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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.9.e67087" ID-Pensoft-Pub="1314-2828-9-e67087" ID-Pensoft-UUID="5464A0159E485DC2AD49727CD812B8EE" ModsDocID="1314-2828-9-e67087" checkinTime="1621628173522" checkinUser="pensoft" docAuthor="Tang, Jiaxin, Liang, Wei, Shi, Haitao, Gao, Caixia, Li, Shuqiang &amp; Zheng, Guo" docDate="2021" docId="9E4911DCC9E45C2E9479CBEDAA697B53" docLanguage="en" docName="BiodivDatJour 9: e67087" docOrigin="Biodiversity Data Journal 9" docPubDate="2021-05-21" docSource="http://dx.doi.org/10.3897/BDJ.9.e67087" docTitle="Opopaea apicalis" docType="treatment" docVersion="2" id="5464A0159E485DC2AD49727CD812B8EE" lastPageNumber="67087" masterDocId="5464A0159E485DC2AD49727CD812B8EE" masterDocTitle="A checklist of spiders from Yongxing Island, South China Sea, with taxonomic notes on four species of goblin spiders" masterLastPageNumber="67087" masterPageNumber="67087" pageNumber="67087" updateTime="1621628173522" updateUser="pensoft">
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<mods:title>A checklist of spiders from Yongxing Island, South China Sea, with taxonomic notes on four species of goblin spiders</mods:title>
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<mods:namePart>Tang, Jiaxin</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang, China</mods:affiliation>
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<mods:namePart>Liang, Wei</mods:namePart>
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<mods:affiliation>Ministry of Education Key Laboratory for Ecology of Tropical Islands, Hainan Normal University, Haikou, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shi, Haitao</mods:namePart>
<mods:affiliation>Ministry of Education Key Laboratory for Ecology of Tropical Islands, Hainan Normal University, Haikou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gao, Caixia</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:namePart>Zheng, Guo</mods:namePart>
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<taxonomicName LSID="9E4911DC-C9E4-5C2E-9479-CBEDAA697B53" authority="(Simon, 1893)" baseAuthorityName="Simon" baseAuthorityYear="1893" class="Arachnida" family="Oonopidae" genus="Opopaea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Opopaea apicalis" order="Araneae" pageId="0" pageNumber="67087" phylum="Arthropoda" rank="species" species="apicalis">Opopaea apicalis (Simon, 1893)</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="67087">Type status:</emphasis>
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individualCount:
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; sex:
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family: Onopidae
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see
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Platnick and
<normalizedToken originalValue="Dupérré">Duperre</normalizedToken>
(2009)
</bibRefCitation>
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Key to Indo-Pacific
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<emphasis id="2494A015B554FFEC2084FEA0FE23FEDF" bold="true" box="[401,486,311,337]" italics="true" pageId="8" pageNumber="27">Galeus</emphasis>
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species
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(adapted from
<bibRefCitation id="727101F6B554FFEC2056FECDFDCAFEFA" author="Compagno" box="[323,527,346,372]" pageId="8" pageNumber="27" refString="Compagno, L. J. V. (1984) FAO Species Catalogue. Vol. 4. Sharks of the World. An annotated and illustrated catalogue of sharks species known to date. Part 2. Carcharhiniformes. FAO Fishery Synopsis (125) Vol. 4, Pt. 2; 251 - 655." type="journal article" year="1984">Compagno, 1984</bibRefCitation>
)
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<paragraph id="165F7C07B554FFEC2182FE3DFA50FE4A" blockId="8.[151,1437,426,1458]" box="[151,1429,426,452]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FE3DFA50FE4A" box="[151,1429,426,452]" pageId="8" pageNumber="27">1a- A crest of enlarged denticles on the preventral caudal margin....................................................................2</keyLead>
</paragraph>
<paragraph id="165F7C07B554FFEC2182FE45FA51FE62" blockId="8.[151,1437,426,1458]" box="[151,1428,466,492]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FE45FA51FE62" box="[151,1428,466,492]" pageId="8" pageNumber="27">1b- No crest of enlarged denticles on the preventral caudal margin..................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD1464A2B554FFEC2182FE6AFA50FDCC" pageId="8" pageNumber="27">
<paragraph id="165F7C07B554FFEC2182FE6AFA51FD99" blockId="8.[151,1437,426,1458]" box="[151,1428,506,535]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FE6AFA51FD99" box="[151,1428,506,535]" pageId="8" pageNumber="27">
2a- 3 broad dark saddles in front of 1st dorsal fin
<emphasis id="2494A015B554FFEC23C0FE69FB67FD99" box="[725,1186,510,535]" italics="true" pageId="8" pageNumber="27">
.........................................
<taxonomicName id="D1E00784B554FFEC2511FE69FB67FD99" ID-CoL="3F4XQ" box="[1028,1186,510,535]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="boardmani">G. boardmani</taxonomicName>
</emphasis>
(southern
<collectingCountry id="6EF73C97B554FFEC240AFE6AFA48FD99" box="[1311,1421,509,535]" name="Australia" pageId="8" pageNumber="27">Australia</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="165F7C07B554FFEC2182FDBFFA50FDCC" blockId="8.[151,1437,426,1458]" box="[151,1429,549,578]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FDBFFA50FDCC" box="[151,1429,549,578]" pageId="8" pageNumber="27">
2b- 10 to 16 narrow bands in saddles in front of 1st dorsal fin................
<taxonomicName id="D1E00784B554FFEC22CAFDBEFBF7FDCC" ID-CoL="4LVQ" box="[991,1074,553,578]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="2494A015B554FFEC22CAFDBEFBF7FDCC" box="[991,1074,553,578]" italics="true" pageId="8" pageNumber="27">Galeus</emphasis>
</taxonomicName>
sp. B (northeastern
<collectingCountry id="6EF73C97B554FFEC240AFDBFFA48FDCC" box="[1311,1421,552,578]" name="Australia" pageId="8" pageNumber="27">Australia</collectingCountry>
)
</keyLead>
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<keyStep id="AD1464A2B554FFEC2182FDC7FA50FD6C" pageId="8" pageNumber="27">
<paragraph id="165F7C07B554FFEC2182FDC7FA51FD1C" blockId="8.[151,1437,426,1458]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FDC7FA51FD1C" pageId="8" pageNumber="27">
3a- Labial furrows very short, confined to mouth corners; snout broadly rounded (pre-oral length much shorter than mouth width)
<emphasis id="2494A015B554FFEC2315FDEEFBB3FD1C" box="[512,1142,633,658]" italics="true" pageId="8" pageNumber="27">......................................................................................</emphasis>
<taxonomicName id="D1E00784B554FFEC2569FDEEFA51FD1C" ID-CoL="3F4YL" authority="(Philippines)" baseAuthorityName="Philippines" box="[1148,1428,632,658]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="2494A015B554FFEC2569FDEEFB30FD1C" box="[1148,1269,633,658]" italics="true" pageId="8" pageNumber="27">G. schultzi</emphasis>
(
<collectingCountry id="6EF73C97B554FFEC2411FDEFFA49FD1C" box="[1284,1420,632,658]" name="Philippines" pageId="8" pageNumber="27">Philippines</collectingCountry>
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<paragraph id="165F7C07B554FFEC2182FD37FA50FD6C" blockId="8.[151,1437,426,1458]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FD37FA50FD6C" pageId="8" pageNumber="27">3b- Labial furrows short but distinct, not confined to mouth corners; snout more pointed (pre-oral length about equal or longer than mouth width)....................................................................................................4</keyLead>
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<keyLead id="AD11DF32B554FFEC2182FD67FA51FC84" box="[151,1428,752,778]" pageId="8" pageNumber="27">
4a- Dorsal fins and sometimes caudal lobes black-tipped
<emphasis id="2494A015B554FFEC2221FD67FB48FC87" box="[820,1165,752,777]" italics="true" pageId="8" pageNumber="27">
...............................
<taxonomicName id="D1E00784B554FFEC2509FD67FB48FC87" ID-CoL="3F4YK" box="[1052,1165,752,777]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="sauteri">G. sauteri</taxonomicName>
</emphasis>
(
<collectingCountry id="6EF73C97B554FFEC2588FD67FAE6FC84" box="[1181,1315,752,778]" name="Philippines" pageId="8" pageNumber="27">Philippines</collectingCountry>
to
<collectingCountry id="6EF73C97B554FFEC245DFD67FA48FC84" box="[1352,1421,752,778]" name="Japan" pageId="8" pageNumber="27">Japan</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="165F7C07B554FFEC2182FC8FFA51FCBC" blockId="8.[151,1437,426,1458]" box="[151,1428,792,818]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FC8FFA51FCBC" box="[151,1428,792,818]" pageId="8" pageNumber="27">4b- Dorsal and caudal fins not black-tipped, usually white edged....................................................................5</keyLead>
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<paragraph id="165F7C07B554FFEC2182FCD7FA51FCD4" blockId="8.[151,1437,426,1458]" box="[151,1428,832,858]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FCD7FA51FCD4" box="[151,1428,832,858]" pageId="8" pageNumber="27">5a- Mouth lining dark, dusky to blackish..........................................................................................................6</keyLead>
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<paragraph id="165F7C07B554FFEC2182FCFFFA51FC0C" blockId="8.[151,1437,426,1458]" box="[151,1428,872,898]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FCFFFA51FC0C" box="[151,1428,872,898]" pageId="8" pageNumber="27">5b- Mouth lining pale, white or light grey.........................................................................................................7</keyLead>
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<keyStep id="AD1464A2B554FFEC2182FC07FA50FB4C" pageId="8" pageNumber="27">
<paragraph id="165F7C07B554FFEC2182FC07FA50FBAC" blockId="8.[151,1437,426,1458]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FC07FA50FBAC" pageId="8" pageNumber="27">
6a- Caudal fin relatively short, lower postventral caudal margin 12.414.5% TL; clasper short and thick (fall- ing well in front of anal origin in mature male, clasper outer length 5.27.3 % TL, its base width 3.04.9 times in outer length); origin of crest of enlarged denticles on upper caudal margin well in front of origin of caudal ventral lobe; precaudal vertebral centra 7478
<emphasis id="2494A015B554FFEC224CFB9FFBE6FBAF" box="[857,1059,1032,1057]" italics="true" pageId="8" pageNumber="27">
...........
<taxonomicName id="D1E00784B554FFEC22BFFB9FFBE6FBAF" ID-CoL="3F4Y2" box="[938,1059,1032,1057]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="gracilis">G. gracilis</taxonomicName>
</emphasis>
(northern
<collectingCountry id="6EF73C97B554FFEC258AFB9FFAC8FBAC" box="[1183,1293,1032,1058]" name="Australia" pageId="8" pageNumber="27">Australia</collectingCountry>
,
<collectingCountry id="6EF73C97B554FFEC240CFB9FFA48FBAC" box="[1305,1421,1032,1058]" name="Indonesia" pageId="8" pageNumber="27">Indonesia</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="165F7C07B554FFEC2182FBA7FA50FB4C" blockId="8.[151,1437,426,1458]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FBA7FA50FB4C" pageId="8" pageNumber="27">
6b- Caudal fin relatively long, lower postventral caudal margin 15.516.2% TL; clasper very long and slender (reaching anal origin in mature male, clasper outer length 9.011.2 % TL, its base width 7.210.0 times in outer length); origin of crest of enlarged denticles on upper caudal margin over level of origin of caudal ventral lobe; precaudal vertebral centra 8186
<emphasis id="2494A015B554FFEC23F6FB3FFB90FB4F" box="[739,1109,1192,1217]" italics="true" pageId="8" pageNumber="27">
.................................
<taxonomicName id="D1E00784B554FFEC22C9FB3FFB90FB4F" ID-CoL="3F4YJ" box="[988,1109,1192,1217]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="priapus">G. priapus</taxonomicName>
</emphasis>
(
<collectingCountry id="6EF73C97B554FFEC2570FB3FFADAFB4C" box="[1125,1311,1192,1218]" name="New Caledonia" pageId="8" pageNumber="27">New Caledonia</collectingCountry>
,
<collectingCountry id="6EF73C97B554FFEC243EFB3FFA48FB4C" box="[1323,1421,1192,1218]" name="Vanuatu" pageId="8" pageNumber="27">Vanuatu</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD1464A2B554FFEC2182FB47FA50FA9C" pageId="8" pageNumber="27">
<paragraph id="165F7C07B554FFEC2182FB47FA51FB64" blockId="8.[151,1437,426,1458]" box="[151,1428,1232,1258]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FB47FA51FB64" box="[151,1428,1232,1258]" pageId="8" pageNumber="27">
7a- Snout very long, pre-oral length over 7 % TL
<emphasis id="2494A015B554FFEC23C9FB47FB48FB67" box="[732,1165,1232,1257]" italics="true" pageId="8" pageNumber="27">...........................................................</emphasis>
<taxonomicName id="D1E00784B554FFEC2585FB47FA51FB64" ID-CoL="3F4Y6" authority="(Japan)" baseAuthorityName="Japan" box="[1168,1428,1232,1258]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="longirostris">
<emphasis id="2494A015B554FFEC2585FB47FAF2FB67" box="[1168,1335,1232,1257]" italics="true" pageId="8" pageNumber="27">G. longirostris</emphasis>
(
<collectingCountry id="6EF73C97B554FFEC2453FB47FA49FB64" box="[1350,1420,1232,1258]" name="Japan" pageId="8" pageNumber="27">Japan</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="165F7C07B554FFEC2182FB6FFA50FA9C" blockId="8.[151,1437,426,1458]" box="[151,1429,1272,1298]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FB6FFA50FA9C" box="[151,1429,1272,1298]" pageId="8" pageNumber="27">7b- Snout moderately elongated, pre-oral length less than 7 % TL...................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD1464A2B554FFEC2182FAB7FA51FA3C" pageId="8" pageNumber="27">
<paragraph id="165F7C07B554FFEC2182FAB7FA51FAEC" blockId="8.[151,1437,426,1458]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FAB7FA51FAEC" pageId="8" pageNumber="27">
8a- Distance from mouth to inner nostril much shorter than length of upper labial furrow; anal-fin base rela- tively short, 810% TL, shorter than pelvic-anal interspace
<emphasis id="2494A015B554FFEC2291FADFFB40FAEF" box="[900,1157,1352,1377]" italics="true" pageId="8" pageNumber="27">...................................</emphasis>
<taxonomicName id="D1E00784B554FFEC259EFADFFA51FAEC" ID-CoL="6K7J7" authority="(Japan)" baseAuthorityName="Japan" box="[1163,1428,1352,1378]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="nipponensis">
<emphasis id="2494A015B554FFEC259EFADFFAF3FAEF" box="[1163,1334,1352,1377]" italics="true" pageId="8" pageNumber="27">G. nipponensis</emphasis>
(
<collectingCountry id="6EF73C97B554FFEC2453FADFFA49FAEC" box="[1350,1420,1352,1378]" name="Japan" pageId="8" pageNumber="27">Japan</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="165F7C07B554FFEC2182FAE7FA51FA3C" blockId="8.[151,1437,426,1458]" pageId="8" pageNumber="27">
<keyLead id="AD11DF32B554FFEC2182FAE7FA51FA3C" pageId="8" pageNumber="27">
8b- Distance from mouth to inner nostril longer than length of upper labial furrow; anal-fin base long and low, about 12% TL, longer than pelvic-anal interspace
<emphasis id="2494A015B554FFEC221AFA0FFB74FA3F" box="[783,1201,1432,1457]" italics="true" pageId="8" pageNumber="27">
......................................
<taxonomicName id="D1E00784B554FFEC253DFA0FFB74FA3F" ID-CoL="3F4XZ" box="[1064,1201,1432,1457]" class="Elasmobranchii" family="Scyliorhinidae" genus="Galeus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="eastmani">G. eastmani</taxonomicName>
</emphasis>
(
<collectingCountry id="6EF73C97B554FFEC25D5FA0FFACDFA3C" box="[1216,1288,1432,1458]" name="China" pageId="8" pageNumber="27">China</collectingCountry>
Sea,
<collectingCountry id="6EF73C97B554FFEC245DFA0FFA48FA3C" box="[1352,1421,1432,1458]" name="Japan" pageId="8" pageNumber="27">Japan</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

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<mods:title id="F08093608CB4B43C4057764015CF51AC">The phylogeny of the species of the genus Agelaia Lepeletier, 1836, one of the basalmost groups of Epiponini, with notes on male genitalia (Hymenoptera: Vespidae; Polistinae)</mods:title>
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<heading id="4D17D319D964FFB2FE65FC389A61FBCD" box="[457,863,1005,1031]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="5">
<taxonomicName id="D1E01FF6D964FFB2FE65FC389A61FBCD" ID-CoL="5TQD5" authority="DeGeer, 1773" authorityName="DeGeer" authorityYear="1773" box="[457,863,1005,1031]" class="Insecta" family="Vespidae" genus="Agelaia" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fulvofasciata">
<emphasis id="2494B867D964FFB2FE65FC389B89FBCD" box="[457,695,1005,1031]" italics="true" pageId="21" pageNumber="22">Agelaia fulvofasciata</emphasis>
DeGeer, 1773
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<paragraph id="165F6475D964FFB2FFC0FBFD9B66FB88" blockId="21.[108,600,1064,1090]" box="[108,600,1064,1090]" pageId="21" pageNumber="22">
[
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<collectingCountry id="6EF724E5D964FFB2FFDFFBFD99DFFB88" box="[115,225,1064,1090]" name="Suriname" pageId="21" pageNumber="22">Surinam</collectingCountry>
]:
<collectingRegion id="D424AA97D964FFB2FF5FFBFD9840FB88" box="[243,382,1064,1090]" country="Suriname" name="Paramaribo" pageId="21" pageNumber="22">Paramaribo</collectingRegion>
,
<specimenCount id="00E6AFFCD964FFB2FE27FBFD98E4FB88" box="[395,474,1064,1090]" count="1" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
(
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)
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.
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DESCRIPTION: (1) paramere about 1.8× longer than wide; spine without hairs; apical angle truncate (
<figureCitation id="8EDB78F0D964FFB2FF72FB53980CFB6A" box="[222,306,1158,1184]" captionStart="FIGURE 2" captionStartId="8.[108,194,1673,1694]" captionTargetBox="[181,1159,1231,1579]" captionTargetPageId="8" captionText="FIGURE 2. Male genitalia, from top to bottom, of Agelaia centralis, Agelaia flavipennis, and Agelaia fulvofasciata. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777786" httpUri="https://zenodo.org/record/10777786/files/figure.png" pageId="21" pageNumber="22">fig. 2K</figureCitation>
); (2) aedeagus long, lobe broad on apical region only, row of teeth extending ventrolaterally, basal teeth stronger than those on apical region; ventral process little projected, pointed (
<figureCitation id="8EDB78F0D964FFB2FF78FB059877FB21" box="[212,329,1232,1259]" captionStart="FIGURE 2" captionStartId="8.[108,194,1673,1694]" captionTargetBox="[181,1159,1231,1579]" captionTargetPageId="8" captionText="FIGURE 2. Male genitalia, from top to bottom, of Agelaia centralis, Agelaia flavipennis, and Agelaia fulvofasciata. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777786" httpUri="https://zenodo.org/record/10777786/files/figure.png" pageId="21" pageNumber="22">fig. 2L, M</figureCitation>
); (3) cuspis long and acute apically, with short, evenly spaced hairs (
<figureCitation id="8EDB78F0D964FFB2FBF7FB059D91FB21" box="[1115,1199,1232,1259]" captionStart="FIGURE 2" captionStartId="8.[108,194,1673,1694]" captionTargetBox="[181,1159,1231,1579]" captionTargetPageId="8" captionText="FIGURE 2. Male genitalia, from top to bottom, of Agelaia centralis, Agelaia flavipennis, and Agelaia fulvofasciata. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777786" httpUri="https://zenodo.org/record/10777786/files/figure.png" pageId="21" pageNumber="22">fig. 2N</figureCitation>
); (4) digitus wide apically, rounded, with scattered hairs; mesal surface with slightly spaced punctures (
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).
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<mods:title id="ECD509BC8ECF36EF55D655124F59AA1C">The phylogeny of the species of the genus Agelaia Lepeletier, 1836, one of the basalmost groups of Epiponini, with notes on male genitalia (Hymenoptera: Vespidae; Polistinae)</mods:title>
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<emphasis id="2494B867D964FFB2FE76FDB29B8DFD4B" box="[474,691,615,641]" italics="true" pageId="21" pageNumber="22">Agelaia flavipennis</emphasis>
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DESCRIPTION: (1) paramere about 1.9× longer than wide; spine without hairs; apical angle truncate (
<figureCitation id="8EDB78F0D964FFB2FF71FCD59815FCD0" box="[221,299,768,794]" captionStart="FIGURE 2" captionStartId="8.[108,194,1673,1694]" captionTargetBox="[181,1159,1231,1579]" captionTargetPageId="8" captionText="FIGURE 2. Male genitalia, from top to bottom, of Agelaia centralis, Agelaia flavipennis, and Agelaia fulvofasciata. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777786" httpUri="https://zenodo.org/record/10777786/files/figure.png" pageId="21" pageNumber="22">fig. 2F</figureCitation>
); (2) aedeagus long, lobe broad on apical region only, slightly pointed at distal end; row of teeth extending ventrolaterally, basal teeth stronger than those on apical region; ventral process little projected, rounded (
<figureCitation id="8EDB78F0D964FFB2FDF4FC9F9BEDFCAF" box="[600,723,842,869]" captionStart="FIGURE 2" captionStartId="8.[108,194,1673,1694]" captionTargetBox="[181,1159,1231,1579]" captionTargetPageId="8" captionText="FIGURE 2. Male genitalia, from top to bottom, of Agelaia centralis, Agelaia flavipennis, and Agelaia fulvofasciata. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777786" httpUri="https://zenodo.org/record/10777786/files/figure.png" pageId="21" pageNumber="22">fig. 2G, H</figureCitation>
); (3) cuspis long and acute apically, with very short and well-spaced hairs (
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); (4) digitus acute apically, pointed, without hairs; mesal surface with evenly spaced punctures (
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).
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Haliday, 1836
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DESCRIPTION: (1) paramere about 1.7× longer than wide; spine with short, dense hairs all over, apical angle truncate (
<figureCitation id="8EDB78F0D964FFB2FE1FF9C49B39F9E1" box="[435,519,1553,1579]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="21" pageNumber="22">fig. 3A</figureCitation>
); (2) aedeagus long, lobe broad on apical region only, row of teeth extending dorsoventrally; the teeth have the same size, from top to bottom, however, those close to basal region are less spaced than those of apical region; ventral process little projected, pointed (
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); (3) cuspis wider apically, pointed, with dense, short hairs (
<figureCitation id="8EDB78F0D964FFB2FB3AF95499ACF90A" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="21" pageNumber="22">fig. 3D</figureCitation>
); (4) digitus wide apically, rounded, with dense hairs; mesal surface with evenly spaced punctures (
<figureCitation id="8EDB78F0D964FFB2FF5DF919987CF92C" box="[241,322,1740,1766]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="21" pageNumber="22">fig. 3E</figureCitation>
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Cameron, 1907
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DESCRIPTION: (1) paramere about 1.7× longer than wide, spine with hairs on margin only; apical angle truncate (
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); (2) aedeagus long; lobe broad on apical region only; row of teeth extending ventrolaterally, basal teeth stronger than those on apical region; ventral process little projected, curved and rounded (
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<heading id="4D17D319D965FFB3FDABFC7F9A1FFC0E" box="[519,801,938,964]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="5">
<taxonomicName id="D1E01FF6D965FFB3FDABFC7F9A1FFC0E" ID-CoL="65QDN" authority="Say, 1873" authorityName="Say" authorityYear="1873" box="[519,801,938,964]" class="Insecta" family="Vespidae" genus="Agelaia" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="areata">
<emphasis id="2494B867D965FFB3FDABFC7F9B93FC0E" box="[519,685,938,964]" italics="true" pageId="20" pageNumber="21">Agelaia areata</emphasis>
Say, 1873
</taxonomicName>
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</paragraph>
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<materialsCitation id="A6886E28D965FFB3FFC0FC3D9A36FBC8" box="[108,776,1000,1026]" collectionCode="AMNH" country="Panama" location="Changuinola Dist." pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1">
<collectingCountry id="6EF724E5D965FFB3FFC0FC3D99F3FBC8" box="[108,205,1000,1026]" name="Panama" pageId="20" pageNumber="21">Panama</collectingCountry>
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Boca Toro,
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<subSubSection id="5EFA37FED965FFB3FF30FBF698B7FAD7" pageId="20" pageNumber="21" type="description">
<paragraph id="165F6475D965FFB3FF30FBF698B7FAD7" blockId="20.[108,1212,1058,1309]" pageId="20" pageNumber="21">
DESCRIPTION: (1) paramere about 2.4× longer than wide; spine long with evanescent hair on apical portion; apical angle truncate (
<figureCitation id="8EDB78F0D965FFB3FDC1FB9D9BFEFBA8" box="[621,704,1096,1122]" captionStart="FIGURE 1" captionStartId="7.[108,194,1666,1687]" captionTargetBox="[232,1116,1238,1594]" captionTargetPageId="7" captionText="FIGURE 1. Male genitalia, from top to bottom, of Agelaia angulata, Agelaia areata, and Agelaia cajennensis. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777784" httpUri="https://zenodo.org/record/10777784/files/figure.png" pageId="20" pageNumber="21">fig. 1F</figureCitation>
); (2) aedeagus long; lobe broad apically, extending to medial region; row of teeth extending ventrolaterally, basal teeth stronger than those on apical region; ventral process little projected and rounded (
<figureCitation id="8EDB78F0D965FFB3FC1CFB479D10FB67" box="[944,1070,1170,1197]" captionStart="FIGURE 1" captionStartId="7.[108,194,1666,1687]" captionTargetBox="[232,1116,1238,1594]" captionTargetPageId="7" captionText="FIGURE 1. Male genitalia, from top to bottom, of Agelaia angulata, Agelaia areata, and Agelaia cajennensis. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777784" httpUri="https://zenodo.org/record/10777784/files/figure.png" pageId="20" pageNumber="21">fig. 1G, H</figureCitation>
); (3) cuspis long and wider apically, with very short, scattered hairs (
<figureCitation id="8EDB78F0D965FFB3FC8AFB6D9A4EFB18" box="[806,880,1208,1234]" captionStart="FIGURE 1" captionStartId="7.[108,194,1666,1687]" captionTargetBox="[232,1116,1238,1594]" captionTargetPageId="7" captionText="FIGURE 1. Male genitalia, from top to bottom, of Agelaia angulata, Agelaia areata, and Agelaia cajennensis. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777784" httpUri="https://zenodo.org/record/10777784/files/figure.png" pageId="20" pageNumber="21">fig. 1I</figureCitation>
); (4) digitus acute apically, pointed, with short hairs extending to medial region; mesal surface with punctures extending to basal region (
<figureCitation id="8EDB78F0D965FFB3FE83FAD79847FAD7" box="[303,377,1282,1309]" captionStart="FIGURE 1" captionStartId="7.[108,194,1666,1687]" captionTargetBox="[232,1116,1238,1594]" captionTargetPageId="7" captionText="FIGURE 1. Male genitalia, from top to bottom, of Agelaia angulata, Agelaia areata, and Agelaia cajennensis. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777784" httpUri="https://zenodo.org/record/10777784/files/figure.png" pageId="20" pageNumber="21">fig. 1J</figureCitation>
).
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<mods:title id="6F352150609008346FE360E665C9071C">The phylogeny of the species of the genus Agelaia Lepeletier, 1836, one of the basalmost groups of Epiponini, with notes on male genitalia (Hymenoptera: Vespidae; Polistinae)</mods:title>
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<taxonomicName id="D1E01FF6D965FFB3FE6BFA8F9A5EFABF" ID-CoL="5TQD4" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[455,864,1370,1397]" class="Insecta" family="Vespidae" genus="Agelaia" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cajennensis">
<emphasis id="2494B867D965FFB3FE6BFA8F9B99FABE" box="[455,679,1370,1396]" italics="true" pageId="20" pageNumber="21">Agelaia cajennensis</emphasis>
Fabricius, 1798
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<collectingCountry id="6EF724E5D965FFB3FFC0FA4D99DDFA78" box="[108,227,1432,1458]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
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,
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DESCRIPTION: (1) paramere ~2× longer than wide; spine without hairs; apical angle truncate (
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); (2) aedeagus long, lobe broad, becoming more constricted on distal end, extending to medial region; row of teeth extending ventrolaterally with stronger teeth on apical region than those on basal region; ventral process little projected ad pointed (
<figureCitation id="8EDB78F0D965FFB3FC3BF9939D09F9AA" box="[919,1079,1606,1632]" captionStart="FIGURE 1" captionStartId="7.[108,194,1666,1687]" captionTargetBox="[232,1116,1238,1594]" captionTargetPageId="7" captionText="FIGURE 1. Male genitalia, from top to bottom, of Agelaia angulata, Agelaia areata, and Agelaia cajennensis. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777784" httpUri="https://zenodo.org/record/10777784/files/figure.png" pageId="20" pageNumber="21">fig. 1L and M</figureCitation>
); (3) cuspis long and acute apically, with short, spaced hairs (
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View file

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DESCRIPTION: (1) paramere about 2.2× longer than wide; spine with short, scattered hairs on base; apical angle truncate (
<figureCitation id="8EDB78F0D965FFB3FE73FD6F9B0DFD1E" box="[479,563,698,724]" captionStart="FIGURE 1" captionStartId="7.[108,194,1666,1687]" captionTargetBox="[232,1116,1238,1594]" captionTargetPageId="7" captionText="FIGURE 1. Male genitalia, from top to bottom, of Agelaia angulata, Agelaia areata, and Agelaia cajennensis. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777784" httpUri="https://zenodo.org/record/10777784/files/figure.png" pageId="20" pageNumber="21">fig. 1A</figureCitation>
); (2) aedeagus long; lobe broad on apical region only; row of teeth extending ventrolaterally, basal teeth stronger than those on apical region; ventral process little projected and rounded (
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); (3) cuspis long and wider apically, with short spaced hairs (
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); (4) digitus wide apically, rounded, with evanescent hairs; mesal surface with evenly and spaced punctures (
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DESCRIPTION: (1) paramere about 1.6× longer than wide; spine without hairs, apical angle truncate (
<figureCitation id="8EDB78F0D966FFB0FF71FEAF980FFE5E" box="[221,305,378,404]" captionStart="FIGURE 4" captionStartId="10.[108,194,1660,1681]" captionTargetBox="[206,1115,1205,1579]" captionTargetPageId="10" captionText="FIGURE 4. Male genitalia, from top to bottom, of Agelaia panamaensis, Agelaia testacea, and Agelaia timida. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777793" httpUri="https://zenodo.org/record/10777793/files/figure.png" pageId="23" pageNumber="24">fig. 4K</figureCitation>
); (2) aedeagus short, wide lobe, broad apically; row of teeth extending dorsoventrally; the teeth are very long, stronger apically than basally; ventral process little projected, acute (
<figureCitation id="8EDB78F0D966FFB0FF16FE11980CFE15" box="[186,306,452,479]" captionStart="FIGURE 4" captionStartId="10.[108,194,1660,1681]" captionTargetBox="[206,1115,1205,1579]" captionTargetPageId="10" captionText="FIGURE 4. Male genitalia, from top to bottom, of Agelaia panamaensis, Agelaia testacea, and Agelaia timida. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777793" httpUri="https://zenodo.org/record/10777793/files/figure.png" pageId="23" pageNumber="24">fig. 4L, M</figureCitation>
); (3) cuspis long and acute apically, with very short, scattered hairs (
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); (4) digitus acute apically, pointed, with scattered short hairs; mesal surface with evenly spaced punctures, extending to basal region (
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DESCRIPTION: (1) paramere ~2× longer than wide; spine with short and closely spaced hairs on first half, apical angle truncate (
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); (2) aedeagus long, lobe broad on apical region only; row of teeth extending dorsoventrally; teeth have the same size, from top to bottom; ventral process little projected, rounded (
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[= Porto Alegre],
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DESCRIPTION: (1) paramere about 2.2× long than wide; spine without hairs, apical angle truncate (
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); (2) aedeagus very long and thin, lobe broad apically only; row of small teeth extending dorsoventrally; teeth have same size, from top to bottom; ventral process little projected, rounded (
<figureCitation id="8EDB78F0D966FFB0FE98FC9D9898FCA8" box="[308,422,840,867]" captionStart="FIGURE 5" captionStartId="11.[108,194,1248,1269]" captionTargetBox="[125,325,722,1206]" captionTargetId="graphics-2122@11.[145,325,722,1206]" captionTargetPageId="11" captionText="FIGURE 5. Male genitalia, from top to bottom, of Agelaia vicina and Agelaia yepocapa. A and F = paramere, lateral view; B and G = aedeagus, ventral view; C and H = aedeagus, lateral view; D and I = cuspis, lateral view; E and J = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777795" httpUri="https://zenodo.org/record/10777795/files/figure.png" pageId="23" pageNumber="24">fig. 5B, C</figureCitation>
); (3) cuspis wider apically, pointed, with evenly spaced, short hairs (
<figureCitation id="8EDB78F0D966FFB0FFD8FCB899F7FC42" box="[116,201,877,904]" captionStart="FIGURE 5" captionStartId="11.[108,194,1248,1269]" captionTargetBox="[125,325,722,1206]" captionTargetId="graphics-2122@11.[145,325,722,1206]" captionTargetPageId="11" captionText="FIGURE 5. Male genitalia, from top to bottom, of Agelaia vicina and Agelaia yepocapa. A and F = paramere, lateral view; B and G = aedeagus, ventral view; C and H = aedeagus, lateral view; D and I = cuspis, lateral view; E and J = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777795" httpUri="https://zenodo.org/record/10777795/files/figure.png" pageId="23" pageNumber="24">fig. 5D</figureCitation>
); (4) digitus pointed apically, rounded, with short, evenly spaced hairs; mesal surface with evenly spaced punctures (
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DESCRIPTION: (1) paramere about 1.7× longer than wide; spine without hairs, apical angle rounded (
<figureCitation id="8EDB78F0D967FFB1FF4DFB7E9808FB0F" box="[225,310,1195,1221]" captionStart="FIGURE 4" captionStartId="10.[108,194,1660,1681]" captionTargetBox="[206,1115,1205,1579]" captionTargetPageId="10" captionText="FIGURE 4. Male genitalia, from top to bottom, of Agelaia panamaensis, Agelaia testacea, and Agelaia timida. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777793" httpUri="https://zenodo.org/record/10777793/files/figure.png" pageId="22" pageNumber="23">fig. 4A</figureCitation>
); (2) aedeagus long, lobe broad apically only; row of teeth extending dorsoventrally; teeth have the same size, from top to bottom; ventral process little projected, rounded (
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); (3) cuspis wider apically, pointed, with spaced and short hairs (
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<emphasis id="2494B867D967FFB1FE47FDB29BA0FD4B" box="[491,670,615,641]" italics="true" pageId="22" pageNumber="23">Agelaia pallipes</emphasis>
Olivier, 1792
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<paragraph id="165F6475D967FFB1FF30FD0E9BCAFC1F" blockId="22.[108,1212,730,981]" pageId="22" pageNumber="23">
DESCRIPTION: (1) paramere about 1.9× longer than wide; spine with short and closely spaced hairs all over, apical angle truncate (
<figureCitation id="8EDB78F0D967FFB1FDD3FCD59BEBFCD0" box="[639,725,768,794]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3K</figureCitation>
); (2) aedeagus long, lobe broad apically, extending medial region; row of teeth extending dorsoventrally; the teeth have the same size, from top to bottom, however, those close to basal region are less spaced than those of apical region; ventral process little projected, rounded (
<figureCitation id="8EDB78F0D967FFB1FD04FCA59A20FC40" box="[680,798,880,906]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3L, M</figureCitation>
); (3) cuspis wider apically, pointed, with spaced and short hairs (
<figureCitation id="8EDB78F0D967FFB1FE6CFC409B2BFC65" box="[448,533,917,943]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3N</figureCitation>
); (4) digitus wider apically, rounded, without hairs; mesal surface with few and evenlyspaced punctures (
<figureCitation id="8EDB78F0D967FFB1FD3DFC6F9BD8FC1F" box="[657,742,954,981]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3O</figureCitation>
).
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<mods:title id="116A14ADCE40AA5E6C5707FA9DACA2D4">The phylogeny of the species of the genus Agelaia Lepeletier, 1836, one of the basalmost groups of Epiponini, with notes on male genitalia (Hymenoptera: Vespidae; Polistinae)</mods:title>
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<emphasis id="2494B867D967FFB1FE5DFF349BA2FF31" box="[497,668,225,251]" italics="true" pageId="22" pageNumber="23">Agelaia ornata</emphasis>
Ducke, 1905
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<materialsCitation id="A6886E28D967FFB1FFC0FEC99B9DFEFD" box="[108,675,284,311]" collectionCode="BMNH" collectorName="La Macarena" country="Colombia" location="Meta" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Meta">
<collectingCountry id="6EF724E5D967FFB1FFC0FEC999DDFEFC" box="[108,227,284,310]" name="Colombia" pageId="22" pageNumber="23">Colombia</collectingCountry>
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,
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,
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(
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<paragraph id="165F6475D967FFB1FF30FE809ACBFDE3" blockId="22.[108,1213,340,554]" pageId="22" pageNumber="23">
DESCRIPTION: (1) paramere about 1.8× long than wide; spine with short and space hairs on first half, apical angle rounded (
<figureCitation id="8EDB78F0D967FFB1FDAEFEAF9B6EFE5E" box="[514,592,378,404]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3F</figureCitation>
); (2) aedeagus long, lobe broad on apical region only, becoming pointed on distal end; row of teeth extending dorsoventrally; the teeth have the same size, from top to bottom; ventral process little projected, rounded (
<figureCitation id="8EDB78F0D967FFB1FCD9FE119AD2FE15" box="[885,1004,452,479]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3G, H</figureCitation>
); (3) cuspis wider apically, pointed, with slightly spaced and very short hairs (
<figureCitation id="8EDB78F0D967FFB1FC9AFE3F9ABEFDCE" box="[822,896,490,516]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3I</figureCitation>
); (4) digitus wide apically, rounded, without hairs; mesal surface with evenly spaced punctures (
<figureCitation id="8EDB78F0D967FFB1FC32FDDA9AD9FDE3" box="[926,999,527,553]" captionStart="FIGURE 3" captionStartId="9.[108,194,1673,1694]" captionTargetBox="[164,341,229,1605]" captionTargetId="graphics-2888@9.[202,334,715,1103]" captionTargetPageId="9" captionText="FIGURE 3. Male genitalia, from top to bottom, of Agelaia multipicta, Agelaia ornata, and Agelaia pallipes. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777790" httpUri="https://zenodo.org/record/10777790/files/figure.png" pageId="22" pageNumber="23">fig. 3J</figureCitation>
).
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<mods:title id="E151CAE9A9FA968FC3C869F632B5BAF7">The phylogeny of the species of the genus Agelaia Lepeletier, 1836, one of the basalmost groups of Epiponini, with notes on male genitalia (Hymenoptera: Vespidae; Polistinae)</mods:title>
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<taxonomicName id="D1E01FF6D967FFB1FE77FA4B9A72FA72" ID-CoL="5TQCW" authority="Fabricius, 1804" authorityName="Fabricius" authorityYear="1804" box="[475,844,1438,1464]" class="Insecta" family="Vespidae" genus="Agelaia" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="testacea">
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Fabricius, 1804
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DESCRIPTION: (1) paramere about 1.8× longer than wide; spine without hairs, apical angle truncate (
<figureCitation id="8EDB78F0D967FFB1FF73F9E3980EF99A" box="[223,304,1590,1616]" captionStart="FIGURE 4" captionStartId="10.[108,194,1660,1681]" captionTargetBox="[206,1115,1205,1579]" captionTargetPageId="10" captionText="FIGURE 4. Male genitalia, from top to bottom, of Agelaia panamaensis, Agelaia testacea, and Agelaia timida. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777793" httpUri="https://zenodo.org/record/10777793/files/figure.png" pageId="22" pageNumber="23">fig. 4F</figureCitation>
); (2) aedeagus long, lobe broad apically only; row of teeth extending dorsoventrally; teeth have the same size, from top to bottom; ventral process little projected, acute (
<figureCitation id="8EDB78F0D967FFB1FFD8F95499D2F951" box="[116,236,1665,1691]" captionStart="FIGURE 4" captionStartId="10.[108,194,1660,1681]" captionTargetBox="[206,1115,1205,1579]" captionTargetPageId="10" captionText="FIGURE 4. Male genitalia, from top to bottom, of Agelaia panamaensis, Agelaia testacea, and Agelaia timida. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777793" httpUri="https://zenodo.org/record/10777793/files/figure.png" pageId="22" pageNumber="23">fig. 4G, H</figureCitation>
); (3) cuspis long and acute apically, with short and evenly spaced hairs (
<figureCitation id="8EDB78F0D967FFB1FB97F9549DBBF951" box="[1083,1157,1665,1691]" captionStart="FIGURE 4" captionStartId="10.[108,194,1660,1681]" captionTargetBox="[206,1115,1205,1579]" captionTargetPageId="10" captionText="FIGURE 4. Male genitalia, from top to bottom, of Agelaia panamaensis, Agelaia testacea, and Agelaia timida. A, F, and K = paramere, lateral view; B, G, and L = aedeagus, ventral view; C, H, and M = aedeagus, lateral view; D, I, and N = cuspis, lateral view; E, J, and O = digitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10777793" httpUri="https://zenodo.org/record/10777793/files/figure.png" pageId="22" pageNumber="23">fig. 4I</figureCitation>
); (4) digitus wide apically, rounded, with short and evenly spaced hairs; mesal surface with evenly spaced punctures (
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).
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KEY TO THE SPECIES Of
<taxonomicName id="D1E01FF6D96CFFBAFD04FEE79A9CFE86" ID-CoL="629D7" ID-ENA="91383" authority="LEPELETIER" authorityName="Lepeletier" authorityYear="1836" box="[680,930,306,332]" class="Insecta" family="Vespidae" genus="Agelaia" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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LEPELETIER
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<paragraph id="165F6475D96CFFBAFF4BFEB29D7FFEB5" blockId="29.[231,1089,359,383]" box="[231,1089,359,383]" pageId="29" pageNumber="30">Modified from Cooper (2000, 2001) and Garcete-Barrett (personal commun.).</paragraph>
<paragraph id="165F6475D96CFFBAFFC0FE6E9A60FE19" blockId="29.[108,862,443,467]" box="[108,862,443,467]" pageId="29" pageNumber="30">This key does not identify subspecies, for which see Richards (1978).</paragraph>
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1. Dorsal pronotal carina obsolete or very weak (
<figureCitation id="8EDB78F0D96CFFBAFD3AFE3C9A3FFDCB" box="[662,769,489,513]" captionStart="FIGURE 7" captionStartId="34.[108,194,1566,1587]" captionTargetBox="[112,1208,224,1540]" captionTargetId="figure-11@34.[112,1208,224,1541]" captionTargetPageId="34" captionText="FIGURE 7. A. Dorsal pronotal carina obselete in Agelaia cajennensis; B. lateral view, pronotal carina obselete in Agelaia cajennensis; C. dorsal pronotal carina developed in Agelaia. flavipennis; D. lateral view pronotal carina developed in Agelaia. flavipennis; E. pterostigma about 1.52× as long as wide in Agelaia lobipleura; F. pterostigma about as long as wide in Agelaia angulata. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10777797" httpUri="https://zenodo.org/record/10777797/files/figure.png" pageId="29" pageNumber="30">fig. 7A, B</figureCitation>
); pterostigma about 1.52× as long as wide (
<figureCitation id="8EDB78F0D96CFFBAFF70FDDF9816FDE8" box="[220,296,522,546]" captionStart="FIGURE 7" captionStartId="34.[108,194,1566,1587]" captionTargetBox="[112,1208,224,1540]" captionTargetId="figure-11@34.[112,1208,224,1541]" captionTargetPageId="34" captionText="FIGURE 7. A. Dorsal pronotal carina obselete in Agelaia cajennensis; B. lateral view, pronotal carina obselete in Agelaia cajennensis; C. dorsal pronotal carina developed in Agelaia. flavipennis; D. lateral view pronotal carina developed in Agelaia. flavipennis; E. pterostigma about 1.52× as long as wide in Agelaia lobipleura; F. pterostigma about as long as wide in Agelaia angulata. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10777797" httpUri="https://zenodo.org/record/10777797/files/figure.png" pageId="29" pageNumber="30">fig. 7E</figureCitation>
). Malar space of female with slightly concave, hairless, reticulate-coriarious area (
<figureCitation id="8EDB78F0D96CFFBAFF0DFDFE99D1FD89" box="[161,239,555,579]" captionStart="FIGURE 8" captionStartId="35.[108,194,1567,1588]" captionTargetBox="[113,1207,224,1540]" captionTargetId="figure-7@35.[113,1207,224,1541]" captionTargetPageId="35" captionText="FIGURE 8. A. Malar space with slightly concave and reticulate-coriarious area in Agelaia cajennensis; B. malar space without a concave area, wholly dull due to fine sculpturation in Agelaia angulata; C. anterior margin of pronotum markedly sinuate below fovea in Agelaia lobipleura; D. malar space about as long as height of antennal socket in Agelaia lobipleura; E. dorsal pronotal carina, very weak; almost entirely yellow in Agelaia acreana; F. mesoscutum without distinct dark stripes in Agelaia acreana. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10777799" httpUri="https://zenodo.org/record/10777799/files/figure.png" pageId="29" pageNumber="30">fig. 8A</figureCitation>
). Male with aedeagus broader, pointed and not widened at apex and clypeus with inconspicuous pubescence. Terga without a posterior, yellowish band............................2
</keyStep>
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</paragraph>
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</treatment>
</document>

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12.
<taxonomicName id="EE6F815964290C9F4F09D15362091DCB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:132501" class="Insecta" family="Formicidae" genus="Oecodoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Oecodoma nigra Smith" order="Hymenoptera" pageId="186" pageNumber="186" phylum="Arthropoda" rank="species" species="nigra">Oecodoma nigra</taxonomicName>
. B.M.
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<paragraph id="74F74E4B735F50CE6131FA87DC9B4D39" pageId="186" pageNumber="186">Female. Length 6 lines. - Black: the mandibles ferruginous, longitudinally striated, and having two blunt teeth at their apex, their base, outer margins and apex black; two longitudinal carinae on the front, which run irregularly and obliquely to the lateral posterior angles of the head, each carina is elevated into a spine or tooth opposite the eyes; the head is longitudinally strigose, and has a number of short spines at the posterior angles.- Thorax: a short curved spine on each side in front, and two rather longer curved ones on the metathorax; the meso- ' thorax and scutellum longitudinally covered with a series of irregular carinae; the legs covered with short brown pubescence, the tarsi obscurely ferruginous. Abdomen globose, covered with irregular points and tubercles; the first node of the peduncle quadrate, and having a number of short points or tubercles; the second node transverse, and similarly tuberculate.</paragraph>
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<paragraph id="4546A6038A9E7D6C675BD57FD09EE478" pageId="186" pageNumber="186">Hab. Brazil; Rio (Tejuca). (Coll. Rev. Hamlet Clark.)</paragraph>
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