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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.85399" ID-Pensoft-Pub="1313-2970-1110-39" ID-Pensoft-UUID="766F9A94793D5793A9BE1E70F575798B" ID-ZooBank="4681CC6DE5F347C6B1D052DEA78BE7C3" ModsDocID="1313-2970-1110-39" checkinTime="1657006630715" checkinUser="pensoft" docAuthor="Sian Man, Nem, Siriboon, Thanit, Lin, Aung, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="202F05341A86581EB8988C1AD67FD947" docLanguage="en" docName="ZooKeys 1110: 39-102" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.85399" docTitle="Carinartemis exacutus" docType="treatment" docVersion="1" id="766F9A94793D5793A9BE1E70F575798B" lastPageNumber="39" masterDocId="766F9A94793D5793A9BE1E70F575798B" masterDocTitle="Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species" masterLastPageNumber="102" masterPageNumber="39" pageNumber="39" updateTime="1657006630715" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sian Man, Nem</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4453-734X</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Siriboon, Thanit</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok, 10110, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Aung</mods:namePart>
<mods:affiliation>Fauna and Flora International, No. 35, 3</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1110</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>39</mods:start>
<mods:end>102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1110.85399</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.85399</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1110-39</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947" httpUri="http://treatment.plazi.org/id/202F05341A86581EB8988C1AD67FD947" lastPageNumber="39" pageId="0" pageNumber="39">
<subSubSection pageId="0" pageNumber="39" type="nomenclature">
<paragraph pageId="0" pageNumber="39">
<taxonomicName LSID="202F0534-1A86-581E-B898-8C1AD67FD947" authority="(Gould, 1856)" baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">Carinartemis exacutus (Gould, 1856)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="description">
<paragraph pageId="0" pageNumber="39">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Approximate sampling locations used in this study. The location name and coordinates are listed in Table 1. Historical type localities of a broad scale, such as Tavoy, Burma, Moulmein, Mergui, and Tenasserim, are not indicated on this map." figureDoi="10.3897/zookeys.1110.85399.figure1" httpUri="https://binary.pensoft.net/fig/710466" pageId="0" pageNumber="39">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Living snails A Discartemon tonywhitteni from near Phra (Buddha) Cave, Tanintharyi Region (shell width ~ 10 mm) B Discartemon paurodeviatus sp. nov. from the type locality (shell width ~ 11 mm) C Haploptychius heliakosus sp. nov. from the type locality (shell height ~ 9 mm) D Carinartemis exacutus from Lun Nga Mountain, Kayin State (shell height ~ 14 mm) E, F Carinartemis sankeyi (E) from Kayon Cave, Mon State (shell height ~ 10 mm) and (F) from Saddan Cave, Mawlamyine, Mon State (shell height ~ 10 mm)." figureDoi="10.3897/zookeys.1110.85399.figure2" httpUri="https://binary.pensoft.net/fig/710467" pageId="0" pageNumber="39">, 2D</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A, B Carinartemis petitii A syntype MCZ 169290 from Tavoy, British Burmah and B specimen NHMUK 1909.3.15.67 from Moulmein C, D Carinartemis exacutus C specimen NHMUK 1888.12.4.771 - 773 from Moulmein with apertural dentition and D specimen CUMZ 13025 from Taung Wine Cave, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure17" httpUri="https://binary.pensoft.net/fig/710482" pageId="0" pageNumber="39">, 17C, D</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Carinartemis exacutus A specimen NHMUK 1906.2.2.198 from Moulmein B specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an, Kayin State C specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39">, 18</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">, 21A, B</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Internal sculpture of genitalia of Carinartemis exacutus, specimen CUMZ 13021 A atrium surface B overview of internal penial wall C arrangement of penial hooks on proximal part of penis D lateral view of penial hooks E arrangement of penial hooks on middle part of penis F lateral view penial hooks G arrangement of longitudinal oblique vaginal folds." figureDoi="10.3897/zookeys.1110.85399.figure22" httpUri="https://binary.pensoft.net/fig/710487" pageId="0" pageNumber="39">, 22</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Radula of specimens A Discartemon tonywhitteni, paratype CUMZ 13001 B Discartemon paurodeviatus sp. nov., paratype CUMZ 13003 C Haploptychius heliakosus sp. nov., paratype CUMZ 13014 D Carinartemis exacutus, specimen CUMZ 13021 E, F Carinartemis sankeyi, specimen CUMZ 13029 E overview of radula and F high magnification of central teeth." figureDoi="10.3897/zookeys.1110.85399.figure24" httpUri="https://binary.pensoft.net/fig/710489" pageId="0" pageNumber="39">, 24D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="reference_group">
<paragraph pageId="0" pageNumber="39">
<taxonomicName authorityName="Gould" authorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">Streptaxis exacutus</taxonomicName>
Gould, 1856: 13. Type locality: Burma [Myanmar].
</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis exacuta" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacuta">Streptaxis exacuta</taxonomicName>
[sic] -
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Brockhaus, Lipsiae" pageId="0" pageNumber="39" refId="B61" refString="Pfeiffer, L, 1859. Monographia Heliceorum Viventium sistens Descriptiones Systematicas et Criticas Omnium Huius Familiae Generum et Specierum Hodie Cognitarum. Volume 4. F.A. Brockhaus, Lipsiae" title="Monographia Heliceorum Viventium sistens Descriptiones Systematicas et Criticas Omnium Huius Familiae Generum et Specierum Hodie Cognitarum. Volume 4. F. A." year="1859">Pfeiffer 1859</bibRefCitation>
: 331.
</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName authorityName="Gould" authorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">Streptaxis exacutus</taxonomicName>
-
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="39" refId="B64" refString="Pfeiffer, L, 1871. Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 4. Verlag von Theodor Fischer Cassel 1-171 pls 109-137. [pp 25-40 pls 118-125 (1871) ; pp 145-171, pls 135-137 (1876). Published in parts, dates follow Johnson (1969)]" title="Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 4. Verlag von Theodor Fischer Cassel 1 - 171 pls 109 - 137. [pp 25 - 40 pls 118 - 125 (1871); pp 145 - 171, pls 135 - 137 (1876). Published in parts, dates follow Johnson (1969)]" year="1871">Pfeiffer 1871</bibRefCitation>
: 30, 31, pl. 115, figs 13, 14.
<bibRefCitation author="Hanley, SCT" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" refId="B33" refString="Hanley, SCT, Theobald, W, 1870. Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London[i-xviii+] 1-65 pp.[160 pls.] [pp 1-18 pls 1-40(1870) ; pp 19-28, pls 41-60 (1872); pp 29-34, pls 61-80 (1873); pp 35-48, pls 81-120 (1874); pp 49-56, pls 121-140 (1875); pp 57-65, pls 141-160 (1876)] [Published in parts, dates after Prashad (1927)]." title="Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London [i-xviii +] 1 - 65 pp. [160 pls.] [pp 1 - 18 pls 1 - 40 (1870); pp 19 - 28, pls 41 - 60 (1872); pp 29 - 34, pls 61 - 80 (1873); pp 35 - 48, pls 81 - 120 (1874); pp 49 - 56, pls 121 - 140 (1875); pp 57 - 65, pls 141 - 160 (1876)] [Published in parts, dates after Prashad (1927)]." year="1870">Hanley and Theobald 1870</bibRefCitation>
: 40, pl. 98, figs 8-10.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="39" refId="B48" refString="Nevill, G, 1878. Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
: 3.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.6534" author="Tryon Jr, GW" journalOrPublisher="Gustav Fischer, Jena" pageId="0" pageNumber="39" refId="B90" refString="Tryon Jr, GW, 1885. Manual of Conchology, Structure and Systematic, with Illustrations of the Species. Volume 1. The Academy of Natural Science of Philadelphia, 364 pp. https://doi.org/10.5962/bhl.title.6534" title="Manual of Conchology, Structure and Systematic, with Illustrations of the Species. Volume 1. The Academy of Natural Science of Philadelphia, 364 pp." url="https://doi.org/10.5962/bhl.title.6534" year="1885">Tryon 1885</bibRefCitation>
: 72, pl. 14, figs 11, 12. Blandford and Godwin-Austin 1908: 8.
</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">Haploptychius exacutus</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Journal of the Society for Bibliography of Natural History" pageId="0" pageNumber="39" refId="B37" refString="Kobelt, W, 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1(12b) (2): 1-211 pls42-71 [pp. 1-96 pls 42-59 (1905) ; 97-211, pls 60-71 (1906)]." title="Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (12 b) (2): 1 - 211 pls 42 - 71 [pp. 1 - 96 pls 42 - 59 (1905); 97 - 211, pls 60 - 71 (1906)]." year="1905">Kobelt 1906</bibRefCitation>
: 142, 143, pl. 57, figs 16-18.
<bibRefCitation author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
: 215.
<bibRefCitation author="Siriboon, T" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="39" pagination="161 - 174" refId="B77" refString="Siriboon, T, Sutcharit, C, Naggs, F, Rowson, B, Panha, S, 2014b. Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae). The Raffles Bulletin of Zoology 62: 161 - 174" title="Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)." volume="62" year="2014 b">Siriboon et al. 2014b</bibRefCitation>
: 169, 171.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="materials_examined">
<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="39">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Myanmar" county="Hpa-an District" latitude="16.842335" location="Blanford" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" specimenCount="73" specimenCount-juvenile="2" stateProvince="Kayin State">
Moulmein: NHMUK 1874.9.3.15 (3 shells) ex. Godwin-Austen. NHMUK 1888.12.4.771-773 (3 shells; Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A, B Carinartemis petitii A syntype MCZ 169290 from Tavoy, British Burmah and B specimen NHMUK 1909.3.15.67 from Moulmein C, D Carinartemis exacutus C specimen NHMUK 1888.12.4.771 - 773 from Moulmein with apertural dentition and D specimen CUMZ 13025 from Taung Wine Cave, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure17" httpUri="https://binary.pensoft.net/fig/710482" pageId="0" pageNumber="39">17C</figureCitation>
). NHMUK 1906.2.2.198 (3 shells +
<specimenCount type="juvenile">2 juveniles</specimenCount>
; Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Carinartemis exacutus A specimen NHMUK 1906.2.2.198 from Moulmein B specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an, Kayin State C specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39">18A</figureCitation>
) ex.
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:55688629ACBB939B5C5B016B0CA3685D" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Blanford" stateProvince="Kayin State">Blanford</location>
collection. NHMUK 1903.7.1.3999 (3 shells) ex.
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:CC3CFFEA6A907B2E4B7222AFB6355061" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Godwin-Austen" stateProvince="Kayin State">Godwin-Austen</location>
collection. Burma: NHMUK1950.12.9.170 (1 shell) ex.
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:C7E56C935EF3600DF4015CB12DA4419B" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Laidlaw" stateProvince="Kayin State">Laidlaw</location>
collection.
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:0F0317CEEC56CBD840A1F3752D4AC998" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Mergui" stateProvince="Kayin State">Mergui</location>
: NHMUK ex.
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:4009D16CBFFF7E46ED2E50012E2E45B7" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Cuming" stateProvince="Kayin State">Cuming</location>
collection (2 shells).
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:3FA5B5748BD172486A6F4F2BD89972D2" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Sadhdan Cave" stateProvince="Kayin State">Sadhdan Cave</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:25DC79E28759BAD4B2D1BBF49C6ED17A" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an Township" stateProvince="Kayin State">Hpa-an Township</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:BB1AE7376015F305B38A923F48686B1A" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an District" stateProvince="Kayin State">Hpa-an District</location>
,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
,
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate degrees="16" direction="north" minutes="44" orientation="latitude" precision="1" seconds="23.4" value="16.739834">16°44'23.4&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="43" orientation="longitude" precision="1" seconds="04.2" value="97.717834">97°43'04.2&quot;E</geoCoordinate>
): CUMZ 13020 (3 shells).
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:BCAAEA2C716C088137BEC3CB6BBD0382" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Bayin Nyi Cave" stateProvince="Kayin State">Bayin Nyi Cave</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:CE6F5E2C14C25A4195A0BEBE89CBE9B0" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an Township" stateProvince="Kayin State">Hpa-an Township</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:35CA49639A9E8134B79AE225ABC969CE" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an District" stateProvince="Kayin State">Hpa-an District</location>
,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
,
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate degrees="16" direction="north" minutes="58" orientation="latitude" precision="1" seconds="10.1" value="16.969473">16°58'10.1&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="29" orientation="longitude" precision="1" seconds="30.6" value="97.49183">97°29'30.6&quot;E</geoCoordinate>
): CUMZ 13021 (8 shells; Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Carinartemis exacutus A specimen NHMUK 1906.2.2.198 from Moulmein B specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an, Kayin State C specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39">18B</figureCitation>
), CUMZ 13022 (
<specimenCount type="generic">15 specimens</specimenCount>
in ethanol).
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:37FCC48805D59B587DE76236A8AE59C7" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Lun Nga Mountain" stateProvince="Kayin State">Lun Nga Mountain</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:3C8431B3F2219D42DF40DB468FB958EB" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an Township" stateProvince="Kayin State">Hpa-an Township</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:A239B6DC27B20137A185F7A9FE5B2273" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an District" stateProvince="Kayin State">Hpa-an District</location>
,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
,
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate degrees="16" direction="north" minutes="44" orientation="latitude" precision="1" seconds="53.2" value="16.748112">16°44'53.2&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="47" orientation="longitude" precision="1" seconds="09.5" value="97.78597">97°47'09.5&quot;E</geoCoordinate>
): CUMZ 13023 (14 shells; Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Carinartemis exacutus A specimen NHMUK 1906.2.2.198 from Moulmein B specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an, Kayin State C specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39">18C</figureCitation>
), CUMZ 13024 (
<specimenCount type="generic">50 specimens</specimenCount>
in ethanol).
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:8C1F29DE3D812638FD7419FECE5CC1EB" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Taung Wine Cave" stateProvince="Kayin State">Taung Wine Cave</location>
, near Thiri Hpa-an Hotel,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:C943BD1A5480D4C202E2F29B758A8FD4" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an Township" stateProvince="Kayin State">Hpa-an Township</location>
,
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:6112C50E65FEAA131D094419A4529BFE" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Hpa-an District" stateProvince="Kayin State">Hpa-an District</location>
,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
,
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate degrees="16" direction="north" minutes="50" orientation="latitude" precision="1" seconds="31.1" value="16.841972">16°50'31.1&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="37" orientation="longitude" precision="1" seconds="18.4" value="97.62178">97°37'18.4&quot;E</geoCoordinate>
): CUMZ 13025 (3 shells; Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A, B Carinartemis petitii A syntype MCZ 169290 from Tavoy, British Burmah and B specimen NHMUK 1909.3.15.67 from Moulmein C, D Carinartemis exacutus C specimen NHMUK 1888.12.4.771 - 773 from Moulmein with apertural dentition and D specimen CUMZ 13025 from Taung Wine Cave, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure17" httpUri="https://binary.pensoft.net/fig/710482" pageId="0" pageNumber="39">17D</figureCitation>
), CUMZ13026 (
<specimenCount type="generic">6 specimens</specimenCount>
in ethanol).
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:3BFAE07DD7BC9E2AA8D9E02E2CF3926C" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Kaw Ka Taung Cave" stateProvince="Kayin State">Kaw Ka Taung Cave</location>
(
<location LSID="urn:lsid:plazi:treatment:202F05341A86581EB8988C1AD67FD947:D653A60708A673F129661365014A36C9" country="Myanmar" county="Hpa-an District" latitude="16.842335" longLatPrecision="1" longitude="97.6197" municipality="Hpa-an Township" name="Golden valley" stateProvince="Kayin State">Golden valley</location>
),
<collectingMunicipality>Hpa-an Township</collectingMunicipality>
,
<collectingCounty>Hpa-an District</collectingCounty>
,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
,
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate degrees="16" direction="north" minutes="50" orientation="latitude" precision="1" seconds="32.4" value="16.842335">16°50'32.4&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="37" orientation="longitude" precision="1" seconds="10.9" value="97.6197">97°37'10.9&quot;E</geoCoordinate>
): CUMZ 13027 (1 shell)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">Figure 18.</emphasis>
<taxonomicName baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">
<emphasis italics="true" pageId="0" pageNumber="39">Carinartemis exacutus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="39">A</emphasis>
specimen NHMUK 1906.2.2.198 from Moulmein
<emphasis bold="true" pageId="0" pageNumber="39">B</emphasis>
specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
<emphasis bold="true" pageId="0" pageNumber="39">C</emphasis>
specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an,
<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
with apertural dentition.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="diagnosis">
<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">
<emphasis italics="true" pageId="0" pageNumber="39">Carinartemis exacutus</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName genus="C." lsidName="C. sankeyi" pageId="0" pageNumber="39" rank="species" species="sankeyi">
<emphasis italics="true" pageId="0" pageNumber="39">C. sankeyi</emphasis>
</taxonomicName>
,
<taxonomicName genus="C." lsidName="C. vesperus" pageId="0" pageNumber="39" rank="species" species="vesperus">
<emphasis italics="true" pageId="0" pageNumber="39">C. vesperus</emphasis>
</taxonomicName>
Siriboon &amp; Panha, 2014, and
<taxonomicName genus="C." lsidName="C. striatus" pageId="0" pageNumber="39" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="39">C. striatus</emphasis>
</taxonomicName>
Siriboon &amp; Panha, 2014 from western Thailand, but it has a larger shell, convex spire, immediately expanded penultimate whorl, semi-ovate aperture, two parietal lamellae, very thickened penial sheath, vas deferens passes through penial sheath, curved portion of vas deferens with very thin connective tissue, and hooks located on irregular trapezoidal penial papillae separated by longitudinal folds. In comparison,
<taxonomicName genus="C." lsidName="C. sankeyi" pageId="0" pageNumber="39" rank="species" species="sankeyi">
<emphasis italics="true" pageId="0" pageNumber="39">C. sankeyi</emphasis>
</taxonomicName>
,
<taxonomicName genus="C." lsidName="C. vesperus" pageId="0" pageNumber="39" rank="species" species="vesperus">
<emphasis italics="true" pageId="0" pageNumber="39">C. vesperus</emphasis>
</taxonomicName>
, and
<taxonomicName genus="C." lsidName="C. striatus" pageId="0" pageNumber="39" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="39">C. striatus</emphasis>
</taxonomicName>
have an elevated spire, regularly expanded penultimate whorl, but
<taxonomicName genus="C." lsidName="C. sankeyi" pageId="0" pageNumber="39" rank="species" species="sankeyi">
<emphasis italics="true" pageId="0" pageNumber="39">C. sankeyi</emphasis>
</taxonomicName>
has fine transverse ridges on the entire shell, subquadrangular aperture, one parietal lamella, slender atrium, thin penial sheath, vas deferens does not pass through a penial sheath, curved portion of vas deferens is without connective tissue, proximal penial hook located on laterally-flattened penial papillae, and distal penial hooks located on laterally compressed penial papillae separated by reticulated folds. In contrast,
<taxonomicName genus="C." lsidName="C. vesperus" pageId="0" pageNumber="39" rank="species" species="vesperus">
<emphasis italics="true" pageId="0" pageNumber="39">C. vesperus</emphasis>
</taxonomicName>
has a subcircular aperture, lacks parietal lamellae, has a less axially deflected last whorl, vas deferens passes through the penial sheath, curved portion of vas deferens is without connective tissue, and penial papillae absent. Meanwhile,
<taxonomicName genus="C." lsidName="C. striatus" pageId="0" pageNumber="39" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="39">C. striatus</emphasis>
</taxonomicName>
has strong transverse ridges over the entire shell, a semi-ovate aperture, one parietal lamella, vas deferens is attached to the distal end of the penial sheath with very thin connective tissue, and hooks are located on papillae without connected longitudinal folds.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="description">
<paragraph pageId="0" pageNumber="39">Description.</paragraph>
<paragraph pageId="0" pageNumber="39">
Shell oblique-heliciform, white, and translucent; whorls 6-
<normalizedToken originalValue="6½">61/2</normalizedToken>
; spire convex with distinct suture. Shell surface glossy with fine transverse ridges, nearly smooth with few transverse ridges near peristome; varices present. Embryonic shell ~
<normalizedToken originalValue="2½">21/2</normalizedToken>
whorls with smooth surface; following whorls intermediately coiled. Shell periphery wide and sharply keeled along nearly the entire penultimate whorl; last whorl axially deflected. Aperture semi-ovate; peristome discontinuous, thickened, expanded, and slightly reflected. Apertural dentition with one strong parietal lamella and sometimes with a second parietal lamella adjoined at a right angle. Umbilicus open and deep (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A, B Carinartemis petitii A syntype MCZ 169290 from Tavoy, British Burmah and B specimen NHMUK 1909.3.15.67 from Moulmein C, D Carinartemis exacutus C specimen NHMUK 1888.12.4.771 - 773 from Moulmein with apertural dentition and D specimen CUMZ 13025 from Taung Wine Cave, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure17" httpUri="https://binary.pensoft.net/fig/710482" pageId="0" pageNumber="39">17C, D</figureCitation>
,
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Carinartemis exacutus A specimen NHMUK 1906.2.2.198 from Moulmein B specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an, Kayin State C specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39">18</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">
<emphasis italics="true" pageId="0" pageNumber="39">Genital organs</emphasis>
.
</emphasis>
Atrium (at) short. Penis (p) a very thin, and long tube. Penial sheath (ps) muscularly enlarged, very thickened and extending entire length of penis; penial sheath retractor muscle (psr) thin, originating near genital orifice, attached to atrium with short and thin connective tissue, and inserting distally on penial sheath (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">21A</figureCitation>
). Vas deferens (vd) passes through a short portion of penial sheath, then extends to curved portion at ~ 1/11 of the penial sheath length before entering penis distally. Curved portion of vas deferens with very thin connective tissue originating at penial sheath (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">21B</figureCitation>
). Penial retractor muscle (pr) thin, very long, inserting at penis and vas deferens junction.
</paragraph>
<paragraph pageId="0" pageNumber="39">
Internal wall of atrium generally smooth with sparse atrial pores (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Internal sculpture of genitalia of Carinartemis exacutus, specimen CUMZ 13021 A atrium surface B overview of internal penial wall C arrangement of penial hooks on proximal part of penis D lateral view of penial hooks E arrangement of penial hooks on middle part of penis F lateral view penial hooks G arrangement of longitudinal oblique vaginal folds." figureDoi="10.3897/zookeys.1110.85399.figure22" httpUri="https://binary.pensoft.net/fig/710487" pageId="0" pageNumber="39">22A</figureCitation>
). Penial wall with scattered pale brownish penial hooks, ~ 20 hooks/200
<normalizedToken originalValue="μm">μm</normalizedToken>
2; hooks located on laterally flattened penial papillae and separated by longitudinal folds (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Internal sculpture of genitalia of Carinartemis exacutus, specimen CUMZ 13021 A atrium surface B overview of internal penial wall C arrangement of penial hooks on proximal part of penis D lateral view of penial hooks E arrangement of penial hooks on middle part of penis F lateral view penial hooks G arrangement of longitudinal oblique vaginal folds." figureDoi="10.3897/zookeys.1110.85399.figure22" httpUri="https://binary.pensoft.net/fig/710487" pageId="0" pageNumber="39">22B-D</figureCitation>
). Penial hooks small (&lt;0.02 mm in length), expanded at base, tips pointed and curved towards genital orifice (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Internal sculpture of genitalia of Carinartemis exacutus, specimen CUMZ 13021 A atrium surface B overview of internal penial wall C arrangement of penial hooks on proximal part of penis D lateral view of penial hooks E arrangement of penial hooks on middle part of penis F lateral view penial hooks G arrangement of longitudinal oblique vaginal folds." figureDoi="10.3897/zookeys.1110.85399.figure22" httpUri="https://binary.pensoft.net/fig/710487" pageId="0" pageNumber="39">22E, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="39">
Vagina (v) very short and ~ 1/12 of penis length. Gametolytic duct (gd) a long tube extending as far as albumin gland; gametolytic sac (gs) ovate. Proximal free oviduct (fo) enlarged then tapering to smaller diameter distally. Oviduct (ov) enlarged and folded; prostate gland inconspicuous and bound to oviduct. Talon (ta) small, short and club shaped. Hermaphroditic duct (hd) bearing long seminal vesicle (sv) ~ 1/2 the length from talon to branching point of seminal vesicle (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">21A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="39">
Vaginal wall with longitudinal oblique vaginal folds, folds with nearly smooth surface and vaginal hook absent (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Internal sculpture of genitalia of Carinartemis exacutus, specimen CUMZ 13021 A atrium surface B overview of internal penial wall C arrangement of penial hooks on proximal part of penis D lateral view of penial hooks E arrangement of penial hooks on middle part of penis F lateral view penial hooks G arrangement of longitudinal oblique vaginal folds." figureDoi="10.3897/zookeys.1110.85399.figure22" httpUri="https://binary.pensoft.net/fig/710487" pageId="0" pageNumber="39">22G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="39" phylum="Marchantiophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="39">Radula</emphasis>
</taxonomicName>
.
</emphasis>
Each row consists of 41-45 teeth with formula (22-20)-1-(20-22). Central tooth is very small with pointed cusp. Lateral and marginal teeth undifferentiated, lanceolate, unicuspid, and lanceolate. Latero-marginal teeth gradually reduced in size, with outermost teeth much smaller and shorter than inner teeth (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Radula of specimens A Discartemon tonywhitteni, paratype CUMZ 13001 B Discartemon paurodeviatus sp. nov., paratype CUMZ 13003 C Haploptychius heliakosus sp. nov., paratype CUMZ 13014 D Carinartemis exacutus, specimen CUMZ 13021 E, F Carinartemis sankeyi, specimen CUMZ 13029 E overview of radula and F high magnification of central teeth." figureDoi="10.3897/zookeys.1110.85399.figure24" httpUri="https://binary.pensoft.net/fig/710489" pageId="0" pageNumber="39">24D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="distribution">
<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="39">This species was collected from five limestone hills in Kayin State, southern Myanmar, in this survey.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="remarks">
<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="39">
This species appears at a high abundance among the limestone karsts in Hpa-an, Kayin State. All the specimens examined from the populations from Lun Nga Mountain and Bayin Nyi Cave have a very small to indistinct upper parietal lamella, and some specimens from Bayin Nyi Cave have strong transverse ridges on almost the entire whorl. Moreover, the Lun Nga Mountain population have a last whorl that is more extended anteriorly and an elongated semi-ovate aperture (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Carinartemis exacutus A specimen NHMUK 1906.2.2.198 from Moulmein B specimen CUMZ 13021 from Bayin Nyi Cave, Hpa-an, Kayin State C specimen CUMZ 13023 from Lun Nga Mountain, Hpa-an, Kayin State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure18" httpUri="https://binary.pensoft.net/fig/710483" pageId="0" pageNumber="39">18C</figureCitation>
). However, the genitalia in all localities show no difference from this locality; thus, we consider them as intraspecific variations within
<taxonomicName genus="C." lsidName="C. exacutus" pageId="0" pageNumber="39" rank="species" species="exacutus">
<emphasis italics="true" pageId="0" pageNumber="39">C. exacutus</emphasis>
</taxonomicName>
. Some specimens from Bayin Nyi Cave have strong transverse ridges on almost the entire whorl.
</paragraph>
<paragraph pageId="0" pageNumber="39">
Although the type locality was listed as
<normalizedToken originalValue="Burma">'Burma'</normalizedToken>
[Myanmar], all the known records and specimens examined in this study were collected from Kayin State. Therefore, the precise type locality of this species is probably southwestern Myanmar in Mon State and Kayin State.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,48 @@
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<taxonomicName id="2BC4B66E5D01CC33FE83767C917D0FD0" ID-CoL="3ZLND" authority="Habermehl, 1923" authorityName="Habermehl" authorityYear="1923" class="Insecta" family="Ichneumonidae" genus="Meloboris" kingdom="Animalia" lsidName="Meloboris neglecta" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="neglecta">Meloboris neglecta (Habermehl, 1923)</taxonomicName>
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<taxonomicName id="2ED019D164EBE9D8DD2065A5C68FE97D" class="Insecta" family="Ichneumonidae" genus="Pseudocymodusa" kingdom="Animalia" lsidName="Pseudocymodusa neglecta" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="neglecta">Pseudocymodusa neglecta</taxonomicName>
Habermehl, 1923
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<mods:title id="5F54A4C667977B3DAC3D85D30B4B1A0B">A review of the Arcuphantes species with the genitalia of arcuatulus - type (Araneae, Linyphiidae)</mods:title>
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<heading id="F371B82109208C50FF6A8764F27FFA4D" bold="true" box="[151,612,1391,1417]" fontSize="11" level="1" pageId="9" pageNumber="507" reason="1">
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<taxonomicName id="6F8674CE09208C50FF6A8764F1D6FA4D" ID-CoL="GC72" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[151,461,1391,1417]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="507" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="9AF2D35F09208C50FF6A8764F1D6FA4D" bold="true" box="[151,461,1391,1417]" italics="true" pageId="9" pageNumber="507">Arcuphantes denticulatus</emphasis>
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<figureCitation id="30BD13C809208C50FF6A8799F11CFA6F" box="[151,263,1426,1451]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="9" pageNumber="507">Figs 2FJ</figureCitation>
,
<figureCitation id="30BD13C809208C50FEEE8799F138FA6F" box="[275,291,1426,1451]" captionStart="FIGURE 6" captionStartId="9.[151,250,1258,1280]" captionTargetBox="[159,1423,193,1241]" captionTargetId="figure@9.[159,1423,193,1242]" captionTargetPageId="9" captionText="FIGURE 6. Arcuphantes denticulatus n. sp. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE, embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270397/files/figure.png" pageId="9" pageNumber="507">6</figureCitation>
,
<figureCitation id="30BD13C809208C50FED38798F170FA6E" box="[302,363,1427,1450]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="9" pageNumber="507">7EF</figureCitation>
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<paragraph id="A8390F4D09208C50FF6A87D1F576F9D3" blockId="9.[151,1436,1498,2027]" pageId="9" pageNumber="507">
<emphasis id="9AF2D35F09208C50FF6A87D1F1DFFA37" bold="true" box="[151,452,1498,1523]" pageId="9" pageNumber="507">Type material examined.</emphasis>
<emphasis id="9AF2D35F09208C50FE3387D1F22FFA36" box="[462,564,1498,1522]" italics="true" pageId="9" pageNumber="507">Holotype</emphasis>
♂.
<materialsCitation id="18EE051009208C50FD9987D0F399F9D2" ID-GBIF-Occurrence="1848253883" collectingDate="1959-08-27" collectorName="W. J. Gertsch" country="United States of America" latitude="47.883335" location="Of" longLatPrecision="1114" longitude="-122.01667" pageId="9" pageNumber="507" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
<collectingCountry id="D0914FDD09208C50FD9987D0F284FA37" box="[612,671,1499,1523]" name="United States of America" pageId="9" pageNumber="507">USA</collectingCountry>
,
<emphasis id="9AF2D35F09208C50FD5087D1F327FA37" bold="true" box="[685,828,1498,1523]" pageId="9" pageNumber="507">
<collectingRegion id="6A42C1AF09208C50FD5087D1F327FA37" box="[685,828,1498,1523]" country="United States of America" name="Washington" pageId="9" pageNumber="507">Washington</collectingRegion>
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:
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northwest
<location id="AD59599609208C50FBD987D1F420FA37" LSID="urn:lsid:plazi:treatment:202FBE5B09208C50FF6A8764F25DF82F:AD59599609208C50FBD987D1F420FA37" box="[1060,1083,1498,1523]" country="United States of America" latitude="47.883335" longLatPrecision="1114" longitude="-122.01667" name="Of" pageId="9" pageNumber="507" stateProvince="Washington">of</location>
Monroe,
<geoCoordinate id="CDB2698A09208C50FB5287D0F53DFA37" box="[1199,1318,1499,1523]" direction="west" orientation="longitude" pageId="9" pageNumber="507" precision="925" value="-122.01667">122°01W</geoCoordinate>
;
<geoCoordinate id="CDB2698A09208C50FAC987D1F58CFA37" box="[1332,1431,1498,1523]" direction="north" orientation="latitude" pageId="9" pageNumber="507" precision="925" value="47.883335">47°53N</geoCoordinate>
,
<date id="DC38298D09208C50FF6A87F5F13AF9D3" box="[151,289,1534,1559]" pageId="9" pageNumber="507" value="1959-08-27">
<collectingDate id="CC7CD06509208C50FF6A87F5F13AF9D3" box="[151,289,1534,1559]" pageId="9" pageNumber="507" value="1959-08-27">27.viii.1959</collectingDate>
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,
<collectorName id="05736A9B09208C50FED187F4F1D9F9D3" box="[300,450,1534,1559]" pageId="9" pageNumber="507">W. J. Gertsch</collectorName>
&amp; V. Roth leg. (AMNH-146).
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<typeStatus id="773DB1EF09208C50FCE187F4F399F9D2" box="[796,898,1535,1558]" pageId="9" pageNumber="507" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
.
<materialsCitation id="18EE051009208C50FC6D87F4F572F9D3" ID-GBIF-Occurrence="1848253884" box="[912,1385,1534,1559]" collectingDate="1959-08-27" collectorName="W. J. Gertsch" country="United States of America" latitude="47.883335" location="Of" longLatPrecision="1114" longitude="-122.01667" pageId="9" pageNumber="507" specimenCount="1" specimenCount-male="1" stateProvince="Washington" typeStatus="holotype">
<specimenCount id="BE80C4C409208C50FC6D87F4F3AFF9D3" box="[912,948,1535,1559]" pageId="9" pageNumber="507" type="male">1♂</specimenCount>
, same data, together with the
<typeStatus id="773DB1EF09208C50FAFE87F5F572F9D3" box="[1283,1385,1534,1559]" pageId="9" pageNumber="507" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609208C50FF3A8429F289F99B" pageId="9" pageNumber="507" type="etymology">
<paragraph id="A8390F4D09208C50FF3A8429F289F99B" blockId="9.[151,1436,1498,2027]" pageId="9" pageNumber="507">
<emphasis id="9AF2D35F09208C50FF3A8429F155F9FF" bold="true" box="[199,334,1570,1595]" pageId="9" pageNumber="507">Etymology.</emphasis>
The species name comes from the Latin words denticulatus, in reference to the fine and close teeth on the central serrated area of the radix.
</paragraph>
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<subSubSection id="E09C5CC609208C50FF3A8461F141F8D7" pageId="9" pageNumber="507" type="diagnosis">
<paragraph id="A8390F4D09208C50FF3A8461F141F8D7" blockId="9.[151,1436,1498,2027]" pageId="9" pageNumber="507">
<emphasis id="9AF2D35F09208C50FF3A8461F15AF947" bold="true" box="[199,321,1642,1667]" pageId="9" pageNumber="507">Diagnosis.</emphasis>
The male
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<emphasis id="9AF2D35F09208C50FE3C8460F1C9F946" box="[449,466,1643,1666]" italics="true" pageId="9" pageNumber="507">A</emphasis>
.
<emphasis id="9AF2D35F09208C50FE1E8461F277F946" box="[483,620,1642,1666]" italics="true" pageId="9" pageNumber="507">denticulatus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="81C16E2409208C50FD888461F2ACF947" box="[629,695,1642,1667]" pageId="9" pageNumber="507" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished by the bifid outer paracymbial arm with a sharp inner apophysis (
<figureCitation id="30BD13C809208C50FE5F8485F21BF963" box="[418,512,1678,1703]" captionStart="FIGURE 6" captionStartId="9.[151,250,1258,1280]" captionTargetBox="[159,1423,193,1241]" captionTargetId="figure@9.[159,1423,193,1242]" captionTargetPageId="9" captionText="FIGURE 6. Arcuphantes denticulatus n. sp. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE, embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270397/files/figure.png" pageId="9" pageNumber="507">Figs 6C</figureCitation>
,
<figureCitation id="30BD13C809208C50FDF28484F22BF963" box="[527,560,1679,1703]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="9" pageNumber="507">7F</figureCitation>
), which is blunt in
<taxonomicName id="6F8674CE09208C50FCEB8485F3B4F962" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[790,943,1678,1702]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="507" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F09208C50FCEB8485F3B4F962" box="[790,943,1678,1702]" italics="true" pageId="9" pageNumber="507">A. arcuatulus</emphasis>
</taxonomicName>
(
<figureCitation id="30BD13C809208C50FC3D8485F404F963" box="[960,1055,1678,1703]" captionStart="FIGURE 4" captionStartId="7.[151,250,1960,1982]" captionTargetBox="[151,1436,840,1908]" captionTargetId="figure@7.[151,1436,840,1911]" captionTargetPageId="7" captionText="FIGURE 4. Arcuphantes arcuatulus. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE. embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270393/files/figure.png" pageId="9" pageNumber="507">Figs 4C</figureCitation>
,
<figureCitation id="30BD13C809208C50FBD08484F449F963" box="[1069,1106,1679,1703]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="9" pageNumber="507">7B</figureCitation>
); the flat terminal apophysis having the anterior projection extending forwards (
<figureCitation id="30BD13C809208C50FD1F84B9F353F90F" box="[738,840,1714,1739]" captionStart="FIGURE 6" captionStartId="9.[151,250,1258,1280]" captionTargetBox="[159,1423,193,1241]" captionTargetId="figure@9.[159,1423,193,1242]" captionTargetPageId="9" captionText="FIGURE 6. Arcuphantes denticulatus n. sp. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE, embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270397/files/figure.png" pageId="9" pageNumber="507">Figs. 6D</figureCitation>
,
<figureCitation id="30BD13C809208C50FCAB84B8F362F90F" box="[854,889,1715,1739]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="9" pageNumber="507">7E</figureCitation>
), and by the central serrated area with fine and close teeth (
<figureCitation id="30BD13C809208C50FEE384DDF18BF92B" box="[286,400,1750,1775]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="9" pageNumber="507">
Figs.
<date id="DC38298D09208C50FEA384DCF16EF92B" box="[350,373,1751,1775]" pageId="9" pageNumber="507">2I</date>
J
</figureCitation>
,
<figureCitation id="30BD13C809208C50FE6184DDF1FAF92B" box="[412,481,1750,1775]" captionStart="FIGURE 6" captionStartId="9.[151,250,1258,1280]" captionTargetBox="[159,1423,193,1241]" captionTargetId="figure@9.[159,1423,193,1242]" captionTargetPageId="9" captionText="FIGURE 6. Arcuphantes denticulatus n. sp. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE, embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270397/files/figure.png" pageId="9" pageNumber="507">6DE</figureCitation>
), while it is uneven in
<taxonomicName id="6F8674CE09208C50FD1F84DDF362F92A" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[738,889,1750,1774]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="507" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F09208C50FD1F84DDF362F92A" box="[738,889,1750,1774]" italics="true" pageId="9" pageNumber="507">A. arcuatulus</emphasis>
</taxonomicName>
and
<taxonomicName id="6F8674CE09208C50FC4D84DDF423F92A" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[944,1080,1750,1774]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="507" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F09208C50FC4D84DDF3DAF929" box="[944,961,1750,1773]" italics="true" pageId="9" pageNumber="507">A</emphasis>
.
<emphasis id="9AF2D35F09208C50FC2D84DCF423F92A" box="[976,1080,1751,1774]" italics="true" pageId="9" pageNumber="507">cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F09208C50FBC284DDF465F92B" bold="true" box="[1087,1150,1750,1775]" pageId="9" pageNumber="507">
<taxonomicNameLabel id="81C16E2409208C50FBC284DDF465F92B" box="[1087,1150,1750,1775]" pageId="9" pageNumber="507" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="30BD13C809208C50FB7384DDF510F92B" box="[1166,1291,1750,1775]" captionStart="FIGURE 4" captionStartId="7.[151,250,1960,1982]" captionTargetBox="[151,1436,840,1908]" captionTargetId="figure@7.[151,1436,840,1911]" captionTargetPageId="7" captionText="FIGURE 4. Arcuphantes arcuatulus. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE. embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270393/files/figure.png" pageId="9" pageNumber="507">Figs 4DE</figureCitation>
,
<figureCitation id="30BD13C809208C50FAEA84DDF546F92B" box="[1303,1373,1750,1775]" captionStart="FIGURE 5" captionStartId="8.[151,250,1905,1927]" captionTargetBox="[171,1415,193,1871]" captionTargetId="figure@8.[171,1415,193,1878]" captionTargetPageId="8" captionText="FIGURE 5. Arcuphantes cruciatus n. sp. AB, male palp, with embolic division removed, A, prolateral; B, retrolateral; CD, embolic division, arrows point to central serrated area of radix, C, ventral; D, dorsal; EH, epigynum, with extensible basal part removed, E, ventral, ventral plate artificially broken; F, dorsal; G, caudal; H, inner, shows epigynal grooves. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270395/files/figure.png" pageId="9" pageNumber="507">5DE</figureCitation>
). The female unknown.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609208C50FF3A8515F12DF803" pageId="9" pageNumber="507" type="description">
<paragraph id="A8390F4D09208C50FF3A8515F355F8BB" blockId="9.[151,1436,1498,2027]" pageId="9" pageNumber="507">
<emphasis id="9AF2D35F09208C50FF3A8515F142F8F3" bold="true" box="[199,345,1822,1847]" pageId="9" pageNumber="507">Description.</emphasis>
<emphasis id="9AF2D35F09208C50FEA28515F182F8F2" box="[351,409,1822,1846]" italics="true" pageId="9" pageNumber="507">Male</emphasis>
(AMNH-146): Total length 2.29. Carapace 0.90 long, 0.75 wide. Abdomen 1.36 long, 0.75 wide. Leg lengths: I 9.02 (2.52+2.95+2.14+1.41); II 8.02 (2.16+2.51+2.18+1.17), III 5.36 (1.63+1.51+1.43+0.79); IV 7.49 (2.04+2.28+1.96+1.21). Tm I 0.22. Female unknown.
</paragraph>
<paragraph id="A8390F4D09208C50FF3A8581F12DF803" blockId="9.[151,1436,1498,2027]" pageId="9" pageNumber="507">
Other somatic and genital characters: see the general description for the
<taxonomicName id="6F8674CE09208C50FC0B8581F498F866" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1014,1155,1930,1954]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F09208C50FC0B8581F498F866" box="[1014,1155,1930,1954]" italics="true" pageId="9" pageNumber="507">Arcuphantes</emphasis>
</taxonomicName>
species with
<taxonomicName id="6F8674CE09208C50FAE18581F588F866" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1308,1427,1930,1954]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="507" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F09208C50FAE18581F588F866" box="[1308,1427,1930,1954]" italics="true" pageId="9" pageNumber="507">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF09208C50FF6A85A4F0DCF803" box="[151,199,1967,1991]" pageId="9" pageNumber="507">type</typeStatus>
genitalia.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609208C50FF3A85D9F25DF82F" box="[199,582,2002,2027]" pageId="9" pageNumber="507" type="distribution">
<paragraph id="A8390F4D09208C50FF3A85D9F25DF82F" blockId="9.[151,1436,1498,2027]" box="[199,582,2002,2027]" pageId="9" pageNumber="507">
<emphasis id="9AF2D35F09208C50FF3A85D9F17BF82F" bold="true" box="[199,352,2002,2027]" pageId="9" pageNumber="507">Distribution.</emphasis>
<collectingCountry id="D0914FDD09208C50FE9A85D8F184F82F" box="[359,415,2003,2027]" name="United States of America" pageId="9" pageNumber="507">USA</collectingCountry>
(
<collectingRegion id="6A42C1AF09208C50FE5085D9F221F82F" box="[429,570,2002,2027]" country="United States of America" name="Washington" pageId="9" pageNumber="507">Washington</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,178 @@
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<mods:title id="306887C526096F50AE6638AA0363D973">A review of the Arcuphantes species with the genitalia of arcuatulus - type (Araneae, Linyphiidae)</mods:title>
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<mods:namePart id="A99B4D8A1E87FC38DD48AE8034124D07">Tu, Lihong</mods:namePart>
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<treatment id="202FBE5B09238C52FF6A84A5F229FE14" ID-DOI="http://doi.org/10.5281/zenodo.5998058" ID-GBIF-Taxon="144235599" ID-Zenodo-Dep="5998058" LSID="urn:lsid:plazi:treatment:202FBE5B09238C52FF6A84A5F229FE14" httpUri="http://treatment.plazi.org/id/202FBE5B09238C52FF6A84A5F229FE14" lastPageId="11" lastPageNumber="509" pageId="10" pageNumber="508">
<subSubSection id="E09C5CC609238C53FF6A84A5F118F92D" pageId="10" pageNumber="508" type="nomenclature">
<paragraph id="A8390F4D09238C53FF6A84A5F28BF90C" blockId="10.[151,656,1710,1770]" box="[151,656,1710,1736]" pageId="10" pageNumber="508">
<heading id="F371B82109238C53FF6A84A5F28BF90C" bold="true" box="[151,656,1710,1736]" fontSize="11" level="1" pageId="10" pageNumber="508" reason="1">
<emphasis id="9AF2D35F09238C53FF6A84A5F28BF90C" bold="true" box="[151,656,1710,1736]" pageId="10" pageNumber="508">
<taxonomicName id="6F8674CE09238C53FF6A84A5F1E2F90C" ID-CoL="GC7Y" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[151,505,1710,1736]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="508" phylum="Arthropoda" rank="species" species="semiorbiculatus" status="sp. nov.">
<emphasis id="9AF2D35F09238C53FF6A84A5F1E2F90C" bold="true" box="[151,505,1710,1736]" italics="true" pageId="10" pageNumber="508">Arcuphantes semiorbiculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81C16E2409238C53FDFD84A5F28BF90C" box="[512,656,1710,1736]" pageId="10" pageNumber="508" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A8390F4D09238C53FF6A84DAF118F92D" blockId="10.[151,656,1710,1770]" box="[151,259,1745,1770]" pageId="10" pageNumber="508">
<figureCitation id="30BD13C809238C53FF6A84DAF118F92D" box="[151,259,1745,1770]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="10" pageNumber="508">Fig. 3GI</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609238C53FF6A8512F121F8BE" pageId="10" pageNumber="508" type="materials_examined">
<paragraph id="A8390F4D09238C53FF6A8512F121F8BE" blockId="10.[151,1436,1816,2022]" pageId="10" pageNumber="508">
<emphasis id="9AF2D35F09238C53FF6A8512F1DBF8F6" bold="true" box="[151,448,1817,1842]" pageId="10" pageNumber="508">Type material examined.</emphasis>
<emphasis id="9AF2D35F09238C53FE3B8512F252F8F6" box="[454,585,1816,1842]" italics="true" pageId="10" pageNumber="508">Holotype ♀</emphasis>
.
<materialsCitation id="18EE051009238C53FDA58512F386F891" ID-GBIF-Occurrence="1848253929" collectingDate="1958-07-11" collectorName="W. J. Gertsch" country="United States of America" latitude="36.15" location="west Of NelsOn Camp" longLatPrecision="1189" longitude="-118.75" municipality="Creek" pageId="10" pageNumber="508" specimenCount="1" stateProvince="California" typeStatus="paratype">
<collectingCountry id="D0914FDD09238C53FDA58512F28FF8F5" box="[600,660,1817,1841]" name="United States of America" pageId="10" pageNumber="508">USA</collectingCountry>
:
<emphasis id="9AF2D35F09238C53FD628512F301F8F6" bold="true" box="[671,794,1817,1842]" pageId="10" pageNumber="508">
<collectingRegion id="6A42C1AF09238C53FD628512F301F8F6" box="[671,794,1817,1842]" country="United States of America" name="California" pageId="10" pageNumber="508">California</collectingRegion>
</emphasis>
: Soda
<collectingMunicipality id="485D953709238C53FC958512F3B5F8F6" box="[872,942,1817,1842]" pageId="10" pageNumber="508">Creek</collectingMunicipality>
,
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,
<geoCoordinate id="CDB2698A09238C53FB468512F500F8F6" box="[1211,1307,1817,1842]" direction="north" orientation="latitude" pageId="10" pageNumber="508" precision="925" value="36.15">36°09N</geoCoordinate>
,
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,
<date id="DC38298D09238C53FF6A8537F103F891" box="[151,280,1852,1877]" pageId="10" pageNumber="508" value="1958-07-11">
<collectingDate id="CC7CD06509238C53FF6A8537F103F891" box="[151,280,1852,1877]" pageId="10" pageNumber="508" value="1958-07-11">11.vii.1958</collectingDate>
</date>
,
<collectorName id="05736A9B09238C53FEDA8535F1DEF891" box="[295,453,1852,1877]" pageId="10" pageNumber="508">W. J. Gertsch</collectorName>
&amp; V. Roth leg. (AMNH-158).
<emphasis id="9AF2D35F09238C53FCCB8535F386F891" box="[822,925,1854,1877]" italics="true" pageId="10" pageNumber="508">
<typeStatus id="773DB1EF09238C53FCCB8535F386F891" box="[822,925,1854,1877]" pageId="10" pageNumber="508" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
.
<materialsCitation id="18EE051009238C53FC528536F121F8BE" ID-GBIF-Occurrence="1848253932" collectingDate="1958-07-11" collectorName="W. J. Gertsch" country="United States of America" latitude="36.15" location="west Of NelsOn Camp" longLatPrecision="1189" longitude="-118.75" municipality="Creek" pageId="10" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
<specimenCount id="BE80C4C409238C53FC528536F3C9F892" box="[943,978,1852,1878]" pageId="10" pageNumber="508" type="female">
1
<emphasis id="9AF2D35F09238C53FC408537F3C9F892" box="[957,978,1852,1878]" italics="true" pageId="10" pageNumber="508"></emphasis>
</specimenCount>
, same data, together with the
<typeStatus id="773DB1EF09238C53FAC78537F587F891" box="[1338,1436,1852,1877]" pageId="10" pageNumber="508" type="holotype">holotype</typeStatus>
(AMNH-158).
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</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609238C53FF3A858EF205F805" pageId="10" pageNumber="508" type="etymology">
<paragraph id="A8390F4D09238C53FF3A858EF205F805" blockId="10.[151,1436,1816,2022]" pageId="10" pageNumber="508">
<emphasis id="9AF2D35F09238C53FF3A858EF155F85A" bold="true" box="[199,334,1925,1950]" pageId="10" pageNumber="508">Etymology.</emphasis>
The species name comes from the Latin word semiorbiculatus, in reference to hemispherical apex of the distal part of the scape.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609238C52FF3A85C6F299FF3C" lastPageId="11" lastPageNumber="509" pageId="10" pageNumber="508" type="diagnosis">
<paragraph id="A8390F4D09238C52FF3A85C6F299FF3C" blockId="10.[151,1436,1816,2022]" lastBlockId="11.[151,1436,151,464]" lastPageId="11" lastPageNumber="509" pageId="10" pageNumber="508">
<emphasis id="9AF2D35F09238C53FF3A85C6F15AF822" bold="true" box="[199,321,1997,2022]" pageId="10" pageNumber="508">Diagnosis.</emphasis>
The female
<taxonomicName id="6F8674CE09238C53FE2C85C5F2BFF821" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[465,676,1997,2021]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="508" phylum="Arthropoda" rank="species" species="semiorbiculatus" status="sp. nov.">
<emphasis id="9AF2D35F09238C53FE2C85C5F2BFF821" box="[465,676,1997,2021]" italics="true" pageId="10" pageNumber="508">A. semiorbiculatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F09238C53FD5185C6F2F7F822" bold="true" box="[684,748,1997,2022]" pageId="10" pageNumber="508">
<taxonomicNameLabel id="81C16E2409238C53FD5185C6F2F7F822" box="[684,748,1997,2022]" pageId="10" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from A.
<taxonomicName id="6F8674CE09238C53FBB885C6F4A7F821" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1093,1212,1997,2021]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="508" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F09238C53FBB885C6F4A7F821" box="[1093,1212,1997,2021]" italics="true" pageId="10" pageNumber="508">arcuatulus</emphasis>
</taxonomicName>
and
<taxonomicName id="6F8674CE09238C53FB0885C5F566F821" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1269,1405,1998,2021]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="508" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F09238C53FB0885C5F566F821" box="[1269,1405,1998,2021]" italics="true" pageId="10" pageNumber="508">A. cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F09238C52FA7885C6F0A3FF74" bold="true" lastPageId="11" lastPageNumber="509" pageId="10" pageNumber="508">
<taxonomicNameLabel id="81C16E2409238C52FA7885C6F0A3FF74" lastPageId="11" lastPageNumber="509" pageId="10" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, by the epigynum triangular in ventral view (
<figureCitation id="30BD13C809228C52FD4A829CF314FF74" box="[695,783,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="11" pageNumber="509">Fig. 3G</figureCitation>
) and bulged in lateral view (
<figureCitation id="30BD13C809228C52FBAE829CF4B6FF74" box="[1107,1197,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="11" pageNumber="509">Fig. 3H</figureCitation>
), the U-shaped scape more expanded than the other two species, and by the distal part of scape having the two lateral lobes fused into a hemisphere shape (
<figureCitation id="30BD13C809228C52FE9382D4F1A7FF3C" box="[366,444,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="11" pageNumber="509">
Fig.
<date id="DC38298D09228C52FE5F82EBF1A7FF3C" box="[418,444,224,248]" pageId="11" pageNumber="509">3I</date>
</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609228C52FF3A830FF12DFE69" pageId="11" pageNumber="509" type="description">
<paragraph id="A8390F4D09228C52FF3A830FF452FEA1" blockId="11.[151,1436,151,464]" pageId="11" pageNumber="509">
<emphasis id="9AF2D35F09228C52FF3A830FF142FED9" bold="true" box="[199,345,260,285]" pageId="11" pageNumber="509">Description.</emphasis>
<emphasis id="9AF2D35F09228C52FE9F830FF1ADFED8" box="[354,438,260,284]" italics="true" pageId="11" pageNumber="509">Female</emphasis>
(AMNH-158): Total length 1.59. Carapace 0.76 long, 0.55 wide. Abdomen 0.92 long, 0.63 wide. Leg lengths: I 2.95 (0.82+1.02+0.69+0.42); II 3.01 (0.83+1.03+0.71+0.44); III 2.63 (0.66+0.98+0.61+0.38); IV 3.38 (0.98+1.11+0.85+0.44). Tm I 0.23. Male unknown.
</paragraph>
<paragraph id="A8390F4D09228C52FF3A8364F12DFE69" blockId="11.[151,1436,151,464]" pageId="11" pageNumber="509">
Other somatic and genital characters: see the general description for the
<taxonomicName id="6F8674CE09228C52FC0B837BF498FE4C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1014,1155,368,392]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F09228C52FC0B837BF498FE4C" box="[1014,1155,368,392]" italics="true" pageId="11" pageNumber="509">Arcuphantes</emphasis>
</taxonomicName>
species with
<taxonomicName id="6F8674CE09228C52FAE1837BF588FE4C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1308,1427,368,392]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="509" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F09228C52FAE1837BF588FE4C" box="[1308,1427,368,392]" italics="true" pageId="11" pageNumber="509">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF09228C52FF6A839EF0DCFE69" box="[151,199,405,429]" pageId="11" pageNumber="509">type</typeStatus>
genitalia.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC609228C52FF3A83BCF229FE14" box="[199,562,439,464]" pageId="11" pageNumber="509" type="distribution">
<paragraph id="A8390F4D09228C52FF3A83BCF229FE14" blockId="11.[151,1436,151,464]" box="[199,562,439,464]" pageId="11" pageNumber="509">
<emphasis id="9AF2D35F09228C52FF3A83BCF17BFE14" bold="true" box="[199,352,439,464]" pageId="11" pageNumber="509">Distribution.</emphasis>
<collectingCountry id="D0914FDD09228C52FE9A83B3F184FE14" box="[359,415,440,464]" name="United States of America" pageId="11" pageNumber="509">USA</collectingCountry>
(
<collectingRegion id="6A42C1AF09228C52FE5083BCF23DFE14" box="[429,550,439,464]" country="United States of America" name="California" pageId="11" pageNumber="509">California</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,75 @@
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<emphasis id="9AF2D35F092B8C5BFF6A862BF129FBFE" bold="true" box="[151,306,1056,1082]" italics="true" pageId="2" pageNumber="500">Arcuphantes</emphasis>
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<emphasis id="9AF2D35F092B8C5BFF6A866DF139FBB8" bold="true" box="[151,290,1126,1148]" pageId="2" pageNumber="500">Type species.</emphasis>
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<emphasis id="9AF2D35F092B8C5BFED4866DF1E3FBB8" box="[297,504,1126,1148]" italics="true" pageId="2" pageNumber="500">Arcuphantes fragilis</emphasis>
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<mods:namePart id="455431D1F35359408C8781BFC12D3E14">Ma, Shuangchun</mods:namePart>
</mods:name>
<mods:name id="EE61D91167EA37DF5FA93E350133651E" type="personal">
<mods:role id="A73E744EDD85D46A35F0B6648571B1BE">
<mods:roleTerm id="8CE77E5D510F39BED1AD098B41A8980E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB3B0893A8D0B5EA848641C5E2B95474">Tu, Lihong</mods:namePart>
</mods:name>
<mods:typeOfResource id="2991D602A4ED7661ABA8274DD84A77E3">text</mods:typeOfResource>
<mods:relatedItem id="0B350EF6B1D2652A40DA7EFB637450A5" type="host">
<mods:titleInfo id="877BD823C120063BD4AF136120DB7002">
<mods:title id="0854631A6B350875C565967335C78555">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="118E5FEE206C79E7AD3B9F93E118876B">
<mods:date id="AE1F12261EAB1360D24AC860B230AD25">2018</mods:date>
<mods:detail id="843CAD6434DFDB6AD92630BD112A06D0" type="volume">
<mods:number id="5C0B085E1B7675301AD480D514FDA1A6">4425</mods:number>
</mods:detail>
<mods:detail id="B82D27094E04CBFDC5E1D618DD65FF64" type="issue">
<mods:number id="1A849722DF8B7043619FE685C79E53F0">3</mods:number>
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<mods:start id="806AB449084463813876D522D6BE5373">498</mods:start>
<mods:end id="62368A9A81E2D596BA0638C8C472E1BA">510</mods:end>
</mods:extent>
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<mods:classification id="2DFE13EE17A70ED2B8ED74687241FC1E">journal article</mods:classification>
<mods:identifier id="A816237F942DB46472D33BF7F3461488" type="CLB-Dataset">29989</mods:identifier>
<mods:identifier id="22477D935B461B92CAA5DCC19406A4B7" type="DOI">10.11646/zootaxa.4425.3.4</mods:identifier>
<mods:identifier id="9ACE19160B44C101510894A1CB1A95A1" type="GBIF-Dataset">9956099a-5e4b-4b2e-b24e-d347a707f5f9</mods:identifier>
<mods:identifier id="6FE8FDB081A9F210CC487A972A856682" type="ISSN">1175-5326</mods:identifier>
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<treatment id="202FBE5B092B8C5DFF6A86C7F33CFC44" ID-DOI="http://doi.org/10.5281/zenodo.5998044" ID-GBIF-Taxon="144235594" ID-Zenodo-Dep="5998044" LSID="urn:lsid:plazi:treatment:202FBE5B092B8C5DFF6A86C7F33CFC44" httpUri="http://treatment.plazi.org/id/202FBE5B092B8C5DFF6A86C7F33CFC44" lastPageId="4" lastPageNumber="502" pageId="2" pageNumber="500">
<subSubSection id="E09C5CC6092B8C5BFF6A86C7F18BFA94" pageId="2" pageNumber="500" type="nomenclature">
<paragraph id="A8390F4D092B8C5BFF6A86C7F343FB23" blockId="2.[151,856,1228,1255]" box="[151,856,1228,1255]" pageId="2" pageNumber="500">
<heading id="F371B821092B8C5BFF6A86C7F343FB23" bold="true" box="[151,856,1228,1255]" fontSize="11" level="1" pageId="2" pageNumber="500" reason="1">
<emphasis id="9AF2D35F092B8C5BFF6A86C7F343FB23" bold="true" box="[151,856,1228,1255]" pageId="2" pageNumber="500">
<taxonomicName id="6F8674CE092B8C5BFF6A86C7F129FB22" ID-CoL="32F4" ID-ENA="430923" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[151,306,1228,1254]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092B8C5BFF6A86C7F129FB22" bold="true" box="[151,306,1228,1254]" italics="true" pageId="2" pageNumber="500">Arcuphantes</emphasis>
</taxonomicName>
species with the genitalia of
<taxonomicName id="6F8674CE092B8C5BFD6B86C7F303FB22" box="[662,792,1228,1254]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092B8C5BFD6B86C7F303FB22" bold="true" box="[662,792,1228,1254]" italics="true" pageId="2" pageNumber="500">arcuatulus</emphasis>
</taxonomicName>
-type
</emphasis>
</heading>
</paragraph>
<paragraph id="A8390F4D092B8C5BFF6A8718F18BFA94" blockId="2.[151,1437,1299,2008]" pageId="2" pageNumber="500">
<emphasis id="9AF2D35F092B8C5BFF6A8718F12DFAE8" bold="true" box="[151,310,1299,1324]" pageId="2" pageNumber="500">Composition.</emphasis>
<taxonomicName id="6F8674CE092B8C5BFEBB871FF339FAE8" authority="(ROewer, 1942)" authorityName="Roewer" authorityYear="1942" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[326,802,1300,1324]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092B8C5BFEBB871FF242FAE8" box="[326,601,1300,1324]" italics="true" pageId="2" pageNumber="500">Arcuphantes arcuatulus</emphasis>
(Roewer, 1942)
</taxonomicName>
,
<taxonomicName id="6F8674CE092B8C5BFCCB871FF3DDFAEF" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[822,966,1300,1323]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092B8C5BFCCB871FF3DDFAEF" box="[822,966,1300,1323]" italics="true" pageId="2" pageNumber="500">A. cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092B8C5BFC2B8718F406FAE8" bold="true" box="[982,1053,1299,1324]" pageId="2" pageNumber="500">
<taxonomicNameLabel id="81C16E24092B8C5BFC2B8718F406FAE8" box="[982,1053,1299,1324]" pageId="2" pageNumber="500" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="6F8674CE092B8C5BFBC9871FF4FEFAE8" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1076,1253,1300,1324]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="9AF2D35F092B8C5BFBC9871FF4FEFAE8" box="[1076,1253,1300,1324]" italics="true" pageId="2" pageNumber="500">A. denticulatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092B8C5BFB098718F527FAE8" bold="true" box="[1268,1340,1299,1324]" pageId="2" pageNumber="500">
<taxonomicNameLabel id="81C16E24092B8C5BFB098718F527FAE8" box="[1268,1340,1299,1324]" pageId="2" pageNumber="500" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="6F8674CE092B8C5BFA79871FF151FA94" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="semiorbiculatus" status="sp. nov.">
<emphasis id="9AF2D35F092B8C5BFA79871FF151FA94" italics="true" pageId="2" pageNumber="500">A. semiorbiculatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092B8C5BFEAC873CF18BFA94" bold="true" box="[337,400,1335,1360]" pageId="2" pageNumber="500">
<taxonomicNameLabel id="81C16E24092B8C5BFEAC873CF18BFA94" box="[337,400,1335,1360]" pageId="2" pageNumber="500" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092B8C5BFF3A8750F18CF918" pageId="2" pageNumber="500" type="diagnosis">
<paragraph id="A8390F4D092B8C5BFF3A8750F0EAFA24" blockId="2.[151,1437,1299,2008]" pageId="2" pageNumber="500">
<emphasis id="9AF2D35F092B8C5BFF3A8750F3E0FAB7" bold="true" box="[199,1019,1371,1396]" pageId="2" pageNumber="500">
Diagnosis of the
<taxonomicName id="6F8674CE092B8C5BFE6E8750F23CFAB0" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[403,551,1371,1396]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092B8C5BFE6E8750F23CFAB0" bold="true" box="[403,551,1371,1396]" italics="true" pageId="2" pageNumber="500">Arcuphantes</emphasis>
</taxonomicName>
species with
<taxonomicName id="6F8674CE092B8C5BFD378750F35DFAB0" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[714,838,1371,1396]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092B8C5BFD378750F35DFAB0" bold="true" box="[714,838,1371,1396]" italics="true" pageId="2" pageNumber="500">arcuatulus</emphasis>
</taxonomicName>
-type genitalia.
</emphasis>
The male palp can be distinguished from that of other
<taxonomicName id="6F8674CE092B8C5BFE90878BF1E1FA5C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[365,506,1408,1432]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092B8C5BFE90878BF1E1FA5C" box="[365,506,1408,1432]" italics="true" pageId="2" pageNumber="500">Arcuphantes</emphasis>
</taxonomicName>
species by a narrow and posteriorly prolonged radix, and an unbranched lamella characteristica with a broom-shaped and strongly sclerotized apex (
<figureCitation id="30BD13C8092B8C5BFC6E87A8F3FFFA78" box="[915,996,1443,1468]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="2" pageNumber="500">
Fig.
<date id="DC38298D092B8C5BFC3487AFF3FFFA78" box="[969,996,1444,1468]" pageId="2" pageNumber="500">2I</date>
</figureCitation>
); the epigynum with a U-shaped scape (
<figureCitation id="30BD13C8092B8C5BFF6287CCF0FFFA24" box="[159,228,1479,1504]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="2" pageNumber="500">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="A8390F4D092B8C5BFF3A87E0F18CF918" blockId="2.[151,1437,1299,2008]" pageId="2" pageNumber="500">
<emphasis id="9AF2D35F092B8C5BFF3A87E0F46CF9C0" bold="true" box="[199,1143,1515,1540]" pageId="2" pageNumber="500">
General description for the
<taxonomicName id="6F8674CE092B8C5BFDE487E0F2B6F9C0" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[537,685,1515,1540]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092B8C5BFDE487E0F2B6F9C0" bold="true" box="[537,685,1515,1540]" italics="true" pageId="2" pageNumber="500">Arcuphantes</emphasis>
</taxonomicName>
species with
<taxonomicName id="6F8674CE092B8C5BFCB387E0F3D1F9C0" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[846,970,1515,1540]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092B8C5BFCB387E0F3D1F9C0" bold="true" box="[846,970,1515,1540]" italics="true" pageId="2" pageNumber="500">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092B8C5BFC2F87E0F41CF9C0" box="[978,1031,1515,1540]" pageId="2" pageNumber="500">type</typeStatus>
genitalia
</emphasis>
(
<figureCitation id="30BD13C8092B8C5BFB7587E0F4D7F9C0" box="[1160,1228,1515,1540]" captionStart="FIGURE 1" captionStartId="3.[151,250,1651,1673]" captionTargetBox="[271,1312,584,1633]" captionTargetId="figure@3.[271,1312,584,1637]" captionTargetPageId="3" captionText="FIGURE 1. Arcuphantes species with the genitalia of arcuatulus-type. A, A. arcuatulus, male, dorsal; B, A. arcuatulus, female, dorsal; C, A. cruciatus n. sp., female, dorsal; D, A. denticulatus n. sp., male, dorsal; E, A. semiorbiculatus n. sp., female, dorsal; F, A. semiorbiculatus n. sp., female, lateral. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270387/files/figure.png" pageId="2" pageNumber="500">Fig. 1</figureCitation>
): medium in size, total length
<quantity id="6F7EA2A8092B8C5BFED08404F1DAF9EC" box="[301,449,1551,1576]" metricMagnitude="-2" metricUnit="m" metricValue="6.603999999999999" metricValueMax="7.747" metricValueMin="5.460999999999999" pageId="2" pageNumber="500" unit="in" value="2.5999999999999996" valueMax="3.05" valueMin="2.15">2.153.05 in</quantity>
male,
<quantity id="6F7EA2A8092B8C5BFDE08404F2AAF9EC" box="[541,689,1551,1576]" metricMagnitude="-2" metricUnit="m" metricValue="4.7498000000000005" metricValueMax="5.460999999999999" metricValueMin="4.038600000000001" pageId="2" pageNumber="500" unit="in" value="1.87" valueMax="2.15" valueMin="1.59">1.592.15 in</quantity>
female. Carapace yellowish brown to dark brown. Each eye surrounded by a black ring. Anterior eye row recurved in dorsal view, posterior eye row straight. Anterior median eyes subcontiguous, lateral eyes adjacent. Abdomen dark in color, with black and white spots on the dorsal surface. Legs long and slender, with black rings. Chaetotaxy: Ti I-IV: 2-2-2-2; Mt I-IV: 1-1-1-1; Tm I about 0.190.23, Tm IV absent. Tracheal system haplotracheate; epiandrous gland spigots present in male
<taxonomicName id="6F8674CE092B8C5BFB8B84ABF51AF973" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1142,1281,1696,1719]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092B8C5BFB8B84ABF49CF973" box="[1142,1159,1696,1719]" italics="true" pageId="2" pageNumber="500">A</emphasis>
.
<emphasis id="9AF2D35F092B8C5BFB6484ABF51AF973" box="[1177,1281,1696,1719]" italics="true" pageId="2" pageNumber="500">cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092B8C5BFAF18494F555F97C" bold="true" box="[1292,1358,1695,1720]" pageId="2" pageNumber="500">
<taxonomicNameLabel id="81C16E24092B8C5BFAF18494F555F97C" box="[1292,1358,1695,1720]" pageId="2" pageNumber="500" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(other species not examined).
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092B8C5AFF3A84E3F3B0FDD3" lastPageId="3" lastPageNumber="501" pageId="2" pageNumber="500" type="description">
<paragraph id="A8390F4D092B8C5AFF3A84E3F549FF10" blockId="2.[151,1437,1299,2008]" lastBlockId="3.[151,1437,151,536]" lastPageId="3" lastPageNumber="501" pageId="2" pageNumber="500">
<emphasis id="9AF2D35F092B8C5BFF3A84E3F121F8C4" box="[199,314,1768,1792]" italics="true" pageId="2" pageNumber="500">Male palp</emphasis>
(
<figureCitation id="30BD13C8092B8C5BFEB784ECF194F8C4" box="[330,399,1767,1792]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="2" pageNumber="500">Fig. 2</figureCitation>
). Both patella and tibia with one macroseta (
<figureCitation id="30BD13C8092B8C5BFC7184ECF3FCF8C4" box="[908,999,1767,1792]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="2" pageNumber="500">Fig. 2G</figureCitation>
). Cymbial retrobasal process wide and blunt (
<figureCitation id="30BD13C8092B8C5BFF188500F159F8E0" box="[229,322,1803,1828]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="2" pageNumber="500">Fig. 2B</figureCitation>
). Paracymbium narrow and somewhat cross-shaped, with distal arm elongated and rolling up, middle and outer arms arising from anterior and posterior margins, respectively, connected by a transverse ridge (
<figureCitation id="30BD13C8092B8C5BFF628558F106F8A8" box="[159,285,1875,1900]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="2" pageNumber="500">Fig. 2GH</figureCitation>
). Distal suprategular apophysis stout, pit hook remnant as a small apophysis pointed forward (
<figureCitation id="30BD13C8092B8C5BFACB8558F58BF8A8" box="[1334,1424,1875,1900]" captionStart="FIGURE 4" captionStartId="7.[151,250,1960,1982]" captionTargetBox="[151,1436,840,1908]" captionTargetId="figure@7.[151,1436,840,1911]" captionTargetPageId="7" captionText="FIGURE 4. Arcuphantes arcuatulus. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE. embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270393/files/figure.png" pageId="2" pageNumber="500">Fig. 4A</figureCitation>
). Embolic division (
<figureCitation id="30BD13C8092B8C5BFE94857CF1A2F854" box="[361,441,1911,1936]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="2" pageNumber="500">
Fig.
<date id="DC38298D092B8C5BFE638573F1A2F854" box="[414,441,1912,1936]" pageId="2" pageNumber="500">2I</date>
</figureCitation>
): radix narrow, posteriorly prolonged with an anterior process and central serrated area, Fickert's gland located anterior part of radix. Embolic membrane with numerous papillae and chitinous basal part. Lamella characteristica about half of the radix in length, unbranched, broom-shaped apex with numerous strong sclerotized teeth. Terminal apophysis as a large sclerite with several processes, covering over embolus. Embolus trunk-like with serrated surface, a large thumb and a pointed apex at each side of embolus proper (
<figureCitation id="30BD13C8092A8C5AFB1682B7F55EFF10" box="[1259,1349,188,213]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="3" pageNumber="501">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="A8390F4D092A8C5AFF3A82EBF3B0FDD3" blockId="3.[151,1437,151,536]" pageId="3" pageNumber="501">
<emphasis id="9AF2D35F092A8C5AFF3A82EBF122FF33" box="[199,313,224,247]" italics="true" pageId="3" pageNumber="501">Epigynum</emphasis>
(
<figureCitation id="30BD13C8092A8C5AFEB682D4F188FF3C" box="[331,403,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="3" pageNumber="501">Fig. 3</figureCitation>
). Protruding out, posteriorly prolonged (
<figureCitation id="30BD13C8092A8C5AFC9682D4F3D8FF3C" box="[875,963,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1651,1673]" captionTargetBox="[271,1312,584,1633]" captionTargetId="figure@3.[271,1312,584,1637]" captionTargetPageId="3" captionText="FIGURE 1. Arcuphantes species with the genitalia of arcuatulus-type. A, A. arcuatulus, male, dorsal; B, A. arcuatulus, female, dorsal; C, A. cruciatus n. sp., female, dorsal; D, A. denticulatus n. sp., male, dorsal; E, A. semiorbiculatus n. sp., female, dorsal; F, A. semiorbiculatus n. sp., female, lateral. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270387/files/figure.png" pageId="3" pageNumber="501">Fig. 1F</figureCitation>
). Epigynal plate in protruding state with paired groove slits extending on dorsal surface (
<figureCitation id="30BD13C8092A8C5AFD47830FF30AFED9" box="[698,785,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1905,1927]" captionTargetBox="[171,1415,193,1871]" captionTargetId="figure@8.[171,1415,193,1878]" captionTargetPageId="8" captionText="FIGURE 5. Arcuphantes cruciatus n. sp. AB, male palp, with embolic division removed, A, prolateral; B, retrolateral; CD, embolic division, arrows point to central serrated area of radix, C, ventral; D, dorsal; EH, epigynum, with extensible basal part removed, E, ventral, ventral plate artificially broken; F, dorsal; G, caudal; H, inner, shows epigynal grooves. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270395/files/figure.png" pageId="3" pageNumber="501">Fig. 5F</figureCitation>
). Integument of basal part wrinkled making it extensible. Median plate on dorsal surface about three times longer than wide, and then the posterior part of epigynum becoming gradually narrowed, passing into a scape, without epigynal cavity. Spermathecae located at posterior half part of median plate. Two chambers with special glands at boundary (
<figureCitation id="30BD13C8092A8C5AFC288364F428FE4C" box="[981,1075,367,392]" captionStart="FIGURE 5" captionStartId="8.[151,250,1905,1927]" captionTargetBox="[171,1415,193,1871]" captionTargetId="figure@8.[171,1415,193,1878]" captionTargetPageId="8" captionText="FIGURE 5. Arcuphantes cruciatus n. sp. AB, male palp, with embolic division removed, A, prolateral; B, retrolateral; CD, embolic division, arrows point to central serrated area of radix, C, ventral; D, dorsal; EH, epigynum, with extensible basal part removed, E, ventral, ventral plate artificially broken; F, dorsal; G, caudal; H, inner, shows epigynal grooves. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270395/files/figure.png" pageId="3" pageNumber="501">Fig. 5H</figureCitation>
), basal part spiraling with one chamber turning upwards and one downwards. Fertilization groove extending forwards along lateral margins of median plate, ending outside epigastric furrow. Scape in U-shaped, with copulatory grooves extending along its dorsal surface, without turning point. Distal part of scape with a pair of lateral lobes, each furnished with a pocket, in which hosting copulatory opening. Neither stretcher nor pit present.
</paragraph>
</subSubSection>
<caption id="FCF95FC5092A8C5AFF6A8478F34BF902" httpUri="https://zenodo.org/record/1270387/files/figure.png" pageId="3" pageNumber="501" startId="3.[151,250,1651,1673]" targetBox="[271,1312,584,1633]" targetPageId="3">
<paragraph id="A8390F4D092A8C5AFF6A8478F34BF902" blockId="3.[151,1436,1651,1734]" pageId="3" pageNumber="501">
<emphasis id="9AF2D35F092A8C5AFF6A8478F109F94D" bold="true" box="[151,274,1651,1673]" pageId="3" pageNumber="501">FIGURE 1.</emphasis>
<taxonomicName id="6F8674CE092A8C5AFEE58478F18CF94D" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[280,407,1651,1673]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092A8C5AFEE58478F18CF94D" box="[280,407,1651,1673]" italics="true" pageId="3" pageNumber="501">Arcuphantes</emphasis>
</taxonomicName>
species with the genitalia of
<taxonomicName id="6F8674CE092A8C5AFD468478F33DF94D" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[699,806,1651,1673]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFD468478F33DF94D" box="[699,806,1651,1673]" italics="true" pageId="3" pageNumber="501">arcuatulus</emphasis>
</taxonomicName>
-type.
<emphasis id="9AF2D35F092A8C5AFC9A8478F362F94D" bold="true" box="[871,889,1651,1673]" pageId="3" pageNumber="501">A</emphasis>
,
<taxonomicName id="6F8674CE092A8C5AFC78847FF417F94D" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[901,1036,1651,1673]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFC78847FF417F94D" box="[901,1036,1651,1673]" italics="true" pageId="3" pageNumber="501">A. arcuatulus</emphasis>
</taxonomicName>
, male, dorsal;
<emphasis id="9AF2D35F092A8C5AFB5D8478F4AAF94D" bold="true" box="[1184,1201,1651,1673]" pageId="3" pageNumber="501">B</emphasis>
,
<taxonomicName id="6F8674CE092A8C5AFB40847FF55FF94D" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[1213,1348,1651,1673]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFB40847FF55FF94D" box="[1213,1348,1651,1673]" italics="true" pageId="3" pageNumber="501">A. arcuatulus</emphasis>
</taxonomicName>
, female, dorsal;
<emphasis id="9AF2D35F092A8C5AFF188499F0ECF96C" bold="true" box="[229,247,1682,1704]" pageId="3" pageNumber="501">C</emphasis>
,
<taxonomicName id="6F8674CE092A8C5AFEFA8499F19FF96C" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[263,388,1682,1704]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092A8C5AFEFA8499F19FF96C" box="[263,388,1682,1704]" italics="true" pageId="3" pageNumber="501">A. cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092A8C5AFE708499F1D2F96C" bold="true" box="[397,457,1682,1704]" pageId="3" pageNumber="501">
<taxonomicNameLabel id="81C16E24092A8C5AFE708499F1D2F96C" box="[397,457,1682,1704]" pageId="3" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, dorsal;
<emphasis id="9AF2D35F092A8C5AFD868499F296F96C" bold="true" box="[635,653,1682,1704]" pageId="3" pageNumber="501">D</emphasis>
,
<taxonomicName id="6F8674CE092A8C5AFD608499F323F96C" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[669,824,1682,1704]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="9AF2D35F092A8C5AFD608499F323F96C" box="[669,824,1682,1704]" italics="true" pageId="3" pageNumber="501">A. denticulatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092A8C5AFCBD8499F366F96C" bold="true" box="[832,893,1682,1704]" pageId="3" pageNumber="501">
<taxonomicNameLabel id="81C16E24092A8C5AFCBD8499F366F96C" box="[832,893,1682,1704]" pageId="3" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, dorsal;
<emphasis id="9AF2D35F092A8C5AFBE68499F437F96C" bold="true" box="[1051,1068,1682,1704]" pageId="3" pageNumber="501">E</emphasis>
,
<taxonomicName id="6F8674CE092A8C5AFBC68499F4E7F96C" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1083,1276,1682,1704]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="semiorbiculatus" status="sp. nov.">
<emphasis id="9AF2D35F092A8C5AFBC68499F4E7F96C" box="[1083,1276,1682,1704]" italics="true" pageId="3" pageNumber="501">A. semiorbiculatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092A8C5AFAF88499F55AF96C" bold="true" box="[1285,1345,1682,1704]" pageId="3" pageNumber="501">
<taxonomicNameLabel id="81C16E24092A8C5AFAF88499F55AF96C" box="[1285,1345,1682,1704]" pageId="3" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, dorsal;
<emphasis id="9AF2D35F092A8C5AFF1F84BBF0F4F902" bold="true" box="[226,239,1712,1734]" pageId="3" pageNumber="501">F</emphasis>
,
<taxonomicName id="6F8674CE092A8C5AFF0184BAF1A1F902" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[252,442,1712,1734]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="semiorbiculatus" status="sp. nov.">
<emphasis id="9AF2D35F092A8C5AFF0184BAF1A1F902" box="[252,442,1712,1734]" italics="true" pageId="3" pageNumber="501">A. semiorbiculatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092A8C5AFE3D84BBF1E2F902" bold="true" box="[448,505,1712,1734]" pageId="3" pageNumber="501">
<taxonomicNameLabel id="81C16E24092A8C5AFE3D84BBF1E2F902" box="[448,505,1712,1734]" pageId="3" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, lateral. [Scale bars, mm]
</paragraph>
</caption>
<subSubSection id="E09C5CC6092A8C5DFF3A84F8F134FC99" lastPageId="4" lastPageNumber="502" pageId="3" pageNumber="501" type="discussion">
<paragraph id="A8390F4D092A8C5DFF3A84F8F138FDF9" blockId="3.[151,1436,1779,2020]" lastBlockId="4.[151,1437,151,896]" lastPageId="4" lastPageNumber="502" pageId="3" pageNumber="501">
<emphasis id="9AF2D35F092A8C5AFF3A84F8F120F8C8" bold="true" box="[199,315,1779,1804]" pageId="3" pageNumber="501">Remarks.</emphasis>
As stated by
<bibRefCitation id="CC1772BC092A8C5AFE2384FFF284F8C8" box="[478,671,1779,1804]" pageId="3" pageNumber="501" refId="ref4170" refString="Ma, S. C., Marusik, Y. M. &amp; Tu, L. H. (2016) A review of the Nearctic linyphiid spider Arcuphantes fragilis (Araneae, Linyphiidae) and closely related species. Zootaxa, 4144 (3), 383 - 396. https: // doi. org / 10.11646 / zootaxa. 4144.3.6" type="journal article">
Ma
<emphasis id="9AF2D35F092A8C5AFDF084FFF25AF8CF" box="[525,577,1779,1803]" italics="true" pageId="3" pageNumber="501">et al</emphasis>
. (2016)
</bibRefCitation>
, genitalia of the Nearctic
<taxonomicName id="6F8674CE092A8C5AFC3084F8F441F8CF" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[973,1114,1779,1803]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092A8C5AFC3084F8F441F8CF" box="[973,1114,1779,1803]" italics="true" pageId="3" pageNumber="501">Arcuphantes</emphasis>
</taxonomicName>
species can be divided into three
<typeStatus id="773DB1EF092A8C5AFF2B8513F10EF8F4" box="[214,277,1816,1840]" pageId="3" pageNumber="501">types</typeStatus>
:
<taxonomicName id="6F8674CE092A8C5AFEDD851CF16AF8EB" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[288,369,1815,1839]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092A8C5AFEDD851CF16AF8EB" box="[288,369,1815,1839]" italics="true" pageId="3" pageNumber="501">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFE858513F1B6F8F4" box="[376,429,1816,1840]" pageId="3" pageNumber="501">type</typeStatus>
,
<taxonomicName id="6F8674CE092A8C5AFE45851CF234F8EB" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[440,559,1815,1839]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFE45851CF234F8EB" box="[440,559,1815,1839]" italics="true" pageId="3" pageNumber="501">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFDCA8513F277F8F4" box="[567,620,1816,1840]" pageId="3" pageNumber="501">type</typeStatus>
, and
<taxonomicName id="6F8674CE092A8C5AFD5B851CF2E9F8EB" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[678,754,1815,1839]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="pictilis">
<emphasis id="9AF2D35F092A8C5AFD5B851CF2E9F8EB" box="[678,754,1815,1839]" italics="true" pageId="3" pageNumber="501">pictilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFD078513F334F8F4" box="[762,815,1816,1840]" pageId="3" pageNumber="501">type</typeStatus>
. The epigyna of the three
<typeStatus id="773DB1EF092A8C5AFBAE8513F495F8F4" box="[1107,1166,1816,1840]" pageId="3" pageNumber="501">types</typeStatus>
are easily distinguished by the shape of the scape: U-shaped in
<taxonomicName id="6F8674CE092A8C5AFDB18530F2D8F897" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[588,707,1851,1875]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFDB18530F2D8F897" box="[588,707,1851,1875]" italics="true" pageId="3" pageNumber="501">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFD378537F2E7F890" box="[714,764,1852,1876]" pageId="3" pageNumber="501">type</typeStatus>
(
<figureCitation id="30BD13C8092A8C5AFCF68530F355F890" box="[779,846,1851,1876]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="3" pageNumber="501">Fig. 3</figureCitation>
), straight in
<taxonomicName id="6F8674CE092A8C5AFC218530F436F897" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[988,1069,1851,1875]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092A8C5AFC218530F436F897" box="[988,1069,1851,1875]" italics="true" pageId="3" pageNumber="501">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFBC88537F47CF890" box="[1077,1127,1852,1876]" pageId="3" pageNumber="501">type</typeStatus>
(
<bibRefCitation id="CC1772BC092A8C5AFB888537F504F890" box="[1141,1311,1851,1876]" pageId="3" pageNumber="501" refId="ref4170" refString="Ma, S. C., Marusik, Y. M. &amp; Tu, L. H. (2016) A review of the Nearctic linyphiid spider Arcuphantes fragilis (Araneae, Linyphiidae) and closely related species. Zootaxa, 4144 (3), 383 - 396. https: // doi. org / 10.11646 / zootaxa. 4144.3.6" type="journal article">
Ma
<emphasis id="9AF2D35F092A8C5AFB5E8537F4C8F897" box="[1187,1235,1851,1875]" italics="true" pageId="3" pageNumber="501">et al</emphasis>
. 2016
</bibRefCitation>
: fig. 4) and S-shaped in
<taxonomicName id="6F8674CE092A8C5AFED68554F16CF8B3" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[299,375,1887,1911]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="pictilis">
<emphasis id="9AF2D35F092A8C5AFED68554F16CF8B3" box="[299,375,1887,1911]" italics="true" pageId="3" pageNumber="501">pictilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFE83856BF1ABF8BC" box="[382,432,1888,1912]" pageId="3" pageNumber="501">type</typeStatus>
(
<bibRefCitation id="CC1772BC092A8C5AFE398554F357F8BC" box="[452,844,1887,1912]" pageId="3" pageNumber="501" refId="ref4098" refString="Chamberlin, R. V. &amp; Ivie, W. (1943) New genera and species of North American linyphiid spiders. Bulletin of the University of Utah, 10, 1 - 39." type="journal article">Chamberlin &amp; Ivie 1943: fig. 36</bibRefCitation>
). Furthermore, the epigyna of
<taxonomicName id="6F8674CE092A8C5AFB448554F511F8B3" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1209,1290,1887,1911]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092A8C5AFB448554F511F8B3" box="[1209,1290,1887,1911]" italics="true" pageId="3" pageNumber="501">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFAEC856BF558F8BC" box="[1297,1347,1888,1912]" pageId="3" pageNumber="501">type</typeStatus>
have a triangular epigynal cavity, which is absent in other two
<typeStatus id="773DB1EF092A8C5AFCE0858FF347F858" box="[797,860,1924,1948]" pageId="3" pageNumber="501">types</typeStatus>
. The male palps of
<taxonomicName id="6F8674CE092A8C5AFBB98588F4A0F85F" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[1092,1211,1923,1947]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFBB98588F4A0F85F" box="[1092,1211,1923,1947]" italics="true" pageId="3" pageNumber="501">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFB3F858FF4EFF858" box="[1218,1268,1924,1948]" pageId="3" pageNumber="501">type</typeStatus>
(
<figureCitation id="30BD13C8092A8C5AFAFB8588F557F858" box="[1286,1356,1923,1948]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="3" pageNumber="501">Fig. 2</figureCitation>
) differ from those of
<taxonomicName id="6F8674CE092A8C5AFEC385ACF194F87B" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[318,399,1959,1983]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092A8C5AFEC385ACF194F87B" box="[318,399,1959,1983]" italics="true" pageId="3" pageNumber="501">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFE6A85A3F1D2F804" box="[407,457,1960,1984]" pageId="3" pageNumber="501">type</typeStatus>
(
<bibRefCitation id="CC1772BC092A8C5AFE2785A3F296F804" box="[474,653,1959,1984]" pageId="3" pageNumber="501" refId="ref4170" refString="Ma, S. C., Marusik, Y. M. &amp; Tu, L. H. (2016) A review of the Nearctic linyphiid spider Arcuphantes fragilis (Araneae, Linyphiidae) and closely related species. Zootaxa, 4144 (3), 383 - 396. https: // doi. org / 10.11646 / zootaxa. 4144.3.6" type="journal article">
Ma
<emphasis id="9AF2D35F092A8C5AFDF185A3F25BF87B" box="[524,576,1959,1983]" italics="true" pageId="3" pageNumber="501">et al</emphasis>
. 2016
</bibRefCitation>
: fig. 1; the male of
<taxonomicName id="6F8674CE092A8C5AFC8785A3F3F3F87B" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[890,1000,1959,1983]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="pictilis">
<emphasis id="9AF2D35F092A8C5AFC8785A3F390F87B" box="[890,907,1960,1983]" italics="true" pageId="3" pageNumber="501">A</emphasis>
.
<emphasis id="9AF2D35F092A8C5AFC6185ACF3F3F87B" box="[924,1000,1959,1983]" italics="true" pageId="3" pageNumber="501">pictilis</emphasis>
</taxonomicName>
remains unknown) as follows: 1) the cymbial retrobasal process in the
<taxonomicName id="6F8674CE092A8C5AFDF185C0F298F827" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[524,643,1995,2019]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092A8C5AFDF185C0F298F827" box="[524,643,1995,2019]" italics="true" pageId="3" pageNumber="501">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092A8C5AFD7785C7F2A7F820" box="[650,700,1996,2020]" pageId="3" pageNumber="501">type</typeStatus>
is wide and blunt, while it is a thumb-like apophysis in that of the
<taxonomicName id="6F8674CE092D8C5DFF6A8293F0F3FF74" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[151,232,152,176]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092D8C5DFF6A8293F0F3FF74" box="[151,232,152,176]" italics="true" pageId="4" pageNumber="502">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFF0D8293F13EFF74" box="[240,293,152,176]" pageId="4" pageNumber="502">type</typeStatus>
; 2) the paracymbium in the
<taxonomicName id="6F8674CE092D8C5DFD9C8293F2C3FF74" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[609,728,152,176]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFD9C8293F2C3FF74" box="[609,728,152,176]" italics="true" pageId="4" pageNumber="502">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFD228293F30AFF74" box="[735,785,152,176]" pageId="4" pageNumber="502">type</typeStatus>
is narrow and elongated, with a middle and an outer arm, whereas that of the
<taxonomicName id="6F8674CE092D8C5DFE8D82B7F1DAFF10" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[368,449,188,212]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092D8C5DFE8D82B7F1DAFF10" box="[368,449,188,212]" italics="true" pageId="4" pageNumber="502">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFE3482B6F1E0FF11" box="[457,507,189,213]" pageId="4" pageNumber="502">type</typeStatus>
is wide with two distal apophyses; 3) the radix in the
<taxonomicName id="6F8674CE092D8C5DFBAE82B7F4D1FF10" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[1107,1226,188,212]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFBAE82B7F4D1FF10" box="[1107,1226,188,212]" italics="true" pageId="4" pageNumber="502">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFB2C82B6F518FF11" box="[1233,1283,189,213]" pageId="4" pageNumber="502">type</typeStatus>
is posteriorly prolonged with a central serrated area, and Fickert's gland is located anteriorly, while in the
<taxonomicName id="6F8674CE092D8C5DFB4C82EBF519FF3C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1201,1282,224,248]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092D8C5DFB4C82EBF519FF3C" box="[1201,1282,224,248]" italics="true" pageId="4" pageNumber="502">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFAF782EBF527FF3C" box="[1290,1340,224,248]" pageId="4" pageNumber="502">type</typeStatus>
it keeps boat-shaped as that in many other micronetine genera (
<bibRefCitation id="CC1772BC092D8C5DFD01830FF45FFED9" box="[764,1092,260,285]" pageId="4" pageNumber="502" refId="ref4300" refString="Saaristo, M. I. &amp; Tanasevitch, A. V. (1996) Redelimitation of the Subfamily Micronetinae Hull, 1920 and the Genus Lepthyphantes Menge, 1866 with Descriptions of Some New Genera (Aranei, Linyphiidae). Berichte naturw. - mediz. Verein Innsbruck, 83, 163 - 186." type="journal article">Saaristo &amp; Tanasevitch 1996</bibRefCitation>
), without central serrated area, and with Fickert's gland located posteriorly; 4) the apex of lamella characteristica in the
<taxonomicName id="6F8674CE092D8C5DFB858323F4F4FE84" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[1144,1263,296,320]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFB858323F4F4FE84" box="[1144,1263,296,320]" italics="true" pageId="4" pageNumber="502">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFB0B8323F533FE84" box="[1270,1320,296,320]" pageId="4" pageNumber="502">type</typeStatus>
is broomshaped with strong sclerotized teeth, while in the
<taxonomicName id="6F8674CE092D8C5DFD3E8347F30FFEA0" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[707,788,332,356]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092D8C5DFD3E8347F30FFEA0" box="[707,788,332,356]" italics="true" pageId="4" pageNumber="502">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFCE68346F356FEA1" box="[795,845,333,357]" pageId="4" pageNumber="502">type</typeStatus>
it is wide and stout with numerous fine denticles; 5) the terminal apophysis in the
<taxonomicName id="6F8674CE092D8C5DFE1C837BF243FE4C" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[481,600,368,392]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFE1C837BF243FE4C" box="[481,600,368,392]" italics="true" pageId="4" pageNumber="502">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFDA2837BF28AFE4C" box="[607,657,368,392]" pageId="4" pageNumber="502">type</typeStatus>
is a large sclerite with several processes, while that of
<taxonomicName id="6F8674CE092D8C5DFB05837BF552FE4C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1272,1353,368,392]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092D8C5DFB05837BF552FE4C" box="[1272,1353,368,392]" italics="true" pageId="4" pageNumber="502">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFAAD837BF599FE4C" box="[1360,1410,368,392]" pageId="4" pageNumber="502">type</typeStatus>
is divided into two parts: a tooth-like anterior terminal apophysis and a waved membranous posterior terminal apophysis; 6) the embolus in both
<typeStatus id="773DB1EF092D8C5DFDCD83B3F270FE14" box="[560,619,440,464]" pageId="4" pageNumber="502">types</typeStatus>
are trunk-like, while in the
<taxonomicName id="6F8674CE092D8C5DFC4683B3F417FE14" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[955,1036,440,464]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="9AF2D35F092D8C5DFC4683B3F417FE14" box="[955,1036,440,464]" italics="true" pageId="4" pageNumber="502">fragilis</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFBEE83B3F45EFE14" box="[1043,1093,440,464]" pageId="4" pageNumber="502">type</typeStatus>
it is S-shaped with a middle depression. Accordingly, each genital
<typeStatus id="773DB1EF092D8C5DFDB983D6F26FFE31" box="[580,628,477,501]" pageId="4" pageNumber="502">type</typeStatus>
consists of a complex of genital characters, and large gaps exist among different genital
<typeStatus id="773DB1EF092D8C5DFEA4800BF18CFDDC" box="[345,407,512,536]" pageId="4" pageNumber="502">types</typeStatus>
. The relationships among the species having different genital
<typeStatus id="773DB1EF092D8C5DFB92800BF4B1FDDC" box="[1135,1194,512,536]" pageId="4" pageNumber="502">types</typeStatus>
need to be tested in future study.
</paragraph>
<paragraph id="A8390F4D092D8C5DFF3A8043F134FC99" blockId="4.[151,1437,151,896]" pageId="4" pageNumber="502">
Four species recognized here having the genitalia of
<taxonomicName id="6F8674CE092D8C5DFCDD8043F38CFDA4" baseAuthorityName="Roewer" baseAuthorityYear="1942" box="[800,919,584,608]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFCDD8043F38CFDA4" box="[800,919,584,608]" italics="true" pageId="4" pageNumber="502">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092D8C5DFC628043F3CFFDA4" box="[927,980,584,608]" pageId="4" pageNumber="502">type</typeStatus>
; one of them is based only on two male specimens and one on two females. Interspecific variation is observed mainly in the characters of the palpal paracymbial apophysis (
<figureCitation id="30BD13C8092D8C5DFE558084F1F6FD6C" box="[424,493,655,680]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="4" pageNumber="502">Fig. 7</figureCitation>
), the central serrated area of the radix (
<figureCitation id="30BD13C8092D8C5DFC558084F40CFD6C" box="[936,1047,655,680]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="4" pageNumber="502">Fig. 2D, I</figureCitation>
) and the terminal apophysis in the male (
<figureCitation id="30BD13C8092D8C5DFF2180BFF13BFD08" box="[220,288,692,717]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="4" pageNumber="502">Fig. 7</figureCitation>
), and the shape of the epigynal scape in the female (
<figureCitation id="30BD13C8092D8C5DFC9B80BFF3E5FD08" box="[870,1022,692,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="4" pageNumber="502">Fig. 3B, E, H</figureCitation>
). The rigorous matching between the male and female genitalia plays an important role in the copulation success in linyphiids (
<bibRefCitation id="CC1772BC092D8C5DFB7080DCF54AFD34" box="[1165,1361,727,752]" pageId="4" pageNumber="502" refId="ref4709" refString="van Helsdingen, P. J. (1965) Sexual behavior of Lepthyphantes leprosus (Ohlert) (Araneida, Linyphiidae), with notes on the function of the genital organs. Zoologische Mededelingen, 41, 15 - 42." type="journal article">Helsdingen 1965</bibRefCitation>
,
<bibRefCitation id="CC1772BC092D8C5DFAA180DCF58CFD34" box="[1372,1431,727,752]" pageId="4" pageNumber="502" refId="ref4752" refString="van Helsdingen, P. J. (1969) A reclassification of the species of Linyphia Latreille based on the functioning of the genitalia (Araneida, Linyphiidae), I. Zoologische Verhandelingen, 105, 1 - 303." type="journal article">1969</bibRefCitation>
;
<bibRefCitation id="CC1772BC092D8C5DFF6A80F6F15DFCD1" box="[151,326,764,789]" pageId="4" pageNumber="502" refId="ref4170" refString="Ma, S. C., Marusik, Y. M. &amp; Tu, L. H. (2016) A review of the Nearctic linyphiid spider Arcuphantes fragilis (Araneae, Linyphiidae) and closely related species. Zootaxa, 4144 (3), 383 - 396. https: // doi. org / 10.11646 / zootaxa. 4144.3.6" type="journal article">
Ma
<emphasis id="9AF2D35F092D8C5DFF3B80F6F0E1FCD0" box="[198,250,764,788]" italics="true" pageId="4" pageNumber="502">et al</emphasis>
. 2016
</bibRefCitation>
), especially in
<taxonomicName id="6F8674CE092D8C5DFE0880F7F299FCD0" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[501,642,764,788]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092D8C5DFE0880F7F299FCD0" box="[501,642,764,788]" italics="true" pageId="4" pageNumber="502">Arcuphantes</emphasis>
</taxonomicName>
species, the close coupling of the male palpal paracymbium and the epigynal scape provides support for the compression of an inflated basal haematodocha that is the key for sperm insemination.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092D8C5DFF3A816CF33CFC44" box="[199,807,871,896]" pageId="4" pageNumber="502" type="distribution">
<paragraph id="A8390F4D092D8C5DFF3A816CF33CFC44" blockId="4.[151,1437,151,896]" box="[199,807,871,896]" pageId="4" pageNumber="502">
<emphasis id="9AF2D35F092D8C5DFF3A816CF17BFC44" bold="true" box="[199,352,871,896]" pageId="4" pageNumber="502">Distribution.</emphasis>
<collectingCountry id="D0914FDD092D8C5DFE9A8163F184FC44" box="[359,415,872,896]" name="United States of America" pageId="4" pageNumber="502">USA</collectingCountry>
(
<collectingRegion id="6A42C1AF092D8C5DFE50816CF22CFC44" box="[429,567,871,896]" country="United States of America" name="Washington" pageId="4" pageNumber="502">Washington</collectingRegion>
,
<collectingRegion id="6A42C1AF092D8C5DFDBF8163F282FC44" box="[578,665,872,896]" country="United States of America" name="Oregon" pageId="4" pageNumber="502">Oregon</collectingRegion>
,
<collectingRegion id="6A42C1AF092D8C5DFD58816CF307FC44" box="[677,796,871,896]" country="United States of America" name="California" pageId="4" pageNumber="502">California</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="C86FEA36A8568C401DD5F784ACF903F5" ID-CLB-Dataset="29989" ID-DOI="10.11646/zootaxa.4425.3.4" ID-GBIF-Dataset="9956099a-5e4b-4b2e-b24e-d347a707f5f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1270385" ID-ZooBank="038364CA-C0C7-4A56-8C23-7E0AC47F69ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528143569523" checkinUser="plazi" docAuthor="Jin, Miao, Ma, Shuangchun &amp; Tu, Lihong" docDate="2018" docId="202FBE5B092D8C5DFF6A81C2F25DF91A" docLanguage="en" docName="zootaxa.4425.3.4.pdf.imf" docOrigin="Zootaxa 4425 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcuphantes arcuatulus Chamberlin &amp; Ivie 1943" docType="treatment" docVersion="7" lastPageNumber="502" masterDocId="DC16C62309298C59FFFD820BF01BFFC4" masterDocTitle="A review of the Arcuphantes species with the genitalia of arcuatulus - type (Araneae, Linyphiidae)" masterLastPageNumber="510" masterPageNumber="498" pageNumber="502" updateTime="1698546615580" updateUser="ExternalLinkService">
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<mods:title id="307B6A8A2B47EEE75DF4D88858C11C62">A review of the Arcuphantes species with the genitalia of arcuatulus - type (Araneae, Linyphiidae)</mods:title>
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<mods:namePart id="F540C2A764919AC791618B0C35F2F372">Jin, Miao</mods:namePart>
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<mods:namePart id="04BC8CA4F91D5C27AC17B02B14008A35">Ma, Shuangchun</mods:namePart>
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<mods:namePart id="C85FCC94A8E090933DB5544A3AA51165">Tu, Lihong</mods:namePart>
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<mods:date id="048ADFEFFA894698C9698C633C394E64">2018</mods:date>
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<treatment id="202FBE5B092D8C5DFF6A81C2F25DF91A" ID-DOI="http://doi.org/10.5281/zenodo.5998046" ID-GBIF-Taxon="144235597" ID-Zenodo-Dep="5998046" LSID="urn:lsid:plazi:treatment:202FBE5B092D8C5DFF6A81C2F25DF91A" httpUri="http://treatment.plazi.org/id/202FBE5B092D8C5DFF6A81C2F25DF91A" lastPageNumber="502" pageId="4" pageNumber="502">
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<paragraph id="A8390F4D092D8C5DFF6A81C2F2E1FC27" blockId="4.[151,762,969,1029]" box="[151,762,969,995]" pageId="4" pageNumber="502">
<heading id="F371B821092D8C5DFF6A81C2F2E1FC27" bold="true" box="[151,762,969,995]" fontSize="11" level="1" pageId="4" pageNumber="502" reason="1">
<taxonomicName id="6F8674CE092D8C5DFF6A81C2F2E1FC27" ID-CoL="GC6N" authority="Chamberlin &amp; Ivie, 1943" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[151,762,969,995]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFF6A81C2F2E1FC27" bold="true" box="[151,762,969,995]" pageId="4" pageNumber="502">
<emphasis id="9AF2D35F092D8C5DFF6A81C2F1A0FC27" bold="true" box="[151,443,969,995]" italics="true" pageId="4" pageNumber="502">Arcuphantes arcuatulus</emphasis>
<bibRefCitation id="CC1772BC092D8C5DFE3F81C2F2E1FC27" box="[450,762,969,995]" pageId="4" pageNumber="502" refId="ref4098" refString="Chamberlin, R. V. &amp; Ivie, W. (1943) New genera and species of North American linyphiid spiders. Bulletin of the University of Utah, 10, 1 - 39." type="journal article">Chamberlin &amp; Ivie, 1943</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A8390F4D092D8C5DFF6A81E7F1D5FBC0" blockId="4.[151,762,969,1029]" box="[151,462,1004,1029]" pageId="4" pageNumber="502">
<figureCitation id="30BD13C8092D8C5DFF6A81E7F109FBC0" box="[151,274,1004,1029]" captionStart="FIGURE 2" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[278,1315,193,1918]" captionTargetId="figure@5.[278,1315,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Male palp. AE. Arcuphantes arcuatulus. A, prolateral; B, ventral; C, retrolateral; D, embolic division, ventral; E, ditto, dorsal; FJ, A. denticulatus n. sp., F, prolateral; G, ventral; H, retrolateral; I, embolic division, ventral; J, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="4" pageNumber="502">Figs 2AE</figureCitation>
,
<figureCitation id="30BD13C8092D8C5DFEE081E7F178FBC0" box="[285,355,1004,1028]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="4" pageNumber="502">3AC</figureCitation>
,
<figureCitation id="30BD13C8092D8C5DFE9381E7F165FBC0" box="[366,382,1004,1028]" captionStart="FIGURE 4" captionStartId="7.[151,250,1960,1982]" captionTargetBox="[151,1436,840,1908]" captionTargetId="figure@7.[151,1436,840,1911]" captionTargetPageId="7" captionText="FIGURE 4. Arcuphantes arcuatulus. AC, male palp, with embolic division removed, A, prolateral; B, ventral; C, retrolateral; DE. embolic division, arrows point to central serrated area of radix, D, ventral; E, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270393/files/figure.png" pageId="4" pageNumber="502">4</figureCitation>
,
<figureCitation id="30BD13C8092D8C5DFE7781E7F1D5FBC0" box="[394,462,1004,1028]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="4" pageNumber="502">7AB</figureCitation>
</paragraph>
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<subSubSection id="E09C5CC6092D8C5DFF6A863FF3BEFB00" pageId="4" pageNumber="502" type="reference_group">
<paragraph id="A8390F4D092D8C5DFF6A863FF43BFB8E" blockId="4.[151,1436,1076,1220]" box="[151,1056,1076,1098]" pageId="4" pageNumber="502">
<treatmentCitationGroup id="88962863092D8C5DFF6A863FF43BFB8E" box="[151,1056,1076,1098]" pageId="4" pageNumber="502">
<taxonomicName id="6F8674CE092D8C5DFF6A863FF238FB8E" authority="Keyserling, 1886: 74" authorityName="Keyserling" authorityPageNumber="74" authorityYear="1886" box="[151,547,1076,1098]" class="Arachnida" family="Linyphiidae" genus="Linyphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="9AF2D35F092D8C5DFF6A863FF15EFB8E" box="[151,325,1076,1098]" italics="true" pageId="4" pageNumber="502">Linyphia arcuata</emphasis>
<treatmentCitation id="2927295C092D8C5DFEB6863FF238FB8E" author="" box="[331,547,1076,1098]" page="74" pageId="4" pageNumber="502">
<bibRefCitation id="CC1772BC092D8C5DFEB6863FF1E6FB8E" box="[331,509,1076,1098]" pageId="4" pageNumber="502" refId="ref4135" refString="Keyserling, G. E. (1886) Die Spinnen Amerikas. Theridiidae. Vol. 2. Verlag von Bauer &amp; Raspe (E. Kuster), Nurnberg, 295 pp." type="book">Keyserling, 1886</bibRefCitation>
: 74
</treatmentCitation>
</taxonomicName>
, pl. 13, f. 173 (Df; preoccupied by Thorell, 1856).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8390F4D092D8C5DFF6A8659F28FFBAC" blockId="4.[151,1436,1076,1220]" box="[151,660,1106,1128]" pageId="4" pageNumber="502">
<taxonomicName id="6F8674CE092D8C5DFF6A8659F28BFBAC" authority="Petrunkevitch, 1911: 244" authorityName="Petrunkevitch" authorityPageNumber="244" authorityYear="1911" box="[151,656,1106,1128]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="9AF2D35F092D8C5DFF6A8659F192FBAC" box="[151,393,1106,1128]" italics="true" pageId="4" pageNumber="502">Lepthyphantes arcuatus</emphasis>
Petrunkevitch, 1911: 244
</taxonomicName>
.
</paragraph>
<paragraph id="A8390F4D092D8C5DFF6A867AF321FB43" blockId="4.[151,1436,1076,1220]" box="[151,826,1137,1159]" pageId="4" pageNumber="502">
<taxonomicName id="6F8674CE092D8C5DFF6A867AF278FB43" authority="Roewer, 1942: 560" authorityName="Roewer" authorityPageNumber="560" authorityYear="1942" box="[151,611,1137,1159]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFF6A867AF186FB43" box="[151,413,1137,1159]" italics="true" pageId="4" pageNumber="502">Lepthyphantes arcuatulus</emphasis>
Roewer, 1942: 560
</taxonomicName>
(replacement name).
</paragraph>
<paragraph id="A8390F4D092D8C5DFF6A869BF3BEFB00" blockId="4.[151,1436,1076,1220]" pageId="4" pageNumber="502">
<treatmentCitationGroup id="88962863092D8C5DFF6A869BF3BEFB00" infered="true" pageId="4" pageNumber="502">
<taxonomicName id="6F8674CE092D8C5DFF6A869BF2DAFB62" authority="Chamberlin &amp; Ivie, 1943: 21" authorityName="Chamberlin &amp; Ivie" authorityPageNumber="21" authorityYear="1943" box="[151,705,1168,1190]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="9AF2D35F092D8C5DFF6A869BF161FB62" box="[151,378,1168,1190]" italics="true" pageId="4" pageNumber="502">Arcuphantes arcuatus</emphasis>
<treatmentCitation id="2927295C092D8C5DFE78869BF2DAFB62" author="" box="[389,705,1168,1190]" page="21" pageId="4" pageNumber="502">
<bibRefCitation id="CC1772BC092D8C5DFE78869BF28CFB62" box="[389,663,1168,1190]" pageId="4" pageNumber="502" refId="ref4098" refString="Chamberlin, R. V. &amp; Ivie, W. (1943) New genera and species of North American linyphiid spiders. Bulletin of the University of Utah, 10, 1 - 39." type="journal article">Chamberlin &amp; Ivie, 1943</bibRefCitation>
: 21
</treatmentCitation>
</taxonomicName>
, f. 37-40 (Tf from
<taxonomicName id="6F8674CE092D8C5DFC61869BF470FB62" ID-CoL="5CTJ" authority=", Dm" authorityName="Dm" box="[924,1131,1168,1190]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092D8C5DFC61869BF42BFB62" box="[924,1072,1168,1190]" italics="true" pageId="4" pageNumber="502">Lepthyphantes</emphasis>
, Dm
</taxonomicName>
; specific name is a primary homonym replaced by Roewer);
<treatmentCitation id="2927295C092D8C5DFDE986A4F30AFB00" author="" box="[532,785,1198,1220]" page="61" pageId="4" pageNumber="502">
<bibRefCitation id="CC1772BC092D8C5DFDE986A4F2F6FB00" box="[532,749,1198,1220]" pageId="4" pageNumber="502" refId="ref4433" refString="Tu, L. H. &amp; Hormiga, G. (2010) The female genitalic morphology of &quot; micronetine &quot; spiders (Araneae, Linyphiidae). Genetica, 138, 59 - 73. https: // doi. org / 10.1007 / s 10709 - 009 - 9368 - 9" type="journal article">Tu &amp; Hormiga, 2010</bibRefCitation>
: 61
</treatmentCitation>
, f. 2a-f, 9a (f).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092D8C5DFF6A86FAF121FA96" pageId="4" pageNumber="502" type="materials_examined">
<paragraph id="A8390F4D092D8C5DFF6A86FAF121FA96" blockId="4.[151,1437,1265,1758]" pageId="4" pageNumber="502">
<materialsCitation id="18EE0510092D8C5DFF6A86FAF12DFAE9" ID-GBIF-Occurrence="1848253933" collectingDate="1935-09-09" collectorName="Chamberlin &amp; Ivie" country="United States of America" location="ArlingtOn" pageId="4" pageNumber="502" specimenCount="1" specimenCount-male="1" stateProvince="Washington">
<emphasis id="9AF2D35F092D8C5DFF6A86FAF192FACE" bold="true" box="[151,393,1265,1290]" pageId="4" pageNumber="502">Material examined.</emphasis>
<specimenCount id="BE80C4C4092D8C5DFE6586F9F1A7FACE" box="[408,444,1266,1290]" pageId="4" pageNumber="502" type="male">1♂</specimenCount>
,
<collectingCountry id="D0914FDD092D8C5DFE2C86F9F216FACE" box="[465,525,1266,1290]" name="United States of America" pageId="4" pageNumber="502">USA</collectingCountry>
:
<emphasis id="9AF2D35F092D8C5DFDDD86FAF2B6FACE" bold="true" box="[544,685,1265,1290]" pageId="4" pageNumber="502">
<collectingRegion id="6A42C1AF092D8C5DFDDD86FAF2B6FACE" box="[544,685,1265,1290]" country="United States of America" name="Washington" pageId="4" pageNumber="502">Washington</collectingRegion>
</emphasis>
:
<location id="AD595996092D8C5DFD3B86FAF32DFACE" LSID="urn:lsid:plazi:treatment:202FBE5B092D8C5DFF6A81C2F25DF91A:AD595996092D8C5DFD3B86FAF32DFACE" box="[710,822,1265,1290]" country="United States of America" name="ArlingtOn" pageId="4" pageNumber="502" stateProvince="Washington">Arlington</location>
, 48°N, 122°W,
<date id="DC38298D092D8C5DFC0286FAF470FACE" box="[1023,1131,1265,1290]" pageId="4" pageNumber="502" value="1935-09-09">
<collectingDate id="CC7CD065092D8C5DFC0286FAF470FACE" box="[1023,1131,1265,1290]" pageId="4" pageNumber="502" value="1935-09-09">9.ix.1935</collectingDate>
</date>
,
<collectorName id="05736A9B092D8C5DFB8286FAF518FACE" box="[1151,1283,1265,1290]" pageId="4" pageNumber="502">Chamberlin</collectorName>
&amp;
<collectorName id="05736A9B092D8C5DFAC586FAF57FFACE" box="[1336,1380,1265,1290]" pageId="4" pageNumber="502">Ivie</collectorName>
leg. (AMNH-161)
</materialsCitation>
;
<materialsCitation id="18EE0510092D8C5DFEB9871EF121FA96" ID-GBIF-Occurrence="1848253934" collectingDate="1935-09-10" collectorName="Chamberlin &amp; Ivie" country="United States of America" location="nOrth Of VancOuver" pageId="4" pageNumber="502" specimenCount="1" specimenCount-female="1" stateProvince="Washington">
<specimenCount id="BE80C4C4092D8C5DFEB9871EF17DFAE9" box="[324,358,1301,1325]" pageId="4" pageNumber="502" type="female">1♀</specimenCount>
<emphasis id="9AF2D35F092D8C5DFE8D871EF1E4FAEA" bold="true" box="[368,511,1301,1326]" pageId="4" pageNumber="502">
<collectingRegion id="6A42C1AF092D8C5DFE8D871EF1E4FAEA" box="[368,511,1301,1326]" country="United States of America" name="Washington" pageId="4" pageNumber="502">Washington</collectingRegion>
</emphasis>
:
<quantity id="6F7EA2A8092D8C5DFDED871DF268FAEA" box="[528,627,1301,1326]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="4" pageNumber="502" unit="mi" value="10.0">10 miles</quantity>
<location id="AD595996092D8C5DFD81871EF347FAEA" LSID="urn:lsid:plazi:treatment:202FBE5B092D8C5DFF6A81C2F25DF91A:AD595996092D8C5DFD81871EF347FAEA" box="[636,860,1301,1326]" country="United States of America" name="nOrth Of VancOuver" pageId="4" pageNumber="502" stateProvince="Washington">north of Vancouver</location>
, 45°N, 122°W,
<date id="DC38298D092D8C5DFBEF871EF497FAEA" box="[1042,1164,1301,1326]" pageId="4" pageNumber="502" value="1935-09-10">
<collectingDate id="CC7CD065092D8C5DFBEF871EF497FAEA" box="[1042,1164,1301,1326]" pageId="4" pageNumber="502" value="1935-09-10">10.ix.1935</collectingDate>
</date>
,
<collectorName id="05736A9B092D8C5DFB67871EF505FAEA" box="[1178,1310,1301,1326]" pageId="4" pageNumber="502">Chamberlin</collectorName>
&amp;
<collectorName id="05736A9B092D8C5DFAC0871EF572FAEA" box="[1341,1385,1301,1326]" pageId="4" pageNumber="502">Ivie</collectorName>
leg. (AMNH-165).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092D8C5DFF3A8756F34DF9C2" pageId="4" pageNumber="502" type="diagnosis">
<paragraph id="A8390F4D092D8C5DFF3A8756F34DF9C2" blockId="4.[151,1437,1265,1758]" pageId="4" pageNumber="502">
<emphasis id="9AF2D35F092D8C5DFF3A8756F15AFAB2" bold="true" box="[199,321,1373,1398]" pageId="4" pageNumber="502">Diagnosis.</emphasis>
The male
<taxonomicName id="6F8674CE092D8C5DFE458755F256FAB1" authorityName="Roewer" authorityYear="1942" box="[440,589,1373,1397]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFE458755F256FAB1" box="[440,589,1373,1397]" italics="true" pageId="4" pageNumber="502">A. arcuatulus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="6F8674CE092D8C5DFC828755F41DFAB1" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[895,1030,1374,1397]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092D8C5DFC828755F38BFAB1" box="[895,912,1374,1397]" italics="true" pageId="4" pageNumber="502">A</emphasis>
.
<emphasis id="9AF2D35F092D8C5DFC638755F41DFAB1" box="[926,1030,1374,1397]" italics="true" pageId="4" pageNumber="502">cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092D8C5DFBF08756F457FAB2" bold="true" box="[1037,1100,1373,1398]" pageId="4" pageNumber="502">
<taxonomicNameLabel id="81C16E24092D8C5DFBF08756F457FAB2" box="[1037,1100,1373,1398]" pageId="4" pageNumber="502" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="6F8674CE092D8C5DFB778755F528FAB1" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1162,1331,1373,1397]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="9AF2D35F092D8C5DFB778755F528FAB1" box="[1162,1331,1373,1397]" italics="true" pageId="4" pageNumber="502">A. denticulatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092D8C5DFAC78756F562FAB2" bold="true" box="[1338,1401,1373,1398]" pageId="4" pageNumber="502">
<taxonomicNameLabel id="81C16E24092D8C5DFAC78756F562FAB2" box="[1338,1401,1373,1398]" pageId="4" pageNumber="502" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the bifid outer paracymbial arm with a blunt inner apophysis (
<figureCitation id="30BD13C8092D8C5DFCA7878AF3A9FA5E" box="[858,946,1409,1434]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="4" pageNumber="502">Fig. 7B</figureCitation>
), and by the terminal apophysis having the anterior process turning backwards (
<figureCitation id="30BD13C8092D8C5DFDCA87AEF28FFA7A" box="[567,660,1445,1470]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="4" pageNumber="502">Fig. 7A</figureCitation>
). The female can be distinguished by the epigynum in ventral view more slender than that of
<taxonomicName id="6F8674CE092D8C5DFE4187C1F25EFA25" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[444,581,1482,1505]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092D8C5DFE4187C1F1D6FA25" box="[444,461,1482,1505]" italics="true" pageId="4" pageNumber="502">A</emphasis>
.
<emphasis id="9AF2D35F092D8C5DFE2087C1F25EFA25" box="[477,581,1482,1505]" italics="true" pageId="4" pageNumber="502">cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092D8C5DFDB087C2F296FA26" bold="true" box="[589,653,1481,1506]" pageId="4" pageNumber="502">
<taxonomicNameLabel id="81C16E24092D8C5DFDB087C2F296FA26" box="[589,653,1481,1506]" pageId="4" pageNumber="502" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the prolonged basal part under the extensible base is almost square, with the parallel lateral margins slightly incurved (
<figureCitation id="30BD13C8092D8C5DFD3387E6F352F9C2" box="[718,841,1517,1542]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="4" pageNumber="502">Fig. 3AC</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092D8C5DFF3A841AF12DF97E" pageId="4" pageNumber="502" type="description">
<paragraph id="A8390F4D092D8C5DFF3A841AF242F9B6" blockId="4.[151,1437,1265,1758]" pageId="4" pageNumber="502">
<emphasis id="9AF2D35F092D8C5DFF3A841AF142F9EE" bold="true" box="[199,345,1553,1578]" pageId="4" pageNumber="502">Description.</emphasis>
<emphasis id="9AF2D35F092D8C5DFE9D841AF181F9ED" box="[352,410,1553,1577]" italics="true" pageId="4" pageNumber="502">Male</emphasis>
(AMNH-161): Total length 1.94. Carapace 0.78 long, 0.63 wide. Abdomen 1.11 long, 0.53 wide.
<emphasis id="9AF2D35F092D8C5DFF23843EF129F989" box="[222,306,1589,1613]" italics="true" pageId="4" pageNumber="502">Female</emphasis>
(AMNH-165): Total length 1.26. Carapace 0.57 long, 0.54 wide. Abdomen 0.75 long, 0.49 wide. Legs broken, data on leg lengths absent.
</paragraph>
<paragraph id="A8390F4D092D8C5DFF3A8476F12DF97E" blockId="4.[151,1437,1265,1758]" pageId="4" pageNumber="502">
Other somatic and genital characters: see the general description for the
<taxonomicName id="6F8674CE092D8C5DFC0B8476F498F951" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1014,1155,1661,1685]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092D8C5DFC0B8476F498F951" box="[1014,1155,1661,1685]" italics="true" pageId="4" pageNumber="502">Arcuphantes</emphasis>
</taxonomicName>
species with
<taxonomicName id="6F8674CE092D8C5DFAE18476F588F951" authorityName="Roewer" authorityYear="1942" box="[1308,1427,1661,1685]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092D8C5DFAE18476F588F951" box="[1308,1427,1661,1685]" italics="true" pageId="4" pageNumber="502">arcuatulus</emphasis>
</taxonomicName>
- type genitalia.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092D8C5DFF3A84CEF25DF91A" box="[199,582,1733,1758]" pageId="4" pageNumber="502" type="distribution">
<paragraph id="A8390F4D092D8C5DFF3A84CEF25DF91A" blockId="4.[151,1437,1265,1758]" box="[199,582,1733,1758]" pageId="4" pageNumber="502">
<emphasis id="9AF2D35F092D8C5DFF3A84CEF17BF91A" bold="true" box="[199,352,1733,1758]" pageId="4" pageNumber="502">Distribution.</emphasis>
<collectingCountry id="D0914FDD092D8C5DFE9A84CDF184F91A" box="[359,415,1734,1758]" name="United States of America" pageId="4" pageNumber="502">USA</collectingCountry>
(
<collectingRegion id="6A42C1AF092D8C5DFE5084CEF221F91A" box="[429,570,1733,1758]" country="United States of America" name="Washington" pageId="4" pageNumber="502">Washington</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,291 @@
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<mods:title id="513F566C1DF3F3C4B64AD5644B62D236">A review of the Arcuphantes species with the genitalia of arcuatulus - type (Araneae, Linyphiidae)</mods:title>
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<treatment id="202FBE5B092D8C5EFF6A852DF20FFD34" ID-DOI="http://doi.org/10.5281/zenodo.5998050" ID-GBIF-Taxon="144235596" ID-Zenodo-Dep="5998050" LSID="urn:lsid:zoobank.org:act:8B526ADC-5673-44F9-93F2-9BD3F5BDCEF6" httpUri="http://treatment.plazi.org/id/202FBE5B092D8C5EFF6A852DF20FFD34" lastPageId="7" lastPageNumber="505" pageId="4" pageNumber="502">
<subSubSection id="E09C5CC6092D8C5DFF6A852DF163F8A6" pageId="4" pageNumber="502" type="nomenclature">
<paragraph id="A8390F4D092D8C5DFF6A852DF25BF884" blockId="4.[151,576,1830,1890]" box="[151,576,1830,1856]" pageId="4" pageNumber="502">
<heading id="F371B821092D8C5DFF6A852DF25BF884" bold="true" box="[151,576,1830,1856]" fontSize="11" level="1" pageId="4" pageNumber="502" reason="1">
<emphasis id="9AF2D35F092D8C5DFF6A852DF25BF884" bold="true" box="[151,576,1830,1856]" pageId="4" pageNumber="502">
<taxonomicName id="6F8674CE092D8C5DFF6A852DF1B2F884" ID-CoL="GC6V" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[151,425,1830,1856]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="502" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092D8C5DFF6A852DF1B2F884" bold="true" box="[151,425,1830,1856]" italics="true" pageId="4" pageNumber="502">Arcuphantes cruciatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81C16E24092D8C5DFE4D852DF25BF884" box="[432,576,1830,1856]" pageId="4" pageNumber="502" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A8390F4D092D8C5DFF6A8542F163F8A6" blockId="4.[151,576,1830,1890]" box="[151,376,1865,1890]" pageId="4" pageNumber="502">
<figureCitation id="30BD13C8092D8C5DFF6A8542F115F8A6" box="[151,270,1865,1890]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="4" pageNumber="502">Figs 3DF</figureCitation>
,
<figureCitation id="30BD13C8092D8C5DFEE78542F132F8A6" box="[282,297,1865,1890]" captionStart="FIGURE 5" captionStartId="8.[151,250,1905,1927]" captionTargetBox="[171,1415,193,1871]" captionTargetId="figure@8.[171,1415,193,1878]" captionTargetPageId="8" captionText="FIGURE 5. Arcuphantes cruciatus n. sp. AB, male palp, with embolic division removed, A, prolateral; B, retrolateral; CD, embolic division, arrows point to central serrated area of radix, C, ventral; D, dorsal; EH, epigynum, with extensible basal part removed, E, ventral, ventral plate artificially broken; F, dorsal; G, caudal; H, inner, shows epigynal grooves. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270395/files/figure.png" pageId="4" pageNumber="502">5</figureCitation>
,
<figureCitation id="30BD13C8092D8C5DFEC88541F163F8A6" box="[309,376,1866,1890]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="4" pageNumber="502">7CD</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092D8C5DFF6A859AF16EF836" pageId="4" pageNumber="502" type="materials_examined">
<paragraph id="A8390F4D092D8C5DFF6A859AF16EF836" blockId="4.[151,1436,1937,2034]" pageId="4" pageNumber="502">
<emphasis id="9AF2D35F092D8C5DFF6A859AF1D2F86E" bold="true" box="[151,457,1937,1962]" pageId="4" pageNumber="502">Type material examined.</emphasis>
<emphasis id="9AF2D35F092D8C5DFE29859AF220F86D" box="[468,571,1937,1961]" italics="true" pageId="4" pageNumber="502">Holotype</emphasis>
♂.
<materialsCitation id="18EE0510092D8C5DFD928599F29EF809" ID-GBIF-Occurrence="1848253931" collectingDate="1959-09-30" country="United States of America" latitude="42.85" location="Sixes" longLatPrecision="1147" longitude="-124.5" pageId="4" pageNumber="502" specimenCount="1" stateProvince="Oregon" typeStatus="paratype">
<collectingCountry id="D0914FDD092D8C5DFD928599F2B0F86E" box="[623,683,1938,1962]" name="United States of America" pageId="4" pageNumber="502">USA</collectingCountry>
:
<emphasis id="9AF2D35F092D8C5DFD47859AF30FF86E" bold="true" box="[698,788,1937,1962]" pageId="4" pageNumber="502">
<collectingRegion id="6A42C1AF092D8C5DFD47859AF30FF86E" box="[698,788,1937,1962]" country="United States of America" name="Oregon" pageId="4" pageNumber="502">Oregon</collectingRegion>
</emphasis>
:
<location id="AD595996092D8C5DFCD5859AF37EF86E" LSID="urn:lsid:plazi:treatment:202FBE5B092D8C5EFF6A852DF20FFD34:AD595996092D8C5DFCD5859AF37EF86E" box="[808,869,1937,1962]" country="United States of America" latitude="42.85" longLatPrecision="1147" longitude="-124.5" name="Sixes" pageId="4" pageNumber="502" stateProvince="Oregon">Sixes</location>
(
<quantity id="6F7EA2A8092D8C5DFC868599F3DEF86E" box="[891,965,1937,1962]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="4" pageNumber="502" unit="mi" value="1.0">1 mile</quantity>
north),
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;
<geoCoordinate id="CDB2698A092D8C5DFB50859AF516F86E" box="[1197,1293,1937,1962]" direction="north" orientation="latitude" pageId="4" pageNumber="502" precision="925" value="42.85">42°51N</geoCoordinate>
,
<date id="DC38298D092D8C5DFAE0859AF58CF86E" box="[1309,1431,1937,1962]" pageId="4" pageNumber="502" value="1959-09-30">
<collectingDate id="CC7CD065092D8C5DFAE0859AF58CF86E" box="[1309,1431,1937,1962]" pageId="4" pageNumber="502" value="1959-09-30">30.ix.1959</collectingDate>
</date>
, Vincent Roth leg. (AMNH-147).
<emphasis id="9AF2D35F092D8C5DFDEE85BDF29EF809" box="[531,645,1974,1997]" italics="true" pageId="4" pageNumber="502">
<typeStatus id="773DB1EF092D8C5DFDEE85BDF29EF809" box="[531,645,1974,1997]" pageId="4" pageNumber="502" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
.
<materialsCitation id="18EE0510092D8C5DFD6985BDF16EF836" ID-GBIF-Occurrence="1848253930" collectingDate="1959-09-30" country="United States of America" latitude="42.85" location="Sixes" longLatPrecision="1147" longitude="-124.5" pageId="4" pageNumber="502" specimenCount="2" specimenCount-female="2" stateProvince="Oregon" typeStatus="holotype">
<specimenCount id="BE80C4C4092D8C5DFD6985BDF2ACF80A" box="[660,695,1972,1998]" pageId="4" pageNumber="502" type="female">
2
<emphasis id="9AF2D35F092D8C5DFD5F85BFF2ACF80A" box="[674,695,1972,1998]" italics="true" pageId="4" pageNumber="502"></emphasis>
</specimenCount>
, same data, together with the
<typeStatus id="773DB1EF092D8C5DFBEF85BEF46FF80A" box="[1042,1140,1973,1998]" pageId="4" pageNumber="502" type="holotype">holotype</typeStatus>
(AMNH-147, one female without epigynum).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="FCF95FC5092C8C5CFF6A859AF321F820" httpUri="https://zenodo.org/record/1270389/files/figure.png" pageId="5" pageNumber="503" startId="5.[151,250,1937,1959]" targetBox="[278,1315,193,1918]" targetPageId="5">
<paragraph id="A8390F4D092C8C5CFF6A859AF321F820" blockId="5.[151,1437,1937,2020]" pageId="5" pageNumber="503">
<emphasis id="9AF2D35F092C8C5CFF6A859AF109F863" bold="true" box="[151,274,1937,1959]" pageId="5" pageNumber="503">FIGURE 2.</emphasis>
Male palp.
<emphasis id="9AF2D35F092C8C5CFE76859AF1A0F863" bold="true" box="[395,443,1937,1959]" pageId="5" pageNumber="503">AE</emphasis>
.
<taxonomicName id="6F8674CE092C8C5CFE3A859AF2ACF863" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[455,695,1937,1959]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="503" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092C8C5CFE3A859AF2ACF863" box="[455,695,1937,1959]" italics="true" pageId="5" pageNumber="503">Arcuphantes arcuatulus</emphasis>
</taxonomicName>
.
<emphasis id="9AF2D35F092C8C5CFD39859AF2CDF863" bold="true" box="[708,726,1937,1959]" pageId="5" pageNumber="503">A</emphasis>
, prolateral;
<emphasis id="9AF2D35F092C8C5CFCAC859AF379F863" bold="true" box="[849,866,1937,1959]" pageId="5" pageNumber="503">B</emphasis>
, ventral;
<emphasis id="9AF2D35F092C8C5CFC3F859AF3CFF863" bold="true" box="[962,980,1937,1959]" pageId="5" pageNumber="503">C</emphasis>
, retrolateral;
<emphasis id="9AF2D35F092C8C5CFBA0859AF474F863" bold="true" box="[1117,1135,1937,1959]" pageId="5" pageNumber="503">D</emphasis>
, embolic division, ventral;
<emphasis id="9AF2D35F092C8C5CFA78859AF58DF863" bold="true" box="[1413,1430,1937,1959]" pageId="5" pageNumber="503">E</emphasis>
, ditto, dorsal;
<emphasis id="9AF2D35F092C8C5CFEE285A4F152F801" bold="true" box="[287,329,1967,1989]" pageId="5" pageNumber="503">FJ</emphasis>
,
<taxonomicName id="6F8674CE092C8C5CFEAB85BBF1F4F801" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[342,495,1967,1989]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="503" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="9AF2D35F092C8C5CFEAB85BBF1F4F801" box="[342,495,1967,1989]" italics="true" pageId="5" pageNumber="503">A. denticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81C16E24092C8C5CFE0A85A4F22AF801" box="[503,561,1967,1989]" pageId="5" pageNumber="503" rank="species">
<emphasis id="9AF2D35F092C8C5CFE0A85A4F22AF801" bold="true" box="[503,561,1967,1989]" pageId="5" pageNumber="503">n. sp.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="9AF2D35F092C8C5CFDC285A4F257F801" bold="true" box="[575,588,1967,1989]" pageId="5" pageNumber="503">F</emphasis>
, prolateral;
<emphasis id="9AF2D35F092C8C5CFD3485A4F2C1F801" bold="true" box="[713,730,1967,1989]" pageId="5" pageNumber="503">G</emphasis>
, ventral;
<emphasis id="9AF2D35F092C8C5CFCC085A4F34AF801" bold="true" box="[829,849,1967,1989]" pageId="5" pageNumber="503">H</emphasis>
, retrolateral;
<emphasis id="9AF2D35F092C8C5CFC2185A4F3FDF801" bold="true" box="[988,998,1967,1989]" pageId="5" pageNumber="503">I</emphasis>
, embolic division, ventral;
<emphasis id="9AF2D35F092C8C5CFAFC85A4F516F801" bold="true" box="[1281,1293,1967,1989]" pageId="5" pageNumber="503">J</emphasis>
, ditto, dorsal. Arrows in D, E, I, J refer to serrate area on radix. [Scale bars, mm]
</paragraph>
</caption>
<caption id="FCF95FC5092F8C5FFF6A859DF1EAF82E" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="6" pageNumber="504" startId="6.[151,250,1942,1964]" targetBox="[215,1372,193,1920]" targetPageId="6">
<paragraph id="A8390F4D092F8C5FFF6A859DF1EAF82E" blockId="6.[151,1436,1942,2026]" pageId="6" pageNumber="504">
<emphasis id="9AF2D35F092F8C5FFF6A859DF10FF868" bold="true" box="[151,276,1942,1964]" pageId="6" pageNumber="504">FIGURE 3.</emphasis>
Epigynum.
<emphasis id="9AF2D35F092F8C5FFE68859DF1DDF868" bold="true" box="[405,454,1942,1964]" pageId="6" pageNumber="504">AC</emphasis>
.
<taxonomicName id="6F8674CE092F8C5FFE28859DF2D3F868" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[469,712,1942,1964]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="504" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092F8C5FFE28859DF2D3F868" box="[469,712,1942,1964]" italics="true" pageId="6" pageNumber="504">Arcuphantes arcuatulus</emphasis>
</taxonomicName>
,
<emphasis id="9AF2D35F092F8C5FFD2B859DF2F3F868" bold="true" box="[726,744,1942,1964]" pageId="6" pageNumber="504">A</emphasis>
, ventral;
<emphasis id="9AF2D35F092F8C5FFCB0859DF345F868" bold="true" box="[845,862,1942,1964]" pageId="6" pageNumber="504">B</emphasis>
, lateral;
<emphasis id="9AF2D35F092F8C5FFC41859DF3D5F868" bold="true" box="[956,974,1942,1964]" pageId="6" pageNumber="504">C</emphasis>
, dorsal, arrow points to the proximal end of fertilization groove;
<emphasis id="9AF2D35F092F8C5FFE9685BEF180F80F" bold="true" box="[363,411,1973,1995]" pageId="6" pageNumber="504">DC</emphasis>
,
<taxonomicName id="6F8674CE092F8C5FFE5785BDF23DF80F" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[426,550,1974,1995]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="504" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092F8C5FFE5785BDF23DF80F" box="[426,550,1974,1995]" italics="true" pageId="6" pageNumber="504">A. cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092F8C5FFDD385BEF272F80F" bold="true" box="[558,617,1973,1995]" pageId="6" pageNumber="504">
<taxonomicNameLabel id="81C16E24092F8C5FFDD385BEF272F80F" box="[558,617,1973,1995]" pageId="6" pageNumber="504" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="9AF2D35F092F8C5FFD8585BEF291F80F" bold="true" box="[632,650,1973,1995]" pageId="6" pageNumber="504">D</emphasis>
, ventral;
<emphasis id="9AF2D35F092F8C5FFD1385BEF2E4F80F" bold="true" box="[750,767,1973,1995]" pageId="6" pageNumber="504">E</emphasis>
, lateral;
<emphasis id="9AF2D35F092F8C5FFCA185BEF372F80F" bold="true" box="[860,873,1973,1995]" pageId="6" pageNumber="504">F</emphasis>
, dorsal;
<emphasis id="9AF2D35F092F8C5FFC3885BEF3F4F80F" bold="true" box="[965,1007,1973,1995]" pageId="6" pageNumber="504">GI</emphasis>
,
<taxonomicName id="6F8674CE092F8C5FFC0085BDF4A5F80F" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1021,1214,1973,1995]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="504" phylum="Arthropoda" rank="species" species="semiorbiculatus" status="sp. nov.">
<emphasis id="9AF2D35F092F8C5FFC0085BDF4A5F80F" box="[1021,1214,1973,1995]" italics="true" pageId="6" pageNumber="504">A. semiorbiculatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092F8C5FFB3885BEF51BF80F" bold="true" box="[1221,1280,1973,1995]" pageId="6" pageNumber="504">
<taxonomicNameLabel id="81C16E24092F8C5FFB3885BEF51BF80F" box="[1221,1280,1973,1995]" pageId="6" pageNumber="504" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, G, ventral;
<emphasis id="9AF2D35F092F8C5FFA7F85BEF58DF80F" bold="true" box="[1410,1430,1973,1995]" pageId="6" pageNumber="504">H</emphasis>
, lateral;
<emphasis id="9AF2D35F092F8C5FFF1E85DFF0F6F82E" bold="true" box="[227,237,2004,2026]" pageId="6" pageNumber="504">I</emphasis>
, dorsal. [Scale bars, mm]
</paragraph>
</caption>
<subSubSection id="E09C5CC6092E8C5EFF3A829CF122FF11" pageId="7" pageNumber="505" type="etymology">
<paragraph id="A8390F4D092E8C5EFF3A829CF122FF11" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="505">
<emphasis id="9AF2D35F092E8C5EFF3A829CF155FF74" bold="true" box="[199,334,151,176]" pageId="7" pageNumber="505">Etymology.</emphasis>
The species name comes from the Latin word cruciatus, in reference to the cross-shaped paracymbium.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092E8C5EFF3A82D4F214FE14" pageId="7" pageNumber="505" type="diagnosis">
<paragraph id="A8390F4D092E8C5EFF3A82D4F214FE14" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="505">
<emphasis id="9AF2D35F092E8C5EFF3A82D4F15AFF3C" bold="true" box="[199,321,223,248]" pageId="7" pageNumber="505">Diagnosis.</emphasis>
The male
<taxonomicName id="6F8674CE092E8C5EFE4482EBF25AFF33" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[441,577,224,247]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092E8C5EFE4482EBF1D1FF33" box="[441,458,224,247]" italics="true" pageId="7" pageNumber="505">A</emphasis>
.
<emphasis id="9AF2D35F092E8C5EFE2482EBF25AFF33" box="[473,577,224,247]" italics="true" pageId="7" pageNumber="505">cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092E8C5EFDB482D4F293FF3C" bold="true" box="[585,648,223,248]" pageId="7" pageNumber="505">
<taxonomicNameLabel id="81C16E24092E8C5EFDB482D4F293FF3C" box="[585,648,223,248]" pageId="7" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="6F8674CE092E8C5EFC4682EBF449FF3C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[955,1106,224,248]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092E8C5EFC4682EBF449FF3C" box="[955,1106,224,248]" italics="true" pageId="7" pageNumber="505">A. arcuatulus</emphasis>
</taxonomicName>
and
<taxonomicName id="6F8674CE092E8C5EFB7482EBF52AFF3C" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[1161,1329,224,248]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="9AF2D35F092E8C5EFB7482EBF52AFF3C" box="[1161,1329,224,248]" italics="true" pageId="7" pageNumber="505">A. denticulatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092E8C5EFAC482D4F563FF3C" bold="true" box="[1337,1400,223,248]" pageId="7" pageNumber="505">
<taxonomicNameLabel id="81C16E24092E8C5EFAC482D4F563FF3C" box="[1337,1400,223,248]" pageId="7" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the thin sclerite-like paracymbial outer arm (
<figureCitation id="30BD13C8092E8C5EFD75830FF2F9FED8" box="[648,738,260,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="7" pageNumber="505">Fig. 7D</figureCitation>
), which is thick and bifid in two other species (
<figureCitation id="30BD13C8092E8C5EFB0D830FF57EFED8" box="[1264,1381,260,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="7" pageNumber="505">Fig. 7B, F</figureCitation>
); the fold at the basal part of the paracymbial middle arm is absent (
<figureCitation id="30BD13C8092E8C5EFCA3832CF3A2FE84" box="[862,953,295,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="7" pageNumber="505">Fig. 7D</figureCitation>
), but present in the other two species (
<figureCitation id="30BD13C8092E8C5EFA93832CF0C2FEA0" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="7" pageNumber="505">Fig. 7B, F</figureCitation>
); and by the anterior part of terminal apophysis bulged (
<figureCitation id="30BD13C8092E8C5EFCAF8347F3B1FEA0" box="[850,938,332,357]" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="7" pageNumber="505">Fig. 7C</figureCitation>
), whereas in two other species it is flat (
<figureCitation id="30BD13C8092E8C5EFA938347F0C6FE4C" captionStart="FIGURE 7" captionStartId="10.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1491]" captionTargetId="figure@10.[151,1436,193,1494]" captionTargetPageId="10" captionText="FIGURE 7. Male palpal embolic division and paracymbium. AB, Arcuphantes arcuatulus, CD, A. cruciatus n. sp., EF, A. denticulatus n. sp., A, C, E, embolic division, anterior, embolus apex artificially broken in E, arrows point to anterior process of terminal apophysis turning backwards in A and extending forward in E, and bulged anterior part of terminal apophysis in C; B, D, F, paracymbium, arrows in B and F point to basal fold of paracymbial middle arm. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270399/files/figure.png" pageId="7" pageNumber="505">Fig. 7A, E</figureCitation>
). The epigynum of
<taxonomicName id="6F8674CE092E8C5EFE46837BF258FE43" authority="Jin &amp; Ma &amp; Tu, 2018" authorityName="Jin &amp; Ma &amp; Tu" authorityYear="2018" box="[443,579,368,391]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="species" species="cruciatus" status="sp. nov.">
<emphasis id="9AF2D35F092E8C5EFE46837BF1D7FE43" box="[443,460,368,391]" italics="true" pageId="7" pageNumber="505">A</emphasis>
.
<emphasis id="9AF2D35F092E8C5EFE26837BF258FE43" box="[475,579,368,391]" italics="true" pageId="7" pageNumber="505">cruciatus</emphasis>
</taxonomicName>
<emphasis id="9AF2D35F092E8C5EFDB68364F290FE4C" bold="true" box="[587,651,367,392]" pageId="7" pageNumber="505">
<taxonomicNameLabel id="81C16E24092E8C5EFDB68364F290FE4C" box="[587,651,367,392]" pageId="7" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is similar to that of
<taxonomicName id="6F8674CE092E8C5EFC8F837BF41CFE4C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[882,1031,368,392]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092E8C5EFC8F837BF41CFE4C" box="[882,1031,368,392]" italics="true" pageId="7" pageNumber="505">A. arcuatulus</emphasis>
</taxonomicName>
, but the appearance of epigynum is more stout than in the latter species; the prolonged basal part under the extensible base is wider than long, with parallel lateral margins (
<figureCitation id="30BD13C8092E8C5EFE5583BCF219FE14" box="[424,514,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[215,1372,193,1920]" captionTargetId="figure@6.[215,1372,193,1921]" captionTargetPageId="6" captionText="FIGURE 3. Epigynum. AC. Arcuphantes arcuatulus, A, ventral; B, lateral; C, dorsal, arrow points to the proximal end of fertilization groove; DC, A. cruciatus n. sp., D, ventral; E, lateral; F, dorsal; GI, A. semiorbiculatus n. sp., G, ventral; H, lateral; I, dorsal. [Scale bars, mm]" httpUri="https://zenodo.org/record/1270391/files/figure.png" pageId="7" pageNumber="505">Fig. 3D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092E8C5EFF3A83D7F12DFD09" pageId="7" pageNumber="505" type="description">
<paragraph id="A8390F4D092E8C5EFF3A83D7F0CEFDDC" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="505">
<emphasis id="9AF2D35F092E8C5EFF3A83D7F142FE31" bold="true" box="[199,345,476,501]" pageId="7" pageNumber="505">Description.</emphasis>
<emphasis id="9AF2D35F092E8C5EFEA283D7F182FE30" box="[351,409,476,500]" italics="true" pageId="7" pageNumber="505">Male</emphasis>
(AMNH-147): Total length 2.21. Carapace 1.05 long, 0.87 wide. Abdomen 1.24 long, 0.60 wide.
</paragraph>
<paragraph id="A8390F4D092E8C5EFF3A802FF23AFD40" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="505">
<emphasis id="9AF2D35F092E8C5EFF3A802FF100FDF8" box="[199,283,548,572]" italics="true" pageId="7" pageNumber="505">Female</emphasis>
(AMNH-147): Total length 1.63. Carapace 0.63 long, 0.59 wide. Abdomen 0.98 long, 0.70 wide. Leg lengths: I 6.53 (1.73+2.10+1.64+1.06); II 5.84 (1.64+1.83+1.46+0.91); III 5.00 (1.35+1.49+1.29+0.87); IV 5.33 (1.69+1.72+1.27+0.65). Tm I 0.19.
</paragraph>
<paragraph id="A8390F4D092E8C5EFF3A8084F12DFD09" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="505">
Other somatic and genital characters: see the general description for the
<taxonomicName id="6F8674CE092E8C5EFC0B809BF498FD6C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1014,1155,656,680]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="9AF2D35F092E8C5EFC0B809BF498FD6C" box="[1014,1155,656,680]" italics="true" pageId="7" pageNumber="505">Arcuphantes</emphasis>
</taxonomicName>
species with
<taxonomicName id="6F8674CE092E8C5EFAE1809BF588FD6C" authorityName="Chamberlin &amp; Ivie" authorityYear="1943" box="[1308,1427,656,680]" class="Arachnida" family="Linyphiidae" genus="Arcuphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="505" phylum="Arthropoda" rank="species" species="arcuatulus">
<emphasis id="9AF2D35F092E8C5EFAE1809BF588FD6C" box="[1308,1427,656,680]" italics="true" pageId="7" pageNumber="505">arcuatulus</emphasis>
</taxonomicName>
-
<typeStatus id="773DB1EF092E8C5EFF6A80BEF0DCFD09" box="[151,199,693,717]" pageId="7" pageNumber="505">type</typeStatus>
genitalia.
</paragraph>
</subSubSection>
<subSubSection id="E09C5CC6092E8C5EFF3A80DCF20FFD34" box="[199,532,727,752]" pageId="7" pageNumber="505" type="distribution">
<paragraph id="A8390F4D092E8C5EFF3A80DCF20FFD34" blockId="7.[151,1437,151,752]" box="[199,532,727,752]" pageId="7" pageNumber="505">
<emphasis id="9AF2D35F092E8C5EFF3A80DCF17BFD34" bold="true" box="[199,352,727,752]" pageId="7" pageNumber="505">Distribution.</emphasis>
<collectingCountry id="D0914FDD092E8C5EFE9A80D3F184FD34" box="[359,415,728,752]" name="United States of America" pageId="7" pageNumber="505">USA</collectingCountry>
(
<collectingRegion id="6A42C1AF092E8C5EFE5380D3F212FD34" box="[430,521,728,752]" country="United States of America" name="Oregon" pageId="7" pageNumber="505">Oregon</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,64 @@
<document ID-DOI="http://dx.doi.org/10.3897/BDJ.6.e27161" ID-PMC="PMC6220116" ID-Pensoft-Pub="1314-2828--27161" ID-PubMed="30425605" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2828--e27161" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="A list of bees from three locations in the Northern Rockies Ecoregion (NRE) of western Montana" checkinTime="1553125790152" checkinUser="pensoft" docAuthor="Reese, Elizabeth G., Burkle, Laura A., Delphia, Casey M. &amp; Griswold, Terry" docDate="2018" docId="202FF5FC3738837074070055A6185BD6" docLanguage="en" docName="BiodivDatJour 6: e27161" docOrigin="Biodiversity Data Journal 6" docSource="http://dx.doi.org/10.3897/BDJ.6.e27161" docTitle="Bombus (Pyrobombus) vagans Smith 1854" docType="treatment" docVersion="5" lastPageNumber="27161" masterDocId="44466B5EFFF9D76DCD4FFF84D505FFCC" masterDocTitle="A list of bees from three locations in the Northern Rockies Ecoregion (NRE) of western Montana" masterLastPageNumber="27161" masterPageNumber="27161" pageNumber="27161" updateTime="1668124804368" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A list of bees from three locations in the Northern Rockies Ecoregion (NRE) of western Montana</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Reese, Elizabeth G.</mods:namePart>
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<mods:role>
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<mods:namePart>Burkle, Laura A.</mods:namePart>
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</mods:role>
<mods:namePart>Delphia, Casey M.</mods:namePart>
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<mods:role>
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</mods:role>
<mods:namePart>Griswold, Terry</mods:namePart>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27161</mods:start>
<mods:end>27161</mods:end>
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<mods:url>http://dx.doi.org/10.3897/BDJ.6.e27161</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.6.e27161</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--27161</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:202FF5FC3738837074070055A6185BD6" httpUri="http://treatment.plazi.org/id/202FF5FC3738837074070055A6185BD6" lastPageNumber="27161" pageId="0" pageNumber="27161">
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<paragraph pageId="0" pageNumber="27161">
<taxonomicName authority="Smith, 1854" authorityName="Smith" authorityYear="1854" class="Insecta" family="Apidae" genus="Bombus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bombus (Pyrobombus) vagans" order="Hymenoptera" pageId="0" pageNumber="27161" phylum="Arthropoda" rank="species" species="vagans" subGenus="Pyrobombus">Bombus (Pyrobombus) vagans Smith, 1854</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27161" type="notes">
<paragraph pageId="0" pageNumber="27161">Notes</paragraph>
<paragraph pageId="0" pageNumber="27161">Collected from the Flathead County site (Table 1, Suppl. material 1)</paragraph>
</subSubSection>
</treatment>
</document>