B5E82955082A8E" bold="true" box="[151,301,621,648]" italics="true" pageId="36" pageNumber="37">Calvariopsis</emphasis>
</taxonomicName>
sp., female,
<collectingCountry id="81AAFFEAF24D141272EAE82A560A2A8E" box="[456,559,622,648]" name="Panama" pageId="36" pageNumber="37">Panama</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24D141273B5E8D5551E2AAD" box="[151,315,657,683]" pageId="36" pageNumber="37" type="description">
<paragraph id="F902BF7AF24D141273B5E8D5551E2AAD" blockId="36.[151,559,621,683]" box="[151,315,657,683]" pageId="36" pageNumber="37">
(
<figureCitation id="6186A3FFF24D141273BDE8D554D02AAD" box="[159,245,657,683]" captionStart="FIGURE 2" captionStartId="4.[151,250,1057,1082]" captionTargetBox="[164,1423,199,1014]" captionTargetId="figure@4.[164,1423,199,1014]" captionTargetPageId="4" captionText="FIGURE 2. Calvariopsis gen. nov., females, dorsal view.A) C. sp., Colombia; B) C. sp., Costa Rica; C) C. sp., Ecuador; D) C. sp., Ecuador; E) C. sp., French Guiana; F) C. sp., Panama; G) C. sp., Paraguay; H) C. sp., Peru. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783686/files/figure.png" pageId="36" pageNumber="37">Figs 2F</figureCitation>
,
<figureCitation id="6186A3FFF24D14127223E8D555172AAD" box="[257,306,657,683]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="36" pageNumber="37">27K</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24D141273B5E89D50B82B85" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="F902BF7AF24D141273B5E89D50402B11" blockId="36.[151,1437,729,1008]" pageId="36" pageNumber="37">
<emphasis id="CBC96368F24D141273B5E89D55A62AF2" bold="true" box="[151,387,729,756]" pageId="36" pageNumber="37">Material examined.</emphasis>
Female (SEMC): “
<collectingCountry id="81AAFFEAF24D14127141E89D56F92AF5" box="[611,732,729,755]" name="Panama" pageId="36" pageNumber="37">PANAMA</collectingCountry>
<collectingRegion id="3B797198F24D141271C7E89D57672AF5" box="[741,834,729,755]" country="Panama" name="Chiriqui" pageId="36" pageNumber="37">Chiriqui</collectingRegion>
Prov. \ La Fortuna, „Río Hornito \ Trail.” 08°42’N, 82°14’W \
<quantity id="3E45129FF24D14127231E8B9554F2B1E" box="[275,362,765,792]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1000.0">1000 m</quantity>
,
<date id="8D0399BAF24D14127254E8B955D12B1E" box="[374,500,765,792]" pageId="36" pageNumber="37" value="1995-06-11">11 VI 1995</date>
\ J. Ashe &amp; R. Brooks #175 \ ex: fogging fungusy log”.
</paragraph>
<paragraph id="F902BF7AF24D141273E5E96650B82B85" blockId="36.[151,1437,729,1008]" pageId="36" pageNumber="37">
<emphasis id="CBC96368F24D141273E5E96655EE2B3A" bold="true" box="[199,459,802,828]" pageId="36" pageNumber="37">Notes on morphology.</emphasis>
Body of moderate size (TL
<quantity id="3E45129FF24D14127026E96557492B3A" box="[772,876,801,828]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="36" pageNumber="37" unit="mm" value="2.85">2.85 mm</quantity>
), oblong, dark brown, apical half of elytra densely covered with erect setae, excitators absent. Gonocoxites wide, subtriangular, with subapical gonostyli (
<figureCitation id="6186A3FFF24D14127604E90151AA2B59" box="[1318,1423,837,863]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="36" pageNumber="37">Fig. 27K</figureCitation>
), bursal sclerite subtriangular, widened in posterior portion, with small subconical outgrowth.
</paragraph>
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<subSubSection id="B1A7ECF1F24D141273E5E9CA556A2BF6" pageId="36" pageNumber="37" type="discussion">
<paragraph id="F902BF7AF24D141273E5E9CA556A2BF6" blockId="36.[151,1437,729,1008]" pageId="36" pageNumber="37">
<emphasis id="CBC96368F24D141273E5E9CA551E2BAE" bold="true" box="[199,315,910,936]" pageId="36" pageNumber="37">Remarks.</emphasis>
Size and morphology of gonocoxites suggest this is a member of group I although it lacks elytral excitators. Instead elytra are covered with unusually long, erect setae. On the map (
<figureCitation id="6186A3FFF24D14127763E9F550B62BCD" box="[1089,1171,945,971]" captionStart="FIGURE 28" captionStartId="38.[151,250,1421,1446]" captionTargetBox="[153,1435,471,1378]" captionTargetId="figure@38.[151,1436,471,1378]" captionTargetPageId="38" captionText="FIGURE 28. Distribution of Calvariopsis gen. nov." httpUri="https://zenodo.org/record/2783808/files/figure.png" pageId="36" pageNumber="37">Fig. 28</figureCitation>
) marked as a female of unknown group.
</paragraph>
</subSubSection>
</treatment>
</document>

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sp., female,
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(
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,
<figureCitation id="6186A3FFF24D14127227EAF9553628D1" box="[261,275,189,215]" captionStart="FIGURE 4" captionStartId="6.[151,250,1424,1449]" captionTargetBox="[178,1410,181,1383]" captionTargetId="figure@6.[177,1411,179,1383]" captionTargetPageId="6" captionText="FIGURE 4. Calvariopsis gen. nov., mouthparts.A–C) C. yanayacuense sp. nov.; D–F) C. sp., group II, female (French Guiana). A, D) mandible; B, E) maxilla; C, F) labium." httpUri="https://zenodo.org/record/2783698/files/figure.png" pageId="36" pageNumber="37">4</figureCitation>
D–E, 27J)
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<emphasis id="CBC96368F24D141273B5EB4155A42926" bold="true" box="[151,385,261,288]" pageId="36" pageNumber="37">Material examined.</emphasis>
<materialsCitation id="49D5B527F24D141272ABEB4154EE2945" ID-GBIF-Occurrence="2239706653" collectingDate="1995-11-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" location="Francaise" municipality="Mont Grand Matoury" pageId="36" pageNumber="37" specimenCount="3" specimenCount-female="3">
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(
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): “
<collectingCountry id="81AAFFEAF24D1412715DEB4156FC2919" box="[639,729,261,287]" name="French Guiana" pageId="36" pageNumber="37">Guyane</collectingCountry>
<location id="FC62E9A1F24D141271C2EB41576E2919" LSID="urn:lsid:plazi:treatment:71140E6CF24D141273B5EADD555F2A1A:FC62E9A1F24D141271C2EB41576E2919" box="[736,843,261,287]" country="French Guiana" municipality="Mont Grand Matoury" name="Francaise" pageId="36" pageNumber="37">Francaise</location>
\
<collectingMunicipality id="19662500F24D14127041EB4250702926" box="[867,1109,261,288]" pageId="36" pageNumber="37">Mont Grand Matoury</collectingMunicipality>
\ 2.–
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<collectingDate id="9D476052F24D141277AEEB4151282926" box="[1164,1293,261,288]" pageId="36" pageNumber="37" value="1995-11-10">11.10.1995</collectingDate>
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\
<collectorName id="5448DAACF24D14127606EB4251B92919" box="[1316,1436,261,288]" pageId="36" pageNumber="37">M. Kocian</collectorName>
lgt.”
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.
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<emphasis id="CBC96368F24D141273E5EB0A55F4296E" bold="true" box="[199,465,334,360]" pageId="36" pageNumber="37">Notes on morphology.</emphasis>
Body very small (TL
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), brownish black, elytra without modifications. Gonocoxites moderately wide, subtriangular, with subapical gonostyli (
<figureCitation id="6186A3FFF24D141270E1EB355001298D" box="[963,1060,369,395]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="36" pageNumber="37">Fig. 27J</figureCitation>
). Bursal sclerite in form of elongated plate, poorly preserved in the specimens studied.
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<emphasis id="CBC96368F24D141273E5EBFE551E29D2" bold="true" box="[199,315,442,468]" pageId="36" pageNumber="37">Remarks.</emphasis>
These are females of either
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<emphasis id="CBC96368F24D141271A4EBFE571A29D5" box="[646,831,441,468]" italics="true" pageId="36" pageNumber="37">C. bituberculata</emphasis>
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<taxonomicNameLabel id="D0FADE13F24D1412706AEBFE578129D2" box="[840,932,442,468]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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or
<taxonomicName id="3EBDC4F9F24D141270ECEBFE507529D5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[974,1104,441,468]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kawense" status="sp. nov.">
<emphasis id="CBC96368F24D141270ECEBFE507529D5" box="[974,1104,441,468]" italics="true" pageId="36" pageNumber="37">C. kawense</emphasis>
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As both species are sympatric and very similar externally, it is impossible to assign females to any of the two species. Undoubtedly a member of group II.
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)
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<emphasis id="CBC96368F24D141273B5EEE955A42CCE" bold="true" box="[151,385,1197,1224]" pageId="36" pageNumber="37">Material examined.</emphasis>
Female (ZMHB): “
<collectingCountry id="81AAFFEAF24D1412717FEEE956ED2CC1" box="[605,712,1197,1223]" name="Paraguay" pageId="36" pageNumber="37">Paraguay</collectingCountry>
\ leg. Fiebrig”.
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<paragraph id="F902BF7AF24D141273E5EE9651AB2D51" blockId="36.[151,1437,1197,1439]" pageId="36" pageNumber="37">
<emphasis id="CBC96368F24D141273E5EE9655E92CEA" bold="true" box="[199,460,1234,1260]" pageId="36" pageNumber="37">Notes on morphology.</emphasis>
Body of moderate size (TL
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), oval, dark brown, covered with yellowish setae. Apical portion of each elytron with two deep concavities: smaller, situated in depressed adsutural area, with a tuft of hooked setae, and slightly larger, situated on a small tubercle. Gonocoxites wide, subtriangular, with subapical gonostyli (
<figureCitation id="6186A3FFF24D1412722DEF7955512D51" box="[271,372,1341,1367]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="36" pageNumber="37">Fig. 27L</figureCitation>
), bursal sclerite subtriangular, widened in posterior portion, with small subconical outgrowth.
</paragraph>
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<subSubSection id="B1A7ECF1F24D141273E5EF26561A2D99" pageId="36" pageNumber="37" type="discussion">
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<emphasis id="CBC96368F24D141273E5EF26551E2D7A" bold="true" box="[199,315,1378,1404]" pageId="36" pageNumber="37">Remarks.</emphasis>
Included in group I due to the following characters: elytra with distinct foveate excitators, wide gonocoxites with subapical gonostyli.
</paragraph>
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112
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<emphasis id="CBC96368F24D141273B5EFB555082E0A" bold="true" box="[151,301,1521,1548]" italics="true" pageId="36" pageNumber="37">Calvariopsis</emphasis>
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sp., female,
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<paragraph id="F902BF7AF24D141273B5EC5155552E29" blockId="36.[151,516,1521,1583]" box="[151,368,1557,1583]" pageId="36" pageNumber="37">
(
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,
<figureCitation id="6186A3FFF24D1412722AEC5155032E29" box="[264,294,1557,1583]" captionStart="FIGURE 8" captionStartId="10.[151,250,1352,1377]" captionTargetBox="[153,1435,181,1302]" captionTargetId="figure@10.[153,1438,181,1302]" captionTargetPageId="10" captionText="FIGURE 8. Calvariopsis gen. nov., group I (A, C, E–F) and group II (B, D), females, apical portion of elytra, excitators. A) C. pittieri; B) C. brasiliense, an arrow indicates patch of modified setae; C) C. sp., Colombia, Gorgona Island; D) C. sp., Ecuador, an arrow indicates patch of modified setae; E) C. sp., Paraguay; F) C. sp., Peru." httpUri="https://zenodo.org/record/2783720/files/figure.png" pageId="36" pageNumber="37">8F</figureCitation>
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<figureCitation id="6186A3FFF24D14127213EC51554D2E29" box="[305,360,1557,1583]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="36" pageNumber="37">27M</figureCitation>
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<emphasis id="CBC96368F24D141273B5EC1955A52E7E" bold="true" box="[151,384,1629,1656]" pageId="36" pageNumber="37">Material examined.</emphasis>
<materialsCitation id="49D5B527F24D141272A4EC1957DE2E9D" ID-GBIF-Occurrence="2239706623" collectingDate="1993-07-26" collectionCode="SEMC" country="Peru" elevation="240" location="Teniente Lopez" pageId="36" pageNumber="37" specimenCount="2" specimenCount-female="2" stateProvince="Loreto">
<specimenCount id="EFBB74F3F24D141272A4EC1955D72E71" box="[390,498,1629,1655]" pageId="36" pageNumber="37" type="female">2 females</specimenCount>
(
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: Dept.
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\
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N
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\ 2°35.66’S, 76°06.92’W \
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\ Richard Leschen #211 \ ex:
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.
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<emphasis id="CBC96368F24D141273E5ECE255EC2EC6" bold="true" box="[199,457,1702,1728]" pageId="36" pageNumber="37">Notes on morphology.</emphasis>
Body of moderate size (TL
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), oblong, dark brown, one specimen with yel- lowish apical portion of elytral suture, apical half of elytra densely covered with erect setae, each elytron with two shallow concavities in apical portion. Gonocoxites wide, subtriangular, with subapical gonostyli (
<figureCitation id="6186A3FFF24D141277FAECA951612F01" box="[1240,1348,1773,1799]" captionStart="FIGURE 27" captionStartId="35.[151,250,1929,1954]" captionTargetBox="[153,1435,195,1889]" captionTargetId="figure@35.[151,1436,195,1889]" captionTargetPageId="35" captionText="FIGURE 27. Calvariopsis gen. nov., females. A–B) C. sp., Bolivia; C–D) C. sp., Colombia; E–G) C. sp., Costa Rica; H) C. sp., Ecuador; I) C. sp., Ecuador; J) C. sp., French Guiana; K) C. sp., Panama; L) C. sp., Paraguay; M) C. sp., Peru. A, H, I–M) genital tract; B, G) bursal sclerite; C), E) ovipositor; D, F) sternite VIII." httpUri="https://zenodo.org/record/2783802/files/figure.png" pageId="36" pageNumber="37">Fig. 27M</figureCitation>
), bursal sclerite elongate, anterior portion oblong oval, small subconical outgrowth present in central portion, posterior portion slightly widened, subtriangular.
</paragraph>
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<emphasis id="CBC96368F24D141273E5ED1E551E2F72" bold="true" box="[199,315,1882,1908]" pageId="36" pageNumber="37">Remarks.</emphasis>
Included in group I due to the following characters: elytra with distinct foveate excitators, wide gonocoxites with subapical gonostyli.
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427
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<mods:title id="0E5E8BBAD50C494D23E06F0D87D8D43E">Zootaxa</mods:title>
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<mods:part id="00BC0B6EA88265C3746DAC5363060356">
<mods:date id="96B540EA769903606500FDE1553EF399">2019</mods:date>
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<mods:number id="0A8FBE2BBDD76A3D70F1E800544BDB82">2019-05-13</mods:number>
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<mods:number id="EF4C294A690B3369F3D4607E2778337E">4604</mods:number>
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<mods:identifier id="28458BDE030BE83AC57F3FFFC412948E" type="DOI">10.11646/zootaxa.4604.1.1</mods:identifier>
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<treatment id="71140E6CF24F141173B5EFBD51B92AFF" ID-GBIF-Taxon="157135221" LSID="urn:lsid:plazi:treatment:71140E6CF24F141173B5EFBD51B92AFF" httpUri="http://treatment.plazi.org/id/71140E6CF24F141173B5EFBD51B92AFF" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection id="B1A7ECF1F24F141073B5EFBD51392E30" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="F902BF7AF24F141073B5EFBD57032E15" blockId="38.[151,1308,1528,1591]" box="[151,806,1528,1555]" pageId="38" pageNumber="39">
<heading id="A24A0816F24F141073B5EFBD57032E15" bold="true" box="[151,806,1528,1555]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="CBC96368F24F141073B5EFBD57032E15" bold="true" box="[151,806,1528,1555]" pageId="38" pageNumber="39">
Key to identification for males of
<taxonomicName id="3EBDC4F9F24F14107104EFBC56902E15" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[550,693,1528,1555]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F24F14107104EFBC56902E15" bold="true" box="[550,693,1528,1555]" italics="true" pageId="38" pageNumber="39">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F24F1410719FEFBD57032E15" box="[701,806,1529,1555]" pageId="38" pageNumber="39" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F902BF7AF24F141073B5EC5851392E30" blockId="38.[151,1308,1528,1591]" box="[151,1308,1564,1591]" pageId="38" pageNumber="39">
Please note that
<emphasis id="CBC96368F24F1410726CEC5855F12E30" box="[334,468,1564,1590]" italics="true" pageId="38" pageNumber="39">Calvaropsis</emphasis>
<taxonomicName id="3EBDC4F9F24F141072F9EC58568D2E31" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[475,680,1564,1591]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sculptipenne">
<emphasis id="CBC96368F24F141072F9EC58564C2E30" box="[475,617,1564,1590]" italics="true" pageId="38" pageNumber="39">sculptipenne</emphasis>
(Pic)
</taxonomicName>
, with only the female known, is not included in the key.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24F141173B5EC2551B92AFF" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="key">
<keyStep id="4249A7DFF24F141073B5EC2551B92E92" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5EC2551B92E71" blockId="38.[151,1437,1633,2037]" box="[151,1436,1633,1655]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EC2551B92E71" box="[151,1436,1633,1655]" pageId="38" pageNumber="39">
1. Sternite IX with paired apodemes (
<figureCitation id="6186A3FFF24F14107110EC2556AD2E71" box="[562,648,1633,1655]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="38" pageNumber="39">Fig. 11D</figureCitation>
)........................................................ 2 (group I)
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5EC3B51B92E92" blockId="38.[151,1437,1633,2037]" box="[151,1436,1662,1685]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EC3B51B92E92" box="[151,1436,1662,1685]" pageId="38" pageNumber="39">
-. Sternite IX with a single apodeme (
<figureCitation id="6186A3FFF24F14107114EC3A56A92E93" box="[566,652,1662,1685]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="38" pageNumber="39">Fig. 21C</figureCitation>
)....................................................... 9 (group II)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24F141073B5ECD851B92EEA" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5ECD851B92EB7" blockId="38.[151,1437,1633,2037]" box="[151,1436,1692,1714]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ECD851B92EB7" box="[151,1436,1692,1714]" pageId="38" pageNumber="39">
2. Basal half of penis distinctly wider than apical half (
<figureCitation id="6186A3FFF24F141071FCECD857132EB4" box="[734,822,1692,1714]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="38" pageNumber="39">Fig. 12A</figureCitation>
)................................
<taxonomicName id="3EBDC4F9F24F141077E0ECD8516F2EB7" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1218,1354,1692,1713]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="panamense" status="sp. nov.">
<emphasis id="CBC96368F24F141077E0ECD8516F2EB7" box="[1218,1354,1692,1713]" italics="true" pageId="38" pageNumber="39">C. panamense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672ECD851B92EB7" bold="true" box="[1360,1436,1692,1713]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672ECD851B92EB7" box="[1360,1436,1692,1713]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5ECFE51B92EEA" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ECFE51B92EEA" pageId="38" pageNumber="39">
-. Basal portion of penis as wide as apical half (
<figureCitation id="6186A3FFF24F141071BFECFD56D02EC9" box="[669,757,1721,1743]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="38" pageNumber="39">Fig. 14A</figureCitation>
) or less than basal 1/3 of penis is slightly wider than apical half (Figs 13A, 15A)...........................................................................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24F141073B5ECB051B92F42" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5ECB051B92F0F" blockId="38.[151,1437,1633,2037]" box="[151,1436,1780,1802]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ECB051B92F0F" box="[151,1436,1780,1802]" pageId="38" pageNumber="39">
3. Penis moderately long, 6× longer than wide (
<figureCitation id="6186A3FFF24F141071B0ECB056CB2F0F" box="[658,750,1780,1802]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="38" pageNumber="39">Figs 14A</figureCitation>
,
<figureCitation id="6186A3FFF24F141071DAECB057012F0C" box="[760,804,1780,1802]" captionStart="FIGURE 15" captionStartId="20.[151,250,1137,1162]" captionTargetBox="[186,1401,251,1088]" captionTargetId="figure@20.[186,1403,272,1088]" captionTargetPageId="20" captionText="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783750/files/figure.png" pageId="38" pageNumber="39">15A</figureCitation>
), with subparallel or subtly sinuate lateral margins............4
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5ED5651B92F42" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ED5651B92F42" pageId="38" pageNumber="39">
-. Penis very long and narrow, more than 8–10× longer than wide (
<figureCitation id="6186A3FFF24F14107068ED5557802F20" box="[842,933,1809,1831]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="38" pageNumber="39">Figs 11A</figureCitation>
,
<figureCitation id="6186A3FFF24F1410708DED5557FE2F21" box="[943,987,1809,1831]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="38" pageNumber="39">13A</figureCitation>
), widened in basal portion and narrowed in the middle...............................................................................................5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24F141073B5ED0851B92F9A" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5ED0851B92F78" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ED0851B92F78" pageId="38" pageNumber="39">
4. Penis with sinuate lateral margins, trigonium narrow, with subparallel sides (
<figureCitation id="6186A3FFF24F141070E0ED08503F2F64" box="[962,1050,1868,1890]" captionStart="FIGURE 15" captionStartId="20.[151,250,1137,1162]" captionTargetBox="[186,1401,251,1088]" captionTargetId="figure@20.[186,1403,272,1088]" captionTargetPageId="20" captionText="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783750/files/figure.png" pageId="38" pageNumber="39">Fig. 15A</figureCitation>
), known only from
<collectingCountry id="81AAFFEAF24F141077FAED08511E2F64" box="[1240,1339,1868,1890]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
.............................................................................................
<taxonomicName id="3EBDC4F9F24F141077CEED2D516C2F79" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1260,1353,1897,1919]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="CBC96368F24F141077CEED2D516C2F79" box="[1260,1353,1897,1919]" italics="true" pageId="38" pageNumber="39">C. pittieri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672ED2D51B92F78" bold="true" box="[1360,1436,1897,1918]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672ED2D51B92F78" box="[1360,1436,1897,1918]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5EDC351B92F9A" blockId="38.[151,1437,1633,2037]" box="[151,1436,1926,1949]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EDC351B92F9A" box="[151,1436,1926,1949]" pageId="38" pageNumber="39">
-. Penis with subparallel sides, trigonium wide, subtriangular (
<figureCitation id="6186A3FFF24F14107001EDC257592F9B" box="[803,892,1926,1949]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="38" pageNumber="39">Fig. 14A</figureCitation>
), known only from
<collectingCountry id="81AAFFEAF24F14107718EDC250A42F9A" box="[1082,1153,1926,1948]" name="Bolivia" pageId="38" pageNumber="39">Bolivia</collectingCountry>
.........
<taxonomicName id="3EBDC4F9F24F141077DCEDC3516F2F9A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1278,1354,1927,1948]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="picta" status="sp. nov.">
<emphasis id="CBC96368F24F141077DCEDC3516F2F9A" box="[1278,1354,1927,1948]" italics="true" pageId="38" pageNumber="39">C. picta</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672EDC351B92F9A" bold="true" box="[1360,1436,1927,1948]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672EDC351B92F9A" box="[1360,1436,1927,1948]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="F902BF7AF24F141073B5EDE051B92FD0" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EDE051B92FD0" pageId="38" pageNumber="39">
<keyStep id="4249A7DFF24F141073B5EDE051B92FD0" pageId="38" pageNumber="39">
5. TL
<quantity id="3E45129FF24F14107225EDE055702FBF" box="[263,341,1956,1977]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="38" pageNumber="39" unit="mm" value="2.3">2.3 mm</quantity>
, median process of tergite VIII not flanked with deep emarginations (Fig.
<figureCitation id="6186A3FFF24F14107719EDE050B42FBC" box="[1083,1169,1956,1978]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="38" pageNumber="39">Fig. 11E</figureCitation>
), known only from
<collectingCountry id="81AAFFEAF24F14107678EDE055022FD1" name="French Guiana" pageId="38" pageNumber="39">French Guiana</collectingCountry>
...........................................................................
<taxonomicName id="3EBDC4F9F24F14107788ED85516C2FD0" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1194,1353,1985,2006]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F24F14107788ED85516C2FD0" box="[1194,1353,1985,2006]" italics="true" pageId="38" pageNumber="39">C. fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672ED8551B92FD0" bold="true" box="[1360,1436,1985,2006]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672ED8551B92FD0" box="[1360,1436,1985,2006]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5ED9B51B92FF2" blockId="38.[151,1437,1633,2037]" box="[151,1436,2014,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ED9B51B92FF2" box="[151,1436,2014,2037]" pageId="38" pageNumber="39">
-. TL
<quantity id="3E45129FF24F14107226ED9A555F2FF2" box="[260,378,2014,2036]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.6" metricValueMin="2.5" pageId="38" pageNumber="39" unit="mm" value="3.05" valueMax="3.6" valueMin="2.5">2.5–3.6 mm</quantity>
, median process of tergite VIII flanked with deep emarginations (
<figureCitation id="6186A3FFF24F141070CEED9A501C2FF2" box="[1004,1081,2014,2036]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="38" pageNumber="39">Figs 9E</figureCitation>
,
<figureCitation id="6186A3FFF24F14107760ED9B504E2FF2" box="[1090,1131,2015,2036]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="38" pageNumber="39">13E</figureCitation>
,
<figureCitation id="6186A3FFF24F14107756ED9B50BA2FF2" box="[1140,1183,2015,2036]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="38" pageNumber="39">17C</figureCitation>
, 18D), known from Andes
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173C0EADE51B928CA" blockId="39.[151,1437,153,762]" pageId="39" pageNumber="40">
(
<collectingCountry id="81AAFFEAF24E141173CBEADE553C28A9" box="[233,281,154,175]" name="Peru" pageId="39" pageNumber="40">Peru</collectingCountry>
,
<collectingCountry id="81AAFFEAF24E14117200EADD555128A9" box="[290,372,153,175]" name="Ecuador" pageId="39" pageNumber="40">Ecuador</collectingCountry>
,
<collectingCountry id="81AAFFEAF24E1411725EEADD55C528A9" box="[380,480,153,175]" name="Venezuela" pageId="39" pageNumber="40">Venezuela</collectingCountry>
).............................................................................. 6
<keyLead id="424C1C4FF24E141173B5EAF351B928CA" box="[151,1436,183,205]" pageId="39" pageNumber="40">
<keyStep id="4249A7DFF24E141173B5EAF351B928CA" box="[151,1436,183,205]" pageId="39" pageNumber="40">
6. Median process of tergite VIII short, flanked by deep, semicircular emarginations (
<figureCitation id="6186A3FFF24E141170DFEAF3507428CB" box="[1021,1105,183,205]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="39" pageNumber="40">Fig. 13E</figureCitation>
)........
<taxonomicName id="3EBDC4F9F24E141177EEEAF3516C28CA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1228,1353,183,204]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="CBC96368F24E141177EEEAF3516C28CA" box="[1228,1353,183,204]" italics="true" pageId="39" pageNumber="40">C. peruviana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EAF351B928CA" bold="true" box="[1360,1436,183,204]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EAF351B928CA" box="[1360,1436,183,204]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EA9151B928EC" blockId="39.[151,1437,153,762]" box="[151,1436,212,234]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EA9151B928EC" box="[151,1436,212,234]" pageId="39" pageNumber="40">
-. Median process of tergite VIII longer, flanked by shallow and wide emarginations (
<figureCitation id="6186A3FFF24E14117723EA90506A28EF" box="[1025,1103,212,234]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="39" pageNumber="40">Figs 9E</figureCitation>
,
<figureCitation id="6186A3FFF24E1411777BEA9050A628EF" box="[1113,1155,212,233]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="39" pageNumber="40">17C</figureCitation>
, 18D)................. 7
</keyLead>
</paragraph>
<keyStep id="4249A7DFF24E141173B5EAB651B92922" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EAB651B92901" blockId="39.[151,1437,153,762]" box="[151,1436,241,264]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EAB651B92901" box="[151,1436,241,264]" pageId="39" pageNumber="40">
7. Median process of tergite VIII wide, short, M-shaped, emarginations without denticles (Fig. 18D).
<taxonomicName id="3EBDC4F9F24E14117785EAB6516F2901" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1191,1354,242,263]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
<emphasis id="CBC96368F24E14117785EAB6516F2901" box="[1191,1354,242,263]" italics="true" pageId="39" pageNumber="40">C. yanayacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EAB651B92901" bold="true" box="[1360,1436,242,263]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EAB651B92901" box="[1360,1436,242,263]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EB4B51B92922" blockId="39.[151,1437,153,762]" box="[151,1436,271,293]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB4B51B92922" box="[151,1436,271,293]" pageId="39" pageNumber="40">-. Median process of tergite VIII longer, emarginations covered with denticles........................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EB6851B92959" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EB6851B92947" blockId="39.[151,1437,153,762]" box="[151,1436,300,322]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB6851B92947" box="[151,1436,300,322]" pageId="39" pageNumber="40">
8. Body oblong oval, with rounded sides (
<figureCitation id="6186A3FFF24E1411717EEB68568D2944" box="[604,680,300,322]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="39" pageNumber="40">Fig. 1A</figureCitation>
), TL/EW 1.6, known only from
<collectingCountry id="81AAFFEAF24E141170C2EB6850152944" box="[992,1072,300,322]" name="Ecuador" pageId="39" pageNumber="40">Ecuador</collectingCountry>
............
<taxonomicName id="3EBDC4F9F24E141177ECEB68516C2947" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1230,1353,300,321]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F24E141177ECEB68516C2947" box="[1230,1353,300,321]" italics="true" pageId="39" pageNumber="40">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB6851B92947" bold="true" box="[1360,1436,300,321]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB6851B92947" box="[1360,1436,300,321]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EB0E51B92959" blockId="39.[151,1437,153,762]" box="[151,1436,329,352]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB0E51B92959" box="[151,1436,329,352]" pageId="39" pageNumber="40">
-. Body elongate, with subparallel sides (
<figureCitation id="6186A3FFF24E14117174EB0D56BA2966" box="[598,671,329,352]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="39" pageNumber="40">Fig. 1L</figureCitation>
), TL/EW 1.8, known only from
<collectingCountry id="81AAFFEAF24E141170F4EB0D501C2959" box="[982,1081,329,351]" name="Venezuela" pageId="39" pageNumber="40">Venezuela</collectingCountry>
.........
<taxonomicName id="3EBDC4F9F24E14117792EB0E516C2958" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1200,1353,329,351]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F24E14117792EB0E516C2958" box="[1200,1353,329,351]" italics="true" pageId="39" pageNumber="40">C. venezuelense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB0E51B92959" bold="true" box="[1360,1436,330,351]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB0E51B92959" box="[1360,1436,330,351]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EB2351B9299F" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EB2351B9297A" blockId="39.[151,1437,153,762]" box="[151,1436,359,381]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB2351B9297A" box="[151,1436,359,381]" pageId="39" pageNumber="40">
9. Last abdominal ventrite with paired tubercles (
<figureCitation id="6186A3FFF24E14117180EB2356C9297A" box="[674,748,359,381]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="39" pageNumber="40">Figs 3F</figureCitation>
,
<figureCitation id="6186A3FFF24E141171D4EB23573B297B" box="[758,798,359,381]" captionStart="FIGURE 20" captionStartId="26.[151,250,1241,1266]" captionTargetBox="[153,1435,361,1198]" captionTargetId="figure@26.[151,1436,361,1198]" captionTargetPageId="26" captionText="FIGURE 20. Calvariopsis bituberculata sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX; E) apex of abdomen. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783774/files/figure.png" pageId="39" pageNumber="40">20E</figureCitation>
)................................
<taxonomicName id="3EBDC4F9F24E1411778FEB23516C297A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1197,1353,359,380]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
<emphasis id="CBC96368F24E1411778FEB23516C297A" box="[1197,1353,359,380]" italics="true" pageId="39" pageNumber="40">C. bituberculata</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB2351B9297A" bold="true" box="[1360,1436,359,380]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB2351B9297A" box="[1360,1436,359,380]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EBC151B9299F" blockId="39.[151,1437,153,762]" box="[151,1436,388,410]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBC151B9299F" box="[151,1436,388,410]" pageId="39" pageNumber="40">-. Last abdominal ventrite without tubercles................................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EBE651B829D2" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EBE651B929B1" blockId="39.[151,1437,153,762]" box="[151,1436,417,439]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBE651B929B1" box="[151,1436,417,439]" pageId="39" pageNumber="40">
10. Tergite VIII without lateral spines, tegmen without basal apodemes (
<figureCitation id="6186A3FFF24E14117051EBE557EB29B1" box="[883,974,417,439]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="39" pageNumber="40">Figs 25B</figureCitation>
,
<figureCitation id="6186A3FFF24E141170FAEBE5502429B1" box="[984,1025,417,439]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="39" pageNumber="40">26C</figureCitation>
)................................11
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EBFB51B829D2" blockId="39.[151,1437,153,762]" box="[151,1437,447,469]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBFB51B829D2" box="[151,1437,447,469]" pageId="39" pageNumber="40">
-. Tergite VIII with lateral spines, tegmen with paired basal apodemes (
<figureCitation id="6186A3FFF24E1411705AEBFB57F629D2" box="[888,979,447,469]" captionStart="FIGURE 20" captionStartId="26.[151,250,1241,1266]" captionTargetBox="[153,1435,361,1198]" captionTargetId="figure@26.[151,1436,361,1198]" captionTargetPageId="26" captionText="FIGURE 20. Calvariopsis bituberculata sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX; E) apex of abdomen. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783774/files/figure.png" pageId="39" pageNumber="40">Figs 20B</figureCitation>
,
<figureCitation id="6186A3FFF24E141170FFEBFB502229D2" box="[989,1031,447,468]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">21B</figureCitation>
,
<figureCitation id="6186A3FFF24E14117733EBFB501E29D2" box="[1041,1083,447,468]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">22B</figureCitation>
,
<figureCitation id="6186A3FFF24E14117767EBFB504B29D3" box="[1093,1134,447,469]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="39" pageNumber="40">23B</figureCitation>
).......................12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EB9851B92A09" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EB9851B929F7" blockId="39.[151,1437,153,762]" box="[151,1436,476,498]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB9851B929F7" box="[151,1436,476,498]" pageId="39" pageNumber="40">
11. Tegmen V-shaped (
<figureCitation id="6186A3FFF24E141172B9EB9855D629F4" box="[411,499,476,498]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="39" pageNumber="40">Fig. 25B</figureCitation>
), penis not curved in lateral view...............................
<taxonomicName id="3EBDC4F9F24E14117795EB98516F29F7" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1207,1354,476,497]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="saopaulense" status="sp. nov.">
<emphasis id="CBC96368F24E14117795EB98516F29F7" box="[1207,1354,476,497]" italics="true" pageId="39" pageNumber="40">C. saopaulense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB9851B929F7" bold="true" box="[1360,1436,476,497]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB9851B929F7" box="[1360,1436,476,497]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EBBE51B92A09" blockId="39.[151,1437,153,762]" box="[151,1436,505,527]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBBE51B92A09" box="[151,1436,505,527]" pageId="39" pageNumber="40">
-. Tegmen subrectangular (
<figureCitation id="6186A3FFF24E141172EDEBBD56022A09" box="[463,551,505,527]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="39" pageNumber="40">Fig. 26C</figureCitation>
), penis curved in lateral view (
<figureCitation id="6186A3FFF24E1411706DEBBD57832A09" box="[847,934,505,527]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="39" pageNumber="40">Fig. 26A</figureCitation>
).........................
<taxonomicName id="3EBDC4F9F24E141177FFEBBE516F2A09" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1245,1354,506,527]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="wittmeri" status="sp. nov.">
<emphasis id="CBC96368F24E141177FFEBBE516F2A09" box="[1245,1354,506,527]" italics="true" pageId="39" pageNumber="40">C. wittmeri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EBBE51B92A09" bold="true" box="[1360,1436,506,527]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EBBE51B92A09" box="[1360,1436,506,527]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5E85351B92A4F" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5E85351B92A2A" blockId="39.[151,1437,153,762]" box="[151,1436,535,557]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E85351B92A2A" box="[151,1436,535,557]" pageId="39" pageNumber="40">
12. Tergite VIII with T-shaped emargination in central portion (
<figureCitation id="6186A3FFF24E1411703CE85357502A2B" box="[798,885,535,557]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="39" pageNumber="40">Fig. 23D</figureCitation>
).............................
<taxonomicName id="3EBDC4F9F24E141177FEE853516C2A2A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1244,1353,535,556]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kawense" status="sp. nov.">
<emphasis id="CBC96368F24E141177FEE853516C2A2A" box="[1244,1353,535,556]" italics="true" pageId="39" pageNumber="40">C. kawense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672E85351B92A2A" bold="true" box="[1360,1436,535,556]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672E85351B92A2A" box="[1360,1436,535,556]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5E87151B92A4F" blockId="39.[151,1437,153,762]" box="[151,1436,564,586]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E87151B92A4F" box="[151,1436,564,586]" pageId="39" pageNumber="40">
-. Tergite VIII with V-shaped emargination in central portion (
<figureCitation id="6186A3FFF24E14117002E87057592A4F" box="[800,892,564,586]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">Figs 21D</figureCitation>
,
<figureCitation id="6186A3FFF24E141170A4E87057952A4F" box="[902,944,564,585]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">22C</figureCitation>
,
<figureCitation id="6186A3FFF24E14117098E87057C12A4C" box="[954,996,564,586]" captionStart="FIGURE 24" captionStartId="30.[151,250,1766,1791]" captionTargetBox="[153,1435,971,1727]" captionTargetId="figure@30.[151,1436,971,1727]" captionTargetPageId="30" captionText="FIGURE 24. Calvariopsis nana sp. nov., male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783786/files/figure.png" pageId="39" pageNumber="40">24B</figureCitation>
)...................................13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5E81651B92AB9" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5E81651B92A83" blockId="39.[151,1437,153,762]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E81651B92A83" pageId="39" pageNumber="40">
13. Trigonium of penis shorter, reaching 1/4 of length of penis, penis with subparallel sides (
<figureCitation id="6186A3FFF24E1411776EE81550822A6E" box="[1100,1191,593,616]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">Fig. 21A</figureCitation>
), emargination of central portion of tergite VIII deep (
<figureCitation id="6186A3FFF24E141172D1E82B566F2A83" box="[499,586,623,645]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">Fig. 21D</figureCitation>
)......................................................
<taxonomicName id="3EBDC4F9F24E141177C0E82B51B92A83" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[1250,1436,623,645]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F24E141177C0E82B51422A82" box="[1250,1383,623,644]" italics="true" pageId="39" pageNumber="40">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5E8C951B92AB9" blockId="39.[151,1437,153,762]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E8C951B92AB9" pageId="39" pageNumber="40">
-. Trigonium of penis relatively long, reaching 1/3 of length of penis, penis with curved sides (
<figureCitation id="6186A3FFF24E1411777DE8C8509E2AA7" box="[1119,1211,652,674]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">Figs 22A</figureCitation>
,
<figureCitation id="6186A3FFF24E141177E4E8C850D72AA4" box="[1222,1266,652,674]" captionStart="FIGURE 24" captionStartId="30.[151,250,1766,1791]" captionTargetBox="[153,1435,971,1727]" captionTargetId="figure@30.[151,1436,971,1727]" captionTargetPageId="30" captionText="FIGURE 24. Calvariopsis nana sp. nov., male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783786/files/figure.png" pageId="39" pageNumber="40">24A</figureCitation>
), emargination of central portion of tergite VIII shallow (
<figureCitation id="6186A3FFF24E1411717AE8ED56972AB9" box="[600,690,681,703]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">Figs 22C</figureCitation>
,
<figureCitation id="6186A3FFF24E1411719EE8EE56C32AC6" box="[700,742,682,704]" captionStart="FIGURE 24" captionStartId="30.[151,250,1766,1791]" captionTargetBox="[153,1435,971,1727]" captionTargetId="figure@30.[151,1436,971,1727]" captionTargetPageId="30" captionText="FIGURE 24. Calvariopsis nana sp. nov., male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783786/files/figure.png" pageId="39" pageNumber="40">24B</figureCitation>
)....................................................... 14
</keyLead>
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<keyStep id="4249A7DFF24E141173B5E88351B92AFF" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5E88351B92ADA" blockId="39.[151,1437,153,762]" box="[151,1436,711,733]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E88351B92ADA" box="[151,1436,711,733]" pageId="39" pageNumber="40">
14. TL
<quantity id="3E45129FF24E14117226E88355752ADA" box="[260,336,711,732]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="39" pageNumber="40" unit="mm" value="1.7">1.7 mm</quantity>
, known from
<collectingCountry id="81AAFFEAF24E141172F4E88356342ADB" box="[470,529,711,733]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
.............................................................
<taxonomicName id="3EBDC4F9F24E141177DCE883516F2ADA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1278,1354,711,732]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="CBC96368F24E141177DCE883516F2ADA" box="[1278,1354,711,732]" italics="true" pageId="39" pageNumber="40">C. nana</emphasis>
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<emphasis id="CBC96368F24E14117672E88351B92ADA" bold="true" box="[1360,1436,711,732]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672E88351B92ADA" box="[1360,1436,711,732]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="F902BF7AF24E141173B5E8A151B92AFF" blockId="39.[151,1437,153,762]" box="[151,1436,740,762]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E8A151B92AFF" box="[151,1436,740,762]" pageId="39" pageNumber="40">
-. TL 2.0 mm, known from
<collectingCountry id="81AAFFEAF24E141172F4E8A0563E2AFC" box="[470,539,740,762]" name="Guyana" pageId="39" pageNumber="40">Guiana</collectingCountry>
........................................................
<taxonomicName id="3EBDC4F9F24E141177EAE8A0516C2AFC" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1224,1353,740,762]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
<emphasis id="CBC96368F24E141177EAE8A0516C2AFC" box="[1224,1353,740,762]" italics="true" pageId="39" pageNumber="40">C. guyanense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672E8A051B92AFF" bold="true" box="[1360,1436,740,761]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672E8A051B92AFF" box="[1360,1436,740,761]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</subSubSection>
</treatment>
</document>

337
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<taxonomicName id="3EBDC4F9F262143D73B5EAA5558C28FA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,425,225,252]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F262143D73B5EAA5558C28FA" bold="true" box="[151,425,225,252]" italics="true" pageId="11" pageNumber="12">Calvariopsis borowieci</emphasis>
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<paragraph id="F902BF7AF262143D73B5EB4155052919" blockId="11.[151,527,225,287]" box="[151,288,261,287]" pageId="11" pageNumber="12">
(
<figureCitation id="6186A3FFF262143D73BDEB4154D92919" box="[159,252,261,287]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="11" pageNumber="12">Figs 1A</figureCitation>
,
<figureCitation id="6186A3FFF262143D722AEB4155322919" box="[264,279,261,287]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73B5EB0A55BF2AC9" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="F902BF7AF262143D73B5EB0A55BF2AC9" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73B5EB0A5560296E" bold="true" box="[151,325,333,360]" pageId="11" pageNumber="12">Type material.</emphasis>
<materialsCitation id="49D5B527F262143D7269EB0954ED298D" ID-GBIF-Occurrence="2239706624" collectingDate="2009-11-30" collectionCode="DBET" collectorName="Lech" country="Ecuador" location="Yanayacu-Rio" municipality="Aliso" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F262143D7269EB0955932961" box="[331,438,333,359]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF262143D7124EB0956732961" box="[518,598,333,359]" pageId="11" pageNumber="12">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF262143D7153EB0956DE2961" box="[625,763,333,359]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
\ road
<location id="FC62E9A1F262143D706AEB0957C22961" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D706AEB0957C22961" box="[840,999,333,359]" country="Ecuador" municipality="Aliso" name="Yanayacu-Rio" pageId="11" pageNumber="12">Yanayacu-Rio</location>
\
<collectingMunicipality id="19662500F262143D70D8EB09501C2961" box="[1018,1081,333,359]" pageId="11" pageNumber="12">Aliso</collectingMunicipality>
,
<date id="8D0399BAF262143D7767EB0950E62961" box="[1093,1219,333,360]" pageId="11" pageNumber="12" value="2009-11-30">
<collectingDate id="9D476052F262143D7767EB0950E62961" box="[1093,1219,333,360]" pageId="11" pageNumber="12" value="2009-11-30">30 XI 2009</collectingDate>
</date>
\ leg.
<collectorName id="5448DAACF262143D762AEB0951652961" box="[1288,1344,333,359]" pageId="11" pageNumber="12">Lech</collectorName>
Borow- iec”
</materialsCitation>
.
<materialsCitation id="49D5B527F262143D73F3EB36571129A9" ID-GBIF-Occurrence="2239706610" collectingDate="2009-12-17" collectionCode="DBET" collectorName="R. Ruta" country="Ecuador" elevation="2100" location="Male" pageId="11" pageNumber="12" specimenCount="8" specimenCount-female="8" stateProvince="Napo" typeStatus="paratype">
<typeStatus id="260601D8F262143D73F3EB365567298A" box="[209,322,370,396]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
.
<location id="FC62E9A1F262143D7268EB3555A1298D" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D7268EB3555A1298D" box="[330,388,369,395]" country="Ecuador" name="Male" pageId="11" pageNumber="12" stateProvince="Napo">Male</location>
(
<collectionCode id="9FAC27BFF262143D72B3EB3555C4298D" box="[401,481,369,395]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 77,
<collectingRegion id="3B797198F262143D71ABEB3656E3298A" box="[649,710,370,396]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Yanayacu \ Station,
<quantity id="3E45129FF262143D77B1EB35510C298D" box="[1171,1321,369,395]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F262143D77B1EB35510C298D" box="[1171,1321,369,395]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, \
<date id="8D0399BAF262143D720AEBD1559129A9" box="[296,436,405,432]" pageId="11" pageNumber="12" value="2009-12-17">
<collectingDate id="9D476052F262143D720AEBD1559129A9" box="[296,436,405,432]" pageId="11" pageNumber="12" value="2009-12-17">17 XII 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF262143D72D2EBD2566929A9" box="[496,588,405,432]" pageId="11" pageNumber="12">R. Ruta</collectorName>
”.
<specimenCount id="EFBB74F3F262143D7142EBD156E929A9" box="[608,716,405,431]" pageId="11" pageNumber="12" type="female">8 females</specimenCount>
(
<collectionCode id="9FAC27BFF262143D71FEEBD1570929A9" box="[732,812,405,431]" pageId="11" pageNumber="12">DBET</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="49D5B527F262143D701CEBD256EF29F1" ID-GBIF-Occurrence="2239706640" collectingDate="9 XII" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" elevation="2125" latitude="-0.6051111" location="Female" longLatPrecision="1" longitude="-77.88583" municipality="Aliso" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="2" stateProvince="Napo" typeStatus="paratype">
same data as holotype.
<location id="FC62E9A1F262143D7766EBD150B329A9" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D7766EBD150B329A9" box="[1092,1174,405,431]" latitude="-0.6051111" longLatPrecision="1" longitude="-77.88583" name="Female" pageId="11" pageNumber="12" stateProvince="Napo">Female</location>
(
<collectionCode id="9FAC27BFF262143D7784EBD150D329A9" box="[1190,1270,405,431]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 64,
<collectingRegion id="3B797198F262143D73B5EBFE54F129D2" box="[151,212,442,468]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Yanayacu \ Station, Bamboo hill trail, \
<quantity id="3E45129FF262143D708BEBFD502629D2" box="[937,1027,441,468]" metricMagnitude="3" metricUnit="m" metricValue="2.125" pageId="11" pageNumber="12" unit="m" value="2125.0">
<elevation id="72905849F262143D708BEBFD502629D2" box="[937,1027,441,468]" metricMagnitude="3" metricUnit="m" metricValue="2.125" pageId="11" pageNumber="12" unit="m" value="2125.0">2125 m</elevation>
</quantity>
,
<geoCoordinate id="9C89D9BDF262143D7733EBFD509129D5" box="[1041,1204,441,467]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="-0.6051111">S 00º36’18.4’’</geoCoordinate>
\
<geoCoordinate id="9C89D9BDF262143D77EDEBFE51A529D2" box="[1231,1408,441,468]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="-77.88583">W 77º53’09.0’’</geoCoordinate>
,
<date id="8D0399BAF262143D76ACEBFD549B29FE" pageId="11" pageNumber="12">
<collectingDate id="9D476052F262143D76ACEBFD549B29FE" pageId="11" pageNumber="12">9 XII</collectingDate>
</date>
\ 2009, leg.
<collectorName id="5448DAACF262143D7275EB9955FC29F1" box="[343,473,477,503]" pageId="11" pageNumber="12">Rafał Ruta</collectorName>
”.
<specimenCount id="EFBB74F3F262143D72D2EB99564529F1" box="[496,608,477,503]" pageId="11" pageNumber="12" type="female">2 females</specimenCount>
(
<collectionCode id="9FAC27BFF262143D7150EB9956E729F1" box="[626,706,477,503]" pageId="11" pageNumber="12">DBET</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="49D5B527F262143D71FAEB9955B32AC9" ID-GBIF-Occurrence="2239706633" collectingDate="1999-11-05" collectingDateMax="2009-12-17" collectingDateMin="1999-11-05" collectionCode="DBET, SEMC" collectorName="L. Borowiec &amp; Rafal Ruta &amp; Z. H. Fallin" country="Ecuador" elevation="2100" location="Female" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
“
<collectingCountry id="81AAFFEAF262143D71C3EB99574B29F1" box="[737,878,477,503]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
,
<collectingRegion id="3B797198F262143D705FEB9A579F29FE" box="[893,954,478,504]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov. \ Cosanga vic., Yanayacu \ Station,
<quantity id="3E45129FF262143D73B5E845550B2A1D" box="[151,302,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F262143D73B5E845550B2A1D" box="[151,302,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’ S, 77°53’W, \
<date id="8D0399BAF262143D7115E84557342A1D" box="[567,785,513,540]" pageId="11" pageNumber="12" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">
<collectingDate id="9D476052F262143D7115E84557342A1D" box="[567,785,513,540]" pageId="11" pageNumber="12" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">23 XI–17 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF262143D7079E84657C82A1D" box="[859,1005,513,540]" pageId="11" pageNumber="12">L. Borowiec</collectorName>
”.
<location id="FC62E9A1F262143D7723E84550762A1D" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D7723E84550762A1D" box="[1025,1107,513,539]" country="Ecuador" name="Female" pageId="11" pageNumber="12" stateProvince="Napo">Female</location>
(
<collectionCode id="9FAC27BFF262143D7741E84550962A1D" box="[1123,1203,513,539]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 19,
<collectingRegion id="3B797198F262143D7642E84651B82A1A" box="[1376,1437,514,540]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Yanayacu \ Station, moist forest near \ the road,
<quantity id="3E45129FF262143D7093E861506D2A39" box="[945,1096,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F262143D7093E861506D2A39" box="[945,1096,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, 27 \
<date id="8D0399BAF262143D765DE86254F72A65" pageId="11" pageNumber="12" value="2009-11">
<collectingDate id="9D476052F262143D765DE86254F72A65" pageId="11" pageNumber="12" value="2009-11">XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF262143D722CE80D55A82A65" box="[270,397,585,611]" pageId="11" pageNumber="12">Rafał Ruta</collectorName>
”. Female (
<collectionCode id="9FAC27BFF262143D7126E80D56722A65" box="[516,599,585,611]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="12" type="Museum">SEMC</collectionCode>
): “
<collectingCountry id="81AAFFEAF262143D7156E80D57262A65" box="[628,771,585,611]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
:
<collectingRegion id="3B797198F262143D702FE80E57692A62" box="[781,844,586,612]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
, Cosanga \ 1.0 km N on Baeza-Tena \ Road then
<quantity id="3E45129FF262143D765BE80D549E2A81" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="km" value="1.5">1.5 km</quantity>
SW on road to \ Sierra Azul,
<quantity id="3E45129FF262143D712CE82956792A8E" box="[526,604,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="11" pageNumber="12" unit="m" value="2150.0">
<elevation id="72905849F262143D712CE82956792A8E" box="[526,604,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="11" pageNumber="12" unit="m" value="2150.0">2150m</elevation>
</quantity>
\ 0°35’24”S, 77°53’19”W \
<date id="8D0399BAF262143D7082E82950742A81" box="[928,1105,621,648]" pageId="11" pageNumber="12" value="1999-11-05" valueMax="1999-11-07" valueMin="1999-11-05">
<collectingDate id="9D476052F262143D7082E82950742A81" box="[928,1105,621,648]" pageId="11" pageNumber="12" value="1999-11-05" valueMax="1999-11-07" valueMin="1999-11-05">5–7 NOV 1999</collectingDate>
</date>
;
<collectorName id="5448DAACF262143D777FE82A50C72A81" box="[1117,1250,621,648]" pageId="11" pageNumber="12">Z. H. Fallin</collectorName>
\ ECU1F99 124
<collectingMethod id="20FCC76DF262143D73B5E8D555502AAD" box="[151,373,657,683]" pageId="11" pageNumber="12">flight intercept trap</collectingMethod>
”. Female (
<collectionCode id="9FAC27BFF262143D72CEE8D556192AAD" box="[492,572,657,683]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 41,
<collectingRegion id="3B797198F262143D71C8E8D657022AAA" box="[746,807,658,684]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Antisana \ trail, Yanayacu,
<date id="8D0399BAF262143D7673E8D551A82AAA" box="[1361,1421,657,684]" pageId="11" pageNumber="12">
<collectingDate id="9D476052F262143D7673E8D551A82AAA" box="[1361,1421,657,684]" pageId="11" pageNumber="12">3 XII</collectingDate>
</date>
\ 2009, leg.
<collectorName id="5448DAACF262143D722FE8F155A92AC9" box="[269,396,693,719]" pageId="11" pageNumber="12">Rafał Ruta</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5E89D563D2B85" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="F902BF7AF262143D73E5E89D563D2B85" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5E89D55672AF2" bold="true" box="[199,322,729,756]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Moderately large (TL
<quantity id="3E45129FF262143D711FE89D56E22AF2" box="[573,711,729,756]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="11" pageNumber="12" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
), resembling black individuals of
<taxonomicName id="3EBDC4F9F262143D771CE89E50DE2AF5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1086,1275,730,756]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
<emphasis id="CBC96368F262143D771CE89E50DE2AF5" box="[1086,1275,730,756]" italics="true" pageId="11" pageNumber="12">C. yanayacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D7623E89E517F2AF2" bold="true" box="[1281,1370,730,756]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D7623E89E517F2AF2" box="[1281,1370,730,756]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Penis very long and narrow, apical plate of tergite VIII with M-shaped median process flanked with shallow emarginations covered with distinct denticles. Genitalia similar to that of
<taxonomicName id="3EBDC4F9F262143D700EE96657C22B3A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[812,999,802,828]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F262143D700EE96657C22B3A" box="[812,999,802,828]" italics="true" pageId="11" pageNumber="12">C. fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D70CCE966506D2B3A" bold="true" box="[1006,1096,802,828]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D70CCE966506D2B3A" box="[1006,1096,802,828]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but median process of tergite VIII is longer and better developed in
<taxonomicName id="3EBDC4F9F262143D7172E90256C42B59" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[592,737,837,864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F262143D7172E90256C42B59" box="[592,737,837,864]" italics="true" pageId="11" pageNumber="12">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D71C8E90257632B66" bold="true" box="[746,838,838,864]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D71C8E90257632B66" box="[746,838,838,864]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Similar to
<taxonomicName id="3EBDC4F9F262143D70EAE90250592B59" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[968,1148,837,864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F262143D70EAE90250592B59" box="[968,1148,837,864]" italics="true" pageId="11" pageNumber="12">C. venezuelense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D77A7E90250C42B66" bold="true" box="[1157,1249,838,864]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D77A7E90250C42B66" box="[1157,1249,838,864]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but body is less elongated and its sides are curved.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5E9C955732F01" pageId="11" pageNumber="12" type="description">
<paragraph id="F902BF7AF262143D73E5E9C9511A2CCE" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5E9C9557C2BAE" bold="true" box="[199,345,909,936]" pageId="11" pageNumber="12">Description.</emphasis>
Male. Body oblong oval, convex, covered with brown, suberect setae. Body uniformly dark brown, venter brown, legs yellowish-brown, antennae yellowish, darkening towards apex. Head very broad, 1.6× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by a 3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, with very subtle punctation. Elytra oblong, without traces of longitudinal ridges, sides subparallel in basal 1/2, curved and regularly converging to apex in posterior 1/2. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
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Male genitalia. Penis (L
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,
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mm,
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A–B) very long and narrow, widest in basal portion, parameroids short, very narrow, curved, rounded at apices, trigonium subtriangular, narrow, pointed at apex. Teg- men (L
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mm,
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) very long, sides diverging, apex with short lateral lobes, covered with setae. Sternite IX (L
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,
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mm,
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) subrectangular, with paired apodemes, apical margin rounded, api- cal portion covered with short, sparse setae. Tergite VIII (L
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mm,
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) with short apical plate, central portion of apical plate produced, M-shaped, with V-shaped emargination at apex, lateral portions of apical margin with a few denticles, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
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mm,
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) with membranous apical plate, apodemes longer than apical plate.
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Female. Body (
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) dark brown to black, with no orange pattern on dorsal portion, TL (n = 5) 2.60–3.10 (2.88) mm. Apical portion of each elytron with two concavities, shallow one and deeper one, situated closer to suture and covered with longer setae, forming a peculiar hooked tuft of setae (
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). Gonocoxites (
<figureCitation id="6186A3FFF262143D760EEC7D51B12E55" box="[1324,1428,1593,1619]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="11" pageNumber="12">Fig. 10A</figureCitation>
) broadly triangular, gonostyli subapical, bursal sclerite (L
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mm,
<figureCitation id="6186A3FFF262143D7732EC1950472E71" box="[1040,1122,1629,1655]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="11" pageNumber="12">Figs 10</figureCitation>
B–C) elongated, widened in posterior portion, with subconical outgrowth in middle.
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<emphasis id="CBC96368F262143D73E5ECE255DA2EC6" bold="true" box="[199,511,1701,1728]" pageId="11" pageNumber="12">Measurements and ratios.</emphasis>
Males (n = 2) TL 2.60–3.13 (2.87) mm, PL 0.50–0.57 (0.53) mm, PW 1.07–1.23 (1.15) mm, EL 2.33–2.67 (2.50) mm, EW 1.60–1.93 (1.77) mm, TL/EW 1.6, PW/PL 2.1–2.2 (2.2), EL/EW 1.4–1.5 (1.4), EL/PL 4.7.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5ED5557BB2F2D" box="[199,926,1809,1836]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="F902BF7AF262143D73E5ED5557BB2F2D" blockId="11.[151,1437,333,1908]" box="[199,926,1809,1836]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5ED5555442F2A" bold="true" box="[199,353,1809,1836]" pageId="11" pageNumber="12">Distribution.</emphasis>
Known from localities in
<collectingRegion id="3B797198F262143D71A5ED5657152F2D" box="[647,816,1809,1836]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo province</collectingRegion>
,
<collectingCountry id="81AAFFEAF262143D7019ED5557BF2F2D" box="[827,922,1809,1835]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5ED7251AF2F75" pageId="11" pageNumber="12" type="etymology">
<paragraph id="F902BF7AF262143D73E5ED7251AF2F75" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5ED72556B2F56" bold="true" box="[199,334,1846,1872]" pageId="11" pageNumber="12">Etymology.</emphasis>
Named after Prof. Lech Borowiec, an expert in
<taxonomicName id="3EBDC4F9F262143D707DED7150BF2F56" authority="" box="[863,1178,1845,1872]" class="Insecta" family="Chrysomelidae" genus="Cassidinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Chrysomelidae">Cassidinae (Chrysomelidae)</taxonomicName>
systematics, my teach- er, member of the expedition to
<collectingCountry id="81AAFFEAF262143D72DCED1D567E2F75" box="[510,603,1881,1907]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
in 2009 and collector of many specimens examined in the present study.
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</treatment>
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260
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<emphasis id="CBC96368F265143A73B5EFFE55FF2DD2" bold="true" box="[151,474,1466,1493]" italics="true" pageId="12" pageNumber="13">Calvariopsis fourgassiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F265143A72C0EFFE56652DD2" box="[482,576,1466,1492]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143A73B5EF9A550E2DFE" box="[151,299,1502,1528]" pageId="12" pageNumber="13" type="description">
<paragraph id="F902BF7AF265143A73B5EF9A550E2DFE" blockId="12.[151,576,1466,1528]" box="[151,299,1502,1528]" pageId="12" pageNumber="13">
(
<figureCitation id="6186A3FFF265143A73BDEF9A54DF2DFE" box="[159,250,1502,1528]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="12" pageNumber="13">Figs 1C</figureCitation>
,
<figureCitation id="6186A3FFF265143A7224EF9A55042DFE" box="[262,289,1502,1528]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143A73B5EC6257742E62" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="F902BF7AF265143A73B5EC6257742E62" blockId="12.[151,1437,1573,2032]" pageId="12" pageNumber="13">
<emphasis id="CBC96368F265143A73B5EC6255602E46" bold="true" box="[151,325,1573,1600]" pageId="12" pageNumber="13">Type material.</emphasis>
<materialsCitation id="49D5B527F265143A7269EC6255062E62" ID-GBIF-Occurrence="2239706639" collectingDate="1995-01-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" county="Fourgassie" location="Francaise" municipality="Montagne Kaw" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F265143A7269EC6255932E46" box="[331,438,1574,1600]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF265143A7124EC6256462E46" box="[518,611,1574,1600]" pageId="12" pageNumber="13">MNHG</collectionCode>
): “
<collectingCountry id="81AAFFEAF265143A715DEC6256FC2E46" box="[639,729,1574,1600]" name="French Guiana" pageId="12" pageNumber="13">Guyane</collectingCountry>
<location id="FC62E9A1F265143A71C2EC62576E2E46" LSID="urn:lsid:plazi:treatment:71140E6CF265143973B5EFFE573A2918:FC62E9A1F265143A71C2EC62576E2E46" box="[736,843,1574,1600]" country="French Guiana" county="Fourgassie" municipality="Montagne Kaw" name="Francaise" pageId="12" pageNumber="13">Francaise</location>
\
<collectingMunicipality id="19662500F265143A7042EC62502B2E46" box="[864,1038,1574,1600]" pageId="12" pageNumber="13">Montagne Kaw</collectingMunicipality>
\
<collectingCounty id="1063C7F6F265143A7701EC6250B92E46" box="[1059,1180,1574,1600]" pageId="12" pageNumber="13">Fourgassié</collectingCounty>
\ 15.9.–
<date id="8D0399BAF265143A77D6EC6151422E46" box="[1268,1383,1573,1600]" pageId="12" pageNumber="13" value="1995-01-10">
<collectingDate id="9D476052F265143A77D6EC6151422E46" box="[1268,1383,1573,1600]" pageId="12" pageNumber="13" value="1995-01-10">1.10.1995</collectingDate>
</date>
\
<collectorName id="5448DAACF265143A765EEC6254CD2E62" pageId="12" pageNumber="13">M. Kocian</collectorName>
lgt.”
</materialsCitation>
.
<materialsCitation id="49D5B527F265143A720CEC0E57682E62" ID-GBIF-Occurrence="2239706629" box="[302,845,1610,1636]" collectingDate="1995-01-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" county="Fourgassie" location="Francaise" municipality="Montagne Kaw" pageId="12" pageNumber="13" specimenCount="1" typeStatus="paratype">
<typeStatus id="260601D8F265143A720CEC0E55B62E62" box="[302,403,1610,1636]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="9FAC27BFF265143A72C1EC0E56652E62" box="[483,576,1610,1636]" pageId="12" pageNumber="13">MNHG</collectionCode>
): same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143A73E5EC2955A22F1E" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="F902BF7AF265143A73E5EC2955A22F1E" blockId="12.[151,1437,1573,2032]" pageId="12" pageNumber="13">
<emphasis id="CBC96368F265143A73E5EC2955672E8E" bold="true" box="[199,322,1645,1672]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Small (TL
<quantity id="3E45129FF265143A72E0EC2A56392E8E" box="[450,540,1646,1672]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="13" unit="mm" value="2.3">2.3 mm</quantity>
), dark brown, with rounded sides of elytra, externally resembling some group II species of
<taxonomicName id="3EBDC4F9F265143A722EECD655BE2EAA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[268,411,1682,1708]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F265143A722EECD655BE2EAA" box="[268,411,1682,1708]" italics="true" pageId="12" pageNumber="13">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F265143A7283ECD656222EAA" box="[417,519,1682,1708]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
Penis very long and narrow, strongly narrowed in its middle, apical plate of tergite VIII with M-shaped, shallowly notched median process and distinct denticles in lateral portions. Genitalia similar to
<taxonomicName id="3EBDC4F9F265143A7396EC9E55662EF2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[180,323,1754,1780]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F265143A7396EC9E55662EF2" box="[180,323,1754,1780]" italics="true" pageId="12" pageNumber="13">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F265143A7268EC9E55862EF2" bold="true" box="[330,419,1754,1780]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="D0FADE13F265143A7268EC9E55862EF2" box="[330,419,1754,1780]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but median process of tergite VIII is distinctly shorter and more weakly developed than in
<taxonomicName id="3EBDC4F9F265143A73B5ECBA55022F1E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,295,1790,1816]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F265143A73B5ECBA55022F1E" box="[151,295,1790,1816]" italics="true" pageId="12" pageNumber="13">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F265143A720CECBA55A22F1E" bold="true" box="[302,391,1790,1816]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="D0FADE13F265143A720CECBA55A22F1E" box="[302,391,1790,1816]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143973E5ED6556EA28D1" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="F902BF7AF265143873E5ED6555312E54" blockId="12.[151,1437,1573,2032]" lastBlockId="14.[151,1437,1484,2014]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<emphasis id="CBC96368F265143A73E5ED65557C2F3A" bold="true" box="[199,345,1825,1852]" pageId="12" pageNumber="13">Description.</emphasis>
Male. Body oval, convex, covered with brown, suberect setae. Head brown with yellowish frons and clypeus, pronotum and elytra dark brown, venter brown, legs yellowish-brown, antennae yellowish. Head very broad, 1.8× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by 3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
<caption id="ADC2EFF2F264143B73B5ED7D55832FE4" httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1849,1874]" targetBox="[178,1410,181,1810]" targetPageId="13">
<paragraph id="F902BF7AF264143B73B5ED7D55832FE4" blockId="13.[151,1437,1849,2018]" pageId="13" pageNumber="14">
<emphasis id="CBC96368F264143B73B5ED7D55062F54" bold="true" box="[151,291,1849,1874]" pageId="13" pageNumber="14">FIGURE 10.</emphasis>
<taxonomicName id="3EBDC4F9F264143B720FED7E558A2F54" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[301,431,1850,1874]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F264143B720FED7E558A2F54" box="[301,431,1850,1874]" italics="true" pageId="13" pageNumber="14">Calvariopsis</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B729AED7E563F2F54" bold="true" box="[440,538,1850,1874]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B729AED7E563F2F54" box="[440,538,1850,1874]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C)
<taxonomicName id="3EBDC4F9F264143B7614ED1B54992F9C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F264143B7614ED1B54992F9C" italics="true" pageId="13" pageNumber="14">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B73E0EDC655362F9C" bold="true" box="[194,275,1922,1946]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B73E0EDC655362F9C" box="[194,275,1922,1946]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D)
<taxonomicName id="3EBDC4F9F264143B7263EDC755812F9C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[321,420,1923,1946]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="CBC96368F264143B7263EDC755812F9C" box="[321,420,1923,1946]" italics="true" pageId="13" pageNumber="14">C. pittieri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B7288EDC655DE2F9C" bold="true" box="[426,507,1922,1946]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B7288EDC655DE2F9C" box="[426,507,1922,1946]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E–H)
<taxonomicName id="3EBDC4F9F264143B7164EDC756C12F9C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[582,740,1923,1946]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="yanyacuense" status="sp. nov.">
<emphasis id="CBC96368F264143B7164EDC756C12F9C" box="[582,740,1923,1946]" italics="true" pageId="13" pageNumber="14">C. yanyacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B71C9EDC6571E2F9C" bold="true" box="[747,827,1922,1946]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B71C9EDC6571E2F9C" box="[747,827,1922,1946]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; I–K)
<taxonomicName id="3EBDC4F9F264143B705DEDC750602F9C" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[895,1093,1921,1946]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F264143B705DEDC750292F9C" box="[895,1036,1922,1946]" italics="true" pageId="13" pageNumber="14">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
, specimen from Sao Paulo; L–M)
<taxonomicName id="3EBDC4F9F264143B73B5EDE355782FB8" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[151,349,1957,1982]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F264143B73B5EDE355012FB8" box="[151,292,1958,1982]" italics="true" pageId="13" pageNumber="14">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
, specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor.
</paragraph>
</caption>
<caption id="ADC2EFF2F267143873B5EF2555D92D9B" httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1377,1402]" targetBox="[153,1435,181,1322]" targetPageId="14">
<paragraph id="F902BF7AF267143873B5EF2555D92D9B" blockId="14.[151,1436,1377,1438]" pageId="14" pageNumber="15">
<emphasis id="CBC96368F267143873B5EF25553A2D7C" bold="true" box="[151,287,1377,1402]" pageId="14" pageNumber="15">FIGURE 11.</emphasis>
<taxonomicName id="3EBDC4F9F26714387207EF2556122D7C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[293,567,1377,1402]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F26714387207EF2556122D7C" box="[293,567,1377,1402]" italics="true" pageId="14" pageNumber="15">Calvariopsis fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F2671438711CEF2656AA2D7C" bold="true" box="[574,655,1378,1402]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="D0FADE13F2671438711CEF2656AA2D7C" box="[574,655,1378,1402]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="F902BF7AF267143873E5EC1857CC2F74" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
Male genitalia. Penis (L
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,
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W
<quantity id="3E45129FF26714387150EC1856862E70" box="[626,675,1628,1654]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="14" pageNumber="15" unit="mm" value="0.07">0.07</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF267143871C7EC1857122E70" box="[741,823,1628,1654]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Figs 11</figureCitation>
A–B) very long and narrow, widest in basal half, wid- ened also in apical portion, parameroids short, narrow, curved, trigonium subtriangular, narrow, pointed at apex. Tegmen (L
<quantity id="3E45129FF2671438723BECE0555A2EB8" box="[281,383,1700,1726]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="14" pageNumber="15" unit="mm" value="0.61">0.61 mm</quantity>
,
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W
<quantity id="3E45129FF26714387289ECE055F92EB8" box="[427,476,1700,1726]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="15" unit="mm" value="0.17">0.17</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2671438713FECE0565A2EB8" box="[541,639,1700,1726]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Fig. 11C</figureCitation>
) very long, sides diverging, apex with short lateral lobes, covered with setae. Sternite IX (L
<quantity id="3E45129FF2671438725EEC8C55C42EE4" box="[380,481,1736,1762]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="15" unit="mm" value="0.29">0.29 mm</quantity>
,
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W
<quantity id="3E45129FF26714387129EC8C56192EE4" box="[523,572,1736,1762]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="15" unit="mm" value="0.21">0.21</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF26714387158EC8C56F92EE4" box="[634,732,1736,1762]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Fig. 11D</figureCitation>
) subrectangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
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,
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W
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</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2671438774FECA850EB2F00" box="[1133,1230,1772,1798]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Fig. 11E</figureCitation>
) with short apical plate, central portion of apical plate produced, with small notch at apex, lateral portions of apical margin with a few denticles in mesal part, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF26714387781ED70512D2F48" box="[1187,1288,1844,1870]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="15" unit="mm" value="0.38">0.38 mm</quantity>
,
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W
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mm,
<figureCitation id="6186A3FFF267143873B5ED1C54D32F74" box="[151,246,1880,1906]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Fig. 11F</figureCitation>
) with membranous apical plate, apodemes longer than apical plate.
</paragraph>
<paragraph id="F902BF7AF267143873E5ED3855BB2F90" blockId="14.[151,1437,1484,2014]" box="[199,414,1916,1942]" pageId="14" pageNumber="15">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF267143873E5EDE450FC2FBC" blockId="14.[151,1437,1484,2014]" box="[199,1241,1952,1978]" pageId="14" pageNumber="15">
Variability.
<typeStatus id="260601D8F26714387268EDE455892FBC" box="[330,428,1952,1978]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
has a small, orange spot along adsutural portion in central part of elytra.
</paragraph>
<paragraph id="F902BF7AF267143973E5ED8056EA28D1" blockId="14.[151,1437,1484,2014]" lastBlockId="15.[151,1436,152,287]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="CBC96368F267143873E5ED8055DA2FD8" bold="true" box="[199,511,1987,2014]" pageId="14" pageNumber="15">Measurements and ratios.</emphasis>
Males (n = 2) TL 2.27–2.30 (2.28) mm, PL 0.47–0.50 (0.48) mm, PW 1.03–1.07 (1.05) mm, EL 1.83–1.93 (1.88) mm, EW 1.47–1.50 (1.48) mm, TL/EW 1.53–1.55 (1.54), PW/PL 2.07–2.29 (2.18), EL/EW 1.22–1.32 (1.27), EL/PL 3.67–4.14 (3.90).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973E5EAA4579028FC" box="[199,949,224,251]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="F902BF7AF266143973E5EAA4579028FC" blockId="15.[151,1436,152,287]" box="[199,949,224,251]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973E5EAA4554428FD" bold="true" box="[199,353,224,251]" pageId="15" pageNumber="16">Distribution.</emphasis>
Known only from its
<typeStatus id="260601D8F2661439717BEAA556AC28FD" box="[601,649,225,251]" pageId="15" pageNumber="16">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF2661439702AEAA4579428FC" box="[776,945,224,250]" name="French Guiana" pageId="15" pageNumber="16">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973E5EB41573A2918" box="[199,799,260,287]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="F902BF7AF266143973E5EB41573A2918" blockId="15.[151,1436,152,287]" box="[199,799,260,287]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973E5EB41556B2919" bold="true" box="[199,334,261,287]" pageId="15" pageNumber="16">Etymology.</emphasis>
Named after
<typeStatus id="260601D8F266143972C5EB4156322919" box="[487,535,261,287]" pageId="15" pageNumber="16">type</typeStatus>
locality of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

202
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<mods:title id="E80B1E27C0AA9A6B3B31480FF91EC6A7">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F266143973B5EB345607298D" box="[151,546,368,395]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="F902BF7AF266143973B5EB345607298D" blockId="15.[151,546,368,431]" box="[151,546,368,395]" pageId="15" pageNumber="16">
<heading id="A24A0816F266143973B5EB345607298D" bold="true" box="[151,546,368,395]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="CBC96368F266143973B5EB345607298D" bold="true" box="[151,546,368,395]" pageId="15" pageNumber="16">
<taxonomicName id="3EBDC4F9F266143973B5EB345598298D" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,445,368,395]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="panamense" status="sp. nov.">
<emphasis id="CBC96368F266143973B5EB345598298D" bold="true" box="[151,445,368,395]" italics="true" pageId="15" pageNumber="16">Calvariopsis panamense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F266143972E7EB355607298D" box="[453,546,369,395]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973B5EBD0550B29A9" box="[151,302,404,431]" pageId="15" pageNumber="16" type="description">
<paragraph id="F902BF7AF266143973B5EBD0550B29A9" blockId="15.[151,546,368,431]" box="[151,302,404,431]" pageId="15" pageNumber="16">
(
<figureCitation id="6186A3FFF266143973BDEBD054DE29A9" box="[159,251,404,431]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="15" pageNumber="16">Figs 1D</figureCitation>
,
<figureCitation id="6186A3FFF2661439722AEBD1550029A9" box="[264,293,405,431]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="15" pageNumber="16">12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973B5EB9956172A1D" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="F902BF7AF266143973B5EB9956172A1D" blockId="15.[151,1437,476,647]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973B5EB99556D29F1" bold="true" box="[151,328,476,503]" pageId="15" pageNumber="16">Type material.</emphasis>
<materialsCitation id="49D5B527F26614397270EB9856172A1D" ID-GBIF-Occurrence="2397829663" collectingDate="1993-06-10" collectionCode="SEMC" country="Panama" location="Old Gamboa Rd." municipality="Gamboa" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Panama" typeStatus="holotype">
<typeStatus id="260601D8F26614397270EB98559829F0" box="[338,445,476,502]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF26614397136EB99564229F1" box="[532,615,477,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="15" pageNumber="16" type="Museum">SEMC</collectionCode>
): “
<collectingCountry id="81AAFFEAF266143971A4EB99572129F1" box="[646,772,477,503]" name="Panama" pageId="15" pageNumber="16">PANAMA</collectingCountry>
:
<collectingRegion id="3B797198F26614397033EB99574F29F1" box="[785,874,477,503]" country="Panama" name="Panama" pageId="15" pageNumber="16">Panama</collectingRegion>
\
<collectingMunicipality id="19662500F266143970A4EB9857CD29F0" box="[902,1000,476,502]" pageId="15" pageNumber="16">Gamboa</collectingMunicipality>
,
<location id="FC62E9A1F266143970D4EB98509E29F0" LSID="urn:lsid:plazi:treatment:71140E6CF266142673B5EB3450432A39:FC62E9A1F266143970D4EB98509E29F0" box="[1014,1211,476,502]" country="Panama" municipality="Gamboa" name="Old Gamboa Rd." pageId="15" pageNumber="16" stateProvince="Panama">Old Gamboa Rd.</location>
\
<date id="8D0399BAF266143977F4EB99515129F1" box="[1238,1396,477,503]" pageId="15" pageNumber="16" value="1993-06-10">
<collectingDate id="9D476052F266143977F4EB99515129F1" box="[1238,1396,477,503]" pageId="15" pageNumber="16" value="1993-06-10">10 June 1993</collectingDate>
</date>
, C Michalski \ ex:
<collectingMethod id="20FCC76DF26614397264E84456012A1D" box="[326,548,512,539]" pageId="15" pageNumber="16">flight intercept trap</collectingMethod>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973E5E86056F82A81" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="F902BF7AF266143973E5E86056F82A81" blockId="15.[151,1437,476,647]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973E5E86055672A39" bold="true" box="[199,322,548,575]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Small (TL
<quantity id="3E45129FF266143972E2E861563C2A39" box="[448,537,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="15" pageNumber="16" unit="mm" value="2.4">2.4 mm</quantity>
), oval, identifiable only on the basis of male genitalia, especially unique shape of penis. Penis with wide basal half and narrow apical part, a condition unknown in other members of the genus. Apical plate of tergite VIII with very short median process.
</paragraph>
</subSubSection>
<caption id="ADC2EFF2F266143973B5EFA455BD2E1A" httpUri="https://zenodo.org/record/3403754/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1504,1529]" targetBox="[153,1435,668,1486]" targetPageId="15">
<paragraph id="F902BF7AF266143973B5EFA455BD2E1A" blockId="15.[151,1436,1504,1565]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973B5EFA4553A2DFF" bold="true" box="[151,287,1504,1529]" pageId="15" pageNumber="16">FIGURE 12.</emphasis>
<emphasis id="CBC96368F26614397206EFA556382DFE" box="[292,541,1505,1529]" italics="true" pageId="15" pageNumber="16">Calvariopsis panamense</emphasis>
<emphasis id="CBC96368F26614397100EFA556572DFF" bold="true" box="[546,626,1505,1529]" pageId="15" pageNumber="16">sp. nov.</emphasis>
, male. A) penis; B) tegmen; C) sternite IX; D) tergite and sternite VIII; E) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
<subSubSection id="B1A7ECF1F266142673E5EC0E56CB29F1" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="F902BF7AF266143973E5EC0E55412FAF" blockId="15.[151,1437,1610,2033]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973E5EC0E557C2E63" bold="true" box="[199,345,1610,1637]" pageId="15" pageNumber="16">Description.</emphasis>
Male. Body oval, convex, covered with yellowish, suberect setae. Head brown with lighter frons and clypeus, pronotum and elytra dark brown, venter brown, legs and antennae yellowish. Head very broad, 1.7 times wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by a 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 0.7–1.0× diameter of a puncture.
</paragraph>
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Male genitalia. Penis (L
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,
<geoCoordinate id="9C89D9BDF26614397174EDF0568C2FCB" box="[598,681,1971,1998]" degrees="0.11" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="555" value="-0.11">
W
<quantity id="3E45129FF2661439715BEDF7568C2FCB" box="[633,681,1971,1997]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="0.11">0.11</quantity>
</geoCoordinate>
mm,
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) moderately long, wide in basal half and narrowed in apical portion, parameroids short, narrow, with subparallel sides, rounded at apices, trigonium subtriangular, rounded at apex. Tegmen (L
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,
<geoCoordinate id="9C89D9BDF2791426716DEADD568428B5" box="[591,673,153,179]" degrees="0.19" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.19">
W
<quantity id="3E45129FF27914267152EADD568428B5" box="[624,673,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="16" pageNumber="17" unit="mm" value="0.19">0.19</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF279142671C1EADD576228B5" box="[739,839,153,179]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="16" pageNumber="17">Fig. 12B</figureCitation>
) short, sides diverging, apex with short lateral lobes, covered with sparse setae. Sternite VIII (L
<quantity id="3E45129FF2791426715FEAF956C128D1" box="[637,740,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="17" unit="mm" value="0.17">0.17 mm</quantity>
,
<geoCoordinate id="9C89D9BDF279142671CDEAF9576728D1" box="[751,834,189,215]" degrees="0.06" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.06">
W
<quantity id="3E45129FF27914267033EAF9576728D1" box="[785,834,189,215]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="mm" value="0.06">0.06</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF279142670A6EAF957CF28D1" box="[900,1002,189,215]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="16" pageNumber="17">Fig. 12D</figureCitation>
) very small, transverse. Sternite IX (L
<quantity id="3E45129FF279142673B5EAA554DA28FD" box="[151,255,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="17" unit="mm" value="0.26">0.26 mm</quantity>
,
<geoCoordinate id="9C89D9BDF2791426722EEAA5554528FD" box="[268,352,225,251]" degrees="0.20" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.2">
W
<quantity id="3E45129FF2791426720DEAA5554528FD" box="[303,352,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="mm" value="0.2">0.20</quantity>
</geoCoordinate>
mm) subtriangular, with paired apodemes, apical margin rounded, apical portion covered with sparse setae. Tergite VIII (L
<quantity id="3E45129FF279142672F4EB4156192919" box="[470,572,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="16" pageNumber="17" unit="mm" value="0.31">0.31 mm</quantity>
,
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W
<quantity id="3E45129FF2791426714AEB4156BC2919" box="[616,665,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="17" unit="mm" value="0.34">0.34</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF279142671FBEB41571B2919" box="[729,830,261,287]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="16" pageNumber="17">Fig. 12D</figureCitation>
) with short apical plate, central portion of apical plate slightly produced, lateral portions of apical margin with numerous small denticles, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF2791426711BEB0956BA2961" box="[569,671,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="17" unit="mm" value="0.33">0.33 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27914267188EB0956D92961" box="[682,764,333,359]" degrees="0.24" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.24">
W
<quantity id="3E45129FF279142671E9EB0956D92961" box="[715,764,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="17" unit="mm" value="0.24">0.24</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2791426701FEB0957BA2961" box="[829,927,333,359]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="16" pageNumber="17">Fig. 12E</figureCitation>
) with membranous apical plate, apodemes as long as apical plate.
</paragraph>
<paragraph id="F902BF7AF279142673E5EBD155BB29A9" blockId="16.[151,1437,153,575]" box="[199,414,405,431]" pageId="16" pageNumber="17">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF279142673E5EBFD56CB29F1" blockId="16.[151,1437,153,575]" pageId="16" pageNumber="17">
<emphasis id="CBC96368F279142673E5EBFD55D929D5" bold="true" box="[199,508,440,467]" pageId="16" pageNumber="17">Measurements and ratios.</emphasis>
Male (n = 1) TL
<quantity id="3E45129FF2791426719CEBFD570129D5" box="[702,804,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="16" pageNumber="17" unit="mm" value="2.37">2.37 mm</quantity>
, PL
<quantity id="3E45129FF27914267077EBFC579E29D5" box="[853,955,440,467]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="mm" value="0.5">0.50 mm</quantity>
, PW
<quantity id="3E45129FF279142670D4EBFD507929D5" box="[1014,1116,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="16" pageNumber="17" unit="mm" value="1.13">1.13 mm</quantity>
, EL
<quantity id="3E45129FF279142677ADEBFD50D029D5" box="[1167,1269,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="16" pageNumber="17" unit="mm" value="2.03">2.03 mm</quantity>
, EW
<quantity id="3E45129FF27914267610EBFC51BD29D5" box="[1330,1432,440,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="16" pageNumber="17" unit="mm" value="1.53">1.53 mm</quantity>
, TL/EW 1.54, PW/PL 2.27, EL/EW 1.33, EL/PL 4.07.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F279142673E5E84457552A1D" box="[199,880,512,539]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="F902BF7AF279142673E5E84457552A1D" blockId="16.[151,1437,153,575]" box="[199,880,512,539]" pageId="16" pageNumber="17">
<emphasis id="CBC96368F279142673E5E84455442A1D" bold="true" box="[199,353,512,539]" pageId="16" pageNumber="17">Distribution.</emphasis>
Known only from the
<typeStatus id="260601D8F27914267142E84556B52A1D" box="[608,656,513,539]" pageId="16" pageNumber="17">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF2791426702DE845574E2A1D" box="[783,875,513,539]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F279142673E5E86150432A39" box="[199,1126,549,575]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="F902BF7AF279142673E5E86150432A39" blockId="16.[151,1437,153,575]" box="[199,1126,549,575]" pageId="16" pageNumber="17">
<emphasis id="CBC96368F279142673E5E861556B2A39" bold="true" box="[199,334,549,575]" pageId="16" pageNumber="17">Etymology.</emphasis>
Named after
<collectingCountry id="81AAFFEAF279142672C5E86156662A39" box="[487,579,549,575]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
, where the
<typeStatus id="260601D8F279142671E0E86156D72A39" box="[706,754,549,575]" pageId="16" pageNumber="17">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7B75A3E2B9BE3BA845B3285C43A28FC9">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F270142F73B5EE65561E2C3D" box="[151,571,1057,1084]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="F902BF7AF270142F73B5EE65561E2C3D" blockId="25.[151,571,1057,1119]" box="[151,571,1057,1084]" pageId="25" pageNumber="26">
<heading id="A24A0816F270142F73B5EE65561E2C3D" bold="true" box="[151,571,1057,1084]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="CBC96368F270142F73B5EE65561E2C3D" bold="true" box="[151,571,1057,1084]" pageId="25" pageNumber="26">
<taxonomicName id="3EBDC4F9F270142F73B5EE6555F02C3A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,469,1057,1084]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
<emphasis id="CBC96368F270142F73B5EE6555F02C3A" bold="true" box="[151,469,1057,1084]" italics="true" pageId="25" pageNumber="26">Calvariopsis bituberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F270142F72FFEE65561E2C3D" box="[477,571,1057,1083]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F270142F73B5EE01558D2C59" box="[151,424,1092,1119]" pageId="25" pageNumber="26" type="description">
<paragraph id="F902BF7AF270142F73B5EE01558D2C59" blockId="25.[151,571,1057,1119]" box="[151,424,1092,1119]" pageId="25" pageNumber="26">
(
<figureCitation id="6186A3FFF270142F73BDEE0154DE2C59" box="[159,251,1093,1119]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="25" pageNumber="26">Figs 1Q</figureCitation>
,
<figureCitation id="6186A3FFF270142F722AEE0155312C59" box="[264,276,1093,1119]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="25" pageNumber="26">3</figureCitation>
D–F, 5B, 20)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F270142F73B5EEC957012CCD" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="F902BF7AF270142F73B5EEC957012CCD" blockId="25.[151,1437,1164,2019]" pageId="25" pageNumber="26">
<emphasis id="CBC96368F270142F73B5EEC955602CA1" bold="true" box="[151,325,1164,1191]" pageId="25" pageNumber="26">Type material.</emphasis>
<materialsCitation id="49D5B527F270142F726FEEC954EE2CCD" ID-GBIF-Occurrence="2239706636" collectingDate="1995-11-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" location="Francaise" municipality="Mont Grand Matoury" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F270142F726FEEC9559D2CA1" box="[333,440,1165,1191]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF270142F712BEEC956432CA1" box="[521,614,1165,1191]" pageId="25" pageNumber="26">MNHG</collectionCode>
): “
<collectingCountry id="81AAFFEAF270142F71A0EEC956F92CA1" box="[642,732,1165,1191]" name="French Guiana" pageId="25" pageNumber="26">Guyane</collectingCountry>
<location id="FC62E9A1F270142F71C1EEC9576B2CA1" LSID="urn:lsid:plazi:treatment:71140E6CF270142C73B5EE6550C82944:FC62E9A1F270142F71C1EEC9576B2CA1" box="[739,846,1165,1191]" country="French Guiana" municipality="Mont Grand Matoury" name="Francaise" pageId="25" pageNumber="26">Francaise</location>
\
<collectingMunicipality id="19662500F270142F7047EECA50722CAE" box="[869,1111,1165,1192]" pageId="25" pageNumber="26">Mont Grand Matoury</collectingMunicipality>
\ 2.–
<date id="8D0399BAF270142F77AFEEC8512B2CA1" box="[1165,1294,1164,1191]" pageId="25" pageNumber="26" value="1995-11-10">
<collectingDate id="9D476052F270142F77AFEEC8512B2CA1" box="[1165,1294,1164,1191]" pageId="25" pageNumber="26" value="1995-11-10">11.10.1995</collectingDate>
</date>
\
<collectorName id="5448DAACF270142F7606EECA51B92CA1" box="[1316,1436,1165,1192]" pageId="25" pageNumber="26">M. Kocian</collectorName>
lgt.”
</materialsCitation>
.
<materialsCitation id="49D5B527F270142F73F4EEF657052CCD" ID-GBIF-Occurrence="2239706620" box="[214,800,1200,1228]" collectingDate="1995-11-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" location="Francaise" municipality="Mont Grand Matoury" pageId="25" pageNumber="26" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
<typeStatus id="260601D8F270142F73F4EEF655622CCA" box="[214,327,1202,1228]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="EFBB74F3F270142F7273EEF455822CCD" box="[337,423,1200,1227]" pageId="25" pageNumber="26" type="male">5 males</specimenCount>
(
<collectionCode id="9FAC27BFF270142F7294EEF556362CCD" box="[438,531,1201,1227]" pageId="25" pageNumber="26">MNHG</collectionCode>
): same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F270142F73E5EE90505E2D31" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="F902BF7AF270142F73E5EE90505E2D31" blockId="25.[151,1437,1164,2019]" pageId="25" pageNumber="26">
<emphasis id="CBC96368F270142F73E5EE9055672CE9" bold="true" box="[199,322,1236,1263]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
Small (TL 1.9–2.0 mm), oval species externally resembling many other species included in group II of
<taxonomicName id="3EBDC4F9F270142F7394EEBD55602D15" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[182,325,1273,1299]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F270142F7394EEBD55602D15" box="[182,325,1273,1299]" italics="true" pageId="25" pageNumber="26">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F270142F726FEEBE55912D12" box="[333,436,1274,1300]" pageId="25" pageNumber="26" rank="genus">gen. nov.</taxonomicNameLabel>
Tegmen with basal apodemes, tergite VIII with narrow but deep U-shaped emargination in median portion of apical plate. Apex of abdominal ventrite 5 with two small tubercles.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F270142C73E5EF04568028FD" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="F902BF7AF270142F73E5EF0457742EA6" blockId="25.[151,1437,1164,2019]" pageId="25" pageNumber="26">
<emphasis id="CBC96368F270142F73E5EF04557C2D5D" bold="true" box="[199,345,1344,1371]" pageId="25" pageNumber="26">Description.</emphasis>
Male. Body oval, convex, covered with brownish, suberect setae. Dorsum brown, head slightly lighter than pronotum and elytra, venter brown, legs, antennae and mouthparts yellowish. Head broad, 1.9× wider than width of interocular space, with subtle punctation, punctures small, granulate, separated by 3.0× diameter of a puncture, eyes large, slightly protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 3.0–4.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles right-angled. Scutellar shield subtriangular, almost impunctate. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture. Abdominal ventrite 5 with two small, subtriangular tubercles (
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,
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) at apex.
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Male genitalia. Penis (L
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,
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W
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mm,
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) relatively wide, with curved sides, parameroids short, curved, with subtriangular apices, trigonium subtriangular, pointed at apex. Tegmen (L
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,
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W
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mm,
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) U-shaped, with apodemes in basal portion, apices of parameres rounded, setose. Sternite IX (L
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,
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W
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mm,
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) elongate oval, with single, relatively short apodeme, apical margin rounded, covered with short, sparse setae. Tergite VIII (L
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,
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W
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mm,
<figureCitation id="6186A3FFF270142F71FCED7D57672F55" box="[734,834,1849,1875]" captionStart="FIGURE 20" captionStartId="26.[151,250,1241,1266]" captionTargetBox="[153,1435,361,1198]" captionTargetId="figure@26.[151,1436,361,1198]" captionTargetPageId="26" captionText="FIGURE 20. Calvariopsis bituberculata sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX; E) apex of abdomen. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783774/files/figure.png" pageId="25" pageNumber="26">Fig. 20C</figureCitation>
) with deep, U-shaped emargination in central portion of apical margin, apical margin subtly setose, apodemes as long as apical plate, slightly diverging. Tergite IX (L
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,
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W
<quantity id="3E45129FF270142F7209EDC555792F9D" box="[299,348,1921,1947]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="25" pageNumber="26" unit="mm" value="0.26">0.26</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF270142F72BFEDC556262F9D" box="[413,515,1921,1947]" captionStart="FIGURE 20" captionStartId="26.[151,250,1241,1266]" captionTargetBox="[153,1435,361,1198]" captionTargetId="figure@26.[151,1436,361,1198]" captionTargetPageId="26" captionText="FIGURE 20. Calvariopsis bituberculata sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX; E) apex of abdomen. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783774/files/figure.png" pageId="25" pageNumber="26">Fig. 20D</figureCitation>
) with membranous and relatively narrow apical plate, apodemes as long as apical plate, strongly diverging.
</paragraph>
<paragraph id="F902BF7AF270142F73E5ED8D55BB2FE5" blockId="25.[151,1437,1164,2019]" box="[199,414,1993,2019]" pageId="25" pageNumber="26">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF273142C73E5EADD568028FD" blockId="26.[151,1436,152,323]" pageId="26" pageNumber="27">
<emphasis id="CBC96368F273142C73E5EADD55DB28B5" bold="true" box="[199,510,152,179]" pageId="26" pageNumber="27">Measurements and ratios.</emphasis>
Males (n = 2) TL 1.93–2.00 (1.97) mm, PL 0.43 (0.43) mm, PW 0.97–1.00 (0.98) mm, EL 1.63–1.70 (1.67) mm, EW 1.30–1.33 (1.31) mm, TL/EW 1.45–1.54 (1.50), PW/PL 2.23–2.31 (2.26), EL/ EW 1.25–1.31 (1.27), EL/PL 3.77–3.92 (3.86).
</paragraph>
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<emphasis id="CBC96368F273142C73E5EB4055442919" bold="true" box="[199,353,260,287]" pageId="26" pageNumber="27">Distribution.</emphasis>
Known only from the
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locality in
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.
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<subSubSection id="B1A7ECF1F273142C73E5EB6D50C82944" box="[199,1261,296,323]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="F902BF7AF273142C73E5EB6D50C82944" blockId="26.[151,1436,152,323]" box="[199,1261,296,323]" pageId="26" pageNumber="27">
<emphasis id="CBC96368F273142C73E5EB6D556B2945" bold="true" box="[199,334,297,323]" pageId="26" pageNumber="27">Etymology.</emphasis>
The name refers to the presence of paired tubercles on the last abdominal ventrite.
</paragraph>
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270
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<taxonomicName id="3EBDC4F9F273142C73B5EF2C56672D84" authority="(Pic, 1916)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[151,578,1384,1411]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brasiliense" status="comb. nov.">
<emphasis id="CBC96368F273142C73B5EF2C55932D85" bold="true" box="[151,438,1384,1411]" italics="true" pageId="26" pageNumber="27">Calvariopsis brasiliense</emphasis>
(
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)
</taxonomicName>
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(
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,
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,
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)
</paragraph>
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=
<taxonomicName id="3EBDC4F9F273142C738EEF9456122DEE" authority="Pic, 1916: 7" authorityName="Pic. Both" authorityPageNumber="7" authorityYear="1916" box="[172,567,1488,1513]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="CBC96368F273142C738EEF9455922DEE" box="[172,439,1488,1512]" italics="true" pageId="26" pageNumber="27">Prionocyphon brasiliensis</emphasis>
<treatmentCitation id="781C996BF273142C729FEF9456122DEE" author="Pic, M." box="[445,567,1488,1513]" page="7" pageId="26" pageNumber="27" year="1916">
<bibRefCitation id="9D2CC28BF273142C729FEF9456052DEF" author="Pic, M." box="[445,544,1488,1513]" pageId="26" pageNumber="27" pagination="1 - 20" refId="ref23088" refString="Pic, M. (1916) Descriptions abregees diverses. Melanges exotico-entomologiques, 20 (1), 1 - 20." type="journal article" year="1916">Pic, 1916</bibRefCitation>
: 7
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F273142C73B5EC5357202EE3" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="F902BF7AF273142C73B5EC5356A42E7F" blockId="26.[151,1437,1558,2017]" pageId="26" pageNumber="27">
<emphasis id="CBC96368F273142C73B5EC5355602E37" bold="true" box="[151,325,1558,1585]" pageId="26" pageNumber="27">Type material.</emphasis>
<materialsCitation id="49D5B527F273142C726EEC5356A42E7F" ID-GBIF-Occurrence="2239706652" collectionCode="MNHN" country="Brazil" location="Pic's" pageId="26" pageNumber="27" specimenCount="2" typeStatus="holotype">
<typeStatus id="260601D8F273142C726EEC5355922E37" box="[332,439,1559,1585]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF273142C712AEC5356402E37" box="[520,613,1559,1585]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
): “Blumenau \
<collectingCountry id="81AAFFEAF273142C702FEC5357712E37" box="[781,852,1559,1585]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
” [handwritten]; “Type” [yellow label, handwritten]; “TYPE” [red label, printed]; “
<taxonomicName id="3EBDC4F9F273142C72CDEC7F56AA2E53" authorityName="Redtenbacher" authorityYear="1858" box="[495,655,1595,1621]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="41" phylum="Arthropoda" rank="genus">Prionocyphon</taxonomicName>
\
<taxonomicName id="3EBDC4F9F273142C718BEC7F57732E53" authority="Pic" authorityName="Pic. Both" authorityYear="1916" box="[681,854,1595,1621]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brasiliensis">brasiliensis Pic</taxonomicName>
” [in
<location id="FC62E9A1F273142C70B3EC7F57E22E53" LSID="urn:lsid:plazi:treatment:71140E6CF273142A73B5EF2C506328D1:FC62E9A1F273142C70B3EC7F57E22E53" box="[913,967,1595,1621]" country="Brazil" name="Pic's" pageId="26" pageNumber="27">Pic’s</location>
hand]; “
<typeStatus id="260601D8F273142C7709EC7F50F02E53" box="[1067,1237,1595,1621]" pageId="26" pageNumber="27" type="holotype">HOLOTYPUS</typeStatus>
\
<taxonomicName id="3EBDC4F9F273142C77CCEC7F51AE2E53" authorityName="Redtenbacher" authorityYear="1858" box="[1262,1419,1595,1621]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="41" phylum="Arthropoda" rank="genus">Prionocyphon</taxonomicName>
\ brasiliensis \
<bibRefCitation id="9D2CC28BF273142C7209EC1B55B22E7F" author="Pic, M." box="[299,407,1631,1657]" pageId="26" pageNumber="27" pagination="1 - 20" refId="ref23088" refString="Pic, M. (1916) Descriptions abregees diverses. Melanges exotico-entomologiques, 20 (1), 1 - 20." type="journal article" year="1916">Pic, 1916</bibRefCitation>
” [red label, printed].
</materialsCitation>
</paragraph>
<paragraph id="F902BF7AF273142C73E5ECC657202EE3" blockId="26.[151,1437,1558,2017]" pageId="26" pageNumber="27">
<emphasis id="CBC96368F273142C73E5ECC656122E9B" bold="true" box="[199,567,1666,1693]" pageId="26" pageNumber="27">Additional material examined.</emphasis>
Male, female (NMPC): “
<collectingRegion id="3B797198F273142C7078ECC757DD2E9B" box="[858,1016,1667,1693]" country="Brazil" name="Sao Paulo" pageId="26" pageNumber="27">SAO PAULO</collectingRegion>
\ BRAS MRÁZ LGT. \ MUS. PRA- GENSE” [printed]. Male (ZMHB): “
<collectingCountry id="81AAFFEAF273142C7111ECE356BE2EC7" box="[563,667,1703,1729]" name="Brazil" pageId="26" pageNumber="27">Brasilien</collectingCountry>
\ Cubatao \
<date id="8D0399BAF273142C7005ECE357592EC7" box="[807,892,1703,1729]" pageId="26" pageNumber="27" value="1927-11-02">2.XI.27</date>
\ E. Schrim S. G.”. Female (NHM): “Ilha Santo Amaro \ nr Santos,
<collectingCountry id="81AAFFEAF273142C7250EC8F559F2EE3" box="[370,442,1739,1765]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
. \ G. E. Bryant \
<date id="8D0399BAF273142C7154EC8F56D22EE3" box="[630,759,1739,1765]" pageId="26" pageNumber="27" value="1912-04-11">11.IV.1912</date>
”.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F273142D73E5ECAA55762F3B" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="F902BF7AF273142D73E5ECAA553128FD" blockId="26.[151,1437,1558,2017]" lastBlockId="27.[151,1437,153,539]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="CBC96368F273142C73E5ECAA55502F0F" bold="true" box="[199,373,1774,1801]" pageId="26" pageNumber="27">Redescription.</emphasis>
Male. Body oblong oval, convex, covered with brownish suberect setae. Body uniformly dark brown, venter brown. Head very broad, 1.8× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by 1.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 1.5–2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, almost impunctate. Angle between pronotum and elytra subtly marked in dorsal outline. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length. Sides slightly curved in apical 1/2, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 1.5× diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF272142D73E5EB41569A2A1D" blockId="27.[151,1437,153,539]" pageId="27" pageNumber="28">
Male genitalia. Penis (L
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,
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W
<quantity id="3E45129FF272142D714FEB4156BB2919" box="[621,670,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="mm" value="0.1">0.10</quantity>
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mm,
<figureCitation id="6186A3FFF272142D71FDEB4157612919" box="[735,836,261,287]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="27" pageNumber="28">Fig. 21A</figureCitation>
) with subtly curved sides, parameroids short, curved, with subtriangular apices, trigonium narrow, short, pointed at apex. Tegmen (L
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,
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W
<quantity id="3E45129FF272142D77BEEB6C50E82945" box="[1180,1229,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="28" unit="mm" value="0.15">0.15</quantity>
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mm,
<figureCitation id="6186A3FFF272142D762FEB6D51552945" box="[1293,1392,297,323]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="27" pageNumber="28">Fig. 21B</figureCitation>
) Ushaped, with apodemes in basal portion, apices of parameres rounded, setose. Sternite IX (L
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,
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W
<quantity id="3E45129FF272142D7610EB0951462961" box="[1330,1379,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="27" pageNumber="28" unit="mm" value="0.17">0.17</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF272142D73B5EB3554D9298D" box="[151,252,369,395]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="27" pageNumber="28">Fig. 21C</figureCitation>
) elongate oval, with single, relatively short apodeme, apical margin rounded, covered with short, sparse setae. Tergite VIII (L
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,
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W
<quantity id="3E45129FF272142D713FEBD1566B29A9" box="[541,590,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="27" pageNumber="28" unit="mm" value="0.37">0.37</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF272142D71B2EBD1573D29A9" box="[656,792,405,431]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="27" pageNumber="28">Figs 21D, E</figureCitation>
) with deep, V-shaped emargination of apical margin, and two distinct denticles in lateral portions of apical margin, apical margin not setose, apodemes as long as apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF272142D728AEB99562B29F1" box="[424,526,477,503]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="28" unit="mm" value="0.33">0.33 mm</quantity>
,
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W
<quantity id="3E45129FF272142D711BEB99564F29F1" box="[569,618,477,503]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="27" pageNumber="28" unit="mm" value="0.34">0.34</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF272142D7188EB99572C29F1" box="[682,777,477,503]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="27" pageNumber="28">Fig. 21F</figureCitation>
) with membranous and relatively narrow apical plate, apo- demes as long as apical plate, strongly diverging.
</paragraph>
<caption id="ADC2EFF2F272142D73B5EFDC57142DD2" httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1432,1457]" targetBox="[153,1435,616,1388]" targetPageId="27">
<paragraph id="F902BF7AF272142D73B5EFDC57142DD2" blockId="27.[151,1437,1432,1493]" pageId="27" pageNumber="28">
<emphasis id="CBC96368F272142D73B5EFDC55042DB7" bold="true" box="[151,289,1432,1457]" pageId="27" pageNumber="28">FIGURE 21.</emphasis>
<taxonomicName id="3EBDC4F9F272142D720AEFDD567C2DB7" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[296,601,1432,1457]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F272142D720AEFDD563A2DB7" box="[296,543,1433,1457]" italics="true" pageId="27" pageNumber="28">Calvariopsis brasiliense</emphasis>
(Pic)
</taxonomicName>
, male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="F902BF7AF272142D73E5EC47559F2EAB" blockId="27.[151,1437,1539,2033]" pageId="27" pageNumber="28">
Female. Body (
<figureCitation id="6186A3FFF272142D7257EC4755E22E1B" box="[373,455,1539,1565]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="27" pageNumber="28">Fig. 1S</figureCitation>
) dark brown, TL (n = 1)
<quantity id="3E45129FF272142D71C4EC4757682E1B" box="[742,845,1539,1565]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="27" pageNumber="28" unit="mm" value="2.6">2.60 mm</quantity>
. Posterior portion of each elytron with shallow, oval depression covered with wide scales, situated close to suture, slightly behind the middle (
<figureCitation id="6186A3FFF272142D77BCEC6350DD2E47" box="[1182,1272,1575,1601]" captionStart="FIGURE 8" captionStartId="10.[151,250,1352,1377]" captionTargetBox="[153,1435,181,1302]" captionTargetId="figure@10.[153,1438,181,1302]" captionTargetPageId="10" captionText="FIGURE 8. Calvariopsis gen. nov., group I (A, C, E–F) and group II (B, D), females, apical portion of elytra, excitators. A) C. pittieri; B) C. brasiliense, an arrow indicates patch of modified setae; C) C. sp., Colombia, Gorgona Island; D) C. sp., Ecuador, an arrow indicates patch of modified setae; E) C. sp., Paraguay; F) C. sp., Peru." httpUri="https://zenodo.org/record/2783720/files/figure.png" pageId="27" pageNumber="28">Fig. 8B</figureCitation>
). Gonocoxites (
<figureCitation id="6186A3FFF272142D7382EC0F54CB2E63" box="[160,238,1611,1637]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="27" pageNumber="28">Fig. 10</figureCitation>
J–L) triangular, gonostyli subapical, bursal sclerite (
<figureCitation id="6186A3FFF272142D7014EC0F57982E63" box="[822,957,1611,1637]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="27" pageNumber="28">Fig. 10M, L</figureCitation>
<quantity id="3E45129FF272142D70E7EC0F500F2E63" box="[965,1066,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="mm" value="0.32">0.32 mm</quantity>
,
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W
<quantity id="3E45129FF272142D7774EC0F50A22E63" box="[1110,1159,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="27" pageNumber="28" unit="mm" value="0.12">0.12</quantity>
</geoCoordinate>
mm) elongated, anterior portion with small, subconical outgrowth, central portion covered with subtle microreticulation resembling scales, posterior portion rod-like.
</paragraph>
<paragraph id="F902BF7AF272142D73E5ECF356CB2EF3" blockId="27.[151,1437,1539,2033]" pageId="27" pageNumber="28">
<emphasis id="CBC96368F272142D73E5ECF355DB2ED7" bold="true" box="[199,510,1718,1745]" pageId="27" pageNumber="28">Measurements and ratios.</emphasis>
Male (n = 1) TL
<quantity id="3E45129FF272142D71E1ECF2570C2ED7" box="[707,809,1718,1745]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="27" pageNumber="28" unit="mm" value="2.35">2.35 mm</quantity>
, PL
<quantity id="3E45129FF272142D707EECF257E62ED7" box="[860,963,1718,1745]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="28" unit="mm" value="0.5">0.50 mm</quantity>
, PW
<quantity id="3E45129FF272142D70DDECF250432ED7" box="[1023,1126,1718,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="27" pageNumber="28" unit="mm" value="1.15">1.15 mm</quantity>
, EL
<quantity id="3E45129FF272142D77B8ECF351242ED7" box="[1178,1281,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" pageId="27" pageNumber="28" unit="mm" value="1.96">1.96 mm</quantity>
, EW
<quantity id="3E45129FF272142D761DECF351B22ED7" box="[1343,1431,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="28" unit="mm" value="1.6">1.6 mm</quantity>
, TL/EW 1.47, PW/PL 2.30, EL/EW 1.23, EL/PL 3.92.
</paragraph>
<paragraph id="F902BF7AF272142D73E5ECBA55762F3B" blockId="27.[151,1437,1539,2033]" pageId="27" pageNumber="28">
<emphasis id="CBC96368F272142D73E5ECBA55682F1F" bold="true" box="[199,333,1790,1817]" pageId="27" pageNumber="28">Variability.</emphasis>
Sternite IX of the specimen from the collection of NMPC has a more rounded apical plate and longer apodeme.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F272142D73E5ED02571D2F67" box="[199,824,1862,1889]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="F902BF7AF272142D73E5ED02571D2F67" blockId="27.[151,1437,1539,2033]" box="[199,824,1862,1889]" pageId="27" pageNumber="28">
<emphasis id="CBC96368F272142D73E5ED0255442F67" bold="true" box="[199,353,1862,1889]" pageId="27" pageNumber="28">Distribution.</emphasis>
Known from three localities in SE
<collectingCountry id="81AAFFEAF272142D71CCED0357112F67" box="[750,820,1863,1889]" name="Brazil" pageId="27" pageNumber="28">Brasil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F272142A73E5ED2F506328D1" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="discussion">
<paragraph id="F902BF7AF272142A73E5ED2F506328D1" blockId="27.[151,1437,1539,2033]" lastBlockId="28.[151,1436,153,215]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="CBC96368F272142D73E5ED2F551E2F83" bold="true" box="[199,315,1899,1925]" pageId="27" pageNumber="28">Remarks.</emphasis>
The species is excluded from
<taxonomicName id="3EBDC4F9F272142D71BEED2F57FB2F83" authority="Redtenbacher" authorityName="Redtenbacher" authorityYear="1858" box="[668,990,1899,1925]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="CBC96368F272142D71BEED2F571E2F83" box="[668,827,1899,1925]" italics="true" pageId="27" pageNumber="28">Prionocyphon</emphasis>
Redtenbacher
</taxonomicName>
as it lacks diagnostic characters of the genus. In
<taxonomicName id="3EBDC4F9F272142D7221EDCB55872FAF" authorityName="Redtenbacher" authorityYear="1858" box="[259,418,1935,1961]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="CBC96368F272142D7221EDCB55872FAF" box="[259,418,1935,1961]" italics="true" pageId="27" pageNumber="28">Prionocyphon</emphasis>
</taxonomicName>
apical labial palpomere arises from side of preapical one (in
<taxonomicName id="3EBDC4F9F272142D7764EDCB50F02FAF" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1094,1237,1935,1961]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F272142D7764EDCB50F02FAF" box="[1094,1237,1935,1961]" italics="true" pageId="27" pageNumber="28">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F272142D77FEEDCB51642FAF" box="[1244,1345,1935,1961]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
it arises from apex), antennomere 3 equal to or shorter than antennomere 2, antennomeres 2 and 3 are together shorter than antennomere 4 (in
<taxonomicName id="3EBDC4F9F272142D7250ED9356242FF7" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[370,513,2007,2033]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F272142D7250ED9356242FF7" box="[370,513,2007,2033]" italics="true" pageId="27" pageNumber="28">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F272142D7129ED9356502FF7" box="[523,629,2007,2033]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
antennomeres 2 and 3 are together longer than antennomere 4) (
<bibRefCitation id="9D2CC28BF272142A7647ED9354F628B5" author="Ruta, R." lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" pagination="341 - 350" refId="ref23240" refString="Ruta, R. (2010) Revision of the genus Calvarium Pic, 1918 (Coleoptera: Scirtidae). Part 1. Redescription of the genus and catalogue of described taxa. Annales Zoologici, 60 (3), 341 - 350." type="journal article" year="2010">Ruta 2010</bibRefCitation>
;
<bibRefCitation id="9D2CC28BF275142A73FEEADD557A28B5" author="Watts, C. H. S." box="[220,351,153,179]" pageId="28" pageNumber="29" pagination="53 - 88" refId="ref23286" refString="Watts, C. H. S. (2010) Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134 (1), 53 - 88. https: // doi. org / 10.1080 / 3721426.2010.10887131" type="journal article" year="2010">Watts 2010</bibRefCitation>
). Features of male and female genitalia of
<taxonomicName id="3EBDC4F9F275142A7011EADD57E728B5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[819,962,153,179]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F275142A7011EADD57E728B5" box="[819,962,153,179]" italics="true" pageId="28" pageNumber="29">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F275142A70EAEADD501528B5" box="[968,1072,153,179]" pageId="28" pageNumber="29" rank="genus">gen. nov.</taxonomicNameLabel>
, i.e. modifications of tergite VIII and shape of bursal sclerite, differ considerably from those known in
<taxonomicName id="3EBDC4F9F275142A7082EAF9501A28D1" authorityName="Redtenbacher" authorityYear="1858" box="[928,1087,189,215]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="CBC96368F275142A7082EAF9501A28D1" box="[928,1087,189,215]" italics="true" pageId="28" pageNumber="29">Prionocyphon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

161
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<mods:title id="E06A13E20B0E05DF3BDBE99617285586">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F274142B73B5E86156252A39" box="[151,512,549,576]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="F902BF7AF274142B73B5E86156252A39" blockId="29.[151,512,549,611]" box="[151,512,549,576]" pageId="29" pageNumber="30">
<heading id="A24A0816F274142B73B5E86156252A39" bold="true" box="[151,512,549,576]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="CBC96368F274142B73B5E86156252A39" bold="true" box="[151,512,549,576]" pageId="29" pageNumber="30">
<taxonomicName id="3EBDC4F9F274142B73B5E86155BE2A46" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,411,549,576]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kawense" status="sp. nov.">
<emphasis id="CBC96368F274142B73B5E86155BE2A46" bold="true" box="[151,411,549,576]" italics="true" pageId="29" pageNumber="30">Calvariopsis kawense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F274142B7280E86156252A39" box="[418,512,549,575]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F274142B73B5E80D55082A65" box="[151,301,585,611]" pageId="29" pageNumber="30" type="description">
<paragraph id="F902BF7AF274142B73B5E80D55082A65" blockId="29.[151,512,549,611]" box="[151,301,585,611]" pageId="29" pageNumber="30">
(
<figureCitation id="6186A3FFF274142B73BDE80D54D92A65" box="[159,252,585,611]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="29" pageNumber="30">Figs 1U</figureCitation>
,
<figureCitation id="6186A3FFF274142B722AE80D55002A65" box="[264,293,585,611]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="29" pageNumber="30">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F274142B73B5E8D554EA2AF5" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="F902BF7AF274142B73B5E8D554EA2AF5" blockId="29.[151,1436,656,863]" pageId="29" pageNumber="30">
<emphasis id="CBC96368F274142B73B5E8D5556D2AAD" bold="true" box="[151,328,656,683]" pageId="29" pageNumber="30">Type material.</emphasis>
Holotype, male (MNHG): “Guyane Francaise \ Montagne Kaw \ Fourgassié \ 15.9.–1.10.1995 \ M. Kocian lgt.”. Paratype, male (MNHG): “Guyane Francaise \ Mont Grand Matoury \ 2.–11.10.1995 \ M. Kocian lgt.”.
</paragraph>
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<paragraph id="F902BF7AF274142B73E5E8B856E12FD3" blockId="29.[151,1436,656,863]" lastBlockId="29.[151,1436,1689,2041]" pageId="29" pageNumber="30">
<subSubSection id="B1A7ECF1F274142B73E5E8B857322B59" pageId="29" pageNumber="30" type="diagnosis">
<emphasis id="CBC96368F274142B73E5E8B855672B11" bold="true" box="[199,322,764,791]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
Small (TL 1.9–2.0 mm), round, blackish species, externally resembling
<emphasis id="CBC96368F274142B77A2E8B9511D2B11" box="[1152,1336,765,791]" italics="true" pageId="29" pageNumber="30">C. bituberculata</emphasis>
<emphasis id="CBC96368F274142B7663E8B951B92B11" bold="true" box="[1345,1436,765,791]" pageId="29" pageNumber="30">sp. nov.</emphasis>
Tegmen with basal apodemes, tergite VIII with T-shaped emargination flanked with two denticles. The shape of the emargination allows certain identification of this species.
</subSubSection>
<subSubSection id="B1A7ECF1F274142B73E5ECDD56E12FD3" pageId="29" pageNumber="30" type="description">
<emphasis id="CBC96368F274142B73E5ECDD557C2EB2" bold="true" box="[199,345,1689,1716]" pageId="29" pageNumber="30">Description.</emphasis>
Male. Body oval, convex, covered with whitish, suberect setae. Body brown, legs, antennae and mouthparts yellowish. Head broad, 1.6× wider than width of interocular space, with subtle punctation, punctures small, not granulate, separated by a 3.0–4.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, not granulate punctation, punctures separated by ca. 3.0–4.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles subtly obtuse. Scutellar shield subtriangular, very finely punctate. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</subSubSection>
</paragraph>
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<paragraph id="F902BF7AF274142B73B5EC6B55282E6D" blockId="29.[151,1436,1583,1644]" pageId="29" pageNumber="30">
<emphasis id="CBC96368F274142B73B5EC6B55052E4E" bold="true" box="[151,288,1583,1608]" pageId="29" pageNumber="30">FIGURE 23.</emphasis>
<emphasis id="CBC96368F274142B7207EC7456262E4E" box="[293,515,1584,1608]" italics="true" pageId="29" pageNumber="30">Calvariopsis kawense</emphasis>
<emphasis id="CBC96368F274142B712BEC74567C2E4E" bold="true" box="[521,601,1584,1608]" pageId="29" pageNumber="30">sp. nov.</emphasis>
, male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
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Male genitalia. Penis (L
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,
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W
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</geoCoordinate>
mm,
<figureCitation id="6186A3FFF274142B71DDED9A574D2FFE" box="[767,872,2014,2040]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="29" pageNumber="30">Fig. 23A</figureCitation>
) relatively wide, with hooked apical portions of parameroids and subtriangular trigonium. Tegmen (L
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,
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W
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</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27714287720EADD504228B5" box="[1026,1127,153,179]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="30" pageNumber="31">Fig. 23B</figureCitation>
) U-shaped, with apodemes in basal portion, apices of parameres with long setae. Sternite IX (L
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,
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W
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mm,
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) subtriangular, with single apodeme and numerous setae on apical margin. Tergite VIII (L
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,
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W
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mm,
<figureCitation id="6186A3FFF277142877F7EAA5511D28FD" box="[1237,1336,225,251]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="30" pageNumber="31">Fig. 23D</figureCitation>
) with Ushaped emargination of central portion of basal margin, lateral portions with short setae, apodemes as long as apical plate, diverging. Tergite IX (L
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,
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W
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mm,
<figureCitation id="6186A3FFF277142871C4EB6D57622945" box="[742,839,297,323]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="30" pageNumber="31">Fig. 23E</figureCitation>
) with membranous and relatively narrow apical plate, apodemes as long as apical plate, diverging.
</paragraph>
<paragraph id="F902BF7AF277142873E5EB3555BB298D" blockId="30.[151,1437,153,575]" box="[199,414,369,395]" pageId="30" pageNumber="31">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF277142873E5EBD1568129F1" blockId="30.[151,1437,153,575]" pageId="30" pageNumber="31">
<emphasis id="CBC96368F277142873E5EBD155DA29A9" bold="true" box="[199,511,404,431]" pageId="30" pageNumber="31">Measurements and ratios.</emphasis>
Males (n = 2) TL 1.90–1.97 (1.93) mm, PL 0.40–0.43 (0.42) mm, PW 0.93–0.97 (0.95) mm, EL 1.63–1.70 (1.67) mm, EW 1.30 (1.30) mm, TL/EW 1.46–1.51 (1.49), PW/PL 2.15–2.42 (2.29), EL/ EW 1.26–1.31 (1.28), EL/PL 3.77–4.25 (4.01).
</paragraph>
<subSubSection id="B1A7ECF1F277142873E5E84450152A1D" box="[199,1072,512,539]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="F902BF7AF277142873E5E84450152A1D" blockId="30.[151,1437,153,575]" box="[199,1072,512,539]" pageId="30" pageNumber="31">
<emphasis id="CBC96368F277142873E5E84455442A1D" bold="true" box="[199,353,512,539]" pageId="30" pageNumber="31">Distribution.</emphasis>
Known from Mt Kaw and Mt Grand Maoury in
<collectingCountry id="81AAFFEAF277142870A0E84550092A1D" box="[898,1068,513,539]" name="French Guiana" pageId="30" pageNumber="31">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F277142873E5E861504D2A39" box="[199,1128,549,575]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="F902BF7AF277142873E5E861504D2A39" blockId="30.[151,1437,153,575]" box="[199,1128,549,575]" pageId="30" pageNumber="31">
<emphasis id="CBC96368F277142873E5E861556B2A39" bold="true" box="[199,334,549,575]" pageId="30" pageNumber="31">Etymology.</emphasis>
Named after Mt Kaw, where the
<typeStatus id="260601D8F277142871E6E86156D12A39" box="[708,756,549,575]" pageId="30" pageNumber="31">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
</treatment>
</document>

198
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<mods:title id="67244188B6D4F13692E98417DC335E40">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<heading id="A24A0816F275142A73B5EB6D563F2945" bold="true" box="[151,538,297,324]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="CBC96368F275142A73B5EB6D563F2945" bold="true" box="[151,538,297,324]" pageId="28" pageNumber="29">
<taxonomicName id="3EBDC4F9F275142A73B5EB6D55902945" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,437,297,324]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
<emphasis id="CBC96368F275142A73B5EB6D55902945" bold="true" box="[151,437,297,324]" italics="true" pageId="28" pageNumber="29">Calvariopsis guyanense</emphasis>
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</heading>
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</subSubSection>
<subSubSection id="B1A7ECF1F275142A73B5EB09550D2961" box="[151,296,333,359]" pageId="28" pageNumber="29" type="description">
<paragraph id="F902BF7AF275142A73B5EB09550D2961" blockId="28.[151,538,297,359]" box="[151,296,333,359]" pageId="28" pageNumber="29">
(
<figureCitation id="6186A3FFF275142A73BDEB0954D22961" box="[159,247,333,359]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="28" pageNumber="29">Figs 1T</figureCitation>
,
<figureCitation id="6186A3FFF275142A7221EB0955052961" box="[259,288,333,359]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="28" pageNumber="29">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F275142A73B5EBD155B829D5" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="F902BF7AF275142A73B5EBD155B829D5" blockId="28.[151,1437,404,899]" pageId="28" pageNumber="29">
<emphasis id="CBC96368F275142A73B5EBD1556329A9" bold="true" box="[151,326,404,431]" pageId="28" pageNumber="29">Type material.</emphasis>
<materialsCitation id="49D5B527F275142A726CEBD155B829D5" ID-GBIF-Occurrence="2239706654" collectionCode="NHM" country="Guyana" location="Kurupukari" municipality="Malaise" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F275142A726CEBD1559C29A9" box="[334,441,405,431]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF275142A712EEBD1567129A9" box="[524,596,405,431]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="28" pageNumber="29" type="Herbarium">NHM</collectionCode>
): “
<collectingCountry id="81AAFFEAF275142A7153EBD156CC29A9" box="[625,745,405,431]" name="Guyana" pageId="28" pageNumber="29">GUYANA</collectingCountry>
ix-xi.1992 \
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\ 4°40’ N: 58°40’ W \
<collectingMunicipality id="19662500F275142A7620EBD1517F29A9" box="[1282,1370,405,431]" pageId="28" pageNumber="29">Malaise</collectingMunicipality>
/FIT \ BMNH{E} 2006-132”.
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</paragraph>
</subSubSection>
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<paragraph id="F902BF7AF275142A73E5EB9857802A39" blockId="28.[151,1437,404,899]" pageId="28" pageNumber="29">
<emphasis id="CBC96368F275142A73E5EB98556729F1" bold="true" box="[199,322,476,503]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
Small (TL 2.0 mm), oval, brown. Tegmen with basal apodemes, tergite VIII with V-shaped emar- gination flanked with two denticles. Male genital features resemble those of
<taxonomicName id="3EBDC4F9F275142A70D7E84550F42A1A" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[1013,1233,513,540]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F275142A70D7E84550B42A1D" box="[1013,1169,513,539]" italics="true" pageId="28" pageNumber="29">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
, but emargination of central portion of tergite VIII is shallower in
<taxonomicName id="3EBDC4F9F275142A718CE86157602A39" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[686,837,549,575]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
<emphasis id="CBC96368F275142A718CE86157602A39" box="[686,837,549,575]" italics="true" pageId="28" pageNumber="29">C. guyanense</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="D0FADE13F275142A706EE86157802A39" box="[844,933,549,575]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F275142B73E5E80C56CB298D" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
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<emphasis id="CBC96368F275142A73E5E80C557C2A65" bold="true" box="[199,345,584,611]" pageId="28" pageNumber="29">Description.</emphasis>
Male. Body oval, convex, covered with whitish, suberect setae. Dorsum brown, posterior portion of pronotum and scutellar shield slightly darker, venter brown, legs and antennae yellowish. Head very broad, 1.8× wider than width of interocular space, with fine punctation, punctures very small, separated by a 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
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<paragraph id="F902BF7AF275142A73B5ECBE55112F30" blockId="28.[151,1436,1786,1847]" pageId="28" pageNumber="29">
<emphasis id="CBC96368F275142A73B5ECBE55052F15" bold="true" box="[151,288,1786,1811]" pageId="28" pageNumber="29">FIGURE 22.</emphasis>
<taxonomicName id="3EBDC4F9F275142A7205ECBE563F2F14" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[295,538,1786,1810]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
<emphasis id="CBC96368F275142A7205ECBE563F2F14" box="[295,538,1786,1810]" italics="true" pageId="28" pageNumber="29">Calvariopsis guyanense</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="D0FADE13F275142A7102ECBF56542F15" box="[544,625,1787,1811]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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, male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
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Male genitalia. Penis (L
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,
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W
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mm,
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) narrow, with curved sides, parameroids short, curved, with subtriangular apices, trigonium subtriangular, short. Tegmen (L
<quantity id="3E45129FF275142A708EEDCC50312FA5" box="[940,1044,1928,1955]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="28" pageNumber="29" unit="mm" value="0.35">0.35 mm</quantity>
,
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W
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mm,
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) U-shaped, with apodemes in basal portion, apices of parameres rounded, not setose. Sternite IX (L
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,
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W
<quantity id="3E45129FF275142A7627EDE951132FC1" box="[1285,1334,1965,1991]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="mm" value="0.12">0.12</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF275142A7655EDE954E22FED" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="28" pageNumber="29">Fig 22B</figureCitation>
) subtriangular, with single, relatively long apodeme, apical margin rounded, covered with short, sparse setae. Tergite VIII (L
<quantity id="3E45129FF274142B7268EADD559728B5" box="[330,434,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="29" pageNumber="30" unit="mm" value="0.31">0.31 mm</quantity>
,
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W
<quantity id="3E45129FF274142B72C0EADD563628B5" box="[482,531,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="29" pageNumber="30" unit="mm" value="0.28">0.28</quantity>
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mm,
<figureCitation id="6186A3FFF274142B717AEADD569228B5" box="[600,695,153,179]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="29" pageNumber="30">Fig 22C</figureCitation>
) with shallow, V-shaped emargination of apical margin, and two distinct denticles in lateral portions of apical margin, apical margin not setose, apodemes as long as apical plate. Tergite IX (L
<quantity id="3E45129FF274142B7217EAA555B828FD" box="[309,413,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="mm" value="0.26">0.26 mm</quantity>
,
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W
<quantity id="3E45129FF274142B72E9EAA555D928FD" box="[459,508,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="29" pageNumber="30" unit="mm" value="0.24">0.24</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF274142B711DEAA556BA28FD" box="[575,671,225,251]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="29" pageNumber="30">Fig 22D</figureCitation>
) with membranous and relatively narrow apical plate, apodemes as long as apical plate, strongly diverging.
</paragraph>
<paragraph id="F902BF7AF274142B73E5EB6D55BB2945" blockId="29.[151,1437,153,467]" box="[199,414,297,323]" pageId="29" pageNumber="30">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF274142B73E5EB0956CB298D" blockId="29.[151,1437,153,467]" pageId="29" pageNumber="30">
<emphasis id="CBC96368F274142B73E5EB0955D82961" bold="true" box="[199,509,332,359]" pageId="29" pageNumber="30">Measurements and ratios.</emphasis>
Male (n = 1) TL 2.00 mm, PL
<quantity id="3E45129FF274142B7077EB09579E2961" box="[853,955,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="mm" value="0.4">0.40 mm</quantity>
, PW
<quantity id="3E45129FF274142B70D4EB0950792961" box="[1014,1116,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="29" pageNumber="30" unit="mm" value="1.03">1.03 mm</quantity>
, EL
<quantity id="3E45129FF274142B77ADEB0950D02961" box="[1167,1269,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="29" pageNumber="30" unit="mm" value="1.67">1.67 mm</quantity>
, EW
<quantity id="3E45129FF274142B7610EB0951BD2961" box="[1330,1432,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="mm" value="1.4">1.40 mm</quantity>
, TL/EW 1.43, PW/PL 2.58, EL/EW 1.19, EL/PL 4.17.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F274142B73E5EBD0574B29A9" box="[199,878,404,431]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="F902BF7AF274142B73E5EBD0574B29A9" blockId="29.[151,1437,153,467]" box="[199,878,404,431]" pageId="29" pageNumber="30">
<emphasis id="CBC96368F274142B73E5EBD0554429A9" bold="true" box="[199,353,404,431]" pageId="29" pageNumber="30">Distribution.</emphasis>
Known only from the
<typeStatus id="260601D8F274142B7142EBD156B529A9" box="[608,656,405,431]" pageId="29" pageNumber="30">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF274142B702DEBD1574F29A9" box="[783,874,405,431]" name="Guyana" pageId="29" pageNumber="30">Guyana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F274142B73E5EBFD504129D5" box="[199,1124,441,467]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="F902BF7AF274142B73E5EBFD504129D5" blockId="29.[151,1437,153,467]" box="[199,1124,441,467]" pageId="29" pageNumber="30">
<emphasis id="CBC96368F274142B73E5EBFD556B29D5" bold="true" box="[199,334,441,467]" pageId="29" pageNumber="30">Etymology.</emphasis>
Named after
<collectingCountry id="81AAFFEAF274142B72C5EBFD566429D5" box="[487,577,441,467]" name="Guyana" pageId="29" pageNumber="30">Guyana</collectingCountry>
, where the
<typeStatus id="260601D8F274142B71E3EBFD56D429D5" box="[705,753,441,467]" pageId="29" pageNumber="30">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
</treatment>
</document>

187
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<heading id="A24A0816F276142973B5E92D560B2B85" bold="true" box="[151,558,873,900]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="CBC96368F276142973B5E92D560B2B85" bold="true" box="[151,558,873,900]" pageId="31" pageNumber="32">
<taxonomicName id="3EBDC4F9F276142973B5E92D55ED2B82" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,456,873,900]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="saopaulense" status="sp. nov.">
<emphasis id="CBC96368F276142973B5E92D55ED2B82" bold="true" box="[151,456,873,900]" italics="true" pageId="31" pageNumber="32">Calvariopsis saopaulense</emphasis>
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</subSubSection>
<subSubSection id="B1A7ECF1F276142973B5E9C955142BA1" box="[151,305,908,935]" pageId="31" pageNumber="32" type="description">
<paragraph id="F902BF7AF276142973B5E9C955142BA1" blockId="31.[151,558,873,935]" box="[151,305,908,935]" pageId="31" pageNumber="32">
(
<figureCitation id="6186A3FFF276142973BDE9C955252BA1" box="[159,256,909,935]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="31" pageNumber="32">Figs 1W</figureCitation>
,
<figureCitation id="6186A3FFF2761429722EE9C8550C2BA1" box="[268,297,908,935]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="31" pageNumber="32">25</figureCitation>
)
</paragraph>
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<subSubSection id="B1A7ECF1F276142973B5E99156612C15" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="F902BF7AF276142973B5E99156612C15" blockId="31.[151,1437,980,2015]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973B5E99155622BE9" bold="true" box="[151,327,980,1007]" pageId="31" pageNumber="32">Type material.</emphasis>
<materialsCitation id="49D5B527F2761429726DE99151022BE9" ID-GBIF-Occurrence="2239706650" box="[335,1319,980,1007]" collectionCode="NMPC" location="SAO PAULO" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="260601D8F2761429726DE991559F2BE9" box="[335,442,981,1007]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
, male (
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): “
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\ BRAS MRÁZ LGT. \ MUS. PRAGENSE”
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.
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<typeStatus id="260601D8F27614297611E99151BD2BE9" box="[1331,1432,981,1007]" pageId="31" pageNumber="32" type="paratype">Paratype</typeStatus>
, male (
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): same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F276142973E5EE5856342CA5" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="F902BF7AF276142973E5EE5856342CA5" blockId="31.[151,1437,980,2015]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973E5EE5855672C31" bold="true" box="[199,322,1052,1079]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
Small (TL
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), blackish, widest in anterior portion of elytra, externally resembling
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<emphasis id="CBC96368F27614297672EE5954F72C5D" italics="true" pageId="31" pageNumber="32">C. wittmeri</emphasis>
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</emphasis>
Identification is possible only on the basis of morphology of male genitalia: trigonium of penis very narrow, tegmen V-shaped (unique in known
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<emphasis id="CBC96368F276142971A7EE2157312C79" box="[645,788,1125,1151]" italics="true" pageId="31" pageNumber="32">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F27614297039EE21575A2C79" box="[795,895,1125,1151]" pageId="31" pageNumber="32" rank="genus">gen. nov</taxonomicNameLabel>
,), without basal apodemes, apical plate of tergite VIII with V-shaped emargination.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F276142973E5EEE856EA2F91" pageId="31" pageNumber="32" type="description">
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<emphasis id="CBC96368F276142973E5EEE8557C2CC1" bold="true" box="[199,345,1196,1223]" pageId="31" pageNumber="32">Description.</emphasis>
Male. Body oblong oval, convex, covered with brownish, suberect setae. Body brown, legs, antennae and mouthparts yellowish. Head very broad, 1.7× wider than width of interocular space, with fine punctation, punctures small, granulate, separated by a 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest slightly before the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation much stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF276142973E5EFB5552F2F01" blockId="31.[151,1437,980,2015]" pageId="31" pageNumber="32">
Male genitalia. Penis (L
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,
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W
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mm,
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) moderately wide, with slightly curved sides, param- eroids long, subtriangular, gradually narrowing, pointed at apices, trigonium very narrow, subtriangular. Tegmen (L
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,
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W
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mm,
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) V-shaped, apices of parameres pointed, covered with setae. Sternite IX (L
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,
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W
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mm,
<figureCitation id="6186A3FFF2761429720CEC1955B12E71" box="[302,404,1628,1655]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="31" pageNumber="32">Fig. 25B</figureCitation>
) small, subrectangular, with single, short apodeme, apical portion of sternite covered with numerous setae. Tergite VIII (L
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,
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W
<quantity id="3E45129FF276142971B8ECC456EE2E9D" box="[666,715,1664,1691]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="32" unit="mm" value="0.25">0.25</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2761429702FECC557572E9D" box="[781,882,1664,1691]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="31" pageNumber="32">Fig. 25C</figureCitation>
) with V-shaped emargination of anterior margin, anterior margin with small denticles and small setae, apodemes as long as apical plate. Tergite IX (L
<quantity id="3E45129FF2761429762DECE151532EB9" box="[1295,1398,1701,1727]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="31" pageNumber="32" unit="mm" value="0.23">0.23 mm</quantity>
,
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W
<quantity id="3E45129FF276142973B5EC8D54ED2EE5" box="[151,200,1737,1763]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="31" pageNumber="32" unit="mm" value="0.21">0.21</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27614297225EC8D554E2EE5" box="[263,363,1736,1763]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="31" pageNumber="32">Fig. 25D</figureCitation>
) with membranous and relatively narrow apical plate, apodemes as long as apical plate, strongly diverging.
</paragraph>
<paragraph id="F902BF7AF276142973E5ED5555BB2F2D" blockId="31.[151,1437,980,2015]" box="[199,414,1809,1835]" pageId="31" pageNumber="32">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF276142973E5ED7156EA2F91" blockId="31.[151,1437,980,2015]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973E5ED7155DA2F49" bold="true" box="[199,511,1844,1871]" pageId="31" pageNumber="32">Measurements and ratios.</emphasis>
Males (n = 2) TL 1.93–2.07 (2.00) mm, PL 0.40–0.43 (0.42) mm, PW 0.90–0.97 (0.93) mm, EL 1.73–1.83 (1.78) mm, EW 1.23–1.33 (1.28) mm, TL/EW 1.55–1.57 (1.56), PW/PL 2.23–2.25 (2.24), EL/EW 1.38–1.41 (1.39), EL/PL 4.23–4.33 (4.28).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F276142973E5EDE457FF2FBD" box="[199,986,1952,1979]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="F902BF7AF276142973E5EDE457FF2FBD" blockId="31.[151,1437,980,2015]" box="[199,986,1952,1979]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973E5EDE455442FBD" bold="true" box="[199,353,1952,1979]" pageId="31" pageNumber="32">Distribution.</emphasis>
Known only from the
<typeStatus id="260601D8F27614297142EDE556B52FBD" box="[608,656,1953,1979]" pageId="31" pageNumber="32">type</typeStatus>
locality in
<collectingRegion id="3B797198F2761429702DEDE557A62FBD" box="[783,899,1953,1979]" country="Brazil" name="Sao Paulo" pageId="31" pageNumber="32">Sao Paulo</collectingRegion>
,
<collectingCountry id="81AAFFEAF276142970ACEDE557F32FBD" box="[910,982,1953,1979]" name="Brazil" pageId="31" pageNumber="32">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F276142973E5ED81505B2FD9" box="[199,1150,1989,2015]" pageId="31" pageNumber="32" type="etymology">
<paragraph id="F902BF7AF276142973E5ED81505B2FD9" blockId="31.[151,1437,980,2015]" box="[199,1150,1989,2015]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973E5ED81556B2FD9" bold="true" box="[199,334,1989,2015]" pageId="31" pageNumber="32">Etymology.</emphasis>
Named after
<collectingRegion id="3B797198F276142972C5ED81567E2FD9" box="[487,603,1989,2015]" country="Brazil" name="Sao Paulo" pageId="31" pageNumber="32">Sao Paulo</collectingRegion>
, where the
<typeStatus id="260601D8F276142971F8ED81572F2FD9" box="[730,778,1989,2015]" pageId="31" pageNumber="32">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
</treatment>
</document>

206
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(
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Figs
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,
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)
</paragraph>
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<emphasis id="CBC96368F277142873B5E8B955602B11" bold="true" box="[151,325,764,791]" pageId="30" pageNumber="31">Type material.</emphasis>
<materialsCitation id="49D5B527F2771428726EE8B9502D2B11" ID-GBIF-Occurrence="2239706651" box="[332,1032,765,791]" collectionCode="CMNH" country="Brazil" location="Brazil" pageId="30" pageNumber="31" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F2771428726EE8B955922B11" box="[332,439,765,791]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
, male (
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): “Taperina \
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\ Acc. No. 2966”.
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<subSubSection id="B1A7ECF1F277142873E5E964557A2BA1" pageId="30" pageNumber="31" type="diagnosis">
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<emphasis id="CBC96368F277142873E5E96455672B3D" bold="true" box="[199,322,800,827]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
The smallest known species of
<taxonomicName id="3EBDC4F9F2771428719BE965576D2B3D" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[697,840,801,827]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F2771428719BE965576D2B3D" box="[697,840,801,827]" italics="true" pageId="30" pageNumber="31">Calvariopsis</emphasis>
</taxonomicName>
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(TL
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), round, blackish, externally resembling
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<emphasis id="CBC96368F27714287239E90155F72B59" box="[283,466,837,863]" italics="true" pageId="30" pageNumber="31">C. bituberculata</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="D0FADE13F277142872F8E90156112B59" box="[474,564,837,863]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Tergite VIII similar to that of
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<emphasis id="CBC96368F277142870A9E90150022B59" box="[907,1063,837,863]" italics="true" pageId="30" pageNumber="31">C. brasiliense</emphasis>
(Pic)
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and
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<emphasis id="CBC96368F277142877BEE90151112B59" box="[1180,1332,837,863]" italics="true" pageId="30" pageNumber="31">C. guyanense</emphasis>
</taxonomicName>
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, apical plate with shallow V-shaped emargination flanked with two spine-like processes, covered with setae on posterolateral angles.
</paragraph>
</subSubSection>
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<emphasis id="CBC96368F277142873B5ECA255052EF9" bold="true" box="[151,288,1766,1791]" pageId="30" pageNumber="31">FIGURE 24.</emphasis>
<taxonomicName id="3EBDC4F9F27714287204ECA255C42EF8" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[294,481,1766,1790]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="CBC96368F27714287204ECA255C42EF8" box="[294,481,1766,1790]" italics="true" pageId="30" pageNumber="31">Calvariopsis nana</emphasis>
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<emphasis id="CBC96368F277142872C5ECA356122EF9" bold="true" box="[487,567,1767,1791]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="D0FADE13F277142872C5ECA356122EF9" box="[487,567,1767,1791]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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, male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm.
</paragraph>
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<subSubSection id="B1A7ECF1F277142973E5ED6856CB2A39" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
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<emphasis id="CBC96368F277142873E5ED68557C2F41" bold="true" box="[199,345,1836,1863]" pageId="30" pageNumber="31">Description.</emphasis>
Male. Body oval, convex, covered with whitish, suberect setae. Body brown, legs, antennae and mouthparts yellowish. Head very broad, 1.8× wider than width of interocular space, with fine punctation, punctures small, granulate, separated by a 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong (left elytron missing in the studied specimen), without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
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Male genitalia. Penis (L
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,
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W
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mm,
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) with curved sides, parameroids short, subtriangular, trigonium subtriangular. Tegmen and sternite IX missing in the studied specimen. Tergite VIII (L
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,
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W
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mm,
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) with shallow V-shaped emargination of apical margin, two sharp denticles in lateral portions of apical plate and short setae on posterolateral portion, apodemes as long as apical plate, slightly diverging. Tergite IX (L
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,
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W
<quantity id="3E45129FF27614297208EBD1557E29A9" box="[298,347,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="31" pageNumber="32" unit="mm" value="0.19">0.19</quantity>
</geoCoordinate>
mm,
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) with membranous apical plate, apical margin covered with short setae, apodemes as long as apical plate, diverging.
</paragraph>
<paragraph id="F902BF7AF276142973E5EB9955BB29F1" blockId="31.[151,1437,153,791]" box="[199,414,477,503]" pageId="31" pageNumber="32">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF276142973E5E84556CB2A39" blockId="31.[151,1437,153,791]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973E5E84555D82A1D" bold="true" box="[199,509,512,539]" pageId="31" pageNumber="32">Measurements and ratios.</emphasis>
Male (n = 1) TL
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, PL
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, PW
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, EL
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, EW
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, TL/EW 1.53, PW/PL 2.25, EL/EW 1.32, EL/PL 3.75.
</paragraph>
</subSubSection>
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<emphasis id="CBC96368F276142973E5E80C55442A65" bold="true" box="[199,353,584,611]" pageId="31" pageNumber="32">Distribution.</emphasis>
Known only from the
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locality in
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.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F276142973E5E82950392A81" box="[199,1052,621,647]" pageId="31" pageNumber="32" type="etymology">
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<emphasis id="CBC96368F276142973E5E829556B2A81" bold="true" box="[199,334,621,647]" pageId="31" pageNumber="32">Etymology.</emphasis>
Latin
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<emphasis id="CBC96368F276142972B5E82A55EA2A81" box="[407,463,622,647]" italics="true" pageId="31" pageNumber="32">nana</emphasis>
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= dwarf; a reference to the small size of the species.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F276142973E5E8D555C42B11" pageId="31" pageNumber="32" type="discussion">
<paragraph id="F902BF7AF276142973E5E8D555C42B11" blockId="31.[151,1437,153,791]" pageId="31" pageNumber="32">
<emphasis id="CBC96368F276142973E5E8D5551E2AAD" bold="true" box="[199,315,657,683]" pageId="31" pageNumber="32">Remarks.</emphasis>
According to
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, the specimen is part of a collection bought from Herbert H. Smith in 1906 and probably also collected by him. Taperina is alternatively spelled Taperinha; this is a plantation on the Amazon,
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downstream from Santarém. The specimen was probably collected in the 1870s, as Smith began exploration of the Amazon basin in 1874.
</paragraph>
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</treatment>
</document>

244
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</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F279142673B5E8F155682AC9" box="[151,333,693,719]" pageId="16" pageNumber="17" type="description">
<paragraph id="F902BF7AF279142673B5E8F155682AC9" blockId="16.[151,532,657,719]" box="[151,333,693,719]" pageId="16" pageNumber="17">
(
<figureCitation id="6186A3FFF279142673BDE8F154C42AC9" box="[159,225,693,719]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="16" pageNumber="17">Figs 1</figureCitation>
E–G, 13)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F279142673B5E8B9516E2B85" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="F902BF7AF279142673B5E8B9516E2B85" blockId="16.[151,1437,764,1763]" pageId="16" pageNumber="17">
<emphasis id="CBC96368F279142673B5E8B9556E2B11" bold="true" box="[151,331,764,791]" pageId="16" pageNumber="17">Type material.</emphasis>
<materialsCitation id="49D5B527F27914267275E8B9508A2B11" ID-GBIF-Occurrence="2239706647" box="[343,1199,764,791]" collectionCode="MNHN" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F27914267275E8B955E72B11" box="[343,450,765,791]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF2791426713CE8B9565E2B11" box="[542,635,765,791]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
): “Callanga \ Perou” [green label, handwritten]
</materialsCitation>
.
<materialsCitation id="49D5B527F279142677E2E8B951622B85" ID-GBIF-Occurrence="2239706628" collectingDate="1935-11-01" collectingDateMax="2007-11-09" collectingDateMin="1935-11-01" collectionCode="MNHN, SEMC" collectorName="D. Brzoska" country="Peru" elevation="1120" location="Male" pageId="16" pageNumber="17" specimenCount="849" specimenCount-male="9" stateProvince="Cusco" typeStatus="paratype">
<typeStatus id="260601D8F279142677E2E8B951142B11" box="[1216,1329,765,791]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="EFBB74F3F27914267662E8B951B92B11" box="[1344,1436,765,791]" pageId="16" pageNumber="17" type="male">9 males</specimenCount>
(
<collectionCode id="9FAC27BFF279142673BDE96554D92B3D" box="[159,252,801,827]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
): same data as holotype.
<location id="FC62E9A1F27914267139E96556702B3D" LSID="urn:lsid:plazi:treatment:71140E6CF279142773B5E8D550642FF1:FC62E9A1F27914267139E96556702B3D" box="[539,597,801,827]" country="Peru" name="Male" pageId="16" pageNumber="17" stateProvince="Cusco">Male</location>
(
<collectionCode id="9FAC27BFF27914267144E965569C2B3D" box="[614,697,801,827]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
): “
<collectingCountry id="81AAFFEAF279142671F5E96557002B3D" box="[727,805,801,827]" name="Peru" pageId="16" pageNumber="17">PERU</collectingCountry>
: Dept.
<collectingRegion id="3B797198F2791426705AE96557E72B3D" box="[888,962,801,827]" country="Peru" name="Cusco" pageId="16" pageNumber="17">Cusco</collectingRegion>
: \ Cock of the Rock Lodge, \ NE Paucar- tambo 13°03.3’S \ 71°32.7’W
<quantity id="3E45129FF279142672D6E90156642B59" box="[500,577,837,863]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="16" pageNumber="17" unit="m" value="1120.0">
<elevation id="72905849F279142672D6E90156642B59" box="[500,577,837,863]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="16" pageNumber="17" unit="m" value="1120.0">1120m</elevation>
</quantity>
\
<date id="8D0399BAF27914267178E90156C82B59" box="[602,749,837,863]" pageId="16" pageNumber="17" value="2007-11-04" valueMax="2007-11-09" valueMin="2007-11-04">
<collectingDate id="9D476052F27914267178E90156C82B59" box="[602,749,837,863]" pageId="16" pageNumber="17" value="2007-11-04" valueMax="2007-11-09" valueMin="2007-11-04">4–9-XI-2007</collectingDate>
</date>
<collectorName id="5448DAACF279142671D7E90157502B59" box="[757,885,837,863]" pageId="16" pageNumber="17">D. Brzoska</collectorName>
\
<collectingMethod id="20FCC76DF279142670ACE90150A82B59" box="[910,1165,837,863]" pageId="16" pageNumber="17">ex flight intercept trap</collectingMethod>
\ PER1B07 001”. Male (
<collectionCode id="9FAC27BFF279142673BDE92D54D72B85" box="[159,242,873,899]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
): “
<collectingCountry id="81AAFFEAF2791426722DE92D55782B85" box="[271,349,873,899]" name="Peru" pageId="16" pageNumber="17">PERU</collectingCountry>
: Jauja Prov. \
<collectingRegion id="3B797198F279142672DAE92D56102B85" box="[504,565,873,899]" country="Peru" name="Junin" pageId="16" pageNumber="17">Junín</collectingRegion>
Dept., Sani \ Beni,
<specimenCount id="EFBB74F3F27914267030E92D577C2B85" box="[786,857,873,899]" pageId="16" pageNumber="17" type="generic">
<quantity id="3E45129FF27914267030E92D57702B85" box="[786,853,873,899]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="16" pageNumber="17" unit="m" value="840.0">
<elevation id="72905849F27914267030E92D57702B85" box="[786,853,873,899]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="16" pageNumber="17" unit="m" value="840.0">840m</elevation>
</quantity>
.
</specimenCount>
\
<date id="8D0399BAF2791426704CE92C50712B85" box="[878,1108,872,899]" pageId="16" pageNumber="17" value="1935-11-01" valueMax="1935-11-05" valueMin="1935-11-01">
<collectingDate id="9D476052F2791426704CE92C50712B85" box="[878,1108,872,899]" pageId="16" pageNumber="17" value="1935-11-01" valueMax="1935-11-05" valueMin="1935-11-01">1–5 November 1935</collectingDate>
</date>
\ Felix Woytkowski”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F279142673E5E9C850E42BE9" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="F902BF7AF279142673E5E9C850E42BE9" blockId="16.[151,1437,764,1763]" pageId="16" pageNumber="17">
<emphasis id="CBC96368F279142673E5E9C855672BA1" bold="true" box="[199,322,908,935]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Extremely variable species of moderate size (TL
<quantity id="3E45129FF27914267058E9C850222BA1" box="[890,1031,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="16" pageNumber="17" unit="mm" value="2.85" valueMax="3.2" valueMin="2.5">2.5–3.2 mm</quantity>
), externally resembling
<taxonomicName id="3EBDC4F9F2791426763AE9C951572BA1" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1304,1394,909,935]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="picta" status="sp. nov.">
<emphasis id="CBC96368F2791426763AE9C951572BA1" box="[1304,1394,909,935]" italics="true" pageId="16" pageNumber="17">C. picta</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27914267659E9C954ED2BCD" bold="true" pageId="16" pageNumber="17">
<taxonomicNameLabel id="D0FADE13F27914267659E9C954ED2BCD" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Identifiable on the basis of male genital characters: penis very long and narrow.Apical plate of tergite VIII with distinct M-shaped notch in median portion, flanked with two deep, semicircular emarginations.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F279142773E5E9BC56EB2FB6" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="F902BF7AF279142673E5E9BC506C2D7D" blockId="16.[151,1437,764,1763]" pageId="16" pageNumber="17">
<emphasis id="CBC96368F279142673E5E9BC557C2C15" bold="true" box="[199,345,1016,1043]" pageId="16" pageNumber="17">Description.</emphasis>
<typeStatus id="260601D8F2791426727DE9BD55EF2C15" box="[351,458,1017,1043]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
, male. Body oval, convex, covered with brown, suberect setae. Head, pronotum, scutellar shield and venter dark brown, elytra brown, each elytron with three orange maculae: oval macula in humeral portion of each elytron, transverse macula in middle portion of elytra and orange adsutural macula in apical part of elytra, legs, antennae, and mouthparts yellowish-brown. Head very broad, 1.7× wider than width of interocular space, with subtle punctation, punctures small, granulate, separated by a 3.0–4.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest in middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 0.5–1.0× diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF279142673E5EFC1512B2E9D" blockId="16.[151,1437,764,1763]" pageId="16" pageNumber="17">
Male genitalia. Penis (L
<quantity id="3E45129FF279142672FCEFC056602D99" box="[478,581,1412,1439]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="17" unit="mm" value="1.05">1.05 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27914267173EFC156812D99" box="[593,676,1413,1439]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.12">
W
<quantity id="3E45129FF27914267151EFC156812D99" box="[627,676,1413,1439]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="17" unit="mm" value="0.12">0.12</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF279142671C5EFC157102D99" box="[743,821,1413,1439]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
A–B) very long and narrow, widened in basal portion, parameroids short, narrow, rounded at apices, curved, trigonium subtriangular, narrow, pointed at apex, curved in lateral view. Tegmen (L
<quantity id="3E45129FF27914267285EF8956292DE1" box="[423,524,1485,1511]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="17" unit="mm" value="0.8">0.80 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27914267135EF89564C2DE1" box="[535,617,1485,1511]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.21">
W
<quantity id="3E45129FF2791426711AEF89564C2DE1" box="[568,617,1485,1511]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="17" unit="mm" value="0.21">0.21</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2791426718BEF89572E2DE1" box="[681,779,1485,1511]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="16" pageNumber="17">Fig. 13C</figureCitation>
) very long, sides subparallel, apex with short lateral lobes, covered with setae. Sternite IX (L
<quantity id="3E45129FF27914267102EFB556A22E0D" box="[544,647,1521,1547]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="17" unit="mm" value="0.32">0.32 mm</quantity>
,
<geoCoordinate id="9C89D9BDF279142671B1EFB556C32E0D" box="[659,742,1520,1547]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.25">
W
<quantity id="3E45129FF27914267197EFB456C32E0D" box="[693,742,1520,1547]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="17" unit="mm" value="0.25">0.25</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF2791426700BEFB557AA2E0D" box="[809,911,1521,1547]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="16" pageNumber="17">Fig. 13D</figureCitation>
) subrectangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
<quantity id="3E45129FF27914267705EC5150AB2E29" box="[1063,1166,1557,1583]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="17" unit="mm" value="0.44">0.44 mm</quantity>
,
<geoCoordinate id="9C89D9BDF279142677B8EC5150CB2E29" box="[1178,1262,1557,1583]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.36">
W
<quantity id="3E45129FF2791426779FEC5150CB2E29" box="[1213,1262,1557,1583]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="16" pageNumber="17" unit="mm" value="0.36">0.36</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27914267613EC5151B12E29" box="[1329,1428,1557,1583]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="16" pageNumber="17">Fig. 13E</figureCitation>
) with short apical plate, central portion of apical margin with two oval emarginations and two short denticles forming M-shaped structure, apical margin subtly setose, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF2791426739AECC5553B2E9D" box="[184,286,1665,1691]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="17" unit="mm" value="0.43">0.43 mm</quantity>
,
<geoCoordinate id="9C89D9BDF2791426720BECC5555E2E9D" box="[297,379,1665,1691]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-0.29">
W
<quantity id="3E45129FF27914267268ECC5555E2E9D" box="[330,379,1665,1691]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="17" unit="mm" value="0.29">0.29</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27914267299ECC5563E2E9D" box="[443,539,1665,1691]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="16" pageNumber="17">Fig. 13F</figureCitation>
) with membranous apical plate, apodemes longer than apical plate.
</paragraph>
<paragraph id="F902BF7AF279142673E5ECE155872EE5" blockId="16.[151,1437,764,1763]" pageId="16" pageNumber="17">Variability. Colouration of body variable, from uniformly orange to completely black. The following colour patterns were observed:</paragraph>
<paragraph id="F902BF7AF279142673B5ED5557582F2D" blockId="16.[151,1437,1809,2016]" box="[151,893,1809,1835]" pageId="16" pageNumber="17">- head light brown, pronotum, scutellar shield and elytra orange;</paragraph>
<paragraph id="F902BF7AF279142673B5ED7155F62F72" blockId="16.[151,1437,1809,2016]" pageId="16" pageNumber="17">
- head, pronotum, and scutellar shield brownish-black, elytra orange with transverse black stripe in posterior half (Dept.
<collectingRegion id="3B797198F27914267229ED1D55712F75" box="[267,340,1881,1907]" country="Peru" name="Cusco" pageId="16" pageNumber="17">Cusco</collectingRegion>
, Callanga);
</paragraph>
<paragraph id="F902BF7AF279142673B5ED3957332F9E" blockId="16.[151,1437,1809,2016]" box="[151,790,1917,1944]" pageId="16" pageNumber="17">- as above, but whole apices of elytra black (Callanga);</paragraph>
<paragraph id="F902BF7AF279142673B5EDE557E02FE6" blockId="16.[151,1437,1809,2016]" pageId="16" pageNumber="17">- head and pronotum light brown, scutellar shield orange, elytra orange with indistinct brown pattern consisting of three elongate maculae in anterior portion of each elytron (Callanga);</paragraph>
<paragraph id="F902BF7AF278142773B5EADD57C628DE" blockId="17.[151,1436,153,396]" pageId="17" pageNumber="18">- head light brown, pronotum light brown with black central portion, elytra orange with arcuate black stripe in basal portion and transverse black stripe in posterior portion (Callanga);</paragraph>
<paragraph id="F902BF7AF278142773B5EAA555B42942" blockId="17.[151,1436,153,396]" pageId="17" pageNumber="18">- head, pronotum, and scutellar shield dark brown, elytra brown with two oval orange maculae in humeral portions, transverse orange maculae in the middle of elytra, and orange adsutural macula in apical part of each elytron (Callanga);</paragraph>
<paragraph id="F902BF7AF278142773B5EB0950722961" blockId="17.[151,1436,153,396]" box="[151,1111,333,359]" pageId="17" pageNumber="18">
- whole dorsum dark brown, except apices of elytra which are orange (Dept.
<collectingRegion id="3B797198F2781427772BEB09506E2961" box="[1033,1099,333,359]" country="Peru" name="Junin" pageId="17" pageNumber="18">Junín</collectingRegion>
);
</paragraph>
<paragraph id="F902BF7AF278142773B5EB355660298A" blockId="17.[151,1436,153,396]" box="[151,581,369,396]" pageId="17" pageNumber="18">- whole dorsum blackish (Callanga).</paragraph>
<caption id="ADC2EFF2F278142773B5ECDE57362ED1" httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1690,1715]" targetBox="[153,1435,396,1639]" targetPageId="17">
<paragraph id="F902BF7AF278142773B5ECDE57362ED1" blockId="17.[151,1436,1690,1751]" pageId="17" pageNumber="18">
<emphasis id="CBC96368F278142773B5ECDE55042EB5" bold="true" box="[151,289,1690,1715]" pageId="17" pageNumber="18">FIGURE 13.</emphasis>
<taxonomicName id="3EBDC4F9F2781427720BECDF563F2EB5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[297,538,1691,1715]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="CBC96368F2781427720BECDF563F2EB5" box="[297,538,1691,1715]" italics="true" pageId="17" pageNumber="18">Calvariopsis peruviana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27814277100ECDF56512EB5" bold="true" box="[546,628,1691,1715]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="D0FADE13F27814277100ECDF56512EB5" box="[546,628,1691,1715]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="F902BF7AF278142773E5ED6D55BB2F45" blockId="17.[151,1436,1833,2040]" box="[199,414,1833,1859]" pageId="17" pageNumber="18">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF278142773E5ED0A56EB2FB6" blockId="17.[151,1436,1833,2040]" pageId="17" pageNumber="18">
<emphasis id="CBC96368F278142773E5ED0A55DB2F6E" bold="true" box="[199,510,1869,1896]" pageId="17" pageNumber="18">Measurements and ratios.</emphasis>
Males (n = 10) TL 2.50–3.20 (2.86) mm, PL 0.47–0.67 (0.58) mm, PW 1.13–1.43 (1.32) mm, EL 2.23–2.63 (2.46) mm, EW 1.63–2.07 (1.85) mm, TL/EW 1.50–1.66 (1.55), PW/PL 2.15–2.47 (2.27), EL/EW 1.27–1.40 (1.34), EL/PL 3.80–4.79 (4.25).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F278142773E5EDFD516D2FD2" box="[199,1352,1977,2004]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="F902BF7AF278142773E5EDFD516D2FD2" blockId="17.[151,1436,1833,2040]" box="[199,1352,1977,2004]" pageId="17" pageNumber="18">
<emphasis id="CBC96368F278142773E5EDFD55442FD2" bold="true" box="[199,353,1977,2004]" pageId="17" pageNumber="18">Distribution.</emphasis>
Known from three quite distant (located
<quantity id="3E45129FF2781427700DEDFD57992FD5" box="[815,956,1977,2004]" metricMagnitude="5" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="17" pageNumber="18" unit="km" value="400.0" valueMax="500.0" valueMin="300.0">300–500 km</quantity>
from each other) localities in
<collectingCountry id="81AAFFEAF2781427762FEDFE51612FD2" box="[1293,1348,1978,2004]" name="Peru" pageId="17" pageNumber="18">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F278142773E5ED9A50642FF1" box="[199,1089,2013,2040]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="F902BF7AF278142773E5ED9A50642FF1" blockId="17.[151,1436,1833,2040]" box="[199,1089,2013,2040]" pageId="17" pageNumber="18">
<emphasis id="CBC96368F278142773E5ED9A556B2FFE" bold="true" box="[199,334,2014,2040]" pageId="17" pageNumber="18">Etymology.</emphasis>
Named after
<collectingCountry id="81AAFFEAF278142772C5ED9A56382FFE" box="[487,541,2014,2040]" name="Peru" pageId="17" pageNumber="18">Peru</collectingCountry>
, where the
<typeStatus id="260601D8F278142771BFED9A56E82FFE" box="[669,717,2014,2040]" pageId="17" pageNumber="18">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
</treatment>
</document>

242
data/71/14/0E/71140E6CF27A142273B5E80D560128FD.xml Normal file
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@ -0,0 +1,242 @@
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<mods:title id="7451A3F8F584C698887E76624D39B531">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<treatment id="71140E6CF27A142273B5E80D560128FD" ID-GBIF-Taxon="157135237" LSID="urn:lsid:plazi:treatment:71140E6CF27A142273B5E80D560128FD" httpUri="http://treatment.plazi.org/id/71140E6CF27A142273B5E80D560128FD" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="B1A7ECF1F27A142573B5E80D55CC2A65" box="[151,489,585,612]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="F902BF7AF27A142573B5E80D55CC2A65" blockId="19.[151,489,585,647]" box="[151,489,585,612]" pageId="19" pageNumber="20">
<heading id="A24A0816F27A142573B5E80D55CC2A65" bold="true" box="[151,489,585,612]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="CBC96368F27A142573B5E80D55CC2A65" bold="true" box="[151,489,585,612]" pageId="19" pageNumber="20">
<taxonomicName id="3EBDC4F9F27A142573B5E80D55A12A65" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,388,585,612]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="CBC96368F27A142573B5E80D55A12A65" bold="true" box="[151,388,585,612]" italics="true" pageId="19" pageNumber="20">Calvariopsis pittieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F27A142572AEE80D55CC2A65" box="[396,489,585,611]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27A142573B5E82955062A81" box="[151,291,620,647]" pageId="19" pageNumber="20" type="description">
<paragraph id="F902BF7AF27A142573B5E82955062A81" blockId="19.[151,489,585,647]" box="[151,291,620,647]" pageId="19" pageNumber="20">
(
<figureCitation id="6186A3FFF27A142573BDE82954D42A81" box="[159,241,621,647]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="19" pageNumber="20">
Figs
<date id="8D0399BAF27A142573FAE82954D42A81" box="[216,241,621,647]" pageId="19" pageNumber="20">1I</date>
</figureCitation>
,
<figureCitation id="6186A3FFF27A142573DFE828553F2A81" box="[253,282,620,647]" captionStart="FIGURE 15" captionStartId="20.[151,250,1137,1162]" captionTargetBox="[186,1401,251,1088]" captionTargetId="figure@20.[186,1403,272,1088]" captionTargetPageId="20" captionText="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783750/files/figure.png" pageId="19" pageNumber="20">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27A142573B5E8F154D22C15" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="F902BF7AF27A142573B5E8F154D22C15" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573B5E8F1556C2AC9" bold="true" box="[151,329,692,719]" pageId="19" pageNumber="20">Type material.</emphasis>
<materialsCitation id="49D5B527F27A14257271E8F157692B3D" ID-GBIF-Occurrence="2239706643" collectingDate="1990-05-14" collectionCode="NHM" collectorName="P. Nac. Henri Pittier &amp; J. G. Davies" country="Venezuela" county="Estado" elevation="400" location="Pozo" municipality="Campo Experim" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Aragua" typeStatus="holotype">
<typeStatus id="260601D8F27A14257271E8F1559B2AC9" box="[339,446,693,719]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF27A14257134E8F1567B2AC9" box="[534,606,693,719]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="19" pageNumber="20" type="Herbarium">NHM</collectionCode>
): “N.
<collectingCountry id="81AAFFEAF27A14257184E8F157742AC9" box="[678,849,693,719]" name="Venezuela" pageId="19" pageNumber="20">VENEZUELA</collectingCountry>
: \
<collectingCounty id="1063C7F6F27A14257053E8F157982AC9" box="[881,957,693,719]" pageId="19" pageNumber="20">Estado</collectingCounty>
<collectingRegion id="3B797198F27A142570E4E8F1503E2AC9" box="[966,1051,693,719]" country="Venezuela" name="Aragua" pageId="19" pageNumber="20">Aragua</collectingRegion>
, \
<collectorName id="5448DAACF27A1425771FE8F1510F2AC9" box="[1085,1322,693,719]" pageId="19" pageNumber="20">P. Nac. Henri Pittier</collectorName>
, \
<collectingMunicipality id="19662500F27A14257669E8F155242AF5" pageId="19" pageNumber="20">Campo Experim</collectingMunicipality>
[e]nt[a]l CENIAP \
<location id="FC62E9A1F27A142572C2E89D563D2AF5" LSID="urn:lsid:plazi:treatment:71140E6CF27A142273B5E80D560128FD:FC62E9A1F27A142572C2E89D563D2AF5" box="[480,536,729,755]" country="Venezuela" county="Estado" municipality="Campo Experim" name="Pozo" pageId="19" pageNumber="20" stateProvince="Aragua">Pozo</location>
del
<location id="FC62E9A1F27A1425716DE89D56BB2AF5" LSID="urn:lsid:plazi:treatment:71140E6CF27A142273B5E80D560128FD:FC62E9A1F27A1425716DE89D56BB2AF5" box="[591,670,729,755]" country="Venezuela" county="Estado" municipality="Campo Experim" name="Diablo" pageId="19" pageNumber="20" stateProvince="Aragua">Diablo</location>
,
<quantity id="3E45129FF27A1425718FE89D56D62AF5" box="[685,755,729,755]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="400.0">
<elevation id="72905849F27A1425718FE89D56D62AF5" box="[685,755,729,755]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="400.0">400m</elevation>
</quantity>
”; “Bristol Univ. Exped. \ Brit. Mus. Nat. Hist. \ 1992-6.”; “TRAY \ 15”; “FOG 17.
<date id="8D0399BAF27A14257297E8B956032B11" box="[437,550,765,791]" pageId="19" pageNumber="20" value="1990-05-14">
<collectingDate id="9D476052F27A14257297E8B956032B11" box="[437,550,765,791]" pageId="19" pageNumber="20" value="1990-05-14">14.v.1990</collectingDate>
</date>
. \
<location id="FC62E9A1F27A14257160E8B956C72B11" LSID="urn:lsid:plazi:treatment:71140E6CF27A142273B5E80D560128FD:FC62E9A1F27A14257160E8B956C72B11" box="[578,738,765,791]" country="Venezuela" county="Estado" municipality="Campo Experim" name="Gallery forest" pageId="19" pageNumber="20" stateProvince="Aragua">Gallery forest</location>
. \ Mangifera indica. \ In fruit.
<collectorName id="5448DAACF27A1425771FE8B950EA2B11" box="[1085,1231,765,791]" pageId="19" pageNumber="20">J. G. Davies</collectorName>
”; “Remainder of \ series at Mus. Inst. \ Zoo. Agricola. UCV: \ Maracay VENEZ”
</materialsCitation>
.
<materialsCitation id="49D5B527F27A14257077E96555792B59" ID-GBIF-Occurrence="2239706632" collectionCode="ZMHB" country="Venezuela" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Aragua" typeStatus="paratype">
<typeStatus id="260601D8F27A14257077E96557E32B3D" box="[853,966,801,827]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
. Male (
<collectionCode id="9FAC27BFF27A14257734E965504B2B3D" box="[1046,1134,801,827]" pageId="19" pageNumber="20">ZMHB</collectionCode>
): “49594”; “Caracas Mor”; “fasciatus \ mihi”
</materialsCitation>
.
<materialsCitation id="49D5B527F27A14257247E90150C82B85" ID-GBIF-Occurrence="2239706627" collectionCode="ZMHB" country="Venezuela" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Aragua" typeStatus="paratype">
<typeStatus id="260601D8F27A14257247E90155EF2B59" box="[357,458,837,863]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="9FAC27BFF27A1425713AE90156552B59" box="[536,624,837,863]" pageId="19" pageNumber="20">ZMHB</collectionCode>
): “49593”; “Caracas Mor”; “bifasciatus \ mihi”; “Hist.-Coll. (
<taxonomicName id="3EBDC4F9F27A14257634E90151B02B66" box="[1302,1429,837,864]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) \ Nr. 49593 \
<taxonomicName id="3EBDC4F9F27A14257205E92D55A52B85" box="[295,384,873,899]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Cyphon</taxonomicName>
bifasciatus Moritz \ Caracas, Moritz \ Zool. Mus. Berlin” [green label, printed]
</materialsCitation>
.
<materialsCitation id="49D5B527F27A142577D5E92D54D62C15" ID-GBIF-Occurrence="2239706641" collectionCode="ZMHB" country="Venezuela" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Aragua" typeStatus="paratype">
<typeStatus id="260601D8F27A142577D5E92D51792B85" box="[1271,1372,873,899]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="9FAC27BFF27A142573BDE9C954D22BA1" box="[159,247,909,935]" pageId="19" pageNumber="20">ZMHB</collectionCode>
): “49601”; “Columb. Moritz”; “Hist.-Coll. (
<taxonomicName id="3EBDC4F9F27A142571CCE9C957482BAE" box="[750,877,909,936]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) \ Nr. 49601 \
<taxonomicName id="3EBDC4F9F27A14257738E9C950562BA1" box="[1050,1139,909,935]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Cyphon</taxonomicName>
spec. \ Columb., Moritz \ Zool. Mus. Berlin” [green label, printed].
<specimenCount id="EFBB74F3F27A142571A1E9F556D72BCD" box="[643,754,945,971]" pageId="19" pageNumber="20" type="female">2 females</specimenCount>
(
<collectionCode id="9FAC27BFF27A14257027E9F557782BCD" box="[773,861,945,971]" pageId="19" pageNumber="20">ZMHB</collectionCode>
): “49590”; “Columb. Moritz”; “obscurus \ mihi”; “Hist.-Coll. (
<taxonomicName id="3EBDC4F9F27A14257205E99155832BF6" box="[295,422,981,1008]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) \ Nr. 49590 \
<taxonomicName id="3EBDC4F9F27A14257161E99157722BE9" authority="Moritz" authorityName="Moritz" box="[579,855,981,1007]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obscurus">Cyphon obscurus Moritz</taxonomicName>
\ Columb., Moritz \ Zool. Mus. Berlin” [green label, printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27A142573E5EE5857EF2C5D" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="F902BF7AF27A142573E5EE5857EF2C5D" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573E5EE5855672C31" bold="true" box="[199,322,1052,1079]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Small (TL
<quantity id="3E45129FF27A142572E1EE59566B2C31" box="[451,590,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="19" pageNumber="20" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
), dark brown species. Penis relatively short, narrowed in the middle, tergite VIII with distinct median process and without emarginations on its sides.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27A142573E5EE2056EB2FD9" pageId="19" pageNumber="20" type="description">
<paragraph id="F902BF7AF27A142573E5EE20506C2D99" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573E5EE20557C2C79" bold="true" box="[199,345,1124,1151]" pageId="19" pageNumber="20">Description.</emphasis>
<typeStatus id="260601D8F27A1425727DEE2155EF2C79" box="[351,458,1125,1151]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
, male. Body oval, convex, covered with whitish, suberect setae. Body dark brown, legs, antennae, and mouthparts yellowish. Head very broad, 1.7× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by 1.0–2.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 0.5–1.0× diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF27A142573E5EFED553A2EB9" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="20">
Male genitalia. Penis (L
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,
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W
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mm,
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) long and narrow, widened in basal half, parameroids short, hooked, pointed at apices, trigonium subtriangular, narrow, pointed at apex. Tegmen (L
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,
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W
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mm,
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) long, sides diverging, apex with wide lobes, covered with setae. Sternite IX (L
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,
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W
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mm,
<figureCitation id="6186A3FFF27A14257652EFB554E22E29" captionStart="FIGURE 15" captionStartId="20.[151,250,1137,1162]" captionTargetBox="[186,1401,251,1088]" captionTargetId="figure@20.[186,1403,272,1088]" captionTargetPageId="20" captionText="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783750/files/figure.png" pageId="19" pageNumber="20">Fig. 15C</figureCitation>
) subtriangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
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,
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W
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mm,
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) with relatively short apical plate, central portion projecting, with two subtriangular denticles, lateral parts of apical margin subtly setose, apodemes longer than apical plate, diverging. Tergite IX (L
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,
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W
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mm,
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) with membranous apical plate, apodemes slightly longer than apical plate.
</paragraph>
<paragraph id="F902BF7AF27A142573E5EC8D55B32F49" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="20">
Female. Body (
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) brown, TL (n = 1)
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. Apical portion of each elytron with two deep concavities and shallow, adsutural depression (
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). Gonocoxites (
<figureCitation id="6186A3FFF27A14257011ECA957BE2F01" box="[819,923,1773,1799]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="19" pageNumber="20">Fig. 10D</figureCitation>
) narrowly triangular, membranous, gonostyli subapical, bursal sclerite present (L
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,
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W
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mm), subtriangular, with small subconical process and wid- ened posterior portion.
</paragraph>
<paragraph id="F902BF7AF27A142573E5ED1D50222F75" blockId="19.[151,1437,692,2015]" box="[199,1031,1881,1907]" pageId="19" pageNumber="20">Variability. Colouration of body variable, from light brown to dark brown.</paragraph>
<paragraph id="F902BF7AF27A142573E5ED3956EB2FD9" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573E5ED3955DA2F91" bold="true" box="[199,511,1916,1943]" pageId="19" pageNumber="20">Measurements and ratios.</emphasis>
Males (n = 4) TL 2.27–2.63 (2.51) mm, PL 0.47–0.57 (0.51) mm, PW 1.10–1.23 (1.15) mm, EL 1.97–2.17 (2.09) mm, EW 1.13–1.27 (1.19) mm, TL/EW 1.42–1.58 (1.54), PW/PL 2.18–2.36 (2.27), EL/EW 1.23–1.31 (1.28), EL/PL 3.82–4.50 (4.13).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27D142273E5EADC560128FD" pageId="20" pageNumber="21" type="distribution">
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<emphasis id="CBC96368F27D142273E5EADC554428B5" bold="true" box="[199,353,152,179]" pageId="20" pageNumber="21">Distribution.</emphasis>
Known from two localities in
<collectingCountry id="81AAFFEAF27D1422719AEADC571528B4" box="[696,816,152,178]" name="Venezuela" pageId="20" pageNumber="21">Venezuela</collectingCountry>
: Caracas (old records) and Henri Pittier National Park.
<emphasis id="CBC96368F27D142273E5EAF9556B28D1" bold="true" box="[199,334,189,215]" pageId="20" pageNumber="21">Etymology.</emphasis>
Named after Henri François Pittier (
<date id="8D0399BAF27D142271DEEAF8575A28D1" box="[764,895,188,215]" pageId="20" pageNumber="21" value="1857" valueMax="1950">1857–1950</date>
), a Swiss-born botanist, who studied flora and fauna of
<collectingCountry id="81AAFFEAF27D142273D8EAA4554828FC" box="[250,365,224,250]" name="Venezuela" pageId="20" pageNumber="21">Venezuela</collectingCountry>
for many years.
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</treatment>
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<paragraph id="F902BF7AF27B142473B5EAF9550B28D1" blockId="18.[151,468,153,215]" box="[151,302,189,215]" pageId="18" pageNumber="19">
(
<figureCitation id="6186A3FFF27B142473BDEAF954DE28D1" box="[159,251,189,215]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="18" pageNumber="19">Figs 1H</figureCitation>
,
<figureCitation id="6186A3FFF27B1424722AEAF9550028D1" box="[264,293,189,215]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27B142473B5EB4257EA2945" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="F902BF7AF27B142473B5EB4257EA2945" blockId="18.[151,1437,261,828]" pageId="18" pageNumber="19">
<emphasis id="CBC96368F27B142473B5EB4255622926" bold="true" box="[151,327,261,288]" pageId="18" pageNumber="19">Type material.</emphasis>
<materialsCitation id="49D5B527F27B14247272EB4155862945" ID-GBIF-Occurrence="2239706622" collectionCode="MNHN" country="Bolivia" location="Collection Leon Fairmaire" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
<typeStatus id="260601D8F27B14247272EB41559E2919" box="[336,443,261,287]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF27B14247133EB41564B2919" box="[529,622,261,287]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="19">MNHN</collectionCode>
): “
<collectingRegion id="3B797198F27B142471A9EB4157052919" box="[651,800,261,287]" country="Bolivia" name="Cochabamba" pageId="18" pageNumber="19">Cochabamba</collectingRegion>
\ (
<collectingCountry id="81AAFFEAF27B14247061EB4157BC2926" box="[835,921,261,288]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
) Germain.”; “MUSEUM PARIS \
<location id="FC62E9A1F27B1424760AEB4155602945" LSID="urn:lsid:plazi:treatment:71140E6CF27B142573B5EADD50E029F1:FC62E9A1F27B1424760AEB4155602945" country="Bolivia" name="Collection Leon Fairmaire" pageId="18" pageNumber="19" stateProvince="Cochabamba">Collection Léon Fairmaire</location>
\ 1906”
</materialsCitation>
.
<materialsCitation id="49D5B527F27B1424728FEB6E57E92945" ID-GBIF-Occurrence="2239706646" box="[429,972,297,324]" collectionCode="MNHN" country="Bolivia" location="Collection Leon Fairmaire" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Cochabamba" typeStatus="paratype">
<typeStatus id="260601D8F27B1424728FEB6E56372942" box="[429,530,298,324]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="9FAC27BFF27B14247140EB6D569A2945" box="[610,703,297,323]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="19">MNHN</collectionCode>
): same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27B142473E5EB0955B229A9" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="F902BF7AF27B142473E5EB0955B229A9" blockId="18.[151,1437,261,828]" pageId="18" pageNumber="19">
<emphasis id="CBC96368F27B142473E5EB095567296E" bold="true" box="[199,322,333,360]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Resembles some colour forms of
<taxonomicName id="3EBDC4F9F27B142471E4EB0A577F2961" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[710,858,334,360]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="CBC96368F27B142471E4EB0A577F2961" box="[710,858,334,360]" italics="true" pageId="18" pageNumber="19">C. peruviana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27B14247043EB0A5799296E" bold="true" box="[865,956,334,360]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="D0FADE13F27B14247043EB0A5799296E" box="[865,956,334,360]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It is separable from similar species on the basis of characters of male genitalia: penis parallel-sided, shorter than in
<taxonomicName id="3EBDC4F9F27B142470F9EB36504A298D" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[987,1135,370,396]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="CBC96368F27B142470F9EB36504A298D" box="[987,1135,370,396]" italics="true" pageId="18" pageNumber="19">C. peruviana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27B14247755EB3650F7298A" bold="true" box="[1143,1234,370,396]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="D0FADE13F27B14247755EB3650F7298A" box="[1143,1234,370,396]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
tergite VIII lacks distinct emarginations.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27B142573E5EBFD56CB29A9" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="F902BF7AF27B142473E5EBFD57F42B3A" blockId="18.[151,1437,261,828]" pageId="18" pageNumber="19">
<emphasis id="CBC96368F27B142473E5EBFD557C29D2" bold="true" box="[199,345,441,468]" pageId="18" pageNumber="19">Description.</emphasis>
<typeStatus id="260601D8F27B14247242EBFD55EE29D5" box="[352,459,441,467]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
, male. Body oval, convex, covered with brownish, suberect setae. Head brown, pronotum orange, elytra orange with brown maculation, each elytron with brown stripe in anterior half, and with brown macula close to outer margin, posterior portion of each elytron with brown, oval macula, close to suture, outer margin of elytra brown, legs, antennae and mouthparts yellowish. Head very broad, 1.6× wider than width of interocular space, with distinct punctation, punctures small, granulate, separated by a 1.5–2.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 1.0–2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, impunctate. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
<caption id="ADC2EFF2F27B142473B5EC8757B22EF9" httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1731,1756]" targetBox="[184,1403,897,1694]" targetPageId="18">
<paragraph id="F902BF7AF27B142473B5EC8757B22EF9" blockId="18.[151,1436,1731,1792]" pageId="18" pageNumber="19">
<emphasis id="CBC96368F27B142473B5EC8755052EDA" bold="true" box="[151,288,1731,1756]" pageId="18" pageNumber="19">FIGURE 14.</emphasis>
<taxonomicName id="3EBDC4F9F27B14247204EC8055C52EDD" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[294,480,1732,1756]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="picta" status="sp. nov.">
<emphasis id="CBC96368F27B14247204EC8055C52EDD" box="[294,480,1732,1756]" italics="true" pageId="18" pageNumber="19">Calvariopsis picta</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27B142472C4EC8056132EDA" bold="true" box="[486,566,1732,1756]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="D0FADE13F27B142472C4EC8056132EDA" box="[486,566,1732,1756]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="F902BF7AF27B142573E5ED6A56DC28D1" blockId="18.[151,1437,1838,2044]" lastBlockId="19.[151,1437,153,503]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Male genitalia (
<figureCitation id="6186A3FFF27B1424725AED6A55F92F4E" box="[376,476,1838,1864]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">Fig. 14F</figureCitation>
). Penis (L
<quantity id="3E45129FF27B14247177ED6A56992F4E" box="[597,700,1838,1864]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="18" pageNumber="19" unit="mm" value="0.68">0.68 mm</quantity>
,
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W
<quantity id="3E45129FF27B142471CBED6A573F2F4E" box="[745,794,1838,1864]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="19" unit="mm" value="0.09">0.09</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27B14247079ED6A57E52F4E" box="[859,960,1838,1864]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">Fig. 14A</figureCitation>
) long, with subparallel sides, parameroids short, relatively wide, rounded at apices, trigonium subtriangular, narrow, pointed at apex. Tegmen (L
<quantity id="3E45129FF27B1424760DED1551BD2F6A" box="[1327,1432,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="18" pageNumber="19" unit="mm" value="0.54">0.54 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27B142473B5ED3254CE2F96" box="[151,235,1910,1936]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="-0.16">
W
<quantity id="3E45129FF27B14247398ED3254CE2F96" box="[186,235,1910,1936]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="mm" value="0.16">0.16</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27B1424720CED3255B72F96" box="[302,402,1910,1936]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">Fig. 14B</figureCitation>
) long, sides diverging, apex with short lateral lobes, covered with setae. Sternite VIII very small, U-shaped (
<figureCitation id="6186A3FFF27B1424727FEDDE55E12FB2" box="[349,452,1946,1972]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">Fig. 14D</figureCitation>
), sternite IX (L
<quantity id="3E45129FF27B14247155EDDE56F82FB2" box="[631,733,1946,1972]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="mm" value="0.3">0.30 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27B142471CAEDDE571F2FB2" box="[744,826,1946,1972]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="-0.17">
W
<quantity id="3E45129FF27B1424702BEDDE571F2FB2" box="[777,826,1946,1972]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="19" unit="mm" value="0.17">0.17</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27B14247059EDDE57FA2FB2" box="[891,991,1946,1972]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">Fig. 14C</figureCitation>
) subrectangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
<quantity id="3E45129FF27B1424774BEDFA50EA2FDE" box="[1129,1231,1982,2008]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="19" unit="mm" value="0.33">0.33 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27B142477F9EDFA510B2FDE" box="[1243,1326,1981,2008]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="-0.35">
W
<quantity id="3E45129FF27B142477DFEDF9510B2FDE" box="[1277,1326,1981,2008]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="19" unit="mm" value="0.35">0.35</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27B14247652EDFA54EC2FFA" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="18" pageNumber="19">Fig. 14D</figureCitation>
) with short apical plate, central portion of apical margin with small indentation, apical margin subtly setose, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF27A1425708BEADD503528B5" box="[937,1040,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="19" pageNumber="20" unit="mm" value="0.36">0.36 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27A1425773EEADD504B28B5" box="[1052,1134,153,179]" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="-0.27">
W
<quantity id="3E45129FF27A1425771FEADD504B28B5" box="[1085,1134,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="19" pageNumber="20" unit="mm" value="0.27">0.27</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27A14257792EADD513728B5" box="[1200,1298,153,179]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="19" pageNumber="20">Fig. 14E</figureCitation>
) with mem- branous apical plate, apodemes as long as apical plate.
</paragraph>
<paragraph id="F902BF7AF27A142573E5EAA550252945" blockId="19.[151,1437,153,503]" pageId="19" pageNumber="20">
Variability. Dorsal pattern very variable.
<typeStatus id="260601D8F27A142571A8EAA556C928FD" box="[650,748,225,251]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
with brownish black dorsum, each elytron with oval adsutural portion in apical part, outer part of anterior half of elytron orange. Certain variation can be noticed in the sternite VII which can have more (
<figureCitation id="6186A3FFF27A142572B7EB6D55D32945" box="[405,502,297,323]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="19" pageNumber="20">Fig. 14F</figureCitation>
) or less (
<figureCitation id="6186A3FFF27A1425717FEB6D56E62945" box="[605,707,297,323]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="19" pageNumber="20">Fig. 14D</figureCitation>
) projecting median process.
</paragraph>
<paragraph id="F902BF7AF27A142573E5EB0955BB2961" blockId="19.[151,1437,153,503]" box="[199,414,333,359]" pageId="19" pageNumber="20">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF27A142573E5EB3556CB29A9" blockId="19.[151,1437,153,503]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573E5EB3555D9298D" bold="true" box="[199,508,368,395]" pageId="19" pageNumber="20">Measurements and ratios.</emphasis>
Male (n = 1) TL
<quantity id="3E45129FF27A1425719CEB355701298D" box="[702,804,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="20" unit="mm" value="2.4">2.40 mm</quantity>
, PL
<quantity id="3E45129FF27A14257077EB35579E298D" box="[853,955,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="19" pageNumber="20" unit="mm" value="0.47">0.47 mm</quantity>
, PW
<quantity id="3E45129FF27A142570D4EB355079298D" box="[1014,1116,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="19" pageNumber="20" unit="mm" value="1.03">1.03 mm</quantity>
, EL
<quantity id="3E45129FF27A142577ADEB3550D0298D" box="[1167,1269,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="19" pageNumber="20" unit="mm" value="2.1">2.10 mm</quantity>
, EW
<quantity id="3E45129FF27A14257610EB3451BD298D" box="[1330,1432,368,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="19" pageNumber="20" unit="mm" value="1.53">1.53 mm</quantity>
, TL/EW 1.57, PW/PL 2.21, EL/EW 1.37, EL/PL 4.50.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27A142573E5EBFC574C29D5" box="[199,873,440,467]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="F902BF7AF27A142573E5EBFC574C29D5" blockId="19.[151,1437,153,503]" box="[199,873,440,467]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573E5EBFC554429D5" bold="true" box="[199,353,440,467]" pageId="19" pageNumber="20">Distribution.</emphasis>
Known only from the
<typeStatus id="260601D8F27A14257142EBFD56B529D5" box="[608,656,441,467]" pageId="19" pageNumber="20">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF27A1425702DEBFD574029D5" box="[783,869,441,467]" name="Bolivia" pageId="19" pageNumber="20">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27A142573E5EB9950E029F1" box="[199,1221,477,503]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="F902BF7AF27A142573E5EB9950E029F1" blockId="19.[151,1437,153,503]" box="[199,1221,477,503]" pageId="19" pageNumber="20">
<emphasis id="CBC96368F27A142573E5EB99556B29F1" bold="true" box="[199,334,477,503]" pageId="19" pageNumber="20">Etymology.</emphasis>
Lat.
<taxonomicName id="3EBDC4F9F27A142572AAEB9A55E529F1" authorityName="Ruta" authorityYear="2019" box="[392,448,478,503]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="CBC96368F27A142572AAEB9A55E529F1" box="[392,448,478,503]" italics="true" pageId="19" pageNumber="20">picta</emphasis>
</taxonomicName>
= painted, of various colours; a reference to colourful dorsal pattern.
</paragraph>
</subSubSection>
</treatment>
</document>

212
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<mods:title id="07EEF13CA09EFB7BBB46C7904FF77A87">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F27C142373B5EE9A56122CFF" box="[151,567,1246,1273]" pageId="21" pageNumber="22" type="nomenclature">
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<heading id="A24A0816F27C142373B5EE9A56122CFF" bold="true" box="[151,567,1246,1273]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="CBC96368F27C142373B5EE9A56122CFF" bold="true" box="[151,567,1246,1273]" pageId="21" pageNumber="22">
<taxonomicName id="3EBDC4F9F27C142373B5EE9A55F72CFF" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,466,1246,1273]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F27C142373B5EE9A55F72CFF" bold="true" box="[151,466,1246,1273]" italics="true" pageId="21" pageNumber="22">Calvariopsis venezuelense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F27C142372F8EE9B56122CFF" box="[474,567,1247,1273]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27C142373B5EF46550F2D1A" box="[151,298,1282,1308]" pageId="21" pageNumber="22" type="description">
<paragraph id="F902BF7AF27C142373B5EF46550F2D1A" blockId="21.[151,567,1246,1308]" box="[151,298,1282,1308]" pageId="21" pageNumber="22">
(
<figureCitation id="6186A3FFF27C142373BDEF4654DD2D1A" box="[159,248,1282,1308]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="21" pageNumber="22">Figs 1L</figureCitation>
,
<figureCitation id="6186A3FFF27C14237227EF4655072D1A" box="[261,290,1282,1308]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="21" pageNumber="22">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27C142373B5EF0E50FB2D8E" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="F902BF7AF27C142373B5EF0E50FB2D8E" blockId="21.[151,1437,1353,2029]" pageId="21" pageNumber="22">
<emphasis id="CBC96368F27C142373B5EF0E55622D62" bold="true" box="[151,327,1353,1380]" pageId="21" pageNumber="22">Type material.</emphasis>
<materialsCitation id="49D5B527F27C14237273EF0E50FB2D8E" ID-GBIF-Occurrence="2239706611" collectingDate="1930-05" collectionCode="SEMC" collectorName="J. Ashe &amp; R. Brooks &amp; R. Hanley" country="Venezuela" elevation="2420" location="Tariba" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Tachira" typeStatus="holotype">
<typeStatus id="260601D8F27C14237273EF0E55992D62" box="[337,444,1354,1380]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
, male (
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): “
<collectingCountry id="81AAFFEAF27C142371A1EF0E57142D62" box="[643,817,1354,1380]" name="Venezuela" pageId="21" pageNumber="22">VENEZUELA</collectingCountry>
:
<collectingRegion id="3B797198F27C1423701FEF0E57B62D62" box="[829,915,1354,1380]" country="Venezuela" name="Tachira" pageId="21" pageNumber="22">Táchira</collectingRegion>
\
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,
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NE,
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<elevation id="72905849F27C14237786EF0E50DE2D63" box="[1188,1275,1354,1381]" metricMagnitude="3" metricUnit="m" metricValue="2.42" pageId="21" pageNumber="22" unit="m" value="2420.0">2420 m</elevation>
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\ 7°58’20’’N, 72°9’15’’W
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<date id="8D0399BAF27C14237201EF2A55A12D8E" box="[291,388,1390,1416]" pageId="21" pageNumber="22" value="1930-05">30 MAY</date>
</collectingDate>
\ 1998;
<collectorName id="5448DAACF27C142372C2EF2B56112D8E" box="[480,564,1390,1417]" pageId="21" pageNumber="22">J. Ashe</collectorName>
,
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,
<collectorName id="5448DAACF27C1423719DEF2B57142D8E" box="[703,817,1390,1417]" pageId="21" pageNumber="22">R. Hanley</collectorName>
\ VEN1ABH98 161 ex: fungusy log”.
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</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27C142373E5EFD551202DF2" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="F902BF7AF27C142373E5EFD551202DF2" blockId="21.[151,1437,1353,2029]" pageId="21" pageNumber="22">
<emphasis id="CBC96368F27C142373E5EFD555672DAA" bold="true" box="[199,322,1425,1452]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Small species (TL
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) with elongated body. Penis distinctly narrowed in the middle of its length, tergite VIII with distinct process in the middle of apical margin, flanked with two emarginations covered with spines. Similar to
<taxonomicName id="3EBDC4F9F27C142372B8EF9F560F2DF2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[410,554,1498,1525]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F27C142372B8EF9F560F2DF2" box="[410,554,1498,1525]" italics="true" pageId="21" pageNumber="22">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27C14237113EF9E56AF2DF2" bold="true" box="[561,650,1498,1524]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="D0FADE13F27C14237113EF9E56AF2DF2" box="[561,650,1498,1524]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but body is more elongated and its sides are subparallel.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27C142073E5EFB956CB2F25" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="F902BF7AF27C142373E5EFB9507B2F13" blockId="21.[151,1437,1353,2029]" pageId="21" pageNumber="22">
<emphasis id="CBC96368F27C142373E5EFB9557C2E1E" bold="true" box="[199,345,1533,1560]" pageId="21" pageNumber="22">Description.</emphasis>
Male. Body elongated, convex, covered with whitish, suberect setae. Body dark brown, legs, antennae, and mouthparts yellowish. Head broad, 1.6× wider than width of interocular space, almost impunctate, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with very fine, granulate punctation, punctures separated by ca. 3.0–4.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation, similar to those on pronotum. Elytra elongate, without traces of longitudinal ridges, subparallel in basal 2/3, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation much stronger than on pronotum, distance between punctures ca. the diameter of a puncture.
</paragraph>
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Male genitalia. Penis (L
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,
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W
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mm,
<figureCitation id="6186A3FFF27C142371F7ED5A571D2F3E" box="[725,824,1822,1848]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="21" pageNumber="22">Fig. 17A</figureCitation>
) long, distinctly widened in basal portion, parameroids short, rounded at apices, slightly curved, trigonium subtriangular, rounded at apex. Tegmen (L
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,
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W
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mm,
<figureCitation id="6186A3FFF27C142373F2ED2255162F86" box="[208,307,1894,1920]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="21" pageNumber="22">Fig. 17B</figureCitation>
) long, sides diverging, apex with lobes covered with setae. Sternite IX (L
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,
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W
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</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27C14237652ED2254E32FA2" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="21" pageNumber="22">Fig. 17B</figureCitation>
) subtriangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
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,
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W
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</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27C1423716CEDEA56972FCE" box="[590,690,1966,1992]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="21" pageNumber="22">Fig. 17C</figureCitation>
) with relatively short apical plate, central portion projecting, with two subtriangular denticles, lateral parts of apical margin with 4–5 sharp denticles on each side, and several smaller ones, apodemes longer than apical plate. Tergite IX (L
<quantity id="3E45129FF27F14207025EADD574A28B5" box="[775,879,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="mm" value="0.3">0.30 mm</quantity>
,
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W
<quantity id="3E45129FF27F142070BFEADD57EB28B5" box="[925,974,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="23" unit="mm" value="0.24">0.24</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF27F14207733EADD505328B5" box="[1041,1142,153,179]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="22" pageNumber="23">Fig. 17D</figureCitation>
) with membranous apical plate, apodemes longer than apical plate.
</paragraph>
<caption id="ADC2EFF2F27F142073B5EC1155112E94" httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1621,1646]" targetBox="[152,1422,215,1572]" targetPageId="22">
<paragraph id="F902BF7AF27F142073B5EC1155112E94" blockId="22.[151,1436,1621,1682]" pageId="22" pageNumber="23">
<emphasis id="CBC96368F27F142073B5EC11553A2E68" bold="true" box="[151,287,1621,1646]" pageId="22" pageNumber="23">FIGURE 17.</emphasis>
<taxonomicName id="3EBDC4F9F27F14207206EC12560A2E68" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[292,559,1622,1646]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F27F14207206EC12560A2E68" box="[292,559,1622,1646]" italics="true" pageId="22" pageNumber="23">Calvariopsis venezuelense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F27F14207117EC1256A12E68" bold="true" box="[565,644,1622,1646]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="D0FADE13F27F14207117EC1256A12E68" box="[565,644,1622,1646]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="F902BF7AF27F142073E5EC8455BB2EDC" blockId="22.[151,1436,1728,1899]" box="[199,414,1728,1754]" pageId="22" pageNumber="23">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF27F142073E5ECA156CB2F25" blockId="22.[151,1436,1728,1899]" pageId="22" pageNumber="23">
<emphasis id="CBC96368F27F142073E5ECA155D92EF9" bold="true" box="[199,508,1764,1791]" pageId="22" pageNumber="23">Measurements and ratios.</emphasis>
Male (n = 1) TL
<quantity id="3E45129FF27F1420719CECA057012EF9" box="[702,804,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="22" pageNumber="23" unit="mm" value="2.63">2.63 mm</quantity>
, PL
<quantity id="3E45129FF27F14207077ECA1579E2EF9" box="[853,955,1765,1791]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="22" pageNumber="23" unit="mm" value="0.43">0.43 mm</quantity>
, PW 1.00 mm, EL
<quantity id="3E45129FF27F142077ADECA150D02EF9" box="[1167,1269,1765,1791]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="22" pageNumber="23" unit="mm" value="2.3">2.30 mm</quantity>
, EW
<quantity id="3E45129FF27F14207610ECA151BD2EF9" box="[1330,1432,1765,1791]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" pageId="22" pageNumber="23" unit="mm" value="1.47">1.47 mm</quantity>
, TL/EW 1.80, PW/PL 2.31, EL/EW 1.57, EL/PL 5.31.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27F142073E5ED6857AF2F40" box="[199,906,1836,1863]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="F902BF7AF27F142073E5ED6857AF2F40" blockId="22.[151,1436,1728,1899]" box="[199,906,1836,1863]" pageId="22" pageNumber="23">
<emphasis id="CBC96368F27F142073E5ED6855442F41" bold="true" box="[199,353,1836,1863]" pageId="22" pageNumber="23">Distribution.</emphasis>
Known only from the
<typeStatus id="260601D8F27F14207142ED6956B52F41" box="[608,656,1837,1863]" pageId="22" pageNumber="23">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF27F1420702DED6857A32F40" box="[783,902,1836,1862]" name="Venezuela" pageId="22" pageNumber="23">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27F142073E5ED1550A52F6C" box="[199,1152,1872,1899]" pageId="22" pageNumber="23" type="etymology">
<paragraph id="F902BF7AF27F142073E5ED1550A52F6C" blockId="22.[151,1436,1728,1899]" box="[199,1152,1872,1899]" pageId="22" pageNumber="23">
<emphasis id="CBC96368F27F142073E5ED15556B2F6D" bold="true" box="[199,334,1873,1899]" pageId="22" pageNumber="23">Etymology.</emphasis>
Named after
<collectingCountry id="81AAFFEAF27F142072C4ED1456792F6C" box="[486,604,1872,1898]" name="Venezuela" pageId="22" pageNumber="23">Venezuela</collectingCountry>
, where the
<typeStatus id="260601D8F27F142071FEED1557292F6D" box="[732,780,1873,1899]" pageId="22" pageNumber="23">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
</treatment>
</document>

160
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<mods:title id="B2FD0781F1F844F230FDDC3936602D14">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<subSubSection id="B1A7ECF1F27D142273B5EF4456C22D1D" box="[151,743,1280,1307]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="F902BF7AF27D142273B5EF4456C22D1D" blockId="20.[151,743,1280,1343]" box="[151,743,1280,1307]" pageId="20" pageNumber="21">
<heading id="A24A0816F27D142273B5EF4456C22D1D" bold="true" box="[151,743,1280,1307]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="CBC96368F27D142273B5EF4456C22D1D" bold="true" box="[151,743,1280,1307]" pageId="20" pageNumber="21">
<taxonomicName id="3EBDC4F9F27D142273B5EF44567C2D1C" authority="(Pic, 1931)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[151,601,1280,1307]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipenne" status="comb. nov.">
<emphasis id="CBC96368F27D142273B5EF4455E82D1D" bold="true" box="[151,461,1280,1307]" italics="true" pageId="20" pageNumber="21">Calvariopsis sculptipenne</emphasis>
(
<bibRefCitation id="9D2CC28BF27D142272FFEF4456742D1C" author="Pic, M." box="[477,593,1280,1306]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F27D14227142EF4556C22D1D" box="[608,743,1281,1307]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27D142273B5EF6055B72D39" box="[151,402,1316,1343]" pageId="20" pageNumber="21" type="description">
<paragraph id="F902BF7AF27D142273B5EF6055B72D39" blockId="20.[151,743,1280,1343]" box="[151,402,1316,1343]" pageId="20" pageNumber="21">
(
<figureCitation id="6186A3FFF27D142273BDEF6054D92D38" box="[159,252,1316,1342]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="20" pageNumber="21">Figs 1K</figureCitation>
,
<figureCitation id="6186A3FFF27D1422722AEF6055302D38" box="[264,277,1316,1342]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="20" pageNumber="21">7</figureCitation>
C–D, 16A)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27D142273B5EF2D56DC2D99" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="F902BF7AF27D142273B5EF2D562F2D87" blockId="20.[151,761,1384,1440]" box="[151,522,1384,1409]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="D9AD9854F27D142273B5EF2D562F2D87" box="[151,522,1384,1409]" pageId="20" pageNumber="21">
<emphasis id="CBC96368F27D142273B5EF2D548D2D86" box="[151,168,1385,1408]" italics="true" pageId="20" pageNumber="21">=</emphasis>
<taxonomicName id="3EBDC4F9F27D1422738DEF2D562F2D87" authority="Pic, 1931: 5" authorityName="Pic" authorityPageNumber="5" authorityYear="1931" box="[175,522,1384,1409]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipennis">
<emphasis id="CBC96368F27D1422738DEF2D55AF2D87" box="[175,394,1385,1409]" italics="true" pageId="20" pageNumber="21">Cyphon sculptipennis</emphasis>
<treatmentCitation id="781C996BF27D142272B2EF2C562F2D87" author="Pic, M." box="[400,522,1384,1409]" page="5" pageId="20" pageNumber="21" year="1931">
<bibRefCitation id="9D2CC28BF27D142272B2EF2C55D62D87" author="Pic, M." box="[400,499,1384,1409]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
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</paragraph>
<paragraph id="F902BF7AF27D142273B5EFCC56DC2D99" blockId="20.[151,761,1384,1440]" box="[151,761,1415,1440]" pageId="20" pageNumber="21">
<emphasis id="CBC96368F27D142273B5EFCC548D2D99" box="[151,168,1416,1439]" italics="true" pageId="20" pageNumber="21">=</emphasis>
<taxonomicName id="3EBDC4F9F27D1422738DEFC356192D99" authority="(Pic, 1931)" authorityName="Pic" authorityYear="1931" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[175,572,1415,1440]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipennis">
<emphasis id="CBC96368F27D1422738DEFC355E72D99" box="[175,450,1415,1439]" italics="true" pageId="20" pageNumber="21">Contacyphon sculptipennis</emphasis>
(
<bibRefCitation id="9D2CC28BF27D142272EDEFC356102D99" author="Pic, M." box="[463,565,1415,1440]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
)
</taxonomicName>
: Zwick
<emphasis id="CBC96368F27D142271AFEFCC56E52D99" box="[653,704,1415,1439]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2013
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27D142273B5EF8A506C2E0A" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="F902BF7AF27D142273B5EF8A506C2E0A" blockId="20.[151,1437,1485,2016]" pageId="20" pageNumber="21">
<emphasis id="CBC96368F27D142273B5EF8A55622DEE" bold="true" box="[151,327,1485,1512]" pageId="20" pageNumber="21">Type material.</emphasis>
<materialsCitation id="49D5B527F27D14227272EF8A506C2E0A" ID-GBIF-Occurrence="2239706645" collectionCode="MNHN" pageId="20" pageNumber="21" specimenCount="2" typeStatus="holotype">
<typeStatus id="260601D8F27D14227272EF8A559E2DEE" box="[336,443,1486,1512]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
, female (
<collectionCode id="9FAC27BFF27D14227104EF8A56A62DEE" box="[550,643,1486,1512]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="21">MNHN</collectionCode>
): “Corumba \ Matt Grosso” [printed label]; “sculptipennis \ n sp” [in Pic’s hand]; “
<typeStatus id="260601D8F27D1422720FEFB655F22E0A" box="[301,471,1522,1548]" pageId="20" pageNumber="21" type="holotype">HOLOTYPUS</typeStatus>
\
<taxonomicName id="3EBDC4F9F27D142272CEEFB656FB2E0A" authorityName="Pic" authorityYear="1931" box="[492,734,1522,1548]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipennis">
<emphasis id="CBC96368F27D142272CEEFB656FB2E0A" box="[492,734,1522,1548]" italics="true" pageId="20" pageNumber="21">Cyphon sculptipennis</emphasis>
</taxonomicName>
\
<bibRefCitation id="9D2CC28BF27D142271D1EFB6577A2E0A" author="Pic, M." box="[755,863,1522,1548]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
” [red label, printed].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27D142373E5EC51570328D1" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="F902BF7AF27D142273E5EC5157062FBA" blockId="20.[151,1437,1485,2016]" pageId="20" pageNumber="21">
<emphasis id="CBC96368F27D142273E5EC5155502E36" bold="true" box="[199,373,1557,1584]" pageId="20" pageNumber="21">Redescription.</emphasis>
Female. Body oblong oval, convex, covered with yellowish suberect setae. Body uniformly dark brown, venter brown. Head very broad, 1.75× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by ca. 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 1.5–2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, with very fine, granulate punctation. Angle between pronotum and elytra subtly marked in dorsal outline. Elytra oblong, without traces of longitudinal ridges, broadest in the middle of their length. Sides slightly curved in apical 1/2, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 0.7–1.0× diameter of a puncture. Apical portion of elytra depressed in adsutural portion, with two deep cavities on each elytron (
<figureCitation id="6186A3FFF27D1422713CED3A56402F9E" box="[542,613,1918,1944]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="20" pageNumber="21">Figs 7</figureCitation>
C–D). Sternite VIII elongate, Y-shaped, with single apodeme. Remaining structures of genital tract missing in the studied specimen.
</paragraph>
<paragraph id="F902BF7AF27D142273E5ED8255A02FE6" blockId="20.[151,1437,1485,2016]" box="[199,389,1990,2016]" pageId="20" pageNumber="21">Male. Unknown.</paragraph>
<paragraph id="F902BF7AF27C142373E5EADD570328D1" blockId="21.[151,1437,152,359]" pageId="21" pageNumber="22">
<emphasis id="CBC96368F27C142373E5EADD562528B5" bold="true" box="[199,512,152,179]" pageId="21" pageNumber="22">Measurements and ratios.</emphasis>
Female (n = 1) TL
<quantity id="3E45129FF27C142371C1EADD576E28B5" box="[739,843,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.83" pageId="21" pageNumber="22" unit="mm" value="2.83">2.83 mm</quantity>
, PL
<quantity id="3E45129FF27C142370A2EADC57C228B5" box="[896,999,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="21" pageNumber="22" unit="mm" value="0.57">0.57 mm</quantity>
, PW
<quantity id="3E45129FF27C14237704EADD50A828B5" box="[1062,1165,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="21" pageNumber="22" unit="mm" value="1.23">1.23 mm</quantity>
, EL
<quantity id="3E45129FF27C142377E1EADD510E28B5" box="[1219,1323,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="21" pageNumber="22" unit="mm" value="2.34">2.34 mm</quantity>
, EW
<quantity id="3E45129FF27C14237649EADD54E028D1" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="21" pageNumber="22" unit="mm" value="1.83">1.83 mm</quantity>
, TL/EW 1.55, PW/PL 2.16, EL/EW 1.28, EL/PL 4.11.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27C142373E5EAA454D22918" pageId="21" pageNumber="22" type="distribution">
<paragraph id="F902BF7AF27C142373E5EAA454D22918" blockId="21.[151,1437,152,359]" pageId="21" pageNumber="22">
<emphasis id="CBC96368F27C142373E5EAA4554428FD" bold="true" box="[199,353,224,251]" pageId="21" pageNumber="22">Distribution.</emphasis>
Known only from the
<typeStatus id="260601D8F27C14237149EAA556BE28FD" box="[619,667,225,251]" pageId="21" pageNumber="22">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF27C14237002EAA4574228FC" box="[800,871,224,250]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
, close to the border with
<collectingCountry id="81AAFFEAF27C142377ADEAA450C128FC" box="[1167,1252,224,250]" name="Bolivia" pageId="21" pageNumber="22">Bolivia</collectingCountry>
, in the Pantanal wetland.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27C142373E5EB6D579A2961" pageId="21" pageNumber="22" type="discussion">
<paragraph id="F902BF7AF27C142373E5EB6D579A2961" blockId="21.[151,1437,152,359]" pageId="21" pageNumber="22">
<emphasis id="CBC96368F27C142373E5EB6D551E2945" bold="true" box="[199,315,297,323]" pageId="21" pageNumber="22">Remarks.</emphasis>
Presence of well developed, foveate excitators in apical portions of elytra, and TL larger than
<quantity id="3E45129FF27C1423766AEB6D51B82945" box="[1352,1437,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="mm" value="2.4">2.4 mm</quantity>
support inclusion of this species in the group I of
<taxonomicName id="3EBDC4F9F27C142371E0EB0857742960" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[706,849,332,358]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F27C142371E0EB0857742960" box="[706,849,332,358]" italics="true" pageId="21" pageNumber="22">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F27C1423707BEB09579A2961" box="[857,959,333,359]" pageId="21" pageNumber="22" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

683
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<mods:title id="FBB816BB8FF66B8830F52D6BEC4C02CB">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<emphasis id="CBC96368F27E142173B5EADD566628B2" bold="true" box="[151,579,153,180]" pageId="23" pageNumber="24">
<taxonomicName id="3EBDC4F9F27E142173B5EADD55F828B2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,477,153,180]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
<emphasis id="CBC96368F27E142173B5EADD55F828B2" bold="true" box="[151,477,153,180]" italics="true" pageId="23" pageNumber="24">Calvariopsis yanayacuense</emphasis>
</taxonomicName>
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</emphasis>
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</paragraph>
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<subSubSection id="B1A7ECF1F27E142173B5EAF9566728D1" box="[151,578,189,216]" pageId="23" pageNumber="24" type="description">
<paragraph id="F902BF7AF27E142173B5EAF9566728D1" blockId="23.[151,579,153,216]" box="[151,578,189,216]" pageId="23" pageNumber="24">
(
<figureCitation id="6186A3FFF27E142173BDEAF954C428D1" box="[159,225,189,215]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="23" pageNumber="24">Figs 1</figureCitation>
M–O, 3A–C, 4A–C, 5A, 6, 18)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27E142173B5EB4257CA2CA5" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="F902BF7AF27E142173B5EB4257CA2CA5" blockId="23.[151,1437,261,1980]" pageId="23" pageNumber="24">
<emphasis id="CBC96368F27E142173B5EB42556D2926" bold="true" box="[151,328,261,288]" pageId="23" pageNumber="24">Type material.</emphasis>
<materialsCitation id="49D5B527F27E14217270EB4155772945" ID-GBIF-Occurrence="2239706619" collectingDate="2009-12-02" collectionCode="DBET" collectorName="Rafal Ruta" county="Ecuador" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="260601D8F27E14217270EB4155982919" box="[338,445,261,287]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
, male (
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): “
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36,
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prov., \
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+
<location id="FC62E9A1F27E1421770DEB4150802919" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E1421770DEB4150802919" box="[1071,1189,261,287]" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\ road,
<date id="8D0399BAF27E14217621EB4151A32919" box="[1283,1414,261,288]" pageId="23" pageNumber="24" value="2009-12-02">
<collectingDate id="9D476052F27E14217621EB4151A32919" box="[1283,1414,261,288]" pageId="23" pageNumber="24" value="2009-12-02">2 XII 2009</collectingDate>
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<collectorName id="5448DAACF27E142173EBEB6D556D2945" box="[201,328,297,323]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
”
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.
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<typeStatus id="260601D8F27E1421727CEB6E55EA2942" box="[350,463,298,324]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
.
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(
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\ road
<collectingMunicipality id="19662500F27E142170BAEB6D50122945" box="[920,1079,297,323]" pageId="23" pageNumber="24">Yanayacu-Rio</collectingMunicipality>
\
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,
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<collectingDate id="9D476052F27E142177BBEB6D513E2945" box="[1177,1307,297,324]" pageId="23" pageNumber="24" value="2009-11-30">30 XI 2009</collectingDate>
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\ leg.
<collectorName id="5448DAACF27E14217646EB6D552F2961" pageId="23" pageNumber="24">Lech Borowiec</collectorName>
”
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.
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<specimenCount id="EFBB74F3F27E1421723DEB0955A12961" box="[287,388,333,360]" pageId="23" pageNumber="24" type="male">15 males</specimenCount>
(
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41,
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prov., \
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vic.,
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\ trail+
<location id="FC62E9A1F27E142177A4EB0950DB2961" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142177A4EB0950DB2961" box="[1158,1278,333,359]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
,
<date id="8D0399BAF27E14217628EB095162296E" box="[1290,1351,333,360]" pageId="23" pageNumber="24">
<collectingDate id="9D476052F27E14217628EB095162296E" box="[1290,1351,333,360]" pageId="23" pageNumber="24">3 XII</collectingDate>
</date>
\ 2009, leg.
<collectorName id="5448DAACF27E142173E5EB355561298D" box="[199,324,369,395]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
”
</materialsCitation>
.
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<specimenCount id="EFBB74F3F27E1421727AEB35559E298D" box="[344,443,369,395]" pageId="23" pageNumber="24" type="male">10 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E142172E8EB35563F298D" box="[458,538,369,395]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217117EB3556B0298D" box="[565,661,369,395]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
4,
<collectingRegion id="3B797198F27E14217195EB3656D1298A" box="[695,756,370,396]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E1421706FEB355796298D" box="[845,947,369,395]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
+
<location id="FC62E9A1F27E142170E0EB35501C298D" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142170E0EB35501C298D" box="[962,1081,369,395]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
, \ collecting along the road, \
<date id="8D0399BAF27E142176A2EB3554D329A9" pageId="23" pageNumber="24" value="2009-11-25">
<collectingDate id="9D476052F27E142176A2EB3554D329A9" pageId="23" pageNumber="24" value="2009-11-25">25 XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E1421720DEBD255AF29A9" box="[303,394,405,432]" pageId="23" pageNumber="24">R. Ruta</collectorName>
”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E142172BCEBD1572029D5" ID-GBIF-Occurrence="2239706618" collectingDate="2009-12-15" collectionCode="DBET" collectorName="L. Borowiec" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E142172BCEBD155CD29A9" box="[414,488,405,431]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(
<collectionCode id="9FAC27BFF27E142172D5EBD1566229A9" box="[503,583,405,431]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217140EBD156E729A9" box="[610,706,405,431]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
76,
<collectingRegion id="3B797198F27E142171D3EBD2570B29B6" box="[753,814,406,432]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E142170AAEBD157CD29A9" box="[904,1000,405,431]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E14217706EBD250B429B6" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217706EBD250B429B6" box="[1060,1169,406,432]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E14217787EBD150D229A9" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217787EBD150D229A9" box="[1189,1271,405,431]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E14217623EBD151BD29A9" box="[1281,1432,405,431]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E14217623EBD151BD29A9" box="[1281,1432,405,431]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, \
<date id="8D0399BAF27E142172B4EBFD560429D5" box="[406,545,441,468]" pageId="23" pageNumber="24" value="2009-12-15">
<collectingDate id="9D476052F27E142172B4EBFD560429D5" box="[406,545,441,468]" pageId="23" pageNumber="24" value="2009-12-15">15 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF27E1421714BEBFE56DE29D5" box="[617,763,441,468]" pageId="23" pageNumber="24">L. Borowiec</collectorName>
”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E1421702CEBFD507E29F1" ID-GBIF-Occurrence="2239706655" collectingDate="2009-12-16" collectionCode="DBET" collectorName="R. Ruta" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="2" specimenCount-male="2" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E1421702CEBFD574129D5" box="[782,868,441,467]" pageId="23" pageNumber="24" type="male">2 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E14217050EBFD57E729D5" box="[882,962,441,467]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E142170FCEBFD501B29D5" box="[990,1086,441,467]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
75,
<collectingRegion id="3B797198F27E1421774CEBFE508E29D2" box="[1134,1195,442,468]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217627EBFD514029D5" box="[1285,1381,441,467]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E142173B5EB9A552129FE" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142173B5EB9A552129FE" box="[151,260,478,504]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E1421723FEB99554A29F1" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E1421723FEB99554A29F1" box="[285,367,477,503]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E1421725EEB99563729F1" box="[380,530,477,503]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E1421725EEB99563729F1" box="[380,530,477,503]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, \
<date id="8D0399BAF27E14217005EB99579329F1" box="[807,950,477,504]" pageId="23" pageNumber="24" value="2009-12-16">
<collectingDate id="9D476052F27E14217005EB99579329F1" box="[807,950,477,504]" pageId="23" pageNumber="24" value="2009-12-16">16 XII 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E142170D6EB9A507429F1" box="[1012,1105,477,504]" pageId="23" pageNumber="24">R. Ruta</collectorName>
”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E14217745EB9955402A39" ID-GBIF-Occurrence="2239706638" collectingDate="2009-12-09" collectionCode="DBET" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="5" specimenCount-male="5" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E14217745EB99509B29F1" box="[1127,1214,477,504]" pageId="23" pageNumber="24" type="male">5 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E142177EDEB99513A29F1" box="[1231,1311,477,503]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E1421761EEB9951B929F1" box="[1340,1436,477,503]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
58,
<collectingRegion id="3B797198F27E142173E2E84654D82A1A" box="[192,253,514,540]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217275E84555922A1D" box="[343,439,513,539]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E142172D1E84656452A1A" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142172D1E84656452A1A" box="[499,608,514,540]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E14217156E84556E32A1D" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217156E84556E32A1D" box="[628,710,513,539]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
, stream trail, \
<quantity id="3E45129FF27E14217048E84550252A1D" box="[874,1024,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E14217048E84550252A1D" box="[874,1024,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, 00°35’S \ 77°53’W,
<date id="8D0399BAF27E142177CFE845514C2A1D" box="[1261,1385,513,540]" pageId="23" pageNumber="24" value="2009-12-09">
<collectingDate id="9D476052F27E142177CFE845514C2A1D" box="[1261,1385,513,540]" pageId="23" pageNumber="24" value="2009-12-09">9 XII 2009</collectingDate>
</date>
, leg. \ Lech Borowiec”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E14217253E86156D92A65" ID-GBIF-Occurrence="2239706621" collectingDate="2009-11" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="5" specimenCount-male="5" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E14217253E86155EC2A39" box="[369,457,549,576]" pageId="23" pageNumber="24" type="male">5 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E142172F8E861560F2A39" box="[474,554,549,575]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217165E86156822A39" box="[583,679,549,575]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
1,
<collectingRegion id="3B797198F27E142171ECE862572E2A46" box="[718,779,550,576]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217049E86157EE2A39" box="[875,971,549,575]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E1421772EE862505C2A46" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E1421772EE862505C2A46" box="[1036,1145,550,576]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E142177B0E86150C12A39" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142177B0E86150C12A39" box="[1170,1252,549,575]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E142177D3E86151AD2A39" box="[1265,1416,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E142177D3E86151AD2A39" box="[1265,1416,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, 23–24 \
<date id="8D0399BAF27E142172FAE80E561D2A65" box="[472,568,585,612]" pageId="23" pageNumber="24" value="2009-11">
<collectingDate id="9D476052F27E142172FAE80E561D2A65" box="[472,568,585,612]" pageId="23" pageNumber="24" value="2009-11">XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E14217156E80D56D62A65" box="[628,755,585,611]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E14217025E80D56B12A81" ID-GBIF-Occurrence="2239706612" collectingDate="2009-12-03" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" county="Ecuador" location="Antisana" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="3" specimenCount-male="3" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E14217025E80D57782A65" box="[775,861,585,611]" pageId="23" pageNumber="24" type="male">3 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E1421704FE80D57982A65" box="[877,957,585,611]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E142170FBE80D501C2A65" box="[985,1081,585,611]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
42,
<collectingRegion id="3B797198F27E14217749E80E508D2A62" box="[1131,1192,586,612]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217626E80D51412A65" box="[1284,1380,585,611]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E142173B5E82954DE2A81" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142173B5E82954DE2A81" box="[151,251,621,647]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Antisana" pageId="23" pageNumber="24" stateProvince="Napo">Antisana</location>
\ trail,
<date id="8D0399BAF27E1421726BE82955E02A81" box="[329,453,621,648]" pageId="23" pageNumber="24" value="2009-12-03">
<collectingDate id="9D476052F27E1421726BE82955E02A81" box="[329,453,621,648]" pageId="23" pageNumber="24" value="2009-12-03">3 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF27E1421712FE82956AF2A81" box="[525,650,621,647]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E142171BCE82A504F2AAD" ID-GBIF-Occurrence="2239706614" collectingDate="2009-11-23" collectingDateMax="2009-12-17" collectingDateMin="2009-11-23" collectionCode="DBET" collectorName="L. Borowiec" country="Ecuador" county="Cosanga" elevation="2100" location="Station" municipality="Yanayacu" pageId="23" pageNumber="24" specimenCount="7" specimenCount-male="7" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E142171BCE82A56D62A81" box="[670,755,621,648]" pageId="23" pageNumber="24" type="male">7 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E14217023E82957742A81" box="[769,849,621,647]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF27E1421704EE82957DC2A81" box="[876,1017,621,647]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
,
<collectingRegion id="3B797198F27E14217726E82A50642A8E" box="[1028,1089,622,648]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov. \
<collectingCounty id="1063C7F6F27E142177B1E82950D62A81" box="[1171,1267,621,647]" pageId="23" pageNumber="24">Cosanga</collectingCounty>
vic.,
<collectingMunicipality id="19662500F27E1421760DE82A51B92A8E" box="[1327,1436,622,648]" pageId="23" pageNumber="24">Yanayacu</collectingMunicipality>
\
<location id="FC62E9A1F27E14217384E8D554DD2AAD" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217384E8D554DD2AAD" box="[166,248,657,683]" country="Ecuador" county="Cosanga" municipality="Yanayacu" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E14217226E8D555BF2AAD" box="[260,410,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E14217226E8D555BF2AAD" box="[260,410,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’ S, 77°53’W, \
<date id="8D0399BAF27E14217185E8D557A72AAD" box="[679,898,657,684]" pageId="23" pageNumber="24" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">
<collectingDate id="9D476052F27E14217185E8D557A72AAD" box="[679,898,657,684]" pageId="23" pageNumber="24" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">23 XI–17 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF27E142170EFE8D650452AAD" box="[973,1120,657,684]" pageId="23" pageNumber="24">L. Borowiec</collectorName>
”
</materialsCitation>
.
<materialsCitation id="49D5B527F27E14217756E8D556A42AF5" ID-GBIF-Occurrence="2239706634" collectingDate="2009-12-05" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" county="Ecuador" elevation="2125" latitude="-0.6051111" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="EFBB74F3F27E14217756E8D550E52AAD" box="[1140,1216,657,683]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(
<collectionCode id="9FAC27BFF27E142177F2E8D551052AAD" box="[1232,1312,657,683]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E1421761EE8D551B92AAD" box="[1340,1436,657,683]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
52,
<collectingRegion id="3B797198F27E142173E1E8F255252AD6" box="[195,256,694,720]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217240E8F155E72AC9" box="[354,450,693,719]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E14217126E8F256542AD6" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217126E8F256542AD6" box="[516,625,694,720]" country="Ecuador" county="Ecuador" latitude="-0.6051111" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E142171A9E8F156F82AC9" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142171A9E8F156F82AC9" box="[651,733,693,719]" country="Ecuador" county="Ecuador" latitude="-0.6051111" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<location id="FC62E9A1F27E142171C8E8F1576C2AC9" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142171C8E8F1576C2AC9" box="[746,841,693,719]" country="Ecuador" county="Ecuador" latitude="-0.6051111" municipality="Cosanga" name="Bamboo" pageId="23" pageNumber="24" stateProvince="Napo">Bamboo</location>
hill trail \ &amp; stream,
<quantity id="3E45129FF27E14217763E8F150BF2AD6" box="[1089,1178,693,720]" metricMagnitude="3" metricUnit="m" metricValue="2.125" pageId="23" pageNumber="24" unit="m" value="2125.0">
<elevation id="72905849F27E14217763E8F150BF2AD6" box="[1089,1178,693,720]" metricMagnitude="3" metricUnit="m" metricValue="2.125" pageId="23" pageNumber="24" unit="m" value="2125.0">2125 m</elevation>
</quantity>
, \
<geoCoordinate id="9C89D9BDF27E1421779AE8F1517D2AC9" box="[1208,1368,693,719]" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-0.6051111">S 00º36’18.4”</geoCoordinate>
/
<geoCoordinate id="9C89D9BDF27E14217650E8F255042AF2" pageId="23" pageNumber="24">W \ 77º53’09.0”</geoCoordinate>
,
<date id="8D0399BAF27E1421720DE89D55942AF5" box="[303,433,729,756]" pageId="23" pageNumber="24" value="2009-12-05">
<collectingDate id="9D476052F27E1421720DE89D55942AF5" box="[303,433,729,756]" pageId="23" pageNumber="24" value="2009-12-05">5 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF27E14217123E89D56A42AF5" box="[513,641,729,755]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E142171A3E89D579B2B11" ID-GBIF-Occurrence="2239706656" collectingDate="2009-11-25" collectionCode="DBET" collectorName="L. Borowiec" country="Ecuador" elevation="1950" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="7" specimenCount-male="7" stateProvince="Napo" typeStatus="paratype">
”.
<specimenCount id="EFBB74F3F27E142171BAE89E56D52AF5" box="[664,752,729,756]" pageId="23" pageNumber="24" type="male">7 males</specimenCount>
(
<collectionCode id="9FAC27BFF27E14217023E89D57742AF5" box="[769,849,729,755]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF27E1421704DE89D57D92AF5" box="[879,1020,729,755]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
,
<collectingRegion id="3B797198F27E14217728E89E50622AF2" box="[1034,1095,730,756]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov. \
<collectingMunicipality id="19662500F27E14217780E89D51222AF5" box="[1186,1287,729,755]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
+
<location id="FC62E9A1F27E14217634E89D51A92AF5" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217634E89D51A92AF5" box="[1302,1420,729,755]" country="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\ collecting along the road \
<quantity id="3E45129FF27E142172E1E8B956722B11" box="[451,599,765,791]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="23" pageNumber="24" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">
<elevation id="72905849F27E142172E1E8B956722B11" box="[451,599,765,791]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="23" pageNumber="24" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900–2000m</elevation>
</quantity>
\
<date id="8D0399BAF27E1421714FE8B956D52B11" box="[621,752,765,792]" pageId="23" pageNumber="24" value="2009-11-25">
<collectingDate id="9D476052F27E1421714FE8B956D52B11" box="[621,752,765,792]" pageId="23" pageNumber="24" value="2009-11-25">25 XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E1421700EE8BA579B2B11" box="[812,958,765,792]" pageId="23" pageNumber="24">L. Borowiec</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E1421709CE8B951AA2B3D" ID-GBIF-Occurrence="2239706644" collectingDate="2009-11-30" collectionCode="DBET" collectorName="L. Borowiec" country="Ecuador" county="Cosanga" elevation="2100" location="Rio Aliso" municipality="Yanayacu St." pageId="23" pageNumber="24" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
”. Male (
<collectionCode id="9FAC27BFF27E14217739E8B9504E2B11" box="[1051,1131,765,791]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF27E142177A5E8B951312B11" box="[1159,1300,765,791]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
,
<collectingRegion id="3B797198F27E14217602E8BA51782B1E" box="[1312,1373,766,792]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov. \
<collectingCounty id="1063C7F6F27E14217384E96555232B3D" box="[166,262,801,827]" pageId="23" pageNumber="24">Cosanga</collectingCounty>
vic.,
<collectingMunicipality id="19662500F27E14217266E96655F22B3D" box="[324,471,801,828]" pageId="23" pageNumber="24">Yanayacu St.</collectingMunicipality>
\ -
<location id="FC62E9A1F27E142172DFE965564D2B3D" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142172DFE965564D2B3D" box="[509,616,801,827]" country="Ecuador" county="Cosanga" municipality="Yanayacu St." name="Rio Aliso" pageId="23" pageNumber="24" stateProvince="Napo">Rio Aliso</location>
road,
<quantity id="3E45129FF27E1421718DE96557662B3D" box="[687,835,801,827]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E1421718DE96557662B3D" box="[687,835,801,827]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
\ 00°35’S/77°53’W,
<date id="8D0399BAF27E1421770DE96550972B3D" box="[1071,1202,801,828]" pageId="23" pageNumber="24" value="2009-11-30">
<collectingDate id="9D476052F27E1421770DE96550972B3D" box="[1071,1202,801,828]" pageId="23" pageNumber="24" value="2009-11-30">30 XI 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF27E142177DFE96651AA2B3D" box="[1277,1423,801,828]" pageId="23" pageNumber="24">L. Borowiec</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E142176ADE96551B92B59" ID-GBIF-Occurrence="2239706637" collectingDate="1992-01-03" collectionCode="SEMC" country="Ecuador" county="Monona" elevation="2250" location="Cuenca" municipality="Santiago" pageId="23" pageNumber="24" specimenCount="1" typeStatus="paratype">
”. Male (
<collectionCode id="9FAC27BFF27E142173C3E90155112B59" box="[225,308,837,863]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="23" pageNumber="24" type="Museum">SEMC</collectionCode>
): “
<collectingCountry id="81AAFFEAF27E14217273E90155C52B59" box="[337,480,837,863]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
:
<collectingCounty id="1063C7F6F27E142172C8E90256622B66" box="[490,583,838,864]" pageId="23" pageNumber="24">Monona</collectingCounty>
\
<collectingMunicipality id="19662500F27E1421717DE90156E12B59" box="[607,708,837,863]" pageId="23" pageNumber="24">Santiago</collectingMunicipality>
,
<quantity id="3E45129FF27E142171F2E90157032B59" box="[720,806,837,863]" metricMagnitude="5" metricUnit="m" metricValue="1.41" pageId="23" pageNumber="24" unit="km" value="141.0">141 km</quantity>
E of \
<location id="FC62E9A1F27E14217057E90157EC2B59" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217057E90157EC2B59" box="[885,969,837,863]" country="Ecuador" county="Monona" municipality="Santiago" name="Cuenca" pageId="23" pageNumber="24">Cuenca</location>
nr. rd. to
<location id="FC62E9A1F27E1421771AE90150BA2B59" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E1421771AE90150BA2B59" box="[1080,1183,837,863]" country="Ecuador" county="Monona" municipality="Santiago" name="Arenillas" pageId="23" pageNumber="24">Arenillas</location>
\
<quantity id="3E45129FF27E14217794E901512B2B66" box="[1206,1294,837,864]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="23" pageNumber="24" unit="m" value="2250.0">
<elevation id="72905849F27E14217794E901512B2B66" box="[1206,1294,837,864]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="23" pageNumber="24" unit="m" value="2250.0">2250 m</elevation>
</quantity>
,
<date id="8D0399BAF27E14217638E90151B92B59" box="[1306,1436,837,864]" pageId="23" pageNumber="24" value="1992-01-03">
<collectingDate id="9D476052F27E14217638E90151B92B59" box="[1306,1436,837,864]" pageId="23" pageNumber="24" value="1992-01-03">3 Jan. 1992</collectingDate>
</date>
</materialsCitation>
\
<materialsCitation id="49D5B527F27E14217384E92D50292BA1" ID-GBIF-Occurrence="2239706649" collectingDate="2009-11-23" collectingDateMax="2009-12-17" collectingDateMin="2009-11-23" collectionCode="DBET" collectorName="C. Carlton &amp; R. Leschen &amp; L. Borowiec" country="Ecuador" county="Cosanga" elevation="2100" location="Station" municipality="Yanayacu" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" stateProvince="Napo" typeStatus="paratype">
<collectorName id="5448DAACF27E14217384E92D553B2B85" box="[166,286,873,899]" pageId="23" pageNumber="24">C. Carlton</collectorName>
,
<collectorName id="5448DAACF27E14217208E92E55822B85" box="[298,423,873,900]" pageId="23" pageNumber="24">R. Leschen</collectorName>
\ #104”.
<specimenCount id="EFBB74F3F27E14217133E92D56582B85" box="[529,637,873,899]" pageId="23" pageNumber="24" type="female">2 females</specimenCount>
(
<collectionCode id="9FAC27BFF27E142171AFE92D56F82B85" box="[653,733,873,899]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF27E142171DBE92D57A32B85" box="[761,902,873,899]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
,
<collectingRegion id="3B797198F27E142170B3E92E57EB2B82" box="[913,974,874,900]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov. \
<collectingCounty id="1063C7F6F27E14217706E92D50A12B85" box="[1060,1156,873,899]" pageId="23" pageNumber="24">Cosanga</collectingCounty>
vic.,
<collectingMunicipality id="19662500F27E142177E0E92E510A2B82" box="[1218,1327,874,900]" pageId="23" pageNumber="24">Yanayacu</collectingMunicipality>
\
<location id="FC62E9A1F27E14217664E92D51BD2B85" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217664E92D51BD2B85" box="[1350,1432,873,899]" country="Ecuador" county="Cosanga" municipality="Yanayacu" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E142173B5E9C9550B2BA1" box="[151,302,909,935]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E142173B5E9C9550B2BA1" box="[151,302,909,935]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’ S, 77°53’W, \
<date id="8D0399BAF27E14217167E9C957032BA1" box="[581,806,909,936]" pageId="23" pageNumber="24" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">
<collectingDate id="9D476052F27E14217167E9C957032BA1" box="[581,806,909,936]" pageId="23" pageNumber="24" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">23 XI–17 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF27E14217055E9CA50292BA1" box="[887,1036,909,936]" pageId="23" pageNumber="24">L. Borowiec</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E1421772EE9C9515F2BCD" ID-GBIF-Occurrence="2239706642" collectingDate="2009-12-16" collectionCode="DBET" collectorName="R. Ruta" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" stateProvince="Napo" typeStatus="paratype">
”.
<specimenCount id="EFBB74F3F27E14217700E9C950B52BA1" box="[1058,1168,909,935]" pageId="23" pageNumber="24" type="female">2 females</specimenCount>
(
<collectionCode id="9FAC27BFF27E14217780E9C950D72BA1" box="[1186,1266,909,935]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217632E9C951552BA1" box="[1296,1392,909,935]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
75,
<collectingRegion id="3B797198F27E142173B5E9F654F12BCA" box="[151,212,946,972]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217213E9F555B42BCD" box="[305,401,945,971]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E142172EDE9F656192BCA" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142172EDE9F656192BCA" box="[463,572,946,972]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E14217171E9F556802BCD" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217171E9F556802BCD" box="[595,677,945,971]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E14217192E9F557622BCD" box="[688,839,945,971]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E14217192E9F557622BCD" box="[688,839,945,971]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, \
<date id="8D0399BAF27E14217777E9F550C72BCD" box="[1109,1250,945,972]" pageId="23" pageNumber="24" value="2009-12-16">
<collectingDate id="9D476052F27E14217777E9F550C72BCD" box="[1109,1250,945,972]" pageId="23" pageNumber="24" value="2009-12-16">16 XII 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E1421763CE9F6515F2BCD" box="[1310,1402,945,972]" pageId="23" pageNumber="24">R. Ruta</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E1421765BE9F5562B2C15" ID-GBIF-Occurrence="2239706635" collectingDate="2009-11" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="8" specimenCount-female="8" stateProvince="Napo" typeStatus="paratype">
”.
<specimenCount id="EFBB74F3F27E142176ACE9F554CB2BE9" pageId="23" pageNumber="24" type="female">8 females</specimenCount>
(
<collectionCode id="9FAC27BFF27E142173DCE991556B2BE9" box="[254,334,981,1007]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217249E99155EE2BE9" box="[363,459,981,1007]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
1,
<collectingRegion id="3B797198F27E142172CDE99256092BF6" box="[495,556,982,1008]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E142171A8E99156CF2BE9" box="[650,746,981,1007]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E1421700BE99257B32BF6" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E1421700BE99257B32BF6" box="[809,918,982,1008]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E1421708CE99150252BE9" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E1421708CE99150252BE9" box="[942,1024,981,1007]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
,
<quantity id="3E45129FF27E1421772EE99150872BE9" box="[1036,1186,981,1007]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E1421772EE99150872BE9" box="[1036,1186,981,1007]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, 23–24 \
<date id="8D0399BAF27E142173D1E9BE55762C15" box="[243,339,1017,1044]" pageId="23" pageNumber="24" value="2009-11">
<collectingDate id="9D476052F27E142173D1E9BE55762C15" box="[243,339,1017,1044]" pageId="23" pageNumber="24" value="2009-11">XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E142172ADE9BD562B2C15" box="[399,526,1017,1043]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E1421712FE9BD564E2C31" ID-GBIF-Occurrence="2239706625" collectingDate="2009-11-25" collectionCode="DBET" collectorName="R. Ruta" country="Ecuador" county="Ecuador" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="6" specimenCount-female="6" stateProvince="Napo" typeStatus="paratype">
”.
<specimenCount id="EFBB74F3F27E14217100E9BD56AB2C15" box="[546,654,1017,1043]" pageId="23" pageNumber="24" type="female">6 females</specimenCount>
(
<collectionCode id="9FAC27BFF27E142171BCE9BD56CB2C15" box="[670,750,1017,1043]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217028E9BD574F2C15" box="[778,874,1017,1043]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
4,
<collectingRegion id="3B797198F27E142170AFE9BE57EF2C12" box="[909,970,1018,1044]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E14217705E9BD50A82C15" box="[1063,1165,1017,1043]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
+
<location id="FC62E9A1F27E142177BEE9BD51362C15" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142177BEE9BD51362C15" box="[1180,1299,1017,1043]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
, \ collecting along the road, \
<date id="8D0399BAF27E14217271EE5955F02C31" box="[339,469,1053,1080]" pageId="23" pageNumber="24" value="2009-11-25">
<collectingDate id="9D476052F27E14217271EE5955F02C31" box="[339,469,1053,1080]" pageId="23" pageNumber="24" value="2009-11-25">25 XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF27E1421712DEE5A564E2C31" box="[527,619,1053,1080]" pageId="23" pageNumber="24">R. Ruta</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E14217148EE5956BC2C5D" ID-GBIF-Occurrence="2239706617" collectingDate="3 XII" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" county="Ecuador" location="Antisana" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="4" specimenCount-female="4" stateProvince="Napo" typeStatus="paratype">
”.
<specimenCount id="EFBB74F3F27E1421715DEE5956CE2C31" box="[639,747,1053,1079]" pageId="23" pageNumber="24" type="female">4 females</specimenCount>
(
<collectionCode id="9FAC27BFF27E142171DBEE59576C2C31" box="[761,841,1053,1079]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E14217047EE5957E02C31" box="[869,965,1053,1079]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
41,
<collectingRegion id="3B797198F27E142170D4EE5A50162C3E" box="[1014,1075,1054,1080]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E142177ACEE5950CB2C31" box="[1166,1262,1053,1079]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E14217608EE5951AB2C31" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217608EE5951AB2C31" box="[1322,1422,1053,1079]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Antisana" pageId="23" pageNumber="24" stateProvince="Napo">Antisana</location>
\ trail+
<location id="FC62E9A1F27E142173FAEE0555752C5D" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142173FAEE0555752C5D" box="[216,336,1089,1115]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
,
<date id="8D0399BAF27E14217278EE0555B02C5A" box="[346,405,1089,1116]" pageId="23" pageNumber="24">
<collectingDate id="9D476052F27E14217278EE0555B02C5A" box="[346,405,1089,1116]" pageId="23" pageNumber="24">3 XII</collectingDate>
</date>
\ 2009, leg.
<collectorName id="5448DAACF27E1421713EEE0556BC2C5D" box="[540,665,1089,1115]" pageId="23" pageNumber="24">Rafał Ruta</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E142171BBEE0555E22C79" ID-GBIF-Occurrence="2239706626" collectingDate="2009-11-30" collectionCode="DBET" collectorName="Lech Borowiec" country="Ecuador" location="Aliso" municipality="Yanayacu-Rio" pageId="23" pageNumber="24" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
”.
<specimenCount id="EFBB74F3F27E1421718EEE0557322C5D" box="[684,791,1089,1115]" pageId="23" pageNumber="24" type="female">3 females</specimenCount>
(
<collectionCode id="9FAC27BFF27E14217006EE0557512C5D" box="[804,884,1089,1115]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF27E142170ADEE05503C2C5D" box="[911,1049,1089,1115]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
\ road
<collectingMunicipality id="19662500F27E14217741EE0551272C5D" box="[1123,1282,1089,1115]" pageId="23" pageNumber="24">Yanayacu-Rio</collectingMunicipality>
\
<location id="FC62E9A1F27E14217636EE0551762C5D" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217636EE0551762C5D" box="[1300,1363,1089,1115]" country="Ecuador" municipality="Yanayacu-Rio" name="Aliso" pageId="23" pageNumber="24">Aliso</location>
,
<date id="8D0399BAF27E1421767FEE0554EA2C79" pageId="23" pageNumber="24" value="2009-11-30">
<collectingDate id="9D476052F27E1421767FEE0554EA2C79" pageId="23" pageNumber="24" value="2009-11-30">30 XI 2009</collectingDate>
</date>
\ leg.
<collectorName id="5448DAACF27E14217237EE2155E22C79" box="[277,455,1125,1151]" pageId="23" pageNumber="24">Lech Borowiec</collectorName>
</materialsCitation>
<materialsCitation id="49D5B527F27E142172E5EE2157CA2CA5" ID-GBIF-Occurrence="2239706609" collectingDate="2009-12-09" collectionCode="DBET" country="Ecuador" county="Ecuador" elevation="2100" location="Yanayacu" municipality="Cosanga" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
”. Female (
<collectionCode id="9FAC27BFF27E1421711FEE2156A82C79" box="[573,653,1125,1151]" pageId="23" pageNumber="24">DBET</collectionCode>
): “
<collectingCounty id="1063C7F6F27E1421718BEE21572C2C79" box="[681,777,1125,1151]" pageId="23" pageNumber="24">Ecuador</collectingCounty>
58,
<collectingRegion id="3B797198F27E14217018EE2257522C86" box="[826,887,1126,1152]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
prov., \
<collectingMunicipality id="19662500F27E142170F6EE2150112C79" box="[980,1076,1125,1151]" pageId="23" pageNumber="24">Cosanga</collectingMunicipality>
vic.,
<location id="FC62E9A1F27E14217753EE2250FB2C86" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E14217753EE2250FB2C86" box="[1137,1246,1126,1152]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="23" pageNumber="24" stateProvince="Napo">Yanayacu</location>
\
<location id="FC62E9A1F27E142177D6EE2151632C79" LSID="urn:lsid:plazi:treatment:71140E6CF27E142F73B5EADD507228D1:FC62E9A1F27E142177D6EE2151632C79" box="[1268,1350,1125,1151]" country="Ecuador" county="Ecuador" municipality="Cosanga" name="Station" pageId="23" pageNumber="24" stateProvince="Napo">Station</location>
, stream trail, \
<quantity id="3E45129FF27E142173C3EECD555D2CA5" box="[225,376,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F27E142173C3EECD555D2CA5" box="[225,376,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, 00°35’S \ 77°53’W,
<date id="8D0399BAF27E14217145EECD56C02CA5" box="[615,741,1161,1188]" pageId="23" pageNumber="24" value="2009-12-09">
<collectingDate id="9D476052F27E14217145EECD56C02CA5" box="[615,741,1161,1188]" pageId="23" pageNumber="24" value="2009-12-09">9 XII 2009</collectingDate>
</date>
, leg. \ Lech Borowiec”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27E142173E5EEE9565D2D35" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="F902BF7AF27E142173E5EEE9565D2D35" blockId="23.[151,1437,261,1980]" pageId="23" pageNumber="24">
<emphasis id="CBC96368F27E142173E5EEE955672CCE" bold="true" box="[199,322,1197,1224]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
The largest (TL 2.8–3.6) known species of
<taxonomicName id="3EBDC4F9F27E1421701FEEE957E92CC1" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[829,972,1197,1223]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F27E1421701FEEE957E92CC1" box="[829,972,1197,1223]" italics="true" pageId="23" pageNumber="24">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F27E142170F7EEEA50652CCE" box="[981,1088,1198,1224]" pageId="23" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
, with dorsal pattern extremely variable, sometimes with contrasting orange spots. Penis is very long and narrow, apical plate of tergite VIII has relatively short median process with deep V-shaped notch (process is M-shaped), flanked with shallow emarginations which are not covered with denticles.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F27E142E73E5EF7956EB2F88" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="F902BF7AF27E142173E5EF7955B92E9A" blockId="23.[151,1437,261,1980]" pageId="23" pageNumber="24">
<emphasis id="CBC96368F27E142173E5EF79557C2D5E" bold="true" box="[199,345,1341,1368]" pageId="23" pageNumber="24">Description.</emphasis>
<typeStatus id="260601D8F27E14217243EF7955E92D51" box="[353,460,1341,1367]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
, male. Body oval, convex, covered with brownish, suberect setae. Body black, apical portion of elytra with orange adsutural spot, antennae and mouthparts brownish, legs yellowish. Head broad, 1.6× wider than width of interocular space, with distinct but fine punctation, punctures small, not granulate, separated by a 2.0–4.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, not granulate punctation, punctures separated by ca. 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest before middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 0.5–1.0× diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF27E142173E5ECE157142FBD" blockId="23.[151,1437,261,1980]" pageId="23" pageNumber="24">
Male genitalia. Penis (L
<quantity id="3E45129FF27E142172F5ECE156182EC6" box="[471,573,1701,1728]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="23" pageNumber="24" unit="mm" value="0.92">0.92 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27E14217165ECE256BD2EB9" box="[583,664,1701,1728]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="555" value="-0.09">
W
<quantity id="3E45129FF27E14217145ECE156BD2EB9" box="[615,664,1701,1727]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="24" unit="mm" value="0.09">0.09</quantity>
</geoCoordinate>
mm, Fig. 18A) very long and narrow, widened in basal portion, para- meroids short, narrow, rounded at apices, curved, trigonium subtriangular, narrow, pointed at apex. Tegmen (L
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,
<geoCoordinate id="9C89D9BDF27E142173F3ECAA55062F01" box="[209,291,1773,1800]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="555" value="-0.24">
W
<quantity id="3E45129FF27E142173D0ECA955062F01" box="[242,291,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="24" unit="mm" value="0.24">0.24</quantity>
</geoCoordinate>
mm, Fig. 18B) very long, sides subparallel, apex with short lateral lobes, covered with setae. Sternite IX (L
<quantity id="3E45129FF27E142173FEED5555672F2A" box="[220,322,1809,1836]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="23" pageNumber="24" unit="mm" value="0.38">0.38 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27E1421726CED5655852F2D" box="[334,416,1809,1836]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="555" value="-0.26">
W
<quantity id="3E45129FF27E1421724DED5555852F2D" box="[367,416,1809,1835]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="23" pageNumber="24" unit="mm" value="0.26">0.26</quantity>
</geoCoordinate>
mm, Fig. 18C) subtriangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
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,
<geoCoordinate id="9C89D9BDF27E14217013ED7257A02F49" box="[817,901,1845,1872]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="555" value="-0.41">
W
<quantity id="3E45129FF27E14217076ED7157A02F49" box="[852,901,1845,1871]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="23" pageNumber="24" unit="mm" value="0.41">0.41</quantity>
</geoCoordinate>
mm, Fig. 18D) with short apical plate and two short and wide denticles forming M-shaped emargination in central portion of apical margin, apical margin subtly setose, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF27E142170D4ED3950792F9E" box="[1014,1116,1917,1944]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="23" pageNumber="24" unit="mm" value="0.42">0.42 mm</quantity>
,
<geoCoordinate id="9C89D9BDF27E14217745ED3A509C2F91" box="[1127,1209,1917,1944]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="555" value="-0.31">
W
<quantity id="3E45129FF27E142177AAED39509C2F91" box="[1160,1209,1917,1943]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="23" pageNumber="24" unit="mm" value="0.31">0.31</quantity>
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mm, Fig. 18E) with membranous apical plate, apodemes as long as apical plate.
</paragraph>
<paragraph id="F902BF7AF271142E73B5EADD503028B5" blockId="24.[151,1045,153,179]" box="[151,1045,153,179]" pageId="24" pageNumber="25">Variability. Dorsal pattern variable. The following colour patterns were noticed:</paragraph>
<paragraph id="F902BF7AF271142E73E5EC3956412F22" blockId="24.[151,1437,1660,2040]" pageId="24" pageNumber="25">
Female. Body (
<figureCitation id="6186A3FFF271142E7250EC3955E72E91" box="[370,450,1661,1687]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="24" pageNumber="25">Fig. 1P</figureCitation>
) dark brown to black, sometimes with lighter, orange suture, TL (n = 5) 3.07–3.67 (3.39) mm. Apical portion of each elytron with two shallow concavities covered with hexagonal microsculpture (
<figureCitation id="6186A3FFF271142E7618ECE451AA2EBD" box="[1338,1423,1696,1723]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="24" pageNumber="25">Fig. 7F</figureCitation>
), the one situated closer to suture is a bit smaller (
<figureCitation id="6186A3FFF271142E718AEC8056CB2ED8" box="[680,750,1732,1758]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="24" pageNumber="25">Figs 7</figureCitation>
E–F). Gonocoxites (
<figureCitation id="6186A3FFF271142E70ECEC8050052ED8" box="[974,1056,1732,1758]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="24" pageNumber="25">Figs 10</figureCitation>
E–F) broadly triangular, gonostyli subapical, bursal sclerite (
<figureCitation id="6186A3FFF271142E729EECA356342F07" box="[444,529,1767,1793]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="24" pageNumber="25">Figs 10</figureCitation>
G–H, L
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,
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W
<quantity id="3E45129FF271142E7021ECA357112F07" box="[771,820,1767,1793]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="mm" value="0.1">0.10</quantity>
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mm) elongated, anterior portion wide, with digitiform outgrowth and rod-like posterior portion.
</paragraph>
<paragraph id="F902BF7AF271142E73E5ED6956EB2F88" blockId="24.[151,1437,1660,2040]" pageId="24" pageNumber="25">
<emphasis id="CBC96368F271142E73E5ED6955DB2F41" bold="true" box="[199,510,1836,1863]" pageId="24" pageNumber="25">Measurements and ratios.</emphasis>
Males (n = 10) TL 2.77–3.60 (3.18) mm, PL 0.50–0.70 (0.60) mm, PW 1.17–1.60 (1.33) mm, EL 2.50–3.20 (2.75) mm, EW 1.73–2.30 (2.00) mm, TL/EW 1.50–1.70 (1.59), PW/PL 2.20–2.30 (2.23), EL/EW 1.30–1.50 (1.38), EL/PL 4.30–5.00 (4.63).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F271142E73E5EDD257642FB7" box="[199,833,1942,1969]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="F902BF7AF271142E73E5EDD257642FB7" blockId="24.[151,1437,1660,2040]" box="[199,833,1942,1969]" pageId="24" pageNumber="25">
<emphasis id="CBC96368F271142E73E5EDD255442FB7" bold="true" box="[199,353,1942,1969]" pageId="24" pageNumber="25">Distribution.</emphasis>
Known from several localities in
<collectingCountry id="81AAFFEAF271142E71FFEDD357192FB7" box="[733,828,1943,1969]" name="Ecuador" pageId="24" pageNumber="25">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F271142E73E5EDFE55AF2FFE" pageId="24" pageNumber="25" type="etymology">
<paragraph id="F902BF7AF271142E73E5EDFE55AF2FFE" blockId="24.[151,1437,1660,2040]" pageId="24" pageNumber="25">
<emphasis id="CBC96368F271142E73E5EDFE556B2FD2" bold="true" box="[199,334,1978,2004]" pageId="24" pageNumber="25">Etymology.</emphasis>
Named after Yanayacu Biological Station (http://www.yanayacu.org/), where the
<typeStatus id="260601D8F271142E77D2EDFF51052FD3" box="[1264,1312,1979,2005]" pageId="24" pageNumber="25">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F270142F73E5EADD507228D1" pageId="25" pageNumber="26" type="discussion">
<paragraph id="F902BF7AF270142F73E5EADD507228D1" blockId="25.[151,1436,152,215]" pageId="25" pageNumber="26">
<emphasis id="CBC96368F270142F73E5EADD551E28B5" bold="true" box="[199,315,153,179]" pageId="25" pageNumber="26">Remarks.</emphasis>
Numerous specimens were collected on trees densely covered with lichens, growing along the road near Yanayacu Biological Station, at the altitude of about
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a.s.l. (
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A–B).
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(Figs 2E, 6)
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obtortum
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Assing, 2015: 1225.
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: Shavrin and Smetana, 2016: 228.
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<paragraph id="B1474549344F5F22A828E5C1E36A399B" pageId="2" pageNumber="57">Material studied.</paragraph>
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China: Sichuan: 1 ♂, Xiaojin County, Jiajin Shan,
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,
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, 2490 m, 30.VII.2016, Zhou, Jiang, Liu &amp; Gao leg. (SNUC).
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<paragraph id="3B7B38712D6C8073A0F1E10F5AE4389A" pageId="2" pageNumber="57">Comment.</paragraph>
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The original description was based on two type specimens from Min Shan, Gansu (
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). It is recorded from Sichuan for the first time. For illustrations of
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see figure 2E and figures 7-8, 15-18, 23-25 in
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. The specimen was sifted from leaf litter near the mountain track in mixed deciduous forests at an altitude of 2490 m (Fig. 6).
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The key given by
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needs to be partially modified by the above discussion of diagnostic characteristics and the incorporation of
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. These partial modifications are mostly related to the South American genera, because specimens of the North American species were not available and are out of the scope of the present work. The item in the key corresponding to the genus
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2009), Di Iorio
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1. Ectospermalege “hat-shaped”, right ventral near lateral margin between sixth and seventh segments. Sides of pronotum with long bristles; posterlateral angles of pronotum rounded; females with apical tufts of hairs in front and middle tibiae. Western
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<taxonomicName id="3EBD4D63E8037C65FC710A5FFB97E802" ID-CoL="GQX" box="[990,1086,818,841]" class="Aves" family="Strigidae" kingdom="Animalia" order="Strigiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Strigidae</taxonomicName>
(secondary)
<emphasis id="CBC9EAF2E8037C65FB650A5FFA32E802" box="[1226,1435,818,841]" italics="true" pageId="15" pageNumber="16">
........
<taxonomicName id="3EBD4D63E8037C65FAAB0A5FFA32E802" ID-CoL="7P5SQ" box="[1284,1435,818,841]" class="Insecta" family="Cimicidae" genus="Hesperocimex" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Hesperocimex</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380A3FFA32E822" blockId="15.[151,1437,754,1737]" box="[151,1435,850,873]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380A3FFA32E822" box="[151,1435,850,873]" pageId="15" pageNumber="16">1’. Ectospermalege dorsal either at middle or on right side on sixth or seventh segments .............................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="42492E45E8037C65FF380A1FFA32E882" pageId="15" pageNumber="16">
<paragraph id="F90236E0E8037C65FF380A1FFA32E8C2" blockId="15.[151,1437,754,1737]" box="[151,1435,882,905]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380A1FFA32E8C2" box="[151,1435,882,905]" pageId="15" pageNumber="16">2. Posterior margin of pronotum slightly concave or convex........................................................................................... 3</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380AFFFA32E882" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380AFFFA32E882" pageId="15" pageNumber="16">
2’. Posterior margin of pronotum strongly convex (see fig.
<quantity id="3E459B05E8037C65FCB90AFFFC97E8E2" box="[790,830,914,937]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="16" unit="in" value="1.0">1 in</quantity>
Hernandez Triana &amp; de la
<bibRefCitation id="9D2C4B11E8037C65FBF30AFFFB66E8E2" author="Cruz" box="[1116,1231,914,937]" pageId="15" pageNumber="16" refString="Hernandez Triana, L. M. &amp; de la Cruz, J. (1994) Nuevo genero nidicola de Cimicidae (Heteroptera) de Cuba. Caribbean Journal of Science, 30 (3 - 4), 198 - 202." type="journal article" year="1994">Cruz 1994</bibRefCitation>
).
<collectingCountry id="81AA7670E8037C65FB4F0AFFFAB2E8E2" box="[1248,1307,914,937]" name="Cuba" pageId="15" pageNumber="16">Cuba</collectingCountry>
. Host:
<taxonomicName id="3EBD4D63E8037C65FACA0AFFFEA8E882" ID-CoL="6L5" class="Aves" family="Apodidae" kingdom="Animalia" order="Apodiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Apodidae</taxonomicName>
<emphasis id="CBC9EAF2E8037C65FEAB0ADFFA32E882" box="[260,1435,946,969]" italics="true" pageId="15" pageNumber="16">............................................................................................................................................................. Alayocimex</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="42492E45E8037C65FF380ABFFA35EF22" pageId="15" pageNumber="16">
<paragraph id="F90236E0E8037C65FF380ABFFA3CEF42" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380ABFFA3CEF42" pageId="15" pageNumber="16">
3. Posterolateral angles of pronotum more or less angulate, with two longer bristles (Lb1 + Lb2 or Lb2 + Lb3) (
<figureCitation id="61862A65E8037C65FAC90ABFFF4BEF42" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="15" pageNumber="16">Figs. 11</figureCitation>
,
<figureCitation id="61862A65E8037C65FF410A9FFEA3EF42" box="[238,266,1010,1033]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,272,1793,1817]" captionTargetBox="[167,1429,571,1778]" captionTargetId="figure@11.[155,1440,555,1789]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Acanthocrios furnarii: 13, antennae, head, and pronotum (female); 14, pronotal margin; 15, male genitalia (laboratory culture, coll. Usinger)." httpUri="https://zenodo.org/record/200082/files/figure.png" pageId="15" pageNumber="16">15</figureCitation>
,
<figureCitation id="61862A65E8037C65FEB90A9FFE9AEF42" box="[278,307,1010,1033]" captionStart="FIGURES 16 – 20" captionStartId="13.[151,275,1649,1673]" captionTargetBox="[159,1415,528,1630]" captionTargetId="figure@13.[147,1432,518,1644]" captionTargetPageId="13" captionText="FIGURES 16 – 20. Psitticimex uritui: 16, antennae, head, and pronotum (female); 17, pronotal margin; 18, male genitalia; 19, female spermalege; 20, tibia III." httpUri="https://zenodo.org/record/200084/files/figure.png" pageId="15" pageNumber="16">18</figureCitation>
); row of bristles in lateral margins of pronotum does not continues to posterior margin .......................... 4
</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380D7FFA35EF22" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380D7FFA35EF22" pageId="15" pageNumber="16">
3’. Posterolateral angles of pronotum rounded; Lb2 distinguisable by its greater length and its more internal base (
<figureCitation id="61862A65E8037C65FAC90D7FFF5BEF02" captionStart="FIGURES 2 – 7" captionStartId="3.[151,269,1937,1961]" captionTargetBox="[158,1427,460,1896]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Cyanolicimex patagonus: 2, antennae, head, and pronotum (female Holotype); 3 – 4, pronotal margin; 5, spermalege (female Holotype); 6, tibia III; 7, male genitalia (Paratype)." httpUri="https://zenodo.org/record/200079/files/figure.png" pageId="15" pageNumber="16">Figs. 5–6</figureCitation>
); row of bristles in lateral margins of pronotum continue to posterior margin (
<figureCitation id="61862A65E8037C65FBE80D5FFB23EF02" box="[1095,1162,1074,1097]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,269,1937,1961]" captionTargetBox="[158,1427,460,1896]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Cyanolicimex patagonus: 2, antennae, head, and pronotum (female Holotype); 3 – 4, pronotal margin; 5, spermalege (female Holotype); 6, tibia III; 7, male genitalia (Paratype)." httpUri="https://zenodo.org/record/200079/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
).
<collectingCountry id="81AA7670E8037C65FB300D5FFAA0EF02" box="[1183,1289,1074,1097]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
(Río Negro). Host:
<taxonomicName id="3EBD4D63E8037C65FEA80D3FFED3EF22" ID-CoL="FBD" box="[263,378,1106,1129]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
(
<taxonomicName id="3EBD4D63E8037C65FE250D3FFD2BEF22" ID-CoL="32NN5" box="[394,642,1106,1129]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="patagonus">
<emphasis id="CBC9EAF2E8037C65FE250D3FFD2BEF22" box="[394,642,1106,1129]" italics="true" pageId="15" pageNumber="16">Cyanoliseus patagonus</emphasis>
</taxonomicName>
)
<emphasis id="CBC9EAF2E8037C65FD230D3FFA35EF22" box="[652,1436,1106,1129]" italics="true" pageId="15" pageNumber="16">............................................................................................... Cyanolicimex</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="42492E45E8037C65FF380D1FFA3CEFE2" pageId="15" pageNumber="16">
<paragraph id="F90236E0E8037C65FF380D1FFA35EFC2" blockId="15.[151,1437,754,1737]" box="[151,1436,1138,1161]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380D1FFA35EFC2" box="[151,1436,1138,1161]" pageId="15" pageNumber="16">
4. Abdominal tergites in females strongly asymmetrical.
<collectingCountry id="81AA7670E8037C65FCA40D1FFC34EFC2" box="[779,925,1138,1161]" name="United States of America" pageId="15" pageNumber="16">United States</collectingCountry>
. Host:
<taxonomicName id="3EBD4D63E8037C65FC470D1FFBE6EFC2" ID-CoL="6L5" box="[1000,1103,1138,1161]" class="Aves" family="Apodidae" kingdom="Animalia" order="Apodiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Apodidae</taxonomicName>
<emphasis id="CBC9EAF2E8037C65FBF70D1FFA35EFC2" box="[1112,1436,1138,1161]" italics="true" pageId="15" pageNumber="16">
..........................
<taxonomicName id="3EBD4D63E8037C65FAA70D1FFA35EFC2" ID-CoL="7Q3HJ" box="[1288,1436,1138,1161]" class="Insecta" family="Cimicidae" genus="Synxenoderus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Synxenoderus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380DFFFA3CEFE2" blockId="15.[151,1437,754,1737]" box="[151,1429,1170,1193]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380DFFFA3CEFE2" box="[151,1429,1170,1193]" pageId="15" pageNumber="16">4’. Abdominal tergites not asymmetrical at right margin and not at all in males ............................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="42492E45E8037C65FF380DDFFA3CEE02" pageId="15" pageNumber="16">
<paragraph id="F90236E0E8037C65FF380DDFFA35EE42" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380DDFFA35EE42" pageId="15" pageNumber="16">
5. Rostrum long (&gt;
<quantity id="3E459B05E8037C65FE2C0DDFFE6CEF82" box="[387,453,1202,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
), reaching beyond apices of middle coxae (pinned specimens); A2 subequal to IS; spermalege mid-dorsal.
<collectingCountry id="81AA7670E8037C65FED50DBFFDA3EFA2" box="[378,522,1234,1257]" name="United States of America" pageId="15" pageNumber="16">United States</collectingCountry>
;
<collectingCountry id="81AA7670E8037C65FDBA0DBFFDC3EFA2" box="[533,618,1234,1257]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
. Hosts: raptor birds [
<taxonomicName id="3EBD4D63E8037C65FCEB0DBFFC67EFA2" ID-CoL="5W6" box="[836,974,1234,1257]" class="Aves" family="Accipitridae" kingdom="Animalia" order="Accipitriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Accipitridae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FC770DBFFBFFEFA2" ID-CoL="7SX" box="[984,1110,1234,1257]" class="Aves" family="Cathartidae" kingdom="Animalia" order="Accipitriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Cathartidae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FBCF0DBFFB71EFA2" ID-CoL="623R8" box="[1120,1240,1234,1257]" class="Aves" family="Falconidae" kingdom="Animalia" order="Falconiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Falconidae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FB4E0DBFFAEAEFA2" ID-CoL="GQX" box="[1249,1347,1234,1257]" class="Aves" family="Strigidae" kingdom="Animalia" order="Strigiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Strigidae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FAE20DBFFF46EE42" ID-CoL="HMY" class="Aves" family="Tytonidae" kingdom="Animalia" order="Strigiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Tytonidae</taxonomicName>
] (primary);
<taxonomicName id="3EBD4D63E8037C65FEC00D9FFE47EE42" ID-CoL="625WG" box="[367,494,1266,1289]" class="Aves" family="Phasianidae" kingdom="Animalia" order="Galliformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Phasianidae</taxonomicName>
(secondary)
<emphasis id="CBC9EAF2E8037C65FDD60D9FFA35EE42" box="[633,1436,1266,1289]" italics="true" pageId="15" pageNumber="16">
...............................................................................................
<taxonomicName id="3EBD4D63E8037C65FB570D9FFA35EE42" ID-CoL="7P53J" box="[1272,1436,1266,1289]" class="Insecta" family="Cimicidae" genus="Haematosiphon" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Haematosiphon</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380C7FFA3CEE02" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380C7FFA3CEE02" pageId="15" pageNumber="16">
5’. Rostrum shorter (&lt;
<quantity id="3E459B05E8037C65FE350C7FFE75EE62" box="[410,476,1298,1321]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
), reaching to or a little beyond apices of front coxae (in pinned and also on slide-mounted specimens); A2 &lt;pIS; spermalege opening on right side ......................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="42492E45E8037C65FF380C3FFA35ED42" pageId="15" pageNumber="16">
<paragraph id="F90236E0E8037C65FF380C3FFA32EEC2" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380C3FFA32EEC2" pageId="15" pageNumber="16">
6. Maximal width of pronotum at middle of pronotal length; lateral margins converging anteriorly and posteriorly (
<figureCitation id="61862A65E8037C65FF600C1FFE81EEC2" box="[207,296,1394,1417]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,272,1793,1817]" captionTargetBox="[167,1429,571,1778]" captionTargetId="figure@11.[155,1440,555,1789]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Acanthocrios furnarii: 13, antennae, head, and pronotum (female); 14, pronotal margin; 15, male genitalia (laboratory culture, coll. Usinger)." httpUri="https://zenodo.org/record/200082/files/figure.png" pageId="15" pageNumber="16">Figs. 14</figureCitation>
,
<figureCitation id="61862A65E8037C65FE9C0C1FFEF9EEC2" box="[307,336,1394,1417]" captionStart="FIGURES 16 – 20" captionStartId="13.[151,275,1649,1673]" captionTargetBox="[159,1415,528,1630]" captionTargetId="figure@13.[147,1432,518,1644]" captionTargetPageId="13" captionText="FIGURES 16 – 20. Psitticimex uritui: 16, antennae, head, and pronotum (female); 17, pronotal margin; 18, male genitalia; 19, female spermalege; 20, tibia III." httpUri="https://zenodo.org/record/200084/files/figure.png" pageId="15" pageNumber="16">17</figureCitation>
); tuft of hairs in apices of front and middle tibiae in both sexes ............................................................ 7
</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380CFFFA35ED42" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380CFFFA35ED42" pageId="15" pageNumber="16">
6’. Pronotum widest at base; lateral margins converging anteriorly (
<figureCitation id="61862A65E8037C65FCD80CFFFC6EEEE2" box="[887,967,1426,1449]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
); Lb1 indistinguisable (
<figureCitation id="61862A65E8037C65FB6A0CFFFABAEEE2" box="[1221,1299,1426,1449]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
); with a first posterior long bristle (Lb3) respect to Lb2; both bristles nearly 9 times pronotal length; paramere not exceeding margin of genital segment (
<figureCitation id="61862A65E8037C65FE670CBFFDB1EEA2" box="[456,536,1490,1513]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,272,1793,1817]" captionTargetBox="[167,1429,571,1778]" captionTargetId="figure@11.[155,1440,555,1789]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Acanthocrios furnarii: 13, antennae, head, and pronotum (female); 14, pronotal margin; 15, male genitalia (laboratory culture, coll. Usinger)." httpUri="https://zenodo.org/record/200082/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
); females with apical tuft of hairs only in front tibiae.
<collectingCountry id="81AA7670E8037C65FBC80CBFFB35EEA2" box="[1127,1180,1490,1513]" name="United States of America" pageId="15" pageNumber="16">USA</collectingCountry>
(introduced?);
<collectingCountry id="81AA7670E8037C65FAEC0CBFFA31EEA2" box="[1347,1432,1490,1513]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
;
<collectingCountry id="81AA7670E8037C65FF680C9FFEA5ED42" box="[199,268,1522,1545]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
;
<collectingCountry id="81AA7670E8037C65FEB90C9FFE2DED42" box="[278,388,1522,1545]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
. Hosts:
<taxonomicName id="3EBD4D63E8037C65FE760C9FFDCCED42" ID-CoL="B3Y" box="[473,613,1522,1545]" class="Aves" family="Hirundinidae" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Hirundinidae</taxonomicName>
(primary),
<taxonomicName id="3EBD4D63E8037C65FD700C9FFCFAED42" ID-CoL="FBD" box="[735,851,1522,1545]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
(?);
<taxonomicName id="3EBD4D63E8037C65FC2A0C9FFBADED42" ID-CoL="625WG" box="[901,1028,1522,1545]" class="Aves" family="Phasianidae" kingdom="Animalia" order="Galliformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Phasianidae</taxonomicName>
(secondary)
<emphasis id="CBC9EAF2E8037C65FB3B0C9FFA35ED42" box="[1172,1436,1522,1545]" italics="true" pageId="15" pageNumber="16">
..................
<taxonomicName id="3EBD4D63E8037C65FABD0C9FFA35ED42" ID-CoL="7PNP8" box="[1298,1436,1522,1545]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Ornithocoris</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="42492E45E8037C65FF380F7FFA35ED82" pageId="15" pageNumber="16">
<paragraph id="F90236E0E8037C65FF380F7FFA35ED22" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380F7FFA35ED22" pageId="15" pageNumber="16">
7. A2&gt; aIS (
<figureCitation id="61862A65E8037C65FE960F7FFE20ED62" box="[313,393,1554,1577]" captionStart="FIGURES 16 – 20" captionStartId="13.[151,275,1649,1673]" captionTargetBox="[159,1415,528,1630]" captionTargetId="figure@13.[147,1432,518,1644]" captionTargetPageId="13" captionText="FIGURES 16 – 20. Psitticimex uritui: 16, antennae, head, and pronotum (female); 17, pronotal margin; 18, male genitalia; 19, female spermalege; 20, tibia III." httpUri="https://zenodo.org/record/200084/files/figure.png" pageId="15" pageNumber="16">Fig. 17</figureCitation>
); ectospermalege on anterior margin of a long internal lobe produced forward from anterior margin of seventh tergite at middle (
<figureCitation id="61862A65E8037C65FD810F5FFD28ED02" box="[558,641,1586,1609]" captionStart="FIGURES 16 – 20" captionStartId="13.[151,275,1649,1673]" captionTargetBox="[159,1415,528,1630]" captionTargetId="figure@13.[147,1432,518,1644]" captionTargetPageId="13" captionText="FIGURES 16 – 20. Psitticimex uritui: 16, antennae, head, and pronotum (female); 17, pronotal margin; 18, male genitalia; 19, female spermalege; 20, tibia III." httpUri="https://zenodo.org/record/200084/files/figure.png" pageId="15" pageNumber="16">Fig. 20</figureCitation>
); paramere long, exceeding side of genital segment (
<figureCitation id="61862A65E8037C65FB6B0F5FFABCED02" box="[1220,1301,1586,1609]" captionStart="FIGURES 16 – 20" captionStartId="13.[151,275,1649,1673]" captionTargetBox="[159,1415,528,1630]" captionTargetId="figure@13.[147,1432,518,1644]" captionTargetPageId="13" captionText="FIGURES 16 – 20. Psitticimex uritui: 16, antennae, head, and pronotum (female); 17, pronotal margin; 18, male genitalia; 19, female spermalege; 20, tibia III." httpUri="https://zenodo.org/record/200084/files/figure.png" pageId="15" pageNumber="16">Fig. 19</figureCitation>
).
<collectingCountry id="81AA7670E8037C65FA840F5FFA30ED02" box="[1323,1433,1586,1609]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
. Host:
<taxonomicName id="3EBD4D63E8037C65FEA80F3FFED3ED22" ID-CoL="FBD" box="[263,378,1618,1641]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
[
<taxonomicName id="3EBD4D63E8037C65FE250F3FFDC6ED22" ID-CoL="4532H" box="[394,623,1618,1641]" class="Aves" family="Psittacidae" genus="Myiopsitta" kingdom="Animalia" order="Psittaciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="monachus">
<emphasis id="CBC9EAF2E8037C65FE250F3FFDC6ED22" box="[394,623,1618,1641]" italics="true" pageId="15" pageNumber="16">Myiopsitta monachus</emphasis>
</taxonomicName>
] (primary)
<emphasis id="CBC9EAF2E8037C65FD450F3FFA35ED22" box="[746,1436,1618,1641]" italics="true" pageId="15" pageNumber="16">..................................................................................... Psitticimex</emphasis>
</keyLead>
</paragraph>
<paragraph id="F90236E0E8037C65FF380F1FFA35ED82" blockId="15.[151,1437,754,1737]" pageId="15" pageNumber="16">
<keyLead id="424C95D5E8037C65FF380F1FFA35ED82" pageId="15" pageNumber="16">
7’. A2 &lt;aIS (
<figureCitation id="61862A65E8037C65FE900F1FFE39EDC2" box="[319,400,1650,1673]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,272,1793,1817]" captionTargetBox="[167,1429,571,1778]" captionTargetId="figure@11.[155,1440,555,1789]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Acanthocrios furnarii: 13, antennae, head, and pronotum (female); 14, pronotal margin; 15, male genitalia (laboratory culture, coll. Usinger)." httpUri="https://zenodo.org/record/200082/files/figure.png" pageId="15" pageNumber="16">Fig. 14</figureCitation>
); ectospermalege not as above; paramere just beyond the margin of genital segment (
<figureCitation id="61862A65E8037C65FA910F1FFA39EDC2" box="[1342,1424,1650,1673]" captionStart="FIGURES 16 – 20" captionStartId="13.[151,275,1649,1673]" captionTargetBox="[159,1415,528,1630]" captionTargetId="figure@13.[147,1432,518,1644]" captionTargetPageId="13" captionText="FIGURES 16 – 20. Psitticimex uritui: 16, antennae, head, and pronotum (female); 17, pronotal margin; 18, male genitalia; 19, female spermalege; 20, tibia III." httpUri="https://zenodo.org/record/200084/files/figure.png" pageId="15" pageNumber="16">Fig. 16</figureCitation>
).
<collectingCountry id="81AA7670E8037C65FF680FFFFE9CEDE2" box="[199,309,1682,1705]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
;
<collectingCountry id="81AA7670E8037C65FE900FFFFE37EDE2" box="[319,414,1682,1705]" name="Uruguay" pageId="15" pageNumber="16">Uruguay</collectingCountry>
. Hosts:
<taxonomicName id="3EBD4D63E8037C65FE5C0FFFFD21EDE2" ID-CoL="3RL" box="[499,648,1682,1705]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
[
<taxonomicName id="3EBD4D63E8037C65FD3A0FFFFC8BEDE2" ID-CoL="9N5" box="[661,802,1682,1705]" class="Aves" family="Emberizidae" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Emberizidae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FC830FFFFC12EDE2" ID-CoL="B3Y" box="[812,955,1682,1705]" class="Aves" family="Hirundinidae" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Hirundinidae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FC690FFFFB8FEDE2" ID-CoL="BCZ" box="[966,1062,1682,1705]" class="Aves" family="Icteridae" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Icteridae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FB9F0FFFFB0AEDE2" ID-CoL="626DC" box="[1072,1187,1682,1705]" class="Aves" family="Passeridae" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Passeridae</taxonomicName>
,
<taxonomicName id="3EBD4D63E8037C65FB010FFFFAEFEDE2" ID-CoL="626S2" box="[1198,1350,1682,1705]" class="Aves" family="Troglodytidae" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Troglodytidae</taxonomicName>
) ..........
<emphasis id="CBC9EAF2E8037C65FF610FDFFA35ED82" box="[206,1436,1714,1737]" italics="true" pageId="15" pageNumber="16">...................................................................................................................................................................Acanthocrios</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

270
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@ -0,0 +1,270 @@
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<mods:title id="22DC5263CED94FAE43164A03B18BA2B8">Insects found in birds’ nests from Argentina. Cyanoliseus patagonus (Vieillot, 1818) [Aves: Psittacidae], with the description of Cyanolicimex patagonicus, gen. n., sp. n., and a key to the genera of Haematosiphoninae (Hemiptera: Cimicidae)</mods:title>
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<paragraph id="F90236E0E8057C63FF3809FAFE6CEBFA" blockId="9.[151,453,151,177]" box="[151,453,151,177]" pageId="9" pageNumber="10">
<heading id="A24A818CE8057C63FF3809FAFE6CEBFA" bold="true" box="[151,453,151,177]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="3EBD4D63E8057C63FF3809FAFE6CEBFA" ID-CoL="7PNP8" authority="Pinto, 1927" authorityName="Pinto" authorityYear="1927" box="[151,453,151,177]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FF3809FAFE6CEBFA" bold="true" box="[151,453,151,177]" pageId="9" pageNumber="10">
<emphasis id="CBC9EAF2E8057C63FF3809FAFE9BEBFA" bold="true" box="[151,306,151,177]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
<bibRefCitation id="9D2C4B11E8057C63FE9709FAFE6CEBFA" author="Pinto" box="[312,453,151,177]" pageId="9" pageNumber="10" refString="Pinto, C. (1927) Ornithocoris toledoi, novo genero e nova especie de percevejo de ave (Hemiptera - Fam. Cimicidae). Revista de Biologia e Hygiene, 1 (2), 17 - 22." type="journal article" year="1927">Pinto, 1927</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F90236E0E8057C63FF38098FFC86EBB2" blockId="9.[151,1435,226,319]" box="[151,815,226,250]" pageId="9" pageNumber="10">
Type-species:
<taxonomicName id="3EBD4D63E8057C63FE9F098FFD27EBB2" authority="Pinto, 1927" authorityName="Pinto" authorityYear="1927" box="[304,654,226,250]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FE9F098FFDA4EBB1" bold="true" box="[304,525,226,250]" italics="true" pageId="9" pageNumber="10">Ornithocoris toledoi</emphasis>
<bibRefCitation id="9D2C4B11E8057C63FDBB098FFD27EBB2" author="Pinto" box="[532,654,226,249]" pageId="9" pageNumber="10" refString="Pinto, C. (1927) Ornithocoris toledoi, novo genero e nova especie de percevejo de ave (Hemiptera - Fam. Cimicidae). Revista de Biologia e Hygiene, 1 (2), 17 - 22." type="journal article" year="1927">Pinto, 1927</bibRefCitation>
</taxonomicName>
[by monotipy]
</paragraph>
</subSubSection>
<subSubSection id="B1A7656BE8057C63FF380865FE08ED1C" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="F90236E0E8057C63FF380865FDA3EA74" blockId="9.[151,1435,226,319]" pageId="9" pageNumber="10">
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<taxonomicName id="3EBD4D63E8057C63FF380865FE53EA54" ID-CoL="7W6LN" authority="Pinto, 1927" authorityName="Pinto" authorityYear="1927" box="[151,506,264,287]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FF380865FEDBEA54" box="[151,370,264,287]" italics="true" pageId="9" pageNumber="10">Ornithocoris toledoi</emphasis>
<bibRefCitation id="9D2C4B11E8057C63FED50865FE53EA54" author="Pinto" box="[378,506,264,287]" pageId="9" pageNumber="10" refString="Pinto, C. (1927) Ornithocoris toledoi, novo genero e nova especie de percevejo de ave (Hemiptera - Fam. Cimicidae). Revista de Biologia e Hygiene, 1 (2), 17 - 22." type="journal article" year="1927">Pinto, 1927</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="781C10F1E8057C63FDAA0865FD7DEA54" author="Ringuelet" box="[517,724,264,287]" page="35" pageId="9" pageNumber="10" year="1948">
<bibRefCitation id="9D2C4B11E8057C63FDAA0865FD04EA54" author="Ringuelet" box="[517,685,264,287]" pageId="9" pageNumber="10" refString="Ringuelet, R. (1948) Zooparasitos de interes veterinario. Su distribucion en la Argentina segun comprobaciones de la Direccion de Patologia Animal 1935 - 1945. Ministerio de Agricultura de la Nacion, Direccion General de Ganaderia, Direccion de Informaciones, Publicacion Miscelanea, (281), 1 - 54." type="journal article" year="1948">Ringuelet 1948</bibRefCitation>
: 35
</treatmentCitation>
[map], 39 [distr.; host]; Jurberg &amp; Milward de
<treatmentCitation id="781C10F1E8057C63FB610865FA26EA54" author="Azevedo" box="[1230,1423,264,287]" page="25" pageId="9" pageNumber="10" year="1982">
<bibRefCitation id="9D2C4B11E8057C63FB610865FAC3EA54" author="Azevedo" box="[1230,1386,264,287]" pageId="9" pageNumber="10" refString="Jurberg, J. &amp; Milward de Azevedo, E. M. V. (1982) Contribuicao para o estudo da morfologia de Ornithocoris toledoi Pinto 1927 (Hemiptera, Cimicidae). Revista Brasileira de Biologia, 42 (1), 255 - 262." type="journal article" year="1982">Azevedo 1982</bibRefCitation>
: 25
</treatmentCitation>
– 262 [re-descr.; morphol.; distr.]
</treatmentCitationGroup>
</paragraph>
<paragraph id="F90236E0E8057C63FF380800FDE5E957" blockId="9.[151,1437,365,1623]" pageId="9" pageNumber="10">
The genus and the species were described from
<collectingCountry id="81AA7670E8057C63FD430800FC94EACC" box="[748,829,365,391]" name="Brazil" pageId="9" pageNumber="10">Brazil</collectingCountry>
: São Paulo (Limeira: Sorocaba) by
<bibRefCitation id="9D2C4B11E8057C63FB560800FA31EACC" author="Pinto" box="[1273,1432,365,391]" pageId="9" pageNumber="10" refString="Pinto, C. (1927) Ornithocoris toledoi, novo genero e nova especie de percevejo de ave (Hemiptera - Fam. Cimicidae). Revista de Biologia e Hygiene, 1 (2), 17 - 22." type="journal article" year="1927">Pinto (1927)</bibRefCitation>
.
<bibRefCitation id="9D2C4B11E8057C63FF3808FFFEE3EAE7" author="Moraes" box="[151,330,402,428]" pageId="9" pageNumber="10" refString="Moraes, R. G. de (1939) Breve nota sobre o Ornithocoris toledoi Pinto, 1927. Revista Medicina-Cirurgica do Brasil, 47 (2), 250 - 255." type="journal article" year="1939">Moraes (1939)</bibRefCitation>
made a re-description and illustrates the specimens found in Minas Gerais: Ponte Nóva by
<bibRefCitation id="9D2C4B11E8057C63FF3808D5FEC9EA99" author="Carvalho" box="[151,352,440,466]" pageId="9" pageNumber="10" refString="Carvalho, J. C. M. (1939) Sobre a biologia de Ornithocoris toledoi Pinto, 1927, percevejo dos galinheiros em Minas Gerais (Hemiptera, Cimicidae). Ceres, 1, 120 - 140." type="journal article" year="1939">Carvalho (1939)</bibRefCitation>
.
<bibRefCitation id="9D2C4B11E8057C63FEC208D5FD8DEA99" author="Usinger" box="[365,548,440,466]" pageId="9" pageNumber="10" refString="Usinger, R. L. (1966) Monograph of Cimicidae (Hemiptera - Heteroptera). The Thomas Say Foundation, 7, xi + 1 - 585." type="journal article" year="1966">Usinger (1966)</bibRefCitation>
states that the
<typeStatus id="26068842E8057C63FD7608D5FCA4EA99" box="[729,781,440,466]" pageId="9" pageNumber="10">type</typeStatus>
of
<taxonomicName id="3EBD4D63E8057C63FC9908D5FC23EA9A" ID-CoL="7W6LN" box="[822,906,440,465]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FC9908D5FC23EA9A" box="[822,906,440,465]" italics="true" pageId="9" pageNumber="10">toledoi</emphasis>
</taxonomicName>
is in the IOC. Twenty-five specimens of
<taxonomicName id="3EBD4D63E8057C63FAD008D5FF41EABD" ID-CoL="7W6LN" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FAD008D5FF41EABD" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
, all of them with data according to the original description (probable
<typeStatus id="26068842E8057C63FBB608B0FB2DEABC" box="[1049,1156,477,503]" pageId="9" pageNumber="10" type="syntype">syntypes</typeStatus>
), are apparently located in the IOC (see below
<emphasis id="CBC9EAF2E8057C63FE300B6FFD92E950" box="[415,571,514,539]" italics="true" pageId="9" pageNumber="10">Acanthocrios</emphasis>
).
</paragraph>
<paragraph id="F90236E0E8057C63FF690B45FEA6E827" blockId="9.[151,1437,365,1623]" pageId="9" pageNumber="10">
According to
<bibRefCitation id="9D2C4B11E8057C63FED80B45FDD1E909" author="Wygodzinsky" box="[375,632,552,578]" pageId="9" pageNumber="10" refString="Wygodzinsky, P. (1951) Notas sobre Cimicidae de la Republica Argentina (Hemiptera). Anales del Instituto de Medicina Regional, Tucuman, 3 (2), 185 - 197." type="journal article" year="1951">Wygodzinsky (1951)</bibRefCitation>
, the genus
<taxonomicName id="3EBD4D63E8057C63FCBD0B45FC00E90A" ID-CoL="7PNP8" box="[786,937,552,577]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FCBD0B45FC00E90A" box="[786,937,552,577]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
can be recognized by the “very long” macroquetae in the posterolateral angles of the pronotum, the presence of two distinct bristles in the middle and hind tibiae, and the dorsal position of the spermalege (= organ of Ribaga).
<taxonomicName id="3EBD4D63E8057C63FB810B1FFB0BE9C0" ID-CoL="7W6LN" box="[1070,1186,626,651]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FB810B1FFB0BE9C0" box="[1070,1186,626,651]" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
have long bristles on the posterolateral angles not longer than 1/3 of the pronotal length [Lb &lt;PL /3] (
<figureCitation id="61862A65E8057C63FBD00BF5FB70E9F9" box="[1151,1241,664,690]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
) (
<bibRefCitation id="9D2C4B11E8057C63FB5C0BF5FF7EE99C" author="Wygodzinsky" pageId="9" pageNumber="10" refString="Wygodzinsky, P. (1951) Notas sobre Cimicidae de la Republica Argentina (Hemiptera). Anales del Instituto de Medicina Regional, Tucuman, 3 (2), 185 - 197." type="journal article" year="1951">Wygodzinsky 1951</bibRefCitation>
). In Usinger’s key (1966),
<taxonomicName id="3EBD4D63E8057C63FDB50BD0FD1AE99D" ID-CoL="7PNP8" box="[538,691,701,726]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FDB50BD0FD1AE99D" box="[538,691,701,726]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
can be recognized by the bristles at the sides of the pronotum being short and dense (
<figureCitation id="61862A65E8057C63FE010B8FFDA1E9B7" box="[430,520,738,764]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
), its larger size, and the pronotum being
<quantity id="3E459B05E8057C63FC580B8FFB94E9B7" box="[1015,1085,738,764]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1.0">1 mm</quantity>
or more wide. In the generic description, nothing is said about the bristles at the sides of the pronotum, and
<taxonomicName id="3EBD4D63E8057C63FBEE0A65FB73E86A" ID-CoL="7PNP8" box="[1089,1242,776,801]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FBEE0A65FB73E86A" box="[1089,1242,776,801]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
is differentiated by the pronotum being 2.5 to 3 times as wide as long [PW = 2.5 PL to PW = 3.0 PL ⊣ PL = PW / 2.5 to PL = PW / 3.0].
</paragraph>
<paragraph id="F90236E0E8057C63FF690A15FAA5EF39" blockId="9.[151,1437,365,1623]" pageId="9" pageNumber="10">
The genus
<taxonomicName id="3EBD4D63E8057C63FEE80A15FE76E8DA" ID-CoL="7PNP8" box="[327,479,888,913]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FEE80A15FE76E8DA" box="[327,479,888,913]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
can be recognized by the smaller and uniformly-spaced pilosity on the dorsum of the head and the pronotum (
<figureCitation id="61862A65E8057C63FE700AF0FDF7E8FC" box="[479,606,925,951]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="9" pageNumber="10">Figs. 9, 11</figureCitation>
); the pronotum trapezoid shape, with its greater width at the base (
<figureCitation id="61862A65E8057C63FAC30AF0FF01E897" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
), wider and longer than in
<emphasis id="CBC9EAF2E8057C63FE5B0AAFFD3CE890" box="[500,661,962,987]" italics="true" pageId="9" pageNumber="10">Acanthocrios</emphasis>
[but PW / PL is similar to the smaller specimens of
<taxonomicName id="3EBD4D63E8057C63FAB40AAFFA34E890" box="[1307,1437,962,987]" class="Arachnida" family="Argasidae" genus="Argas" kingdom="Animalia" order="Ixodida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="furnarii">
<emphasis id="CBC9EAF2E8057C63FAB40AAFFA34E890" box="[1307,1437,962,987]" italics="true" pageId="9" pageNumber="10">A. furnarii</emphasis>
</taxonomicName>
(Table 1)]; the two long bristles located at the posterolateral angles of the pronotum: Lb2, with its base near the margin (
<figureCitation id="61862A65E8057C63FE8B0D60FED3EF6C" box="[292,378,1037,1063]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
); and a second long bristle (Lb3), represented by the first posterior lateral bristle respect to Lb2 (Lb1 absent, and Lb2 longer than Lb3) (
<figureCitation id="61862A65E8057C63FD1B0D5FFCA4EF07" box="[692,781,1074,1100]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,270,1327,1351]" captionTargetBox="[153,1429,192,1319]" captionTargetId="figure@10.[151,1436,183,1324]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Ornitocoris toledoi: 9, antennae, head, and pronotum (female); 10, pronotal margin; 11 head (detail); 12, male genitalia." httpUri="https://zenodo.org/record/200081/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
). In males in
<taxonomicName id="3EBD4D63E8057C63FC1D0D5FFBE5EF00" ID-CoL="7PNP8" box="[946,1100,1074,1099]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FC1D0D5FFBE5EF00" box="[946,1100,1074,1099]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
, the front and middle tibiae have a well-developed apical tuft of hair (
<figureCitation id="61862A65E8057C63FD2F0D35FD71EF39" box="[640,728,1112,1138]" captionStart="FIGURES 21 – 27" captionStartId="14.[151,270,1463,1487]" captionTargetBox="[155,1433,311,1431]" captionTargetId="figure@14.[151,1436,306,1442]" captionTargetPageId="14" captionText="FIGURES 21 – 27. Apical tuft of hairs of tibiae: 21 – 24: Acanthocrios furnarii: 21 – 22, male: 21, tibia I; 22, tibia II; 23 – 24, female: 23, tibia I; 24, tibia II; 26 – 27, Ornithocoris toledoi (male), tibia II and III; 27, Cyanolicimex patagonicus (male), tibia I." httpUri="https://zenodo.org/record/200085/files/figure.png" pageId="9" pageNumber="10">Fig. 26</figureCitation>
); the females have a tuft only in the front tibiae.
</paragraph>
<paragraph id="F90236E0E8057C63FF690D10FAC6EE89" blockId="9.[151,1437,365,1623]" pageId="9" pageNumber="10">
<taxonomicName id="3EBD4D63E8057C63FF690D10FDD7EFDC" ID-CoL="7W6LM" authority="Usinger, 1959" authorityName="Usinger" authorityYear="1959" box="[198,638,1149,1175]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="CBC9EAF2E8057C63FF690D10FE60EFDD" box="[198,457,1149,1174]" italics="true" pageId="9" pageNumber="10">Ornithocoris pallidus</emphasis>
<bibRefCitation id="9D2C4B11E8057C63FE7B0D10FDD7EFDC" author="Usinger" box="[468,638,1149,1175]" pageId="9" pageNumber="10" refString="Usinger, R. L. (1959) New species of Cimicidae (Hemiptera). Entomologist, 92, 218 - 222. [cited by Usinger 1966]" type="journal article" year="1959">Usinger, 1959</bibRefCitation>
</taxonomicName>
was described by
<bibRefCitation id="9D2C4B11E8057C63FCC60D10FB8AEFDC" author="Usinger" box="[873,1059,1149,1175]" pageId="9" pageNumber="10" refString="Usinger, R. L. (1959) New species of Cimicidae (Hemiptera). Entomologist, 92, 218 - 222. [cited by Usinger 1966]" type="journal article" year="1959">Usinger (1959)</bibRefCitation>
, and redescribed in 1966. The pronotum is nearly twice as wide as the head (PW / HW ≥ 1.8) in
<taxonomicName id="3EBD4D63E8057C63FC080DCEFBB4EFF7" ID-CoL="7W6LN" box="[935,1053,1187,1212]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FC080DCEFBB4EFF7" box="[935,1053,1187,1212]" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
(female: HW 0
<quantity id="3E459B05E8057C63FB700DCFFAE2EFF7" box="[1247,1355,1186,1212]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="mm" value="0.8">0.80 mm</quantity>
, PW =
<quantity id="3E459B05E8057C63FF380DA5FEADEFA9" box="[151,260,1224,1250]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="9" pageNumber="10" unit="mm" value="1.45">1.45 mm</quantity>
), and about 1 2/3 as wide as the head (PW / HW = 1.6–1.7) in
<taxonomicName id="3EBD4D63E8057C63FC4A0DA5FB4AEFAA" ID-CoL="7W6LM" box="[997,1251,1224,1249]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="CBC9EAF2E8057C63FC4A0DA5FB4AEFAA" box="[997,1251,1224,1249]" italics="true" pageId="9" pageNumber="10">Ornithocoris pallidus</emphasis>
</taxonomicName>
(female: HW = 0.71, PW = 1.25) (
<bibRefCitation id="9D2C4B11E8057C63FE2B0D80FD9FEE4C" author="Usinger" box="[388,566,1261,1287]" pageId="9" pageNumber="10" refString="Usinger, R. L. (1966) Monograph of Cimicidae (Hemiptera - Heteroptera). The Thomas Say Foundation, 7, xi + 1 - 585." type="journal article" year="1966">Usinger 1966</bibRefCitation>
). With the respective measurements given by
<bibRefCitation id="9D2C4B11E8057C63FBD20D80FA9EEE4C" author="Moraes" box="[1149,1335,1261,1287]" pageId="9" pageNumber="10" refString="Moraes, R. G. de (1939) Breve nota sobre o Ornithocoris toledoi Pinto, 1927. Revista Medicina-Cirurgica do Brasil, 47 (2), 250 - 255." type="journal article" year="1939">Moraes (1939)</bibRefCitation>
and the specimens examined of
<taxonomicName id="3EBD4D63E8057C63FE1E0C7EFD8FEE67" ID-CoL="7W6LN" box="[433,550,1299,1324]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FE1E0C7EFD8FEE67" box="[433,550,1299,1324]" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
(Table 1), PW/HW ranges between 2.10 and 2.45. The other characteristics to separate both species of
<taxonomicName id="3EBD4D63E8057C63FE7E0C55FDC0EE1A" ID-CoL="7PNP8" box="[465,617,1336,1361]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FE7E0C55FDC0EE1A" box="[465,617,1336,1361]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
seem to be the color, paler in
<taxonomicName id="3EBD4D63E8057C63FC6A0C55FBEFEE1A" ID-CoL="7W6LM" box="[965,1094,1336,1361]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="CBC9EAF2E8057C63FC6A0C55FBEFEE1A" box="[965,1094,1336,1361]" italics="true" pageId="9" pageNumber="10">O. pallidus</emphasis>
</taxonomicName>
and darker in
<taxonomicName id="3EBD4D63E8057C63FB410C55FACBEE1A" ID-CoL="7W6LN" box="[1262,1378,1336,1361]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FB410C55FACBEE1A" box="[1262,1378,1336,1361]" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
; and the size, smaller in
<taxonomicName id="3EBD4D63E8057C63FED30C30FD14EE3C" ID-CoL="7W6LM" authority="Usinger 1966" authorityName="Usinger" authorityYear="1966" box="[380,701,1373,1399]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="CBC9EAF2E8057C63FED30C30FE56EE3D" box="[380,511,1373,1398]" italics="true" pageId="9" pageNumber="10">O. pallidus</emphasis>
(
<bibRefCitation id="9D2C4B11E8057C63FDA10C30FD1AEE3C" author="Usinger" box="[526,691,1373,1399]" pageId="9" pageNumber="10" refString="Usinger, R. L. (1966) Monograph of Cimicidae (Hemiptera - Heteroptera). The Thomas Say Foundation, 7, xi + 1 - 585." type="journal article" year="1966">Usinger 1966</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="3EBD4D63E8057C63FD670C30FCE2EE3D" ID-CoL="7W6LM" box="[712,843,1373,1398]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="CBC9EAF2E8057C63FD670C30FCE2EE3D" box="[712,843,1373,1398]" italics="true" pageId="9" pageNumber="10">O. pallidus</emphasis>
</taxonomicName>
was described from Viçoza, Minas Gerais, distant only
<quantity id="3E459B05E8057C63FF790CEFFE95EED7" box="[214,316,1410,1436]" metricMagnitude="4" metricUnit="m" metricValue="3.78" pageId="9" pageNumber="10" unit="km" value="37.8">37.8 km</quantity>
from Ponte Nóva, where dark brown specimens of
<taxonomicName id="3EBD4D63E8057C63FC6D0CEEFB96EED7" ID-CoL="7W6LN" box="[962,1087,1411,1436]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FC6D0CEEFB96EED7" box="[962,1087,1411,1436]" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
were found (
<bibRefCitation id="9D2C4B11E8057C63FB490CEFFA39EED7" author="Moraes" box="[1254,1424,1410,1436]" pageId="9" pageNumber="10" refString="Moraes, R. G. de (1939) Breve nota sobre o Ornithocoris toledoi Pinto, 1927. Revista Medicina-Cirurgica do Brasil, 47 (2), 250 - 255." type="journal article" year="1939">Moraes 1939</bibRefCitation>
). Specimens of
<taxonomicName id="3EBD4D63E8057C63FE920CC5FE1BEE8A" ID-CoL="7W6LN" box="[317,434,1448,1473]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="toledoi">
<emphasis id="CBC9EAF2E8057C63FE920CC5FE1BEE8A" box="[317,434,1448,1473]" italics="true" pageId="9" pageNumber="10">O. toledoi</emphasis>
</taxonomicName>
were also collected from Ponte Nóva in 1957 by Usinger (
<bibRefCitation id="9D2C4B11E8057C63FBCC0CC5FACAEE89" author="Sakamoto" box="[1123,1379,1448,1474]" pageId="9" pageNumber="10" refString="Sakamoto, J. M., Feinstein, J. &amp; Rasgon, J. L. (2006) Wolbachia infections in the Cimicidae: museum specimens as an untapped resource for endosymbiont surveys. Applied and Environmental Microbiology, 72, 3161 - 3167." type="journal article" year="2006">
Sakamoto
<emphasis id="CBC9EAF2E8057C63FB4E0CC5FABCEE8A" box="[1249,1301,1448,1473]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. 2006
</bibRefCitation>
).
</paragraph>
<paragraph id="F90236E0E8057C63FF690CA0FE08ED1C" blockId="9.[151,1437,365,1623]" pageId="9" pageNumber="10">
The insects given by
<bibRefCitation id="9D2C4B11E8057C63FE120CA0FD2FEEAC" box="[445,646,1485,1511]" pageId="9" pageNumber="10" refString="Otto, M. A., Da Silva, A. S., Zanette, R. A., Schmitt, L. &amp; Monteiro, S. G. (2008) Infestacao por Ornithocoris sp. em residencias do Municipio de Santa Maria, Rio Grande do Sul, Brasil. Revista Brasileira de Parasitologia Veterinaria, 17 (Suplemento 1), 99 - 101." type="journal article">
Otto
<emphasis id="CBC9EAF2E8057C63FE570CA0FD83EEAD" box="[504,554,1485,1510]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. (2008)
</bibRefCitation>
[= “
<taxonomicName id="3EBD4D63E8057C63FD130CA0FCF9EEAD" ID-CoL="7PNP8" box="[700,848,1485,1510]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8057C63FD130CA0FCF9EEAD" box="[700,848,1485,1510]" italics="true" pageId="9" pageNumber="10">Ornithocoris</emphasis>
</taxonomicName>
sp.”] in figure 1 do not belong to
<taxonomicName id="3EBD4D63E8057C63FB730CA0FAFFEEAC" ID-CoL="7NGSF" box="[1244,1366,1485,1511]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Cimicidae</taxonomicName>
. They are nymphs with pterothecae in the meso- and metathorax, and the rostrum reaches the posterior margin of the middle coxae. The identification was done by the shape of fecal spots in furniture using [only] the work of
<bibRefCitation id="9D2C4B11E8057C63FF380F50FE34ED1C" box="[151,413,1597,1623]" pageId="9" pageNumber="10" refString="Schofield, C. J., Williams, N. G., Kirk, M. L., Garcia Zapata, M. &amp; Marsden, P. D. (1986) A key for identifying faecal smears to detect domestic infestations of triatomine bugs. Revista da Sociedade Brasileira de Medicina Tropical, 19 (1), 5 - 8." type="journal article">
Schofield
<emphasis id="CBC9EAF2E8057C63FEA40F50FE94ED1D" box="[267,317,1597,1622]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. (1986)
</bibRefCitation>
!
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="08649C89A83AB7B5C16C069976ED3738" ID-CLB-Dataset="46910" ID-DOI="10.5281/zenodo.200077" ID-GBIF-Dataset="bd65ba8e-803f-4025-900d-c620a8c8ec9d" ID-ISSN="1175-5326" ID-Zenodo-Dep="200077" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460352922709" checkinUser="plazi" docAuthor="Iorio, Osvaldo Di, Turienzo, Paola, Masello, Juan &amp; Carpintero, Diego L." docDate="2010" docId="711487F6E8097C62FF380CDFFB4EE9D2" docLanguage="en" docName="zt02728p022.pdf" docOrigin="Zootaxa 2728" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyanolicimex patagonicus Carpintero, Di Iorio, Masello &amp; Turienzo, sp. n." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="8D2DFF8EE80C7C6AFFAF096DFFA9EB4B" masterDocTitle="Insects found in birds’ nests from Argentina. Cyanoliseus patagonus (Vieillot, 1818) [Aves: Psittacidae], with the description of Cyanolicimex patagonicus, gen. n., sp. n., and a key to the genera of Haematosiphoninae (Hemiptera: Cimicidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="6" updateTime="1698293739268" updateUser="ExternalLinkService">
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<mods:title id="00761C27B222210018FB5D5245E21ED1">Insects found in birds’ nests from Argentina. Cyanoliseus patagonus (Vieillot, 1818) [Aves: Psittacidae], with the description of Cyanolicimex patagonicus, gen. n., sp. n., and a key to the genera of Haematosiphoninae (Hemiptera: Cimicidae)</mods:title>
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<mods:namePart id="231B64A1FFD1BFA3777477E4076CC78F">Iorio, Osvaldo Di</mods:namePart>
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<mods:namePart id="035977AE3A03EEDC77F2A348221A0412">Masello, Juan</mods:namePart>
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<mods:namePart id="71FC2AE494E31FAA1AA16523A857CB78">Carpintero, Diego L.</mods:namePart>
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<paragraph id="F90236E0E8097C6FFF380CDFFB83EE87" blockId="5.[151,1066,1458,1484]" box="[151,1066,1458,1484]" pageId="5" pageNumber="6">
<heading id="A24A818CE8097C6FFF380CDFFB83EE87" bold="true" box="[151,1066,1458,1484]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="CBC9EAF2E8097C6FFF380CDFFB83EE87" bold="true" box="[151,1066,1458,1484]" pageId="5" pageNumber="6">
<taxonomicName id="3EBD4D63E8097C6FFF380CDFFC49EE87" ID-CoL="7STKW" authority="Carpintero, Di Iorio, Masello &amp; Turienzo" authorityName="Carpintero, Di Iorio, Masello &amp; Turienzo" box="[151,992,1458,1484]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="patagonicus" status="sp. nov.">
<emphasis id="CBC9EAF2E8097C6FFF380CDFFE7FEE87" bold="true" box="[151,470,1458,1484]" italics="true" pageId="5" pageNumber="6">Cyanolicimex patagonicus</emphasis>
Carpintero, Di Iorio, Masello &amp; Turienzo
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<taxonomicNameLabel id="D0FA5789E8097C6FFC470CDFFB83EE87" box="[1000,1066,1458,1484]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
<paragraph id="F90236E0E8097C6FFF380C97FE5CED1A" blockId="5.[151,1436,1530,1617]" pageId="5" pageNumber="6">
=
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Psitticimex
<taxonomicName id="3EBD4D63E8097C6FFE860C97FECDED5A" box="[297,356,1530,1553]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="uritui">uritui</taxonomicName>
</emphasis>
[
<emphasis id="CBC9EAF2E8097C6FFEDB0C97FE35ED5A" box="[372,412,1530,1553]" italics="true" pageId="5" pageNumber="6">non</emphasis>
Lent &amp; Abalos, 1946]:
<bibRefCitation id="9D2C4B11E8097C6FFD360C97FC6FED5A" author="Masello" box="[665,966,1530,1553]" pageId="5" pageNumber="6" refString="Masello, J. F. &amp; Quillfeldt, P. (2004 b) Are haematological parameters related to body condition, ornamentation and breeding success in wild burrowing parrots Cyanoliseus patagonus?. Journal of Avian Biology, 35, 445 - 454." type="journal article" year="2004" yearSuffix="b">Masello &amp; Quillfeldt 2004b</bibRefCitation>
: 451 [distr.; host];
<bibRefCitation id="9D2C4B11E8097C6FFB220C97FAC3ED5A" author="Masello" box="[1165,1386,1530,1553]" pageId="5" pageNumber="6" refString="Masello, J. F., Pagnossin, M. L., Sommer, C. &amp; Quillfeldt, P. (2006 a) Population size, provisioning frequency, flock size and foraging range at the largest known colony of Psittaciformes: the Burrowing Parrots of the north-eastern Patagonian coastal cliffs. Emu, 106, 69 - 79." type="journal article" year="2006" yearSuffix="a">
Masello
<emphasis id="CBC9EAF2E8097C6FFB460C97FAB1ED5A" box="[1257,1304,1530,1553]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2006a
</bibRefCitation>
: 522 [distr.: host]; Turienzo &amp; Di Iorio 2007: 34 [cat.; host; distr.; refs.],
<figureCitation id="61862A65E8097C6FFC350F77FC70ED7A" box="[922,985,1562,1585]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,269,1937,1961]" captionTargetBox="[158,1427,460,1896]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Cyanolicimex patagonus: 2, antennae, head, and pronotum (female Holotype); 3 – 4, pronotal margin; 5, spermalege (female Holotype); 6, tibia III; 7, male genitalia (Paratype)." httpUri="https://zenodo.org/record/200079/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
[distr.; host]; Di Iorio
<emphasis id="CBC9EAF2E8097C6FFB630F77FB55ED7A" box="[1228,1276,1562,1585]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2008: Table 7 [host; distr.; refs.], 20 [biol.]
</paragraph>
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Female
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: general coloration mostly brownish, with lateral margins of pronotum, hemelytral pads, legs, and antennae (excepting the apex of second segment), pale. Disks of head, pronotum, mesonotum, and hemelytral pads with long, sparse, and fine bristles.
</paragraph>
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Head
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wide (= pIS),
<quantity id="3E459B05E8097C6FFDB20F9DFD2CEC41" box="[541,645,1776,1802]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="5" pageNumber="6" unit="mm" value="0.81">0.81 mm</quantity>
long; posterior interocular space 4.47 times as wide as an eye (
<quantity id="3E459B05E8097C6FFAC80F9DFF6CEC64" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="6" unit="mm" value="0.17">0.17 mm</quantity>
wide at the base). Antennae
<quantity id="3E459B05E8097C6FFD8D0E78FD25EC64" box="[546,652,1813,1839]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="5" pageNumber="6" unit="mm" value="2.13">2.13 mm</quantity>
long [A1,
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; A2, 0.821; A3,
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; A4,
<quantity id="3E459B05E8097C6FFABD0E78FA38EC64" box="[1298,1425,1813,1839]" metricMagnitude="-4" metricUnit="m" metricValue="4.95" pageId="5" pageNumber="6" unit="mm" value="0.495">0.495 mm</quantity>
]. Rostrum does not surpass posterior margin of fore coxae (slide-mounted).
</paragraph>
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<emphasis id="CBC9EAF2E80A7C6CFF380DA1FE89EFAF" bold="true" box="[151,288,1228,1252]" pageId="6" pageNumber="7">FIGURE 8.</emphasis>
Phylogeny of the
<taxonomicName id="3EBD4D63E80A7C6CFE5C0DA1FD7FEFA8" box="[499,726,1228,1251]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Haematosiphoninae">Haematosiphoninae</taxonomicName>
, with the incorporation of
<emphasis id="CBC9EAF2E80A7C6CFBA60DA1FB0DEFA8" box="[1033,1188,1228,1251]" italics="true" pageId="6" pageNumber="7">Cyanolicimex</emphasis>
as the sister genus of
<emphasis id="CBC9EAF2E80A7C6CFF380D81FEA4EE48" box="[151,269,1260,1283]" italics="true" pageId="6" pageNumber="7">Psitticimex</emphasis>
.
</paragraph>
</caption>
<paragraph id="F90236E0E80A7C6CFF690C4AFB6DED52" blockId="6.[151,1437,1319,2037]" pageId="6" pageNumber="7">
Pronotum
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wide,
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long; PW / PL = 2.85; hind margins of pronotum rounded; bristles of sides regularly spaced, approximately of same length (
<quantity id="3E459B05E80A7C6CFCB40C26FC21EE2E" box="[795,904,1355,1381]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="0.35">0.35 mm</quantity>
) all along lateral margins; Lb2 longer (
<quantity id="3E459B05E80A7C6CFAF70C26FF6FEEC2" metricMagnitude="-4" metricUnit="m" metricValue="3.96" pageId="6" pageNumber="7" unit="mm" value="0.396">0.396 mm</quantity>
) than lateral bristles (
<figureCitation id="61862A65E80A7C6CFE6C0C02FD9DEEC2" box="[451,564,1391,1417]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,269,1937,1961]" captionTargetBox="[158,1427,460,1896]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Cyanolicimex patagonus: 2, antennae, head, and pronotum (female Holotype); 3 – 4, pronotal margin; 5, spermalege (female Holotype); 6, tibia III; 7, male genitalia (Paratype)." httpUri="https://zenodo.org/record/200079/files/figure.png" pageId="6" pageNumber="7">Figs. 3–4</figureCitation>
); both anterior and posterior lateral bristles (respect to Lb2)
<quantity id="3E459B05E80A7C6CFB5B0C02FAF5EEC2" box="[1268,1372,1391,1417]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="0.35">0.35 mm</quantity>
(Lb1 not distinguishable from remaining lateral bristles, although it will correspond to first anterior lateral bristle respect to Lb2) (
<figureCitation id="61862A65E80A7C6CFEF20CDAFE7BEE9A" box="[349,466,1463,1489]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,269,1937,1961]" captionTargetBox="[158,1427,460,1896]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Cyanolicimex patagonus: 2, antennae, head, and pronotum (female Holotype); 3 – 4, pronotal margin; 5, spermalege (female Holotype); 6, tibia III; 7, male genitalia (Paratype)." httpUri="https://zenodo.org/record/200079/files/figure.png" pageId="6" pageNumber="7">Figs. 3–4</figureCitation>
). Mesonotum slightly raised along middle. Abdomen above with long bristles on basal end of each segment, extending well beyond edge. Legs long and slender; hind ones are the longest (
<quantity id="3E459B05E80A7C6CFF300C92FEA7ED52" box="[159,270,1535,1561]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="6" pageNumber="7" unit="mm" value="2.18">2.18 mm</quantity>
). Front tibiae with a very small apical tuft of hair. All tarsi very long and slender.
</paragraph>
<paragraph id="F90236E0E80A7C6CFF690F4EFABDED2B" blockId="6.[151,1437,1319,2037]" pageId="6" pageNumber="7">
Male
<typeStatus id="26068842E80A7C6CFEA40F4EFEDFED76" box="[267,374,1571,1597]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
: all tibiae with long, erect, and thick bristles; anterior ones slightly thinner. Small tuft of hair in apices of front (
<figureCitation id="61862A65E80A7C6CFE0C0F2BFE51ED2B" box="[419,504,1606,1632]" captionStart="FIGURES 21 – 27" captionStartId="14.[151,270,1463,1487]" captionTargetBox="[155,1433,311,1431]" captionTargetId="figure@14.[151,1436,306,1442]" captionTargetPageId="14" captionText="FIGURES 21 – 27. Apical tuft of hairs of tibiae: 21 – 24: Acanthocrios furnarii: 21 – 22, male: 21, tibia I; 22, tibia II; 23 – 24, female: 23, tibia I; 24, tibia II; 26 – 27, Ornithocoris toledoi (male), tibia II and III; 27, Cyanolicimex patagonicus (male), tibia I." httpUri="https://zenodo.org/record/200085/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
) and middle tibiae. Genital segment strongly asymmetrical (
<figureCitation id="61862A65E80A7C6CFB110F2BFAAFED2B" box="[1214,1286,1606,1632]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,269,1937,1961]" captionTargetBox="[158,1427,460,1896]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Cyanolicimex patagonus: 2, antennae, head, and pronotum (female Holotype); 3 – 4, pronotal margin; 5, spermalege (female Holotype); 6, tibia III; 7, male genitalia (Paratype)." httpUri="https://zenodo.org/record/200079/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1A7656BE80A7C6DFF690F05FB54EBB1" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="F90236E0E80A7C6CFF690F05FD6CECC3" blockId="6.[151,1437,1319,2037]" pageId="6" pageNumber="7">
<emphasis id="CBC9EAF2E80A7C6CFF690F05FE6BEDC9" bold="true" box="[198,450,1640,1666]" pageId="6" pageNumber="7">Material examined.</emphasis>
<collectingCountry id="81AA7670E80A7C6CFE7E0F05FD28EDC9" box="[465,641,1640,1666]" name="Argentina" pageId="6" pageNumber="7">ARGENTINA</collectingCountry>
:
<emphasis id="CBC9EAF2E80A7C6CFD3C0F05FCB0EDC9" bold="true" box="[659,793,1640,1666]" pageId="6" pageNumber="7">Río Negro</emphasis>
: El Cóndor,
<date id="8D031020E80A7C6CFC690F05FB9EEDC9" box="[966,1079,1640,1666]" pageId="6" pageNumber="7" value="2004-12">XII-2004</date>
, J.F. Masello leg.,
<specimenCount id="EFBBFD69E80A7C6CFA800F05FA35EDC9" box="[1327,1436,1640,1666]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
<typeStatus id="26068842E80A7C6CFF380FE3FE9AEDE3" box="[151,307,1678,1704]" pageId="6" pageNumber="7" type="holotype">HOLOTYPE</typeStatus>
[
<collectionCode id="9FACAE25E80A7C6CFEED0FE3FE08EDE3" box="[322,417,1678,1704]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="6" pageNumber="7">MACN</collectionCode>
] (slide-mounted), from nest of
<taxonomicName id="3EBD4D63E80A7C6CFCB80FE3FB85EDEC" box="[791,1068,1678,1703]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="patagonus">
<emphasis id="CBC9EAF2E80A7C6CFCB80FE3FB85EDEC" box="[791,1068,1678,1703]" italics="true" pageId="6" pageNumber="7">Cyanoliseus patagonus</emphasis>
</taxonomicName>
;
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south of El Cóndor,
<date id="8D031020E80A7C6CFADA0FE3FEAEED86" pageId="6" pageNumber="7" value="1999-12-21">21- XII-1999</date>
, J.F. Masello leg.,
<specimenCount id="EFBBFD69E80A7C6CFE4D0FDEFD98ED86" box="[482,561,1715,1741]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
Allotype [
<collectionCode id="9FACAE25E80A7C6CFD010FDEFD5AED86" box="[686,755,1715,1741]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="7">ZMB</collectionCode>
] (slide-mounted),
<specimenCount id="EFBBFD69E80A7C6CFC640FDEFB9BED86" box="[971,1074,1715,1741]" pageId="6" pageNumber="7" type="female">3 female</specimenCount>
<typeStatus id="26068842E80A7C6CFB960FDEFB02ED86" box="[1081,1195,1715,1741]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
[
<collectionCode id="9FACAE25E80A7C6CFB160FDEFB57ED86" box="[1209,1278,1715,1741]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="7">ZMB</collectionCode>
] (pinned), on
<taxonomicName id="3EBD4D63E80A7C6CFF380FB4FE04EDB9" box="[151,429,1753,1778]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="patagonus">
<emphasis id="CBC9EAF2E80A7C6CFF380FB4FE04EDB9" box="[151,429,1753,1778]" italics="true" pageId="6" pageNumber="7">Cyanoliseus patagonus</emphasis>
</taxonomicName>
, 1 nymph V
<typeStatus id="26068842E80A7C6CFDE30FB5FD1DEDB9" box="[588,692,1752,1778]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
[
<collectionCode id="9FACAE25E80A7C6CFD6C0FB5FCA2EDB9" box="[707,779,1752,1778]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="7">ZMB</collectionCode>
], # 931(58) (pinned), “
<emphasis id="CBC9EAF2E80A7C6CFBB00FB4FB47EDB9" box="[1055,1262,1753,1778]" italics="true" pageId="6" pageNumber="7">
Psitticimex
<taxonomicName id="3EBD4D63E80A7C6CFB030FB4FB47EDB9" authority="Lent &amp; Ab." authorityName="Lent &amp; Ab." box="[1196,1262,1753,1778]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="uritui">uritui</taxonomicName>
</emphasis>
(Lent &amp; Ab.), det. J. Deckert 2001”, 1 nymph IV
<typeStatus id="26068842E80A7C6CFD9A0F93FD32EC53" box="[565,667,1790,1816]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
[
<collectionCode id="9FACAE25E80A7C6CFD050F93FD58EC53" box="[682,753,1790,1816]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="7">ZMB</collectionCode>
], # 927(17) (pinned); same locality,
<date id="8D031020E80A7C6CFB340F93FA81EC53" box="[1179,1320,1790,1816]" pageId="6" pageNumber="7" value="1999-11-27">27-XI-1999</date>
, 1 nymph II
<typeStatus id="26068842E80A7C6CFF1E0E4EFEB1EC76" box="[177,280,1827,1853]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
[
<collectionCode id="9FACAE25E80A7C6CFE890E4EFEC2EC76" box="[294,363,1827,1853]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="7">ZMB</collectionCode>
] (pinned); same locality,
<date id="8D031020E80A7C6CFD3C0E4EFCB2EC76" box="[659,795,1827,1853]" pageId="6" pageNumber="7" value="1999-12-07">7-XII-1999</date>
, 1 nymph IV
<typeStatus id="26068842E80A7C6CFC140E4EFB88EC76" box="[955,1057,1827,1853]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
[
<collectionCode id="9FACAE25E80A7C6CFB9F0E4EFBDCEC76" box="[1072,1141,1827,1853]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="7">ZMB</collectionCode>
] (pinned), “Bñg + Tlen”; El Cóndor,
<date id="8D031020E80A7C6CFE8D0E25FE3CEC29" box="[290,405,1864,1890]" pageId="6" pageNumber="7" value="2009-12">XII-2009</date>
, J.F. Masello leg., 5 nymphs IV
<typeStatus id="26068842E80A7C6CFCB10E25FC3BEC29" box="[798,914,1864,1890]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
[ODI], one male
<typeStatus id="26068842E80A7C6CFBC60E25FB7BEC29" box="[1129,1234,1864,1890]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
[ODI], all slidemounted, from nests of
<taxonomicName id="3EBD4D63E80A7C6CFE010E03FD17ECCC" box="[430,702,1902,1927]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="patagonus">
<emphasis id="CBC9EAF2E80A7C6CFE010E03FD17ECCC" box="[430,702,1902,1927]" italics="true" pageId="6" pageNumber="7">Cyanoliseus patagonus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F90236E0E80A7C6DFF690EFFFB54EBB1" blockId="6.[151,1437,1319,2037]" lastBlockId="7.[151,1437,152,656]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="CBC9EAF2E80A7C6CFF690EFFFE6DECE7" bold="true" box="[198,452,1938,1964]" pageId="6" pageNumber="7">Known distribution.</emphasis>
<collectingCountry id="81AA7670E80A7C6CFE650EFEFDD3ECE6" box="[458,634,1939,1965]" name="Argentina" pageId="6" pageNumber="7">ARGENTINA</collectingCountry>
:
<emphasis id="CBC9EAF2E80A7C6CFD2B0EFFFCAAECE7" bold="true" box="[644,771,1938,1964]" pageId="6" pageNumber="7">Río Negro</emphasis>
: Atlantic coast, easternmost kilometer of the colony (
<geoCoordinate id="9C895027E80A7C6CFADB0EFEFF65EC9A" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-41.05">41º 3’ S</geoCoordinate>
,
<geoCoordinate id="9C895027E80A7C6CFF740EDAFEF5EC9A" box="[219,348,1975,2001]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-62.8">62º 48’ W</geoCoordinate>
) (
<bibRefCitation id="9D2C4B11E80A7C6CFED90EDAFD71EC9A" author="Masello" box="[374,728,1975,2001]" pageId="6" pageNumber="7" refString="Masello, J. F. &amp; Quillfeldt, P. (2004 b) Are haematological parameters related to body condition, ornamentation and breeding success in wild burrowing parrots Cyanoliseus patagonus?. Journal of Avian Biology, 35, 445 - 454." type="journal article" year="2004" yearSuffix="b">Masello &amp; Quillfeldt 2004b</bibRefCitation>
);
<quantity id="3E459B05E80A7C6CFD410EDAFC84EC9A" box="[750,813,1975,2001]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="km" value="3.0">3 km</quantity>
west of the mouth of the Río Negro River,
<quantity id="3E459B05E80A7C6CFAE10EDAFA34EC9A" box="[1358,1437,1975,2001]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="km" value="30.0">30 km</quantity>
southeast of Viedma (
<geoCoordinate id="9C895027E80A7C6CFE090EB6FDB7ECBE" box="[422,542,2011,2037]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-41.066666">41º 04’ S</geoCoordinate>
,
<geoCoordinate id="9C895027E80A7C6CFD820EB6FD07ECBE" box="[557,686,2011,2037]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-62.833332">62º 50’ W</geoCoordinate>
) (
<bibRefCitation id="9D2C4B11E80A7C6CFD680EB6FC66ECBE" author="Masello" box="[711,975,2011,2037]" pageId="6" pageNumber="7" refString="Masello, J. F., Pagnossin, M. L., Sommer, C. &amp; Quillfeldt, P. (2006 a) Population size, provisioning frequency, flock size and foraging range at the largest known colony of Psittaciformes: the Burrowing Parrots of the north-eastern Patagonian coastal cliffs. Emu, 106, 69 - 79." type="journal article" year="2006" yearSuffix="a">
Masello
<emphasis id="CBC9EAF2E80A7C6CFC9A0EB6FCC5ECBF" box="[821,876,2011,2036]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2006a
</bibRefCitation>
). Further explorations are needed for determining if
<taxonomicName id="3EBD4D63E80B7C6DFEFE09F5FD33EBFA" box="[337,666,152,177]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="CBC9EAF2E80B7C6DFEFE09F5FD33EBFA" box="[337,666,152,177]" italics="true" pageId="7" pageNumber="8">Cyanolicimex patagonicus</emphasis>
</taxonomicName>
is restricted to the type-locality and/or only to the subspecies
<taxonomicName id="3EBD4D63E80B7C6DFF3809D1FD9CEB9E" box="[151,565,188,213]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="subSpecies" species="patagonus" subSpecies="patagonus">
<emphasis id="CBC9EAF2E80B7C6DFF3809D1FD9CEB9E" box="[151,565,188,213]" italics="true" pageId="7" pageNumber="8">Cyanoliseus patagonus patagonus</emphasis>
</taxonomicName>
. This is the southernmost known limit of the
<taxonomicName id="3EBD4D63E80B7C6DFBCE09D1FAE6EB9D" box="[1121,1359,188,214]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Haematosiphoninae">Haematosiphoninae</taxonomicName>
in the Western Hemisphere (
<figureCitation id="61862A65E80B7C6DFE34098DFE76EBB1" box="[411,479,224,250]" captionStart="FIGURE 1" captionStartId="1.[151,255,1908,1932]" captionTargetBox="[214,1396,433,1890]" captionTargetId="figure@1.[207,1412,421,1903]" captionTargetPageId="1" captionText="FIGURE 1. Geographic distribution of Cyanoliseus patagonus in Argentina (modified from Darrieu 1980). a) Buenos Aires province, according to Narosky &amp; Di Giacomo (1993); b) La Pampa province, according to Siegenthaler et al. (2004); c) breeding and non breeding areas, according to InfoNatura (2005). Parrot colony of El Condor (type locality of Cyanolicimex patagonicus) is indicated by a red area." httpUri="https://zenodo.org/record/200078/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
), and the third
<taxonomicName id="3EBD4D63E80B7C6DFD3B098DFCD4EBB1" box="[660,893,224,250]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Haematosiphoninae">Haematosiphoninae</taxonomicName>
with a
<taxonomicName id="3EBD4D63E80B7C6DFC7B098DFBFAEBB1" box="[980,1107,224,250]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
bird as a host.
</paragraph>
</subSubSection>
<subSubSection id="B1A7656BE80B7C62FF69086EFB4EE9D2" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="etymology">
<paragraph id="F90236E0E80B7C6DFF69086EFAFBEA55" blockId="7.[151,1437,152,656]" box="[198,1362,259,286]" pageId="7" pageNumber="8">
<emphasis id="CBC9EAF2E80B7C6DFF69086EFEFAEA56" bold="true" box="[198,339,259,285]" pageId="7" pageNumber="8">Etymology.</emphasis>
Specific epithet refers to the Argentinean region where this species was found (
<figureCitation id="61862A65E80B7C6DFB540869FAE9EA55" box="[1275,1344,260,286]" captionStart="FIGURE 1" captionStartId="1.[151,255,1908,1932]" captionTargetBox="[214,1396,433,1890]" captionTargetId="figure@1.[207,1412,421,1903]" captionTargetPageId="1" captionText="FIGURE 1. Geographic distribution of Cyanoliseus patagonus in Argentina (modified from Darrieu 1980). a) Buenos Aires province, according to Narosky &amp; Di Giacomo (1993); b) La Pampa province, according to Siegenthaler et al. (2004); c) breeding and non breeding areas, according to InfoNatura (2005). Parrot colony of El Condor (type locality of Cyanolicimex patagonicus) is indicated by a red area." httpUri="https://zenodo.org/record/200078/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F90236E0E80B7C6DFF690845FC84E903" blockId="7.[151,1437,152,656]" pageId="7" pageNumber="8">
<emphasis id="CBC9EAF2E80B7C6DFF690845FCC9EA08" bold="true" box="[198,864,296,323]" pageId="7" pageNumber="8">
Bionomics of the host and
<emphasis id="CBC9EAF2E80B7C6DFDB70844FCC9EA08" bold="true" box="[536,864,297,323]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="3EBD4D63E80B7C6DFDB70844FCF5EA08" box="[536,860,297,323]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="patagonicus">Cyanolicimex patagonicus</taxonomicName>
.
</emphasis>
</emphasis>
The cimicid bugs live inside crevices along the face of the sandstone cliff where the Burrowing Parrots dig their burrows. From there, they access the burrows of the parrots and their breeding chambers. During the first years of bug collection (1999, 2001, 2003, 2004), the numbers of bugs were not quantified, but they were very high around all nests of the study sector at the Burrowing Parrot colony (for details and precise location of the study sector see Masello &amp; Quillfeldt 2002, 2004a). In contrast, the cimicid bug was very scarce in the same parrot colony during December, 2009 (Masello pers. obs.). The summer
<date id="8D031020E80B7C6DFD9A0B64FD12E968" box="[565,699,521,547]" pageId="7" pageNumber="8" value="2009" valueMax="2010">2009–2010</date>
was very hot and dry, and the collected specimens died soon after they were extracted from the crevices and burrows.
</paragraph>
<paragraph id="F90236E0E80B7C6DFF690B39FBC3E9DB" blockId="7.[151,1437,152,656]" pageId="7" pageNumber="8">
The entire sequence of a breeding season of
<taxonomicName id="3EBD4D63E80B7C6DFD730B39FC47E926" box="[732,1006,596,621]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="patagonus">
<emphasis id="CBC9EAF2E80B7C6DFD730B39FC47E926" box="[732,1006,596,621]" italics="true" pageId="7" pageNumber="8">Cyanoliseus patagonus</emphasis>
</taxonomicName>
at El Cóndor (Masello &amp; Quillfeldt 2002,
<date id="8D031020E80B7C6DFF4F0B1BFEC5E9DB" box="[224,364,630,656]" pageId="7" pageNumber="8" value="2004" valueMax="2008">2004a 2008</date>
, Masello
<emphasis id="CBC9EAF2E80B7C6DFE710B1AFDB9E9DB" box="[478,528,631,656]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2006) can be, on average, summarized as follows.
</paragraph>
<paragraph id="F90236E0E80B7C6DFF380BD1FDFDE9B3" blockId="7.[151,1043,700,934]" pageId="7" pageNumber="8">1) Return of the migrating birds to the colony: August-September. 2) Copulations: September–October.</paragraph>
<paragraph id="F90236E0E80B7C6DFF380A6CFD63E850" blockId="7.[151,1043,700,934]" box="[151,714,769,795]" pageId="7" pageNumber="8">3) Incubation of the eggs: October–November.</paragraph>
<paragraph id="F90236E0E80B7C6DFF380A49FDD4E8ED" blockId="7.[151,1043,700,934]" pageId="7" pageNumber="8">4) Hatching of the nestlings: end of October through the end of November. 5) Nestling period: end of October through the end of January. 6) Fledgling: end of December through the end of January. 7) Migration (partial): February–March.</paragraph>
<paragraph id="F90236E0E80B7C6DFF690ABCFDDAEFCB" blockId="7.[151,1438,977,2011]" pageId="7" pageNumber="8">
Therefore, the feeding period of
<taxonomicName id="3EBD4D63E80B7C6DFDCF0ABCFCBFE8A1" box="[608,790,977,1002]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="CBC9EAF2E80B7C6DFDCF0ABCFCBFE8A1" box="[608,790,977,1002]" italics="true" pageId="7" pageNumber="8">C. patagonicus</emphasis>
</taxonomicName>
is restricted when the burrows are occupied by the parrots, i.e., from August −September through the end of January (fledgling of the nestlings). As the breeding pairs spend the night inside the nests during the nestling period (
<bibRefCitation id="9D2C4B11E80B7C6DFC050D71FB00EF7D" author="Masello" box="[938,1193,1052,1078]" pageId="7" pageNumber="8" refString="Masello, J. F., Sommer, C. &amp; Quillfeldt, P. (2006 c) Full house: the Burrowing Parrots of Patagonia. PsittaScene, 18 (2), 3 - 7." type="journal article" year="2006" yearSuffix="c">
Masello
<emphasis id="CBC9EAF2E80B7C6DFBBA0D71FBE3EF7E" box="[1045,1098,1052,1077]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2006c
</bibRefCitation>
), both nestlings and adults can be food sources for
<taxonomicName id="3EBD4D63E80B7C6DFDAD0D2CFD1DEF11" box="[514,692,1089,1114]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="CBC9EAF2E80B7C6DFDAD0D2CFD1DEF11" box="[514,692,1089,1114]" italics="true" pageId="7" pageNumber="8">C. patagonicus</emphasis>
</taxonomicName>
. After this, the bugs are deprived of food for at least 6 months (from February through the end of July).
</paragraph>
<paragraph id="F90236E0E80B7C6DFF690DE1FD6AEE2B" blockId="7.[151,1438,977,2011]" pageId="7" pageNumber="8">
In this way, the bionomics of
<taxonomicName id="3EBD4D63E80B7C6DFD850DE1FD74EFEE" box="[554,733,1164,1189]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="CBC9EAF2E80B7C6DFD850DE1FD74EFEE" box="[554,733,1164,1189]" italics="true" pageId="7" pageNumber="8">C. patagonicus</emphasis>
</taxonomicName>
are similar to other species of
<taxonomicName id="3EBD4D63E80B7C6DFBFF0DE1FB60EFED" box="[1104,1225,1164,1190]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Cimicidae</taxonomicName>
from the nests of swallows in the Northern Hemisphere [
<taxonomicName id="3EBD4D63E80B7C6DFDC50DDCFC58EF80" authority="Horvath, 1912" authorityName="Horvath" authorityYear="1912" box="[618,1009,1201,1227]" class="Insecta" family="Cimicidae" genus="Oeciacus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="CBC9EAF2E80B7C6DFDC50DDCFC97EF81" box="[618,830,1201,1226]" italics="true" pageId="7" pageNumber="8">Oeciacus vicarius</emphasis>
Horvath, 1912
</taxonomicName>
from North
<collectingCountry id="81AA7670E80B7C6DFB280DDCFB46EF80" box="[1159,1263,1201,1227]" name="United States of America" pageId="7" pageNumber="8">America</collectingCountry>
, and
<taxonomicName id="3EBD4D63E80B7C6DFA810DDCFE67EFBB" authority="Lamark, 1816" authorityName="Lamark" authorityYear="1816" class="Insecta" family="Cimicidae" genus="Oeciacus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hirundinis">
<emphasis id="CBC9EAF2E80B7C6DFA810DDCFEB9EFBB" italics="true" pageId="7" pageNumber="8">Oeciacus hirundinis</emphasis>
(Lamark, 1816)
</taxonomicName>
from Europe]. Where colonies are reoccupied each year, the three-month breeding period of the cliff swallow
<taxonomicName id="3EBD4D63E80B7C6DFE5A0D91FBA4EE5D" authority="Vieillot, 1817" authorityName="Vieillot" authorityYear="1817" box="[501,1037,1276,1302]" class="Aves" family="Hirundinidae" genus="Petrochelidon" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pyrrhonota">
<emphasis id="CBC9EAF2E80B7C6DFE5A0D91FC97EE5E" box="[501,830,1276,1301]" italics="true" pageId="7" pageNumber="8">Petrochelidon pyrrhonota</emphasis>
(Vieillot, 1817)
</taxonomicName>
[Aves:
<taxonomicName id="3EBD4D63E80B7C6DFBDE0D91FAB5EE5D" box="[1137,1308,1276,1302]" class="Aves" family="Hirundinidae" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Hirundinidae</taxonomicName>
] limits its specialized ectoparasite,
<taxonomicName id="3EBD4D63E80B7C6DFE670C4CFDF9EE71" box="[456,592,1313,1338]" class="Insecta" family="Cimicidae" genus="Oeciacus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="CBC9EAF2E80B7C6DFE670C4CFDF9EE71" box="[456,592,1313,1338]" italics="true" pageId="7" pageNumber="8">O. vicarius</emphasis>
</taxonomicName>
, to a short feeding period before host migration, and a food resource deprivation that lasts nine months (
<bibRefCitation id="9D2C4B11E80B7C6DFD9E0C2BFD1FEE2B" author="Loye" box="[561,694,1350,1376]" pageId="7" pageNumber="8" refString="Loye, J. E. (1985) The life history and ecology of the cliff swallow bug, Oeciacus vicarius (Hemiptera: Cimicidae). Cab. ORSTOM, serie Entomologie Medique et Parasitologie, 23 (2), 133 - 139." type="journal article" year="1985">Loye 1985</bibRefCitation>
).
</paragraph>
<paragraph id="F90236E0E80B7C6DFF690C01FB51ED50" blockId="7.[151,1438,977,2011]" pageId="7" pageNumber="8">
Semi-collapsed and abandoned burrows of
<taxonomicName id="3EBD4D63E80B7C6DFD660C01FCCEEECE" box="[713,871,1388,1413]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="patagonus">
<emphasis id="CBC9EAF2E80B7C6DFD660C01FCCEEECE" box="[713,871,1388,1413]" italics="true" pageId="7" pageNumber="8">C. patagonus</emphasis>
</taxonomicName>
are also used by other bird species for nesting. Nevertheless, there are only few records in the literature (Appendix II), probably because of the inaccessibility of the places chosen by the Burrowing Parrots. Observations by one of the authors (J.F.M.) permit the addition of new records (Appendix II). These other bird species may be potential hosts of
<taxonomicName id="3EBD4D63E80B7C6DFA2F0CB1FE8FED51" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="CBC9EAF2E80B7C6DFA2F0CB1FE8FED51" italics="true" pageId="7" pageNumber="8">C. patagonicus</emphasis>
</taxonomicName>
, especially if some of them use the burrows for roosting during autumn and winter.
</paragraph>
<paragraph id="F90236E0E80B7C6DFF690F4BFACEEDB0" blockId="7.[151,1438,977,2011]" pageId="7" pageNumber="8">
Noteworthily, the geographically-distant species
<taxonomicName id="3EBD4D63E80B7C6DFCB60F4AFAEFED0B" authority="Swainson, 1827" authorityName="Swainson" authorityYear="1827" box="[793,1350,1574,1600]" class="Aves" family="Psittacidae" genus="Rhynchopsitta" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachyrhyncha">
<emphasis id="CBC9EAF2E80B7C6DFCB60F4AFBC2ED0B" box="[793,1131,1575,1600]" italics="true" pageId="7" pageNumber="8">Rhynchopsitta pachyrhyncha</emphasis>
(Swainson, 1827)
</taxonomicName>
[Aves:
<taxonomicName id="3EBD4D63E80B7C6DFF380F21FEBFED2D" box="[151,278,1612,1638]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
] from
<collectingCountry id="81AA7670E80B7C6DFECC0F21FE12ED2D" box="[355,443,1612,1638]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
has a similar-associated ectoparasitic fauna (
<tableCitation id="B43F035BE80B7C6DFC610F21FB83ED2D" box="[974,1066,1612,1638]" captionStart="TABLE 2" captionStartId="8.[151,239,153,177]" captionText="TABLE 2. Known ectoparasitic fauna associated to two geographical distant species of parrots. References: [1] Wolffhügel (1912); [2] Mey et al. (2002); [3] Stone et al. (2005); [4] Blank et al. (2007); [5] Present work." pageId="7" pageNumber="8">Table 2</tableCitation>
). The nests of
<taxonomicName id="3EBD4D63E80B7C6DFB750F21FA35ED2E" box="[1242,1436,1612,1637]" class="Aves" family="Psittacidae" genus="Rhynchopsitta" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachyrhyncha">
<emphasis id="CBC9EAF2E80B7C6DFB750F21FA35ED2E" box="[1242,1436,1612,1637]" italics="true" pageId="7" pageNumber="8">R. pachyrhyncha</emphasis>
</taxonomicName>
are made inside natural cavities or by using old woodpecker holes, forming high-density clusters of nesting pairs sharing nest trees, with a maximum of three nesting pairs per tree. Clusters contained a mean of 11.5 breeding pairs [5 nests/ha]. Thirty-three (68%) from a total of 48 monitored nest trees were reused for at least 2 nesting seasons; a mean annual reuse was 62 ± 0.08% nest trees [range 56–71%] (
<bibRefCitation id="9D2C4B11E80B7C6DFC540F8CFAF2EDB0" author="Monterrubio-Rico" box="[1019,1371,1761,1787]" pageId="7" pageNumber="8" refString="Monterrubio-Rico, T. C., Cruz-Nieto, J., Enkerlin-Hoeflich, E., Venegas-Holguin, D., Tellez-Garcia, D. &amp; Marin-Togo, C. (2006) Gregarious nesting behavior of thick-billed parrots (Rhynchopsitta pachyrhyncha) in aspen stands. The Wilson Journal of Ornithology, 118 (2), 237 - 243." type="journal article" year="2006">
Monterrubio-Rico
<emphasis id="CBC9EAF2E80B7C6DFB760F8CFAA4EDB1" box="[1241,1293,1761,1786]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2006
</bibRefCitation>
).
</paragraph>
<paragraph id="F90236E0E80B7C6DFF690E6BFEF1EC90" blockId="7.[151,1438,977,2011]" pageId="7" pageNumber="8">
In contrast,
<taxonomicName id="3EBD4D63E80B7C6DFEFB0E6AFCB7EC6B" authority="Boddaert, 1873" authorityName="Boddaert" authorityYear="1873" box="[340,798,1798,1824]" class="Aves" family="Psittacidae" genus="Myiopsitta" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="monachus">
<emphasis id="CBC9EAF2E80B7C6DFEFB0E6AFDF8EC6B" box="[340,593,1799,1824]" italics="true" pageId="7" pageNumber="8">Myiopsitta monachus</emphasis>
(Boddaert, 1873)
</taxonomicName>
[Aves:
<taxonomicName id="3EBD4D63E80B7C6DFCD60E6BFC51EC6B" box="[889,1016,1798,1824]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
] is a nonmigratory
<taxonomicName id="3EBD4D63E80B7C6DFB4E0E6BFACAEC6B" box="[1249,1379,1798,1824]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
, that uses its nest year-round for roosting and breeding (
<bibRefCitation id="9D2C4B11E80B7C6DFCA40E41FBABEC0D" author="Navarro" box="[779,1026,1836,1862]" pageId="7" pageNumber="8" refString="Navarro, J. L., Martella, M. B. &amp; Bucher, E. H. (1992) Breeding season and productivity of monk parakeets in Cordoba, Argentina. Wilson Bulletin, 104 (3), 413 - 424." type="journal article" year="1992">
Navarro
<emphasis id="CBC9EAF2E80B7C6DFCD50E41FC19EC0E" box="[890,944,1836,1861]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 1992
</bibRefCitation>
). In this way, the food sources of
<emphasis id="CBC9EAF2E80B7C6DFF380E3CFECCEC21" box="[151,357,1873,1898]" italics="true" pageId="7" pageNumber="8">
Psitticimex
<taxonomicName id="3EBD4D63E80B7C6DFE8C0E3CFECCEC21" authority="Lent &amp; Abalos, 1946" authorityName="Lent &amp; Abalos" authorityYear="1946" box="[291,357,1873,1898]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="uritui">uritui</taxonomicName>
</emphasis>
(Lent &amp; Abalos, 1946) [
<taxonomicName id="3EBD4D63E80B7C6DFD220E3CFCBBEC20" box="[653,786,1873,1899]" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
:
<taxonomicName id="3EBD4D63E80B7C6DFCB10E3CFC35EC20" box="[798,924,1873,1899]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Cimicidae</taxonomicName>
] are available all the time. The nests of
<taxonomicName id="3EBD4D63E80B7C6DFAD30E3CFEBBECDB" class="Aves" family="Psittacidae" genus="Myiopsitta" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="monachus">
<emphasis id="CBC9EAF2E80B7C6DFAD30E3CFEBBECDB" italics="true" pageId="7" pageNumber="8">M. monachus</emphasis>
</taxonomicName>
from western and northern
<collectingCountry id="81AA7670E80B7C6DFDC60E1BFD4BECDB" box="[617,738,1910,1936]" name="Argentina" pageId="7" pageNumber="8">Argentina</collectingCountry>
are also colonized by great numbers of
<taxonomicName id="3EBD4D63E80B7C6DFB7E0E1AFED5ECFD" authority="Keirans, Radovsky" authorityName="Keirans, Radovsky" class="Arachnida" family="Argasidae" genus="Argas" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="monachus">
<emphasis id="CBC9EAF2E80B7C6DFB7E0E1AFA35ECDB" box="[1233,1436,1911,1936]" italics="true" pageId="7" pageNumber="8">Argas monachus</emphasis>
Keirans, Radovsky
</taxonomicName>
, &amp; Clifford, 1973 [Acari:
<taxonomicName id="3EBD4D63E80B7C6DFD170EF1FC9EECFD" box="[696,823,1948,1974]" class="Arachnida" family="Argasidae" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Argasidae</taxonomicName>
], exclusively specific to this parrot (Turienzo &amp; Di Iorio pers. obs.).
</paragraph>
<caption id="ADC26668E8047C62FF3809F4FA90EB9A" pageId="8" pageNumber="9">
<paragraph id="F90236E0E8047C62FF3809F4FA90EB9A" blockId="8.[151,1436,153,665]" pageId="8" pageNumber="9">
<emphasis id="CBC9EAF2E8047C62FF3809F4FEA6EBFA" bold="true" box="[151,271,153,177]" pageId="8" pageNumber="9">TABLE 2.</emphasis>
Known ectoparasitic fauna associated to two geographical distant species of parrots. References: [1] Wolffhügel (1912); [2] Mey
<emphasis id="CBC9EAF2E8047C62FE5709D7FD8EEB9A" box="[504,551,186,209]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2002); [3] Stone
<emphasis id="CBC9EAF2E8047C62FD5D09D7FC89EB9A" box="[754,800,186,209]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2005); [4] Blank
<emphasis id="CBC9EAF2E8047C62FC4009D7FBB7EB9A" box="[1007,1054,186,209]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2007); [5] Present work.
</paragraph>
</caption>
<paragraph id="F90236E0E8047C62FDB7099DFAB3EA4C" blockId="8.[151,1436,153,665]" box="[536,1306,240,263]" pageId="8" pageNumber="9">
<emphasis id="CBC9EAF2E8047C62FDB7099DFAB3EA4C" box="[536,1306,240,263]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FDB7099DFCE7EA4C" box="[536,846,240,263]" class="Aves" family="Psittacidae" genus="Rhynchopsitta" kingdom="Animalia" order="Psittaciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pachyrhyncha">Rhynchopsitta pachyrhyncha</taxonomicName>
<taxonomicName id="3EBD4D63E8047C62FB8D099DFAB3EA4C" box="[1058,1306,240,263]" class="Aves" family="Psittacidae" genus="Cyanoliseus" kingdom="Animalia" order="Psittaciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="patagonus">Cyanoliseus patagonus</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="F90236E0E8047C62FDD00875FBA6EA13" blockId="8.[151,1436,153,665]" pageId="8" pageNumber="9">
<collectingCountry id="81AA7670E8047C62FDD00875FD41EA64" box="[639,744,280,303]" name="Mexico" pageId="8" pageNumber="9">MEXICO</collectingCountry>
<collectingCountry id="81AA7670E8047C62FBFF0875FB45EA64" box="[1104,1260,280,303]" name="Argentina" pageId="8" pageNumber="9">ARGENTINA</collectingCountry>
<taxonomicName id="3EBD4D63E8047C62FF0F082CFEC1EA13" box="[160,360,321,344]" class="Insecta" kingdom="Animalia" order="Siphonaptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="order">SIPHONAPTERA</taxonomicName>
<emphasis id="CBC9EAF2E8047C62FE7B082DFBA6EA13" bold="true" box="[468,1039,320,344]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FE7B082DFD25EA13" box="[468,652,320,344]" class="Insecta" family="Ceratophyllidae" kingdom="Animalia" order="Siphonaptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Ceratophyllidae</taxonomicName>
<taxonomicName id="3EBD4D63E8047C62FC09082DFBA6EA13" box="[934,1039,320,344]" class="Insecta" family="Pulicidae" kingdom="Animalia" order="Siphonaptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Pulicidae</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="F90236E0E8047C62FE7B0804FB09EACB" blockId="8.[151,1436,153,665]" box="[468,1184,361,384]" pageId="8" pageNumber="9">
<emphasis id="CBC9EAF2E8047C62FE7B0804FD6AEACB" box="[468,707,361,384]" italics="true" pageId="8" pageNumber="9">Psyttopsylla mexicana</emphasis>
[3]
<taxonomicName id="3EBD4D63E8047C62FC090804FBD3EACB" box="[934,1146,361,384]" class="Insecta" family="Tungidae" genus="Hectopsylla" kingdom="Animalia" order="Siphonaptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="psittaci">
<emphasis id="CBC9EAF2E8047C62FC090804FBD3EACB" box="[934,1146,361,384]" italics="true" pageId="8" pageNumber="9">Hectopsylla psittaci</emphasis>
</taxonomicName>
[1]
</paragraph>
<paragraph id="F90236E0E8047C62FC0908FCFB37EAE3" blockId="8.[151,1436,153,665]" box="[934,1182,401,424]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FC0908FCFBD0EAE3" box="[934,1145,401,424]" class="Insecta" family="Tungidae" genus="Hectopsylla" kingdom="Animalia" order="Siphonaptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="narium">
<emphasis id="CBC9EAF2E8047C62FC0908FCFBD0EAE3" box="[934,1145,401,424]" italics="true" pageId="8" pageNumber="9">Hectopsylla narium</emphasis>
</taxonomicName>
[4]
</paragraph>
<paragraph id="F90236E0E8047C62FF0F08D4FBE8EA9A" blockId="8.[151,1436,153,665]" box="[160,1089,441,465]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FF0F08D4FEC1EA9B" box="[160,360,441,464]" class="Insecta" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="order">PHTHIRAPTERA</taxonomicName>
<taxonomicName id="3EBD4D63E8047C62FE7B08D4FBE8EA9A" box="[468,1089,441,465]" class="Insecta" family="Menoponidae" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">
<emphasis id="CBC9EAF2E8047C62FE7B08D4FBE8EA9A" bold="true" box="[468,1089,441,465]" pageId="8" pageNumber="9">Menoponidae Menoponidae</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8047C62FE7B088FFB47EAB3" blockId="8.[151,1436,153,665]" box="[468,1262,481,505]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FE7B088FFD29EAB2" box="[468,640,482,505]" class="Insecta" family="Menoponidae" genus="Heteromenopon" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8047C62FE7B088FFD29EAB2" box="[468,640,482,505]" italics="true" pageId="8" pageNumber="9">Heteromenopon</emphasis>
</taxonomicName>
sp. [3]
<taxonomicName id="3EBD4D63E8047C62FC09088FFB6EEAB2" box="[934,1223,482,505]" class="Insecta" family="Menoponidae" genus="Heteromenopon" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrurum">
<emphasis id="CBC9EAF2E8047C62FC09088FFB6EEAB2" box="[934,1223,482,505]" italics="true" pageId="8" pageNumber="9">Heteromenopon macrurum</emphasis>
</taxonomicName>
[2]
</paragraph>
<paragraph id="F90236E0E8047C62FE7B0B67FBE9E96A" blockId="8.[151,1436,153,665]" box="[468,1088,521,545]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FE7B0B67FDDAE96A" box="[468,627,522,545]" class="Insecta" family="Menoponidae" genus="Psittacobrosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8047C62FE7B0B67FDDAE96A" box="[468,627,522,545]" italics="true" pageId="8" pageNumber="9">Psittacobrosus</emphasis>
</taxonomicName>
sp. [3]
<taxonomicName id="3EBD4D63E8047C62FC090B64FBE9E96A" box="[934,1088,521,545]" class="Insecta" family="Philopteridae" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">
<emphasis id="CBC9EAF2E8047C62FC090B64FBE9E96A" bold="true" box="[934,1088,521,545]" pageId="8" pageNumber="9">Philopteridae</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8047C62FC090B5FFBB0E93A" blockId="8.[151,1436,153,665]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FC090B5FFB75E902" box="[934,1244,562,585]" class="Insecta" family="Philopteridae" genus="Paragoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="CBC9EAF2E8047C62FC090B5FFB75E902" box="[934,1244,562,585]" italics="true" pageId="8" pageNumber="9">Paragoniocotes meridionalis</emphasis>
</taxonomicName>
[2]
<taxonomicName id="3EBD4D63E8047C62FF0F0B37FE90E93A" box="[160,313,602,625]" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="order">HEMIPTERA</taxonomicName>
<taxonomicName id="3EBD4D63E8047C62FE7B0B34FBB0E93A" box="[468,1049,601,625]" class="Insecta" family="Cimicidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">
<emphasis id="CBC9EAF2E8047C62FE7B0B34FBB0E93A" bold="true" box="[468,1049,601,625]" pageId="8" pageNumber="9">Cimicidae Cimicidae</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8047C62FE7B0BEFFB4EE9D2" blockId="8.[151,1436,153,665]" box="[468,1255,642,665]" pageId="8" pageNumber="9">
<taxonomicName id="3EBD4D63E8047C62FE7B0BEFFDF7E9D2" box="[468,606,642,665]" class="Insecta" family="Cimicidae" genus="Ornithocoris" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="CBC9EAF2E8047C62FE7B0BEFFDF7E9D2" box="[468,606,642,665]" italics="true" pageId="8" pageNumber="9">Ornithocoris</emphasis>
</taxonomicName>
sp. [3]
<taxonomicName id="3EBD4D63E8047C62FC090BEFFB68E9D2" box="[934,1217,642,665]" class="Insecta" family="Cimicidae" genus="Cyanolicimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="CBC9EAF2E8047C62FC090BEFFB68E9D2" box="[934,1217,642,665]" italics="true" pageId="8" pageNumber="9">Cyanolicimex patagonicus</emphasis>
</taxonomicName>
[5]
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="09AC45DF7BD96F7BE5A7EFAC7625630F">Insects found in birds’ nests from Argentina. Cyanoliseus patagonus (Vieillot, 1818) [Aves: Psittacidae], with the description of Cyanolicimex patagonicus, gen. n., sp. n., and a key to the genera of Haematosiphoninae (Hemiptera: Cimicidae)</mods:title>
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<emphasis id="CBC9EAF2E8197C7FFF38098FFE5DEBB1" bold="true" box="[151,500,226,250]" italics="true" pageId="21" pageNumber="22">Cyanoliseus patagonus andinus</emphasis>
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<emphasis id="CBC9EAF2E8197C7FFF38082FFEFBEA11" bold="true" box="[151,338,322,346]" italics="true" pageId="21" pageNumber="22">Coragyps atratus</emphasis>
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=
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: Castellanos 1932.
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:
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: Valle de Los Reartes, Barranca de Las Lagunas, in abandoned nest burrows (Castellanos 1932).
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<emphasis id="CBC9EAF2E8197C7FFF380B0FFDC8E931" bold="true" box="[151,609,610,634]" italics="true" pageId="21" pageNumber="22">Geranoaetus melanoleucus melanoleucus</emphasis>
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:
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: El Cóndor, in semi-collapsed and abandoned burrows.
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jota
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:
<emphasis id="CBC9EAF2E8197C7FFEEF0B8FFE1DE9B1" bold="true" box="[320,436,738,762]" pageId="21" pageNumber="22">Río Negro</emphasis>
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<taxonomicName id="3EBD4D63E8197C7FFF380A4FFD90E872" authority="Sharpe, 1875" authorityName="Sharpe" authorityYear="1875" box="[151,569,802,826]" class="Aves" family="Falconidae" genus="Falco" kingdom="Animalia" order="Falconiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="peregrinus" subSpecies="cassini">
<emphasis id="CBC9EAF2E8197C7FFF380A4FFE0CE871" bold="true" box="[151,421,802,826]" italics="true" pageId="21" pageNumber="22">Falco peregrinus cassini</emphasis>
Sharpe, 1875
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:
<emphasis id="CBC9EAF2E8197C7FFEEF0A2FFE1DE811" bold="true" box="[320,436,834,858]" pageId="21" pageNumber="22">Río Negro</emphasis>
: El Cóndor, in semi-collapsed and abandoned burrows.
</paragraph>
<paragraph id="F90236E0E8197C7FFF380A0FFDB9E832" blockId="21.[151,1436,578,1242]" box="[151,528,866,890]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380A0FFDB9E832" authority="Vieillot, 1816" authorityName="Vieillot" authorityYear="1816" box="[151,528,866,890]" class="Aves" family="Falconidae" genus="Milvago" kingdom="Animalia" order="Falconiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="chimango">
<emphasis id="CBC9EAF2E8197C7FFF380A0FFECCE831" bold="true" box="[151,357,866,890]" italics="true" pageId="21" pageNumber="22">Milvago chimango</emphasis>
(Vieillot, 1816)
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380AEFFBADE8D2" blockId="21.[151,1436,578,1242]" box="[151,1028,898,922]" pageId="21" pageNumber="22">
<collectingCountry id="81AA7670E8197C7FFF380AEFFE9EE8D2" box="[151,311,898,921]" name="Argentina" pageId="21" pageNumber="22">ARGENTINA</collectingCountry>
:
<emphasis id="CBC9EAF2E8197C7FFEEF0AEFFE1DE8D1" bold="true" box="[320,436,898,922]" pageId="21" pageNumber="22">Río Negro</emphasis>
: El Cóndor, in semi-collapsed and abandoned burrows.
</paragraph>
<paragraph id="F90236E0E8197C7FFF380ACFFEF3E8F2" blockId="21.[151,1436,578,1242]" box="[151,346,930,953]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380ACFFEF3E8F2" box="[151,346,930,953]" class="Aves" family="Hirundinidae" kingdom="Animalia" order="Passeriformes" pageId="21" pageNumber="22" phylum="Chordata" rank="family">HIRUNDINIDAE</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380AAFFE77E892" blockId="21.[151,1436,578,1242]" box="[151,478,962,986]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380AAFFE77E892" authority="Baird, 1865" authorityName="Baird" authorityYear="1865" box="[151,478,962,986]" class="Aves" family="Hirundinidae" genus="Progne" kingdom="Animalia" order="Passeriformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="modesta">
<emphasis id="CBC9EAF2E8197C7FFF380AAFFEEEE891" bold="true" box="[151,327,962,986]" italics="true" pageId="21" pageNumber="22">Progne modesta</emphasis>
(Baird, 1865)
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380A8FFB90EF52" blockId="21.[151,1436,578,1242]" pageId="21" pageNumber="22">
<collectingCountry id="81AA7670E8197C7FFF380A8FFE9EE8B2" box="[151,311,994,1017]" name="Argentina" pageId="21" pageNumber="22">ARGENTINA</collectingCountry>
: without locality (Sclater &amp; Hudson 1887, Hudson 1920);
<emphasis id="CBC9EAF2E8197C7FFC070A8FFB97E8B1" bold="true" box="[936,1086,994,1018]" pageId="21" pageNumber="22">Buenos Aires</emphasis>
: Carmen de Patagones (Fraga &amp; Narosky 1985);
<emphasis id="CBC9EAF2E8197C7FFEDB0D6FFE4EEF51" bold="true" box="[372,487,1026,1050]" pageId="21" pageNumber="22">Río Negro</emphasis>
: El Cóndor, in semi-collapsed and abandoned burrows.
</paragraph>
<paragraph id="F90236E0E8197C7FFF380D4FFEA9EF72" blockId="21.[151,1436,578,1242]" box="[151,256,1058,1081]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380D4FFEA9EF72" box="[151,256,1058,1081]" class="Aves" family="Picidae" kingdom="Animalia" order="Piciformes" pageId="21" pageNumber="22" phylum="Chordata" rank="family">PICIDAE</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380D2FFDB5EF12" blockId="21.[151,1436,578,1242]" box="[151,540,1090,1114]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380D2FFDB5EF12" authority="Vieillot, 1818" authorityName="Vieillot" authorityYear="1818" box="[151,540,1090,1114]" class="Aves" family="Picidae" genus="Colaptes" kingdom="Animalia" order="Piciformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="campestris">
<emphasis id="CBC9EAF2E8197C7FFF380D2FFED8EF11" bold="true" box="[151,369,1090,1114]" italics="true" pageId="21" pageNumber="22">Colaptes campestris</emphasis>
(Vieillot, 1818)
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380D0FFBADEF32" blockId="21.[151,1436,578,1242]" box="[151,1028,1122,1146]" pageId="21" pageNumber="22">
<collectingCountry id="81AA7670E8197C7FFF380D0FFE9EEF32" box="[151,311,1122,1145]" name="Argentina" pageId="21" pageNumber="22">ARGENTINA</collectingCountry>
:
<emphasis id="CBC9EAF2E8197C7FFEEF0D0FFE1DEF31" bold="true" box="[320,436,1122,1146]" pageId="21" pageNumber="22">Río Negro</emphasis>
: El Cóndor, in semi-collapsed and abandoned burrows.
</paragraph>
<paragraph id="F90236E0E8197C7FFF380DEFFE98EFD2" blockId="21.[151,1436,578,1242]" box="[151,305,1154,1177]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380DEFFE98EFD2" box="[151,305,1154,1177]" class="Aves" family="Tytonidae" kingdom="Animalia" order="Strigiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="family">TYTONIDAE</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380DCFFE4BEFF2" blockId="21.[151,1436,578,1242]" box="[151,482,1186,1210]" pageId="21" pageNumber="22">
<taxonomicName id="3EBD4D63E8197C7FFF380DCFFE4BEFF2" authority="Gray, 1878" authorityName="Gray" authorityYear="1878" box="[151,482,1186,1210]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="alba" subSpecies="tuidara">
<emphasis id="CBC9EAF2E8197C7FFF380DCFFEFDEFF1" bold="true" box="[151,340,1186,1210]" italics="true" pageId="21" pageNumber="22">Tyto alba tuidara</emphasis>
(Gray, 1878)
</taxonomicName>
</paragraph>
<paragraph id="F90236E0E8197C7FFF380DAFFBADEF92" blockId="21.[151,1436,578,1242]" box="[151,1028,1218,1242]" pageId="21" pageNumber="22">
<collectingCountry id="81AA7670E8197C7FFF380DAFFE9EEF92" box="[151,311,1218,1241]" name="Argentina" pageId="21" pageNumber="22">ARGENTINA</collectingCountry>
:
<emphasis id="CBC9EAF2E8197C7FFEEF0DAFFE1DEF91" bold="true" box="[320,436,1218,1242]" pageId="21" pageNumber="22">Río Negro</emphasis>
: El Cóndor, in semi-collapsed and abandoned burrows.
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165
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<paragraph pageId="18" pageNumber="19">Taxon classification Animalia Hymenoptera Chrysididae</paragraph>
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<paragraph pageId="18" pageNumber="19">
Genus
<taxonomicName authority="Spinola, 1806" authorityName="Spinola" authorityYear="1806" class="Insecta" family="Chrysididae" genus="Elampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Elampus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Elampus Spinola, 1806</taxonomicName>
Figs 37, 38-46
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<paragraph pageId="19" pageNumber="20">
<taxonomicName class="Insecta" family="Chrysididae" genus="Elampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Elampus" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="19" pageNumber="20" start="start">Elampus</pageBreakToken>
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Spinola, 1806: 10.
</paragraph>
<paragraph pageId="19" pageNumber="20">
<taxonomicName class="Insecta" family="Chrysididae" genus="Ellampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ellampus" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Ellampus</taxonomicName>
Agassiz, 1846: 136.
</paragraph>
<paragraph pageId="19" pageNumber="20">
<taxonomicName class="Insecta" family="Chrysididae" genus="Notozus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Notozus" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Notozus</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1853: 351.
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<subSubSection pageId="19" pageNumber="20" type="note">
<paragraph pageId="19" pageNumber="20">Note.</paragraph>
<paragraph pageId="19" pageNumber="20">
This genus has been treated as a subgenus of
<taxonomicName class="Insecta" family="Hymenoptera" genus="Omalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Omalus" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Omalus</taxonomicName>
by some authors (e.g.
<bibRefCitation author="Linsenmaier, W" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="95" pageNumber="96" pagination="1 - 232" title="Revision der Familie Chrysididae (Hymenoptera) mit besonderer Bruecksichtigung der europaeischen Spezies." volume="32" year="1959">Linsenmaier 1959</bibRefCitation>
,
<bibRefCitation author="Linsenmaier, W" journalOrPublisher="Veroeffentlichungen aus dem Natur-Museum Luzern" pageId="95" pageNumber="96" pagination="1 - 139" title="Die Goldwespen der Schweiz." volume="9" year="1997">1997</bibRefCitation>
). It is well characterised by the shape of the metascutellum, which has a large tongue-like projection dorsally (Fig. 38). The posterior margin of T3 is usually extended into a horseshoe-shaped or falcate rim forming truncation (Figs 39-40, 44-46). The female has a row of dense and erect setae along the genal margin (Fig. 41). These setae are replaced by long irregularly placed bristles in the male. The hosts are ground-nesting crabronid wasps, such as
<taxonomicName class="Insecta" family="Crabronidae" genus="Mimesa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mimesa" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Mimesa</taxonomicName>
Shuckard and
<taxonomicName class="Insecta" family="Crabronidae" genus="Mimumesa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mimumesa" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Mimumesa</taxonomicName>
Malloch (
<bibRefCitation author="Kimsey, LS" journalOrPublisher="Oxford Press, New York" pageId="94" pageNumber="95" title="[1990] The Chrysidid Wasps of the World." year="1991">Kimsey and Bohart 1991</bibRefCitation>
). The genus is distributed in the Palearctic Region (more than 40 species), North America (8 species), Africa (7 species) and South America (3 species) (
<bibRefCitation author="Kimsey, LS" journalOrPublisher="Oxford Press, New York" pageId="94" pageNumber="95" title="[1990] The Chrysidid Wasps of the World." year="1991">Kimsey and Bohart 1991</bibRefCitation>
,
<bibRefCitation author="Linsenmaier, W" journalOrPublisher="Entomofauna, Supplement" pageId="95" pageNumber="96" pagination="1 - 281" title="Die Goldwespen Nordafrikas (Hymenoptera, Chrysididae)." volume="10" year="1999">Linsenmaier 1999</bibRefCitation>
,
<bibRefCitation author="Madl, M" journalOrPublisher="Linzer biologische Beitraege" pageId="95" pageNumber="96" pagination="5 - 169" title="A catalogue of the Chrysididae (Hymenoptera: Chrysidoidea) of the Ethiopian Region excluding Malagasy Subregion." volume="44" year="2012">Madl and Rosa 2012</bibRefCitation>
). A total of 12 species have been found in Europe (
<bibRefCitation author="Rosa, P" editor="Mitroiu, M-D" journalOrPublisher="Zootaxa" pageId="97" pageNumber="98" title="Fauna Europaea: Hymenoptera, version 2.5." url="http://www.faunaeur.org" year="2012">Rosa and Soon 2012</bibRefCitation>
), and three of these occur in the Nordic and Baltic countries (
<bibRefCitation author="Paukkunen, J" journalOrPublisher="Zootaxa" pageId="97" pageNumber="98" pagination="1 - 67" title="Faunistic review of the cuckoo wasps of Fennoscandia, Denmark and the Baltic countries (Hymenoptera: Chrysididae)." url="10.11646/zootaxa.3864.1.1" volume="3864" year="2014">Paukkunen et al. 2014</bibRefCitation>
).
</paragraph>
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<paragraph pageId="19" pageNumber="20">
Figure 37.
<taxonomicName class="Insecta" family="Chrysididae" genus="Elampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Elampus panzeri" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="panzeri">Elampus panzeri</taxonomicName>
♀. Scale 1 mm.
</paragraph>
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Key to
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species of the Nordic and Baltic countries
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(Fabricius)
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(
<normalizedToken originalValue="Mocsáry">Mocsary</normalizedToken>
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<treatment id="7114A27EE35D0B82FE1C45797A6B58E9" ID-DOI="http://doi.org/10.5281/zenodo.10915929" ID-GBIF-Taxon="224779208" ID-Zenodo-Dep="10915929" LSID="urn:lsid:plazi:treatment:7114A27EE35D0B82FE1C45797A6B58E9" httpUri="http://treatment.plazi.org/id/7114A27EE35D0B82FE1C45797A6B58E9">
<subSubSection id="9579972AAB16EACDF0BEAD35BA9E6C72" type="nomenclature">
<paragraph id="E0E2B15C78B4670DCDDAEA6BEAA24888">
<taxonomicName id="2DCA7375C6AF040E9A2CD8075ADC62F2" ID-CoL="YXZN" authority="(Roxb.) G. Klotz" authorityName="G. Klotz" baseAuthorityName="Roxb." class="Magnoliopsida" family="Rosaceae" genus="Cotoneaster" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="integrifolius">
<emphasis id="CCF90C1473982B16E9C1B2B8C5D8A202" italics="true">Cotoneaster integrifolius</emphasis>
(Roxb.) G. Klotz
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1E14330FB696CEF736A77D866E096B2D" type="vernacular_names">
<paragraph id="230A9945718D3BD92CF68821620ABE98">
<vernacularName id="8A6AEAE278479377D76627325302F1E3">Ganzrandige Zwergmispel</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="5CF1EEFD2E5313C20BE7AE3B96CCCDDA" type="multiple">
<paragraph id="F2F202CBAF684B3533E061CBA602EBDE">
Art ISFS: 121850 Checklist: 1013460
<taxonomicName id="165809BA907DCB4090186814A41C3127" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
<taxonomicName id="3E56EF3229F1BE92646FB52E46282F36" class="Magnoliopsida" family="Rosaceae" genus="Cotoneaster" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="genus">Cotoneaster</taxonomicName>
<taxonomicName id="A22C01DD456706350BE18F8BBAE10208" authority="(Roxb.) G. Klotz" authorityName="G. Klotz" baseAuthorityName="Roxb." class="Magnoliopsida" family="Rosaceae" genus="Cotoneaster" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="integrifolius">Cotoneaster integrifolius (Roxb.) G. Klotz</taxonomicName>
</paragraph>
<paragraph id="786FC399FC3625BCD17C906A0D24FDFA">
<heading id="18CD44938D9453F38F42F35AD805205C">Zusammenfassung</heading>
KEINE ANGABE
</paragraph>
<paragraph id="DE3CC31BF5D77ADA802A9F030A07BF26">
<heading id="980415301A72CED266AC90D9E3386559">
<normalizedToken id="101B53E27092EA49258D3C6D503722EC" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="ADBF5BFB1E7908552CDBFE714259D861">
<heading id="8345FA91B9FA0DF1A957FABCAB872F54">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="268D0774F0BA8EEA38A79BC7527A02AA" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="8B8E42A09914E0A4011A7186CA72F22D">
<table id="B8E5850FE4C99A366FE45C937794BEBC" class="table" inLine="true">
<tr id="966AB8B0B0D5969E8D01C9F8AF7FD559">
<td id="462716553810AAFAC83F4334B3D520C2">
<tbody id="70385BCDC082C8105F11BD227ABED260">KEINE ANGABE</tbody>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="AA6D079035812B8CFF25DFF4F79A54AC" type="biology_ecology">
<paragraph id="ACC82AAB02FE9AC315F1F971DC780266">
<normalizedToken id="54120CD4F0B15DE151A66A029B048F2D" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="8C34B1CE2DEDD7CC34342469C1F54769" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="1C0C4A8B3956242E3D634719B23B6E82">
<table id="F4EDBFB0D1CA1A5A1D22ED840D700BDB" class="table" inLine="true">
<tbody id="17FDCE70A828711B938016E52D4D6817">
<tr id="5C1BA3D61BD5848C200FE7506A27558C">
<th id="5F117DFFE6226C8C66929716F4B51FFA" colspan="2">
<emphasis id="B5309DD73AEDA76D3B2E1BBE8D99A8E5" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="FA9ABF568F3099FB9C9AF5287384C723" colspan="2">
<emphasis id="C1D4615AE84A8AF17EEB90A076DB7981" bold="true">Klimafaktoren</emphasis>
</th>
<th id="7B76618E513A8D6B2E0362E14033DD94" colspan="2">
<emphasis id="5AFFD407D51C03123AC731657D1E2511" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="ED3089852896FBE85A9A027CD50FF547">
<td id="8C696A71FE0FBF015CBDD2FE3E50B595">Feuchtezahl F</td>
<td id="1FF1A81FDADB34A05D9A7F96895552F3" style="text-align:left;width:100px">--</td>
<td id="9B31C8E625B1BF63AC234701BEE5E7ED">Lichtzahl L</td>
<td id="822BBEFA111B4F9069AFE9BE3BE689A5" style="text-align:left;width:60px">--</td>
<td id="50D48D45F6BC20A8C919B2F94BC133BC">Salzzeichen</td>
<td id="030C7891B2849EC5F601A5C660AB6E30" style="text-align:left;width:60px">--</td>
</tr>
<tr id="235A66BC8F4966781971A0EAEF4DD19D">
<td id="B6B2D8DC60582F2AA5478AC16194FF67">Reaktionszahl R</td>
<td id="13D6E07C75DFB52C0C257E108AA5FCB1" style="text-align:left">--</td>
<td id="352D8B5D345D65F94FD57FBC843D203A">Temperaturzahl T</td>
<td id="5C1C8464396CA6645C4A7CCAE3F5EC7A" style="text-align:left">--</td>
</tr>
<tr id="ED3413A3229C07ACF9BB2F1DC53B4097">
<td id="44C73A9E3D20A5A93C8A0EBF3AF2440F">
<normalizedToken id="E3B165439F963596BDDBA4CA5A08522D" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="BC790B30712A022A94236C31347CF3A6" style="text-align:left">--</td>
<td id="98FFA8426BD02F02F70AEBF3CA97A5F9">
<normalizedToken id="8967A683F960228AB666DD892E60C969" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="0CFA2EF5A5CBE2ED30330195D3858FAC" style="text-align:left">--</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="2E2459D802BF87BD3DC0BC164C422EAA" type="reference_group">
<paragraph id="1ACDE254835B9F4FA481B0B981FCEBE5">
<heading id="DAB254BB48DD8C05F33F5EFFCEA8CE64">Nomenklatur</heading>
</paragraph>
<paragraph id="A581A9C253656AFF58882D593A27E63C">
<treatmentCitation id="4A9965788C12E083214EF508BAD397BD">
<heading id="02D664610C330AF4EBA46968852A2210">
<normalizedToken id="4E5A9345A900D72215758306995C9C23" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="0D345CCA61B99DD8AC763A161571C664" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="72B089FB7A545F9342F32ECD6CC921D3" ID-CoL="YXZN" authority="(Roxb.) G. Klotz" authorityName="G. Klotz" baseAuthorityName="Roxb." class="Magnoliopsida" family="Rosaceae" genus="Cotoneaster" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="integrifolius">
<emphasis id="ADE9C13BD0FB5537CF422C3864DD799D" italics="true">Cotoneaster integrifolius</emphasis>
(Roxb.) G. Klotz
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="FA9C1106C90065891F872292CB6194FB" type="vernacular_names">
<paragraph id="48F46278F2C55EB13EEF675AF6476CC4">
Volksname Deutscher Name:
<vernacularName id="0B4A0A8185236A6CE5AD87C3594ABFD2" language="deu">Ganzrandige Zwergmispel</vernacularName>
Nom
<normalizedToken id="BA1039E27DB8FB26B9BB1E82D6994D26" originalValue="français">francais</normalizedToken>
:
<vernacularName id="DB2C5226D6B13C10C4C1D60B4A166BC4" language="fra">
<taxonomicName id="C90E7A78C3CE72148E6576E878ED13F9" class="Magnoliopsida" family="Rosaceae" genus="Cotoneaster" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="genus">
<normalizedToken id="EE178286BA6AC0FE06D388DF457886DB" originalValue="Cotonéaster">Cotoneaster</normalizedToken>
</taxonomicName>
<normalizedToken id="6921FCD3E2B63539716D5D0145CAC21C" originalValue="à">a</normalizedToken>
feuilles de thym
</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="18367D2A7283C7F0CBC54DDBA147D78F" type="multiple">
<paragraph id="D6D1131A5C48507EC162A261682010F3">Nome italiano: --</paragraph>
</subSubSection>
<subSubSection id="55346B9FAF2EFB6B18FCC7566DAE81E5" type="reference_group">
<paragraph id="4EA8806BFF67EC77AC2682400CE64B0F">
<normalizedToken id="0CB65E73300B2663668B5827247A5AEB" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="15AF58B00241B0777E42A1BD0659E9B7">
<table id="9D11EF2FDF7B36FF12E1ADA00C91477C" class="table table-condensed" inLine="true">
<tbody id="CCA2CDF56EF41138A53AE0D503E6D432">
<tr id="45478A3FA8D4AC676606FCBCA2862366">
<th id="46F430294394D9162DE09107CFB45DFA" width="80px">Relation</th>
<th id="C74645CE8A845CAC0D4C27819758EDD9">Nom</th>
<th id="5E8FA455E69E76E1C7B5E6B042CD60B4">Referenzwerke</th>
<th id="5B716A98A504BFC138953F973651382E" width="90px">No</th>
</tr>
<tr id="7D595846D0C3E03D9644BA22E0F70F00">
<td id="D58437AE8D22149E598BED5F4D245D1E" class="taxon-relation">=</td>
<td id="E2018A393CFBB4F00977F070A091FDA3">
<taxonomicName id="819567B870DE8DD1B65D0B9A07921C6C" authority="(Roxb.) G. Klotz" authorityName="G. Klotz" baseAuthorityName="Roxb." class="Magnoliopsida" family="Rosaceae" genus="Cotoneaster" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="integrifolius">Cotoneaster integrifolius (Roxb.) G. Klotz</taxonomicName>
</td>
<td id="4AC54214D5338E386BED2ECC52BC7FDA">
<treatmentCitation id="DABD453201F34A7CB0B1F7A211DABB65">
<bibRefCitation id="053F0E0EAB1A2B1BA7D0F75B268326AA" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
</treatmentCitation>
</td>
<td id="E2579C7E73F8322B81390744AA918DF9">121850</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="E7F128558E6063F162ACBC68FAA23623">
<tableNote id="E864D5AA9E3FA682879C5C93AEE0D149">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="6237791B345FCF3D33E0DA52D00C479C" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="9AB6D33A5E7B6C56144742CE48C8FECC" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="4377FF87BFAECF7CF673F04C1E535A22" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="6AB9C248CBB9975AA72B0DE011F466CA" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="97146CCDB522422F90E7E41BF0E0D645" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="56D5CF3CDD194C3922F99CC13CF3A321" type="multiple">
<paragraph id="2DA742017C1631B3F5A6D1BF4579CA47">
Kommentare aus der
<bibRefCitation id="CB5143F40E85245F11DA9A7D4D76C26C" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
Neues Taxon:
<normalizedToken id="10AD2CB7AA6C59DF619C566FAEF2E89F" originalValue="Gegenüber">Gegenueber</normalizedToken>
SISF-2 neu aufgenommener Neophyt. Aus Kultur verwildert. Checklist
</paragraph>
</subSubSection>
<subSubSection id="ADA58BEB511EEB0F63D316697D73EBB8" type="conservation">
<paragraph id="7E6291FE990769F1D10793DA27FEBD57">
<heading id="1338B9EC2EB6A31FFD659C25B366AC01">Status Indigenat</heading>
: Kultivierter Neophyt: nach dem Jahr
<quantity id="E4C372B0D7BC3C8A26949AA75BFAF71E" metricMagnitude="1" metricUnit="m" metricValue="3.81" unit="in" value="1500.0">1500 in</quantity>
der Schweiz aufgetreten
</paragraph>
<paragraph id="32233816A1C9CC82245BBEE7FA9F3E01">
<emphasis id="40FC18CEDE3D914F7841CAB99FF3627F" bold="true">
<heading id="ECD3FCF10F11899EB0F1A422B69F28EF">
Liste der
<normalizedToken id="D3877B64E434BB2E3A3CB6015BFC809D" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="9517AFBE6AC5EA891F06C1CF84A9E91A" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
</subSubSection>
</treatment>
</document>

218
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.935.51414" ID-GBIF-Dataset="96c93bf9-a3f5-4a1a-b7e5-01d652e80d4f" ID-PMC="PMC7256064" ID-Pensoft-Pub="1313-2970-935-37" ID-Pensoft-UUID="87CF7856B9B3578091EE1A493EB198D1" ID-PubMed="32508501" ID-ZooBank="E09A085DF71640DE9B243E1E0BC01BA0" ModsDocID="1313-2970-935-37" checkinTime="1590112776923" checkinUser="pensoft" docAuthor="Chen, Zhe-Yu &amp; Wu, Min" docDate="2020" docId="7114A84B87085039B69ACAFA11B9AA6F" docLanguage="en" docName="ZooKeys 935: 37-46" docOrigin="ZooKeys 935" docSource="http://dx.doi.org/10.3897/zookeys.935.51414" docTitle="Sinoarinia Chen &amp; Wu 2020, gen. nov." docType="treatment" docUuid="97E91F22-0BD2-4A64-A6A9-2811065ED0F4" docUuidSource="ZooBank" docVersion="4" id="87CF7856B9B3578091EE1A493EB198D1" lastPageNumber="37" masterDocId="87CF7856B9B3578091EE1A493EB198D1" masterDocTitle="Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae)" masterLastPageNumber="46" masterPageNumber="37" pageNumber="37" updateTime="1668168876055" updateUser="ExternalLinkService">
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<mods:title>Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Zhe-Yu</mods:namePart>
<mods:affiliation>College of food science and engineering, Wuhan Polytechnic University, Wuhan 430023, China &amp; School of Life Sciences, Nanjing University, Xianlindadao 163, Nanjing 210023, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4150-8906</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wu, Min</mods:namePart>
<mods:affiliation>School of Life Sciences, Nanjing University, Xianlindadao 163, Nanjing 210023, China</mods:affiliation>
<mods:nameIdentifier type="email">minwu1969@aliyun.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>935</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>37</mods:start>
<mods:end>46</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.935.51414</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.935.51414</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-935-37</mods:identifier>
<mods:identifier type="ZooBank">E09A085DF71640DE9B243E1E0BC01BA0</mods:identifier>
<mods:identifier type="Pensoft-UUID">87CF7856B9B3578091EE1A493EB198D1</mods:identifier>
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<treatment ID-GBIF-Taxon="164248516" LSID="urn:lsid:zoobank.org:act:97E91F22-0BD2-4A64-A6A9-2811065ED0F4" httpUri="http://treatment.plazi.org/id/7114A84B87085039B69ACAFA11B9AA6F" lastPageNumber="37" pageId="0" pageNumber="37">
<subSubSection pageId="0" pageNumber="37" type="nomenclature">
<paragraph pageId="0" pageNumber="37">
<taxonomicName LSID="7114A84B-8708-5039-B69A-CAFA11B9AA6F" authority="Chen &amp; Wu" authorityName="Chen &amp; Wu" authorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus" status="gen. nov.">Sinoarinia Chen &amp; Wu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="37">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="type species">
<paragraph pageId="0" pageNumber="37">Type species.</paragraph>
<paragraph pageId="0" pageNumber="37">
<taxonomicName authorityName="Chen, gen. et" authorityYear="2020" class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia feii" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="feii">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia feii</emphasis>
</taxonomicName>
Chen, sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="diagnosis">
<paragraph pageId="0" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="37">Shell minute, dextral, thin, translucent, cylindrical. Penultimate whorl strongly constricted. Last whorl strongly ascending, covering part of penultimate whorl. Protoconch oblique. Apical whorls depressed, with strong or weak ribs. Aperture rounded; peristome double, expanded but not reflected. Columellar lamella not visible from aperture but internally present; one parietal lamella and one palatal plica extending inwards from last whorl to penultimate whorl (in type species). Palatal plica visible through semitransparent parietal wall. Constriction absent or very weak where lamellae and plica terminate. Umbilicus closed, forming a chink.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="etymology">
<paragraph pageId="0" pageNumber="37">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="37">
Combination of the Latin prefix sino- (= China) with
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Arinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Arinia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="distribution">
<paragraph pageId="0" pageNumber="37">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="37">Southern China (Hubei, Guizhou), northern Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="remarks">
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="37">
The new genus can be distinguished from the sympatric diplommatinid genera (
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Arinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Arinia</emphasis>
</taxonomicName>
,
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Diplommatina</emphasis>
</taxonomicName>
Benson, 1849 and
<taxonomicName authorityName="Kobelt &amp; Moellendorff" authorityYear="1900" class="Gastropoda" family="Diplommatinidae" genus="Gastroptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastroptychia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Gastroptychia</emphasis>
</taxonomicName>
Kobelt &amp;
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1900) by its constricted penultimate whorl and the ascending last whorl (
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="37" refId="B8" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." year="1902">Kobelt 1902</bibRefCitation>
;
<bibRefCitation author="Zilch, A" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="37" refId="B20" refString="Zilch, A, 1953. Die Typen und Typoide des Natur-Museums Senckenberg, 9: Mollusca, Cyclophoridae, Diplommatininae. Archiv fuer Molluskenkunde 82(1/3): 1-47." title="Die Typen und Typoide des Natur-Museums Senckenberg, 9: Mollusca, Cyclophoridae, Diplommatininae. Archiv fuer Molluskenkunde 82 (1 / 3): 1 - 47." year="1953">Zilch 1953</bibRefCitation>
;
<bibRefCitation author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="37" pagination="87 - 138" refId="B16" refString="Vermeulen, JJ, 1996. Notes on the non-marine molluscs of the island of Borneo 8, the genus Arinia; additions to the genera Diplommatina and Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae). Basteria 60 (4-6): 87 - 138" title="Notes on the non-marine molluscs of the island of Borneo 8, the genus Arinia; additions to the genera Diplommatina and Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae)." volume="60" year="1996">Vermeulen 1996</bibRefCitation>
;
<bibRefCitation author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="37" pagination="81 - 92" refId="B17" refString="Vermeulen, JJ, Phung, LC, Truong, QT, 2007. New species of terrestrial molluscs (Caenogastropoda, Pupinidae and Pulmonata, Vertiginidae) of the Hon Chong-Ha Tien limestone hills, Southern Vietnam. Basteria 71 (1-3): 81 - 92" title="New species of terrestrial molluscs (Caenogastropoda, Pupinidae and Pulmonata, Vertiginidae) of the Hon Chong-Ha Tien limestone hills, Southern Vietnam." volume="71" year="2007">Vermeulen et al. 2007</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5252/zoosystema2018v40a3" author="Pall-Gergely, B" journalOrPublisher="Zoosystema" pageId="0" pageNumber="37" pagination="59 - 66" refId="B13" refString="Pall-Gergely, B, Hunyadi, A, 2018. Four new cyclophoroid species from Thailand and Laos (Gastropoda: Caenogastropoda: Alycaeidae, Diplommatinidae, Pupinidae). Zoosystema 40 (3): 59 - 66, DOI: https://doi.org/10.5252/zoosystema2018v40a3" title="Four new cyclophoroid species from Thailand and Laos (Gastropoda: Caenogastropoda: Alycaeidae, Diplommatinidae, Pupinidae)." url="https://doi.org/10.5252/zoosystema2018v40a3" volume="40" year="2018">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
and Hunyadi 2018
</bibRefCitation>
). In general,
<taxonomicName authorityName="Chen &amp; Wu" authorityYear="2020" class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia</emphasis>
</taxonomicName>
gen. nov. has a cylindrical shell, depressed apex, and deeply incised lamellae and plica, which differs from the conical shell with an exposed columellar lamella in
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Diplommatina</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kobelt &amp; Moellendorff" authorityYear="1900" class="Gastropoda" family="Diplommatinidae" genus="Gastroptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastroptychia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Gastroptychia</emphasis>
</taxonomicName>
.
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Arinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Arinia</emphasis>
</taxonomicName>
usually lacks internal apertural barriers (
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="37" refId="B8" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." year="1902">Kobelt 1902</bibRefCitation>
;
<bibRefCitation author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="37" pagination="87 - 138" refId="B16" refString="Vermeulen, JJ, 1996. Notes on the non-marine molluscs of the island of Borneo 8, the genus Arinia; additions to the genera Diplommatina and Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae). Basteria 60 (4-6): 87 - 138" title="Notes on the non-marine molluscs of the island of Borneo 8, the genus Arinia; additions to the genera Diplommatina and Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae)." volume="60" year="1996">Vermeulen 1996</bibRefCitation>
;
<bibRefCitation author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="37" pagination="81 - 92" refId="B17" refString="Vermeulen, JJ, Phung, LC, Truong, QT, 2007. New species of terrestrial molluscs (Caenogastropoda, Pupinidae and Pulmonata, Vertiginidae) of the Hon Chong-Ha Tien limestone hills, Southern Vietnam. Basteria 71 (1-3): 81 - 92" title="New species of terrestrial molluscs (Caenogastropoda, Pupinidae and Pulmonata, Vertiginidae) of the Hon Chong-Ha Tien limestone hills, Southern Vietnam." volume="71" year="2007">Vermeulen et al. 2007</bibRefCitation>
), which are well developed in
<taxonomicName authorityName="Chen &amp; Wu" authorityYear="2020" class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia</emphasis>
</taxonomicName>
gen. nov. The constricted structure of the new genus is similar to that found in the non-sympatric diplommatinid genus
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Diancta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diancta" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Diancta</emphasis>
</taxonomicName>
E. von Martens, 1864, which differs from the new genus by its mostly sinistral coiling and conical whorls as showed in
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Diancta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diancta" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Diancta</emphasis>
</taxonomicName>
(s. str.) (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.487.8463" author="Neubert, E" journalOrPublisher="ZooKeys" pageId="0" pageNumber="37" pagination="1 - 85" refId="B11" refString="Neubert, E, Bouchet, P, 2015. The Diplommatinidae of Fiji-a hotspot of Pacific land snail biodiversity (Caenogastropoda, Cyclophoroidea). ZooKeys 487: 1 - 85, DOI: https://doi.org/10.3897/zookeys.487.8463" title="The Diplommatinidae of Fiji-a hotspot of Pacific land snail biodiversity (Caenogastropoda, Cyclophoroidea)." url="https://doi.org/10.3897/zookeys.487.8463" volume="487" year="2015">Neubert and Bouchet 2015</bibRefCitation>
), or by the constricted penultimate whorl which is not partially covered by the last whorl, as showed in
<taxonomicName authority="" class="Gastropoda" family="Diplommatinidae" genus="Diancta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diancta (Paradiancta)" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="subGenus" subGenus="Paradiancta">Diancta (Paradiancta)</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1895 (
<bibRefCitation author="Egorov, R" journalOrPublisher="Treasure of Russian Shells, Supplement" pageId="0" pageNumber="37" pagination="1 - 62" refId="B3" refString="Egorov, R, 2013. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. Treasure of Russian Shells, Supplement 3 (3): 1 - 62" title="A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae." volume="3" year="2013">Egorov 2013</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="37">
Three species,
<taxonomicName authorityName="Y.Li, W.-C.Zhou &amp; T.-C.Luo" authorityYear="2005" class="Gastropoda" family="Diplommatinidae" genus="Arinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arinia mirifica" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="mirifica">
<emphasis italics="true" pageId="0" pageNumber="37">Arinia mirifica</emphasis>
</taxonomicName>
Li, Zhuo &amp; Luo, 2005,
<taxonomicName lsidName="A. maolanensis" pageId="0" pageNumber="37" rank="species" species="maolanensis">
<emphasis italics="true" pageId="0" pageNumber="37">A. maolanensis</emphasis>
</taxonomicName>
Zhang, Chen &amp; Zhuo, 2013, and
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina aesopus" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="aesopus">
<emphasis italics="true" pageId="0" pageNumber="37">Diplommatina aesopus</emphasis>
</taxonomicName>
Bavay &amp; Dautzenberg, 1904, which some authors currently assign to
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Arinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Arinia</emphasis>
</taxonomicName>
(
<bibRefCitation author="Li, Y" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="37" pagination="67 - 69" refId="B9" refString="Li, Y, Zhou, W, Luo, T, 2005. A new species of Cyclophoridae from Guizhou Province, China, (Prosobranchia: Mesogastropoda: Cyclophoridae). Acta Zootaxonomica Sinica 30 (1): 67 - 69" title="A new species of Cyclophoridae from Guizhou Province, China, (Prosobranchia: Mesogastropoda: Cyclophoridae)." volume="30" year="2005">Li et al. 2005</bibRefCitation>
;
<bibRefCitation author="Zhang, W" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="37" pagination="773 - 775" refId="B19" refString="Zhang, W, Chen, D, Zhou, W, 2013. A new species of the genus Arinia H. and A. Adams from China (Prosobranchia: Caenogastropoda: Diplommatinidae). Acta Zootaxonomica Sinica 38 (4): 773 - 775" title="A new species of the genus Arinia H. and A. Adams from China (Prosobranchia: Caenogastropoda: Diplommatinidae)." volume="38" year="2013">Zhang et al. 2013</bibRefCitation>
) and
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Diplommatina</emphasis>
</taxonomicName>
(Bavay &amp; Dautzenberg, 1904), are here transferred to
<taxonomicName authorityName="Chen &amp; Wu" authorityYear="2020" class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="included species">
<paragraph pageId="0" pageNumber="37">Included species.</paragraph>
<paragraph pageId="0" pageNumber="37">
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia aesopus" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="aesopus">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia aesopus</emphasis>
</taxonomicName>
(Bavay &amp; Dautzenberg, 1904) comb. nov.,
<taxonomicName authorityName="Chen, gen. et" authorityYear="2020" class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia feii" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="feii">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia feii</emphasis>
</taxonomicName>
Chen sp. nov.,
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia maolanensis" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="maolanensis">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia maolanensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Zhang, W" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="37" pagination="773 - 775" refId="B19" refString="Zhang, W, Chen, D, Zhou, W, 2013. A new species of the genus Arinia H. and A. Adams from China (Prosobranchia: Caenogastropoda: Diplommatinidae). Acta Zootaxonomica Sinica 38 (4): 773 - 775" title="A new species of the genus Arinia H. and A. Adams from China (Prosobranchia: Caenogastropoda: Diplommatinidae)." volume="38" year="2013">Zhang et al., 2013</bibRefCitation>
) comb. nov., and
<taxonomicName class="Gastropoda" family="Diplommatinidae" genus="Sinoarinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoarinia mirifica" order="Architaenioglossa" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="mirifica">
<emphasis italics="true" pageId="0" pageNumber="37">Sinoarinia mirifica</emphasis>
</taxonomicName>
(
<bibRefCitation author="Li, Y" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="37" pagination="67 - 69" refId="B9" refString="Li, Y, Zhou, W, Luo, T, 2005. A new species of Cyclophoridae from Guizhou Province, China, (Prosobranchia: Mesogastropoda: Cyclophoridae). Acta Zootaxonomica Sinica 30 (1): 67 - 69" title="A new species of Cyclophoridae from Guizhou Province, China, (Prosobranchia: Mesogastropoda: Cyclophoridae)." volume="30" year="2005">Li et al., 2005</bibRefCitation>
) comb. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="vernacular name">
<paragraph pageId="0" pageNumber="37">Vernacular name.</paragraph>
<paragraph pageId="0" pageNumber="37">
<normalizedToken originalValue="华阿勇螺属">华阿勇螺属</normalizedToken>
</paragraph>
</subSubSection>
</treatment>
</document>