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<document id="26792399BCBEF199444FC26E3E2C083B" ID-DOI="10.11646/zootaxa.4018.2.7" ID-GBIF-Dataset="a4d79715-1e98-4536-9c94-29069d2d71ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="237549" ID-ZooBank="63B06BDB-DE4C-4B34-8AB5-CE233FC58029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461256482198" checkinUser="plazi" docAuthor="Sites, Robert W. &amp; Reynoso-Velasco, Daniel" docDate="2015" docId="203D141B5221FFCF55885026040AFD8F" docLanguage="en" docName="zt04018p291.pdf" docOrigin="Zootaxa 4018 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ambrysus stali La Rivers" docType="treatment" docVersion="7" lastPageNumber="289" masterDocId="DC046C635229FFC5551F551A0114FFD8" masterDocTitle="Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica" masterLastPageNumber="291" masterPageNumber="279" pageNumber="287" updateTime="1698608226834" updateUser="plazi">
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<mods:title id="D1765C7E5F7C568C98BE232DC306312C">Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica</mods:title>
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<treatment id="203D141B5221FFCF55885026040AFD8F" ID-DOI="http://doi.org/10.5281/zenodo.6119253" ID-GBIF-Taxon="119636079" ID-Zenodo-Dep="6119253" LSID="urn:lsid:plazi:treatment:203D141B5221FFCF55885026040AFD8F" httpUri="http://treatment.plazi.org/id/203D141B5221FFCF55885026040AFD8F" lastPageId="10" lastPageNumber="289" pageId="8" pageNumber="287">
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<heading id="F36312615221FFCD5588502600D7FA8E" bold="true" box="[151,451,1340,1366]" fontSize="11" level="1" pageId="8" pageNumber="287" reason="1">
<taxonomicName id="6F94DE8E5221FFCD5588502600D7FA8E" ID-CoL="7QM66" authority="La Rivers" authorityName="La Rivers" box="[151,451,1340,1366]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5221FFCD5588502600D7FA8E" bold="true" box="[151,451,1340,1366]" pageId="8" pageNumber="287">
<emphasis id="9AE0791F5221FFCD558850260051FA8E" bold="true" box="[151,325,1340,1366]" italics="true" pageId="8" pageNumber="287">Ambrysus stali</emphasis>
La Rivers
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A82BA50D5221FFCD55885045001AFAA0" blockId="8.[151,451,1340,1400]" box="[151,270,1375,1400]" pageId="8" pageNumber="287">
(
<figureCitation id="30AFB9885221FFCD5580504501FEFAAF" box="[159,234,1375,1400]" captionStart="FIGURE 1" captionStartId="2.[151,250,1189,1211]" captionTargetBox="[151,1436,198,1167]" captionTargetId="figure@2.[151,1436,198,1167]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species in the Ambrysus stali species complex." httpUri="https://zenodo.org/record/237550/files/figure.png" pageId="8" pageNumber="287">Figs. 1</figureCitation>
,
<figureCitation id="30AFB9885221FFCD55E750450012FAA0" box="[248,262,1375,1400]" captionStart="FIGURE 6" captionStartId="9.[151,250,1547,1569]" captionTargetBox="[160,1420,304,1503]" captionTargetId="figure@9.[160,1420,304,1504]" captionTargetPageId="9" captionText="FIGURE 6. Structures of Ambrysus stali. (A) dorsal habitus of male, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237555/files/figure.png" pageId="8" pageNumber="287">6</figureCitation>
)
</paragraph>
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<subSubSection id="E08EF6865221FFCD558850BC0223FA03" pageId="8" pageNumber="287" type="reference_group">
<paragraph id="A82BA50D5221FFCD558850BC0223FA03" blockId="8.[151,870,1446,1499]" pageId="8" pageNumber="287">
<taxonomicName id="6F94DE8E5221FFCD558850BC00C7FA64" ID-CoL="7QM66" authority="La Rivers 1962" authorityName="La Rivers" authorityYear="1962" box="[151,467,1446,1468]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5221FFCD558850BC0038FA64" box="[151,300,1446,1468]" italics="true" pageId="8" pageNumber="287">Ambrysus stali</emphasis>
<bibRefCitation id="CC05D8FC5221FFCD542C50BD00C7FA64" author="La" box="[307,467,1446,1468]" pageId="8" pageNumber="287" refString="La Rivers, I. (1962) A new Ambrysus from South America (Hemiptera, Naucoridae). Bulletin of the Southern California Academy of Sciences, 61 (3), 185 - 187." type="journal article" year="1962">La Rivers 1962</bibRefCitation>
</taxonomicName>
: Bull. S. Cal. Acad. Sci. Bull. 185–187.
<taxonomicName id="6F94DE8E5221FFCD558850DF0317FA03" ID-CoL="7QM6X" authority="De Carlo 1968" authorityName="De Carlo" authorityYear="1968" box="[151,515,1477,1499]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="bourquini">
<emphasis id="9AE0791F5221FFCD558850DF0070FA03" box="[151,356,1477,1499]" italics="true" pageId="8" pageNumber="287">Ambrysus bourquini</emphasis>
<bibRefCitation id="CC05D8FC5221FFCD547550DC0317FA03" author="De" box="[362,515,1477,1499]" pageId="8" pageNumber="287" refString="De Carlo, J. A. (1968) Descripcion de especies nuevas de los generos Ambrysus, Cryphocricos y Heleocoris. Allotypus de Cryphocricos daguerrei (Hemiptera - Naucoridae). Revista de la Sociedad Entomologica Argentina, 30, 99 - 104 (1967)." type="journal article" year="1968">
<collectingCountry id="D083E59D5221FFCD547550DC0093FA03" box="[362,391,1478,1499]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="287">De</collectingCountry>
Carlo 1968
</bibRefCitation>
</taxonomicName>
: Rev. Soc. Entomol. Arg. 100.
</paragraph>
</subSubSection>
<subSubSection id="E08EF6865221FFCC5588531D0534FF20" lastPageId="9" lastPageNumber="288" pageId="8" pageNumber="287" type="discussion">
<paragraph id="A82BA50D5221FFCD5588531D05FFF968" blockId="8.[151,1437,1543,2036]" pageId="8" pageNumber="287">
<emphasis id="9AE0791F5221FFCD5588531D0000F9F8" bold="true" box="[151,276,1543,1568]" pageId="8" pageNumber="287">Discussion</emphasis>
.
<bibRefCitation id="CC05D8FC5221FFCD5435531200E3F9F8" box="[298,503,1543,1568]" pageId="8" pageNumber="287" refString="La Rivers, I. (1962) A new Ambrysus from South America (Hemiptera, Naucoridae). Bulletin of the Southern California Academy of Sciences, 61 (3), 185 - 187." type="journal article">La Rivers (1962)</bibRefCitation>
considered this species to be closely related to
<taxonomicName id="6F94DE8E5221FFCD5156531205F8F9F8" box="[1097,1260,1543,1568]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="oblongulus">
<emphasis id="9AE0791F5221FFCD5156531205F8F9F8" box="[1097,1260,1543,1568]" italics="true" pageId="8" pageNumber="287">A. oblongulus</emphasis>
</taxonomicName>
and presented characteristics to distinguish them. The male accessory genitalic process of tergum VI (
<figureCitation id="30AFB9885221FFCD51A9533105EBF99C" box="[1206,1279,1579,1604]" captionStart="FIGURE 6" captionStartId="9.[151,250,1547,1569]" captionTargetBox="[160,1420,304,1503]" captionTargetId="figure@9.[160,1420,304,1504]" captionTargetPageId="9" captionText="FIGURE 6. Structures of Ambrysus stali. (A) dorsal habitus of male, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237555/files/figure.png" pageId="8" pageNumber="287">Fig. 6</figureCitation>
B) is slightly narrower than in other members of the
<taxonomicName id="6F94DE8E5221FFCD5755534A038DF9B0" box="[586,665,1615,1640]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5221FFCD5755534A038DF9B0" box="[586,665,1615,1640]" italics="true" pageId="8" pageNumber="287">A. stali</emphasis>
</taxonomicName>
complex, although intraspecific variation is evident. Also distinctive is that the longitudinal stripes of the pronotum are less linear and more coalescent than in other members of the complex, and the hemelytra are uniformly brown or dark brown without lighter mottling (
<figureCitation id="30AFB9885221FFCD519B538D05DCF968" box="[1156,1224,1687,1712]" captionStart="FIGURE 6" captionStartId="9.[151,250,1547,1569]" captionTargetBox="[160,1420,304,1503]" captionTargetId="figure@9.[160,1420,304,1504]" captionTargetPageId="9" captionText="FIGURE 6. Structures of Ambrysus stali. (A) dorsal habitus of male, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237555/files/figure.png" pageId="8" pageNumber="287">Fig. 6</figureCitation>
A).
</paragraph>
<paragraph id="A82BA50D5221FFCD55D853A1040AF8C4" blockId="8.[151,1437,1543,2036]" pageId="8" pageNumber="287">
Reported here are the first records of
<taxonomicName id="6F94DE8E5221FFCD577953A603A0F90C" box="[614,692,1723,1748]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5221FFCD577953A603A0F90C" box="[614,692,1723,1748]" italics="true" pageId="8" pageNumber="287">A. stali</emphasis>
</taxonomicName>
from
<collectingCountry id="D083E59D5221FFCD57E753A60247F90C" box="[760,851,1724,1748]" name="Guyana" pageId="8" pageNumber="287">Guyana</collectingCountry>
.
<bibRefCitation id="CC05D8FC5221FFCD564253A102E3F90C" author="Nieser" box="[861,1015,1723,1748]" pageId="8" pageNumber="287" refString="Nieser, N. (1975) The water bugs of the Guyana region. Studies on the Fauna of Suriname and Other Guyanas, 59, 1 - 310. http: // dx. doi. org / 10.1007 / 978 - 94 - 017 - 7118 - 4 _ 1" type="journal article" year="1975">Nieser (1975)</bibRefCitation>
indicated that this species is common in
<collectingCountry id="D083E59D5221FFCD55AB53C5000BF920" box="[180,287,1759,1784]" name="Suriname" pageId="8" pageNumber="287">Suriname</collectingCountry>
and occurs in small, shaded streams where it occurs in &quot;barriers of branches and trees in moderate to quite strong currents (3– over
<quantity id="6F6C08E85221FFCD54F6521E0330F8C4" box="[489,548,1795,1820]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="287" unit="m" value="10.0">10 m</quantity>
/min.)&quot;, and less commonly at the margins on vegetation and debris.
</paragraph>
<paragraph id="A82BA50D5221FFCD55D852320489F82C" blockId="8.[151,1437,1543,2036]" pageId="8" pageNumber="287">
In papers concerning the fauna of
<collectingCountry id="D083E59D5221FFCD5754523D03A9F898" box="[587,701,1831,1856]" name="Argentina" pageId="8" pageNumber="287">Argentina</collectingCountry>
,
<bibRefCitation id="CC05D8FC5221FFCD57D652320299F898" author="Lopez" box="[713,909,1831,1856]" pageId="8" pageNumber="287" refString="Lopez Ruf, M. L. (1987) Nuevas citas para la Argentina de especies de Limnocoridae y Naucoridae (Heteroptera). Physis, 45 (109), 76." type="journal article" year="1987">López Ruf (1987</bibRefCitation>
,
<bibRefCitation id="CC05D8FC5221FFCD5686523202C7F898" author="Ruf" box="[921,979,1832,1856]" pageId="8" pageNumber="287" refString="Lopez Ruf, M. L. (2007) Notas sobre Naucoroidea (Hemiptera: Naucoridae). 3 ra. Serie. Estudios con microscopio electronico de barrido: Corion de los huevos de Ambrysus (Ambrysus) attenuatus Montandon, Ambrysus (Ambrysus) acutangulus Montandon y Ambrysus (Ambrysus) stali La Rivers. Lundiana, 8 (1), 9 - 12." type="journal article" year="2007">2007</bibRefCitation>
) and Mazzucconi et al. (2009) indicated the presence of
<taxonomicName id="6F94DE8E5221FFCD545352560089F8BC" box="[332,413,1867,1892]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5221FFCD545352560089F8BC" box="[332,413,1867,1892]" italics="true" pageId="8" pageNumber="287">A. stali</emphasis>
</taxonomicName>
in Misiones Province, which is in the northeastern part of the country.
<bibRefCitation id="CC05D8FC5221FFCD51D052560488F8BC" box="[1231,1436,1867,1892]" pageId="8" pageNumber="287" refString="Lopez Ruf, M. L. (2007) Notas sobre Naucoroidea (Hemiptera: Naucoridae). 3 ra. Serie. Estudios con microscopio electronico de barrido: Corion de los huevos de Ambrysus (Ambrysus) attenuatus Montandon, Ambrysus (Ambrysus) acutangulus Montandon y Ambrysus (Ambrysus) stali La Rivers. Lundiana, 8 (1), 9 - 12." type="journal article">López Ruf (2007)</bibRefCitation>
also provided a description of egg structure of the
<collectingCountry id="D083E59D5221FFCD57FB52750247F850" box="[740,851,1903,1928]" name="Argentina" pageId="8" pageNumber="287">Argentina</collectingCountry>
population. We have been unable to confirm their identifications because voucher specimens are not present at Museo de La Plata or Museo
<collectingCountry id="D083E59D5221FFCD51DF52890425F874" box="[1216,1329,1939,1964]" name="Argentina" pageId="8" pageNumber="287">Argentino</collectingCountry>
Ciencias Naturales. An undescribed species of
<taxonomicName id="6F94DE8E5221FFCD572452AD03BDF808" box="[571,681,1975,2000]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F5221FFCD572452AD03BDF808" box="[571,681,1975,2000]" italics="true" pageId="8" pageNumber="287">Ambrysus</emphasis>
</taxonomicName>
occurs nearby in
<collectingCountry id="D083E59D5221FFCD567152A202C2F808" box="[878,982,1976,2000]" name="Paraguay" pageId="8" pageNumber="287">Paraguay</collectingCountry>
that bears superficial resemblance to the
<taxonomicName id="6F94DE8E5221FFCD558852C601FFF82C" box="[151,235,2011,2036]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5221FFCD558852C601FFF82C" box="[151,235,2011,2036]" italics="true" pageId="8" pageNumber="287">A. stali</emphasis>
</taxonomicName>
complex; however, male genitalic features indicate that it is clearly does not belong to this complex.
</paragraph>
<paragraph id="A82BA50D5220FFCC558855820534FF20" blockId="9.[151,1436,151,248]" pageId="9" pageNumber="288">
Because the
<collectingCountry id="D083E59D5220FFCC5434558D008EFF68" box="[299,410,151,176]" name="Argentina" pageId="9" pageNumber="288">Argentina</collectingCountry>
records are widely separated from all other records of the species and a similar species occurs in that region, we suspect that the
<collectingCountry id="D083E59D5220FFCC577955A603C1FF0D" box="[614,725,188,213]" name="Argentina" pageId="9" pageNumber="288">Argentina</collectingCountry>
material is probably not conspecific with
<taxonomicName id="6F94DE8E5220FFCC51B255A605E8FF0D" box="[1197,1276,188,213]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="288" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5220FFCC51B255A605E8FF0D" box="[1197,1276,188,213]" italics="true" pageId="9" pageNumber="288">A. stali</emphasis>
</taxonomicName>
.
<emphasis id="9AE0791F5220FFCC55D855C50071FF20" bold="true" box="[199,357,223,248]" pageId="9" pageNumber="288">
<typeStatus id="772F1BAF5220FFCC55D855C50016FF20" box="[199,258,223,248]" pageId="9" pageNumber="288">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D083E59D5220FFCC547355FA0346FF20" box="[364,594,224,248]" name="French Guiana" pageId="9" pageNumber="288">FRENCH GUIANA</collectingCountry>
, St. Laurent.
<typeStatus id="772F1BAF5220FFCC57F755C50244FF20" box="[744,848,223,248]" pageId="9" pageNumber="288" type="holotype">Holotype</typeStatus>
is housed at CAS.
</paragraph>
</subSubSection>
<caption id="FCEBF5855220FFCC55885311044CF987" httpUri="https://zenodo.org/record/237555/files/figure.png" pageId="9" pageNumber="288" targetBox="[160,1420,304,1503]" targetPageId="9">
<paragraph id="A82BA50D5220FFCC55885311044CF987" blockId="9.[151,1436,1547,1631]" pageId="9" pageNumber="288">
<emphasis id="9AE0791F5220FFCC558853110007F9F8" bold="true" box="[151,275,1547,1569]" pageId="9" pageNumber="288">FIGURE 6.</emphasis>
Structures of
<taxonomicName id="6F94DE8E5220FFCC54BD5311032CF9F9" box="[418,568,1547,1569]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="288" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F5220FFCC54BD5311032CF9F9" box="[418,568,1547,1569]" italics="true" pageId="9" pageNumber="288">Ambrysus stali</emphasis>
</taxonomicName>
. (A) dorsal habitus of male, (B) 6th and 7th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (setae have been removed to reveal shape).
</paragraph>
</caption>
<subSubSection id="E08EF6865220FFCF55D8539E040AFD8F" lastPageId="10" lastPageNumber="289" pageId="9" pageNumber="288" type="materials_examined">
<paragraph id="A82BA50D5220FFCF55D8539E05B0FE35" blockId="9.[151,1436,1668,2010]" lastBlockId="10.[151,1437,151,599]" lastPageId="10" lastPageNumber="289" pageId="9" pageNumber="288">
<emphasis id="9AE0791F5220FFCC55D8539E00BFF945" bold="true" box="[199,427,1668,1693]" pageId="9" pageNumber="288">Material examined.</emphasis>
,
<emphasis id="9AE0791F5220FFCC54A8539E03B6F945" bold="true" box="[439,674,1668,1693]" pageId="9" pageNumber="288">
<collectingCountry id="D083E59D5220FFCC54A8539E03B6F945" box="[439,674,1668,1693]" name="French Guiana" pageId="9" pageNumber="288">FRENCH GUIANA</collectingCountry>
</emphasis>
: St. Laurent [du Maroni],
<date id="DC2A83CD5220FFCC56A0539E0542F945" box="[959,1110,1668,1693]" pageId="9" pageNumber="288" value="1937-10">October 1937</date>
, H. E. Hinton, Cal. Acad. Sci. Coll. /
<collectingCountry id="D083E59D5220FFCC55FB53B30016F918" box="[228,258,1705,1728]" name="Iran" pageId="9" pageNumber="288">Ira</collectingCountry>
La Rivers Collection, Bequeathed to the California Academy of Sciences—1978 / California Academy of Sciences
<typeStatus id="772F1BAF5220FFCC55E253D60027F93C" box="[253,307,1740,1764]" pageId="9" pageNumber="288">Type</typeStatus>
No. 13412 /
<taxonomicName id="6F94DE8E5220FFCC54DB53D103CFF93C" authority="La Rivers" authorityName="La Rivers" box="[452,731,1739,1764]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="288" phylum="Arthropoda" rank="species" species="stali">Ambrysus stäli La Rivers</taxonomicName>
<typeStatus id="772F1BAF5220FFCC57FD53D60267F93C" box="[738,883,1740,1764]" pageId="9" pageNumber="288" type="holotype">HOLOTYPE</typeStatus>
(♂,
<collectionCode id="CE853DC85220FFCC56B953D602F4F93C" LSID="urn:lsid:biocol.org:col:34700" box="[934,992,1740,1764]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="9" pageNumber="288">CAS</collectionCode>
).
<emphasis id="9AE0791F5220FFCC56ED53D1057CF93C" bold="true" box="[1010,1128,1739,1764]" pageId="9" pageNumber="288">
<collectingCountry id="D083E59D5220FFCC56ED53D1057CF93C" box="[1010,1128,1739,1764]" name="Guyana" pageId="9" pageNumber="288">GUYANA</collectingCountry>
</emphasis>
: REGION 9, Parabara, elev
<quantity id="6F6C08E85220FFCC558853F501F5F8DE" box="[151,225,1775,1799]" metricMagnitude="2" metricUnit="m" metricValue="2.7399999999999998" pageId="9" pageNumber="288" unit="m" value="274.0">274 m</quantity>
, 2º 6' 29.52&quot;, -59 º13' 39.183&quot;,
<date id="DC2A83CD5220FFCC575553F50203F8DF" box="[586,791,1774,1799]" pageId="9" pageNumber="288" value="2013-11-03">3 November 2013</date>
, Short, Isaacs, Salisbury / GY–1103–02A, Small flowing creek with sand-detritus margins &amp; leaf packs-stick jams (
<specimenCount id="BE926E845220FFCC562352080274F8F2" box="[828,864,1810,1834]" pageId="9" pageNumber="288" type="generic">2♂</specimenCount>
, 2♀, UMC); Kusad Mts., Basecamp, 2.811917, - 59.866833, 163,
<date id="DC2A83CD5220FFCC544F522F031FF895" box="[336,523,1845,1870]" pageId="9" pageNumber="288" value="2013-10-23">23 October 2013</date>
, A. E. Short, small sandy stream with detritus, GY
<date id="DC2A83CD5220FFCC5126522F05ABF895" box="[1081,1215,1845,1869]" pageId="9" pageNumber="288" value="1023-01-13">13–1023–01</date>
A (
<specimenCount id="BE926E845220FFCC51FA522F041CF895" box="[1253,1288,1845,1869]" pageId="9" pageNumber="288" type="generic">1♂</specimenCount>
, 1–5th instar, UMC); Region 9, Kusad Mts., Basecamp, 2.811917, -59.866833,
<quantity id="6F6C08E85220FFCC567D524202BEF8A8" box="[866,938,1880,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="9" pageNumber="288" unit="m" value="163.0">163 m</quantity>
,
<date id="DC2A83CD5220FFCC56AB5242057AF8A8" box="[948,1134,1880,1905]" pageId="9" pageNumber="288" value="2013-10-23">23 October 2013</date>
, A. E. Short, pool in mostly dried streambed GY
<date id="DC2A83CD5220FFCC546252660310F84C" box="[381,516,1916,1940]" pageId="9" pageNumber="288" value="1023-02-13">13–1023–02</date>
A (
<specimenCount id="BE926E845220FFCC573252660344F84C" box="[557,592,1916,1940]" pageId="9" pageNumber="288" type="generic">1♂</specimenCount>
, UMC); N. Parabara, creek by basecamp (Bototo wau creek), 2.181800, - 59.338432,
<quantity id="6F6C08E85220FFCC540352850073F86E" box="[284,359,1951,1975]" metricMagnitude="2" metricUnit="m" metricValue="2.8899999999999997" pageId="9" pageNumber="288" unit="m" value="289.0">289 m</quantity>
,
<date id="DC2A83CD5220FFCC546952850323F86F" box="[374,567,1950,1975]" pageId="9" pageNumber="288" value="2013-10-31">31 October 2013</date>
, Short, Isaacs, Salisbury, stream margins, GY
<date id="DC2A83CD5220FFCC515C528505AAF86F" box="[1091,1214,1951,1975]" pageId="9" pageNumber="288" value="1031-01-13">13-1031-01</date>
A (
<specimenCount id="BE926E845220FFCC51F85285041EF86F" box="[1255,1290,1951,1975]" pageId="9" pageNumber="288" type="generic">1♂</specimenCount>
, 1♀, UMC).
<emphasis id="9AE0791F5220FFCC558852DB0039F802" bold="true" box="[151,301,1985,2010]" pageId="9" pageNumber="288">
<collectingCountry id="D083E59D5220FFCC558852DB0039F802" box="[151,301,1985,2010]" name="Suriname" pageId="9" pageNumber="288">SURINAME</collectingCountry>
</emphasis>
: SIPALIWINI, Camp 1, Kutari river,
<geoCoordinate id="CDA0C3CA5220FFCC57F652D8027CF802" box="[745,872,1986,2010]" direction="north" orientation="latitude" pageId="9" pageNumber="288" precision="1" value="2.1828833">2°10.973'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA5220FFCC566752DB051CF802" box="[888,1032,1985,2010]" direction="west" orientation="longitude" pageId="9" pageNumber="288" precision="1" value="-56.78725">56°47.235'W</geoCoordinate>
,
<quantity id="6F6C08E85220FFCC510752D80571F801" box="[1048,1125,1986,2010]" metricMagnitude="2" metricUnit="m" metricValue="2.38" pageId="9" pageNumber="288" unit="m" value="238.0">238 m</quantity>
,
<date id="DC2A83CD5220FFCC516B52D8043BF802" box="[1140,1327,1986,2010]" pageId="9" pageNumber="288" value="2010-08-22">22 August 2010</date>
, Short &amp; Kadosoe, forest swamp, SR10-0822-02A (
<specimenCount id="BE926E845223FFCF576C55820382FF68" box="[627,662,152,176]" pageId="10" pageNumber="289" type="generic">1♂</specimenCount>
, 2♀, UMC); Camp 1, Kutari river, 2.175350, -56.787399,
<quantity id="6F6C08E85223FFCF503555820460FF77" box="[1322,1396,152,176]" metricMagnitude="2" metricUnit="m" metricValue="2.28" pageId="10" pageNumber="289" unit="m" value="228.0">228 m</quantity>
,
<date id="DC2A83CD5223FFCF509E55820033FF0B" pageId="10" pageNumber="289" value="2010-08-20">20 August 2010</date>
, Short &amp; Kadosoe, forest stream, SR10-0820-01A (
<specimenCount id="BE926E845223FFCF567E55A10292FF0B" box="[865,902,187,211]" pageId="10" pageNumber="289" type="generic">4♂</specimenCount>
, UMC); Camp 4 (low): Kasikasima, 2.977310, - 55.384998,
<quantity id="6F6C08E85223FFCF540755C50074FF2E" box="[280,352,223,247]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="289" unit="m" value="200.0">200 m</quantity>
,
<date id="DC2A83CD5223FFCF547555C50300FF2F" box="[362,532,222,247]" pageId="10" pageNumber="289" value="2012-03-20">20 March 2012</date>
, A. E. Short, sandy stream, SR12-0320-02A (
<specimenCount id="BE926E845223FFCF511955C5053EFF2F" box="[1030,1066,223,247]" pageId="10" pageNumber="289" type="generic">2♂</specimenCount>
, UMC); Central
<collectingCountry id="D083E59D5223FFCF51FD55C4045FFF2F" box="[1250,1355,222,247]" name="Suriname" pageId="10" pageNumber="289">Suriname</collectingCountry>
Nature Reserve: Tafelberg Summit, near Augustus Creek Camp, 3.926667, -56.188332,
<quantity id="6F6C08E85223FFCF513B541B057BFEC1" box="[1060,1135,257,282]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="289" unit="m" value="600.0">600 m</quantity>
,
<date id="DC2A83CD5223FFCF516254180420FEC2" box="[1149,1332,258,282]" pageId="10" pageNumber="289" value="2013-08-17">17 August 2013</date>
, Short &amp; Bloom, forested detrital pools and creeks on trail into Arrowhead basin, SR13-0817-01A (1♀, UMC); Central
<collectingCountry id="D083E59D5223FFCF5588545D0014FEB8" box="[151,256,327,352]" name="Suriname" pageId="10" pageNumber="289">Suriname</collectingCountry>
Nature Reserve, Tafelberg Summit, near Augustus Creek Camp, 3.926667, -56.188332,
<quantity id="6F6C08E85223FFCF51CA545D040AFE87" box="[1237,1310,327,352]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="289" unit="m" value="600.0">600 m</quantity>
,
<date id="DC2A83CD5223FFCF5035545201C5FE5B" pageId="10" pageNumber="289" value="2013-08-17">17 August 2013</date>
, SR13–0817–03A (
<specimenCount id="BE926E845223FFCF54B2547100C6FE5B" box="[429,466,363,387]" pageId="10" pageNumber="289" type="generic">4♂</specimenCount>
, 5♀, UMC); Central
<collectingCountry id="D083E59D5223FFCF57DD5471023FFE5C" box="[706,811,363,388]" name="Suriname" pageId="10" pageNumber="289">Suriname</collectingCountry>
Nature Reserve, Tafelberg Summit, near
<collectingCountry id="D083E59D5223FFCF51E254710446FE5C" box="[1277,1362,363,388]" name="Cayman Islands" pageId="10" pageNumber="289">Caiman</collectingCountry>
Creek Camp, 3.899033, -56.180817,
<quantity id="6F6C08E85223FFCF54F05495032EFE7E" box="[495,570,399,423]" metricMagnitude="2" metricUnit="m" metricValue="7.33" pageId="10" pageNumber="289" unit="m" value="733.0">733 m</quantity>
,
<date id="DC2A83CD5223FFCF575654950214FE7F" box="[585,768,399,423]" pageId="10" pageNumber="289" value="2013-08-19">19 August 2013</date>
, Short &amp; Bloom, Caimen Creek, pools and creek margins, SR13-0819-04B (
<specimenCount id="BE926E845223FFCF544254A80094FE12" box="[349,384,434,458]" pageId="10" pageNumber="289" type="generic">1♂</specimenCount>
).
<emphasis id="9AE0791F5223FFCF548854AB0336FE12" bold="true" box="[407,546,433,458]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF548854AB0336FE12" box="[407,546,433,458]" name="Trinidad and Tobago" pageId="10" pageNumber="289">TRINIDAD</collectingCountry>
</emphasis>
: bet. Four Roads &amp; Carmichael,
<date id="DC2A83CD5223FFCF568654AB050CFE12" box="[921,1048,433,458]" pageId="10" pageNumber="289" value="2005-07-05">
5 VII
<quantity id="6F6C08E85223FFCF56C054AB050CFE12" box="[991,1048,433,458]" metricMagnitude="5" metricUnit="m" metricValue="6.11191056" pageId="10" pageNumber="289" unit="ft" value="2005220.0">2005</quantity>
</date>
, 220 ft, unnamed stream,
<geoCoordinate id="CDA0C3CA5223FFCF502254A8048CFE12" box="[1341,1432,434,458]" direction="north" orientation="latitude" pageId="10" pageNumber="289" precision="925" value="10.5">10º30'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA5223FFCF558854CE01E1FE35" box="[151,245,468,493]" direction="west" orientation="longitude" pageId="10" pageNumber="289" precision="925" value="-61.216667">61º13'W</geoCoordinate>
/ (underside of same label) WDS-A-1647 / William D. Shepard, leg. (
<specimenCount id="BE926E845223FFCF56E954CF050DFE35" box="[1014,1049,469,493]" pageId="10" pageNumber="289" type="generic">1♂</specimenCount>
, 1♀, UMC).
</paragraph>
<paragraph id="A82BA50D5223FFCF55D854ED040AFD8F" blockId="10.[151,1437,151,599]" pageId="10" pageNumber="289">
<emphasis id="9AE0791F5223FFCF55D854ED00BDFDC8" bold="true" box="[199,425,503,528]" pageId="10" pageNumber="289">Published records.</emphasis>
<bibRefCitation id="CC05D8FC5223FFCF54A654E2039FFDC8" box="[441,651,503,528]" pageId="10" pageNumber="289" refString="La Rivers, I. (1962) A new Ambrysus from South America (Hemiptera, Naucoridae). Bulletin of the Southern California Academy of Sciences, 61 (3), 185 - 187." type="journal article">La Rivers (1962)</bibRefCitation>
:
<emphasis id="9AE0791F5223FFCF578054ED021CFDC8" bold="true" box="[671,776,503,528]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF578054ED021CFDC8" box="[671,776,503,528]" name="Brazil" pageId="10" pageNumber="289">BRAZIL</collectingCountry>
</emphasis>
, Pará;
<emphasis id="9AE0791F5223FFCF567754ED0577FDC8" bold="true" box="[872,1123,503,528]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF567754ED054AFDC8" box="[872,1118,503,528]" name="French Guiana" pageId="10" pageNumber="289">FRENCH GUIANA</collectingCountry>
,
</emphasis>
St. Laurent [du Maroni]; [
<emphasis id="9AE0791F5223FFCF55BF57010022FDEC" bold="true" box="[160,310,539,564]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF55BF57010022FDEC" box="[160,310,539,564]" name="Suriname" pageId="10" pageNumber="289">SURINAME</collectingCountry>
</emphasis>
];
<emphasis id="9AE0791F5223FFCF5451570100CDFDEC" bold="true" box="[334,473,539,564]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF5451570100CDFDEC" box="[334,473,539,564]" name="Trinidad and Tobago" pageId="10" pageNumber="289">TRINIDAD</collectingCountry>
</emphasis>
.
<bibRefCitation id="CC05D8FC5223FFCF54F8570603B2FDEB" box="[487,678,539,564]" pageId="10" pageNumber="289" refString="La Rivers, I. &amp; Nieser, N. (1972) The Ambrysus of Surinam, with the description of a new species, A. bifidus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 29, 1 - 8." type="journal article">La Rivers (1972)</bibRefCitation>
,
<bibRefCitation id="CC05D8FC5223FFCF57AD57010259FDEC" author="Nieser" box="[690,845,539,564]" pageId="10" pageNumber="289" refString="Nieser, N. (1975) The water bugs of the Guyana region. Studies on the Fauna of Suriname and Other Guyanas, 59, 1 - 310. http: // dx. doi. org / 10.1007 / 978 - 94 - 017 - 7118 - 4 _ 1" type="journal article" year="1975">Nieser (1975)</bibRefCitation>
:
<emphasis id="9AE0791F5223FFCF5646570102D7FDEC" bold="true" box="[857,963,539,564]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF5646570102D7FDEC" box="[857,963,539,564]" name="Brazil" pageId="10" pageNumber="289">BRAZIL</collectingCountry>
</emphasis>
, Amazonas and Pará states;
<emphasis id="9AE0791F5223FFCF51E057010481FDEC" bold="true" box="[1279,1429,539,564]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF51E057010481FDEC" box="[1279,1429,539,564]" name="Suriname" pageId="10" pageNumber="289">SURINAME</collectingCountry>
</emphasis>
, Brokopondo, Marowijne, and Saramacca Districts.
<bibRefCitation id="CC05D8FC5223FFCF57D6572402CBFD8F" author="Araujo" box="[713,991,574,599]" pageId="10" pageNumber="289" refString="Araujo, Y. &amp; Beserra, P. (2007) Biodiversidad de invertebrados consumidos por las etnias Yanomani y Yekuana del Alto Orinoco, Venezuela. Interciencia, 32 (5), 318 - 323." type="journal article" year="2007">Araujo &amp; Beserra (2007)</bibRefCitation>
:
<emphasis id="9AE0791F5223FFCF56F657240581FD8F" bold="true" box="[1001,1173,574,599]" pageId="10" pageNumber="289">
<collectingCountry id="D083E59D5223FFCF56F657240581FD8F" box="[1001,1173,574,599]" name="Venezuela" pageId="10" pageNumber="289">VENEZUELA</collectingCountry>
</emphasis>
, Amazonas.
</paragraph>
</subSubSection>
</treatment>
</document>

164
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<heading id="F36312615222FFCE5588558D00EBFF6A" bold="true" box="[151,511,151,178]" fontSize="11" level="1" pageId="11" pageNumber="290" reason="1">
<taxonomicName id="6F94DE8E5222FFCE5588558D00EBFF6A" ID-CoL="7QM69" authority="La Rivers" authorityName="La Rivers" box="[151,511,151,178]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5222FFCE5588558D00EBFF6A" bold="true" box="[151,511,151,178]" pageId="11" pageNumber="290">
<emphasis id="9AE0791F5222FFCE5588558D0069FF69" bold="true" box="[151,381,151,177]" italics="true" pageId="11" pageNumber="290">Ambrysus tricuspis</emphasis>
La Rivers
</emphasis>
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<paragraph id="A82BA50D5222FFCE558855A00006FF0B" blockId="11.[151,511,151,211]" box="[151,274,186,211]" pageId="11" pageNumber="290">
(
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,
<figureCitation id="30AFB9885222FFCE55E555A1001DFF0B" box="[250,265,187,211]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1931]" captionTargetBox="[166,1426,653,1862]" captionTargetId="figure@10.[166,1426,653,1862]" captionTargetPageId="10" captionText="FIGURE 7. Structures of Ambrysus tricuspis (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of holotype showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing)." httpUri="https://zenodo.org/record/237556/files/figure.png" pageId="11" pageNumber="290">7</figureCitation>
)
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<subSubSection id="E08EF6865222FFCE558854190264FEC1" box="[151,880,259,281]" pageId="11" pageNumber="290" type="reference_group">
<paragraph id="A82BA50D5222FFCE558854190264FEC1" blockId="11.[151,880,259,281]" box="[151,880,259,281]" pageId="11" pageNumber="290">
<taxonomicName id="6F94DE8E5222FFCE5588541900EAFEC1" ID-CoL="7QM69" authority="La Rivers 1974" authorityName="La Rivers" authorityYear="1974" box="[151,510,259,281]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5222FFCE55885419004CFEC1" box="[151,344,259,281]" italics="true" pageId="11" pageNumber="290">Ambrysus tricuspis</emphasis>
<bibRefCitation id="CC05D8FC5222FFCE5441541E00EAFEC1" author="La" box="[350,510,259,281]" pageId="11" pageNumber="290" refString="La Rivers, I. (1974) Catalogue of taxa described in the family Naucoridae (Hemiptera) supplement no. 1: Corrections, emendations and additions, with descriptions of new species. Biological Society of Nevada Occasional Papers, 38, 1 - 17." type="journal article" year="1974">La Rivers 1974</bibRefCitation>
</taxonomicName>
: Biol. Soc. Nevada Occas. Pap. 2–3.
</paragraph>
</subSubSection>
<subSubSection id="E08EF6865222FFCE558854240459FCF4" pageId="11" pageNumber="290" type="discussion">
<paragraph id="A82BA50D5222FFCE558854240459FCF4" blockId="11.[151,1437,318,956]" pageId="11" pageNumber="290">
<emphasis id="9AE0791F5222FFCE558854240001FE8F" bold="true" box="[151,277,318,343]" pageId="11" pageNumber="290">Discussion</emphasis>
. This species is known only from the
<typeStatus id="772F1BAF5222FFCE57DB54240232FE8F" box="[708,806,318,343]" pageId="11" pageNumber="290" type="holotype">holotype</typeStatus>
and allotype, which were collected together in northern
<collectingCountry id="D083E59D5222FFCE55885479001DFEA4" box="[151,265,355,380]" name="Colombia" pageId="11" pageNumber="290">Colombia</collectingCountry>
. No information regarding habitat or biology was given in the original description. It differs from
<taxonomicName id="6F94DE8E5222FFCE509B547E01F7FE47" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5222FFCE509B547E01F7FE47" italics="true" pageId="11" pageNumber="290">A. bifidus</emphasis>
</taxonomicName>
by the unbifurcated median lobe of the subgenital plate (
<figureCitation id="30AFB9885222FFCE567C549C02B2FE47" box="[867,934,390,415]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1931]" captionTargetBox="[166,1426,653,1862]" captionTargetId="figure@10.[166,1426,653,1862]" captionTargetPageId="10" captionText="FIGURE 7. Structures of Ambrysus tricuspis (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of holotype showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing)." httpUri="https://zenodo.org/record/237556/files/figure.png" pageId="11" pageNumber="290">Fig. 7</figureCitation>
E). Although
<bibRefCitation id="CC05D8FC5222FFCE5121549205E9FE47" box="[1086,1277,390,415]" pageId="11" pageNumber="290" refString="La Rivers, I. (1974) Catalogue of taxa described in the family Naucoridae (Hemiptera) supplement no. 1: Corrections, emendations and additions, with descriptions of new species. Biological Society of Nevada Occasional Papers, 38, 1 - 17." type="journal article">La Rivers (1974)</bibRefCitation>
suggested the male accessory genitalic process is more massive and elongated laterally (
<figureCitation id="30AFB9885222FFCE56FC54B10532FE1B" box="[995,1062,427,452]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1931]" captionTargetBox="[166,1426,653,1862]" captionTargetId="figure@10.[166,1426,653,1862]" captionTargetPageId="10" captionText="FIGURE 7. Structures of Ambrysus tricuspis (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) terminal abdominal sterna of holotype showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing)." httpUri="https://zenodo.org/record/237556/files/figure.png" pageId="11" pageNumber="290">Fig. 7</figureCitation>
B) than in
<taxonomicName id="6F94DE8E5222FFCE51BE54B60419FE1C" box="[1185,1293,427,452]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5222FFCE51BE54B60419FE1C" box="[1185,1293,427,452]" italics="true" pageId="11" pageNumber="290">A. bifidus</emphasis>
</taxonomicName>
, this feature exhibits intraspecific variation in other species of
<taxonomicName id="6F94DE8E5222FFCE57CB54D40256FE3F" box="[724,834,462,487]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F5222FFCE57CB54D40256FE3F" box="[724,834,462,487]" italics="true" pageId="11" pageNumber="290">Ambrysus</emphasis>
</taxonomicName>
, including within this species complex. Specifically,
<bibRefCitation id="CC05D8FC5222FFCE558854E90026FDD4" author="Nieser" box="[151,306,499,524]" pageId="11" pageNumber="290" refString="Nieser, N. (1975) The water bugs of the Guyana region. Studies on the Fauna of Suriname and Other Guyanas, 59, 1 - 310. http: // dx. doi. org / 10.1007 / 978 - 94 - 017 - 7118 - 4 _ 1" type="journal article" year="1975">Nieser (1975)</bibRefCitation>
documented sufficient intraspecific variation in the shape of the accessory genitalic process among
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of
<taxonomicName id="6F94DE8E5222FFCE5439570D0085FDF7" box="[294,401,534,559]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5222FFCE5439570D0085FDF7" box="[294,401,534,559]" italics="true" pageId="11" pageNumber="290">A. bifidus</emphasis>
</taxonomicName>
from
<collectingCountry id="D083E59D5222FFCE54CA570C0354FDF7" box="[469,576,534,559]" name="Suriname" pageId="11" pageNumber="290">Suriname</collectingCountry>
that its precise shape should not be considered a reliable diagnostic attribute. La Rivers made no attempt to distinguish this species specifically from
<taxonomicName id="6F94DE8E5222FFCE56D757260523FD8C" box="[968,1079,571,596]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F5222FFCE56D757260523FD8C" box="[968,1079,571,596]" italics="true" pageId="11" pageNumber="290">A. scolius</emphasis>
</taxonomicName>
, but thought the &quot;three-pointed subgenital plate&quot; and &quot;exaggerated dog's head outline&quot; of the male accessory genitalic process would satisfactorily distinguish
<taxonomicName id="6F94DE8E5222FFCE5403579E008AFD43" box="[284,414,644,667]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5222FFCE5403579E008AFD43" box="[284,414,644,667]" italics="true" pageId="11" pageNumber="290">A. tricuspis</emphasis>
</taxonomicName>
from all congeners. Although we are unable to find reliable characters to distinguish
<taxonomicName id="6F94DE8E5222FFCE509B579E01E3FD67" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5222FFCE509B579E01E3FD67" italics="true" pageId="11" pageNumber="290">A. tricuspis</emphasis>
</taxonomicName>
from
<taxonomicName id="6F94DE8E5222FFCE545D57BD00A6FD67" box="[322,434,678,703]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F5222FFCE545D57BD00A6FD67" box="[322,434,678,703]" italics="true" pageId="11" pageNumber="290">A. scolius</emphasis>
</taxonomicName>
, the slight difference in shape of the male accessory genitalic process, the geographic distance between the known ranges, and the paucity of specimens are enough that we chose not to synonymize these species at this time. More specimens of both species are needed to determine the extent of intraspecific variation in these features, which will then enable us to more effectively address the validity of this species.
</paragraph>
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<subSubSection id="E08EF6865222FFCE55D8562C05BBFC63" pageId="11" pageNumber="290" type="materials_examined">
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<emphasis id="9AE0791F5222FFCE55D8562C0071FC97" bold="true" box="[199,357,822,847]" pageId="11" pageNumber="290">
<typeStatus id="772F1BAF5222FFCE55D8562C0016FC97" box="[199,258,822,847]" pageId="11" pageNumber="290">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D083E59D5222FFCE5473562D0310FC97" box="[364,516,823,847]" name="Colombia" pageId="11" pageNumber="290">COLOMBIA</collectingCountry>
, Bolivar Department.
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specimen is housed at
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.
</paragraph>
<paragraph id="A82BA50D5222FFCE55D8564105BBFC63" blockId="11.[151,1437,318,956]" pageId="11" pageNumber="290">
<emphasis id="9AE0791F5222FFCE55D856410349FCAC" bold="true" box="[199,605,859,884]" pageId="11" pageNumber="290">
Material examined.
<collectingCountry id="D083E59D5222FFCE54A056410349FCAC" box="[447,605,859,884]" name="Colombia" pageId="11" pageNumber="290">COLOMBIA</collectingCountry>
</emphasis>
: Bolivar, San Marcos / (reverse side of locality label)
<date id="DC2A83CD5222FFCE51FC56410464FCAB" box="[1251,1392,859,884]" pageId="11" pageNumber="290" value="1964-08-08">1964 Aug 8</date>
,
<collectingCountry id="D083E59D5222FFCE509F5641048CFCAB" box="[1408,1432,859,883]" name="Iceland" pageId="11" pageNumber="290">O</collectingCountry>
. Pinada /
<taxonomicName id="6F94DE8E5222FFCE55E7566400C5FC4F" box="[248,465,894,919]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="tricuspis">Ambrysus tricuspis</taxonomicName>
<typeStatus id="772F1BAF5222FFCE54C85664032BFC4F" box="[471,575,894,919]" pageId="11" pageNumber="290" type="holotype">Holotype</typeStatus>
/ California Academy of Sciences
<typeStatus id="772F1BAF5222FFCE56DA569A02E9FC40" box="[965,1021,896,920]" pageId="11" pageNumber="290">Type</typeStatus>
No. 13406 /
<collectingCountry id="D083E59D5222FFCE5191569A05B9FC4F" box="[1166,1197,896,919]" name="Iran" pageId="11" pageNumber="290">Ira</collectingCountry>
La Rivers Collection Bequeathed to the California Academy of Sciences (♀,
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); same data, Allotype (♂,
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).
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="6F94DE8E5228FFC4558852380376F8E4" ID-CoL="7QM3R" authority="La Rivers &amp; Nieser" authorityName="La Rivers &amp; Nieser" box="[151,610,1826,1852]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="280" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5228FFC4558852380376F8E4" bold="true" box="[151,610,1826,1852]" pageId="1" pageNumber="280">
<emphasis id="9AE0791F5228FFC455885238007DF8E4" bold="true" box="[151,361,1826,1852]" italics="true" pageId="1" pageNumber="280">Ambrysus bifidus</emphasis>
La Rivers &amp; Nieser
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A82BA50D5228FFC45588525F0006F886" blockId="1.[151,610,1826,1886]" box="[151,274,1861,1886]" pageId="1" pageNumber="280">
(
<figureCitation id="30AFB9885228FFC45580525F001CF886" box="[159,264,1861,1886]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1189,1211]" captionStartId-1="3.[151,250,1509,1531]" captionTargetBox-0="[151,1436,198,1167]" captionTargetBox-1="[151,1436,193,1486]" captionTargetId-0="figure@2.[151,1436,198,1167]" captionTargetId-1="figure@3.[151,1436,193,1487]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Distribution of species in the Ambrysus stali species complex." captionText-1="FIGURE 2. Structures of Ambrysus bifidus paratypes. (A) dorsal habitus of female, (B) 6 th and 7 th abdominal terga of male, (C) 8 th abdominal tergum of male, (D) ventral and left sides of phallosoma, (E) male pygophore (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing). agp = accessory genitalic process, lvl = left ventral lobe of phallosoma, pct = proctiger, pp = pseudoparamere, pr = paramere, pyg = pygophore." httpUri-0="https://zenodo.org/record/237550/files/figure.png" httpUri-1="https://zenodo.org/record/237551/files/figure.png" pageId="1" pageNumber="280">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08EF6865228FFC4558852970503F81A" pageId="1" pageNumber="280" type="reference_group">
<paragraph id="A82BA50D5228FFC45588529702D4F87B" blockId="1.[151,1047,1933,1986]" box="[151,960,1933,1955]" pageId="1" pageNumber="280">
<taxonomicName id="6F94DE8E5228FFC455885297035AF87B" ID-CoL="7QM3R" authority="La Rivers &amp; Nieser 1972" authorityName="La Rivers &amp; Nieser" authorityYear="1972" box="[151,590,1933,1955]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="280" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5228FFC4558852970052F87B" box="[151,326,1933,1955]" italics="true" pageId="1" pageNumber="280">Ambrysus bifidus</emphasis>
<bibRefCitation id="CC05D8FC5228FFC454535294035AF87B" author="La" box="[332,590,1933,1955]" pageId="1" pageNumber="280" refString="La Rivers, I. &amp; Nieser, N. (1972) The Ambrysus of Surinam, with the description of a new species, A. bifidus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 29, 1 - 8." type="journal article" year="1972">La Rivers &amp; Nieser 1972</bibRefCitation>
</taxonomicName>
: Biol. Soc. Nevada Occas. Pap. 4–8.
</paragraph>
<paragraph id="A82BA50D5228FFC4558852B60503F81A" blockId="1.[151,1047,1933,1986]" box="[151,1047,1964,1986]" pageId="1" pageNumber="280">
<taxonomicName id="6F94DE8E5228FFC4558852B600C4F81A" ID-CoL="7QM3R" authority="Nieser 1975" authorityName="Nieser" authorityYear="1975" box="[151,464,1964,1986]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="280" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5228FFC4558852B60052F81A" box="[151,326,1964,1986]" italics="true" pageId="1" pageNumber="280">Ambrysus bifidus</emphasis>
:
<bibRefCitation id="CC05D8FC5228FFC4544C52B600C4F81A" author="Nieser" box="[339,464,1964,1986]" pageId="1" pageNumber="280" refString="Nieser, N. (1975) The water bugs of the Guyana region. Studies on the Fauna of Suriname and Other Guyanas, 59, 1 - 310. http: // dx. doi. org / 10.1007 / 978 - 94 - 017 - 7118 - 4 _ 1" type="journal article" year="1975">Nieser 1975</bibRefCitation>
</taxonomicName>
, Stud. Fauna
<collectingCountry id="D083E59D5228FFC4574552B603AFF81A" box="[602,699,1964,1986]" name="Suriname" pageId="1" pageNumber="280">Suriname</collectingCountry>
Guyanas 56–57 (suppl. descript.).
</paragraph>
</subSubSection>
<caption id="FCEBF585522BFFC7558851BF029AFB63" box="[151,910,1189,1211]" httpUri="https://zenodo.org/record/237550/files/figure.png" pageId="2" pageNumber="281" targetBox="[151,1436,198,1167]" targetPageId="2">
<paragraph id="A82BA50D522BFFC7558851BF029AFB63" blockId="2.[151,910,1189,1211]" box="[151,910,1189,1211]" pageId="2" pageNumber="281">
<emphasis id="9AE0791F522BFFC7558851BF0007FB62" bold="true" box="[151,275,1189,1211]" pageId="2" pageNumber="281">FIGURE 1.</emphasis>
Distribution of species in the
<taxonomicName id="6F94DE8E522BFFC7575B51BF03CDFB63" box="[580,729,1189,1211]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F522BFFC7575B51BF03CDFB63" box="[580,729,1189,1211]" italics="true" pageId="2" pageNumber="281">Ambrysus stali</emphasis>
</taxonomicName>
species complex.
</paragraph>
</caption>
<subSubSection id="E08EF686522BFFC7558851FB050EF972" pageId="2" pageNumber="281" type="discussion">
<paragraph id="A82BA50D522BFFC7558851FB00B6FA2E" blockId="2.[151,1437,1249,1958]" pageId="2" pageNumber="281">
<emphasis id="9AE0791F522BFFC7558851FB0001FB22" bold="true" box="[151,277,1249,1274]" pageId="2" pageNumber="281">Discussion</emphasis>
. This species is known from
<collectingCountry id="D083E59D522BFFC7574351FB03D3FB22" box="[604,711,1249,1274]" name="Suriname" pageId="2" pageNumber="281">Suriname</collectingCountry>
and northern
<collectingCountry id="D083E59D522BFFC7567D51FB02B2FB22" box="[866,934,1249,1274]" name="Brazil" pageId="2" pageNumber="281">Brazil</collectingCountry>
(
<bibRefCitation id="CC05D8FC522BFFC756AA51FB05C6FB21" author="La" box="[949,1234,1249,1274]" pageId="2" pageNumber="281" refString="La Rivers, I. &amp; Nieser, N. (1972) The Ambrysus of Surinam, with the description of a new species, A. bifidus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 29, 1 - 8." type="journal article" year="1972">La Rivers &amp; Nieser 1972</bibRefCitation>
). Reported here is the first record from
<collectingCountry id="D083E59D522BFFC7549D501C00C9FAC6" box="[386,477,1286,1310]" name="Guyana" pageId="2" pageNumber="281">Guyana</collectingCountry>
. This species was distinguished from
<taxonomicName id="6F94DE8E522BFFC75699501C02E7FAC6" box="[902,1011,1285,1310]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F522BFFC75699501C02E7FAC6" box="[902,1011,1285,1310]" italics="true" pageId="2" pageNumber="281">A. scolius</emphasis>
</taxonomicName>
by a &quot;slight bifurcation&quot; of the apical median lobe of the female subgenital plate (
<figureCitation id="30AFB988522BFFC7579B503303D2FA99" box="[644,710,1321,1346]" captionStart="FIGURE 2" captionStartId="3.[151,250,1509,1531]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@3.[151,1436,193,1487]" captionTargetPageId="3" captionText="FIGURE 2. Structures of Ambrysus bifidus paratypes. (A) dorsal habitus of female, (B) 6 th and 7 th abdominal terga of male, (C) 8 th abdominal tergum of male, (D) ventral and left sides of phallosoma, (E) male pygophore (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing). agp = accessory genitalic process, lvl = left ventral lobe of phallosoma, pct = proctiger, pp = pseudoparamere, pr = paramere, pyg = pygophore." httpUri="https://zenodo.org/record/237551/files/figure.png" pageId="2" pageNumber="281">Fig. 2</figureCitation>
F) (
<bibRefCitation id="CC05D8FC522BFFC757F15033051AFA99" author="La" box="[750,1038,1321,1346]" pageId="2" pageNumber="281" refString="La Rivers, I. &amp; Nieser, N. (1972) The Ambrysus of Surinam, with the description of a new species, A. bifidus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 29, 1 - 8." type="journal article" year="1972">La Rivers &amp; Nieser 1972</bibRefCitation>
), and from
<taxonomicName id="6F94DE8E522BFFC751905030041AFA99" box="[1167,1294,1322,1345]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F522BFFC751905030041AFA99" box="[1167,1294,1322,1345]" italics="true" pageId="2" pageNumber="281">A. tricuspis</emphasis>
</taxonomicName>
by the same bifurcation of the subgenital plate and also the male accessory genitalic process less massive or elongated laterally (
<figureCitation id="30AFB988522BFFC75580506B01F6FA51" box="[159,226,1393,1418]" captionStart="FIGURE 2" captionStartId="3.[151,250,1509,1531]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@3.[151,1436,193,1487]" captionTargetPageId="3" captionText="FIGURE 2. Structures of Ambrysus bifidus paratypes. (A) dorsal habitus of female, (B) 6 th and 7 th abdominal terga of male, (C) 8 th abdominal tergum of male, (D) ventral and left sides of phallosoma, (E) male pygophore (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing). agp = accessory genitalic process, lvl = left ventral lobe of phallosoma, pct = proctiger, pp = pseudoparamere, pr = paramere, pyg = pygophore." httpUri="https://zenodo.org/record/237551/files/figure.png" pageId="2" pageNumber="281">Fig. 2</figureCitation>
B) (
<bibRefCitation id="CC05D8FC522BFFC75414506B00A8FA51" author="La" box="[267,444,1393,1418]" pageId="2" pageNumber="281" refString="La Rivers, I. (1974) Catalogue of taxa described in the family Naucoridae (Hemiptera) supplement no. 1: Corrections, emendations and additions, with descriptions of new species. Biological Society of Nevada Occasional Papers, 38, 1 - 17." type="journal article" year="1974">La Rivers 1974</bibRefCitation>
).
<bibRefCitation id="CC05D8FC522BFFC754D0506B037DFA52" author="Nieser" box="[463,617,1393,1418]" pageId="2" pageNumber="281" refString="Nieser, N. (1975) The water bugs of the Guyana region. Studies on the Fauna of Suriname and Other Guyanas, 59, 1 - 310. http: // dx. doi. org / 10.1007 / 978 - 94 - 017 - 7118 - 4 _ 1" type="journal article" year="1975">Nieser (1975)</bibRefCitation>
provided a supplemental description in his treatment of the water bugs of the
<collectingCountry id="D083E59D522BFFC755DF508C0003FA76" box="[192,279,1430,1454]" name="Guyana" pageId="2" pageNumber="281">Guyana</collectingCountry>
region, in which he documented the variation in the shape of the accessory genitalic process of tergum VI.
<taxonomicName id="6F94DE8E522BFFC755DA50A00026FA0A" box="[197,306,1465,1490]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F522BFFC755DA50A00026FA0A" box="[197,306,1465,1490]" italics="true" pageId="2" pageNumber="281">A. bifidus</emphasis>
</taxonomicName>
can be distinguished from the other species in this complex by the bifurcated median lobe of the female subgenital plate.
</paragraph>
<paragraph id="A82BA50D522BFFC755D8531B050EF972" blockId="2.[151,1437,1249,1958]" pageId="2" pageNumber="281">
The habitat of the
<typeStatus id="772F1BAF522BFFC754BC531800C7F9C2" box="[419,467,1538,1562]" pageId="2" pageNumber="281">type</typeStatus>
series was given in the original description as &quot;floating plant-debris upstream of a barrier of tree branches, etc., velocity of current
<quantity id="6F6C08E8522BFFC757D5533F03EDF9E6" box="[714,761,1573,1598]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="281" unit="m" value="5.0">5 m</quantity>
/min.&quot; In
<collectingCountry id="D083E59D522BFFC75677533F02C2F9E6" box="[872,982,1573,1598]" name="Suriname" pageId="2" pageNumber="281">Suriname</collectingCountry>
,
<emphasis id="9AE0791F522BFFC756FB533C05AFF9E6" box="[996,1211,1573,1598]" italics="true" pageId="2" pageNumber="281">
<taxonomicName id="6F94DE8E522BFFC756FB533C0540F9E6" box="[996,1108,1573,1598]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="bifidus">A. bifidus</taxonomicName>
,
<taxonomicName id="6F94DE8E522BFFC7517D533C05A3F9E6" box="[1122,1207,1573,1598]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="stali">A. stali</taxonomicName>
,
</emphasis>
and
<taxonomicName id="6F94DE8E522BFFC751E8533C01F4F9BA" authority="La Rivers" authorityName="La Rivers" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="usingeri">
<emphasis id="9AE0791F522BFFC751E8533C0461F9E5" box="[1271,1397,1574,1597]" italics="true" pageId="2" pageNumber="281">A. usingeri</emphasis>
La Rivers
</taxonomicName>
were considered to be common in small streams in savannah woodlands with dark water low in electrolytes (
<bibRefCitation id="CC05D8FC522BFFC75580537400D3F95E" author="La" box="[159,455,1645,1670]" pageId="2" pageNumber="281" refString="La Rivers, I. &amp; Nieser, N. (1972) The Ambrysus of Surinam, with the description of a new species, A. bifidus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 29, 1 - 8." type="journal article" year="1972">La Rivers &amp; Nieser 1972</bibRefCitation>
).
<taxonomicName id="6F94DE8E522BFFC754C35374035DF95E" box="[476,585,1645,1670]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F522BFFC754C35374035DF95E" box="[476,585,1645,1670]" italics="true" pageId="2" pageNumber="281">A. bifidus</emphasis>
</taxonomicName>
was collected with
<taxonomicName id="6F94DE8E522BFFC7562E53740297F95E" box="[817,899,1645,1670]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F522BFFC7562E53740297F95E" box="[817,899,1645,1670]" italics="true" pageId="2" pageNumber="281">A. stali</emphasis>
</taxonomicName>
in
<collectingCountry id="D083E59D522BFFC756B45377051EF95E" box="[939,1034,1645,1670]" name="Suriname" pageId="2" pageNumber="281">Surinam</collectingCountry>
mostly in barriers of branches and dead leaves or at the bases of trees standing in water where the current is strong.
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522BFFC755D853AF0529F87D" pageId="2" pageNumber="281" type="materials_examined">
<paragraph id="A82BA50D522BFFC755D853AF0518F916" blockId="2.[151,1437,1249,1958]" box="[199,1036,1717,1742]" pageId="2" pageNumber="281">
<emphasis id="9AE0791F522BFFC755D853AF0071F916" bold="true" box="[199,357,1717,1742]" pageId="2" pageNumber="281">
<typeStatus id="772F1BAF522BFFC755D853AF0016F916" box="[199,258,1717,1742]" pageId="2" pageNumber="281">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D083E59D522BFFC7547353AC0315F916" box="[364,513,1718,1742]" name="Suriname" pageId="2" pageNumber="281">SURINAME</collectingCountry>
, Zanderijsavanne.
<typeStatus id="772F1BAF522BFFC757CC53AF022FF916" box="[723,827,1717,1742]" pageId="2" pageNumber="281" type="holotype">Holotype</typeStatus>
is housed at
<collectionCode id="CE853DC8522BFFC756D153AC051CF916" LSID="urn:lsid:biocol.org:col:34700" box="[974,1032,1718,1742]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="2" pageNumber="281">CAS</collectionCode>
.
</paragraph>
<paragraph id="A82BA50D522BFFC755D853C303AFF886" blockId="2.[151,1437,1249,1958]" pageId="2" pageNumber="281">
<emphasis id="9AE0791F522BFFC755D853C30325F92A" bold="true" box="[199,561,1753,1778]" pageId="2" pageNumber="281">
Material examined.
<collectingCountry id="D083E59D522BFFC754A653C30325F92A" box="[441,561,1753,1778]" name="Guyana" pageId="2" pageNumber="281">GUYANA</collectingCountry>
</emphasis>
: Pirara Ranch,
<geoCoordinate id="CDA0C3CA522BFFC757FC53C30243F929" box="[739,855,1753,1777]" direction="north" orientation="latitude" pageId="2" pageNumber="281" precision="92" value="3.3683333">3º 22.1' N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522BFFC7567C53C302FDF929" box="[867,1001,1753,1778]" direction="west" orientation="longitude" pageId="2" pageNumber="281" precision="92" value="-59.675">59º 40.5' W</geoCoordinate>
, Pirara River,
<date id="DC2A83CD522BFFC7519353C3040AF92A" box="[1164,1310,1753,1778]" pageId="2" pageNumber="281" value="1995-04-25">25 Apr 1995</date>
, colln #19, PJ Spangler &amp; SA Perry (1♀, UMC).
<emphasis id="9AE0791F522BFFC7575C53E703CFF8CE" bold="true" box="[579,731,1789,1814]" pageId="2" pageNumber="281">
<collectingCountry id="D083E59D522BFFC7575C53E703CFF8CE" box="[579,731,1789,1814]" name="Suriname" pageId="2" pageNumber="281">SURINAME</collectingCountry>
</emphasis>
: Zanderijsavanna
<taxonomicName id="6F94DE8E522BFFC756B453E70521F8CE" authority="Cr." authorityName="Cr." box="[939,1077,1789,1814]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="genus">Carolina Cr.</taxonomicName>
/ (reverse side of locality label)
<date id="DC2A83CD522BFFC75588523B0026F8E1" box="[151,306,1825,1850]" pageId="2" pageNumber="281" value="1969-08-18">1969 Aug 18</date>
, N. Nieser /
<taxonomicName id="6F94DE8E522BFFC754D3523B0383F8E2" box="[460,663,1825,1850]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="bifidus">Ambrysus bifidus</taxonomicName>
L&amp;N 1972
<typeStatus id="772F1BAF522BFFC756365238029FF8E2" box="[809,907,1826,1850]" pageId="2" pageNumber="281" type="paratype">Paratype</typeStatus>
/
<collectingCountry id="D083E59D522BFFC756B7523802D3F8E1" box="[936,967,1826,1849]" name="Iran" pageId="2" pageNumber="281">Ira</collectingCountry>
La Rivers Collection Bequeathed to the California Academy of Sciences (
<specimenCount id="BE926E84522BFFC7570C525C0323F886" box="[531,567,1862,1886]" pageId="2" pageNumber="281" type="generic">1♂</specimenCount>
, 1♀,
<collectionCode id="CE853DC8522BFFC7576B525C03BBF886" LSID="urn:lsid:biocol.org:col:34700" box="[628,687,1862,1886]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="2" pageNumber="281">CAS</collectionCode>
).
</paragraph>
<paragraph id="A82BA50D522BFFC755D852730529F87D" blockId="2.[151,1437,1249,1958]" pageId="2" pageNumber="281">
<emphasis id="9AE0791F522BFFC755D8527300BCF85A" bold="true" box="[199,424,1897,1922]" pageId="2" pageNumber="281">Published records.</emphasis>
(
<bibRefCitation id="CC05D8FC522BFFC754A5527003F3F85A" author="La" box="[442,743,1897,1922]" pageId="2" pageNumber="281" refString="La Rivers, I. &amp; Nieser, N. (1972) The Ambrysus of Surinam, with the description of a new species, A. bifidus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 29, 1 - 8." type="journal article" year="1972">La Rivers &amp; Nieser 1972</bibRefCitation>
):
<emphasis id="9AE0791F522BFFC757E25273027CF85A" bold="true" box="[765,872,1897,1922]" pageId="2" pageNumber="281">
<collectingCountry id="D083E59D522BFFC757E25273027CF85A" box="[765,872,1897,1922]" name="Brazil" pageId="2" pageNumber="281">BRAZIL</collectingCountry>
</emphasis>
, Amazonas, Pará;
<emphasis id="9AE0791F522BFFC7515A527305DFF85A" bold="true" box="[1093,1227,1897,1922]" pageId="2" pageNumber="281">
<collectingCountry id="D083E59D522BFFC7515A527305DFF85A" box="[1093,1227,1897,1922]" name="Suriname" pageId="2" pageNumber="281">SURINAM</collectingCountry>
</emphasis>
, Zanderijsavanne. (
<bibRefCitation id="CC05D8FC522BFFC7558152970086F87E" author="Pereira" box="[158,402,1933,1958]" pageId="2" pageNumber="281" refString="Pereira, D. L. V. &amp; Melo, A. L. de (2007) Heteropteros (Insecta) aquaticos e semi-aquaticos de Pitinga, Amazonas, Brasil. Acta Amazonica, 37 (4), 643 - 648. http: // dx. doi. org / 10.1590 / S 0044 - 59672007000400021" type="journal article" year="2007">Pereira &amp; Melo 2007</bibRefCitation>
):
<collectingCountry id="D083E59D522BFFC754BA52970304F87E" box="[421,528,1933,1958]" name="Brazil" pageId="2" pageNumber="281">
<emphasis id="9AE0791F522BFFC754BA52970304F87E" bold="true" box="[421,528,1933,1958]" pageId="2" pageNumber="281">BRAZIL</emphasis>
</collectingCountry>
, Amazonas, Waimiri-Atroari Biological Reserve.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="A06B322A9E66FFC5580E5BBAA9CA4188">Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica</mods:title>
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<subSubSection id="E08EF686522AFFC6558853A2003AF92C" pageId="3" pageNumber="282" type="nomenclature">
<paragraph id="A82BA50D522AFFC6558853A2020CF90B" blockId="3.[151,792,1720,1780]" box="[151,792,1720,1747]" pageId="3" pageNumber="282">
<heading id="F3631261522AFFC6558853A2020CF90B" bold="true" box="[151,792,1720,1747]" fontSize="11" level="1" pageId="3" pageNumber="282" reason="1">
<emphasis id="9AE0791F522AFFC6558853A2020CF90B" bold="true" box="[151,792,1720,1747]" pageId="3" pageNumber="282">
<taxonomicName id="6F94DE8E522AFFC6558853A20357F90B" ID-CoL="7QM54" authority="Sites and Reynoso" authorityName="Sites and Reynoso" box="[151,579,1720,1747]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="maya" status="sp. nov.">
<emphasis id="9AE0791F522AFFC6558853A2004CF90A" bold="true" box="[151,344,1720,1746]" italics="true" pageId="3" pageNumber="282">Ambrysus maya</emphasis>
Sites and Reynoso
</taxonomicName>
,
<taxonomicNameLabel id="81D3C464522AFFC6574F53A3020CF90B" box="[592,792,1721,1747]" pageId="3" pageNumber="282" rank="species">NEW SPECIES</taxonomicNameLabel>
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<paragraph id="A82BA50D522AFFC6558853C1003AF92C" blockId="3.[151,792,1720,1780]" box="[151,302,1755,1780]" pageId="3" pageNumber="282">
(
<figureCitation id="30AFB988522AFFC6558053C101F8F92C" box="[159,236,1755,1780]" captionStart="FIGURE 1" captionStartId="2.[151,250,1189,1211]" captionTargetBox="[151,1436,198,1167]" captionTargetId="figure@2.[151,1436,198,1167]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species in the Ambrysus stali species complex." httpUri="https://zenodo.org/record/237550/files/figure.png" pageId="3" pageNumber="282">Figs. 1</figureCitation>
,
<figureCitation id="30AFB988522AFFC655E553C60032F92C" box="[250,294,1756,1780]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1560,1582]" captionStartId-1="5.[151,250,1896,1918]" captionTargetBox-0="[151,1436,193,1538]" captionTargetBox-1="[151,1436,1271,1850]" captionTargetId-0="figure@4.[151,1436,193,1538]" captionTargetId-1="figure@5.[151,1436,1271,1850]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." captionText-1="FIGURE 4. Ambrysus maya n. sp. (A) Río Bravo, the type locality with habitat inset, (B) live specimen in Río Bravo." httpUri-0="https://zenodo.org/record/237552/files/figure.png" httpUri-1="https://zenodo.org/record/237553/files/figure.png" pageId="3" pageNumber="282">3–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522AFFC3558852390228FD28" lastPageId="6" lastPageNumber="285" pageId="3" pageNumber="282" type="description">
<paragraph id="A82BA50D522AFFC65588523905FEF828" blockId="3.[151,1437,1827,2032]" pageId="3" pageNumber="282">
<emphasis id="9AE0791F522AFFC6558852390333F8E4" bold="true" box="[151,551,1827,1852]" pageId="3" pageNumber="282">Description. Macropterous male.</emphasis>
<typeStatus id="772F1BAF522AFFC6572D523E03DDF8E4" box="[562,713,1828,1852]" pageId="3" pageNumber="282" type="holotype">HOLOTYPE</typeStatus>
, length 7.84; maximum width 4.40.
<typeStatus id="772F1BAF522AFFC65168523E05F0F8E4" box="[1143,1252,1828,1852]" pageId="3" pageNumber="282" type="paratype">Paratypes</typeStatus>
(n = 2), length 7.84–7.92 (mean = 7.88); maximum width 4.32–4.52 (mean = 4.42). General shape fusiform; widest across abdominal segment III (
<figureCitation id="30AFB988522AFFC654AF527100E2F85C" box="[432,502,1899,1924]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="3" pageNumber="282">Fig. 3</figureCitation>
A). Overall dorsal coloration anteriorly pale yellow with dark brown pattern on forelegs, head, and pronotum (
<figureCitation id="30AFB988522AFFC6571C5295035FF870" box="[515,587,1935,1960]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="3" pageNumber="282">Fig. 3</figureCitation>
A); anterior pale yellow has greenish tone in live specimens (
<figureCitation id="30AFB988522AFFC6502E5295046EF870" box="[1329,1402,1935,1960]" captionStart="FIGURE 4" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[151,1436,1271,1850]" captionTargetId="figure@5.[151,1436,1271,1850]" captionTargetPageId="5" captionText="FIGURE 4. Ambrysus maya n. sp. (A) Río Bravo, the type locality with habitat inset, (B) live specimen in Río Bravo." httpUri="https://zenodo.org/record/237553/files/figure.png" pageId="3" pageNumber="282">Fig. 4</figureCitation>
B); posteriorly with dark brown scutellum and hemelytra. Exposed connexiva with alternating creamy white and dark checkered appearance. Dorsal surface coarsely punctate. Ventrally, mostly brown, lighter centrally.
</paragraph>
<caption id="FCEBF585522DFFC1558853020089F952" httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="4" pageNumber="283" targetBox="[151,1436,193,1538]" targetPageId="4">
<paragraph id="A82BA50D522DFFC1558853020089F952" blockId="4.[151,1436,1560,1674]" pageId="4" pageNumber="283">
<emphasis id="9AE0791F522DFFC1558853020006F9F5" bold="true" box="[151,274,1560,1582]" pageId="4" pageNumber="283">FIGURE 3.</emphasis>
Structures of
<taxonomicName id="6F94DE8E522DFFC154BB53020351F9F6" box="[420,581,1560,1582]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="maya" status="sp. nov.">
<emphasis id="9AE0791F522DFFC154BB53020351F9F6" box="[420,581,1560,1582]" italics="true" pageId="4" pageNumber="283">Ambrysus maya</emphasis>
</taxonomicName>
<emphasis id="9AE0791F522DFFC1575353020393F9F5" bold="true" box="[588,647,1560,1581]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="81D3C464522DFFC1575353020393F9F5" box="[588,647,1560,1581]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A) dorsal habitus of holotype, (B) 6th and 7th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape).
</paragraph>
</caption>
<paragraph id="A82BA50D522DFFC155D853B5028AF8A4" blockId="4.[151,1437,1711,2024]" pageId="4" pageNumber="283">
<emphasis id="9AE0791F522DFFC155D853B50010F910" box="[199,260,1711,1736]" italics="true" pageId="4" pageNumber="283">Head</emphasis>
. Head length 1.36; maximum width 1.98. Eyes convergent, synthlipsis 0.72; thin band of cuticle posterior to eye; eyes flat and not raised above level of vertex, slightly raised above level of pronotum. Anterior margin between eyes convex, extending anteriorly in front of eyes 7% of head length; posterior margin between eyes strongly convex, extending posteriorly 30% of head length. Labrum width 1.8 x length, evenly rounded. Labium brown, with three visible segments, extending 0.48 beyond labrum. Antennal segment proportions 2:9:11:6, length 0.54, elongate hairs on segment 4 and distal 2/3 of 3.
</paragraph>
<paragraph id="A82BA50D522DFFC055D8529D00FBFDE4" blockId="4.[151,1437,1711,2024]" lastBlockId="5.[151,1437,151,1221]" lastPageId="5" lastPageNumber="284" pageId="4" pageNumber="283">
<emphasis id="9AE0791F522DFFC155D8529D000CF878" box="[199,280,1927,1952]" italics="true" pageId="4" pageNumber="283">Thorax</emphasis>
. Pronotum mostly pale yellowish brown with dark brown punctures, anteromedial brown maculation, irregular longitudinal brown stripes, some longitudinal stripes enhanced with dark punctures; slightly rugose; lateral margins convex; sparse erect setae; explanate, strongly convergent anteriorly; posterior margin nearly straight; anterior margin deeply concave receiving convex posterior margin of head; transverse line of punctures defining anterior margin of band in posterior 1/4; posterolateral corners obtuse, rounded; width 3.2 x length; length at midline 1.22; maximum width at posterolateral corners 3.92. Prothorax ventrally brown and pruinose medially, with yellow glabrous lateral band; apices of propleura meeting broadly at midline, separated from level of prosternellum. Probasisternum brown, with sharp medial carina and generally appearing pruinose, row of elongate pale hairs lateral to carina. Prosternellum light brown, extending beneath apices of propleura. Elongate setae along posterior margin of propleuron. Scutellum mostly brown with yellow anterior corners and apex, triangular, densely punctate, width 1.9 x length, width 2.60, length 1.36. Hemelytra densely punctate. Claval commissure length 0.72. Embolium length 2.30, greatest width 0.62; lateral margin slightly convex in proximal half, more pronounced in distal half.
<collectingCountry id="D083E59D522CFFC0540A54C60059FE2D" box="[277,333,476,501]" name="India" pageId="5" pageNumber="284">Hind</collectingCountry>
wings well developed. Mesobasisternum with midventral longitudinal tumescence with sulcus on midline; tumescence terminating posteriorly as acute mesosternellum. Metasternellum (=metaxyphus) transverse, subtriangular, with apex acute.
</paragraph>
<paragraph id="A82BA50D522CFFC055D85753007DFB1C" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="284">
<emphasis id="9AE0791F522CFFC055D8575301E8FDB8" box="[199,252,585,608]" italics="true" pageId="5" pageNumber="284">Legs</emphasis>
. All leg segments pale yellowish brown, darker distally on fore and hind legs. Profemur pale yellowish brown with dark brown along anterior margin continuous to apex; dark brown posterior marking extending anteriorly, becoming discontinuous where it approaches or meets anterior margin dark marking; row of 10–12 combs of short brown spines in basal half of posterior margin; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with inner surface flattened and with spatulate setae; tarsus immovable, onesegmented; pretarsal claw single, minute, triangular. Procoxa with dark brown basal marking on lateral to anterior surfaces, cluster of stout brown anteromedial spines. Mesocoxa partially recessed into thorax. Metacoxa with longitudinal sulcus that can accommodate flexed metafemur. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal 1/3 and single on mesofemur, combs of 1–4 spines irregularly spaced along full length of metafemur. Mesotibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines, dorsolateral and dorsomedial rows include combs of up to 4 spines; stout spines at apical rim dorsally, two transverse rows of stout spines at apex of ventral margin. Metatibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines, transverse rows of stout spines at apex of ventral margin. Meso- and metatibiae and -tarsi with long, pale swimming hairs; hairs sparse on mesotibia and -tarsus, profuse on metatibia and -tarsus. Meso- and metapretarsi with paired claws slender, curved in distal half, with basal tooth. Leg measurements as follows: foreleg, femur 1.86, tibia 1.60, tarsus 0.32; middle leg, femur 1.88, tibia 1.64, tarsomeres 1–3, 0.10, 0.28, 0.36; hind leg, femur 2.32, tibia 2.72, tarsomeres 1– 3, 0.16, 0.52, 0.47.
</paragraph>
<caption id="FCEBF585522CFFC0558852720452F8A6" box="[151,1350,1896,1918]" httpUri="https://zenodo.org/record/237553/files/figure.png" pageId="5" pageNumber="284" targetBox="[151,1436,1271,1850]" targetPageId="5">
<paragraph id="A82BA50D522CFFC0558852720452F8A6" blockId="5.[151,1350,1896,1918]" box="[151,1350,1896,1918]" pageId="5" pageNumber="284">
<emphasis id="9AE0791F522CFFC0558852720006F8A5" bold="true" box="[151,274,1896,1918]" pageId="5" pageNumber="284">FIGURE 4.</emphasis>
<taxonomicName id="6F94DE8E522CFFC05406527200AEF8A6" box="[281,442,1896,1918]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="maya" status="sp. nov.">
<emphasis id="9AE0791F522CFFC05406527200AEF8A6" box="[281,442,1896,1918]" italics="true" pageId="5" pageNumber="284">Ambrysus maya</emphasis>
</taxonomicName>
<emphasis id="9AE0791F522CFFC054DF527200EDF8A5" bold="true" box="[448,505,1896,1917]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="81D3C464522CFFC054DF527200EDF8A5" box="[448,505,1896,1917]" pageId="5" pageNumber="284" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A) Río Bravo, the type locality with habitat inset, (B) live specimen in Río Bravo.
</paragraph>
</caption>
<paragraph id="A82BA50D522FFFC355D8558D0571FDE4" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D8558D0025FF68" box="[199,305,151,176]" italics="true" pageId="6" pageNumber="285">Abdomen</emphasis>
. Abdomen dorsally with connexiva of III–VII exposed and with checkered appearance (
<figureCitation id="30AFB988522FFFC3502B558D046FFF68" box="[1332,1403,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[151,1436,1271,1850]" captionTargetId="figure@5.[151,1436,1271,1850]" captionTargetPageId="5" captionText="FIGURE 4. Ambrysus maya n. sp. (A) Río Bravo, the type locality with habitat inset, (B) live specimen in Río Bravo." httpUri="https://zenodo.org/record/237553/files/figure.png" pageId="6" pageNumber="285">Fig. 4</figureCitation>
B), posterior half of III–VII dark brown, anterior half of III–V creamy white and of VI–VII brown; VIII mostly brown; lateral margin with row of pale setae, group of trichobothria on dorsal surface near posterolateral corners. Posterolateral corner of III slightly acuminate, IV–VII spinose. Accessory genitalic process of VI broad, apical margin flattened with corners broadly rounded, broadly prolonged laterally (
<figureCitation id="30AFB988522FFFC35136543D0565FE98" box="[1065,1137,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="6" pageNumber="285">Fig. 3</figureCitation>
B). Medial lobes of VIII (pseudoparameres) with apical corners rounded, distal margin flat (
<figureCitation id="30AFB988522FFFC356B6545602FBFEBC" box="[937,1007,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="6" pageNumber="285">Fig. 3</figureCitation>
D). Ventrally mostly brown laterally, brownish yellow medially, pruinose, with pile of fine hairs. Laterosternites III–VI with yellow near middle of lateral margin. Lateral margin with thin glabrous band. Glabrous patches on laterosternites II–VII. Mediosternite V with posterior margin slightly asymmetrical. Phallosoma with left ventral lobe reflexed dorsad, unarmed with teeth; right ventral lobe appressed to body of phallosoma, unarmed (
<figureCitation id="30AFB988522FFFC3568E54C602C0FE2C" box="[913,980,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="6" pageNumber="285">Fig. 3</figureCitation>
C). Parameres subquadrate, pedunculate, symmetrical, mesal margin slightly convex with corners broadly rounded (
<figureCitation id="30AFB988522FFFC3511454E50547FDC0" box="[1035,1107,511,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="6" pageNumber="285">Fig. 3</figureCitation>
E). Pygophore with anterior margin between parameres slightly produced at midline, posterior apex with fine setae.
</paragraph>
<paragraph id="A82BA50D522FFFC355D8575D0228FD28" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D8575D00D1FDB8" bold="true" box="[199,453,583,608]" pageId="6" pageNumber="285">Macropterous female</emphasis>
.
<typeStatus id="772F1BAF522FFFC354CB57520355FDB8" box="[468,577,584,608]" pageId="6" pageNumber="285" type="paratype">Paratypes</typeStatus>
(n = 11), length 7.68–8.96 (mean = 8.45); maximum width 4.36–4.72 (mean = 4.55). Similar to male in general structure and coloration except as follows: Mediosternite V symmetrical. Mediosternite VII (subgenital plate) with medial convexity and distolateral production (
<figureCitation id="30AFB988522FFFC3516D579505A1FD70" box="[1138,1205,655,680]" captionStart="FIGURE 3" captionStartId="4.[151,250,1560,1582]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure@4.[151,1436,193,1538]" captionTargetPageId="4" captionText="FIGURE 3. Structures of Ambrysus maya n. sp. (A) dorsal habitus of holotype, (B) 6 th and 7 th abdominal terga of male, (C) ventral and left sides of phallosoma, (D) 8 th abdominal tergum showing medial pseudoparameres, (E) male pygophore with paired parameres (phallosoma removed), (F) terminal abdominal sterna of female showing subgenital plate (setae have been removed to reveal shape)." httpUri="https://zenodo.org/record/237552/files/figure.png" pageId="6" pageNumber="285">Fig. 3</figureCitation>
F). Laterosternite VI with posteromedial corner on left side produced into well-developed spine directed posteroventrad, posteromedial corner on right side produced, tuberculate, deflexed ventrad.
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522FFFC355D857E600CCFB1C" pageId="6" pageNumber="285" type="diagnosis">
<paragraph id="A82BA50D522FFFC355D857E60407FC58" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D857E60349FCCD" bold="true" box="[199,605,764,789]" pageId="6" pageNumber="285">Diagnosis and comparative notes</emphasis>
. This species can be distinguished from other members of the species complex by the significantly narrower fore femora (t=3.803, df=15, p=0.002) and its noncoincident geographic range. It can be distinguished from all other naucorids in Mesoamerica by its small size, fusiform body shape, and dark longitudinal markings of the pronotum and dark markings on the dorsal surface of the profemora.
</paragraph>
<paragraph id="A82BA50D522FFFC355D8569600CCFB1C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D8569600A4FC7D" bold="true" box="[199,432,908,933]" pageId="6" pageNumber="285">Habitat description</emphasis>
. At the
<typeStatus id="772F1BAF522FFFC3571256970329FC7D" box="[525,573,909,933]" pageId="6" pageNumber="285">type</typeStatus>
locality, specimens were collected from marginal vegetation including dead leaves, grasses, and other vegetation (
<figureCitation id="30AFB988522FFFC3575656B5039BFC10" box="[585,655,943,968]" captionStart="FIGURE 4" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[151,1436,1271,1850]" captionTargetId="figure@5.[151,1436,1271,1850]" captionTargetPageId="5" captionText="FIGURE 4. Ambrysus maya n. sp. (A) Río Bravo, the type locality with habitat inset, (B) live specimen in Río Bravo." httpUri="https://zenodo.org/record/237553/files/figure.png" pageId="6" pageNumber="285">Fig. 4</figureCitation>
A). The river was shallow (ca.
<quantity id="6F6C08E8522FFFC356E856AA0531FC10" box="[1015,1061,944,968]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="285" unit="m" value="1.0">1 m</quantity>
) with cobble substrate and swift current. In the riffles were the naucorids
<taxonomicName id="6F94DE8E522FFFC3577F56CE02EEFC35" authority="Montandon" authorityName="Montandon" box="[608,1018,980,1005]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="circumcinctus">
<emphasis id="9AE0791F522FFFC3577F56CE0266FC34" box="[608,882,980,1005]" italics="true" pageId="6" pageNumber="285">Ambrysus circumcinctus</emphasis>
Montandon
</taxonomicName>
and
<taxonomicName id="6F94DE8E522FFFC3512F56CF048CFC35" authority="Champion" authorityName="Champion" box="[1072,1432,980,1005]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="9AE0791F522FFFC3512F56CF040DFC35" box="[1072,1305,980,1005]" italics="true" pageId="6" pageNumber="285">Limnocoris insularis</emphasis>
Champion
</taxonomicName>
, and the byrrhoid beetles
<taxonomicName id="6F94DE8E522FFFC354A956ED0291FBC8" authority="LeConte" authorityName="LeConte" box="[438,901,1015,1040]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="9AE0791F522FFFC354A956ED0391FBC8" box="[438,645,1015,1040]" italics="true" pageId="6" pageNumber="285">Helichus suturalis</emphasis>
LeConte (Dryopidae)
</taxonomicName>
,
<taxonomicName id="6F94DE8E522FFFC3568C56ED05F8FBC8" authority="Sharp" authorityName="Sharp" box="[915,1260,1015,1040]" class="Insecta" family="Elmidae" genus="Phanocerus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="clavicornis">
<emphasis id="9AE0791F522FFFC3568C56ED058AFBC8" box="[915,1182,1015,1040]" italics="true" pageId="6" pageNumber="285">Phanocerus clavicornis</emphasis>
Sharp
</taxonomicName>
,
<taxonomicName id="6F94DE8E522FFFC351E556ED01F9FBED" authority="Sharp" authorityName="Sharp" class="Insecta" family="Elmidae" genus="Xenelmis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="bufo">
<emphasis id="9AE0791F522FFFC351E556ED0488FBC8" box="[1274,1436,1015,1040]" italics="true" pageId="6" pageNumber="285">Xenelmis bufo</emphasis>
(Sharp)
</taxonomicName>
,
<taxonomicName id="6F94DE8E522FFFC355E651060067FBED" box="[249,371,1052,1077]" class="Insecta" family="Elmidae" genus="Heterelmis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC355E651060067FBED" box="[249,371,1052,1077]" italics="true" pageId="6" pageNumber="285">Heterelmis</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="6F94DE8E522FFFC354B751060359FBED" box="[424,589,1052,1077]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC354B751060359FBED" box="[424,589,1052,1077]" italics="true" pageId="6" pageNumber="285">Hexacylloepus</emphasis>
</taxonomicName>
sp., and
<taxonomicName id="6F94DE8E522FFFC357AC51060274FBED" box="[691,864,1052,1077]" class="Insecta" family="Elmidae" genus="Microcylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC357AC51060274FBED" box="[691,864,1052,1077]" italics="true" pageId="6" pageNumber="285">Microcylloepus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="6F94DE8E522FFFC35689510602EEFBED" box="[918,1018,1052,1077]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
). Specimens of
<taxonomicName id="6F94DE8E522FFFC351B35106044AFBEC" box="[1196,1374,1052,1077]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="9AE0791F522FFFC351B35106044AFBEC" box="[1196,1374,1052,1077]" italics="true" pageId="6" pageNumber="285">Ambrysus maya</emphasis>
</taxonomicName>
from the Sibun River in
<collectingCountry id="D083E59D522FFFC35474512500A7FB80" box="[363,435,1087,1112]" name="Belize" pageId="6" pageNumber="285">Belize</collectingCountry>
were collected mostly from vegetational debris caught on branches or along the margins. Other insect species in the Sibun River included
<taxonomicName id="6F94DE8E522FFFC357D1517E0219FBA3" box="[718,781,1124,1147]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC357D1517E0219FBA3" box="[718,781,1124,1147]" italics="true" pageId="6" pageNumber="285">Argia</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="6F94DE8E522FFFC35658517E02EBFBA5" box="[839,1023,1124,1149]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="285" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
);
<taxonomicName id="6F94DE8E522FFFC3510A517E01CFFB78" authority="Sharp" authorityName="Sharp" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="subSpecies" species="oscillator" subSpecies="laevipennis">
<emphasis id="9AE0791F522FFFC3510A517E0488FBA5" box="[1045,1436,1124,1149]" italics="true" pageId="6" pageNumber="285">Laccophilus oscillator laevipennis</emphasis>
Sharp
</taxonomicName>
,
<taxonomicName id="6F94DE8E522FFFC355F951930325FB78" authority="Say" authorityName="Say" box="[230,561,1159,1184]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="9AE0791F522FFFC355F95193007FFB78" box="[230,363,1161,1184]" italics="true" pageId="6" pageNumber="285">L. proximus</emphasis>
Say (Dytiscidae)
</taxonomicName>
;
<taxonomicName id="6F94DE8E522FFFC35723519D02A9FB78" authority="Wiedemann" authorityName="Wiedemann" box="[572,957,1159,1184]" class="Insecta" family="Culicidae" genus="Anopheles" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="albimanus">
<emphasis id="9AE0791F522FFFC35723519D0239FB78" box="[572,813,1159,1184]" italics="true" pageId="6" pageNumber="285">Anopheles albimanus</emphasis>
Wiedemann
</taxonomicName>
, and
<taxonomicName id="6F94DE8E522FFFC356E8519D0488FB78" authority="Dyar" authorityName="Dyar" box="[1015,1436,1159,1184]" class="Insecta" family="Culicidae" genus="Chagasia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="bathana">
<emphasis id="9AE0791F522FFFC356E8519D05D2FB78" box="[1015,1222,1159,1184]" italics="true" pageId="6" pageNumber="285">Chagasia bathana</emphasis>
(Dyar) (Culicidae)
</taxonomicName>
(R. Vogtsberger, pers. com.).
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522FFFC355D851D50212F964" pageId="6" pageNumber="285" type="discussion">
<paragraph id="A82BA50D522FFFC355D851D50212F964" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D851D50058FB30" bold="true" box="[199,332,1231,1256]" pageId="6" pageNumber="285">Discussion.</emphasis>
Although the Central American countries south of
<collectingCountry id="D083E59D522FFFC356BF51D5050DFB30" box="[928,1049,1231,1256]" name="Guatemala" pageId="6" pageNumber="285">Guatemala</collectingCountry>
have been well-collected and the fauna studied by many scientists over the past century beginning with Champion (1901),
<taxonomicName id="6F94DE8E522FFFC351BD51EE0415FAD4" box="[1186,1281,1268,1292]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="9AE0791F522FFFC351BD51EE0415FAD4" box="[1186,1281,1268,1292]" italics="true" pageId="6" pageNumber="285">A. maya</emphasis>
</taxonomicName>
has not been reported from there. Thus, this species appears to be widely separated from
<taxonomicName id="6F94DE8E522FFFC356E350020568FAE8" box="[1020,1148,1304,1328]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F522FFFC356E350020568FAE8" box="[1020,1148,1304,1328]" italics="true" pageId="6" pageNumber="285">A. tricuspis</emphasis>
</taxonomicName>
in
<collectingCountry id="D083E59D522FFFC351BB500D0401FAE8" box="[1188,1301,1303,1328]" name="Colombia" pageId="6" pageNumber="285">Colombia</collectingCountry>
, the nearest known member of the
<taxonomicName id="6F94DE8E522FFFC35487502600FCFA8D" box="[408,488,1340,1365]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="9AE0791F522FFFC35487502600FCFA8D" box="[408,488,1340,1365]" italics="true" pageId="6" pageNumber="285">A. stali</emphasis>
</taxonomicName>
complex (
<figureCitation id="30AFB988522FFFC3577F502603B6FA8C" box="[608,674,1340,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1189,1211]" captionTargetBox="[151,1436,198,1167]" captionTargetId="figure@2.[151,1436,198,1167]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species in the Ambrysus stali species complex." httpUri="https://zenodo.org/record/237550/files/figure.png" pageId="6" pageNumber="285">Fig. 1</figureCitation>
). The majority of insects in Central
<collectingCountry id="D083E59D522FFFC3515E502605B5FA8D" box="[1089,1185,1340,1365]" name="United States of America" pageId="6" pageNumber="285">America</collectingCountry>
are suggested to have originated from taxonomic elements of northern South
<collectingCountry id="D083E59D522FFFC3563B50450290FAA0" box="[804,900,1375,1400]" name="United States of America" pageId="6" pageNumber="285">America</collectingCountry>
(
<bibRefCitation id="CC05D8FC522FFFC356875045052CFAA0" author="Kimsey" box="[920,1080,1375,1400]" pageId="6" pageNumber="285" refString="Kimsey, L. S. (1992) Biogeography of the Panamanian region, from an insect perspective, In: Quintero, D. &amp; Aiello, A. (Eds.), Insects in Panama and Mesoamerica: Selected Studies. Oxford University Press, Oxford, U. K., pp. 14 - 24." type="book chapter" year="1992">Kimsey 1992</bibRefCitation>
).
<taxonomicName id="6F94DE8E522FFFC3514E5045041CFAA0" box="[1105,1288,1375,1400]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="9AE0791F522FFFC3514E5045041CFAA0" box="[1105,1288,1375,1400]" italics="true" pageId="6" pageNumber="285">Ambrysus maya</emphasis>
</taxonomicName>
follows this pattern since most of the diversity of this species complex is distributed throughout the
<collectingCountry id="D083E59D522FFFC35171509E05D1FA44" box="[1134,1221,1412,1436]" name="Guyana" pageId="6" pageNumber="285">Guyana</collectingCountry>
Shield region with a single incursion into Mesoamerica. Alternatively, the ancestor of this complex might have moved southward from the Mexican geographic center of
<taxonomicName id="6F94DE8E522FFFC3576D50D603F5FA3D" box="[626,737,1484,1509]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC3576D50D603F5FA3D" box="[626,737,1484,1509]" italics="true" pageId="6" pageNumber="285">Ambrysus</emphasis>
</taxonomicName>
diversity, and subsequently became discontinuous with the current species through vicariance. Substantially more geological time for either scenario to have occurred than was previously considered was shown recently.
<bibRefCitation id="CC05D8FC522FFFC357AB530F028EF9F5" box="[692,922,1556,1581]" pageId="6" pageNumber="285" refString="Montes, C., Cardona, A., Jaramillo, C., Pardo, A., Silva, J. C., Valencia, V., Ayala, C., Perez-Angel, L. C., Rodriguez-Parra, L. A., Ramirez, V. &amp; Nino, H. (2015) Middle Miocene closure of the Central American Seaway. Science, 348, 226 - 229. http: // dx. doi. org / 10.1126 / science. aaa 2815" type="journal article">Montes et al. (2015)</bibRefCitation>
provided evidence that the Central American land bridge formed in the Miocene epoch 10–12 million years earlier than the 3 million year Pliocene model that had been widely accepted. Moreover, a phylogenetic analysis of New World naucorid groups with attention to divergence times may be able to evaluate invasion patterns. The diversity of
<taxonomicName id="6F94DE8E522FFFC356F35365054FF940" box="[1004,1115,1663,1688]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC356F35365054FF940" box="[1004,1115,1663,1688]" italics="true" pageId="6" pageNumber="285">Ambrysus</emphasis>
</taxonomicName>
is greatest in North
<collectingCountry id="D083E59D522FFFC3502353650488F940" box="[1340,1436,1663,1688]" name="United States of America" pageId="6" pageNumber="285">America</collectingCountry>
and of
<taxonomicName id="6F94DE8E522FFFC355FB53BE0068F965" box="[228,380,1700,1725]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC355FB53BE0068F965" box="[228,380,1700,1725]" italics="true" pageId="6" pageNumber="285">Cryphocricos</emphasis>
</taxonomicName>
and
<taxonomicName id="6F94DE8E522FFFC354AC53BF0326F964" box="[435,562,1701,1724]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="9AE0791F522FFFC354AC53BF0326F964" box="[435,562,1701,1724]" italics="true" pageId="6" pageNumber="285">Limnocoris</emphasis>
</taxonomicName>
in South
<collectingCountry id="D083E59D522FFFC3578053BE03EBF965" box="[671,767,1700,1725]" name="United States of America" pageId="6" pageNumber="285">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522FFFC355D853DD0202F84C" pageId="6" pageNumber="285" type="etymology">
<paragraph id="A82BA50D522FFFC355D853DD03D3F8DD" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D853DD0052F938" bold="true" box="[199,326,1735,1760]" pageId="6" pageNumber="285">Etymology</emphasis>
. The specific epithet is a noun in apposition in reference to the
<taxonomicName id="6F94DE8E522FFFC3510853D30543F939" box="[1047,1111,1737,1761]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="290" phylum="Arthropoda" rank="species" species="maya">Maya</taxonomicName>
civilization, which occupied this region of Mesoamerica until the 17th century.
</paragraph>
<paragraph id="A82BA50D522FFFC355D852150202F84C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D85215004FF8F0" bold="true" box="[199,347,1807,1832]" pageId="6" pageNumber="285">Repositories</emphasis>
. The
<typeStatus id="772F1BAF522FFFC35485521500E8F8F0" box="[410,508,1807,1832]" pageId="6" pageNumber="285" type="holotype">holotype</typeStatus>
and some
<typeStatus id="772F1BAF522FFFC3576B520B03CBF8F1" box="[628,735,1809,1833]" pageId="6" pageNumber="285" type="paratype">paratypes</typeStatus>
are deposited in the Enns Entomology Museum (University of Missouri); additional
<typeStatus id="772F1BAF522FFFC35497522F00E0F895" box="[392,500,1845,1869]" pageId="6" pageNumber="285" type="paratype">paratypes</typeStatus>
are deposited in Midwestern State University (Wichita Falls), Universidad del Valle de
<collectingCountry id="D083E59D522FFFC355A7524D0025F8A8" box="[184,305,1879,1904]" name="Guatemala" pageId="6" pageNumber="285">Guatemala</collectingCountry>
(
<collectingCountry id="D083E59D522FFFC3545F524D00AFF8A8" box="[320,443,1879,1904]" name="Guatemala" pageId="6" pageNumber="285">Guatemala</collectingCountry>
City), Universidad Nacional Autónoma de
<collectingCountry id="D083E59D522FFFC356BB524D02EEF8A8" box="[932,1018,1879,1904]" name="Mexico" pageId="6" pageNumber="285">México</collectingCountry>
(
<collectingCountry id="D083E59D522FFFC35117524D0574F8A8" box="[1032,1120,1879,1904]" name="Mexico" pageId="6" pageNumber="285">Mexico</collectingCountry>
City), and the
<collectingCountry id="D083E59D522FFFC35018524D0488F8A8" box="[1287,1436,1879,1904]" name="United States of America" pageId="6" pageNumber="285">United States</collectingCountry>
National Museum of Natural History (Washington D.C.).
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522FFFC255D852BA02ADFE2C" lastPageId="7" lastPageNumber="286" pageId="6" pageNumber="285" type="materials_examined">
<paragraph id="A82BA50D522FFFC255D852BA02ADFE2C" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,151,501]" lastPageId="7" lastPageNumber="286" pageId="6" pageNumber="285">
<emphasis id="9AE0791F522FFFC355D852BA00FBF861" bold="true" box="[199,495,1952,1977]" pageId="6" pageNumber="285">
<typeStatus id="772F1BAF522FFFC355D852BA0016F861" box="[199,258,1952,1977]" pageId="6" pageNumber="285">Type</typeStatus>
material examined
</emphasis>
.
<typeStatus id="772F1BAF522FFFC3571F52BA0380F860" box="[512,660,1952,1976]" pageId="6" pageNumber="285" type="holotype">HOLOTYPE</typeStatus>
♂:
<emphasis id="9AE0791F522FFFC357D952BA023AF861" bold="true" box="[710,814,1952,1977]" pageId="6" pageNumber="285">
<collectingCountry id="D083E59D522FFFC357D952BA023AF861" box="[710,814,1952,1977]" name="Belize" pageId="6" pageNumber="285">BELIZE</collectingCountry>
</emphasis>
:
<emphasis id="9AE0791F522FFFC3565E52BA055EF861" bold="true" box="[833,1098,1952,1977]" pageId="6" pageNumber="285">Orange Walk District</emphasis>
: Río Bravo ca.
<quantity id="6F6C08E8522FFFC3501652BA0445F861" box="[1289,1361,1952,1977]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="6" pageNumber="285" unit="mi" value="10.0">10 mi</quantity>
. N. of Gallon Jug at Cedar Crossing,
<date id="DC2A83CD522FFFC354F052DE0384F804" box="[495,656,1988,2013]" pageId="6" pageNumber="285" value="2014-04-19">19 April 2014</date>
, 44 m,
<geoCoordinate id="CDA0C3CA522FFFC357FD52DE0263F805" box="[738,887,1988,2013]" direction="north" orientation="latitude" pageId="6" pageNumber="285" precision="1" value="17.69025">17°41.415´N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522FFFC3569D52DE050EF805" box="[898,1050,1988,2013]" direction="west" orientation="longitude" pageId="6" pageNumber="285" precision="1" value="-89.02692">89°01.615´W</geoCoordinate>
, R. Sites &amp; W. Shepard, vegetated margins, L-1740;
<typeStatus id="772F1BAF522EFFC2546C55820306FF68" box="[371,530,152,176]" pageId="7" pageNumber="286" type="paratype">PARATYPES</typeStatus>
: same data as
<typeStatus id="772F1BAF522EFFC257C7558D022EFF68" box="[728,826,151,176]" pageId="7" pageNumber="286" type="holotype">holotype</typeStatus>
(1♀);
<emphasis id="9AE0791F522EFFC25687558D055FFF68" bold="true" box="[920,1099,151,176]" pageId="7" pageNumber="286">
<collectingCountry id="D083E59D522EFFC25687558D02F4FF68" box="[920,992,151,176]" name="Belize" pageId="7" pageNumber="286">Belize</collectingCountry>
District
</emphasis>
: Sibun River,
<geoCoordinate id="CDA0C3CA522EFFC25015558D0483FF68" box="[1290,1431,151,176]" direction="north" orientation="latitude" pageId="7" pageNumber="286" precision="1" value="17.4115">17º24.690'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522EFFC2558855A60022FF0D" box="[151,310,188,213]" direction="west" orientation="longitude" pageId="7" pageNumber="286" precision="1" value="-88.42146">088º25.288'W</geoCoordinate>
,
<date id="DC2A83CD522EFFC2545555A600CDFF0C" box="[330,473,188,212]" pageId="7" pageNumber="286" value="2009-07-01">1 VII 2009</date>
, bamboo &amp; branches, coll J. H. Adams (1♀); same data except
<geoCoordinate id="CDA0C3CA522EFFC2501555A6048CFF0C" box="[1290,1432,188,212]" direction="north" orientation="latitude" pageId="7" pageNumber="286" precision="1" value="17.404684">17º24.281'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522EFFC2558855C50022FF20" box="[151,310,223,248]" direction="west" orientation="longitude" pageId="7" pageNumber="286" precision="1" value="-88.43353">088º26.012'W</geoCoordinate>
(1♀); same data except
<geoCoordinate id="CDA0C3CA522EFFC2576055FA0219FF20" box="[639,781,224,248]" direction="north" orientation="latitude" pageId="7" pageNumber="286" precision="1" value="17.4117">17º24.702'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522EFFC2563C55C502D6FF20" box="[803,962,223,248]" direction="west" orientation="longitude" pageId="7" pageNumber="286" precision="1" value="-88.421616">088º25.297'W</geoCoordinate>
(
<specimenCount id="BE926E84522EFFC256C155FA0517FF20" box="[990,1027,224,248]" pageId="7" pageNumber="286" type="generic">1♂</specimenCount>
); same data except
<geoCoordinate id="CDA0C3CA522EFFC2501455C5048CFF20" box="[1291,1432,223,248]" direction="north" orientation="latitude" pageId="7" pageNumber="286" precision="1" value="17.268534">17º16.112'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522EFFC25588541E0022FEC5" box="[151,310,260,285]" direction="west" orientation="longitude" pageId="7" pageNumber="286" precision="1" value="-88.571266">088º34.276'W</geoCoordinate>
,
<date id="DC2A83CD522EFFC2545C541E00DDFEC4" box="[323,457,260,284]" pageId="7" pageNumber="286" value="2009-06-29">29 VI 2009</date>
, submerged stones (1♀);
<emphasis id="9AE0791F522EFFC257F0541E0284FEC5" bold="true" box="[751,912,260,285]" pageId="7" pageNumber="286">Cayo District</emphasis>
: Sibun River, 0338582N, 1916372W, detritus patch,
<date id="DC2A83CD522EFFC255FB543D0082FE98" box="[228,406,295,320]" pageId="7" pageNumber="286" value="2009-06-26" valueMax="2009-06-30" valueMin="2009-06-26">26–30 VI 2009</date>
, coll J. H. Adams (1♀); same data except 0333330N, 1912078W (1♀); same data except 0348708N, 1925658W (1♀); same data except 0338622N, 1916554W (1♀); same data except 033283N, 1911970W (1♀).
<emphasis id="9AE0791F522EFFC25443547503ABFE50" bold="true" box="[348,703,367,392]" pageId="7" pageNumber="286">
<collectingCountry id="D083E59D522EFFC2544354750303FE50" box="[348,535,367,392]" name="Guatemala" pageId="7" pageNumber="286">GUATEMALA</collectingCountry>
: Alta Verapaz
</emphasis>
: Lag[una]. Lachua,
<date id="DC2A83CD522EFFC256BC546A053AFE50" box="[931,1070,367,392]" pageId="7" pageNumber="286" value="1995-07-19">19 VII 1995</date>
, J. C. Schuster (1♀).
<emphasis id="9AE0791F522EFFC25003547501EDFE75" bold="true" pageId="7" pageNumber="286">
<collectingCountry id="D083E59D522EFFC2500354750483FE50" box="[1308,1431,367,392]" name="Mexico" pageId="7" pageNumber="286">MEXICO</collectingCountry>
: Chiapas
</emphasis>
: Chajal Arroyo Manzana,
<date id="DC2A83CD522EFFC25738548E03BEFE74" box="[551,682,404,428]" pageId="7" pageNumber="286" value="2008-12-09">9-XII-2008</date>
, G. Mendez / Colección del Instituto de Biología,
<collectionCode id="CE853DC8522EFFC251FD548E0423FE74" box="[1250,1335,404,428]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="7" pageNumber="286">UNAM</collectionCode>
<collectingCountry id="D083E59D522EFFC25020548E048CFE75" box="[1343,1432,404,429]" name="Mexico" pageId="7" pageNumber="286">México</collectingCountry>
, D.F. (1♀);
<emphasis id="9AE0791F522EFFC2540754AD0063FE08" bold="true" box="[280,375,439,464]" pageId="7" pageNumber="286">Tabasco</emphasis>
: Río Carrizal, Est. No. 7,
<date id="DC2A83CD522EFFC257AD54AD022DFE08" box="[690,825,439,464]" pageId="7" pageNumber="286" value="1996-06-10">10-VI-1996</date>
,
<geoCoordinate id="CDA0C3CA522EFFC2565854AD02D3FE08" box="[839,967,439,464]" direction="north" orientation="latitude" pageId="7" pageNumber="286" precision="9" value="17.953833">17º57.23'N</geoCoordinate>
,
<geoCoordinate id="CDA0C3CA522EFFC256C954A2054DFE08" box="[982,1113,440,464]" direction="west" orientation="longitude" pageId="7" pageNumber="286" precision="9" value="-93.1195">93º07.17'W</geoCoordinate>
, J. Bueno, A. Contreras, R. Barba / Colección del Instituto de Biología,
<collectionCode id="CE853DC8522EFFC2579954C603CFFE2C" box="[646,731,476,500]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="7" pageNumber="286">UNAM</collectionCode>
<collectingCountry id="D083E59D522EFFC257FD54C6022EFE2D" box="[738,826,476,501]" name="Mexico" pageId="7" pageNumber="286">México</collectingCountry>
, D.F. (
<specimenCount id="BE926E84522EFFC2569954C602BFFE2C" box="[902,939,476,500]" pageId="7" pageNumber="286" type="generic">1♂</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

221
data/20/3D/14/203D141B522EFFCD5588524102CBFB30.xml Normal file
View file

@ -0,0 +1,221 @@
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<mods:title id="1623595252C0A9B30577A9F2487366A7">Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica</mods:title>
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<treatment id="203D141B522EFFCD5588524102CBFB30" ID-DOI="http://doi.org/10.5281/zenodo.6119247" ID-GBIF-Taxon="119636075" ID-Zenodo-Dep="6119247" LSID="urn:lsid:plazi:treatment:203D141B522EFFCD5588524102CBFB30" httpUri="http://treatment.plazi.org/id/203D141B522EFFCD5588524102CBFB30" lastPageId="8" lastPageNumber="287" pageId="7" pageNumber="286">
<subSubSection id="E08EF686522EFFC2558852410006F84F" pageId="7" pageNumber="286" type="nomenclature">
<paragraph id="A82BA50D522EFFC25588524100F8F8AE" blockId="7.[151,492,1883,1943]" box="[151,492,1883,1910]" pageId="7" pageNumber="286">
<heading id="F3631261522EFFC25588524100F8F8AE" bold="true" box="[151,492,1883,1910]" fontSize="11" level="1" pageId="7" pageNumber="286" reason="1">
<taxonomicName id="6F94DE8E522EFFC25588524100F8F8AE" ID-CoL="7QM5X" authority="La Rivers" authorityName="La Rivers" box="[151,492,1883,1910]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F522EFFC25588524100F8F8AE" bold="true" box="[151,492,1883,1910]" pageId="7" pageNumber="286">
<emphasis id="9AE0791F522EFFC255885241007EF8AD" bold="true" box="[151,362,1883,1909]" italics="true" pageId="7" pageNumber="286">Ambrysus scolius</emphasis>
La Rivers
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A82BA50D522EFFC2558852640006F84F" blockId="7.[151,492,1883,1943]" box="[151,274,1918,1943]" pageId="7" pageNumber="286">
(
<figureCitation id="30AFB988522EFFC25580526401F8F84F" box="[159,236,1918,1943]" captionStart="FIGURE 1" captionStartId="2.[151,250,1189,1211]" captionTargetBox="[151,1436,198,1167]" captionTargetId="figure@2.[151,1436,198,1167]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species in the Ambrysus stali species complex." httpUri="https://zenodo.org/record/237550/files/figure.png" pageId="7" pageNumber="286">Figs. 1</figureCitation>
,
<figureCitation id="30AFB988522EFFC255E55264001CF84F" box="[250,264,1918,1943]" captionStart="FIGURE 5" captionStartId="7.[151,250,1764,1786]" captionTargetBox="[160,1420,555,1717]" captionTargetId="figure@7.[160,1420,555,1718]" captionTargetPageId="7" captionText="FIGURE 5. Ambrysus scolius female paratype. (A) dorsal habitus, (B) terminal abdominal sterna showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing)." httpUri="https://zenodo.org/record/237554/files/figure.png" pageId="7" pageNumber="286">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08EF686522EFFC2558852DA03F3F80E" box="[151,743,1984,2006]" pageId="7" pageNumber="286" type="reference_group">
<paragraph id="A82BA50D522EFFC2558852DA03F3F80E" blockId="7.[151,743,1984,2006]" box="[151,743,1984,2006]" pageId="7" pageNumber="286">
<taxonomicName id="6F94DE8E522EFFC2558852DA00F9F80E" ID-CoL="7QM5X" authority="La Rivers 1970" authorityName="La Rivers" authorityYear="1970" box="[151,493,1984,2006]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F522EFFC2558852DA0053F80E" box="[151,327,1984,2006]" italics="true" pageId="7" pageNumber="286">Ambrysus scolius</emphasis>
<bibRefCitation id="CC05D8FC522EFFC2545252DB00F9F80E" author="La" box="[333,493,1984,2006]" pageId="7" pageNumber="286" refString="La Rivers, I. (1970) A new Ambrysus from Venezuela (Hemiptera, Naucoridae). The Great Basin Naturalist, 30 (1), 1 - 4." type="journal article" year="1970">La Rivers 1970</bibRefCitation>
</taxonomicName>
: Great Basin Natur. 1–4.
</paragraph>
</subSubSection>
<subSubSection id="E08EF6865221FFCD5588558D0591FC1F" pageId="8" pageNumber="287" type="discussion">
<paragraph id="A82BA50D5221FFCD5588558D0580FE2D" blockId="8.[151,1437,151,1257]" pageId="8" pageNumber="287">
<emphasis id="9AE0791F5221FFCD5588558D0001FF68" bold="true" box="[151,277,151,176]" pageId="8" pageNumber="287">Discussion</emphasis>
. This species is known by only three females from the
<collectingCountry id="D083E59D5221FFCD568F558202F3FF68" box="[912,999,152,176]" name="Guyana" pageId="8" pageNumber="287">Guyana</collectingCountry>
Shield countries of
<collectingCountry id="D083E59D5221FFCD51D05582043DFF68" box="[1231,1321,152,176]" name="Guyana" pageId="8" pageNumber="287">Guyana</collectingCountry>
,
<collectingCountry id="D083E59D5221FFCD5029558D048CFF68" box="[1334,1432,151,176]" name="Trinidad and Tobago" pageId="8" pageNumber="287">Trinidad</collectingCountry>
, and
<collectingCountry id="D083E59D5221FFCD55D755A6002BFF0D" box="[200,319,188,213]" name="Venezuela" pageId="8" pageNumber="287">Venezuela</collectingCountry>
. Males are not known. Although we have been unable to locate the
<typeStatus id="772F1BAF5221FFCD515455A605A4FF0D" box="[1099,1200,188,213]" pageId="8" pageNumber="287" type="holotype">holotype</typeStatus>
, the
<typeStatus id="772F1BAF5221FFCD51F855A70447FF0D" box="[1255,1363,189,213]" pageId="8" pageNumber="287" type="paratype">paratypes</typeStatus>
in the CAS (
<figureCitation id="30AFB9885221FFCD55C155C50036FF20" box="[222,290,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1764,1786]" captionTargetBox="[160,1420,555,1717]" captionTargetId="figure@7.[160,1420,555,1718]" captionTargetPageId="7" captionText="FIGURE 5. Ambrysus scolius female paratype. (A) dorsal habitus, (B) terminal abdominal sterna showing subgenital plate with posterior and lateral margins outlined in black (terminal terga missing)." httpUri="https://zenodo.org/record/237554/files/figure.png" pageId="8" pageNumber="287">Fig. 5</figureCitation>
) that we examined do not resemble the habitus figure of the
<typeStatus id="772F1BAF5221FFCD56C055C50555FF20" box="[991,1089,223,248]" pageId="8" pageNumber="287" type="holotype">holotype</typeStatus>
in the original description.
<bibRefCitation id="CC05D8FC5221FFCD506055FA0021FEC4" pageId="8" pageNumber="287" refString="La Rivers, I. (1970) A new Ambrysus from Venezuela (Hemiptera, Naucoridae). The Great Basin Naturalist, 30 (1), 1 - 4." type="journal article">La Rivers (1970)</bibRefCitation>
gave the sigmoid posterior margin of laterosternite VI (La Rivers' segment V) as the diagnostic feature of this species. We have examined the two
<typeStatus id="772F1BAF5221FFCD57D05432022EFE98" box="[719,826,296,320]" pageId="8" pageNumber="287" type="paratype">paratypes</typeStatus>
housed at the CAS and considered this feature in both anterior/posterior and dorsal/ventral planes, but are uncertain how to interpret the sigmoid nature of the margin and how it differs interspecifically. In addition, the left laterosternite VI has a well-developed posteromedial spine directed posteroventrad, whereas the right laterosternite is developed differently in the two
<typeStatus id="772F1BAF5221FFCD51CE548F0455FE75" box="[1233,1345,405,429]" pageId="8" pageNumber="287" type="paratype">paratypes</typeStatus>
: In one specimen the posterior margin is more strongly developed medially and becomes deflexed ventrad at the corner, whereas the other specimen has the posteromedial corner tuberculate and deflexed ventrad.
</paragraph>
<paragraph id="A82BA50D5221FFCD55D8571A049CFD70" blockId="8.[151,1437,151,1257]" pageId="8" pageNumber="287">
<bibRefCitation id="CC05D8FC5221FFCD55D8571A009CFDC0" box="[199,392,511,536]" pageId="8" pageNumber="287" refString="La Rivers, I. (1970) A new Ambrysus from Venezuela (Hemiptera, Naucoridae). The Great Basin Naturalist, 30 (1), 1 - 4." type="journal article">La Rivers (1970)</bibRefCitation>
described
<taxonomicName id="6F94DE8E5221FFCD571C571A0365FDC0" box="[515,625,511,536]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F5221FFCD571C571A0365FDC0" box="[515,625,511,536]" italics="true" pageId="8" pageNumber="287">A. scolius</emphasis>
</taxonomicName>
without knowledge of the related species he would describe in the next few years; thus, he did not know if the sigmoid posterior margin of laterosternite VI would be a unique feature. However, he made no mention of its condition in the original descriptions of
<taxonomicName id="6F94DE8E5221FFCD56ED57520549FDB8" box="[1010,1117,583,608]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="9AE0791F5221FFCD56ED57520549FDB8" box="[1010,1117,583,608]" italics="true" pageId="8" pageNumber="287">A. bifidus</emphasis>
</taxonomicName>
and
<taxonomicName id="6F94DE8E5221FFCD518C57520407FDB8" box="[1171,1299,584,608]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5221FFCD518C57520407FDB8" box="[1171,1299,584,608]" italics="true" pageId="8" pageNumber="287">A. tricuspis</emphasis>
</taxonomicName>
. Because
<taxonomicName id="6F94DE8E5221FFCD509B575201F1FD5D" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F5221FFCD509B575201F1FD5D" italics="true" pageId="8" pageNumber="287">A. scolius</emphasis>
</taxonomicName>
and
<taxonomicName id="6F94DE8E5221FFCD5438577600B9FD5C" box="[295,429,620,644]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5221FFCD5438577600B9FD5C" box="[295,429,620,644]" italics="true" pageId="8" pageNumber="287">A. tricuspis</emphasis>
</taxonomicName>
have the same subgenital plate shape and our examination reveals nearly the same condition for the posterior margin of laterosternite VI, we are not able to satisfactorily distinguish between them.
</paragraph>
<paragraph id="A82BA50D5221FFCD55D857AF0591FC1F" blockId="8.[151,1437,151,1257]" pageId="8" pageNumber="287">
We provisionally determined eight specimens including four females from northwestern Pará,
<collectingCountry id="D083E59D5221FFCD500D57AE0442FD15" box="[1298,1366,692,717]" name="Brazil" pageId="8" pageNumber="287">Brazil</collectingCountry>
as
<taxonomicName id="6F94DE8E5221FFCD509B57AE01F1FD28" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F5221FFCD509B57AE01F1FD28" italics="true" pageId="8" pageNumber="287">A. scolius</emphasis>
</taxonomicName>
. However, there is substantial variation in the condition of the female subgenital plate and laterosternite VI. Specifically, the subgenital plate posterior margin varies from straight to strongly convex, but is not bifurcated. The right laterosternite VI varies from a flat, unmodified posterior margin to bearing a short posteromedian spine. This suggests that considerable intraspecific variation in these features might also exist in other species in the complex.
<collectingCountry id="D083E59D5221FFCD5588567201A2FC58" box="[151,182,872,896]" name="American Samoa" pageId="8" pageNumber="287">As</collectingCountry>
such, species determinations can be equivocal for most of these species and series of specimens should be collected when possible to establish the limits of expression of these features. For these specimens from
<collectingCountry id="D083E59D5221FFCD504E5696048CFC7D" box="[1361,1432,908,933]" name="Brazil" pageId="8" pageNumber="287">Brazil</collectingCountry>
, geographic proximity suggests conspecificity with
<taxonomicName id="6F94DE8E5221FFCD57CD56AA022BFC10" box="[722,831,943,968]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="scolius">
<emphasis id="9AE0791F5221FFCD57CD56AA022BFC10" box="[722,831,943,968]" italics="true" pageId="8" pageNumber="287">A. scolius</emphasis>
</taxonomicName>
rather than with
<taxonomicName id="6F94DE8E5221FFCD56E156AA056AFC10" box="[1022,1150,944,968]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="9AE0791F5221FFCD56E156AA056AFC10" box="[1022,1150,944,968]" italics="true" pageId="8" pageNumber="287">A. tricuspis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E08EF6865221FFCD55D856CE02CBFB30" pageId="8" pageNumber="287" type="materials_examined">
<paragraph id="A82BA50D5221FFCD55D856CE00F6FBC8" blockId="8.[151,1437,151,1257]" pageId="8" pageNumber="287">
<emphasis id="9AE0791F5221FFCD55D856CE007DFC35" bold="true" box="[199,361,980,1005]" pageId="8" pageNumber="287">
<typeStatus id="772F1BAF5221FFCD55D856CE0016FC35" box="[199,258,980,1005]" pageId="8" pageNumber="287">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D083E59D5221FFCD546C56CE0308FC34" box="[371,540,980,1004]" name="Venezuela" pageId="8" pageNumber="287">VENEZUELA</collectingCountry>
, Monagas,
<quantity id="6F6C08E85221FFCD57BC56CE03F9FC35" box="[675,749,980,1005]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="287" unit="km" value="42.0">42 km</quantity>
SE Maturin. Repository of
<typeStatus id="772F1BAF5221FFCD512B56CE0582FC35" box="[1076,1174,980,1005]" pageId="8" pageNumber="287" type="holotype">holotype</typeStatus>
unknown; two female
<typeStatus id="772F1BAF5221FFCD558856E20016FBC8" box="[151,258,1016,1040]" pageId="8" pageNumber="287" type="paratype">paratypes</typeStatus>
are housed at
<collectionCode id="CE853DC85221FFCD54BB56E200CAFBC8" LSID="urn:lsid:biocol.org:col:34700" box="[420,478,1016,1040]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="8" pageNumber="287">CAS</collectionCode>
.
</paragraph>
<paragraph id="A82BA50D5221FFCD55D8510602DDFB1C" blockId="8.[151,1437,151,1257]" pageId="8" pageNumber="287">
<emphasis id="9AE0791F5221FFCD55D851060331FBED" bold="true" box="[199,549,1052,1077]" pageId="8" pageNumber="287">
Material examined.
<collectingCountry id="D083E59D5221FFCD54A551060331FBED" box="[442,549,1052,1077]" name="Brazil" pageId="8" pageNumber="287">BRAZIL</collectingCountry>
</emphasis>
: Pará state, 378, Rio Mapuera, Igarapé do Aracu,
<date id="DC2A83CD5221FFCD517B510605EBFBED" box="[1124,1279,1052,1077]" pageId="8" pageNumber="287" value="1986-06-19">19 June 1986</date>
, Py-Daniel &amp; Barbosa (
<specimenCount id="BE926E845221FFCD5419515A003FFB80" box="[262,299,1088,1112]" pageId="8" pageNumber="287" type="generic">2♂</specimenCount>
, 2♀,
<collectionCode id="CE853DC85221FFCD5472515A00A4FB80" box="[365,432,1088,1112]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="8" pageNumber="287">INPA</collectionCode>
;
<specimenCount id="BE926E845221FFCD54A1515A00F7FB80" box="[446,483,1088,1112]" pageId="8" pageNumber="287" type="generic">2♂</specimenCount>
, 2♀, UMC).
<emphasis id="9AE0791F5221FFCD579F5125026DFB80" bold="true" box="[640,889,1087,1112]" pageId="8" pageNumber="287">
<collectingCountry id="D083E59D5221FFCD579F51250260FB80" box="[640,884,1087,1112]" name="Guyana" pageId="8" pageNumber="287">BRITISH GUIANA</collectingCountry>
:
</emphasis>
Honey Camp Cr.,
<date id="DC2A83CD5221FFCD5148515A05E5FB80" box="[1111,1265,1088,1112]" pageId="8" pageNumber="287" value="1937-10-24">Oct. 24 1937</date>
, S. Harris /
<taxonomicName id="6F94DE8E5221FFCD509E515A01F1FBA5" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="scolius">A. scolius</taxonomicName>
♀
<typeStatus id="772F1BAF5221FFCD540D517F0060FBA5" box="[274,372,1125,1149]" pageId="8" pageNumber="287" type="paratype">Paratype</typeStatus>
/
<collectingCountry id="D083E59D5221FFCD548C517F00A6FBA4" box="[403,434,1125,1148]" name="Iran" pageId="8" pageNumber="287">Ira</collectingCountry>
La Rivers Collection Bequeathed to the California Academy of Sciences (♀,
<collectionCode id="CE853DC85221FFCD504A517E0484FBA4" LSID="urn:lsid:biocol.org:col:34700" box="[1365,1424,1124,1148]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="8" pageNumber="287">CAS</collectionCode>
).
<emphasis id="9AE0791F5221FFCD5588519D0030FB78" bold="true" box="[151,292,1159,1184]" pageId="8" pageNumber="287">
<collectingCountry id="D083E59D5221FFCD5588519D0030FB78" box="[151,292,1159,1184]" name="Trinidad and Tobago" pageId="8" pageNumber="287">TRINIDAD</collectingCountry>
</emphasis>
: B.W.I., Port of
<collectingCountry id="D083E59D5221FFCD54EA519D032CFB78" box="[501,568,1159,1184]" name="Spain" pageId="8" pageNumber="287">Spain</collectingCountry>
,
<date id="DC2A83CD5221FFCD5757519303CBFB78" box="[584,735,1159,1184]" pageId="8" pageNumber="287" value="1931-11-05">Nov. 5 1931</date>
, W. E. Broadway /
<taxonomicName id="6F94DE8E5221FFCD56CA5192055FFB78" box="[981,1099,1159,1184]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="scolius">A. scolius</taxonomicName>
♀
<typeStatus id="772F1BAF5221FFCD5165519305C8FB79" box="[1146,1244,1161,1185]" pageId="8" pageNumber="287" type="paratype">Paratype</typeStatus>
/
<collectingCountry id="D083E59D5221FFCD51E251930408FB78" box="[1277,1308,1161,1184]" name="Iran" pageId="8" pageNumber="287">Ira</collectingCountry>
La Rivers Collection Bequeathed to the California Academy of Sciences (♀,
<collectionCode id="CE853DC85221FFCD569D51B602A9FB1C" LSID="urn:lsid:biocol.org:col:34700" box="[898,957,1196,1220]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="8" pageNumber="287">CAS</collectionCode>
).
</paragraph>
<paragraph id="A82BA50D5221FFCD55D851D502CBFB30" blockId="8.[151,1437,151,1257]" box="[199,991,1231,1257]" pageId="8" pageNumber="287">
<emphasis id="9AE0791F5221FFCD55D851D500B1FB30" bold="true" box="[199,421,1231,1256]" pageId="8" pageNumber="287">Published records.</emphasis>
(
<bibRefCitation id="CC05D8FC5221FFCD54AC51CA0372FB30" author="La" box="[435,614,1231,1256]" pageId="8" pageNumber="287" refString="La Rivers, I. (1970) A new Ambrysus from Venezuela (Hemiptera, Naucoridae). The Great Basin Naturalist, 30 (1), 1 - 4." type="journal article" year="1970">La Rivers 1970</bibRefCitation>
):
<emphasis id="9AE0791F5221FFCD576551D5023CFB30" bold="true" box="[634,808,1231,1256]" pageId="8" pageNumber="287">
<collectingCountry id="D083E59D5221FFCD576551D5023CFB30" box="[634,808,1231,1256]" name="Venezuela" pageId="8" pageNumber="287">VENEZUELA</collectingCountry>
</emphasis>
, Monagas state.
</paragraph>
</subSubSection>
</treatment>
</document>

605
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View file

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<taxonomicName id="F7525A001229825A399187472C131CB1" ID-CoL="3M7ZD" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">
<emphasis id="E2E1724970B9B22ABA4D3D8188A8E5F3" italics="true">Holcus lanatus</emphasis>
L.
</taxonomicName>
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<subSubSection id="81C282BDD0840D37426C0E4572D96E17" type="vernacular_names">
<paragraph id="6E3BBACD1EAD13BFA01119778411B821">
<vernacularName id="8C245AA61164646EDDBB11B41CA6C96D">Wolliges Honiggras</vernacularName>
</paragraph>
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<subSubSection id="BE643A553D954CE3C97E196E0E8F52FC" type="multiple">
<paragraph id="75C29FEB6493275C017D43DEC979F1F7">
Art ISFS: 206800 Checklist: 1023630
<taxonomicName id="ADCF6E1A380185FD5572AFD4C672558F" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
<taxonomicName id="F50EB7F4B4420178999C29B9B7AE8187" class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="genus">Holcus</taxonomicName>
<taxonomicName id="ED95F1A61C22BE039B69E1384F28360A" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
</paragraph>
<paragraph id="4BD1497192945EE3C497067945253EA2">
<heading id="B9292F3A49A62DFA1DC911D2A5D50B72">Zusammenfassung</heading>
</paragraph>
</subSubSection>
<subSubSection id="28A1B45C4010B218CDAD1057539680FB" type="description">
<paragraph id="1BCC38AE90FBCC0BE6358EC4199D5C22">
Artbeschreibung
<emphasis id="0E98CCE86BEEB7A34D91A38BA104E135" italics="true">
(nach
<bibRefCitation id="C0CCCDBDE4F0FB28D4DA28131885CE1D" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<quantity id="5C4F7E72EF5464F81A9F68786556B10A" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" unit="cm" value="55.0" valueMax="80.0" valueMin="30.0">30-80 cm</quantity>
hoch,
<emphasis id="877DD4C6FFC3B44BABCB704841FED63B" bold="true">meist dicht weichhaarig</emphasis>
.
<normalizedToken id="4B243B5A046F6F56F194A213C610F8BA" originalValue="Blätter">Blaetter</normalizedToken>
<quantity id="53FFCB099C8BB644DF5D76D37016C322" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5-8 mm</quantity>
breit, nicht rau.
<normalizedToken id="7FB85ED4BA6F97C3FFCF1976F136F6D1" originalValue="Blatthäutchen">Blatthaeutchen</normalizedToken>
ca.
<quantity id="D5C6F14E2608891E74851216A844C117" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
lang, behaart.
<emphasis id="04BDE5B1A7702918B80F5BA9F04F5F44" bold="true">Blattscheiden etwas aufgeblasen</emphasis>
.
<normalizedToken id="3CDF3A07FBDD621376390E2D24E5B3A8" originalValue="Blütenstand">Bluetenstand</normalizedToken>
eine bis
<quantity id="C8FE2313193C55229ABB3EA5848FD6E1" metricMagnitude="-1" metricUnit="m" metricValue="2.0" unit="cm" value="20.0">20 cm</quantity>
lange,
<normalizedToken id="17053DE0AF23351ECADCD565E851021C" originalValue="knäuelige">knaeuelige</normalizedToken>
Rispe.
<normalizedToken id="F9B5562045F85DDCDAA8E2829D1F680D" originalValue="Ährchen">Aehrchen</normalizedToken>
2
<normalizedToken id="CD6FBBCAD76B811772AB67678A9CB3CA" originalValue="blütig">bluetig</normalizedToken>
, beide
<normalizedToken id="D218459E3DB3B7B4076EA3DF3891EDDE" originalValue="Blüten">Blueten</normalizedToken>
in die 2
<normalizedToken id="7C696974611CA3677823B41B32F5792F" originalValue="Hüllspelzen">Huellspelzen</normalizedToken>
eingeschlossen,
<emphasis id="B93B468C9DDB91EC262FB47DC270B287" bold="true">
diese aussen behaart, meist
<normalizedToken id="1DAC193C763AC0511B81AF3AEDAF7EED" originalValue="rötlich">roetlich</normalizedToken>
. Deckspelze der oberen
<normalizedToken id="C72352AB478E720AE1F74F0D3DFDBE14" originalValue="Blüte">Bluete</normalizedToken>
mit kurzer, hakig gebogener, das
<normalizedToken id="3E827A8E628B8CCB0230B198CFEC278D" originalValue="Ährchen">Aehrchen</normalizedToken>
kaum
<normalizedToken id="0A2D13F899DECF5EBC8EBB30CF09927F" originalValue="überragender">ueberragender</normalizedToken>
Granne
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="BAA0F9C3142C86F2734AFD1F99E6091D" type="biology_ecology">
<paragraph id="25B20A23F0E7DB9D68DA96B071F15214">
<normalizedToken id="34D58464197A6A0BEA137880C8E5CBA4" originalValue="Blütezeit">Bluetezeit</normalizedToken>
<emphasis id="22EB2EED6AD9E58FB7BD18EBF3CA8D30" italics="true">
(nach
<bibRefCitation id="467136F13D6BEB1B8783F79749D5C564" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: 5-8
</paragraph>
<paragraph id="8D24BE8833DB02CFC07F456D7BA1344D">
Standort und Verbreitung in der Schweiz
<emphasis id="9E70390C23470B162BC7A5843D8071F3" italics="true">
(nach
<bibRefCitation id="322708E3D9D8EDA02DB342C835EE6B63" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: Fettwiesen, Weiden, Flachmoore / kollin-montan(-subalpin) / CH
</paragraph>
</subSubSection>
<subSubSection id="89734D2A82C48B9840D065757F692128" type="distribution">
<paragraph id="1FF470E6692E43DDD7A94A96F31F3BA4">
Verbreitung global
<emphasis id="9B75D8A4E586533F7EE33F2BE0527ADC" italics="true">
(nach
<bibRefCitation id="B8B9FEAF533F93ED1559E7B4CA4F6084" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="744069500709CE9234292C2A4560B47F" originalValue="Europäisch">Europaeisch</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="F60B29B65FEBE4DF6CE277B3111BAD14" type="biology_ecology">
<paragraph id="D7B4FD7C37E49A7AD84DDAA11CE59581">
<normalizedToken id="346EBFEB2AEAA6888C7B247639A2C6BE" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte
<emphasis id="92C2936B65790553F50F3B0AD9D1A78F" italics="true">
(nach
<bibRefCitation id="20110EC9EE0836C0B4F4CB0D67A33242" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. 2010</bibRefCitation>
)
</emphasis>
3w33-43 + 3.h.2n=14
</paragraph>
</subSubSection>
<subSubSection id="E638C53EA7073392ACECED7966BE2FA2" type="multiple">
<paragraph id="9BDEB35D994E68B1FEB984BCCDC0F017">Status</paragraph>
</subSubSection>
<subSubSection id="70497655711CF4F968BCE39EC1ADCCE6" type="conservation">
<paragraph id="BBC1B6387A04BB372254C73923E82517">
<heading id="CDDBD71FD0729766755BCC7F315ADA71">Status IUCN</heading>
: Nicht
<normalizedToken id="2B3BD866F19F2B55AD7B5122C7E38C86" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="6C9C44644D2844329A551EE6C540F0D2" type="multiple">
<paragraph id="99383646FD8ECFC1CD9A57A058FFB060">Anatomie</paragraph>
<paragraph id="46D8C6FF5765B524F297D77D11342470">
<heading id="0225A56990EFAA8218E4AA42DFEB2510">Zusammenfassung</heading>
der Blattanatomie Obere Epidermiszellen gleich gross wie untere. Epidermis mit Papillen. Epidermiszellen aussen verholzt.
<normalizedToken id="820F39115E604DF9901F3FA855B2DDBE" originalValue="Leitbündelhülle">Leitbuendelhuelle</normalizedToken>
verholzt.
</paragraph>
<paragraph id="F14D725001245D9EFEEDC20BD3A44562">
<heading id="295B559DF2C3E77FB3B6520785C36847">Zusammenfassung der Stammanatomie</heading>
</paragraph>
<paragraph id="127A63350650AFE5C9C0F0B68C9E2BE1">
Umriss rund mit Rippen.
<normalizedToken id="7C8D37A6957CD949A1AC91B7DB8D9589" originalValue="Leitbündel">Leitbuendel</normalizedToken>
in einer Reihe. Kleine Interzellularen, oft dreieckig. Epidermiszellen verholzt. Epidermiszellen mit Papillen. Chlorenchyma in tangential
<normalizedToken id="ACDB24B410B40EC7587C252BF6F9E676" originalValue="verlängerten">verlaengerten</normalizedToken>
Gruppen.
</paragraph>
<paragraph id="8D160DB713ED0EEE5C966943EC8210A4">
<heading id="E02A8573F3F67A43FC79FA2894ED2167">Beschreibung (Englisch)</heading>
</paragraph>
<paragraph id="21B106E5928335E0CF1266426E3A3FE2">
Culm-diameter
<quantity id="BD1925BB294387EA21EA0424224BC6A4" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2-5 mm</quantity>
, wall thin, radius of culm in relation to wall thickness approximately 1:0.25 or &lt;0.25. Outline circular, with rips. Culm-center hollow and surrounded by a few thin-walled, not lignified cells. Epidermis with hairs. Epidermis-cells thin-walled all around. Large vascular bundles arranged in one peripheral row. Chlorenchyma in round, oval, square or rectangular groups. Sclerenchyma in a large, peripheral continuous belt (&gt; 3 cells). Cells thick-walled. Girders square, rectangular or conic. Small sclerenchymatic sheath with 1-2 cells around vascular bundles. Largest vessels in vascular bundles in lateral position. Largest vessel in the bundle 20-50
<normalizedToken id="19D1BE3FA6AB50DF2D90E6B61CEBD10F" originalValue="μm">μm</normalizedToken>
. Cavities (intercellulars) between parenchyma- cells present, small, often triangular. Distinct cavities (intercellulars) in the protoxylem area of vascular bundles.
</paragraph>
<paragraph id="92068007295C69F3AD08935A6A314E6A">
<heading id="C9D5A9B621343DF2991A0D1674874DA4">
<normalizedToken id="04EB6F849F87276534C15F171FD2BE3A" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="08FC517A6A0BFBFB4C29100C2D7E53C8">
Lebensform
<normalizedToken id="D39799857661725783E8CF2ECB719326" originalValue="Mehrjähriger">Mehrjaehriger</normalizedToken>
Hemikryptophyt
</paragraph>
<paragraph id="EA2454CA8E64BCE4DA733AEFAE265129">
<heading id="532F5EEEFF93C93F09E9D8AC1C74146B">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="58438133624211554CC533E3E1C2DCA0" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="07EB3A93B770D0582407A2B431A4CB68">
<table id="0FAE814C5662BB0921E25CBE26320BAD" class="table" inLine="true">
<tbody id="EE18B1836570101F6814689CA8891DE4">
<tr id="AD1DE7DFE390F2AD7D5B6393414560D0">
<td id="4D683A559A9A85DD59B84E3284F147A6">
<emphasis id="CB779C88AC572E5C0D7857F78EAE2FA2" bold="true">
4.5.1 - Talfettwiesen (Fromentalwiese) (
<emphasis id="2F78C05ECEC3B34C33321B3C41973068" italics="true">Arrhenatherion</emphasis>
)
</emphasis>
</td>
</tr>
<tr id="695FEEC9B620B5A57120D9FBD7A093D8">
<td id="D2A3A7DB9DC01A49AAA4132D9AF1DB18">
4.5.2 - Bergfettwiese (Goldhaferwiese) (
<emphasis id="0F8AD4935AAA903426C237D68509580A" italics="true">Polygono-Trisetion</emphasis>
)
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="B2167819E57151A2DCE35E05E51C6B6D">
<tableNote id="70D1193ED2ED9BFD4D448F481D786956">
<emphasis id="FC2A9084C5B21FF0C6C1B773FDF3FED4" bold="true">fett</emphasis>
<emphasis id="7D0F81FB7C60D807BFF947AC3FA0B4D4" italics="true">
Dominante Art, welche das Aussehen des Lebensraumes
<normalizedToken id="8FFE8DB6C33A498EF5EB591E2C49D5AE" originalValue="mitprägt">mitpraegt</normalizedToken>
</emphasis>
<emphasis id="1FBC2CB539A130C9569779C28A828BE0" italics="true">Charakterart</emphasis>
<emphasis id="DB5B436F8E4EFE985A3D55287D3E89E5" italics="true">Weniger strikt an den Lebensraum gebundene Art</emphasis>
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="2355AE6A8F805B1B52361A2A243DEC36" type="biology_ecology">
<paragraph id="31D570892E65160A60A1D5E870C84FCF">
<normalizedToken id="F27DB5F75F778AB44048FB94D08148E0" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="763791DF93A782EBC073F7CE064F7E6B" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="2FEB385560751148DCA4FDA6C79FB102">
<table id="2F2B7921CF59B092A46332877A7CDBF0" class="table" inLine="true">
<tbody id="5E186B4DB7213330BB8CFD95BF4B716B">
<tr id="602F8CA87424B2F717E0864D47206BAE">
<th id="0FB013A24E6EBEADC4B45D6CD5F68E07" colspan="2">
<emphasis id="BFC9D533DF447821E206E26C3646BA48" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="59CAEBA9D1A432B6C858371551CDB01D" colspan="2">
<emphasis id="23FC3E9B6BA167A437107A009968E53B" bold="true">Klimafaktoren</emphasis>
</th>
<th id="80E3275868C18269B74A900364676666" colspan="2">
<emphasis id="3E09B083CFBC1D2DE94BFBDFF5C4DAC1" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="199A5ACD9840C1FE470D22A56407D3F4">
<td id="FE6CE3EC27942B310293E2BEB6DB8F00">Feuchtezahl F</td>
<td id="16374841E54D4A5C0AF192792B902AEE" style="text-align:left;width:100px">
<normalizedToken id="3A231ACF6B60129EEF4D5287B8112393" originalValue="mässig">maessig</normalizedToken>
feucht; Feuchtigkeit
<normalizedToken id="79EC5A781F4D20394B301FE292139025" originalValue="mässig">maessig</normalizedToken>
wechselnd (
<normalizedToken id="6A361E9D5430DCAEA3EE37E4B6703E63" originalValue="±">+/-</normalizedToken>
1-2 Stufen)
</td>
<td id="AAE5D09B6FDDABD8231230AB2B074474">Lichtzahl L</td>
<td id="9E832ABAEF7903C765F89EFAFB8F0CD8" style="text-align:left;width:60px">hell</td>
<td id="9C12E09092210E4CAF15423EC02430FD">Salzzeichen</td>
<td id="FCCEA7EA62809872218E7660D29FB05C" style="text-align:left;width:60px">--</td>
</tr>
<tr id="47BC54BED289A161541D1A9135DF3332">
<td id="6517DDB39EA2CF6474773340989F112F">Reaktionszahl R</td>
<td id="33DE6A4AB7EE6F0968994CC9CB8DC5B8" style="text-align:left">schwach sauer bis neutral (pH 4.5-7.5)</td>
<td id="1432FEB20790659BD7B80BB778A85493">Temperaturzahl T</td>
<td id="F938F4C87718AD67C8C2668AE827E31A" style="text-align:left">unter-montan und ober-kollin</td>
</tr>
<tr id="5DA334E24C61879190A6BC60256B734C">
<td id="06350C5891A78C6A1ECC5B00E4BA3F56">
<normalizedToken id="D5C192DFF48E2882527E0E1520CDAD73" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="168CCB454161E1FCE822009F856E3536" style="text-align:left">
<normalizedToken id="85160F1061D1F6C5B98790EA62A5D89B" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="18A2E00B90E281C880A9B690123C0480" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
bis
<normalizedToken id="FE0CEDD347E95A3A08AE41526F9EC54D" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="79BCF8DB42B14D90BCFE86D540443DFE" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
<td id="6DD63F92DB3F29CBD2B10C82314638D5">
<normalizedToken id="11E54D88E7B94B1ADCC45A255CC57A56" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="01A457D13773050577C5690020333868" style="text-align:left">
subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
<normalizedToken id="28A9CCF4C6274CADDDEB12726ABDB8B4" originalValue="mässige">maessige</normalizedToken>
Temperaturschwankungen und
<normalizedToken id="39156231BECD7EBFA0C578699156A299" originalValue="mässig">maessig</normalizedToken>
tiefe Wintertemperaturen)
</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="4B86B2E871ABF92B7A00A132242A0251" type="reference_group">
<paragraph id="FEF27A65738985D520E3677037715424">
<heading id="EF298B66361B40B056836E490FF12FBF">Nomenklatur</heading>
</paragraph>
<paragraph id="A125EC198C16B902E6DD010629AA550C">
<treatmentCitation id="9C799F0E340075E2747E77FD509C1CA7">
<heading id="1469E868B5808AD6313DA4BC1D68ED1B">
<normalizedToken id="01037FD1C32C92BFE6B8A9C2004CCB0E" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="190C89C470220865618620774945495B" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="6A114F1E5A7E2B7F119A3A0376815A02" ID-CoL="3M7ZD" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">
<emphasis id="94C7F9BC24CE9A3D24FF3872D8A5AE6B" italics="true">Holcus lanatus</emphasis>
L.
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="A45D21928029EB3EC80CC52379A3E268" type="vernacular_names">
<paragraph id="13612F77592C06774C7E60E9510A7DFD">
Volksname Deutscher Name:
<vernacularName id="034E89CB703AB04A446C7A6A0C182D38" language="deu">Wolliges Honiggras</vernacularName>
Nom
<normalizedToken id="E37E59C4AE6680517790A5FECDFF6FAB" originalValue="français">francais</normalizedToken>
:
<vernacularName id="40B1AB6398A6950C8C63E0EDB62B3DD0" language="fra">Houque laineuse</vernacularName>
Nome italiano:
<vernacularName id="A6DD873E6241F0CD15E207BB9983DA2A" language="ita">Bambagione pubescente</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="5EF6A088999AF93F5AD216B139E9254D" type="reference_group">
<paragraph id="7AE054F5ECAB885817C4E2C5AD82D513">
<normalizedToken id="11C4077B0CE15AA4B148B88E957CE0AA" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="C76752044C366C24C7FCB5F63A644C4D">
<table id="B955258227A50247E471C2C1A23840B9" class="table table-condensed" inLine="true">
<tbody id="B4EBB31C956AB69EFD15C32C3ACD03D0">
<tr id="2996E4459798D361EC8DFF35C140F8E7">
<th id="A2DE3B004603B7EE8B3FC04CA57148B3" width="80px">Relation</th>
<th id="B6052EA93C84C8C002EFD68146D731DB">Nom</th>
<th id="37537CF71D11A3971FAB0D750E1A4C96">Referenzwerke</th>
<th id="3B5E721D9F39EDD1D579989595503CE8" width="90px">No</th>
</tr>
<tr id="E0E2DE3278F61822EE6C256EC6C7685B">
<td id="E3D8FB227C532DDEDBDB3D42D1F9C336" class="taxon-relation">=</td>
<td id="841795804DC726EC848083AC93BA1C5C">
<taxonomicName id="4BA655CA7ACFF9BB0A834813371D1B10" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
</td>
<td id="1AA48AEFD656EC197B4A4F6170237D4D">
<treatmentCitation id="EF64F0654782F371CC89D3E972307193">
<bibRefCitation id="599571FD6EC8B12DF68DD4228A1AAB3A" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
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<td id="6418763141BAE651F3F2F29101BF6619">206800</td>
</tr>
<tr id="49C1C49378E6DC2F87B47607B7086E80">
<td id="87531A97AA904254C2B6B71F6308397D" class="taxon-relation">=</td>
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<taxonomicName id="A9025D5D9C4B7FEB0353BC2F733C6A59" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
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<td id="472FAFF7D07CDBE95379586749DE06DC">
<treatmentCitation id="D0BCF1898402391FCCACBA4F8B3925C6">
<bibRefCitation id="CEDAD89990F5703AE7C7193E8AEE2AC7">Flora Helvetica 2001</bibRefCitation>
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<td id="8C6844F50E4D5805F1D1ED53DEB1659E">2727</td>
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<tr id="3A42676426E9D9CB56CB879891CD8164">
<td id="39472D32D80A09CF491E7313935AAF64" class="taxon-relation">=</td>
<td id="5A3EF1730021B2D5D27028ED6B38CF7D">
<taxonomicName id="BE32BFC1C9A84B933F8B937E88719799" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
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<td id="573A86D5A6F45E926F8F630EE98E2A9C">
<treatmentCitation id="BE85D3F4530DEB07F4702A83330B513E">
<bibRefCitation id="3C58E6FBAD53690C8E7A0747DBFD2BC4">Flora Helvetica 2012</bibRefCitation>
</treatmentCitation>
</td>
<td id="D48B816AFA82A0642042B263FB928A84">2907</td>
</tr>
<tr id="A33357CCB38EA16FE7FFA457BBC61D42">
<td id="90C10EE8D163C2EBCD6E741F866F4E0D" class="taxon-relation">=</td>
<td id="3EED877D2A7693EF4C5347177326F810">
<taxonomicName id="1424308CD9E15466D17A8FDCE78FBF0C" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
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<td id="59D5AC4C648FEF7C7E585479E66D3CB7">
<treatmentCitation id="EFB3E1CBFCA205C6D96AC33EB6CF2C8C" httpUri="http://treatment.plazi.org/id/D8D5BAB4AB5D3F66E6717746E86F9134">
<bibRefCitation id="9982922D2C7CFC5FD3CED20C72F0F8BB">Flora Helvetica 2018</bibRefCitation>
</treatmentCitation>
</td>
<td id="1EC6D594E3C51DB8C2BA1740839754BE">2907</td>
</tr>
<tr id="D834531EF94B91D69F301217CFF62B26">
<td id="DC2F292A5C3684519327DF30C6278238" class="taxon-relation">=</td>
<td id="ABCF970480DC4A38924A18AF68CF914C">
<taxonomicName id="83B74620CFFB87F14223EF66BBD46B5C" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
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<td id="23C376175F306654AF9F9EB299B064B3">
<treatmentCitation id="FE2304637BBBF1E52D3FA8CD0602384E">
<bibRefCitation id="0A4EFB55BA1FB29A2D2CB7EA77EBC977">Index synonymique 1996</bibRefCitation>
</treatmentCitation>
</td>
<td id="F1A7C7BBDBAA2662D53F869FCDF9A06A">206800</td>
</tr>
<tr id="79F4801A10C477340615E38C37EC542E">
<td id="1B9E80235B5C2BFA212D9C636AFF9F8A" class="taxon-relation">=</td>
<td id="226C8869DF8CE8D35AF70E7D5B88B99C">
<taxonomicName id="3FFA56E88F4508E1FF7DDDB6A33088EA" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
</td>
<td id="61C43821340DDE4DAA6B99ED9EBDB8AE">
<treatmentCitation id="AC44EB413AE77BB3E9619AB3494F88B2">
<bibRefCitation id="F9721CF4894BCB141A8443108DD40644">Landolt 1977</bibRefCitation>
</treatmentCitation>
</td>
<td id="9A44CFA9B4244017129F29B06E38110E">238</td>
</tr>
<tr id="A072EC2825056A7C5A770989040494C7">
<td id="3AFCFFF933258E68E0093845844FC3C8" class="taxon-relation">=</td>
<td id="13F7F82F014445D9B6E72464770D8473">
<taxonomicName id="206F969CCF9AAE400AB52528A67F9902" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
</td>
<td id="EACEC9C903ECDD5FC33F1DC1937498A1">
<treatmentCitation id="AA87318E603CB0B032DEFB071B073BD1">
<bibRefCitation id="72A3B825D8A30B4FE487CAB95A8ACB2E">Landolt 1991</bibRefCitation>
</treatmentCitation>
</td>
<td id="8E7CABEDCBC997233AB65156845ABCDB">219</td>
</tr>
<tr id="453F489D87C5F09F5C73A44103AB0FF2">
<td id="8C550A0E6D9DE5EEF8280513C093FA17" class="taxon-relation">=</td>
<td id="638BAC406B83119776D6EA9B73BF458E">
<taxonomicName id="2B43737E519CC2B5E66A201571CB2526" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
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<td id="524E4B22EF68BB86725A5A6F186A499B">
<treatmentCitation id="D58D06FD49B92F8DFF289C7A124077D8">
<bibRefCitation id="BCD781E0B6AACE3B9AED34724F30AB46">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="0F34F7181A11B5A698AC1F6E4D2C17E9">206800</td>
</tr>
<tr id="EDBAE7FA05A559C95CBD94851D9A227E">
<td id="8219423F9EC981EB701C7BF855A19E3E" class="taxon-relation">=</td>
<td id="286AC9259ACA2DCFD1E8CEF346E51C33">
<taxonomicName id="5F98339C52FB03C3C4F0EA17CC8D7246" authority="L." authorityName="L." class="Liliopsida" family="Poaceae" genus="Holcus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="lanatus">Holcus lanatus L.</taxonomicName>
</td>
<td id="BA6B2A31D32C4A0409F6F21F112AD108">
<treatmentCitation id="A62321291B06B7C2C2E56C1C91948E0C">
<bibRefCitation id="3485903805FA537652B2F554B063B3DA">Welten &amp; Sutter 1982</bibRefCitation>
</treatmentCitation>
</td>
<td id="0A9BFCFF7B977AD5C937700AF2811F7B">2301</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="9554FB2A7A7060A68ADA4D05E819B64F">
<tableNote id="9596E98B01D08E4A608D49D02E203297">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="A9569C2488E9B60A8AD9DE4E28EFFFE7" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="514E16C9C6EC3D6777D61BB4FDD780E2" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="97FFF8E3ED4B4CA9506FAF9E94A49F99" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="B03F474D6EDD44FB4BF2A61312EEBE71" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="2DCD54127591E0FEAD6DD1D97F88EF7A" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="BFBDD20F42AEE10CF691D14551EE7407" type="conservation">
<paragraph id="E64668304437BAA1AC8D521F24750FEA">
<heading id="65E3079AC1884E8AA455534BDE80AFBC">Status Indigenat</heading>
: Indigen
</paragraph>
<paragraph id="3E69CFF83E58523CC0D502B79B72C53D">
<emphasis id="D7CD0EC4A55C099E8365E392813ED597" bold="true">
<heading id="5D855D261DB032C7E3B5A9996F0C0C68">
Liste der
<normalizedToken id="E3FAA85180B7098465F916DF37D145DA" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="C15DA84AD702A188DBB074B757AE8460" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
<paragraph id="9B751C3BB539FB8E97C907786901FFFB">
<heading id="8C331ECACC19C6EA83104C257C841E1E">Status Rote Liste national 2016</heading>
</paragraph>
<paragraph id="6728081080E3B62F6A19E650DDE3AA7E">
<heading id="9A1D9E1FEDF2AAFA1215E3B4BC9E9A84">Status IUCN</heading>
: Nicht
<normalizedToken id="5B75C8493D48C2151E4D625BA72F7E1C" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
<paragraph id="D91C9482E93B76649427B54EC4B78210">
<heading id="E314219411A8A660D7FBDB243FB99142">
<normalizedToken id="1CFFEC0FBE404517510F78F7629EB361" originalValue="Zusätzliche">Zusaetzliche</normalizedToken>
Informationen
</heading>
Kriterien IUCN: --
</paragraph>
<paragraph id="1A2E0F1098E8E4E67F8993867D0B40E9">
<heading id="E804455F653517713331EF8EC743E59A">Status Rote Liste regional 2019</heading>
</paragraph>
<paragraph id="72CADA7E3C74D38ABE4859AA1F23B3B4">
<table id="E13E46935FE7AF0473D0153CA86BBB8E" class="table table-condensed" inLine="true">
<tbody id="7B6FE36744FFBFAE7D15592F105EB472">
<tr id="F1BF03CF59D463ED16491098730344C7">
<th id="E113BA38E22A08B28E800C85A67860ED">Biogeografische Regionen</th>
<th id="4189C63437A33FC2FB65737FFF210960">Status</th>
<th id="054B7448AA131E8C6D76828675BE6A05">Kriterien IUCN</th>
</tr>
<tr id="8D3FB64236FFEB0D959CA2A07698FDAD">
<td id="C1C15DE351C0916E396C6D34875AE231" class="regional_status">Jura (JU)</td>
<td id="8D9B2EB4FFDA3A65876A78BF5F44FAC2">
nicht
<normalizedToken id="31EB6F369F759F895914AEBB03CCD742" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="69AABCD3C8126815B49E7AA7F9E798C3">
<td id="9C062940D7E6EDDB6C8F599BA27B3486" class="regional_status">Mittelland (MP)</td>
<td id="6CC81336EA7847558FB730D266BC8811">
nicht
<normalizedToken id="D25DBFAA275B5916782A590C6F79AA39" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="796673F89240FE1AAC3A7BC5409DC640">
<td id="A5C4B49BC7A50B9A70983692B85D974D" class="regional_status">Alpennordflanke (NA)</td>
<td id="F18E57685BD2127EDF16F8259FDB4BF1">
nicht
<normalizedToken id="426A4B7BE408458391B7B1CC31483284" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="0E3054250BFBD8B37604C773B5B11071">
<td id="8FDE77AF7C0083C928D31F6BA3116FDE" class="regional_status">
<normalizedToken id="DE2E8DB2BE8C8E924EF3528C715D8030" originalValue="Alpensüdflanke">Alpensuedflanke</normalizedToken>
(SA)
</td>
<td id="ED652C47C5EBF379BB72C46DB38C5F5E">
nicht
<normalizedToken id="662F2E7B2D50B1BAD86EDAD2DFE3C314" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="E38C319BD29A5DFBB39FA27D98FF021F">
<td id="9A36B257F0765D414411EF0B37696F96" class="regional_status">
<normalizedToken id="A3FFE5E372C2FB75FABD6A8EC9B0D2C0" originalValue="Östliche">Oestliche</normalizedToken>
Zentralalpen (EA)
</td>
<td id="29FF0670D94CBC0273F718F9BF4387B6">
nicht
<normalizedToken id="70AA8DC09F54D8372BDB8AADF8A37E23" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="5919C3A9C871DC726C36589F0597DCC5">
<td id="431619BCE5BABA4446928BFEF073897C" class="regional_status">Westliche Zentralalpen (WA)</td>
<td id="5BCB0878A05C7314515B423F13B03705">
nicht
<normalizedToken id="4D0F85B598F2B3EC0B5B35F91D53E2BF" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="F1BCB72C3550587B3CE5D40991EDDE13">
<heading id="2B8B3748C895D0786A7A2BCDD7309853">
Status nationale
<normalizedToken id="375A1DB15BB1CFE570CC58810C595F32" originalValue="Priorität">Prioritaet</normalizedToken>
/Verantwortung
</heading>
</paragraph>
<paragraph id="E4DA002A3EBB87EEAE5AF904BCFC3527">
<table id="E70C4BC175BAE361B309EA96C8E5AE60" class="table" inLine="true">
<tr id="0B12A96F05C8E6E64A7EC252B21A7E11">
<td id="83239C8D21A16294DA1E706FE042FF45">
Keine nationale
<normalizedToken id="9BC62CD5371A977377EA0F9C4EEA8D87" originalValue="Priorität">Prioritaet</normalizedToken>
oder internationale Verantwortung
</td>
</tr>
</table>
</paragraph>
<paragraph id="219038D5E90D510AD3A921EC8A6BF2F4">
<heading id="760100F2139F382008176434D5828AA8">Schutzstatus</heading>
</paragraph>
<paragraph id="D25E967A1308D121ADF9EFD7CFC3F77D">
<table id="7F7E1D5EA745AE0517679A46FB83789C" class="table" inLine="true">
<tr id="D9F63A82F93ED8816D9F60D10FE5F659">
<td id="D22A9BD5785F5EEB8821367A1A18DB48">Kein internationaler, nationaler oder kantonaler Schutz</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

113
data/20/3D/58/203D585ACE97EE265E01552B60E4BB4C.xml Normal file
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@ -0,0 +1,113 @@
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Subtribe
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<taxonomicName genus="Brenthides" lsidName="Brenthides" pageId="247" pageNumber="248" rank="genus">Brenthides</taxonomicName>
Billberg, 1820a: 40 [stem: Brent-]. Type genus:
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Fabricius, 1787 [as
<taxonomicName class="Insecta" family="Brentidae" genus="Brenthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brenthus" order="Coleoptera" pageId="247" pageNumber="248" phylum="Arthropoda" rank="genus">Brenthus</taxonomicName>
, unjustified emendation of type genus name by Illiger (1801), not in prevailing usage].
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<heading id="F36381EAFFF4257FFC7F200ABB44FAEB" box="[961,1307,1333,1360]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="597" reason="2">
Genus
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<emphasis id="9AE0EA94FFF4257FFBB0200ABAD8FAF4" box="[1038,1159,1333,1359]" italics="true" pageId="4" pageNumber="597">Coccoglypta</emphasis>
Pilsbry, 1895
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<taxonomicName id="6F944D05FFF4257FFC932053BAF5FA3F" authority="Pilsbry, 1895: 211" authorityName="Pilsbry" authorityPageNumber="211" authorityYear="1895" box="[813,1194,1388,1412]" class="Gastropoda" family="Bradybaenidae" genus="Eulota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="597" phylum="Mollusca" rank="subGenus" subGenus="Coccoglypta">
<emphasis id="9AE0EA94FFF4257FFC932053BD35FA3F" box="[813,874,1388,1412]" italics="true" pageId="4" pageNumber="597">Eulota</emphasis>
(
<emphasis id="9AE0EA94FFF4257FFCC62053BDBAFA3F" box="[888,997,1388,1412]" italics="true" pageId="4" pageNumber="597">Coccoglypta</emphasis>
) Pilsbry, 1895: 211
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.
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<emphasis id="9AE0EA94FFF4257FFC9320A6BDCCFA0A" box="[813,915,1433,1457]" italics="true" pageId="4" pageNumber="597">Coccoglypta</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFF4257FFC1920A5BA73FA0A" author="YEN T. - C." box="[935,1068,1433,1457]" page="153" pageId="4" pageNumber="597" year="1939">
<bibRefCitation id="CC054B77FFF4257FFC1920A5BA73FA0A" author="YEN T. - C." box="[935,1068,1433,1457]" pageId="4" pageNumber="597" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen 1939: 153</bibRefCitation>
</treatmentCitation>
. —
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<bibRefCitation id="CC054B77FFF4257FFBEF20A6BB0FFA0A" author="SCHILEYKO A. A." box="[1105,1360,1433,1457]" pageId="4" pageNumber="597" pagination="1627 - 1763" refId="ref8091" refString="SCHILEYKO A. A. 2004. - Treatise on Recent terrestrial pulmonate molluscs Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1763." type="journal article" year="2004">
Schileyko 2004:
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, fig. 2164.
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<subSubSection id="E08E650DFFF4257FFC9320F9BBFDFA64" box="[813,1442,1478,1503]" pageId="4" pageNumber="597" type="type_taxon">
<paragraph id="A82B3686FFF4257FFC9320F9BBFDFA64" blockId="4.[813,1442,1478,1503]" box="[813,1442,1478,1503]" pageId="4" pageNumber="597">
<typeStatus id="772F8824FFF4257FFC9320F9BD02FA60" box="[813,861,1478,1502]" pageId="4" pageNumber="597">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="6F944D05FFF4257FFC6920F9BB50FA65" authority="Heude, 1886" authorityName="Heude" authorityYear="1886" box="[983,1295,1478,1503]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="597" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF4257FFC6920F9BADAFA65" box="[983,1157,1478,1502]" italics="true" pageId="4" pageNumber="597">Helix pinchoniana</emphasis>
<bibRefCitation id="CC054B77FFF4257FFB3220F8BB50FA65" author="HEUDE R. P. M." box="[1164,1295,1478,1503]" pageId="4" pageNumber="597" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
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, by monotypy.
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<paragraph id="A82B3686FFF4257FFC9320C9BA52F851" blockId="4.[813,1456,1526,2026]" pageId="4" pageNumber="597">DIAGNOSIS. — Shell dextral, depressed-conical, with 5-6 slightly convex whorls; body whorl rounded or slightly keeled; colour yellowish to greenish and light brown; protoconch without notable sculpture, seemingly smooth, or slightly pitted or “hammered”; teleoconch with irregular, rough wrinkles and strong tubercles, occasionally with fine spiral grooves; sculpture on ventral surface sometimes weaker than on dorsal side; aperture rounded to suboval, strongly oblique to shell axis; peristome expanded but not reflected; parietal callus weak, only indicated by fine whitish calcareous layer; umbilicus open, moderately wide to rather narrow, shows all whorls. Penis cylindrical or flattened, moderately long, internally with fine longitudinal pilasters or reticulated sculpture; no epiphallus observed; vas deferens slender, retractor muscle attached on the junction of penis and vas deferens; vagina short, internally with widely-spaced, sometimes converging longitudinal folds; stylophore of moderate size, accessory sac of about same size and shape, each of two alveolar; mucus glands large, distinct or forming a single gland mass, entering accessory sac independently; spermathecal stalk extremely long, bursa oval, small.</paragraph>
</subSubSection>
<caption id="FCEB660EFFF5257EFF3A2324BEAEF9FF" ID-DOI="http://doi.org/10.5281/zenodo.3726112" ID-Zenodo-Dep="3726112" httpUri="https://zenodo.org/record/3726112/files/figure.png" pageId="5" pageNumber="598" startId="5.[132,143,1563,1581]" targetBox="[132,1356,220,1516]" targetPageId="5">
<paragraph id="A82B3686FFF5257EFF3A2324BEAEF9FF" blockId="5.[132,1455,1563,1604]" pageId="5" pageNumber="598">FIG. 1. — Phylogenetic tree using Bayesian inference with mitochondrial 16S and CO1 genes. Numbers above or below branches indicate Bayesian posterior probabilities.</paragraph>
</caption>
<subSubSection id="E08E650DFFF5257EFF3A23ECBFFCF856" pageId="5" pageNumber="598" type="discussion">
<paragraph id="A82B3686FFF5257EFF3A23ECBEB7F957" blockId="5.[132,775,1747,2029]" box="[132,232,1747,1773]" pageId="5" pageNumber="598">REMARKS</paragraph>
<paragraph id="A82B3686FFF5257EFF3A23CBBFFCF856" blockId="5.[132,775,1747,2029]" pageId="5" pageNumber="598">
<taxonomicName id="6F944D05FFF5257EFF3A23CBBEA2F8B5" authorityName="Pilsbry" authorityYear="1895" box="[132,253,1780,1806]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF5257EFF3A23CBBEA2F8B5" box="[132,253,1780,1806]" italics="true" pageId="5" pageNumber="598">Coccoglypta</emphasis>
</taxonomicName>
was erected as a section of
<taxonomicName id="6F944D05FFF5257EFDA023CBBEE3F895" authority="W. Hartmann, 1841" authorityName="W. Hartmann" authorityYear="1841" class="Gastropoda" family="Bradybaenidae" genus="Eulota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF5257EFDA023CBBC3DF8B5" box="[542,610,1780,1806]" italics="true" pageId="5" pageNumber="598">Eulota</emphasis>
W. Hartmann, 1841
</taxonomicName>
for two species, namely “
<taxonomicName id="6F944D05FFF5257EFE6A222BBC92F895" authority="Heude" authorityName="Heude" box="[468,717,1812,1838]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="dimidiata">
<emphasis id="9AE0EA94FFF5257EFE6A222BBC23F895" box="[468,636,1812,1838]" italics="true" pageId="5" pageNumber="598">Helix dimidiata</emphasis>
Heude
</taxonomicName>
” and “
<taxonomicName id="6F944D05FFF5257EFF2E220CBFF4F8F6" authority="Heude" authorityName="Heude" authorityYear="1886" box="[144,427,1843,1869]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF5257EFF2E220CBF0BF8F6" box="[144,340,1843,1869]" italics="true" pageId="5" pageNumber="598">Helix pinchoniana</emphasis>
Heude
</taxonomicName>
”. Pilsbry (1895) designated the former species as the
<typeStatus id="772F8824FFF5257EFEDD226BBFCEF8D5" box="[355,401,1876,1902]" pageId="5" pageNumber="598">type</typeStatus>
species. Heude, however, has never described a species with the name dimidiata (see
<bibRefCitation id="CC054B77FFF5257EFD13224CBEE1F816" author="JOHNSON R. J." pageId="5" pageNumber="598" refId="ref7571" refString="JOHNSON R. J. 1973. - Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, 111 p. https: // doi. org / 10.5962 / bhl. title. 141074" type="book" year="1973">Johnson 1973</bibRefCitation>
).
<bibRefCitation id="CC054B77FFF5257EFF6922ABBF0AF815" author="YEN T. - C." box="[215,341,1939,1966]" pageId="5" pageNumber="598" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen (1939)</bibRefCitation>
and
<bibRefCitation id="CC054B77FFF5257EFE2F22ACBC46F815" author="ZILCH A." box="[401,537,1939,1966]" pageId="5" pageNumber="598" refId="ref8700" refString="ZILCH A. 1959 - 1960. - Handbuch der Paleozoologie, 6 (2) Euthyneura. Gebruder Borntraeger, Berlin-Nikolassee, 834 p." type="book" year="1959">
Zilch (
<date id="DC2A1046FFF5257EFE6022ACBC46F815" box="[478,537,1939,1966]" pageId="5" pageNumber="598" value="1959" valueMax="1960">1959</date>
</bibRefCitation>
-1960) corrected this mistake and clarified
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species of
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<emphasis id="9AE0EA94FFF5257EFD99228CBCFAF876" box="[551,677,1971,1997]" italics="true" pageId="5" pageNumber="598">Coccoglypta</emphasis>
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as
<taxonomicName id="6F944D05FFF5257EFD72228CBFC1F856" authority="Heude, 1886" authorityName="Heude" authorityYear="1886" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF5257EFD72228CBF5AF856" italics="true" pageId="5" pageNumber="598">Helix pinchoniana</emphasis>
<bibRefCitation id="CC054B77FFF5257EFEB222ECBFC1F856" author="HEUDE R. P. M." box="[268,414,2003,2029]" pageId="5" pageNumber="598" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
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Subfamily
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<paragraph id="A82B3686FFF4257FFF3A234BBEB7F935" blockId="4.[132,776,1652,2030]" box="[132,232,1652,1678]" pageId="4" pageNumber="597">REMARKS</paragraph>
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and
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are traditionally distinguished on the basis of the absence of the dart sac and mucous glands in the former and their presence in latter. The molecular phylogeny of
<bibRefCitation id="CC054B77FFF4257FFE1023CBBC39F8B5" author="WADE C. M. &amp; MORDAN P. B. &amp; NAGGS F." box="[430,614,1780,1806]" pageId="4" pageNumber="597" pagination="593 - 610" refId="ref8410" refString="WADE C. M., MORDAN P. B. &amp; NAGGS F. 2006. - Evolutionary relationships among the Pulmonate land snails and slugs (Pulmonata, Stylommatophora). Biological Journal of the Linnean Society 87: 593 - 610. https: // doi. org / 10.1111 / j. 1095 - 8312.2006.00596. x" type="journal article" year="2006">
Wade
<emphasis id="9AE0EA94FFF4257FFE4E23CABC7EF8B5" box="[496,545,1780,1806]" italics="true" pageId="4" pageNumber="597">et al.</emphasis>
(2006
</bibRefCitation>
,
<bibRefCitation id="CC054B77FFF4257FFDCC23CBBCF6F8B5" author="WADE C. M. &amp; HUDELOT C. &amp; DAVISON A. &amp; NAGGS F. &amp; MORDAN P. B." box="[626,681,1780,1806]" pageId="4" pageNumber="597" pagination="411 - 415" refId="ref8471" refString="WADE C. M., HUDELOT C., DAVISON A., NAGGS F. &amp; MORDAN P. B. 2007. - Molecular phylogeny of the helicoid land snails (Pulmonata: Stylommatophora: Helicoidea), with special emphasis on the Camaenidae. Journal of Molluscan Studies 73: 411 - 415. https: // doi. org / 10.1093 / mollus / eym 030" type="journal article" year="2007">2007</bibRefCitation>
) showed that the dart sac was lost multiple times during the evolution of the
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.
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and
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form a single clade, but neither of them is monophyletic. Therefore
<bibRefCitation id="CC054B77FFF4257FFF3A224CBFCEF835" author="GITTENBERGER E. &amp; HAMANN T. D. &amp; ASAMI T." box="[132,401,1907,1934]" pageId="4" pageNumber="597" pagination="34005" refId="ref7250" refString="GITTENBERGER E., HAMANN T. D. &amp; ASAMI T. 2012. - Chiral Speciation in Terrestrial Pulmonate Snails. PLoS ONE 7 (4): e 34005. https: // doi. org / 10.1371 / journal. pone. 0034005" type="journal article" year="2012">
Gittenberger
<emphasis id="9AE0EA94FFF4257FFEAC224BBF1CF836" box="[274,323,1907,1933]" italics="true" pageId="4" pageNumber="597">et al.</emphasis>
(2012)
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formally treated the
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as a junior synonym of
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.
<bibRefCitation id="CC054B77FFF4257FFDA022ACBD58F815" author="BOUCHET P. &amp; ROCROI J. - P. &amp; HAUSDORF B. &amp; KAIM A. &amp; KANO Y. &amp; ZEL A. &amp; PARKHAEV P. &amp; SCHRODL M. &amp; STRONG E. E." box="[542,775,1939,1966]" pageId="4" pageNumber="597" pagination="1 - 526" refId="ref6999" refString="BOUCHET P., ROCROI J. - P., HAUSDORF B., KAIM A., KANO Y., NUT- ZEL A., PARKHAEV P., SCHRODL M. &amp; STRONG E. E. 2017. - Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1 - 2): 1 - 526." type="journal article" year="2017">
Bouchet
<emphasis id="9AE0EA94FFF4257FFD3E22ABBCEAF816" box="[640,693,1939,1965]" italics="true" pageId="4" pageNumber="597">et al.</emphasis>
(2017)
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retained the subfamily
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under
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. This system is followed here.
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<treatment id="203D8790FFF52573FC1123ECBF46F856" ID-DOI="http://doi.org/10.5281/zenodo.3729510" ID-GBIF-Taxon="163283974" ID-Zenodo-Dep="3729510" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856" httpUri="http://treatment.plazi.org/id/203D8790FFF52573FC1123ECBF46F856" lastPageId="8" lastPageNumber="601" pageId="5" pageNumber="598">
<subSubSection id="E08E650DFFF5257EFC1123ECBB72F955" box="[943,1325,1747,1774]" pageId="5" pageNumber="598" type="nomenclature">
<paragraph id="A82B3686FFF5257EFC1123ECBB72F955" blockId="5.[943,1325,1747,1806]" box="[943,1325,1747,1774]" pageId="5" pageNumber="598">
<heading id="F36381EAFFF5257EFC1123ECBB72F955" box="[943,1325,1747,1774]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="598" reason="2">
<taxonomicName id="6F944D05FFF5257EFC1123ECBAB7F955" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[943,1256,1747,1774]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="liui" status="sp. nov.">
<emphasis id="9AE0EA94FFF5257EFC1123ECBA04F955" bold="true" box="[943,1115,1747,1774]" italics="true" pageId="5" pageNumber="598">Coccoglypta liui</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="81D357EFFFF5257EFB4C23EBBB72F955" box="[1266,1325,1748,1774]" pageId="5" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF5257EFBAF23CBBBC9F8F9" pageId="5" pageNumber="598" type="description">
<paragraph id="A82B3686FFF5257EFBAF23CBBA94F8B5" blockId="5.[943,1325,1747,1806]" box="[1041,1227,1780,1806]" pageId="5" pageNumber="598">
(
<figureCitation id="30AF2A03FFF5257EFBA623CBBAD6F8B5" box="[1048,1161,1780,1806]" captionStart="FIG" captionStartId="6.[132,143,1815,1833]" captionTargetBox="[132,1455,215,1771]" captionTargetId="figure@6.[132,1455,214,1775]" captionTargetPageId="6" captionText="FIG. 2. — Shells of Coccoglypta Pilsbry, 1895 species: A-D, Coccoglypta liui Páll-Gergely, n. sp. (holotype); E-F, syntype of Coccoglypta pinchoniana (USNM 472156); I-L, Coccoglypta cf. pinchoniana from Emei Shan (SMF 42574); M, second specimen of Coccoglypta cf. pinchoniana from the same sample as previ- ous; N, protoconch of the same shell as I-L. Scale bar: 30 mm; N, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3726114" httpUri="https://zenodo.org/record/3726114/files/figure.png" pageId="5" pageNumber="598">Figs 2A-D</figureCitation>
;
<figureCitation id="30AF2A03FFF5257EFB2623CBBAF9F8B5" box="[1176,1190,1780,1806]" captionStart="FIG" captionStartId="7.[132,143,1273,1291]" captionTargetBox="[165,1416,232,1205]" captionTargetId="figure@7.[132,1455,216,1236]" captionTargetPageId="7" captionText="FIG. 3. — Reproductive anatomy of Coccoglypta liui Páll-Gergely, n. sp. (holotype, HNHM 103475): A, entire genitalia; B, schematic drawing of the inner structure of the dart sac. Scale bar: A, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3726116" httpUri="https://zenodo.org/record/3726116/files/figure.png" pageId="5" pageNumber="598">3</figureCitation>
,
<figureCitation id="30AF2A03FFF5257EFB0A23CBBA9DF8B5" box="[1204,1218,1780,1806]" captionStart="FIG" captionStartId="8.[133,144,953,971]" captionTargetBox="[132,1455,215,894]" captionTargetId="figure@8.[132,1455,215,894]" captionTargetPageId="8" captionText="FIG. 4. — Inner structure of the genitalia of Coccoglypta liui Páll-Gergely, n. sp. (holotype, HNHM 103475): A, penis; B, vagina. Genital atrium situated leftwards in both images. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3726118" httpUri="https://zenodo.org/record/3726118/files/figure.png" pageId="5" pageNumber="598">4</figureCitation>
)
</paragraph>
<paragraph id="A82B3686FFF5257EFCF8220FBBC9F8F9" blockId="5.[838,1430,1840,1858]" box="[838,1430,1840,1858]" pageId="5" pageNumber="598">
<uri id="DC053A84FFF5257EFCF8220FBBC9F8F9" box="[838,1430,1840,1858]" pageId="5" pageNumber="598">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF5257CFC932257BEACF9ED" lastPageId="7" lastPageNumber="600" pageId="5" pageNumber="598" type="materials_examined">
<paragraph id="A82B3686FFF5257EFC932257BB23F80E" blockId="5.[813,1455,1896,2027]" pageId="5" pageNumber="598">
<collectionCode id="CE85AE43FFF5257EFC932257BD03F8C6" box="[813,860,1896,1920]" pageId="5" pageNumber="598">TYPE</collectionCode>
<collectionCode id="CE85AE43FFF5257EFCE12252BDE8F8C5" box="[863,951,1901,1918]" pageId="5" pageNumber="598">MATERIAL</collectionCode>
. —
<materialsCitation id="18FC3CDBFFF5257EFC632257BB23F80E" ID-GBIF-Occurrence="2580042133" collectingDate="2014-10-27" collectionCode="TYPE, MATERIAL" collectorName="Okubo" country="China" county="Chengdu Shi" elevation="1200" location="Qingcheng Houshan" municipality="Dujiangyan Shi" pageId="5" pageNumber="598" specimenCode="HNHM 103475" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="9AE0EA94FFF5257EFC632257BA68F83B" bold="true" box="[989,1079,1896,1920]" pageId="5" pageNumber="598">
<typeStatus id="772F8824FFF5257EFC632257BA68F83B" box="[989,1079,1896,1920]" pageId="5" pageNumber="598" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCountry id="D0837616FFF5257EFBFF2257BA22F83B" box="[1089,1149,1896,1920]" name="China" pageId="5" pageNumber="598">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF5257EFB3B2257BA8EF83B" box="[1157,1233,1896,1920]" country="China" name="Sichuan" pageId="5" pageNumber="598">Sichuan</collectingRegion>
,
<collectingCounty id="414A4E0AFFF5257EFB672257BB0BF83B" box="[1241,1364,1896,1920]" pageId="5" pageNumber="598">Chengdu Shi</collectingCounty>
,
<collectingMunicipality id="484FACFCFFF5257EFAE52257BD2BF820" pageId="5" pageNumber="598">Dujiangyan Shi</collectingMunicipality>
,
<location id="AD4B605DFFF5257EFCC222BCBA1BF820" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF5257EFCC222BCBA1BF820" box="[892,1092,1923,1947]" country="China" county="Chengdu Shi" municipality="Dujiangyan Shi" name="Qingcheng Houshan" pageId="5" pageNumber="598" stateProvince="Sichuan">Qingcheng Houshan</location>
;
<elevation id="23B9D1B5FFF5257EFBF322BCBA99F820" box="[1101,1222,1923,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="598" unit="m" value="1200.0">
<quantity id="6F6C9B63FFF5257EFBF322BCBAC9F821" box="[1101,1174,1923,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="598" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="05615350FFF5257EFB7122BCBB4CF820" box="[1231,1299,1923,1947]" pageId="5" pageNumber="598">Okubo</collectorName>
leg.;
<date id="DC2A1046FFF5257EFAFD22BCBBF3F820" box="[1347,1452,1923,1947]" pageId="5" pageNumber="598" value="2014-10-27">
<collectingDate id="CC6EE9AEFFF5257EFAFD22BCBBF3F820" box="[1347,1452,1923,1947]" pageId="5" pageNumber="598" value="2014-10-27">27.X.2014</collectingDate>
</date>
;
<specimenCode id="F8329EFDFFF5257EFC9322A1BD8AF80E" box="[813,981,1949,1973]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="598">HNHM 103475</specimenCode>
; D =
<quantity id="6F6C9B63FFF5257EFBB222A2BA36F80E" box="[1036,1129,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="5" pageNumber="598" unit="mm" value="32.9">32.9 mm</quantity>
, H =
<quantity id="6F6C9B63FFF5257EFB1F22A2BAA3F80E" box="[1185,1276,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="2.06" pageId="5" pageNumber="598" unit="mm" value="20.6">20.6 mm</quantity>
(
<figureCitation id="30AF2A03FFF5257EFAB422A2BB1DF80E" box="[1290,1346,1949,1973]" captionStart="FIG" captionStartId="6.[132,143,1815,1833]" captionTargetBox="[132,1455,215,1771]" captionTargetId="figure@6.[132,1455,214,1775]" captionTargetPageId="6" captionText="FIG. 2. — Shells of Coccoglypta Pilsbry, 1895 species: A-D, Coccoglypta liui Páll-Gergely, n. sp. (holotype); E-F, syntype of Coccoglypta pinchoniana (USNM 472156); I-L, Coccoglypta cf. pinchoniana from Emei Shan (SMF 42574); M, second specimen of Coccoglypta cf. pinchoniana from the same sample as previ- ous; N, protoconch of the same shell as I-L. Scale bar: 30 mm; N, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3726114" httpUri="https://zenodo.org/record/3726114/files/figure.png" pageId="5" pageNumber="598">Fig. 2</figureCitation>
A-D).
</materialsCitation>
</paragraph>
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<materialsCitation id="18FC3CDBFFF5257EFC932288BB75F851" ID-GBIF-Occurrence="2580042135" collectorName="Chengdu Shi &amp; Dujiangyan Shi &amp; Qingcheng Shan &amp; Buyun Cableway" country="China" location="Sichuan" pageId="5" pageNumber="598" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="9AE0EA94FFF5257EFC932288BDD1F874" bold="true" box="[813,910,1975,1999]" pageId="5" pageNumber="598">
<typeStatus id="772F8824FFF5257EFC932288BDD1F874" box="[813,910,1975,1999]" pageId="5" pageNumber="598" type="paratype">Paratypes</typeStatus>
</emphasis>
. 2015/50;
<collectingCountry id="D0837616FFF5257EFC412287BA60F86B" box="[1023,1087,1976,2000]" name="China" pageId="5" pageNumber="598">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF5257EFBF52287BAC2F86B" box="[1099,1181,1976,2000]" country="China" name="Sichuan" pageId="5" pageNumber="598">Sichuan</collectingRegion>
,
<collectorName id="05615350FFF5257EFB172287BB6FF86B" box="[1193,1328,1976,2000]" pageId="5" pageNumber="598">Chengdu Shi</collectorName>
,
<collectorName id="05615350FFF5257EFA852287BD10F851" pageId="5" pageNumber="598">Dujiangyan Shi</collectorName>
,
<collectorName id="05615350FFF5257EFCE922EDBDA4F851" box="[855,1019,2002,2026]" pageId="5" pageNumber="598">Qingcheng Shan</collectorName>
,
<collectorName id="05615350FFF5257EFBBD22ECBAF8F851" box="[1027,1191,2002,2027]" pageId="5" pageNumber="598">Buyun Cableway</collectorName>
upper station
</materialsCitation>
–
<materialsCitation id="18FC3CDBFFF5257DFAFE22EDBF97F80F" ID-GBIF-Occurrence="2580042132" lastPageId="6" lastPageNumber="599" pageId="5" pageNumber="598">
Shangqing- gong;
<quantity id="6F6C9B63FFF6257DFF7E22BEBF53F822" box="[192,268,1921,1945]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="599" unit="m" value="1150.0">1150 m</quantity>
a.s.l.;
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,
<geoCoordinate id="CDA05041FFF6257DFE6A22BEBC01F822" box="[468,606,1921,1945]" degrees="103" direction="east" minutes="33.714" orientation="longitude" pageId="6" pageNumber="599" precision="1" value="103.5619">103°33.714’E</geoCoordinate>
; A. Hunyadi leg.;
<date id="DC2A1046FFF6257DFF3A22A3BEA6F80F" box="[132,249,1948,1972]" pageId="6" pageNumber="599" value="2015-06-06">
06.
<collectionCode id="CE85AE43FFF6257DFF1D22A3BE9FF80F" box="[163,192,1948,1972]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="599">VI</collectionCode>
.2015
</date>
; coll. HA/
<specimenCount id="BE92FD0FFFF6257DFEDD22A3BF97F80F" box="[355,456,1948,1972]" pageId="6" pageNumber="599" type="generic" typeStatus="paratype">1 paratype</specimenCount>
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF6257DFE4E22A3BB7EF821" ID-GBIF-Occurrence="2580042137" collectingDate="2015-06-04" collectionCode="VI" collectorName="A. Hunyadi" country="China" county="Chengdu Shi" elevation="1530" latitude="30.946434" location="Qingcheng Houshan" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" pageId="6" pageNumber="599" specimenCount="1" specimenCount-adult="1" stateProvince="Sichuan" typeStatus="paratype">
2015/45;
<collectingRegion id="6A50F864FFF6257DFDEF22A3BCFDF80F" box="[593,674,1948,1972]" country="China" name="Sichuan" pageId="6" pageNumber="599">Sichuan</collectingRegion>
,
<collectingCounty id="414A4E0AFFF6257DFD1222A3BEF8F875" pageId="6" pageNumber="599">Chengdu Shi</collectingCounty>
,
<collectingMunicipality id="484FACFCFFF6257DFF0D2289BF0CF875" box="[179,339,1974,1998]" pageId="6" pageNumber="599">Dujiangyan Shi</collectingMunicipality>
,
<location id="AD4B605DFFF6257DFEE02289BC6CF875" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF6257DFEE02289BC6CF875" box="[350,563,1974,1998]" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Qingcheng Houshan" pageId="6" pageNumber="599" stateProvince="Sichuan">Qingcheng Houshan</location>
,
<location id="AD4B605DFFF6257DFD802289BCEAF875" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF6257DFD802289BCEAF875" box="[574,693,1974,1998]" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Taian Zhen" pageId="6" pageNumber="599" stateProvince="Sichuan">Taian Zhen</location>
,
<location id="AD4B605DFFF6257DFD7F2289BEEDF852" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF6257DFD7F2289BEEDF852" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Baiyun Cun" pageId="6" pageNumber="599" stateProvince="Sichuan">Baiyun Cun</location>
,
<location id="AD4B605DFFF6257DFF0322EEBF67F852" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF6257DFF0322EEBF67F852" box="[189,312,2001,2025]" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Jiusengdong" pageId="6" pageNumber="599" stateProvince="Sichuan">Jiusengdong</location>
;
<elevation id="23B9D1B5FFF6257DFEFC22EFBF9CF852" box="[322,451,2000,2025]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="6" pageNumber="599" unit="m" value="1530.0">
<quantity id="6F6C9B63FFF6257DFEFC22EFBFD0F853" box="[322,399,2000,2024]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="6" pageNumber="599" unit="m" value="1530.0">1530 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="CDA05041FFF6257DFE7022EFBC10F853" box="[462,591,2000,2024]" degrees="30" direction="north" minutes="56.786" orientation="latitude" pageId="6" pageNumber="599" precision="1" value="30.946434">30°56.786’N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFF6257DFDE222EFBCB9F853" box="[604,742,2000,2024]" degrees="103" direction="east" minutes="28.587" orientation="longitude" pageId="6" pageNumber="599" precision="1" value="103.47645">103°28.587’E</geoCoordinate>
;
<collectorName id="05615350FFF6257DFD4F22EEBDDDF822" box="[753,898,1921,2025]" pageId="6" pageNumber="599">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFF6257DFC0C22BEBA7BF822" box="[946,1060,1921,1945]" pageId="6" pageNumber="599" value="2015-06-04">
<collectingDate id="CC6EE9AEFFF6257DFC0C22BEBA7BF822" box="[946,1060,1921,1945]" pageId="6" pageNumber="599" value="2015-06-04">
04.
<collectionCode id="CE85AE43FFF6257DFC6E22BEBDB3F822" box="[976,1004,1921,1945]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="599">VI</collectionCode>
.2015
</collectingDate>
</date>
; coll. HA/
<specimenCount id="BE92FD0FFFF6257DFB3422BEBA97F822" box="[1162,1224,1921,1945]" pageId="6" pageNumber="599" type="adult">1 adult</specimenCount>
<typeStatus id="772F8824FFF6257DFB7222BDBB7EF821" box="[1228,1313,1922,1946]" pageId="6" pageNumber="599" type="paratype">paratype</typeStatus>
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF6257DFAF822BEBD8BF852" ID-GBIF-Occurrence="2580042134" collectingDate="2018-04-07" collectorName="Liu Zheng-Ping" country="China" latitude="30.916666" location="Qingcheng Houshan" longLatPrecision="925" longitude="103.49444" pageId="6" pageNumber="599" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
A1;
<collectingCountry id="D0837616FFF6257DFAD322BEBBF4F822" box="[1389,1451,1921,1945]" name="China" pageId="6" pageNumber="599">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF6257DFC9322A3BD88F80F" box="[813,983,1948,1972]" country="China" name="Sichuan" pageId="6" pageNumber="599">Sichuan Province</collectingRegion>
,
<location id="AD4B605DFFF6257DFC6122A3BAF1F80F" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF6257DFC6122A3BAF1F80F" box="[991,1198,1948,1972]" country="China" latitude="30.916666" longLatPrecision="925" longitude="103.49444" name="Qingcheng Houshan" pageId="6" pageNumber="599" stateProvince="Sichuan">Qingcheng Houshan</location>
;
<emphasis id="9AE0EA94FFF6257DFB0822A3BA99F808" box="[1206,1222,1948,1971]" italics="true" pageId="6" pageNumber="599">c.</emphasis>
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,
<geoCoordinate id="CDA05041FFF6257DFA9422A3BBF3F80F" box="[1322,1452,1948,1972]" degrees="103" direction="east" minutes="29" orientation="longitude" pageId="6" pageNumber="599" precision="15" seconds="40" value="103.49444">103°29’40”E</geoCoordinate>
;
<collectorName id="05615350FFF6257DFC932289BD98F875" box="[813,967,1974,1998]" pageId="6" pageNumber="599">Liu Zheng-Ping</collectorName>
leg.;
<date id="DC2A1046FFF6257DFC472289BA36F875" box="[1017,1129,1974,1998]" pageId="6" pageNumber="599" value="2018-04-07">
<collectingDate id="CC6EE9AEFFF6257DFC472289BA36F875" box="[1017,1129,1974,1998]" pageId="6" pageNumber="599" value="2018-04-07">07.IV.2018</collectingDate>
</date>
; SYS m001013 (
<typeStatus id="772F8824FFF6257DFAB72289BB01F875" box="[1289,1374,1974,1998]" pageId="6" pageNumber="599" type="paratype">paratype</typeStatus>
used for molecular study)
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF6257CFC4322EEBD5CFAD6" ID-GBIF-Occurrence="2580042143" collectionCode="HBUMM" collectorName="Liu Zheng-Ping" country="China" lastPageId="7" lastPageNumber="600" location="Shuijing Dong" municipality="Qingcheng Houshan" pageId="6" pageNumber="599" specimenCode="HBUMM10005" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
<collectingMunicipality id="484FACFCFFF6257DFC4322EEBA92F852" box="[1021,1229,2001,2025]" pageId="6" pageNumber="599">Qingcheng Houshan</collectingMunicipality>
,
<location id="AD4B605DFFF6257DFB6922EEBB38F852" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF6257DFB6922EEBB38F852" box="[1239,1383,2001,2025]" country="China" municipality="Qingcheng Houshan" name="Shuijing Dong" pageId="6" pageNumber="599" stateProvince="Sichuan">Shuijing Dong</location>
[水晶洞], 2016;
<collectorName id="05615350FFF7257CFE9B206ABFE4FAD6" box="[293,443,1365,1389]" pageId="7" pageNumber="600">Liu Zheng-Ping</collectorName>
leg.;
<specimenCode id="F8329EFDFFF7257CFE55206ABCD1FAD6" box="[491,654,1365,1389]" collectionCode="HBUMM" pageId="7" pageNumber="600">HBUMM10005</specimenCode>
; empty shell
</materialsCitation>
.
</paragraph>
<caption id="FCEB660EFFF6257DFF3A2228BD4AF8EC" ID-DOI="http://doi.org/10.5281/zenodo.3726114" ID-Zenodo-Dep="3726114" httpUri="https://zenodo.org/record/3726114/files/figure.png" pageId="6" pageNumber="599" startId="6.[132,143,1815,1833]" targetBox="[132,1455,215,1771]" targetPageId="6">
<paragraph id="A82B3686FFF6257DFF3A2228BD4AF8EC" pageId="6" pageNumber="599">
FIG. 2. — Shells of
<emphasis id="9AE0EA94FFF6257DFE9B2228BFD1F892" box="[293,398,1815,1833]" italics="true" pageId="6" pageNumber="599">Coccoglypta</emphasis>
Pilsbry, 1895 species:
<emphasis id="9AE0EA94FFF6257DFDED2228BC2AF892" bold="true" box="[595,629,1815,1833]" pageId="6" pageNumber="599">A -D</emphasis>
,
<emphasis id="9AE0EA94FFF6257DFD3E2228BD58F892" box="[640,775,1815,1833]" italics="true" pageId="6" pageNumber="599">Coccoglypta liui</emphasis>
Páll-Gergely, n. sp. (holotype);
<emphasis id="9AE0EA94FFF6257DFBAE2228BA71F892" bold="true" box="[1040,1070,1815,1833]" pageId="6" pageNumber="599">E -F</emphasis>
, syntype of
<emphasis id="9AE0EA94FFF6257DFB262228BB32F892" box="[1176,1389,1815,1833]" italics="true" pageId="6" pageNumber="599">Coccoglypta pinchoniana</emphasis>
(USNM 472156);
<emphasis id="9AE0EA94FFF6257DFF6C2211BEB5F8FB" bold="true" box="[210,234,1838,1856]" pageId="6" pageNumber="599">I -L</emphasis>
,
<emphasis id="9AE0EA94FFF6257DFF4B2211BF01F8FB" box="[245,350,1838,1856]" italics="true" pageId="6" pageNumber="599">Coccoglypta</emphasis>
cf.
<emphasis id="9AE0EA94FFF6257DFEC02211BFBCF8FB" box="[382,483,1838,1856]" italics="true" pageId="6" pageNumber="599">pinchoniana</emphasis>
from Emei Shan (SMF 42574);
<emphasis id="9AE0EA94FFF6257DFD562211BCA6F8FB" bold="true" box="[744,761,1838,1856]" pageId="6" pageNumber="599">M</emphasis>
, second specimen of
<emphasis id="9AE0EA94FFF6257DFC0D2211BA43F8FB" box="[947,1052,1838,1856]" italics="true" pageId="6" pageNumber="599">Coccoglypta</emphasis>
cf.
<emphasis id="9AE0EA94FFF6257DFB822211BAFEF8FB" box="[1084,1185,1838,1856]" italics="true" pageId="6" pageNumber="599">pinchoniana</emphasis>
from the same sample as previ- ous;
<emphasis id="9AE0EA94FFF6257DFF13227ABEE4F8EC" bold="true" box="[173,187,1861,1879]" pageId="6" pageNumber="599">N</emphasis>
, protoconch of the same shell as
<emphasis id="9AE0EA94FFF6257DFE6B227ABFB2F8EC" bold="true" box="[469,493,1861,1879]" pageId="6" pageNumber="599">I -L</emphasis>
. Scale bar: 30 mm; N, not to scale.
</paragraph>
</caption>
<caption id="FCEB660EFFF7257CFF3A21C6BFF2FA99" ID-DOI="http://doi.org/10.5281/zenodo.3726116" ID-Zenodo-Dep="3726116" httpUri="https://zenodo.org/record/3726116/files/figure.png" pageId="7" pageNumber="600" startId="7.[132,143,1273,1291]" targetBox="[165,1416,232,1205]" targetPageId="7">
<paragraph id="A82B3686FFF7257CFF3A21C6BFF2FA99" pageId="7" pageNumber="600">
FIG. 3. — Reproductive anatomy of
<emphasis id="9AE0EA94FFF7257CFE1621C6BC72FAB0" box="[424,557,1273,1291]" italics="true" pageId="7" pageNumber="600">Coccoglypta liui</emphasis>
Páll-Gergely, n. sp. (holotype, HNHM 103475):
<emphasis id="9AE0EA94FFF7257CFC0D21C6BD9FFAB0" bold="true" box="[947,960,1273,1291]" pageId="7" pageNumber="600">A</emphasis>
, entire genitalia;
<emphasis id="9AE0EA94FFF7257CFBF221C6BA06FAB0" bold="true" box="[1100,1113,1273,1291]" pageId="7" pageNumber="600">B</emphasis>
, schematic drawing of the inner structure of the dart sac. Scale bar: A, 5 mm.
</paragraph>
</caption>
<paragraph id="A82B3686FFF7257CFF3A20BBBEACF9ED" blockId="7.[132,783,1412,1622]" pageId="7" pageNumber="600">
OTHER MATERIAL EXAMINED. —
<materialsCitation id="18FC3CDBFFF7257CFE0B20BABEF3FA69" ID-GBIF-Occurrence="2580042147" country="China" location="Sichuan" pageId="7" pageNumber="600" specimenCount="1" stateProvince="Sichuan">
2015/50;
<collectingCountry id="D0837616FFF7257CFDAC20BABC10FA26" box="[530,591,1413,1437]" name="China" pageId="7" pageNumber="600">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF7257CFDE920BABCF9FA26" box="[599,678,1413,1437]" country="China" name="Sichuan" pageId="7" pageNumber="600">Sichuan</collectingRegion>
, Chengdu Shi, Dujiangyan Shi, Qingcheng Shan, Buyun Cableway upper station
</materialsCitation>
–
<materialsCitation id="18FC3CDBFFF7257CFF7D2085BD54FA57" ID-GBIF-Occurrence="2580042128" collectingDate="2015-06-06" collectorName="A. Hunyadi" country="China" elevation="1150" latitude="30.91305" location="Shangqinggong" longLatPrecision="1" longitude="103.5619" pageId="7" pageNumber="600" specimenCount="7" specimenCount-juvenile="7" stateProvince="Sichuan">
<location id="AD4B605DFFF7257CFF7D2085BF01FA69" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF7257CFF7D2085BF01FA69" box="[195,350,1466,1490]" country="China" latitude="30.91305" longLatPrecision="1" longitude="103.5619" name="Shangqinggong" pageId="7" pageNumber="600" stateProvince="Sichuan">Shangqinggong</location>
;
<elevation id="23B9D1B5FFF7257CFED82085BFBCFA69" box="[358,483,1466,1490]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="600" unit="m" value="1150.0">
<quantity id="6F6C9B63FFF7257CFED82085BFEEFA6A" box="[358,433,1466,1490]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="600" unit="m" value="1150.0">1150 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="CDA05041FFF7257CFE532085BC32FA6A" box="[493,621,1466,1490]" degrees="30" direction="north" minutes="54.783" orientation="latitude" pageId="7" pageNumber="600" precision="1" value="30.91305">30°54.783’N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFF7257CFDC62085BD5CFA69" box="[632,771,1466,1490]" degrees="103" direction="east" minutes="33.714" orientation="longitude" pageId="7" pageNumber="600" precision="1" value="103.5619">103°33.714’E</geoCoordinate>
;
<collectorName id="05615350FFF7257CFF3A20EBBEA6FA57" box="[132,249,1492,1516]" pageId="7" pageNumber="600">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFF7257CFE8F20EBBFF9FA57" box="[305,422,1492,1516]" pageId="7" pageNumber="600" value="2015-06-06">
<collectingDate id="CC6EE9AEFFF7257CFE8F20EBBFF9FA57" box="[305,422,1492,1516]" pageId="7" pageNumber="600" value="2015-06-06">06.VI.2015</collectingDate>
</date>
; coll. HA/
<specimenCount id="BE92FD0FFFF7257CFDA920EABC26FA57" box="[535,633,1492,1516]" pageId="7" pageNumber="600" type="juvenile">7 juvenile</specimenCount>
/broken shells
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF7257CFF1B20D0BEB0F9ED" ID-GBIF-Occurrence="2580042136" collectingDate="2015-06-04" collectorName="A. Hunyadi" country="China" county="Chengdu Shi" elevation="1530" latitude="30.946434" location="Qingcheng Houshan" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" pageId="7" pageNumber="600" specimenCount="2" specimenCount-juvenile="2" stateProvince="Sichuan">
2015/45;
<collectingRegion id="6A50F864FFF7257CFEB920D0BF07F9BC" box="[263,344,1519,1543]" country="China" name="Sichuan" pageId="7" pageNumber="600">Sichuan</collectingRegion>
,
<collectingCounty id="414A4E0AFFF7257CFEDD20D0BFB8F9BC" box="[355,487,1519,1543]" pageId="7" pageNumber="600">Chengdu Shi</collectingCounty>
,
<collectingMunicipality id="484FACFCFFF7257CFE4C20D0BCD1F9BC" box="[498,654,1519,1543]" pageId="7" pageNumber="600">Dujiangyan Shi</collectingMunicipality>
,
<location id="AD4B605DFFF7257CFD2420D0BEBEF99A" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF7257CFD2420D0BEBEF99A" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Qingcheng Houshan" pageId="7" pageNumber="600" stateProvince="Sichuan">Qingcheng Houshan</location>
,
<location id="AD4B605DFFF7257CFF522336BF3CF99A" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF7257CFF522336BF3CF99A" box="[236,355,1545,1569]" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Taian Zhen" pageId="7" pageNumber="600" stateProvince="Sichuan">Taian Zhen</location>
,
<location id="AD4B605DFFF7257CFED02336BFB5F99A" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF7257CFED02336BFB5F99A" box="[366,490,1545,1569]" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Baiyun Cun" pageId="7" pageNumber="600" stateProvince="Sichuan">Baiyun Cun</location>
,
<location id="AD4B605DFFF7257CFE482336BC2DF99A" LSID="urn:lsid:plazi:treatment:203D8790FFF52573FC1123ECBF46F856:AD4B605DFFF7257CFE482336BC2DF99A" box="[502,626,1545,1569]" country="China" county="Chengdu Shi" latitude="30.946434" longLatPrecision="1" longitude="103.47645" municipality="Dujiangyan Shi" name="Jiusengdong" pageId="7" pageNumber="600" stateProvince="Sichuan">Jiusengdong</location>
;
<elevation id="23B9D1B5FFF7257CFDC32336BD5DF99A" box="[637,770,1545,1569]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="7" pageNumber="600" unit="m" value="1530.0">
<quantity id="6F6C9B63FFF7257CFDC32336BC93F99A" box="[637,716,1545,1569]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="7" pageNumber="600" unit="m" value="1530.0">1530 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="CDA05041FFF7257CFF3A231CBF5FF980" box="[132,256,1571,1595]" degrees="30" direction="north" minutes="56.786" orientation="latitude" pageId="7" pageNumber="600" precision="1" value="30.946434">30°56.786’N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFF7257CFEB5231CBFCEF980" box="[267,401,1571,1595]" degrees="103" direction="east" minutes="28.587" orientation="longitude" pageId="7" pageNumber="600" precision="1" value="103.47645">103°28.587’E</geoCoordinate>
;
<collectorName id="05615350FFF7257CFE27231BBC58F987" box="[409,519,1572,1596]" pageId="7" pageNumber="600">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFF7257CFD89231CBCF7F980" box="[567,680,1571,1595]" pageId="7" pageNumber="600" value="2015-06-04">
<collectingDate id="CC6EE9AEFFF7257CFD89231CBCF7F980" box="[567,680,1571,1595]" pageId="7" pageNumber="600" value="2015-06-04">04.VI.2015</collectingDate>
</date>
; coll. HA/
<specimenCount id="BE92FD0FFFF7257CFF3A2301BEB0F9ED" box="[132,239,1598,1622]" pageId="7" pageNumber="600" type="juvenile">2 juveniles</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF7257CFF3A2352BEA3F900" pageId="7" pageNumber="600" type="etymology">
<paragraph id="A82B3686FFF7257CFF3A2352BEA3F900" blockId="7.[132,775,1645,1723]" pageId="7" pageNumber="600">ETYMOLOGY. — This new species is named after Liu Zheng-Ping, who has made important contributions to the discovery of this new species.</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF7257CFF3A23EDBF82F8BE" pageId="7" pageNumber="600" type="distribution">
<paragraph id="A82B3686FFF7257CFF3A23EDBF82F8BE" blockId="7.[132,775,1746,1797]" pageId="7" pageNumber="600">
TYPE LOCALITY. —
<collectingCountry id="D0837616FFF7257CFE8023EDBF22F951" box="[318,381,1746,1770]" name="China" pageId="7" pageNumber="600">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF7257CFE3823EDBF88F951" box="[390,471,1746,1770]" country="China" name="Sichuan" pageId="7" pageNumber="600">Sichuan</collectingRegion>
, Chengdu Shi, Dujiangyan Shi, Qingcheng Houshan,
<quantity id="6F6C9B63FFF7257CFEE023D2BFF4F8BF" box="[350,427,1773,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="600" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF7257CFF3A2223BF5EF8D2" pageId="7" pageNumber="600" type="diagnosis">
<paragraph id="A82B3686FFF7257CFF3A2223BF5EF8D2" blockId="7.[132,775,1820,1897]" pageId="7" pageNumber="600">
DIAGNOSIS. — A
<taxonomicName id="6F944D05FFF7257CFE8A2223BFFEF88F" authorityName="Pilsbry" authorityYear="1895" box="[308,417,1820,1844]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="600" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF7257CFE8A2223BFFEF88F" box="[308,417,1820,1844]" italics="true" pageId="7" pageNumber="600">Coccoglypta</emphasis>
</taxonomicName>
species with a rounded body whorl, greenish to yellowish colour and weak sculpture on the ventral shell surface.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF72573FF3A22ACBCA1FAAB" lastPageId="8" lastPageNumber="601" pageId="7" pageNumber="600" type="description">
<paragraph id="A82B3686FFF7257CFF3A22ACBF4AF810" blockId="7.[132,775,1939,2030]" box="[132,277,1939,1965]" pageId="7" pageNumber="600">DESCRIPTION</paragraph>
<paragraph id="A82B3686FFF7257CFF3A228CBA5EF836" blockId="7.[132,775,1939,2030]" lastBlockId="7.[813,1462,1365,1933]" pageId="7" pageNumber="600">Shell dextral, rather large, light brown (dorsal surface) to greenish or yellowish (ventral surface, and dorsal side behind the aperture); colour changing below keel, and some light colouration indicating some growth lines; shell depressed globular, body whorl rounded or with very slightly marked keel (if present, mostly visible from ventral view of the body whorl); entire shell consisting of 5.25-5.75 whorls, separated by rather shallow suture, sometimes indicated by a whitish line; protoconch consisting of 1.5-1.75 whorls, matt, rather smooth, dorsal side of teleoconch dominated by irregular, rather rough wrinkles and tubercles (tubercles appearing after the first 1.5 whorls of teleoconch); some fine spiral grooves also visible; ventral surface without or with much less prominent tubercles, rather “hammered”, irregularly wrinkled and spirally grooved; aperture strongly oblique to shell axis, white; peristome expanded (mostly in basal and umbilical direction) but not reflected; parietal callus weak, only indicated by fine whitish calcareous layer, which is transparent in fresh shells; umbilicus open, moderately wide, showing all whorls.</paragraph>
<paragraph id="A82B3686FFF7257CFC93228BBDE1F876" blockId="7.[813,1331,1972,2030]" box="[813,958,1972,1997]" pageId="7" pageNumber="600">
<emphasis id="9AE0EA94FFF7257CFC93228BBDE1F876" box="[813,958,1972,1997]" italics="true" pageId="7" pageNumber="600">Measurements</emphasis>
</paragraph>
<paragraph id="A82B3686FFF72573FC9322EBBFDCFBEB" blockId="7.[813,1331,1972,2030]" lastBlockId="8.[132,775,1078,1296]" lastPageId="8" lastPageNumber="601" pageId="7" pageNumber="600">
D =
<quantity id="6F6C9B63FFF7257CFCE022ECBDA0F855" box="[862,1023,2003,2030]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.29" metricValueMin="2.91" pageId="7" pageNumber="600" unit="mm" value="31.0" valueMax="32.9" valueMin="29.1">29.1-32.9 mm</quantity>
, H =
<quantity id="6F6C9B63FFF7257CFB8322ECBA84F855" box="[1085,1243,2003,2030]" metricMagnitude="-2" metricUnit="m" metricValue="1.835" metricValueMax="2.06" metricValueMin="1.61" pageId="7" pageNumber="600" unit="mm" value="18.35" valueMax="20.6" valueMin="16.1">16.1-20.6 mm</quantity>
(n = 3).
<emphasis id="9AE0EA94FFF82573FF3A2109BFDCFBEB" box="[132,387,1078,1104]" italics="true" pageId="8" pageNumber="601">Remarks on the genitalia</emphasis>
</paragraph>
<caption id="FCEB660EFFF82573FF3B2686BFC5FC59" ID-DOI="http://doi.org/10.5281/zenodo.3726118" ID-Zenodo-Dep="3726118" httpUri="https://zenodo.org/record/3726118/files/figure.png" pageId="8" pageNumber="601" startId="8.[133,144,953,971]" targetBox="[132,1455,215,894]" targetPageId="8">
<paragraph id="A82B3686FFF82573FF3B2686BFC5FC59" blockId="8.[133,1455,953,994]" pageId="8" pageNumber="601">
FIG. 4. — Inner structure of the genitalia of
<taxonomicName id="6F944D05FFF82573FE572686BC83FC70" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[489,732,953,971]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="liui" status="sp. nov.">
<emphasis id="9AE0EA94FFF82573FE572686BC30FC70" box="[489,623,953,971]" italics="true" pageId="8" pageNumber="601">Coccoglypta liui</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="81D357EFFFF82573FD5D2686BD4CFC70" box="[739,787,953,971]" pageId="8" pageNumber="601" rank="species">n. sp.</taxonomicNameLabel>
(holotype, HNHM 103475):
<emphasis id="9AE0EA94FFF82573FC442685BA58FC77" bold="true" box="[1018,1031,954,972]" pageId="8" pageNumber="601">A</emphasis>
, penis;
<emphasis id="9AE0EA94FFF82573FBF62685BA0AFC77" bold="true" box="[1096,1109,954,972]" pageId="8" pageNumber="601">B</emphasis>
, vagina. Genital atrium situated leftwards in both images. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="A82B3686FFF82573FF3A2168BCA1FAAB" blockId="8.[132,775,1078,1296]" pageId="8" pageNumber="601">
From a single specimen anatomically examined from Qingcheng Houshan (
<figureCitation id="30AF2A03FFF82573FF492149BF0CFB2B" box="[247,339,1142,1168]" captionStart-0="FIG" captionStart-1="FIG" captionStartId-0="7.[132,143,1273,1291]" captionStartId-1="8.[133,144,953,971]" captionTargetBox-0="[165,1416,232,1205]" captionTargetBox-1="[132,1455,215,894]" captionTargetId-0="figure@7.[132,1455,216,1236]" captionTargetId-1="figure@8.[132,1455,215,894]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIG. 3. — Reproductive anatomy of Coccoglypta liui Páll-Gergely, n. sp. (holotype, HNHM 103475): A, entire genitalia; B, schematic drawing of the inner structure of the dart sac. Scale bar: A, 5 mm." captionText-1="FIG. 4. — Inner structure of the genitalia of Coccoglypta liui Páll-Gergely, n. sp. (holotype, HNHM 103475): A, penis; B, vagina. Genital atrium situated leftwards in both images. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3726116" figureDoi-1="http://doi.org/10.5281/zenodo.3726118" httpUri-0="https://zenodo.org/record/3726116/files/figure.png" httpUri-1="https://zenodo.org/record/3726118/files/figure.png" pageId="8" pageNumber="601">Figs 3-4</figureCitation>
). Right ommatophoral retractor crossed vagina and penis. Inner wall of the penis finely reticulated (
<figureCitation id="30AF2A03FFF82573FF332189BE82FB6B" box="[141,221,1206,1232]" captionStart="FIG" captionStartId="8.[133,144,953,971]" captionTargetBox="[132,1455,215,894]" captionTargetId="figure@8.[132,1455,215,894]" captionTargetPageId="8" captionText="FIG. 4. — Inner structure of the genitalia of Coccoglypta liui Páll-Gergely, n. sp. (holotype, HNHM 103475): A, penis; B, vagina. Genital atrium situated leftwards in both images. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3726118" httpUri="https://zenodo.org/record/3726118/files/figure.png" pageId="8" pageNumber="601">Fig. 4A</figureCitation>
), produced by the perpendicular projections of the slender longitudinal folds. Inner wall of vagina with widelyspaced, sometimes converging longitudinal folds (
<figureCitation id="30AF2A03FFF82573FD2121CABCB2FAAB" box="[671,749,1269,1296]" captionStart="FIG" captionStartId="8.[133,144,953,971]" captionTargetBox="[132,1455,215,894]" captionTargetId="figure@8.[132,1455,215,894]" captionTargetPageId="8" captionText="FIG. 4. — Inner structure of the genitalia of Coccoglypta liui Páll-Gergely, n. sp. (holotype, HNHM 103475): A, penis; B, vagina. Genital atrium situated leftwards in both images. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3726118" httpUri="https://zenodo.org/record/3726118/files/figure.png" pageId="8" pageNumber="601">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FF3A2008BFC1F9B2" pageId="8" pageNumber="601" type="diagnosis">
<paragraph id="A82B3686FFF82573FF3A2008BFC1F9B2" blockId="8.[132,776,1335,1545]" pageId="8" pageNumber="601">
DIFFERENTIAL DIAGNOSIS. —
<taxonomicName id="6F944D05FFF82573FE132008BC9AFAF4" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[429,709,1335,1359]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="liui" status="sp. nov.">
<emphasis id="9AE0EA94FFF82573FE132008BC1CFAF4" box="[429,579,1335,1359]" italics="true" pageId="8" pageNumber="601">Coccoglypta liui</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="81D357EFFFF82573FD6E2007BD57FAF4" box="[720,776,1336,1359]" pageId="8" pageNumber="601" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="6F944D05FFF82573FF44206DBFD5FAD1" authorityName="Heude" authorityYear="1886" box="[250,394,1362,1386]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF82573FF44206DBFD5FAD1" box="[250,394,1362,1386]" italics="true" pageId="8" pageNumber="601">C. pinchoniana</emphasis>
</taxonomicName>
, which also occurs sympatrically, by the usually narrower umbilicus, weaker sculpture of the ventral surface, the lighter colour, and the less keeled body whorl. The ventral side of
<taxonomicName id="6F944D05FFF82573FF21209DBF6EFA02" authorityName="Heude" authorityYear="1886" box="[159,305,1441,1465]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF82573FF21209DBF6EFA02" box="[159,305,1441,1465]" italics="true" pageId="8" pageNumber="601">C. pinchoniana</emphasis>
</taxonomicName>
is dominated by ribs and tubercles, but also has fine spiral grooves. However, the ventral shell surface of
<taxonomicName id="6F944D05FFF82573FD242083BF7AFA55" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="liui" status="sp. nov.">
<emphasis id="9AE0EA94FFF82573FD242083BEFAFA55" italics="true" pageId="8" pageNumber="601">Coccoglypta liui</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="81D357EFFFF82573FE8E20E8BF38FA55" box="[304,359,1495,1518]" pageId="8" pageNumber="601" rank="species">n. sp.</taxonomicNameLabel>
has much less prominent tubercles and is dominated by spiral grooves.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FF3A230FBD59F9D8" pageId="8" pageNumber="601" type="distribution">
<paragraph id="A82B3686FFF82573FF3A230FBD59F9D8" blockId="8.[132,775,1584,1635]" pageId="8" pageNumber="601">
DISTRIBUTION. — So far this species is known only from Qingcheng Houshan and Qingcheng Shan in
<collectingRegion id="6A50F864FFF82573FE772374BC2EF9D8" box="[457,625,1611,1635]" country="China" name="Sichuan" pageId="8" pageNumber="601">Sichuan Province</collectingRegion>
,
<collectingCountry id="D0837616FFF82573FDC42374BCE9F9D8" box="[634,694,1611,1635]" name="China" pageId="8" pageNumber="601">China</collectingCountry>
(
<figureCitation id="30AF2A03FFF82573FD7C2374BCA6F9D8" box="[706,761,1611,1635]" captionStart="FIG" captionStartId="9.[132,143,1206,1224]" captionTargetBox="[244,1340,215,1147]" captionTargetId="figure@9.[244,1340,214,1147]" captionTargetPageId="9" captionText="FIG. 5. — Map of Sichuan showing the localities of Coccoglypta Pilsbry, 1895 species. Dashed line shows the edge of the Tibetan Plateau. Coccoglypta pinchoniana (Heude, 1886) was described from Chengdu, and it was known from the Emeishan, and a fossil record from Chongqing. Now we report this species and C. liui Páll-Gergely, n. sp. from Qingcheng Shan and Qingcheng Houshan (black area)." figureDoi="http://doi.org/10.5281/zenodo.3726120" httpUri="https://zenodo.org/record/3726120/files/figure.png" pageId="8" pageNumber="601">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FF3A23ACBF46F856" pageId="8" pageNumber="601" type="discussion">
<paragraph id="A82B3686FFF82573FF3A23ACBEB7F917" blockId="8.[132,776,1683,2029]" box="[132,232,1683,1709]" pageId="8" pageNumber="601">REMARKS</paragraph>
<paragraph id="A82B3686FFF82573FF3A238BBF46F856" blockId="8.[132,776,1683,2029]" pageId="8" pageNumber="601">
Our observations on the genitalia generally match with those of
<bibRefCitation id="CC054B77FFF82573FF1D23EBBF3CF955" author="SCHILEYKO A. A." box="[163,355,1748,1774]" pageId="8" pageNumber="601" pagination="1627 - 1763" refId="ref8091" refString="SCHILEYKO A. A. 2004. - Treatise on Recent terrestrial pulmonate molluscs Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1763." type="journal article" year="2004">Schileyko (2004)</bibRefCitation>
, based on a
<taxonomicName id="6F944D05FFF82573FE4C23EBBCC7F955" authorityName="Heude" authorityYear="1886" box="[498,664,1748,1774]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF82573FE4C23EBBCC7F955" box="[498,664,1748,1774]" italics="true" pageId="8" pageNumber="601">C. pinchoniana</emphasis>
</taxonomicName>
specimen collected in Emei Shan. The notable differences are the following: 1) the mucus glands form a single oval mass, not separated clearly as on
<bibRefCitation id="CC054B77FFF82573FE36220CBC0BF8F5" author="SCHILEYKO A. A." box="[392,596,1843,1870]" pageId="8" pageNumber="601" pagination="1627 - 1763" refId="ref8091" refString="SCHILEYKO A. A. 2004. - Treatise on Recent terrestrial pulmonate molluscs Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1763." type="journal article" year="2004">Schileyko’s (2004)</bibRefCitation>
drawing; 2) we have not found a twisted part of the spermathecal stalk in a capsule; and 3) the bursa copulatrix is smaller and more elongated than the specimen examined by
<bibRefCitation id="CC054B77FFF82573FDF622ACBD5DF815" author="SCHILEYKO A. A." box="[584,770,1939,1966]" pageId="8" pageNumber="601" pagination="1627 - 1763" refId="ref8091" refString="SCHILEYKO A. A. 2004. - Treatise on Recent terrestrial pulmonate molluscs Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1763." type="journal article" year="2004">Schileyko (2004)</bibRefCitation>
. It is yet unclear which of these differences represents speciesspecific traits.
</paragraph>
</subSubSection>
</treatment>
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<heading id="F36381EAFFF82573FC2D210ABB16FBEA" box="[915,1353,1077,1105]" level="2" pageId="8" pageNumber="601">
<taxonomicName id="6F944D05FFF82573FC2D210ABB16FBEA" authority="(Heude, 1886)" baseAuthorityName="Heude" baseAuthorityYear="1886" box="[915,1353,1077,1105]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF82573FC2D210ABAFEFBEB" bold="true" box="[915,1185,1077,1104]" italics="true" pageId="8" pageNumber="601">Coccoglypta pinchoniana</emphasis>
(
<bibRefCitation id="CC054B77FFF82573FB0E2109BB1EFBEA" author="HEUDE R. P. M." box="[1200,1345,1078,1105]" pageId="8" pageNumber="601" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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(
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)
</paragraph>
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<subSubSection id="E08E650DFFF82573FC9321B0BAE2FA9D" pageId="8" pageNumber="601" type="reference_group">
<paragraph id="A82B3686FFF82573FC9321B0BAC3FB1C" blockId="8.[813,1180,1167,1191]" box="[813,1180,1167,1191]" pageId="8" pageNumber="601">
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<taxonomicName id="6F944D05FFF82573FC9321B0BAC7FB1C" authority="Heude, 1886: 213" authorityName="Heude" authorityPageNumber="213" authorityYear="1886" box="[813,1176,1167,1191]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF82573FC9321B0BD84FB1C" box="[813,987,1167,1191]" italics="true" pageId="8" pageNumber="601">Helix pinchoniana</emphasis>
<bibRefCitation id="CC054B77FFF82573FC5C21B0BAC7FB1C" author="HEUDE R. P. M." box="[994,1176,1167,1191]" pageId="8" pageNumber="601" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886: 213</bibRefCitation>
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.
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<emphasis id="9AE0EA94FFF82573FC932181BA49FB6D" box="[813,1046,1214,1238]" italics="true" pageId="8" pageNumber="601">Coccoglypta pinchoniana</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFF82573FB8E2180BAAEFB6D" author="YEN T. - C." box="[1072,1265,1214,1238]" page="335" pageId="8" pageNumber="601" year="1936">
<bibRefCitation id="CC054B77FFF82573FB8E2180BAAEFB6D" author="YEN T. - C." box="[1072,1265,1214,1238]" pageId="8" pageNumber="601" pagination="331 - 342" refId="ref8619" refString="YEN T. - C. 1936. - Some Quaternary gastropods from eastern Szechwan. Bulletin of the Geological Society of China 15 (3): 331 - 342. https: // doi. org / 10.1111 / j. 1755 - 6724.1936. mp 15003005. x" type="journal article" year="1936">Yen 1936: 335-336</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="29351097FFF82573FB422180BB3BFB6D" author="YEN T. - C." box="[1276,1380,1214,1239]" page="153" pageId="8" pageNumber="601" year="1939">
<bibRefCitation id="CC054B77FFF82573FB422180BB3BFB6D" author="YEN T. - C." box="[1276,1380,1214,1239]" pageId="8" pageNumber="601" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">1939: 153</bibRefCitation>
</treatmentCitation>
, pl. 15, fig. 64. — Zilch 1960: 640-641, fig. 2242. —
<treatmentCitation id="29351097FFF82573FB4F21E6BDDCFAB0" author="CHEN D-N. &amp; GAO J. - X." page="158" pageId="8" pageNumber="601" year="1987">
<bibRefCitation id="CC054B77FFF82573FB4F21E6BDDCFAB0" author="CHEN D-N. &amp; GAO J. - X." pageId="8" pageNumber="601" refId="ref7112" refString="CHEN D-N. &amp; GAO J. - X. 1987. - Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing, China. 186 p. (in Chinese)" type="book" year="1987">Chen &amp; Gao 1987: 158-159</bibRefCitation>
</treatmentCitation>
, fig. 204. —
<treatmentCitation id="29351097FFF82573FBB521CCBB7DFAB0" author="SCHILEYKO A. A." box="[1035,1314,1267,1292]" page="1678" pageId="8" pageNumber="601" year="2004">
<bibRefCitation id="CC054B77FFF82573FBB521CCBB7DFAB0" author="SCHILEYKO A. A." box="[1035,1314,1267,1292]" pageId="8" pageNumber="601" pagination="1627 - 1763" refId="ref8091" refString="SCHILEYKO A. A. 2004. - Treatise on Recent terrestrial pulmonate molluscs Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1763." type="journal article" year="2004">
Schileyko 2004:
<date id="DC2A1046FFF82573FB0C21CCBB7DFAB0" box="[1202,1314,1267,1291]" pageId="8" pageNumber="601" value="1678" valueMax="1679">1678-1679</date>
</bibRefCitation>
</treatmentCitation>
, fig. 2164. —
<treatmentCitation id="29351097FFF82573FC932031BA3FFA9D" author="CHEN D. - N. &amp; ZHANG G. - Q." box="[813,1120,1294,1318]" page="371" pageId="8" pageNumber="601" year="2004">
<bibRefCitation id="CC054B77FFF82573FC932031BA3FFA9D" author="CHEN D. - N. &amp; ZHANG G. - Q." box="[813,1120,1294,1318]" pageId="8" pageNumber="601" refId="ref7148" refString="CHEN D. - N. &amp; ZHANG G. - Q. 2004. - Fauna Sinica (Invertebrata Vol. 37 Mollusca: Gastropoda: Stylommatophora: Bradybaenidae). Science Press, Beijing, China, 482 p. (in Chinese)." type="book" year="2004">Chen &amp; Zhang 2004: 371-372</bibRefCitation>
</treatmentCitation>
, fig. 370.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FC932000BAC9F95B" pageId="8" pageNumber="601" type="materials_examined">
<paragraph id="A82B3686FFF82573FC932000BDE0FACA" blockId="8.[813,1455,1343,1394]" pageId="8" pageNumber="601">
<collectionCode id="CE85AE43FFF82573FC932000BD3FFAEF" box="[813,864,1343,1367]" pageId="8" pageNumber="601">TYPE</collectionCode>
<collectionCode id="CE85AE43FFF82573FCD7207BBA63FAEE" box="[873,1084,1348,1365]" pageId="8" pageNumber="601">MATERIAL EXAMINED</collectionCode>
. —
<materialsCitation id="18FC3CDBFFF82573FBD02000BDE4FACA" ID-GBIF-Occurrence="2580042142" collectionCode="USNM, MCZ" pageId="8" pageNumber="601" specimenCode="USNM 472156, MCZ 167190" specimenCount="1" typeStatus="syntype">
<emphasis id="9AE0EA94FFF82573FBD02000BA93FAEC" bold="true" box="[1134,1228,1343,1367]" pageId="8" pageNumber="601">
<typeStatus id="772F8824FFF82573FBD02000BA93FAEC" box="[1134,1228,1343,1367]" pageId="8" pageNumber="601" type="syntype">Syntypes</typeStatus>
</emphasis>
.
<specimenCode id="F8329EFDFFF82573FB652000BBDEFAEC" box="[1243,1409,1343,1367]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="601" type="Museum">USNM 472156</specimenCode>
and
<specimenCode id="F8329EFDFFF82573FC932065BDE4FACA" box="[813,955,1369,1394]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="601" type="Museum">MCZ 167190</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="A82B3686FFF82573FC9320B5BAC9F95B" blockId="8.[813,1456,1418,1760]" pageId="8" pageNumber="601">
ADDITIONAL
<collectionCode id="CE85AE43FFF82573FC0920B0BAD5FA1B" box="[951,1162,1423,1440]" pageId="8" pageNumber="601">MATERIAL EXAMINED</collectionCode>
. —
<materialsCitation id="18FC3CDBFFF82573FB0320B4BAD2FA06" ID-GBIF-Occurrence="2580042150" collectionCode="SMF" collectorName="Schmacker" location="Sytchuan" municipality="Berg Omi" pageId="8" pageNumber="601" specimenCount="1">
“
<collectingMunicipality id="484FACFCFFF82573FB7820B4BB6CFA18" box="[1222,1331,1419,1443]" pageId="8" pageNumber="601">Berg Omi</collectingMunicipality>
,
<location id="AD4B605DFFF82573FAFE20B4BBFDFA18" LSID="urn:lsid:plazi:treatment:203D8790FFF82572FC2D210ABB3FF994:AD4B605DFFF82573FAFE20B4BBFDFA18" box="[1344,1442,1419,1443]" municipality="Berg Omi" name="Sytchuan" pageId="8" pageNumber="601">Sytchuan</location>
”;
<collectorName id="05615350FFF82573FC93209ABDC7FA06" box="[813,920,1445,1469]" pageId="8" pageNumber="601">Schmacker</collectorName>
leg. 1893;
<collectionCode id="CE85AE43FFF82573FBBE209ABA6EFA06" box="[1024,1073,1445,1469]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="601">SMF</collectionCode>
42574/2
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF82573FB0D209ABD88FA49" ID-GBIF-Occurrence="2580042145" collectorName="Dujiangyan Shi &amp; Qingcheng Shan &amp; Buyun Cableway" country="China" location="Sichuan" pageId="8" pageNumber="601" specimenCount="1" stateProvince="Sichuan">
2015/50;
<collectingCountry id="D0837616FFF82573FAAD209ABB0DFA06" box="[1299,1362,1445,1469]" name="China" pageId="8" pageNumber="601">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF82573FAE5209ABBF3FA06" box="[1371,1452,1445,1469]" country="China" name="Sichuan" pageId="8" pageNumber="601">Sichuan</collectingRegion>
, Chengdu Shi,
<collectorName id="05615350FFF82573FC7E2080BA3FFA6C" box="[960,1120,1471,1495]" pageId="8" pageNumber="601">Dujiangyan Shi</collectorName>
,
<collectorName id="05615350FFF82573FBD22080BB49FA6C" box="[1132,1302,1471,1495]" pageId="8" pageNumber="601">Qingcheng Shan</collectorName>
,
<collectorName id="05615350FFF82573FA9C20FFBD0CFA49" pageId="8" pageNumber="601">Buyun Cableway</collectorName>
felső állomás
</materialsCitation>
–
<materialsCitation id="18FC3CDBFFF82573FC4D20E5BB74F9B6" ID-GBIF-Occurrence="2580042144" collectingDate="2015-06-06" collectionCode="VI" collectorName="A. Hunyadi" country="China" elevation="1150" latitude="30.91305" location="Shangqinggong" longLatPrecision="1" longitude="103.5619" pageId="8" pageNumber="601" specimenCount="1" stateProvince="Sichuan">
<location id="AD4B605DFFF82573FC4D20E5BAD1FA49" LSID="urn:lsid:plazi:treatment:203D8790FFF82572FC2D210ABB3FF994:AD4B605DFFF82573FC4D20E5BAD1FA49" box="[1011,1166,1498,1522]" country="China" latitude="30.91305" longLatPrecision="1" longitude="103.5619" name="Shangqinggong" pageId="8" pageNumber="601" stateProvince="Sichuan">Shangqinggong</location>
;
<elevation id="23B9D1B5FFF82573FB2720E5BB43FA49" box="[1177,1308,1498,1522]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="601" unit="m" value="1150.0">
<quantity id="6F6C9B63FFF82573FB2720E5BAB8FA4A" box="[1177,1255,1498,1522]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="601" unit="m" value="1150.0">1150 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="CDA05041FFF82573FA9620E5BBF6FA4A" box="[1320,1449,1498,1522]" degrees="30" direction="north" minutes="54.783" orientation="latitude" pageId="8" pageNumber="601" precision="1" value="30.91305">30°54.783’N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFF82573FC9320CABDF0F9B6" box="[813,943,1525,1549]" degrees="103" direction="east" minutes="33.714" orientation="longitude" pageId="8" pageNumber="601" precision="1" value="103.5619">103°33.714’E</geoCoordinate>
;
<collectorName id="05615350FFF82573FC0920CABA7CF9B7" box="[951,1059,1524,1549]" pageId="8" pageNumber="601">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFF82573FBEF20CBBAE0F9B7" box="[1105,1215,1524,1548]" pageId="8" pageNumber="601" value="2015-06-06">
<collectingDate id="CC6EE9AEFFF82573FBEF20CBBAE0F9B7" box="[1105,1215,1524,1548]" pageId="8" pageNumber="601" value="2015-06-06">
06.
<collectionCode id="CE85AE43FFF82573FBD320CBBAD7F9B7" box="[1133,1160,1524,1548]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="601">VI</collectionCode>
.2015
</collectingDate>
</date>
, coll. HA/1
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF82573FAF120CBBA7CF9E7" ID-GBIF-Occurrence="2580042151" collectingDate="2015-06-06" collectionCode="VI" collectorName="Chaoyangdong &amp; Shangqinggong &amp; A. Hunyadi" country="China" county="Chengdu Shi" elevation="50" location="Qingcheng Shan" municipality="Dujiangyan Shi" pageId="8" pageNumber="601" specimenCount="1" stateProvince="Sichuan">
2015/51a;
<collectingCountry id="D0837616FFF82573FC932330BD33F99C" box="[813,876,1551,1575]" name="China" pageId="8" pageNumber="601">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF82573FCC82330BD98F99C" box="[886,967,1551,1575]" country="China" name="Sichuan" pageId="8" pageNumber="601">Sichuan</collectingRegion>
,
<collectingCounty id="414A4E0AFFF82573FC6F2330BA0AF99C" box="[977,1109,1551,1575]" pageId="8" pageNumber="601">Chengdu Shi</collectingCounty>
,
<collectingMunicipality id="484FACFCFFF82573FBE12330BAA5F99C" box="[1119,1274,1551,1575]" pageId="8" pageNumber="601">Dujiangyan Shi</collectingMunicipality>
,
<location id="AD4B605DFFF82573FABB2330BBF4F99C" LSID="urn:lsid:plazi:treatment:203D8790FFF82572FC2D210ABB3FF994:AD4B605DFFF82573FABB2330BBF4F99C" box="[1285,1451,1551,1575]" country="China" county="Chengdu Shi" municipality="Dujiangyan Shi" name="Qingcheng Shan" pageId="8" pageNumber="601" stateProvince="Sichuan">Qingcheng Shan</location>
,
<collectorName id="05615350FFF82573FC932316BD9BF9FA" box="[813,964,1577,1601]" pageId="8" pageNumber="601">Chaoyangdong</collectorName>
<quantity id="6F6C9B63FFF82573FC722316BA5EF9FA" box="[972,1025,1577,1601]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="601" unit="m" value="50.0">
<elevation id="23B9D1B5FFF82573FC722316BA5EF9FA" box="[972,1025,1577,1601]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="601" unit="m" value="50.0">50 m</elevation>
</quantity>
towards
<collectorName id="05615350FFF82573FBE12316BAA3F9FA" box="[1119,1276,1577,1601]" pageId="8" pageNumber="601">Shangqinggong</collectorName>
;
<collectorName id="05615350FFF82573FAB62315BB21F9FA" box="[1288,1406,1577,1602]" pageId="8" pageNumber="601">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFF82573FC93237BBDFAF9E7" box="[813,933,1604,1628]" pageId="8" pageNumber="601" value="2015-06-06">
<collectingDate id="CC6EE9AEFFF82573FC93237BBDFAF9E7" box="[813,933,1604,1628]" pageId="8" pageNumber="601" value="2015-06-06">
06.
<collectionCode id="CE85AE43FFF82573FCF2237BBD35F9E7" box="[844,874,1604,1628]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="601">VI</collectionCode>
.2015
</collectingDate>
</date>
; coll. HA/2
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF82573FBEF237BBDC1F910" ID-GBIF-Occurrence="2580042149" collectingDate="2015-06-07" collectionCode="VI" collectorName="A. Hunyadi" country="China" county="Chengdu Shi" elevation="940" latitude="30.89555" location="Qingcheng Shan" longLatPrecision="1" longitude="103.55145" municipality="Dujiangyan Shi" pageId="8" pageNumber="601" specimenCount="1" stateProvince="Sichuan">
2015/55;
<collectingCountry id="D0837616FFF82573FB08237BBAA8F9E7" box="[1206,1271,1604,1628]" name="China" pageId="8" pageNumber="601">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF82573FABA237BBB07F9E7" box="[1284,1368,1604,1628]" country="China" name="Sichuan" pageId="8" pageNumber="601">Sichuan</collectingRegion>
,
<collectingCounty id="414A4E0AFFF82573FADA237BBD2DF9CD" pageId="8" pageNumber="601">Chengdu Shi</collectingCounty>
,
<collectingMunicipality id="484FACFCFFF82573FCC12361BA7FF9CD" box="[895,1056,1630,1654]" pageId="8" pageNumber="601">Dujiangyan Shi</collectingMunicipality>
,
<location id="AD4B605DFFF82573FB932361BA84F9CD" LSID="urn:lsid:plazi:treatment:203D8790FFF82572FC2D210ABB3FF994:AD4B605DFFF82573FB932361BA84F9CD" box="[1069,1243,1630,1654]" country="China" county="Chengdu Shi" latitude="30.89555" longLatPrecision="1" longitude="103.55145" municipality="Dujiangyan Shi" name="Qingcheng Shan" pageId="8" pageNumber="601" stateProvince="Sichuan">Qingcheng Shan</location>
,
<location id="AD4B605DFFF82573FB582361BB3FF9CD" LSID="urn:lsid:plazi:treatment:203D8790FFF82572FC2D210ABB3FF994:AD4B605DFFF82573FB582361BB3FF9CD" box="[1254,1376,1630,1654]" country="China" county="Chengdu Shi" latitude="30.89555" longLatPrecision="1" longitude="103.55145" municipality="Dujiangyan Shi" name="Jinbian Yan" pageId="8" pageNumber="601" stateProvince="Sichuan">Jinbian Yan</location>
;
<elevation id="23B9D1B5FFF82573FAD22361BD05F92A" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="8" pageNumber="601" unit="m" value="940.0">
<quantity id="6F6C9B63FFF82573FAD22361BBF0F9CD" box="[1388,1455,1630,1654]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="8" pageNumber="601" unit="m" value="940.0">940 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="CDA05041FFF82573FCD82346BDB8F92B" box="[870,999,1657,1681]" degrees="30" direction="north" minutes="53.733" orientation="latitude" pageId="8" pageNumber="601" precision="1" value="30.89555">30°53.733’N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFF82573FC4B2346BA20F92A" box="[1013,1151,1657,1681]" degrees="103" direction="east" minutes="33.087" orientation="longitude" pageId="8" pageNumber="601" precision="1" value="103.55145">103°33.087’E</geoCoordinate>
;
<collectorName id="05615350FFF82573FB352346BAA0F92A" box="[1163,1279,1657,1681]" pageId="8" pageNumber="601">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFF82573FA882346BBF4F92A" box="[1334,1451,1657,1681]" pageId="8" pageNumber="601" value="2015-06-07">
<collectingDate id="CC6EE9AEFFF82573FA882346BBF4F92A" box="[1334,1451,1657,1681]" pageId="8" pageNumber="601" value="2015-06-07">
07.
<collectionCode id="CE85AE43FFF82573FAEA2346BB2EF92A" box="[1364,1393,1657,1681]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="601">VI</collectionCode>
.2015
</collectingDate>
</date>
; coll. HA/1
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFF82573FC7523ACBACDF95B" ID-GBIF-Occurrence="2580042146" collectingDate="2018-01-13" collectorName="Liu Zheng-Ping" country="China" county="Chongzhou Shi" location="Fengqishan" municipality="Jiezi Zhen" pageId="8" pageNumber="601" specimenCount="1" stateProvince="Sichuan">
<collectingCountry id="D0837616FFF82573FC7523ACBA53F910" box="[971,1036,1683,1707]" name="China" pageId="8" pageNumber="601">China</collectingCountry>
,
<collectingRegion id="6A50F864FFF82573FBA623ACBA94F910" box="[1048,1227,1683,1707]" country="China" name="Sichuan" pageId="8" pageNumber="601">Sichuan Province</collectingRegion>
,
<collectingCounty id="414A4E0AFFF82573FB6923ACBB26F910" box="[1239,1401,1683,1707]" pageId="8" pageNumber="601">Chongzhou Shi</collectingCounty>
,
<collectingMunicipality id="484FACFCFFF82573FA3B23ACBD3BF97D" pageId="8" pageNumber="601">Jiezi Zhen</collectingMunicipality>
,
<location id="AD4B605DFFF82573FCD32391BD80F97D" LSID="urn:lsid:plazi:treatment:203D8790FFF82572FC2D210ABB3FF994:AD4B605DFFF82573FCD32391BD80F97D" box="[877,991,1710,1734]" country="China" county="Chongzhou Shi" municipality="Jiezi Zhen" name="Fengqishan" pageId="8" pageNumber="601" stateProvince="Sichuan">Fengqishan</location>
;
<collectorName id="05615350FFF82573FC562391BADCF97D" box="[1000,1155,1710,1734]" pageId="8" pageNumber="601">Liu Zheng-Ping</collectorName>
leg.;
<date id="DC2A1046FFF82573FB082391BB46F97D" box="[1206,1305,1710,1734]" pageId="8" pageNumber="601" value="2018-01-13">
<collectingDate id="CC6EE9AEFFF82573FB082391BB46F97D" box="[1206,1305,1710,1734]" pageId="8" pageNumber="601" value="2018-01-13">13.I.2018</collectingDate>
</date>
; SYS m001014 (specimen used for molecular study)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FC9323C6BBEFF8AA" box="[813,1456,1785,1809]" pageId="8" pageNumber="601" type="distribution">
<paragraph id="A82B3686FFF82573FC9323C6BBEFF8AA" blockId="8.[813,1456,1785,1809]" box="[813,1456,1785,1809]" pageId="8" pageNumber="601">
TYPE LOCALITY. — Tchen-tou-fou (Chengdu city,
<collectingRegion id="6A50F864FFF82573FB4123C6BBF9F8AA" box="[1279,1446,1785,1809]" country="China" name="Sichuan" pageId="8" pageNumber="601">Sichuan Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FC93220EBDACF8C4" pageId="8" pageNumber="601" type="diagnosis">
<paragraph id="A82B3686FFF82573FC93220EBDACF8C4" blockId="8.[813,1456,1841,1919]" pageId="8" pageNumber="601">
DIAGNOSIS. — A
<taxonomicName id="6F944D05FFF82573FC5D220DBA0CF8F1" authorityName="Pilsbry" authorityYear="1895" box="[995,1107,1842,1866]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF82573FC5D220DBA0CF8F1" box="[995,1107,1842,1866]" italics="true" pageId="8" pageNumber="601">Coccoglypta</emphasis>
</taxonomicName>
species with a keeled body whorl, light brown, greenish to greyish colour and strong sculpture on the ventral shell surface.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF82573FC9322A0BA84F857" pageId="8" pageNumber="601" type="distribution">
<paragraph id="A82B3686FFF82573FC9322A0BA84F857" blockId="8.[813,1455,1951,2028]" pageId="8" pageNumber="601">
DISTRIBUTION. — This species is more widely distributed than
<taxonomicName id="6F944D05FFF82573FC932285BDB0F869" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[813,1007,1978,2002]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="601" phylum="Mollusca" rank="species" species="liui" status="sp. nov.">
<emphasis id="9AE0EA94FFF82573FC932285BD32F869" box="[813,877,1978,2002]" italics="true" pageId="8" pageNumber="601">C. liui</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="81D357EFFFF82573FC442285BA6DF86A" box="[1018,1074,1978,2001]" pageId="8" pageNumber="601" rank="species">n. sp.</taxonomicNameLabel>
Namely, it was collected on the Emei Shan, the Qingcheng Shan and Fengqishan.
</paragraph>
</subSubSection>
<caption id="FCEB660EFFF92572FF3A2189BD0EFB4D" ID-DOI="http://doi.org/10.5281/zenodo.3726120" ID-Zenodo-Dep="3726120" httpUri="https://zenodo.org/record/3726120/files/figure.png" pageId="9" pageNumber="602" startId="9.[132,143,1206,1224]" targetBox="[244,1340,215,1147]" targetPageId="9">
<paragraph id="A82B3686FFF92572FF3A2189BD0EFB4D" pageId="9" pageNumber="602">
FIG. 5. — Map of Sichuan showing the localities of
<taxonomicName id="6F944D05FFF92572FD972189BD5DFB73" authority="Pilsbry, 1895" authorityName="Pilsbry" authorityYear="1895" box="[553,770,1206,1224]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF92572FD972189BCCDFB73" box="[553,658,1206,1224]" italics="true" pageId="9" pageNumber="602">Coccoglypta</emphasis>
Pilsbry, 1895
</taxonomicName>
species. Dashed line shows the edge of the Tibetan Plateau.
<taxonomicName id="6F944D05FFF92572FABF2189BF74FB64" authority="(Heude, 1886)" baseAuthorityName="Heude" baseAuthorityYear="1886" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF92572FABF2189BEEFFB64" italics="true" pageId="9" pageNumber="602">Coccoglypta pinchoniana</emphasis>
(
<bibRefCitation id="CC054B77FFF92572FF0521F2BF78FB64" author="HEUDE R. P. M." box="[187,295,1229,1247]" pageId="9" pageNumber="602" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
)
</taxonomicName>
was described from Chengdu, and it was known from the Emeishan, and a fossil record from Chongqing. Now we report this species and
<taxonomicName id="6F944D05FFF92572FF3A21DBBF7FFB4D" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[132,288,1252,1270]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="species" species="liui" status="sp. nov.">
<emphasis id="9AE0EA94FFF92572FF3A21DBBEECFB4D" box="[132,179,1252,1270]" italics="true" pageId="9" pageNumber="602">C. liui</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="81D357EFFFF92572FE9921DBBF08FB4D" box="[295,343,1252,1270]" pageId="9" pageNumber="602" rank="species">n. sp.</taxonomicNameLabel>
from Qingcheng Shan and Qingcheng Houshan (black area).
</paragraph>
</caption>
<subSubSection id="E08E650DFFF92572FF3A20ABBB3FF994" pageId="9" pageNumber="602" type="description">
<paragraph id="A82B3686FFF92572FF3A20ABBF8CFA16" blockId="9.[132,776,1428,1934]" box="[132,467,1428,1454]" pageId="9" pageNumber="602">COMPLEMENT OF DESCRIPTION</paragraph>
<paragraph id="A82B3686FFF92572FF3A208ABD57F836" blockId="9.[132,776,1428,1934]" pageId="9" pageNumber="602">Shell dextral, rather large, light brown, greenish to greyish, dorsal and ventral surface of similar colour; shell depressed globular or conical, body whorl slightly or prominently keeled; entire shell consists of 5.25-5.75 whorls, separated by rather shallow suture; protoconch consisting of 1.5-1.75 whorls, matt, rather smooth, dorsal side of teleoconch dominated by irregular, very rough wrinkles and strong tubercles (tubercles appearing after the first 1.5 whorls of teleoconch); some fine spiral grooves also visible, especially close to the suture; ventral surface similar to dorsal, although with slightly weaker sculpture; aperture strongly oblique to shell axis, white; peristome expanded (mostly in basal and umbilical direction) but not reflected; parietal callus weak, only indicated by fine whitish calcareous layer, which is transparent in fresh shells; umbilicus open, moderately wide to rather narrow, showing all whorls.</paragraph>
<paragraph id="A82B3686FFF92572FF3A228BBF4AF876" blockId="9.[132,651,1972,2030]" box="[132,277,1972,1997]" pageId="9" pageNumber="602">
<emphasis id="9AE0EA94FFF92572FF3A228BBF4AF876" box="[132,277,1972,1997]" italics="true" pageId="9" pageNumber="602">Measurements</emphasis>
</paragraph>
<paragraph id="A82B3686FFF92572FF3A22EBBDCEFA16" blockId="9.[132,651,1972,2030]" box="[132,913,1428,2030]" lastBlockId="9.[813,1455,1428,1583]" pageId="9" pageNumber="602">
D =
<quantity id="6F6C9B63FFF92572FF0822ECBF08F855" box="[182,343,2003,2030]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" metricValueMax="3.19" metricValueMin="2.57" pageId="9" pageNumber="602" unit="mm" value="28.799999999999997" valueMax="31.9" valueMin="25.7">25.7-31.9 mm</quantity>
, H = 15.0-
<quantity id="6F6C9B63FFF92572FE7122ECBC6CF855" box="[463,563,2003,2030]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="9" pageNumber="602" unit="mm" value="17.5">17.5 mm</quantity>
(n = 6). REMARKS
</paragraph>
<paragraph id="A82B3686FFF92572FC93208ABB3FF994" blockId="9.[813,1455,1428,1583]" pageId="9" pageNumber="602">
This species was originally described from Chengdu city,
<collectingRegion id="6A50F864FFF92572FC9320EABDB7FA54" box="[813,1000,1493,1519]" country="China" name="Sichuan" pageId="9" pageNumber="602">Sichuan Province</collectingRegion>
(
<bibRefCitation id="CC054B77FFF92572FC4620EABAD2FA4B" author="HEUDE R. P. M." box="[1016,1165,1493,1520]" pageId="9" pageNumber="602" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
). It is known as a subfossil record from
<collectingRegion id="6A50F864FFF92572FC0520CABA66F9B4" box="[955,1081,1525,1551]" country="China" name="Chongqing" pageId="9" pageNumber="602">Chongqing</collectingRegion>
town (
<bibRefCitation id="CC054B77FFF92572FB3020CABAA7F9B4" author="YEN T. - C." box="[1166,1272,1525,1551]" pageId="9" pageNumber="602" pagination="331 - 342" refId="ref8619" refString="YEN T. - C. 1936. - Some Quaternary gastropods from eastern Szechwan. Bulletin of the Geological Society of China 15 (3): 331 - 342. https: // doi. org / 10.1111 / j. 1755 - 6724.1936. mp 15003005. x" type="journal article" year="1936">Yen 1936</bibRefCitation>
). This species is considered a vulnerable species (
<bibRefCitation id="CC054B77FFF92572FB37232ABB0FF994" author="WANG S. &amp; XIE Y." box="[1161,1360,1556,1583]" pageId="9" pageNumber="602" pagination="368 - 369" refId="ref8542" refString="WANG S. &amp; XIE Y. 2005. - China Species Red List (Vol. III, Invertebrates). Higher Education Press, Beijing: 368 - 369 p." type="book chapter" year="2005">Wang &amp; Xie 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

249
data/20/3D/87/203D8790FFF92570FCE4234CBF3BFBEB.xml Normal file
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<subSubSection id="E08E650DFFF92572FCE4234CBBDDF935" box="[858,1410,1651,1679]" pageId="9" pageNumber="602" type="nomenclature">
<paragraph id="A82B3686FFF92572FCE4234CBBDDF935" blockId="9.[858,1410,1651,1711]" box="[858,1410,1651,1679]" pageId="9" pageNumber="602">
<heading id="F36381EAFFF92572FCE4234CBBDDF935" box="[858,1410,1651,1679]" level="2" pageId="9" pageNumber="602">
<taxonomicName id="6F944D05FFF92572FCE4234CBB44F935" authority="(Deshayes, 1870)" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Deshayes" baseAuthorityYear="1870" box="[858,1307,1651,1678]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="species" species="arbusticola" status="comb. nov.">
<emphasis id="9AE0EA94FFF92572FCE4234CBA07F935" bold="true" box="[858,1112,1651,1678]" italics="true" pageId="9" pageNumber="602">Coccoglypta arbusticola</emphasis>
(
<bibRefCitation id="CC054B77FFF92572FBD8234BBB4CF935" author="DESHAYES G. P." box="[1126,1299,1652,1678]" pageId="9" pageNumber="602" pagination="19 - 27" refId="ref7195" refString="DESHAYES G. P. 1870. - Diagnoses d'especes nouvelles de mollusques terrestres et fluviatiles de la principaute de Moupin, Thibet oriental envoyees au Museum d'Histoire naturelle de Paris par M. l'Abbe Armand David missionnaire. Bulletin des Nouvelles Archives du Museum d'Histoire naturelle de Paris 6: 19 - 27." type="journal article" year="1870">Deshayes, 1870</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="81D357EFFFF92572FA9F234ABBDDF935" box="[1313,1410,1652,1679]" pageId="9" pageNumber="602" rank="species">n. comb.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF92572FB9023ABBAF1F914" box="[1070,1198,1684,1711]" pageId="9" pageNumber="602" type="description">
<paragraph id="A82B3686FFF92572FB9023ABBAF1F914" blockId="9.[858,1410,1651,1711]" box="[1070,1198,1684,1711]" pageId="9" pageNumber="602">
(
<figureCitation id="30AF2A03FFF92572FB8923ABBAFAF915" box="[1079,1189,1684,1710]" captionStart="FIG" captionStartId="10.[132,143,1964,1982]" captionTargetBox="[165,1412,215,1902]" captionTargetId="figure@10.[165,1412,215,1907]" captionTargetPageId="10" captionText="FIG. 6. — Potential members of the genus Coccoglypta Pilsbry,1895:A-D, syntype of Coccoglypta arbusticola (Deshayes,1870) n. comb.(MNHN-IM-2000-34192); E-H, “Bradybaena” chrysomphala (Möllendorff, 1899),SMF 9159 (lectotype); I-L, Coccoglypta cf. leprosula n. comb. (Heude, 1885) from the Guanwushan Forest Farm. Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.3726122" httpUri="https://zenodo.org/record/3726122/files/figure.png" pageId="9" pageNumber="602">Fig. 6A-D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF92572FC9323E9BBEFF8EB" pageId="9" pageNumber="602" type="reference_group">
<paragraph id="A82B3686FFF92572FC9323E9BAC6F955" blockId="9.[813,1177,1750,1774]" box="[813,1177,1750,1774]" pageId="9" pageNumber="602">
<treatmentCitationGroup id="888411A8FFF92572FC9323E9BAC6F955" box="[813,1177,1750,1774]" pageId="9" pageNumber="602">
<treatmentCitation id="29351097FFF92572FC9323E9BACBF955" author="DESHAYES G. P." box="[813,1172,1750,1774]" page="20" pageId="9" pageNumber="602" year="1870">
<taxonomicName id="6F944D05FFF92572FC9323E9BACBF955" authority="Deshayes, 1870: 20" authorityName="Deshayes" authorityPageNumber="20" authorityYear="1870" box="[813,1172,1750,1774]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="species" species="arbusticola">
<emphasis id="9AE0EA94FFF92572FC9323E9BD94F955" box="[813,971,1750,1774]" italics="true" pageId="9" pageNumber="602">Helix arbusticola</emphasis>
<bibRefCitation id="CC054B77FFF92572FC6C23E9BACBF955" author="DESHAYES G. P." box="[978,1172,1750,1774]" pageId="9" pageNumber="602" pagination="19 - 27" refId="ref7195" refString="DESHAYES G. P. 1870. - Diagnoses d'especes nouvelles de mollusques terrestres et fluviatiles de la principaute de Moupin, Thibet oriental envoyees au Museum d'Histoire naturelle de Paris par M. l'Abbe Armand David missionnaire. Bulletin des Nouvelles Archives du Museum d'Histoire naturelle de Paris 6: 19 - 27." type="journal article" year="1870">Deshayes, 1870: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A82B3686FFF92572FC932237BBF0F8A4" blockId="9.[813,1455,1799,1823]" box="[813,1455,1799,1823]" pageId="9" pageNumber="602">
<treatmentCitationGroup id="888411A8FFF92572FC932237BBF0F8A4" box="[813,1455,1799,1823]" pageId="9" pageNumber="602">
non
<taxonomicName id="6F944D05FFF92572FCE92238BACEF8A4" authority="arbusticola" box="[855,1169,1799,1823]" class="Gastropoda" family="Bradybaenidae" genus="Bradybaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="subSpecies" species="arbusticola" subSpecies="arbusticola">
<emphasis id="9AE0EA94FFF92572FCE92238BACEF8A4" box="[855,1169,1799,1823]" italics="true" pageId="9" pageNumber="602">Bradybaena arbusticola arbusticola</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFF92572FB182237BB6EF8A4" author="YEN T. - C." box="[1190,1329,1799,1823]" page="135" pageId="9" pageNumber="602" year="1939">
<bibRefCitation id="CC054B77FFF92572FB182237BB6EF8A4" author="YEN T. - C." box="[1190,1329,1799,1823]" pageId="9" pageNumber="602" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen 1939: 135</bibRefCitation>
</treatmentCitation>
, pl. 14, fig. 2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A82B3686FFF92572FC932206BBEFF8EB" blockId="9.[813,1456,1848,1872]" box="[813,1456,1848,1872]" pageId="9" pageNumber="602">
<treatmentCitationGroup id="888411A8FFF92572FC932206BBEFF8EB" box="[813,1456,1848,1872]" pageId="9" pageNumber="602">
non
<taxonomicName id="6F944D05FFF92572FCE82207BA7BF8EB" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Deshayes" baseAuthorityYear="1870" box="[854,1060,1848,1872]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="602" phylum="Mollusca" rank="species" species="arbusticola">
<emphasis id="9AE0EA94FFF92572FCE82207BA7BF8EB" box="[854,1060,1848,1872]" italics="true" pageId="9" pageNumber="602">Coccoglypta arbusticola</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFF92572FB872207BB05F8EB" author="CHEN D. - N. &amp; ZHANG G. - Q." box="[1081,1370,1848,1872]" page="155" pageId="9" pageNumber="602" year="2004">
<bibRefCitation id="CC054B77FFF92572FB872207BB05F8EB" author="CHEN D. - N. &amp; ZHANG G. - Q." box="[1081,1370,1848,1872]" pageId="9" pageNumber="602" refId="ref7148" refString="CHEN D. - N. &amp; ZHANG G. - Q. 2004. - Fauna Sinica (Invertebrata Vol. 37 Mollusca: Gastropoda: Stylommatophora: Bradybaenidae). Science Press, Beijing, China, 482 p. (in Chinese)." type="book" year="2004">Chen &amp; Zhang 2004: 155-156</bibRefCitation>
</treatmentCitation>
, fig. 123.
</treatmentCitationGroup>
</paragraph>
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<paragraph id="A82B3686FFF92572FC932254BBF0F825" blockId="9.[813,1455,1899,1951]" pageId="9" pageNumber="602">
<materialsCitation id="18FC3CDBFFF92572FC932254BBF6F825" ID-GBIF-Occurrence="2580042139" collectionCode="TYPE, MATERIAL, EXAMINED" country="China" location="l'Abbe David" municipality="Moupin" pageId="9" pageNumber="602" specimenCount="1" stateProvince="Sichuan" typeStatus="syntype">
<collectionCode id="CE85AE43FFF92572FC932254BD00F83B" box="[813,863,1899,1923]" pageId="9" pageNumber="602">TYPE</collectionCode>
<collectionCode id="CE85AE43FFF92572FCD9224FBA69F83A" box="[871,1078,1904,1921]" pageId="9" pageNumber="602">MATERIAL EXAMINED</collectionCode>
. —
<emphasis id="9AE0EA94FFF92572FBD92254BAF8F838" bold="true" box="[1127,1191,1899,1923]" pageId="9" pageNumber="602">
<collectingCountry id="D0837616FFF92572FBD92254BAF8F838" box="[1127,1191,1899,1923]" name="China" pageId="9" pageNumber="602">China</collectingCountry>
</emphasis>
.
<collectingMunicipality id="484FACFCFFF92572FB0B2253BB57F83F" box="[1205,1288,1900,1924]" pageId="9" pageNumber="602">Moupin</collectingMunicipality>
[Muping Zhen,
<collectingRegion id="6A50F864FFF92572FC9322B9BDDFF825" box="[813,896,1926,1950]" country="China" name="Sichuan" pageId="9" pageNumber="602">Sichuan</collectingRegion>
],
<location id="AD4B605DFFF92572FC3022B9BA4FF825" LSID="urn:lsid:plazi:treatment:203D8790FFF92570FCE4234CBF3BFBEB:AD4B605DFFF92572FC3022B9BA4FF825" box="[910,1040,1926,1950]" country="China" municipality="Moupin" name="l'Abbe David" pageId="9" pageNumber="602" stateProvince="Sichuan">l’Abbé David</location>
, 1869,
<typeStatus id="772F8824FFF92572FBE822B8BAFCF824" box="[1110,1187,1927,1951]" pageId="9" pageNumber="602" type="syntype">syntype</typeStatus>
, MNHN-IM-2000-34192
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.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFF92572FC932288BDFEF851" pageId="9" pageNumber="602" type="distribution">
<paragraph id="A82B3686FFF92572FC932288BDFEF851" blockId="9.[813,1455,1975,2026]" pageId="9" pageNumber="602">TYPE LOCALITY. — “Principauté de Moupin, Thibet oriental” (from title).</paragraph>
</subSubSection>
<caption id="FCEB660EFFFA2571FF3A2293BF0CF857" ID-DOI="http://doi.org/10.5281/zenodo.3726122" ID-Zenodo-Dep="3726122" httpUri="https://zenodo.org/record/3726122/files/figure.png" pageId="10" pageNumber="603" startId="10.[132,143,1964,1982]" targetBox="[165,1412,215,1902]" targetPageId="10">
<paragraph id="A82B3686FFFA2571FF3A2293BF0CF857" pageId="10" pageNumber="603">
FIG. 6. — Potential members of the genus
<emphasis id="9AE0EA94FFFA2571FE662293BC1EF805" box="[472,577,1964,1982]" italics="true" pageId="10" pageNumber="603">Coccoglypta</emphasis>
Pilsbry,1895:
<emphasis id="9AE0EA94FFFA2571FD082292BC87F804" bold="true" box="[694,728,1964,1983]" pageId="10" pageNumber="603">A -D</emphasis>
, syntype of
<emphasis id="9AE0EA94FFFA2571FC842293BA5FF805" box="[826,1024,1964,1982]" italics="true" pageId="10" pageNumber="603">Coccoglypta arbusticola</emphasis>
(Deshayes,1870) n. comb.(MNHN-IM-2000-34192);
<emphasis id="9AE0EA94FFFA2571FF3A22FCBEF9F86E" bold="true" box="[132,166,1987,2005]" pageId="10" pageNumber="603">E -H</emphasis>
, “
<emphasis id="9AE0EA94FFFA2571FF0922FCBF44F86E" box="[183,283,1987,2005]" italics="true" pageId="10" pageNumber="603">Bradybaena</emphasis>
<emphasis id="9AE0EA94FFFA2571FEA522FCBF7CF86E" bold="true" box="[283,291,1987,2005]" pageId="10" pageNumber="603">”</emphasis>
<emphasis id="9AE0EA94FFFA2571FE9922FCBFC4F86E" box="[295,411,1987,2005]" italics="true" pageId="10" pageNumber="603">chrysomphala</emphasis>
(Möllendorff, 1899),SMF 9159 (lectotype);
<emphasis id="9AE0EA94FFFA2571FCBE22FCBD47F86E" bold="true" box="[768,792,1987,2005]" pageId="10" pageNumber="603">I -L</emphasis>
,
<emphasis id="9AE0EA94FFFA2571FC9F22FCBDD5F86E" box="[801,906,1987,2005]" italics="true" pageId="10" pageNumber="603">Coccoglypta</emphasis>
cf.
<emphasis id="9AE0EA94FFFA2571FC1722FCBDABF86E" box="[937,1012,1987,2005]" italics="true" pageId="10" pageNumber="603">leprosula</emphasis>
n. comb. (Heude, 1885) from the Guanwushan Forest Farm. Scale bar: 30 mm.
</paragraph>
</caption>
<subSubSection id="E08E650DFFFB2570FF3A25E6BF3BFBEB" pageId="11" pageNumber="604" type="discussion">
<paragraph id="A82B3686FFFB2570FF3A25E6BEB7FF4A" blockId="11.[132,776,217,1104]" box="[132,232,217,243]" pageId="11" pageNumber="604">REMARKS</paragraph>
<paragraph id="A82B3686FFFB2570FF3A25C6BF36FE48" blockId="11.[132,776,217,1104]" pageId="11" pageNumber="604">
This species has previously been assigned to the genus
<taxonomicName id="6F944D05FFFB2570FF3A2426BF5EFE88" authorityName="H.Beck" authorityYear="1837" box="[132,257,281,307]" class="Gastropoda" family="Bradybaenidae" genus="Bradybaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFB2570FF3A2426BF5EFE88" box="[132,257,281,307]" italics="true" pageId="11" pageNumber="604">Bradybaena</emphasis>
</taxonomicName>
, however it surely does not belong to that genus in its present concept. The type species of
<taxonomicName id="6F944D05FFFB2570FDC22406BCA0FEE8" authorityName="H.Beck" authorityYear="1837" box="[636,767,313,339]" class="Gastropoda" family="Bradybaenidae" genus="Bradybaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFB2570FDC22406BCA0FEE8" box="[636,767,313,339]" italics="true" pageId="11" pageNumber="604">Bradybaena</emphasis>
</taxonomicName>
,
<taxonomicName id="6F944D05FFFB2570FF3A2466BFC6FEC8" authority="(Rang, 1831)" baseAuthorityName="Rang" baseAuthorityYear="1831" box="[132,409,345,371]" class="Gastropoda" family="Bradybaenidae" genus="Bradybaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="similaris">
<emphasis id="9AE0EA94FFFB2570FF3A2466BEA2FEC8" box="[132,253,345,371]" italics="true" pageId="11" pageNumber="604">B. similaris</emphasis>
(Rang, 1831)
</taxonomicName>
, has a small (
<emphasis id="9AE0EA94FFFB2570FD992466BC66FEC9" box="[551,569,345,370]" italics="true" pageId="11" pageNumber="604">c.</emphasis>
<quantity id="6F6C9B63FFFB2570FD802466BC27FEC8" box="[574,632,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="604" unit="cm" value="1.0">1 cm</quantity>
), fragile shell without any distinctive sculpture. However,
<taxonomicName id="6F944D05FFFB2570FDCF2446BD58FE29" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Deshayes" baseAuthorityYear="1870" box="[625,775,376,403]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="arbusticola" status="comb. nov.">
<emphasis id="9AE0EA94FFFB2570FDCF2446BD58FE29" box="[625,775,376,403]" italics="true" pageId="11" pageNumber="604">C. arbusticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81D357EFFFFB2570FF3A24A6BEB4FE09" box="[132,235,408,435]" pageId="11" pageNumber="604" rank="species">n. comb.</taxonomicNameLabel>
is larger (shell diameter of
<typeStatus id="772F8824FFFB2570FD9824A6BCDFFE08" box="[550,640,409,435]" pageId="11" pageNumber="604" type="syntype">syntype</typeStatus>
:
<quantity id="6F6C9B63FFFB2570FD3224A7BCA4FE08" box="[652,763,408,435]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="11" pageNumber="604" unit="mm" value="24.5">24.5 mm</quantity>
), and has a thick, finely mamillated shell, reminiscent of those of
<taxonomicName id="6F944D05FFFB2570FF5A24E7BF3EFE49" authorityName="Pilsbry" authorityYear="1895" box="[228,353,472,498]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFB2570FF5A24E7BF3EFE49" box="[228,353,472,498]" italics="true" pageId="11" pageNumber="604">Coccoglypta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A82B3686FFFB2570FF2524C6BC20FC8A" blockId="11.[132,776,217,1104]" pageId="11" pageNumber="604">
We have examined the
<typeStatus id="772F8824FFFB2570FE1224C7BC4BFDA9" box="[428,532,504,530]" pageId="11" pageNumber="604" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="6F944D05FFFB2570FD8024C7BFAEFD89" authority="Mollendorff, 1899" authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Eulota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="subSpecies" species="arbusticola" subSpecies="chrysomphala">
<emphasis id="9AE0EA94FFFB2570FD8024C7BF4AFD89" italics="true" pageId="11" pageNumber="604">Eulota arbusticola chrysomphala</emphasis>
<bibRefCitation id="CC054B77FFFB2570FEA12727BFAEFD89" author="MOLLENDORFF O. &amp; VON" box="[287,497,536,562]" pageId="11" pageNumber="604" pagination="46 - 144" refId="ref7681" refString="MOLLENDORFF O. VON. 1899. - Binnen-Mollusken aus Westchina und Central-Asien Ezhegodnik Zoologicheskago muzeia (Annals of the Zoological Museum of the Saint Petersburg Academy of Sciences) 4 (1): 46 - 144." type="journal article" year="1899">Möllendorff, 1899</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="CC054B77FFFB2570FD902727BEFFFDE9" author="MOLLENDORFF O. &amp; VON" pageId="11" pageNumber="604" pagination="46 - 144" refId="ref7681" refString="MOLLENDORFF O. VON. 1899. - Binnen-Mollusken aus Westchina und Central-Asien Ezhegodnik Zoologicheskago muzeia (Annals of the Zoological Museum of the Saint Petersburg Academy of Sciences) 4 (1): 46 - 144." type="journal article" year="1899">Möllendorff, 1899: 70</bibRefCitation>
and
<bibRefCitation id="CC054B77FFFB2570FF6C2706BF2CFDE8" author="YEN T. - C." box="[210,371,568,595]" pageId="11" pageNumber="604" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen, 1939: 135</bibRefCitation>
) in the Senckenberg Museum (
<collectingCountry id="D0837616FFFB2570FD022707BD5CFDE9" box="[700,771,568,594]" name="China" pageId="11" pageNumber="604">China</collectingCountry>
: W. Sy-tshuan, Fu-bien-ho, SMF 9159, figs 6E-H). It had a light brown, very finely wrinkled and extremely finely spirally grooved shell, without any signs of mamillae. The aperture is also comparatively much larger in
<emphasis id="9AE0EA94FFFB2570FDA12788BCF2FD6A" box="[543,685,695,721]" italics="true" pageId="11" pageNumber="604">chrysomphala</emphasis>
than in
<taxonomicName id="6F944D05FFFB2570FF3A27E8BEADFD4A" baseAuthorityName="Deshayes" baseAuthorityYear="1870" box="[132,242,727,753]" class="Gastropoda" family="Bradybaenidae" genus="Eulota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="arbusticola">
<emphasis id="9AE0EA94FFFB2570FF3A27E8BEADFD4A" box="[132,242,727,753]" italics="true" pageId="11" pageNumber="604">arbusticola</emphasis>
</taxonomicName>
. Thus, “
<taxonomicName id="6F944D05FFFB2570FEF527E8BF9BFD4A" authorityName="H.Beck" authorityYear="1837" box="[331,452,727,753]" class="Gastropoda" family="Bradybaenidae" genus="Bradybaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFB2570FEF527E8BF9BFD4A" box="[331,452,727,753]" italics="true" pageId="11" pageNumber="604">Bradybaena</emphasis>
</taxonomicName>
”
<emphasis id="9AE0EA94FFFB2570FE6A27E8BC01FD4A" box="[468,606,727,753]" italics="true" pageId="11" pageNumber="604">chrysomphala</emphasis>
and
<taxonomicName id="6F944D05FFFB2570FD2E27E8BEABFCAA" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Deshayes" baseAuthorityYear="1870" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="arbusticola" status="comb. nov.">
<emphasis id="9AE0EA94FFFB2570FD2E27E8BEABFCAA" italics="true" pageId="11" pageNumber="604">Coccoglypta arbusticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81D357EFFFFB2570FF4427C7BF06FCAA" box="[250,345,759,786]" pageId="11" pageNumber="604" rank="species">n. comb.</taxonomicNameLabel>
cannot be subspecies of the same species and must be considered as two distinct species.
</paragraph>
<paragraph id="A82B3686FFFB2570FF252608BF3BFBEB" blockId="11.[132,776,217,1104]" pageId="11" pageNumber="604">
The single shell in the Senckenberg Museum identified as
<taxonomicName id="6F944D05FFFB2570FF1F2667BF6CFCCA" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Deshayes" baseAuthorityYear="1870" box="[161,307,855,881]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="arbusticola">
<emphasis id="9AE0EA94FFFB2570FF1F2667BF6CFCCA" box="[161,307,855,881]" italics="true" pageId="11" pageNumber="604">B. arbusticola</emphasis>
</taxonomicName>
and figured by
<bibRefCitation id="CC054B77FFFB2570FE5D2667BC01FCC9" author="YEN T. - C." box="[483,606,855,882]" pageId="11" pageNumber="604" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen (1939)</bibRefCitation>
has a narrower umbilicus and more rapidly growing whorls than the
<typeStatus id="772F8824FFFB2570FD192648BC88FC2A" box="[679,727,887,913]" pageId="11" pageNumber="604">type</typeStatus>
, and there are also no signs of a mamillated sculpture. Therefore, we here exclude it from the present species. Its true identity remains unknown. Furthermore, the shell figured in
<bibRefCitation id="CC054B77FFFB2570FD1126E9BF44FBAA" author="CHEN D. - N. &amp; ZHANG G. - Q." pageId="11" pageNumber="604" refId="ref7148" refString="CHEN D. - N. &amp; ZHANG G. - Q. 2004. - Fauna Sinica (Invertebrata Vol. 37 Mollusca: Gastropoda: Stylommatophora: Bradybaenidae). Science Press, Beijing, China, 482 p. (in Chinese)." type="book" year="2004">Chen &amp; Zhang (2004)</bibRefCitation>
also belong to another species, because it has a narrower umbilicus, a dark spiral band, and a more strongly expanded peristome.
</paragraph>
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</treatment>
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Genus
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<emphasis id="9AE0EA94FFFB2570FBAF2628BAD9FC8A" box="[1041,1158,791,817]" italics="true" pageId="11" pageNumber="604">Hemiplecta</emphasis>
Albers, 1850
</taxonomicName>
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<typeStatus id="772F8824FFFB2570FC93266FBD02FCDE" box="[813,861,848,872]" pageId="11" pageNumber="604">TYPE</typeStatus>
SPECIES. —
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Martens
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Albers
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1860).
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<emphasis id="9AE0EA94FFFB2570FF6E21AABFF3FB0B" bold="true" box="[208,428,1173,1200]" italics="true" pageId="11" pageNumber="604">Coccoglypta billiana</emphasis>
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As for the previous species,
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did not mention
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specimens in American museums, and we could not examine the
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. Since the sculpture is also mamillated, the species also potentially belongs to
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<paragraph id="A82B3686FFFB2570FF75236BBC9EF9D5" blockId="11.[203,705,1620,1679]" box="[203,705,1620,1647]" pageId="11" pageNumber="604">
<heading id="F36381EAFFFB2570FF75236BBC9EF9D5" box="[203,705,1620,1647]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="604" reason="2">
<taxonomicName id="6F944D05FFFB2570FF75236BBC05F9D4" authority="(Heude, 1885)" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[203,602,1620,1647]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="leprosula" status="comb. nov.">
<emphasis id="9AE0EA94FFFB2570FF75236BBFEDF9D4" bold="true" box="[203,434,1620,1647]" italics="true" pageId="11" pageNumber="604">Coccoglypta leprosula</emphasis>
(Heude, 1885)
</taxonomicName>
<taxonomicNameLabel id="81D357EFFFFB2570FDDF236ABC9EF9D5" box="[609,705,1620,1647]" pageId="11" pageNumber="604" rank="species">n. comb.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FE33234BBFA1F934" box="[397,510,1652,1679]" pageId="11" pageNumber="604" type="description">
<paragraph id="A82B3686FFFB2570FE33234BBFA1F934" blockId="11.[203,705,1620,1679]" box="[397,510,1652,1679]" pageId="11" pageNumber="604">
(
<figureCitation id="30AF2A03FFFB2570FE28234BBFA9F935" box="[406,502,1652,1678]" captionStart="FIG" captionStartId="10.[132,143,1964,1982]" captionTargetBox="[165,1412,215,1902]" captionTargetId="figure@10.[165,1412,215,1907]" captionTargetPageId="10" captionText="FIG. 6. — Potential members of the genus Coccoglypta Pilsbry,1895:A-D, syntype of Coccoglypta arbusticola (Deshayes,1870) n. comb.(MNHN-IM-2000-34192); E-H, “Bradybaena” chrysomphala (Möllendorff, 1899),SMF 9159 (lectotype); I-L, Coccoglypta cf. leprosula n. comb. (Heude, 1885) from the Guanwushan Forest Farm. Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.3726122" httpUri="https://zenodo.org/record/3726122/files/figure.png" pageId="11" pageNumber="604">Fig. 6I-L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FF3A2393BCB0F899" pageId="11" pageNumber="604" type="reference_group">
<paragraph id="A82B3686FFFB2570FF3A2393BC06F97F" blockId="11.[132,601,1708,1779]" box="[132,601,1708,1732]" pageId="11" pageNumber="604">
<treatmentCitationGroup id="888411A8FFFB2570FF3A2393BC06F97F" box="[132,601,1708,1732]" pageId="11" pageNumber="604">
<treatmentCitation id="29351097FFFB2570FF3A2393BF9CF97F" author="HEUDE R. P. M." box="[132,451,1708,1732]" page="106" pageId="11" pageNumber="604" year="1885">
<taxonomicName id="6F944D05FFFB2570FF3A2393BF9CF97F" authority="Heude, 1885 a: 106" authorityName="Heude" authorityPageNumber="106" authorityYear="1885" box="[132,451,1708,1732]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="leprosa">
<emphasis id="9AE0EA94FFFB2570FF3A2393BEA2F97F" box="[132,253,1708,1732]" italics="true" pageId="11" pageNumber="604">Helix leprosa</emphasis>
<bibRefCitation id="CC054B77FFFB2570FEBD2393BF9CF97F" author="HEUDE R. P. M." box="[259,451,1708,1732]" pageId="11" pageNumber="604" pagination="89 - 132" refId="ref7394" refString="HEUDE R. P. M. 1885 a. - Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de Jesus. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu. III, Mission Catholique, Chang-Hai, 89 - 132 p." type="book chapter" year="1885">Heude, 1885a: 106</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 27, fig. 15.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A82B3686FFFB2570FF3A23E4BF8EF948" blockId="11.[132,601,1708,1779]" box="[132,465,1755,1779]" pageId="11" pageNumber="604">
<treatmentCitationGroup id="888411A8FFFB2570FF3A23E4BF8EF948" box="[132,465,1755,1779]" pageId="11" pageNumber="604">
<treatmentCitation id="29351097FFFB2570FF3A23E4BF92F948" author="HEUDE R. P. M." box="[132,461,1755,1779]" page="43" pageId="11" pageNumber="604" year="1885">
<taxonomicName id="6F944D05FFFB2570FF3A23E4BF92F948" authority="Heude, 1885 b: 43" authorityName="Heude" authorityPageNumber="43" authorityYear="1885" box="[132,461,1755,1779]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="leprosula">
<emphasis id="9AE0EA94FFFB2570FF3A23E4BF4FF948" box="[132,272,1755,1779]" italics="true" pageId="11" pageNumber="604">Helix leprosula</emphasis>
<bibRefCitation id="CC054B77FFFB2570FEA823E4BF92F948" author="HEUDE R. P. M." box="[278,461,1755,1779]" pageId="11" pageNumber="604" pagination="42 - 43" refId="ref7445" refString="HEUDE R. P. M. 1885 b. - Rectifications de nomenclature. Journal de Conchyliologie 33: 42 - 43." type="journal article" year="1885">Heude, 1885b: 43</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A82B3686FFFB2570FF3A2235BCB0F899" blockId="11.[132,751,1802,1826]" box="[132,751,1802,1826]" pageId="11" pageNumber="604">
<treatmentCitationGroup id="888411A8FFFB2570FF3A2235BCB0F899" box="[132,751,1802,1826]" pageId="11" pageNumber="604">
<taxonomicName id="6F944D05FFFB2570FF3A2235BF29F899" authorityName="Heude" authorityYear="1885" box="[132,374,1802,1826]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="leprosula" subGenus="Satsuma">
<emphasis id="9AE0EA94FFFB2570FF3A2235BEE8F899" box="[132,183,1802,1826]" italics="true" pageId="11" pageNumber="604">Helix</emphasis>
(
<emphasis id="9AE0EA94FFFB2570FF782235BF4AF89A" box="[198,277,1802,1825]" italics="true" pageId="11" pageNumber="604">Satsuma</emphasis>
)
<emphasis id="9AE0EA94FFFB2570FE9A2235BF29F899" box="[292,374,1802,1826]" italics="true" pageId="11" pageNumber="604">leprosula</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFFB2570FE302235BC69F899" author="TRYON G. W." box="[398,566,1802,1826]" page="220" pageId="11" pageNumber="604" year="1887">
<bibRefCitation id="CC054B77FFFB2570FE302235BC69F899" author="TRYON G. W." box="[398,566,1802,1826]" pageId="11" pageNumber="604" refId="ref8264" refString="TRYON G. W. 1887. - Manual of Conchology; Structural and Systematic. With Illustrations of the Species. Second series: Pulmonata. Volume III. Helicidae, Volume I., Published by the Author, Philadelphia, 313 p." type="book" year="1887">Tryon 1887: 220</bibRefCitation>
</treatmentCitation>
, pl. 51, figs 80-82.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FF3A2206BFF2F800" pageId="11" pageNumber="604" type="materials_examined">
<paragraph id="A82B3686FFFB2570FF3A2206BFF2F800" blockId="11.[132,776,1849,1979]" pageId="11" pageNumber="604">
<materialsCitation id="18FC3CDBFFFB2570FF3A2206BF26F8EA" ID-GBIF-Occurrence="2580042141" box="[132,377,1849,1873]" collectionCode="MATERIAL, EXAMINED" collectorName="A. Hunyadi &amp; M. Szekeres" country="China" county="Mianyang Shi" elevation="1590" latitude="31.94411" location="Wudu Zhen" longLatPrecision="1" longitude="104.733635" municipality="Shi" pageId="11" pageNumber="604" specimenCount="1" specimenCount-adult="1" specimenCount-juvenile="2" stateProvince="Sichuan">
<collectionCode id="CE85AE43FFFB2570FF3A2206BEB9F8F4" box="[132,230,1849,1873]" pageId="11" pageNumber="604">MATERIAL</collectionCode>
<collectionCode id="CE85AE43FFFB2570FF502201BF0DF8F4" box="[238,338,1854,1871]" pageId="11" pageNumber="604">EXAMINED</collectionCode>
. —
</materialsCitation>
<materialsCitation id="18FC3CDBFFFB2570FE3F2206BF1FF81A" ID-GBIF-Occurrence="2580042129" collectionCode="MATERIAL, EXAMINED" collectorName="A. Hunyadi &amp; M. Szekeres" country="China" county="Mianyang Shi" elevation="1590" latitude="31.94411" location="Wudu Zhen" longLatPrecision="1" longitude="104.733635" municipality="Shi" pageId="11" pageNumber="604" specimenCount="1" specimenCount-adult="1" specimenCount-juvenile="2" stateProvince="Sichuan">
<emphasis id="9AE0EA94FFFB2570FE3F2206BFE0F8EA" bold="true" box="[385,447,1849,1873]" pageId="11" pageNumber="604">
<collectingCountry id="D0837616FFFB2570FE3F2206BFE0F8EA" box="[385,447,1849,1873]" name="China" pageId="11" pageNumber="604">China</collectingCountry>
</emphasis>
.
<collectingRegion id="6A50F864FFFB2570FE732205BC41F8E9" box="[461,542,1850,1874]" country="China" name="Sichuan" pageId="11" pageNumber="604">Sichuan</collectingRegion>
,
<collectingCounty id="414A4E0AFFFB2570FD942205BCE9F8E9" box="[554,694,1850,1874]" pageId="11" pageNumber="604">Mianyang Shi</collectingCounty>
, Jiangy- ou
<collectingMunicipality id="484FACFCFFFB2570FF18226BBE96F8D7" box="[166,201,1876,1900]" pageId="11" pageNumber="604">Shi</collectingMunicipality>
,
<location id="AD4B605DFFFB2570FF6B226BBF0CF8D7" LSID="urn:lsid:plazi:treatment:203D8790FFFB2570FF75236BBA43FD29:AD4B605DFFFB2570FF6B226BBF0CF8D7" box="[213,339,1876,1900]" country="China" county="Mianyang Shi" latitude="31.94411" longLatPrecision="1" longitude="104.733635" municipality="Shi" name="Wudu Zhen" pageId="11" pageNumber="604" stateProvince="Sichuan">Wudu Zhen</location>
,
<location id="AD4B605DFFFB2570FEE1226BBC38F8D7" LSID="urn:lsid:plazi:treatment:203D8790FFFB2570FF75236BBA43FD29:AD4B605DFFFB2570FEE1226BBC38F8D7" box="[351,615,1876,1900]" country="China" county="Mianyang Shi" latitude="31.94411" longLatPrecision="1" longitude="104.733635" municipality="Shi" name="Guanwushan Forest Farm" pageId="11" pageNumber="604" stateProvince="Sichuan">
<location id="AD4B605DFFFB2570FEE1226BBC76F8D7" LSID="urn:lsid:plazi:treatment:203D8790FFFB2570FF75236BBA43FD29:AD4B605DFFFB2570FEE1226BBC76F8D7" box="[351,553,1876,1900]" country="China" county="Mianyang Shi" latitude="31.94411" longLatPrecision="1" longitude="104.733635" municipality="Shi" name="Guanwushan Forest" pageId="11" pageNumber="604" stateProvince="Sichuan">Guanwushan Forest</location>
Farm
</location>
;
<geoCoordinate id="CDA05041FFFB2570FDCD226BBD5FF8D7" box="[627,768,1876,1900]" degrees="31" direction="north" minutes="56" orientation="latitude" pageId="11" pageNumber="604" precision="1" seconds="38.8" value="31.94411">31°56’38.8”N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFFB2570FF3A2250BF51F83C" box="[132,270,1903,1927]" degrees="104" direction="east" minutes="44" orientation="longitude" pageId="11" pageNumber="604" precision="1" seconds="1.1" value="104.733635">104°44’1.1”E</geoCoordinate>
;
<quantity id="6F6C9B63FFFB2570FEA52251BF32F83D" box="[283,365,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="11" pageNumber="604" unit="m" value="1590.0">
<elevation id="23B9D1B5FFFB2570FEA52251BF32F83D" box="[283,365,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="11" pageNumber="604" unit="m" value="1590.0">1590 m</elevation>
</quantity>
;
<collectorName id="05615350FFFB2570FEC72250BFB0F83D" box="[377,495,1902,1927]" pageId="11" pageNumber="604">A. Hunyadi</collectorName>
&amp;
<collectorName id="05615350FFFB2570FDAA2250BCD3F83D" box="[532,652,1902,1926]" pageId="11" pageNumber="604">M. Szekeres</collectorName>
leg.;
<date id="DC2A1046FFFB2570FD7B2250BC80F83C" box="[709,735,1903,1927]" pageId="11" pageNumber="604" value="2015-06-21" valueMax="2015-06-22" valueMin="2015-06-21">21</date>
-22.
<collectionCode id="CE85AE43FFFB2570FF3A22B6BEFFF81A" box="[132,160,1929,1953]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="604">VI</collectionCode>
.2015; coll. HA;
</materialsCitation>
<materialsCitation id="18FC3CDBFFFB2570FEF822B6BFF7F800" ID-GBIF-Occurrence="2580042127" collectionCode="VI" country="China" pageId="11" pageNumber="604" specimenCount="3" specimenCount-adult="1" specimenCount-juvenile="2">
<specimenCount id="BE92FD0FFFFB2570FEF822B6BFD3F81A" box="[326,396,1929,1953]" pageId="11" pageNumber="604" type="adult">1 adult</specimenCount>
+
<specimenCount id="BE92FD0FFFFB2570FE1922B6BC58F81A" box="[423,519,1929,1953]" pageId="11" pageNumber="604" type="juvenile">2 juvenile</specimenCount>
shells (
<taxonomicName id="6F944D05FFFB2570FDEC22B6BEE4F800" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Heude" baseAuthorityYear="1885" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="leprosula" status="comb. nov.">
<emphasis id="9AE0EA94FFFB2570FDEC22B6BCE0F81A" box="[594,703,1929,1953]" italics="true" pageId="11" pageNumber="604">Coccoglypta</emphasis>
cf.
<emphasis id="9AE0EA94FFFB2570FD5A22B6BEE4F800" italics="true" pageId="11" pageNumber="604">leprosula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81D357EFFFFB2570FF7C229BBF43F800" box="[194,284,1955,1979]" pageId="11" pageNumber="604" rank="species">n. comb.</taxonomicNameLabel>
,
<figureCitation id="30AF2A03FFFB2570FE98229CBF21F800" box="[294,382,1955,1979]" captionStart="FIG" captionStartId="10.[132,143,1964,1982]" captionTargetBox="[165,1412,215,1902]" captionTargetId="figure@10.[165,1412,215,1907]" captionTargetPageId="10" captionText="FIG. 6. — Potential members of the genus Coccoglypta Pilsbry,1895:A-D, syntype of Coccoglypta arbusticola (Deshayes,1870) n. comb.(MNHN-IM-2000-34192); E-H, “Bradybaena” chrysomphala (Möllendorff, 1899),SMF 9159 (lectotype); I-L, Coccoglypta cf. leprosula n. comb. (Heude, 1885) from the Guanwushan Forest Farm. Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.3726122" httpUri="https://zenodo.org/record/3726122/files/figure.png" pageId="11" pageNumber="604">Figures 6</figureCitation>
I-L)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FF3A22ECBD57F850" box="[132,776,2003,2027]" pageId="11" pageNumber="604" type="distribution">
<paragraph id="A82B3686FFFB2570FF3A22ECBD57F850" blockId="11.[132,776,2003,2027]" box="[132,776,2003,2027]" pageId="11" pageNumber="604">
<typeStatus id="772F8824FFFB2570FF3A22ECBEECF853" box="[132,179,2003,2027]" pageId="11" pageNumber="604">TYPE</typeStatus>
LOCALITY. — “Tchen K’eou” (
<location id="AD4B605DFFFB2570FE7222ECBC6DF850" LSID="urn:lsid:plazi:treatment:203D8790FFFB2570FF75236BBA43FD29:AD4B605DFFFB2570FE7222ECBC6DF850" box="[460,562,2003,2027]" name="Chengkou" pageId="11" pageNumber="604" stateProvince="Chongqing">Chengkou</location>
,
<collectingRegion id="6A50F864FFFB2570FD8422ECBCA1F850" box="[570,766,2003,2027]" country="China" name="Chongqing" pageId="11" pageNumber="604">Chongqing Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FC9325E6BA43FD29" pageId="11" pageNumber="604" type="discussion">
<paragraph id="A82B3686FFFB2570FC9325E6BDCEFF4A" blockId="11.[813,1456,217,658]" box="[813,913,217,243]" pageId="11" pageNumber="604">REMARKS</paragraph>
<paragraph id="A82B3686FFFB2570FC9325C6BB62FE08" blockId="11.[813,1456,217,658]" pageId="11" pageNumber="604">
<bibRefCitation id="CC054B77FFFB2570FC9325C6BD8CFEAF" author="JOHNSON R. J." box="[813,979,249,276]" pageId="11" pageNumber="604" refId="ref7571" refString="JOHNSON R. J. 1973. - Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, 111 p. https: // doi. org / 10.5962 / bhl. title. 141074" type="book" year="1973">Johnson (1973)</bibRefCitation>
did not mention
<typeStatus id="772F8824FFFB2570FB3725C5BAE8FEAF" box="[1161,1207,250,276]" pageId="11" pageNumber="604">type</typeStatus>
specimens in American museums, and we could not contact the
<collectingRegion id="6A50F864FFFB2570FB682426BB7CFE88" box="[1238,1315,281,307]" country="China" name="Beijing" pageId="11" pageNumber="604">Beijing</collectingRegion>
Natural History Museum, where some of Heude’s
<typeStatus id="772F8824FFFB2570FB662406BB50FEE8" box="[1240,1295,313,339]" pageId="11" pageNumber="604">types</typeStatus>
are deposited. Therefore, we rely on the original description only. Accordingly, the shell is
<quantity id="6F6C9B63FFFB2570FC602446BA74FE28" box="[990,1067,377,403]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="604" unit="mm" value="26.0">26 mm</quantity>
wide, narrowly umbilicated, and the sculpture is finely tuberculated, as in
<taxonomicName id="6F944D05FFFB2570FB0324A7BB69FE09" authorityName="Pilsbry" authorityYear="1895" box="[1213,1334,408,434]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFB2570FB0324A7BB69FE09" box="[1213,1334,408,434]" italics="true" pageId="11" pageNumber="604">Coccoglypta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A82B3686FFFB2570FC932487BA43FD29" blockId="11.[813,1456,217,658]" pageId="11" pageNumber="604">
The specimen we examined matches the original description, but the identification is certainly doubtful. It was collected
<emphasis id="9AE0EA94FFFB2570FC9324C6BD62FDA9" box="[813,829,505,530]" italics="true" pageId="11" pageNumber="604">c.</emphasis>
<quantity id="6F6C9B63FFFB2570FCFA24C7BDC8FDA9" box="[836,919,504,530]" metricMagnitude="5" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="604" unit="km" value="370.0">370 km</quantity>
west from the
<typeStatus id="772F8824FFFB2570FB8824C6BA3BFDA8" box="[1078,1124,505,531]" pageId="11" pageNumber="604">type</typeStatus>
locality of “
<taxonomicName id="6F944D05FFFB2570FB5824C6BB62FDA8" box="[1254,1341,505,531]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFB2570FB5824C6BB62FDA8" box="[1254,1341,505,531]" italics="true" pageId="11" pageNumber="604">Satsuma</emphasis>
</taxonomicName>
”
<taxonomicName id="6F944D05FFFB2570FAF124C7BBF6FDA9" authorityName="Páll-Gergely &amp; Hunyadi &amp; Chen &amp; Lyu" authorityYear="2019" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[1359,1449,504,530]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="leprosula">
<emphasis id="9AE0EA94FFFB2570FAF124C7BBF6FDA9" box="[1359,1449,504,530]" italics="true" pageId="11" pageNumber="604">leprosula</emphasis>
</taxonomicName>
. However, it must be kept in mind that the
<typeStatus id="772F8824FFFB2570FAAE2726BB61FD88" box="[1296,1342,537,563]" pageId="11" pageNumber="604">type</typeStatus>
locality is just a rough estimate, Heude received specimens from other missionaries who collected shells during their travels across large geographic areas.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C32DEDABE0E3F0446FD0444622C0431C">A review of the genus Coccoglypta Pilsbry, 1895 (Gastropoda: Pulmonata: Camaenidae)</mods:title>
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<paragraph id="A82B3686FFFB2570FC812689BBC2FC6A" blockId="11.[813,1455,848,1009]" box="[831,1437,950,977]" pageId="11" pageNumber="604">
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<taxonomicName id="6F944D05FFFB2570FC812689BBC2FC6A" authority="(Gredler, 1885)" authorityName="Albers" authorityYear="1850" baseAuthorityName="Gredler" baseAuthorityYear="1885" box="[831,1437,950,977]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="scrobiculata">
<emphasis id="9AE0EA94FFFB2570FC812689BDE4FC6A" bold="true" box="[831,955,950,977]" italics="true" pageId="11" pageNumber="604">Hemiplecta</emphasis>
(?)
<emphasis id="9AE0EA94FFFB2570FC5A2689BAB2FC6A" bold="true" box="[996,1261,950,977]" italics="true" pageId="11" pageNumber="604">scrobiculata scrobiculata</emphasis>
(
<bibRefCitation id="CC054B77FFFB2570FB422688BBCAFC6A" author="GREDLER V." box="[1276,1429,950,977]" pageId="11" pageNumber="604" pagination="219 - 235" refId="ref7297" refString="GREDLER V. 1885. - Zur Conchylien-Fauna von China. VII. Stuck. Jahrbucher der Deutschen Malakozoologischen Gesellschaft 12: 219 - 235." type="journal article" year="1885">Gredler, 1885</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="E08E650DFFFB2570FB8B26E8BAF8FC4A" box="[1077,1191,982,1009]" pageId="11" pageNumber="604" type="description">
<paragraph id="A82B3686FFFB2570FB8B26E8BAF8FC4A" blockId="11.[813,1455,848,1009]" box="[1077,1191,982,1009]" pageId="11" pageNumber="604">
(
<figureCitation id="30AF2A03FFFB2570FB8026E9BAC1FC4A" box="[1086,1182,982,1009]" captionStart="FIG" captionStartId="12.[132,143,1985,2003]" captionTargetBox="[198,1389,215,1919]" captionTargetId="figure@12.[198,1389,214,1919]" captionTargetPageId="12" captionText="FIG. 7. — Shells of “Hemiplecta” species:A-D, Hemiplecta scrobiculata hupeiana (Gredler, 1887) (syntype,SMF 44672);E-I, Hemiplecta scrobiculata scrobiculata (Gredler, 1885), SMF 42575; J-H, Hemiplecta laotica (Möllendorff, 1899) (holotype, SMF 226681). Scale bars: 10 mm; I-J not to scale." figureDoi="http://doi.org/10.5281/zenodo.3726126" httpUri="https://zenodo.org/record/3726126/files/figure.png" pageId="11" pageNumber="604">Fig. 7E-I</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FC93212EBA27FB7B" pageId="11" pageNumber="604" type="reference_group">
<paragraph id="A82B3686FFFB2570FC93212EBA1BFBF8" blockId="11.[813,1456,1041,1141]" pageId="11" pageNumber="604">
<treatmentCitationGroup id="888411A8FFFB2570FC93212EBA1BFBF8" pageId="11" pageNumber="604">
<treatmentCitation id="29351097FFFB2570FC93212EBA84FB92" author="GREDLER V." box="[813,1243,1041,1065]" page="221" pageId="11" pageNumber="604" year="1885">
<taxonomicName id="6F944D05FFFB2570FC93212EBA84FB92" authority="Gredler, 1885: 220 - 221" authorityName="Gredler" authorityPageNumber="220 - 221" authorityYear="1885" box="[813,1243,1041,1065]" class="Gastropoda" family="Zonitidae" genus="Zonites" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="scrobiculatus">
<emphasis id="9AE0EA94FFFB2570FC93212EBDB3FB92" box="[813,1004,1041,1065]" italics="true" pageId="11" pageNumber="604">Zonites scrobiculatus</emphasis>
<bibRefCitation id="CC054B77FFFB2570FC4F212EBA84FB92" author="GREDLER V." box="[1009,1243,1041,1065]" pageId="11" pageNumber="604" pagination="219 - 235" refId="ref7297" refString="GREDLER V. 1885. - Zur Conchylien-Fauna von China. VII. Stuck. Jahrbucher der Deutschen Malakozoologischen Gesellschaft 12: 219 - 235." type="journal article" year="1885">Gredler, 1885: 220-221</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 6, fig. 2. —
<treatmentCitation id="29351097FFFB2570FAC9212EBA1FFBF8" author="BACHMANN O. &amp; GREDLER V." page="416" pageId="11" pageNumber="604" year="1894">
<bibRefCitation id="CC054B77FFFB2570FAC9212EBA1FFBF8" author="BACHMANN O. &amp; GREDLER V." pageId="11" pageNumber="604" pagination="415 - 429" refId="ref6963" refString="BACHMANN O. &amp; GREDLER V. 1894. - Zur Conchylien-Fauna von China. XVIII. Stuck. Annalen des K. K. Naturhistorischen Hofmuseums 9 (3): 415 - 429." type="journal article" year="1894">Bachmann &amp; Gredler 1894: 416</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="A82B3686FFFB2570FC932163BBEFFBCF" blockId="11.[813,1456,1041,1141]" box="[813,1456,1116,1141]" pageId="11" pageNumber="604">
<treatmentCitationGroup id="888411A8FFFB2570FC932163BBEFFBCF" box="[813,1456,1116,1141]" pageId="11" pageNumber="604">
<taxonomicName id="6F944D05FFFB2570FC932163BA22FBCF" authority="scrobiculata" authorityName="scrobiculata (Gredler" authorityYear="1885" box="[813,1149,1116,1140]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="scrobiculata">
<emphasis id="9AE0EA94FFFB2570FC932163BA22FBCF" box="[813,1149,1116,1140]" italics="true" pageId="11" pageNumber="604">Coccoglypta scrobiculata scrobiculata</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFFB2570FB2D2162BB7EFBCF" author="YEN T. - C." box="[1171,1313,1116,1141]" page="153" pageId="11" pageNumber="604" year="1939">
<bibRefCitation id="CC054B77FFFB2570FB2D2162BB7EFBCF" author="YEN T. - C." box="[1171,1313,1116,1141]" pageId="11" pageNumber="604" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen 1939: 153</bibRefCitation>
</treatmentCitation>
, pl. 15, fig. 62.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A82B3686FFFB2570FC9321B2BA27FB7B" blockId="11.[813,1455,1165,1292]" pageId="11" pageNumber="604">
<treatmentCitationGroup id="888411A8FFFB2570FC9321B2BA27FB7B" pageId="11" pageNumber="604">
<taxonomicName id="6F944D05FFFB2570FC9321B2BA55FB1E" baseAuthorityName="Gredler" baseAuthorityYear="1885" box="[813,1034,1165,1189]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="604" phylum="Mollusca" rank="species" species="scrobiculata">
<emphasis id="9AE0EA94FFFB2570FC9321B2BA55FB1E" box="[813,1034,1165,1189]" italics="true" pageId="11" pageNumber="604">Coccoglypta scrobiculata</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFFB2570FB9E21B1BB42FB1D" author="CHEN D. - N. &amp; ZHANG G. - Q." box="[1056,1309,1166,1190]" page="370" pageId="11" pageNumber="604" year="2004">
<bibRefCitation id="CC054B77FFFB2570FB9E21B1BB42FB1D" author="CHEN D. - N. &amp; ZHANG G. - Q." box="[1056,1309,1166,1190]" pageId="11" pageNumber="604" refId="ref7148" refString="CHEN D. - N. &amp; ZHANG G. - Q. 2004. - Fauna Sinica (Invertebrata Vol. 37 Mollusca: Gastropoda: Stylommatophora: Bradybaenidae). Science Press, Beijing, China, 482 p. (in Chinese)." type="book" year="2004">Chen &amp; Zhang 2004: 370</bibRefCitation>
</treatmentCitation>
, fig. 368 (treating
<emphasis id="9AE0EA94FFFB2570FCEC2197BDC1FB7B" box="[850,926,1192,1216]" italics="true" pageId="11" pageNumber="604">hupeina</emphasis>
as a junior synonym).
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="E08E650DFFFB2570FC9321E6BAF6FAB0" pageId="11" pageNumber="604" type="materials_examined">
<paragraph id="A82B3686FFFB2570FC9321E6BAF6FAB0" blockId="11.[813,1455,1165,1292]" pageId="11" pageNumber="604">
<collectionCode id="CE85AE43FFFB2570FC9321E6BDC7FB54" box="[813,920,1241,1265]" pageId="11" pageNumber="604">MATERIAL</collectionCode>
<collectionCode id="CE85AE43FFFB2570FC1A21E1BA4EFB54" box="[932,1041,1246,1263]" pageId="11" pageNumber="604">EXAMINED</collectionCode>
. —
<materialsCitation id="18FC3CDBFFFB2570FBF721E6BAFAFAB0" ID-GBIF-Occurrence="2580042131" pageId="11" pageNumber="604">
<emphasis id="9AE0EA94FFFB2570FBF721E6BAD2FB4A" bold="true" box="[1097,1165,1241,1265]" pageId="11" pageNumber="604">
<collectingCountry id="D0837616FFFB2570FBF721E6BAD2FB4A" box="[1097,1165,1241,1265]" name="China" pageId="11" pageNumber="604">China</collectingCountry>
</emphasis>
.
<collectingRegion id="6A50F864FFFB2570FB1E21E5BAB0FB4A" box="[1184,1263,1242,1265]" country="China" name="Hunan" pageId="11" pageNumber="604">Hunan</collectingRegion>
, Heng-san-hsien, coll. O. v. Möllendorff,
<collectionCode id="CE85AE43FFFB2570FBA821CBBA18FAB7" box="[1046,1095,1268,1292]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="604">SMF</collectionCode>
42575/3
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2570FC93201BBDDDFAEC" pageId="11" pageNumber="604" type="distribution">
<paragraph id="A82B3686FFFB2570FC93201BBDDDFAEC" blockId="11.[813,1455,1316,1367]" pageId="11" pageNumber="604">
<typeStatus id="772F8824FFFB2570FC93201BBD00FA81" box="[813,863,1316,1340]" pageId="11" pageNumber="604">TYPE</typeStatus>
LOCALITY. — “
<collectingMunicipality id="484FACFCFFFB2570FC40201ABA08FA86" box="[1022,1111,1317,1341]" pageId="11" pageNumber="604">Hen-san</collectingMunicipality>
, im
<location id="AD4B605DFFFB2570FB39201ABA81FA86" LSID="urn:lsid:plazi:treatment:203D8790FFFB2576FC812689BEAAFE88:AD4B605DFFFB2570FB39201ABA81FA86" box="[1159,1246,1317,1341]" municipality="Hen-san" name="Districte" pageId="11" pageNumber="604" stateProvince="Hunan">Districte</location>
von Hen-kiou-fu in
<collectingRegion id="6A50F864FFFB2570FC932000BD2AFAEC" box="[813,885,1343,1367]" country="China" name="Hunan" pageId="11" pageNumber="604">Hunan</collectingRegion>
”.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFB2576FC93204BBEAAFE88" lastPageId="13" lastPageNumber="606" pageId="11" pageNumber="604" type="description">
<paragraph id="A82B3686FFFB2570FC93204BBDE2FA36" blockId="11.[813,1456,1396,1934]" box="[813,957,1396,1422]" pageId="11" pageNumber="604">DESCRIPTION</paragraph>
<paragraph id="A82B3686FFFB2570FC9320AABAF8F836" blockId="11.[813,1456,1396,1934]" pageId="11" pageNumber="604">Shell dextral, medium-sized, dark yellowish-slightly greenish on both ventral and dorsal sides; shell depressed globularconical with a rounded body whorl; entire shell consisting of 4.75-5 whorls, separated by rather shallow suture; protoconch consisting of 1.75-2 whorls, very finely mamillated with recognisable spiral striation; dorsal side of teleoconch dominated by irregular, wavy ribs (waviness seemingly caused by spiral lines occurring only at back of ribs); area between middle line of body whorl (from apertural view) and ventral side (except for umbilicus region) glossy, with much weaker sculpture than on dorsal side or inside umbilicus; aperture oblique to shell axis, whitish in one specimen, but yellow in the other two; peristome not expanded; parietal callus weak, only indicated by fine whitish calcareous transparent layer; umbilicus open, rather narrow, deep, showing all whorls.</paragraph>
<paragraph id="A82B3686FFFB2570FC93228BBDE1F876" blockId="11.[813,1331,1972,2030]" box="[813,958,1972,1997]" pageId="11" pageNumber="604">
<emphasis id="9AE0EA94FFFB2570FC93228BBDE1F876" box="[813,958,1972,1997]" italics="true" pageId="11" pageNumber="604">Measurements</emphasis>
</paragraph>
<paragraph id="A82B3686FFFB2576FC9322EBBEB7FF4A" blockId="11.[813,1331,1972,2030]" lastBlockId="13.[132,775,217,307]" lastPageId="13" lastPageNumber="606" pageId="11" pageNumber="604">
D =
<quantity id="6F6C9B63FFFB2570FCE022ECBDA0F855" box="[862,1023,2003,2030]" metricMagnitude="-2" metricUnit="m" metricValue="1.9149999999999998" metricValueMax="1.9899999999999998" metricValueMin="1.8399999999999999" pageId="11" pageNumber="604" unit="mm" value="19.15" valueMax="19.9" valueMin="18.4">18.4-19.9 mm</quantity>
, H = 10.7-12.0 mm (n = 3). REMARKS
</paragraph>
<caption id="FCEB660EFFFC2577FF3A22FEBA87F851" ID-DOI="http://doi.org/10.5281/zenodo.3726126" ID-Zenodo-Dep="3726126" httpUri="https://zenodo.org/record/3726126/files/figure.png" pageId="12" pageNumber="605" startId="12.[132,143,1985,2003]" targetBox="[198,1389,215,1919]" targetPageId="12">
<paragraph id="A82B3686FFFC2577FF3A22FEBA87F851" blockId="12.[132,1456,1851,2026]" pageId="12" pageNumber="605">
FIG. 7. — Shells of “
<taxonomicName id="6F944D05FFFC2577FE9822FEBFDBF868" authorityName="Albers" authorityYear="1850" box="[294,388,1985,2003]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="605" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFC2577FE9822FEBFDBF868" box="[294,388,1985,2003]" italics="true" pageId="12" pageNumber="605">Hemiplecta</emphasis>
</taxonomicName>
” species:
<emphasis id="9AE0EA94FFFC2577FE6422FEBFA3F868" bold="true" box="[474,508,1985,2003]" pageId="12" pageNumber="605">A -D</emphasis>
,
<taxonomicName id="6F944D05FFFC2577FDB822FEBDC5F868" authority="(Gredler, 1887)" baseAuthorityName="Gredler" baseAuthorityYear="1887" box="[518,922,1985,2003]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="605" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="hupeiana">
<emphasis id="9AE0EA94FFFC2577FDB822FEBD42F868" box="[518,797,1985,2003]" italics="true" pageId="12" pageNumber="605">Hemiplecta scrobiculata hupeiana</emphasis>
(
<bibRefCitation id="CC054B77FFFC2577FC9822FEBDC9F868" author="GREDLER V." box="[806,918,1985,2003]" pageId="12" pageNumber="605" pagination="343 - 369" refId="ref7323" refString="GREDLER V. 1887. - Beschreibung neuer Arten und Varietaten. Jahrbucher der Deutschen Malakozoologischen Gesellschaft 14: 343 - 369." type="journal article" year="1887">Gredler, 1887</bibRefCitation>
)
</taxonomicName>
(syntype,SMF 44672);
<emphasis id="9AE0EA94FFFC2577FBE222FEBA29F868" bold="true" box="[1116,1142,1985,2003]" pageId="12" pageNumber="605">E -I</emphasis>
,
<taxonomicName id="6F944D05FFFC2577FBC122FEBF5EF851" authority="(Gredler, 1885)" authorityName="scrobiculata (Gredler" authorityYear="1885" baseAuthorityName="Gredler" baseAuthorityYear="1885" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="605" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="scrobiculata">
<emphasis id="9AE0EA94FFFC2577FBC122FEBBEFF868" box="[1151,1456,1985,2003]" italics="true" pageId="12" pageNumber="605">Hemiplecta scrobiculata scrobiculata</emphasis>
(
<bibRefCitation id="CC054B77FFFC2577FF3722E8BEA3F851" author="GREDLER V." box="[137,252,2007,2026]" pageId="12" pageNumber="605" pagination="219 - 235" refId="ref7297" refString="GREDLER V. 1885. - Zur Conchylien-Fauna von China. VII. Stuck. Jahrbucher der Deutschen Malakozoologischen Gesellschaft 12: 219 - 235." type="journal article" year="1885">Gredler, 1885</bibRefCitation>
)
</taxonomicName>
, SMF 42575;
<emphasis id="9AE0EA94FFFC2577FECD22E7BFCDF851" bold="true" box="[371,402,2008,2026]" pageId="12" pageNumber="605">J -H</emphasis>
,
<taxonomicName id="6F944D05FFFC2577FE2222E7BC88F851" authority="(Mollendorff, 1899)" baseAuthorityName="Mollendorff" baseAuthorityYear="1899" box="[412,727,2008,2026]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="605" phylum="Mollusca" rank="species" species="laotica">
<emphasis id="9AE0EA94FFFC2577FE2222E7BC69F851" box="[412,566,2008,2026]" italics="true" pageId="12" pageNumber="605">Hemiplecta laotica</emphasis>
(
<bibRefCitation id="CC054B77FFFC2577FDFE22E7BC8CF851" author="MOLLENDORFF O. &amp; VON" box="[576,723,2008,2026]" pageId="12" pageNumber="605" pagination="46 - 144" refId="ref7681" refString="MOLLENDORFF O. VON. 1899. - Binnen-Mollusken aus Westchina und Central-Asien Ezhegodnik Zoologicheskago muzeia (Annals of the Zoological Museum of the Saint Petersburg Academy of Sciences) 4 (1): 46 - 144." type="journal article" year="1899">Möllendorff, 1899</bibRefCitation>
)
</taxonomicName>
(holotype, SMF 226681). Scale bars: 10 mm; I-J not to scale.
</paragraph>
</caption>
<paragraph id="A82B3686FFFD2576FF3A25C6BEAAFE88" blockId="13.[132,775,217,307]" pageId="13" pageNumber="606">
<bibRefCitation id="CC054B77FFFD2576FF3A25C6BF50FEAF" author="ZILCH A." box="[132,271,249,276]" pageId="13" pageNumber="606" pagination="171 - 228" refId="ref8726" refString="ZILCH A. 1974. - Vinzenz Gredler und die Erforschung der Weichtiere Chinas durch Franziskaner aus Tirol. Archiv fur Molluskenkunde 104: 171 - 228." type="journal article" year="1974">Zilch (1974)</bibRefCitation>
found a single
<typeStatus id="772F8824FFFD2576FE0825C5BC57FEAF" box="[438,520,250,276]" pageId="13" pageNumber="606" type="syntype">syntype</typeStatus>
in Bolzano in Gredler’s collection.
</paragraph>
</subSubSection>
</treatment>
</document>

235
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@ -0,0 +1,235 @@
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<treatment id="203D8790FFFD2576FF1B2447BCA9FBCA" ID-DOI="http://doi.org/10.5281/zenodo.3730379" ID-GBIF-Taxon="163283963" ID-Zenodo-Dep="3730379" LSID="urn:lsid:plazi:treatment:203D8790FFFD2576FF1B2447BCA9FBCA" httpUri="http://treatment.plazi.org/id/203D8790FFFD2576FF1B2447BCA9FBCA" lastPageNumber="606" pageId="13" pageNumber="606">
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<paragraph id="A82B3686FFFD2576FF1B2447BCB9FE28" blockId="13.[132,776,376,489]" box="[165,742,376,403]" pageId="13" pageNumber="606">
<heading id="F36381EAFFFD2576FF1B2447BCB9FE28" box="[165,742,376,403]" level="2" pageId="13" pageNumber="606">
<taxonomicName id="6F944D05FFFD2576FF1B2447BCB9FE28" authorityName="Albers" authorityYear="1850" baseAuthorityName="Schileyko" baseAuthorityYear="2003" box="[165,742,376,403]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="606" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="hupeiana">
<emphasis id="9AE0EA94FFFD2576FF1B2447BF7EFE28" bold="true" box="[165,289,376,403]" italics="true" pageId="13" pageNumber="606">Hemiplecta</emphasis>
(?)
<emphasis id="9AE0EA94FFFD2576FEF42447BC68FE28" bold="true" box="[330,567,376,403]" italics="true" pageId="13" pageNumber="606">scrobiculata hupeiana</emphasis>
(
<bibRefCitation id="CC054B77FFFD2576FDFB2446BC81FE28" author="GREDLER V." box="[581,734,377,403]" pageId="13" pageNumber="606" pagination="343 - 369" refId="ref7323" refString="GREDLER V. 1887. - Beschreibung neuer Arten und Varietaten. Jahrbucher der Deutschen Malakozoologischen Gesellschaft 14: 343 - 369." type="journal article" year="1887">Gredler, 1887</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FE3824A6BC59FE08" box="[390,518,408,435]" pageId="13" pageNumber="606" type="description">
<paragraph id="A82B3686FFFD2576FE3824A6BC59FE08" blockId="13.[132,776,376,489]" box="[390,518,408,435]" pageId="13" pageNumber="606">
(
<figureCitation id="30AF2A03FFFD2576FE3124A7BFA2FE08" box="[399,509,408,435]" captionStart="FIG" captionStartId="12.[132,143,1985,2003]" captionTargetBox="[198,1389,215,1919]" captionTargetId="figure@12.[198,1389,214,1919]" captionTargetPageId="12" captionText="FIG. 7. — Shells of “Hemiplecta” species:A-D, Hemiplecta scrobiculata hupeiana (Gredler, 1887) (syntype,SMF 44672);E-I, Hemiplecta scrobiculata scrobiculata (Gredler, 1885), SMF 42575; J-H, Hemiplecta laotica (Möllendorff, 1899) (holotype, SMF 226681). Scale bars: 10 mm; I-J not to scale." figureDoi="http://doi.org/10.5281/zenodo.3726126" httpUri="https://zenodo.org/record/3726126/files/figure.png" pageId="13" pageNumber="606">Fig. 7A-D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FF3A24EDBF8FFD88" pageId="13" pageNumber="606" type="reference_group">
<paragraph id="A82B3686FFFD2576FF3A24EDBD57FE52" blockId="13.[132,776,376,489]" box="[132,776,465,489]" pageId="13" pageNumber="606">
<treatmentCitationGroup id="888411A8FFFD2576FF3A24EDBD57FE52" box="[132,776,465,489]" pageId="13" pageNumber="606">
<taxonomicName id="6F944D05FFFD2576FF3A24EDBE98FE52" ID-CoL="7PLG3" box="[132,199,466,489]" class="Gastropoda" family="Zonitidae" genus="Zonites" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="606" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFD2576FF3A24EDBE98FE52" box="[132,199,466,489]" italics="true" pageId="13" pageNumber="606">Zonites</emphasis>
</taxonomicName>
(
<taxonomicName id="6F944D05FFFD2576FF6A24EDBF43FE52" authorityName="Sowerby" authorityYear="1842" box="[212,284,466,489]" class="Gastropoda" family="Camaenidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="606" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFFD2576FF6A24EDBF43FE52" box="[212,284,466,489]" italics="true" pageId="13" pageNumber="606">Nanina</emphasis>
</taxonomicName>
?)
<treatmentCitation id="29351097FFFD2576FE8E24EEBD5BFE52" author="GREDLER V." box="[304,772,465,489]" page="345" pageId="13" pageNumber="606" year="1887">
<taxonomicName id="6F944D05FFFD2576FE8E24EEBD5BFE52" authority="Gredler, 1887: 344 - 345" authorityName="Gredler" authorityPageNumber="344 - 345" authorityYear="1887" box="[304,772,465,489]" class="Gastropoda" family="Zonitidae" genus="Zonites" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="606" phylum="Mollusca" rank="variety" species="scrobiculatus" variety="hupeina">
<emphasis id="9AE0EA94FFFD2576FE8E24EEBFFBFE52" box="[304,420,465,489]" italics="true" pageId="13" pageNumber="606">scrobiculatus</emphasis>
var.
<emphasis id="9AE0EA94FFFD2576FE7124EEBC45FE52" box="[463,538,465,489]" italics="true" pageId="13" pageNumber="606">hupeina</emphasis>
<bibRefCitation id="CC054B77FFFD2576FDA024EEBD5BFE52" author="GREDLER V." box="[542,772,465,489]" pageId="13" pageNumber="606" pagination="343 - 369" refId="ref7323" refString="GREDLER V. 1887. - Beschreibung neuer Arten und Varietaten. Jahrbucher der Deutschen Malakozoologischen Gesellschaft 14: 343 - 369." type="journal article" year="1887">Gredler, 1887: 344-345</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A82B3686FFFD2576FF3A273EBF8FFD88" blockId="13.[132,776,513,941]" pageId="13" pageNumber="606">
<treatmentCitationGroup id="888411A8FFFD2576FF3A273EBF8FFD88" pageId="13" pageNumber="606">
<taxonomicName id="6F944D05FFFD2576FF3A273EBFEAFDA2" baseAuthorityName="Gredler" baseAuthorityYear="1887" box="[132,437,513,537]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="606" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="hupeina">
<emphasis id="9AE0EA94FFFD2576FF3A273EBFEAFDA2" box="[132,437,513,537]" italics="true" pageId="13" pageNumber="606">Coccoglypta scrobiculata hupeina</emphasis>
</taxonomicName>
–
<treatmentCitation id="29351097FFFD2576FE75273EBC06FDA2" author="YEN T. - C." box="[459,601,513,537]" page="153" pageId="13" pageNumber="606" year="1939">
<bibRefCitation id="CC054B77FFFD2576FE75273EBC06FDA2" author="YEN T. - C." box="[459,601,513,537]" pageId="13" pageNumber="606" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen 1939: 153</bibRefCitation>
</treatmentCitation>
, pl. 15, fig. 63. —
<treatmentCitation id="29351097FFFD2576FF3A2724BF2CFD88" author="CHEN D-N. &amp; GAO J. - X." box="[132,371,539,564]" page="159" pageId="13" pageNumber="606" year="1987">
<bibRefCitation id="CC054B77FFFD2576FF3A2724BF2CFD88" author="CHEN D-N. &amp; GAO J. - X." box="[132,371,539,564]" pageId="13" pageNumber="606" refId="ref7112" refString="CHEN D-N. &amp; GAO J. - X. 1987. - Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing, China. 186 p. (in Chinese)" type="book" year="1987">Chen &amp; Gao 1987: 159</bibRefCitation>
</treatmentCitation>
, fig. 205.
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</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FF3A277ABFB7FCFB" pageId="13" pageNumber="606" type="materials_examined">
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<collectionCode id="CE85AE43FFFD2576FF3A277ABEEAFDE1" box="[132,181,581,605]" pageId="13" pageNumber="606">TYPE</collectionCode>
<collectionCode id="CE85AE43FFFD2576FF022775BFDBFDE0" box="[188,388,586,603]" pageId="13" pageNumber="606">MATERIAL EXAMINED</collectionCode>
. —
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<emphasis id="9AE0EA94FFFD2576FE0C277ABFAFFDE6" bold="true" box="[434,496,581,605]" pageId="13" pageNumber="606">
<collectingCountry id="D0837616FFFD2576FE0C277ABFAFFDE6" box="[434,496,581,605]" name="China" pageId="13" pageNumber="606">China</collectingCountry>
</emphasis>
.
<collectorName id="05615350FFFD2576FE43277ABC1CFDE6" box="[509,579,581,605]" pageId="13" pageNumber="606">Hu-bei</collectorName>
(Hupei), coll.
<collectorName id="05615350FFFD2576FD68277ABEA0FDCC" pageId="13" pageNumber="606">O. v. Möllendorff</collectorName>
, ex coll.
<collectorName id="05615350FFFD2576FEE52760BFF7FDCC" box="[347,424,607,631]" pageId="13" pageNumber="606">Gredler</collectorName>
,
<collectionCode id="CE85AE43FFFD2576FE0D275FBFBAFDC3" box="[435,485,608,632]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="606">SMF</collectionCode>
42576/
<specimenCount id="BE92FD0FFFFD2576FD892760BCFCFDC3" box="[567,675,607,632]" pageId="13" pageNumber="606" type="generic" typeStatus="syntypes">2 syntypes</specimenCount>
</materialsCitation>
. —
<materialsCitation id="18FC3CDBFFFD2576FD71275FBF3BFD7C" ID-GBIF-Occurrence="2580042148" collectionCode="SMF" collectorName="Hupe &amp; G. Naegele &amp; Gredler &amp; O. Boettger" country="China" location="Hubei" pageId="13" pageNumber="606" specimenCount="2" stateProvince="Hubei" typeStatus="syntype">
<collectorName id="05615350FFFD2576FD71275FBD58FDC3" box="[719,775,608,632]" pageId="13" pageNumber="606">Hupé</collectorName>
(Hupei), coll.
<collectorName id="05615350FFFD2576FEB02745BF22FD29" box="[270,381,634,658]" pageId="13" pageNumber="606">G. Naegele</collectorName>
, ex coll.
<collectorName id="05615350FFFD2576FE6E2745BC44FD29" box="[464,539,634,658]" pageId="13" pageNumber="606">Gredler</collectorName>
,
<collectionCode id="CE85AE43FFFD2576FD9B2745BC09FD29" box="[549,598,634,658]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="606">SMF</collectionCode>
50091/
<specimenCount id="BE92FD0FFFFD2576FD162745BD5CFD29" box="[680,771,634,658]" pageId="13" pageNumber="606" type="generic" typeStatus="syntype">1 syntype</specimenCount>
; China:
<collectingRegion id="6A50F864FFFD2576FF6E27ABBF50FD17" box="[208,271,660,684]" country="China" name="Hubei" pageId="13" pageNumber="606">Hubei</collectingRegion>
(Hupei), coll.
<collectorName id="05615350FFFD2576FE1927ABBC40FD16" box="[423,543,660,685]" pageId="13" pageNumber="606">O. Boettger</collectorName>
, ex coll.
<collectorName id="05615350FFFD2576FDC427ABBC9AFD17" box="[634,709,660,684]" pageId="13" pageNumber="606">Gredler</collectorName>
1888,
<collectionCode id="CE85AE43FFFD2576FF3A2790BEEAFD7C" box="[132,181,687,711]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="606">SMF</collectionCode>
44672/
<specimenCount id="BE92FD0FFFFD2576FEB62790BF3BFD7C" box="[264,356,687,711]" pageId="13" pageNumber="606" type="generic" typeStatus="syntype">1 syntype</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="A82B3686FFFD2576FF3A27E7BFB7FCFB" blockId="13.[132,776,513,941]" pageId="13" pageNumber="606">
ADDITIONAL
<collectionCode id="CE85AE43FFFD2576FEBA27E2BF96FD55" box="[260,457,733,750]" pageId="13" pageNumber="606">MATERIAL EXAMINED</collectionCode>
. —
<materialsCitation id="18FC3CDBFFFD2576FE4C27E7BFBBFCFB" ID-GBIF-Occurrence="2580042140" collectingDate="2010-11-05" collectorName="Mufucun SW &amp; A. Hunyadi" country="China" county="Enshi Tujiazu Miaozu Zizhizhou" latitude="30.433817" location="Enshi Daxiagu" longLatPrecision="1" longitude="109.171" municipality="Enshi Shi" pageId="13" pageNumber="606" specimenCount="1" stateProvince="Hubei">
<emphasis id="9AE0EA94FFFD2576FE4C27E7BC6FFD4B" bold="true" box="[498,560,728,752]" pageId="13" pageNumber="606">
<collectingCountry id="D0837616FFFD2576FE4C27E7BC6FFD4B" box="[498,560,728,752]" name="China" pageId="13" pageNumber="606">China</collectingCountry>
</emphasis>
.
<collectingRegion id="6A50F864FFFD2576FD8527E7BC24FD4B" box="[571,635,728,752]" country="China" name="Hubei" pageId="13" pageNumber="606">Hubei</collectingRegion>
,
<collectingCounty id="414A4E0AFFFD2576FD3A27E7BF65FCB0" pageId="13" pageNumber="606">Enshi Tujiazu Miaozu Zizhizhou</collectingCounty>
,
<collectingMunicipality id="484FACFCFFFD2576FEFC27CCBFFEFCB0" box="[322,417,755,779]" pageId="13" pageNumber="606">Enshi Shi</collectingMunicipality>
,
<collectorName id="05615350FFFD2576FE1427CCBC6FFCB0" box="[426,560,755,779]" pageId="13" pageNumber="606">Mufucun SW</collectorName>
<quantity id="6F6C9B63FFFD2576FD8B27CCBC36FCB0" box="[565,617,755,779]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="606" unit="km" value="3.0">3 km</quantity>
,
<location id="AD4B605DFFFD2576FDCC27CCBD5BFCB0" LSID="urn:lsid:plazi:treatment:203D8790FFFD2576FF1B2447BCA9FBCA:AD4B605DFFFD2576FDCC27CCBD5BFCB0" box="[626,772,755,779]" country="China" county="Enshi Tujiazu Miaozu Zizhizhou" latitude="30.433817" longLatPrecision="1" longitude="109.171" municipality="Enshi Shi" name="Enshi Daxiagu" pageId="13" pageNumber="606" stateProvince="Hubei">Enshi Daxiagu</location>
,
<location id="AD4B605DFFFD2576FF3A2632BF35FC9E" LSID="urn:lsid:plazi:treatment:203D8790FFFD2576FF1B2447BCA9FBCA:AD4B605DFFFD2576FF3A2632BF35FC9E" box="[132,362,781,805]" country="China" county="Enshi Tujiazu Miaozu Zizhizhou" latitude="30.433817" longLatPrecision="1" longitude="109.171" municipality="Enshi Shi" name="Mother-child Affection" pageId="13" pageNumber="606" stateProvince="Hubei">Mother-child Affection</location>
;
<geoCoordinate id="CDA05041FFFD2576FECD2632BFABFC9E" box="[371,500,781,805]" degrees="30" direction="north" minutes="26.029" orientation="latitude" pageId="13" pageNumber="606" precision="1" value="30.433817">30°26.029’N</geoCoordinate>
,
<geoCoordinate id="CDA05041FFFD2576FDBE2632BCD5FC9E" box="[512,650,781,805]" degrees="109" direction="east" minutes="10.260" orientation="longitude" pageId="13" pageNumber="606" precision="1" value="109.171">109°10.260’E</geoCoordinate>
;
<collectorName id="05615350FFFD2576FD2A2632BD58FC9E" box="[660,775,781,805]" pageId="13" pageNumber="606">A. Hunyadi</collectorName>
leg.;
<date id="DC2A1046FFFD2576FF0D2617BF79FCFB" box="[179,294,808,832]" pageId="13" pageNumber="606" value="2010-11-05">
<collectingDate id="CC6EE9AEFFFD2576FF0D2617BF79FCFB" box="[179,294,808,832]" pageId="13" pageNumber="606" value="2010-11-05">05.XI.2010</collectingDate>
</date>
; coll. HA/10 shells
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FF3A266EBFCAFCD2" box="[132,405,849,873]" pageId="13" pageNumber="606" type="distribution">
<paragraph id="A82B3686FFFD2576FF3A266EBFCAFCD2" blockId="13.[132,776,513,941]" box="[132,405,849,873]" pageId="13" pageNumber="606">
<typeStatus id="772F8824FFFD2576FF3A266EBEEAFCDD" box="[132,181,849,873]" pageId="13" pageNumber="606">TYPE</typeStatus>
LOCALITY. — “Hupé”.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FF3A2645BD57FC16" pageId="13" pageNumber="606" type="diagnosis">
<paragraph id="A82B3686FFFD2576FF3A2645BD57FC16" blockId="13.[132,776,513,941]" pageId="13" pageNumber="606">
DIFFERENTIAL DIAGNOSIS. —
<taxonomicName id="6F944D05FFFD2576FE272644BC98FC28" baseAuthorityName="Gredler" baseAuthorityYear="1887" box="[409,711,891,915]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="606" phylum="Mollusca" rank="subSpecies" species="scrobiculata" subSpecies="hupeiana">
<emphasis id="9AE0EA94FFFD2576FE272644BC98FC28" box="[409,711,891,915]" italics="true" pageId="13" pageNumber="606">Hemiplecta scrobiculata hupeiana</emphasis>
</taxonomicName>
differs from the nominotypical subspecies by the larger, slightly keeled shell.
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FF3A26FBBD58FC66" box="[132,775,963,989]" pageId="13" pageNumber="606" type="description">
<paragraph id="A82B3686FFFD2576FF3A26FBBD58FC66" blockId="13.[132,775,963,989]" box="[132,775,963,989]" pageId="13" pageNumber="606">
MEASUREMENTS. — D:
<quantity id="6F6C9B63FFFD2576FEDC26FCBFA5FC66" box="[354,506,963,989]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" metricValueMax="2.33" metricValueMin="2.01" pageId="13" pageNumber="606" unit="mm" value="21.7" valueMax="23.3" valueMin="20.1">20.1-23.3 mm</quantity>
, H:
<quantity id="6F6C9B63FFFD2576FD9D26FCBCE6FC66" box="[547,697,963,989]" metricMagnitude="-2" metricUnit="m" metricValue="1.165" metricValueMax="1.24" metricValueMin="1.09" pageId="13" pageNumber="606" unit="mm" value="11.65" valueMax="12.4" valueMin="10.9">10.9-12.4 mm</quantity>
(n = 4).
</paragraph>
</subSubSection>
<subSubSection id="E08E650DFFFD2576FF3A2129BCA9FBCA" pageId="13" pageNumber="606" type="discussion">
<paragraph id="A82B3686FFFD2576FF3A2129BEB7FB95" blockId="13.[132,775,1046,1137]" box="[132,232,1046,1072]" pageId="13" pageNumber="606">REMARKS</paragraph>
<paragraph id="A82B3686FFFD2576FF3A2109BCA9FBCA" blockId="13.[132,775,1046,1137]" pageId="13" pageNumber="606">
The examined
<typeStatus id="772F8824FFFD2576FE9C2108BF22FBEA" box="[290,381,1079,1105]" pageId="13" pageNumber="606" type="syntype">syntypes</typeStatus>
were taken from Gredler’s collection to the Senckenberg Museum by Adolf Zilch (
<bibRefCitation id="CC054B77FFFD2576FDD02169BCB9FBCB" author="ZILCH A." box="[622,742,1110,1136]" pageId="13" pageNumber="606" pagination="171 - 228" refId="ref8726" refString="ZILCH A. 1974. - Vinzenz Gredler und die Erforschung der Weichtiere Chinas durch Franziskaner aus Tirol. Archiv fur Molluskenkunde 104: 171 - 228." type="journal article" year="1974">Zilch 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

90
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Linnaeus
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,
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1
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: 156. 1753
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.
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<paragraph id="6513199ACFCF8BC52858304ACCCA73E4" pageNumber="438">&quot;Habitat in Europa meridionali.&quot; RCN: 1242.</paragraph>
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<paragraph id="3BC5548A98B764CDBBCA7A97BCB5A4AF" pageNumber="438">
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(
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in
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281: 210. 1967): Herb. Linn. No. 218.26 (
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.
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<paragraph id="40981FAA4EAD952E2FB617F80318EAF8" pageNumber="438">
Current name:
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<emphasis id="67BC7B26F18C35BA5050E8E76BC5E81C" bold="true" italics="true" pageNumber="438">Convolvulus althaeoides</emphasis>
</emphasis>
L.
</taxonomicName>
(
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).
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<treatment LSID="urn:lsid:plazi:treatment:203DFB62AA405B81A470ADD7B939349A" httpUri="http://treatment.plazi.org/id/203DFB62AA405B81A470ADD7B939349A" lastPageNumber="47537" pageId="0" pageNumber="47537">
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<taxonomicName LSID="203DFB62-AA40-5B81-A470-ADD7B939349A" authority="Lacepede, 1802" authorityName="Lacepede" authorityYear="1802" class="Actinopterygii" family="Chaetodontidae" genus="Chaetodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chaetodon lunula" order="Perciformes" pageId="0" pageNumber="47537" phylum="Chordata" rank="species" species="lunula">
Chaetodon lunula (
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, 1802)
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</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47537" type="materials_examined">
<paragraph pageId="0" pageNumber="47537">Materials</paragraph>
<paragraph pageId="0" pageNumber="47537">
<materialsCitation country="Malaysia" location="Redang islands" pageId="0" pageNumber="47537" specimenCount="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="47537">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="47537">Other material</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="47537">Occurrence:</emphasis>
occurrenceID: BDJ_12482_62;
<emphasis bold="true" pageId="0" pageNumber="47537">Location:</emphasis>
country:
<collectingCountry pageId="0" pageNumber="47537">Malaysia</collectingCountry>
; locality:
<location LSID="urn:lsid:plazi:treatment:203DFB62AA405B81A470ADD7B939349A:0A1EC07D047284914E68634581F44695" country="Malaysia" name="Redang islands" pageId="0" pageNumber="47537">Redang islands</location>
;
<emphasis bold="true" pageId="0" pageNumber="47537">Identification:</emphasis>
identifiedBy:
<determinerName pageId="0" pageNumber="47537">Loh KH and Du Jianguo</determinerName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47537" type="notes">
<paragraph pageId="0" pageNumber="47537">Notes</paragraph>
<paragraph pageId="0" pageNumber="47537">
<bibRefCitation author="Yusuf, Y. B." journalOrPublisher="Department of Fisheries, Kuala Lumpur and KUSTEM, Kuala Terengganu, Malaysia" pageId="0" pageNumber="47537" refId="B5376898" refString="Yusuf, Y. B., Mohd-Norizam, M., Ali, A. B., Zaidnuddin, I., 2001. Coral reef fish of some selected sites at Pulau Redang Marine Park, Terenggganu: A brief study. Department of Fisheries, Kuala Lumpur and KUSTEM, Kuala Terengganu, Malaysia" title="Coral reef fish of some selected sites at Pulau Redang Marine Park, Terenggganu: A brief study" year="2001">Yusuf et al. 2001</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>