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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23685" ID-PMC="PMC5904534" ID-Pensoft-Pub="1313-2970-745-101" ID-Pensoft-UUID="047DFFCAFFA5F32EA97C873F4708943F" ID-PubMed="29670449" ID-Zenodo-Dep="1222435" ID-ZooBank="8785DC6BC2A244FD94B6243EB07C717F" ModsDocID="1313-2970-745-101" checkinTime="1521799180220" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. &amp; Branham, Marc A." docDate="2018" docId="39CD1809998A57C0AB92940D37848B23" docLanguage="en" docName="ZooKeys 745: 101-378" docOrigin="ZooKeys 745" docPubDate="2018-03-22" docSource="http://dx.doi.org/10.3897/zookeys.745.23685" docTitle="Augoderia boliviana Endrodi 1981" docType="treatment" docVersion="3" id="047DFFCAFFA5F32EA97C873F4708943F" lastPageNumber="127" masterDocId="047DFFCAFFA5F32EA97C873F4708943F" masterDocTitle="Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)" masterLastPageNumber="378" masterPageNumber="101" pageNumber="127" updateTime="1668165599527" updateUser="ExternalLinkService">
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<mods:title>Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moore, Matthew R.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
<mods:nameIdentifier type="email">cyclocephala@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cave, Ronald D.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Branham, Marc A.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
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<taxonomicName LSID="39CD1809-998A-57C0-AB92-940D37848B23" authority="Endrodi, 1981" authorityName="Endrodi" authorityYear="1981" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia boliviana" order="Coleoptera" pageId="26" pageNumber="127" phylum="Arthropoda" rank="species" species="boliviana">
Augoderia boliviana
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, 1981
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<taxonomicName authorityName="Endrodi" authorityYear="1981" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia boliviana" order="Coleoptera" pageId="26" pageNumber="127" phylum="Arthropoda" rank="species" species="boliviana">Augoderia boliviana</taxonomicName>
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
, 1981: 198 [original combination].
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="127" type="types">
<paragraph pageId="26" pageNumber="127">Types.</paragraph>
<paragraph pageId="26" pageNumber="127">
Holotype ♂ at ZSMC (
<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B320" refString="Endrodi, S, 1981. Neue und seltene Dynastinen aus Suedamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici 73: 197-202." title="Neue und seltene Dynastinen aus Suedamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici 73: 197 - 202." year="1981">
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
1981
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="127" type="distribution">
<paragraph pageId="26" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="26" pageNumber="127">BOLIVIA: Santa Cruz.</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="127" type="references">
<paragraph pageId="26" pageNumber="127">References.</paragraph>
<paragraph pageId="26" pageNumber="127">
<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B320" refString="Endrodi, S, 1981. Neue und seltene Dynastinen aus Suedamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici 73: 197-202." title="Neue und seltene Dynastinen aus Suedamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici 73: 197 - 202." year="1981">
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
1981
</bibRefCitation>
,
<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
,
<bibRefCitation author="Ponchel, Y" pageId="214" pageNumber="315" refId="B907" refString="Ponchel, Y, 2009. Deux nouvelles especes de Cyclocephalini de Guyane francaise (ColeopteraDynastidae). L'Entomologiste 65: 183-185." title="Deux nouvelles especes de Cyclocephalini de Guyane francaise (ColeopteraDynastidae). L'Entomologiste 65: 183 - 185." year="2009">Ponchel 2009</bibRefCitation>
,
<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik 2005</bibRefCitation>
,
<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B659" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma.X Supplement 5, Pilsen, 278 pp." title="Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Pilsen, 278 pp." year="2012">2012</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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.
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<subSubSection id="353F28CDCC632D2F2EF66CED238ECA2C" type="description">
<paragraph id="B313E931B3858C27DAFAE9F7AE183436" pageNumber="153">Das mir vorliegende ungefluegelte Weibchen ist 11 Mm. lang (die Arbeiter 9 — 10 Mm.), dem Arbeiter sehr aehnlich, die Pronotumdornen viel kuerzer, die Basalflaeche des Metanotum verkehrt-trapezfoermig, etwas kuerzer als vorne breit, hinten mit zwei sehr kleinen Zaehnen, sie geht ohne deutliche Grenze in die von oben nach unten concave, ziemlich quadratische abschuessige Flaeche ueber.</paragraph>
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<paragraph id="89657195B8E1AA2AF5534AA19EB6EF9E" pageNumber="153">Camerun (Dr. Brauns, Berliner Museum), Cap Lopez am Suedostende des Golfs von Guinea (Dr. Brauns).</paragraph>
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Linnaeus
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,
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2
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: 795. 1753
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.
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<paragraph id="CECDEDE6451DD1DE1C80BF6E42EEC939" pageNumber="610">&quot;Habitat in Insula Carolina Balthici.&quot; RCN: 5817.</paragraph>
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<paragraph id="C00A2159D89B3C7C460E8478043BC9EA" pageNumber="610">
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(van Raamsdonk in Wisskirchen in
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108: 105. 1997): Herb. Linn. No. 950.1 (
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)
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.
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<paragraph id="369282B9D37EE072250E51EFF94D9E6D" pageNumber="610">
Current name:
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<emphasis id="F337E08D2954AB08D48B6068DF0F9572" bold="true" italics="true" pageNumber="610">Lactuca quercina</emphasis>
L.
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(
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).
</paragraph>
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<paragraph id="4531E2B2B1A37CE2655064FE086778A1" pageNumber="610">
<emphasis id="ABD93029DAD306A90CB6FF69A4925374" italics="true" pageNumber="610">Note:</emphasis>
Stearn (in
<emphasis id="9CF15CD23C52D48E444C96D24D4F19B0" italics="true" pageNumber="610">Biol. J. Linn. Soc.</emphasis>
5: 11. 1973), in an account of
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<normalizedToken id="13A6A4377BD908A0A1DC832F7071A8D0" originalValue="Öland">Oeland</normalizedToken>
and Gotland journey of 1741, treated Lilla
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, near Gotland, as the restricted type locality, and noted the existence of 950.1 (LINN). In his paper, he attributed restricted type localities irrespective of whether any material existed in LINN and, where specimens do exist, he does not refer to any of them as type specimens. The collection subsequently designated as the type shows no annotation associating it with Gotland.
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<taxonomicName id="C1F54637F8A25BC91974C30400C5B708" LSID="39CD8A30-AB3D-5E09-A01E-3188B448E6F3" authority="sp." class="Gastropoda" family="Arcidae" genus="Bentharca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bentharca" order="Vetigastropoda" pageId="0" pageNumber="105444" phylum="Mollusca" rank="genus">Bentharca sp.</taxonomicName>
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<paragraph id="6C1F409CA6C3B7B4399E45FF5246469A" pageId="0" pageNumber="105444">Native status</paragraph>
<paragraph id="182971FD8B827521794956090F914B6D" pageId="0" pageNumber="105444">Found on sandy mud bottoms in upper sublittoral zones.</paragraph>
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<paragraph id="24475139984800A0F65A95FE66811EC6" pageId="0" pageNumber="105444">Distribution</paragraph>
<paragraph id="8A5B68A68433E07DA14BC974CE73D070" pageId="0" pageNumber="105444">Bohai Sea of China.</paragraph>
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<paragraph id="34979D75CC2D11750907D27ACA6FD216" pageId="0" pageNumber="105444">Notes</paragraph>
<paragraph id="8C33BA7973AEFD48841C0390DA2296A6" pageId="0" pageNumber="105444">
This species is very similar to
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spp. in shell characters including radial riblets and is covered with lamellate epidermis at interstices, cardinal plate and umbo.
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<paragraph id="F17C54D889C8260EE2DE4E0726D3D5B0" pageId="0" pageNumber="105444">Diagnosis</paragraph>
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Shell minute (2.3
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0.02 mm in length, 1.5
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0.04 mm in height), elongate squarish, thin, moderately inflated (Fig.
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). Antero-posterior margin round, anterior narrower and shorter than posterior; posterior hardly oblique, sides weakly angulated at upper part. Umbo corroded, situated in front of dorsal margin. Surface with 33 radial riblets of various prominence and strong concentric ribs near the umbo; covered with lamellate epidermis at interstices; inner surface margin is toothed. With a depression at the ventral margin. Cardinal plate with 3 teeth on anterior arc and 4 on posterior arc.
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174
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<paragraph id="9CEE9BA14C2B79DE31323E3CAAA653E3" pageId="14" pageNumber="35">Taxon classification Animalia Polydesmida Paradoxosomatidae</paragraph>
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<taxonomicName id="1A962C345A3225E2D9EEC486ACF956F0" ID-CoL="67CR7" authority="Golovatch, 2013" authorityName="Golovatch" authorityYear="2013" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha rosea" order="Polydesmida" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="rosea">Antheromorpha rosea Golovatch, 2013</taxonomicName>
Figs 13, 14, 15, 21
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rosea
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Golovatch, 2013a: 23 (D).
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<taxonomicName id="85D82B5A50854BA6DC375BA289E21323" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha rosea" order="Polydesmida" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="rosea">Antheromorpha rosea</taxonomicName>
-
<bibRefCitation id="CDEB79A2353F01377040C7F7017A4F53" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="28" pageNumber="49" pagination="307 - 332" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIV." volume="22" year="2013 b">Golovatch 2013b</bibRefCitation>
: 308 (D);
<bibRefCitation id="79EFFBAA89A3F708B1E6D1A0D7FFCD91" author="Nguyen, AD" journalOrPublisher="Check List" pageId="28" pageNumber="49" pagination="1132 - 1353" title="A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida)." url="http://www.checklist.org.br/getpdf?SL107-12" volume="9" year="2013">Nguyen and Sierwald 2013</bibRefCitation>
: 1235 (M).
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<paragraph id="C70B2B6F1E13E1277457934741A77AB8" pageId="15" pageNumber="36">Material examined.</paragraph>
<paragraph id="F83D0178D44D4EF2AEE95204DB4D3EDA" pageId="15" pageNumber="36">
5 ♂, 9 ♀ (CUMZ), 1 ♂, 1 ♀ (ZMUM
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3057), 1 ♂, 1 ♀ (ZMUC), 1 ♂, 1 ♀ (NHMW), Thailand, Chiang Mai Province, Mae Rim District, Queen Sirikit Botanic Garden,
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,
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, ca 640 m a.s.l., 25.09.2014, leg. N. Likhitrakrn. 2 ♂, 30 ♀ (CUMZ), same District, Pong Yang, Ban Muang Kham,
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,
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,
<geoCoordinate id="4CFA9928D33D16BD74337E6FFF0FA5F6" direction="east" orientation="longitude" precision="15" value="98.82972">98°49'47&quot;E</geoCoordinate>
, ca 590 m a.s.l., 24.10.2009, leg. N. Likhitrakarn. 1 ♂ (CUMZ), same Province, Chiang Dao District, Wat Tam Pha Plong,
<geoCoordinate id="735D9DC41C663976B85071A46C1208BE" direction="east" orientation="longitude" precision="15" value="19.40361">19°24'13&quot;E</geoCoordinate>
,
<geoCoordinate id="E56EF622224FC384BD5E17B0BA8018B8" direction="east" orientation="longitude" precision="15" value="98.92111">98°55'16&quot;E</geoCoordinate>
, 470 m a.s.l., 28.09.2010, leg. N. Likhitrakarn. 1 ♂ (CUMZ), same Province, Mae Taeng District, Cave Buathong,
<geoCoordinate id="BFB57D139779112E35D30C0E30ABD391" direction="north" orientation="latitude" precision="1" value="19.075294">19° 4'31.06&quot;N</geoCoordinate>
,
<geoCoordinate id="635D367C04B5B98BCA0653E34A43F837" direction="east" orientation="longitude" precision="1" value="99.08596">99° 5'9.45&quot;E</geoCoordinate>
, ca 530 m a.s.l., 22.11.2012, leg. N. Likhitrakarn.
</paragraph>
</subSubSection>
<subSubSection id="24702303D160F78F38004F204BFE1123" lastPageId="18" lastPageNumber="39" pageId="16" pageNumber="37" type="descriptive notes">
<paragraph id="B8DDECF4B79912A10A371690DA27449C" pageId="16" pageNumber="37">
<pageBreakToken id="D93DBACF98D8BA031BFAB9E2478B1FDB" pageId="16" pageNumber="37" start="start">Descriptive</pageBreakToken>
notes.
</paragraph>
<paragraph id="D4A78C55A2F8B8AEDA05F196CF658EDB" pageId="16" pageNumber="37">Length 33.5-38.0 (♂) or 34.0-44.5 mm (♀), width of midbody pro- and metazonae 2.6-3.5 and 4.4-5.0 mm (♂), 3.2-4.2 and 4.9-5.8 mm (♀), respectively.</paragraph>
<paragraph id="89CF01385DF76B17365C6084CAECA6C8" pageId="16" pageNumber="37">
Coloration of live animals pinkish (Fig. 13A), with an anterior black band on metaterga and collum; head and antennae blackish, legs dark to light brown; coloration in alcohol, after six months of preservation, faded to light pinkish to pale yellowish (Fig. 13
<normalizedToken id="A8C327368BD695F986BAB7CD7512A503" originalValue="B–H">B-H</normalizedToken>
), with a dark brown to blackish brown band on anterior metaterga and collum; head and antennae blackish to light brown, legs light brown to pale yellowish.
</paragraph>
<caption id="54F74BFC1F578428EE969AF8612B503D" pageId="16" pageNumber="37">
<paragraph id="98E0E17E44B0351749B85AE822A14CF1" pageId="16" pageNumber="37">
Figure 13.
<taxonomicName id="ABC1561E8E9B391A3688724F09A239C3" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha rosea" order="Polydesmida" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="rosea">Antheromorpha rosea</taxonomicName>
Golovatch, 2013, ♂ from Queen Sirikit Botanic Garden. A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively
<normalizedToken id="6FE800E1630F6094A01B2CC254AA171E" originalValue="F–H">F-H</normalizedToken>
posterior part of body, lateral, dorsal and ventral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<paragraph id="452ED9875A47E57A9F3AB2E8EC3D4302" lastPageId="18" lastPageNumber="39" pageId="17" pageNumber="38">
<pageBreakToken id="3236594027A016ECF9198ADA41C381D1" pageId="17" pageNumber="38" start="start">Antennae</pageBreakToken>
long (Fig. 13A, C), extending behind metaterga 3 when stretched dorsally (♂, ♀). In width, head &lt;segment 3 = 4 &lt;collum &lt;segment 2 &lt;5-17 (♂, ♀), gently and gradually tapering thereafter (Fig. 13B). Paraterga very strongly developed (Fig. 13
<normalizedToken id="AF272E9D077411A2623E5D2EA97EB2A2" originalValue="B–H">B-H</normalizedToken>
), mostly slightly upturned, all lying below dorsum, set at about upper 1/3 of midbody height, caudal corner almost to fully pointed, increasingly acutangular, from narrowly rounded to nearly pointed, especially strongly so in segment 15, thereafter slightly curved mesad (Fig. 13B, D, F). Pleurosternal carinae complete crests with a sharp caudal tooth in segment 2, likewise a sharp caudal tooth in segments 3 and 4, a small, mostly sharp tooth until segment 16 (♂, ♀) (Fig. 13C, E). Sterna delicately
<pageBreakToken id="0E424C3E368C42C91B703AEB7D12A196" pageId="18" pageNumber="39" start="start">and</pageBreakToken>
sparsely setose, without modifications, but with a pair of small, rounded, fully separated cones between ♂ coxae 4 (Fig. 13I, J).
</paragraph>
</subSubSection>
<subSubSection id="2388C246EAD7AEA42BDDFCA3B4780114" lastPageId="19" lastPageNumber="40" pageId="18" pageNumber="39" type="remarks">
<paragraph id="8EE888B9C2DC3D1E4EA5A207A43AD780" pageId="18" pageNumber="39">Remarks.</paragraph>
<paragraph id="910658D93886BCCA8D2FDD0ED917BDB3" lastPageId="19" lastPageNumber="40" pageId="18" pageNumber="39">
The available descriptions (
<bibRefCitation id="94696CBA2396E3E38E039E4AC08F1CB5" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="28" pageNumber="49" pagination="1 - 31" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIII." volume="22" year="2013 a">Golovatch 2013a</bibRefCitation>
,
<bibRefCitation id="31936BD5762BF9F887459BA497EEA2B5" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="28" pageNumber="49" pagination="307 - 332" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIV." volume="22" year="2013 b">2013b</bibRefCitation>
) of this species were sufficiently detailed to necessitate only a few notes on variation and some new illustrations (Figs 13-15) to show coloration, certain structural details and the gonopod conformation based on new material. This species was described from the ♂ holotype (kept in Senckenberg Museum Frankfurt, Germany) from Gaoligong Shan Moutains, south of Pianma,
<geoCoordinate id="F49A09C5E0338045097D0B214E692474" direction="north" orientation="latitude" precision="925" value="25.966667">25°58'N</geoCoordinate>
,
<geoCoordinate id="790F45F9C0D4874378AEEBD5C67FCAEF" direction="east" orientation="longitude" precision="925" value="98.666664">98°40'E</geoCoordinate>
, 1600-1700 m a.s.l., Yunnan Province, China (
<bibRefCitation id="1422A56F893EEAE7DA7906B1D42568AC" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="28" pageNumber="49" pagination="1 - 31" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIII." volume="22" year="2013 a">Golovatch 2013a</bibRefCitation>
), a little later reported (1 ♂, 1 ♀, deposited in the National Natural
<pageBreakToken id="EDDF01ECF654789E1C9FBDD68CA528D8" pageId="19" pageNumber="40" start="start">History</pageBreakToken>
Museum, Sofia, Bulgaria) nearly from the same place (
<bibRefCitation id="946D5C3B2507FF8DB63EFA84E1CCB717" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="28" pageNumber="49" pagination="307 - 332" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIV." volume="22" year="2013 b">Golovatch 2013b</bibRefCitation>
). Even though both these Yunnan localities (Fig. 21) lie far away (ca 730 air-km) from the new Thai records, even despite minor variations, the species identity is beyond doubt.
</paragraph>
<caption id="F4B2BCC3C4981A8A1742D5CF2C4A5B62" pageId="19" pageNumber="40">
<paragraph id="90A48D2B696AEB413315D9B56CED9ECC" pageId="19" pageNumber="40">
Figure 14.
<taxonomicName id="7184466F7D6BA11488BD7CD78F0A8931" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha rosea" order="Polydesmida" pageId="19" pageNumber="40" phylum="Arthropoda" rank="species" species="rosea">Antheromorpha rosea</taxonomicName>
Golovatch, 2013, ♂ from Queen Sirikit Botanic Garden. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.4 mm.
</paragraph>
</caption>
<caption id="0B130C1E7A9905DF81DBD47DE527D939" pageId="19" pageNumber="40">
<paragraph id="593468DA36059279C2196316DBBFC712" pageId="19" pageNumber="40">
Figure 15.
<taxonomicName id="433F9BEEA9AD238CA9838CF72FE2879F" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha rosea" order="Polydesmida" pageId="19" pageNumber="40" phylum="Arthropoda" rank="species" species="rosea">Antheromorpha rosea</taxonomicName>
Golovatch, 2013, ♂ from Queen Sirikit Botanic Garden. A, B right gonopod, mesal and lateral views, respectively
<normalizedToken id="613065BB0E28D1352C671E91D671E8A0" originalValue="C–F">C-F</normalizedToken>
distal part of right gonopod, subcaudal, suboral, submesal and sublateral views, respectively. Scale bar: 0.2 mm.
</paragraph>
</caption>
<paragraph id="B8090BF566075B5A64F3E5170F4CBE31" pageId="19" pageNumber="40">
At least in Thailand, adult
<taxonomicName id="8D2E6BB74D8A33A9BCF244BB800A77C2" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha rosea" order="Polydesmida" pageId="19" pageNumber="40" phylum="Arthropoda" rank="species" species="rosea">Antheromorpha rosea</taxonomicName>
have been found to occur every year only for one or two weeks in September or October, disappearing thereafter.
</paragraph>
</subSubSection>
</treatment>
</document>

253
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<mods:title>The tremendous diversity of Labiobaetis Novikova &amp; Kluge in Indonesia (Ephemeroptera, Baetidae)</mods:title>
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<treatment ID-GBIF-Taxon="160631150" LSID="urn:lsid:zoobank.org:act:5EAF4EA0-6625-48E2-9EC9-079ED6C55504" httpUri="http://treatment.plazi.org/id/39CDDCC103A650E9ABDAF1D5E92FFFAF" lastPageNumber="1" pageId="0" pageNumber="1">
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12.
<taxonomicName LSID="39CDDCC1-03A6-50E9-ABDA-F1D5E92FFFAF" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis pakpak" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pakpak">Labiobaetis pakpak</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Figures 20</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Labiobaetis pakpak sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape." figureDoi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure21">21</figureCitation>
,
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Habitus, larvae, dorsal view: a Labiobaetis weifangae sp. nov. b Labiobaetis itineris sp. nov., without head c Labiobaetis lubu sp. nov. d Labiobaetis pakpak sp. nov." figureDoi="10.3897/zookeys.895.38576.figure48" httpUri="https://binary.pensoft.net/fig/361939" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure48">48d</figureCitation>
,
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Distribution of Labiobaetis in Indonesia." figureDoi="10.3897/zookeys.895.38576.figure55" httpUri="https://binary.pensoft.net/fig/361946" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure55">55a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combination of characters: A) dorsal surface of labrum with submarginal arc of 18-19 clavate setae; B) labial palp segment II with large, lobed distomedial protuberance, segment III slightly pentagonal; C) left mandible without setae at apex of mola; D) fore femur rather slender, length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with a row of 12-15 spine-like setae and many fine, simple setae; E) paraproct distally expanded, with&gt; 40 stout marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Figs 20</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Labiobaetis pakpak sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape." figureDoi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure21">21</figureCitation>
,
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Habitus, larvae, dorsal view: a Labiobaetis weifangae sp. nov. b Labiobaetis itineris sp. nov., without head c Labiobaetis lubu sp. nov. d Labiobaetis pakpak sp. nov." figureDoi="10.3897/zookeys.895.38576.figure48" httpUri="https://binary.pensoft.net/fig/361939" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure48">48d</figureCitation>
). Body length 3.7 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
. Head, thorax, and abdomen dorsally brown, head and thorax with bright median, dorsal suture. Head, thorax, and abdomen ventrally light brown, legs transparent, caudal filaments light brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
with scape and pedicel subcylindrical, without distolateral process at scape; flagellum with lanceolate spines and fine, simple setae on apex of each segment.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Fig. 20a</figureCitation>
). Rectangular, length 0.7
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with long, fine, simple setae scattered over surface; submarginal arc of setae composed of 18-19 long, clavate setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid setae; ventral surface with five short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Fig. 20b, c</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Fig. 20d, e</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 3 + 3 denticles and one minute intermediate denticle. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hypopharynx</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Fig. 20f</figureCitation>
). Lingua approx. as long as superlingua. Lingua longer than broad; medial tuft of stout setae well developed; distal half laterally expanded. Superlingua straight; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Fig. 20g</figureCitation>
). Galea-lacinia with two simple, robust apical setae under crown. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and 3-4 medium, simple setae. Maxillary palp 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of galea-lacinia; two segmented; palp segment II 1.3
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment rounded, with slight excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pakpak sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure20">Fig. 20h, i</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with 4-6 spine-like setae increasing in length distally; apex with two long and one medium, robust setae; outer margin with 4-6 long, spine-like setae increasing in length distally; ventral surface with short, fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and a row of three or four medium, simple setae in anteromedial area; dorsally with a row of three long, spine-like setae near inner margin. Labial palp with segment I 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with large, lobed distomedial protuberance; distomedial protuberance 0.8
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; inner and outer margin with short, fine, simple setae; dorsally with two long, spine-like, simple setae near outer margin. Segment III slightly pentagonal; apex slightly truncate; length 1.2
<normalizedToken originalValue="×">x</normalizedToken>
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<caption doi="10.3897/zookeys.895.38576.figure20" httpUri="https://binary.pensoft.net/fig/361910" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis pakpak" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pakpak">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis pakpak</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
Labrum
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
Right mandible
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
Right prostheca
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
Left mandible
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
Left prostheca
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">Hypopharynx</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
Maxilla
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
Labium
<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
Apex of paraglossa.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Hind wing pads</emphasis>
absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Labiobaetis pakpak sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape." figureDoi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure21">Fig. 21a, b</figureCitation>
). Ratio of foreleg segments 1.3:1.0:0.5:0.2.
<emphasis italics="true" pageId="0" pageNumber="1">Femur</emphasis>
. Length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with a row 12-15 spine-like setae and many fine, simple setae; length of setae 0.23
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded; with 1-2 pairs of curved, spine-like setae and some short, stout setae. Many stout, lanceolate setae scattered along the ventral margin; femoral patch poorly developed.
<emphasis italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
. Dorsal margin with a row of short, stout setae (not always present) and fine, simple setae. Ventral margin with a row of short, spine-like setae, on apex some stout, spine-like, partially bipectinate setae and a tuft of fine, simple setae. Patellotibial suture present on basal 1/3.
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
. Dorsal margin with a row of fine, simple setae. Ventral margin with a row of curved, spine-like setae. Tarsal claw with one row of 10-11 denticles; distally pointed; with three stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tergum</emphasis>
(
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Labiobaetis pakpak sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape." figureDoi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure21">Fig. 21c</figureCitation>
). Surface with rows of U-shaped scale bases and scattered micropores. Posterior margin of tergum IV with triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gills</emphasis>
(
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Labiobaetis pakpak sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape." figureDoi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure21">Fig. 21d</figureCitation>
). Present on segments
<normalizedToken originalValue="II–VII">II-VII</normalizedToken>
. Margin with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Gill IV as long as length of segments V and VI combined. Gill VII as long as length of segments VIII and 1/2 IX combined.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Labiobaetis pakpak sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape." figureDoi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure21">Fig. 21e</figureCitation>
). Distally expanded, with&gt; 40 stout marginal spines. Surface scattered with U-shaped scale bases and micropores. Cercotractor with numerous small marginal spines.
</paragraph>
<caption doi="10.3897/zookeys.895.38576.figure21" httpUri="https://binary.pensoft.net/fig/361911" pageId="0" pageNumber="1" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis pakpak" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pakpak">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis pakpak</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
Foreleg
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
Fore claw
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
Tergum IV
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
Gill IV
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
Paraproct
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
Scape.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Dedicated to the indigenous Pakpak people from Sumatra.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Indonesia: Sumatra.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="1">The specimens were collected in forest at an altitude of 490 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type-material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Larva (on slide, GBIFCH 00592219), Indonesia, Sumatra Barat, Sijunjung / Muara area, forest, 488 m, 10.XI.2011,
<geoCoordinate degrees="00" direction="south" minutes="40.10" orientation="latitude" precision="9" value="-0.6683333">00°40.10'S</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="07.26" orientation="longitude" precision="9" value="101.121">101°07.26'E</geoCoordinate>
, M. Balke leg. (UN7). Temporary deposited in MZL before definitely housed in MZB.
<emphasis bold="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
10 larvae (2 on slides, GBIFCH 00592221, GBIFCH 00592220, 3 in alcohol, GBIFCH 00235851, GBIFCH 00235781, GBIFCH 00515344, deposited in MZL; 5 in alcohol, GBIFCH 00515345, deposited in ZSM), same data as holotype.
</paragraph>
</subSubSection>
</treatment>
</document>