|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| Body length (mm) | +465 | +278 | +270 | +250–320 | +120 | +73 | +
| Body width (mm) | +13.3 | +10.1 | +9 | +10–11 | +6 | +3 | +
| Segment number | +176 | +160 | +126 | +126–148 | +114 | +nd | +
| Spermathecal pores | +6/7/8/9 | +6/7/8/9 | +6/7/8/9 | +6/7/8/9 | +6/7/8/9 | +6/7/8/9 | +
| First dorsal pore | +12/13 | +12/13 | +13/14 | +12/13 | +12/13 | +12/13 | +
| Setal number | +||||||
| vii | +68 | +53 | +40 | +nd | +nd | +nd | +
| xx | +101 | +81 | +49 | +65–84 | +nd | +nd | +
| Between male pores | +26 | +25 | +13 | +10–20 | +29 | +nd | +
| Genital markings | +||||||
| Preclitellum | +absent | +absent | +absent | +median in VIII | +absent | +absent | +
| Postclitellum | +absent | +crescent shape | +group of small circular papillae on XVIII | +absent | +paired at 17/18, 18/19 | +paired at XVIII | +
| Spermathecae | +large sac | +oval | +oval | +spherical | +flatten | +spherical | +
| Diverticulum | +slender | +small ovate | +long slender | +tubular | +tubular and coiled | +convoluted | +
| Prostate gland | +XVII–XXII | +XVI–XXI | +XV–XX | +XV–XXIV | +XVII–XX | +nd | +
| Intestinal caeca | +simple | +simple | +simple | +simple | +simple | +simple | +
| Type locality | +Thailand | +Thailand | +Sumatra, Indonesia | +South Andaman Islands | +Burma | +Thailand | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|---|---|
| Body length (mm) | +314 | +332 | +145 | +1 metre | +170 | +>86 | +137 | +75–135 | +
| Body width (mm) | +8.7 | +11.1 | +4–9 | +8 | +7 | +6 | +7 | +4.5–7 | +
| Segment number | +132 | +198 | +133 | +370 | +138 | +>120 | +90 | +103–123 | +
| Spermathecal pores | +5/6/7/8/9 | +5/6/7/8/9 | +5/6/7/8/9 | +5/6/7/8/9 | +5/6/7/8/9 | +5/6/7/8/9 | +5/6/7/8/9 | +5/6/7/8/9 | +
| First dorsal pore | +12/13 | +12/13 | +12/13 | +10/11 | +12/13 | +12/13 | +nd | +12/13 | +
| Setal number | +||||||||
| vii | +71 | +108 | +33–43 | +96–114 | +63–71 | +41 | +nd | +60-70 | +
| xx | +68 | +105 | +58–72 | +96–106 | +96 | +89–102 | +nd | +75 | +
| Between male pores | +19 | +23 | +13 | +10 | +29 | +31 | +3 | +2 | +
| Genital markings | +||||||||
| Preclitellum | +absent | +absent | +absent | +absent | +absent | +absent | +absent | +absent | +
| Postclitellum | +paired on 19/20–24/25 | +paired on XVII, XIX–XXI | +absent | +absent | +paired on 18/19–20/21 | +three trios on 18/19–20/21 | +cluster on XVIII | +absent | +
| Spermathecae | +large sac | +large sac | +oval | +sacciform | +spherical | +nd | +oval | +pear-shaped | +
| Diverticulum | +loosely looped | +zigzaged | +moniliform | +clavate | +saccular | +slender | +intimately looped | +bent | +
| Prostate gland | +XVII–XXI | +XVI–XIX | +XVII–XX | +XVIII | +XVIII | +XVIII | +XVII–XX | +XVII–XIX | +
| Intestinal caeca | +simple | +simple | +simple | +simple | +simple | +simple | +simple | +simple | +
| Type locality | +Thailand | +Thailand | +India | +Mekong river, Vietnam | +Burma | +Burma | +Indonesia | +Andaman Islands | +
|
+3.2 - Alluvionen und
+ |
+
|
+3.2.1.1 - Alluvionen mit krautiger Pioniervegetation (
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+203500 | +
| = | +
+ |
+
+ |
+2353 | +
| = | +
+ |
+
+ |
+2341 | +
| = | +
+ |
+
+ |
+2341 | +
| = | +
+ |
+
+ |
+203500 | +
| = | +
+ |
+
+ |
+3329 | +
| = | +
+ |
+
+ |
+2663 | +
| = | +
+ |
+
+ |
+203500 | +
| = | +
+ |
+
+ |
+1992 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(01.01.2018) | +
|
+ |
+-- | +
|
+ |
+L - Leitart | +weitere Informationen | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +0 - unbedeutend, keine Bindung. | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+391250 | +
| = | +
+ |
+
+ |
+391250 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +regional beziehungsweise in der Schweiz ausgestorben (Regionally Extinct) | +|
| Mittelland (MP) | +vom Aussterben bedroht (Critically Endangered) | +A3c; C1 | +
| Alpennordflanke (NA) | +verletzlich (Vulnerable) | +C2a(i) | +
|
+ |
+
+stark
+ |
+C2a(i) | +
|
+ |
+verletzlich (Vulnerable) | +C2a(i) | +
| Westliche Zentralalpen (WA) | +verletzlich (Vulnerable) | +C2a(i) | +
|
+ |
+
+4 -
+ |
+
|
+ |
+
+1 -
+ |
+
|
+ |
+1 - Gering | +
|
+ |
+
+1 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(01.01.2018) | +
|
+ |
+-- | + + +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+sehr warm-kollin (nur an
+ |
+||
|
+ |
+
+ |
+
+ |
+kontinental (sehr niedrige Luftfeuchtigkeit, sehr grosse Temperaturschwankungen, kalte Winter) | +||
|
+1.
+ |
+|
|
+2. Alle
+ |
+
+ |
+
|
+2*.
+ |
+|
|
+3.
+ |
+
+ |
+
|
+3*.
+ |
+
+ |
+
|
+1*.
+ |
+|
|
+4. Krone vor dem
+ |
+
+ |
+
|
+4*. Kronenspitze vor dem
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+|
|---|---|---|---|
|
+ |
+
+ |
+
+ |
+|
| Inflorescence | +monochasial | +monochasial | +thyrsoid or | +
| monochasial | +|||
| Glandular | +absent | +absent | +present | +
| trichomes | +|||
| 3-radiate | +present only on absent | +absent | +|
| trichomes | +the stems | +||
| Simple | +absent | +present | +present | +
| trichomes | +|||
| Bracteoles | +8-12 mm long | +3-4 mm long | +2-4 mm long | +
|
+1.
+ |
+|
|
+2.
+ |
+
+ |
+
|
+2*.
+ |
+|
|
+3. Stengel steif aufrecht; Rand des
+ |
+
+ |
+
|
+3*. Stengel bogig aufsteigend, stets
+ |
+
+ |
+
|
+1*.
+ |
+|
|
+4. Frucht den Fruchtschlauch nicht
+ |
+
+ |
+
|
+4*. Frucht den Fruchtschlauch ganz
+ |
+
+ |
+
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+235000 | +
| = | +
+ |
+
+ |
+751 | +
| = | +
+ |
+
+ |
+986 | +
| = | +
+ |
+
+ |
+986 | +
| = | +
+ |
+
+ |
+235000 | +
| = | +
+ |
+
+ |
+1253 | +
| = | +
+ |
+
+ |
+1071 | +
| = | +
+ |
+
+ |
+235000 | +
| = | +
+ |
+
+ |
+578 | +
| Species | +Location of preroal organ | +Shape of preoral lobe | +Oral ridges | +Accessory plates | +Mouthhook | +No. of comb-like processes | +|||
|---|---|---|---|---|---|---|---|---|---|
| Number | +Posterior margins | +Number | +Posterior margins | +Ventral surface | +Length b (mm) | +||||
|
+ |
+Lateral to MH | +Long, narrow, with 3-4 petal-like lobes adjacent to preoral organ | +12-14 | +Finely serrate or entire | +15-17; mostly in one series | +Finely serrate or entire | +Concave, Rough | +0.22 | +Absent | +
|
+ |
+Anterolateral to MH | +Long, narrow, split apically, extending posterior to preoral organ | +14-15 | +Entire or undulant | +27-36, covering a much larger area than oral ridges | +Dentate | +Concave, eroded | +0.21-0.23 | +Absent | +
|
+ |
+Anterolateral to MH | +Long, broad, extending posterior to preoral organ | +13-15 | +Fringed | +Present; apparently in one series | +Fringed | +Concave, medial carina, smooth | +0.16-0.17 | +Absent | +
|
+ |
+Lateral to MH | +Long, narrow, with 3-5 single or bifid secondary lobes adjacent to preoral organ | +19-20 | +Dentate with long moderately spaced projections | +Numerous plates | +Fringed | +Concave, apparently smooth | +0.21-0.22 | +Absent | +
|
+ |
+Anterior to MH | +Short, irregular-rounded lobe, smaller than preoral organ | +12-14 | +Irregularly dentate and entire | +14-20; plates in one series | +Fringed | +Concave, eroded | +0.10-0.13 | +3-5 | +
|
+ |
+? | +? | +13-15 | +? | +? | +? | +? | +? | +? | +
|
+ |
+Anterior to MH | +Short, narrow, extends to posterior middle of preoral organ | +16-19 | +Fringed, 3-4 posterior ridges entire | +~ 36; medial and posterior plates in two series | +Fringed | +Concave, Smooth | +0.12-0.15 | +6-8 | +
|
+ |
+Anterior to MH | +Short, broad, irregular shape, larger than preoral organ | +22-23 | +Densely fringed, posterior ridges dentate | +Numerous plates, overlapping with oral ridges | +Fringed | +Concave, medial carina, smooth | +0.18-0.20 | +Absent | +
|
+ |
+Lateral to MH | +Long, narrow | +18-23 | +Fringed | +Present; apparently in one series | +Fringed | +Concave, smooth | +0.24-0.25 | +Absent | +
|
+ |
+Anterior to MH | +Short-elongate, narrow, extends partially posterior to preoral organ | +13-16 | +Dentate with long closely spaced projections, 2-3 posterior ridges entire | +Numerous plates; plates in two or more series | +Fringed | +Concave, eroded | +0.16-0.17 | +7-9 | +
| Species | +Prothoracic spiracle | +Dorsal spinule pattern | +||
|---|---|---|---|---|
| No. of tubules | +Apical width (mm) | +Thoracic segment | +Abdominal segment | +|
| No. of rows | +No. of rows | +|||
|
+ |
+24-27 | +0.35-0.36 | +T1 5-7; T2 4-5; T3 absent | +Absent on A1-A8 | +
|
+ |
+17-21 | +0.28-0.33 | +T1 3; T2 3; T3 absent | +Absent on A1-A8 | +
|
+ |
+16-21 | +0.22-0.24 | +T1 9-11; T2 3-5; T3 1-2 | +Absent on A1-A8 | +
|
+ |
+20-24 | +0.32-0.35 | +T1 5-7; T2 3-4; T3 1 | +Absent on A1-A8 | +
|
+ |
+12-18 | +0.19-0.24 | +T1 6-7; T2 2-5; T3 absent | +Absent on A1-A8 | +
|
+ |
+20-22 | +? | +Present on T2-T3 with minute spinules | +? | +
|
+ |
+18-21 | +0.26-0.34 | +T1 3-5; T2 3-5; T3 1-2 | +Absent on A1-A8 | +
|
+ |
+23-29 | +0.28-0.35 | +T1 2; T2 5-6; T3 2-3 | +A1 3; absent on A2-A8 | +
|
+ |
+22-30 | +0.41-0.44 | +T1 3; T2 4-5; T3 4 | +A1 2; absent on A2-A8 | +
|
+ |
+12-17 | +0.23-0.28 | +T1 5; T2 3; T3 absent | +Absent on A1-A8 | +
| Species | +Posterior spiracle (SP-I and SP-IV) | +Anal lobe | +|||
|---|---|---|---|---|---|
| Length of spiracular opening (μm) | +No. of trunks | +No. of tips | +Basal width (μm) | +||
|
+ |
+94-101 | +SP-I 4-9; SP-IV 3-7 | +SP-I 12-21; SP-IV 10-15 | +SP-I 9-12; SP-IV 9-10 | +Grooved, entire | +
|
+ |
+76-89 | +SP-I 5-8; SP IV 4-7 | +SP I 10-18; SP IV 7-17 | +SP I 7-13; SP IV 5-7 | +Entire | +
|
+ |
+58-73 | +SP-I 14-18; SP IV 14-20 | +SP I 33-51; SP IV 31-39 | +SP I 20-30; SP IV 16-28 | +Entire | +
|
+ |
+69-80 | +SP-I 9-12; SP IV 9-12 | +SP I 13-27; SP IV 16-23 | +SP I 12-18; SP IV 14-15 | +Entire | +
|
+ |
+56-77 | +SP-I 9-15; SP-IV 8-15 | +SP-I 21-33; SP-IV 17-31 | +SP-I 14-28; SP-IV 12-21 | +Entire | +
|
+ |
+~100 | +SPI ~8-9; SP IV ~7-8 | +SPI 12; SP IV 11 | +? | +? | +
|
+ |
+83-108 | +SP-I 8-11; SP IV 4-12 | +SP I 9-26; SP IV 8-24 | +SP I 9-15; SP IV 7-12 | +Entire | +
|
+ |
+84-97 | +SP- I 9-11; SP-IV 7-11 | +SP-I 12-20; SP-IV 13-16 | +SPI 12-15; SP IV 9-19 | +Entire | +
|
+ |
+122-145 | +SP- I 5-11; SP-IV 7-10 | +SP-I 9-20; SP-IV 14-21 | +SPI 8-11; SP IV 9-12 | +Entire | +
|
+ |
+69-80 | +SP- I 13-18; SP-IV 13-17 | +SP-I 19-34; SP-IV 25-40 | +SPI 29-36; SP IV 23-34 | +Entire | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +schattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+sehr warm-kollin (nur an
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+158800 | +
| = | +
+ |
+
+ |
+2844 | +
| = | +
+ |
+
+ |
+2423 | +
| = | +
+ |
+
+ |
+2423 | +
| = | +
+ |
+
+ |
+158800 | +
| = | +
+ |
+
+ |
+643 | +
| = | +
+ |
+
+ |
+561 | +
| = | +
+ |
+
+ |
+158800 | +
| = | +
+ |
+
+ |
+2075 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +nicht anwendbar (Not Applicable) | +|
| Mittelland (MP) | +verletzlich (Vulnerable) | +D2 | +
| Alpennordflanke (NA) | +-- | +|
|
+ |
+
+potenziell
+ |
+B2a | +
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+
+ |
+|
|
+ |
+
+ |
+(23.01.2013) | +
|
+ |
+
+ |
+
+ |
+||||
|---|---|---|---|---|---|---|
|
+Male n
+ |
+
+Female n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+|
| AFW | +||||||
| Range Mean SD | +0.14–0.15 0.14 0.002 | +0.14–0.16 0.15 0.005 | +0.14–0.18 0.16 0.01 | +0.14–0.18 0.16 0.01 | +0.14–0.17 0.15 0.01 | +0.14–0.17 0.16 0.01 | +
| LW | +||||||
| Range Mean SD | +0.51–0.59 0.55 0.02 | +0.52–0.57 0.55 0.01 | +0.55–0.74 0.65 0.04 | +0.57–0.73 0.65 0.04 | +0.60–0.71 0.67 0.03 | +0.61–0.73 0.68 0.03 | +
| CL | +||||||
| Range Mean SD | +0.14–0.17 0.16 0.01 | +0.14–0.17 0.15 0.01 | +0.14–0.19 0.17 0.01 | +0.14–0.20 0.17 0.01 | +0.14–0.17 0.15 0.01 | +0.14–0.17 0.16 0.01 | +
| CW | +||||||
| Range Mean SD | +0.62–0.71 0.66 0.03 | +0.63–0.70 0.67 0.02 | +0.62–0.86 0.73 0.04 | +0.68–0.83 0.74 0.04 | +0.69–0.79 0.74 0.03 | +0.71–0.82 0.77 0.03 | +
| DD | +||||||
| Range Mean SD | +0.64–0.73 0.68 0.02 | +0.67–0.73 0.70 0.02 | +0.69–0.95 0.80 0.04 | +0.74–0.93 0.81 0.04 | +0.74–0.86 0.80 0.03 | +0.76–0.90 0.84 0.04 | +
| DDV | +||||||
| Range Mean SD | +0.25–0.30 0.28 0.01 | +0.26–0.30 0.28 0.01 | +0.27–0.38 0.32 0.02 | +0.28–0.38 0.32 0.02 | +0.29–0.36 0.33 0.02 | +0.29–0.36 0.34 0.02 | +
| DEL | +||||||
| Range Mean SD | +0.42–0.48 0.45 0.02 | +0.43–0.48 0.45 0.02 | +0.44–0.54 0.48 0.02 | +0.43–0.54 0.49 0.02 | +0.47–0.51 0.49 0.01 | +0.46–0.55 0.50 0.02 | +
| HL | +||||||
| Range Mean SD | +0.64–0.75 0.70 0.03 | +0.69–0.76 0.72 0.03 | +0.67–0.88 0.76 0.04 | +0.67–0.89 0.76 0.05 | +0.69–0.80 0.74 0.03 | +0.69–0.84 0.77 0.03 | +
| DHW | +||||||
| Range Mean SD | +1.40–1.55 1.46 0.04 | +1.40–1.53 1.47 0.05 | +1.47–1.83 1.64 0.7 | +1.55–1.81 1.66 0.07 | +1.55–1.75 1.67 0.05 | +1.59–1.82 1.73 0.06 | +
| VHW | +||||||
| Range Mean SD | +1.37–1.52 1.44 0.03 | +1.39–1.53 1.45 0.05 | +1.48–1.81 1.62 0.07 | +1.52–1.89 1.65 0.07 | +1.55–1.73 1.64 0.05 | +1.54–1.79 1.71 0.06 | +
|
+ |
+
+ |
+
+ |
+||||
|---|---|---|---|---|---|---|
|
+Male n
+ |
+
+Female n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+|
| PPH | +||||||
| Range Mean SD | +0.24–0.38 0.32 0.04 | +0.21–0.40 0.33 0.05 | +0.36–0.72 0.55 0.08 | +0.40–0.69 0.57 0.07 | +0.57–0.73 0.66 0.04 | +0.57–0.74 0.67 0.05 | +
| PL | +||||||
| Range Mean SD | +0.76–0.91 0.83 0.04 | +0.76–0.85 0.79 0.03 | +0.81–1.07 0.94 0.06 | +0.76–1.07 0.92 0.06 | +0.86–0.99 0.92 0.04 | +0.84–1.00 0.92 0.04 | +
| PW | +||||||
| Range Mean SD | +1.80–2.12 2.02 0.08 | +1.93–2.13 2.03 0.07 | +2.02–2.62 2.31 0.11 | +2.12–2.57 2.32 0.11 | +2.24–2.52 2.40 0.07 | +2.24–2.67 2.47 0.11 | +
| BH | +||||||
| Range Mean SD | +1.81–2.00 1.90 0.04 | +1.88–2.03 1.93 0.06 | +1.90–2.33 2.08 0.08 | +1.88–2.26 2.13 0.08 | +2.05–2.30 2.15 0.06 | +2.10–2.45 2.26 0.08 | +
| EL | +||||||
| Range Mean SD | +2.97–3.44 3.25 0.13 | +3.18–3.57 3.39 0.13 | +3.15–4.00 3.55 0.20 | +3.14–4.18 3.64 0.22 | +3.39–4.12 3.68 0.16 | +3.45–4.18 3.85 0.18 | +
| EH | +||||||
| Range Mean SD | +1.45–1.64 1.54 0.06 | +1.46–1.64 1.56 0.06 | +1.44–1.87 1.70 0.08 | +1.51–1.94 1.73 0.08 | +1.57–1.96 1.79 0.09 | +1.67–2.02 1.83 0.09 | +
| EW | +||||||
| Range Mean SD | +2.14–2.38 2.25 0.05 | +2.12–2.31 2.23 0.06 | +2.24–2.94 2.58 0.13 | +2.38–2.97 2.59 0.13 | +2.52–2.96 2.73 0.11 | +2.50–3.03 2.82 0.13 | +
| EPH | +||||||
| Range Mean SD | +0.10–0.17 0.12 0.02 | +0.10–0.17 0.13 0.02 | +0.10–0.52 0.29 0.10 | +0.10–0.62 0.36 0.15 | +0.26–0.36 0.31 0.02 | +0.27–0.39 0.35 0.03 | +
| PTL | +||||||
| Range Mean SD | +0.70–0.80 0.75 0.03 | +0.70–0.81 0.74 0.03 | +0.76–1.02 0.84 0.05 | +0.76–1.02 0.84 0.05 | +0.79–0.93 0.83 0.03 | +0.79–0.93 0.85 0.03 | +
| PTW | +||||||
| Range Mean SD | +0.26–0.30 0.28 0.01 | +0.21–0.26 0.23 0.01 | +0.29–0.39 0.34 0.02 | +0.25–0.33 0.28 0.02 | +0.31–0.36 0.33 0.01 | +0.26–0.39 0.29 0.02 | +
| PFL | +||||||
| Range Mean SD | +1.07–1.19 1.12 0.03 | +1.07–1.17 1.11 0.03 | +1.15–1.50 1.30 0.07 | +1.14–1.46 1.28 0.07 | +1.28–1.40 1.32 0.03 | +1.22–1.44 1.34 0.05 | +
|
+ |
+
+ |
+
+ |
+||||
|---|---|---|---|---|---|---|
|
+Male n
+ |
+
+Female n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+|
| PFW | +||||||
| Range Mean SD | +0.31–0.36 0.35 0.01 | +0.31–0.38 0.34 0.02 | +0.33–0.43 0.38 0.02 | +0.36–0.43 0.39 0.02 | +0.36–0.40 0.38 0.01 | +0.36–0.42 0.39 0.02 | +
| HTL | +||||||
| Range Mean SD | +0.61–0.71 0.66 0.03 | +0.64–0.70 0.67 0.02 | +0.69–0.89 0.79 0.04 | +0.71–0.89 0.82 0.05 | +0.73–0.84 0.80 0.03 | +0.74–0.90 0.83 0.04 | +
| HTW | +||||||
| Range Mean SD | +0.45–0.49 0.47 0.01 | +0.45–0.50 0.47 0.02 | +0.48–0.64 0.55 0.03 | +0.50–0.64 0.55 0.03 | +0.52–0.62 0.56 0.03 | +0.52–0.64 0.57 0.03 | +
| HFL | +||||||
| Range Mean SD | +0.63–0.71 0.67 0.02 | +0.62–0.71 0.67 0.03 | +0.69–0.91 0.77 0.04 | +0.71–0.88 0.79 0.04 | +0.76–0.85 0.81 0.02 | +0.74–0.90 0.84 0.04 | +
| HFW | +||||||
| Range Mean SD | +0.37–0.41 0.39 0.01 | +0.34–0.44 0.39 0.02 | +0.41–0.50 0.45 0.02 | +0.38–0.50 0.46 0.03 | +0.41–0.47 0.44 0.02 | +0.43–0.50 0.46 0.02 | +
| MTL | +||||||
| Range Mean SD | +0.40–0.46 0.42 0.02 | +0.45–0.60 0.54 0.03 | +0.48–0.60 0.53 0.03 | +|||
| MTW | +||||||
| Range Mean SD | +0.22–0.26 0.24 0.01 | +0.24–0.36 0.30 0.03 | +0.26–0.31 0.27 0.02 | +|||
| SBL | +||||||
| Range Mean SD | +4.49–5.06 4.78 0.16 | +4.68–5.09 4.89 0.16 | +4.78–5.89 5.26 0.26 | +4.69–5.98 5.34 0.30 | +4.99–5.76 5.34 0.18 | +5.04–5.97 5.54 0.22 | +
| SBL/BH | +||||||
| Range Mean SD | +2.42–2.16 2.52 0.06 | +2.46–2.68 2.53 0.06 | +2.39–2.73 2.53 0.09 | +2.25–2.97 2.51 0.11 | +2.43–2.62 2.49 0.05 | +2.34–2.58 2.46 0.07 | +
| SBL/EW | +||||||
| Range Mean SD | +2.01–2.20 2.13 0.05 | +2.08–2.23 2.18 0.04 | +1.93–2.28 2.03 0.07 | +1.89–2.32 2.06 0.08 | +1.89–2.09 1.95 0.04 | +1.88–2.16 1.96 0.06 | +
| EL/EW | +||||||
| Range Mean SD | +1.33–1.52 1.44 0.05 | +1.41–1.55 1.50 0.04 | +1.28–1.51 1.38 0.06 | +1.27–1.63 1.40 0.06 | +1.30–1.42 1.34 0.03 | +1.30–1.53 1.36 0.05 | +
|
+ |
+
+ |
+
+ |
+||||
|---|---|---|---|---|---|---|
|
+Male n
+ |
+
+Female n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+
+Male (n
+ |
+
+Female (n
+ |
+|
| PTL/PTW | +||||||
| Range Mean SD | +2.39–2.91 2.69 0.16 | +2.93–3.56 3.17 0.20 | +2.19–2.80 2.48 0.15 | +2.58–3.36 2.96 0.15 | +2.24–2.69 2.50 0.11 | +2.30–3.14 2.91 0.17 | +
| PW/PL | +||||||
| Range Mean SD | +2.23–2.69 2.42 0.11 | +2.44–2.72 2.55 0.09 | +2.24–2.72 2.45 0.11 | +2.31–2.87 2.52 0.10 | +2.48–2.80 2.62 0.09 | +2.39–3.03 2.68 0.15 | +
| HW/HL | +||||||
| Range Mean SD | +1.96–2.22 2.08 0.07 | +1.96–2.14 2.04 0.06 | +1.98–2.42 2.16 0.09 | +1.93–2.47 2.18 0.10 | +2.13–2.40 2.25 0.07 | +2.11–2.40 2.26 0.08 | +
| EW/PW | +||||||
| Range Mean SD | +1.06–1.16 1.11 0.02 | +1.09–1.14 1.11 0.01 | +1.00–1.17 1.12 0.02 | +1.02–1.17 1.12 0.03 | +1.10–1.18 1.13 0.02 | +1.02–1.23 1.15 0.04 | +
| PW/HW | +||||||
| Range Mean SD | +1.25–1.48 1.38 0.05 | +1.33–1.41 1.38 0.02 | +1.36–1.48 1.41 0.02 | +1.32–1.47 1.40 0.03 | +1.42–1.47 1.44 0.01 | +1.34–1.56 1.43 0.05 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +schattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+187800 | +
| = | +
+ |
+
+ |
+1384 | +
| = | +
+ |
+
+ |
+1065 | +
| = | +
+ |
+
+ |
+1065 | +
| = | +
+ |
+
+ |
+187800 | +
| = | +
+ |
+
+ |
+1896 | +
| = | +
+ |
+
+ |
+1557 | +
| = | +
+ |
+
+ |
+187800 | +
| = | +
+ |
+
+ |
+935 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +verletzlich (Vulnerable) | +D2 | +
| Mittelland (MP) | +nicht anwendbar (Not Applicable) | +|
| Alpennordflanke (NA) | +
+stark
+ |
+C2a(i) | +
|
+ |
+
+potenziell
+ |
+C2a(i) | +
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +
+stark
+ |
+D | +
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+|||||
|---|---|---|---|---|---|
|
+ |
+
+ |
+||||
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
| CL | ++ | +4 | +ML | +F | +|
| Mc | +G | +||||
| Af | ++ | +10 | +ML | +B | +|
| Mc | +A,C,D,E | +||||
| Oc | ++ | +3 | +ML | +OCm | +|
| Mc | +OCp | +||||
| me or Mc | +OCa | +||||
| Di + De | ++ | +4 | +ML | +Di1,De1 | +|
| Mc | +Di2,De2 | +||||
| DL | ++ | +6 | +ML | +DL1,DL5 | +|
| Mc | +DL2,DL3,DL4,DL6 | +||||
| L + So | ++ | +10 | +ML | +So1,L1,L4 | +|
| Mc | +L2,L3 | +||||
| me | +So3 to 6 | +||||
| mi | +L3’ | +||||
| Vi: 6 | +|||||
| Ve: 11 | +|||||
| Labrum: 4/2.4 | +|||||
| Labium: 11, 0x | +|||||
| Ant.I, II: 7, 12 | +|||||
| Ant.III: 17+5s (a4 absent) | +|||||
| Ant.IV: 12 mou+or+i+8S | +|||||
|
+ |
+|||||
|
+ |
+
+ |
+
+ |
+|||
| Th.I | +1 2 | +1 | +0 3 | +6 | +13 19 | +
| Th.II | +3 3+S | +3+S+ms | +3 2 7 | +6 | +12 19 | +
| Th.III | +3 3+S | +3+S | +3 2 8 | +6 | +11 18 | +
| Abd.I | +2 3+S | +2 | +3 VT: 4 | +||
| Abd.II | +2 3+S | +2 | +3 Ve: 5 (Ve1 present) | +||
| Abd.III | +2 3+S | +2 | +4 Ve: 5 or 6; Fu: 6me, 0mi | +||
| Abd.IV | +2 2+S | +3 | +8-9 Ve: 8; VL: 4 | +||
| Abd.V | +3+3 | +7+S | +1 Ag: 3; VL: 1 | +||
| Abd.VI | +
+ |
+Ve: 14-15; An: 2 | +|||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| 1618191619 | +2 | +
| 1346910121315 | +4 | +
|
+ |
+|
| 21161819211821 | +3 | +
| 18211619MSLBWM | +
+ |
+
| MSLBWM | +
+ |
+
|
+ |
+|
| 5 | +|
| 1215111412 | +
+ |
+
| 3698593 | +6 | +
|
+ |
+
+ |
+
| 879 | +
+ |
+
| SK | +
| Saskatchewan: 54.4188°, -108.944° 53.9804°, -106.28° 49.5978°, -109.9231° | +
|
+ |
+
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +0 - unbedeutend, keine Bindung. | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+438495 | +
| = | +
+ |
+
+ |
+438495 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +-- | +|
|
+ |
+
+stark
+ |
+D | +
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+ |
+
+3 - Mittlere nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+1 -
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+1.
+ |
+
+ |
+
|
+1*.
+ |
+|
|
+2.
+ |
+
+ |
+
|
+2*.
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +sehr trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+1.3.3 - Kalkarme Quellflur (
+ |
+
|
+6.1.4 - Hartholz-Auenwald (
+ |
+
|
+6.6.1 - Tannen-Fichtenwald (
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +schattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+251400 | +
| = | +
+ |
+
+ |
+846 | +
| = | +
+ |
+
+ |
+1382 | +
| = | +
+ |
+
+ |
+1382 | +
| = | +
+ |
+
+ |
+251400 | +
| = | +
+ |
+
+ |
+2331 | +
| = | +
+ |
+
+ |
+1895 | +
| = | +
+ |
+
+ |
+251400 | +
| = | +
+ |
+
+ |
+1263 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| Species | +TII | +TIII | +AI | +AII | +AIII | +AIVant. | +AIVmed. | +AIV pos. | +Ref. | +
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+19 | +14 | +6 | +5 | +1 | +6 | +4 | +3 | +1 | +
|
+ |
+24 | +17 | +9 | +5 | +1 | +9 | +4 | +4 | +1,2 | +
|
+ |
+13 | +11 | +5 | +4 | +1 | +4 | +5 | +4 | +3 | +
|
+ |
+22 | +15 | +9 | +5 | +1 | +8 | +4 | +4 | +2 | +
|
+ |
+15 | +15 | +6 | +5 | +1 | +5 | +4 | +3 | +4 | +
|
+ |
+14 | +14 | +6 | +4 | +1 | +4 | +4 | +3 | +4 | +
| Features | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| Ratio height/width of RR-ring<1.7 | ++ | ++ | ++ | ++ | ++ | +− | +
| Ratio height/width of IBr2<1.4 | ++ | ++ | ++ | +− | ++ | ++ | +
| Brs bearing P1 | +8–10 | +5–15 | +5–10 | +10–14 | +10–16 | +4 | +
| Number of knobby processess on IBrs 1 | +0 | +0–9 | +0 | +?–10 | +9 | +? | +
| Number of knobby processes on IBrs 2 | +4 | +4–101 | +9 | +?–10 | +0 | +? | +
| Knobby processes on Brs 1–2 present | ++ | +− | +− | +− | +− | +− | +
| Knobbles arranged in nearly parallel ridges | +− | +− | +− | +− | ++ | ++ | +
| Cover plates | ++ | ++ | ++ | +− | ++ | ++ | +
| Most frequent distal arm pattern a b c+d e f+g h i | ++ | +−/+2 | +− | +− | +− | +−/+ | +
| Numerous needle-like spines on external surface of IBrs and Brs | ++ | +− | +− | +− | +− | +− | +
| Pinnule absent in every third Br (distal arm) | ++ | +−/+3 | +− | +− | +− | +− | +
| Side plates in pinnules | ++ | +− | +− | +− | ++ | +? | +
| X-shaped tube feet plates present | ++ | ++ | ++ | ++ | +− | +? | +
| Saccules present | +− | +− | +− | +− | ++ | +? | +
| Deep ligament depression in mesistele synarthries | +− | ++ | ++ | ++ | ++ | +− | +
| Attachment by root-like radix | ++ | ++4 | ++ | +−5 | ++ | +− | +
| Pn2 | +Pn3 | +Pn4 | +Pn5 | +Pn6 | +Pn7 | +Pn8 | +|
|---|---|---|---|---|---|---|---|
| P1 on Br8 | +1.07 | +1.24 | +1.32 | +1.11 | +1.00 | +0.78 | +— | +
| P1 on Br9 | +1.00 | +1.14 | +1.14 | +0.98 | +0.80 | +0.71 | +— | +
| P1 on Br9? | +1.23 | +1.47 | +1.34 | +1.19 | +1.12 | +0.93 | +0.49 | +
| P1 on Br9 | +1.02 | +1.25 | +1.08 | +0.87 | +0.85 | +0.64 | +0.43 | +
|
+ |
+
+ |
+
+ |
+
|---|---|---|
|
+ |
+1.670–1.720 | +1.650–1.800 | +
|
+ |
+0.340–0.390 | +0.360–0.390 | +
|
+ |
+0.695–0.720 | +0.800–0.850 | +
|
+ |
+0.190–0.210 | +0.230–0.280 | +
|
+ |
+0.080–0.100 | +0.120–0.130 | +
|
+ |
+0.120–0.130 | +0.140–0.150 | +
|
+ |
+0.165–0.170 | +0.160–0.170 | +
|
+ |
+0.125–0.135 | +0.130 | +
|
+ |
+0.035–0.045 | +0.030–0.0340 | +
|
+ |
+0.070 | +0.070–0.075 | +
|
+ |
+0.520–0.540 | +0.620 | +
|
+ |
+0.750–0760 | +0.950–0.970 | +
|
+ |
+0.150–0.155 | +- | +
|
+ |
+0.025–0.030 | +0.025–0.035 | +
|
+ |
+
+ |
+
+ |
+
|---|---|---|
| Apterous viviparous females | +||
| Spinal scleroites | +two rows and larger than the other ones | +one row and of the same size as the other ones | +
| Abdominal pleural setae | +visibly longer than spinal ones | +of the same length as other ones | +
| ABD I–V longest setae | +0.027 mm | +0.050 mm | +
| ANT VI basal setae | +4 | +5–6 | +
| III TIBIAE setae | +not of the same length, longest ones up to 0.115 mm | +of the same length, longest ones about 0.075 mm | +
| HT II setae | +with two long dorsal setae, much longer than other tarsal setae | +all setae of the same length | +
| PT/BASE | +0.25–0.32 | +0.17–0.25 | +
| URS/ANT III | +0.33–0.36 | +0.26–0.33 | +
| URS/ANT VI | +0.35–0.40 | +0.41–0.42 | +
| Alate viviparous females | +||
| Spinal scleroites | +two rows and larger than the other ones | +one row and of the same size as the other ones | +
| Abdominal pleural setae | +visibly longer than spinal ones | +of the same length as other ones | +
| ANT VI basal setae | +4 | +5–6 | +
| Head longest setae | +0.070 mm | +0.110 mm | +
| III TIBIAE setae | +not of the same length, longest ones up to 0.120 mm | +of the same length, longest ones about 0.090 mm | +
| Species | +Type host | +Locality (Country) | +Males | +||||
|---|---|---|---|---|---|---|---|
| Spicule (µm) | +Spicule tip | +Caudal Papillae distribution (pre:ad:post cloacal) | +Unpaired post cloacal papilla | +Clocal lip | +|||
|
+ |
+
+ |
+Fernando de Noronha, state of Pernambuco (Brazil) | +80–100 | +sharp | +1:1:1 | +No | +Smooth | +
|
+ |
+
+ |
+Toledos, state of Minas Gerais (Brazil) | +100–140 | +sharp | +1:0:3 | +No | +Echinate | +
|
+ |
+
+ |
+Province of La Rioja, (Argentina) | +90–110 | +sharp | +1:0:2 | +Yes | +Echinate | +
|
+ |
+
+ |
+Exu, state of Pernam- buco (Brazil) | +57–71 | +blunt | +1:1:2 | +Yes | +Smooth | +
|
+ |
+
+ |
+Province of San Juan (Argentina) | +80–180 | +sharp | +2:0:2 | +No | +Echinate | +
|
+ |
+
+ |
+Miranda, State of Mato Grosso do Sul (Brazil) | +40–58 | +sharp | +0:2:1 | +No | +Echinate | +
|
+ |
+
+ |
+State of Rio de Janeiro (Brazil) | +54–68 | +sharp | +0:2:1 | +Yes | +Smooth | +
|
+ |
+
+ |
+Rio de Janeiro, State of Rio de Janeiro (Brazil) | +82–100 | +Sharp | +2:1:1 | +Yes | +Smooth | +
|
+ |
+
+ |
+João Alfredo, State of Pernambuco (Brazil) | +54–68 | +Sharp | +1:1:1 | +Yes | +Smooth | +
|
+ |
+
+ |
+Caxiuanã National Forest, State of Pará (Brazil) | +53–75 | +sharp | +1:1:1 | +No | +Echinate | +
|
+ |
+
+ |
+Petrolina, State of Per- nambuco (Brazil) | +110–123 | +sharp | +1:0:3 | +No | +Smooth | +
| Species | +Females | +Reference | +|||
|---|---|---|---|---|---|
| Ovary location | +Vulva | +Tail terminus | +Eggs hell | +||
|
+ |
+Pre-bulbar | +Equatorial | +Spike stout | +Thin, smooth | +
+ |
+
|
+ |
+Pre-bulbar | +Equatorial | +Spike stout | +Thick, punctated | +
+ |
+
|
+ |
+Post bulbar | +Post equatorial | +Spike stout | +Thin, punctated | +
+ |
+
|
+ |
+Post bulbar | +equatorial | +Spike stout | +Thick, punctated | +
+ |
+
|
+ |
+Post bulbar | +equatorial | +Spike stout | +Thin, punctated | +
+ |
+
|
+ |
+Post bulbar | +Pre equatorial | +Conical | +Thin | +
+ |
+
|
+ |
+Pre bulbar | +Pre equatorial | +Spike stout | +Thin, smooth | +
+ |
+
|
+ |
+Pre bulbar | +Equatorial | +Spike stout | +- | +
+ |
+
|
+ |
+Pre bulbar | +Equatorial | +Spike stout | +- | +
+ |
+
|
+ |
+Post bulbar | +Pre equatorial | +Conical | +Thick | +
+ |
+
|
+ |
+Pre bulbar | +equatorial | +Spike stout | +Thick, punctated | +Current study | +
| Species | +Type host | +Country | +
+ |
+
+ |
+
+ |
+||
|---|---|---|---|---|---|---|---|
| (Biogeographical realm) | +
+ |
+
+ |
+
+ |
+
+ |
+|||
|
+ |
+
+ |
+Botswana (Afrotropical) | +79–98 | +Sharp | +Stout spike | +Analate, shell smooth | +
+ |
+
|
+ |
+
+ |
+USA (Nearctic) | +53–76 | +Sharp | +Stout spike | +Analate, shell smooth | +
+ |
+
|
+ |
+
+ |
+India (Oriental) | +60–75 | +Sharp | +Stout spike | +Analate, shell punctate | +
+ |
+
|
+ |
+
+ |
+Kazakhstan (Palearctic) | +139–176 | +Sharp | +Thin spike | +Analate, shell smooth | +
+ |
+
|
+ |
+
+ |
+India (Oriental) | +79–92 | +Blunt | +Stout spike | +Not described | +
+ |
+
|
+ |
+
+ |
+Brazil (Neotropical) | +110–123 | +Sharp | +Stout spike | +Analate, shell punctate | +Current study | +
|
+ |
+
+ |
+
+ |
+
+ |
+|
|---|---|---|---|---|
|
+ |
+
+11
+ |
+
+7
+ |
+plus 4 (0.36) | +paene 8 (0.67) | +
|
+ |
+15.00 (1.00) | +8.40 (0.56) | +5.60 (0.37) | +9.00 (0.60) | +
|
+ |
+15.52 (1.00) | +8.40 (0.54) | +5.04 (0.32) | +8.88 (0.57) | +
|
+ |
+18.80 (1.00) | +10.50 (0.56) | +5.70 (0.30) | +10.70 (0.57) | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| P1 | +P2 | +P3 | +
|---|
|
+ |
+|||
|---|---|---|---|
|
+ |
+
+ |
+♀ | +J | +
|
+Mixed Samples - BHCB 179859
+ |
+8 0 | +0 0 | +|
|
+BHCB 179940
+ |
+9 | +0 | +|
|
+BHCB 179940
+ |
+0 | +0 | +|
|
+BHCB 179868
+ |
+7 | +0 | +|
|
+BHCB 179868
+ |
+0 | +0 | +|
|
+BHCB 179912
+ |
+10 | +0 | +|
|
+BHCB 179912
+ |
+0 | +0 | +|
|
+BHCB-187650
+ |
+9 | +0 | +|
|
+BHCB-187650
+ |
+0 | +0 | +|
|
+BHCB-187625
+ |
+6 | +0 | +|
|
+BHCB-187625
+ |
+0 | +0 | +|
|
+BHCB-187612
+ |
+10 | +0 | +|
|
+BHCB-187612
+ |
+0 | +0 | +|
|
+BHCB-187663
+ |
+8 | +0 | +|
|
+BHCB-187663
+ |
+0 | +0 | +|
|
+BHCB-187641
+ |
+0 | +9 | +|
|
+ |
+
+BHCB 179921
+ |
+0 | +0 | +
|
+BHCB 179935
+ |
+0 | +3 | +|
|
+BHCB 179910
+ |
+0 | +0 | +|
|
+BHCB 179839
+ |
+1 | +0 | +|
|
+BHCB-187626
+ |
+2 | +0 | +|
|
+BHCB-187632
+ |
+1 | +1 | +|
|
+BHCB-187631
+ |
+1 0 | +0 0 | +|
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+404450 | +
|
+ |
+
+ |
+
+ |
+|
|---|---|---|---|
| Standard length (SL) in mm | +10.0–12.6 (12.6) | +||
|
+ |
+|||
| Head length (HL) | +20–21.9 (21.4) | +21.0 | +0.6 | +
| Predorsal length | +66.1–71.4 (69.6) | +69.6 | +1.4 | +
| Preanal length | +52.5–56.3 (54.5) | +54.5 | +1.3 | +
| Body depth at vent | +15.4–19.0 (19.0) | +17.3 | +1.1 | +
| Caudal peduncle depth | +7.7–9.0 (8.7) | +8.4 | +0.4 | +
|
+ |
+|||
| Eye diameter | +28.0–31.8 (29.6) | +30.0 | +1.8 | +
| Snout length | +20.0–24.0 (22.2) | +22.6 | +1.4 | +
| Character | +
+ |
+
|---|---|
| Sex | +Male | +
| SVL | +415.0 | +
| TaL | +50.0* | +
| TL | +465.0 | +
| HL | +15.3 | +
| HW | +10.8 | +
| HH | +6.1 | +
| IO | +6.3 | +
| EN | +3.3 | +
| IN | +4.1 | +
| ED | +2.6 | +
| SnL | +6.0 | +
|
+ |
+|
| ASR | +17 | +
| MSR | +17 | +
| PSR | +15 | +
| Keeling | +6 dorsal scale rows on the posterior body third feebly keeled | +
| VEN | +241 | +
| PrVEN | +2 | +
| Ventral notched | +Yes | +
| Ventral keeled | +No | +
| SC | +26* | +
| divided | +Yes | +
| Cloacal | +Single | +
| Loreal | +1/1 | +
| Loreal entering orbit | +Yes | +
| SL | +8/8 | +
| entering orbit | +3,4,5 | +
| largest SL | +6/6 | +
| IL | +10/10 | +
| IL in contact with 1st chin shield | +1−5 | +
| PreOc | +1/1 | +
| PostOc | +2/2 | +
| Temporal scales | +|
| anterior | +2/2 | +
| posterior | +3/3 | +
| Scales around paraparietal | +5/7 | +
| Scales between parietals | +4 | +
| Nuchal band | +Absent | +
| Body blotches | +87 (yellow blotches) | +
| Tail blotches | +15* | +
| Belly pattern | +uniform grey cream | +
| Ventral tail pattern | +grey cream | +
| First body blotch position (at VEN) | +13 | +
| First blotch width (vertebral scales) | +0.5 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
|
+ |
+Yellowish grey, corticolous, loosely adnate | +Whitish grey, saxicolous, loosely adnate | +Greyish green to light brown, corticolous or saxicolous, moderately adnate | +Yellowish green to yellowish brown, corticolous, adnate to a little extend | +Glaucous white, Grey to darker grey, corticolous, rarely saxicolous, loosely adnate | +Grey to grey green, corticolous, rarely saxicolous, loosely adnate | +
|
+ |
+Rotund to crenate | +Rounded to undulate | +Rounded to subrounded | +Subrounded, flat to plane | +Rotund | +Rotund | +
|
+ |
+Yellow | +Yellow | +Yellow | +Yellow | +
+Normally white, yellow colour in lower medulla is reported by
+ |
+White in upper part and yellow in lower part | +
|
+ |
+Eciliate | +Eciliate | +Eciliate | +Ciliate | +Ciliate | +Ciliate | +
|
+ |
+Isidia laminal, dense to simple coralloid | +Isidia coarse, pustular, hollow | +Isidia slender, dense, not pustular | +Isidia fine, sparse, cylindrical | +Sorediate | +Sorediate | +
| Chemistry (medulla) | +K: + yellow turning darker, C: + slight yellow-red, KC: -, P: - | +K: + yellow turning red, C: -, KC: -, P: + yellow | +K: -, C: + red, KC: + red, P: - | +K: -, C: -, KC: -, P: - | +K: -, C: + rose-red, KC: + red, P: - | +K: -; C: + pink, KC: + red. P: - | +
|
+ |
+Galbinic acid, entothein, atranorin | +Salazinic acid, atranorin, consalazinic acid | +Atranorin, entothein & gyrophoric acid | +Atranorin and vulpinic acid | +Atranorin and gyrophoric acid | +Atranorin, gyrophoric acid and an unknown pigment | +
| Specimens | +
+holotype
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Code | +Parameters | +mm % Mean | +Min | +Max | +S.D. Mean Min Max S.D. | +
| TOT | +Total length (mm) | +391 100 392.4 | +354 | +446 | +33.4 251.5 100 335 117.6 | +
| PD1 | +Pre-first dorsal length | +175.6 44.9 43.2 | +41.7 | +44.9 | +1.2 44.5 43.2 46.6 1.3 | +
| PD2 | +Pre-second dorsal length | +245 62.7 61.5 | +60.1 | +62.9 | +1.3 63.1 60.4 65.1 1.7 | +
| PRC | +Precaudal length | +281 71.9 72 | +71.3 | +73.3 | +0.8 73.4 71.1 79 3 | +
| PP1 | +Prepectoral length | +76 19.4 18.7 | +17.8 | +19.4 | +0.6 20.3 19.2 21.5 0.9 | +
| PP2 | +Prepelvic length | +154.8 39.6 38.3 | +35.8 | +39.6 | +1.7 39.7 37.2 42.1 1.9 | +
| PAL | +Preanal length | +224.5 57.4 56 | +54.2 | +57.4 | +1.2 56.1 54.2 58.4 1.8 | +
| SVL | +Snout-vent length | +174.8 44.7 42.5 | +41.1 | +44.7 | +1.5 43.1 40.7 44.8 1.6 | +
| POB | +Preorbital length (direct) | +31.9 8.2 7.6 | +7.1 | +8.2 | +0.4 7.4 6.6 7.8 0.5 | +
| EYL | +Eye length | +12.9 3.3 3.2 | +2.6 | +3.5 | +0.3 3.4 3.2 3.7 0.2 | +
| EYH | +Eye height | +3.2 0.8 0.8 | +0.6 | +1 | +0.2 1.5 0.9 2.1 0.5 | +
| INO | +Interorbital space | +25.7 6.6 6.5 | +6 | +7 | +0.4 6.5 6 6.8 0.3 | +
| PSP | +Prespiracular length | +48.4 12.4 11.8 | +11.2 | +12.4 | +0.5 11.6 10.6 12.2 0.6 | +
| SPL | +Spiracle length | +2.5 0.6 0.7 | +0.5 | +0.8 | +0.1 0.9 0.6 1.2 0.2 | +
| PRN | +Prenarial length | +19.6 5 4.5 | +3.9 | +5 | +0.4 4 3 4.5 0.6 | +
| NOW | +Nostril width | +9 2.3 2.3 | +2.2 | +2.3 | +0.1 2.4 2.2 2.7 0.2 | +
| INW | +Internarial space | +10.9 2.8 2.9 | +2.7 | +3.2 | +0.2 2.6 2.4 3 0.2 | +
| ANF | +Anterior nasal flap length | +3.4 0.9 0.8 | +0.8 | +0.9 | +0.1 0.9 0.6 1.2 0.3 | +
| POR | +Preoral length | +30.9 7.9 7.4 | +6.9 | +7.9 | +0.4 7.1 6.4 7.8 0.6 | +
| MOW | +Mouth width | +30.4 7.8 7.6 | +6.9 | +8.1 | +0.6 7.3 7 8.1 0.5 | +
| MOL | +Mouth length | +12.5 3.2 3.1 | +2.8 | +3.6 | +0.3 3.3 3 3.8 0.3 | +
| ULA | +Upper labial furrow length | +6.4 1.6 1.7 | +1.4 | +2.1 | +0.3 2 1.6 2.4 0.3 | +
| LLA | +Lower labial furrow length | +11.2 2.9 2.5 | +2.1 | +2.9 | +0.3 2.2 1.9 2.4 0.2 | +
| PG1 | +Prebranchial length | +65.6 16.8 15.7 | +14.9 | +16.8 | +0.8 15.8 13.8 16.7 1.1 | +
| HDL | +Head length | +79.5 20.3 19.4 | +18.4 | +20.3 | +0.8 20.7 18.3 22.1 1.4 | +
| GS1 | +First gill slit height | +7.6 1.9 1.8 | +1.6 | +2 | +0.2 1.5 1.2 1.8 0.2 | +
| GS5 | +Fifth gill slit height | +5.5 1.4 1.3 | +1.1 | +1.6 | +0.2 1.1 0.6 2.1 0.6 | +
| D1A | +First dorsal anterior margin | +34.2 8.7 9.4 | +8.4 | +10.1 | +0.8 9.1 8.4 10 0.6 | +
| D1B | +First dorsal base | +22.4 5.7 6 | +5.6 | +6.4 | +0.4 6.9 5.7 7.9 0.9 | +
| DIH | +First dorsal height | +14.9 3.8 3.8 | +3.5 | +4.3 | +0.3 4.3 3.8 4.8 0.4 | +
| D1I | +First dorsal inner margin | +9.2 2.4 2.4 | +2.3 | +2.5 | +0.1 2.5 2.1 2.7 0.2 | +
| D1P | +First dorsal posterior margin | +12.4 3.2 3.3 | +3.1 | +3.4 | +0.1 3.3 2.6 3.9 0.4 | +
| IDS | +Interdorsal space | +51.2 13.1 13.2 | +12.4 | +13.7 | +0.5 12.4 11.2 13.4 0.7 | +
| D2A | +Second dorsal Anterior margin | +31.2 8 8.7 | +7.7 | +9.6 | +0.8 7.8 7.4 8.3 0.3 | +
| D2B | +Second dorsal base | +22.9 5.9 6 | +5.3 | +6.5 | +0.5 5.9 5.4 6.7 0.5 | +
| D2H | +Second dorsal height | +11.3 2.9 3.3 | +2.8 | +3.8 | +0.5 3.6 3.3 3.9 0.3 | +
| D2I | +Second dorsal inner margin | +8.6 2.2 2.4 | +2 | +2.7 | +0.3 2.4 2.2 2.5 0.1 | +
| D2P | +Second dorsal posterior margin | +12.4 3.2 3.2 | +2.8 | +4 | +0.5 3 2.7 3.3 0.3 | +
| P1A | +Pectoral anterior margin | +42.3 10.8 11.3 | +10.3 | +12 | +0.7 10.7 9.7 11.1 0.6 | +
| P1B | +Pectoral base | +23.9 6.1 5.6 | +5.3 | +6.1 | +0.4 5.9 5.2 6.4 0.4 | +
| P1I | +Pectoral inner margin | +22.8 5.8 6 | +5.3 | +6.7 | +0.6 5.6 4.6 6.9 0.8 | +
| P1P | +Pectoral posterior margin | +33.1 | +8.5 | +8.7 | +8 | +9.5 | +0.6 | +7.8 | +7 | +8.9 | +0.8 | +
|---|---|---|---|---|---|---|---|---|---|---|---|
| PPS | +Pectoral-pelvic space | +55.6 | +14.2 | +13.6 | +10.9 | +16.6 | +2.1 | +15.4 | +14.3 | +16.4 | +0.7 | +
| P2A | +Pelvic anterior margin | +26.6 | +6.8 | +6.1 | +5.4 | +6.8 | +0.7 | +5.8 | +4.8 | +6.4 | +0.6 | +
| P2B | +Pelvic base | +36.2 | +9.3 | +8.3 | +7.6 | +9.3 | +0.7 | +7.9 | +7.1 | +9.7 | +0.9 | +
| P2L | +Pelvic length | +49.9 | +12.7 | +12 | +10.6 | +12.9 | +1 | +10.8 | +10.3 | +11.3 | +0.4 | +
| P2I | +Pelvic inner margin length | +11.8 | +3 | +3.6 | +2.8 | +4.2 | +0.7 | +3.1 | +2.2 | +4.5 | +0.8 | +
| P2P | +Pelvic posterior margin length | +30.2 | +7.7 | +7.6 | +6.5 | +8.6 | +1 | +5.8 | +4.8 | +7.5 | +1.1 | +
| PAS | +Pelvic-anal space | +39.5 | +10.1 | +10.6 | +10 | +11.7 | +0.8 | +9 | +8 | +10.3 | +1 | +
| ANA | +Anal anterior margin | +24.6 | +6.3 | +6.7 | +6.2 | +7.8 | +0.7 | +7.5 | +7 | +9 | +0.8 | +
| ANB | +Anal base | +34.5 | +8.8 | +9.3 | +7.9 | +10.2 | +0.9 | +10.9 | +10.3 | +11.6 | +0.4 | +
| ANL | +Anal length | +41.7 | +10.7 | +10.8 | +9.6 | +12 | +0.9 | +12.8 | +12.4 | +13.8 | +0.6 | +
| ANI | +Anal Inner margin | +6.9 | +1.8 | +1.7 | +1.3 | +1.9 | +0.2 | +1.8 | +1.5 | +2.1 | +0.2 | +
| ANP | +Anal posterior margin | +23 | +5.9 | +5.4 | +4.6 | +5.9 | +0.6 | +6.9 | +6.4 | +7.3 | +0.5 | +
| DCS | +Dorsal-caudal space | +17 | +4.4 | +3.6 | +3 | +4.4 | +0.5 | +3.5 | +2.9 | +4.2 | +0.4 | +
| ACS | +Anal-caudal space | +24.6 | +6.3 | +6.9 | +6.1 | +8.1 | +0.9 | +6.3 | +5.8 | +7.6 | +0.7 | +
| CPH | +Caudal peduncle height | +12.4 | +3.2 | +3.2 | +2.8 | +3.6 | +0.3 | +3.1 | +3 | +3.3 | +0.1 | +
| CPW | +Caudal peduncle width | +8.7 | +2.2 | +2 | +1.6 | +2.7 | +0.5 | +2.4 | +2.1 | +2.7 | +0.3 | +
| CDM | +Dorsal caudal margin | +105 | +26.8 | +27.5 | +25.9 | +28.3 | +1 | +26.8 | +25.4 | +27.9 | +1.1 | +
| CPV | +Preventral caudal margin | +29.7 | +7.6 | +8 | +7.3 | +8.6 | +0.5 | +8.9 | +8.2 | +10.2 | +0.7 | +
| CST | +Subterminal caudal margin | +1.3 | +0.3 | +0.3 | +0.2 | +0.4 | +0.1 | +4.1 | +3.6 | +4.8 | +0.4 | +
| CLO | +Clasper outer length | +38.3 | +9.8 | +10 | +9 | +11.2 | +1.1 | +6.7 | +5.2 | +7.3 | +1 | +
| CLI | +Clasper inner length | +41.7 | +10.7 | +10.9 | +9.8 | +12.3 | +1.3 | +9.9 | +8.2 | +10.8 | +1.2 | +
| CLB | +Clasper base width | +5.4 | +1.4 | +1.2 | +1 | +1.4 | +0.2 | +1.6 | +1.5 | +1.7 | +0.1 | +
| HDH | +Head height | +27.3 | +7 | +7.7 | +7 | +8.5 | +0.7 | +7.1 | +6.3 | +8.7 | +0.9 | +
| HDW | +Head width | +35.7 | +9.1 | +9.1 | +8.3 | +9.9 | +0.6 | +9.9 | +8.9 | +11.6 | +1 | +
| ANH | +Anal height | +12.6 | +3.2 | +3.3 | +2.9 | +4.3 | +0.6 | +3.3 | +3 | +3.5 | +0.2 | +
| CTR | +Terminal caudal margin | +17 | +4.3 | +4.7 | +3.9 | +5.3 | +0.6 | +4.2 | +3.9 | +4.5 | +0.2 | +
|
+Ratios
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Holo mean mini | +max | +Holo mean mini | +max | +mean mini max | +|
| 28/46 | +Head lenght /pectoral-pelvic space 1.4 1.4 1.1 | +1.7 | +1.4 1.3 1.1 | +1.5 | +1.3 1.1 1.5 | +
| 5/9 | +pre-first dorsal length / prepelvic lgth 1.1 1.1 1.1 | +1.2 | +1.1 1.1 1.1 | +1.2 | +1.1 1.1 1.2 | +
| 5/36 | +pre-first dorsal length / interdorsal space 3.4 3.4 3 | +4.1 | +3.6 3.5 3.3 | +3.6 | +3.6 3.3 4.1 | +
| 6/10 | +pre-2nd dorsal length / preanal lgth 1.1 1.1 1.1 | +1.1 | +1.1 1.1 1.1 | +1.2 | +1.1 1.1 1.2 | +
| 52/54 | +Pelvic-anal space / anal base length 1.1 1.1 1 | +1.4 | +0.9 0.9 0.8 | +0.9 | +0.8 0.7 0.9 | +
| 52/59 | +pelvic-anal space / anal-caudal space 1.6 1.5 1.1 | +1.7 | +1.6 1.5 1.3 | +1.6 | +1.4 1.3 1.6 | +
| 59/54 | +anal-caudal space / anal base length 0.7 0.8 0.6 | +1 | +0.6 0.6 0.5 | +0.7 | +0.6 0.5 0.7 | +
| 22/23 | +Preoral length / mouth width 1 1 0.9 | +1.1 | +1 0.9 0.8 | +1 | +1 0.8 1.1 | +
| 22/13 | +preoral lenght / eye length 2.4 2.3 2 | +2.8 | +2.4 2.1 2 | +2.4 | +2.1 1.8 2.4 | +
| 8/18 | +Prepectoral lenght / prenarial length 3.9 4.2 3.9 | +4.5 | +5.1 5.5 5.1 | +6.5 | +5.2 4.4 6.5 | +
| 18/13 | +Premarial lenght / eye length 1.5 1.4 1.1 | +1.7 | +1.3 1.1 0.9 | +1.3 | +1.2 0.9 1.3 | +
| 28/13 | +Head length / eye length 6.1 6.1 5.3 | +7.1 | +6.7 6.2 5.7 | +6.7 | +6.1 5.7 6.7 | +
| 23/24 | +Mouth width / mouth length 2.4 2.5 1.9 | +2.9 | +2.5 2.3 1.8 | +2.6 | +2.2 1.8 2.6 | +
| 33/39 | +1st dorsal height / second dorsal height 1.3 1.2 0.9 | +1.6 | +1.5 1.2 1.1 | +1.5 | +1.2 1.1 1.5 | +
| 36/54 | +Interdorsal space / anal base length 1.5 1.4 1.2 | +1.7 | +1.2 1.2 1.1 | +1.2 | +1.1 1.1 1.2 | +
| 60/61 | +caudal peduncle heigh / peduncle width 1.4 1.7 1.3 | +2 | +1.1 1.2 1.1 | +1.5 | +1.3 1.1 1.6 | +
| 65/67 | +Clapser outer length / clapser base width 7.2 8.6 7.2 | +10 | +4.8 4.8 | +4.9 | +4.2 3 4.9 | +
|
+ |
+
+ |
+
+ |
+|
|---|---|---|---|
| Holotype 10 Paratypes | +4 types NW Australia | +2 specimens Indonesia | +|
| Monospondylous centra | +38 35–38 | +33 - 36 | +34–37 | +
| Precaudal centra | +85 81–86 | +74–78 | +78–79 | +
| Total number of centra | +? 134–144 | +130–134 | +128–131 | +
| Tooth rows in upper jaw | +60 57–60 | +54–57 | +56 | +
| Tooth rows in lower jaw | +60 55–64 | +54–62 | +
| Variable | +Males (n = 9) | +Females (n = 7) | +
|---|---|---|
| SVL Head length | +52.8 ± 6.4(42.7–61.7) 12.5 ± 1.3(10.8–15.0) | +55.3 ± 3.3(50.82–59.9) 12.6 ± 0.5(12.1–13.3) | +
| Head width | +10.8 ± 0.9(9.4–12.4) | +10.7 ± 0.53(10.0–11.6) | +
| Arm length Tibial length | +15.9 ± 1.8(13.4–17.9) 11.1 ± 1.10(9.3–12.3) | +16.4 ± 0.44(15.5–16.9) 10.9 ± 0.48(10.0–11.5) | +
| Foot length | +19.1 ± 1.3(18.0–22.0) | +16.0 ± 0.63(15.0–16.8) | +
| Axilla-groin distance Midbody scales | +21.6 ± 3.2(16.3–27.2) 67.2 ± 2.4(64–71) | +24.7 ± 2.3(21.5–27.9) 67.1 ± 1.0(66–69) | +
| Dorsal scales | +84.7 ± 3.7(78–90) | +83.8 ± 2.4(82–88) | +
| Ventral scales Pre-cloacal pores | +78.5 ± 4.5(74–89) 7–9 | +81.4 ± 3.4(77–87) 0 | +
| Infradigital lamellae (3rd to hand) Infradigital lamellae (4th to foot) | +18.4 ± 0.7 (17–19) 23.2 ± 1.5 (22–26) | +17.6 ± 0.8 (17–19) 22.7 ± 1.1(21–24) | +
| Supralabial scales Infralabial scales | +8–10 7–10 | +9–11 8–10 | +
|
+ |
+56 | +60 | +60–66 | +57–64 | +3–4 | +4–7 | +no | +evident | +yes? | +
+Verrastro
+ |
+
|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+48.7 | +48.3 | +32–42 | +35–37 | +0–3 | +5–6 | +yes | +evident | +yes | +Avila 2003, this study | +
|
+ |
+59.9 | +61.7 | +64–71 | +78–90 | +0 | +7–9 | +no | +absent | +yes | +This study | +
|
+ |
+69 | +84 | +56–74 | +54–75 | +0 | +6 | +no | +absent | +no | +Etheridge, 2000 | +
|
+ |
+64 | +72 | +68–92 | +79–118 | +0 | +5–11 | +yes | +absent | +yes | +Etheridge, 2000 | +
|
+ |
+60 | +70 | +67–79 | +75–86 | +0 | +7–10 | +no | +absent | +no | +
+Verrastro
+ |
+
|
+ |
+68 | +61 | +73–81 | +92–105 | +0 | +5–9 | +yes | +absent | +yes | +Cei, 1974; Etheridge, 2000 | +
|
+ |
+57 | +63 | +61–79 | +72–93 | +0–6 | +6–11 | +yes | +absent | +yes | +
+Etheridge, 2000; Laspiur
+ |
+
|
+ |
+68 | +76 | +58–77 | +69–97 | +0 | +6–10 | +yes | +absent | +yes | +Etheridge, 2000 | +
|
+ |
+65 | +77 | +49–68 | +58–78 | +0–6 | +5–10 | +yes | +absent | +yes | +Etheridge, 2000 | +
|
+ |
+60 | +59.1 | +38–58 | +43–67 | +0–6 | +4–8 | +yes | +strong | +yes | +This study | +
| Taxon 1 | +Taxon 2 | +Distance in % | +
|---|---|---|
|
+ |
+
+ |
+1.23 | +
|
+ |
+
+ |
+1.51 | +
|
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+2.84 | +
|
+ |
+
+ |
+4.03 | +
|
+ |
+
+ |
+4.20 | +
|
+ |
+
+ |
+4.52 | +
|
+ |
+
+ |
+4.81 | +
|
+ |
+
+ |
+9.26 | +
|
+ |
+
+ |
+9.26 | +
|
+ |
+
+ |
+9.51 | +
|
+ |
+
+ |
+9.51 | +
|
+ |
+
+ |
+9.75 | +
|
+ |
+
+ |
+9.75 | +
|
+ |
+
+ |
+9.75 | +
|
+ |
+
+ |
+9.88 | +
|
+ |
+
+ |
+9.88 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| SVL | +||||||
| TL | +||||||
| OT | +||||||
| TrK | +||||||
| HW | +||||||
| HH | +||||||
| HL | +||||||
| EN | +||||||
| IN | +||||||
| EYE | +||||||
| EAR | +||||||
| FA | +||||||
| CS | +
| Diagnostic characters | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|
| corniculi developed | +yes (mostly), Figs 1, 3, 21, 22 | +yes | +not | +not (mostly) | +
| index of length of scape to pedicel | +1.6–3.2, Figs 4, 23 | +1.5–3.3 | +2.3–3.5 | +3.1–4.7 | +
| shape of pedicel | +globular, Figs 4, 23 | +not globular, asymmetrical | +globular | +globular | +
| first flagellomere put under rigid bristles | +not, Figs 4, 23 | +not | +yes | +not | +
| index of wing length to width | +2.4–3.3, Figs 18, 35 | +2.3–2.8 | +2.4–2.8 | +2.4–2.7 | +
| medial wing angle | +90–180°, Figs 18, 35 | +180–222° | +146–210° | +122–180° | +
| ejaculatory apodeme (basiphallus) flattened | +in dorsal view, Figs 14, 38 | +in dorsal view | +in dorsal view | +in lateral view | +
|
+sabre- or sickle- shaped outer paired convergent protubernces (phallomeres) of aedeagal
+ |
+not developed, Figs 14, 36, 38 | +developed | +developed | +not developed | +
|
+glenoid blades (paired appendages of distiphallus) of aedeagal
+ |
+full or partially developed, Figs 14, 36, 38 | +not developed | +not developed | +not developed or in a very reduced form | +
|
+lance- or dagger- shaped inner parallel and divergent protuberances (phallomeres) of aedeagal
+ |
+not developed, Figs 14, 36, 38 | +not developed | +not developed | +full or partially developed | +
| furca developed | +yes, Figs 14, 17, 34, 36, 38 | +not | +not | +yes | +
| number of retinaculi | +7–14, Figs 19, 20, 37, 42 | +5–35 | +5–12 | +17–30 | +
|
+ |
+Ježek sp. nov. | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+Lanceolate, linear- lanceolate with protracted round to capitate apices | +22–42 | +2.5–5 | +12–14 | +Round to rectangular | +N-CA | +~50 | +Rounded; covered by the hymen | +Current study | +
|
+ |
+Linear (longest valve); lanceolate to elliptical (Small valves); Protracted towards the subcapitate ends | +8–45 | +2.5–4.5 | +9–12 | +Not equidistance, Irregularly distributed | +CA | +23–27 | +-- | +
+ |
+
|
+ |
+Lanceolate with rostrate to subcapitate ends | +7–46 | +2.5–5.5 | +10–12 | +Wider separation between 2 central fibulae | +CA | +26–30 | +-- | +
+ |
+
|
+ |
+Linear-lanceolate to linear with protracted rostrate to subcapitate ends | +10.3– 48.6 | +2.8–4 | +12–17 | +Irregularly distributed and connected with one or two transapical costae | +CA | +27–35 | +Rounded to slightly elongated | +
+ |
+
|
+ |
+Narrowly lanceolate with weakly protracted ends | +14.5– 44.6 | +3–3.7 | +8–10 | +Irregularly distributed along the keel | +CA | +23–25 | +-- | +
+ |
+
|
+ |
+Valves elliptical, lanceolate, linear- lanceolate to linear | +3–60 | +2–4.5 | +8–16 | +Block shaped | +CA | +19–32 | +-- | +
+ |
+
|
+ |
+Valves lanceolate, linear lanceolate to linear | +5–40 | +2–4.5 | +7–10 | +Block shaped | +N-CA | +20 | +-- | +
+ |
+
|
+4.2.2 -
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+163100 | +
| = | +
+ |
+
+ |
+1838 | +
| = | +
+ |
+
+ |
+1779 | +
| = | +
+ |
+
+ |
+1779 | +
| = | +
+ |
+
+ |
+163100 | +
| = | +
+ |
+
+ |
+2713 | +
| = | +
+ |
+
+ |
+2201 | +
| = | +
+ |
+
+ |
+163100 | +
| = | +
+ |
+
+ |
+164100 | +
| = | +
+ |
+
+ |
+1535 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +
+ |
+|
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
| Westliche Zentralalpen (WA) | +
+potenziell
+ |
+B2b(iii) | +
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+
+in
+ |
+
+ |
+(Caporiacco), | +
+ |
+sp. nov., | +
|---|---|---|---|---|---|
| Fe Pt | +Ti | +Mt | +Ta | +Total | +|
|
+ |
+|||||
| Male | +|||||
| I 2.15 0.95 | +1.80 | +1.90 | +1.40 | +8.20 | +|
| II 2.10 0.90 | +1.60 | +1.85 | +1.30 | +7.75 | +|
| III 2.00 0.85 | +1.50 | +2.00 | +1.15 | +7.50 | +|
| IV 2.70 1.00 | +2.20 | +3.40 | +1.55 | +10.85 | +|
| Female | +|||||
| I 2.60 1.15 | +2.10 | +2.00 | +1.45 | +9.30 | +|
| II 2.50 1.10 | +1.90 | +1.95 | +1.35 | +8.80 | +|
| III 2.35 1.05 | +1.75 | +2.20 | +1.20 | +8.55 | +|
| IV 3.20 1.20 | +2.65 | +4.00 | +1.75 | +12.80 | +|
|
+ |
+|||||
| Male | +|||||
| I 2.40 1.10 | +1.95 | +2.15 | +1.35 | +8.95 | +|
| II 2.30 1.05 | +1.70 | +2.05 | +1.20 | +8.30 | +|
| III 2.25 1.00 | +1.55 | +2.20 | +1.05 | +8.05 | +|
| IV 2.90 1.15 | +2.25 | +3.45 | +1.45 | +11.20 | +|
| Female | +|||||
| I 2.75 1.30 | +2.15 | +2.20 | +1.40 | +9.80 | +|
| II 2.65 1.25 | +1.90 | +2.10 | +1.35 | +9.25 | +|
| III 2.50 1.15 | +1.75 | +2.35 | +1.20 | +8.95 | +|
| IV 3.40 1.35 | +2.50 | +3.80 | +1.60 | +12.65 | +|
|
+ |
+|||||
| Male | +|||||
| I 2.30 1.05 | +1.90 | +2.10 | +1.40 | +8.75 | +|
| II 2.25 1.00 | +1.70 | +2.00 | +1.30 | +8.25 | +|
| III 2.15 0.95 | +1.55 | +2.20 | +1.20 | +8.05 | +|
| IV 2.90 1.10 | +2.30 | +3.60 | +1.60 | +11.50 | +|
| Female | +|||||
| I 2.70 1.25 | +2.15 | +2.15 | +1.50 | +9.75 | +|
| II 2.65 1.20 | +1.95 | +2.10 | +1.45 | +9.35 | +|
| III 2.50 1.10 | +1.85 | +2.40 | +1.35 | +9.20 | +|
| IV 3.40 1.30 | +2.75 | +4.30 | +1.85 | +13.60 | +|
|
+ |
+|||||
| Female | +|||||
| I 1.95 0.95 | +1.55 | +1.50 | +1.10 | +7.05 | +|
| II 1.90 0.90 | +1.40 | +1.50 | +1.05 | +6.75 | +|
| III 1.80 0.90 | +1.30 | +1.65 | +0.95 | +6.60 | +|
| IV 2.40 1.00 | +2.00 | +2.90 | +1.30 | +9.60 | +
|
+ |
+
+ |
+|
|---|---|---|
| Dorsal surface of carapace | +Regions gently convex; postorbital region elevated not sunken (Figs. 1B, C, 3A, B, 4A, 8A, B); epigastric, mesogastric and postorbital cristae prominent, lined with numerous granules (Figs. 1A, 2, 8A, B); posterolateral and posterior carapace regions without transverse submarginal grooves (Figs. 1A, 2, 8A, B) | +Regions almost flat; postorbital region prominently sunken (Fig. 5C); epigastric, mesogastric and postorbital cristae short, not distinct, lined only with scattered granules (Figs. 5A, 6A, 8C, D); posterolateral and posterior carapace regions with distinct transverse submarginal grooves which connect to grooves in sub-branchial region (Figs. 5A, 6A, D, 8C, D) | +
| Anterolateral margin | +First and second teeth not prominently flattened dorsoventrally, with short, low, median longitudinal ridge (Figs. 1A, 2, 4A, 8A, B) | +First and second teeth prominently flattened dorsoventrally, appears foliaceous, without discernible longitudinal ridge (Figs. 5A, 6A, 8C, D) | +
| Frontal and orbital regions | +Inner angle of supraorbital margin low; postorbital region not prominently sunken in (Figs. 1A, B, 2, 3A, 4A, 8A, B) | +Inner angle of supraorbital margin prominent; postorbital region prominently sunken in (Figs. 5A, C, 6A, 8C, D) | +
| Sub-branchial region | +With indistinct groups of low granules not arranged into ridges (Fig. 9A) | +With numerous medially eroded granules distinctly arranged into ridges to form grooves (Fig. 9B) | +
| Chelipeds | +Surfaces smooth or with scattered granules, without ridges, pits or eroded depressions; dorsal margin of chela rounded without ridges although shallow longitudinal subdorsal groove may be present; merus without submedian or subdistal teeth; inner distal tooth of carpus low, dentiform; dense setae evenly covering and obscuring almost all surfaces except for outer and inner surfaces of palm and fingers; outer surface of larger palm of both sexes almost completely glabrous (Figs. 2, 3C, D). | +Surfaces with eroded depressions of varying sizes, setae covers all surfaces except for ridges; dorsal margin of chela with 2 prominent ridges; carpus and merus with prominent ridges; merus with a prominent sharp submedian tooth and a subdistal tooth; inner distal tooth of carpus large, lobiform; setae dense but covering only flattened and depressed areas of articles, ridges glabrous, clearly visible; on chela, only fingers and ridges glabrous; outer surfaces of palms of both sexes never completely glabrous (Figs. 5, 6E) | +
| Ambulatory legs | +Surfaces smooth or with scattered low granules; articles not crested, without ridges; merus of first to third legs with low subdistal tooth; dense setae evenly covering and obscuring almost all surfaces (Figs. 2, 10A, B) | +Surfaces with eroded depressions of varying sizes; merus, carpus and proposed with dorsal crest; merus of first to third legs with prominent subdorsal ridge, subdistal tooth large; carpus and propodus with prominent submedian ridges; setae dense but covering only flattened and depressed areas of articles, ridges glabrous, clearly visible (Figs. 5, 6F, G) | +
| Surfaces of anterior thoracic sternum and abdomen | +With scattered granules but without depressions (Figs. 1C, 3B) | +With numerous eroded depressions of varying sizes (Figs. 5B, 6H, 7A) | +
| G1 | +Relatively short, stout with tip prominently dilated, appearing flared (Fig. 4C, D) | +Relatively slender, longer, distal part tapering to sharp tip (Fig. 7B-D) | +
|
+ |
+fig. | +C. | +— | +Lanchester | +1900: | +743. | +— | +Johnson | +1963: | +288. | +— | +Goh | +
+ |
+
+ |
+
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 1990: 30, 37. — Davie 2002: 416. | +||||||||||||||
|
+“
+ |
+||||||||||||||
|
+ |
+
| Species and subjective synonyms | +Original combination | +Type locality | +
|---|---|---|
|
+31 species previously assigned to
+ |
+||
|
+ |
+
+ |
+between Brittany and Ireland, Atlantic Ocean | +
|
+ |
+
+ |
+South Shetland Islands, off Antarctic Peninsula | +
|
+ |
+
+ |
+off Sand Key, Florida, Atlantic Ocean | +
|
+ |
+
+ |
+western parts of the Bering Sea | +
|
+ |
+
+ |
+Brazil, Atlantic Ocean | +
|
+ |
+
+ |
+Taiwan, Pacific Ocean | +
|
+ |
+
+ |
+Singapore and Philippines, Pacific Ocean | +
|
+ |
+
+ |
+Mediterranean Sea | +
|
+ |
+Madeira, Atlantic Ocean | +|
|
+ |
+Cannes, France, Atlantic Ocean | +|
|
+ |
+Gulf of Naples, Mediterranean Sea | +|
|
+ |
+Philippines, Pacific Ocean | +|
|
+ |
+Gulf of Naples, Mediterranean Sea | +|
|
+ |
+
+ |
+Cameroon, Africa, Atlantic Ocean | +
|
+ |
+
+ |
+Loyalty Islands, New Caledonia, Pacific Ocean | +
|
+ |
+
+ |
+The Bismarck Archipelago, Pacific Ocean | +
|
+ |
+
+ |
+Tuamotu Islands, Pacific Ocean | +
|
+ |
+
+ |
+Ganjam Coast, India | +
|
+ |
+
+ |
+Okhotsk Sea | +
|
+ |
+
+ |
+Mediterranean Sea | +
|
+ |
+
+ |
+Amboina in Indonesia | +
|
+ |
+
+ |
+Virginia, Atlantic Ocean | +
|
+ |
+
+ |
+Keiskama Estuary, South Africa | +
|
+ |
+
+ |
+Indonesia, Pacific Ocean | +
|
+ |
+
+ |
+Philippines, Pacific Ocean | +
|
+ |
+
+ |
+Galapagos, Pacific Ocean | +
|
+ |
+
+ |
+Central Japan, Pacific Ocean | +
|
+ |
+
+ |
+north of the Aleutian Islands, Bering Sea | +
|
+ |
+
+ |
+Hawaii, Pacific Ocean | +
|
+ |
+
+ |
+West Africa, Atlantic Ocean | +
|
+ |
+
+ |
+Nova Scotia, eastern Canada, Atlantic Ocean | +
|
+ |
+
+ |
+California, Pacific Ocean | +
|
+ |
+
+ |
+off western Mexico, Pacific Ocean | +
|
+ |
+
+ |
+Florida Keys, Caribbean Sea, Atlantic Ocean | +
|
+ |
+
+ |
+Madeira, Atlantic Ocean | +
|
+ |
+
+ |
+Hawaii, Pacific Ocean | +
| Three species added by the present study | +||
|
+ |
+
+ |
+Malay Archipelago, Pacific Ocean | +
|
+ |
+
+ |
+Ryukyu Islands, southern Japan, Pacific Ocean | +
|
+ |
+
+ |
+Ryukyu Islands, southern Japan, Pacific Ocean | +
| Colony | +Head length to end of postclypeus | +Postclypeal length | +Max. head width | +Pronotal width | +Hind tibia length | +Fore-tibia width: length ratio | +
|---|---|---|---|---|---|---|
| Holotype | +||||||
| GU105 | +||||||
| GU106 | +||||||
| GU753 | +||||||
| GU754 | +||||||
| GU783 | +||||||
| GU784 | +||||||
| GU785 | +||||||
| GU786 | +||||||
| GU787 | +||||||
| GU788 | +||||||
| PU505 | +||||||
| Range (n=132) | +
| Femur | +Patella | +Tibia | +Metatarsus | +Tarsus | +Total | +|
|---|---|---|---|---|---|---|
| Palp | +2.61 (3.19) | +1.41 (1.78) | +1.83 (2.36) | +— | +0.72 (2.25) | +6.57 (9.58) | +
| Leg I | +4.13 (4.31) | +2.26 (2.77) | +3.36 (3.41) | +3.27 (2.66) | +1.99 (2.04) | +15.01 (15.19) | +
| Leg II Leg III Leg IV | +3.81 (4.13) 3.63 (3.46) 4.71 (4.62) | +2.03 (2.47) 1.71 (1.89) 2.14 (2.50) | +3.02 (2.88) 2.58 (2.45) 3.67 (3.59) | +3.07 (2.70) 3.73 (3.02) 5.18 (4.66) | +1.94 (2.03) 2.03 (2.03) 2.65 (2.25) | +13.87 (14.21) 13.68 (12.85) 18.35 (17.62) | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +1 | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+sehr
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Character | +
+ |
+
+ |
+||
|---|---|---|---|---|
|
+This study
+ |
+
+ |
+
+This study
+ |
+
+ |
+|
| Dorsal-fin spines | +XII | +XII | +XII | +XII-XIII | +
| Dorsal-fin soft rays | +21 | +21 | +14 | +14 | +
| Pectoral-fin soft rays | +17 | +17 | +17 | +- | +
| Pelvic-fin spines | +I | +I | +I | +- | +
| Pelvic-fin soft rays | +5 | +5 | +5 | +- | +
| Anal-fin spines | +III | +III | +III | +III | +
| Anal-fin soft rays | +8 | +8 | +7 | +7-9 | +
| Caudal-fin rays | +18 | +17 | +17-20 | +- | +
| Gill rakers | +5 + 15 | +- | +- | +- | +
| Measurements | +
+ |
+
+ |
+
|---|---|---|
| Total length [mm] | +335 | +148-197 | +
| Standard length [mm] | +300 | +116-159 | +
|
+ |
+||
| Body depth | +40.0 | +42.70-49.10 | +
| Head length | +26.6 | +32.70-36.80 | +
| Inter orbital wide | +9.0 | +7.70-8.20 | +
| Pre orbital length | +8.3 | +9.40-10.60 | +
| Post orbital length | +12.6 | +13.80-14.70 | +
| Eye diameter | +7.3 | +10.06-11.48 | +
| Snout length | +9.3 | +44.03-60.34 | +
| Caudal peduncle length | +14.0 | +12.30-12.93 | +
| Dorsal-fin base length | +60.6 | +55.35-61.48 | +
| largest 5th dorsal-fin length | +11.0 | +16.98-18.03 | +
| Pectoral-fin base length | +7.0 | +- | +
| Pectoral-fin length | +20.6 | +29.56-33.61 | +
| Pelvic-fin base length | +19.6 | +5.03-5.74 | +
| Pelvic-fin length | +20.3 | +22.64-27.05 | +
| Anal-fin base length | +11.3 | +15.09-14.75 | +
| Anal-fin length | +14.3 | +22.41-22.13 | +
| Caudal-fin base length | +13.0 | +11.95-13.11 | +
| Caudal-fin length | +20.3 | +30.17-30.30 | +
| Pre dorsal length | +35.0 | +- | +
| Pre pectoral length | +27.6 | +- | +
| Pre pelvic length | +32.3 | +- | +
| Pre anal length | +60.6 | +- | +
| - | +USNM 442696 | +USNM 442071 | +
+CIUFES 2414 (
+ |
+All types | +
|---|---|---|---|---|
| SL | +19.5 | +28.6 | +22.4 | +16.4-22.4 | +
| Sex | +male | +female | +male | +
+ |
+
| Morphometrics in % SL | +||||
| Eye diameter | +9.2 | +8.4 | +8.7 | +8.6-9.15 | +
| Jaw length | +9.7 | +10.1 | +9.6 | +9.6-11.5 | +
| Snout length | +6.2 | +5.6 | +5.8 | +5.2-7.0 | +
| Head length | +23.6 | +29.7 | +27.9 | +27.9-29.5 | +
| Postorbital length | +
+ |
+14 | +13.5 | +13.5-16.6 | +
| Depth at first dorsal-fin origin | +17.4 | +18.5 | +16.3 | +15.8-16.5 | +
| Depth at anal-fin origin | +17.4 | +17.1 | +17.3 | +14.6-17.3 | +
| Least caudal peduncle depth | +
+ |
+11.2 | +12.5 | +10.4-12.5 | +
| Caudal peduncle length | +
+ |
+17.5 | +22.9 | +18.8-23.0 | +
| Caudal-fin length | +23.4 | +22.21 | +26.3 | +26.3-27.8 | +
| Pectoral-fin length | +
+ |
+26.2 | +20.9 | +19.8-25.0 | +
| First dorsal-fin elements | +VII | +VII | +VII | +VII | +
| Second dorsal-fin elements | +
+ |
+
+ |
+I,10 | +I,10 | +
| Anal-fin elements | +I,10 | +I,10 | +I,10 | +I,10 | +
| Pectoral-fin rays | +19 | +19 | +18/19 | +18-19 | +
| Caudal-fin rays segmented, branched | +
+17,
+ |
+17,14 | +17;14 | +17;14 | +
| Pelvic-fin elements | +I,V | +I,V | +I,V | +I,V | +
| Length of fifth pelvic-fin ray relative to fourth | +
+ |
+1/2 | +1/2 | +1/2 | +
| Vertebrae - precaudal + caudal | +? | +11+16 | +11+16 | +11+16 | +
| Dorsal-fin pterygiophore pattern | +? | +3-221110 | +3-221110 | +3-221110 | +
| Anal-fin pterygiophores inserted before haemal arch | +? | +2 | +2 | +2 | +
| Pelvic fins 1-4 | +branched, no fleshy tips | +branched, no fleshy tips | +branched, no fleshy tips | +branched, no fleshy tips | +
| Lateral scale rows | +43 | +47 | +40-45* | +40-47 | +
| Tranverse scale rows | +11 | +14 | +10** | +10-13** | +
| URCA-H | +SVL | +TL | +Body annuli | +Tail annuli | +Segments Pores | +SL | +IL | +Sex | +HW | +BW | +
|---|---|---|---|---|---|---|---|---|---|---|
| 3540* | +263 | +41.2 | +257 | +34 | +20/21 6 | +3 | +3 | +F | +4.9 | +6.7 | +
| 3543 | +231 | +70 | +264 | +6/autotomy | +21/24 6 | +3 | +3 | +F | +5 | +7 | +
| 3541 | +252 | +34.8 | +256 | +30 | +20/21 6 | +3 | +3 | +M | +5 | +6.5 | +
| 3542 | +258 | +254 | +6/autotomy | +20/24 6 | +3 | +3 | +M | +4.8 | +7.1 | +|
| 3544 | +233 | +256 | +6/autotomy | +21/23 6 | +3 | +3 | +M | +4.2 | +5.9 | +|
| 3546 | +244 | +252 | +6/autotomy | +20/21 6 | +3 | +3 | +? | +5.2 | +6.6 | +|
| 3552 | +252 | +11.5 | +258 | +6/autotomy | +22/23 6 | +3 | +3 | +M | +4.8 | +7.3 | +
| 3551 | +278 | +252 | +6/autotomy | +20/21 6 | +3 | +3 | +F | +5.8 | +7.6 | +
| SL | +IL | +Sulci | +AS | +PR | +Dorsal Coloration | +|
|---|---|---|---|---|---|---|
|
+ |
+4 | +3 | +50 | +– | +yes | +Dark brown | +
|
+ |
+3 | +3–4 | +? | +4–6 | +? | +Checkered black and white | +
|
+ |
+4 | +3 | +48 | +6 | +yes | +Brown with light sutures | +
|
+ |
+3 | +3 | +42 | +6 | +yes | +Creamy with center of scales dark brown | +
|
+ |
+3 | +3 | +? | +6–10 | +no | +Reddish brown | +
|
+ |
+3 | +3 | +40 | +5–7 | +yes | +Dark brown | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| First chamber length | +431.2 | +92.32 | +―― | +++ | +++ | +0 | +
| Proloculus nominal diameter | +192.8 | +44.16 | +0 | +++ | +++ | +―― | +
| Deuteroloculus ratio | +1.080 | +0.0797 | +- | +0 | +0 | +0 | +
| Initial marginal radius | +255.3 | +51.51 | +0 | +++ | +++ | +―― | +
| Marginal radius increase | +0.113 | +0.0132 | +- | +―― | +0 | +++ | +
| Spiral chamber height increase | +3.7 | +0.68 | +0 | +++ | +++ | +0 | +
| Initial spiral chamber height | +77.1 | +23.95 | +0 | +++ | +++ | +- | +
| Backbend angle | +0.563 | +0.1055 | +0 | +―― | +0 | +++ | +
| Initial chamber length | +287.2 | +78.67 | +- | +++ | +++ | +0 | +
| Chamber length increase | +0.033 | +0.0117 | +0 | +―― | +0 | +++ | +
| Perimeter ratio | +1.302 | +0.0758 | +0 | +―― | +0 | +++ | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| First chamber length | +655.5 | +116.47 | +++ | +++ | +++ | +++ | +
| Proloculus nominal diameter | +213.6 | +43.52 | +0 | +++ | +++ | +―― | +
| Deuteroloculus ratio | +1.317 | +0.1965 | ++ | +0 | +++ | ++ | +
| Initial marginal radius | +292.5 | +50.03 | +0 | +++ | +++ | +0 | +
| Marginal radius increase | +0.136 | +0.0151 | ++ | +0 | +++ | +++ | +
| Spiral chamber height increase | +3.2 | +1.35 | +0 | +0 | +0 | +0 | +
| Initial spiral chamber height | +93.0 | +33.38 | +0 | +++ | +++ | +0 | +
| Backbend angle | +0.632 | +0.0866 | +0 | +0 | ++ | +++ | +
| Initial chamber length | +434.4 | +104.66 | ++ | +++ | +++ | +++ | +
| Chamber length increase | +0.042 | +0.0187 | +0 | +0 | +0 | +++ | +
| Perimeter ratio | +1.435 | +0.1299 | +0 | +0 | ++ | +++ | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
|---|---|---|---|---|
| First chamber length | +369.0 | +165.69 | +++ | +0 | +
| Proloculus nominal diameter | +146.3 | +67.08 | +0 | +―― | +
| Deuteroloculus ratio | +1.166 | +0.1684 | +++ | +++ | +
| Initial marginal radius | +192.0 | +88.59 | +0 | +―― | +
| Marginal radius increase | +0.126 | +0.0191 | +0 | +++ | +
| Spiral chamber height increase | +2.8 | +1.15 | +++ | +―― | +
| Initial spiral chamber height | +62.5 | +32.84 | +0 | +―― | +
| Backbend angle | +0.636 | +0.1120 | +0 | +++ | +
| Initial chamber length | +224.0 | +126.60 | +++ | +0 | +
| Chamber length increase | +0.048 | +0.0220 | +0 | +++ | +
| Perimeter ratio | +1.392 | +0.1130 | +0 | +++ | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
|---|---|---|---|---|
| First chamber length | +243.3 | +38.59 | +―― | +―― | +
| Proloculus nominal diameter | +113.0 | +25.84 | +0 | +―― | +
| Deuteroloculus ratio | +0.888 | +0.1258 | +―― | +0 | +
| Initial marginal radius | +140.1 | +28.76 | +0 | +―― | +
| Marginal radius increase | +0.094 | +0.0195 | +0 | +0 | +
| Spiral chamber height increase | +1.7 | +0.07 | +―― | +―― | +
| Initial spiral chamber height | +46.5 | +10.02 | +0 | +―― | +
| Backbend angle | +0.495 | +0.1320 | +0 | ++ | +
| Initial chamber length | +139.0 | +17.17 | +―― | +―― | +
| Chamber length increase | +0.028 | +0.0175 | +0 | +0 | +
| Perimeter ratio | +1.277 | +0.1076 | +0 | +0 | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| First chamber length | +218.2 | +53.81 | +―― | +―― | +0 | +―― | +
| Proloculus nominal diameter | +99.0 | +28.91 | +―― | +―― | +0 | +―― | +
| Deuteroloculus ratio | +0.950 | +0.1376 | +0 | +―― | +- | +0 | +
| Initial marginal radius | +126.8 | +32.48 | +―― | +―― | +0 | +―― | +
| Marginal radius increase | +0.099 | +0.0172 | +0 | +―― | +―― | +++ | +
| Spiral chamber height increase | +1.7 | +0.30 | +―― | +0 | +0 | +―― | +
| Initial spiral chamber height | +35.5 | +17.71 | +―― | +―― | +0 | +―― | +
| Backbend angle | +0.445 | +0.1165 | +0 | +- | +―― | +++ | +
| Initial chamber length | +151.9 | +26.76 | +―― | +―― | +0 | +―― | +
| Chamber length increase | +0.025 | +0.0149 | +0 | +0 | +―― | +0 | +
| Perimeter ratio | +1.263 | +0.0859 | +0 | +- | +―― | ++ | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| First chamber length | +252.0 | +92.62 | +―― | +―― | +0 | +―― | +
| Proloculus nominal diameter | +92.2 | +29.93 | +―― | +―― | +0 | +―― | +
| Deuteroloculus ratio | +1.191 | +0.1834 | +0 | +0 | ++ | +++ | +
| Initial marginal radius | +116.9 | +44.90 | +―― | +―― | +0 | +―― | +
| Marginal radius increase | +0.133 | +0.0193 | +++ | +0 | +++ | +++ | +
| Spiral chamber height increase | +2.0 | +0.96 | +―― | +0 | +0 | +―― | +
| Initial spiral chamber height | +38.5 | +15.47 | +―― | +―― | +0 | +―― | +
| Backbend angle | +0.687 | +0.1008 | +++ | +0 | +++ | +++ | +
| Initial chamber length | +129.1 | +43.24 | +―― | +―― | +0 | +―― | +
| Chamber length increase | +0.060 | +0.0202 | +++ | +0 | +++ | +++ | +
| Perimeter ratio | +1.448 | +0.0809 | +++ | +0 | +++ | +++ | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
|---|---|---|---|---|
| First chamber length | +448.7 | +220.50 | +0 | +++ | +
| Proloculus nominal diameter | +334.2 | +152.84 | +++ | +++ | +
| Deuteroloculus ratio | +0.999 | +0.1297 | +―― | +0 | +
| Initial marginal radius | +355.9 | +143.51 | +++ | +++ | +
| Marginal radius increase | +0.062 | +0.0081 | +―― | +0 | +
| Spiral chamber height increase | +3.8 | +1.47 | +++ | +++ | +
| Initial spiral chamber height | +111.0 | +58.49 | +++ | +++ | +
| Backbend angle | +0.164 | +0.0513 | +―― | ++ | +
| Initial chamber length | +246.6 | +125.21 | +0 | +++ | +
| Chamber length increase | +0.009 | +0.0065 | +―― | +0 | +
| Perimeter ratio | +1.101 | +0.0595 | +―― | +0 | +
|
+ |
+Mean | +SD | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| First chamber length | +454.7 | +221.33 | +0 | +―― | +++ | +++ | +
| Proloculus nominal diameter | +335.0 | +151.10 | +++ | +++ | +++ | +++ | +
| Deuteroloculus ratio | +1.003 | +0.1318 | +0 | +- | +―― | +0 | +
| Initial marginal radius | +360.2 | +144.63 | +++ | +0 | +++ | +++ | +
| Marginal radius increase | +0.062 | +0.0080 | +―― | +―― | +―― | +―― | +
| Spiral chamber height increase | +3.8 | +1.45 | +0 | +0 | +++ | +++ | +
| Initial spiral chamber height | +112.9 | +59.18 | ++ | +0 | +++ | +++ | +
| Backbend angle | +0.167 | +0.0527 | +―― | +―― | +―― | +―― | +
| Initial chamber length | +253.2 | +131.05 | +0 | +―― | +++ | +++ | +
| Chamber length increase | +0.009 | +0.0064 | +―― | +―― | +―― | +0 | +
| Perimeter ratio | +1.102 | +0.0591 | +―― | +―― | +―― | +―― | +
| Peloponnisos | +Central Greece | +Ipiros | +Ionian Is. | +Thessalia | +N Sporadhes | +Makedhonia | +Thracian Is. | +Kikladhes | +Kriti | +
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +||||||||
|
+ |
+X | +
| Species | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| ♂ (n=3) | +♀(n=3) | +♂ (n=2) | +♀(n=3) | +♂ (n=2) | +♂ (n=2) | +♀(n=1) | +♂ (n=1) | +♀(n=1) | +♂ (n=6) | +♀(n=2) | +♂ (n=2) | +♀(n=2) | +|
| Body | +18–19,5 | +22–24 | +18,5–19,5 | +20–23 | +15–16 | +18–20 | +20 | +22 | +20 | +20–23,5 | +17,5–19 | +21–22,5 | +19 | +
| Pronotum | +3,5 | +4–4,5 | +4–4,5 | +4 | +3–4 | +4 | +4 | +4,5 | +4 | +4–4,5 | +3,5–4 | +4 | +4,5 | +
| Fore femora. | +16 | +16,5–17,5 | +17 | +16–17 | +16 | +15–16 | +15,5 | +17 | +14 | +15–17 | +15 | +14,5–18 | +16 | +
| Mid femora. | +16–17 | +16–17 | +16,5–17 | +16 | +16,5–18 | +15–16 | +15 | +17,5 | +15 | +14,5–19 | +15–15,5 | +14–18,5 | +15,5 | +
| Hind femora. | +25 | +26–28 | +26–27 | +27 | +26 | +23–23,5 | +25 | +27 | +26 | +24,5–28 | +24–26 | +24,5–29 | +27 | +
| Fore tibia | +18 | +17,5–18,5 | +20 | +18 | +19 | +17–18 | +16,5 | +18 | +16 | +17–20 | +17–17,5 | +17–20 | +17,5 | +
| Mid tibia | +18–19 | +17,5–19 | +20–21 | +18 | +18,5–19 | +18 | +17 | +18,5 | +17 | +17–19 | +17 | +17,5–20 | +17 | +
| Hind tibia | +32,5–34 | +28–35 | +34 | +34 | +31,5–32 | +25–30 | +31 | +31 | +30 | +31–38 | +32–34,5 | +34–36 | +32 | +
| Hind tarsus | +12,5–13 | +13 | +12–12,5 | +12–13 | +12–12,5 | +11–11,5 | +12 | +12 | +11 | +11–13 | +12–13 | +13 | +10 | +
| 1°art. hind tarsus | +6 | +7 | +6 | +6 | +6 | +5–6 | +5,5 | +6,5 | +5 | +6 | +6–7,5 | +5,5 | +|
| Ovipositor | +13–15 | +12–14 | +15 | +19 | +12- 13 | +19 | +|||||||
| Denticles | +18 | +16 | +20 | +20 | +19 | +16 | +|||||||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| 1 | +
+Metafemoral pubescence with horizontal hairline (Fig.
+ |
+2 | +
| - | +
+Metafemoral pubescence extended with oblique hairline, covering basal two-thirds to three-quarters of femur.
+ |
+
+ |
+
| 2 | +
+Median lobe of aedeagus slender, ca. 3 times longer than wide, attenuated towards apex with nearly straight lateral margins (Fig.
+ |
+
+ |
+
| - | +
+Median lobe of aedeagus wide, ca. 2.2-2.5 times longer than wide, triangular and attenuated towards apex in apical part, nearly parallel-sided in basal part (Fig.
+ |
+3 | +
| 3 | +
+Colouration of elytra dark, with yellowish spot anteromesally around scutellar shield (Fig.
+ |
+
+ |
+
| - | +
+Colouration of elytra variable, yellowish to black but elytra without yellowish spot anteromesally around scutellar shield (Fig.
+ |
+
+ |
+
| Position | +1b | +2 a | +3 c | +|||
|---|---|---|---|---|---|---|
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
| 2 | +4.71, | +81.2 | +4.67, d (7.3) | +81.1 | +4.70, | +81.2 | +
| d (7.3) | +d (7.3) | +|||||
| 3 | +3.97, q | +75.4 | +3.93, q (7.3) | +75.2 | +3.96, td | +75.3 | +
| (7.3) | +(5.4, 7.7) | +|||||
| 4 | +2.86, dd | +27.3 | +2.83, dd | +27.3 | +2.86, dd | +27.4 | +
| (5.5, 16.5) | +(5.5, 16.3) | +(5.4, 16.2) | +||||
| 2.75, dd | +2.69, dd | +2.72, dd | +||||
| (7.7, 16.5) | +(7.8, 16.3) | +(5.4, 16.2) | +||||
| 5 | +158.8 | +157.2 | +158.7 | +|||
| 6 | +6.10, s | +96.0 | +6.06, s | +95.9 | +6.09, s | +95.8 | +
| 7 | +157.3 | +155.7 | +157.3 | +|||
| 8 | +95.9 | +95.7 | +95.8 | +|||
| 9 | +155.5 | +155.4 | +157.3 | +|||
| 10 | +100.8 | +100.7 | +100.8 | +|||
| 1′ | +131.6 | +131.5 | +131.6 | +|||
| 2′ | +6.76, | +115.0 | +6.73, d (2.0) | +114.9 | +6.76, | +115.0 | +
| d (2.0) | +d (2.1) | +|||||
| 3′ | +146.2 | +146.1 | +146.2 | +|||
| 4′ | +146.4 | +146.3 | +146.4 | +|||
| 5′ | +6.73, | +116.1 | +6.69, d (8.1) | +115.9 | +6.72, | +116.0 | +
| d (7.8) | +d (8.1) | +|||||
| 6′ | +6.66, dd | +119.6 | +6.62, dd | +119.5 | +6.65, dd | +119.6 | +
| (2.3, 8.3) | +(2.0, 8.1) | +(2.1, 8.1) | +||||
| 1′′ | +115.0 | +114.9 | +114.9 | +|||
| 2′′ | +131.3 | +131.2 | +131.2 | +|||
| 3′′ | +2.78, q | +26.2 | +2.75, q (7.3) | +26.2 | +2.76, p | +26.2 | +
| (7.3) | +(7.0) | +|||||
| 4′′ | +1.11, | +21.5 | +1.07, d (7.1) | +21.4 | +1.10, | +21.5 | +
| d (7.0) | +d (7.0) | +|||||
| 5′′ | +1.13, | +21.6 | +1.10, d (7.1) | +21.5 | +1.12, | +21.6 | +
| d (7.0) | +d (7.0) | +|||||
| C= NH | +11.93d | +147.6 | +11.31 | +147.6 | +11.43 | +147.6 | +
| 1’’’ | +3.81, | +41.9 | +3.77, d (7.0) | +41.8 | +3.34, | +40.4 | +
| d (6.9) | +d (6.9) | +|||||
| 2’’’ | +5.26, t | +120.1 | +5.23, t (7.3) | +120.1 | +1.75, dd | +42.4 | +
| (6.9) | +(6.9, 8.2) | +|||||
| 3’’’ | +138.5 | +138.4 | +70.9 | +|||
| 4’’’ | +1.69, s | +18.0 | +1.66, s | +17.9 | +1.23, | +29.5 | +
| d (5.0) | +||||||
| 5’’’ | +1.73, s | +25.7 | +1.70, s | +25.6 | +1.23, | +29.5 | +
| d (5.0) | +||||||
| 1’’’’ | +4.35, | +102.0 | +4.33, s | +102.0 | +4.35, | +102.0 | +
| d (1.5) | +d (1.7) | +|||||
| 2’’’’ | +3.54, | +72.0 | +3.51, br s | +71.9 | +3.53, q | +72.0 | +
| d (3.4) | +(1.7) | +|||||
| 3’’’’ | +3.57, dd | +72.3 | +3.55, br s | +72.1 | +3.56, | +72.2 | +
| (3.4, 9.4) | +d (9.5) | +|||||
| 4’’’’ | +3.33, | +73.9 | +3.28, | +73.8 | +3.33, | +73.9 | +
| d (9.4) | +d (10.0) | +d (7.4) | +||||
| 5’’’’ | +3.65, dq | +70.3 | +3.62, dt | +70.3 | +3.64, dd | +70.3 | +
| (6.3, 12.4) | +(3.4, 6.1) | +(6.2, 9.5) | +||||
| 6’’’’ | +1.25, | +17.9 | +1.21, d (6.1) | +17.8 | +1.24, | +17.9 | +
| d (6.3) | +d (6.2) | +|||||
| 1′′ -NH | +11.65d | +11.23 | +11.31 | +
| Fig. 11Figs 4, 7 | +cuneata | +
| Fig. 40Figs 52, 58Fig. 33Fig. 56 | +laminata | +
|
+ |
+
+ |
+
| Figs 4, 7 | +|
| Fig. 8Figs 7-8 | +
+ |
+
| Fig. 3Figs 3-4 | +
+ |
+
| Figs 47-48 | +
+ |
+
| Figs 28, 32, 36, 40, 55Figs 3356 | +|
| Figs 14, 19, 24, 43, 47, 60 | +|
| Figs 28, 32 | +|
| Fig. 36 | +|
| Fig. 32 | +
+ |
+
|
+Fig. 28
+ |
+
+ |
+
| Fig. 38 | +
+ |
+
| Figs 41, 56 | +|
| Fig. 42Fig. 42Figs 40, 42 | +
+ |
+
| Fig. 57Fig. 58Fig. 59Figs 55, 59 | +
+ |
+
|
+ |
+
+ |
+
| Figs 20, 45, 64Figs 19, 62 | +|
| Fig. 62Fig. 65Figs 62, 65 | +
+ |
+
| Figs 19, 43, 60Figs 14, 24Figs 17, 23, 46 | +|
| Fig. 19Fig. 20Fig. 23 | +
+ |
+
| Fig. 43Figs 16, 26, 45 | +|
| Fig. 60Fig. 61 | +
+ |
+
| Figs 14, 24, 43Figs 17, 46 | +|
| Fig. 43 | +
+ |
+
| Figs 14, 24Figs 14-15 | +|
| Fig. 14Fig. 14 | +
+ |
+
| Fig. 24Fig. 24 | +
+ |
+
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+63100 | +
| = | +
+ |
+
+ |
+2815 | +
| = | +
+ |
+
+ |
+2996 | +
| = | +
+ |
+
+ |
+2996 | +
| = | +
+ |
+
+ |
+63100 | +
| = | +
+ |
+
+ |
+161 | +
| = | +
+ |
+
+ |
+152 | +
| = | +
+ |
+
+ |
+63100 | +
| = | +
+ |
+
+ |
+2361 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+potenziell
+ |
+B2b(iii) | +
| Mittelland (MP) | +
+potenziell
+ |
+B2b(iii) | +
| Alpennordflanke (NA) | +
+potenziell
+ |
+B2b(iii) | +
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
| Westliche Zentralalpen (WA) | +
+potenziell
+ |
+B2b(iii) | +
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(01.01.2012) | +
|
+ |
+-- | +
|
+ |
+
+ |
+
+ |
+|
|---|---|---|---|
| Standard length (mm) | +23.6–33.4 | +28.6 | +— | +
|
+ |
+|||
| Depth at dorsal-fin origin | +30.3–34.0 | +30.7 | +1.0 | +
| Snout to dorsal-fin origin | +51.8–54.3 | +52.7 | +0.5 | +
| Snout to pectoral-fin origin | +24.6–26.2 | +24.9 | +0.3 | +
| Snout to pelvic-fin origin | +45.9–46.9 | +46.7 | +0.3 | +
| Snout to anal-fin origin | +61.9–63.4 | +62.0 | +0.4 | +
| Caudal-peduncle depth | +10.0–10.8 | +10.6 | +0.2 | +
| Caudal-peduncle length | +11.7–13.1 | +12.3 | +0.3 | +
| Pectoral-fin length | +20.5–25.5 | +23.4 | +1.0 | +
| Pelvic-fin length | +15.2–17.9 | +17.2 | +0.6 | +
| Pelvic-fin origin to anal-fin origin | +16.6–18.2 | +16.8 | +0.5 | +
| Dorsal-fin length | +26.2–30.6 | +29.0 | +0.9 | +
| Dorsal-fin base length | +12.8–14.1 | +13.9 | +0.4 | +
| Anal-fin length | +18.2–21.4 | +20.8 | +0.9 | +
| Anal-fin base length | +28.5–31.0 | +29.7 | +0.5 | +
| Eye to dorsal-fin origin | +37.9–40.3 | +38.1 | +0.6 | +
| Dorsal-fin origin to caudal-fin base | +49.5–52.0 | +51.6 | +0.5 | +
| Head depth | +23.8–25.7 | +25.0 | +0.3 | +
| Head length | +24.1–24.9 | +24.8 | +0.3 | +
|
+ |
+|||
| Horizontal eye diameter | +41.8–46.6 | +45.6 | +1.3 | +
| Snout length | +26.7–28.8 | +28.1 | +0.4 | +
| Interorbital width | +33.8–36.4 | +35.2 | +0.5 | +
| Upper jaw length | +43.3–46.6 | +43.7 | +0.9 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+179200 | +
| = | +
+ |
+
+ |
+179200 | +
| = | +
+ |
+
+ |
+2792 | +
| = | +
+ |
+
+ |
+179200 | +
| 1 | +Large species: body length more than 17.5 mm and forewing length more than 15.0 mm in female; clypeus basally distinctly emarginate; between the notaulices shining apically, not punctate and without carina; metasomal tergum 2 basally with one smooth area | +
+ |
+
| - | +Small species: body length less than 15.0 mm and forewing length less than 14.0 mm in female; clypeus basally indistinctly emarginate; between the notaulices usually not shining, with punctures or carina; metasomal tergum 2 basally without one smooth area | +2 | +
| 2 | +
+Clypeus slightly emarginate apically; anterior slop of metasoma tergum 1 rather gentle; tergum 2 with a very narrow
+ |
+
+ |
+
| - | +
+Clypeus usually widely rounded or truncated apically; anterior slop of metasoma tergum 1 steep; tergum 2 without narrow
+ |
+3 | +
| 3 | +Metasomal terga 2-4 with a reflex apical lamella, respectively | +4 | +
| - | +Metasomal terga 2-3 usually with a lamella, respectively (sometimes just tergum 2 with lamella) | +5 | +
| 4 | +
+Propodeum with lateral ridges and lateral face with superficial and irregular large punctures in the upper portion; clypeus and mesosoma usually entirely black; stigma of forewing brown to dark brown (
+ |
+
+ |
+
| - | +
+Propodeum without lateral ridges and lateral face weakly reticulated; usually two-third of clypeus apicaly yellow and mesosoma with rich yellow markings; stigma of forewing amber yellow (
+ |
+
+ |
+
| 5 | +In female, mandible with four teeth; in male, mandible with three teeth and the apical teeth very long; scutellum and metanotum without obviously median furrow; tegula usually entirely black; the punctures on the metasoma tergum 1 bigger and sparser, sternum 2 without yellow apical band; in male, outer face of fore femur with a broad yellow line | +
+ |
+
| - | +Mandible with two or three teeth in female and three or four teeth in male, the apical teeth not long in both sexes; scutellum and metanotum with median furrow; tegula usually with yellow or brownish colour; the punctures on the metasoma tergum 1 smaller and denser, sternum 2 usually with yellow apical band; in male, outer face of fore femur usually entirely black | +6 | +
| 6 | +In frontal view, face including clypeus flattened in female; frons, clypeus, mesoscutum and scutellum densely with longitudinal carinae in the female; male antennal articles 8-10 distinctly with tyloids covering their lower faces extensively; terminal article stumpy, in profile slightly longer than breadth, and obliquely truncated on apical margin (Kim, 2005) | +7 | +
| - | +
+In frontal view, face including clypeus not flattened in female; frons, clypeus, mesoscutum and scutellum densely with punctures, which only partially form
+ |
+
+ |
+
| 7 | +Vertex without yellow line; mesosoma black, except pronotum with a pair of yellow dots | +
+ |
+
| - | +Vertex with a pair of short oblique yellow line; pronotum, tegula, mesepisternum, scutellum and metanotum with yellow markings, respectively | +
+ |
+
| - | +n | +min-max | +
+ |
+s | +cv | +
+ |
+
|---|---|---|---|---|---|---|
| clmin | +7 | +4.71-6.21 | +5.58 | +0.46 | +8.3 | +5.12-6.04 | +
| clmax | +8 | +5.80-8.44 | +6.86 | +0.89 | +12.9 | +5.97-7.75 | +
| ccd | +8 | +5.46-8.35 | +6.40 | +20.90 | +14.1 | +5.49-7.30 | +
| septa | +9 | +42-56 | +46.6 | +4.67 | +10.0 | +42-51 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+337750 | +
| KEINE ANGABE | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+|
|
+ |
+|
| 241 | +
+ |
+
| 228229230231 | +
+ |
+
| Holotype | +paratypes | +|
|---|---|---|
| male | +males (7) | +females (9) | +
| Percent of standard length | +||
| Percent of head length | +||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+364600 | +
| = | +
+ |
+
+ |
+583 | +
| = | +
+ |
+
+ |
+761 | +
| = | +
+ |
+
+ |
+761 | +
| = | +
+ |
+
+ |
+364600 | +
| = | +
+ |
+
+ |
+804 | +
| = | +
+ |
+
+ |
+707 | +
| = | +
+ |
+
+ |
+364600 | +
| = | +
+ |
+
+ |
+118 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+Teilweise
+ |
+(12.03.1973) | +
|
+ |
+
+Teilweise
+ |
+(01.04.2013) | +
|
+ |
+
+Teilweise
+ |
+(01.07.2009) | +
|
+ |
+-- | +|
|
+ |
+
+Teilweise
+ |
+(03.12.1964) | +
|
+ |
+
+Teilweise
+ |
+(01.10.2017) | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+16550 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +nicht beurteilt (Not Evaluated) | +|
| Mittelland (MP) | +nicht beurteilt (Not Evaluated) | +|
| Alpennordflanke (NA) | +nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
| Westliche Zentralalpen (WA) | +nicht beurteilt (Not Evaluated) | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+sehr
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+
|
+ |
+
| KEINE ANGABE | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+1702 | +
| = | +
+ |
+
+ |
+1702 | +
| KEINE ANGABE | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+2.3 - Feuchtwiesen (
+ |
+
|
+4.5.1 - Talfettwiesen (Fromentalwiese) (
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+117700 | +
| = | +
+ |
+
+ |
+2841 | +
| = | +
+ |
+
+ |
+2421 | +
| = | +
+ |
+
+ |
+2421 | +
| = | +
+ |
+
+ |
+117700 | +
| = | +
+ |
+
+ |
+641 | +
| = | +
+ |
+
+ |
+559 | +
| < | +
+ |
+
+ |
+117750 | +
| = | +
+ |
+
+ |
+117700 | +
| = | +
+ |
+
+ |
+2066 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+potenziell
+ |
+A3c | +
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| KEINE ANGABE | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +frisch | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+236700 | +
| = | +
+ |
+
+ |
+2127 | +
| = | +
+ |
+
+ |
+236700 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Characters | +
+ |
+
+ |
+
|---|---|---|
| Longitudinal stripe on midline from Th. III-Abd. II | +absent | +present# | +
| Pigment on Th. II-III laterally | +present | +absent# | +
| Denticle on labral papilla | +present | +absent# | +
| R chaeta on labial base | +not duplicate | +duplicate# | +
| Medio-sublateral mac on Th. II | +4 (rarely 5) | +2-3* | +
| Mac on Abd. I | +10-11 | +13# | +
| Mac m3ei on Abd. II | +absent | +present# | +
| Mac on Abd. IV centrally | +8 | +5* | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +feucht | +Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +unter-subalpin und ober-montan | +||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||