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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.392.6328" ID-PMC="PMC3974435" ID-Pensoft-Pub="1313-2970-392-1" ID-Pensoft-UUID="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" ID-PubMed="24715782" ID-Zenodo-Dep="578044" ModsDocID="1313-2970-392-1" checkinTime="1451246138192" checkinUser="pensoft" docAuthor="Breure, Abraham S. H. &amp; Ablett, Jonathan D." docDate="2014" docId="200A1B4CAE1B7239234BFF7887B06B86" docLanguage="en" docName="ZooKeys 392: 1-367" docOrigin="ZooKeys 392" docPubDate="2014-03-21" docSource="http://dx.doi.org/10.3897/zookeys.392.6328" docTitle="Bulimus cretaceus Pfeiffer 1855" docType="treatment" docVersion="5" id="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" lastPageNumber="44" masterDocId="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" masterDocTitle="Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London" masterLastPageNumber="367" masterPageNumber="1" pageNumber="44" updateTime="1668158078298" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Breure, Abraham S. H.</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">bbreure@xs4all.nl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Natural History Museum, Division of Higher Invertebrates, London, SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-03-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>392</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>367</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.392.6328</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.392.6328</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-392-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFCF5A59FFB1FF93FFF10B43FFAFFFF5</mods:identifier>
<mods:identifier type="Zenodo-Dep">578044</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152052046" LSID="urn:lsid:plazi:treatment:200A1B4CAE1B7239234BFF7887B06B86" httpUri="http://treatment.plazi.org/id/200A1B4CAE1B7239234BFF7887B06B86" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection pageId="43" pageNumber="44" type="nomenclature">
<paragraph pageId="43" pageNumber="44">
<taxonomicName LSID="200A1B4C-AE1B-7239-234B-FF7887B06B86" authority="Pfeiffer, 1855" authorityName="Pfeiffer" authorityYear="1855" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Bulimus cretaceus" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cretaceus">Bulimus cretaceus Pfeiffer, 1855</taxonomicName>
<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Scutalus species. A-B Scutalus baroni (Fulton, 1896), lectotype NHMUK 1896.6.23.1 (H = 12.7) C-F Scutalus cretaceus (Pfeiffer, 1855) C-D lectotype NHMUK 1975388 (H = 35.2) E-F lectotype of Bulimulus (Drymaeus) baroni Fulton, 1897 NHMUK 1897.8.3.173 (H = 34.3) G Scutalus chiletensis Weyrauch, 1967, paratype NHMUK 1975384 (H = 34.2) H Scutalus grandiventris Weyrauch, 1960, paratype NHMUK 1975386 (H = 54.1) I-J Scutalus latecolumellaris Preston, 1909, lectotype NHMUK 1922.2.24.39 (H = 54.0). All enlarged." httpUri="https://binary.pensoft.net/fig/26739" pageId="43" pageNumber="44">Figs 65C-F</figureCitation>
<figureCitation captionStart="Figure L15" captionStartId="F15L" captionText="Figure L 15. Labels. i Bulinus conspersus Sowerby I, 1833 ii Bulimus cotopaxiensis Pfeiffer, 1853 iii Bulimus cretaceus Pfeiffer, 1855 iv Bulinus crichtoni Broderip, 1836 v Helix culminea d'Orbigny, 1835." httpUri="https://binary.pensoft.net/fig/26766" pageId="43" pageNumber="44">, L15iii</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="reference_group">
<paragraph pageId="43" pageNumber="44">
<taxonomicName authorityName="Pfeiffer" authorityYear="1855" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cretaceus">Bulimus cretaceus</taxonomicName>
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Novitates conchologicae" pageId="187" pageNumber="188" pagination="122 - 126" refId="B121" refString="Pfeiffer, L, 1855g. Descriptions of sixteen new species of Helicea, from the collection of H. Cuming Esq. In: Proceedings of the Zoological Society of London.: 122 - 126" title="Descriptions of sixteen new species of Helicea, from the collection of H. Cuming Esq." volumeTitle="Proceedings of the Zoological Society of London." year="1855 g">Pfeiffer 1855g</bibRefCitation>
: 123;
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="181" pageNumber="182" pagination="1 - 215" refId="B15" refString="Breure, ASH, 1979. Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen Leiden 168: 1 - 215" title="Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca)." volume="168" year="1979">Breure 1979</bibRefCitation>
: 83.
</paragraph>
<paragraph pageId="43" pageNumber="44">
<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1855" class="Gastropoda" family="Bulimulidae" genus="Scutalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cretaceus" subGenus="Scutalus">Scutalus (Scutalus) cretaceus</taxonomicName>
(Pfeiffer);
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="181" pageNumber="182" pagination="1 - 255" refId="B14" refString="Breure, ASH, 1978. Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische Verhandelingen Leiden 164: 1 - 255" title="Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda)." volume="164" year="1978">Breure 1978</bibRefCitation>
: 166, figs 277-278, pl. 8 fig. 7 (lectotype designation).
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="type locality">
<paragraph pageId="43" pageNumber="44">Type locality.</paragraph>
<paragraph pageId="43" pageNumber="44">
&quot;Eastern Islands [sic, see remarks] (
<emphasis italics="true" pageId="43" pageNumber="44">Captain Keppell</emphasis>
)&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="label">
<paragraph pageId="43" pageNumber="44">Label.</paragraph>
<paragraph pageId="43" pageNumber="44">
&quot;No locality but I / had the from Cap. Keppell / who got them in the / Eastern Islands but
<normalizedToken originalValue="don’t">don't</normalizedToken>
/ recollect&quot;, taxon label in
<normalizedToken originalValue="Pfeiffer’s">Pfeiffer's</normalizedToken>
handwriting. M.C. label style IV.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="dimensions">
<paragraph pageId="43" pageNumber="44">Dimensions.</paragraph>
<paragraph pageId="43" pageNumber="44">&quot;Long. 36, diam. 18 mill.&quot;; figured specimen herein H 35.2, D 23.2, W 6.2.</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="type material">
<paragraph pageId="43" pageNumber="44">Type material.</paragraph>
<paragraph pageId="43" pageNumber="44">NHMUK 1975388, lectotype; 1975389, one paralectotype, ex Keppell (Cuming coll.).</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="remarks">
<paragraph pageId="43" pageNumber="44">Remarks.</paragraph>
<paragraph pageId="43" pageNumber="44">
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Manual of Conchology" pageId="188" pageNumber="189" pagination="1 - 302" refId="B152" refString="Pilsbry, HA, 1901-1902. Oriental bulimoid Helicidae; Odontostominae; Cerionidae. Manual of Conchology 14: 1 - 302" title="Oriental bulimoid Helicidae; Odontostominae; Cerionidae." volume="14" year="1901 - 1902">Pilsbry (1902 [1901-1902]</bibRefCitation>
: 141) was the first to recognize that this species was from Peru. The type locality is here restricted to Dept. Cajamarca,
<normalizedToken originalValue="Río">Rio</normalizedToken>
Jequetepeque, near Tembladera (
<emphasis bold="true" pageId="43" pageNumber="44">restr. n.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="current systematic position">
<paragraph pageId="43" pageNumber="44">Current systematic position.</paragraph>
<paragraph pageId="43" pageNumber="44">
<taxonomicName authorityName="Tryon" authorityYear="1867" class="Gastropoda" family="Bulimulidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="family">Bulimulidae</taxonomicName>
,
<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1855" class="Gastropoda" family="Bulimulidae" genus="Scutalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cretaceus">
<emphasis italics="true" pageId="43" pageNumber="44">Scutalus cretaceus</emphasis>
</taxonomicName>
(Pfeiffer, 1855).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.2.e1069" ID-PMC="PMC4040401" ID-Pensoft-Pub="1314-2828--1069" ID-PubMed="24891826" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1314-2828--e1069" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Chromosome studies in the aquatic monocots of Myanmar: A brief review with additional records" checkinTime="1553126765086" checkinUser="pensoft" docAuthor="Ito, Yu &amp; Tanaka, Nobuyuki" docDate="2014" docId="200A318D888CFABB93A316BEDD168E92" docLanguage="en" docName="BiodivDatJour 2: e1069" docOrigin="Biodiversity Data Journal 2" docSource="http://dx.doi.org/10.3897/BDJ.2.e1069" docTitle="Potamogeton distinctus A. Benn. x P. nodosus Poir." docType="treatment" docVersion="4" lastPageNumber="1069" masterDocId="FFC9F745FFB17061A804FFC5546CFFEE" masterDocTitle="Chromosome studies in the aquatic monocots of Myanmar: A brief review with additional records" masterLastPageNumber="1069" masterPageNumber="1069" pageNumber="1069" updateTime="1668122537107" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Chromosome studies in the aquatic monocots of Myanmar: A brief review with additional records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ito, Yu</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tanaka, Nobuyuki</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1069</mods:start>
<mods:end>1069</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.2.e1069</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.2.e1069</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--1069</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:200A318D888CFABB93A316BEDD168E92" httpUri="http://treatment.plazi.org/id/200A318D888CFABB93A316BEDD168E92" lastPageNumber="1069" pageId="0" pageNumber="1069">
<subSubSection pageId="0" pageNumber="1069" type="nomenclature">
<paragraph pageId="0" pageNumber="1069">
<taxonomicName class="Liliopsida" family="Potamogetonaceae" genus="Potamogeton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Potamogeton distinctus A. Benn. x P. nodosus Poir." order="Alismatales" pageId="0" pageNumber="1069" phylum="Tracheophyta" rank="species" species="distinctus A. Benn. x P. nodosus Poir.">
Potamogeton distinctus A. Benn.
<normalizedToken originalValue="×">x</normalizedToken>
P. nodosus Poir.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1069" type="materials_examined">
<paragraph pageId="0" pageNumber="1069">Materials</paragraph>
<paragraph pageId="0" pageNumber="1069">
<materialsCitation collectingDate="2008-12-04" collectionCode="MBK, RAF, TI" collectorName="Y. Ito" country="Myanmar" latitude="20.457779" location="Myanmar" longLatPrecision="21" longitude="96.84361" pageId="0" pageNumber="1069" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1069">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="1069">Y. Ito</collectorName>
; Location: country:
<collectingCountry pageId="0" pageNumber="1069">Myanmar</collectingCountry>
; stateProvince: Shan; verbatimLocality: Inle Lake; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="15" value="20.457779">20°27'28&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="east" orientation="longitude" precision="15" value="96.84361">96°50'37&quot;E</geoCoordinate>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="1069" value="2008-12-04">4 Dec 2008</collectingDate>
; Record Level: collectionID: N. Tanaka &amp; al. 080662; institutionCode:
<collectionCode pageId="0" pageNumber="1069">MBK, RAF, TI</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1069" type="notes">
<paragraph pageId="0" pageNumber="1069">Notes</paragraph>
<paragraph pageId="0" pageNumber="1069">Chromosome counts: 2n = 52 (Fig. 5; obtained in this study). The chromosome count for this taxon is new to science.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Linnaeus, Carolus</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:originInfo>
<mods:dateIssued>1758</mods:dateIssued>
<mods:publisher>Laurentius Salvius</mods:publisher>
<mods:place>
<mods:placeTerm>Stockholm</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:location>
<mods:url>https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf</mods:url>
</mods:location>
<mods:classification>book</mods:classification>
<mods:identifier type="ZooBank">2C6327E1-5560-4DB4-B9CA-76A0FA03D975</mods:identifier>
<mods:identifier type="DOI">https://doi.org/10.5962/bhl.title.542</mods:identifier>
<mods:identifier type="Zenodo-Dep">3922206</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3918060" ID-GBIF-Taxon="164810244" ID-Zenodo-Dep="3918060" LSID="urn:lsid:zoobank.org:act:D81D1215-0F71-41F4-B346-29A5B3419D48" httpUri="http://treatment.plazi.org/id/200A5D6245681C60037975BF32C41B9D" lastPageNumber="137" pageNumber="136" sourcePageUrl="https://www.biodiversitylibrary.org/page/727041">
<subSubSection type="nomenclature">
<paragraph pageNumber="136">
<taxonomicName LSID="urn:lsid:zoobank.org:act:D81D1215-0F71-41F4-B346-29A5B3419D48" authorityName="Linnaeus" authorityYear="1758" class="Aves" family="Laridae" genus="Larus" kingdom="Animalia" order="Charadriiformes" phylum="Chordata" rank="species" species="parasiticus" status="spec. nov." zbkClass="Aves" zbkKingdom="Animalia" zbkOrder="Anseres">Larus parasiticus</taxonomicName>
[
<taxonomicNameLabel rank="species">spec. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="136">L. rectricibus duabus intermediis longissimis.</paragraph>
<paragraph pageNumber="136">
Sterna rectricibus maximis nigris.
<emphasis italics="true">Fn. svec.</emphasis>
129.
<emphasis italics="true">It.</emphasis>
Wgot. 182.
<emphasis italics="true">Act. Stockh.</emphasis>
1753.
<emphasis italics="true">p.</emphasis>
291.
</paragraph>
<footnote>
<paragraph pageNumber="136">
Pulli
<emphasis italics="true">primi anni plurumque in his grisei sunt; unde speciei multiplicatio.</emphasis>
</paragraph>
</footnote>
<paragraph pageNumber="137">
<pageBreakToken pageNumber="137">Avis</pageBreakToken>
norvegica Kyuffa.
<emphasis italics="true">Oel. mus.</emphasis>
1.
<emphasis italics="true">s.</emphasis>
11.
<emphasis italics="true">n.</emphasis>
20.
</paragraph>
<paragraph pageNumber="137">
Truen s. Fur.
<emphasis italics="true">Barth. act.</emphasis>
1.
<emphasis italics="true">p.</emphasis>
91.
</paragraph>
<paragraph pageNumber="137">
Coprotheres.
<emphasis italics="true">Raj. av.</emphasis>
127.
<emphasis italics="true">n.</emphasis>
2.
<emphasis italics="true">Martens. spitz.</emphasis>
63.
<emphasis italics="true">t. L. f. D.</emphasis>
</paragraph>
<paragraph pageNumber="137">
Avis arctica.
<emphasis italics="true">Edw. av.</emphasis>
148.
<emphasis italics="true">t.</emphasis>
148. &amp;
<emphasis italics="true">t.</emphasis>
149.
<emphasis italics="true">f.</emphasis>
2.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageNumber="137">
<emphasis italics="true">Habitat intra tropicum</emphasis>
Cancri, Europae, Americae, Asiae.
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="137">
<emphasis italics="true">Piscaturae ineptus agitat congeneres, vomituque ejectum</emphasis>
cibum arripit.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="79C607A05B76572022F19E89734996D0">Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region</mods:title>
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<mods:namePart id="E46E91990112C4ED2A83B1339AF87B16">Tarasov, Sergey I.</mods:namePart>
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<mods:date id="88F8E8F062F641D79602038D8CC22DA7">2010</mods:date>
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<treatment id="200A8781FFD8F35500F091D09137204E" ID-DOI="http://doi.org/10.5281/zenodo.6208722" ID-GBIF-Taxon="119393974" ID-Zenodo-Dep="6208722" LSID="urn:lsid:plazi:treatment:200A8781FFD8F35500F091D09137204E" httpUri="http://treatment.plazi.org/id/200A8781FFD8F35500F091D09137204E" lastPageId="2" lastPageNumber="46" pageId="1" pageNumber="46">
<subSubSection id="E0B9651CFFD8F35600F091D09033242E" box="[151,617,1271,1296]" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="A81C3697FFD8F35600F091D09033242E" blockId="1.[151,1325,1271,1359]" box="[151,617,1271,1296]" pageId="1" pageNumber="46">
<heading id="F35481FBFFD8F35600F091D09033242E" bold="true" box="[151,617,1271,1296]" fontSize="10" level="2" pageId="1" pageNumber="46" reason="2">
<taxonomicName id="6FA34D14FFD8F35600F091D09033242E" ID-CoL="49MKG" authority="Boucomont, 1914" authorityName="Boucomont" authorityYear="1914" box="[151,617,1271,1296]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F35600F091D09033242E" bold="true" box="[151,617,1271,1296]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600F091D093C72431" bold="true" box="[151,413,1272,1296]" italics="true" pageId="1" pageNumber="46">Onthophagus gracilipes</emphasis>
<bibRefCitation id="CC324B66FFD8F35601C391DF9033242E" author="Boucomont" box="[420,617,1271,1295]" pageId="1" pageNumber="46" refString="Boucomont, A. (1914) Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 46, 210 - 243." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600F090309777246E" pageId="1" pageNumber="46" type="reference_group">
<paragraph id="A81C3697FFD8F35600F090309777246E" blockId="1.[151,1325,1271,1359]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="88B311B9FFD8F35600F090309777246E" pageId="1" pageNumber="46">
<taxonomicName id="6FA34D14FFD8F35600F0903090C9240E" ID-CoL="49MKG" authority="Boucomont, 1914: 219" authorityName="Boucomont" authorityPageNumber="219" authorityYear="1914" box="[151,659,1304,1327]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F35600F0903093C2240E" box="[151,408,1304,1327]" italics="true" pageId="1" pageNumber="46">Onthophagus gracilipes</emphasis>
<treatmentCitation id="29021086FFD8F35601F9903090C9240E" author="Boucomont" box="[414,659,1304,1327]" page="219" pageId="1" pageNumber="46" year="1914">
<bibRefCitation id="CC324B66FFD8F35601F99030903B240E" author="Boucomont" box="[414,609,1304,1327]" pageId="1" pageNumber="46" refString="Boucomont, A. (1914) Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 46, 210 - 243." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="77188835FFD8F35602C69030908A240E" box="[673,720,1304,1327]" pageId="1" pageNumber="46">type</typeStatus>
locality: Carin Cheba [
<collectingCountry id="D0B47607FFD8F35603AF9030966D240E" box="[968,1079,1304,1327]" name="Myanmar" pageId="1" pageNumber="46">Myanmar</collectingCountry>
])
<taxonomicName id="6FA34D14FFD8F35600F09010914D246E" ID-CoL="49MDS" authority="Frey, 1971: 95" authorityName="Frey" authorityPageNumber="95" authorityYear="1971" box="[151,791,1336,1359]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis" subGenus="Digithonthophagus">
<emphasis id="9AD7EA85FFD8F35600F09010902F246E" box="[151,629,1336,1359]" italics="true" pageId="1" pageNumber="46">Onthophagus (Digithonthophagus) laosensis</emphasis>
<treatmentCitation id="29021086FFD8F356021B9010914D246E" author="Frey" box="[636,791,1336,1359]" page="95" pageId="1" pageNumber="46" year="1971">
<bibRefCitation id="CC324B66FFD8F356021B901090AB246E" author="Frey" box="[636,753,1336,1359]" pageId="1" pageNumber="46" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey, 1971</bibRefCitation>
: 95
</treatmentCitation>
</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F356037A901F913A246E" box="[797,864,1335,1359]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F356037A901F913A246E" bold="true" box="[797,864,1335,1359]" pageId="1" pageNumber="46">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFD8F356030A901091C7246E" box="[877,925,1336,1359]" pageId="1" pageNumber="46">type</typeStatus>
locality:
<collectingCountry id="D0B47607FFD8F35604669010966D246E" box="[1025,1079,1336,1359]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
,
<collectingCountry id="D0B47607FFD8F35604259010962A246E" box="[1090,1136,1336,1359]" name="Iceland" pageId="1" pageNumber="46">Sala</collectingCountry>
Nam Chan
<collectingCountry id="D0B47607FFD8F35604979010977F246E" box="[1264,1317,1336,1359]" name="China" pageId="1" pageNumber="46">Chin</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600F0905F961726EE" pageId="1" pageNumber="46" type="materials_examined">
<paragraph id="A81C3697FFD8F35600F0905F973024AE" blockId="1.[151,1436,1399,2031]" box="[151,1386,1399,1423]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600F0905F93EB24AE" bold="true" box="[151,433,1399,1423]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F35600F0905F929524AE" box="[151,207,1399,1423]" pageId="1" pageNumber="46">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="77188835FFD8F35601D09050904524AE" box="[439,543,1400,1423]" pageId="1" pageNumber="46" type="lectotype">
<emphasis id="9AD7EA85FFD8F35601D09050904524AE" box="[439,543,1400,1423]" italics="true" pageId="1" pageNumber="46">Lectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F35602259050909724AE" box="[578,717,1400,1423]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F35602259050909724AE" box="[578,717,1400,1423]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F35602259050900224AE" box="[578,600,1400,1423]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD8F35602BC9050916A24AE" LSID="urn:lsid:biocol.org:col:34988" box="[731,816,1400,1423]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="46">MNHN</collectionCode>
)
<emphasis id="9AD7EA85FFD8F3560359905F91B524AE" bold="true" box="[830,1007,1399,1423]" pageId="1" pageNumber="46">here designated</emphasis>
, male bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFD8F35600A290B09696248E" blockId="1.[151,1436,1399,2031]" box="[197,1228,1432,1455]" pageId="1" pageNumber="46">
1. White: Carin Cheba [printed]
<quantity id="6F5B9B72FFD8F356024790B090C5248E" box="[544,671,1432,1455]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="46" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500–1000m</quantity>
[handwritten] L. Fea [printed]
<date id="DC1D1057FFD8F356038B90B0966F248E" box="[1004,1077,1432,1455]" pageId="1" pageNumber="46" value="1987-12">XII-87</date>
[handwritten]
</paragraph>
<paragraph id="A81C3697FFD8F35600A29090901024EE" blockId="1.[151,1436,1399,2031]" box="[197,586,1464,1487]" pageId="1" pageNumber="46">2. White, printed: Mus. Civ. Genova</paragraph>
<paragraph id="A81C3697FFD8F35600A290F091D724CE" blockId="1.[151,1436,1399,2031]" box="[197,909,1496,1519]" pageId="1" pageNumber="46">
3. White, printed: MUSÉUM
<collectionCode id="CEB2AE52FFD8F356026490F0901624CE" box="[515,588,1496,1519]" pageId="1" pageNumber="46">PARIS</collectionCode>
1936
<collectionCode id="CEB2AE52FFD8F35602E890F0908224CE" box="[655,728,1496,1519]" pageId="1" pageNumber="46">COLL</collectionCode>
. BOUCOMONT
</paragraph>
<paragraph id="A81C3697FFD8F35600A290D093E8272E" blockId="1.[151,1436,1399,2031]" box="[197,434,1528,1551]" pageId="1" pageNumber="46">
4. Red, printed:
<typeStatus id="77188835FFD8F356011690D093E8272E" box="[369,434,1528,1551]" pageId="1" pageNumber="46">Typus</typeStatus>
</paragraph>
<paragraph id="A81C3697FFD8F35600A2933091FE2708" blockId="1.[151,1436,1399,2031]" box="[197,932,1559,1583]" pageId="1" pageNumber="46">
5. White: Boucomont det. 191 [printed] 4 onthoph.
<taxonomicName id="6FA34D14FFD8F356028C93309109270E" box="[747,851,1560,1583]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">gracilipes</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F356033E933091D6270E" box="[857,908,1560,1583]" pageId="1" pageNumber="46" rank="species">n.sp.</taxonomicNameLabel>
♂
</paragraph>
<paragraph id="A81C3697FFD8F35600A293109689276E" blockId="1.[151,1436,1399,2031]" box="[197,1235,1592,1615]" pageId="1" pageNumber="46">
6. Orange printed:
<typeStatus id="77188835FFD8F35601E99310905F276E" box="[398,517,1592,1615]" pageId="1" pageNumber="46" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="6FA34D14FFD8F356026C9310961E276E" authority="Boucomont, 1914 S. Tarasov" authorityName="Boucomont, 1914 S. Tarasov" box="[523,1092,1592,1615]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
Onthophagus gracilipes
<bibRefCitation id="CC324B66FFD8F356036893109197276E" author="Boucomont" box="[783,973,1592,1615]" pageId="1" pageNumber="46" refString="Boucomont, A. (1914) Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 46, 210 - 243." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
S. Tarasov
</taxonomicName>
design. 2009
</paragraph>
<paragraph id="A81C3697FFD8F35600A293709132278E" blockId="1.[151,1436,1399,2031]" pageId="1" pageNumber="46">
Also, 1Ψ
<typeStatus id="77188835FFD8F356014D937093E7274E" box="[298,445,1624,1647]" pageId="1" pageNumber="46" type="paralectotype">
<emphasis id="9AD7EA85FFD8F356014D937093E7274E" box="[298,445,1624,1647]" italics="true" pageId="1" pageNumber="46">paralectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F356018793709031274E" box="[480,619,1624,1647]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F356018793709031274E" box="[480,619,1624,1647]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F3560187937093AC274E" box="[480,502,1624,1647]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD8F356021F93709097274E" LSID="urn:lsid:biocol.org:col:34988" box="[632,717,1624,1647]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="46">MNHN</collectionCode>
), bearing labels identical to
<typeStatus id="77188835FFD8F356039D93709606274E" box="[1018,1116,1624,1647]" pageId="1" pageNumber="46" type="lectotype">lectotype</typeStatus>
with the exception of the first label “Carin Ghecu
<quantity id="6F5B9B72FFD8F35601109350905227AE" box="[375,520,1656,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="46" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400m</quantity>
L. Fea
<date id="DC1D1057FFD8F35602079350909E27AE" box="[608,708,1656,1679]" pageId="1" pageNumber="46" value="1988-02" valueMax="1988-03-31" valueMin="1988-02-00">II–III.88</date>
”. The
<typeStatus id="77188835FFD8F356036A9350912927AE" box="[781,883,1656,1679]" pageId="1" pageNumber="46" type="lectotype">lectotype</typeStatus>
is designated in order to preserve the stability of zoological nomenclature according to the article 74.7 (
<bibRefCitation id="CC324B66FFD8F35602BA93B09106278E" author="ICZN" box="[733,860,1688,1711]" pageId="1" pageNumber="46" refString="ICZN (1999) International Code of Zoological Nomenclature (4 th Edition). International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">
<collectionCode id="CEB2AE52FFD8F35602BA93B09147278E" box="[733,797,1688,1711]" pageId="1" pageNumber="46">ICZN</collectionCode>
1999
</bibRefCitation>
).
</paragraph>
<paragraph id="A81C3697FFD8F35600A2939091F627EE" blockId="1.[151,1436,1399,2031]" box="[197,940,1720,1743]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F35600A29390937D27EE" box="[197,295,1720,1743]" pageId="1" pageNumber="46" type="holotype">
<emphasis id="9AD7EA85FFD8F35600A29390937D27EE" box="[197,295,1720,1743]" italics="true" pageId="1" pageNumber="46">Holotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F356012D9390939727EE" box="[330,461,1720,1743]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFD8F356012D9390939727EE" box="[330,461,1720,1743]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356012D9390933A27EE" box="[330,352,1720,1743]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
(NHB), female bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFD8F35600A293F0908527CE" blockId="1.[151,1436,1399,2031]" box="[197,735,1752,1775]" pageId="1" pageNumber="46">
1. White, handwritten:
<collectingCountry id="D0B47607FFD8F35601DC93F093A927CE" box="[443,499,1752,1775]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
:
<collectingCountry id="D0B47607FFD8F356019A93F0907127CE" box="[509,555,1752,1775]" name="Iceland" pageId="1" pageNumber="46">Sala</collectingCountry>
Nam Chan
<collectingCountry id="D0B47607FFD8F35602CC93F0908527CE" box="[683,735,1752,1775]" name="China" pageId="1" pageNumber="46">Chin</collectingCountry>
</paragraph>
<paragraph id="A81C3697FFD8F35600A293D0903F262E" blockId="1.[151,1436,1399,2031]" box="[197,613,1784,1807]" pageId="1" pageNumber="46">
2. White printed: VITALIS
<date id="DC1D1057FFD8F356018A93D0903F262E" box="[493,613,1784,1807]" pageId="1" pageNumber="46" value="1919" valueMax="1920">1919–1920</date>
</paragraph>
<paragraph id="A81C3697FFD8F35600A292309725260E" blockId="1.[151,1436,1399,2031]" box="[197,1407,1815,1839]" pageId="1" pageNumber="46">
3. White:
<taxonomicName id="6FA34D14FFD8F3560148923090CE260E" authority="(Digitonthophagus)" baseAuthorityName="Digitonthophagus" box="[303,660,1816,1839]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">Onthophagus (Digitonthophagus)</taxonomicName>
n.sp. Ψ [handwritten] Dr. V. Balthasar det. [handwritten] 60. [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A2921093EE266E" blockId="1.[151,1436,1399,2031]" box="[197,436,1848,1871]" pageId="1" pageNumber="46">
4. Red, printed:
<collectionCode id="CEB2AE52FFD8F3560116921093EE266E" box="[369,436,1848,1871]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F3560116921093EE266E" box="[369,436,1848,1871]" pageId="1" pageNumber="46">TYPE</typeStatus>
</collectionCode>
</paragraph>
<paragraph id="A81C3697FFD8F35600A29270937926AE" blockId="1.[151,1436,1399,2031]" pageId="1" pageNumber="46">
5. White:
<collectionCode id="CEB2AE52FFD8F356015792709328264E" box="[304,370,1880,1903]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F356015792709328264E" box="[304,370,1880,1903]" pageId="1" pageNumber="46">TYPE</typeStatus>
</collectionCode>
Ψ [handwritten]
<taxonomicName id="6FA34D14FFD8F356024B927090E0264E" box="[556,698,1880,1903]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
[printed]
<taxonomicName id="6FA34D14FFD8F356034092709614264E" box="[807,1102,1880,1903]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">Digitonthophagus laosensis</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F3560432927096D2264E" box="[1109,1160,1880,1903]" pageId="1" pageNumber="46" rank="species">n.sp.</taxonomicNameLabel>
[handwritten] det.G.Frey, 196 [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A292B09743268E" blockId="1.[151,1436,1399,2031]" box="[197,1305,1944,1967]" pageId="1" pageNumber="46">
6. Orange:
<typeStatus id="77188835FFD8F356015B92B09393268E" box="[316,457,1944,1967]" pageId="1" pageNumber="46" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="6FA34D14FFD8F35601A892B09166268E" authority="Frey, 1971" authorityName="Frey" authorityYear="1971" box="[463,828,1944,1967]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis">
Onthophagus laosensis
<bibRefCitation id="CC324B66FFD8F35602AC92B09166268E" author="Frey" box="[715,828,1944,1967]" pageId="1" pageNumber="46" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey, 1971</bibRefCitation>
</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A29290961726EE" blockId="1.[151,1436,1399,2031]" box="[197,1101,1976,1999]" pageId="1" pageNumber="46">
7. White:
<taxonomicName id="6FA34D14FFD8F35601489290902B26EE" authority="Bouc." authorityName="Bouc." box="[303,625,1976,1999]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">Onthophagus gracilipes Bouc.</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35500A292FF9137204E" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46" type="discussion">
<paragraph id="A81C3697FFD8F35500A292FF9137204E" blockId="1.[151,1436,1399,2031]" lastBlockId="2.[151,1437,152,367]" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600A292FF936926CE" bold="true" box="[197,307,2007,2031]" pageId="1" pageNumber="46">Remarks.</emphasis>
It seems that
<bibRefCitation id="CC324B66FFD8F35601AF92F0901D26CE" author="Frey" box="[456,583,2008,2031]" pageId="1" pageNumber="46" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey (1971)</bibRefCitation>
did not thoroughly examine the description of
<taxonomicName id="6FA34D14FFD8F356042792F0969626CE" box="[1088,1228,2008,2031]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F356042792F0969626CE" box="[1088,1228,2008,2031]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356042792F0960C26CE" box="[1088,1110,2008,2031]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
and did not see the
<typeStatus id="77188835FFDBF35500F095B0929F218E" box="[151,197,152,175]" pageId="2" pageNumber="47">type</typeStatus>
series of the former. In the original description he compares
<taxonomicName id="6FA34D14FFDBF355032D95B09191218E" box="[842,971,152,175]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFDBF355032D95B09191218E" box="[842,971,152,175]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355032D95B0913A218E" box="[842,864,152,175]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
with
<taxonomicName id="6FA34D14FFDBF355046095B0976E218E" authority="Balthasar, 1933" authorityName="Balthasar" authorityYear="1933" box="[1031,1332,152,175]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="troniceki">
<emphasis id="9AD7EA85FFDBF355046095B096DC218E" box="[1031,1158,152,175]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355046095B09647218E" box="[1031,1053,152,175]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. troniceki
</emphasis>
Balthasar, 1933
</taxonomicName>
. The
<typeStatus id="77188835FFDBF355050995B097C6218E" box="[1390,1436,152,175]" pageId="2" pageNumber="47">type</typeStatus>
series of Frey’s
<taxonomicName id="6FA34D14FFDBF35501589590939821EE" box="[319,450,184,207]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFDBF35501589590939821EE" box="[319,450,184,207]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35501589590930F21EE" box="[319,341,184,207]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
includes only a female
<typeStatus id="77188835FFDBF35502DA9590914721EE" box="[701,797,184,207]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
, however that was not an obstacle for identification because the
<typeStatus id="77188835FFDBF35500D995F092B621CE" box="[190,236,216,239]" pageId="2" pageNumber="47">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFDBF355013395F093BA21CE" box="[340,480,216,239]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFDBF355013395F093BA21CE" box="[340,480,216,239]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355013395F0933021CE" box="[340,362,216,239]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
includes a female. I synonymize these two names because the
<typeStatus id="77188835FFDBF35504E495F096EB21CE" box="[1155,1201,216,239]" pageId="2" pageNumber="47">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFDBF355057E95F097C621CE" box="[1305,1436,216,239]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFDBF355057E95F097C621CE" box="[1305,1436,216,239]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355057E95F0977521CE" box="[1305,1327,216,239]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="6FA34D14FFDBF35500A295D0930B202E" box="[197,337,248,271]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="parvidens">
<emphasis id="9AD7EA85FFDBF35500A295D0930B202E" box="[197,337,248,271]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35500A295D09281202E" box="[197,219,248,271]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. parvidens
</emphasis>
</taxonomicName>
are conspecific based on their same principle diagnostic character states of external morphology: the unique color pattern of the elytra and body, the unique groundplan of head, and the identical pattern of pronotum punctation. Additionally I have examined large material of
<taxonomicName id="6FA34D14FFDBF355036B941091CD206E" box="[780,919,312,335]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFDBF355036B941091CD206E" box="[780,919,312,335]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355036B94109178206E" box="[780,802,312,335]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
from other localities in Indochina and these also match the two series of both names in the characters detailed above.
</paragraph>
</subSubSection>
</treatment>
</document>

277
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<subSubSection id="E0B9651CFFD8F35600F0943093D8204E" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="A81C3697FFD8F35600F09430903B200E" blockId="1.[151,1436,279,495]" box="[151,609,279,304]" pageId="1" pageNumber="46">
<heading id="F35481FBFFD8F35600F09430903B200E" bold="true" box="[151,609,279,304]" fontSize="10" level="2" pageId="1" pageNumber="46" reason="2">
<taxonomicName id="6FA34D14FFD8F35600F09430903B200E" ID-CoL="7XMM7" authority="Boucomont, 1921" authorityName="Boucomont" authorityYear="1921" box="[151,609,279,304]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9AD7EA85FFD8F35600F09430903B200E" bold="true" box="[151,609,279,304]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600F0943093CC2011" bold="true" box="[151,406,280,304]" italics="true" pageId="1" pageNumber="46">Onthophagus cognatus</emphasis>
<bibRefCitation id="CC324B66FFD8F35601FA943F903B200E" author="Boucomont" box="[413,609,279,303]" pageId="1" pageNumber="46" refString="Boucomont, A. &amp; Gillet, J. (1921) Faune entomologique de l'Indochine francaise, volume 4. Famille Scarabaeidae Laparosticti (Coleopteres). Imprimerie Nouvelle Albert Portail, Saigon, 76 pp." type="book" year="1921">Boucomont, 1921</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A81C3697FFD8F35600F0941093D8204E" blockId="1.[151,1436,279,495]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="88B311B9FFD8F35600F0941093D8204E" pageId="1" pageNumber="46">
<taxonomicName id="6FA34D14FFD8F35600F094109042206E" ID-CoL="7XMM7" authority="Boucomont" authorityName="Boucomont" box="[151,536,312,335]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9AD7EA85FFD8F35600F0941093CB206E" box="[151,401,312,335]" italics="true" pageId="1" pageNumber="46">Onthophagus cognatus</emphasis>
Boucomont
</taxonomicName>
(in
<treatmentCitation id="29021086FFD8F356022C941091F0206E" author="Boucomont" box="[587,938,312,335]" page="30" pageId="1" pageNumber="46" year="1921">
<bibRefCitation id="CC324B66FFD8F356022C941091DB206E" author="Boucomont" box="[587,897,312,335]" pageId="1" pageNumber="46" refString="Boucomont, A. &amp; Gillet, J. (1921) Faune entomologique de l'Indochine francaise, volume 4. Famille Scarabaeidae Laparosticti (Coleopteres). Imprimerie Nouvelle Albert Portail, Saigon, 76 pp." type="book" year="1921">Boucomont &amp; Gillet), 1921</bibRefCitation>
: 30
</treatmentCitation>
(
<typeStatus id="77188835FFD8F35603DC941091B0206E" box="[955,1002,312,335]" pageId="1" pageNumber="46">type</typeStatus>
locality:
<collectingCountry id="D0B47607FFD8F3560430941096D4206E" box="[1111,1166,312,335]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
, prov. Tran Ninh [
<collectingCountry id="D0B47607FFD8F3560538941097C2206E" box="[1375,1432,312,335]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
, Xieng Khouang])
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600F09450901420CE" pageId="1" pageNumber="46" type="reference_group">
<paragraph id="A81C3697FFD8F35600F09450965B20AE" blockId="1.[151,1436,279,495]" box="[151,1025,375,399]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="88B311B9FFD8F35600F09450965B20AE" box="[151,1025,375,399]" pageId="1" pageNumber="46">
<taxonomicName id="6FA34D14FFD8F35600F09450900A20AE" ID-CoL="7XNDW" authority="Arrow, 1931: 188" authorityName="Arrow" authorityPageNumber="188" authorityYear="1931" box="[151,592,376,399]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="9AD7EA85FFD8F35600F0945093D720AE" box="[151,397,376,399]" italics="true" pageId="1" pageNumber="46">Onthophagus jucundus</emphasis>
<treatmentCitation id="29021086FFD8F35601F39450900A20AE" author="Arrow" box="[404,592,376,399]" page="188" pageId="1" pageNumber="46" year="1931">
<bibRefCitation id="CC324B66FFD8F35601F39450904420AE" author="Arrow" box="[404,542,376,399]" pageId="1" pageNumber="46" refString="Arrow, G. J. (1931) The Fauna of British India III, Coprinae. Taylor &amp; Francis, London, 428 pp." type="book" year="1931">Arrow, 1931</bibRefCitation>
: 188
</treatmentCitation>
</taxonomicName>
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<emphasis id="9AD7EA85FFD8F3560230945F90C020AE" bold="true" box="[599,666,375,399]" pageId="1" pageNumber="46">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFD8F35602CF9450908D20AE" box="[680,727,376,399]" pageId="1" pageNumber="46">type</typeStatus>
locality:
<collectingCountry id="D0B47607FFD8F356035B9450912920AE" box="[828,883,376,399]" name="India" pageId="1" pageNumber="46">India</collectingCountry>
Or Manipur)
</treatmentCitationGroup>
</paragraph>
<paragraph id="A81C3697FFD8F35600F094B091EC208E" blockId="1.[151,1436,279,495]" box="[151,950,408,431]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="88B311B9FFD8F35600F094B091EC208E" box="[151,950,408,431]" pageId="1" pageNumber="46">
<taxonomicName id="6FA34D14FFD8F35600F094B09387208E" ID-CoL="7XNDW" authority="Arrow" authorityName="Arrow" box="[151,477,408,431]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="9AD7EA85FFD8F35600F094B093D7208E" box="[151,397,408,431]" italics="true" pageId="1" pageNumber="46">Onthophagus jucundus</emphasis>
Arrow
</taxonomicName>
;
<treatmentCitation id="29021086FFD8F356018094B090E0208E" author="Bacchus" box="[487,698,408,431]" page="104" pageId="1" pageNumber="46" year="1978">
<bibRefCitation id="CC324B66FFD8F356018094B090D2208E" author="Bacchus" box="[487,648,408,431]" pageId="1" pageNumber="46" refString="Bacchus, M. E. (1978) A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History), Entomology Series, 37, 97 - 115." type="journal article" year="1978">Bacchus, 1978</bibRefCitation>
: 104
</treatmentCitation>
(
<typeStatus id="77188835FFD8F35602AF94B09176208E" box="[712,812,408,431]" pageId="1" pageNumber="46" type="lectotype">lectotype</typeStatus>
designation)
</treatmentCitationGroup>
</paragraph>
<paragraph id="A81C3697FFD8F35600F09490901420CE" blockId="1.[151,1436,279,495]" pageId="1" pageNumber="46">
<taxonomicName id="6FA34D14FFD8F35600F09490967620EE" ID-CoL="7XMK5" authority="Masumoto, Ochi &amp; Hanboonong, 2008" authorityName="Masumoto, Ochi &amp; Hanboonong" authorityYear="2008" box="[151,1068,440,463]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="chulapornus" subGenus="Serrophorus">
<emphasis id="9AD7EA85FFD8F35600F09490900420EE" box="[151,606,440,463]" italics="true" pageId="1" pageNumber="46">Onthophagus (Serrophorus) chulapornus</emphasis>
Masumoto,
<bibRefCitation id="CC324B66FFD8F35602949490967620EE" author="Ochi" box="[755,1068,440,463]" pageId="1" pageNumber="46" refString="Masumoto, K., Ochi, T. &amp; Hanboonsong, Y. (2008) New species of the genus Onthophagus (Coleoptera: Scarabaeidae) from Thailand, part 4. Descriptions of five new species, revival of a suppressed specific name and seven new distributional records. Entomological Review of Japan, 63, 155 - 169." type="journal article" year="2008">Ochi &amp; Hanboonong, 2008</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F356045C949F962720EE" box="[1083,1149,439,463]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F356045C949F962720EE" bold="true" box="[1083,1149,439,463]" pageId="1" pageNumber="46">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFD8F35604F49490969820EE" box="[1171,1218,440,463]" pageId="1" pageNumber="46">type</typeStatus>
locality:
<collectingCountry id="D0B47607FFD8F3560550949097C220EE" box="[1335,1432,440,463]" name="Thailand" pageId="1" pageNumber="46">Thailand</collectingCountry>
, Chaiyaphum Prov., Chulaporn Dam)
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600F0973F9609228E" pageId="1" pageNumber="46" type="materials_examined">
<paragraph id="A81C3697FFD8F35600F0973F9655236E" blockId="1.[151,1437,535,1199]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600F0973F93E8230E" bold="true" box="[151,434,535,559]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F35600F0973F9295230E" box="[151,207,535,559]" pageId="1" pageNumber="46">Type</typeStatus>
material examined.
</emphasis>
Two
<typeStatus id="77188835FFD8F356018897309013230E" box="[495,585,536,559]" pageId="1" pageNumber="46" type="syntype">
<emphasis id="9AD7EA85FFD8F356018897309013230E" box="[495,585,536,559]" italics="true" pageId="1" pageNumber="46">syntypes</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F3560209973090B5230E" box="[622,751,536,559]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9AD7EA85FFD8F3560209973090B5230E" box="[622,751,536,559]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F3560209973090DE230E" box="[622,644,536,559]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. cognatus
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD8F35602999730910F230E" LSID="urn:lsid:biocol.org:col:34988" box="[766,853,536,559]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="46">MNHN</collectionCode>
), male and female bearing geographical label “
<collectingCountry id="D0B47607FFD8F3560529973097C2230E" box="[1358,1432,536,559]" name="Laos" pageId="1" pageNumber="46">LAOS</collectingCountry>
, prov. Tran Ninh” and indicated by A. Boucomont as “
<taxonomicName id="6FA34D14FFD8F35602B397109190236E" box="[724,970,568,591]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">Onthophagus cognatus</taxonomicName>
n.sp”.
</paragraph>
<paragraph id="A81C3697FFD8F35600A2977091FB234E" blockId="1.[151,1437,535,1199]" box="[197,929,600,623]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F35600A297709377234E" box="[197,301,600,623]" pageId="1" pageNumber="46" type="lectotype">
<emphasis id="9AD7EA85FFD8F35600A297709377234E" box="[197,301,600,623]" italics="true" pageId="1" pageNumber="46">Lectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F35601379770938B234E" box="[336,465,600,623]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="9AD7EA85FFD8F35601379770938B234E" box="[336,465,600,623]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F35601379770933C234E" box="[336,358,600,623]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. jucundus
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD8F35601B89770906E234E" box="[479,564,600,623]" pageId="1" pageNumber="46">BMNH</collectionCode>
), female bearing following labels:
</paragraph>
<paragraph id="A81C3697FFD8F35600A29750905D23AE" blockId="1.[151,1437,535,1199]" box="[197,519,632,655]" pageId="1" pageNumber="46">
1. Round label:
<typeStatus id="77188835FFD8F35601089750905D23AE" box="[367,519,632,655]" pageId="1" pageNumber="46" type="lectotype">LECTOTYPE</typeStatus>
</paragraph>
<paragraph id="A81C3697FFD8F35600A297B09385238E" blockId="1.[151,1437,535,1199]" box="[197,479,664,687]" pageId="1" pageNumber="46">2. White, printed: Doherty</paragraph>
<paragraph id="A81C3697FFD8F35600A29790915823EE" blockId="1.[151,1437,535,1199]" box="[197,770,696,719]" pageId="1" pageNumber="46">3. Yellow, handwritten: [first character illegible] 4311</paragraph>
<paragraph id="A81C3697FFD8F35600A297F0901E23CE" blockId="1.[151,1437,535,1199]" box="[197,580,728,751]" pageId="1" pageNumber="46">
4. White, printed:
<collectingCountry id="D0B47607FFD8F35601E097F093E423CE" box="[391,446,728,751]" name="India" pageId="1" pageNumber="46">India</collectingCountry>
Or Manipur
</paragraph>
<paragraph id="A81C3697FFD8F35600A297D0900F222E" blockId="1.[151,1437,535,1199]" box="[197,597,760,783]" pageId="1" pageNumber="46">5. White, printed: Fry Coll 1905-100.</paragraph>
<paragraph id="A81C3697FFD8F35600A29630916A220E" blockId="1.[151,1437,535,1199]" box="[197,816,792,815]" pageId="1" pageNumber="46">
6. White, handwritten:
<taxonomicName id="6FA34D14FFD8F35601DC963090EE220E" box="[443,692,792,815]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">Onthophagus jucundus</taxonomicName>
.
<typeStatus id="77188835FFD8F35602D9963090B6220E" box="[702,748,792,815]" pageId="1" pageNumber="46">type</typeStatus>
arrow
</paragraph>
<paragraph id="A81C3697FFD8F35600A2961093C9224E" blockId="1.[151,1437,535,1199]" pageId="1" pageNumber="46">
7. White:
<taxonomicName id="6FA34D14FFD8F356015E9610938B226E" box="[313,465,824,847]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
Ψ
<taxonomicName id="6FA34D14FFD8F3560192961090F2226E" authority="Arrow" authorityName="Arrow" box="[501,680,824,847]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">jucundus arrow</taxonomicName>
[handwritten], M.E. Bacchus det 1975 [printed] 5 [handwritten].
<typeStatus id="77188835FFD8F35600F096709375224E" box="[151,303,856,879]" pageId="1" pageNumber="46" type="lectotype">LECTOTYPE</typeStatus>
[printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A29650961E22AE" blockId="1.[151,1437,535,1199]" box="[197,1092,888,911]" pageId="1" pageNumber="46">
8. White:
<taxonomicName id="6FA34D14FFD8F35601489650903322AE" authority="Bouc." authorityName="Bouc." box="[303,617,888,911]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">Onthophagus cognatus Bouc.</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A296B09609228E" blockId="1.[151,1437,535,1199]" box="[197,1107,919,943]" pageId="1" pageNumber="46">
Also, 1Ψ
<typeStatus id="77188835FFD8F356014D96B093E7228E" box="[298,445,920,943]" pageId="1" pageNumber="46" type="paralectotype">
<emphasis id="9AD7EA85FFD8F356014D96B093E7228E" box="[298,445,920,943]" italics="true" pageId="1" pageNumber="46">paralectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F356018796B0903B228E" box="[480,609,920,943]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="9AD7EA85FFD8F356018796B0903B228E" box="[480,609,920,943]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356018796B093AC228E" box="[480,502,920,943]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. jucundus
</emphasis>
</taxonomicName>
bearing geographical label “Birmah Momeit”.
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600A2969F9613258E" pageId="1" pageNumber="46" type="discussion">
<paragraph id="A81C3697FFD8F35600A2969F9613258E" blockId="1.[151,1437,535,1199]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600A2969F936022EE" bold="true" box="[197,314,951,975]" pageId="1" pageNumber="46">Remarks.</emphasis>
I have examined large material of
<taxonomicName id="6FA34D14FFD8F35602A99690910322EE" box="[718,857,952,975]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9AD7EA85FFD8F35602A99690910322EE" box="[718,857,952,975]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F35602A9969090BF22EE" box="[718,741,952,975]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. cognatus
</emphasis>
</taxonomicName>
and similar species from Indochina.
<taxonomicName id="6FA34D14FFD8F3560563969092A222CE" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9AD7EA85FFD8F3560563969092A222CE" italics="true" pageId="1" pageNumber="46">Onthophagus cognatus</emphasis>
</taxonomicName>
is widespread in Indochina and can be easily separated from similar species by external characters and structure of the aedeagus. Although I have not examined
<typeStatus id="77188835FFD8F35602C296D0908E252E" box="[677,724,1016,1039]" pageId="1" pageNumber="46">type</typeStatus>
specimens of
<taxonomicName id="6FA34D14FFD8F356031596D09296250E" authority="Masumoto, Ochi &amp; Hanboonsong, 2008" authorityName="Masumoto, Ochi &amp; Hanboonsong" authorityYear="2008" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="chulapornus">
<emphasis id="9AD7EA85FFD8F356031596D09646252E" box="[882,1052,1016,1039]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356031596D091D2252E" box="[882,904,1016,1039]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. chulapornus
</emphasis>
<bibRefCitation id="CC324B66FFD8F356044396D09296250E" author="Masumoto" pageId="1" pageNumber="46" refString="Masumoto, K., Ochi, T. &amp; Hanboonsong, Y. (2008) New species of the genus Onthophagus (Coleoptera: Scarabaeidae) from Thailand, part 4. Descriptions of five new species, revival of a suppressed specific name and seven new distributional records. Entomological Review of Japan, 63, 155 - 169." type="journal article" year="2008">Masumoto, Ochi &amp; Hanboonsong, 2008</bibRefCitation>
</taxonomicName>
, the original description of this species is well illustrated with a photograph of the
<typeStatus id="77188835FFD8F3560429913096F6250E" box="[1102,1196,1048,1071]" pageId="1" pageNumber="46" type="holotype">holotype</typeStatus>
and illustration of the aedeagus. These illustrations match the character states seen in
<taxonomicName id="6FA34D14FFD8F356032B9110918A256E" box="[844,976,1080,1103]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<collectingCountry id="D0B47607FFD8F356032B91109105256E" box="[844,863,1080,1103]" name="Iceland" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F356032B91109105256E" box="[844,863,1080,1103]" italics="true" pageId="1" pageNumber="46">O</emphasis>
</collectingCountry>
.
<emphasis id="9AD7EA85FFD8F35603099110918A256E" box="[878,976,1080,1103]" italics="true" pageId="1" pageNumber="46">cognatus</emphasis>
</taxonomicName>
and lead me to conclude that these names should be synonymized. It seems that
<taxonomicName id="6FA34D14FFD8F356024B91709088254E" box="[556,722,1112,1135]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="chulapornus">
<emphasis id="9AD7EA85FFD8F356024B91709088254E" box="[556,722,1112,1135]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356024B91709018254E" box="[556,578,1112,1135]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. chulapornus
</emphasis>
</taxonomicName>
was described as a new species simply because the authors did not know the identity of
<taxonomicName id="6FA34D14FFD8F35601199150905925AE" box="[382,515,1144,1167]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9AD7EA85FFD8F35601199150905925AE" box="[382,515,1144,1167]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F3560119915093CF25AE" box="[382,405,1144,1167]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. cognatus
</emphasis>
</taxonomicName>
. Likewise, the
<typeStatus id="77188835FFD8F35602CA9150914B25AE" box="[685,785,1144,1167]" pageId="1" pageNumber="46" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F3560350915091E725AE" box="[823,957,1144,1167]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="jucundus">
<collectingCountry id="D0B47607FFD8F35603509150911025AE" box="[823,842,1144,1167]" name="Iceland" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35603509150911025AE" box="[823,842,1144,1167]" italics="true" pageId="1" pageNumber="46">O</emphasis>
</collectingCountry>
.
<emphasis id="9AD7EA85FFD8F356033D915091E725AE" box="[858,957,1144,1167]" italics="true" pageId="1" pageNumber="46">jucundus</emphasis>
</taxonomicName>
was examined and matched
<taxonomicName id="6FA34D14FFD8F356049D9150972525AE" box="[1274,1407,1144,1167]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cognatus">
<collectingCountry id="D0B47607FFD8F356049D9150975725AE" box="[1274,1293,1144,1167]" name="Iceland" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F356049D9150975725AE" box="[1274,1293,1144,1167]" italics="true" pageId="1" pageNumber="46">O</emphasis>
</collectingCountry>
.
<emphasis id="9AD7EA85FFD8F356057B9150972525AE" box="[1308,1407,1144,1167]" italics="true" pageId="1" pageNumber="46">cognatus</emphasis>
</taxonomicName>
in diagnostic external and aedeagal character states and these names are here synonymized.
</paragraph>
</subSubSection>
</treatment>
</document>

255
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<treatment id="200A8781FFD9F35600F09073977E21EE" ID-DOI="http://doi.org/10.5281/zenodo.6208717" ID-GBIF-Taxon="119393972" ID-Zenodo-Dep="6208717" LSID="urn:lsid:plazi:treatment:200A8781FFD9F35600F09073977E21EE" httpUri="http://treatment.plazi.org/id/200A8781FFD9F35600F09073977E21EE" lastPageId="1" lastPageNumber="45" pageId="0" pageNumber="45">
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<paragraph id="A81C3697FFD9F35700F0907390672452" blockId="0.[151,1120,1371,1459]" box="[151,573,1371,1395]" pageId="0" pageNumber="45">
<heading id="F35481FBFFD9F35700F0907390672452" bold="true" box="[151,573,1371,1395]" fontSize="10" level="2" pageId="0" pageNumber="45" reason="2">
<taxonomicName id="6FA34D14FFD9F35700F0907390672452" ID-CoL="49MHB" authority="Paulian, 1931" authorityName="Paulian" authorityYear="1931" box="[151,573,1371,1395]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="9AD7EA85FFD9F35700F0907390672452" bold="true" box="[151,573,1371,1395]" pageId="0" pageNumber="45">
<emphasis id="9AD7EA85FFD9F35700F0907393C62452" bold="true" box="[151,412,1371,1395]" italics="true" pageId="0" pageNumber="45">Onthophagus clermonti</emphasis>
<bibRefCitation id="CC324B66FFD9F35701C4907390672452" author="Paulian" box="[419,573,1371,1395]" pageId="0" pageNumber="45" refString="Paulian, R. (1931) Coprophages nouveaux pour la faune de Hoa-Binh (Tonkin). Bulletin de la Societe entomologique de France, 36, 270 - 272." type="journal article" year="1931">Paulian, 1931</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD9F35700F09054963A2492" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="A81C3697FFD9F35700F09054963A2492" blockId="0.[151,1120,1371,1459]" pageId="0" pageNumber="45">
<treatmentCitationGroup id="88B311B9FFD9F35700F09054963A2492" pageId="0" pageNumber="45">
<taxonomicName id="6FA34D14FFD9F35700F09054903924B2" ID-CoL="49MHB" authority="Paulian, 1931: 271" authorityName="Paulian" authorityPageNumber="271" authorityYear="1931" box="[151,611,1404,1427]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="9AD7EA85FFD9F35700F0905493C924B2" box="[151,403,1404,1427]" italics="true" pageId="0" pageNumber="45">Onthophagus clermonti</emphasis>
<treatmentCitation id="29021086FFD9F35701FD9054903924B2" author="Paulian" box="[410,611,1404,1427]" page="271" pageId="0" pageNumber="45" year="1931">
<bibRefCitation id="CC324B66FFD9F35701FD9054906A24B2" author="Paulian" box="[410,560,1404,1427]" pageId="0" pageNumber="45" refString="Paulian, R. (1931) Coprophages nouveaux pour la faune de Hoa-Binh (Tonkin). Bulletin de la Societe entomologique de France, 36, 270 - 272." type="journal article" year="1931">Paulian, 1931</bibRefCitation>
: 271
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="77188835FFD9F3570216905490FA24B2" box="[625,672,1404,1427]" pageId="0" pageNumber="45">type</typeStatus>
locality: Hoa-Binh,
<collectingCountry id="D0B47607FFD9F357031C9054919C24B2" box="[891,966,1404,1427]" name="Vietnam" pageId="0" pageNumber="45">Tonkin</collectingCountry>
[
<collectingCountry id="D0B47607FFD9F35703B49054966F24B2" box="[979,1077,1404,1427]" name="Vietnam" pageId="0" pageNumber="45">Vietnam</collectingCountry>
])
<taxonomicName id="6FA34D14FFD9F35700F090B490622492" ID-CoL="49PKK" authority="Frey, 1971: 98" authorityName="Frey" authorityPageNumber="98" authorityYear="1971" box="[151,568,1436,1459]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="parvidens">
<emphasis id="9AD7EA85FFD9F35700F090B493CC2492" box="[151,406,1436,1459]" italics="true" pageId="0" pageNumber="45">Onthophagus parvidens</emphasis>
<treatmentCitation id="29021086FFD9F35701FA90B490622492" author="Frey" box="[413,568,1436,1459]" page="98" pageId="0" pageNumber="45" year="1971">
<bibRefCitation id="CC324B66FFD9F35701FA90B490482492" author="Frey" box="[413,530,1436,1459]" pageId="0" pageNumber="45" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey, 1971</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD9F357025990B390DB2492" box="[574,641,1435,1459]" pageId="0" pageNumber="45">
<emphasis id="9AD7EA85FFD9F357025990B390DB2492" bold="true" box="[574,641,1435,1459]" pageId="0" pageNumber="45">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFD9F35702E890B490E42492" box="[655,702,1436,1459]" pageId="0" pageNumber="45">type</typeStatus>
locality: Hoa-Binh,
<collectingCountry id="D0B47607FFD9F35703FE90B491BE2492" box="[921,996,1436,1459]" name="Vietnam" pageId="0" pageNumber="45">Tonkin</collectingCountry>
[
<collectingCountry id="D0B47607FFD9F357039690B496082492" box="[1009,1106,1436,1459]" name="Vietnam" pageId="0" pageNumber="45">Vietnam</collectingCountry>
])
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD9F35700F090F3969E2672" pageId="0" pageNumber="45" type="materials_examined">
<paragraph id="A81C3697FFD9F35700F090F396E624D2" blockId="0.[151,1436,1499,2035]" box="[151,1212,1499,1523]" pageId="0" pageNumber="45">
<emphasis id="9AD7EA85FFD9F35700F090F393EB24D2" bold="true" box="[151,433,1499,1523]" pageId="0" pageNumber="45">
<typeStatus id="77188835FFD9F35700F090F3929524D2" box="[151,207,1499,1523]" pageId="0" pageNumber="45">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="77188835FFD9F35701D090F4904324D2" box="[439,537,1500,1523]" pageId="0" pageNumber="45" type="holotype">
<emphasis id="9AD7EA85FFD9F35701D090F4904324D2" box="[439,537,1500,1523]" italics="true" pageId="0" pageNumber="45">Holotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD9F357025B90F4909924D2" box="[572,707,1500,1523]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="9AD7EA85FFD9F357025B90F4909924D2" box="[572,707,1500,1523]" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F357025B90F4900824D2" box="[572,594,1500,1523]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. clermonti
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD9F35702B690F4917224D2" LSID="urn:lsid:biocol.org:col:34988" box="[721,808,1500,1523]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="45">MNHN</collectionCode>
), female bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFD9F35700A290D491E82732" blockId="0.[151,1436,1499,2035]" box="[197,946,1532,1555]" pageId="0" pageNumber="45">
1. White, printed: Hoa-Binh (
<collectingCountry id="D0B47607FFD9F357019990D490172732" box="[510,589,1532,1555]" name="Vietnam" pageId="0" pageNumber="45">Tonkin</collectingCountry>
) (A.de Cooman) Coll.J.Clermont
</paragraph>
<paragraph id="A81C3697FFD9F35700A2933493EE2712" blockId="0.[151,1436,1499,2035]" box="[197,436,1564,1587]" pageId="0" pageNumber="45">
2. Red, printed:
<collectionCode id="CEB2AE52FFD9F3570116933493EE2712" box="[369,436,1564,1587]" pageId="0" pageNumber="45">
<typeStatus id="77188835FFD9F3570116933493EE2712" box="[369,436,1564,1587]" pageId="0" pageNumber="45">TYPE</typeStatus>
</collectionCode>
</paragraph>
<paragraph id="A81C3697FFD9F35700A2931490062772" blockId="0.[151,1436,1499,2035]" box="[197,604,1596,1619]" pageId="0" pageNumber="45">
3. White, handwritten:
<taxonomicName id="6FA34D14FFD9F35701DC9314907D2772" box="[443,551,1596,1619]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">Clermonti</taxonomicName>
<typeStatus id="77188835FFD9F3570249931490062772" box="[558,604,1596,1619]" pageId="0" pageNumber="45">type</typeStatus>
</paragraph>
<paragraph id="A81C3697FFD9F35700A2937496E42752" blockId="0.[151,1436,1499,2035]" box="[197,1214,1628,1651]" pageId="0" pageNumber="45">
4. Orange:
<typeStatus id="77188835FFD9F357015B937493932752" box="[316,457,1628,1651]" pageId="0" pageNumber="45" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="6FA34D14FFD9F35701A8937490F52752" authority="Paulian" authorityName="Paulian" box="[463,687,1628,1651]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<collectingCountry id="D0B47607FFD9F35701A8937493BF2752" box="[463,485,1628,1651]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. clermonti Paulian
</taxonomicName>
, 931 [handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFD9F35700A2935491C427B2" blockId="0.[151,1436,1499,2035]" box="[197,926,1660,1683]" pageId="0" pageNumber="45">
<typeStatus id="77188835FFD9F35700A29354937D27B2" box="[197,295,1660,1683]" pageId="0" pageNumber="45" type="holotype">
<emphasis id="9AD7EA85FFD9F35700A29354937D27B2" box="[197,295,1660,1683]" italics="true" pageId="0" pageNumber="45">Holotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD9F357012D9354938E27B2" box="[330,468,1660,1683]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="parvidens">
<emphasis id="9AD7EA85FFD9F357012D9354938E27B2" box="[330,468,1660,1683]" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F357012D9354933A27B2" box="[330,352,1660,1683]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. parvidens
</emphasis>
</taxonomicName>
(NHB), male bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFD9F35700A293B496642792" blockId="0.[151,1436,1499,2035]" box="[197,1086,1692,1715]" pageId="0" pageNumber="45">
1. White, printed: HOA-BINH
<collectingCountry id="D0B47607FFD9F357027593B490222792" box="[530,632,1692,1715]" name="Vietnam" pageId="0" pageNumber="45">TONKIN</collectingCountry>
VII. [handwritten] 1934 A.
<collectingCountry id="D0B47607FFD9F35703FB93B491982792" box="[924,962,1692,1715]" name="South Georgia and the South Sandwich Islands" pageId="0" pageNumber="45">DE</collectingCountry>
COOMAN
</paragraph>
<paragraph id="A81C3697FFD9F35700A29394938027F2" blockId="0.[151,1436,1499,2035]" box="[197,474,1724,1747]" pageId="0" pageNumber="45">
2. Red, handwritten:
<typeStatus id="77188835FFD9F35701C29394938027F2" box="[421,474,1724,1747]" pageId="0" pageNumber="45">Type</typeStatus>
</paragraph>
<paragraph id="A81C3697FFD9F35700A293F496F527D2" blockId="0.[151,1436,1499,2035]" box="[197,1199,1755,1779]" pageId="0" pageNumber="45">
3. White, handwritten:
<typeStatus id="77188835FFD9F35701DC93F493AA27D2" box="[443,496,1756,1779]" pageId="0" pageNumber="45">Type</typeStatus>
<taxonomicName id="6FA34D14FFD9F357019093F490DF27D2" box="[503,645,1756,1779]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
[printed]
<taxonomicName id="6FA34D14FFD9F357029793F4910227D2" box="[752,856,1756,1779]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="parvidens" status="sp. nov.">parvidens</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD9F357033993F491CB27D2" box="[862,913,1756,1779]" pageId="0" pageNumber="45" rank="species">n.sp.</taxonomicNameLabel>
♂ det.G.Frey,196 [printed]
</paragraph>
<paragraph id="A81C3697FFD9F35700A293D496F62632" blockId="0.[151,1436,1499,2035]" box="[197,1196,1788,1811]" pageId="0" pageNumber="45">
4. Orange:
<typeStatus id="77188835FFD9F357015B93D493932632" box="[316,457,1788,1811]" pageId="0" pageNumber="45" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="6FA34D14FFD9F35701A893D4908A2632" authority="Frey, 1971" authorityName="Frey" authorityYear="1971" box="[463,720,1788,1811]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="parvidens">
<collectingCountry id="D0B47607FFD9F35701A893D493BF2632" box="[463,485,1788,1811]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. parvidens
<bibRefCitation id="CC324B66FFD9F357023993D4908A2632" author="Frey" box="[606,720,1788,1811]" pageId="0" pageNumber="45" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey, 1971</bibRefCitation>
</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFD9F35700A2923496072612" blockId="0.[151,1436,1499,2035]" box="[197,1117,1820,1843]" pageId="0" pageNumber="45">
5. White:
<taxonomicName id="6FA34D14FFD9F3570148923490DA2612" authority="Paulian" authorityName="Paulian" box="[303,640,1820,1843]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">Onthophagus clermonti Paulian</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFD9F35700A29213969E2672" blockId="0.[151,1436,1499,2035]" box="[197,1220,1851,1875]" pageId="0" pageNumber="45">
<emphasis id="9AD7EA85FFD9F35700A2921390792672" bold="true" box="[197,547,1851,1875]" pageId="0" pageNumber="45">Additional material examined:</emphasis>
<specimenCount id="BEA5FD1EFFD9F357024E92149012266C" box="[553,584,1851,1875]" pageId="0" pageNumber="45" type="generic">1♂</specimenCount>
,
<collectingCountry id="D0B47607FFD9F3570232921490E82672" box="[597,690,1852,1875]" name="Vietnam" pageId="0" pageNumber="45">Vietnam</collectingCountry>
, Hoa Binh Lac Tho, leg. A. de Cooman (
<collectionCode id="CEB2AE52FFD9F357040F921496E72672" LSID="urn:lsid:biocol.org:col:34988" box="[1128,1213,1852,1875]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="45">MNHN</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD9F35600A29273977E21EE" lastPageId="1" lastPageNumber="46" pageId="0" pageNumber="45" type="discussion">
<paragraph id="A81C3697FFD9F35600A29273977E21EE" blockId="0.[151,1436,1499,2035]" lastBlockId="1.[151,1436,152,207]" lastPageId="1" lastPageNumber="46" pageId="0" pageNumber="45">
<emphasis id="9AD7EA85FFD9F35700A2927393692652" bold="true" box="[197,307,1883,1907]" pageId="0" pageNumber="45">Remarks.</emphasis>
Apparently, Frey did not take into account the insufficient and obscure description of
<taxonomicName id="6FA34D14FFD9F35704B6927497022652" box="[1233,1368,1884,1907]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="9AD7EA85FFD9F35704B6927497022652" box="[1233,1368,1884,1907]" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F35704B6927496BD2652" box="[1233,1255,1884,1907]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. clermonti
</emphasis>
</taxonomicName>
based on one female and did not examine the
<typeStatus id="77188835FFD9F3570221925490FE26B2" box="[582,676,1916,1939]" pageId="0" pageNumber="45" type="holotype">holotype</typeStatus>
of the former. In the original description (
<bibRefCitation id="CC324B66FFD9F3570416925496B926B2" author="Frey" box="[1137,1251,1916,1939]" pageId="0" pageNumber="45" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey 1971</bibRefCitation>
) he compares
<taxonomicName id="6FA34D14FFD9F35705E59254935E2692" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="parvidens">
<emphasis id="9AD7EA85FFD9F35705E59254935E2692" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F35705E5925497C226B2" box="[1410,1432,1916,1939]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. parvidens
</emphasis>
</taxonomicName>
with
<taxonomicName id="6FA34D14FFD9F357012292B490E92692" authority="Lansberge, 1883" authorityName="Lansberge" authorityYear="1883" box="[325,691,1948,1971]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="deflexicollis">
<emphasis id="9AD7EA85FFD9F357012292B493B52692" box="[325,495,1948,1971]" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F357012292B493062692" box="[325,348,1948,1971]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. deflexicollis
</emphasis>
Lansberge, 1883
</taxonomicName>
. The
<typeStatus id="77188835FFD9F357029692B4917B2692" box="[753,801,1948,1971]" pageId="0" pageNumber="45">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFD9F35703E892B496412692" box="[911,1051,1948,1971]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="9AD7EA85FFD9F35703E892B496412692" box="[911,1051,1948,1971]" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F35703E892B491FC2692" box="[911,934,1948,1971]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. clermonti
</emphasis>
</taxonomicName>
is comprised only of the
<typeStatus id="77188835FFD9F357055C92B497C62692" box="[1339,1436,1948,1971]" pageId="0" pageNumber="45" type="holotype">holotype</typeStatus>
female, therefore it is not possible to examine the critical structures of the aedeagus for species identification in
<taxonomicName id="6FA34D14FFD9F35700F092F4937326D2" box="[151,297,2012,2035]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="9AD7EA85FFD9F35700F092F4937326D2" box="[151,297,2012,2035]" italics="true" pageId="0" pageNumber="45">Onthophagus</emphasis>
</taxonomicName>
. However, I conclude that the
<typeStatus id="77188835FFD9F357021A92F490BD26D2" box="[637,743,2012,2035]" pageId="0" pageNumber="45" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="6FA34D14FFD9F357036A92F491C226D2" box="[781,920,2012,2035]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="9AD7EA85FFD9F357036A92F491C226D2" box="[781,920,2012,2035]" italics="true" pageId="0" pageNumber="45">
<collectingCountry id="D0B47607FFD9F357036A92F4917926D2" box="[781,803,2012,2035]" name="Iceland" pageId="0" pageNumber="45">O</collectingCountry>
. clermonti
</emphasis>
</taxonomicName>
and
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<emphasis id="9AD7EA85FFD9F35703A992F4960626D2" box="[974,1116,2012,2035]" italics="true" pageId="0" pageNumber="45">
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. parvidens
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are conspecific because they share the same principle diagnostic character states of external morphology (e.g. the groundplan of the head, coloration, pronotum punctation, etc.). Moreover, both
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series were found at the same locality Hoa-Binh in
<collectingCountry id="D0B47607FFD8F35604A59590977A21EE" box="[1218,1312,184,207]" name="Vietnam" pageId="1" pageNumber="46">Vietnam</collectingCountry>
.
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186
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<emphasis id="9AD7EA85FFDBF35500F0925093CC26B1" bold="true" box="[151,406,1912,1936]" italics="true" pageId="2" pageNumber="47">Onthophagus vladimiri</emphasis>
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<emphasis id="9AD7EA85FFDBF35500F092B093D5268E" box="[151,399,1944,1967]" italics="true" pageId="2" pageNumber="47">Onthophagus vladimiri</emphasis>
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: 687
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(
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locality: Süd
<collectingCountry id="D0B47607FFDBF355037792B0910D268E" box="[784,855,1944,1967]" name="India" pageId="2" pageNumber="47">Indien</collectingCountry>
, Anamalai Hills, Cinchona)
<taxonomicName id="6FA34D14FFDBF35500F0929090F026EE" ID-CoL="7XMQW" authority="Balthasar, 1974: 183" authorityName="Balthasar" authorityPageNumber="183" authorityYear="1974" box="[151,682,1976,1999]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="anamalaiensis">
<emphasis id="9AD7EA85FFDBF35500F09290939D26EE" box="[151,455,1976,1999]" italics="true" pageId="2" pageNumber="47">Onthophagus anamalaiensis</emphasis>
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: 183
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<emphasis id="9AD7EA85FFDBF35502D6929F90AE26EE" bold="true" box="[689,756,1975,1999]" pageId="2" pageNumber="47">syn.n.</emphasis>
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(
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locality: S.
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, Anamalai Hills, Cinchona)
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material examined.
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One male and one female
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of
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. vladimiri
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(NMP) bearing the following two geographical labels:
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<paragraph id="A81C3697FFDAF35400A295F0902321CE" blockId="3.[151,1436,151,559]" box="[197,633,216,239]" pageId="3" pageNumber="48">
1. Anamalai Hills Cinchona
<quantity id="6F5B9B72FFDAF354019295F0901E21CE" box="[501,580,216,239]" metricMagnitude="3" metricUnit="m" metricValue="1.0668" pageId="3" pageNumber="48" unit="ft" value="3500.0">3500 ft</quantity>
.V.56
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2. Süd
<collectingCountry id="D0B47607FFDAF354016895D09309202E" box="[271,339,248,271]" name="India" pageId="3" pageNumber="48">Indien</collectingCountry>
leg. Nathan
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<emphasis id="9AD7EA85FFDAF35400A29430937D200E" box="[197,295,280,303]" italics="true" pageId="3" pageNumber="48">
<typeStatus id="77188835FFDAF35400A29430937D200E" box="[197,295,280,303]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
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of
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<emphasis id="9AD7EA85FFDAF354012D9430905F200E" box="[330,517,280,303]" italics="true" pageId="3" pageNumber="48">
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. anamalaiensis
</emphasis>
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(
<collectionCode id="CEB2AE52FFDAF354027494309008200E" box="[531,594,280,303]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="3" pageNumber="48">NMP</collectionCode>
), male bearing the following labels:
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<paragraph id="A81C3697FFDAF35400A294109131206E" blockId="3.[151,1436,151,559]" box="[197,875,312,335]" pageId="3" pageNumber="48">
1. White, printed: Anamalai Hills Cinchona S.
<collectingCountry id="D0B47607FFDAF35402C894109083206E" box="[687,729,312,335]" name="India" pageId="3" pageNumber="48">Ind</collectingCountry>
.
<quantity id="6F5B9B72FFDAF354028494109168206E" box="[739,818,312,335]" metricMagnitude="3" metricUnit="m" metricValue="1.0668" pageId="3" pageNumber="48" unit="ft" value="3500.0">3500 ft</quantity>
.1959
</paragraph>
<paragraph id="A81C3697FFDAF35400A2947096C6204E" blockId="3.[151,1436,151,559]" box="[197,1180,344,367]" pageId="3" pageNumber="48">
2. Rose: Onth
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[handwritten] Allotypus [printed, crossed with hand]
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<paragraph id="A81C3697FFDAF35400A29450962020AE" blockId="3.[151,1436,151,559]" box="[197,1146,376,399]" pageId="3" pageNumber="48">
3. White:
<taxonomicName id="6FA34D14FFDAF3540148945090C720AE" authority="Frey, 1957" authorityName="Frey" authorityYear="1957" box="[303,669,376,399]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="vladimiri">
Onthophagus vladimiri
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[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
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<subSubSection id="E0B9651CFFDAF35400A294BF93A4230E" pageId="3" pageNumber="48" type="discussion">
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<emphasis id="9AD7EA85FFDAF35400A294BF936D208E" bold="true" box="[197,311,407,431]" pageId="3" pageNumber="48">Remarks.</emphasis>
The examined
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of
<taxonomicName id="6FA34D14FFDAF354020C94B09174208E" box="[619,814,408,431]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="anamalaiensis">
<emphasis id="9AD7EA85FFDAF354020C94B09174208E" box="[619,814,408,431]" italics="true" pageId="3" pageNumber="48">
<collectingCountry id="D0B47607FFDAF354020C94B090D8208E" box="[619,642,408,431]" name="Iceland" pageId="3" pageNumber="48">O</collectingCountry>
. anamalaiensis
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is conspecific with the
<typeStatus id="77188835FFDAF354045F94B09632208E" box="[1080,1128,408,431]" pageId="3" pageNumber="48">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFDAF35404B294B097C6208E" box="[1237,1436,408,431]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDAF35404B294B097C6208E" box="[1237,1436,408,431]" italics="true" pageId="3" pageNumber="48">
<collectingCountry id="D0B47607FFDAF35404B294B096B6208E" box="[1237,1260,408,431]" name="Iceland" pageId="3" pageNumber="48">O</collectingCountry>
. pseudojavanus
</emphasis>
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because they share the same aedeagus and principle diagnostic characters of the external morphology (structure of head and vertex horns, pronotum punctation, body coloration, etc.). Moreover, both
<typeStatus id="77188835FFDAF354039B94F0967120CE" box="[1020,1067,472,495]" pageId="3" pageNumber="48">type</typeStatus>
series were collected at the same locality in southern
<collectingCountry id="D0B47607FFDAF354011394D093EA232E" box="[372,432,504,527]" name="India" pageId="3" pageNumber="48">India</collectingCountry>
. It seems that
<taxonomicName id="6FA34D14FFDAF354022894D0914A232E" box="[591,784,504,527]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="anamalaiensis">
<emphasis id="9AD7EA85FFDAF354022894D0914A232E" box="[591,784,504,527]" italics="true" pageId="3" pageNumber="48">
<collectingCountry id="D0B47607FFDAF354022894D0903C232E" box="[591,614,504,527]" name="Iceland" pageId="3" pageNumber="48">O</collectingCountry>
. anamalaiensis
</emphasis>
</taxonomicName>
was described as a new species because the author did not know the identity of
<taxonomicName id="6FA34D14FFDAF3540112973093AD230E" box="[373,503,536,559]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="vladimiri">
<emphasis id="9AD7EA85FFDAF3540112973093AD230E" box="[373,503,536,559]" italics="true" pageId="3" pageNumber="48">
<collectingCountry id="D0B47607FFDAF3540112973093D1230E" box="[373,395,536,559]" name="Iceland" pageId="3" pageNumber="48">O</collectingCountry>
. vladimiri
</emphasis>
</taxonomicName>
.
</paragraph>
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<treatment id="200A8781FFDBF35500F09030972C260E" ID-DOI="http://doi.org/10.5281/zenodo.6208726" ID-GBIF-Taxon="119393970" ID-Zenodo-Dep="6208726" LSID="urn:lsid:plazi:treatment:200A8781FFDBF35500F09030972C260E" httpUri="http://treatment.plazi.org/id/200A8781FFDBF35500F09030972C260E" lastPageNumber="47" pageId="2" pageNumber="47">
<subSubSection id="E0B9651CFFDBF35500F09030902C240E" box="[151,630,1303,1328]" pageId="2" pageNumber="47" type="nomenclature">
<paragraph id="A81C3697FFDBF35500F09030902C240E" blockId="2.[151,1342,1303,1391]" box="[151,630,1303,1328]" pageId="2" pageNumber="47">
<heading id="F35481FBFFDBF35500F09030902C240E" bold="true" box="[151,630,1303,1328]" fontSize="10" level="2" pageId="2" pageNumber="47" reason="2">
<taxonomicName id="6FA34D14FFDBF35500F09030902C240E" ID-CoL="49MRB" authority="Paulian, 1931" authorityName="Paulian" authorityYear="1931" box="[151,630,1303,1328]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF35500F09030902C240E" bold="true" box="[151,630,1303,1328]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500F09030938C2411" bold="true" box="[151,470,1304,1328]" italics="true" pageId="2" pageNumber="47">Onthophagus pseudojavanus</emphasis>
<bibRefCitation id="CC324B66FFDBF35501BB903F902C240E" author="Paulian" box="[476,630,1303,1327]" pageId="2" pageNumber="47" refString="Paulian, R. (1931) Coprophages nouveaux pour la faune de Hoa-Binh (Tonkin). Bulletin de la Societe entomologique de France, 36, 270 - 272." type="journal article" year="1931">Paulian, 1931</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFDBF35500F090109764244E" pageId="2" pageNumber="47" type="reference_group">
<paragraph id="A81C3697FFDBF35500F090109764244E" blockId="2.[151,1342,1303,1391]" pageId="2" pageNumber="47">
<treatmentCitationGroup id="88B311B9FFDBF35500F090109764244E" pageId="2" pageNumber="47">
<taxonomicName id="6FA34D14FFDBF35500F0901090C1246E" ID-CoL="49MRB" authority="Paulian, 1931: 272" authorityName="Paulian" authorityPageNumber="272" authorityYear="1931" box="[151,667,1336,1359]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF35500F090109396246E" box="[151,460,1336,1359]" italics="true" pageId="2" pageNumber="47">Onthophagus pseudojavanus</emphasis>
<treatmentCitation id="29021086FFDBF35501B5901090C1246E" author="Paulian" box="[466,667,1336,1359]" page="272" pageId="2" pageNumber="47" year="1931">
<bibRefCitation id="CC324B66FFDBF35501B590109032246E" author="Paulian" box="[466,616,1336,1359]" pageId="2" pageNumber="47" refString="Paulian, R. (1931) Coprophages nouveaux pour la faune de Hoa-Binh (Tonkin). Bulletin de la Societe entomologique de France, 36, 270 - 272." type="journal article" year="1931">Paulian, 1931</bibRefCitation>
: 272
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="77188835FFDBF35502CE90109082246E" box="[681,728,1336,1359]" pageId="2" pageNumber="47">type</typeStatus>
locality: Hoa Binh,
<collectingCountry id="D0B47607FFDBF35503D6901091A6246E" box="[945,1020,1336,1359]" name="Vietnam" pageId="2" pageNumber="47">Tonkin</collectingCountry>
[
<collectingCountry id="D0B47607FFDBF355046E90109631246E" box="[1033,1131,1336,1359]" name="Vietnam" pageId="2" pageNumber="47">Vietnam</collectingCountry>
])
<taxonomicName id="6FA34D14FFDBF35500F09070903E244E" authority="Masumoto, 1989: 93" authorityName="Masumoto" authorityPageNumber="93" authorityYear="1989" box="[151,612,1368,1391]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="demaak">
<emphasis id="9AD7EA85FFDBF35500F0907093DA244E" box="[151,384,1368,1391]" italics="true" pageId="2" pageNumber="47">Onthophagus demaak</emphasis>
<treatmentCitation id="29021086FFDBF35501E09070903E244E" author="Masumoto" box="[391,612,1368,1391]" page="93" pageId="2" pageNumber="47" year="1989">
<bibRefCitation id="CC324B66FFDBF35501E090709065244E" author="Masumoto" box="[391,575,1368,1391]" pageId="2" pageNumber="47" refString="Masumoto, K. (1989) Coprophagid beetles from Northwest Thailand IV. The Entomological Review of Japan, 44, 87 - 96" type="journal article" year="1989">Masumoto, 1989</bibRefCitation>
: 93
</treatmentCitation>
</taxonomicName>
<taxonomicNameLabel id="81E457FEFFDBF355020C907F90F4244E" box="[619,686,1367,1391]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF355020C907F90F4244E" bold="true" box="[619,686,1367,1391]" pageId="2" pageNumber="47">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFDBF35502DC907090B0244E" box="[699,746,1368,1391]" pageId="2" pageNumber="47">type</typeStatus>
locality: Doi Suthep/Pui, Chiang Mai Prov. [
<collectingCountry id="D0B47607FFDBF35504AD9070976A244E" box="[1226,1328,1368,1391]" name="Thailand" pageId="2" pageNumber="47">Thailand</collectingCountry>
])
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFDBF35500F090BF960A27EE" pageId="2" pageNumber="47" type="materials_examined">
<paragraph id="A81C3697FFDBF35500F090BF9685248E" blockId="2.[151,1436,1431,1839]" box="[151,1247,1431,1455]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500F090BF93EB248E" bold="true" box="[151,433,1431,1455]" pageId="2" pageNumber="47">
<typeStatus id="77188835FFDBF35500F090BF9295248E" box="[151,207,1431,1455]" pageId="2" pageNumber="47">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="77188835FFDBF35501D090B09043248E" box="[439,537,1432,1455]" pageId="2" pageNumber="47" type="holotype">
<emphasis id="9AD7EA85FFDBF35501D090B09043248E" box="[439,537,1432,1455]" italics="true" pageId="2" pageNumber="47">Holotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF355025B90B090A6248E" box="[572,764,1432,1455]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF355025B90B090A6248E" box="[572,764,1432,1455]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355025B90B09008248E" box="[572,594,1432,1455]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. pseudojavanus
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFDBF355036E90B0913A248E" LSID="urn:lsid:biocol.org:col:34988" box="[777,864,1432,1455]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
), male bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFDBF35500A2909091E824EE" blockId="2.[151,1436,1431,1839]" box="[197,946,1464,1487]" pageId="2" pageNumber="47">
1. White, printed: Hoa-Binh (
<collectingCountry id="D0B47607FFDBF35501999090901724EE" box="[510,589,1464,1487]" name="Vietnam" pageId="2" pageNumber="47">Tonkin</collectingCountry>
) (A.de Cooman) Coll.J.Clermont
</paragraph>
<paragraph id="A81C3697FFDBF35500A290F093EE24CE" blockId="2.[151,1436,1431,1839]" box="[197,436,1496,1519]" pageId="2" pageNumber="47">
2. Red, printed:
<collectionCode id="CEB2AE52FFDBF355011690F093EE24CE" box="[369,436,1496,1519]" pageId="2" pageNumber="47">
<typeStatus id="77188835FFDBF355011690F093EE24CE" box="[369,436,1496,1519]" pageId="2" pageNumber="47">TYPE</typeStatus>
</collectionCode>
</paragraph>
<paragraph id="A81C3697FFDBF35500A290D0913F272E" blockId="2.[151,1436,1431,1839]" box="[197,869,1528,1551]" pageId="2" pageNumber="47">
3. White, handwritten:
<taxonomicName id="6FA34D14FFDBF35501DC90D09013272E" box="[443,585,1528,1551]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
sp.? V. de javanus Lansb.
</paragraph>
<paragraph id="A81C3697FFDBF35500A2933090F7270E" blockId="2.[151,1436,1431,1839]" box="[197,685,1560,1583]" pageId="2" pageNumber="47">
4. White, handwritten:
<taxonomicName id="6FA34D14FFDBF35501DC93309023270E" box="[443,633,1560,1583]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<collectingCountry id="D0B47607FFDBF35501DC9330938B270E" box="[443,465,1560,1583]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. pseudojavanus
</taxonomicName>
<typeStatus id="77188835FFDBF3550218933090F7270E" box="[639,685,1560,1583]" pageId="2" pageNumber="47">type</typeStatus>
</paragraph>
<paragraph id="A81C3697FFDBF35500A293109720276E" blockId="2.[151,1436,1431,1839]" box="[197,1402,1592,1615]" pageId="2" pageNumber="47">
4. Orange:
<typeStatus id="77188835FFDBF355015B93109393276E" box="[316,457,1592,1615]" pageId="2" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="6FA34D14FFDBF35501A8931091C7276E" authority="Paulian, 1931" authorityName="Paulian" authorityYear="1931" box="[463,925,1592,1615]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF35501A89310915E276E" box="[463,772,1592,1615]" italics="true" pageId="2" pageNumber="47">Onthophagus pseudojavanus</emphasis>
<bibRefCitation id="CC324B66FFDBF355036C931091C7276E" author="Paulian" box="[779,925,1592,1615]" pageId="2" pageNumber="47" refString="Paulian, R. (1931) Coprophages nouveaux pour la faune de Hoa-Binh (Tonkin). Bulletin de la Societe entomologique de France, 36, 270 - 272." type="journal article" year="1931">Paulian, 1931</bibRefCitation>
</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFDBF35500A2937093D627AE" blockId="2.[151,1436,1431,1839]" pageId="2" pageNumber="47">
<typeStatus id="77188835FFDBF35500A293709372274E" box="[197,296,1624,1647]" pageId="2" pageNumber="47" type="paratype">
<emphasis id="9AD7EA85FFDBF35500A293709372274E" box="[197,296,1624,1647]" italics="true" pageId="2" pageNumber="47">Paratype</emphasis>
</typeStatus>
male of
<taxonomicName id="6FA34D14FFDBF35501EF937093A7274E" box="[392,509,1624,1647]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="demaak">
<emphasis id="9AD7EA85FFDBF35501EF937093A7274E" box="[392,509,1624,1647]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35501EF937093C4274E" box="[392,414,1624,1647]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. demaak
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFDBF355026B9370903E274E" LSID="urn:lsid:biocol.org:col:34988" box="[524,612,1624,1647]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
), with locality label “Doi Suthep/Pui Chiang Mai Prov.
<date id="DC1D1057FFDBF35504A5937097C6274E" box="[1218,1436,1624,1647]" pageId="2" pageNumber="47" value="1988-07-30" valueMax="1988-08-05" valueMin="1988-07-30">30.VII.–5.VIII.1988</date>
K. MASUMOTO leg.”
</paragraph>
<paragraph id="A81C3697FFDBF35500A293BF960A27EE" blockId="2.[151,1436,1431,1839]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500A293BF9079278E" bold="true" box="[197,547,1687,1711]" pageId="2" pageNumber="47">Additional material examined:</emphasis>
<specimenCount id="BEA5FD1EFFDBF355024E93B090122788" box="[553,584,1687,1711]" pageId="2" pageNumber="47" type="generic">1♂</specimenCount>
and 1Ψ (
<collectionCode id="CEB2AE52FFDBF35502CE93B090A4278E" LSID="urn:lsid:biocol.org:col:34988" box="[681,766,1688,1711]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
) bearing the same geographical label as that of the
<typeStatus id="77188835FFDBF355054593B09725278E" box="[1314,1407,1688,1711]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF35500F0939093B527EE" authority="Paulian, 1931" authorityName="Paulian" authorityYear="1931" box="[151,495,1720,1743]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF35500F09390930F27EE" box="[151,341,1720,1743]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35500F0939092F727EE" box="[151,173,1720,1743]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. pseudojavanus
</emphasis>
<bibRefCitation id="CC324B66FFDBF355013B939093B527EE" author="Paulian" box="[348,495,1720,1743]" pageId="2" pageNumber="47" refString="Paulian, R. (1931) Coprophages nouveaux pour la faune de Hoa-Binh (Tonkin). Bulletin de la Societe entomologique de France, 36, 270 - 272." type="journal article" year="1931">Paulian, 1931</bibRefCitation>
</taxonomicName>
and indicated by R. Paulian as “
<taxonomicName id="6FA34D14FFDBF355032D9390965227EE" box="[842,1032,1720,1743]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<collectingCountry id="D0B47607FFDBF355032D9390913A27EE" box="[842,864,1720,1743]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. pseudojavanus
</taxonomicName>
? m.”.
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFDBF35500A293FF972C260E" pageId="2" pageNumber="47" type="discussion">
<paragraph id="A81C3697FFDBF35500A293FF972C260E" blockId="2.[151,1436,1431,1839]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500A293FF936F27CE" bold="true" box="[197,309,1751,1775]" pageId="2" pageNumber="47">Remarks.</emphasis>
The examined
<typeStatus id="77188835FFDBF355018793F0906427CE" box="[480,574,1752,1775]" pageId="2" pageNumber="47" type="paratype">paratype</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF355020393F0908327CE" box="[612,729,1752,1775]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="demaak">
<emphasis id="9AD7EA85FFDBF355020393F0908327CE" box="[612,729,1752,1775]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355020393F0902027CE" box="[612,634,1752,1775]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. demaak
</emphasis>
</taxonomicName>
is conspecific with the
<typeStatus id="77188835FFDBF355038793F0965527CE" box="[992,1039,1752,1775]" pageId="2" pageNumber="47">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFDBF355041C93F0976427CE" box="[1147,1342,1752,1775]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF355041C93F0976427CE" box="[1147,1342,1752,1775]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355041C93F096C827CE" box="[1147,1170,1752,1775]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. pseudojavanus
</emphasis>
</taxonomicName>
because they share the same aedeagus, structure of the head, vertex horn, pronotum punctation and body coloration. It seems that
<taxonomicName id="6FA34D14FFDBF35500F092309350260E" box="[151,266,1816,1839]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="demaak">
<emphasis id="9AD7EA85FFDBF35500F092309350260E" box="[151,266,1816,1839]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35500F0923092F7260E" box="[151,173,1816,1839]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. demaak
</emphasis>
</taxonomicName>
was described as a new species simply because the author did not know the identity of
<taxonomicName id="6FA34D14FFDBF35504D792309734260E" box="[1200,1390,1816,1839]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="9AD7EA85FFDBF35504D792309734260E" box="[1200,1390,1816,1839]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35504D79230969C260E" box="[1200,1222,1816,1839]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. pseudojavanus
</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="6A54D00B610D2350A9CB84BA1037546D" ID-DOI="10.5281/zenodo.197274" ID-GBIF-Dataset="c6a45517-5e2d-496e-a76c-091740a56475" ID-ISSN="1175-5326" ID-Zenodo-Dep="197274" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460263975273" checkinUser="plazi" docAuthor="Tarasov, Sergey I." docDate="2010" docId="200A8781FFDBF35500F0949091A725EE" docLanguage="en" docName="zt02566p048.pdf" docOrigin="Zootaxa 2566" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus orientalis Harold 1868" docType="treatment" docVersion="8" lastPageNumber="47" masterDocId="DC33FFF9FFD9F35700679528925A2121" masterDocTitle="Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region" masterLastPageNumber="48" masterPageNumber="45" pageNumber="47" updateTime="1698245222659" updateUser="plazi">
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<mods:title id="96A351AC2D84E4CB0A487C77BC46B2DD">Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region</mods:title>
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<mods:date id="FCE4CEDEFC45352360BE9894BCECD623">2010</mods:date>
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<subSubSection id="E0B9651CFFDBF35500F09490906E20EE" box="[151,564,439,464]" pageId="2" pageNumber="47" type="nomenclature">
<paragraph id="A81C3697FFDBF35500F09490906E20EE" blockId="2.[151,1436,439,591]" box="[151,564,439,464]" pageId="2" pageNumber="47">
<heading id="F35481FBFFDBF35500F09490906E20EE" bold="true" box="[151,564,439,464]" fontSize="10" level="2" pageId="2" pageNumber="47" reason="2">
<taxonomicName id="6FA34D14FFDBF35500F09490906E20EE" ID-CoL="7XNRC" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[151,564,439,464]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35500F09490906E20EE" bold="true" box="[151,564,439,464]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500F0949093C020F1" bold="true" box="[151,410,440,464]" italics="true" pageId="2" pageNumber="47">Onthophagus orientalis</emphasis>
<bibRefCitation id="CC324B66FFDBF35501C6949F906E20EE" author="Harold" box="[417,564,439,463]" pageId="2" pageNumber="47" refString="Harold, E. von (1868) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 3, 80 - 86." type="journal article" year="1868">Harold, 1868</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFDBF35500F094F091B3236E" pageId="2" pageNumber="47" type="reference_group">
<paragraph id="A81C3697FFDBF35500F094F09072232E" blockId="2.[151,1436,439,591]" pageId="2" pageNumber="47">
<treatmentCitationGroup id="88B311B9FFDBF35500F094F09072232E" pageId="2" pageNumber="47">
<taxonomicName id="6FA34D14FFDBF35500F094F0900F20CE" ID-CoL="7XNRC" authority="Harold, 1868: 83" authorityName="Harold" authorityPageNumber="83" authorityYear="1868" box="[151,597,472,495]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35500F094F093CE20CE" box="[151,404,472,495]" italics="true" pageId="2" pageNumber="47">Onthophagus orientalis</emphasis>
<treatmentCitation id="29021086FFDBF35501FA94F0900F20CE" author="Harold" box="[413,597,472,495]" page="83" pageId="2" pageNumber="47" year="1868">
<bibRefCitation id="CC324B66FFDBF35501FA94F0907520CE" author="Harold" box="[413,559,472,495]" pageId="2" pageNumber="47" refString="Harold, E. von (1868) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 3, 80 - 86." type="journal article" year="1868">Harold, 1868</bibRefCitation>
: 83
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="77188835FFDBF355020294F090CE20CE" box="[613,660,472,495]" pageId="2" pageNumber="47">type</typeStatus>
locality: original description “
<collectingCountry id="D0B47607FFDBF35503B894F0960E20CE" box="[991,1108,472,495]" name="Hong Kong" pageId="2" pageNumber="47">Hongkong</collectingCountry>
, Bengal”; lecotype designated by
<bibRefCitation id="CC324B66FFDBF355008F94D093D9232E" author="Zunino" box="[232,387,504,527]" pageId="2" pageNumber="47" refString="Zunino, M. (1976) Revisione delle species paleartiche del sottogenere Onthophagus (sensu stricto) Latr. I tipi di H. W. Bates, L. Fairmaire, E. von Harold, G. van Lansberge, S. A. de Marseul, L. Reiche e D. Sharp nel Museum National d'Histoire Naturelle di Parigi. Bollettino del Museo di Zoologia dell'Universit a di Torino, 4, 71 - 110." type="journal article" year="1976">Zunino (1976)</bibRefCitation>
“
<collectingCountry id="D0B47607FFDBF35501F494D0904C232E" box="[403,534,504,527]" name="Hong Kong" pageId="2" pageNumber="47">Hong Kong</collectingCountry>
”)
</treatmentCitationGroup>
</paragraph>
<paragraph id="A81C3697FFDBF35500F0973091FC230E" blockId="2.[151,1436,439,591]" box="[151,934,536,559]" pageId="2" pageNumber="47">
<treatmentCitationGroup id="88B311B9FFDBF35500F0973091FC230E" box="[151,934,536,559]" pageId="2" pageNumber="47">
<taxonomicName id="6FA34D14FFDBF35500F0973093B2230E" ID-CoL="7XNRC" authority="Harold" authorityName="Harold" box="[151,488,536,559]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35500F0973093C9230E" box="[151,403,536,559]" italics="true" pageId="2" pageNumber="47">Onthophagus orientalis</emphasis>
Harold
</taxonomicName>
;
<treatmentCitation id="29021086FFDBF3550195973090F1230E" author="Zunino" box="[498,683,536,559]" page="79" pageId="2" pageNumber="47" year="1976">
<bibRefCitation id="CC324B66FFDBF3550195973090DF230E" author="Zunino" box="[498,645,536,559]" pageId="2" pageNumber="47" refString="Zunino, M. (1976) Revisione delle species paleartiche del sottogenere Onthophagus (sensu stricto) Latr. I tipi di H. W. Bates, L. Fairmaire, E. von Harold, G. van Lansberge, S. A. de Marseul, L. Reiche e D. Sharp nel Museum National d'Histoire Naturelle di Parigi. Bollettino del Museo di Zoologia dell'Universit a di Torino, 4, 71 - 110." type="journal article" year="1976">Zunino, 1976</bibRefCitation>
: 79
</treatmentCitation>
(
<typeStatus id="77188835FFDBF35502DF97309146230E" box="[696,796,536,559]" pageId="2" pageNumber="47" type="lectotype">lectotype</typeStatus>
designation)
</treatmentCitationGroup>
</paragraph>
<paragraph id="A81C3697FFDBF35500F0971091B3236E" blockId="2.[151,1436,439,591]" box="[151,1001,567,591]" pageId="2" pageNumber="47">
<treatmentCitationGroup id="88B311B9FFDBF35500F0971091B3236E" box="[151,1001,567,591]" pageId="2" pageNumber="47">
<taxonomicName id="6FA34D14FFDBF35500F097109063236E" ID-CoL="7XNQR" authority="Paulian, 1978: 40" authorityName="Paulian" authorityPageNumber="40" authorityYear="1978" box="[151,569,568,591]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF35500F09710932C236E" box="[151,374,568,591]" italics="true" pageId="2" pageNumber="47">Onthophagus perroti</emphasis>
<treatmentCitation id="29021086FFDBF355011A97109063236E" author="Paulian" box="[381,569,568,591]" page="40" pageId="2" pageNumber="47" year="1978">
<bibRefCitation id="CC324B66FFDBF355011A97109049236E" author="Paulian" box="[381,531,568,591]" pageId="2" pageNumber="47" refString="Paulian, R. (1978) Quelques nouveaux Coleopteres Scarabaeidae du Vietnam. Bulletin de la Societe entomologique de France, 83, 39 - 42." type="journal article" year="1978">Paulian, 1978</bibRefCitation>
: 40
</treatmentCitation>
</taxonomicName>
<taxonomicNameLabel id="81E457FEFFDBF3550258971F90D8236E" box="[575,642,567,591]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF3550258971F90D8236E" bold="true" box="[575,642,567,591]" pageId="2" pageNumber="47">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFDBF35502E8971090E4236E" box="[655,702,568,591]" pageId="2" pageNumber="47">type</typeStatus>
locality: Saigon [
<collectingCountry id="D0B47607FFDBF355031D97109181236E" box="[890,987,568,591]" name="Vietnam" pageId="2" pageNumber="47">Vietnam</collectingCountry>
])
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFDBF35500F0975F91D7256E" pageId="2" pageNumber="47" type="materials_examined">
<paragraph id="A81C3697FFDBF35500F0975F96F723AE" blockId="2.[151,1436,631,1231]" box="[151,1197,631,655]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500F0975F93EB23AE" bold="true" box="[151,433,631,655]" pageId="2" pageNumber="47">
<typeStatus id="77188835FFDBF35500F0975F929523AE" box="[151,207,631,655]" pageId="2" pageNumber="47">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="77188835FFDBF35501D09750904523AE" box="[439,543,632,655]" pageId="2" pageNumber="47" type="lectotype">
<emphasis id="9AD7EA85FFDBF35501D09750904523AE" box="[439,543,632,655]" italics="true" pageId="2" pageNumber="47">Lectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF35502259750909323AE" box="[578,713,632,655]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35502259750909323AE" box="[578,713,632,655]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35502259750900223AE" box="[578,600,632,655]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. orientalis
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFDBF35502B09750917423AE" LSID="urn:lsid:biocol.org:col:34988" box="[727,814,632,655]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
), male bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFDBF35500A297B09038238E" blockId="2.[151,1436,631,1231]" box="[197,610,664,687]" pageId="2" pageNumber="47">1. White, printed: Ex Musaeo E Harold</paragraph>
<paragraph id="A81C3697FFDBF35500A29790905E23EE" blockId="2.[151,1436,631,1231]" box="[197,516,696,719]" pageId="2" pageNumber="47">
2. White, printed:
<collectingCountry id="D0B47607FFDBF35501E09790905E23EE" box="[391,516,696,719]" name="Hong Kong" pageId="2" pageNumber="47">Hong Kong</collectingCountry>
</paragraph>
<paragraph id="A81C3697FFDBF35500A297F09313222E" blockId="2.[151,1436,631,1231]" pageId="2" pageNumber="47">
3. Red: Lecto [handwritten]
<collectionCode id="CEB2AE52FFDBF355019F97F0901723CE" box="[504,589,728,751]" pageId="2" pageNumber="47">
<typeStatus id="77188835FFDBF355019F97F0901723CE" box="[504,589,728,751]" pageId="2" pageNumber="47">TYPUS</typeStatus>
</collectionCode>
[printed]
<taxonomicName id="6FA34D14FFDBF35502DE97F091B223CE" authority="Har." authorityName="Har." box="[697,1000,728,751]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">Onthophagus orientalis Har.</taxonomicName>
[handwritten] M. ZUNINO 19 [printed] 76 [handwritten]
</paragraph>
<paragraph id="A81C3697FFDBF35500A296309189220E" blockId="2.[151,1436,631,1231]" box="[197,979,791,815]" pageId="2" pageNumber="47">
Also, 1Ψ
<typeStatus id="77188835FFDBF355014D963093E7220E" box="[298,445,792,815]" pageId="2" pageNumber="47" type="paralectotype">
<emphasis id="9AD7EA85FFDBF355014D963093E7220E" box="[298,445,792,815]" italics="true" pageId="2" pageNumber="47">paralectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF35501879630903D220E" box="[480,615,792,815]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35501879630903D220E" box="[480,615,792,815]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF3550187963093AC220E" box="[480,502,792,815]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. orientalis
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFDBF355021296309090220E" LSID="urn:lsid:biocol.org:col:34988" box="[629,714,792,815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
), from the same locality.
</paragraph>
<paragraph id="A81C3697FFDBF35500A2961091C2226E" blockId="2.[151,1436,631,1231]" box="[197,920,824,847]" pageId="2" pageNumber="47">
<typeStatus id="77188835FFDBF35500A29610937D226E" box="[197,295,824,847]" pageId="2" pageNumber="47" type="holotype">
<emphasis id="9AD7EA85FFDBF35500A29610937D226E" box="[197,295,824,847]" italics="true" pageId="2" pageNumber="47">Holotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF355012D961093EE226E" box="[330,436,824,847]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF355012D961093EE226E" box="[330,436,824,847]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355012D9610933A226E" box="[330,352,824,847]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. perroti
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFDBF35501A596109043226E" LSID="urn:lsid:biocol.org:col:34988" box="[450,537,824,847]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
), male bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFDBF35500A296709647224E" blockId="2.[151,1436,631,1231]" box="[197,1053,856,879]" pageId="2" pageNumber="47">
1. White, printed: MUSÉUM
<collectionCode id="CEB2AE52FFDBF355026496709016224E" box="[515,588,856,879]" pageId="2" pageNumber="47">PARIS</collectionCode>
SAIGON 1950 [handwritten] J. BARBIER
</paragraph>
<paragraph id="A81C3697FFDBF35500A2965091B922AE" blockId="2.[151,1436,631,1231]" box="[197,995,888,911]" pageId="2" pageNumber="47">
2. White:
<taxonomicName id="6FA34D14FFDBF35501489650905122AE" box="[303,523,888,911]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti" status="sp. nov.">Onthophagus perroti</taxonomicName>
<taxonomicNameLabel id="81E457FEFFDBF35502769650901E22AE" box="[529,580,888,911]" pageId="2" pageNumber="47" rank="species">n.sp.</taxonomicNameLabel>
[handwritten] R. Paulian det. [printed]
</paragraph>
<paragraph id="A81C3697FFDBF35500A296B093A4228E" blockId="2.[151,1436,631,1231]" box="[197,510,920,943]" pageId="2" pageNumber="47">
3. Red, printed:
<typeStatus id="77188835FFDBF355011696B093A4228E" box="[369,510,920,943]" pageId="2" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
</paragraph>
<paragraph id="A81C3697FFDBF35500A2969096D422EE" blockId="2.[151,1436,631,1231]" box="[197,1166,952,975]" pageId="2" pageNumber="47">
4. Orange:
<taxonomicName id="6FA34D14FFDBF355015B969090EB22EE" authority="Paulian, 1978" authorityName="Paulian" authorityYear="1978" box="[316,689,952,975]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
Onthophagus perroti
<bibRefCitation id="CC324B66FFDBF3550278969090EB22EE" author="Paulian" box="[543,689,952,975]" pageId="2" pageNumber="47" refString="Paulian, R. (1978) Quelques nouveaux Coleopteres Scarabaeidae du Vietnam. Bulletin de la Societe entomologique de France, 83, 39 - 42." type="journal article" year="1978">Paulian, 1978</bibRefCitation>
</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFDBF35500A296F0966D22CE" blockId="2.[151,1436,631,1231]" box="[197,1079,984,1007]" pageId="2" pageNumber="47">
5. White:
<taxonomicName id="6FA34D14FFDBF355014896F0900122CE" authority="Har." authorityName="Har." box="[303,603,984,1007]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">Onthophagus orientalis Har.</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFDBF35500A296DF91D7256E" blockId="2.[151,1436,631,1231]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500A296DF9072252E" bold="true" box="[197,552,1015,1039]" pageId="2" pageNumber="47">Additional material examined:</emphasis>
<specimenCount id="BEA5FD1EFFDBF355024896D090152528" box="[559,591,1015,1039]" pageId="2" pageNumber="47" type="generic">1♂</specimenCount>
(
<collectionCode id="CEB2AE52FFDBF355023996D090E9252E" LSID="urn:lsid:biocol.org:col:34988" box="[606,691,1016,1039]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
) bearing the same geographical label as the
<typeStatus id="77188835FFDBF35504F096D096AC252E" box="[1175,1270,1016,1039]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
and labeled by someone later as a
<typeStatus id="77188835FFDBF355010D91309392250E" box="[362,456,1048,1071]" pageId="2" pageNumber="47" type="paratype">paratype</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF3550189913090A7250E" authority="Paulian, 1978" authorityName="Paulian" authorityYear="1978" box="[494,765,1048,1071]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF355018991309001250E" box="[494,603,1048,1071]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35501899130905F250E" box="[494,517,1048,1071]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. perroti
</emphasis>
<bibRefCitation id="CC324B66FFDBF3550204913090A7250E" author="Paulian" box="[611,765,1048,1071]" pageId="2" pageNumber="47" refString="Paulian, R. (1978) Quelques nouveaux Coleopteres Scarabaeidae du Vietnam. Bulletin de la Societe entomologique de France, 83, 39 - 42." type="journal article" year="1978">Paulian, 1978</bibRefCitation>
</taxonomicName>
. The original description of
<taxonomicName id="6FA34D14FFDBF3550450913096FF250E" box="[1079,1189,1048,1071]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF3550450913096FF250E" box="[1079,1189,1048,1071]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355045091309614250E" box="[1079,1102,1048,1071]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. perroti
</emphasis>
</taxonomicName>
was based on a single specimen (
<bibRefCitation id="CC324B66FFDBF355016E911093C2256E" author="Paulian" box="[265,408,1080,1103]" pageId="2" pageNumber="47" refString="Paulian, R. (1978) Quelques nouveaux Coleopteres Scarabaeidae du Vietnam. Bulletin de la Societe entomologique de France, 83, 39 - 42." type="journal article" year="1978">Paulian 1978</bibRefCitation>
) and therefore this
<typeStatus id="77188835FFDBF355020F9110909E256E" box="[616,708,1080,1103]" pageId="2" pageNumber="47" type="paratype">paratype</typeStatus>
label is erroneous.
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFDBF35500A2917F91A725EE" pageId="2" pageNumber="47" type="discussion">
<paragraph id="A81C3697FFDBF35500A2917F91A725EE" blockId="2.[151,1436,631,1231]" pageId="2" pageNumber="47">
<emphasis id="9AD7EA85FFDBF35500A2917F936D254E" bold="true" box="[197,311,1111,1135]" pageId="2" pageNumber="47">Remarks.</emphasis>
Based on my observations,
<taxonomicName id="6FA34D14FFDBF3550215917090BB254E" box="[626,737,1112,1135]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF3550215917090BB254E" box="[626,737,1112,1135]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF3550215917090D3254E" box="[626,649,1112,1135]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. perroti
</emphasis>
</taxonomicName>
represents individuals exhibiting the minor form of the earlier described and common Oriental species
<taxonomicName id="6FA34D14FFDBF35502289150912825AE" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[591,882,1144,1167]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35502289150908325AE" box="[591,729,1144,1167]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35502289150903F25AE" box="[591,613,1144,1167]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. orientalis
</emphasis>
<bibRefCitation id="CC324B66FFDBF35502879150912825AE" author="Harold" box="[736,882,1144,1167]" pageId="2" pageNumber="47" refString="Harold, E. von (1868) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 3, 80 - 86." type="journal article" year="1868">Harold, 1868</bibRefCitation>
</taxonomicName>
. The character states of external morphology of
<taxonomicName id="6FA34D14FFDBF35505E5915092BA258E" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF35505E5915092BA258E" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35505E5915097C225AE" box="[1410,1432,1144,1167]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. perroti
</emphasis>
</taxonomicName>
(e.g. vertex horn, pronotum punctation, etc.) are compatible with the range of variation of
<taxonomicName id="6FA34D14FFDBF35504C291B09771258E" box="[1189,1323,1176,1199]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF35504C291B09771258E" box="[1189,1323,1176,1199]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35504C291B096E1258E" box="[1189,1211,1176,1199]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. orientalis
</emphasis>
</taxonomicName>
. Morover, the
<typeStatus id="77188835FFDBF35500D99190934125EE" box="[190,283,1208,1231]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF3550158919093F225EE" box="[319,424,1208,1231]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="perroti">
<emphasis id="9AD7EA85FFDBF3550158919093F225EE" box="[319,424,1208,1231]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35501589190930F25EE" box="[319,341,1208,1231]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. perroti
</emphasis>
</taxonomicName>
has the same aedeagus as the
<typeStatus id="77188835FFDBF355028D9190911625EE" box="[746,844,1208,1231]" pageId="2" pageNumber="47" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="6FA34D14FFDBF3550317919091AC25EE" box="[880,1014,1208,1231]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="9AD7EA85FFDBF3550317919091AC25EE" box="[880,1014,1208,1231]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF3550317919091DC25EE" box="[880,902,1208,1231]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. orientalis
</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="16C238926974D3E4055E247C3511D2DB" pageId="292" pageNumber="293" type="nomenclature">
<paragraph id="3CF428B381142A3273400BEB8F7855EF" pageId="292" pageNumber="293">
Genus
<taxonomicName id="425C74DE5B2D40EF642BBF48C228D811" ID-CoL="8KV2W" ID-ENA="2219866" LSID="200A9045-7E5F-1B15-95B4-04B3E32424A8" authority="LeConte, 1852" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Akephorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Akephorus" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">Akephorus LeConte, 1852</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="B0A0A5FCDEAE1B02589B3CF162B8DEF8" pageId="292" pageNumber="293" type="reference_group">
<paragraph id="DF03038C8FC9C43971E8D16450E0E222" pageId="292" pageNumber="293">
<taxonomicName id="B3BE70F43594BDD1DF1B222C6A6365A1" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Akephorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">Akephorus</taxonomicName>
LeConte, 1852a: 194. Type species:
<taxonomicName id="A2CD3AA4B75052368FF4DF76EC1C33F8" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Akephorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="marinus">
<emphasis id="E7E9F649E2F846A8C549226DD7CCF4EA" italics="true" pageId="292" pageNumber="293">Akephorus marinus</emphasis>
</taxonomicName>
LeConte, 1852 by monotypy. Etymology. From the Greek
<emphasis id="A56F53F5DB321F76C9B8F4D4616D02D3" italics="true" pageId="292" pageNumber="293">ake</emphasis>
(point) and
<emphasis id="CD91D9A7E47B211B95997304EBEA35FF" italics="true" pageId="292" pageNumber="293">phoro</emphasis>
(to bear, carry), possibly alluding to the acute apical spur of the protibia of the adult (&quot;
<emphasis id="44989A98B84C33B4B289D800A401D029" italics="true" pageId="292" pageNumber="293">tibiae anticae spinis terminalibus longissimis</emphasis>
&quot;) [masculine]. Note.
<taxonomicName id="3582878EE4B74590BA90B82FAEDACE03" authorityName="Leconte" authorityYear="1857" class="Insecta" family="Carabidae" genus="Acephorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="0B051E2C580E5794D2FBB40242E9366B" italics="true" pageId="292" pageNumber="293">Acephorus</emphasis>
</taxonomicName>
is an incorrect subsequent spelling for
<taxonomicName id="F4D8D765EFE76A78AB009A0AAEC8DF9B" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Akephorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="1CA3895F89CB3EFD04283811DEB3CF56" italics="true" pageId="292" pageNumber="293">Akephorus</emphasis>
</taxonomicName>
, first used by LeConte (1853c: 396), not in prevailing usage.
</paragraph>
</subSubSection>
<subSubSection id="C03CEAFF4CD3E85F719BCE30DB212F87" pageId="292" pageNumber="293" type="diversity">
<paragraph id="503FFE6E25568B0D0987A62FF57372C9" pageId="292" pageNumber="293">Diversity.</paragraph>
<paragraph id="1947DA305B5FC8326F65D427D777014D" pageId="292" pageNumber="293">Two species restricted to the seashore of the Pacific in North America.</paragraph>
</subSubSection>
<subSubSection id="EED56F0A313009EE46AA69AEC496D87D" pageId="292" pageNumber="293" type="identification">
<paragraph id="5AC13E7B809DFF702108EB5927ECFFF3" pageId="292" pageNumber="293">Identification.</paragraph>
<paragraph id="26A642A6839EEE548570707D9E1F0A66" pageId="292" pageNumber="293">
The two species can be differentiated using
<normalizedToken id="05AB8CAD1197C5399E73F9A08E0B23E3" originalValue="Lindroth’s">Lindroth's</normalizedToken>
(1961a) and
<normalizedToken id="7FBAC427DB5833E59ECCF2000547EF4D" originalValue="Bousquet’s">Bousquet's</normalizedToken>
(1988a) keys to
<taxonomicName id="68335A6763BC2F84D5439ED81A334A94" authorityName="Bonelli" authorityYear="1810" class="Hexapoda" family="Carabidae" genus="Dyschirius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="DA855E0531AC4EFA3B22D211C4A008BE" italics="true" pageId="292" pageNumber="293">Dyschirius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F73125FCA6C6F253BBD5664116F0B489" pageId="292" pageNumber="293" type="taxonomic note">
<paragraph id="C8ED4C92E32DFE11F724EEA184104A41" pageId="292" pageNumber="293">Taxonomic Note.</paragraph>
<paragraph id="798BA22FC580DEB1D2F816B7062DC6EE" pageId="292" pageNumber="293">
This taxon has been considered a subgenus of
<taxonomicName id="5A68D8E66D40235E730547C0E4BB0008" authorityName="Bonelli" authorityYear="1810" class="Hexapoda" family="Carabidae" genus="Dyschirius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="75C2B3A5235527F7071875A5BEDDC4A5" italics="true" pageId="292" pageNumber="293">Dyschirius</emphasis>
</taxonomicName>
by Lindroth (1961a) and a distinct genus by Fedorenko (1996). The phylogenetic analysis of Fedorenko (1996) did not yield clear evidence as to the position of the group. Furthermore there are little structural characteristics for members of
<taxonomicName id="161E7FE8F6C5686E43D770481E921DC5" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Akephorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="16A9B4209D36F486A2DCD9E63357BAE0" italics="true" pageId="292" pageNumber="293">Akephorus</emphasis>
</taxonomicName>
. The presence of conspicuous microsculpture on the body is also found in a few
<taxonomicName id="F6723FB0F7DEC8CBF01CF468CA2319D2" authorityName="Bonelli" authorityYear="1810" class="Hexapoda" family="Carabidae" genus="Dyschirius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="6C2ECE085CF7152666DF98CAC766BDF4" italics="true" pageId="292" pageNumber="293">Dyschirius</emphasis>
</taxonomicName>
groups, such as the
<emphasis id="B0084F1A620287E9DA7DC6482092F3DF" italics="true" pageId="292" pageNumber="293">exochus</emphasis>
group, phenetically similar to the remaining
<taxonomicName id="81B1FAB494D688F424812904B25CC4B8" authorityName="Bonelli" authorityYear="1810" class="Hexapoda" family="Carabidae" genus="Dyschirius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="FC0F79E49C428496069A02687C9EF10F" italics="true" pageId="292" pageNumber="293">Dyschirius</emphasis>
</taxonomicName>
. The body shape, however, is distinctive. With the current phylogenetic knowledge of the tribe, I see no harm in retaining this taxon as a distinct genus.
</paragraph>
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