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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.9.e67134" ID-PMC="PMC8486760" ID-Pensoft-Pub="1314-2828-9-e67134" ID-Pensoft-UUID="54EEB1B3D94E5239B847CC1AD9587A36" ID-PubMed="34690511" ID-ZooBank="A1151C0727B74F31BC4B6809DA6F87CD" ModsDocID="1314-2828-9-e67134" checkinTime="1632515415000" checkinUser="pensoft" docAuthor="Lindemann, Jon Peder, Soli, Geir &amp; Kjaerandsen, Jostein" docDate="2021" docId="392331B5C1015BB1B99EC0EFE79A4E3C" docLanguage="en" docName="BiodivDatJour 9: e67134" docOrigin="Biodiversity Data Journal 9" docPubDate="2021-09-24" docSource="http://dx.doi.org/10.3897/BDJ.9.e67134" docTitle="Exechia breviflagellata Lindemann 2021, sp. n." docType="treatment" docUuid="952B7008-5F0F-4182-BD34-B96083EA4ACE" docUuidSource="ZooBank" docVersion="2" id="54EEB1B3D94E5239B847CC1AD9587A36" lastPageNumber="67134" masterDocId="54EEB1B3D94E5239B847CC1AD9587A36" masterDocTitle="Revision of the Exechia parva group (Diptera: Mycetophilidae)" masterLastPageNumber="67134" masterPageNumber="67134" pageNumber="67134" updateTime="1668126095059" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Exechia parva group (Diptera: Mycetophilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lindemann, Jon Peder</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6001-7910</mods:nameIdentifier>
<mods:affiliation>UiT - The Arctic University of Norway, Tromso, Norway</mods:affiliation>
<mods:nameIdentifier type="email">jon.p.lindemann@uit.no</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soli, Geir</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5301-6995</mods:nameIdentifier>
<mods:affiliation>Natural History Museum, Oslo, Norway</mods:affiliation>
<mods:nameIdentifier type="email">geir.soli@nhm.uio.no</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kjaerandsen, Jostein</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3104-073X</mods:nameIdentifier>
<mods:affiliation>UiT - The Arctic University of Norway, Tromso, Norway</mods:affiliation>
<mods:nameIdentifier type="email">jostein.kjarandsen@uit.no</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-09-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>67134</mods:start>
<mods:end>67134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.9.e67134</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.9.e67134</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-9-e67134</mods:identifier>
<mods:identifier type="ZooBank">A1151C0727B74F31BC4B6809DA6F87CD</mods:identifier>
<mods:identifier type="Pensoft-UUID">54EEB1B3D94E5239B847CC1AD9587A36</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:952B7008-5F0F-4182-BD34-B96083EA4ACE" httpUri="http://treatment.plazi.org/id/392331B5C1015BB1B99EC0EFE79A4E3C" lastPageNumber="67134" pageId="0" pageNumber="67134">
<subSubSection pageId="0" pageNumber="67134" type="nomenclature">
<paragraph pageId="0" pageNumber="67134">
<taxonomicName LSID="urn:lsid:zoobank.org:act:952b7008-5f0f-4182-bd34-b96083ea4ace" authority="Lindemann" authorityName="Lindemann" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Exechia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exechia breviflagellata" order="Diptera" pageId="0" pageNumber="67134" phylum="Arthropoda" rank="species" species="breviflagellata" status="sp. n.">Exechia breviflagellata Lindemann</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="67134">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67134" type="materials_examined">
<paragraph pageId="0" pageNumber="67134">Materials</paragraph>
<paragraph pageId="0" pageNumber="67134">
<materialsCitation collectingDate="2014-07-24" collectingMethod="Intercept trap, Intercept Trap" collectionCode="BIOUG" collectorName="BIObus 2014" country="Canada" county="Occurrence" elevation="0" latitude="60.748" location="Location" longitude="-137.513" municipality="Pinned" pageId="0" pageNumber="67134" specimenCode="BIOUG 27487 - H 10" specimenCount="1" specimenCount-male="1" stateProvince="Yukon Territory" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="67134">Type status:</emphasis>
<materialsCitation collectingDate="2014-07-24" collectingMethod="Intercept trap, Intercept Trap" collectionCode="BIOUG" collectorName="BIObus 2014" country="Canada" county="Occurrence" elevation="0" latitude="60.748" location="Location" longitude="-137.513" municipality="Pinned" specimenCode="BIOUG 27487 - H 10" specimenCount="1" specimenCount-male="1" stateProvince="Yukon Territory" typeStatus="Holotype">
<typeStatus pageId="0" pageNumber="67134">Holotype</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="67134">
<collectingCounty>Occurrence</collectingCounty>
:
</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="67134">BIOUG27487-H10</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="67134">BIObus 2014</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="67134" type="generic">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="67134">male</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="67134">adult</specimenType>
; preparations:
<collectingMunicipality>Pinned</collectingMunicipality>
, with genitalia in glycerine in separate microvial;
<emphasis bold="true" pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:541FB50AF63F5A69661E9229AF9C170C" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Location" stateProvince="Yukon Territory">Location</location>
:
</emphasis>
country:
<collectingCountry name="Canada" pageId="0" pageNumber="67134">Canada</collectingCountry>
; stateProvince:
<collectingRegion country="Canada" name="Yukon">Yukon Territory</collectingRegion>
; locality:
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:F1889D25EC7D87199B0106C27BCC2CA0" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Kluane National Park and Reserve, Dezadeash River Trail" pageId="0" pageNumber="67134" stateProvince="Yukon Territory">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:D254B9D190B88D676F1A73A179236028" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Kluane National Park" stateProvince="Yukon Territory">Kluane National Park</location>
and
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:AA86B3D465CE27EB13B8C93E5BF36013" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Reserve" stateProvince="Yukon Territory">Reserve</location>
,
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:4680D5BFB590968EB2320255FD8B7115" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Dezadeash River Trail" stateProvince="Yukon Territory">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:A35C77FE6E9816AD9AD6C0EC27DB61A7" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Dezadeash River" stateProvince="Yukon Territory">
Dezadeash
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:F26E32CD386F56A2AC5EBB22203D6D21" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="River Trail" stateProvince="Yukon Territory">River Trail</location>
</location>
</location>
</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="67134">
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.82" unit="m" value="582.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.82" unit="m" value="582.0">582 m</elevation>
</quantity>
</elevation>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="67134" value="60.748">60.748</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="67134" value="-137.513">-137.513</geoCoordinate>
;
<emphasis bold="true" pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:E320C8052E47564D2DAF59E24E57125A" country="Canada" county="Occurrence" latitude="60.748" longitude="-137.513" municipality="Pinned" name="Event" stateProvince="Yukon Territory">Event</location>
:
</emphasis>
samplingProtocol:
<collectingMethod pageId="0" pageNumber="67134">Intercept trap</collectingMethod>
; eventDate:
<collectingDate pageId="0" pageNumber="67134" value="2014-07-24">2014-07-24</collectingDate>
; habitat: Wetland; fieldNotes: 1
<collectingMethod>Intercept Trap</collectingMethod>
|cold and overcast on day of collection|Wetland with grasses and shrubs;
<emphasis bold="true" pageId="0" pageNumber="67134">Record Level:</emphasis>
institutionCode: CBG
<materialsCitation collectingDate="2013-04-28" collectingMethod="Malaise Trap" collectionCode="BIOUG" collectorName="Chris Johnstone" country="Canada" county="Occurrence" elevation="0" latitude="44.7418" location="Female" longitude="-79.8501" municipality="Chris Johnstone" pageId="0" pageNumber="67134" specimenCode="BIOUG 09240 - B 08" specimenCount="1" specimenCount-female="1" stateProvince="Ontario" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="67134">Type status:</emphasis>
<materialsCitation collectingDate="2013-04-28" collectingMethod="Malaise Trap" collectionCode="BIOUG" collectorName="Chris Johnstone" country="Canada" county="Occurrence" elevation="0" latitude="44.7418" location="Female" longitude="-79.8501" municipality="Chris Johnstone" specimenCode="BIOUG 09240 - B 08" specimenCount="1" specimenCount-female="1" stateProvince="Ontario" typeStatus="Paratype">
<typeStatus pageId="0" pageNumber="67134">Paratype</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="67134">
<collectingCounty>Occurrence</collectingCounty>
:
</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="67134">BIOUG09240-B08</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="67134">
<collectingMunicipality>Chris Johnstone</collectingMunicipality>
</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="67134" type="generic">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:E068EA5726E83B5FB99BAFE0BFBA2531" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Female" stateProvince="Ontario">Female</location>
</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="67134">adult</specimenType>
; preparations:
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:4D1FD1D5CDE2D8947B707426090091ED" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Pinned" stateProvince="Ontario">Pinned</location>
, with genitalia in glycerine in separate microvial;
<emphasis bold="true" pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:418992BC4E58B137DF6E79F29269E88A" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Location" stateProvince="Ontario">Location</location>
:
</emphasis>
country:
<collectingCountry name="Canada" pageId="0" pageNumber="67134">Canada</collectingCountry>
; stateProvince:
<collectingRegion country="Canada" name="Ontario">Ontario</collectingRegion>
; locality:
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:D15411B2FB5ADC98ABA2C55FFFC76883" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Georgian Bay Islands National Park, Administration Office, 901 Wye Valley Rd." pageId="0" pageNumber="67134" stateProvince="Ontario">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:9BB63FB01C9FB36C215E880A5FCE9459" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Georgian Bay Islands National Park" stateProvince="Ontario">Georgian Bay Islands National Park</location>
,
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:3310D8AE8694C0C0DD80AB5294331421" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Administration Office" stateProvince="Ontario">Administration Office</location>
, 901
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:BC2E0EB24D7A0AE6390271D07B5DCA80" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Wye Valley Rd." stateProvince="Ontario">Wye Valley Rd.</location>
</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="67134">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">190 m</elevation>
</quantity>
</elevation>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="67134" value="44.7418">44.7418</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="67134" value="-79.8501">-79.8501</geoCoordinate>
;
<emphasis bold="true" pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:A354A4D0D6FF71033CD166DFE366ACBC" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Event" stateProvince="Ontario">Event</location>
:
</emphasis>
samplingProtocol:
<collectingMethod pageId="0" pageNumber="67134">Malaise Trap</collectingMethod>
; eventDate:
<collectingDate pageId="0" pageNumber="67134" value="2013-04-28">2013-04-28</collectingDate>
; habitat: Wetland; fieldNotes: Marsh;
<emphasis bold="true" pageId="0" pageNumber="67134">Record Level:</emphasis>
institutionCode: CBG
<materialsCitation collectingDate="2013-05-23" collectingMethod="Malaise Trap, Malaise traps" collectionCode="BIOUG" collectorName="Chris Johnstone" country="Canada" county="Occurrence" elevation="0" latitude="44.7418" location="Pinned" longitude="-79.8501" municipality="Chris Johnstone" pageId="0" pageNumber="67134" specimenCode="BIOUG 10611 - B 11" specimenCount="1" specimenCount-male="1" stateProvince="Ontario" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="67134">Type status:</emphasis>
<materialsCitation collectingDate="2013-05-23" collectingMethod="Malaise Trap, Malaise traps" collectionCode="BIOUG" collectorName="Chris Johnstone" country="Canada" county="Occurrence" elevation="0" latitude="44.7418" location="Pinned" longitude="-79.8501" municipality="Chris Johnstone" specimenCode="BIOUG 10611 - B 11" specimenCount="1" specimenCount-male="1" stateProvince="Ontario" typeStatus="Paratype">
<typeStatus pageId="0" pageNumber="67134">Paratype</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="67134">
<collectingCounty>Occurrence</collectingCounty>
:
</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="67134">BIOUG10611-B11</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="67134">
<collectingMunicipality>Chris Johnstone</collectingMunicipality>
</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="67134" type="generic">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="67134">male</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="67134">adult</specimenType>
; preparations:
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:DCACFB4EDF9F480C7B4C94D38E0E34D7" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Pinned" stateProvince="Ontario">Pinned</location>
, with genitalia in glycerine in separate microvial;
<emphasis bold="true" pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:3423D1988D46F7F51EB0E1A69D4B9E09" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Location" stateProvince="Ontario">Location</location>
:
</emphasis>
country:
<collectingCountry name="Canada" pageId="0" pageNumber="67134">Canada</collectingCountry>
; stateProvince:
<collectingRegion country="Canada" name="Ontario">Ontario</collectingRegion>
; locality:
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:990DD928A1FF66EAB7DF08875DCC6E21" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Georgian Bay Islands National Park, Administration Office, 901 Wye Valley Rd." pageId="0" pageNumber="67134" stateProvince="Ontario">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:1BCA80E2B8C8B60993BB71DD355C4A12" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Georgian Bay Islands National Park" stateProvince="Ontario">Georgian Bay Islands National Park</location>
,
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:1DE263552F8EBE0B1744D091A74CC6D2" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Administration Office" stateProvince="Ontario">Administration Office</location>
, 901
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:3C9609A031CA9E13B1B9762B811EEABB" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Wye Valley Rd." stateProvince="Ontario">Wye Valley Rd.</location>
</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="67134">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">190 m</elevation>
</quantity>
</elevation>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="67134" value="44.7418">44.7418</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="67134" value="-79.8501">-79.8501</geoCoordinate>
;
<emphasis bold="true" pageId="0" pageNumber="67134">
<location LSID="urn:lsid:plazi:treatment:392331B5C1015BB1B99EC0EFE79A4E3C:A343744DFEA3AF57E8DE50E91A297A74" country="Canada" county="Occurrence" latitude="44.7418" longitude="-79.8501" municipality="Chris Johnstone" name="Event" stateProvince="Ontario">Event</location>
:
</emphasis>
samplingProtocol:
<collectingMethod pageId="0" pageNumber="67134">Malaise Trap</collectingMethod>
; eventDate:
<collectingDate pageId="0" pageNumber="67134" value="2013-05-23">2013-05-23</collectingDate>
; habitat: Wetland; fieldNotes: 2
<collectingMethod>Malaise traps</collectingMethod>
|Marsh;
<emphasis bold="true" pageId="0" pageNumber="67134">Record Level:</emphasis>
institutionCode: CBG
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67134" type="description">
<paragraph pageId="0" pageNumber="67134">Description</paragraph>
<paragraph pageId="0" pageNumber="67134">
Male (n = 2): Body length 2.9-3.2 mm. Wing length 2.3-2.5 mm.
<emphasis bold="true" pageId="0" pageNumber="67134">Colouration</emphasis>
(Dry specimen). Head dark brown; face and clypeus brown; labellum pale brown; palpus whitish-yellow. Antenna with scape and pedicel brown; flagellum brown. Scutum, lateral sclerites and propleura brown; halteres whitish-yellow. Legs whitish-yellow. Abdomen brown. Terminalia yellow.
<emphasis bold="true" pageId="0" pageNumber="67134">Head</emphasis>
. Frons and vertex covered with pale setae. Clypeus covered with few (15-17) pale setae, evenly distributed. Antenna short, 1.45-1.5 times as long as length from vertex to ventral margin of clypeus; flagellomeres quadrate, with sixth flagellomere 0.8-0.9 times as long as wide.
<emphasis bold="true" pageId="0" pageNumber="67134">Thorax</emphasis>
. Scutum covered with short pale brown setae.
<emphasis bold="true" pageId="0" pageNumber="67134">Legs</emphasis>
. Fore leg with tibia 0.96-1.0 times as long as first tarsomere. Mid-tibia with 20-21 anterior, 3-4 posterodorsal, 7-9 posterior and (n = 1) 2 posteroventral bristles. Hind tibia with 5 anterodorsal, 4-5 posterodorsal and 4-5 posterior bristles.
<emphasis bold="true" pageId="0" pageNumber="67134">Wings</emphasis>
. Vein r-m 2.46-2.6 times longer than stem of M-fork.
<emphasis bold="true" pageId="0" pageNumber="67134">Abdomen.</emphasis>
Tergites covered with pale brown setae.
<emphasis bold="true" pageId="0" pageNumber="67134">Terminalia</emphasis>
(Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322758" pageId="0" pageNumber="67134">a</figureCitation>
,
<figureCitation captionStartId="F7322759" pageId="0" pageNumber="67134">b</figureCitation>
,
<figureCitation captionStartId="F7322760" pageId="0" pageNumber="67134">c</figureCitation>
). Each part of divided tergite IX with about 4-5 setae, apical seta stout. Gonocoxites evenly covered with setae, except on basoventral third (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322758" pageId="0" pageNumber="67134">a</figureCitation>
,
<figureCitation captionStartId="F7322759" pageId="0" pageNumber="67134">b</figureCitation>
). GL with length 0.55-0.61 of gonocoxite width, apico-internal margin slightly angled exteriorly, basal third or fourth covered with setae, apex with 3-4 setae (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322758" pageId="0" pageNumber="67134">a</figureCitation>
,
<figureCitation captionStartId="F7322759" pageId="0" pageNumber="67134">b</figureCitation>
). Aedaegal guides short with acute apex (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322758" pageId="0" pageNumber="67134">a</figureCitation>
,
<figureCitation captionStartId="F7322759" pageId="0" pageNumber="67134">b</figureCitation>
). Hypandrium covered with 16-19 setae, apical pair reaching about half of the GL (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322758" pageId="0" pageNumber="67134">a</figureCitation>
,
<figureCitation captionStartId="F7322759" pageId="0" pageNumber="67134">b</figureCitation>
). Hypandrial lobe with each branch slender, evenly tapering. Gonostylus (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322760" pageId="0" pageNumber="67134">c</figureCitation>
) with DB 1.37-1.42 times longer than broad, short and round; apical lobe well defined, short and broad, 0.18-0.2 times as long as the total DB length, apex rounded; evenly covered with setae on dorsal side, except on most apical part; external margin evenly rounded, with row of 4-5 elongate setae. VB round, with 2 small setae. IB apically with 1 seta close to apex and pair of setae one-third from apex. MB with apex acute, 1 seta close to base.
</paragraph>
<paragraph pageId="0" pageNumber="67134">
Female (n = 1): Body length 3.1 mm. Wing length 2.7 mm.
<emphasis bold="true" pageId="0" pageNumber="67134">Colouration</emphasis>
(Dry specimen). Head, face and clypeus dark brown; labellum pale brown; palpus yellow. Antenna with scape and pedicel yellow; flagellum pale brown. Scutum, lateral sclerites and propleura brown; halteres whitish-yellow. Legs whitish-yellow. Abdomen brown, tergites II-VI with paler lateral areas, not extending notably dorsally. Terminalia yellow.
<emphasis bold="true" pageId="0" pageNumber="67134">Head</emphasis>
. Frons and vertex and clypeus covered with pale setae. Antenna short, 1.4 times as long as length from vertex to ventral margin of clypeus; flagellomeres broader than long, with sixth flagellomere 0.7 as long as wide.
<emphasis bold="true" pageId="0" pageNumber="67134">Thorax</emphasis>
. Scutum covered with pale brown setae.
<emphasis bold="true" pageId="0" pageNumber="67134">Legs</emphasis>
. Fore leg with tibia as long as first tarsomere. Hind tibia with 6 anterodorsal and 6 posterodorsal bristles.
<emphasis bold="true" pageId="0" pageNumber="67134">Wings</emphasis>
. Vein r-m 2.8 times longer than stem of M-fork.
<emphasis bold="true" pageId="0" pageNumber="67134">Abdomen.</emphasis>
Tergites covered with pale brown setae.
<emphasis bold="true" pageId="0" pageNumber="67134">Terminalia</emphasis>
(Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322761" pageId="0" pageNumber="67134">d</figureCitation>
,
<figureCitation captionStartId="F7322762" pageId="0" pageNumber="67134">e</figureCitation>
). Cerci with apical segment 0.7 as long as basal segment. Tergite VIII with apicolateral margin virtually straight (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322761" pageId="0" pageNumber="67134">d</figureCitation>
). Sternite VII with apicoventral margin acuminate. Sternite VIII (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322762" pageId="0" pageNumber="67134">e</figureCitation>
) with hypogynal valves separated by wide v-shaped cleft with depth about one-sixth of sternite VIII and hypogynium length; apical seta about 0.54 times as long as sternite VIII length. Gonapophysis IX with basolateral part expanding in relatively obtuse angle; spermathecal eminence in ventral view appears cross-shaped, but with lateral branches slightly curved distally; gonapophysis IX apically with about 4 small setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67134" type="diagnosis">
<paragraph pageId="0" pageNumber="67134">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="67134">
Distinguished from
<taxonomicName class="Insecta" family="Araceae" kingdom="Animalia" lsidName="E. brevilobata" order="Diptera" pageId="0" pageNumber="67134" phylum="Arthropoda" rank="species" species="brevilobata">
<emphasis italics="true" pageId="0" pageNumber="67134">E. brevilobata</emphasis>
</taxonomicName>
in having the gonostylus with the dorsal branch only 1.37-1.42 times longer than broad, with its apical lobe broader and shorter, 0.18-0.2 of the total dorsal branch length (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322760" pageId="0" pageNumber="67134">c</figureCitation>
); from
<taxonomicName class="Insecta" family="Araceae" kingdom="Animalia" lsidName="E. sphaerata" order="Diptera" pageId="0" pageNumber="67134" phylum="Arthropoda" rank="species" species="sphaerata">
<emphasis italics="true" pageId="0" pageNumber="67134">E. sphaerata</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" family="Araceae" kingdom="Animalia" lsidName="E. repandoides" order="Diptera" pageId="0" pageNumber="67134" phylum="Arthropoda" rank="species" species="repandoides">
<emphasis italics="true" pageId="0" pageNumber="67134">E. repandoides</emphasis>
</taxonomicName>
in having shorter antennae, only 1.4-1.5 times as long as length from vertex to ventral margin of clypeus, in combination with the gonostylus with the apical lobe of the dorsal branch well defined (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322760" pageId="0" pageNumber="67134">c</figureCitation>
); from other species in the
<taxonomicName class="Insecta" family="Araceae" kingdom="Animalia" lsidName="E. parva" order="Diptera" pageId="0" pageNumber="67134" phylum="Arthropoda" rank="species" species="parva">
<emphasis italics="true" pageId="0" pageNumber="67134">E. parva</emphasis>
</taxonomicName>
group in having shorter antennae, in combination with the apico-internal margin of the gonocoxal lobe slightly angled exteriorly (Fig.
<figureCitation captionStartId="F6406432" pageId="0" pageNumber="67134">12</figureCitation>
<figureCitation captionStartId="F7322758" pageId="0" pageNumber="67134">a</figureCitation>
,
<figureCitation captionStartId="F7322759" pageId="0" pageNumber="67134">b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67134" type="etymology">
<paragraph pageId="0" pageNumber="67134">Etymology</paragraph>
<paragraph pageId="0" pageNumber="67134">
From Latin
<emphasis italics="true" pageId="0" pageNumber="67134">brevis</emphasis>
, short and
<emphasis italics="true" pageId="0" pageNumber="67134">flagellum</emphasis>
, whip, relating to the short antennae of the species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67134" type="distribution">
<paragraph pageId="0" pageNumber="67134">Distribution</paragraph>
<paragraph pageId="0" pageNumber="67134">
Nearctic, Canada (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F6415657" captionText="Figure 13. Distribution map of species in the E. parva group occurring in the Nearctic Region, based on localities from studied material, type material and DNA-barcoded material. Different species are represented by individual colours. Altitudes are indicated in shades from grey (lowland) to white (mountains)." figureDoi="10.3897/BDJ.9.e67134.figure13" httpUri="https://binary.pensoft.net/fig/483998" pageId="0" pageNumber="67134">13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67134" type="biology">
<paragraph pageId="0" pageNumber="67134">Biology</paragraph>
<paragraph pageId="0" pageNumber="67134">Adults collected in wetland habitats.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="227E7FDC445641C421947F6ADC709A4A">Occurrence of the millipede genus Piccola Attems, 1953 in China (Diplopoda: Polydesmida: Paradoxosomatidae)</mods:title>
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<paragraph id="B13536A4FFCDFFF5D8886C28FE64FD8E" blockId="1.[151,468,570,630]" box="[151,468,570,596]" pageId="1" pageNumber="404">
<heading id="EA7D81C8FFCDFFF5D8886C28FE64FD8E" bold="true" box="[151,468,570,596]" fontSize="11" level="1" pageId="1" pageNumber="404" reason="1">
<emphasis id="83FEEAB6FFCDFFF5D8886C28FE64FD8E" bold="true" box="[151,468,570,596]" pageId="1" pageNumber="404">
<taxonomicName id="768A4D27FFCDFFF5D8886C28FEC5FD8E" ID-CoL="4HPYR" box="[151,373,570,596]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFCDFFF5D8886C28FEC5FD8E" bold="true" box="[151,373,570,596]" italics="true" pageId="1" pageNumber="404">Piccola golovatchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98CD57CDFFCDFFF5D9636C28FE64FD8E" box="[380,468,570,596]" pageId="1" pageNumber="404" rank="species">sp. nov</taxonomicNameLabel>
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</heading>
</paragraph>
<paragraph id="B13536A4FFCDFFF5D8886C4FFF49FDAC" blockId="1.[151,468,570,630]" box="[151,249,605,630]" pageId="1" pageNumber="404">
<figureCitation id="29B12A21FFCDFFF5D8886C4FFF49FDAC" box="[151,249,605,630]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2. P" captionStart-2="FIGURE 3. P" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1996,2018]" captionStartId-1="2.[151,250,1741,1763]" captionStartId-2="3.[151,249,1907,1929]" captionStartId-3="4.[151,250,1742,1764]" captionTargetBox-0="[224,1362,934,1974]" captionTargetBox-1="[291,1295,193,1719]" captionTargetBox-2="[155,1426,1278,1885]" captionTargetBox-3="[151,1436,193,1720]" captionTargetId-0="figure@1.[224,1363,934,1975]" captionTargetId-1="figure@2.[291,1296,193,1719]" captionTargetId-2="figure@3.[150,1436,1273,1886]" captionTargetId-3="figure@4.[151,1436,193,1721]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Environmental photographs of P. golovatchi sp. nov., A male. B – C females." captionText-1="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." captionText-2="FIGURE 3. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A &amp; D right gonopod, lateral and mesal views, respectively. B &amp; C telopodite of right gonopod, lateral and mesal views, respectively. Designations: sph solenophore, sl solenomere. Scale bar: A &amp; D, 1.0 mm; B &amp; C, 2.0 mm." captionText-3="FIGURE 4. Distribution of the genus Piccola. 1. P. golovatchi sp. nov. 2. P. banana 3. P. corrugata; 4. P. debilis; 5. P. minuscula; 6. P. odontopyga; 7. P. spadix." httpUri-0="https://zenodo.org/record/236843/files/figure.png" httpUri-1="https://zenodo.org/record/236844/files/figure.png" httpUri-2="https://zenodo.org/record/236845/files/figure.png" httpUri-3="https://zenodo.org/record/236846/files/figure.png" pageId="1" pageNumber="404">Figs 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F990652FFFCDFFF5D8886CB7FE00FC94" pageId="1" pageNumber="404" type="materials_examined">
<paragraph id="B13536A4FFCDFFF5D8886CB7FE00FC94" blockId="1.[151,1437,677,846]" pageId="1" pageNumber="404">
<emphasis id="83FEEAB6FFCDFFF5D8886CB7FE31FD64" bold="true" box="[151,385,677,702]" pageId="1" pageNumber="404">Material examined.</emphasis>
<typeStatus id="6E318806FFCDFFF5D9966CB7FE41FD64" box="[393,497,677,702]" pageId="1" pageNumber="404" type="holotype">Holotype</typeStatus>
male (
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, Guangxi, Baise City, Tianlin County, Langping Town, Cave Shizikou Dadong, 24°36’49.79’N’,
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,
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,
<date id="C5341064FFCDFFF5DB4A6CD8FC78FD38" box="[853,968,714,738]" pageId="1" pageNumber="404" value="2014-01-19">2014-I-19</date>
, leg. Tian Mingyi.
<typeStatus id="6E318806FFCDFFF5DCB86CD8FAA8FD38" box="[1191,1304,714,738]" pageId="1" pageNumber="404" type="paratype">Paratypes</typeStatus>
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(
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), same locality and collecting data as the
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.
<specimenCount id="A78CFD2DFFCDFFF5DBE06CFCFBE0FCDC" box="[1023,1104,749,774]" pageId="1" pageNumber="404" type="male">1 male</specimenCount>
,
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(
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<specimenCount id="A78CFD2DFFCDFFF5D9DA6D03FD83FCF0" box="[453,563,785,810]" pageId="1" pageNumber="404" type="female">5 females</specimenCount>
,
<specimenCount id="A78CFD2DFFCDFFF5DA216D00FDCCFCF0" box="[574,636,785,810]" pageId="1" pageNumber="404" type="generic">3 juv.</specimenCount>
(
<collectionCode id="D79BAE61FFCDFFF5DA956D00FD6AFCF0" box="[650,730,786,810]" pageId="1" pageNumber="404">SCAU</collectionCode>
), same locality,
<date id="C5341064FFCDFFF5DB926D00FBA4FCF0" box="[909,1044,786,810]" pageId="1" pageNumber="404" value="2014-06-13">2014-VI-13</date>
, leg. Tian Mingyi, Liu Weixin, Yin Haomin &amp; Luo Xiaozhu.
</paragraph>
</subSubSection>
<caption id="E5F5662CFFCDFFF5D88869DEFBAEF838" box="[151,1054,1996,2018]" httpUri="https://zenodo.org/record/236843/files/figure.png" pageId="1" pageNumber="404" targetBox="[224,1362,934,1974]" targetPageId="1">
<paragraph id="B13536A4FFCDFFF5D88869DEFBAEF838" blockId="1.[151,1054,1996,2018]" box="[151,1054,1996,2018]" pageId="1" pageNumber="404">
<emphasis id="83FEEAB6FFCDFFF5D88869DEFEA3F83B" bold="true" box="[151,275,1996,2018]" pageId="1" pageNumber="404">FIGURE 1.</emphasis>
Environmental photographs of
<taxonomicName id="768A4D27FFCDFFF5DA4B69DFFD68F838" box="[596,728,1996,2018]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFCDFFF5DA4B69DFFD68F838" box="[596,728,1996,2018]" italics="true" pageId="1" pageNumber="404">P. golovatchi</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFCDFFF5DAC169DFFC9FF838" bold="true" box="[734,815,1997,2018]" pageId="1" pageNumber="404">
<taxonomicNameLabel id="98CD57CDFFCDFFF5DAC169DFFC9FF838" box="[734,815,1997,2018]" pageId="1" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="83FEEAB6FFCDFFF5DB2369DEFCFEF83B" bold="true" box="[828,846,1996,2017]" pageId="1" pageNumber="404">A</emphasis>
male.
<emphasis id="83FEEAB6FFCDFFF5DB8C69DEFC72F838" bold="true" box="[915,962,1996,2018]" pageId="1" pageNumber="404">B–C</emphasis>
females.
</paragraph>
</caption>
<caption id="E5F5662CFFCEFFF6D88868DFFA83F8FA" httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="2" pageNumber="405" targetBox="[291,1295,193,1719]" targetPageId="2">
<paragraph id="B13536A4FFCEFFF6D88868DFFA83F8FA" blockId="2.[151,1437,1741,1824]" pageId="2" pageNumber="405">
<emphasis id="83FEEAB6FFCEFFF6D88868DFFEA5F938" bold="true" box="[151,277,1741,1763]" pageId="2" pageNumber="405">FIGURE 2.</emphasis>
<taxonomicName id="768A4D27FFCEFFF6D90268DCFE14F939" box="[285,420,1741,1763]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFCEFFF6D90268DCFE14F939" box="[285,420,1741,1763]" italics="true" pageId="2" pageNumber="405">P. golovatchi</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFCEFFF6D9B368DFFE4FF938" bold="true" box="[428,511,1741,1762]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="98CD57CDFFCEFFF6D9B368DFFE4FF938" box="[428,511,1741,1762]" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ paratype from cave Shizikou Dadong.
<emphasis id="83FEEAB6FFCEFFF6DBA368DFFC5DF939" bold="true" box="[956,1005,1741,1763]" pageId="2" pageNumber="405">A–C</emphasis>
anterior part of body, ventral, lateral and dorsal views, respectively.
<emphasis id="83FEEAB6FFCEFFF6D9B068FEFE72F8DB" bold="true" box="[431,450,1772,1793]" pageId="2" pageNumber="405">D</emphasis>
gonopods
<emphasis id="83FEEAB6FFCEFFF6DA2A68FFFDC6F8D8" box="[565,630,1773,1794]" italics="true" pageId="2" pageNumber="405">in situ</emphasis>
, ventral view.
<emphasis id="83FEEAB6FFCEFFF6DB0D68FEFC8EF8DB" bold="true" box="[786,830,1772,1794]" pageId="2" pageNumber="405">E–F</emphasis>
midbody segments, lateral and dorsal views, respectively.
<emphasis id="83FEEAB6FFCEFFF6D8886918FF71F8C5" bold="true" box="[151,193,1802,1824]" pageId="2" pageNumber="405">G–I</emphasis>
posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A–C, E–I, 1.0 mm; D, 0.5 mm.
</paragraph>
</caption>
<subSubSection id="F990652FFFCEFFF6D8D86955FB26F85E" pageId="2" pageNumber="405" type="etymology">
<paragraph id="B13536A4FFCEFFF6D8D86955FB26F85E" blockId="2.[151,1436,1863,2032]" pageId="2" pageNumber="405">
<emphasis id="83FEEAB6FFCEFFF6D8D86955FEFEF8BA" bold="true" box="[199,334,1863,1888]" pageId="2" pageNumber="405">Etymology.</emphasis>
In honor of Prof. Dr. Sergei I. Golovatch, Institute for Problems of Ecology and Evolution, Russian Academy of Science, Moscow,
<collectingCountry id="C99D7634FFCEFFF6DA466979FD16F85E" box="[601,678,1899,1924]" name="Russia" pageId="2" pageNumber="405">Russia</collectingCountry>
, a prominent expert on millipede taxonomy.
</paragraph>
</subSubSection>
<subSubSection id="F990652FFFCEFFF6D8D8699DFD9BF811" pageId="2" pageNumber="405" type="diagnosis">
<paragraph id="B13536A4FFCEFFF6D8D8699DFD9BF811" blockId="2.[151,1436,1863,2032]" pageId="2" pageNumber="405">
<emphasis id="83FEEAB6FFCEFFF6D8D8699DFEF1F872" bold="true" box="[199,321,1935,1960]" pageId="2" pageNumber="405">Diagnosis.</emphasis>
Differs from congeners by having sternal processes present between coxae
<quantity id="76729B41FFCEFFF6DC876982FB50F872" box="[1176,1248,1935,1960]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.778" metricValueMin="0.762" pageId="2" pageNumber="405" unit="in" value="5.0" valueMax="7.0" valueMin="3.0">3–7 in</quantity>
♂, as well as by certain details of gonopod structure.
</paragraph>
</subSubSection>
<subSubSection id="F990652FFFCEFFF7D8D869C5FD56FBA6" lastPageId="3" lastPageNumber="406" pageId="2" pageNumber="405" type="description">
<paragraph id="B13536A4FFCEFFF7D8D869C5FA22FE9A" blockId="2.[151,1436,1863,2032]" lastBlockId="3.[151,1437,151,1221]" lastPageId="3" lastPageNumber="406" pageId="2" pageNumber="405">
<emphasis id="83FEEAB6FFCEFFF6D8D869C5FEE9F82A" bold="true" box="[199,345,2007,2032]" pageId="2" pageNumber="405">Description.</emphasis>
Length
<emphasis id="83FEEAB6FFCEFFF6D9AA69CBFE60F82A" box="[437,464,2009,2032]" italics="true" pageId="2" pageNumber="405">ca</emphasis>
28.0–32.0 (♂) or 24.0–28.0 mm (♀), width of pro- and metazonae 1.5–1.8 and 2.0–2.2 (♂) or 1.8–2.2 and
<quantity id="76729B41FFCFFFF7D96E6E8AFE49FF75" box="[369,505,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="3" pageNumber="406" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2–2.8 mm</quantity>
(♀), respectively.
<typeStatus id="6E318806FFCFFFF7DAD46E85FC83FF6A" box="[715,819,151,176]" pageId="3" pageNumber="406" type="holotype">Holotype</typeStatus>
30.0 mm long, 1.8 and
<quantity id="76729B41FFCFFFF7DC5D6E8AFB29FF75" box="[1090,1177,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="406" unit="mm" value="2.2">2.2 mm</quantity>
wide on midbody pro- and metazonae, respectively. In width, head &lt;collum&gt; segment 2&gt; 3&gt; 4 &lt;5 = 18&gt; 19&gt; telson. Coloration of living animals (
<figureCitation id="29B12A21FFCFFFF7D9506ECDFE25FF22" box="[335,405,223,248]" captionStart="FIGURE 1" captionStartId="1.[151,250,1996,2018]" captionTargetBox="[224,1362,934,1974]" captionTargetId="figure@1.[224,1363,934,1975]" captionTargetPageId="1" captionText="FIGURE 1. Environmental photographs of P. golovatchi sp. nov., A male. B – C females." httpUri="https://zenodo.org/record/236843/files/figure.png" pageId="3" pageNumber="406">Figs 1</figureCitation>
A–C) rather uniformly yellowish to pallid. Antennomeres 5–7 brownish. Head sparsely setose, epicranial suture distinct (
<figureCitation id="29B12A21FFCFFFF7DA106F16FDE3FEC6" box="[527,595,260,285]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Fig. 2</figureCitation>
A). Antennae rather long and slender, reaching back to segment 5 (♂) or 4 (♀) when stretched dorsally, antennomeres 5 and 6 each with a compact apicodorsal group of bacilliform sensilla.
</paragraph>
<paragraph id="B13536A4FFCFFFF7D8D86F5FFD2EFCE2" blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="406">
Tegument very strongly shining, prozonae faintly microalveolate; surface of metaterga finely rugulose (
<figureCitation id="29B12A21FFCFFFF7DD706F5EFF16FE52" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Fig. 2</figureCitation>
C). Collum with 3+3 setigerous tubercles at front margin; paraterga small, but evident, rounded (
<figureCitation id="29B12A21FFCFFFF7DD126F7DFAE7FE52" box="[1293,1367,367,392]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
A–C). Metaterga 2–19 each with 1+1 setigerous tubercles in anterior row, occasionally metaterga 18 and 19 with 2+2 setigerous tubercles. Paraterga 2 strongly developed, directed downward (
<figureCitation id="29B12A21FFCFFFF7DBEB6FA5FB8FFE0A" box="[1012,1087,439,464]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
A–C). Paraterga 3 and 4 very narrow and small (
<figureCitation id="29B12A21FFCFFFF7D96B6FCEFE0FFE2E" box="[372,447,476,500]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
B–C); paraterga 5–18 developed more strongly in ♂ than in ♀, calluses very thin in poreless segments, slightly thicker and sinuate in dorsal view in caudal 1/3 (marking ozopore position) in porebearing ones (
<figureCitation id="29B12A21FFCFFFF7D9266C36FE32FDE6" box="[313,386,548,573]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
E–F); paraterga 19 nearly suppressed, but its ozopores clear (
<figureCitation id="29B12A21FFCFFFF7DC5C6C36FB3DFDE6" box="[1091,1165,548,573]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
H–I). Ozopores entirely lateral, lying inside an ovoid groove about 1/
<quantity id="76729B41FFCFFFF7DA826C5AFD7AFDBA" box="[669,714,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="406" unit="in" value="3.0">3 in</quantity>
front of caudal paratergal corner (
<figureCitation id="29B12A21FFCFFFF7DC486C55FB2FFDBA" box="[1111,1183,583,608]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
E–F, H–I). Transverse sulcus incomplete on metaterga 5–7, more evident and complete on metaterga 8–18, reaching bases of paraterga, evident and rather deep, faintly striate at bottom (
<figureCitation id="29B12A21FFCFFFF7DAD76C9DFCA0FD72" box="[712,784,655,680]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
C, F, I). Stricture between pro- and metazona very clearly ribbed. Epiproct with a pair of paramedian tubercles near midway, tip evidently emarginate, pre-apical unusually strongly developed. Hypoproct roundly subtrapeziform, caudal setae distinctly separated, borne on evident knobs with a rounded lobe in between (
<figureCitation id="29B12A21FFCFFFF7DA0A6CEEFDEFFCCE" box="[533,607,764,789]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
H–I). Pleurosternal carinae poorly developed, only visible on segment 2 both in ♂ and ♀ (
<figureCitation id="29B12A21FFCFFFF7D9436D0DFE10FCE2" box="[348,416,799,824]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Fig. 2</figureCitation>
B). Axial line missing.
</paragraph>
<paragraph id="B13536A4FFCFFFF7D8D86D56FC12FC12" blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="406">
Sterna sparsely setose, cross-impressions very deep. A large, subquadrate, setose tubercle between coxae
<quantity id="76729B41FFCFFFF7DD6E6D56FA2CFC87" box="[1393,1436,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="406" unit="in" value="3.0">3 in</quantity>
♂; an evident, setose, subtrapeziform process between coxae
<quantity id="76729B41FFCFFFF7DB7F6D7AFC3EFC5A" box="[864,910,872,896]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="406" unit="in" value="4.0">4 in</quantity>
♂; an ovoid setose tubercle between each of coxae
<quantity id="76729B41FFCFFFF7D8FF6D9EFE98FC7F" box="[224,296,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="3" pageNumber="406" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">5–7 in</quantity>
♂ (
<figureCitation id="29B12A21FFCFFFF7D94A6D9EFE29FC7E" box="[341,409,908,933]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Fig. 2</figureCitation>
A). Legs 1 short, following ones increasingly longer and more slenderer towards telson,
<emphasis id="83FEEAB6FFCFFFF7DD9E6D9FFA2BFC7E" box="[1409,1435,909,932]" italics="true" pageId="3" pageNumber="406">ca</emphasis>
2.5–3.0 (♂) or 2.0–2.5 (♀) times as long as body height (
<figureCitation id="29B12A21FFCFFFF7DB0A6DBDFCECFC12" box="[789,860,943,968]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1741,1763]" captionTargetBox="[291,1295,193,1719]" captionTargetId="figure@2.[291,1296,193,1719]" captionTargetPageId="2" captionText="FIGURE 2. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A – C anterior part of body, ventral, lateral and dorsal views, respectively. D gonopods in situ, ventral view. E – F midbody segments, lateral and dorsal views, respectively. G – I posterior part of body, ventral, lateral and dorsal views, respectively. Scale bars: A – C, E – I, 1.0 mm; D, 0.5 mm." httpUri="https://zenodo.org/record/236844/files/figure.png" pageId="3" pageNumber="406">Figs 2</figureCitation>
A–B).
</paragraph>
<paragraph id="B13536A4FFCFFFF7D8D86DC6FD56FBA6" blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="406">
Gonopods (
<figureCitation id="29B12A21FFCFFFF7D9516DC6FE28FC36" box="[334,408,980,1005]" captionStart="FIGURE 3. P" captionStartId="3.[151,249,1907,1929]" captionTargetBox="[155,1426,1278,1885]" captionTargetId="figure@3.[150,1436,1273,1886]" captionTargetPageId="3" captionText="FIGURE 3. P. golovatchi sp. nov., ♂ paratype from cave Shizikou Dadong. A &amp; D right gonopod, lateral and mesal views, respectively. B &amp; C telopodite of right gonopod, lateral and mesal views, respectively. Designations: sph solenophore, sl solenomere. Scale bar: A &amp; D, 1.0 mm; B &amp; C, 2.0 mm." httpUri="https://zenodo.org/record/236845/files/figure.png" pageId="3" pageNumber="406">Figs 3</figureCitation>
A–D) suberect. Coxite subcylindrical, poorly setose distodorsally, about 1/3 as long as telopodite. Prefemoral portion rather short, about 1/4 as long as acropodite, densely setose. Femorite long, slender, slightly curved dorsad. No obvious demarcation sulcus between femur and postfemur. Solenophore (
<emphasis id="83FEEAB6FFCFFFF7DD0B6A0EFA8EFBEF" bold="true" box="[1300,1342,1052,1077]" pageId="3" pageNumber="406">sph</emphasis>
) clearly coiled, divided into two distinct, irregularly shaped lobes apically. Seminal groove running entirely on mesal side of femorite before moving onto free solenomere (
<emphasis id="83FEEAB6FFCFFFF7DADC6A76FD66FBA7" bold="true" box="[707,726,1124,1149]" pageId="3" pageNumber="406">sl</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="F990652FFFCFFFF7D8D86A95FE13FB1F" pageId="3" pageNumber="406" type="discussion">
<paragraph id="B13536A4FFCFFFF7D8D86A95FE13FB1F" blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="406">
<emphasis id="83FEEAB6FFCFFFF7D8D86A95FE8BFB7A" bold="true" box="[199,315,1159,1184]" pageId="3" pageNumber="406">Remarks.</emphasis>
Bacause of its pallid tegument and remarkably enlongated antennae and legs,
<taxonomicName id="768A4D27FFCFFFF7DCB66A9BFA8BFB7A" box="[1193,1339,1160,1184]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFCFFFF7DCB66A9BFA8BFB7A" box="[1193,1339,1160,1184]" italics="true" pageId="3" pageNumber="406">P. golovatchi</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFCFFFF7DD5D6A95FA2CFB7A" bold="true" box="[1346,1436,1159,1184]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="98CD57CDFFCFFFF7DD5D6A95FA2CFB7A" box="[1346,1436,1159,1184]" pageId="3" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is probably a troglobite.
</paragraph>
</subSubSection>
<caption id="E5F5662CFFCFFFF7D8886961FD62F81C" httpUri="https://zenodo.org/record/236845/files/figure.png" pageId="3" pageNumber="406" targetBox="[155,1426,1278,1885]" targetPageId="3">
<paragraph id="B13536A4FFCFFFF7D8886961FD62F81C" blockId="3.[151,1436,1907,1990]" pageId="3" pageNumber="406">
<emphasis id="83FEEAB6FFCFFFF7D8886961FEA4F852" bold="true" box="[151,276,1907,1929]" pageId="3" pageNumber="406">FIGURE 3.</emphasis>
<taxonomicName id="768A4D27FFCFFFF7D9036966FE11F853" box="[284,417,1907,1929]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFCFFFF7D9036966FE11F853" box="[284,417,1907,1929]" italics="true" pageId="3" pageNumber="406">P. golovatchi</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFCFFFF7D9B66961FE4BF852" bold="true" box="[425,507,1907,1928]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="98CD57CDFFCFFFF7D9B66961FE4BF852" box="[425,507,1907,1928]" pageId="3" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ paratype from cave Shizikou Dadong.
<emphasis id="83FEEAB6FFCFFFF7DBAB6961FC4DF852" bold="true" box="[948,1021,1907,1929]" pageId="3" pageNumber="406">A &amp; D</emphasis>
right gonopod, lateral and mesal views, respectively.
<emphasis id="83FEEAB6FFCFFFF7D93F6980FEDCF872" bold="true" box="[288,364,1938,1960]" pageId="3" pageNumber="406">B &amp; C</emphasis>
telopodite of right gonopod, lateral and mesal views, respectively. Designations:
<emphasis id="83FEEAB6FFCFFFF7DCD06980FB45F87D" bold="true" box="[1231,1269,1938,1959]" pageId="3" pageNumber="406">sph</emphasis>
solenophore,
<emphasis id="83FEEAB6FFCFFFF7DD946980FA2CF87D" bold="true" box="[1419,1436,1938,1959]" pageId="3" pageNumber="406">sl</emphasis>
solenomere. Scale bar: A &amp; D, 1.0 mm; B &amp; C, 2.0 mm.
</paragraph>
</caption>
<caption id="E5F5662CFFC8FFF0D88868DCFD8DF8D9" httpUri="https://zenodo.org/record/236846/files/figure.png" pageId="4" pageNumber="407" targetBox="[151,1436,193,1720]" targetPageId="4">
<paragraph id="B13536A4FFC8FFF0D88868DCFD8DF8D9" blockId="4.[151,1434,1742,1795]" pageId="4" pageNumber="407">
<emphasis id="83FEEAB6FFC8FFF0D88868DCFEA6F939" bold="true" box="[151,278,1742,1764]" pageId="4" pageNumber="407">FIGURE 4.</emphasis>
Distribution of the genus
<taxonomicName id="768A4D27FFC8FFF0DA2D68DCFDCFF93E" box="[562,639,1742,1764]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="83FEEAB6FFC8FFF0DA2D68DCFDCFF93E" box="[562,639,1742,1764]" italics="true" pageId="4" pageNumber="407">Piccola</emphasis>
</taxonomicName>
.
<emphasis id="83FEEAB6FFC8FFF0DA8F68DCFD13F939" bold="true" box="[656,675,1742,1763]" pageId="4" pageNumber="407">1.</emphasis>
<taxonomicName id="768A4D27FFC8FFF0DAB268DDFC85F93E" box="[685,821,1742,1764]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFC8FFF0DAB268DDFC85F93E" box="[685,821,1742,1764]" italics="true" pageId="4" pageNumber="407">P. golovatchi</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFC8FFF0DB2068DDFC01F939" bold="true" box="[831,945,1742,1764]" pageId="4" pageNumber="407">
<taxonomicNameLabel id="98CD57CDFFC8FFF0DB2068DDFC23F93E" box="[831,915,1743,1764]" pageId="4" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
2.
</emphasis>
<taxonomicName id="768A4D27FFC8FFF0DBA468DDFB94F93E" box="[955,1060,1742,1764]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="banana">
<emphasis id="83FEEAB6FFC8FFF0DBA468DDFB94F93E" box="[955,1060,1742,1764]" italics="true" pageId="4" pageNumber="407">P. banana</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFC8FFF0DC3268DCFBF0F939" bold="true" box="[1069,1088,1742,1763]" pageId="4" pageNumber="407">3.</emphasis>
<taxonomicName id="768A4D27FFC8FFF0DC5468DDFB7CF93E" box="[1099,1228,1743,1764]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="corrugata">
<emphasis id="83FEEAB6FFC8FFF0DC5468DDFB7CF93E" box="[1099,1228,1743,1764]" italics="true" pageId="4" pageNumber="407">P. corrugata</emphasis>
</taxonomicName>
;
<emphasis id="83FEEAB6FFC8FFF0DCC168DCFB41F939" bold="true" box="[1246,1265,1742,1763]" pageId="4" pageNumber="407">4.</emphasis>
<taxonomicName id="768A4D27FFC8FFF0DCE468DDFAEBF93E" box="[1275,1371,1742,1764]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="83FEEAB6FFC8FFF0DCE468DDFAEBF93E" box="[1275,1371,1742,1764]" italics="true" pageId="4" pageNumber="407">P. debilis</emphasis>
</taxonomicName>
;
<emphasis id="83FEEAB6FFC8FFF0DD7368DDFACFF93E" bold="true" box="[1388,1407,1743,1764]" pageId="4" pageNumber="407">5.</emphasis>
<taxonomicName id="768A4D27FFC8FFF0DD9668DDFF4FF8D9" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="minuscula">
<emphasis id="83FEEAB6FFC8FFF0DD9668DDFF4FF8D9" italics="true" pageId="4" pageNumber="407">P. minuscula</emphasis>
</taxonomicName>
;
<emphasis id="83FEEAB6FFC8FFF0D91368FFFEAFF8D8" bold="true" box="[268,287,1773,1794]" pageId="4" pageNumber="407">6.</emphasis>
<taxonomicName id="768A4D27FFC8FFF0D93A68FCFE05F8D9" box="[293,437,1773,1795]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="odontopyga">
<emphasis id="83FEEAB6FFC8FFF0D93A68FCFE05F8D9" box="[293,437,1773,1795]" italics="true" pageId="4" pageNumber="407">P. odontopyga</emphasis>
</taxonomicName>
;
<emphasis id="83FEEAB6FFC8FFF0D9DC68FFFE66F8D8" bold="true" box="[451,470,1773,1794]" pageId="4" pageNumber="407">7.</emphasis>
<taxonomicName id="768A4D27FFC8FFF0D9C368FCFD87F8D9" box="[476,567,1773,1795]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="spadix">
<emphasis id="83FEEAB6FFC8FFF0D9C368FCFD87F8D9" box="[476,567,1773,1795]" italics="true" pageId="4" pageNumber="407">P. spadix</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="F990652FFFC8FFF1D8D8693DFBE0FF0E" lastPageId="5" lastPageNumber="408" pageId="4" pageNumber="407" type="distribution">
<paragraph id="B13536A4FFC8FFF0D8D8693DFBA8F892" blockId="4.[151,1436,1839,2008]" box="[199,1048,1839,1864]" pageId="4" pageNumber="407">
<emphasis id="83FEEAB6FFC8FFF0D8D8693DFED0F892" bold="true" box="[199,352,1839,1864]" pageId="4" pageNumber="407">Distribution.</emphasis>
<collectingCountry id="C99D7634FFC8FFF0D978693DFE1AF892" box="[359,426,1839,1864]" name="China" pageId="4" pageNumber="407">China</collectingCountry>
(Guangxi). Known only from the
<typeStatus id="6E318806FFC8FFF0DB346922FCEBF892" box="[811,859,1840,1864]" pageId="4" pageNumber="407">type</typeStatus>
locality (
<figureCitation id="29B12A21FFC8FFF0DBD9693DFBBBF892" box="[966,1035,1839,1864]" captionStart="FIGURE 4" captionStartId="4.[151,250,1742,1764]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@4.[151,1436,193,1721]" captionTargetPageId="4" captionText="FIGURE 4. Distribution of the genus Piccola. 1. P. golovatchi sp. nov. 2. P. banana 3. P. corrugata; 4. P. debilis; 5. P. minuscula; 6. P. odontopyga; 7. P. spadix." httpUri="https://zenodo.org/record/236846/files/figure.png" pageId="4" pageNumber="407">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="B13536A4FFC8FFF1D8D86941FBE0FF0E" blockId="4.[151,1436,1839,2008]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="408" pageId="4" pageNumber="407">
Shizikou Dadong is a large and rather long limestone cave (about 1600 meters long). It is a beautiful show cave, having some big halls, and numerous fantastic rock formations. Its underground river belongs to the Hongshui River drainage. Shizikou Dadong holds a rich fauna. Apart from
<taxonomicName id="768A4D27FFC8FFF0DC11698EFB55F86E" box="[1038,1253,1948,1972]" class="Diplopoda" family="Paradoxosomatidae" genus="Piccola" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="83FEEAB6FFC8FFF0DC11698EFB55F86E" box="[1038,1253,1948,1972]" italics="true" pageId="4" pageNumber="407">Piccola golovatchi</emphasis>
</taxonomicName>
<emphasis id="83FEEAB6FFC8FFF0DCEF6989FAFDF86E" bold="true" box="[1264,1357,1947,1972]" pageId="4" pageNumber="407">
<taxonomicNameLabel id="98CD57CDFFC8FFF0DCEF6989FAFDF86E" box="[1264,1357,1947,1972]" pageId="4" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, cavedwelling animals living in this cave are also
<emphasis id="83FEEAB6FFC8FFF0DA8869D2FC60F802" box="[663,976,1984,2008]" italics="true" pageId="4" pageNumber="407">Sinocylocheilus tianlinensis</emphasis>
<bibRefCitation id="D51B4B55FFC8FFF0DBC669ADFB4EF802" author="Zhou" box="[985,1278,1983,2008]" pageId="4" pageNumber="407" refString="Zhou, J., Zhang, C. G. &amp; He, A. Y. (2004) A new species of the genus Sinocyclocheilus from Guangxi, China (Cypriniformes, Cyprinidae). Acta Zootaxonomica Sinica, 29 (3), 591 - 594. [in Chinese]" type="journal article" year="2004">Zhou, Zhang &amp; He, 2004</bibRefCitation>
, a completely blind and highly modified cave fish,
<taxonomicName id="768A4D27FFC9FFF1DA276E8AFD46FF6A" box="[568,758,152,176]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="83FEEAB6FFC9FFF1DA276E8AFD46FF6A" box="[568,758,152,176]" italics="true" pageId="5" pageNumber="408">Giraffaphaenops</emphasis>
</taxonomicName>
sp., a extremely adapted blind ground beetle (
<bibRefCitation id="D51B4B55FFC9FFF1DD1E6E8AFA20FF6A" author="Deuve" box="[1281,1424,152,176]" pageId="5" pageNumber="408" refString="Deuve, T. (2002) Deux remarquables Trechinae anophtalmes des cavites souterraines du Guangxi nord-occidental, Chine (Coleoptera, Trechidae). Bullentin de la Societe entomologique de France, 107 (5), 515 - 523." type="journal article" year="2002">Deuve 2002</bibRefCitation>
), diplopods of the genera
<taxonomicName id="768A4D27FFC9FFF1D9BA6EAEFD90FF0E" box="[421,544,188,212]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="83FEEAB6FFC9FFF1D9BA6EAEFD90FF0E" box="[421,544,188,212]" italics="true" pageId="5" pageNumber="408">Glyphiulus</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="768A4D27FFC9FFF1DA626EAEFC82FF0E" box="[637,818,188,212]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="83FEEAB6FFC9FFF1DA626EAEFC82FF0E" box="[637,818,188,212]" italics="true" pageId="5" pageNumber="408">Eutrichodesmus</emphasis>
</taxonomicName>
sp., bats and spiders, etc.
</paragraph>
</subSubSection>
</treatment>
</document>